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<document id="048CC5C437B74876D7692067EDBAF980" ID-DOI="10.5281/zenodo.203285" ID-GBIF-Dataset="f31dbef8-f48b-4a18-815d-1442ccaf5d15" ID-ISSN="1175-5326" ID-Zenodo-Dep="203285" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460340654504" checkinUser="plazi" docAuthor="Starý, Jaroslav" docDate="2011" docId="537D87AFFF8A696A2AABFF27FBACFEF8" docLanguage="en" docName="zt02999p062.pdf" docOrigin="Zootaxa 2999" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Molophilus (Molophilus) tjederi Stary 1968" docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="AF44FFD7FF9A697A2A3CFFB0FFA6FFC6" masterDocTitle="Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)" masterLastPageNumber="62" masterPageNumber="45" pageNumber="61" updateTime="1698260995331" updateUser="plazi">
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<mods:title id="925EC6A7B92DFF43D78DC219CDEED285">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<mods:namePart id="21DD55E5F3545338B127F1079183BDD3">Starý, Jaroslav</mods:namePart>
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<mods:date id="5503E6DE88097318AD7566DF814AF8D6">2011</mods:date>
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<taxonomicName id="1CD44D3AFF8A696A2AABFF27FD08FF77" ID-CoL="43XQW" authority="Stary, 1968" authorityName="Stary" authorityYear="1968" box="[151,686,151,177]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="61" phylum="Arthropoda" rank="species" species="tjederi" subGenus="Molophilus">
<emphasis id="E9A0EAABFF8A696A2AABFF27FD08FF77" bold="true" box="[151,686,151,177]" pageId="16" pageNumber="61">
<emphasis id="E9A0EAABFF8A696A2AABFF27FDBCFF77" bold="true" box="[151,538,151,177]" italics="true" pageId="16" pageNumber="61">Molophilus (Molophilus) tjederi</emphasis>
<bibRefCitation id="BF454B48FF8A696A281EFF27FD08FF77" author="Stary" box="[546,686,151,177]" pageId="16" pageNumber="61" refString="Stary, J. (1968) Eine neue Molophilus - Art aus der Slowakei. Acta entomologica bohemoslovaca, 65, 239 - 240." type="journal article" year="1968">Starý, 1968</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<emphasis id="E9A0EAABFF8A696A2AABFF6EFE69FF31" bold="true" box="[151,463,222,247]" pageId="16" pageNumber="61">
Material examined.
<collectingCountry id="A3C37629FF8A696A2BB2FF6FFE6CFF31" box="[398,458,223,247]" name="Italy" pageId="16" pageNumber="61">Italy</collectingCountry>
:
</emphasis>
Calabria: Marcellina (CS) env., Abatemarco shores,
<date id="AF6A1079FF8A696A2E07FF6EFB01FF30" box="[1083,1191,222,246]" pageId="16" pageNumber="61" value="1996-06-03">3.vi.1996</date>
, 5 3, 1 Ƥ (J. Martinovský leg.) (
<collectionCode id="BDC5AE7CFF8A696A2B0DFEB2FEC3FEDC" box="[305,357,258,282]" pageId="16" pageNumber="61">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF8A696A2AFBFE96FBACFEF8" box="[199,1034,294,319]" pageId="16" pageNumber="61" type="distribution">
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<emphasis id="E9A0EAABFF8A696A2AFBFE96FEC6FEF9" bold="true" box="[199,352,294,319]" pageId="16" pageNumber="61">Distribution.</emphasis>
<collectingCountry id="A3C37629FF8A696A2B5BFE96FE6CFEF8" box="[359,458,294,318]" name="Bulgaria" pageId="16" pageNumber="61">Bulgaria</collectingCountry>
,
<collectingCountry id="A3C37629FF8A696A2BE9FE96FD9BFEF8" box="[469,573,294,318]" name="Romania" pageId="16" pageNumber="61">Romania</collectingCountry>
,
<collectingCountry id="A3C37629FF8A696A2874FE96FD0BFEF8" box="[584,685,294,318]" name="Slovakia" pageId="16" pageNumber="61">Slovakia</collectingCountry>
,
<collectingCountry id="A3C37629FF8A696A2885FE96FCB1FEF8" box="[697,791,294,318]" name="Ukraine" pageId="16" pageNumber="61">Ukraine</collectingCountry>
. First record for
<collectingCountry id="A3C37629FF8A696A29EDFE96FBA0FEF8" box="[977,1030,294,318]" name="Italy" pageId="16" pageNumber="61">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="694E1FCD586F5BF2AF9CFF77D9BB5E3E">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<heading id="802381D5FF9069702AABFA8CFCA8FA90" bold="true" box="[151,782,1340,1366]" fontSize="11" level="1" pageId="10" pageNumber="55" reason="1">
<emphasis id="E9A0EAABFF9069702AABFA8CFCA8FA90" bold="true" box="[151,782,1340,1366]" pageId="10" pageNumber="55">
<taxonomicName id="1CD44D3AFF9069702AABFA8CFD9EFA90" ID-CoL="43XKB" box="[151,568,1340,1366]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9069702AABFA8CFD9EFA90" bold="true" box="[151,568,1340,1366]" italics="true" pageId="10" pageNumber="55">Molophilus (Molophilus) pleuralis</emphasis>
</taxonomicName>
de
<bibRefCitation id="BF454B48FF9069702858FA8CFCA8FA90" author="Meijere" box="[612,782,1340,1366]" pageId="10" pageNumber="55" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere, 1920</bibRefCitation>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9069702AABFAEFFE9EFAB1" blockId="10.[151,782,1340,1399]" box="[151,312,1375,1399]" pageId="10" pageNumber="55">
<figureCitation id="43EF2A3CFF9069702AABFAEFFE9EFAB1" box="[151,312,1375,1399]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="10" pageNumber="55">Figures 24, 25</figureCitation>
</paragraph>
<paragraph id="DB6B36B9FF9069702AABFA17FB99FA7B" blockId="10.[151,1087,1447,1469]" box="[151,1087,1447,1469]" pageId="10" pageNumber="55">
<treatmentCitationGroup id="FBC41197FF9069702AABFA17FB99FA7B" box="[151,1087,1447,1469]" pageId="10" pageNumber="55">
<taxonomicName id="1CD44D3AFF9069702AABFA17FECCFA7B" ID-CoL="43XKB" box="[151,362,1447,1469]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="E9A0EAABFF9069702AABFA17FECCFA7B" box="[151,362,1447,1469]" italics="true" pageId="10" pageNumber="55">Molophilus pleuralis</emphasis>
</taxonomicName>
de
<treatmentCitation id="5A7510A8FF9069702BB3FA17FDE5FA7B" author="Meijere" box="[399,579,1447,1469]" page="60" pageId="10" pageNumber="55" year="1920">
<bibRefCitation id="BF454B48FF9069702BB3FA17FD87FA7A" author="Meijere" box="[399,545,1447,1469]" pageId="10" pageNumber="55" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere, 1920</bibRefCitation>
: 60
</treatmentCitation>
(description), Pl. 4, Figs 53a–b (male terminalia).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="DB6B36B9FF9069712AABFA5AFC9CFF12" blockId="10.[151,1437,1513,2006]" lastBlockId="11.[151,1436,151,391]" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55">
<emphasis id="E9A0EAABFF9069702AABFA5AFE1BF9C4" bold="true" box="[151,445,1513,1538]" pageId="10" pageNumber="55">
<typeStatus id="046F881BFF9069702AABFA5AFF74F9C4" box="[151,210,1514,1538]" pageId="10" pageNumber="55">Type</typeStatus>
material examined.
</emphasis>
The species was described from an unspecified number of specimens from “Amsterdam, Diemen, Haarlem, Abcoude, Hilversum, Uitgeest, Leimuiden, Zwammerdam” (de
<bibRefCitation id="BF454B48FF9069702E7BF9BDFB45F9E3" author="Meijere" box="[1095,1251,1549,1573]" pageId="10" pageNumber="55" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere 1920</bibRefCitation>
: 60). Altogether 32
<typeStatus id="046F881BFF9069702A80F982FEB8F98C" box="[188,286,1586,1610]" pageId="10" pageNumber="55" type="syntype">syntypes</typeStatus>
(25 3, 5 Ƥ,
<specimenCount id="CDD2FD30FF9069702B84F981FDE3F98F" box="[440,581,1585,1609]" pageId="10" pageNumber="55" type="generic">2 specimens</specimenCount>
without abdomen) are now deposited in
<collectionCode id="BDC5AE7CFF9069702E23F982FBD2F98C" LSID="urn:lsid:biocol.org:col:34975" box="[1055,1140,1586,1610]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="55">ZMAN</collectionCode>
, and one male
<typeStatus id="046F881BFF9069702F1AF982FADBF98C" box="[1318,1405,1586,1610]" pageId="10" pageNumber="55" type="syntype">syntype</typeStatus>
in
<collectionCode id="BDC5AE7CFF9069702AABF9E6FF4EF9AB" box="[151,232,1622,1645]" pageId="10" pageNumber="55">ZMHB</collectionCode>
(de
<bibRefCitation id="BF454B48FF9069702B25F9E6FE37F9AB" author="Jong" box="[281,401,1621,1646]" pageId="10" pageNumber="55" refString="Jong, H. de (2000) The types of Diptera described by J. C. H. de Meijere. Backhuys Publishers, Leiden, viii + 271 pp." type="book" year="2000">Jong 2000</bibRefCitation>
: 170–171). I have examined four male
<typeStatus id="046F881BFF9069702974F9E6FC0CF9A8" box="[840,938,1622,1646]" pageId="10" pageNumber="55" type="syntype">syntypes</typeStatus>
from
<collectionCode id="BDC5AE7CFF90697029D2F9E6FBE2F9A8" LSID="urn:lsid:biocol.org:col:34975" box="[1006,1092,1622,1646]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="55">ZMAN</collectionCode>
, all belonging to
<taxonomicName id="1CD44D3AFF9069702F39F9E7FF6CF954" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9069702F39F9E7FF6CF954" italics="true" pageId="10" pageNumber="55">M. (M.) pleuralis</emphasis>
</taxonomicName>
, including one labelled (but not published) as
<typeStatus id="046F881BFF90697028ECF9C9FC91F957" box="[720,823,1657,1681]" pageId="10" pageNumber="55" type="lectotype">lectotype</typeStatus>
by Br. Theowald. The latter is here designated accordingly.
<emphasis id="E9A0EAABFF9069702AD9F92EFEFDF970" bold="true" box="[229,347,1694,1718]" pageId="10" pageNumber="55">
<typeStatus id="046F881BFF9069702AD9F92EFEFDF970" box="[229,347,1694,1718]" pageId="10" pageNumber="55" type="lectotype">Lectotype</typeStatus>
</emphasis>
3 (by present designation):
<collectingCountry id="A3C37629FF90697028FAF92DFCF6F973" box="[710,848,1693,1717]" name="Netherlands" pageId="10" pageNumber="55">Netherlands</collectingCountry>
, Amsterdam,
<date id="AF6A1079FF90697029C1F92DFBCCF973" box="[1021,1130,1693,1717]" pageId="10" pageNumber="55" value="1919-05-11">11.v.1919</date>
(J.C.H. de Meijere leg.) (
<collectionCode id="BDC5AE7CFF9069702AA3F971FF51F91F" LSID="urn:lsid:biocol.org:col:34975" box="[159,247,1729,1753]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="55">ZMAN</collectionCode>
), labelled “Amsterdam /
<date id="AF6A1079FF9069702833F971FDC3F91F" box="[527,613,1729,1753]" pageId="10" pageNumber="55" value="1919-05-11">11.V.19</date>
/ de Meijere” (partly printed, date handwritten), a paper square with “C”, “
<typeStatus id="046F881BFF9069702A9DF955FEB5F93B" box="[161,275,1765,1789]" pageId="10" pageNumber="55" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="1CD44D3AFF9069702B11F956FE10F93B" box="[301,438,1765,1789]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis">M. pleuralis</taxonomicName>
/ n. sp. / des. Theow.” (handwritten, red), “
<taxonomicName id="1CD44D3AFF9069702993F955FB95F93B" box="[943,1075,1765,1789]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="genus">Molophilus</taxonomicName>
/
<taxonomicName id="1CD44D3AFF9069702E71F955FB08F93B" box="[1101,1198,1765,1789]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis">pleuralis</taxonomicName>
/ de
<bibRefCitation id="BF454B48FF9069702ED0F955FA2DF93B" author="Meijere" box="[1260,1419,1765,1789]" pageId="10" pageNumber="55" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere, 1920</bibRefCitation>
/
<collectionCode id="BDC5AE7CFF9069702AABF8BAFF4FF8E4" LSID="urn:lsid:biocol.org:col:34975" box="[151,233,1802,1826]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="55">ZMAN</collectionCode>
<typeStatus id="046F881BFF9069702ACDF8BAFE87F8E4" box="[241,289,1802,1826]" pageId="10" pageNumber="55">type</typeStatus>
<collectionCode id="BDC5AE7CFF9069702B14F8B9FECEF8E7" box="[296,360,1801,1825]" pageId="10" pageNumber="55">DIPT</collectionCode>
.0671.22” (printed, red). Labelled as
<typeStatus id="046F881BFF9069702938F8B9FCCDF8E7" box="[772,875,1801,1825]" pageId="10" pageNumber="55" type="lectotype">lectotype</typeStatus>
by the present author (“
<typeStatus id="046F881BFF9069702E44F8B9FABAF8E7" box="[1144,1308,1801,1825]" pageId="10" pageNumber="55" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="1CD44D3AFF9069702F0EF8B9FF1EF883" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="genus">Molophilus</taxonomicName>
/
<taxonomicName id="1CD44D3AFF9069702AF3F89DFE89F883" box="[207,303,1837,1861]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="pleuralis">pleuralis</taxonomicName>
de Meijere 3 / J. Starý 2010”; printed red label). The specimen is micro-pinned on a piece of foam, with only left fore and mid legs attached; apex of abdomen cut off. Terminalia mounted by de Meijere in
<collectingCountry id="A3C37629FF9069702F74F8E1FA3AF8AF" box="[1352,1436,1873,1897]" name="Canada" pageId="10" pageNumber="55">Canada</collectingCountry>
balsam on a celluloid slide, pinned with the specimen.
<emphasis id="E9A0EAABFF90697028CAF8C6FC09F848" bold="true" box="[758,943,1910,1934]" pageId="10" pageNumber="55">
<typeStatus id="046F881BFF90697028CAF8C6FC0CF848" box="[758,938,1910,1934]" pageId="10" pageNumber="55" type="paralectotype">Paralectotypes</typeStatus>
:
</emphasis>
<collectingCountry id="A3C37629FF9069702989F8C5FBE7F84B" box="[949,1089,1909,1933]" name="Netherlands" pageId="10" pageNumber="55">Netherlands</collectingCountry>
: Diemen,
<date id="AF6A1079FF9069702E8CF8C5FA8AF84B" box="[1200,1324,1909,1933]" pageId="10" pageNumber="55" value="1918-08-03">3.viii.1918</date>
, 1 3; Uitgeest,
<date id="AF6A1079FF9069702ADCF829FEFAF877" box="[224,348,1945,1969]" pageId="10" pageNumber="55" value="1918-08-01">1.viii.1918</date>
, 1 3; Haarlem, ±
<date id="AF6A1079FF906970281FF829FD0BF877" box="[547,685,1945,1969]" pageId="10" pageNumber="55" value="1919-08-18">18.viii.1919</date>
, 1 3 (all J.C.H. de Meijere leg.) (all
<collectionCode id="BDC5AE7CFF9069702E6FF829FB0CF877" LSID="urn:lsid:biocol.org:col:34975" box="[1107,1194,1945,1969]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="55">ZMAN</collectionCode>
). All specimens have red
<typeStatus id="046F881BFF9069702AF9F80EFE83F810" box="[197,293,1982,2006]" pageId="10" pageNumber="55" type="paratype">paratype</typeStatus>
labels and labels with numbers “0671.17”, “0671.12”, and “0671.25”, respectively. All specimens subsequently labelled as
<typeStatus id="046F881BFF9169712B92FF27FDF6FF69" box="[430,592,151,175]" pageId="11" pageNumber="56" type="paralectotype">paralectotypes</typeStatus>
by the present author. The
<typeStatus id="046F881BFF91697129BEFF27FC4FFF69" box="[898,1001,151,175]" pageId="11" pageNumber="56" type="lectotype">lectotype</typeStatus>
is designated here to stabilize the concept of the name with respect to
<taxonomicName id="1CD44D3AFF9169712838FF0DFD52FF12" box="[516,756,188,212]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9169712838FF0DFD52FF12" box="[516,756,188,212]" italics="true" pageId="11" pageNumber="56">M. (M.) brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF91697128C7FF0CFC9CFF12" bold="true" box="[763,826,188,212]" pageId="11" pageNumber="56">
<taxonomicNameLabel id="F29357D0FF91697128C7FF0CFC9CFF12" box="[763,826,188,212]" pageId="11" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="DB6B36B9FF9169712AFBFF6FFB25FEF9" blockId="11.[151,1436,151,391]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBFF6FFE55FF3E" bold="true" box="[199,499,223,248]" pageId="11" pageNumber="56">Other material examined</emphasis>
(9 3, 1 Ƥ):
<emphasis id="E9A0EAABFF9169712840FF6FFD7FFF3E" bold="true" box="[636,729,223,248]" pageId="11" pageNumber="56">
<collectingCountry id="A3C37629FF9169712840FF6FFD72FF3E" box="[636,724,223,248]" name="Greece" pageId="11" pageNumber="56">Greece</collectingCountry>
:
</emphasis>
Pieria Region: Eginio env., Aliakmonas delta, cut-off meander,
<date id="AF6A1079FF9169712AABFEB4FEA5FEDA" box="[151,259,260,284]" pageId="11" pageNumber="56" value="2007-06-04">4.vi.2007</date>
, 1 3 (J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9169712BD2FEB4FD84FEDA" box="[494,546,260,284]" pageId="11" pageNumber="56">JSO</collectionCode>
). Crete: Khania Region: Kavros,
<date id="AF6A1079FF91697129A2FEB4FBA9FEDA" box="[926,1039,260,284]" pageId="11" pageNumber="56" value="2004-05-19">19.v.2004</date>
, 1 3 (J. Roháček &amp; M. Vála leg.); Kavros, marsh,
<date id="AF6A1079FF9169712B74FE97FE1EFEF9" box="[328,440,295,319]" pageId="11" pageNumber="56" value="2004-05-20">20.v.2004</date>
, 6 3, 1 Ƥ; Kavros,
<date id="AF6A1079FF91697128A8FE97FCA2FEF9" box="[660,772,295,319]" pageId="11" pageNumber="56" value="2004-05-24">24.v.2004</date>
, 1 3 (both J. Starý leg.) (all
<collectionCode id="BDC5AE7CFF9169712E7FFE97FBD0FEF9" box="[1091,1142,295,319]" pageId="11" pageNumber="56">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBFEFBFB8DFE41" pageId="11" pageNumber="56" type="distribution">
<paragraph id="DB6B36B9FF9169712AFBFEFBFB8DFE41" blockId="11.[151,1436,151,391]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBFEFBFEC6FEA2" bold="true" box="[199,352,331,356]" pageId="11" pageNumber="56">Distribution.</emphasis>
Widely distributed in the Palaearctic: Europe, including
<collectingCountry id="A3C37629FF91697129D6FEFCFB98FEA2" box="[1002,1086,332,356]" name="Cyprus" pageId="11" pageNumber="56">Cyprus</collectingCountry>
;
<collectingCountry id="A3C37629FF9169712E70FEFCFB3BFEA2" box="[1100,1181,332,356]" name="Turkey" pageId="11" pageNumber="56">Turkey</collectingCountry>
, Transcaucasia,
<collectingCountry id="A3C37629FF9169712F6BFEFCFA3EFEA2" box="[1367,1432,332,356]" name="Israel" pageId="11" pageNumber="56">Israel</collectingCountry>
,
<collectingCountry id="A3C37629FF9169712AABFEC0FF61FE41" box="[151,199,368,391]" name="Iran" pageId="11" pageNumber="56">Iran</collectingCountry>
, Middle Asia,
<collectingCountry id="A3C37629FF9169712B57FEDFFE10FE41" box="[363,438,367,391]" name="Russia" pageId="11" pageNumber="56">Russia</collectingCountry>
as far as Far East. First records for
<collectingCountry id="A3C37629FF9169712979FEDFFC32FE41" box="[837,916,367,391]" name="Greece" pageId="11" pageNumber="56">Greece</collectingCountry>
(incl. Crete).
</paragraph>
</subSubSection>
</treatment>
</document>

138
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<mods:title id="FF3208EEA820FA2A72333E52C34754F4">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<subSubSection id="93CE6532FF9169712AABFE6CFE91FDD0" pageId="11" pageNumber="56" type="nomenclature">
<paragraph id="DB6B36B9FF9169712AABFE6CFD33FE33" blockId="11.[151,661,475,534]" box="[151,661,475,502]" pageId="11" pageNumber="56">
<heading id="802381D5FF9169712AABFE6CFD33FE33" bold="true" box="[151,661,475,502]" fontSize="11" level="1" pageId="11" pageNumber="56" reason="1">
<emphasis id="E9A0EAABFF9169712AABFE6CFD33FE33" bold="true" box="[151,661,475,502]" pageId="11" pageNumber="56">
<taxonomicName id="1CD44D3AFF9169712AABFE6CFDEDFE30" ID-CoL="43X35" box="[151,587,476,502]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="calabricus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9169712AABFE6CFDEDFE30" bold="true" box="[151,587,476,502]" italics="true" pageId="11" pageNumber="56">Molophilus (Molophilus) calabricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF916971286FFE6BFD33FE33" box="[595,661,475,501]" pageId="11" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9169712AABFE4EFE91FDD0" blockId="11.[151,661,475,534]" box="[151,311,510,534]" pageId="11" pageNumber="56">
<figureCitation id="43EF2A3CFF9169712AABFE4EFE91FDD0" box="[151,311,510,534]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="11" pageNumber="56">Figures 26–29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AABFDF6FB91FD0C" pageId="11" pageNumber="56" type="diagnosis">
<paragraph id="DB6B36B9FF9169712AABFDF6FB91FD0C" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AABFDF6FEB7FD99" bold="true" box="[151,273,582,607]" pageId="11" pageNumber="56">Diagnosis.</emphasis>
Small species within
<taxonomicName id="1CD44D3AFF916971283BFDF6FD20FD98" box="[519,646,582,606]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF916971283BFDF6FD20FD98" box="[519,646,582,606]" italics="true" pageId="11" pageNumber="56">Molophilus</emphasis>
</taxonomicName>
. Body brown, patterned with pale yellow and dark brown, locally suffused with sparse greyish pruinosity. Gonocoxite of male terminalia with thumb-like dorsal lobe and triangular lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, with terminal hook curved laterally, pointed at tip. Inner gonostylus undulate, obtuse at apex. Wing length 3.6–3.8 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBFD66FE33FA78" pageId="11" pageNumber="56" type="description">
<paragraph id="DB6B36B9FF9169712AFBFD66FD3BFCF1" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBFD66FE0CFD29" bold="true" box="[199,426,726,751]" pageId="11" pageNumber="56">Description. Male.</emphasis>
Head. Antenna short, hardly extending to wing base. Scape brown, pedicel yellow. Flagellomeres (except for yellowed flagellomere 1) infuscated, brown, ovoid to short-ovoid, with longest verticils exceeding length of their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9169712AFBFCF2FAD0FB90" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">Thorax brown, patterned with pale yellow and dark brown, locally suffused with sparse greyish pruinosity. Prescutum and scutum, greyish brown, bordered with pale yellow on paratergite and around prescutal pit; scutal lobe yellowed posterolaterally. Scutellum yellowed medially, greyish brown on margins. Postscutellum greyish brown, yellowed anterolaterally. Pleuron locally shiny, with distinct dark brown longitudinal stripe extending from pronotum over anepisternum and fading out on anepimeron; pale yellow areas on dorsopleural membrane and basalare and elongate yellow marking, or streak, on katepisternum contrasting with its dark brown lower part. Wing tinged with yellowish brown; venation generally as for genus. Stem of halter yellow, knob brown. Legs with coxae, trochanters and bases of femora obscure yellow (hind femur almost entirely so), rest of legs dark brown.</paragraph>
<paragraph id="DB6B36B9FF9169712AFBFBD2FA30FA5C" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
Abdomen darker than thorax, dark brown. Male terminalia (
<figureCitation id="43EF2A3CFF916971294CFBD2FC54FBBC" box="[880,1010,1122,1146]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="11" pageNumber="56">Figs 26–29</figureCitation>
) pale brown. Dorsal portion of gonocoxite rather short, with moderately large, thumb-like dorsal lobe and long, triangular lateral lobe; tips of both level. Lateral excision slight, very shallow. Ventral lobe of gonocoxite slender and very long, longer than body of gonocoxite proximal to it, narrowing into thin filament, especially when seen ventrally, extending considerably beyond dorsal lobe. Both gonostyli darkly pigmented distally. Outer gonostylus shorter than inner one, variously twisted, terminating in sharp hook curved laterally, reaching to about half length of inner gonostylus. The latter slightly extending beyond tip of ventral lobe, undulate, its obtuse, down-curved apex serrate at dorsal margin. Aedeagus slender, of moderate length, bent ventrally at about one fourth its length, then gradually tapered and very slightly upturned. Aedeagal plate small, generally oval in ventral/ventrocaudal aspect, without any spines or teeth.
</paragraph>
<paragraph id="DB6B36B9FF9169712AFBFA17FE33FA78" blockId="11.[151,1437,582,1759]" box="[199,405,1446,1471]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBFA17FEB8FA79" bold="true" box="[199,286,1447,1471]" pageId="11" pageNumber="56">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBFA7AFC6CF9C0" pageId="11" pageNumber="56" type="materials_examined">
<paragraph id="DB6B36B9FF9169712AFBFA7AFC6CF9C0" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBFA7AFD85FA25" bold="true" box="[199,547,1482,1507]" pageId="11" pageNumber="56">
Material examined.
<typeStatus id="046F881BFF9169712B8AFA7BFD85FA25" box="[438,547,1483,1507]" pageId="11" pageNumber="56" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF9169712872FA7AFD24FA24" box="[590,642,1482,1506]" name="Italy" pageId="11" pageNumber="56">Italy</collectingCountry>
, Calabria, Marcellina (CS) env., Abatemarco shores,
<date id="AF6A1079FF9169712EEFFA7AFA9BFA24" box="[1235,1341,1482,1506]" pageId="11" pageNumber="56" value="1996-06-03">3.vi.1996</date>
(J. Martinovský leg.) (
<collectionCode id="BDC5AE7CFF9169712B7CFA5EFE30F9C0" box="[320,406,1518,1542]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="11" pageNumber="56">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF9169712B94FA5FFDB8F9C1" bold="true" box="[424,542,1519,1543]" pageId="11" pageNumber="56">
<typeStatus id="046F881BFF9169712B94FA5FFDBFF9C1" box="[424,537,1519,1543]" pageId="11" pageNumber="56" type="paratype">Paratype</typeStatus>
:
</emphasis>
1 3, same data as for
<typeStatus id="046F881BFF9169712925FA5EFCDDF9C0" box="[793,891,1518,1542]" pageId="11" pageNumber="56" type="holotype">holotype</typeStatus>
(
<collectionCode id="BDC5AE7CFF91697129B6FA5EFC18F9C0" box="[906,958,1518,1542]" pageId="11" pageNumber="56">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBF9A3FB61F988" pageId="11" pageNumber="56" type="etymology">
<paragraph id="DB6B36B9FF9169712AFBF9A3FB61F988" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBF9A3FEE8F9ED" bold="true" box="[199,334,1555,1579]" pageId="11" pageNumber="56">Etymology.</emphasis>
The name of this new species,
<taxonomicName id="1CD44D3AFF9169712893F9A3FC83F9ED" box="[687,805,1555,1579]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="calabricus">
<emphasis id="E9A0EAABFF9169712893F9A3FC83F9ED" box="[687,805,1555,1579]" italics="true" pageId="11" pageNumber="56">calabricus</emphasis>
</taxonomicName>
, is derived from the Italian province Calabria where the
<typeStatus id="046F881BFF9169712AABF987FF61F989" box="[151,199,1591,1615]" pageId="11" pageNumber="56">type</typeStatus>
specimens come from. It is to be deemed to be a latinized adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBF9EAFC7FF97C" pageId="11" pageNumber="56" type="discussion">
<paragraph id="DB6B36B9FF9169712AFBF9EAFC7FF97C" blockId="11.[151,1437,582,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBF9EAFEEAF9B5" bold="true" box="[199,332,1626,1651]" pageId="11" pageNumber="56">Discussion.</emphasis>
This new species is distantly related to
<taxonomicName id="1CD44D3AFF9169712931F9ECFB30F9B4" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" box="[781,1174,1626,1651]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="ochraceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9169712931F9ECFC78F9B5" box="[781,990,1627,1651]" italics="true" pageId="11" pageNumber="56">M. (M.) ochraceus</emphasis>
(
<bibRefCitation id="BF454B48FF91697129D1F9EAFB28F9B4" author="Meigen" box="[1005,1166,1626,1650]" pageId="11" pageNumber="56" refString="Meigen, J. W. (1818) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Aachen, 1, i - xxxvi + 1 - 333." type="journal article" year="1818">Meigen, 1818</bibRefCitation>
)
</taxonomicName>
, a common and widely distributed West Palaearctic species, from which it differs by nearly every detail of the male terminalia. The Transcaucasian
<taxonomicName id="1CD44D3AFF9169712B29F914FD72F97C" authority="Savchenko, 1976" authorityName="Savchenko" authorityYear="1976" box="[277,724,1698,1723]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="subochraceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9169712B29F914FDABF97D" box="[277,525,1699,1723]" italics="true" pageId="11" pageNumber="56">M. (M.) subochraceus</emphasis>
Savchenko, 1976
</taxonomicName>
stands still more apart.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9169712AFBF976FE06F918" box="[199,416,1734,1759]" pageId="11" pageNumber="56" type="distribution">
<paragraph id="DB6B36B9FF9169712AFBF976FE06F918" blockId="11.[151,1437,582,1759]" box="[199,416,1734,1759]" pageId="11" pageNumber="56">
<emphasis id="E9A0EAABFF9169712AFBF976FEC6F919" bold="true" box="[199,352,1734,1759]" pageId="11" pageNumber="56">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9169712B5BF976FE3AF918" box="[359,412,1734,1758]" name="Italy" pageId="11" pageNumber="56">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

221
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<mods:title id="3AF3735C19A363D3FFACB15D61C7A603">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<treatment id="537D87AFFF9369732AABFF27FD37F89E" ID-DOI="http://doi.org/10.5281/zenodo.6193286" ID-GBIF-Taxon="119406423" ID-Zenodo-Dep="6193286" LSID="urn:lsid:plazi:treatment:537D87AFFF9369732AABFF27FD37F89E" httpUri="http://treatment.plazi.org/id/537D87AFFF9369732AABFF27FD37F89E" lastPageNumber="54" pageId="9" pageNumber="54">
<subSubSection id="93CE6532FF9369732AABFF27FE9EFF14" pageId="9" pageNumber="54" type="nomenclature">
<paragraph id="DB6B36B9FF9369732AABFF27FD1EFF77" blockId="9.[151,696,151,210]" box="[151,696,151,177]" pageId="9" pageNumber="54">
<heading id="802381D5FF9369732AABFF27FD1EFF77" bold="true" box="[151,696,151,177]" fontSize="11" level="1" pageId="9" pageNumber="54" reason="1">
<emphasis id="E9A0EAABFF9369732AABFF27FD1EFF77" bold="true" box="[151,696,151,177]" pageId="9" pageNumber="54">
<taxonomicName id="1CD44D3AFF9369732AABFF27FDC9FF77" ID-CoL="43X2N" box="[151,623,151,177]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732AABFF27FDC9FF77" bold="true" box="[151,623,151,177]" italics="true" pageId="9" pageNumber="54">Molophilus (Molophilus) brevifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF936973284AFF27FD1EFF77" box="[630,696,151,177]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9369732AABFF0AFE9EFF14" blockId="9.[151,696,151,210]" box="[151,312,186,210]" pageId="9" pageNumber="54">
<figureCitation id="43EF2A3CFF9369732AABFF0AFE9EFF14" box="[151,312,186,210]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="9" pageNumber="54">Figures 21–23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AABFEB2FF6CFE6C" pageId="9" pageNumber="54" type="diagnosis">
<paragraph id="DB6B36B9FF9369732AABFEB2FF6CFE6C" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AABFEB2FEB7FEDD" bold="true" box="[151,273,258,283]" pageId="9" pageNumber="54">Diagnosis.</emphasis>
Medium-sized species within
<taxonomicName id="1CD44D3AFF936973284EFEB3FD57FEDD" box="[626,753,259,283]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF936973284EFEB3FD57FEDD" box="[626,753,259,283]" italics="true" pageId="9" pageNumber="54">Molophilus</emphasis>
</taxonomicName>
. Body yellowish brown, patterned with yellow and brown, locally suffused with sparse, pale greyish pruinosity. Gonocoxite of male terminalia with thumb-like dorsal lobe and triangular lateral lobe. Both gonostyli very short, outer (dorsal) gonostylus longer than inner (ventral) gonostylus, generally hook-shaped, curved mesally, pointed at tip. Inner gonostylus conical, pale. Wing length 4.0–4.8 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AFBFE06FE33FB46" pageId="9" pageNumber="54" type="description">
<paragraph id="DB6B36B9FF9369732AFBFE06FB54FE35" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBFE06FE02FE09" bold="true" box="[199,420,438,463]" pageId="9" pageNumber="54">Description. Male.</emphasis>
Head. Antenna short, not reaching wing base. Basal segments brown. Flagellomeres dark brown, ovoid to short-ovoid, with longest verticils about twice as long as their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9369732AFBFE4EFBAEFCDE" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">Thorax generally yellowish brown, patterned with yellow and brown. Prescutum and scutum pale brown, locally suffused with sparse pale greyish pruinosity, yellowed laterally; scutal lobe yellowed medially and posterolaterally. Scutellum mostly yellow, darker laterally. Postscutellum pale brown, yellowed laterally. Pleuron obscure yellow, with more or less distinct, sometimes diffuse, brown longitudinal stripe extending from pronotum over anepisternum sometimes as far as base of halter; lower part of katepisternum and meron also darkened. Wing tinged with yellowish brown; compared to other species fork M3-M4 very short, mostly much shorter than its petiole (M3+4). Halter yellow, slightly darker on proximal part of knob. Legs with coxae and trochanters obscure yellow to yellowish brown, bases of femora yellowish brown, rest of legs dark brown.</paragraph>
<paragraph id="DB6B36B9FF9369732AFBFC94FD29FB9A" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
Abdomen darker than thorax, yellowish brown to brown. Male terminalia (
<figureCitation id="43EF2A3CFF9369732E30FC94FB2DFCFA" box="[1036,1163,804,828]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="9" pageNumber="54">Figs 21–23</figureCitation>
) yellow. Dorsal portion of gonocoxite moderate in length, with conspicuous, thumb-like dorsal lobe and lateral lobe roughly triangular. Lateral excision wide and shallow. Ventral lobe of gonocoxite short, subacute at tip, extending beyond apex of dorsal lobe. Both gonostyli very short compared to other species. Outer gonostylus hook-shaped, darkly pigmented distally, curved mesally and pointed at tip. Both gonostyli extending same distance, but outer gonostylus longer due to its curvature. Inner gonostylus conical, pale throughout, its subacute apex approximately level with other appendages. Aedeagus long and very slender, bisinuous in lateral view, with tip down-curved, slightly expanded before apex, therefore lanceolate in dorsal/ventral aspect. Aedeagal plate small, generally short-oval in ventral/ventrocaudal aspect, without any spines or teeth.
