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15 isolated teeth; Catahoula Formation;
15 isolated teeth;
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<mods:title id="B957192687BE7307931C3F1E813A898B">A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)</mods:title>
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<taxonomicName id="4C7DE2D4FF89FF82FDF996FAFCFA282C" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[599,891,1001,1028]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="nitidistylus" status="sp. nov.">
<emphasis id="B9094545FF89FF82FDF996FAFCFA282C" bold="true" box="[599,891,1001,1028]" italics="true" pageId="81" pageNumber="82">Sclerocoelus nitidistylus</emphasis>
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<figureCitation id="134685D2FF89FF82FD089123FC872862" box="[678,774,1072,1098]" captionStart="Fig" captionStartId="128.[189,232,1929,1955]" captionTargetBox="[189,1399,265,1874]" captionTargetId="figure-51@128.[189,1399,265,1874]" captionTargetPageId="128" captionText="Fig. 1. Sclerocoelus Marshall, 1995, live specimens. A. Sclerocoelus nitidistylus sp. nov. near an Eciton bivouac in Alajuela Province, Costa Rica. B. Probably Sclerocoelus copiosus sp. nov. from Bellavista Cloud Forest Reserve, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.14980856" httpUri="https://zenodo.org/record/14980856/files/figure.png" pageId="81" pageNumber="82">Figs 1A</figureCitation>
,
<figureCitation id="134685D2FF89FF82FCBD9123FCB62862" box="[787,823,1072,1098]" captionStart="Fig" captionStartId="135.[189,232,1850,1876]" captionTargetBox="[189,1399,277,1815]" captionTargetId="figure-86@135.[189,1399,265,1815]" captionTargetPageId="135" captionText="Fig. 8. Sclerocoelus Marshall, 1995, left wings, ventral view. A. Sclerocoelus nebulosus sp. nov. B. Sclerocoelus nitidistylus sp. nov. C. Sclerocoelus ocellatus sp. nov. D. Sclerocoelus paranebulosus sp. nov. E. Sclerocoelus pararegularis sp. nov. F. Sclerocoelus parasordipes sp. nov. G. Sclerocoelus plumiseta (Duda, 1925) H. Sclerocoelus punensis sp. nov. I. Sclerocoelus puyensis sp. nov.J. Sclerocoelus rectangularis (Malloch, 1914). Scale bars: AJ=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14980872" httpUri="https://zenodo.org/record/14980872/files/figure.png" pageId="81" pageNumber="82">8B</figureCitation>
,
<figureCitation id="134685D2FF89FF82FCEA9123FC0F2862" box="[836,910,1072,1098]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="223.[189,232,1860,1886]" captionStartId-1="224.[189,232,1858,1884]" captionStartId-2="225.[189,232,1927,1953]" captionTargetBox-0="[189,1399,265,1788]" captionTargetBox-1="[197,1395,267,1802]" captionTargetBox-2="[215,1379,266,1883]" captionTargetId-0="figure-66@223.[189,1399,265,1830]" captionTargetId-1="figure-88@224.[189,1399,265,1830]" captionTargetId-2="figure-49@225.[210,1381,251,1904]" captionTargetPageId-0="223" captionTargetPageId-1="224" captionTargetPageId-2="225" captionText-0="Fig. 96. Sclerocoelus nitidistylus sp. nov. A. Male paratype habitus, lateral view (debu00125692). B. Male paratype terminalia, posterior view. C. Male paratype terminalia, lateral view. Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." captionText-1="Fig. 97. Sclerocoelus nitidistylus sp. nov., paratype. A. Aedeagus and associated structures, lateral view. B. Male S5 and genital pouch, ventral view. C. Subcercal complex, posterior view. Abbreviations: bp =basiphallus; dp =distiphallus; epi= epiphallus; hyp= hypandrium; pg =postgonite; ph =phallapodeme; sc (subcostal); se = subepandrial sclerite; ss=surstylus. Scale bars: A= 0.13 mm; B=0.25 mm; C= 0.10 mm." captionText-2="Fig. 98. Sclerocoelus nitidistylus sp. nov., paratype. A. Female terminalia, dorsal view. B. Female terminalia, ventral view. C. Female terminalia, lateral view. D. Spermatheca. Scale bars:AC =0.25 mm; D =0.03 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14981080" figureDoi-1="http://doi.org/10.5281/zenodo.14981082" figureDoi-2="http://doi.org/10.5281/zenodo.14981084" httpUri-0="https://zenodo.org/record/14981080/files/figure.png" httpUri-1="https://zenodo.org/record/14981082/files/figure.png" httpUri-2="https://zenodo.org/record/14981084/files/figure.png" pageId="81" pageNumber="82">9698</figureCitation>
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<emphasis id="B9094545FF89FF82FF139161FEC528A4" bold="true" box="[189,324,1138,1164]" pageId="81" pageNumber="82">Etymology</emphasis>
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This name refers to the shining outer surface of the male surstylus (from the Latin
<emphasis id="B9094545FF89FF82FB0691B3FB792892" box="[1192,1272,1184,1210]" italics="true" pageId="81" pageNumber="82">nitidus</emphasis>
, meaning shiny, polished).
