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<mods:title id="0E3C15D59628E1A23E8DADD05E56C4E5">A revision of Chinese endemic Neptis hesione Leech, 1890 with a new species described from Hainan Island (Lepidoptera, Nymphalidae)</mods:title>
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(
<figureCitation id="97DC2A72413EFF949FAF6847FF7EE262" box="[159,237,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1597,1621]" captionTargetBox="[155,1432,187,1569]" captionTargetId="figure-22@2.[151,1435,181,1574]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of all members of Neptis hesione-group." pageId="3" pageNumber="434">Figs. 1</figureCitation>
<figureCitation id="97DC2A72413EFF949FAF6847FF7EE262" box="[159,237,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1597,1621]" captionTargetBox="[155,1432,187,1569]" captionTargetId="figure-22@2.[151,1435,181,1574]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of all members of Neptis hesione-group." figureDoi="http://doi.org/10.5281/zenodo.13235012" httpUri="https://zenodo.org/record/13235012/files/figure.png" pageId="3" pageNumber="434">Figs. 1</figureCitation>
for adults, 3 for genitalia, 4 for diagnosis, 5 for distribution, 6 for phylogeny)
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@ -131,11 +134,11 @@ leg. (NK1; will be deposited at
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<emphasis id="3D93EAE5413EFF949FF768F6FE36E292" bold="true" box="[199,421,1194,1221]" pageId="3" pageNumber="434">Description: Male.</emphasis>
(
<figureCitation id="97DC2A72413EFF949E8468F7FE66E292" box="[436,501,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1597,1621]" captionTargetBox="[155,1432,187,1569]" captionTargetId="figure-22@2.[151,1435,181,1574]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of all members of Neptis hesione-group." pageId="3" pageNumber="434">Fig. 1</figureCitation>
<figureCitation id="97DC2A72413EFF949E8468F7FE66E292" box="[436,501,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1597,1621]" captionTargetBox="[155,1432,187,1569]" captionTargetId="figure-22@2.[151,1435,181,1574]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of all members of Neptis hesione-group." figureDoi="http://doi.org/10.5281/zenodo.13235012" httpUri="https://zenodo.org/record/13235012/files/figure.png" pageId="3" pageNumber="434">Fig. 1</figureCitation>
) Ground color black on upperside and brown on underside. Wing markings yellow, tinged with orange in upperside and lighter in underside. Forewing upperside: postdiscal spots composed of three bands, lower clavate band from basal part of spaces 1a to 1b, middle round band from spaces 1b to 3, upper ellipse band from spaces 4 to 6; lower and midle postdiscal bands almost contact with each other; subcostal small spots present in spaces 8 and 9, beyond upper postdiscal brand; cell streak with basal part laterally dusted and with distal part pointed. Hindwing upperside: discal band wide, basally whitish and turned into gray when passing through speculum; postdiscal band narrow, interrupted by black veins. Forewing underside: wing patterns similar to upperside but yellow markings broader; a submarginal yellow fascia running from spaces 8 to 3, ending at middle postdiscal brand; another submarginal fascia, outside preceding one, passing from costa to dorsum, tinged with lilac. Hindwing underside: subbase decorated with a yellow streak of an irregular shape; discal band wider, with a black line embedded in its lower edge; postdiscal band wider accompanied by yellow submarginal fascia, with their distal portions overlaid with lilac; ground color between discal and postdiscal bands dark-brown. Speculum on hindwing upperside and corresponding polished area on forewing underside dark. Fringes chequered black and white.
<emphasis id="3D93EAE5413EFF949FD46A23FE0DE0CF" bold="true" box="[228,414,1662,1689]" pageId="3" pageNumber="434">Male Genitalia.</emphasis>
(
<figureCitation id="97DC2A72413EFF949E9E6A23FE62E0CE" box="[430,497,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,970,994]" captionTargetBox="[155,1428,661,940]" captionTargetId="figure-35@4.[151,1436,660,947]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Neptis kejiaae sp. nov. le = lateral left view, ds = dorsal view, vs = ventral view." pageId="3" pageNumber="434">Fig. 3</figureCitation>
<figureCitation id="97DC2A72413EFF949E9E6A23FE62E0CE" box="[430,497,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,970,994]" captionTargetBox="[155,1428,661,940]" captionTargetId="figure-35@4.[151,1436,660,947]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of Neptis kejiaae sp. nov. le = lateral left view, ds = dorsal view, vs = ventral view." figureDoi="http://doi.org/10.5281/zenodo.13235016" httpUri="https://zenodo.org/record/13235016/files/figure.png" pageId="3" pageNumber="434">Fig. 3</figureCitation>
) Vinculum: slender, triple as long as saccus, ventrally broad. Saccus: from the part connected to vinculum, continuously narrowed forward, ending with a sharp point. Tegumen: dorsally elliptical, cephalic part protruding. Gnathos: vertically elongated, ventral portion integrated and protruding caudally. Uncus: straight, elongated into a pointed hook, slightly down-curved. Valva: more or less tongue-shaped with ampulla and harpe terminally differentiated; harpe wide, blade-like, gradually curved upward into a blunt end; ampulla fused with costa, a hump present in middle rear, caudal end of ampulla narrower and somewhat shorter than harpe with inconspicuous upper teeth. Juxta: bilateral parts expansive posteriorly, elliptical. Phallus: almost straight; basally expansive and terminal end forming a sharp upward hook; cornutus with tiny spins in expanded caudal part.