</paragraph>
<paragraph id="DB6B36B9FF9369732AFBFBD8FE33FB46" blockId="9.[151,1437,258,1881]" box="[199,405,1128,1152]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBFBD8FEB8FB46" bold="true" box="[199,286,1128,1152]" pageId="9" pageNumber="54">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AFBFB3CFD88FAF2" pageId="9" pageNumber="54" type="materials_examined">
<paragraph id="DB6B36B9FF9369732AFBFB3CFD88FAF2" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBFB3CFD8BFB63" bold="true" box="[199,557,1164,1189]" pageId="9" pageNumber="54">
Material examined.
<typeStatus id="046F881BFF9369732BFCFB3DFD8BFB63" box="[448,557,1165,1189]" pageId="9" pageNumber="54" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF936973285EFB3CFD61FB62" box="[610,711,1164,1188]" name="Slovakia" pageId="9" pageNumber="54">Slovakia</collectingCountry>
, Novosad (distr. Trebišov),
<date id="AF6A1079FF9369732E33FB3CFBD8FB62" box="[1039,1150,1164,1188]" pageId="9" pageNumber="54" value="1980-05-19">19.v.1980</date>
(J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9369732F07FB3CFA37FB62" box="[1339,1425,1164,1188]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="9" pageNumber="54">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF9369732AABFB00FEA9FB0E" bold="true" box="[151,271,1200,1224]" pageId="9" pageNumber="54">
<typeStatus id="046F881BFF9369732AABFB00FEA9FB0E" box="[151,271,1200,1224]" pageId="9" pageNumber="54" type="paratype">Paratypes</typeStatus>
</emphasis>
(11 3):
<emphasis id="E9A0EAABFF9369732B48FB1FFD9CFB0E" bold="true" box="[372,570,1199,1224]" pageId="9" pageNumber="54">
<collectingCountry id="A3C37629FF9369732B48FB1FFD93FB0E" box="[372,565,1199,1224]" name="Czech Republic" pageId="9" pageNumber="54">Czech Republic</collectingCountry>
:
</emphasis>
Moravia: Nové Mlýny (distr. Břeclav).
<date id="AF6A1079FF9369732E3EFB00FBC3FB0E" box="[1026,1125,1200,1224]" pageId="9" pageNumber="54" value="1978-05-08">8.v.1978</date>
, 6 3 [listed as
<taxonomicName id="1CD44D3AFF9369732F28FB00FAD1FB0E" box="[1300,1399,1200,1224]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="E9A0EAABFF9369732F28FB00FAD1FB0E" box="[1300,1399,1200,1224]" italics="true" pageId="9" pageNumber="54">pleuralis</emphasis>
</taxonomicName>
by
<bibRefCitation id="BF454B48FF9369732AABFB64FEB3FB2A" author="Stary" box="[151,277,1236,1260]" pageId="9" pageNumber="54" refString="Stary, J. (1998) Trichoceridae, Limoniidae, Pediciidae, Cylindrotomidae. In: Rozkosny, R. &amp; Vanhara, J. (Eds), Diptera of the Palava Biosphere Reserve of UNESCO, I. Folia Facultatis Scientiarium Naturalium Universitatis Masarykianae Brunensis, Biologia 99, pp. 21 - 29." type="journal article" year="1998">Starý 1998</bibRefCitation>
]; Brod nad Dyjí (distr. Břeclav),
<date id="AF6A1079FF93697328BEFB64FD43FB2A" box="[642,741,1236,1260]" pageId="9" pageNumber="54" value="1976-05-09">9.v.1976</date>
, 2 3 (all J. Starý leg.) (
<collectionCode id="BDC5AE7CFF93697329DBFB64FBBFFB2A" box="[999,1049,1236,1260]" pageId="9" pageNumber="54">JSO</collectionCode>
,
<collectionCode id="BDC5AE7CFF9369732E18FB64FBDEFB2A" box="[1060,1144,1236,1260]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="9" pageNumber="54">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF9369732EB6FB64FB5CFB2B" bold="true" box="[1162,1274,1236,1261]" pageId="9" pageNumber="54">
<collectingCountry id="A3C37629FF9369732EB6FB64FB53FB2B" box="[1162,1269,1236,1261]" name="Slovakia" pageId="9" pageNumber="54">Slovakia</collectingCountry>
:
</emphasis>
Lendak (distr. Poprad), Belá valley,
<date id="AF6A1079FF9369732BB4FB48FE5FFAD6" box="[392,505,1272,1296]" pageId="9" pageNumber="54" value="1976-05-29">29.v.1976</date>
, 1 3; Slovenský kras, Silická planina,
<date id="AF6A1079FF9369732996FB48FB8AFAD6" box="[938,1068,1272,1296]" pageId="9" pageNumber="54" value="1970-07-10">10.vii.1970</date>
, 2 3 [listed as
<taxonomicName id="1CD44D3AFF9369732EE8FB48FA9EFAD6" box="[1236,1336,1272,1296]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="E9A0EAABFF9369732EE8FB48FA9EFAD6" box="[1236,1336,1272,1296]" italics="true" pageId="9" pageNumber="54">pleuralis</emphasis>
</taxonomicName>
by
<bibRefCitation id="BF454B48FF9369732F5DFB48FF74FAF2" author="Stary" pageId="9" pageNumber="54" refString="Stary, J. (1974) Nachtrage und Berichtigungen zur Limoniinen-Fauna der Tschechoslowakei (Tipulidae, Diptera). C asopis Slezskeho muzea A, 23, 123 - 143." type="journal article" year="1974">Starý 1974</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9369732AE1FAACFEBEFAF2" author="Stary" box="[221,280,1308,1332]" pageId="9" pageNumber="54" refString="Stary, J. (1984) Celad: Limoniidae. In: Cepelak, J. (Ed.), Diptera of Slovakia. Veda-vydavate lstvo Slovenskej akademie vied, pp. 25 - 47. (in Slovak, Russian and English summaries)." type="book chapter" year="1984">1984</bibRefCitation>
] (all J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9369732BD2FAACFD84FAF2" box="[494,546,1308,1332]" pageId="9" pageNumber="54">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AFBFAF0FF51FA65" pageId="9" pageNumber="54" type="etymology">
<paragraph id="DB6B36B9FF9369732AFBFAF0FF51FA65" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBFAF0FEE8FA9E" bold="true" box="[199,334,1344,1368]" pageId="9" pageNumber="54">Etymology.</emphasis>
The compound name of this new species,
<taxonomicName id="1CD44D3AFF9369732901FAF0FC76FA9E" box="[829,976,1344,1368]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus">
<emphasis id="E9A0EAABFF9369732901FAF0FC76FA9E" box="[829,976,1344,1368]" italics="true" pageId="9" pageNumber="54">brevifurcatus</emphasis>
</taxonomicName>
, derives from the Latin brevis (= short) and furca (= the fork), referring to its unusually short fork M3-M
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the wing venation. An adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AFBFA1EFF49F8F2" pageId="9" pageNumber="54" type="discussion">
<paragraph id="DB6B36B9FF9369732AFBFA1EFF49F8F2" blockId="9.[151,1437,258,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBFA1EFEEAFA01" bold="true" box="[199,332,1454,1479]" pageId="9" pageNumber="54">Discussion.</emphasis>
This new species and the following species treated below are distinctive in having a thumb-like dorsal lobe (see e.g.
<figureCitation id="43EF2A3CFF9369732B41FA63FD81FA2D" box="[381,551,1491,1515]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="9" pageNumber="54">Figs 21, 24, 26</figureCitation>
,
<figureCitation id="43EF2A3CFF936973280FFA63FDDBFA2D" box="[563,637,1491,1515]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="9" pageNumber="54">30, 34</figureCitation>
).
<taxonomicName id="1CD44D3AFF93697328ADFA64FC24FA2D" box="[657,898,1491,1515]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF93697328ADFA64FC24FA2D" box="[657,898,1491,1515]" italics="true" pageId="9" pageNumber="54">M. (M.) brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF93697329B5FA63FC6FFA2D" bold="true" box="[905,969,1491,1515]" pageId="9" pageNumber="54">
<taxonomicNameLabel id="F29357D0FF93697329B5FA63FC6FFA2D" box="[905,969,1491,1515]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is closely related to, and much resembles externally
<taxonomicName id="1CD44D3AFF9369732B31FA48FE68F9C9" box="[269,462,1527,1551]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732B31FA48FE68F9C9" box="[269,462,1527,1551]" italics="true" pageId="9" pageNumber="54">M. (M.) pleuralis</emphasis>
</taxonomicName>
de
<bibRefCitation id="BF454B48FF9369732BC9FA47FD33F9C9" author="Meijere" box="[501,661,1527,1551]" pageId="9" pageNumber="54" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere, 1920</bibRefCitation>
, with which it shares extremely short gonostyli and, in fact, the structure of the male terminalia in general. The gonocoxite of
<taxonomicName id="1CD44D3AFF9369732929F9ACFBA2F9F5" box="[789,1028,1563,1587]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732929F9ACFBA2F9F5" box="[789,1028,1563,1587]" italics="true" pageId="9" pageNumber="54">M. (M.) brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9369732E37F9ABFBECF9F5" bold="true" box="[1035,1098,1563,1587]" pageId="9" pageNumber="54">
<taxonomicNameLabel id="F29357D0FF9369732E37F9ABFBECF9F5" box="[1035,1098,1563,1587]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is, however, markedly shorter and broader than that of
<taxonomicName id="1CD44D3AFF9369732B94F9F0FDCEF991" box="[424,616,1599,1623]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732B94F9F0FDCEF991" box="[424,616,1599,1623]" italics="true" pageId="9" pageNumber="54">M. (M.) pleuralis</emphasis>
</taxonomicName>
, with all the appendages better developed. Overall shape of the gonocoxite is definitely a salient character. The lateral expansion of the aedeagus just before its tip does not occur in
<taxonomicName id="1CD44D3AFF9369732F41F9D4FE94F959" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732F41F9D4FE94F959" italics="true" pageId="9" pageNumber="54">M. (M.) pleuralis</emphasis>
</taxonomicName>
, and the aedeagal plate is short-oval in
<taxonomicName id="1CD44D3AFF93697328CBF938FC4EF959" box="[759,1000,1671,1695]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF93697328CBF938FC4EF959" box="[759,1000,1671,1695]" italics="true" pageId="9" pageNumber="54">M. (M.) brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF93697329D3F937FB89F959" bold="true" box="[1007,1071,1671,1695]" pageId="9" pageNumber="54">
<taxonomicNameLabel id="F29357D0FF93697329D3F937FB89F959" box="[1007,1071,1671,1695]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
but circular in
<taxonomicName id="1CD44D3AFF9369732EE7F938FA3AF959" box="[1243,1436,1671,1695]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732EE7F938FA3AF959" box="[1243,1436,1671,1695]" italics="true" pageId="9" pageNumber="54">M. (M.) pleuralis</emphasis>
</taxonomicName>
(cf.
<figureCitation id="43EF2A3CFF9369732AF8F91BFE62F905" box="[196,452,1707,1731]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="9" pageNumber="54">Figs 21–23 and 24, 25</figureCitation>
). Surprisingly, the two species significantly differ from each other externally in that the fork M3-M4 is very short in
<taxonomicName id="1CD44D3AFF9369732BF1F960FD1BF921" box="[461,701,1743,1767]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732BF1F960FD1BF921" box="[461,701,1743,1767]" italics="true" pageId="9" pageNumber="54">M. (M.) brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF93697328F8F97FFCA2F921" bold="true" box="[708,772,1743,1767]" pageId="9" pageNumber="54">
<taxonomicNameLabel id="F29357D0FF93697328F8F97FFCA2F921" box="[708,772,1743,1767]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, mostly much shorter that its petiole (M3+4), whereas, in
<taxonomicName id="1CD44D3AFF9369732F41F960FE95F8C8" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="pleuralis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9369732F41F960FE95F8C8" italics="true" pageId="9" pageNumber="54">M. (M.) pleuralis</emphasis>
</taxonomicName>
, this fork is distinctly longer, with M3 and M4 both longer than M3+4, as is usual for other
<taxonomicName id="1CD44D3AFF9369732F20F946FA3DF8C8" box="[1308,1435,1782,1806]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9369732F20F946FA3DF8C8" box="[1308,1435,1782,1806]" italics="true" pageId="9" pageNumber="54">Molophilus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9369732AFBF8F0FD37F89E" box="[199,657,1856,1881]" pageId="9" pageNumber="54" type="distribution">
<paragraph id="DB6B36B9FF9369732AFBF8F0FD37F89E" blockId="9.[151,1437,258,1881]" box="[199,657,1856,1881]" pageId="9" pageNumber="54">
<emphasis id="E9A0EAABFF9369732AFBF8F0FEC6F89F" bold="true" box="[199,352,1856,1881]" pageId="9" pageNumber="54">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9369732B5BF8F0FDBAF89E" box="[359,540,1856,1880]" name="Czech Republic" pageId="9" pageNumber="54">Czech Republic</collectingCountry>
,
<collectingCountry id="A3C37629FF936973281BF8F0FD2BF89E" box="[551,653,1856,1880]" name="Slovakia" pageId="9" pageNumber="54">Slovakia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

315
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<emphasis id="E9A0EAABFF9469742AABFF27FD01FF77" bold="true" box="[151,679,151,177]" pageId="14" pageNumber="59">
<taxonomicName id="1CD44D3AFF9469742AABFF27FDFBFF77" ID-CoL="43XPG" box="[151,605,151,177]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742AABFF27FDFBFF77" bold="true" box="[151,605,151,177]" italics="true" pageId="14" pageNumber="59">Molophilus (Molophilus) suboccultus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="DB6B36B9FF9469742AABFF0AFE91FF14" blockId="14.[151,1437,151,1949]" box="[151,311,186,210]" pageId="14" pageNumber="59">
<figureCitation id="43EF2A3CFF9469742AABFF0AFE91FF14" box="[151,311,186,210]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="14" pageNumber="59">Figures 34–36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AABFF6EFB10FEA7" pageId="14" pageNumber="59" type="diagnosis">
<paragraph id="DB6B36B9FF9469742AABFF6EFB10FEA7" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AABFF6EFEB7FF31" bold="true" box="[151,273,222,247]" pageId="14" pageNumber="59">Diagnosis.</emphasis>
Small species within
<taxonomicName id="1CD44D3AFF9469742839FF6EFD22FF30" box="[517,644,222,246]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9469742839FF6EFD22FF30" box="[517,644,222,246]" italics="true" pageId="14" pageNumber="59">Molophilus</emphasis>
</taxonomicName>
. Body black, locally suffused with sparse greyish pruinosity, patterned with yellow only lateral to prescutal pit. Gonocoxite of male terminalia with thumb-like dorsal lobe and conical lateral lobe. Outer (dorsal) gonostylus much shorter than inner (ventral) gonostylus, triangular, flattened. Inner gonostylus sickle-shaped, subacute at apex, with serrate concave margin. Wing length 3.3–4.1 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AFBFEDCFD47FBC5" pageId="14" pageNumber="59" type="description">
<paragraph id="DB6B36B9FF9469742AFBFEDCFB25FE6F" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFEDCFE00FE43" bold="true" box="[199,422,364,389]" pageId="14" pageNumber="59">Description. Male.</emphasis>
Head. Antenna short, not reaching wing base, dark brown to almost black throughout. Flagellomeres ovoid, with longest verticils exceeding length of their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9469742AFBFE03FBEEFD9F" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">Thorax black, shiny, locally suffused with sparse greyish pruinosity. Prescutum and scutum almost black, with yellow area lateral to prescutal pit, including adjacent part of paratergite; slightly paler, greyish area between scutal lobes. Scutellum and postscutellum black, the former sometimes slightly paler medially. Pleuron black, slightly paler on dorsopleural membrane. Wing narrow, membrane infuscated; venation generally as for genus. Stem of halter slightly infuscated, knob pale yellow. Legs deep dark brown to black throughout.</paragraph>
<paragraph id="DB6B36B9FF9469742AFBFDD4FCABFC19" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
Abdomen deep dark brown to black, including male terminalia (
<figureCitation id="43EF2A3CFF946974298EFDD4FB9EFDBA" box="[946,1080,612,636]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="14" pageNumber="59">Figs 34–36</figureCitation>
). Dorsal portion of gonocoxite moderate in length. Dorsal lobe comparatively small and narrow, with small tubercle at inner side near base. Lateral lobe conspicuously elongated, conical, narrowly rounded at tip, with row of short setae dorsally. Lateral lobe distinctly exceeding dorsal lobe. Lateral excision moderately deep and wide. Ventral lobe of gonocoxite generally stout and moderately long, its broadly rounded tip slightly extending beyond apices of both dorsal lobe and outer gonostylus. Outer gonostylus much shorter than inner one, generally triangular, flattened and less pigmented, reaching to about half length of inner gonostylus. The latter darkly pigmented, exceeding ventral lobe by about half its length, generally sickle-shaped, straight in proximal half, then curved downwards and serrate at its concave margin, tapered to subacute tip. Aedeagus of moderate length and breadth, slightly bent ventrally, then gradually tapered to very slightly upturned tip. Aedeagal plate of moderate size, generally oval in ventral/ventrocaudal aspect, with extremely slender, blackened median spine.
</paragraph>
<paragraph id="DB6B36B9FF9469742AFBFC5BFD47FBC5" blockId="14.[151,1437,151,1949]" box="[199,737,1003,1027]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFC5BFEB8FBC5" bold="true" box="[199,286,1003,1027]" pageId="14" pageNumber="59">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AFBFBBEFAC1FA4F" pageId="14" pageNumber="59" type="materials_examined">
<paragraph id="DB6B36B9FF9469742AFBFBBEFB49FB56" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFBBEFD81FBE0" bold="true" box="[199,551,1038,1063]" pageId="14" pageNumber="59">
Material examined.
<typeStatus id="046F881BFF9469742B86FBBEFD81FBE0" box="[442,551,1038,1062]" pageId="14" pageNumber="59" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF946974286AFBBEFD1EFBE0" box="[598,696,1038,1062]" name="Portugal" pageId="14" pageNumber="59">Portugal</collectingCountry>
, Algarve, Serra de Monchique Mts, Foia Mt., springs &amp; marshes (
<quantity id="1C2C9B5CFF9469742AA2FB81FEB9FB8F" box="[158,287,1073,1097]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.7" metricValueMin="7.8" pageId="14" pageNumber="59" unit="m" value="825.0" valueMax="870.0" valueMin="780.0">780–870 m</quantity>
) (
<geoCoordinate id="BEE0507EFF9469742B0AFB81FE35FB8F" box="[310,403,1073,1097]" direction="north" orientation="latitude" pageId="14" pageNumber="59" precision="925" value="37.316666">37°19'N</geoCoordinate>
/
<geoCoordinate id="BEE0507EFF9469742BA1FB81FE52FB8F" box="[413,500,1073,1097]" direction="west" orientation="longitude" pageId="14" pageNumber="59" precision="925" value="-8.6">8°36'W</geoCoordinate>
),
<date id="AF6A1079FF946974283BFB81FDC1FB8F" box="[519,615,1073,1097]" pageId="14" pageNumber="59" value="2010-05-02">2.v.2010</date>
(M. Vála leg.) (
<collectionCode id="BDC5AE7CFF9469742925FB81FCC9FB8F" box="[793,879,1073,1097]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="14" pageNumber="59">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF94697429BCFB82FC5EFB8C" bold="true" box="[896,1016,1074,1098]" pageId="14" pageNumber="59">
<typeStatus id="046F881BFF94697429BCFB82FC5EFB8C" box="[896,1016,1074,1098]" pageId="14" pageNumber="59" type="paratype">Paratypes</typeStatus>
</emphasis>
(4 3, 1 Ƥ):
<emphasis id="E9A0EAABFF9469742EBDFB82FB54FB8C" bold="true" box="[1153,1266,1074,1098]" pageId="14" pageNumber="59">
<collectingCountry id="A3C37629FF9469742EBDFB82FB48FB8C" box="[1153,1262,1074,1098]" name="Portugal" pageId="14" pageNumber="59">Portugal</collectingCountry>
:
</emphasis>
Algarve: Serra de Monchique Mts, Foia Mt., springs &amp; marshes (
<quantity id="1C2C9B5CFF94697428F2FBE5FCF7FBAB" box="[718,849,1109,1133]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.7" metricValueMin="7.8" pageId="14" pageNumber="59" unit="m" value="825.0" valueMax="870.0" valueMin="780.0">780–870 m</quantity>
) (
<geoCoordinate id="BEE0507EFF9469742955FBE5FC61FBAB" box="[873,967,1109,1133]" direction="north" orientation="latitude" pageId="14" pageNumber="59" precision="925" value="37.316666">37°19'N</geoCoordinate>
/
<geoCoordinate id="BEE0507EFF94697429ECFBE5FB81FBAB" box="[976,1063,1109,1133]" direction="west" orientation="longitude" pageId="14" pageNumber="59" precision="925" value="-8.6">8°36'W</geoCoordinate>
),
<date id="AF6A1079FF9469742E07FBE5FB38FBAB" box="[1083,1182,1109,1133]" pageId="14" pageNumber="59" value="2010-05-01">1.v.2010</date>
, 1 3,
<date id="AF6A1079FF9469742ED8FBE5FB54FBAB" box="[1252,1266,1109,1133]" pageId="14" pageNumber="59" value="2010-05-02">2</date>
.v.2010, 2 3, 1 Ƥ; Monchique, 1.5 km E, brook (
<quantity id="1C2C9B5CFF9469742830FBC8FDF3FB56" box="[524,597,1144,1168]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="14" pageNumber="59" unit="m" value="430.0">430 m</quantity>
) (
<geoCoordinate id="BEE0507EFF9469742851FBC8FD6CFB56" box="[621,714,1144,1168]" direction="north" orientation="latitude" pageId="14" pageNumber="59" precision="925" value="37.316666">37°19'N</geoCoordinate>
/
<geoCoordinate id="BEE0507EFF94697428EFFBC8FC8CFB56" box="[723,810,1144,1168]" direction="west" orientation="longitude" pageId="14" pageNumber="59" precision="925" value="-8.533334">8°32'W</geoCoordinate>
),
<date id="AF6A1079FF9469742902FBC8FC06FB56" box="[830,928,1144,1168]" pageId="14" pageNumber="59" value="2010-05-01">1.v.2010</date>
, 1 3 (all M. Vála leg.) (
<collectionCode id="BDC5AE7CFF9469742E92FBC8FB44FB56" box="[1198,1250,1144,1168]" pageId="14" pageNumber="59">JSO</collectionCode>
).
</paragraph>
<paragraph id="DB6B36B9FF9469742AFBFB2DFAC1FA4F" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFB2DFDB2FB73" bold="true" box="[199,532,1180,1205]" pageId="14" pageNumber="59">
Excluded from
<typeStatus id="046F881BFF9469742BB4FB2DFE1DFB73" box="[392,443,1181,1205]" pageId="14" pageNumber="59">type</typeStatus>
series:
</emphasis>
<collectingCountry id="A3C37629FF946974281CFB2CFDC4FB72" box="[544,610,1180,1204]" name="Spain" pageId="14" pageNumber="59">Spain</collectingCountry>
, Andalucía, Algeciras, 26.–30.iv.
<date id="AF6A1079FF94697429D8FB2CFBF8FB72" box="[996,1118,1180,1204]" pageId="14" pageNumber="59" value="1925-01-03">1925, 1 3</date>
(H. Zerny leg.) (
<collectionCode id="BDC5AE7CFF9469742F12FB2CFA37FB72" box="[1326,1425,1180,1204]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="59">NHMW</collectionCode>
).
<bibRefCitation id="BF454B48FF9469742AABFB0FFE32FB11" author="Lackschewitz" box="[151,404,1215,1239]" pageId="14" pageNumber="59" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a)</bibRefCitation>
listed the following material under
<taxonomicName id="1CD44D3AFF9469742978FB71FC6AFB1E" box="[836,972,1216,1240]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="E9A0EAABFF9469742978FB71FC6AFB1E" box="[836,972,1216,1240]" italics="true" pageId="14" pageNumber="59">M. occultus</emphasis>
</taxonomicName>
: “
<collectingCountry id="A3C37629FF94697429D6FB0FFBEFFB11" box="[1002,1097,1215,1239]" name="Spain" pageId="14" pageNumber="59">Spanien</collectingCountry>
, Andalusien, Algeciras,
<date id="AF6A1079FF9469742F57FB0FFF7BFB3D" pageId="14" pageNumber="59">26.– 30.IV.</date>
, 12.–20.
<date id="AF6A1079FF9469742B06FB53FE08FB3D" box="[314,430,1251,1275]" pageId="14" pageNumber="59" value="1925-05-03">V.1925, 3</date>
, Ƥ (Zerny)“. I can confirm the male terminalia of the first specimen mentioned above as being identical with
<taxonomicName id="1CD44D3AFF9469742BBEFAB8FDCEFAD8" box="[386,616,1286,1311]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742BBEFAB8FDCEFAD8" box="[386,616,1286,1311]" italics="true" pageId="14" pageNumber="59">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF946974284DFAB7FD14FAD9" bold="true" box="[625,690,1287,1311]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF946974284DFAB7FD14FAD9" box="[625,690,1287,1311]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and the same is probably true for other specimens of Lackschewitz’s series. The body colouration of the specimen, however, is not black but greyish brown, as it is in
<taxonomicName id="1CD44D3AFF9469742F78FA9BFF52FAA0" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742F78FA9BFF52FAA0" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
(even if a certain degree of discolouration is taken into consideration). More material would be needed to solve the problem, so the above-listed specimen is excluded from the
<typeStatus id="046F881BFF946974299FFAC2FC75FA4C" box="[931,979,1394,1418]" pageId="14" pageNumber="59">type</typeStatus>
series of
<taxonomicName id="1CD44D3AFF9469742E7CFAC2FA87FA4F" box="[1088,1313,1393,1417]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742E7CFAC2FA87FA4F" box="[1088,1313,1393,1417]" italics="true" pageId="14" pageNumber="59">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742F14FAC1FAC1FA4F" bold="true" box="[1320,1383,1393,1417]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742F14FAC1FAC1FA4F" box="[1320,1383,1393,1417]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AFBFA25FDA3FA09" pageId="14" pageNumber="59" type="etymology">
<paragraph id="DB6B36B9FF9469742AFBFA25FDA3FA09" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFA25FEE8FA6B" bold="true" box="[199,334,1429,1453]" pageId="14" pageNumber="59">Etymology.</emphasis>
The name of this new species,
<taxonomicName id="1CD44D3AFF9469742890FA25FC96FA6B" box="[684,816,1429,1453]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus">
<emphasis id="E9A0EAABFF9469742890FA25FC96FA6B" box="[684,816,1429,1453]" italics="true" pageId="14" pageNumber="59">suboccultus</emphasis>
</taxonomicName>
, indicates its close relationship to
<taxonomicName id="1CD44D3AFF9469742E8EFA26FACAFA6B" box="[1202,1388,1429,1453]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742E8EFA26FACAFA6B" box="[1202,1388,1429,1453]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
. An adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AFBFA6BFBE7F8BF" pageId="14" pageNumber="59" type="discussion">
<paragraph id="DB6B36B9FF9469742AFBFA6BFBE7F8BF" blockId="14.[151,1437,151,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBFA6BFEEAFA32" bold="true" box="[199,332,1499,1524]" pageId="14" pageNumber="59">Discussion.</emphasis>
The new species is closely related to
<taxonomicName id="1CD44D3AFF94697428C8FA6CFC08FA35" box="[756,942,1499,1523]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF94697428C8FA6CFC08FA35" box="[756,942,1499,1523]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
de
<bibRefCitation id="BF454B48FF94697429EBFA6BFBD1FA35" author="Meijere" box="[983,1143,1499,1523]" pageId="14" pageNumber="59" refString="Meijere, J. C. H. de (1918) Neue hollandische Dipteren. Tijdschrift voor Entomologie, 60, 128 - 141." type="journal article" year="1918">Meijere, 1918</bibRefCitation>
. As with the other species pairs discussed in this paper (
<taxonomicName id="1CD44D3AFF9469742BE3FA4FFDE4F9D1" box="[479,578,1535,1559]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov.">
<emphasis id="E9A0EAABFF9469742BE3FA4FFDE4F9D1" box="[479,578,1535,1559]" italics="true" pageId="14" pageNumber="59">creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742874FA4FFD21F9D1" bold="true" box="[584,647,1535,1559]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742874FA4FFD21F9D1" box="[584,647,1535,1559]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
/
<taxonomicName id="1CD44D3AFF946974289EFA4FFCADF9D1" box="[674,779,1535,1559]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov.">
<emphasis id="E9A0EAABFF946974289EFA4FFCADF9D1" box="[674,779,1535,1559]" italics="true" pageId="14" pageNumber="59">cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF946974292EFA4FFCF7F9D1" bold="true" box="[786,849,1535,1559]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF946974292EFA4FFCF7F9D1" box="[786,849,1535,1559]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="1CD44D3AFF9469742962FA4FFC66F9D1" box="[862,960,1535,1559]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov.">
<emphasis id="E9A0EAABFF9469742962FA4FFC66F9D1" box="[862,960,1535,1559]" italics="true" pageId="14" pageNumber="59">carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF94697429FAFA4FFBA3F9D1" bold="true" box="[966,1029,1535,1559]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF94697429FAFA4FFBA3F9D1" box="[966,1029,1535,1559]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
/
<taxonomicName id="1CD44D3AFF9469742E26F9B0FB37F9D1" box="[1050,1169,1536,1559]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="E9A0EAABFF9469742E26F9B0FB37F9D1" box="[1050,1169,1536,1559]" italics="true" pageId="14" pageNumber="59">nigrescens</emphasis>
</taxonomicName>
,
<taxonomicName id="1CD44D3AFF9469742EA2FA4FFB5EF9D1" box="[1182,1272,1535,1559]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="E9A0EAABFF9469742EA2FA4FFB5EF9D1" box="[1182,1272,1535,1559]" italics="true" pageId="14" pageNumber="59">ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742EC3FA4FFAFEF9D1" bold="true" box="[1279,1368,1535,1559]" pageId="14" pageNumber="59">nom. n.</emphasis>
/
<taxonomicName id="1CD44D3AFF9469742F51F9B0FF56F9FC" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="E9A0EAABFF9469742F51F9B0FF56F9FC" italics="true" pageId="14" pageNumber="59">propinquus</emphasis>
</taxonomicName>
,
<taxonomicName id="1CD44D3AFF9469742AC1F992FE37F9FC" box="[253,401,1570,1594]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov.">
<emphasis id="E9A0EAABFF9469742AC1F992FE37F9FC" box="[253,401,1570,1594]" italics="true" pageId="14" pageNumber="59">brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742BABF993FE70F9FD" bold="true" box="[407,470,1571,1595]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742BABF993FE70F9FD" box="[407,470,1571,1595]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
/
<taxonomicName id="1CD44D3AFF9469742BD6F992FDEBF9FC" box="[490,589,1570,1594]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="E9A0EAABFF9469742BD6F992FDEBF9FC" box="[490,589,1570,1594]" italics="true" pageId="14" pageNumber="59">pleuralis</emphasis>
</taxonomicName>
,
<taxonomicName id="1CD44D3AFF9469742867F993FD64F9FC" box="[603,706,1571,1594]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="E9A0EAABFF9469742867F993FD64F9FC" box="[603,706,1571,1594]" italics="true" pageId="14" pageNumber="59">testaceus</emphasis>
</taxonomicName>
/
<taxonomicName id="1CD44D3AFF94697428EAF993FC81F9FC" box="[726,807,1571,1594]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="E9A0EAABFF94697428EAF993FC81F9FC" box="[726,807,1571,1594]" italics="true" pageId="14" pageNumber="59">griseus</emphasis>
</taxonomicName>
)
<taxonomicName id="1CD44D3AFF946974290BF993FBBEF9FC" box="[823,1048,1570,1594]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF946974290BF993FBBEF9FC" box="[823,1048,1570,1594]" italics="true" pageId="14" pageNumber="59">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742E22F993FBFBF9FD" bold="true" box="[1054,1117,1571,1595]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742E22F993FBFBF9FD" box="[1054,1117,1571,1595]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
much resembles its counterpart in the structure of the male terminalia, being, however, distinguished at first sight by its black body colouration (greyish brown in
<taxonomicName id="1CD44D3AFF9469742B52F9D9FE6DF947" box="[366,459,1641,1665]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="E9A0EAABFF9469742B52F9D9FE6DF947" box="[366,459,1641,1665]" italics="true" pageId="14" pageNumber="59">occultus</emphasis>
</taxonomicName>
). It also differs externally from
<taxonomicName id="1CD44D3AFF9469742978F9DAFBA3F947" box="[836,1029,1641,1665]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742978F9DAFBA3F947" box="[836,1029,1641,1665]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
by having narrower wings. In the structure of the male terminalia, there are numerous distinguishing details between the two. In general, the gonocoxite of
<taxonomicName id="1CD44D3AFF9469742AC3F901FE46F90E" box="[255,480,1712,1736]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742AC3F901FE46F90E" box="[255,480,1712,1736]" italics="true" pageId="14" pageNumber="59">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742BDAF900FD83F90E" bold="true" box="[486,549,1712,1736]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742BDAF900FD83F90E" box="[486,549,1712,1736]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is slightly longer and more slender than that of
<taxonomicName id="1CD44D3AFF9469742E05F901FB54F90E" box="[1081,1266,1712,1736]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742E05F901FB54F90E" box="[1081,1266,1712,1736]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
(the difference is similar to that between
<taxonomicName id="1CD44D3AFF9469742BFAF964FD8FF92A" box="[454,553,1748,1772]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="E9A0EAABFF9469742BFAF964FD8FF92A" box="[454,553,1748,1772]" italics="true" pageId="14" pageNumber="59">pleuralis</emphasis>
</taxonomicName>
and
<taxonomicName id="1CD44D3AFF9469742859F964FD5FF92A" box="[613,761,1748,1772]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="brevifurcatus" status="sp. nov.">
<emphasis id="E9A0EAABFF9469742859F964FD5FF92A" box="[613,761,1748,1772]" italics="true" pageId="14" pageNumber="59">brevifurcatus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF946974293FF964FCE3F92A" bold="true" box="[771,837,1748,1772]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF946974293FF964FCE3F92A" box="[771,837,1748,1772]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, respectively). Furthermore, the new species differs from
<taxonomicName id="1CD44D3AFF9469742AE8F948FE2BF8C9" box="[212,397,1783,1807]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742AE8F948FE2BF8C9" box="[212,397,1783,1807]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
by having the tubercle at the inner side of the dorsal lobe smaller, the lateral lobe shorter, and the ventral lobe only slightly extending beyond the tip of the outer gonostylus (distinctly so in
<taxonomicName id="1CD44D3AFF9469742EC4F8ABFAF3F8F5" box="[1272,1365,1819,1843]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="E9A0EAABFF9469742EC4F8ABFAF3F8F5" box="[1272,1365,1819,1843]" italics="true" pageId="14" pageNumber="59">occultus</emphasis>
</taxonomicName>
). The median spine of the aedeagal plate is shorter and extremely slender in
<taxonomicName id="1CD44D3AFF94697429F4F88FFB09F890" box="[968,1199,1854,1878]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF94697429F4F88FFB09F890" box="[968,1199,1854,1878]" italics="true" pageId="14" pageNumber="59">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9469742E85F88EFB5DF890" bold="true" box="[1209,1275,1854,1878]" pageId="14" pageNumber="59">
<taxonomicNameLabel id="F29357D0FF9469742E85F88EFB5DF890" box="[1209,1275,1854,1878]" pageId="14" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, whereas it is almost twice as long in
<taxonomicName id="1CD44D3AFF9469742B9CF8D2FDFDF8BF" box="[416,603,1889,1913]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9469742B9CF8D2FDFDF8BF" box="[416,603,1889,1913]" italics="true" pageId="14" pageNumber="59">M. (M.) occultus</emphasis>
</taxonomicName>
and not so slender (cf.