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<emphasis id="B9094545FF89FF82FF139016FE2A2937" bold="true" box="[189,427,1285,1311]" pageId="81" pageNumber="82">Material examined</emphasis>
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<emphasis id="B9094545FF89FF82FF449027FEDD2966" bold="true" box="[234,348,1332,1358]" pageId="81" pageNumber="82">
<typeStatus id="54C627F5FF89FF82FF449027FEDD2966" box="[234,348,1332,1358]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
</emphasis>
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<paragraph id="8BC29957FF89FF82FF139044FD8029BD" blockId="81.[189,1399,1332,1429]" pageId="81" pageNumber="82">
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</date>
;
<collectorName id="2688FC81FF89FF82FF139068FEE029BC" box="[189,353,1402,1429]" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<collectionCode id="ED6C0192FF89FF82FE0D9068FE7C29BD" box="[419,509,1403,1429]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC29957FF89FF82FF4490AFFEE629FE" blockId="81.[189,1399,1468,2024]" box="[234,359,1468,1494]" pageId="81" pageNumber="82">
<emphasis id="B9094545FF89FF82FF4490AFFEE629FE" bold="true" box="[234,359,1468,1494]" pageId="81" pageNumber="82">
<typeStatus id="54C627F5FF89FF82FF4490AFFEE629FE" box="[234,359,1468,1494]" pageId="81" pageNumber="82" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC29957FF89FF81FF1390CCFDFE2803" blockId="81.[189,1399,1468,2024]" lastBlockId="82.[189,1399,265,1067]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<materialsCitation id="3B15930AFF89FF82FF1390CCFE8C2A69" collectedFrom="from refuse pile under Eciton burchellii bivouac" collectingDate="2023-04-21" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="1800" latitude="10.254194" location="Albergue Pozo Verde" longLatPrecision="1" longitude="-84.37178" municipality="Quesada" pageId="81" pageNumber="82" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<collectingCountry id="F36AD9C7FF89FF82FF1390CCFEEB29D1" box="[189,362,1503,1529]" name="Costa Rica" pageId="81" pageNumber="82">COSTA RICA</collectingCountry>
<emphasis id="B9094545FF89FF82FE3C90CCFE7D29D1" bold="true" box="[402,508,1503,1529]" pageId="81" pageNumber="82">
<collectingRegion id="49B957B5FF89FF82FE3C90CCFE7D29D1" box="[402,508,1503,1529]" country="Costa Rica" name="Alajuela" pageId="81" pageNumber="82">Alajuela</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF89FF82FDB190CCFDE829D2" box="[543,617,1503,1530]" count="2" pageId="81" pageNumber="82" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF89FF82FDD590CCFD6229D1" box="[635,739,1503,1529]" pageId="81" pageNumber="82">Quesada</collectingMunicipality>
,
<location id="8EA2CF8CFF89FF82FD5C90CCFC7B29D1" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FD5C90CCFC7B29D1" box="[754,1018,1503,1530]" country="Costa Rica" latitude="10.254194" longLatPrecision="1" longitude="-84.37178" municipality="Quesada" name="Albergue Pozo Verde" pageId="81" pageNumber="82" stateProvince="Alajuela">Albergue Pozo Verde</location>
;
<elevation id="00507E64FF89FF82FBA590CCFB2F29D1" box="[1035,1198,1503,1530]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="81" pageNumber="82" unit="m" value="1800.0">
<quantity id="4C8534B2FF89FF82FBA590CCFBE829D2" box="[1035,1129,1503,1530]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="81" pageNumber="82" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE49FF90FF89FF82FB6F90CDFAF329D2" box="[1217,1394,1502,1530]" degrees="10" direction="north" minutes="15" orientation="latitude" pageId="81" pageNumber="82" precision="1" seconds="15.1" value="10.254194">10°1515.1″ N</geoCoordinate>
,
<geoCoordinate id="EE49FF90FF89FF82FF139312FEF32A35" box="[189,370,1537,1565]" degrees="84" direction="west" minutes="22" orientation="longitude" pageId="81" pageNumber="82" precision="1" seconds="18.4" value="-84.37178">84°2218.4″ W</geoCoordinate>
;
<date id="FFC3BF97FF89FF82FE2F9310FDA52A35" box="[385,548,1539,1565]" pageId="81" pageNumber="82" value="2023-04-21">
<collectingDate id="EF87467FFF89FF82FE2F9310FDA52A35" box="[385,548,1539,1565]" pageId="81" pageNumber="82" value="2023-04-21">21 Apr. 2023</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FD9D9310FD5B2A35" box="[563,730,1539,1565]" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<collectedFrom id="3861DAAAFF89FF82FC8A9310FAF32A35" box="[804,1394,1539,1565]" pageId="81" pageNumber="82">
from refuse pile under
<taxonomicName id="4C7DE2D4FF89FF82FB909310FA862A35" authorityName="Westwood" authorityYear="1842" box="[1086,1287,1539,1565]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="burchellii">
<emphasis id="B9094545FF89FF82FB909310FA862A35" box="[1086,1287,1539,1565]" italics="true" pageId="81" pageNumber="82">Eciton burchellii</emphasis>
</taxonomicName>
bivouac
</collectedFrom>
;
<collectionCode id="ED6C0192FF89FF82FF139334FE8C2A69" box="[189,269,1575,1601]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FE829335FCB02A68" box="[300,817,1574,1601]" collectingDate="2023-04-21" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1800" latitude="10.254194" location="Albergue Pozo Verde" longLatPrecision="1" longitude="-84.37178" municipality="Quesada" pageId="81" pageNumber="82" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FE829335FEF52A69" box="[300,372,1574,1601]" count="2" pageId="81" pageNumber="82" type="male">2 ♂♂</specimenCount>
; same data as for preceding;
<collectionCode id="ED6C0192FF89FF82FD759335FCB02A68" box="[731,817,1574,1600]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FCFF9335FE282A4B" collectingDate="2023-04-21" collectionCode="MNCR" collectorName="W. Porras" country="Costa Rica" elevation="1800" latitude="10.254194" location="Albergue Pozo Verde" longLatPrecision="1" longitude="-84.37178" municipality="Quesada" pageId="81" pageNumber="82" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FCFF9335FC192A69" box="[849,920,1574,1601]" count="2" pageId="81" pageNumber="82" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF89FF82FC099335FC6F2A69" box="[935,1006,1574,1601]" count="3" pageId="81" pageNumber="82" type="female">3 ♀♀</specimenCount>
; same data as for preceding;
<collectorName id="2688FC81FF89FF82FAF89334FE892A4C" pageId="81" pageNumber="82">W. Porras</collectorName>
leg.;
<collectionCode id="ED6C0192FF89FF82FEE1935AFE282A4B" box="[335,425,1609,1635]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF89FF82FE60935AFD9C2AAF" collectingDate="1985-04-10" collectionCode="CNCI" collectorName="L. Masner" country="Costa Rica" elevation="500" location="Braulio Carrillo National Park" pageId="81" pageNumber="82" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
<emphasis id="B9094545FF89FF82FE60935AFDB42A4B" bold="true" box="[462,565,1609,1635]" pageId="81" pageNumber="82">