</paragraph>
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@ -147,10 +150,10 @@ The new species can be distinguished from other members in the
<emphasis id="3D93EAE5413EFF949B2E6BC3FB07E1EE" box="[1054,1172,1951,1977]" italics="true" pageId="3" pageNumber="434">N. hesione</emphasis>
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-group by the following characters (
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<figureCitation id="97DC2A72413EFF949E286B9FFECEE18A" box="[280,349,1987,2013]" captionStart="FIGURE 4" captionStartId="4.[151,250,1446,1470]" captionTargetBox="[159,1428,1019,1420]" captionTargetId="figure-60@4.[151,1436,1019,1423]" captionTargetPageId="4" captionText="FIGURE 4. Diagnosis of Neptis kejiaae sp. nov. and other species in N. hesione-group." figureDoi="http://doi.org/10.5281/zenodo.13235018" httpUri="https://zenodo.org/record/13235018/files/figure.png" pageId="3" pageNumber="434">Fig. 4</figureCitation>
):
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<emphasis id="3D93EAE54139FF939FA76E3FFE80E42C" bold="true" box="[151,275,611,635]" pageId="4" pageNumber="435">FIGURE 2.</emphasis>
Holotype of
@ -160,7 +163,7 @@ Holotype of
and the related literatures.
</paragraph>
</caption>
<caption id="5B98667F4139FF939FA76F96FB6AE5B4" box="[151,1273,970,995]" pageId="4" pageNumber="435" startId="4.[151,250,970,994]" targetBox="[155,1428,661,940]" targetPageId="4" targetType="figure">
<caption id="5B98667F4139FF939FA76F96FB6AE5B4" ID-DOI="http://doi.org/10.5281/zenodo.13235016" ID-Zenodo-Dep="13235016" box="[151,1273,970,995]" httpUri="https://zenodo.org/record/13235016/files/figure.png" pageId="4" pageNumber="435" startId="4.[151,250,970,994]" targetBox="[155,1428,661,940]" targetPageId="4" targetType="figure">
<paragraph id="0F5836F74139FF939FA76F96FB6AE5B4" blockId="4.[151,1273,970,995]" box="[151,1273,970,995]" pageId="4" pageNumber="435">
<emphasis id="3D93EAE54139FF939FA76F96FE80E5B5" bold="true" box="[151,275,970,994]" pageId="4" pageNumber="435">FIGURE 3.</emphasis>
Male genitalia of
@ -173,7 +176,7 @@ Male genitalia of
le = lateral left view, ds = dorsal view, vs = ventral view.
</paragraph>
</caption>
<caption id="5B98667F4139FF939FA769FAFB80E3E8" box="[151,1043,1446,1471]" pageId="4" pageNumber="435" startId="4.[151,250,1446,1470]" targetBox="[159,1428,1019,1420]" targetPageId="4" targetType="figure">
<caption id="5B98667F4139FF939FA769FAFB80E3E8" ID-DOI="http://doi.org/10.5281/zenodo.13235018" ID-Zenodo-Dep="13235018" box="[151,1043,1446,1471]" httpUri="https://zenodo.org/record/13235018/files/figure.png" pageId="4" pageNumber="435" startId="4.[151,250,1446,1470]" targetBox="[159,1428,1019,1420]" targetPageId="4" targetType="figure">
<paragraph id="0F5836F74139FF939FA769FAFB80E3E8" blockId="4.[151,1043,1446,1471]" box="[151,1043,1446,1471]" pageId="4" pageNumber="435">
<emphasis id="3D93EAE54139FF939FA769FAFE80E3E8" bold="true" box="[151,275,1446,1471]" pageId="4" pageNumber="435">FIGURE 4.</emphasis>
Diagnosis of
@ -258,7 +261,7 @@ are sympatric with
).
</paragraph>
</subSubSection>
<caption id="5B98667F4138FF929FA76E86FD97E541" pageId="5" pageNumber="436" startId="5.[151,250,730,754]" targetBox="[151,1435,181,705]" targetPageId="5" targetType="figure">
<caption id="5B98667F4138FF929FA76E86FD97E541" ID-DOI="http://doi.org/10.5281/zenodo.13235020" ID-Zenodo-Dep="13235020" httpUri="https://zenodo.org/record/13235020/files/figure.png" pageId="5" pageNumber="436" startId="5.[151,250,730,754]" targetBox="[151,1435,181,705]" targetPageId="5" targetType="figure">
<paragraph id="0F5836F74138FF929FA76E86FD97E541" blockId="5.[151,1436,730,790]" pageId="5" pageNumber="436">
<emphasis id="3D93EAE54138FF929FA76E86FE84E4A5" bold="true" box="[151,279,730,754]" pageId="5" pageNumber="436">FIGURE 5.</emphasis>
Distributional map of
@ -268,7 +271,7 @@ Distributional map of
-group. The type localities are represented by the letter T. Source from references in Material and methods.
</paragraph>
</caption>
<caption id="5B98667F4138FF929FA769E7FB26E3A0" pageId="5" pageNumber="436" startId="5.[151,250,1467,1491]" targetBox="[151,1436,815,1443]" targetPageId="5" targetType="figure">
<caption id="5B98667F4138FF929FA769E7FB26E3A0" ID-DOI="http://doi.org/10.5281/zenodo.13235022" ID-Zenodo-Dep="13235022" httpUri="https://zenodo.org/record/13235022/files/figure.png" pageId="5" pageNumber="436" startId="5.[151,250,1467,1491]" targetBox="[151,1436,815,1443]" targetPageId="5" targetType="figure">
<paragraph id="0F5836F74138FF929FA769E7FB26E3A0" blockId="5.[151,1436,1467,1527]" pageId="5" pageNumber="436">
<emphasis id="3D93EAE54138FF929FA769E7FE87E384" bold="true" box="[151,276,1467,1491]" pageId="5" pageNumber="436">FIGURE 6.</emphasis>
Phylogenetic tree of