<figureCitation id="43EF2A3CFF946974295DF8D1FB92F8BF" box="[865,1076,1889,1913]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="14" pageNumber="59">Figs 34–36 and 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9469742AFBF834FE68F85B" box="[199,462,1924,1949]" pageId="14" pageNumber="59" type="distribution">
<paragraph id="DB6B36B9FF9469742AFBF834FE68F85B" blockId="14.[151,1437,151,1949]" box="[199,462,1924,1949]" pageId="14" pageNumber="59">
<emphasis id="E9A0EAABFF9469742AFBF834FEC6F85B" bold="true" box="[199,352,1924,1949]" pageId="14" pageNumber="59">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9469742B5BF835FE6CF85B" box="[359,458,1925,1949]" name="Portugal" pageId="14" pageNumber="59">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

75
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<mods:title id="2B24C6E139D358FC6D159287C1DAAC36">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<subSubSection id="93CE6532FF9569752AABF952FD5DF93A" box="[151,763,1762,1788]" pageId="15" pageNumber="60" type="nomenclature">
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<heading id="802381D5FF9569752AABF952FD5DF93A" bold="true" box="[151,763,1762,1788]" fontSize="11" level="1" pageId="15" pageNumber="60" reason="1">
<taxonomicName id="1CD44D3AFF9569752AABF952FD5DF93A" ID-CoL="43XQT" authority="Hancock, 2005" authorityName="Hancock" authorityYear="2005" box="[151,763,1762,1788]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="60" phylum="Arthropoda" rank="species" species="tirolensis" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9569752AABF952FD5DF93A" bold="true" box="[151,763,1762,1788]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AABF952FD9BF93A" bold="true" box="[151,573,1762,1788]" italics="true" pageId="15" pageNumber="60">Molophilus (Molophilus) tirolensis</emphasis>
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</emphasis>
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</paragraph>
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<subSubSection id="93CE6532FF9569752AABF899FCB0F84F" pageId="15" pageNumber="60" type="materials_examined">
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<emphasis id="E9A0EAABFF9569752AABF899FE68F884" bold="true" box="[151,462,1833,1858]" pageId="15" pageNumber="60">
Material examined.
<collectingCountry id="A3C37629FF9569752BB0F89AFE6EF884" box="[396,456,1834,1858]" name="Italy" pageId="15" pageNumber="60">Italy</collectingCountry>
:
</emphasis>
Piemonte: Montelupo Albese (CN), Rio Marroni (
<quantity id="1C2C9B5CFF9569752E25F899FBC1F887" box="[1049,1127,1833,1857]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="15" pageNumber="60" unit="m" value="530.0">530 m</quantity>
),
<date id="AF6A1079FF9569752E42F899FB51F887" box="[1150,1271,1833,1857]" pageId="15" pageNumber="60" value="2001-04-24">24.iv.2001</date>
, 5 3; Rodello (CN), Rio S. Rocco (
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),
<date id="AF6A1079FF9569752BDDF8FDFDFCF8A3" box="[481,602,1869,1893]" pageId="15" pageNumber="60" value="2001-04-24">24.iv.2001</date>
, 4 3, 1 Ƥ (both G.B. Delmastro leg.) (all
<collectionCode id="BDC5AE7CFF9569752E16F8FDFBFAF8A3" box="[1066,1116,1869,1893]" pageId="15" pageNumber="60">JSO</collectionCode>
; dried from ethanol).
<emphasis id="E9A0EAABFF9569752AFBF8C1FEC6F84C" bold="true" box="[199,352,1905,1930]" pageId="15" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9569752B5BF8C1FE1AF84F" box="[359,444,1905,1929]" name="Austria" pageId="15" pageNumber="60">Austria</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752BFBF8C2FDB1F84F" box="[455,535,1906,1929]" name="France" pageId="15" pageNumber="60">France</collectingCountry>
. First records for
<collectingCountry id="A3C37629FF95697528E1F8C1FCB4F84F" box="[733,786,1905,1929]" name="Italy" pageId="15" pageNumber="60">Italy</collectingCountry>
.
</paragraph>
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79
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Material examined.
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:
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Steiermark: Admont env., Weng im Gesäuse, Enns River,
<date id="AF6A1079FF9569752EB9F99EFB59F980" box="[1157,1279,1582,1606]" pageId="15" pageNumber="60" value="2006-06-12">12.vi.2006</date>
, 2 3 (J. Starý leg.) (
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).
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,
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,
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,
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,
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. First record for
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.
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Material examined.
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Piemonte: Fabbrica Curone (AL), Caldirola, Rio dell’Olmo (
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, 4 3, 1 Ƥ (G.B. Delmastro leg.) (
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Described from Toscana,
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<taxonomicName id="1CD44D3AFF9569752AABFC7EFC83FC2E" ID-CoL="43XDS" authority="Lackschewitz, 1925" authorityName="Lackschewitz" authorityYear="1925" box="[151,805,974,1000]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="60" phylum="Arthropoda" rank="species" species="maurus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9569752AABFC7EFC83FC2E" bold="true" box="[151,805,974,1000]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AABFC7EFD8DFC2E" bold="true" box="[151,555,974,1000]" italics="true" pageId="15" pageNumber="60">Molophilus (Molophilus) maurus</emphasis>
Lackschewitz, 1925
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="93CE6532FF9569752AABFBA5FEFDFB97" pageId="15" pageNumber="60" type="materials_examined">
<paragraph id="DB6B36B9FF9569752AABFBA5FEFDFB97" blockId="15.[151,1436,1045,1177]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AABFBA5FE55FBE8" bold="true" box="[151,499,1045,1070]" pageId="15" pageNumber="60">
Material examined.
<collectingCountry id="A3C37629FF9569752BACFBA5FE49FBE8" box="[400,495,1045,1070]" name="Austria" pageId="15" pageNumber="60">Austria</collectingCountry>
:
</emphasis>
Steiermark: Admont env., Hall-Grieshof, boggy meadows,
<date id="AF6A1079FF9569752E93FBA5FA8FFBEB" box="[1199,1321,1045,1069]" pageId="15" pageNumber="60" value="2006-06-15">15.vi.2006</date>
, 10 3 (J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9569752B26FB89FEE8FB97" box="[282,334,1081,1105]" pageId="15" pageNumber="60">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9569752AFBFBEDFEC1FB5F" pageId="15" pageNumber="60" type="distribution">
<paragraph id="DB6B36B9FF9569752AFBFBEDFEC1FB5F" blockId="15.[151,1436,1045,1177]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AFBFBEDFEC6FBB0" bold="true" box="[199,352,1117,1142]" pageId="15" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9569752B54FBEDFDB8FBB3" box="[360,542,1117,1141]" name="Czech Republic" pageId="15" pageNumber="60">Czech Republic</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752816FBEDFDD3FBB3" box="[554,629,1117,1141]" name="Latvia" pageId="15" pageNumber="60">Latvia</collectingCountry>
,
<collectingCountry id="A3C37629FF95697528BDFBEDFD49FBB3" box="[641,751,1117,1141]" name="Lithuania" pageId="15" pageNumber="60">Lithuania</collectingCountry>
,
<collectingCountry id="A3C37629FF95697528C7FBEDFCEAFBB3" box="[763,844,1117,1141]" name="Poland" pageId="15" pageNumber="60">Poland</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752964FBEDFC1BFBB3" box="[856,957,1117,1141]" name="Slovakia" pageId="15" pageNumber="60">Slovakia</collectingCountry>
,
<collectingCountry id="A3C37629FF95697529F5FBEDFBF4FBB3" box="[969,1106,1117,1141]" name="Switzerland" pageId="15" pageNumber="60">Switzerland</collectingCountry>
;
<collectingCountry id="A3C37629FF9569752E63FBEDFB08FBB3" box="[1119,1198,1117,1141]" name="Russia" pageId="15" pageNumber="60">Russia</collectingCountry>
: European part. First record for
<collectingCountry id="A3C37629FF9569752B31FB31FEC5FB5F" box="[269,355,1153,1177]" name="Austria" pageId="15" pageNumber="60">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

79
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<taxonomicName id="1CD44D3AFF9569752AABFD64FD7EFD2B" ID-CoL="43XBW" authority="Mendl, 1973" authorityName="Mendl" authorityYear="1973" box="[151,728,723,750]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="60" phylum="Arthropoda" rank="species" species="klementi" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9569752AABFD64FD7EFD2B" bold="true" box="[151,728,723,750]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AABFD64FD90FD28" bold="true" box="[151,566,724,750]" italics="true" pageId="15" pageNumber="60">Molophilus (Molophilus) klementi</emphasis>
<bibRefCitation id="BF454B48FF9569752801FD63FD7EFD2B" author="Mendl" box="[573,728,723,749]" pageId="15" pageNumber="60" refString="Mendl, H. (1973) Drei neue Eriopterini aus dem Allgau (Diptera, Tipulidae). Nachrichtenblatt der Bayerischen Entomologen, 22, 65 - 74." type="journal article" year="1973">Mendl, 1973</bibRefCitation>
</emphasis>
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<emphasis id="E9A0EAABFF9569752AABFCAAFE54FCF5" bold="true" box="[151,498,794,819]" pageId="15" pageNumber="60">
Material examined.
<collectingCountry id="A3C37629FF9569752BA9FCABFE4BFCF5" box="[405,493,795,819]" name="France" pageId="15" pageNumber="60">France</collectingCountry>
:
</emphasis>
Alpes-Maritimes, Le Bar-sur-Loup, 5.5 km NNE (N Saut-du-Loup) (
<quantity id="1C2C9B5CFF9569752F00FCAAFA29FCF4" box="[1340,1423,794,818]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="60" unit="m" value="380.0">380 m</quantity>
),
<date id="AF6A1079FF9569752AABFC8EFEB7FC90" box="[151,273,830,854]" pageId="15" pageNumber="60" value="1994-06-14">14.vi.1994</date>
, 2 3, 1 Ƥ (J.-P. Haenni &amp; C. Dufour leg.) (
<collectionCode id="BDC5AE7CFF95697528C8FC8EFC8EFC90" box="[756,808,830,854]" pageId="15" pageNumber="60">JSO</collectionCode>
).
</paragraph>
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<subSubSection id="93CE6532FF9569752AFBFCD2FBE1FCBC" box="[199,1095,866,891]" pageId="15" pageNumber="60" type="distribution">
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<emphasis id="E9A0EAABFF9569752AFBFCD2FEC6FCBD" bold="true" box="[199,352,866,891]" pageId="15" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9569752B5BFCD2FE1AFCBC" box="[359,444,866,890]" name="Austria" pageId="15" pageNumber="60">Austria</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752BFBFCD2FD96FCBC" box="[455,560,866,890]" name="Germany" pageId="15" pageNumber="60">Germany</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752800FCD2FD02FCBC" box="[572,676,866,890]" name="Romania" pageId="15" pageNumber="60">Romania</collectingCountry>
,
<collectingCountry id="A3C37629FF9569752893FCD2FC9EFCBC" box="[687,824,866,890]" name="Switzerland" pageId="15" pageNumber="60">Switzerland</collectingCountry>
. First record for
<collectingCountry id="A3C37629FF95697529CFFCD3FBE5FCBC" box="[1011,1091,867,890]" name="France" pageId="15" pageNumber="60">France</collectingCountry>
.
</paragraph>
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89
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<taxonomicName id="1CD44D3AFF9569752AABFE69FD6DFE34" ID-CoL="43X26" authority="Goetghebuer" authorityName="Goetghebuer" box="[151,715,472,499]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="60" phylum="Arthropoda" rank="species" species="bifidus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9569752AABFE69FD87FE35" bold="true" box="[151,545,473,499]" italics="true" pageId="15" pageNumber="60">Molophilus (Molophilus) bifidus</emphasis>
Goetghebuer
</taxonomicName>
in
<bibRefCitation id="BF454B48FF95697528CEFE68FBC1FE34" author="Goetghebuer" box="[754,1127,472,498]" pageId="15" pageNumber="60" refString="Goetghebuer, M. &amp; Tonnoir, A. (1920) Catalogue raisonne des Tipulidae de Belgique. Bulletin de la Societe entomologique de Belgique, 2, 104 - 112, 131 - 147." type="journal article" year="1920">Goetghebuer &amp; Tonnoir, 1920</bibRefCitation>
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<emphasis id="E9A0EAABFF9569752AABFDAFFE40FDFE" bold="true" box="[151,486,543,568]" pageId="15" pageNumber="60">
Material examined.
<collectingCountry id="A3C37629FF9569752BB6FDAFFE47FDFE" box="[394,481,543,568]" name="Greece" pageId="15" pageNumber="60">Greece</collectingCountry>
:
</emphasis>
Region Larisa: Karia,
<quantity id="1C2C9B5CFF95697528D7FD90FC83FDF1" box="[747,805,543,567]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="60" unit="km" value="7.0">7 km</quantity>
E (
<quantity id="1C2C9B5CFF9569752972FDAFFC3DFDF1" box="[846,923,543,567]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="15" pageNumber="60" unit="m" value="830.0">830 m</quantity>
),
<date id="AF6A1079FF956975298CFDAFFBBBFDF1" box="[944,1053,543,567]" pageId="15" pageNumber="60" value="2007-06-03">3.vi.2007</date>
, 2 3 (J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9569752F3AFDAFFA9CFDF1" box="[1286,1338,543,567]" pageId="15" pageNumber="60">JSO</collectionCode>
).
<emphasis id="E9A0EAABFF9569752F72FDAFFA3AFDFE" bold="true" box="[1358,1436,543,568]" pageId="15" pageNumber="60">
<collectingCountry id="A3C37629FF9569752F72FDAFFA31FDFE" box="[1358,1431,543,568]" name="Spain" pageId="15" pageNumber="60">Spain</collectingCountry>
:
</emphasis>
Andalucía: Coin, Rio Seco,
<date id="AF6A1079FF9569752BEDFDF4FD9AFD9A" box="[465,572,580,604]" pageId="15" pageNumber="60" value="2005-04-07">7.iv.2005</date>
, 1 3 (J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9569752921FDF4FCF7FD9A" box="[797,849,580,604]" pageId="15" pageNumber="60">JSO</collectionCode>
).
</paragraph>
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<subSubSection id="93CE6532FF9569752AFBFDD7FBD8FDB9" box="[199,1150,615,640]" pageId="15" pageNumber="60" type="distribution">
<paragraph id="DB6B36B9FF9569752AFBFDD7FBD8FDB9" blockId="15.[151,1436,543,640]" box="[199,1150,615,640]" pageId="15" pageNumber="60">
<emphasis id="E9A0EAABFF9569752AFBFDD7FEC6FD46" bold="true" box="[199,352,615,640]" pageId="15" pageNumber="60">Distribution.</emphasis>
Widely distributed in Europe;
<collectingCountry id="A3C37629FF9569752886FDD8FD4DFDB9" box="[698,747,616,639]" name="Iran" pageId="15" pageNumber="60">Iran</collectingCountry>
. First records for
<collectingCountry id="A3C37629FF956975298DFDD7FBA6FDB9" box="[945,1024,615,639]" name="Greece" pageId="15" pageNumber="60">Greece</collectingCountry>
and
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.
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<emphasis id="E9A0EAABFF9569752AABFF6EFE5BFF3E" bold="true" box="[151,509,222,248]" italics="true" pageId="15" pageNumber="60">Molophilus (Molophilus) ater</emphasis>
(
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)
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<emphasis id="E9A0EAABFF9569752AABFE95FDB9FEFB" bold="true" box="[151,543,293,318]" pageId="15" pageNumber="60">
Material examined.
<collectingCountry id="A3C37629FF9569752BBAFE95FDBFFEFB" box="[390,537,293,317]" name="Kazakhstan" pageId="15" pageNumber="60">Kazakhstan</collectingCountry>
:
</emphasis>
Altay Mts, Katon-Karagay N. Park, Bereľ, Yazovka, W shore of Yazovoye lake (
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), 1 3 (Z. Hradílek leg.) (
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<emphasis id="E9A0EAABFF9569752AFBFEDDFEC6FE40" bold="true" box="[199,352,365,390]" pageId="15" pageNumber="60">Distribution.</emphasis>
Widely distributed in Europe;
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: European part, Siberia. First record for
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.
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Lackschewitz, 1940
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stat. n.
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<figureCitation id="43EF2A3CFF9669762AABFF0AFE91FF14" box="[151,311,186,210]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="12" pageNumber="57">Figures 30–32</figureCitation>
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<taxonomicName id="1CD44D3AFF9669762AABFEB2FEC5FEDE" ID-CoL="43X28" box="[151,355,258,280]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="bifilatus">
<emphasis id="E9A0EAABFF9669762AABFEB2FEC5FEDE" box="[151,355,258,280]" italics="true" pageId="12" pageNumber="57">Molophilus bifilatus</emphasis>
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<taxonomicName id="1CD44D3AFF9669762B56FEB3FD4CFEDE" authority="Lackschewitz, 1940a: 22" authorityName="Lackschewitz" authorityPageNumber="22" authorityYear="1940" box="[362,746,258,280]" pageId="12" pageNumber="57" rank="variety" variety="testacea">
var.
<emphasis id="E9A0EAABFF9669762BA9FEB3FE4EFEDE" box="[405,488,259,280]" italics="true" pageId="12" pageNumber="57">testacea</emphasis>
<treatmentCitation id="5A7510A8FF9669762BD2FEB2FD4CFEDE" author="Lackschewitz" box="[494,746,258,280]" page="22" pageId="12" pageNumber="57" year="1940">
<bibRefCitation id="BF454B48FF9669762BD2FEB2FD6EFEDE" author="Lackschewitz" box="[494,712,258,280]" pageId="12" pageNumber="57" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz, 1940a</bibRefCitation>
: 22
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(diagnosis).
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<emphasis id="E9A0EAABFF9669762AABFEF5FEB7FE98" bold="true" box="[151,273,325,350]" pageId="12" pageNumber="57">Diagnosis.</emphasis>
Medium-sized species within
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. Body generally pale brown, patterned with yellow and darker brown, locally suffused with greyish pruinosity. Gonocoxite of male terminalia with thumb-like dorsal lobe and triangular lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, generally hook-shaped, curved mesally, pointed at tip. Inner gonostylus undulate, subacute at apex, with indistinct serrrate flanges. Wing length 5.0–5.4 mm.
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<emphasis id="E9A0EAABFF9669762AFBFE47FE64FDD6" bold="true" box="[199,450,503,528]" pageId="12" pageNumber="57">Redescription. Male.</emphasis>
Head. Antenna of moderate length, slightly extending beyond wing base. Basal segments brown. Flagellomeres dark brown, generally ovoid, with verticils slightly exceeding length of their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9669762AFBFDD3FE25FCB3" blockId="12.[151,1437,325,2029]" pageId="12" pageNumber="57">Thorax generally pale brown, patterned with yellow and darker brown, locally suffused with greyish pruinosity. Prescutum and scutum greyish brown, bordered with yellow laterally on paratergite and around prescutal pit; scutal lobe yellowed anteromedially and posterolaterally. Scutellum mostly yellow, brown laterally. Postscutellum brown, yellowed laterally. Pleuron pale brown, with darker brown patch, or longitudinal stripe, on anepisternum, fading out posteriorly on anepimeron; this stripe surrounded with yellow markings on dorsopleural membrane, basalare and dorsal part of katepisternum. Wing membrane tinged with yellowish brown; venation generally as for genus. Stem of halter dark yellow, knob pale yellow. Legs yellow to yellowish brown, tips of femora and tibiae and entire tarsi darkened.</paragraph>
<paragraph id="DB6B36B9FF9669762AFBFC31FAB3FB71" blockId="12.[151,1437,325,2029]" pageId="12" pageNumber="57">
Abdomen brown. Male terminalia (
<figureCitation id="43EF2A3CFF9669762860FC31FD79FC5F" box="[604,735,897,921]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="12" pageNumber="57">Figs 30–32</figureCitation>
) yellowish brown. Dorsal portion of gonocoxite of moderate length, with conspicuous, long, thumb-like dorsal lobe and triangular lateral lobe; tip of the latter reaching to about half length of the former. Lateral excision wide and shallow. Ventral lobe of gonocoxite slender and long, drawn out into long point, slightly extending beyond apex of dorsal lobe. Both gonostyli darkly pigmented. Outer gonostylus much shorter than inner one, hook-shaped, curved mesally and pointed at tip, reaching just beyond base of inner gonostylus. The latter long, powerful, exceeding tip of ventral lobe by about half of its length, flattened, curved at base, then undulate, with subacute tip slightly upturned and with indistinct, pale, serrate flanges on both sides along mid-length. Aedeagus long and slender, almost straight or slightly undulate in lateral view. Aedeagal plate moderate in size, generally short-oval in ventral/ventrocaudal aspect, without any spines or teeth.
</paragraph>
<paragraph id="DB6B36B9FF9669762AFBFB73FD47FB1C" blockId="12.[151,1437,325,2029]" box="[199,737,1218,1243]" pageId="12" pageNumber="57">
<emphasis id="E9A0EAABFF9669762AFBFB73FEB8FB1D" bold="true" box="[199,286,1219,1243]" pageId="12" pageNumber="57">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9669762AFBFB56FE9AF82B" pageId="12" pageNumber="57" type="materials_examined">
<paragraph id="DB6B36B9FF9669762AFBFB56FECCF9F9" blockId="12.[151,1437,325,2029]" pageId="12" pageNumber="57">
<emphasis id="E9A0EAABFF9669762AFBFB56FE4BFB39" bold="true" box="[199,493,1254,1279]" pageId="12" pageNumber="57">
<typeStatus id="046F881BFF9669762AFBFB56FEA4FB38" box="[199,258,1254,1278]" pageId="12" pageNumber="57">Type</typeStatus>
material examined.
</emphasis>
<bibRefCitation id="BF454B48FF9669762BC8FB56FCB3FB38" author="Lackschewitz" box="[500,789,1254,1278]" pageId="12" pageNumber="57" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a: 23)</bibRefCitation>
based his “
<taxonomicName id="1CD44D3AFF96697629AFFB56FB86FB38" box="[915,1056,1254,1278]" pageId="12" pageNumber="57" rank="variety" variety="testacea">
var.
<emphasis id="E9A0EAABFF96697629F8FB57FB86FB38" box="[964,1056,1255,1278]" italics="true" pageId="12" pageNumber="57">testacea</emphasis>
</taxonomicName>
“ of
<taxonomicName id="1CD44D3AFF9669762E6DFB57FB77FB38" box="[1105,1233,1254,1278]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="bifilatus">
<emphasis id="E9A0EAABFF9669762E6DFB57FB77FB38" box="[1105,1233,1254,1278]" italics="true" pageId="12" pageNumber="57">M. bifilatus</emphasis>
</taxonomicName>
(=
<taxonomicName id="1CD44D3AFF9669762ECBFB57FAEEFB38" box="[1271,1352,1255,1278]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="E9A0EAABFF9669762ECBFB57FAEEFB38" box="[1271,1352,1255,1278]" italics="true" pageId="12" pageNumber="57">griseus</emphasis>
</taxonomicName>
) on the following material: “
<collectingCountry id="A3C37629FF9669762BBCFABAFE79FAE4" box="[384,479,1290,1314]" name="Spain" pageId="12" pageNumber="57">Spanien</collectingCountry>
, Aragonien, Albarracin, 22.–30.
<date id="AF6A1079FF9669762975FABAFC6CFAE4" box="[841,970,1290,1314]" pageId="12" pageNumber="57" value="1924-06-03">VI.1924, 3</date>
(Zerny)”. The single sex symbol without a numeral was used by
<bibRefCitation id="BF454B48FF9669762BA5FA9EFD21FA80" author="Lackschewitz" box="[409,647,1326,1350]" pageId="12" pageNumber="57" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a</bibRefCitation>
&amp;
<bibRefCitation id="BF454B48FF9669762896FA9EFD1FFA80" author="Lackschewitz" box="[682,697,1326,1350]" pageId="12" pageNumber="57" refString="Lackschewitz, P. (1940 b) Die palaarktischen Limnophilinen, Anisomerinen und Pediciinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 68 - 122." type="journal article" year="1940" yearSuffix="b">b</bibRefCitation>
) to indicate a single specimen (see also Starý 2007: 26).
<emphasis id="E9A0EAABFF9669762F13FA9EFA3AFA80" bold="true" box="[1327,1436,1326,1350]" pageId="12" pageNumber="57">
<typeStatus id="046F881BFF9669762F13FA9EFA3AFA80" box="[1327,1436,1326,1350]" pageId="12" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF9669762A81FAE1FF59FAAF" box="[189,255,1361,1385]" name="Spain" pageId="12" pageNumber="57">Spain</collectingCountry>
, Aragón, Albarracin,
<date id="AF6A1079FF9669762BC8FAE1FD3BFAAF" box="[500,669,1361,1385]" pageId="12" pageNumber="57" value="1924-06-22" valueMax="1924-06-30" valueMin="1924-06-22">22.–30.vi.1924</date>
(H. Zerny leg.) (
<collectionCode id="BDC5AE7CFF9669762961FAE1FC19FAAF" box="[861,959,1361,1385]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="57">NHMW</collectionCode>
), labelled “Aragon Albarracin /
<date id="AF6A1079FF9669762F16FAE1FA3AFAAF" box="[1322,1436,1361,1385]" pageId="12" pageNumber="57">22.–30.VI</date>
’24 Zerny” (printed), “
<taxonomicName id="1CD44D3AFF9669762BA4FAC5FD26FA4B" box="[408,640,1397,1421]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="bifilatus">Molophilus bifilatus</taxonomicName>
/ var. Verr. / det. Lacksch.” (printed, white, with “var.” handwritten). The name
<emphasis id="E9A0EAABFF9669762B31FA2AFECFFA77" box="[269,361,1434,1457]" italics="true" pageId="12" pageNumber="57">testacea</emphasis>
was not given on the identification label by Lackschewitz, yet other circumstances clearly indicate that this is the specimen in question. Labelled as
<typeStatus id="046F881BFF96697628CDFA0CFCF5FA12" box="[753,851,1468,1492]" pageId="12" pageNumber="57" type="holotype">holotype</typeStatus>
by the present author (“
<typeStatus id="046F881BFF9669762E62FA0CFB50FA12" box="[1118,1270,1468,1492]" pageId="12" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="1CD44D3AFF9669762F30FA0CFA2BFA12" box="[1292,1421,1468,1492]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="genus">Molophilus</taxonomicName>
/
<taxonomicName id="1CD44D3AFF9669762AABFA51FECFFA3E" authority="Lacksch." authorityName="Lacksch." box="[151,361,1504,1528]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="testaceus">testaceus Lacksch.</taxonomicName>
stat. n. 3 / J. Starý 2010”; printed red label). The specimen is pinned, with only right hind leg and abdomen missing. Terminalia mounted by Lackschewitz in
<collectingCountry id="A3C37629FF966976294BF9B4FC6DF9DA" box="[887,971,1540,1564]" name="Canada" pageId="12" pageNumber="57">Canada</collectingCountry>
balsam between celluloid slides, pinned with the specimen.
</paragraph>
<paragraph id="DB6B36B9FF9669762AFBF9FBFE9AF82B" blockId="12.[151,1437,325,2029]" pageId="12" pageNumber="57">
<emphasis id="E9A0EAABFF9669762AFBF9FBFE51F9A2" bold="true" box="[199,503,1611,1636]" pageId="12" pageNumber="57">Other material examined</emphasis>
(49 3, 8 Ƥ):
<emphasis id="E9A0EAABFF96697628ABF9FCFCAEF9A2" bold="true" box="[663,776,1612,1636]" pageId="12" pageNumber="57">
<collectingCountry id="A3C37629FF96697628ABF9FCFCA5F9A2" box="[663,771,1612,1636]" name="Portugal" pageId="12" pageNumber="57">Portugal</collectingCountry>
:
</emphasis>
Algarve: Vila do Bispo,
<quantity id="1C2C9B5CFF9669762E1BF9FBFBC7F9A5" box="[1063,1121,1611,1635]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="57" unit="km" value="4.0">4 km</quantity>
NE, Lagoa Funda (
<quantity id="1C2C9B5CFF9669762F7EF9FBFA29F9A5" box="[1346,1423,1611,1635]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="12" pageNumber="57" unit="m" value="126.0">126 m</quantity>
), marsh,
<date id="AF6A1079FF9669762AD5F9DFFEC7F941" box="[233,353,1647,1671]" pageId="12" pageNumber="57" value="2010-04-30">30.iv.2010</date>
, 5 3, 4 Ƥ,
<date id="AF6A1079FF9669762BE7F9DFFD98F941" box="[475,574,1647,1671]" pageId="12" pageNumber="57" value="2010-05-03">3.v.2010</date>
, 6 3, 1 Ƥ [1 3+Ƥ in copula]; Monte Ruivo,
<quantity id="1C2C9B5CFF9669762E0CF9DFFBCFF941" box="[1072,1129,1647,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="57" unit="km" value="2.0">2 km</quantity>
E (nr. Poldra Mt.) (
<quantity id="1C2C9B5CFF9669762F78F9DFFA29F941" box="[1348,1423,1647,1671]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="57" unit="m" value="180.0">180 m</quantity>
), marsh,
<date id="AF6A1079FF9669762AD0F923FEE8F96D" box="[236,334,1683,1707]" pageId="12" pageNumber="57" value="2010-05-03">3.v.2010</date>
, 1 3; Serra de Monchique Mts, Foia Mt. (
<quantity id="1C2C9B5CFF9669762905F923FC19F96D" box="[825,959,1683,1707]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.7" metricValueMin="7.8" pageId="12" pageNumber="57" unit="m" value="825.0" valueMax="870.0" valueMin="780.0">780–870 m</quantity>
), springs &amp; marshes,
<date id="AF6A1079FF9669762E87F923FABBF96D" box="[1211,1309,1683,1707]" pageId="12" pageNumber="57" value="2010-05-02">2.v.2010</date>
, 1 3; Rasmalho, 1–2.5 km N, Boina River &amp; tributary (
<quantity id="1C2C9B5CFF96697628A1F907FCA3F909" box="[669,773,1719,1743]" metricMagnitude="1" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="12" pageNumber="57" unit="m" value="62.5" valueMax="75.0" valueMin="50.0">50–75 m</quantity>
),
<date id="AF6A1079FF9669762926F907FC35F909" box="[794,915,1719,1743]" pageId="12" pageNumber="57" value="2010-04-28">28.iv.2010</date>
, 1 3,
<date id="AF6A1079FF96697629EBF907FC52F909" box="[983,1012,1719,1743]" pageId="12" pageNumber="57" value="2010-04-29">29</date>
.iv.2010, 1 3,
<date id="AF6A1079FF9669762EA9F907FB05F909" box="[1173,1187,1719,1743]" pageId="12" pageNumber="57" value="2010-05-02">2</date>
.v.2010, 4 3; Porto de Lagos, 1.5 km NW, Boina River (
<quantity id="1C2C9B5CFF9669762831F96AFDEFF934" box="[525,585,1754,1778]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="57" unit="m" value="25.0">25 m</quantity>
),
<date id="AF6A1079FF9669762861F96AFD73F934" box="[605,725,1754,1778]" pageId="12" pageNumber="57" value="2010-04-29">29.iv.2010</date>
, 1 3 (all J. Starý leg); Ladeira do Vau,
<quantity id="1C2C9B5CFF9669762EB4F96AFB66F934" box="[1160,1216,1754,1778]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="57" unit="km" value="2.0">2 km</quantity>
N, Boina River (
<quantity id="1C2C9B5CFF9669762F43F96AFF16F8D0" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="57" unit="m" value="25.0">25 m</quantity>
), river shores,
<date id="AF6A1079FF9669762B63F94EFE7EF8D0" box="[351,472,1790,1814]" pageId="12" pageNumber="57" value="2010-04-27">27.iv.2010</date>
, 2 3, 1 Ƥ (J. Roháček leg.); Tavira,
<quantity id="1C2C9B5CFF96697629B1F94EFC6FF8D0" box="[909,969,1790,1814]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="57" unit="km" value="3.0">3 km</quantity>
N, Rio Ségua valley (
<quantity id="1C2C9B5CFF9669762EEDF94EFAB4F8D0" box="[1233,1298,1790,1814]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="57" unit="m" value="24.0">24 m</quantity>
), drainages,
<date id="AF6A1079FF9669762AABF892FEA7F8FC" box="[151,257,1826,1850]" pageId="12" pageNumber="57" value="2009-04-01">1.iv.2009</date>
, 1 3, 1 Ƥ; Azinhal env., Barreiros,
<quantity id="1C2C9B5CFF96697628AFF892FD6BF8FC" box="[659,717,1826,1850]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="57" unit="km" value="1.0">1 km</quantity>
S, Beliche River (
<quantity id="1C2C9B5CFF96697629A1F892FC7DF8FC" box="[925,987,1826,1850]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="57" unit="m" value="75.0">75 m</quantity>
),
<date id="AF6A1079FF96697629CCF892FBFCF8FC" box="[1008,1114,1826,1850]" pageId="12" pageNumber="57" value="2009-04-02">2.iv.2009</date>
, 3 3; Monte Gordo,
<quantity id="1C2C9B5CFF9669762F7BF892FA27F8FC" box="[1351,1409,1826,1850]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="57" unit="km" value="1.0">1 km</quantity>
N (
<quantity id="1C2C9B5CFF9669762A9CF8F6FF7AF898" box="[160,220,1862,1886]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="57" unit="m" value="13.0">13 m</quantity>
), saltmarsh,
<date id="AF6A1079FF9669762B54F8F6FE74F898" box="[360,466,1862,1886]" pageId="12" pageNumber="57" value="2009-04-04">4.iv.2009</date>
, 1 3 (all J. Starý leg.) (all
<collectionCode id="BDC5AE7CFF96697628CAF8F6FC8FF898" box="[758,809,1862,1886]" pageId="12" pageNumber="57">JSO</collectionCode>
).