<collectingRegion id="49B957B5FF89FF82FE60935AFDB42A4B" box="[462,565,1609,1635]" country="Costa Rica" name="Cartago" pageId="81" pageNumber="82">Cartago</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF89FF82FDFD935AFD052A4C" box="[595,644,1609,1636]" count="1" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
;
<location id="8EA2CF8CFF89FF82FD3D935AFB832A4B" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FD3D935AFB832A4B" box="[659,1026,1609,1635]" country="Costa Rica" name="Braulio Carrillo National Park" pageId="81" pageNumber="82" stateProvince="Cartago">Braulio Carrillo National Park</location>
;
<elevation id="00507E64FF89FF82FBBE935AFB1E2A4B" box="[1040,1183,1609,1636]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="81" pageNumber="82" unit="m" value="500.0">
<quantity id="4C8534B2FF89FF82FBBE935AFBDD2A4C" box="[1040,1116,1609,1636]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="81" pageNumber="82" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FB00935AFACE2A4C" box="[1198,1359,1609,1636]" pageId="81" pageNumber="82" value="1985-04-10">
<collectingDate id="EF87467FFF89FF82FB00935AFACE2A4C" box="[1198,1359,1609,1636]" pageId="81" pageNumber="82" value="1985-04-10">10 Apr. 1985</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FAF39359FE942AAF" pageId="81" pageNumber="82">L. Masner</collectorName>
leg.;
<location id="8EA2CF8CFF89FF82FEF9937EFE4A2AAF" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FEF9937EFE4A2AAF" box="[343,459,1645,1671]" country="Costa Rica" name="rainforest" pageId="81" pageNumber="82" stateProvince="Cartago">rainforest</location>
;
<collectionCode id="ED6C0192FF89FF82FE79937EFD9C2AAF" box="[471,541,1645,1671]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="81" pageNumber="82">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FD99937EFBD92A82" collectingDate="1985-04-11" collectionCode="CNCI" collectorName="H. Goulet &amp; L. Masner" country="Costa Rica" elevation="1450" location="Braulio Carrillo National Park" pageId="81" pageNumber="82" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FD99937EFDE52AAF" box="[567,612,1645,1671]" count="1" pageId="81" pageNumber="82" type="female">1 ♀</specimenCount>
;
<location id="8EA2CF8CFF89FF82FDDD937EFC5A2AAF" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FDDD937EFC5A2AAF" box="[627,987,1645,1671]" country="Costa Rica" name="Braulio Carrillo National Park" pageId="81" pageNumber="82" stateProvince="Cartago">Braulio Carrillo National Park</location>
;
<elevation id="00507E64FF89FF82FC49937FFB482AAF" box="[999,1225,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="81" pageNumber="82" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<quantity id="4C8534B2FF89FF82FC49937FFB082AAF" box="[999,1161,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="81" pageNumber="82" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">14001500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FB79937EFAF32AAF" box="[1239,1394,1644,1671]" pageId="81" pageNumber="82" value="1985-04-11">
<collectingDate id="EF87467FFF89FF82FB79937EFAF32AAF" box="[1239,1394,1644,1671]" pageId="81" pageNumber="82" value="1985-04-11">11 Apr. 1985</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FF139382FEB22A82" box="[189,307,1680,1707]" pageId="81" pageNumber="82">H. Goulet</collectorName>
and
<collectorName id="2688FC81FF89FF82FEC19382FE682A83" box="[367,489,1681,1707]" pageId="81" pageNumber="82">L. Masner</collectorName>
leg.;
<location id="8EA2CF8CFF89FF82FD819383FC982A82" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FD819383FC982A82" box="[559,793,1680,1706]" country="Costa Rica" name="cool moist riverbed" pageId="81" pageNumber="82" stateProvince="Cartago">cool moist riverbed</location>
,
<location id="8EA2CF8CFF89FF82FC899383FB802A83" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FC899383FB802A83" box="[807,1025,1680,1707]" country="Costa Rica" name="Selva Premontana" pageId="81" pageNumber="82" stateProvince="Cartago">Selva Premontana</location>
;
<collectionCode id="ED6C0192FF89FF82FBA09383FBD92A82" box="[1038,1112,1680,1706]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="81" pageNumber="82">CNCI</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF89FF82FBD29383FDE12AD9" collectingDate="1996-02-12" collectingDateMax="1996-02-15" collectingDateMin="1996-02-12" collectingMethod="carrion traps" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="1050" location="Cacao Field Station" municipality="Guanacaste National Park" pageId="81" pageNumber="82" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B9094545FF89FF82FBD29383FA8E2A82" bold="true" box="[1148,1295,1680,1706]" pageId="81" pageNumber="82">
<collectingRegion id="49B957B5FF89FF82FBD29383FA8E2A82" box="[1148,1295,1680,1706]" country="Costa Rica" name="Guanacaste" pageId="81" pageNumber="82">Guanacaste</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF89FF82FA859383FAF02A83" box="[1323,1393,1680,1707]" count="3" pageId="81" pageNumber="82" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF89FF82FF1393A0FE852AE6" box="[189,260,1715,1742]" count="4" pageId="81" pageNumber="82" type="female">4 ♀♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF89FF82FEB993A0FDD22AE5" box="[279,595,1715,1741]" pageId="81" pageNumber="82">Guanacaste National Park</collectingMunicipality>
,
<location id="8EA2CF8CFF89FF82FDCC93A0FCD72AE5" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FDCC93A0FCD72AE5" box="[610,854,1715,1741]" country="Costa Rica" municipality="Guanacaste National Park" name="Cacao Field Station" pageId="81" pageNumber="82" stateProvince="Guanacaste">Cacao Field Station</location>
;
<elevation id="00507E64FF89FF82FCC893A0FBCE2AE5" box="[870,1103,1715,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="81" pageNumber="82" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">
<quantity id="4C8534B2FF89FF82FCC893A0FB8A2AE6" box="[870,1035,1715,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="81" pageNumber="82" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">10001100 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FBCE93A0FAB32AE5" box="[1120,1330,1715,1742]" pageId="81" pageNumber="82" value="1996-02-12" valueMax="1996-02-15" valueMin="1996-02-12">
<collectingDate id="EF87467FFF89FF82FBCE93A0FAB32AE5" box="[1120,1330,1715,1742]" pageId="81" pageNumber="82" value="1996-02-12" valueMax="1996-02-15" valueMin="1996-02-12">1215 Feb. 1996</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FAEC93A0FEA42AD9" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<collectingMethod id="523CE140FF89FF82FEC493C4FD822AD9" box="[362,515,1751,1777]" pageId="81" pageNumber="82">carrion traps</collectingMethod>
;