<emphasis id="E9A0EAABFF9669762907F8F5FC2FF898" bold="true" box="[827,905,1861,1886]" pageId="12" pageNumber="57">
<collectingCountry id="A3C37629FF9669762907F8F5FC22F898" box="[827,900,1861,1886]" name="Spain" pageId="12" pageNumber="57">Spain</collectingCountry>
:
</emphasis>
Andalucía: Ronda, Rio Grande,
<date id="AF6A1079FF9669762EC9F8F6FAC6F898" box="[1269,1376,1862,1886]" pageId="12" pageNumber="57" value="2005-04-05">5.iv.2005</date>
, 1 3; Coin, Rio Seco,
<date id="AF6A1079FF9669762B66F8D9FE73F847" box="[346,469,1897,1921]" pageId="12" pageNumber="57" value="2005-03-30">30.iii.2005</date>
, 1 3,
<date id="AF6A1079FF9669762818F8D9FD94F847" box="[548,562,1897,1921]" pageId="12" pageNumber="57" value="2005-04-07">7</date>
.iv.2005, 3 3; Casapalma, confluence of Rio Grande and Rio Guadalhorce,
<date id="AF6A1079FF9669762AABF83DFEA7F863" box="[151,257,1933,1957]" pageId="12" pageNumber="57" value="2005-04-01">1.iv.2005</date>
, 8 3; Cártama, Rio Guadalhorce,
<date id="AF6A1079FF96697628BDF83DFD5AF863" box="[641,764,1933,1957]" pageId="12" pageNumber="57" value="2005-03-31">31.iii.2005</date>
, 1 3, 1 Ƥ,
<date id="AF6A1079FF9669762946F83DFC45F863" box="[890,995,1933,1957]" pageId="12" pageNumber="57" value="2005-04-01">1.iv.2005</date>
, 5 3; El Romeral (nr. Cártama), watercourse,
<date id="AF6A1079FF9669762ACCF801FECAF80F" box="[240,364,1969,1993]" pageId="12" pageNumber="57" value="2005-03-31">31.iii.2005</date>
, 2 3 (all J. Starý leg.); Sierra Nevada Mts, Trevélez (
<quantity id="1C2C9B5CFF96697629E5F801FB9AF80F" box="[985,1084,1969,1993]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="57" unit="m" value="1.5">1,500 m</quantity>
),
<date id="AF6A1079FF9669762E6EF801FB7AF80F" box="[1106,1244,1969,1993]" pageId="12" pageNumber="57" value="2006-08-15">15.viii.2006</date>
, 1 3 (M. Barták leg.) (all
<collectionCode id="BDC5AE7CFF9669762AC0F865FE89F82B" box="[252,303,2005,2029]" pageId="12" pageNumber="57">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9769772AFBFF27FD49F89B" pageId="13" pageNumber="58" type="discussion">
<paragraph id="DB6B36B9FF9769772AFBFF27FE29FEF9" blockId="13.[151,1436,151,319]" pageId="13" pageNumber="58">
<emphasis id="E9A0EAABFF9769772AFBFF27FEEAFF76" bold="true" box="[199,332,151,176]" pageId="13" pageNumber="58">Discussion.</emphasis>
<bibRefCitation id="BF454B48FF9769772B6AFF27FDF4FF69" author="Lackschewitz" box="[342,594,151,175]" pageId="13" pageNumber="58" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a)</bibRefCitation>
established this species as “
<taxonomicName id="1CD44D3AFF97697729A1FF27FB89FF76" box="[925,1071,151,176]" pageId="13" pageNumber="58" rank="variety" status="var. nov." variety="testacea">
var.
<emphasis id="E9A0EAABFF97697729EFFF29FB89FF76" box="[979,1071,153,176]" italics="true" pageId="13" pageNumber="58">testacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9769772E05FF28FB25FF69" box="[1081,1155,151,175]" pageId="13" pageNumber="58" rank="variety">n. var.</taxonomicNameLabel>
” of
<taxonomicName id="1CD44D3AFF9769772E86FF29FF69FF12" authority="Verrall, 1886" authorityName="Verrall" authorityYear="1886" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="bifilatus">
<emphasis id="E9A0EAABFF9769772E86FF29FA98FF76" box="[1210,1342,152,176]" italics="true" pageId="13" pageNumber="58">M. bifilatus</emphasis>
Verrall, 1886
</taxonomicName>
[=
<taxonomicName id="1CD44D3AFF9769772AC0FF0DFD90FF12" authority="Meigen, 1804" authorityName="Meigen" authorityYear="1804" box="[252,566,188,212]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="E9A0EAABFF9769772AC0FF0DFED3FF12" box="[252,373,189,212]" italics="true" pageId="13" pageNumber="58">M. griseus</emphasis>
(
<bibRefCitation id="BF454B48FF9769772BBAFF0CFD8AFF12" author="Meigen" box="[390,556,188,212]" pageId="13" pageNumber="58" refString="Meigen, J. W. (1804) Klassifikazion und Beschreibung der europaischen zweiflugeligen Insekten (Diptera Linn.). Braunschweig, 1, i - xxviii + 1 - 152, 2, i - vi + 153 - 314." type="journal article" year="1804">Meigen, 1804</bibRefCitation>
)
</taxonomicName>
]. He (
<bibRefCitation id="BF454B48FF9769772840FF0CFC33FF12" author="Lackschewitz" box="[636,917,188,212]" pageId="13" pageNumber="58" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz 1940a: 22</bibRefCitation>
) described the difference in the shape of the inner gonostylus between his variety and
<taxonomicName id="1CD44D3AFF9769772858FF51FCB7FF3E" box="[612,785,224,248]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772858FF51FCB7FF3E" box="[612,785,224,248]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
, but apparently did not consider the character important at the species level. The name
<emphasis id="E9A0EAABFF9769772BECFEB5FD8AFEDA" box="[464,556,261,284]" italics="true" pageId="13" pageNumber="58">testacea</emphasis>
is subspecific in terms of Article 45.6.4. of
<bibRefCitation id="BF454B48FF9769772E2AFEB4FB0CFEDA" author="ICZN" box="[1046,1194,260,284]" pageId="13" pageNumber="58" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th edition. The International Trust for Zological Nomenclature, c / o The Natural History Museum, London, xxix + 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
, and the taxon is here raised to species rank.
</paragraph>
<caption id="8FAB6631FF9769772AABFAB0FE18FA76" httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="13" pageNumber="58" targetBox="[206,1406,407,1211]" targetPageId="13">
<paragraph id="DB6B36B9FF9769772AABFAB0FE18FA76" blockId="13.[151,1437,1280,1456]" pageId="13" pageNumber="58">
<emphasis id="E9A0EAABFF9769772AABFAB0FEF3FAD0" bold="true" box="[151,341,1280,1303]" pageId="13" pageNumber="58">FIGURES 30–37.</emphasis>
Male terminalia. 30–32—
<taxonomicName id="1CD44D3AFF976977285AFAB2FCBCFAD1" box="[614,794,1282,1303]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF976977285AFAB2FCBCFAD1" box="[614,794,1282,1303]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
(Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33—
<taxonomicName id="1CD44D3AFF97697728F7FA91FCCFFAF0" box="[715,873,1312,1334]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF97697728F7FA91FCCFFAF0" box="[715,873,1312,1334]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
(Czech Republic: Moravičany), aedeagal complex, lateral. 34–36—
<taxonomicName id="1CD44D3AFF9769772B18FA8FFE53FA92" box="[292,501,1342,1364]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="suboccultus" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772B18FA8FFE53FA92" box="[292,501,1342,1364]" italics="true" pageId="13" pageNumber="58">M. (M.) suboccultus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9769772BC2FA8FFD9FFA92" bold="true" box="[510,569,1343,1364]" pageId="13" pageNumber="58">
<taxonomicNameLabel id="F29357D0FF9769772BC2FA8FFD9FFA92" box="[510,569,1343,1364]" pageId="13" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37––
<taxonomicName id="1CD44D3AFF9769772BB5FAEEFD97FAB5" box="[393,561,1373,1395]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772BB5FAEEFD97FAB5" box="[393,561,1373,1395]" italics="true" pageId="13" pageNumber="58">M. (M.) occultus</emphasis>
</taxonomicName>
(Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe.
</paragraph>
</caption>
<paragraph id="DB6B36B9FF9769772AF9FA6DFD49F89B" blockId="13.[151,1436,1501,2030]" pageId="13" pageNumber="58">
Both in general appearance and the structure of the male terminalia,
<taxonomicName id="1CD44D3AFF97697729FCFA6EFB22FA33" box="[960,1156,1501,1525]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF97697729FCFA6EFB22FA33" box="[960,1156,1501,1525]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
much resembles
<taxonomicName id="1CD44D3AFF9769772F7AFA6EFF4EF9DF" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772F7AFA6EFF4EF9DF" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
. It differs from the latter by various details of the male terminalia, such as overall shape of the gonocoxite (more slender in
<taxonomicName id="1CD44D3AFF9769772B6BF996FE1BF9FB" box="[343,445,1574,1597]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="E9A0EAABFF9769772B6BF996FE1BF9FB" box="[343,445,1574,1597]" italics="true" pageId="13" pageNumber="58">testaceus</emphasis>
</taxonomicName>
), the size of the dorsal lobe (somewhat smaller in
<taxonomicName id="1CD44D3AFF97697729CBF996FBFBF9FB" box="[1015,1117,1574,1597]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="E9A0EAABFF97697729CBF996FBFBF9FB" box="[1015,1117,1574,1597]" italics="true" pageId="13" pageNumber="58">testaceus</emphasis>
</taxonomicName>
), the length of the aedeagus (shorter in
<taxonomicName id="1CD44D3AFF9769772B2FF9FAFEDCF9A7" box="[275,378,1610,1633]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="E9A0EAABFF9769772B2FF9FAFEDCF9A7" box="[275,378,1610,1633]" italics="true" pageId="13" pageNumber="58">testaceus</emphasis>
</taxonomicName>
), and especially by the shape of the inner gonostylus, which, in
<taxonomicName id="1CD44D3AFF9769772E6FF9FAFAA5F9A7" box="[1107,1283,1609,1633]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772E6FF9FAFAA5F9A7" box="[1107,1283,1609,1633]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
, is extremely powerful, with distinct, pale flanges on both margins along mid-length, whereas, in
<taxonomicName id="1CD44D3AFF9769772E75F9DEFAA8F943" box="[1097,1294,1645,1669]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772E75F9DEFAA8F943" box="[1097,1294,1645,1669]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
, it is shorter and more slender, with flanges that are not very distinct (cf.
<figureCitation id="43EF2A3CFF9769772909F921FBA0F96F" box="[821,1030,1681,1705]" captionStart="FIGURES 30 – 37" captionStartId="13.[151,264,1280,1303]" captionTargetBox="[206,1406,407,1211]" captionTargetId="figure@13.[151,1436,355,1259]" captionTargetPageId="13" captionText="FIGURES 30 – 37. Male terminalia. 30 – 32 — M. (M.) testaceus (Spain: Casapalma), general view, lateral (30), aedeagal complex, lateral (31), and aedeagal plate, ventral (32). 33 — M. (M.) griseus (Czech Republic: Moravičany), aedeagal complex, lateral. 34 – 36 — M. (M.) suboccultus sp. n. (holotype), general view, lateral (34), aedeagal complex, lateral (35), and aedeagal plate, ventral (36). 37 –– M. (M.) occultus (Czech Republic: Jeseníky Mts, Skřítek), aedeagal complex, lateral. Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; dl—dorsal lobe; ig—inner gonostylus; le—lateral excision; ll—lateral lobe; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203290/files/figure.png" pageId="13" pageNumber="58">Figs 30–32 and 33</figureCitation>
). Similarly,
<taxonomicName id="1CD44D3AFF9769772EB2F922FAF7F96F" box="[1166,1361,1681,1705]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772EB2F922FAF7F96F" box="[1166,1361,1681,1705]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
seems to be closely related to
<taxonomicName id="1CD44D3AFF9769772B9FF906FCEEF90B" authority="Savchenko, 1976" authorityName="Savchenko" authorityYear="1976" box="[419,840,1717,1741]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="subgriseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772B9FF906FDDDF90B" box="[419,635,1717,1741]" italics="true" pageId="13" pageNumber="58">M. (M.) subgriseus</emphasis>
Savchenko, 1976
</taxonomicName>
, from
<collectingCountry id="A3C37629FF97697729A8F905FBB5F90B" box="[916,1043,1717,1741]" name="Azerbaijan" pageId="13" pageNumber="58">Azerbaijan</collectingCountry>
. In the latter species, however, the lateral lobe of the gonocoxite is elongate, darkly pigmented at tip and the inner gonostylus is strongly bent at base, directed ventrally (cf. Savchenko 1976,
<figureCitation id="43EF2A3CFF976977285FF94DFD0FF8D3" box="[611,681,1789,1813]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="13" pageNumber="58">Fig. 7</figureCitation>
).
<taxonomicName id="1CD44D3AFF97697728FCF94EFC2EF8D3" box="[704,904,1789,1813]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF97697728FCF94EFC2EF8D3" box="[704,904,1789,1813]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
and
<taxonomicName id="1CD44D3AFF97697729F8F94EFBD1F8D3" box="[964,1143,1789,1813]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF97697729F8F94EFBD1F8D3" box="[964,1143,1789,1813]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
also slightly differ in the body colouration. Whereas the latter has rather an ochre cast, the former tends to be yellowish brown to pale brown. This makes
<taxonomicName id="1CD44D3AFF9769772B48F8F6FD9EF89B" box="[372,568,1861,1885]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="testaceus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772B48F8F6FD9EF89B" box="[372,568,1861,1885]" italics="true" pageId="13" pageNumber="58">M. (M.) testaceus</emphasis>
</taxonomicName>
appear brighter.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9769772AF9F8D8FBD4F82B" pageId="13" pageNumber="58" type="distribution">
<paragraph id="DB6B36B9FF9769772AF9F8D8FBD4F82B" blockId="13.[151,1436,1501,2030]" pageId="13" pageNumber="58">
<emphasis id="E9A0EAABFF9769772AF9F8D8FEF8F847" bold="true" box="[197,350,1896,1921]" pageId="13" pageNumber="58">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9769772B59F8D9FE0EF847" box="[357,424,1897,1921]" name="Spain" pageId="13" pageNumber="58">Spain</collectingCountry>
. First records for
<collectingCountry id="A3C37629FF976977284CF8D9FD75F847" box="[624,723,1897,1921]" name="Portugal" pageId="13" pageNumber="58">Portugal</collectingCountry>
. If Lackschewitz’s (1940a)
<taxonomicName id="1CD44D3AFF9769772E36F8DAFB2DF847" box="[1034,1163,1897,1921]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="bifilatus">
<emphasis id="E9A0EAABFF9769772E36F8DAFB2DF847" box="[1034,1163,1897,1921]" italics="true" pageId="13" pageNumber="58">M. bifilatus</emphasis>
</taxonomicName>
<taxonomicName id="1CD44D3AFF9769772EAFF8DAFAB9F847" box="[1171,1311,1898,1921]" pageId="13" pageNumber="58" rank="variety" variety="testacea">
var.
<emphasis id="E9A0EAABFF9769772EFFF8DAFAB9F847" box="[1219,1311,1898,1921]" italics="true" pageId="13" pageNumber="58">testacea</emphasis>
</taxonomicName>
is the only record for
<taxonomicName id="1CD44D3AFF9769772B2CF83EFE67F863" box="[272,449,1933,1957]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9769772B2CF83EFE67F863" box="[272,449,1933,1957]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
from the continental
<collectingCountry id="A3C37629FF9769772884F83DFD5EF863" box="[696,760,1933,1957]" name="Spain" pageId="13" pageNumber="58">Spain</collectingCountry>
in the recent checklist (
<bibRefCitation id="BF454B48FF9769772E35F83DFB4DF863" author="Eiroa" box="[1033,1259,1933,1957]" pageId="13" pageNumber="58" refString="Eiroa, E. &amp; Baez, M. (2002) Limoniidae. In: Carles-Tolra, M. (Ed.), Catalogo de los Diptera de Espana, Portugal y Andorra (Insecta). Monografia Sociedad Entomologica Aragonesa, vol. 8, Zaragoza, pp. 54 - 57. (in Spanish)." type="journal article" year="2002">Eiroa &amp; Báez 2002</bibRefCitation>
), then the latter species should be excluded from the list. On the other hand, the record of
<taxonomicName id="1CD44D3AFF97697729EBF802FB20F80F" box="[983,1158,1969,1993]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="griseus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF97697729EBF802FB20F80F" box="[983,1158,1969,1993]" italics="true" pageId="13" pageNumber="58">M. (M.) griseus</emphasis>
</taxonomicName>
from the Canary Islands (
<bibRefCitation id="BF454B48FF9769772AA2F865FEDFF82B" author="Eiroa" box="[158,377,2005,2029]" pageId="13" pageNumber="58" refString="Eiroa, E. &amp; Baez, M. (2002) Limoniidae. In: Carles-Tolra, M. (Ed.), Catalogo de los Diptera de Espana, Portugal y Andorra (Insecta). Monografia Sociedad Entomologica Aragonesa, vol. 8, Zaragoza, pp. 54 - 57. (in Spanish)." type="journal article" year="2002">Eiroa &amp; Báez 2002</bibRefCitation>
) seems to be correct based on the figure by de
<bibRefCitation id="BF454B48FF97697729BBF866FBC9F82B" author="Jong" box="[903,1135,2005,2030]" pageId="13" pageNumber="58" refString="Jong, H. de (1987) Keys for the identification of the Tipuloidea (Insecta, Diptera) recorded for the Canary Islands. Eos, 63, 73 - 92." type="journal article" year="1987">
Jong (1987,
<figureCitation id="43EF2A3CFF9769772E2FF865FBC0F82B" box="[1043,1126,2005,2029]" captionStart="FIGURES 21 – 29" captionStartId="10.[151,264,1140,1163]" captionTargetBox="[191,1414,223,1092]" captionTargetId="figure@10.[151,1436,199,1118]" captionTargetPageId="10" captionText="FIGURES 21 – 29. Male terminalia. 21 – 23 — M. (M.) brevifurcatus sp. n. (paratype; Slovakia: Silická planina), general view, lateral (21), aedeagal complex, lateral (22), and aedeagal plate, ventral (23). 24 – 25 — M. (M.) pleuralis (Slovakia: Stankovany), general view, lateral (24) and aedeagal complex, lateral (25). 26 – 29 –– M. (M.) calabricus sp. n. (holotype), general view, lateral (26), distal part of gonocoxite, ventral (27), aedeagal complex, lateral (28), and aedeagal plate, ventral (29). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203289/files/figure.png" pageId="13" pageNumber="58">Fig. 29</figureCitation>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:classification id="A62DC0B7FD1C6B65D9339FDEA46104F4">journal article</mods:classification>
<mods:identifier id="C0D55C07A2F5E0C68F17C5FC246CC828" type="DOI">10.5281/zenodo.203285</mods:identifier>
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<mods:identifier id="88E4FA7F1AFA393BCC89FC3C40012DFE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EB2AE720427ECEA474099833E78FC8CA" type="Zenodo-Dep">203285</mods:identifier>
</mods:mods>
<treatment id="537D87AFFF9869792AABFAD4FE66FC62" ID-DOI="http://doi.org/10.5281/zenodo.6193274" ID-GBIF-Taxon="119406412" ID-Zenodo-Dep="6193274" LSID="urn:lsid:plazi:treatment:537D87AFFF9869792AABFAD4FE66FC62" httpUri="http://treatment.plazi.org/id/537D87AFFF9869792AABFAD4FE66FC62" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
<subSubSection id="93CE6532FF9869782AABFAD4FEBAFA59" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="DB6B36B9FF9869782AABFAD4FD20FABB" blockId="2.[151,646,1379,1439]" box="[151,646,1379,1406]" pageId="2" pageNumber="47">
<heading id="802381D5FF9869782AABFAD4FD20FABB" bold="true" box="[151,646,1379,1406]" fontSize="11" level="1" pageId="2" pageNumber="47" reason="1">
<emphasis id="E9A0EAABFF9869782AABFAD4FD20FABB" bold="true" box="[151,646,1379,1406]" pageId="2" pageNumber="47">
<taxonomicName id="1CD44D3AFF9869782AABFAD4FD9BFAB8" ID-CoL="43X52" box="[151,573,1380,1406]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9869782AABFAD4FD9BFAB8" bold="true" box="[151,573,1380,1406]" italics="true" pageId="2" pageNumber="47">Molophilus (Molophilus) cypricola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9869782878FAD3FD20FABB" box="[580,646,1379,1405]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9869782AABFA37FEBAFA59" blockId="2.[151,646,1379,1439]" box="[151,284,1415,1439]" pageId="2" pageNumber="47">
<figureCitation id="43EF2A3CFF9869782AABFA37FEBAFA59" box="[151,284,1415,1439]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1185,1208]" captionTargetBox="[211,1427,318,1120]" captionTargetId="figure@2.[151,1436,303,1163]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Male terminalia. 1 – 3 — M. (M.) creticola sp. n. (paratype; Greece: Crete: Kakopetros), general view, lateral (1), aedeagal complex, lateral (2), and aedeagal plate, ventral (3). 4 – 6 — M. (M.) cypricola sp. n. (paratype; Cyprus: Kakopetria), general view, lateral (4), aedeagal complex, lateral (5), and aedeagal plate, ventral (6). Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; ig—inner gonostylus; le—lateral excision; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203286/files/figure.png" pageId="2" pageNumber="47">Figures 4–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9869782AABFA7EFDC2F994" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="DB6B36B9FF9869782AABFA7EFDC2F994" blockId="2.[151,1437,1486,2015]" pageId="2" pageNumber="47">
<emphasis id="E9A0EAABFF9869782AABFA7EFEB7FA21" bold="true" box="[151,273,1486,1511]" pageId="2" pageNumber="47">Diagnosis.</emphasis>
Medium-sized species within
<taxonomicName id="1CD44D3AFF986978284EFA7FFD57FA21" box="[626,753,1487,1511]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF986978284EFA7FFD57FA21" box="[626,753,1487,1511]" italics="true" pageId="2" pageNumber="47">Molophilus</emphasis>
</taxonomicName>
. Body dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, setulose, subacute at tip. Inner gonostylus stouter, obtuse at apex. Wing length 4.6–4.9 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9869792AFBF9EEFE33FE09" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47" type="description">
<paragraph id="DB6B36B9FF9869782AFBF9EEFE9EF906" blockId="2.[151,1437,1486,2015]" pageId="2" pageNumber="47">
<emphasis id="E9A0EAABFF9869782AFBF9EEFE0EF9B1" bold="true" box="[199,424,1630,1655]" pageId="2" pageNumber="47">Description. Male.</emphasis>
Head. Antenna of moderate length, slightly extending beyond wing base, dark brown throughout. Flagellomeres generally ovoid to long-ovoid, with longest verticils about twice as long as their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9869782AFBF97AFD70F851" blockId="2.[151,1437,1486,2015]" pageId="2" pageNumber="47">Thorax dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Prescutum and scutum dark greyish brown, with yellow area lateral to prescutal pit; scutal lobe yellowed posterolaterally. Scutellum yellowed medially, otherwise dark brown. Postscutellum dark greyish brown, yellowed anterolaterally. Pleuron dark greyish brown, yellowed on dorsopleural membrane and basalare. Wing membrane infuscated; venation generally as for genus. Halter pale yellow throughout. Legs with coxae, trochanters and bases of femora yellowish brown, rest of legs darkened, brown to dark brown.</paragraph>
<paragraph id="DB6B36B9FF9869792AFBF812FD2CFE6A" blockId="2.[151,1437,1486,2015]" lastBlockId="3.[151,1436,151,932]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
Abdomen dark brown. Male terminalia (
<figureCitation id="43EF2A3CFF98697828B7F812FD49F87C" box="[651,751,1954,1978]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1185,1208]" captionTargetBox="[211,1427,318,1120]" captionTargetId="figure@2.[151,1436,303,1163]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Male terminalia. 1 – 3 — M. (M.) creticola sp. n. (paratype; Greece: Crete: Kakopetros), general view, lateral (1), aedeagal complex, lateral (2), and aedeagal plate, ventral (3). 4 – 6 — M. (M.) cypricola sp. n. (paratype; Cyprus: Kakopetria), general view, lateral (4), aedeagal complex, lateral (5), and aedeagal plate, ventral (6). Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; ig—inner gonostylus; le—lateral excision; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203286/files/figure.png" pageId="2" pageNumber="47">Figs 4–6</figureCitation>
) dark brown. Dorsal portion of gonocoxite rather long, without dorsal and lateral lobe, subquadrangular in lateral view, parallel-sided and obliquely truncate at apex. Broad, darkened tooth at posteroventral edge of gonocoxite, directed mesally. Lateral excision wide. Ventral lobe of gonocoxite stout and comparatively short, conical, rounded at tip, much shorter than dorsal portion of gonocoxite. Both gonostyli darkly pigmented. Outer gonostylus slender, generally straight or slightly curved ventrally, setulose, subacute at tip, reaching to about two thirds length of inner gonostylus. The latter stouter, exceeding ventral lobe by about two thirds its length, slightly curved ventrally, narrowing distally to obtuse apex. Aedeagus of moderate length and breadth, slightly expanded at about one fourth its length in lateral view and bent ventrally, then gradually tapered to slightly upturned tip. Aedeagal plate large, generally oval in ventral/ventrocaudal aspect, with rather long, stout, curved, blackened median spine.
</paragraph>
<paragraph id="DB6B36B9FF9969792AFBFE08FE33FE09" blockId="3.[151,1436,151,932]" box="[199,405,439,464]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFE08FEB8FE16" bold="true" box="[199,286,440,464]" pageId="3" pageNumber="48">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBFE6BFDB9FD99" pageId="3" pageNumber="48" type="materials_examined">
<paragraph id="DB6B36B9FF9969792AFBFE6BFDB9FD99" blockId="3.[151,1436,151,932]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFE6BFD81FE32" bold="true" box="[199,551,475,500]" pageId="3" pageNumber="48">
Material examined.
<typeStatus id="046F881BFF9969792B86FE6CFD81FE32" box="[442,551,476,500]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF9969792869FE6CFD0FFE32" box="[597,681,476,500]" name="Cyprus" pageId="3" pageNumber="48">Cyprus</collectingCountry>
, Troodos Mts,
<quantity id="1C2C9B5CFF9969792969FE6CFC29FE32" box="[853,911,476,500]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="48" unit="km" value="4.0">4 km</quantity>
SW Kakopetria (
<quantity id="1C2C9B5CFF9969792E69FE6CFB7CFE32" box="[1109,1242,476,500]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="3" pageNumber="48" unit="m" value="925.0" valueMax="950.0" valueMin="900.0">900–950 m</quantity>
),
<date id="AF6A1079FF9969792ECCFE6CFAC9FE32" box="[1264,1391,476,500]" pageId="3" pageNumber="48" value="1992-07-29">29.vii.1992</date>
(H. de Jong leg.) (
<collectionCode id="BDC5AE7CFF9969792B04FE4FFE37FDD1" LSID="urn:lsid:biocol.org:col:34975" box="[312,401,511,535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="3" pageNumber="48">ZMAN</collectionCode>
).
<emphasis id="E9A0EAABFF9969792B99FDB0FDBBFDDE" bold="true" box="[421,541,512,536]" pageId="3" pageNumber="48">
<typeStatus id="046F881BFF9969792B99FDB0FDBBFDDE" box="[421,541,512,536]" pageId="3" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
(3 3):
<emphasis id="E9A0EAABFF996979284FFE4FFD70FDDE" bold="true" box="[627,726,511,536]" pageId="3" pageNumber="48">
<collectingCountry id="A3C37629FF996979284FFE4FFD76FDDE" box="[627,720,511,536]" name="Cyprus" pageId="3" pageNumber="48">Cyprus</collectingCountry>
:
</emphasis>
Troodos Mts, Kakopetria,
<quantity id="1C2C9B5CFF9969792E34FE4FFBE4FDD1" box="[1032,1090,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="48" unit="km" value="2.0">2 km</quantity>
SW, at brook,
<date id="AF6A1079FF9969792ED1FE4FFAC2FDD1" box="[1261,1380,511,535]" pageId="3" pageNumber="48" value="2002-04-11">11.iv.2002</date>
, 1 3 (J. Roháček leg.); Troodos Mts,
<quantity id="1C2C9B5CFF996979283FFD94FD9AFDFA" box="[515,572,548,572]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="48" unit="km" value="4.0">4 km</quantity>
SW Kakopetria (
<quantity id="1C2C9B5CFF996979293CFD94FC22FDFA" box="[768,900,548,572]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="3" pageNumber="48" unit="m" value="925.0" valueMax="950.0" valueMin="900.0">900–950 m</quantity>
),
<date id="AF6A1079FF99697929A5FD94FBBCFDFA" box="[921,1050,548,572]" pageId="3" pageNumber="48" value="1992-07-29">29.vii.1992</date>
, 1 3, (
<quantity id="1C2C9B5CFF9969792E5BFD94FB4DFDFA" box="[1127,1259,548,572]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="48" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</quantity>
),
<date id="AF6A1079FF9969792F3CFD94FA27FDFA" box="[1280,1409,548,572]" pageId="3" pageNumber="48" value="1992-07-30">30.vii.1992</date>
, 1 3 (H. de Jong leg.) (
<collectionCode id="BDC5AE7CFF9969792B41FDF7FE09FD99" box="[381,431,583,607]" pageId="3" pageNumber="48">JSO</collectionCode>
,
<collectionCode id="BDC5AE7CFF9969792B87FDF7FDB5FD99" LSID="urn:lsid:biocol.org:col:34975" box="[443,531,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="3" pageNumber="48">ZMAN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBFDDCFDDFFD0A" pageId="3" pageNumber="48" type="etymology">
<paragraph id="DB6B36B9FF9969792AFBFDDCFDDFFD0A" blockId="3.[151,1436,151,932]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFDDCFEE8FD42" bold="true" box="[199,334,620,644]" pageId="3" pageNumber="48">Etymology.</emphasis>
The compound name of this new species,
<taxonomicName id="1CD44D3AFF996979290BFDDCFC07FD42" box="[823,929,620,644]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="cypricola">
<emphasis id="E9A0EAABFF996979290BFDDCFC07FD42" box="[823,929,620,644]" italics="true" pageId="3" pageNumber="48">cypricola</emphasis>
</taxonomicName>
, refers to its occurrence in
<collectingCountry id="A3C37629FF9969792EE0FDDCFA96FD42" box="[1244,1328,620,644]" name="Cyprus" pageId="3" pageNumber="48">Cyprus</collectingCountry>
; the final component of the name derives from the Latin incola (= the dweller, inhabitant). A noun in nominative singular, standing in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBFD67FE97FCB9" pageId="3" pageNumber="48" type="discussion">
<paragraph id="DB6B36B9FF9969792AFBFD67FE97FCB9" blockId="3.[151,1436,151,932]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFD67FEEAFD36" bold="true" box="[199,332,727,752]" pageId="3" pageNumber="48">Discussion.</emphasis>
<taxonomicName id="1CD44D3AFF9969792B6FFD69FDBDFD36" box="[339,539,728,752]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9969792B6FFD69FDBDFD36" box="[339,539,728,752]" italics="true" pageId="3" pageNumber="48">M. (M.) cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF996979281EFD68FDC4FD36" bold="true" box="[546,610,728,752]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="F29357D0FF996979281EFD68FDC4FD36" box="[546,610,728,752]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is closest to
<taxonomicName id="1CD44D3AFF99697928C8FD69FC12FD36" box="[756,948,728,752]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF99697928C8FD69FC12FD36" box="[756,948,728,752]" italics="true" pageId="3" pageNumber="48">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9969792980FD68FC5AFD36" bold="true" box="[956,1020,728,752]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="F29357D0FF9969792980FD68FC5AFD36" box="[956,1020,728,752]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
The differences are specified in Discussion of the latter. Both species were first recognised by H. Mendl, but not formally described by him. Years ago, he sent me his unnamed drawings. Whereas his specimens of
<taxonomicName id="1CD44D3AFF996979296BFC91FBBFFCFE" box="[855,1049,800,824]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF996979296BFC91FBBFFCFE" box="[855,1049,800,824]" italics="true" pageId="3" pageNumber="48">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9969792E1EFC90FBC5FCFE" bold="true" box="[1058,1123,800,824]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="F29357D0FF9969792E1EFC90FBC5FCFE" box="[1058,1123,800,824]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are deposited in his collection, now in ZFMK (see
<typeStatus id="046F881BFF9969792B8EFCF5FDBBFC9B" box="[434,541,837,861]" pageId="3" pageNumber="48" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="1CD44D3AFF9969792879FCF4FD0EFC9A" box="[581,680,836,860]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov.">
<emphasis id="E9A0EAABFF9969792879FCF4FD0EFC9A" box="[581,680,836,860]" italics="true" pageId="3" pageNumber="48">creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9969792893FCF4FD56FC9A" bold="true" box="[687,752,836,860]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="F29357D0FF9969792893FCF4FD56FC9A" box="[687,752,836,860]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
),
<taxonomicName id="1CD44D3AFF9969792934FCF5FC77FC9A" box="[776,977,836,860]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9969792934FCF5FC77FC9A" box="[776,977,836,860]" italics="true" pageId="3" pageNumber="48">M. (M.) cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF99697929E5FCF4FBBFFC9A" bold="true" box="[985,1049,836,860]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="F29357D0FF99697929E5FCF4FBBFFC9A" box="[985,1049,836,860]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
was not traced there (N. Dorchin, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBFC3BFE66FC62" box="[199,448,907,932]" pageId="3" pageNumber="48" type="distribution">
<paragraph id="DB6B36B9FF9969792AFBFC3BFE66FC62" blockId="3.[151,1436,151,932]" box="[199,448,907,932]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFC3BFEC6FC62" bold="true" box="[199,352,907,932]" pageId="3" pageNumber="48">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9969792B5BFC3CFE1AFC62" box="[359,444,908,932]" name="Cyprus" pageId="3" pageNumber="48">Cyprus</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

208
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<mods:namePart id="2A1183BE872B90010AC99619FB67047F">Starý, Jaroslav</mods:namePart>
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<mods:typeOfResource id="B6D38877740C84146B7A5E206DE1B243">text</mods:typeOfResource>
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<mods:title id="F4A57FEFB64890B0C4EA1E6A1004D8FE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8A3EE35E37C13AF54C9072DF6011E76A">
<mods:date id="4547E8F0DFCF86A618051DB2209EB70A">2011</mods:date>
<mods:detail id="4BB704D7B55D87477D19F78D909668DC" type="volume">
<mods:number id="B21261895107F914728931FD6A5146AB">2999</mods:number>
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<mods:start id="489D86365365B7666F7E10FD348F261D">45</mods:start>
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<mods:classification id="C90414330FB88D0DF983E63044BBEB91">journal article</mods:classification>
<mods:identifier id="4E000F942B8E1CAA02181359E72BFC67" type="DOI">10.5281/zenodo.203285</mods:identifier>
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<treatment id="537D87AFFF99697E2AABFC5FFE68FD61" ID-DOI="http://doi.org/10.5281/zenodo.6193276" ID-GBIF-Taxon="119406413" ID-Zenodo-Dep="6193276" LSID="urn:lsid:plazi:treatment:537D87AFFF99697E2AABFC5FFE68FD61" httpUri="http://treatment.plazi.org/id/537D87AFFF99697E2AABFC5FFE68FD61" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
<subSubSection id="93CE6532FF9969792AABFC5FFEBAFBEC" pageId="3" pageNumber="48" type="nomenclature">
<paragraph id="DB6B36B9FF9969792AABFC5FFDD9FBCF" blockId="3.[151,639,1007,1066]" box="[151,639,1007,1033]" pageId="3" pageNumber="48">
<heading id="802381D5FF9969792AABFC5FFDD9FBCF" bold="true" box="[151,639,1007,1033]" fontSize="11" level="1" pageId="3" pageNumber="48" reason="1">
<emphasis id="E9A0EAABFF9969792AABFC5FFDD9FBCF" bold="true" box="[151,639,1007,1033]" pageId="3" pageNumber="48">
<taxonomicName id="1CD44D3AFF9969792AABFC5FFD90FBCF" ID-CoL="43X3D" box="[151,566,1007,1033]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9969792AABFC5FFD90FBCF" bold="true" box="[151,566,1007,1033]" italics="true" pageId="3" pageNumber="48">Molophilus (Molophilus) carbonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9969792801FC5FFDD9FBCF" box="[573,639,1007,1033]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9969792AABFBA2FEBAFBEC" blockId="3.[151,639,1007,1066]" box="[151,284,1042,1066]" pageId="3" pageNumber="48">
<figureCitation id="43EF2A3CFF9969792AABFBA2FEBAFBEC" box="[151,284,1042,1066]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="3" pageNumber="48">Figures 7–9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AABFBE3FC6BFB11" pageId="3" pageNumber="48" type="diagnosis">
<paragraph id="DB6B36B9FF9969792AABFBE3FC6BFB11" blockId="3.[151,1436,1107,2032]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AABFBE3FEB7FBAA" bold="true" box="[151,273,1107,1132]" pageId="3" pageNumber="48">Diagnosis.</emphasis>
Small species within
<taxonomicName id="1CD44D3AFF9969792832FBE4FD2BFBAA" box="[526,653,1108,1132]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9969792832FBE4FD2BFBAA" box="[526,653,1108,1132]" italics="true" pageId="3" pageNumber="48">Molophilus</emphasis>
</taxonomicName>
. Body deep dark brown, almost blackish, restrictedly patterned with yellow, mostly shiny, locally suffused with greyish pruinosity. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, pointed at tip. Inner gonostylus somewhat stouter, shortly bifurcate at apex. Wing length 3.4–3.8 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBFB53FD47F86E" pageId="3" pageNumber="48" type="description">
<paragraph id="DB6B36B9FF9969792AFBFB53FC4CFAE7" blockId="3.[151,1436,1107,2032]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBFB53FE0EFB3A" bold="true" box="[199,424,1251,1276]" pageId="3" pageNumber="48">Description. Male.</emphasis>
Head. Antenna short, not reaching wing base, dark brown throughout. Flagellomeres ovoid, with longest verticils subequal in length to their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9969792AFBFA9CFCD0F9DA" blockId="3.[151,1436,1107,2032]" pageId="3" pageNumber="48">Thorax deep dark brown, almost blackish, restrictedly patterned with yellow, mostly shiny, locally suffused with sparse greyish pruinosity. Prescutum and scutum deep dark greyish brown, more shiny laterally, with yellow area lateral to prescutal pit, including adjacent part of paratergite; scutal lobe yellowish brown posterolaterally. Scutellum mostly dark brown, sometimes paler, yellowish brown, medially. Postscutellum deep dark greyish brown. Pleuron deep dark brown, mostly shiny, dorsopleural membrane and basalare paler, obscure yellow. Wing membrane infuscated; venation generally as for genus. Stem of halter infuscated, knob pale yellow. Legs with coxae, trochanters and bases of femora brown, rest of legs darker.</paragraph>
<paragraph id="DB6B36B9FF9969792AFBF998FACBF842" blockId="3.[151,1436,1107,2032]" pageId="3" pageNumber="48">
Abdomen deep dark brown. Male terminalia (
<figureCitation id="43EF2A3CFF99697928F0F998FC97F986" box="[716,817,1576,1600]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="3" pageNumber="48">Figs 7–9</figureCitation>
) deep dark brown. Dorsal portion of gonocoxite rather short, without dorsal and lateral lobe, subquadrangular in lateral view, somewhat expanded posterodorsally, broadly rounded at apex, without noticeable process at posteroventral edge. Lateral excision comparatively deep and moderately wide. Ventral lobe of gonocoxite generally slender and long, its somewhat truncate tip level with apex of dorsal portion of gonocoxite. Both gonostyli smooth and pale. Outer gonostylus slender, generally straight, pointed at tip, reaching to about three fourths length of inner gonostylus. The latter exceeding ventral lobe by about one third its length, bent ventrally and tapered to slender, gradually narrowing, slightly upturned distal part with shortly bifurcate apex. Aedeagus of moderate length and breadth, widely and evenly arched at mid-length, with both ends nearly aligned and distal part gradually tapered to slightly upturned tip. Aedeagal plate moderate in size, generally triangular in ventral/ventrocaudal aspect, with rounded apex and barely distinct tooth anterolaterally.
</paragraph>
<paragraph id="DB6B36B9FF9969792AFBF820FD47F86E" blockId="3.[151,1436,1107,2032]" box="[199,737,1936,1960]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBF820FEB8F86E" bold="true" box="[199,286,1936,1960]" pageId="3" pageNumber="48">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9969792AFBF803FBDFF836" pageId="3" pageNumber="48" type="materials_examined">
<paragraph id="DB6B36B9FF9969792AFBF803FBDFF836" blockId="3.[151,1436,1107,2032]" pageId="3" pageNumber="48">
<emphasis id="E9A0EAABFF9969792AFBF803FD82F80A" bold="true" box="[199,548,1971,1996]" pageId="3" pageNumber="48">
Material examined.
<typeStatus id="046F881BFF9969792B8BF804FD82F80A" box="[439,548,1972,1996]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF996979286DF804FD15F80A" box="[593,691,1972,1996]" name="Portugal" pageId="3" pageNumber="48">Portugal</collectingCountry>
, Algarve, Serra de Monchique Mts, Alcaria do Peso (
<quantity id="1C2C9B5CFF9969792F37F804FA29F80A" box="[1291,1423,1972,1996]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="3" pageNumber="48" unit="m" value="475.0" valueMax="500.0" valueMin="450.0">450–500 m</quantity>
),
<date id="AF6A1079FF9969792AABF868FF58F836" box="[151,254,2008,2032]" pageId="3" pageNumber="48" value="2009-04-03">3.iv.2009</date>
(J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9969792B97F868FDA7F836" box="[427,513,2008,2032]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="3" pageNumber="48">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF996979282FF868FD32F836" bold="true" box="[531,660,2008,2032]" pageId="3" pageNumber="48">
<typeStatus id="046F881BFF996979282FF868FD29F836" box="[531,655,2008,2032]" pageId="3" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 3, 1 Ƥ, same data as for
<typeStatus id="046F881BFF99697929FBF868FB8FF836" box="[967,1065,2008,2032]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
(
<collectionCode id="BDC5AE7CFF9969792E05F868FBCBF836" box="[1081,1133,2008,2032]" pageId="3" pageNumber="48">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9E697E2AFBFF28FDB3FF12" pageId="4" pageNumber="49" type="etymology">
<paragraph id="DB6B36B9FF9E697E2AFBFF28FDB3FF12" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="49">
<emphasis id="E9A0EAABFF9E697E2AFBFF28FEE8FF76" bold="true" box="[199,334,152,176]" pageId="4" pageNumber="49">Etymology.</emphasis>
The name of the new species,
<taxonomicName id="1CD44D3AFF9E697E2882FF28FC86FF76" box="[702,800,152,176]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis">
<emphasis id="E9A0EAABFF9E697E2882FF28FC86FF76" box="[702,800,152,176]" italics="true" pageId="4" pageNumber="49">carbonis</emphasis>
</taxonomicName>
, refers to its blackish body colouration (Latin carbo = coal). A noun in genitive singular.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9E697E2AFBFF6FFC9CFD42" pageId="4" pageNumber="49" type="discussion">
<paragraph id="DB6B36B9FF9E697E2AFBFF6FFC9CFD42" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="49">
<emphasis id="E9A0EAABFF9E697E2AFBFF6FFEEAFF3E" bold="true" box="[199,332,223,248]" pageId="4" pageNumber="49">Discussion.</emphasis>
<taxonomicName id="1CD44D3AFF9E697E2B6BFF51FDB9FF3E" box="[343,543,224,248]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2B6BFF51FDB9FF3E" box="[343,543,224,248]" italics="true" pageId="4" pageNumber="49">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E2817FF50FDC8FF3E" bold="true" box="[555,622,224,248]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E2817FF50FDC8FF3E" box="[555,622,224,248]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is closest to
<taxonomicName id="1CD44D3AFF9E697E292CFF51FB79FF31" authority="Lackschewitz, 1940" authorityName="Lackschewitz" authorityYear="1940" box="[784,1247,223,248]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E292CFF51FC4BFF3E" box="[784,1005,224,248]" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
Lackschewitz, 1940
</taxonomicName>
, described from
<collectingCountry id="A3C37629FF9E697E2AABFEB4FF7CFEDA" box="[151,218,260,284]" name="Spain" pageId="4" pageNumber="49">Spain</collectingCountry>
. Whereas the two species only slightly differ in the shape of the gonocoxite (dorsal portion subquadrangular in
<taxonomicName id="1CD44D3AFF9E697E2A88FE98FEB0FE86" box="[180,278,296,320]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov.">
<emphasis id="E9A0EAABFF9E697E2A88FE98FEB0FE86" box="[180,278,296,320]" italics="true" pageId="4" pageNumber="49">carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E2B22FE98FEF8FE86" bold="true" box="[286,350,296,320]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E2B22FE98FEF8FE86" box="[286,350,296,320]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, broadly rounded in
<taxonomicName id="1CD44D3AFF9E697E2877FE99FD64FE86" box="[587,706,297,320]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="E9A0EAABFF9E697E2877FE99FD64FE86" box="[587,706,297,320]" italics="true" pageId="4" pageNumber="49">nigrescens</emphasis>
</taxonomicName>
, cf.
<figureCitation id="43EF2A3CFF9E697E28C9FE97FC36FEF9" box="[757,912,295,319]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="4" pageNumber="49">Figs 7 and 10</figureCitation>
) and the curvature of the gonostyli, the difference in the shape of the aedeagus is quite profound. In
<taxonomicName id="1CD44D3AFF9E697E2939FEFDFC63FEA2" box="[773,965,332,356]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2939FEFDFC63FEA2" box="[773,965,332,356]" italics="true" pageId="4" pageNumber="49">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E29F1FEFCFBABFEA2" bold="true" box="[973,1037,332,356]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E29F1FEFCFBABFEA2" box="[973,1037,332,356]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the aedeagus is widely and evenly arched at mid-length, with both ends approximately in alignment (
<figureCitation id="43EF2A3CFF9E697E29B0FEDFFC74FE41" box="[908,978,367,391]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="4" pageNumber="49">Fig. 8</figureCitation>
). In
<taxonomicName id="1CD44D3AFF9E697E2E34FEC1FB46FE4E" box="[1032,1248,368,392]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2E34FEC1FB46FE4E" box="[1032,1248,368,392]" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
</taxonomicName>
, the aedeagus is markedly modified, being unevenly curved and expanded at mid-length, with an erect, even retrorse tooth at the distal bend ventrally, and the apical part quite straight (
<figureCitation id="43EF2A3CFF9E697E2934FE07FCFCFE09" box="[776,858,439,463]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="4" pageNumber="49">Fig. 11</figureCitation>
). This peculiar shape of the aedeagus clearly separates
<taxonomicName id="1CD44D3AFF9E697E2AEFFE6DFE01FE32" box="[211,423,476,500]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2AEFFE6DFE01FE32" box="[211,423,476,500]" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
</taxonomicName>
from
<taxonomicName id="1CD44D3AFF9E697E2BD0FE6DFD0DFE32" box="[492,683,476,500]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2BD0FE6DFD0DFE32" box="[492,683,476,500]" italics="true" pageId="4" pageNumber="49">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E288EFE6CFD54FE32" bold="true" box="[690,754,476,500]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E288EFE6CFD54FE32" box="[690,754,476,500]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
The aedeagus of
<taxonomicName id="1CD44D3AFF9E697E298BFE6DFB2DFE32" box="[951,1163,476,500]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E298BFE6DFB2DFE32" box="[951,1163,476,500]" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="BF454B48FF9E697E2F66FE6CFEF1FDD1" author="Lackschewitz" pageId="4" pageNumber="49" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a)</bibRefCitation>
as being bayonet-shaped, but has not been adequately illustrated.
<taxonomicName id="1CD44D3AFF9E697E2E03FDB1FB59FDDE" box="[1087,1279,512,536]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2E03FDB1FB59FDDE" box="[1087,1279,512,536]" italics="true" pageId="4" pageNumber="49">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E2F3BFDB0FAE0FDDE" bold="true" box="[1287,1350,512,536]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E2F3BFDB0FAE0FDDE" box="[1287,1350,512,536]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1CD44D3AFF9E697E2F41FDB1FEEFFDFA" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2F41FDB1FEEFFDFA" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
</taxonomicName>
appear to be much similar in external characters, including colouration, although the examined
<typeStatus id="046F881BFF9E697E2AABFDF8FF74FDA6" box="[151,210,584,608]" pageId="4" pageNumber="49">types</typeStatus>
of
<taxonomicName id="1CD44D3AFF9E697E2AC5FDF9FE69FDA6" box="[249,463,584,608]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2AC5FDF9FE69FDA6" box="[249,463,584,608]" italics="true" pageId="4" pageNumber="49">M. (M.) nigrescens</emphasis>
</taxonomicName>
are now somewhat faded. The only possible external difference between the two species is in size;
<taxonomicName id="1CD44D3AFF9E697E2B06FDDDFE5FFD42" box="[314,505,620,644]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2B06FDDDFE5FFD42" box="[314,505,620,644]" italics="true" pageId="4" pageNumber="49">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9E697E283CFDDCFDE6FD42" bold="true" box="[512,576,620,644]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="F29357D0FF9E697E283CFDDCFDE6FD42" box="[512,576,620,644]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
appears to be smaller.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9E697E2AFBFD3FFE68FD61" box="[199,462,655,680]" pageId="4" pageNumber="49" type="distribution">
<paragraph id="DB6B36B9FF9E697E2AFBFD3FFE68FD61" blockId="4.[151,1437,151,680]" box="[199,462,655,680]" pageId="4" pageNumber="49">
<emphasis id="E9A0EAABFF9E697E2AFBFD3FFEC6FD6E" bold="true" box="[199,352,655,680]" pageId="4" pageNumber="49">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9E697E2B5BFD3FFE6CFD61" box="[359,458,655,679]" name="Portugal" pageId="4" pageNumber="49">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

258
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<mods:title id="4834520F7B85798FA09A240D1D9F1F07">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<treatment id="537D87AFFF9B69782AABFF6DFDB2FF31" ID-DOI="http://doi.org/10.5281/zenodo.6193272" ID-GBIF-Taxon="119406410" ID-Zenodo-Dep="6193272" LSID="urn:lsid:plazi:treatment:537D87AFFF9B69782AABFF6DFDB2FF31" httpUri="http://treatment.plazi.org/id/537D87AFFF9B69782AABFF6DFDB2FF31" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
<subSubSection id="93CE6532FF9B697B2AABFF6DFEBAFED1" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="DB6B36B9FF9B697B2AABFF6DFDD9FF31" blockId="1.[151,639,221,279]" box="[151,639,221,247]" pageId="1" pageNumber="46">
<heading id="802381D5FF9B697B2AABFF6DFDD9FF31" bold="true" box="[151,639,221,247]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
<emphasis id="E9A0EAABFF9B697B2AABFF6DFDD9FF31" bold="true" box="[151,639,221,247]" pageId="1" pageNumber="46">
<taxonomicName id="1CD44D3AFF9B697B2AABFF6DFD90FF31" ID-CoL="43X4N" box="[151,566,221,247]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2AABFF6DFD90FF31" bold="true" box="[151,566,221,247]" italics="true" pageId="1" pageNumber="46">Molophilus (Molophilus) creticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9B697B2801FF6DFDD9FF31" box="[573,639,221,247]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9B697B2AABFF4FFEBAFED1" blockId="1.[151,639,221,279]" box="[151,284,255,279]" pageId="1" pageNumber="46">
<figureCitation id="43EF2A3CFF9B697B2AABFF4FFEBAFED1" box="[151,284,255,279]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1185,1208]" captionTargetBox="[211,1427,318,1120]" captionTargetId="figure@2.[151,1436,303,1163]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Male terminalia. 1 – 3 — M. (M.) creticola sp. n. (paratype; Greece: Crete: Kakopetros), general view, lateral (1), aedeagal complex, lateral (2), and aedeagal plate, ventral (3). 4 – 6 — M. (M.) cypricola sp. n. (paratype; Cyprus: Kakopetria), general view, lateral (4), aedeagal complex, lateral (5), and aedeagal plate, ventral (6). Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; ig—inner gonostylus; le—lateral excision; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203286/files/figure.png" pageId="1" pageNumber="46">Figures 1–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9B697B2AABFEF7FDC2FE0A" pageId="1" pageNumber="46" type="diagnosis">
<paragraph id="DB6B36B9FF9B697B2AABFEF7FDC2FE0A" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AABFEF7FEB7FEA6" bold="true" box="[151,273,327,352]" pageId="1" pageNumber="46">Diagnosis.</emphasis>
Medium-sized species within
<taxonomicName id="1CD44D3AFF9B697B284EFEF8FD57FEA6" box="[626,753,328,352]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9B697B284EFEF8FD57FEA6" box="[626,753,328,352]" italics="true" pageId="1" pageNumber="46">Molophilus</emphasis>
</taxonomicName>
. Body dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, glabrous, pointed at tip. Inner gonostylus much stouter, obtuse at apex. Wing length 3.7–5.8 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9B697B2AFBFE67FD47FB5A" pageId="1" pageNumber="46" type="description">
<paragraph id="DB6B36B9FF9B697B2AFBFE67FEA1FDFE" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AFBFE67FE05FE36" bold="true" box="[199,419,471,496]" pageId="1" pageNumber="46">Description. Male.</emphasis>
Head. Antenna of moderate length, extending about to wing base, dark brown throughout. Flagellomeres generally ovoid to long-ovoid, with longest verticils slightly exceeding length of their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9B697B2AFBFDF4FBA4FCC9" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">Thorax dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Prescutum and scutum dark greyish brown, with yellow area lateral to prescutal pit; scutal lobe yellowed anteromedially and posterolaterally. Scutellum mostly yellow, dark greyish brown anteriorly. Postscutellum dark greyish brown, narrowly yellowed laterally. Pleuron dark greyish brown, yellowed on dorsopleural membrane and basalare. Wing membrane infuscated; venation generally as for genus. Halter pale yellow throughout. Legs with coxae, trochanters and bases of femora yellowish brown, rest of legs darkened, brown to dark brown.</paragraph>
<paragraph id="DB6B36B9FF9B697B2AFBFCACFBF1FBBE" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">
Abdomen dark brown. Male terminalia (
<figureCitation id="43EF2A3CFF9B697B28B5FCACFD4AFCF2" box="[649,748,796,820]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1185,1208]" captionTargetBox="[211,1427,318,1120]" captionTargetId="figure@2.[151,1436,303,1163]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Male terminalia. 1 – 3 — M. (M.) creticola sp. n. (paratype; Greece: Crete: Kakopetros), general view, lateral (1), aedeagal complex, lateral (2), and aedeagal plate, ventral (3). 4 – 6 — M. (M.) cypricola sp. n. (paratype; Cyprus: Kakopetria), general view, lateral (4), aedeagal complex, lateral (5), and aedeagal plate, ventral (6). Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; ig—inner gonostylus; le—lateral excision; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203286/files/figure.png" pageId="1" pageNumber="46">Figs 1–3</figureCitation>
) dark brown. Dorsal portion of gonocoxite rather short, without dorsal and lateral lobe, subquadrangular in lateral view, slightly narrowed distally, obliquely truncate at apex. Short, darkened, rounded tooth at posteroventral edge of gonocoxite, directed mesally. Lateral excision rather wide. Ventral lobe of gonocoxite generally short and stout, conical, with subacute tip, much shorter than dorsal portion of gonocoxite. Both gonostyli mostly pale, only weakly pigmented distally. Outer gonostylus slender, glabrous, generally straight or slightly curved dorsally, pointed at tip, reaching to about three fourths length of inner gonostylus. The latter much stouter, exceeding ventral lobe by about two thirds its length, more or less straight, narrowing distally to obtuse apex. Aedeagus of moderate length and breadth, expanded before mid-length in lateral view and bent ventrally, then gradually tapered to slightly upturned tip. Aedeagal plate large, generally oval in ventral/ventrocaudal aspect, with short but comparatively stout, curved, blackened median spine.
</paragraph>
<paragraph id="DB6B36B9FF9B697B2AFBFB34FD47FB5A" blockId="1.[151,1437,327,2008]" box="[199,737,1156,1180]" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AFBFB34FEB8FB5A" bold="true" box="[199,286,1156,1180]" pageId="1" pageNumber="46">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9B697B2AFBFB17FBA8FA51" pageId="1" pageNumber="46" type="materials_examined">
<paragraph id="DB6B36B9FF9B697B2AFBFB17FBA8FA51" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AFBFB17FD83FB06" bold="true" box="[199,549,1191,1216]" pageId="1" pageNumber="46">
Material examined.
<typeStatus id="046F881BFF9B697B2B84FB18FD83FB06" box="[440,549,1192,1216]" pageId="1" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF9B697B286DFB17FD05FB79" box="[593,675,1191,1215]" name="Greece" pageId="1" pageNumber="46">Greece</collectingCountry>
, Crete, Khania Region, Kakopetros, brook,
<date id="AF6A1079FF9B697B2EACFB17FB58FB79" box="[1168,1278,1191,1215]" pageId="1" pageNumber="46" value="2004-05-23">23.v.2004</date>
(J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9B697B2AA3FB7CFF53FB22" box="[159,245,1228,1252]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="1" pageNumber="46">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF9B697B2B3BFB7CFED9FB22" bold="true" box="[263,383,1228,1252]" pageId="1" pageNumber="46">
<typeStatus id="046F881BFF9B697B2B3BFB7CFED9FB22" box="[263,383,1228,1252]" pageId="1" pageNumber="46" type="paratype">Paratypes</typeStatus>
</emphasis>
(35 3, 23 Ƥ):
<emphasis id="E9A0EAABFF9B697B281AFB7BFD25FB22" bold="true" box="[550,643,1227,1252]" pageId="1" pageNumber="46">
<collectingCountry id="A3C37629FF9B697B281AFB7BFDD8FB22" box="[550,638,1227,1252]" name="Greece" pageId="1" pageNumber="46">Greece</collectingCountry>
:
</emphasis>
Crete: Khania Region: Kakopetros (
<quantity id="1C2C9B5CFF9B697B2E20FB7CFBCEFB22" box="[1052,1128,1228,1252]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="46" unit="m" value="400.0">400 m</quantity>
),
<date id="AF6A1079FF9B697B2E40FB7CFB4AFB22" box="[1148,1260,1228,1252]" pageId="1" pageNumber="46" value="1979-05-13">13.v.1979</date>
, 1 3, 1 Ƥ; Kotsifiana (
<quantity id="1C2C9B5CFF9B697B2ACDFB5FFE9BFAC1" box="[241,317,1263,1287]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="46" unit="m" value="500.0">500 m</quantity>
),
<date id="AF6A1079FF9B697B2B6EFB5FFE64FAC1" box="[338,450,1263,1287]" pageId="1" pageNumber="46" value="1977-05-20">20.v.1977</date>
, 2 3, 1 Ƥ; Fassas valley W Chliaro (
<quantity id="1C2C9B5CFF9B697B295FFB5FFC4EFAC1" box="[867,1000,1263,1287]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="1" pageNumber="46" unit="m" value="285.0" valueMax="310.0" valueMin="260.0">260–310 m</quantity>
), 18.–20.v.
<date id="AF6A1079FF9B697B2E5EFB5FFB73FACE" box="[1122,1237,1263,1288]" pageId="1" pageNumber="46" value="1977-01-03">1977, 1 3</date>
, 3 Ƥ (all H. Malicky leg.) (all
<collectionCode id="BDC5AE7CFF9B697B2B00FAA5FE2BFAEA" LSID="urn:lsid:biocol.org:col:34613" box="[316,397,1301,1324]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="46">ZFMK</collectionCode>
; in ethanol); Kakopetros, brook,
<date id="AF6A1079FF9B697B2931FAA4FCDBFAEA" box="[781,893,1300,1324]" pageId="1" pageNumber="46" value="2004-05-23">23.v.2004</date>
, 10 3, 4 Ƥ; Voukolies, brook,
<date id="AF6A1079FF9B697B2ED4FAA4FAFFFAEA" box="[1256,1369,1300,1324]" pageId="1" pageNumber="46" value="2004-05-23">23.v.2004</date>
, 1 3; Alikianos, Xekollimenos River,
<date id="AF6A1079FF9B697B283CFA87FDD7FA89" box="[512,625,1335,1359]" pageId="1" pageNumber="46" value="2004-05-14">14.v.2004</date>
, 3 Ƥ,
<date id="AF6A1079FF9B697B2888FA87FC82FA89" box="[692,804,1335,1359]" pageId="1" pageNumber="46" value="2004-05-17">17.v.2004</date>
, 1 3; Prases,
<quantity id="1C2C9B5CFF9B697B2981FA87FC50FA89" box="[957,1014,1335,1359]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="46" unit="km" value="2.0">2 km</quantity>
W,
<date id="AF6A1079FF9B697B2E1EFA87FB35FA89" box="[1058,1171,1335,1359]" pageId="1" pageNumber="46" value="2004-05-21">21.v.2004</date>
, 8 3, 3 Ƥ; Prases,
<quantity id="1C2C9B5CFF9B697B2F5FFA87FA3AFA89" box="[1379,1436,1335,1359]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="46" unit="km" value="4.0">4 km</quantity>
SW,
<date id="AF6A1079FF9B697B2AF1FAECFE9BFAB2" box="[205,317,1372,1396]" pageId="1" pageNumber="46" value="2004-05-18">18.v.2004</date>
, 2 Ƥ,
<date id="AF6A1079FF9B697B2BB9FAECFE53FAB2" box="[389,501,1372,1396]" pageId="1" pageNumber="46" value="2004-05-21">21.v.2004</date>
, 2 3, 1 Ƥ; Samaria gorge,
<date id="AF6A1079FF9B697B2914FAECFC3FFAB2" box="[808,921,1372,1396]" pageId="1" pageNumber="46" value="2004-05-14">14.v.2004</date>
, 1 3; Stilos, Kiliaris River,
<date id="AF6A1079FF9B697B2EEAFAECFAEEFAB2" box="[1238,1352,1372,1396]" pageId="1" pageNumber="46" value="2004-05-13">13.v.2004</date>
, 5 3, 1 Ƥ; Episkopi, Petres River,
<date id="AF6A1079FF9B697B2BFFFACFFD95FA51" box="[451,563,1407,1431]" pageId="1" pageNumber="46" value="2004-05-16">16.v.2004</date>
, 3 3, 4 Ƥ (all J. Starý leg.) (
<collectionCode id="BDC5AE7CFF9B697B294CFACFFC05FA51" box="[880,931,1407,1431]" pageId="1" pageNumber="46">JSO</collectionCode>
,
<collectionCode id="BDC5AE7CFF9B697B2992FACFFBA4FA51" box="[942,1026,1407,1431]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="1" pageNumber="46">SMOC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9B697B2AFBFA14FD98F9C2" pageId="1" pageNumber="46" type="etymology">
<paragraph id="DB6B36B9FF9B697B2AFBFA14FD98F9C2" blockId="1.[151,1437,327,2008]" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AFBFA14FEE8FA7A" bold="true" box="[199,334,1444,1468]" pageId="1" pageNumber="46">Etymology.</emphasis>
The compound name of this new species,
<taxonomicName id="1CD44D3AFF9B697B2919FA14FC21FA7A" box="[805,903,1444,1468]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola">
<emphasis id="E9A0EAABFF9B697B2919FA14FC21FA7A" box="[805,903,1444,1468]" italics="true" pageId="1" pageNumber="46">creticola</emphasis>
</taxonomicName>
, refers to its occurrence in Crete; the final component of the name derives from the Latin incola (= the dweller, inhabitant). A noun in nominative singular, standing in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9B69782AFBF9BFFCA1FF12" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46" type="discussion">
<paragraph id="DB6B36B9FF9B69782AFBF9BFFCA1FF12" blockId="1.[151,1437,327,2008]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
<emphasis id="E9A0EAABFF9B697B2AFBF9BFFEEAF9EE" bold="true" box="[199,332,1551,1576]" pageId="1" pageNumber="46">Discussion.</emphasis>
The male terminalia of this new species have the dorsal portion of the gonocoxite of generally simple structure, without the dorsal and lateral lobes, thus fitting in the group of species more or less closely related to
<taxonomicName id="1CD44D3AFF9B697B2A8BF9E9FE4DF9A9" authority="Tonnoir" authorityName="Tonnoir" box="[183,491,1623,1648]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="undulatus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2A8BF9E9FE2FF9B6" box="[183,393,1624,1648]" italics="true" pageId="1" pageNumber="46">M. (M.) undulatus</emphasis>
Tonnoir
</taxonomicName>
in
<bibRefCitation id="BF454B48FF9B697B282AF9E7FCD2F9A9" author="Goetghebuer" box="[534,884,1623,1647]" pageId="1" pageNumber="46" refString="Goetghebuer, M. &amp; Tonnoir, A. (1920) Catalogue raisonne des Tipulidae de Belgique. Bulletin de la Societe entomologique de Belgique, 2, 104 - 112, 131 - 147." type="journal article" year="1920">Goetghebuer &amp; Tonnoir, 1920</bibRefCitation>
; most of them having been described relatively recently (
<bibRefCitation id="BF454B48FF9B697B2B3EF9CCFE07F952" author="Caspers" box="[258,417,1660,1684]" pageId="1" pageNumber="46" refString="Caspers, N. (1980) Molophilus franzi sp. n., eine neue Limoniide aus dem Hunsruck (Diptera, Nematocera). Spixiana, 3, 137 - 139." type="journal article" year="1980">Caspers 1980</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9B697B2B93F9CCFD9BF952" author="Mendl" box="[431,573,1660,1684]" pageId="1" pageNumber="46" refString="Mendl, H. (1986) Eine neue Molophilus - Art von Kreta (Diptera Nematocera: Limoniidae). Articulata, 2, 235 - 237." type="journal article" year="1986">Mendl 1986</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9B697B2877F9CCFCAFF952" author="Savchenko" box="[587,777,1660,1684]" pageId="1" pageNumber="46" refString="Savchenko, E. N. (1986) Three new species of the Limoniidae (Diptera) from the USSR fauna. Vestnik zoologii, 1986, 29 - 35. (in Russian, English summary)." type="journal article" year="1986">Savchenko 1986</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9B697B2924F9CCFC31F952" author="Stary" box="[792,919,1660,1684]" pageId="1" pageNumber="46" refString="Stary, J. (1978) Zwei neue europaische Arten aus der Verwandtschaft von Molophilus undulatus (Diptera, Limoniidae). Acta entomologica bohemoslovaca, 75, 344 - 348." type="journal article" year="1978">Starý 1978</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9B697B2999F9CCFC46F952" author="Stary" box="[933,992,1660,1684]" pageId="1" pageNumber="46" refString="Stary, J. (1992) Three new species of the eriopterine Limoniidae from Czechoslovakia (Diptera). C asopis Slezskeho muzea A, 41, 105 - 112." type="journal article" year="1992">1992</bibRefCitation>
,
<bibRefCitation id="BF454B48FF9B697B29D1F9CCFAA2F952" author="Stary" box="[1005,1284,1660,1684]" pageId="1" pageNumber="46" refString="Stary, J. &amp; Freidberg, A. (2007) The Limoniidae of Israel. Israel Journal of Entomology, 37, 301 - 357." type="journal article" year="2007">Starý &amp; Freidberg 2007</bibRefCitation>
). Besides the body colouration and some less conspicuous external traits they differ from each other by various details of the male terminalia, the most indicative of these being the structure of the aedeagus and aedeagal plate (
<bibRefCitation id="BF454B48FF9B697B2F33F974FA36F91A" author="Stary" box="[1295,1424,1732,1756]" pageId="1" pageNumber="46" refString="Stary, J. (1992) Three new species of the eriopterine Limoniidae from Czechoslovakia (Diptera). C asopis Slezskeho muzea A, 41, 105 - 112." type="journal article" year="1992">Starý 1992</bibRefCitation>
).