<collectionCode id="ED6C0192FF89FF82FDBE93C4FDE12AD9" box="[528,608,1751,1777]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FDD293C4FE922B3C" collectingDate="1996-02-13" collectingDateMax="1996-02-15" collectingDateMin="1996-02-13" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="700" location="Cacao Field Station" municipality="Guanacaste National Park" pageId="81" pageNumber="82" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FDD293C4FD2B2AD9" box="[636,682,1751,1777]" count="1" pageId="81" pageNumber="82" type="female">1 ♀</specimenCount>
; same data as for preceding;
<elevation id="00507E64FF89FF82FBA493C4FB162AD9" box="[1034,1175,1751,1777]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="81" pageNumber="82" unit="m" value="700.0">
<quantity id="4C8534B2FF89FF82FBA493C4FBD42AD9" box="[1034,1109,1751,1777]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="81" pageNumber="82" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FB0B93C5FAF32AD8" box="[1189,1394,1750,1777]" pageId="81" pageNumber="82" value="1996-02-13" valueMax="1996-02-15" valueMin="1996-02-13">
<collectingDate id="EF87467FFF89FF82FB0B93C5FAF32AD8" box="[1189,1394,1750,1777]" pageId="81" pageNumber="82" value="1996-02-13" valueMax="1996-02-15" valueMin="1996-02-13">1315 Feb. 1996</collectingDate>
</date>
;
<collectionCode id="ED6C0192FF89FF82FF1393E9FE922B3C" box="[189,275,1786,1812]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FE9D93E9FE7A2B1F" collectingDate="1996-02-15" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1150" location="Cacao Field Station" municipality="Guanacaste National Park" pageId="81" pageNumber="82" specimenCount="2" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FE9D93E9FEFA2B3C" box="[307,379,1786,1812]" count="2" pageId="81" pageNumber="82" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF89FF82FE2593E9FD472B3C" box="[395,710,1786,1812]" pageId="81" pageNumber="82">Guanacaste National Park</collectingMunicipality>
,
<location id="8EA2CF8CFF89FF82FD7A93E9FC472B3C" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FD7A93E9FC472B3C" box="[724,966,1786,1812]" country="Costa Rica" municipality="Guanacaste National Park" name="Cacao Field Station" pageId="81" pageNumber="82" stateProvince="Guanacaste">Cacao Field Station</location>
;
<elevation id="00507E64FF89FF82FC7B93E9FB3C2B3C" box="[981,1213,1786,1812]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="81" pageNumber="82" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">
<quantity id="4C8534B2FF89FF82FC7B93E9FBFB2B3C" box="[981,1146,1786,1812]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="81" pageNumber="82" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">11001200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FB6093E9FAF32B3C" box="[1230,1394,1786,1813]" pageId="81" pageNumber="82" value="1996-02-15">
<collectingDate id="EF87467FFF89FF82FB6093E9FAF32B3C" box="[1230,1394,1786,1813]" pageId="81" pageNumber="82" value="1996-02-15">15 Feb. 1996</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FF13920EFEE02B1F" box="[189,353,1821,1847]" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<collectionCode id="ED6C0192FF89FF82FE0B920EFE7A2B1F" box="[421,507,1821,1847]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FDB8920EFD632B73" collectingDate="1996-02-11" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="300" location="Guanacaste National Park" municipality="Guanacaste National Park" pageId="81" pageNumber="82" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FDB8920EFDC72B10" box="[534,582,1821,1848]" count="1" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
;
<location id="8EA2CF8CFF89FF82FDFD920EFC082B1F" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FDFD920EFC082B1F" box="[595,905,1821,1847]" country="Costa Rica" municipality="Guanacaste National Park" name="Guanacaste National Park" pageId="81" pageNumber="82" stateProvince="Guanacaste">Guanacaste National Park</location>
,
<location id="8EA2CF8CFF89FF82FC3B920EFBF92B1F" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FC3B920EFBF92B1F" box="[917,1144,1821,1848]" country="Costa Rica" municipality="Guanacaste National Park" name="Santa Rosa Station" pageId="81" pageNumber="82" stateProvince="Guanacaste">Santa Rosa Station</location>
;
<elevation id="00507E64FF89FF82FB2B920EFA912B1F" box="[1157,1296,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="81" pageNumber="82" unit="m" value="300.0">
<quantity id="4C8534B2FF89FF82FB2B920EFB512B10" box="[1157,1232,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="81" pageNumber="82" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FAB1920EFF7A2B72" pageId="81" pageNumber="82" value="1996-02-11">
<collectingDate id="EF87467FFF89FF82FAB1920EFF7A2B72" pageId="81" pageNumber="82" value="1996-02-11">11 Feb. 1996</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FEA89252FE282B72" box="[262,425,1856,1883]" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<location id="8EA2CF8CFF89FF82FE449253FDFC2B72" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FE449253FDFC2B72" box="[490,637,1856,1882]" country="Costa Rica" municipality="Guanacaste National Park" name="dry riverbed" pageId="81" pageNumber="82" stateProvince="Guanacaste">dry riverbed</location>
;
<collectionCode id="ED6C0192FF89FF82FD269252FD632B73" box="[648,738,1857,1883]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF89FF82FCAF9253FD712B56" collectingDate="2002-02-21" collectionCode="DEBU" country="Costa Rica" elevation="2000" latitude="10.183333" location="Heredia" longLatPrecision="1298" longitude="-84.11667" pageId="81" pageNumber="82" specimenCount="4" specimenCount-female="4" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="B9094545FF89FF82FCAF9253FCE42B72" bold="true" box="[769,869,1856,1882]" pageId="81" pageNumber="82">
<collectingRegion id="49B957B5FF89FF82FCAF9253FCE42B72" box="[769,869,1856,1882]" country="Costa Rica" name="Heredia" pageId="81" pageNumber="82">Heredia</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF89FF82FCD39252FC3E2B73" box="[893,959,1857,1883]" count="4" pageId="81" pageNumber="82" type="female">4 ♀♀</specimenCount>
;
<locationDeviation id="9A708075FF89FF82FC629253FB7C2B72" box="[972,1277,1856,1883]" pageId="81" pageNumber="82">
<quantity id="4C8534B2FF89FF82FC629253FB862B72" box="[972,1031,1856,1882]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="81" pageNumber="82" unit="km" value="6.0">6 km</quantity>
ENE of Vera Blanca
</locationDeviation>
;
<geoCoordinate id="EE49FF90FF89FF82FAA6922CFAF32B73" box="[1288,1394,1855,1883]" degrees="10" direction="north" minutes="11" orientation="latitude" pageId="81" pageNumber="82" precision="925" value="10.183333">10°11 N</geoCoordinate>
,
<geoCoordinate id="EE49FF90FF89FF82FF139271FEB02B56" box="[189,305,1890,1918]" degrees="84" direction="west" minutes="07" orientation="longitude" pageId="81" pageNumber="82" precision="925" value="-84.11667">84°07 W</geoCoordinate>
;