<taxonomicName id="1CD44D3AFF9B697B2AABF959FEFEF8C6" box="[151,344,1768,1792]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2AABF959FEFEF8C6" box="[151,344,1768,1792]" italics="true" pageId="1" pageNumber="46">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B2B5CF958FE06F8C6" bold="true" box="[352,416,1768,1792]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B2B5CF958FE06F8C6" box="[352,416,1768,1792]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
appears closest to
<taxonomicName id="1CD44D3AFF9B697B284AF959FC98F8C6" box="[630,830,1768,1792]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B284AF959FC98F8C6" box="[630,830,1768,1792]" italics="true" pageId="1" pageNumber="46">M. (M.) cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B297AF958FC20F8C6" bold="true" box="[838,902,1768,1792]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B297AF958FC20F8C6" box="[838,902,1768,1792]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
below, being practically identical with it externally. In
<taxonomicName id="1CD44D3AFF9B697B2AC1F8BDFE67F8E2" box="[253,449,1804,1828]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2AC1F8BDFE67F8E2" box="[253,449,1804,1828]" italics="true" pageId="1" pageNumber="46">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B2BF6F8BCFDAAF8E2" bold="true" box="[458,524,1804,1828]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B2BF6F8BCFDAAF8E2" box="[458,524,1804,1828]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the dorsal portion of the gonocoxite is shorter and somewhat narrowed distally (longer and parallel-sided in
<taxonomicName id="1CD44D3AFF9B697B2BE0F880FDE0F88E" box="[476,582,1840,1864]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov.">
<emphasis id="E9A0EAABFF9B697B2BE0F880FDE0F88E" box="[476,582,1840,1864]" italics="true" pageId="1" pageNumber="46">cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B2872F880FD28F88E" bold="true" box="[590,654,1840,1864]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B2872F880FD28F88E" box="[590,654,1840,1864]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), and the ventral lobe is broader at its base and more sharpened at its tip. The gonostyli of the two species differ in many traits: they are almost pale and generally shorter in
<taxonomicName id="1CD44D3AFF9B697B2F2FF8E5FF71F856" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2F2FF8E5FF71F856" italics="true" pageId="1" pageNumber="46">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B2AE3F8C8FE86F856" bold="true" box="[223,288,1912,1936]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B2AE3F8C8FE86F856" box="[223,288,1912,1936]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(darkly pigmented in
<taxonomicName id="1CD44D3AFF9B697B2823F8C8FD2FF856" box="[543,649,1912,1936]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov.">
<emphasis id="E9A0EAABFF9B697B2823F8C8FD2FF856" box="[543,649,1912,1936]" italics="true" pageId="1" pageNumber="46">cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B28ADF8C8FD74F856" bold="true" box="[657,722,1912,1936]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B28ADF8C8FD74F856" box="[657,722,1912,1936]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), exhibiting further differences in shape. In addition, the outer gonostylus is glabrous in
<taxonomicName id="1CD44D3AFF9B697B2B8FF82DFDD7F872" box="[435,625,1948,1972]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2B8FF82DFDD7F872" box="[435,625,1948,1972]" italics="true" pageId="1" pageNumber="46">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B284BF82CFD10F872" bold="true" box="[631,694,1948,1972]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B284BF82CFD10F872" box="[631,694,1948,1972]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, whereas it is microscopically setulose in
<taxonomicName id="1CD44D3AFF9B697B2EB6F82DFAF7F872" box="[1162,1361,1948,1972]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9B697B2EB6F82DFAF7F872" box="[1162,1361,1948,1972]" italics="true" pageId="1" pageNumber="46">M. (M.) cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9B697B2F6AF82CFA33F872" bold="true" box="[1366,1429,1948,1972]" pageId="1" pageNumber="46">
<taxonomicNameLabel id="F29357D0FF9B697B2F6AF82CFA33F872" box="[1366,1429,1948,1972]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, which appears to be a diagnostic difference. The aedeagal complex likewise shows differences, chiefly in the median spine of the aedeagal plate, which is distinctly shorter in
<taxonomicName id="1CD44D3AFF986978295BFF29FB80FF76" box="[871,1062,152,176]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="creticola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF986978295BFF29FB80FF76" box="[871,1062,152,176]" italics="true" pageId="2" pageNumber="47">M. (M.) creticola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9869782E10FF28FBCAFF76" bold="true" box="[1068,1132,152,176]" pageId="2" pageNumber="47">
<taxonomicNameLabel id="F29357D0FF9869782E10FF28FBCAFF76" box="[1068,1132,152,176]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(cf.
<figureCitation id="43EF2A3CFF9869782EA2FF27FAC4FF69" box="[1182,1378,151,175]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1185,1208]" captionTargetBox="[211,1427,318,1120]" captionTargetId="figure@2.[151,1436,303,1163]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Male terminalia. 1 – 3 — M. (M.) creticola sp. n. (paratype; Greece: Crete: Kakopetros), general view, lateral (1), aedeagal complex, lateral (2), and aedeagal plate, ventral (3). 4 – 6 — M. (M.) cypricola sp. n. (paratype; Cyprus: Kakopetria), general view, lateral (4), aedeagal complex, lateral (5), and aedeagal plate, ventral (6). Scale bar 0.5 mm. aed—aedeagus; aedpl—aedeagal plate; ig—inner gonostylus; le—lateral excision; og—outer gonostylus; vl—ventral lobe." httpUri="https://zenodo.org/record/203286/files/figure.png" pageId="2" pageNumber="47">Figs 1–3 and 4–6</figureCitation>
). For other details, see Discussion of
<taxonomicName id="1CD44D3AFF9869782BC5FF0DFD67FF12" box="[505,705,188,212]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="cypricola" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9869782BC5FF0DFD67FF12" box="[505,705,188,212]" italics="true" pageId="2" pageNumber="47">M. (M.) cypricola</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF98697828FBFF0CFCA1FF12" bold="true" box="[711,775,188,212]" pageId="2" pageNumber="47">
<taxonomicNameLabel id="F29357D0FF98697828FBFF0CFCA1FF12" box="[711,775,188,212]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9869782AFBFF6FFDB2FF31" box="[199,532,223,248]" pageId="2" pageNumber="47" type="distribution">
<paragraph id="DB6B36B9FF9869782AFBFF6FFDB2FF31" blockId="2.[151,1436,151,248]" box="[199,532,223,248]" pageId="2" pageNumber="47">
<emphasis id="E9A0EAABFF9869782AFBFF6FFEC6FF3E" bold="true" box="[199,352,223,248]" pageId="2" pageNumber="47">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9869782B5BFF6FFE10FF31" box="[359,438,223,247]" name="Greece" pageId="2" pageNumber="47">Greece</collectingCountry>
(Crete).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="E9A0EAABFF9D697D2AABFF27FD53FF77" bold="true" box="[151,757,151,177]" pageId="7" pageNumber="52">
<taxonomicName id="1CD44D3AFF9D697D2AABFF27FD88FF77" ID-CoL="6RNW7" box="[151,558,151,177]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9D697D2AABFF27FD88FF77" bold="true" box="[151,558,151,177]" italics="true" pageId="7" pageNumber="52">Molophilus (Molophilus) ibericus</emphasis>
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nom. n., stat. n.
</emphasis>
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</paragraph>
<paragraph id="DB6B36B9FF9D697D2AABFF0AFEC4FF14" blockId="7.[151,757,151,210]" box="[151,354,186,210]" pageId="7" pageNumber="52">
<figureCitation id="43EF2A3CFF9D697D2AABFF0AFEC4FF14" box="[151,354,186,210]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="7" pageNumber="52">Figures 17, 19, 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9D697D2AABFEB2FCFFFEDE" box="[151,857,258,280]" pageId="7" pageNumber="52" type="reference_group">
<paragraph id="DB6B36B9FF9D697D2AABFEB2FCFFFEDE" blockId="7.[151,857,258,280]" box="[151,857,258,280]" pageId="7" pageNumber="52">
<treatmentCitationGroup id="FBC41197FF9D697D2AABFEB2FCFFFEDE" box="[151,857,258,280]" pageId="7" pageNumber="52">
<taxonomicName id="1CD44D3AFF9D697D2AABFEB2FEFDFEDE" ID-CoL="43X8X" box="[151,347,258,280]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="gladius">
<emphasis id="E9A0EAABFF9D697D2AABFEB2FEFDFEDE" box="[151,347,258,280]" italics="true" pageId="7" pageNumber="52">Molophilus gladius</emphasis>
</taxonomicName>
<taxonomicName id="1CD44D3AFF9D697D2B5DFEB3FD7DFEDE" authority="Lackschewitz 1940a: 23" authorityName="Lackschewitz" authorityPageNumber="23" authorityYear="1940" box="[353,731,258,280]" pageId="7" pageNumber="52" rank="variety" variety="obscura">
var.
<emphasis id="E9A0EAABFF9D697D2BB1FEB2FE79FEDE" box="[397,479,258,280]" italics="true" pageId="7" pageNumber="52">obscura</emphasis>
<treatmentCitation id="5A7510A8FF9D697D2BD9FEB2FD7DFEDE" author="Lackschewitz" box="[485,731,258,280]" page="23" pageId="7" pageNumber="52" year="1940">
<bibRefCitation id="BF454B48FF9D697D2BD9FEB2FD1EFEDE" author="Lackschewitz" box="[485,696,258,280]" pageId="7" pageNumber="52" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz 1940a</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
(diagnosis).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9D697D2AABFEF5FDC9FE0F" pageId="7" pageNumber="52" type="diagnosis">
<paragraph id="DB6B36B9FF9D697D2AABFEF5FDC9FE0F" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">
<emphasis id="E9A0EAABFF9D697D2AABFEF5FEB7FE98" bold="true" box="[151,273,325,350]" pageId="7" pageNumber="52">Diagnosis.</emphasis>
Rather small species within
<taxonomicName id="1CD44D3AFF9D697D2869FEF5FD73FE9B" box="[597,725,325,349]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9D697D2869FEF5FD73FE9B" box="[597,725,325,349]" italics="true" pageId="7" pageNumber="52">Molophilus</emphasis>
</taxonomicName>
. Body dark brown, restrictedly patterned with yellow, suffused with greyish pruinosity. Gonocoxite of male terminalia without dorsal lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, spike-shaped, subacute at tip. Inner gonostylus tapered and curved downwards distally, pointed at apex. Wing length 3.3–4.6 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9D697D2AFBFE65FD47FB7B" pageId="7" pageNumber="52" type="description">
<paragraph id="DB6B36B9FF9D697D2AFBFE65FF42FDF3" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">
<emphasis id="E9A0EAABFF9D697D2AFBFE65FE60FE2B" bold="true" box="[199,454,469,494]" pageId="7" pageNumber="52">Redescription. Male.</emphasis>
Head. Antenna (
<figureCitation id="43EF2A3CFF9D697D28A6FE65FD54FE2B" box="[666,754,469,493]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="7" pageNumber="52">Fig. 17</figureCitation>
) short, not reaching wing base, dark brown throughout. Flagellomeres ovoid, with longest verticils slightly exceeding their respective segments; pubescence short and sparse.
</paragraph>
<paragraph id="DB6B36B9FF9D697D2AFBFDF1FB80FCCB" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">Thorax dark brown, restrictedly patterned with yellow, suffused with dense greyish pruinosity. Prescutum and scutum dark greyish brown, with yellow area lateral to prescutal pit, including adjacent part of paratergite; scutal lobe obscure yellow posterolaterally. Scutellum bright yellow, dark greyish brown on margins. Postscutellum dark greyish brown, yellowed anterolaterally. Pleuron dark greyish brown, yellowed on dorsopleural membrane and basalare. Wing membrane slightly infuscated; venation generally as for genus. Halter dark yellow, with paler knob. Legs with coxae, trochanters and bases of femora brown, rest of legs dark brown.</paragraph>
<paragraph id="DB6B36B9FF9D697D2AFBFCA9FD5FFB5F" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">
Abdomen dark greyish brown. Male terminalia (
<figureCitation id="43EF2A3CFF9D697D28D5FCA9FCCDFCF7" box="[745,875,793,817]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="7" pageNumber="52">Figs 19, 20</figureCitation>
) light brown. Dorsal portion of gonocoxite rather short, without dorsal lobe, only expanded posterodorsally and obliquely truncate apically in lateral view, with distinct, setiferous lobule at posteroventral edge. Lateral lobe short, not extending beyond other lobes. Lateral excision rather wide and comparatively shallow. Ventral lobe of gonocoxite of moderate length and breadth, slightly extending beyond apex of dorsal portion of gonocoxite. Both gonostyli darkly pigmented. Outer gonostylus spikeshaped, gently curved ventrally, provided with a few spinules distally, subacute at tip, reaching to about three fourths length of inner gonostylus. The latter smooth, exceeding ventral lobe by about one third its length, tapered and hook-shaped distally, pointed at apex. Aedeagus comparatively short and stout, curved, with middle part arching dorsally, conspicuously expanded in lateral aspect, and with apex drawn out into slender point extending ventrocaudally. Aedeagal plate moderate in size, generally oval in ventral/ventrocaudal aspect, with three dark, slightly curved spines: a median and two posterolateral spines.
</paragraph>
<paragraph id="DB6B36B9FF9D697D2AFBFB15FD47FB7B" blockId="7.[151,1437,325,2005]" box="[199,737,1189,1213]" pageId="7" pageNumber="52">
<emphasis id="E9A0EAABFF9D697D2AFBFB15FEB8FB7B" bold="true" box="[199,286,1189,1213]" pageId="7" pageNumber="52">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9D69722AFBFB7AFBC4FF31" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="materials_examined">
<paragraph id="DB6B36B9FF9D697D2AFBFB7AFC9EF93B" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">
<emphasis id="E9A0EAABFF9D697D2AFBFB7AFE48FB24" bold="true" box="[199,494,1225,1250]" pageId="7" pageNumber="52">
<typeStatus id="046F881BFF9D697D2AFBFB7AFEA4FB24" box="[199,258,1226,1250]" pageId="7" pageNumber="52">Type</typeStatus>
material examined.
</emphasis>
<bibRefCitation id="BF454B48FF9D697D2BC9FB79FCB1FB27" author="Lackschewitz" box="[501,791,1225,1249]" pageId="7" pageNumber="52" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a: 24)</bibRefCitation>
based his “
<taxonomicName id="1CD44D3AFF9D697D29AAFB79FB84FB27" box="[918,1058,1225,1249]" pageId="7" pageNumber="52" rank="variety" variety="obscura">
var.
<emphasis id="E9A0EAABFF9D697D29F4FB79FB84FB27" box="[968,1058,1225,1249]" italics="true" pageId="7" pageNumber="52">obscura</emphasis>
</taxonomicName>
” of
<taxonomicName id="1CD44D3AFF9D697D2E6FFB7AFB6DFB27" box="[1107,1227,1225,1249]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="gladius">
<emphasis id="E9A0EAABFF9D697D2E6FFB7AFB6DFB27" box="[1107,1227,1225,1249]" italics="true" pageId="7" pageNumber="52">M. gladius</emphasis>
</taxonomicName>
(=
<taxonomicName id="1CD44D3AFF9D697D2ECDFB7AFAC9FB27" box="[1265,1391,1226,1249]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="E9A0EAABFF9D697D2ECDFB7AFAC9FB27" box="[1265,1391,1226,1249]" italics="true" pageId="7" pageNumber="52">propinquus</emphasis>
</taxonomicName>
) on the following material: “
<collectingCountry id="A3C37629FF9D697D2B8FFB5DFDB4FAC3" box="[435,530,1261,1285]" name="Spain" pageId="7" pageNumber="52">Spanien</collectingCountry>
, Aragonien, Albarracin, 22.–30.
<date id="AF6A1079FF9D697D29BAFB5DFBAFFAC3" box="[902,1033,1261,1285]" pageId="7" pageNumber="52" value="1924-06-03">VI.1924, 3</date>
ƤƤ
<typeStatus id="046F881BFF9D697D2E75FB5EFB28FAC0" box="[1097,1166,1262,1286]" pageId="7" pageNumber="52">Typen</typeStatus>
(Zerny);
<collectingCountry id="A3C37629FF9D697D2F3DFB5DFACDFAC3" box="[1281,1387,1261,1285]" name="Morocco" pageId="7" pageNumber="52">Marokko</collectingCountry>
, Gr. Atlas, Tachdirt (
<quantity id="1C2C9B5CFF9D697D2B71FAA1FE4BFAEF" box="[333,493,1297,1321]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="7" pageNumber="52" unit="m" value="2550.0" valueMax="2900.0" valueMin="2200.0">2200–2900 m</quantity>
), 11.–19.VII.
<date id="AF6A1079FF9D697D28BCFAA1FD52FAEF" box="[640,756,1297,1321]" pageId="7" pageNumber="52" value="1933-04-03">1933, 4 3</date>
2 Ƥ (Zerny)”. Since, however,
<bibRefCitation id="BF454B48FF9D697D2E65FAA1FAF7FAEF" author="Lackschewitz" box="[1113,1361,1297,1321]" pageId="7" pageNumber="52" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a)</bibRefCitation>
nominated as
<typeStatus id="046F881BFF9D697D2AC7FA86FE90FA88" box="[251,310,1334,1358]" pageId="7" pageNumber="52">types</typeStatus>
only the one male (see note under
<typeStatus id="046F881BFF9D697D28F8FA86FD5DFA88" box="[708,763,1334,1358]" pageId="7" pageNumber="52">Type</typeStatus>
material examined of
<taxonomicName id="1CD44D3AFF9D697D29C7FA86FBC4FA8B" box="[1019,1122,1334,1357]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="E9A0EAABFF9D697D29C7FA86FBC4FA8B" box="[1019,1122,1334,1357]" italics="true" pageId="7" pageNumber="52">testaceus</emphasis>
</taxonomicName>
for the interpretation of the single male symbol) and an unspecified number of females from
<collectingCountry id="A3C37629FF9D697D29B2FAE9FC76FAB7" box="[910,976,1369,1393]" name="Spain" pageId="7" pageNumber="52">Spain</collectingCountry>
, only these are
<typeStatus id="046F881BFF9D697D2EB6FAEAFB56FAB4" box="[1162,1264,1370,1394]" pageId="7" pageNumber="52" type="syntype">syntypes</typeStatus>
; the Moroccan specimens are excluded from the
<typeStatus id="046F881BFF9D697D282CFACEFDE6FA50" box="[528,576,1406,1430]" pageId="7" pageNumber="52">type</typeStatus>
series (
<bibRefCitation id="BF454B48FF9D697D28ABFACDFC6EFA53" author="ICZN" box="[663,968,1405,1429]" pageId="7" pageNumber="52" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th edition. The International Trust for Zological Nomenclature, c / o The Natural History Museum, London, xxix + 306 pp." type="book" year="1999">
<collectionCode id="BDC5AE7CFF9D697D28ABFACDFD7CFA53" box="[663,730,1405,1429]" pageId="7" pageNumber="52">ICZN</collectionCode>
1999, Article 72.4.6
</bibRefCitation>
). At present, 1 3 and 1 Ƥ from “
<collectingCountry id="A3C37629FF9D697D2F05FACDFA3EFA53" box="[1337,1432,1405,1429]" name="Spain" pageId="7" pageNumber="52">Spanien</collectingCountry>
, Aragonien, Albarracin” and 4 3 and 2 Ƥ from “
<collectingCountry id="A3C37629FF9D697D2893FA11FCBDFA7F" box="[687,795,1441,1465]" name="Morocco" pageId="7" pageNumber="52">Marokko</collectingCountry>
, Gr. Atlas, Tachdirt” are deposited in
<collectionCode id="BDC5AE7CFF9D697D2EFEFA12FAB8FA7F" box="[1218,1310,1442,1465]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="52">NHMW</collectionCode>
(P. Sehnal, pers. comm.). I have examined the Spanish and two Moroccan males, and I here select the Spanish male as
<typeStatus id="046F881BFF9D697D2F60FA75FF6DF9C4" pageId="7" pageNumber="52" type="lectotype">lectotype</typeStatus>
.
<emphasis id="E9A0EAABFF9D697D2AE9FA5AFEEDF9C4" bold="true" box="[213,331,1514,1538]" pageId="7" pageNumber="52">
<typeStatus id="046F881BFF9D697D2AE9FA5AFEEDF9C4" box="[213,331,1514,1538]" pageId="7" pageNumber="52" type="lectotype">Lectotype</typeStatus>
</emphasis>
3 (by present designation):
<collectingCountry id="A3C37629FF9D697D28B3FA59FD77F9C7" box="[655,721,1513,1537]" name="Spain" pageId="7" pageNumber="52">Spain</collectingCountry>
, Aragón, Albarracin,
<date id="AF6A1079FF9D697D29FAFA59FBC8F9C7" box="[966,1134,1513,1537]" pageId="7" pageNumber="52" value="1924-06-22" valueMax="1924-06-30" valueMin="1924-06-22">22.–30.vi.1924</date>
(H. Zerny leg.) (
<collectionCode id="BDC5AE7CFF9D697D2F12FA59FA36F9C7" box="[1326,1424,1513,1537]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="52">NHMW</collectionCode>
), labelled “Aragon Albarracin /
<date id="AF6A1079FF9D697D2BD6F9BDFDFAF9E3" box="[490,604,1549,1573]" pageId="7" pageNumber="52">22.–30.VI</date>
’24 Zerny” (printed), “Mol.
<taxonomicName id="1CD44D3AFF9D697D29A0F9BDFB96F9E3" authority="Meij." authorityName="Meij." box="[924,1072,1549,1573]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="gladius">gladius Meij.</taxonomicName>
/
<taxonomicName id="1CD44D3AFF9D697D2E78F9BDFB3DF9E3" box="[1092,1179,1549,1573]" class="Insecta" family="Limoniidae" genus="Erioptera" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="obscura" status="var. nov.">obscura</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9D697D2E9EF9BEFB41F9E3" box="[1186,1255,1550,1573]" pageId="7" pageNumber="52" rank="variety">n. var.</taxonomicNameLabel>
/ det. Lacksch.” (printed, orange). Labelled as
<typeStatus id="046F881BFF9D697D2BCEF981FDFFF98F" box="[498,601,1585,1609]" pageId="7" pageNumber="52" type="lectotype">lectotype</typeStatus>
by the present author (“
<typeStatus id="046F881BFF9D697D294CF981FBB2F98F" box="[880,1044,1585,1609]" pageId="7" pageNumber="52" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="1CD44D3AFF9D697D2E13F981FB16F98F" box="[1071,1200,1585,1609]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">Molophilus</taxonomicName>
/ obscurus Lacksch. stat. n. 3 / J. Starý 2010”; printed red label) and identified as
<taxonomicName id="1CD44D3AFF9D697D2958F9E6FB86F9AB" box="[868,1056,1621,1645]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9D697D2958F9E6FB86F9AB" box="[868,1056,1621,1645]" italics="true" pageId="7" pageNumber="52">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9D697D2E15F9E5FB23F9AB" bold="true" box="[1065,1157,1621,1645]" pageId="7" pageNumber="52">nom. n.</emphasis>
The specimen is micropinned on a piece of foam, with left hind and right fore and hind legs missing; apex of abdomen is cut off. Terminalia mounted by Lackschewitz in
<collectingCountry id="A3C37629FF9D697D283AF92DFDFCF973" box="[518,602,1693,1717]" name="Canada" pageId="7" pageNumber="52">Canada</collectingCountry>
balsam between celluloid slides, pinned with the specimen. The two nontype Moroccan males are listed below under Other material examined. The
<typeStatus id="046F881BFF9D697D29D8F971FBEDF91F" box="[996,1099,1729,1753]" pageId="7" pageNumber="52" type="lectotype">lectotype</typeStatus>
is designated here to stabilize the concept of the name with respect to
<taxonomicName id="1CD44D3AFF9D697D2869F956FC97F93B" box="[597,817,1765,1789]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9D697D2869F956FC97F93B" box="[597,817,1765,1789]" italics="true" pageId="7" pageNumber="52">M. (M.) propinquus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DB6B36B9FF9D697D2AF9F8B9FA3AF813" blockId="7.[151,1437,325,2005]" pageId="7" pageNumber="52">
<emphasis id="E9A0EAABFF9D697D2AF9F8B9FE55F8E4" bold="true" box="[197,499,1801,1826]" pageId="7" pageNumber="52">Other material examined</emphasis>
(75 3, 1 Ƥ):
<emphasis id="E9A0EAABFF9D697D28B3F8B9FD7BF8E4" bold="true" box="[655,733,1801,1826]" pageId="7" pageNumber="52">
<collectingCountry id="A3C37629FF9D697D28B3F8B9FD7EF8E4" box="[655,728,1801,1826]" name="Spain" pageId="7" pageNumber="52">Spain</collectingCountry>
:
</emphasis>
Andalucía: Ronda, Rio Grande,
<date id="AF6A1079FF9D697D2E73F8B9FB1FF8E7" box="[1103,1209,1801,1825]" pageId="7" pageNumber="52" value="2005-04-05">5.iv.2005</date>
, 1 3; El Burgo, Rio Turón,
<date id="AF6A1079FF9D697D2AD5F89DFEF5F883" box="[233,339,1837,1861]" pageId="7" pageNumber="52" value="2005-04-05">5.iv.2005</date>
, 3 3; Guaro (nr. Coin), brook,
<date id="AF6A1079FF9D697D2889F89DFC86F883" box="[693,800,1837,1861]" pageId="7" pageNumber="52" value="2005-04-04">4.iv.2005</date>
, 12 3; Coin, Rio Seco,
<date id="AF6A1079FF9D697D2E0DF89DFB0AF883" box="[1073,1196,1837,1861]" pageId="7" pageNumber="52" value="2005-03-30">30.iii.2005</date>
, 7 3,
<date id="AF6A1079FF9D697D2ECFF89DFAA4F883" box="[1267,1282,1837,1861]" pageId="7" pageNumber="52" value="2005-04-04">4</date>
.iv.2005, 7 3,
<date id="AF6A1079FF9D697D2AABF8E1FF03F8AF" box="[151,165,1873,1897]" pageId="7" pageNumber="52" value="2005-04-07">7</date>
.iv.2005, 19 3; Alhaurin el Grande,
<quantity id="1C2C9B5CFF9D697D2871F8E1FD2CF8AF" box="[589,650,1873,1897]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="52" unit="km" value="5.0">5 km</quantity>
E,
<date id="AF6A1079FF9D697D288AF8E1FC94F8AF" box="[694,818,1873,1897]" pageId="7" pageNumber="52" value="2005-03-30">30.iii.2005</date>
, 1 3; Fuengirola, Rio de Fuengirola,
<date id="AF6A1079FF9D697D2ED1F8E1FAFEF8AF" box="[1261,1368,1873,1897]" pageId="7" pageNumber="52" value="2005-04-06">6.iv.2005</date>
, 2 3; Benalmádena,
<date id="AF6A1079FF9D697D2B03F8C5FE1AF84B" box="[319,444,1909,1933]" pageId="7" pageNumber="52" value="2005-03-28">28.iii.2005</date>
, 13 3,
<date id="AF6A1079FF9D697D2828F8C5FD84F84B" box="[532,546,1909,1933]" pageId="7" pageNumber="52" value="2005-04-09">9</date>
.iv.2005, 3 3; El. Romeral (nr. Cártama), watercourse,
<date id="AF6A1079FF9D697D2EA0F8C5FABEF84B" box="[1180,1304,1909,1933]" pageId="7" pageNumber="52" value="2005-03-31">31.iii.2005</date>
, 1 3 (all J. Starý leg.) (all
<collectionCode id="BDC5AE7CFF9D697D2B76F829FEDBF877" box="[330,381,1945,1969]" pageId="7" pageNumber="52">JSO</collectionCode>
); Sierra Nevada Mts, Veleta road (
<quantity id="1C2C9B5CFF9D697D292AF829FC61F877" box="[790,967,1945,1969]" metricMagnitude="0" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="52" unit="m" value="1.65" valueMax="1.8" valueMin="1.5">1,500–1,800 m</quantity>
),
<date id="AF6A1079FF9D697D29E3F829FBC7F877" box="[991,1121,1945,1969]" pageId="7" pageNumber="52" value="1977-07-10">10.vii.1977</date>
, 1 3, 1 Ƥ (H. Aspöck, H. Rausch &amp; P. Ressl leg.) (
<collectionCode id="BDC5AE7CFF9D697D2B86F80DFDABF813" LSID="urn:lsid:biocol.org:col:34613" box="[442,525,1981,2005]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="7" pageNumber="52">ZFMK</collectionCode>
; in ethanol); Baza env.,
<date id="AF6A1079FF9D697D2914F80DFC3EF813" box="[808,920,1981,2005]" pageId="7" pageNumber="52" value="1979-05-12">12.v.1979</date>
, 1 3 (J. Roháček leg.) (
<collectionCode id="BDC5AE7CFF9D697D2E8FF80DFB41F813" box="[1203,1255,1981,2005]" pageId="7" pageNumber="52">JSO</collectionCode>
).
<emphasis id="E9A0EAABFF9D697D2EC0F80DFAC9F813" bold="true" box="[1276,1391,1981,2005]" pageId="7" pageNumber="52">
<collectingCountry id="A3C37629FF9D697D2EC0F80DFACFF813" box="[1276,1385,1981,2005]" name="Morocco" pageId="7" pageNumber="52">Morocco</collectingCountry>
:
</emphasis>
Gr.
</paragraph>
<paragraph id="DB6B36B9FF9269722AABFF27FBC4FF31" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
Atlas, Tachdirt (
<quantity id="1C2C9B5CFF9269722B70FF27FE5FFF69" box="[332,505,151,175]" metricMagnitude="0" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="8" pageNumber="53" unit="m" value="2.55" valueMax="2.9" valueMin="2.2">2,200–2,900 m</quantity>
), 11.–19.vii.
<date id="AF6A1079FF92697228BEFF27FD53FF76" box="[642,757,151,176]" pageId="8" pageNumber="53" value="1933-02-03">1933, 2 3</date>
(H. Zerny leg.) (
<collectionCode id="BDC5AE7CFF9269722988FF27FBB5FF69" box="[948,1043,151,175]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="53">NHMW</collectionCode>
) [listed as “
<taxonomicName id="1CD44D3AFF9269722EA4FF27FA82FF76" box="[1176,1316,151,176]" pageId="8" pageNumber="53" rank="variety" variety="obscura">
var.
<emphasis id="E9A0EAABFF9269722EF6FF28FA82FF76" box="[1226,1316,152,176]" italics="true" pageId="8" pageNumber="53">obscura</emphasis>
</taxonomicName>
” by
<bibRefCitation id="BF454B48FF9269722F67FF27FEE9FF12" author="Lackschewitz" pageId="8" pageNumber="53" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz 1940a</bibRefCitation>
; non-types (see above)]; Ht Atlas, Massif Toubkal, Imlil,
<quantity id="1C2C9B5CFF9269722E27FF0CFBCFFF12" box="[1051,1129,188,212]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="53" unit="km" value="17.0">17 km</quantity>
S Asni (
<quantity id="1C2C9B5CFF9269722EE6FF0CFA29FF12" box="[1242,1423,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="8" pageNumber="53" unit="m" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1,700–1,900 m</quantity>
),
<date id="AF6A1079FF9269722AABFF6FFEADFF31" box="[151,267,223,247]" pageId="8" pageNumber="53" value="1977-07-07">7.vii.1977</date>
, 2 3 (H. &amp; T. v. Oorschot, E. Houkes &amp; P. Oosterbroek leg.) (
<collectionCode id="BDC5AE7CFF92697229FDFF6FFBBEFF31" LSID="urn:lsid:biocol.org:col:34975" box="[961,1048,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="8" pageNumber="53">ZMAN</collectionCode>
,
<collectionCode id="BDC5AE7CFF9269722E1FFF6FFBF0FF31" box="[1059,1110,223,247]" pageId="8" pageNumber="53">JSO</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9269722AFBFEB4FF57FE41" pageId="8" pageNumber="53" type="etymology">
<paragraph id="DB6B36B9FF9269722AFBFEB4FF57FE41" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
<emphasis id="E9A0EAABFF9269722AFBFEB4FEE8FEDA" bold="true" box="[199,334,260,284]" pageId="8" pageNumber="53">Etymology.</emphasis>
The name of this species,
<taxonomicName id="1CD44D3AFF9269722840FEB4FD70FEDA" box="[636,726,260,284]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="E9A0EAABFF9269722840FEB4FD70FEDA" box="[636,726,260,284]" italics="true" pageId="8" pageNumber="53">ibericus</emphasis>
</taxonomicName>
, was proposed by H. Mendl for his undescribed species, referring to the Iberian Peninsula where Mendl’s specimens (now in ZFMK, see Other material examined) come from, as well as most other material examined here. The name is to be deemed to be a latinized adjective in nominative singular
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9269722AF9FE23FE11FD42" pageId="8" pageNumber="53" type="discussion">
<paragraph id="DB6B36B9FF9269722AF9FE23FE11FD42" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
<emphasis id="E9A0EAABFF9269722AF9FE23FEECFE6A" bold="true" box="[197,330,403,428]" pageId="8" pageNumber="53">Discussion.</emphasis>
<bibRefCitation id="BF454B48FF9269722B6EFE24FDEDFE6A" author="Lackschewitz" box="[338,587,404,428]" pageId="8" pageNumber="53" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a)</bibRefCitation>
established this species as “
<taxonomicName id="1CD44D3AFF92697229B7FE24FBBEFE6A" box="[907,1048,404,428]" pageId="8" pageNumber="53" rank="variety" status="var. nov." variety="obscura">
var.