<elevation id="00507E64FF89FF82FEEE9277FE5F2B56" box="[320,478,1892,1918]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="81" pageNumber="82" unit="m" value="2000.0">
<quantity id="4C8534B2FF89FF82FEEE9277FE1D2B56" box="[320,412,1892,1918]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="81" pageNumber="82" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FE409277FD102B56" box="[494,657,1892,1918]" pageId="81" pageNumber="82" value="2002-02-21">
<collectingDate id="EF87467FFF89FF82FE409277FD102B56" box="[494,657,1892,1918]" pageId="81" pageNumber="82" value="2002-02-21">21 Feb. 2002</collectingDate>
</date>
;
<collectionCode id="ED6C0192FF89FF82FD0E9277FD712B56" box="[672,752,1892,1918]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FCA09277FBDE2B8A" collectingDate="1996-03-03" collectingDateMax="1996-03-04" collectingDateMin="1996-03-03" collectingMethod="yellow pans" collectionCode="DEBU" collectorName="L. Masner" country="Costa Rica" location="near border of Braulio Carrillo National Park" municipality="Moravia" pageId="81" pageNumber="82" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FCA09277FCBE2B56" box="[782,831,1892,1918]" count="1" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7B52DEFF89FF82FCE09277FCFF2B56" box="[846,894,1892,1918]" count="1" pageId="81" pageNumber="82" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF89FF82FC219277FC742B56" box="[911,1013,1892,1918]" pageId="81" pageNumber="82">Moravia</collectingMunicipality>
,
<location id="8EA2CF8CFF89FF82FBAA9277FEE52B89" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FBAA9277FEE52B89" country="Costa Rica" municipality="Moravia" name="near border of Braulio Carrillo National Park" pageId="81" pageNumber="82" stateProvince="Heredia">near border of Braulio Carrillo National Park</location>
;
<date id="FFC3BF97FF89FF82FEDE9294FDA02B89" box="[368,545,1927,1954]" pageId="81" pageNumber="82" value="1996-03-03" valueMax="1996-03-04" valueMin="1996-03-03">
<collectingDate id="EF87467FFF89FF82FEDE9294FDA02B89" box="[368,545,1927,1954]" pageId="81" pageNumber="82" value="1996-03-03" valueMax="1996-03-04" valueMin="1996-03-03">34 Mar. 1996</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FD80929BFD292B8A" box="[558,680,1928,1954]" pageId="81" pageNumber="82">L. Masner</collectorName>
leg.;
<location id="8EA2CF8CFF89FF82FD429294FCE12B89" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FD429294FCE12B89" box="[748,864,1927,1953]" country="Costa Rica" municipality="Moravia" name="creek bed" pageId="81" pageNumber="82" stateProvince="Heredia">creek bed</location>
,
<collectingMethod id="523CE140FF89FF82FCC39294FC7F2B8A" box="[877,1022,1927,1954]" pageId="81" pageNumber="82">yellow pans</collectingMethod>
;
<collectionCode id="ED6C0192FF89FF82FBA5929BFBDE2B8A" box="[1035,1119,1928,1954]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF89FF82FB2C9294FB422BED" collectingDate="2000-06-13" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="81" pageNumber="82" specimenCount="6" specimenCount-male="6" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B9094545FF89FF82FB2C9294FA902B89" bold="true" box="[1154,1297,1927,1953]" pageId="81" pageNumber="82">
<collectingRegion id="49B957B5FF89FF82FB2C9294FA902B89" box="[1154,1297,1927,1953]" country="Costa Rica" name="Puntarenas" pageId="81" pageNumber="82">Puntarenas</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF89FF82FA859294FAF02B8A" box="[1323,1393,1927,1954]" count="6" pageId="81" pageNumber="82" type="male">6 ♂♂</specimenCount>
;
<location id="8EA2CF8CFF89FF82FF1392B9FDB42BED" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF89FF82FF1392B9FDB42BED" box="[189,565,1962,1989]" country="Costa Rica" name="Monteverde Biological Reserve" pageId="81" pageNumber="82" stateProvince="Puntarenas">Monteverde Biological Reserve</location>
;
<elevation id="00507E64FF89FF82FDEF92B9FD5B2BEC" box="[577,730,1962,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="m" value="1500.0">
<quantity id="4C8534B2FF89FF82FDEF92B9FD1B2BED" box="[577,666,1962,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF89FF82FD4992B8FC002BED" box="[743,897,1963,1989]" pageId="81" pageNumber="82" value="2000-06-13">
<collectingDate id="EF87467FFF89FF82FD4992B8FC002BED" box="[743,897,1963,1989]" pageId="81" pageNumber="82" value="2000-06-13">13 Jun. 2000</collectingDate>
</date>
;
<collectorName id="2688FC81FF89FF82FC2092B8FBB02BEC" box="[910,1073,1962,1989]" pageId="81" pageNumber="82">S.A. Marshall</collectorName>
leg.;
<collectionCode id="ED6C0192FF89FF82FBDD92B8FB422BED" box="[1139,1219,1963,1989]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FB7292B9FDE52BC0" collectingDate="2000-06-13" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="81" pageNumber="82" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FB7292B9FAA12BED" box="[1244,1312,1962,1989]" count="6" pageId="81" pageNumber="82" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF89FF82FA8292B8FAEE2BED" box="[1324,1391,1963,1989]" count="3" pageId="81" pageNumber="82" type="female">3 ♀♀</specimenCount>
; same data as for preceding;
<collectionCode id="ED6C0192FF89FF82FDA092DDFDE52BC0" box="[526,612,1998,2024]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="81" pageNumber="82">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF82FD2F92DDFA8E2BC0" box="[641,1295,1998,2024]" collectingDate="2000-06-13" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="81" pageNumber="82" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FD2F92DDFD332BC0" box="[641,690,1998,2024]" count="1" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
; same data as for preceding;
<collectorName id="2688FC81FF89FF82FBBF92DDFBFB2BC0" box="[1041,1146,1998,2024]" pageId="81" pageNumber="82">M. Buck</collectorName>
leg.;
<collectionCode id="ED6C0192FF89FF82FB1192DDFA8E2BC0" box="[1215,1295,1998,2024]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="81" pageNumber="82">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF89FF81FA8592DDFB972D0B" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" lastPageId="82" lastPageNumber="83" location="cloud forest" pageId="81" pageNumber="82" specimenCount="12" specimenCount-female="3" specimenCount-male="9" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF89FF82FA8592DDFAF02BC0" box="[1323,1393,1998,2024]" count="9" pageId="81" pageNumber="82" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FF13941AFE812D0B" box="[189,256,265,291]" count="3" pageId="82" pageNumber="83" type="female">3 ♀♀</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FDF7941AFC9F2D0B" box="[601,798,265,291]" pageId="82" pageNumber="83" value="2000-06-11" valueMax="2000-06-13" valueMin="2000-06-11">
<collectingDate id="EF87467FFF8AFF81FDF7941AFC9F2D0B" box="[601,798,265,291]" pageId="82" pageNumber="83" value="2000-06-11" valueMax="2000-06-13" valueMin="2000-06-11">1113 Jun. 2000</collectingDate>
</date>
;
<location id="8EA2CF8CFF8AFF81FC85941AFC3A2D0B" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FC85941AFC3A2D0B" box="[811,955,265,291]" country="Costa Rica" name="cloud forest" pageId="82" pageNumber="83" stateProvince="Puntarenas">cloud forest</location>
;
<collectionCode id="ED6C0192FF8AFF81FC68941AFB972D0B" box="[966,1046,265,291]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FB9E941AFE412D6E" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="MNCR" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FB9E941AFB022D0B" box="[1072,1155,265,291]" count="10" pageId="82" pageNumber="83" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FB3E941AFB522D0B" box="[1168,1235,265,291]" count="4" pageId="82" pageNumber="83" type="female">4 ♀♀</specimenCount>
; same data as for preceding;
<collectionCode id="ED6C0192FF8AFF81FEC4943FFE412D6E" box="[362,448,300,326]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FE77943FFA862D6F" box="[473,1287,300,327]" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="13" specimenCount-female="3" specimenCount-male="10" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FE77943FFDAA2D6F" box="[473,555,300,327]" count="10" pageId="82" pageNumber="83" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FD96943FFDFA2D6F" box="[568,635,300,327]" count="3" pageId="82" pageNumber="83" type="female">3 ♀♀</specimenCount>
; same data as for preceding;
<collectorName id="2688FC81FF8AFF81FC7F943FFBF42D6E" box="[977,1141,300,326]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<collectionCode id="ED6C0192FF8AFF81FB19943EFA862D6F" box="[1207,1287,301,327]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FA8E943FFD392D42" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="14" specimenCount-female="3" specimenCount-male="11" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FA8E943FFAF02D6F" box="[1312,1393,300,327]" count="11" pageId="82" pageNumber="83" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FF139443FE832D42" box="[189,258,336,362]" count="3" pageId="82" pageNumber="83" type="female">3 ♀♀</specimenCount>
; same data as for preceding;
<collectionCode id="ED6C0192FF8AFF81FDCC9443FD392D42" box="[610,696,336,362]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FD7A9443FE362DA5" collectedFrom="on dung" collectingDate="2000-06-12" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FD7A9443FC822D42" box="[724,771,336,362]" count="1" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FBCD9443FA812D42" box="[1123,1280,336,362]" pageId="82" pageNumber="83" value="2000-06-12">
<collectingDate id="EF87467FFF8AFF81FBCD9443FA812D42" box="[1123,1280,336,362]" pageId="82" pageNumber="83" value="2000-06-12">12 Jun. 2000</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FAA09443FAF72D41" box="[1294,1398,335,362]" pageId="82" pageNumber="83">M. Buck</collectorName>
leg.;
<collectedFrom id="3861DAAAFF8AFF81FF569460FEDB2DA5" box="[248,346,371,397]" pageId="82" pageNumber="83">on dung</collectedFrom>
;
<collectionCode id="ED6C0192FF8AFF81FEC99460FE362DA5" box="[359,439,371,397]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FE619461FE8C2D98" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectingMethod="sweep" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="69" specimenCount-female="19" specimenCount-male="50" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FE619461FDA32DA5" box="[463,546,370,397]" count="50" pageId="82" pageNumber="83" type="male">50 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FD819460FD002DA5" box="[559,641,371,397]" count="19" pageId="82" pageNumber="83" type="female">19 ♀♀</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FC769460FBF02DA5" box="[984,1137,371,397]" pageId="82" pageNumber="83" value="2000-06-11">
<collectingDate id="EF87467FFF8AFF81FC769460FBF02DA5" box="[984,1137,371,397]" pageId="82" pageNumber="83" value="2000-06-11">11 Jun. 2000</collectingDate>
</date>
,
<collectorName id="2688FC81FF8AFF81FBD29460FB622DA5" box="[1148,1251,371,397]" pageId="82" pageNumber="83">M. Buck</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FA8B9460FAF32DA5" box="[1317,1394,371,397]" pageId="82" pageNumber="83">sweep</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FF139485FE8C2D98" box="[189,269,406,432]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FE8B9485FCFF2D98" box="[293,894,406,433]" collectingDate="2000-06-11" collectingDateMax="2000-06-13" collectingDateMin="2000-06-11" collectionCode="MNCR" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="70" specimenCount-female="19" specimenCount-male="51" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FE8B9485FEF62D98" box="[293,375,406,433]" count="51" pageId="82" pageNumber="83" type="male">51 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FE2D9485FE552D98" box="[387,468,406,432]" count="19" pageId="82" pageNumber="83" type="female">19 ♀♀</specimenCount>
; same data as for preceding;
<collectionCode id="ED6C0192FF8AFF81FC869485FCFF2D98" box="[808,894,406,432]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FC399485FDF32DFC" collectingDate="2000-06-12" collectingDateMax="2000-06-13" collectingDateMin="2000-06-12" collectingMethod="pans along stream" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="8" specimenCount-male="8" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FC399485FC5A2D98" box="[919,987,406,432]" count="8" pageId="82" pageNumber="83" type="male">8 ♂♂</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FA839485FEB32DFB" pageId="82" pageNumber="83" value="2000-06-12" valueMax="2000-06-13" valueMin="2000-06-12">
<collectingDate id="EF87467FFF8AFF81FA839485FEB32DFB" pageId="82" pageNumber="83" value="2000-06-12" valueMax="2000-06-13" valueMin="2000-06-12">1213 Jun. 2000</collectingDate>
</date>
;
<collectingMethod id="523CE140FF8AFF81FE9194A9FD972DFC" box="[319,534,441,468]" pageId="82" pageNumber="83">pans along stream</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FD8C94A9FDF32DFC" box="[546,626,442,468]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FD2594AAFEF12DDE" collectingDate="2000-06-13" collectingDateMax="2000-06-14" collectingDateMin="2000-06-13" collectingMethod="pans along stream" collectionCode="MNCR" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="6" specimenCount-male="6" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FD2594AAFD512DFC" box="[651,720,441,468]" count="6" pageId="82" pageNumber="83" type="male">6 ♂♂</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FB8894AAFB6D2DFB" box="[1062,1260,441,468]" pageId="82" pageNumber="83" value="2000-06-13" valueMax="2000-06-14" valueMin="2000-06-13">
<collectingDate id="EF87467FFF8AFF81FB8894AAFB6D2DFB" box="[1062,1260,441,468]" pageId="82" pageNumber="83" value="2000-06-13" valueMax="2000-06-14" valueMin="2000-06-13">1314 Jun. 2000</collectingDate>