<emphasis id="E9A0EAABFF9269722982FE24FBBEFE6A" box="[958,1048,404,428]" italics="true" pageId="8" pageNumber="53">obscura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9269722E1CFE25FBCFFE6A" box="[1056,1129,404,428]" pageId="8" pageNumber="53" rank="variety">n. var.</taxonomicNameLabel>
” of
<taxonomicName id="1CD44D3AFF9269722EA6FE25FAB5FE6A" box="[1178,1299,404,428]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="gladius">
<emphasis id="E9A0EAABFF9269722EA6FE25FAB5FE6A" box="[1178,1299,404,428]" italics="true" pageId="8" pageNumber="53">M. gladius</emphasis>
</taxonomicName>
de
<bibRefCitation id="BF454B48FF9269722F02FE24FF69FE09" author="Meijere" pageId="8" pageNumber="53" refString="Meijere, J. C. H. de (1920) Studien uber palaearktische, vorwiegend hollandische, Limnobiiden, insbesondere uber ihre Kopulationsorgane (Fortsetzung). Tijdschrift voor Entomologie, 63, 46 - 86." type="journal article" year="1920">Meijere, 1920</bibRefCitation>
[=
<taxonomicName id="1CD44D3AFF9269722AC3FE09FDF1FE09" authority="Egger, 1863" authorityName="Egger" authorityYear="1863" box="[255,599,439,464]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="E9A0EAABFF9269722AC3FE09FE01FE16" box="[255,423,441,464]" italics="true" pageId="8" pageNumber="53">M. propinquus</emphasis>
(
<bibRefCitation id="BF454B48FF9269722B86FE07FDE8FE09" author="Egger" box="[442,590,439,463]" pageId="8" pageNumber="53" refString="Egger, J. (1863) Dipterologische Beitrage. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 13, 1101 - 1110." type="journal article" year="1863">Egger, 1863</bibRefCitation>
)
</taxonomicName>
]. The name
<taxonomicName id="1CD44D3AFF92697228D0FE08FCE0FE16" box="[748,838,440,464]" class="Insecta" family="Limoniidae" genus="Erioptera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="E9A0EAABFF92697228D0FE08FCE0FE16" box="[748,838,440,464]" italics="true" pageId="8" pageNumber="53">obscura</emphasis>
</taxonomicName>
is subspecific in terms of Article 45.6.4. of
<bibRefCitation id="BF454B48FF9269722F67FE07FF45FE32" author="ICZN" pageId="8" pageNumber="53" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th edition. The International Trust for Zological Nomenclature, c / o The Natural History Museum, London, xxix + 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
, and the taxon is here raised to species rank. The name, however, is a junior secondary homonym of
<taxonomicName id="1CD44D3AFF9269722F77FE6DFE64FDD1" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" class="Insecta" family="Limoniidae" genus="Erioptera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="E9A0EAABFF9269722F77FE6DFEB8FDDE" italics="true" pageId="8" pageNumber="53">Erioptera obscura</emphasis>
<bibRefCitation id="BF454B48FF9269722B1AFE4FFE64FDD1" author="Meigen" box="[294,450,511,535]" pageId="8" pageNumber="53" refString="Meigen, J. W. (1818) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Aachen, 1, i - xxxvi + 1 - 333." type="journal article" year="1818">Meigen, 1818</bibRefCitation>
</taxonomicName>
(now in
<taxonomicName id="1CD44D3AFF9269722815FDB0FD0FFDDE" box="[553,681,512,536]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9269722815FDB0FD0FFDDE" box="[553,681,512,536]" italics="true" pageId="8" pageNumber="53">Molophilus</emphasis>
</taxonomicName>
). Hence, a new replacement name is proposed here,
<taxonomicName id="1CD44D3AFF9269722EC2FDB0FAFEFDDE" box="[1278,1368,512,536]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="E9A0EAABFF9269722EC2FDB0FAFEFDDE" box="[1278,1368,512,536]" italics="true" pageId="8" pageNumber="53">ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9269722F5CFDB0FF08FDFA" bold="true" pageId="8" pageNumber="53">nom. n.</emphasis>
, based on Mendl’s manuscript name (see Etymology). H. Mendl considered his
<taxonomicName id="1CD44D3AFF9269722E74FD94FB04FDFA" box="[1096,1186,548,572]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="E9A0EAABFF9269722E74FD94FB04FDFA" box="[1096,1186,548,572]" italics="true" pageId="8" pageNumber="53">ibericus</emphasis>
</taxonomicName>
to be a valid species; similarly,
<bibRefCitation id="BF454B48FF9269722B37FDF7FD20FD99" author="Savchenko" box="[267,646,583,607]" pageId="8" pageNumber="53" refString="Savchenko, E. N. (1982) Limoniidae (Eriopterinae). Fauna Ukrainy 14 (3). Akademia Nauk Ukrainskoi SSR, Naukova Dumka, Kiev, 335 pp. (In Ukrainian)." type="book" year="1982">Savchenko (1982: 212, footnote)</bibRefCitation>
suggested Lackschewitz’s “
<taxonomicName id="1CD44D3AFF92697229F2FDF7FBF8FDA6" box="[974,1118,583,608]" pageId="8" pageNumber="53" rank="variety" variety="obscura">
var.
<emphasis id="E9A0EAABFF9269722E38FDF8FBF8FDA6" box="[1028,1118,584,608]" italics="true" pageId="8" pageNumber="53">obscura</emphasis>
</taxonomicName>
” as being possibly distinct from
<taxonomicName id="1CD44D3AFF9269722AE8FDDDFE16FD42" box="[212,432,620,644]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722AE8FDDDFE16FD42" box="[212,432,620,644]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9269722AF9FD21FE6AF9B2" pageId="8" pageNumber="53" type="reference_group">
<paragraph id="DB6B36B9FF9269722AF9FD21FC38FB9E" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
<taxonomicName id="1CD44D3AFF9269722AF9FD21FE07FD6E" ID-CoL="43XKX" box="[197,417,656,680]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722AF9FD21FE07FD6E" box="[197,417,656,680]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
, a common and widely distributed Palaearctic species, and its allies constitute a distinctive species complex distinguished by the structure of the male terminalia, primarily the ventrally curved aedeagus and the aedeagal plate provided with three spines subequal in size.
<taxonomicName id="1CD44D3AFF9269722940FD69FA91FD29" ID-CoL="43WXS" authority="Nielsen, 1963" authorityName="Nielsen" authorityYear="1963" box="[892,1335,727,752]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="alexanderianus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722940FD69FB2BFD36" box="[892,1165,728,752]" italics="true" pageId="8" pageNumber="53">M. (M.) alexanderianus</emphasis>
<bibRefCitation id="BF454B48FF9269722EABFD67FA91FD29" author="Nielsen" box="[1175,1335,727,751]" pageId="8" pageNumber="53" refString="Nielsen, P. (1963) Records and descriptions of Nematocera from Afghanistan. Stuttgarter Beitrage zur Naturkunde, No. 118, 1 - 8." type="book chapter" year="1963">Nielsen, 1963</bibRefCitation>
</taxonomicName>
(Middle Asia),
<taxonomicName id="1CD44D3AFF9269722ADFFD4DFD0DFCD2" ID-CoL="43XL3" authority="Mendl, 1973" authorityName="Mendl" authorityYear="1973" box="[227,683,764,788]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="pseudopropinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722ADFFD4DFDB4FCD2" box="[227,530,764,788]" italics="true" pageId="8" pageNumber="53">M. (M.) pseudopropinquus</emphasis>
<bibRefCitation id="BF454B48FF9269722826FD4CFD0DFCD2" author="Mendl" box="[538,683,764,788]" pageId="8" pageNumber="53" refString="Mendl, H. (1973) Drei neue Eriopterini aus dem Allgau (Diptera, Tipulidae). Nachrichtenblatt der Bayerischen Entomologen, 22, 65 - 74." type="journal article" year="1973">Mendl, 1973</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="A3C37629FF9269722887FD4CFCAAFCD2" box="[699,780,764,788]" name="France" pageId="8" pageNumber="53">France</collectingCountry>
,
<collectingCountry id="A3C37629FF9269722924FD4CFC27FCD2" box="[792,897,764,788]" name="Germany" pageId="8" pageNumber="53">Germany</collectingCountry>
,
<collectingCountry id="A3C37629FF92697229B2FD4CFC50FCD2" box="[910,1014,764,788]" name="Romania" pageId="8" pageNumber="53">Romania</collectingCountry>
,
<collectingCountry id="A3C37629FF9269722E3FFD4CFBCEFCD2" box="[1027,1128,764,788]" name="Slovakia" pageId="8" pageNumber="53">Slovakia</collectingCountry>
,
<collectingCountry id="A3C37629FF9269722E49FD4CFB59FCD2" box="[1141,1279,764,788]" name="Switzerland" pageId="8" pageNumber="53">Switzerland</collectingCountry>
), and
<taxonomicName id="1CD44D3AFF9269722F78FD4DFE67FCF1" ID-CoL="43XKL" authority="Savchenko, 1982" authorityName="Savchenko" authorityYear="1982" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ponticus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722F78FD4DFF50FCFE" italics="true" pageId="8" pageNumber="53">M. (M.) ponticus</emphasis>
<bibRefCitation id="BF454B48FF9269722AC2FCAFFE67FCF1" author="Savchenko" box="[254,449,799,823]" pageId="8" pageNumber="53" refString="Savchenko, E. N. (1982) Limoniidae (Eriopterinae). Fauna Ukrainy 14 (3). Akademia Nauk Ukrainskoi SSR, Naukova Dumka, Kiev, 335 pp. (In Ukrainian)." type="book" year="1982">Savchenko, 1982</bibRefCitation>
</taxonomicName>
(Crimea, North Caucasus, Transcaucasia) belong here and are considered distinct species based on details of the male terminalia. Since, however, a certain genital variation may occur within each species, in particular for the widespread
<taxonomicName id="1CD44D3AFF926972287FFCD9FC85FC46" ID-CoL="43XKX" box="[579,803,872,896]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF926972287FFCD9FC85FC46" box="[579,803,872,896]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
, all the species are more significantly differentiated by their male antennae, which differ considerably among the species by their overall length and the length and density of both verticils and pubescence on the flagellomeres. Some other forms, such as
<taxonomicName id="1CD44D3AFF9269722E08FC01FA3AFC01" ID-CoL="8CW5S" authority="Alexander, 1934" authorityName="Alexander" authorityYear="1934" box="[1076,1436,943,968]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="triacanthus">
<emphasis id="E9A0EAABFF9269722E08FC01FB7FFC0E" box="[1076,1241,944,968]" italics="true" pageId="8" pageNumber="53">M. triacanthus</emphasis>
<bibRefCitation id="BF454B48FF9269722EDEFC1FFA3AFC01" author="Alexander" box="[1250,1436,943,967]" pageId="8" pageNumber="53" refString="Alexander, C. P. (1934) New or little-known Tipulidae from eastern Asia (Diptera), XVIII. The Philippine Journal of Science, 53, 267 - 300." type="journal article" year="1934">Alexander, 1934</bibRefCitation>
</taxonomicName>
(Russian Far East,
<collectingCountry id="A3C37629FF9269722B57FC64FE5CFC2A" box="[363,506,980,1004]" name="North Korea" pageId="8" pageNumber="53">North Korea</collectingCountry>
,
<collectingCountry id="A3C37629FF926972283AFC64FDE1FC2A" box="[518,583,980,1004]" name="Japan" pageId="8" pageNumber="53">Japan</collectingCountry>
) and
<taxonomicName id="1CD44D3AFF92697228B9FC65FBE9FC2A" ID-CoL="8CW5H" authority="Alexander, 1957" authorityName="Alexander" authorityYear="1957" box="[645,1103,980,1004]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="debilispinus">
<emphasis id="E9A0EAABFF92697228B9FC65FC94FC2A" box="[645,818,980,1004]" italics="true" pageId="8" pageNumber="53">M. debilispinus</emphasis>
<bibRefCitation id="BF454B48FF9269722906FC64FC55FC2A" author="Alexander" box="[826,1011,980,1004]" pageId="8" pageNumber="53" refString="Alexander, C. P. (1957) Records and descriptions of Japanese Tipulidae (Diptera), Part V. The crane-flies of Honshu, I. The Philippine Journal of Science, 85, 349 - 388." type="journal article" year="1957">Alexander, 1957</bibRefCitation>
(
<collectingCountry id="A3C37629FF9269722E3EFC64FBE1FC2A" box="[1026,1095,980,1004]" name="Japan" pageId="8" pageNumber="53">Japan</collectingCountry>
)
</taxonomicName>
, are insufficiently known and currently treated as subspecies of
<taxonomicName id="1CD44D3AFF9269722822FC49FD58FBD6" ID-CoL="43XKX" box="[542,766,1016,1040]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722822FC49FD58FBD6" box="[542,766,1016,1040]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="BF454B48FF9269722903FC47FBB6FBC9" author="Oosterbroek" box="[831,1040,1015,1039]" pageId="8" pageNumber="53" refString="Oosterbroek, P. (2010) Catalogue of the Craneflies of the World (Insecta, Diptera, Nematocera, Tipuloidea). Version 20 Feb 2010 [online]. Available from http: // ip 30. eti. uva. nl / ccw /" type="journal article" year="2010">Oosterbroek 2010</bibRefCitation>
). A form illustrated by
<bibRefCitation id="BF454B48FF9269722F1DFC47FEF2FBF2" author="Savchenko" pageId="8" pageNumber="53" refString="Savchenko, E. N. (1982) Limoniidae (Eriopterinae). Fauna Ukrainy 14 (3). Akademia Nauk Ukrainskoi SSR, Naukova Dumka, Kiev, 335 pp. (In Ukrainian)." type="book" year="1982">Savchenko (1982, Fig. 98/4)</bibRefCitation>
as „
<emphasis id="E9A0EAABFF9269722BB9FBADFE02FBF2" box="[389,420,1053,1076]" italics="true" pageId="8" pageNumber="53">M.</emphasis>
(s. str.) sp.
<taxonomicName id="1CD44D3AFF9269722818FBACFCA0FBF2" ID-CoL="43XG9" authority="Meijere" authorityName="Meijere" box="[548,774,1052,1076]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus">
cf.
<emphasis id="E9A0EAABFF9269722874FBACFD00FBF2" box="[584,678,1052,1076]" italics="true" pageId="8" pageNumber="53">occultus</emphasis>
Meijere
</taxonomicName>
“ is of doubtful taxonomic status, but could represent a separate taxon, having in fact nothing in common with
<taxonomicName id="1CD44D3AFF92697228E0FBF1FC31FB9E" ID-CoL="43XG9" box="[732,919,1088,1112]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="occultus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF92697228E0FBF1FC31FB9E" box="[732,919,1088,1112]" italics="true" pageId="8" pageNumber="53">M. (M.) occultus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DB6B36B9FF9269722AFBFBD5FE6AF9B2" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
<taxonomicName id="1CD44D3AFF9269722AFBFBD5FED9FBBA" ID-CoL="6RNW7" box="[199,383,1124,1148]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722AFBFBD5FED9FBBA" box="[199,383,1124,1148]" italics="true" pageId="8" pageNumber="53">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9269722BBBFBD4FE44FBBA" bold="true" box="[391,482,1124,1148]" pageId="8" pageNumber="53">nom. n.</emphasis>
clearly fits in this species complex. It differs from
<taxonomicName id="1CD44D3AFF9269722E18FBD5FAA6FBBA" ID-CoL="43XKX" box="[1060,1280,1124,1148]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722E18FBD5FAA6FBBA" box="[1060,1280,1124,1148]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
externally by its darker body colouration and somewhat smaller size, having the antennae shorter, with moderately long verticils and only sparse and short pubescence; the antennae of
<taxonomicName id="1CD44D3AFF92697228C2FB1DFC7CFB02" ID-CoL="43XKX" box="[766,986,1196,1220]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF92697228C2FB1DFC7CFB02" box="[766,986,1196,1220]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
are longer, with somewhat longer verticils and the pubescence conspicuous, long, suberect and dense (cf.
<figureCitation id="43EF2A3CFF92697229BDFB60FB80FB2E" box="[897,1062,1232,1256]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="8" pageNumber="53">Figs 17 and 18</figureCitation>
) [the antennae of
<taxonomicName id="1CD44D3AFF9269722ED3FB60FA3AFB2E" ID-CoL="43WXS" box="[1263,1436,1232,1256]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="alexanderianus">
<emphasis id="E9A0EAABFF9269722ED3FB60FA3AFB2E" box="[1263,1436,1232,1256]" italics="true" pageId="8" pageNumber="53">alexanderianus</emphasis>
</taxonomicName>
are still longer, with flagellomeres subcylindrical, verticils indistinct, and pubescence conspicuous (see
<bibRefCitation id="BF454B48FF9269722F1DFB44FF74FAF6" author="Savchenko" pageId="8" pageNumber="53" refString="Savchenko, E. N. (1982) Limoniidae (Eriopterinae). Fauna Ukrainy 14 (3). Akademia Nauk Ukrainskoi SSR, Naukova Dumka, Kiev, 335 pp. (In Ukrainian)." type="book" year="1982">Savchenko 1982</bibRefCitation>
, Fig. 87/4); those of
<taxonomicName id="1CD44D3AFF9269722B87FAA8FD2FFAF6" ID-CoL="43XL3" box="[443,649,1304,1328]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="pseudopropinquus">
<emphasis id="E9A0EAABFF9269722B87FAA8FD2FFAF6" box="[443,649,1304,1328]" italics="true" pageId="8" pageNumber="53">pseudopropinquus</emphasis>
</taxonomicName>
are moderately long, with verticils twice the length of their respective segments and pubescence conspicuous; the antennae of
<taxonomicName id="1CD44D3AFF9269722923FA8DFCD8FA92" ID-CoL="43XKL" box="[799,894,1341,1364]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis id="E9A0EAABFF9269722923FA8DFCD8FA92" box="[799,894,1341,1364]" italics="true" pageId="8" pageNumber="53">ponticus</emphasis>
</taxonomicName>
are short, long-verticilate and inconspicuously pubescent (see
<bibRefCitation id="BF454B48FF9269722B7DFAD0FE5BFABE" author="Savchenko" box="[321,509,1376,1400]" pageId="8" pageNumber="53" refString="Savchenko, E. N. (1982) Limoniidae (Eriopterinae). Fauna Ukrainy 14 (3). Akademia Nauk Ukrainskoi SSR, Naukova Dumka, Kiev, 335 pp. (In Ukrainian)." type="book" year="1982">Savchenko 1982</bibRefCitation>
, Fig. 87/3)]. In the structure of the male terminalia,
<taxonomicName id="1CD44D3AFF9269722E01FAD1FB55FABE" ID-CoL="6RNW7" box="[1085,1267,1376,1400]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722E01FAD1FB55FABE" box="[1085,1267,1376,1400]" italics="true" pageId="8" pageNumber="53">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9269722EC6FAD0FAF5FABE" bold="true" box="[1274,1363,1376,1400]" pageId="8" pageNumber="53">nom. n.</emphasis>
is distinguished by having the lateral lobe short, not protruding (
<figureCitation id="43EF2A3CFF9269722970FA34FC04FA5A" box="[844,930,1412,1436]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="8" pageNumber="53">Fig. 19</figureCitation>
), whereas, in
<taxonomicName id="1CD44D3AFF9269722E75FA35FA8AFA5A" ID-CoL="43XKX" box="[1097,1324,1412,1436]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722E75FA35FA8AFA5A" box="[1097,1324,1412,1436]" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
, this part extends posteriorly beyond other lobes of the gonocoxite. The outer gonostylus is shorter and subacute at the tip in
<taxonomicName id="1CD44D3AFF9269722AABFA7DFEF7FA22" ID-CoL="6RNW7" box="[151,337,1484,1508]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722AABFA7DFEF7FA22" box="[151,337,1484,1508]" italics="true" pageId="8" pageNumber="53">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9269722B65FA7CFE13FA22" bold="true" box="[345,437,1484,1508]" pageId="8" pageNumber="53">nom. n.</emphasis>
(longer and rather obtuse at tip in
<taxonomicName id="1CD44D3AFF926972297FFA7DFC67FA22" ID-CoL="43XKX" box="[835,961,1485,1508]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="E9A0EAABFF926972297FFA7DFC67FA22" box="[835,961,1485,1508]" italics="true" pageId="8" pageNumber="53">propinquus</emphasis>
</taxonomicName>
, with abundant, black spinules or spinoid setulae present not only at the apex but also at the inner margin along most of the length of the gonostylus), and the inner gonostylus is more slender and less conspicuously curved distally. The aedeagus is more expanded in the lateral view in
<taxonomicName id="1CD44D3AFF9269722B1AF989FE46F996" ID-CoL="6RNW7" box="[294,480,1592,1616]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722B1AF989FE46F996" box="[294,480,1592,1616]" italics="true" pageId="8" pageNumber="53">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9269722BD5F988FDE2F996" bold="true" box="[489,580,1592,1616]" pageId="8" pageNumber="53">nom. n.</emphasis>
, with a longer narrowed point and the spines on the aedeagal plate may be somewhat less pronounced.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9269722AFBF9CFFE81F919" pageId="8" pageNumber="53" type="distribution">
<paragraph id="DB6B36B9FF9269722AFBF9CFFE81F919" blockId="8.[151,1437,151,1759]" pageId="8" pageNumber="53">
<emphasis id="E9A0EAABFF9269722AFBF9CFFEC6F95E" bold="true" box="[199,352,1663,1688]" pageId="8" pageNumber="53">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9269722B56F9CFFE0AF951" box="[362,428,1663,1687]" name="Spain" pageId="8" pageNumber="53">Spain</collectingCountry>
,
<collectingCountry id="A3C37629FF9269722B86F931FD85F95E" box="[442,547,1665,1688]" name="Morocco" pageId="8" pageNumber="53">Morocco</collectingCountry>
. If Lackschewitz’s (1940a)
<taxonomicName id="1CD44D3AFF926972295FF931FC78F95E" box="[867,990,1664,1688]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="gladius">
<emphasis id="E9A0EAABFF926972295FF931FC78F95E" box="[867,990,1664,1688]" italics="true" pageId="8" pageNumber="53">M. gladius</emphasis>
</taxonomicName>
<taxonomicName id="1CD44D3AFF92697229DBF931FBD2F95E" box="[999,1140,1664,1688]" pageId="8" pageNumber="53" rank="variety" variety="obscura">
var.
<emphasis id="E9A0EAABFF9269722E26F930FBD2F95E" box="[1050,1140,1664,1688]" italics="true" pageId="8" pageNumber="53">obscura</emphasis>
</taxonomicName>
is the only record for
<taxonomicName id="1CD44D3AFF9269722F41F931FEEBF97A" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9269722F41F931FEEBF97A" italics="true" pageId="8" pageNumber="53">M. (M.) propinquus</emphasis>
</taxonomicName>
from
<collectingCountry id="A3C37629FF9269722BAEF914FE74F97A" box="[402,466,1700,1724]" name="Spain" pageId="8" pageNumber="53">Spain</collectingCountry>
in the recent checklist (
<bibRefCitation id="BF454B48FF92697228E7F914FC1EF97A" author="Eiroa" box="[731,952,1700,1724]" pageId="8" pageNumber="53" refString="Eiroa, E. &amp; Baez, M. (2002) Limoniidae. In: Carles-Tolra, M. (Ed.), Catalogo de los Diptera de Espana, Portugal y Andorra (Insecta). Monografia Sociedad Entomologica Aragonesa, vol. 8, Zaragoza, pp. 54 - 57. (in Spanish)." type="journal article" year="2002">Eiroa &amp; Báez 2002</bibRefCitation>
), then the latter species should be excluded from the list.
</paragraph>
</subSubSection>
</treatment>
</document>

200
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<paragraph id="DB6B36B9FF9E697E2AABF90BFCEEF912" blockId="4.[151,840,1722,1782]" box="[151,840,1722,1749]" pageId="4" pageNumber="49">
<heading id="802381D5FF9E697E2AABF90BFCEEF912" bold="true" box="[151,840,1722,1749]" fontSize="11" level="1" pageId="4" pageNumber="49" reason="1">
<taxonomicName id="1CD44D3AFF9E697E2AABF90BFCEEF912" ID-CoL="43XFL" authority="Lackschewitz, 1940" authorityName="Lackschewitz" authorityYear="1940" box="[151,840,1722,1749]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9E697E2AABF90BFCEEF912" bold="true" box="[151,840,1722,1749]" pageId="4" pageNumber="49">
<emphasis id="E9A0EAABFF9E697E2AABF90BFDE9F913" bold="true" box="[151,591,1723,1749]" italics="true" pageId="4" pageNumber="49">Molophilus (Molophilus) nigrescens</emphasis>
Lackschewitz, 1940
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9E697E2AABF96EFE9EF930" blockId="4.[151,840,1722,1782]" box="[151,312,1758,1782]" pageId="4" pageNumber="49">
<figureCitation id="43EF2A3CFF9E697E2AABF96EFE9EF930" box="[151,312,1758,1782]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="4" pageNumber="49">Figures 10–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9E697E2AABF896FBD4F8FD" box="[151,1138,1829,1852]" pageId="4" pageNumber="49" type="reference_group">
<paragraph id="DB6B36B9FF9E697E2AABF896FBD4F8FD" blockId="4.[151,1138,1829,1852]" box="[151,1138,1829,1852]" pageId="4" pageNumber="49">
<treatmentCitationGroup id="FBC41197FF9E697E2AABF896FBD4F8FD" box="[151,1138,1829,1852]" pageId="4" pageNumber="49">
<taxonomicName id="1CD44D3AFF9E697E2AABF896FDDEF8FD" ID-CoL="43XFL" authority="Lackschewitz 1940a: 17" authorityName="Lackschewitz" authorityPageNumber="17" authorityYear="1940" box="[151,632,1829,1852]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="E9A0EAABFF9E697E2AABF896FEDAF8FA" box="[151,380,1830,1852]" italics="true" pageId="4" pageNumber="49">Molophilus nigrescens</emphasis>
<treatmentCitation id="5A7510A8FF9E697E2BBEF895FDDEF8FD" author="Lackschewitz" box="[386,632,1829,1851]" page="17" pageId="4" pageNumber="49" year="1940">
<bibRefCitation id="BF454B48FF9E697E2BBEF895FDF3F8FD" author="Lackschewitz" box="[386,597,1829,1851]" pageId="4" pageNumber="49" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz 1940a</bibRefCitation>
: 17
</treatmentCitation>
</taxonomicName>
(description), Pl. 1,
<figureCitation id="43EF2A3CFF9E697E297BF895FC37F8FD" box="[839,913,1829,1851]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="4" pageNumber="49">Figs 10</figureCitation>
a–c (male terminalia).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9E697E2AABF8D8FCCCF82A" pageId="4" pageNumber="49" type="diagnosis">
<paragraph id="DB6B36B9FF9E697E2AABF8D8FCCCF82A" blockId="4.[151,1436,1896,2028]" pageId="4" pageNumber="49">
<emphasis id="E9A0EAABFF9E697E2AABF8D8FEB7F847" bold="true" box="[151,273,1896,1921]" pageId="4" pageNumber="49">Diagnosis.</emphasis>
Medium-sized species within
<taxonomicName id="1CD44D3AFF9E697E2854F8D8FD41F846" box="[616,743,1896,1920]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9E697E2854F8D8FD41F846" box="[616,743,1896,1920]" italics="true" pageId="4" pageNumber="49">Molophilus</emphasis>
</taxonomicName>
. Body dark brown, restrictedly patterned with yellow, mostly shiny, locally suffused with greyish pruinosity. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, pointed at tip. Inner gonostylus somewhat stouter, shortly bifurcate at apex. Wing length 4.7–5.3 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697F2AFBFF27FCEAFE32" pageId="5" pageNumber="50" type="description">
<paragraph id="DB6B36B9FF9F697F2AFBFF27FCEAFE32" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AFBFF27FDF6FF76" bold="true" box="[199,592,151,176]" pageId="5" pageNumber="50">Redescription of male terminalia</emphasis>
(
<figureCitation id="43EF2A3CFF9F697F2862FF27FD47FF69" box="[606,737,151,175]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="5" pageNumber="50">Figs 10–12</figureCitation>
). Dorsal portion of gonocoxite rather short, without dorsal and lateral lobe, broadly rounded at apex in lateral view, without noticeable process at posteroventral edge. Lateral excision comparatively deep and moderately wide. Ventral lobe of gonocoxite generally slender and long, its somewhat narrowed and rounded tip level with apex of dorsal portion of gonocoxite. Both gonostyli smooth and pale. Outer gonostylus slender, generally straight, pointed at tip, reaching about to three fourths length of inner gonostylus. The latter exceeding ventral lobe by about one third its length, bent ventrally and tapered to slender distal part with shortly bifurcate apex. Aedeagus of moderate length and breadth, roughly producing bayonet-like shape, unevenly arched and expanded at mid-length, with erect, even retrorse tooth at distal bend ventrally and with apical part quite straight. Aedeagal plate moderate in size, generally triangular in ventral/ventrocaudal aspect, with rounded apex and barely distinct tooth laterally at mid-length.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697F2AFBFDB0FB34FC0E" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="DB6B36B9FF9F697F2AFBFDB0FB34FC0E" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AFBFDB0FE52FDDE" bold="true" box="[199,500,511,536]" pageId="5" pageNumber="50">
<typeStatus id="046F881BFF9F697F2AFBFDB0FEA4FDDE" box="[199,258,512,536]" pageId="5" pageNumber="50">Type</typeStatus>
material examined.
</emphasis>
The species was described from an unspecified number of specimens [“
<collectingCountry id="A3C37629FF9F697F2F0AFE4FFA31FDD1" box="[1334,1431,511,535]" name="Spain" pageId="5" pageNumber="50">Spanien</collectingCountry>
, Andalusien, Algeciras, 26.IV.–27.
<date id="AF6A1079FF9F697F282EFD94FD20FDFA" box="[530,646,548,572]" pageId="5" pageNumber="50" value="1925-05-03">V.1925, 3</date>
ƤƤ in Mehrzahl (Zerny).” (
<bibRefCitation id="BF454B48FF9F697F29E7FD94FB54FDFA" author="Lackschewitz" box="[987,1266,548,572]" pageId="5" pageNumber="50" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz 1940a: 18</bibRefCitation>
)]. Altogether 5 3 and 5 Ƥ are now deposited in
<collectionCode id="BDC5AE7CFF9F697F283DFDF8FDFBFD99" box="[513,605,584,607]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="50">NHMW</collectionCode>
(P. Sehnal, pers. comm.). I have examined four male
<typeStatus id="046F881BFF9F697F2E88FDF8FABFFDA6" box="[1204,1305,584,608]" pageId="5" pageNumber="50" type="syntype">syntypes</typeStatus>
, all belonging to
<taxonomicName id="1CD44D3AFF9F697F2AE3FDDDFE12FD42" box="[223,436,620,644]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F2AE3FDDDFE12FD42" box="[223,436,620,644]" italics="true" pageId="5" pageNumber="50">M. (M.) nigrescens</emphasis>
</taxonomicName>
, and I here select as
<typeStatus id="046F881BFF9F697F28A3FDDCFCA0FD42" box="[671,774,620,644]" pageId="5" pageNumber="50" type="lectotype">lectotype</typeStatus>
the one dissected and illustrated by me (
<figureCitation id="43EF2A3CFF9F697F2EECFDDCFAF2FD42" box="[1232,1364,620,644]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="5" pageNumber="50">Figs 10–12</figureCitation>
).
<emphasis id="E9A0EAABFF9F697F2F5BFDDCFF44FD6E" bold="true" pageId="5" pageNumber="50">
<typeStatus id="046F881BFF9F697F2F5BFDDCFF44FD6E" pageId="5" pageNumber="50" type="lectotype">Lectotype</typeStatus>
</emphasis>
3 (by present designation):
<collectingCountry id="A3C37629FF9F697F2819FD3FFDC1FD61" box="[549,615,655,679]" name="Spain" pageId="5" pageNumber="50">Spain</collectingCountry>
, Andalucía, Algeciras,
<date id="AF6A1079FF9F697F2953FD3FFBABFD61" box="[879,1037,655,679]" pageId="5" pageNumber="50" value="1925-05-22" valueMax="1925-05-27" valueMin="1925-05-22">22.–27.v.1925</date>
(H. Zerny leg.) (
<collectionCode id="BDC5AE7CFF9F697F2EF2FD3FFA96FD61" box="[1230,1328,655,679]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="50">NHMW</collectionCode>
), labelled “Andalusia / Algeciras /
<date id="AF6A1079FF9F697F2B91FD04FDB3FD0A" box="[429,533,692,716]" pageId="5" pageNumber="50">22.–27.V</date>
’25 Zerny” (printed), “
<taxonomicName id="1CD44D3AFF9F697F2927FD04FC1FFD0A" box="[795,953,692,716]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrescens">M. nigrescens</taxonomicName>
/ nov. sp. / det. Laksch.” (printed, orange). Labelled as
<typeStatus id="046F881BFF9F697F2B22FD67FE22FD29" box="[286,388,727,751]" pageId="5" pageNumber="50" type="lectotype">lectotype</typeStatus>
by the present author (“
<typeStatus id="046F881BFF9F697F28ACFD67FC92FD29" box="[656,820,727,751]" pageId="5" pageNumber="50" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="1CD44D3AFF9F697F2976FD67FC6DFD29" box="[842,971,727,751]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">Molophilus</taxonomicName>
/
<taxonomicName id="1CD44D3AFF9F697F29DDFD67FB62FD29" authority="Lacksch." authorityName="Lacksch." box="[993,1220,727,751]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrescens">nigrescens Lacksch.</taxonomicName>
3 / J. Starý 2010”; printed red label). The specimen is pinned, practically intact, with only apex of abdomen cut off. Terminalia placed in a sealed plastic tube with glycerine, pinned with the specimen.
<emphasis id="E9A0EAABFF9F697F294AFC90FB96FCFE" bold="true" box="[886,1072,800,824]" pageId="5" pageNumber="50">
<typeStatus id="046F881BFF9F697F294AFC90FB8CFCFE" box="[886,1066,800,824]" pageId="5" pageNumber="50" type="paralectotype">Paralectotypes</typeStatus>
:
</emphasis>
3 3, micro-pinned on a piece of foam or pinned, with the same labelling as for
<typeStatus id="046F881BFF9F697F2890FCF4FCB3FC9A" box="[684,789,836,860]" pageId="5" pageNumber="50" type="lectotype">lectotype</typeStatus>
, except for dates (“
<date id="AF6A1079FF9F697F29CDFCF4FBC0FC9A" box="[1009,1126,836,860]" pageId="5" pageNumber="50">28.–30.IV</date>
‘25”, “1.–
<date id="AF6A1079FF9F697F2EE7FCF4FAB3FC9A" box="[1243,1301,836,860]" pageId="5" pageNumber="50">10.V</date>
‘25”, “
<date id="AF6A1079FF9F697F2F56FCF4FF68FCB9" pageId="5" pageNumber="50">12.– 20.V</date>
‘25”), accordingly labelled as
<typeStatus id="046F881BFF9F697F280DFCD7FD75FCB9" box="[561,723,871,895]" pageId="5" pageNumber="50" type="paralectotype">paralectotypes</typeStatus>
by the present author. Two
<typeStatus id="046F881BFF9F697F2E2BFCD7FB1FFCB9" box="[1047,1209,871,895]" pageId="5" pageNumber="50" type="paralectotype">paralectotypes</typeStatus>
have the terminalia mounted by Lackschewitz in
<collectingCountry id="A3C37629FF9F697F2BDDFC3CFD93FC62" box="[481,565,908,932]" name="Canada" pageId="5" pageNumber="50">Canada</collectingCountry>
balsam between celluloid slides, pinned with the specimens. The
<typeStatus id="046F881BFF9F697F2F27FC3CFA24FC62" box="[1307,1410,908,932]" pageId="5" pageNumber="50" type="lectotype">lectotype</typeStatus>
is designated here to stabilize the concept of the name with respect to
<taxonomicName id="1CD44D3AFF9F697F29B0FC01FBEDFC0E" box="[908,1099,944,968]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F29B0FC01FBEDFC0E" box="[908,1099,944,968]" italics="true" pageId="5" pageNumber="50">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9F697F2E6EFC00FB34FC0E" bold="true" box="[1106,1170,944,968]" pageId="5" pageNumber="50">
<taxonomicNameLabel id="F29357D0FF9F697F2E6EFC00FB34FC0E" box="[1106,1170,944,968]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697F2AFBFC63FBADFBF2" pageId="5" pageNumber="50" type="discussion">
<paragraph id="DB6B36B9FF9F697F2AFBFC63FBADFBF2" blockId="5.[151,1437,151,1112]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AFBFC63FEEAFC2A" bold="true" box="[199,332,979,1004]" pageId="5" pageNumber="50">Discussion.</emphasis>
The differences of
<taxonomicName id="1CD44D3AFF9F697F281BFC65FD5AFC2A" box="[551,764,980,1004]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F281BFC65FD5AFC2A" box="[551,764,980,1004]" italics="true" pageId="5" pageNumber="50">M. (M.) nigrescens</emphasis>
</taxonomicName>
from
<taxonomicName id="1CD44D3AFF9F697F297CFC65FBA6FC2A" box="[832,1024,980,1004]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F297CFC65FBA6FC2A" box="[832,1024,980,1004]" italics="true" pageId="5" pageNumber="50">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9F697F2E3BFC64FBE0FC2A" bold="true" box="[1031,1094,980,1004]" pageId="5" pageNumber="50">
<taxonomicNameLabel id="F29357D0FF9F697F2E3BFC64FBE0FC2A" box="[1031,1094,980,1004]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and other details are specified in Discussion of
<taxonomicName id="1CD44D3AFF9F697F2B68FC49FDB5FBD6" box="[340,531,1016,1040]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="carbonis" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F2B68FC49FDB5FBD6" box="[340,531,1016,1040]" italics="true" pageId="5" pageNumber="50">M. (M.) carbonis</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9F697F2826FC48FDFFFBD6" bold="true" box="[538,601,1016,1040]" pageId="5" pageNumber="50">
<taxonomicNameLabel id="F29357D0FF9F697F2826FC48FDFFFBD6" box="[538,601,1016,1040]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It should be emphasized that the inner gonostylus of
<taxonomicName id="1CD44D3AFF9F697F2E92FC49FA25FBD6" box="[1198,1411,1016,1040]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nigrescens" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F2E92FC49FA25FBD6" box="[1198,1411,1016,1040]" italics="true" pageId="5" pageNumber="50">M. (M.) nigrescens</emphasis>
</taxonomicName>
is pale, not darkly pigmented as illustrated by
<bibRefCitation id="BF454B48FF9F697F28BEFBACFCD2FBF2" author="Lackschewitz" box="[642,884,1052,1076]" pageId="5" pageNumber="50" refString="Lackschewitz, P. (1940 a) Die palaarktischen Rhamphidiinen und Eriopterinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien, 50 (1939), 1 - 67." type="journal article" year="1940" yearSuffix="a">Lackschewitz (1940a</bibRefCitation>
,
<figureCitation id="43EF2A3CFF9F697F2943FBACFC74FBF2" box="[895,978,1052,1076]" captionStart="FIGURES 7 – 12" captionStartId="4.[151,264,1559,1582]" captionTargetBox="[211,1394,760,1512]" captionTargetId="figure@4.[151,1436,735,1537]" captionTargetPageId="4" captionText="FIGURES 7 – 12. Male terminalia. 7 – 9 — M. (M.) carbonis sp. n. (paratype), general view, lateral (7), aedeagal complex, lateral (8), and aedeagal plate, ventral (9). 10 – 12 — M. (M.) nigrescens (lectotype), general view, lateral (10), aedeagal complex, lateral (11), and aedeagal plate, ventral (12). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/203287/files/figure.png" pageId="5" pageNumber="50">Figs 10</figureCitation>
a–b).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697F2AFBFB8FFE08FB91" box="[199,430,1087,1112]" pageId="5" pageNumber="50" type="distribution">
<paragraph id="DB6B36B9FF9F697F2AFBFB8FFE08FB91" blockId="5.[151,1437,151,1112]" box="[199,430,1087,1112]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AFBFB8FFEC6FB9E" bold="true" box="[199,352,1087,1112]" pageId="5" pageNumber="50">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9F697F2B5BFB8FFE0CFB91" box="[359,426,1087,1111]" name="Spain" pageId="5" pageNumber="50">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E459C2F1A09B824245D3B6C600D92286" ID-DOI="10.5281/zenodo.203285" ID-GBIF-Dataset="f31dbef8-f48b-4a18-815d-1442ccaf5d15" ID-ISSN="1175-5326" ID-Zenodo-Dep="203285" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460340654504" checkinUser="plazi" docAuthor="Starý, Jaroslav" docDate="2011" docId="537D87AFFF9F697C2AABFB11FE57F82E" docLanguage="en" docName="zt02999p062.pdf" docOrigin="Zootaxa 2999" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Molophilus (Molophilus) hradileki Starý, 2011, sp. n." docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="AF44FFD7FF9A697A2A3CFFB0FFA6FFC6" masterDocTitle="Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)" masterLastPageNumber="62" masterPageNumber="45" pageNumber="50" updateTime="1698260995331" updateUser="plazi">
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<mods:title id="66948532E098A12CD2121179EC901A65">Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae)</mods:title>
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<mods:date id="32BAF966A36075079873119E4CED341F">2011</mods:date>
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<treatment id="537D87AFFF9F697C2AABFB11FE57F82E" ID-DOI="http://doi.org/10.5281/zenodo.6193282" ID-GBIF-Taxon="119406409" ID-Zenodo-Dep="6193282" LSID="urn:lsid:plazi:treatment:537D87AFFF9F697C2AABFB11FE57F82E" httpUri="http://treatment.plazi.org/id/537D87AFFF9F697C2AABFB11FE57F82E" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
<subSubSection id="93CE6532FF9F697F2AABFB11FE91FB1A" pageId="5" pageNumber="50" type="nomenclature">
<paragraph id="DB6B36B9FF9F697F2AABFB11FD22FB7C" blockId="5.[151,644,1184,1244]" box="[151,644,1184,1211]" pageId="5" pageNumber="50">
<heading id="802381D5FF9F697F2AABFB11FD22FB7C" bold="true" box="[151,644,1184,1211]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
<emphasis id="E9A0EAABFF9F697F2AABFB11FD22FB7C" bold="true" box="[151,644,1184,1211]" pageId="5" pageNumber="50">
<taxonomicName id="1CD44D3AFF9F697F2AABFB11FD9DFB7D" ID-CoL="73NVX" box="[151,571,1185,1211]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="hradileki" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9F697F2AABFB11FD9DFB7D" bold="true" box="[151,571,1185,1211]" italics="true" pageId="5" pageNumber="50">Molophilus (Molophilus) hradileki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29357D0FF9F697F287EFB10FD22FB7C" box="[578,644,1184,1210]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6B36B9FF9F697F2AABFB74FE91FB1A" blockId="5.[151,644,1184,1244]" box="[151,311,1220,1244]" pageId="5" pageNumber="50">
<figureCitation id="43EF2A3CFF9F697F2AABFB74FE91FB1A" box="[151,311,1220,1244]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="5" pageNumber="50">Figures 13–16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697F2AABFABBFE5FFA56" pageId="5" pageNumber="50" type="diagnosis">
<paragraph id="DB6B36B9FF9F697F2AABFABBFE5FFA56" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AABFABBFEB7FAE2" bold="true" box="[151,273,1291,1316]" pageId="5" pageNumber="50">Diagnosis.</emphasis>
Rather large species within
<taxonomicName id="1CD44D3AFF9F697F286CFABCFD76FAE2" box="[592,720,1292,1316]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="E9A0EAABFF9F697F286CFABCFD76FAE2" box="[592,720,1292,1316]" italics="true" pageId="5" pageNumber="50">Molophilus</emphasis>
</taxonomicName>
. Body orange yellow, mostly shiny, patterned with pale yellow. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, setulose, pointed at tip. Inner gonostylus stouter and much longer, obtuse, spatuliform at apex. Wing length 3.8–5.4 mm.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9F697C2AFBFA2BFD47FEA2" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="description">
<paragraph id="DB6B36B9FF9F697F2AFBFA2BFD3BFA3B" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="50">
<emphasis id="E9A0EAABFF9F697F2AFBFA2BFE05FA72" bold="true" box="[199,419,1435,1460]" pageId="5" pageNumber="50">Description. Male.</emphasis>
Head. Antenna of moderate length, extending about to wing base. Basal segments yellowish brown. Flagellomeres infuscated, brown to dark brown, ovoid to long-ovoid, with longest verticils slightly exceeding length of their respective segments.
</paragraph>
<paragraph id="DB6B36B9FF9F697F2AFBF9B7FB2CF912" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="50">Thorax orange yellow, mostly shiny, patterned with pale yellow. Prescutum and scutum yellowish brown medially, with pale yellow area laterally around prescutal pit; scutal lobe pale yellow posterolaterally. Scutellum pale yellow. Postscutellum orange yellow, pale yellow anterolaterally. Pleuron orange yellow, mostly shiny, patterned with pale yellow on dorsopleural membrane, basalare and restrictedly elsewhere. Wing membrane tinged with yellow; venation generally as for genus. Halter pale yellow throughout. Legs yellow, with distal halves of femora, tips of tibiae and most of tarsi (except for yellow proximal two thirds of metatarsus) darkened.</paragraph>
<paragraph id="DB6B36B9FF9F697C2AFBF950FE74FEF9" blockId="5.[151,1437,1291,2036]" lastBlockId="6.[151,1436,151,356]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
Abdomen somewhat darker than thorax, yellowish brown dorsally. Male terminalia (
<figureCitation id="43EF2A3CFF9F697F2EB5F950FAADF93E" box="[1161,1291,1760,1784]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="5" pageNumber="50">Figs 13–16</figureCitation>
) orange yellow. Dorsal portion of gonocoxite moderately long, without dorsal and lateral lobe, tapered conically to obtuse apex in lateral view. Two blackened crests at dorsal inner margin at about two thirds length of gonocoxite; pale, glabrous, finger-like process at posteroventral edge of gonocoxite, directed mesally (
<figureCitation id="43EF2A3CFF9F697F2E6CF8FCFB75F8A2" box="[1104,1235,1868,1892]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="5" pageNumber="50">Figs 13, 14</figureCitation>
). Lateral excision comparatively deep and narrow. Ventral lobe of gonocoxite generally slender and long, its rounded tip not extending as far as apex of dorsal portion of gonocoxite. Both gonostyli darkly pigmented distally. Outer gonostylus tapered from broad base to slender pointed spine, slightly curved dorsally, provided with setulae and reaching to about half length of inner gonostylus. The latter long, exceeding ventral lobe by more than half its length, more or less parallel-sided, undulate, curved ventrally in distal part, obtuse at apex when viewed laterally, but expanded, spatuliform in ventral aspect. Aedeagus of moderate length and breadth, expanded before mid-length in lateral view and slightly bent ventrally, then gradually tapering to slightly upturned tip. Aedeagal plate large, generally oval in ventral/ventrocaudal aspect, with very long, slender, blackened median spine and with microscopically serrate margin posterolaterally.
</paragraph>
<paragraph id="DB6B36B9FF9C697C2AFBFEFCFD47FEA2" blockId="6.[151,1436,151,356]" box="[199,737,332,356]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AFBFEFCFEB8FEA2" bold="true" box="[199,286,332,356]" pageId="6" pageNumber="51">Female</emphasis>
resembling male in general appearance.
</paragraph>
</subSubSection>
<caption id="8FAB6631FF9C697C2AABFAD4FAE8FA10" httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="6" pageNumber="51" targetBox="[207,1434,440,1351]" targetPageId="6">
<paragraph id="DB6B36B9FF9C697C2AABFAD4FAE8FA10" blockId="6.[151,1437,1380,1495]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AABFAD4FEF3FABF" bold="true" box="[151,341,1380,1403]" pageId="6" pageNumber="51">FIGURES 13–20.</emphasis>
Male terminalia and male antennae. 13–16—
<taxonomicName id="1CD44D3AFF9C697C2914FAD5FC7FFABC" box="[808,985,1380,1402]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="hradileki" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2914FAD5FC7FFABC" box="[808,985,1380,1402]" italics="true" pageId="6" pageNumber="51">M. (M.) hradileki</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9C697C29DCFAD5FBBFFABC" bold="true" box="[992,1049,1381,1402]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="F29357D0FF9C697C29DCFAD5FBBFFABC" box="[992,1049,1381,1402]" pageId="6" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17–18—male antenna,
<taxonomicName id="1CD44D3AFF9C697C2FBCFA34FEBBFA7E" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2FBCFA34FEBBFA7E" italics="true" pageId="6" pageNumber="51">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9C697C2B1AFA12FEDFFA71" bold="true" box="[294,377,1442,1463]" pageId="6" pageNumber="51">nom. n.</emphasis>
(Spain: Fuengirola) (17) and
<taxonomicName id="1CD44D3AFF9C697C288EFA13FCDBFA7E" box="[690,893,1442,1464]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="propinquus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C288EFA13FCDBFA7E" box="[690,893,1442,1464]" italics="true" pageId="6" pageNumber="51">M. (M.) propinquus</emphasis>
</taxonomicName>
(Czech Republic: Lobodice) (18). 19–20––
<taxonomicName id="1CD44D3AFF9C697C2F70FA13FF4FFA10" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="ibericus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2F70FA13FF4FFA10" italics="true" pageId="6" pageNumber="51">M. (M.) ibericus</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9C697C2AD3FA71FEE6FA10" bold="true" box="[239,320,1473,1494]" pageId="6" pageNumber="51">nom. n.</emphasis>
(Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm.
</paragraph>
</caption>
<subSubSection id="93CE6532FF9C697C2AFBFA4CFD3CF99A" pageId="6" pageNumber="51" type="materials_examined">
<paragraph id="DB6B36B9FF9C697C2AFBFA4CFD3CF99A" blockId="6.[151,1436,1531,2025]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AFBFA4CFD8FF9D3" bold="true" box="[199,553,1532,1557]" pageId="6" pageNumber="51">
Material examined.
<typeStatus id="046F881BFF9C697C2B80FA4DFD8FF9D3" box="[444,553,1533,1557]" pageId="6" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
3:
<collectingCountry id="A3C37629FF9C697C2860FA4CFD44F9D2" box="[604,738,1532,1556]" name="Kazakhstan" pageId="6" pageNumber="51">Kazakhstan</collectingCountry>
, Altay Mts, Katon-Karagay N. Park, Yazovka, Belaya Bereľ valley, Ser-Sembai (
<quantity id="1C2C9B5CFF9C697C2B46F990FD80F9FE" box="[378,550,1568,1592]" metricMagnitude="0" metricUnit="m" metricValue="1.595" metricValueMax="1.61" metricValueMin="1.58" pageId="6" pageNumber="51" unit="m" value="1.595" valueMax="1.61" valueMin="1.58">1,580–1,610 m</quantity>
) (
<geoCoordinate id="BEE0507EFF9C697C2802F990FD3AF9FE" box="[574,668,1568,1592]" direction="north" orientation="latitude" pageId="6" pageNumber="51" precision="925" value="49.616665">49°37'N</geoCoordinate>
/
<geoCoordinate id="BEE0507EFF9C697C289AF990FCA7F9FE" box="[678,769,1568,1592]" direction="east" orientation="longitude" pageId="6" pageNumber="51" precision="925" value="86.46667">86°28'E</geoCoordinate>
),
<date id="AF6A1079FF9C697C2929F990FC2BF9FE" box="[789,909,1568,1592]" pageId="6" pageNumber="51" value="2007-06-28">28.vi.2007</date>
(Z. Hradílek leg.) (
<collectionCode id="BDC5AE7CFF9C697C2E56F990FB66F9FE" box="[1130,1216,1568,1592]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="6" pageNumber="51">SMOC</collectionCode>
).
<emphasis id="E9A0EAABFF9C697C2EEFF991FAF2F9FF" bold="true" box="[1235,1364,1569,1593]" pageId="6" pageNumber="51">
<typeStatus id="046F881BFF9C697C2EEFF991FAE9F9FF" box="[1235,1359,1569,1593]" pageId="6" pageNumber="51" type="paratype">Paratypes</typeStatus>
:
</emphasis>
19 3, 2 Ƥ, same data as for
<typeStatus id="046F881BFF9C697C2BB7F9F4FE4BF99A" box="[395,493,1604,1628]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
(
<collectionCode id="BDC5AE7CFF9C697C2BC0F9F4FD89F99A" box="[508,559,1604,1628]" pageId="6" pageNumber="51">JSO</collectionCode>
,
<collectionCode id="BDC5AE7CFF9C697C2806F9F4FD28F99A" box="[570,654,1604,1628]" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="6" pageNumber="51">SMOC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9C697C2AFBF9D9FCC5F90E" pageId="6" pageNumber="51" type="etymology">
<paragraph id="DB6B36B9FF9C697C2AFBF9D9FCC5F90E" blockId="6.[151,1436,1531,2025]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AFBF9D9FEE8F947" bold="true" box="[199,334,1641,1665]" pageId="6" pageNumber="51">Etymology.</emphasis>
The new species is named in honour of its collector, Dr. Zbynĕk Hradílek Ph.D. (Univerzita Palackého, Olomouc,
<collectingCountry id="A3C37629FF9C697C2BADF93CFDEFF962" box="[401,585,1676,1700]" name="Czech Republic" pageId="6" pageNumber="51">Czech Republic</collectingCountry>
), a bryologist at the Department of Botany, in thanks for highly interesting cranefly material from
<collectingCountry id="A3C37629FF9C697C2BA6F900FD86F90E" box="[410,544,1712,1736]" name="Kazakhstan" pageId="6" pageNumber="51">Kazakhstan</collectingCountry>
. A noun in genitive singular.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9C697C2AFBF964FE3BF802" pageId="6" pageNumber="51" type="discussion">
<paragraph id="DB6B36B9FF9C697C2AFBF964FE3BF802" blockId="6.[151,1436,1531,2025]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AFBF964FEEAF92B" bold="true" box="[199,332,1748,1773]" pageId="6" pageNumber="51">Discussion.</emphasis>
Based on a distally tapered gonocoxite, the overall shape of the gonostyli, and/or the structure of the aedeagal plate the new species is similar to
<taxonomicName id="1CD44D3AFF9C697C288DF949FCF9F8D6" box="[689,863,1784,1808]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="medius" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C288DF949FCF9F8D6" box="[689,863,1784,1808]" italics="true" pageId="6" pageNumber="51">M. (M.) medius</emphasis>
</taxonomicName>
de
<bibRefCitation id="BF454B48FF9C697C29B5F948FB8CF8D6" author="Meijere" box="[905,1066,1784,1808]" pageId="6" pageNumber="51" refString="Meijere, J. C. H. de (1918) Neue hollandische Dipteren. Tijdschrift voor Entomologie, 60, 128 - 141." type="journal article" year="1918">Meijere, 1918</bibRefCitation>
. It may be compared, and is distantly related to
<taxonomicName id="1CD44D3AFF9C697C2B6FF8AEFD55F8F2" authority="Lackschewitz, 1925" authorityName="Lackschewitz" authorityYear="1925" box="[339,755,1820,1845]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="maurus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2B6FF8AEFDAEF8F3" box="[339,520,1821,1845]" italics="true" pageId="6" pageNumber="51">M. (M.) maurus</emphasis>
Lackschewitz, 1925
</taxonomicName>
and its allies, such as
<taxonomicName id="1CD44D3AFF9C697C29C4F8AEFA3EF8F2" authority="Savchenko, 1978" authorityName="Savchenko" authorityYear="1978" box="[1016,1432,1820,1845]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="urodontus" subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C29C4F8AEFB6CF8F3" box="[1016,1226,1821,1845]" italics="true" pageId="6" pageNumber="51">M. (M.) urodontus</emphasis>
<bibRefCitation id="BF454B48FF9C697C2EEEF8ACFA3EF8F2" author="Savchenko" box="[1234,1432,1820,1844]" pageId="6" pageNumber="51" refString="Savchenko, E. N. (1978) New and little-known species of Palaearctic Limoniidae (Diptera). III. subfam. Eriopterinae, genus Molophilus Curt. Entomologicheskoe Obozrenie, 57, 399 - 415 (in Russian)." type="journal article" year="1978">Savchenko, 1978</bibRefCitation>
</taxonomicName>
, and subspecies
<emphasis id="E9A0EAABFF9C697C2B78F8F0FE65F89E" box="[324,451,1856,1880]" italics="true" pageId="6" pageNumber="51">brachyurus</emphasis>
<bibRefCitation id="BF454B48FF9C697C2BF6F8F0FD2CF89E" author="Savchenko" box="[458,650,1856,1880]" pageId="6" pageNumber="51" refString="Savchenko, E. N. (1978) New and little-known species of Palaearctic Limoniidae (Diptera). III. subfam. Eriopterinae, genus Molophilus Curt. Entomologicheskoe Obozrenie, 57, 399 - 415 (in Russian)." type="journal article" year="1978">Savchenko, 1978</bibRefCitation>
and
<emphasis id="E9A0EAABFF9C697C2883F8F0FC91F89E" box="[703,823,1856,1880]" italics="true" pageId="6" pageNumber="51">cephaliger</emphasis>
<bibRefCitation id="BF454B48FF9C697C2901F8F0FC5BF89E" author="Savchenko" box="[829,1021,1856,1880]" pageId="6" pageNumber="51" refString="Savchenko, E. N. (1978) New and little-known species of Palaearctic Limoniidae (Diptera). III. subfam. Eriopterinae, genus Molophilus Curt. Entomologicheskoe Obozrenie, 57, 399 - 415 (in Russian)." type="journal article" year="1978">Savchenko, 1978</bibRefCitation>
(both apparently represent valid species), all differing externally from
<taxonomicName id="1CD44D3AFF9C697C2822F8D6FD45F8BB" box="[542,739,1893,1917]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="hradileki" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2822F8D6FD45F8BB" box="[542,739,1893,1917]" italics="true" pageId="6" pageNumber="51">M. (M.) hradileki</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9C697C28D7F8D5FC8DF8BB" bold="true" box="[747,811,1893,1917]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="F29357D0FF9C697C28D7F8D5FC8DF8BB" box="[747,811,1893,1917]" pageId="6" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by darker body colouration.
<taxonomicName id="1CD44D3AFF9C697C2E49F8D6FA9FF8BB" box="[1141,1337,1893,1917]" class="Insecta" family="Limoniidae" genus="Molophilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="hradileki" status="sp. nov." subGenus="Molophilus">
<emphasis id="E9A0EAABFF9C697C2E49F8D6FA9FF8BB" box="[1141,1337,1893,1917]" italics="true" pageId="6" pageNumber="51">M. (M.) hradileki</emphasis>
</taxonomicName>
<emphasis id="E9A0EAABFF9C697C2F7DF8D5FA24F8BB" bold="true" box="[1345,1410,1893,1917]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="F29357D0FF9C697C2F7DF8D5FA24F8BB" box="[1345,1410,1893,1917]" pageId="6" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is peculiar in having two blackened crests at the dorsal inner margin of the gonocoxite (
<figureCitation id="43EF2A3CFF9C697C2E6AF838FB71F866" box="[1110,1239,1928,1952]" captionStart="FIGURES 13 – 20" captionStartId="6.[151,264,1380,1403]" captionTargetBox="[207,1434,440,1351]" captionTargetId="figure@6.[151,1436,389,1367]" captionTargetPageId="6" captionText="FIGURES 13 – 20. Male terminalia and male antennae. 13 – 16 — M. (M.) hradileki sp. n. (paratype), general view, lateral (13), distal part of gonocoxite, dorsal (14), aedeagal complex, lateral (15), and aedeagal plate, ventral (16). 17 – 18 — male antenna, M. (M.) ibericus nom. n. (Spain: Fuengirola) (17) and M. (M.) propinquus (Czech Republic: Lobodice) (18). 19 – 20 –– M. (M.) ibericus nom. n. (Spain: Fuengirola), general view, lateral (19) and aedeagal complex, lateral (20). Scale bars 0.5 mm." httpUri="https://zenodo.org/record/203288/files/figure.png" pageId="6" pageNumber="51">Figs 13, 14</figureCitation>
), a character otherwise unknown to me.
</paragraph>
</subSubSection>
<subSubSection id="93CE6532FF9C697C2AFBF860FE57F82E" box="[199,497,2000,2025]" pageId="6" pageNumber="51" type="distribution">
<paragraph id="DB6B36B9FF9C697C2AFBF860FE57F82E" blockId="6.[151,1436,1531,2025]" box="[199,497,2000,2025]" pageId="6" pageNumber="51">
<emphasis id="E9A0EAABFF9C697C2AFBF860FEC6F82F" bold="true" box="[199,352,2000,2025]" pageId="6" pageNumber="51">Distribution.</emphasis>
<collectingCountry id="A3C37629FF9C697C2B5BF860FE4BF82E" box="[359,493,2000,2024]" name="Kazakhstan" pageId="6" pageNumber="51">Kazakhstan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

138
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<mods:title>New records of Braconinae (Hymenoptera, Braconidae) from South Korea</mods:title>
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<mods:namePart>Samartsev, Konstantin</mods:namePart>
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<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034, Russia</mods:affiliation>
<mods:nameIdentifier type="email">ksamartsev@gmail.com</mods:nameIdentifier>
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<mods:namePart>Ku, Deok-Seo</mods:namePart>
<mods:affiliation>The Science Museum of Natural Enemies, Geochang, 50147, South Korea</mods:affiliation>
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<treatment ID-GBIF-Taxon="183203181" LSID="urn:lsid:plazi:treatment:537DC83F88D6526F9B2BD13C8473A106" httpUri="http://treatment.plazi.org/id/537DC83F88D6526F9B2BD13C8473A106" lastPageNumber="21" pageId="0" pageNumber="21">
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<taxonomicName ID-CoL="N4FC" LSID="537DC83F-88D6-526F-9B2B-D13C8473A106" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Insecta" family="Braconidae" genus="Bracon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bracon (Rostrobracon) urinator" order="Hymenoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="urinator" subGenus="Rostrobracon">Bracon (Rostrobracon) urinator (Fabricius, 1798)</taxonomicName>
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<subSubSection pageId="0" pageNumber="21" type="description">
<paragraph pageId="0" pageNumber="21">
<figureCitation captionStart="Figures A5–A8" captionStartId="F12" captionText="Figures A 5 - A 8. Collecting localities of the material on the species new to South Korea. Point numbers correspond with numbers in brackets in text." figureDoi="10.3897/jhr.83.63353.figuresa5-a8" httpUri="https://binary.pensoft.net/fig/559776" pageId="0" pageNumber="21">Fig. A8</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="21" type="materials_examined">
<paragraph pageId="0" pageNumber="21">Material.</paragraph>
<paragraph pageId="0" pageNumber="21">
<materialsCitation collectingDate="1995-08-10" collectorName="S. H. Lee" country="South Korea" county="Jeju-si" location="Hanlla Mountain" municipality="Odeung-dong" specimenCount="1" specimenCount-female="1" stateProvince="Jeju-do">
<collectingCountry name="South Korea">South Korea</collectingCountry>
-
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<collectingRegion country="South Korea" name="Jeju">Jeju-do</collectingRegion>
</emphasis>
•
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;
<collectingCounty>Jeju-si</collectingCounty>
, [89]
<collectingMunicipality>Odeung-dong</collectingMunicipality>
,
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;
<collectingDate value="1995-08-10">10 Aug. 1995</collectingDate>
;
<collectorName>S.H. Lee</collectorName>
leg.; NIBR 13
</materialsCitation>
.
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</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="distribution">
<paragraph pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="21">
Afghanistan. Caucasus. Central Asia. China: Liaoning, Shaanxi, Shandong, Zhejiang. Cyprus. Europe: Eastern, Northern, Southern, and Western Europe. Iran. Israel. Kazakhstan. Mongolia. North Africa: Algeria, Canary Islands, Egypt, Tunisia. Russia: Eastern Siberia: Buryatia Republic, Zabaikalskiy Territory; European part; Far East: Primorskiy Territory; Ural (
<bibRefCitation author="Kostromina, TS" journalOrPublisher="Proceedings of the Russian Entomological Society" pageId="0" pageNumber="21" pagination="47 - 50" refId="B4" refString="Kostromina, TS, 2010. To the knowledge of the cyclostome braconid wasps (Hymenoptera: Braconidae) of South Urals. Proceedings of the Russian Entomological Society 81 (2): 47 - 50" title="To the knowledge of the cyclostome braconid wasps (Hymenoptera: Braconidae) of South Urals." volume="81" year="2010">Kostromina 2010</bibRefCitation>
). Saudi Arabia. South Korea (new record). Syria. Turkey.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="remarks">
<paragraph pageId="0" pageNumber="21">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Tobias" authorityYear="1957" class="Insecta" family="Braconidae" genus="Rostrobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rostrobracon" order="Hymenoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Rostrobracon</emphasis>
</taxonomicName>
Tobias, 1957 is considered here a valid subgenus, because its synonymisation with
<taxonomicName authorityName="Tobias" authorityYear="1957" class="Insecta" family="Braconidae" genus="Cyanopterobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyanopterobracon" order="Hymenoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Cyanopterobracon</emphasis>
</taxonomicName>
Tobias, 1957 was not justified (
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2012.21" author="Papp, J" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="21" pagination="1 - 154" refId="B15" refString="Papp, J, 2012. A revision of the Bracon Fabricius species in Wesmael's collection deposited in Brussels (Hymenoptera, Braconidae, Braconinae). European Journal of Taxonomy 21: 1 - 154, DOI: https://doi.org/10.5852/ejt.2012.21" title="A revision of the Bracon Fabricius species in Wesmael's collection deposited in Brussels (Hymenoptera, Braconidae, Braconinae)." url="https://doi.org/10.5852/ejt.2012.21" volume="21" year="2012">Papp 2012</bibRefCitation>
). On the contrary, the latter subgenus differs by the less elongate eyes, about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide in lateral view (in
<taxonomicName authorityName="Tobias" authorityYear="1957" class="Insecta" family="Braconidae" genus="Rostrobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rostrobracon" order="Hymenoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Rostrobracon</emphasis>
</taxonomicName>
, ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
), the long malar space, ca. 0.5
<normalizedToken originalValue="×">x</normalizedToken>
longitudinal diameter of eye in anterior view (ca. 0.25
<normalizedToken originalValue="×">x</normalizedToken>
), the elongate, 1.3-1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as high, mesosoma (robust, 1.1-1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as high) with evenly convex median lobe of mesoscutum (the median lobe dorsally flattened in anterior part), the deep second metasomal suture (mostly shallow in
<taxonomicName authorityName="Tobias" authorityYear="1957" class="Insecta" family="Braconidae" genus="Rostrobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rostrobracon" order="Hymenoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Rostrobracon</emphasis>
</taxonomicName>
), and the shorter ovipositor sheath, ca. as long as hind tibia (ca. 2
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as long as hind tibia in
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<emphasis italics="true" pageId="0" pageNumber="21">Rostrobracon</emphasis>
</taxonomicName>
).
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