</date>
;
<collectingMethod id="523CE140FF8AFF81FB5694A9FE8E2DDF" pageId="82" pageNumber="83">pans along stream</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FEB494CFFEF12DDE" box="[282,368,476,502]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FE2994CFFA962DDF" box="[391,1303,476,503]" collectingDate="2000-06-14" collectingDateMax="2000-06-14" collectingDateMin="2000-06-13" collectingMethod="treefall sweep and pans" collectionCode="DEBU" collectorName="M. Buck" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FE2994CFFE342DDF" box="[391,437,476,503]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FCAB94CFFC1C2DDE" box="[773,925,476,503]" pageId="82" pageNumber="83" value="2000-06-14">
<collectingDate id="EF87467FFF8AFF81FCAB94CFFC1C2DDE" box="[773,925,476,503]" pageId="82" pageNumber="83" value="2000-06-14">14 Jun. 2000</collectingDate>
</date>
;
<collectingMethod id="523CE140FF8AFF81FC0694CFFB3D2DDF" box="[936,1212,476,503]" pageId="82" pageNumber="83">treefall sweep and pans</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FB6994CEFA962DDF" box="[1223,1303,477,503]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FA8094CEFBF62E32" collectingDate="1998-05-26" collectingDateMax="2000-06-14" collectingDateMin="2000-06-13" collectingMethod="sweep" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="1500" location="Monteverde Biological Reserve" pageId="82" pageNumber="83" specimenCount="11" specimenCount-female="4" specimenCount-male="7" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FA8094CEFAF02DDF" box="[1326,1393,477,503]" count="7" pageId="82" pageNumber="83" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FF139713FF7F2E32" box="[189,254,512,538]" count="4" pageId="82" pageNumber="83" type="female">4 ♀♀</specimenCount>
; same data as for preceding;
<date id="FFC3BF97FF8AFF81FDE494ECFD672E32" box="[586,742,511,538]" pageId="82" pageNumber="83" value="1998-05-26">
<collectingDate id="EF87467FFF8AFF81FDE494ECFD672E32" box="[586,742,511,538]" pageId="82" pageNumber="83" value="1998-05-26">26 May 1998</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FD5E9713FC132E31" box="[752,914,511,538]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FC7E9713FB9C2E32" box="[976,1053,512,538]" pageId="82" pageNumber="83">sweep</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FB899713FBF62E32" box="[1063,1143,512,538]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FB229713FBB82E15" collectingDate="1998-05-26" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" location="Monteverde Biological Station" pageId="82" pageNumber="83" specimenCount="13" specimenCount-male="13" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FB229713FB5F2E32" box="[1164,1246,512,538]" count="13" pageId="82" pageNumber="83" type="male">13 ♂♂</specimenCount>
;
<location id="8EA2CF8CFF8AFF81FB4794ECFE182E15" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB4794ECFE182E15" country="Costa Rica" name="Monteverde Biological Station" pageId="82" pageNumber="83" stateProvince="Puntarenas">Monteverde Biological Station</location>
,
<location id="8EA2CF8CFF8AFF81FE069730FDA82E15" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FE069730FDA82E15" box="[424,553,547,573]" country="Costa Rica" name="lower trail" pageId="82" pageNumber="83" stateProvince="Puntarenas">lower trail</location>
;
<date id="FFC3BF97FF8AFF81FD979730FD602E15" box="[569,737,547,573]" pageId="82" pageNumber="83" value="1998-05-26">
<collectingDate id="EF87467FFF8AFF81FD979730FD602E15" box="[569,737,547,573]" pageId="82" pageNumber="83" value="1998-05-26">26 May 1998</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FD5F9730FC182E15" box="[753,921,547,573]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<collectionCode id="ED6C0192FF8AFF81FC4D9730FBB82E15" box="[995,1081,547,573]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FBF49730FEDD2EAB" collectingDate="1988-02-24" collectingDateMax="1988-03-02" collectingDateMin="1988-02-24" collectingMethod="pan traps" collectionCode="ROME" collectorName="B. Hubley" country="Costa Rica" elevation="1539" latitude="10.3" location="Monteverde" longLatPrecision="1298" longitude="-84.8" pageId="82" pageNumber="83" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FBF49730FB0D2E15" box="[1114,1164,547,573]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FB339730FB4F2E15" box="[1181,1230,547,573]" count="1" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
;
<location id="8EA2CF8CFF8AFF81FB4F9730FAF22E15" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB4F9730FAF22E15" box="[1249,1395,547,573]" country="Costa Rica" latitude="10.3" longLatPrecision="1298" longitude="-84.8" name="Monteverde" pageId="82" pageNumber="83" stateProvince="Puntarenas">Monteverde</location>
;
<geoCoordinate id="EE49FF90FF8AFF81FF139757FEAB2E49" box="[189,298,580,609]" degrees="10" direction="north" minutes="18" orientation="latitude" pageId="82" pageNumber="83" precision="925" value="10.3">10°18 N</geoCoordinate>
,
<geoCoordinate id="EE49FF90FF8AFF81FE969757FE2B2E49" box="[312,426,580,609]" degrees="84" direction="west" minutes="48" orientation="longitude" pageId="82" pageNumber="83" precision="925" value="-84.8">84°48 W</geoCoordinate>
;
<elevation id="00507E64FF8AFF81FE169755FDD22E48" box="[440,595,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.539" pageId="82" pageNumber="83" unit="m" value="1539.0">
<quantity id="4C8534B2FF8AFF81FE169755FD932E49" box="[440,530,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.539" pageId="82" pageNumber="83" unit="m" value="1539.0">1539 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FDCC9755FCDF2E48" box="[610,862,582,609]" pageId="82" pageNumber="83" value="1988-02-24" valueMax="1988-03-02" valueMin="1988-02-24">
<collectingDate id="EF87467FFF8AFF81FDCC9755FCDF2E48" box="[610,862,582,609]" pageId="82" pageNumber="83" value="1988-02-24" valueMax="1988-03-02" valueMin="1988-02-24">24 Feb.2 Mar. 1988</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FCC29754FC642E48" box="[876,997,582,609]" pageId="82" pageNumber="83">B. Hubley</collectorName>
leg.;
<location id="8EA2CF8CFF8AFF81FB849755FABE2E48" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB849755FABE2E48" box="[1066,1343,582,608]" country="Costa Rica" latitude="10.3" longLatPrecision="1298" longitude="-84.8" name="1 ° tropical cloud forest" pageId="82" pageNumber="83" stateProvince="Puntarenas">1° tropical cloud forest</location>
,
<collectingMethod id="523CE140FF8AFF81FAE29754FF7B2EAC" pageId="82" pageNumber="83">pan traps</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FEA9977AFEDD2EAB" box="[263,348,617,643]" collectionName="Canada, Ontario, Toronto, Royal Ontario Museum" country="Canada" httpUri="http://grbio.org/cool/2yxb-9en0" name="Royal Ontario Museum - Entomology" pageId="82" pageNumber="83" type="Museum">ROME</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FED7977AFE062E8F" collectingDate="1987-05-24" collectionCode="DEBU" collectorName="Norrbom &amp; Max" country="Costa Rica" latitude="10.3" location="Pension Queteal" longLatPrecision="1298" longitude="-84.816666" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FED7977AFE2B2EAC" box="[377,426,617,644]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FE16977AFDC82EAB" box="[440,585,617,643]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
,
<location id="8EA2CF8CFF8AFF81FDF8977AFC9A2EAB" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FDF8977AFC9A2EAB" box="[598,795,617,643]" country="Costa Rica" latitude="10.3" longLatPrecision="1298" longitude="-84.816666" municipality="Monteverde" name="Pension Queteal" pageId="82" pageNumber="83" stateProvince="Puntarenas">Pension Queteal</location>
;
<geoCoordinate id="EE49FF90FF8AFF81FC87977BFC162EAC" box="[809,919,616,644]" degrees="10" direction="north" minutes="18" orientation="latitude" pageId="82" pageNumber="83" precision="925" value="10.3">10°18 N</geoCoordinate>
,
<geoCoordinate id="EE49FF90FF8AFF81FC0A977BFB992EAC" box="[932,1048,616,644]" degrees="84" direction="west" minutes="49" orientation="longitude" pageId="82" pageNumber="83" precision="925" value="-84.816666">84°49 W</geoCoordinate>
;
<date id="FFC3BF97FF8AFF81FB88977AFB482EAB" box="[1062,1225,617,644]" pageId="82" pageNumber="83" value="1987-05-24">
<collectingDate id="EF87467FFF8AFF81FB88977AFB482EAB" box="[1062,1225,617,644]" pageId="82" pageNumber="83" value="1987-05-24">24 May 1987</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FB79977AFAC22EAB" box="[1239,1347,617,643]" pageId="82" pageNumber="83">Norrbom</collectorName>
and
<collectorName id="2688FC81FF8AFF81FF13979EFF722E8F" box="[189,243,653,679]" pageId="82" pageNumber="83">Max</collectorName>
leg.;
<collectionCode id="ED6C0192FF8AFF81FE99979EFE062E8F" box="[311,391,653,679]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FE0C979EFED62EE2" collectingDate="1983-07-11" collectingDateMax="1983-07-18" collectingDateMin="1983-07-11" collectingMethod="fruit pitfall" collectionCode="CNCI" collectorName="D. H. Lindeman" country="Costa Rica" elevation="1500" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FE0C979EFE502E8F" box="[418,465,653,679]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FE71979EFDA22E8F" box="[479,547,653,679]" count="2" pageId="82" pageNumber="83" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FD9C979FFD452E8E" box="[562,708,652,678]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<elevation id="00507E64FF8AFF81FD7E979FFCEA2E8E" box="[720,875,652,679]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">
<quantity id="4C8534B2FF8AFF81FD7E979FFCAB2E8F" box="[720,810,652,679]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FCD4979EFBBB2E8F" box="[890,1082,652,679]" pageId="82" pageNumber="83" value="1983-07-11" valueMax="1983-07-18" valueMin="1983-07-11">
<collectingDate id="EF87467FFF8AFF81FCD4979EFBBB2E8F" box="[890,1082,652,679]" pageId="82" pageNumber="83" value="1983-07-11" valueMax="1983-07-18" valueMin="1983-07-11">1118 Jul. 1983</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FBE6979EFA812E8E" box="[1096,1280,652,679]" pageId="82" pageNumber="83">D.H. Lindeman</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FAEA979FFE862EE2" pageId="82" pageNumber="83">fruit pitfall</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FEBF97A3FED62EE2" box="[273,343,688,714]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="82" pageNumber="83">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FEC097A3FAE72EE2" box="[366,1382,687,714]" collectingDate="1988-02-29" collectionCode="DEBU" collectorName="Mason &amp; Wood" country="Costa Rica" elevation="1500" location="rainforest" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FEC097A3FE1D2EE2" box="[366,412,688,714]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FE0997A3FDB82EE2" box="[423,569,688,714]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<elevation id="00507E64FF8AFF81FDED97BCFD5B2EE2" box="[579,730,687,714]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">
<quantity id="4C8534B2FF8AFF81FDED97BCFD1A2EE2" box="[579,667,687,714]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FD4897A3FC002EE2" box="[742,897,688,714]" pageId="82" pageNumber="83" value="1988-02-29">
<collectingDate id="EF87467FFF8AFF81FD4897A3FC002EE2" box="[742,897,688,714]" pageId="82" pageNumber="83" value="1988-02-29">29 Feb. 1988</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FC2297A3FC5D2EE2" box="[908,988,688,714]" pageId="82" pageNumber="83">Mason</collectorName>
and
<collectorName id="2688FC81FF8AFF81FBBC97A3FBD62EE2" box="[1042,1111,688,714]" pageId="82" pageNumber="83">Wood</collectorName>
leg.;
<location id="8EA2CF8CFF8AFF81FB3997A3FA8A2EE2" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB3997A3FA8A2EE2" box="[1175,1291,688,714]" country="Costa Rica" municipality="Monteverde" name="rainforest" pageId="82" pageNumber="83" stateProvince="Puntarenas">rainforest</location>
;
<collectionCode id="ED6C0192FF8AFF81FAB897A3FAE72EE2" box="[1302,1382,688,714]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FF1397C0FA962EC5" box="[189,1303,723,750]" collectingDate="1991-02-24" collectingDateMax="1991-02-27" collectingDateMin="1991-02-24" collectingMethod="trail sweep" collectionCode="DEBU" collectorName="B. J. Sinclair" country="Costa Rica" elevation="1650" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FF1397C0FE802EC5" box="[189,257,723,749]" count="2" pageId="82" pageNumber="83" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FEA397C0FE1E2EC5" box="[269,415,723,749]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<elevation id="00507E64FF8AFF81FE0797C0FD0B2EC5" box="[425,650,723,750]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="82" pageNumber="83" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<quantity id="4C8534B2FF8AFF81FE0797C0FDCA2EC5" box="[425,587,723,750]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="82" pageNumber="83" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">15001800 m</quantity>
a.s.l.
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;
<date id="FFC3BF97FF8AFF81FD3897C0FCDF2EC5" box="[662,862,723,749]" pageId="82" pageNumber="83" value="1991-02-24" valueMax="1991-02-27" valueMin="1991-02-24">
<collectingDate id="EF87467FFF8AFF81FD3897C0FCDF2EC5" box="[662,862,723,749]" pageId="82" pageNumber="83" value="1991-02-24" valueMax="1991-02-27" valueMin="1991-02-24">2427 Feb. 1991</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FCC797C0FC782EC5" box="[873,1017,723,749]" pageId="82" pageNumber="83">B.J. Sinclair</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FB9797C0FB3D2EC5" box="[1081,1212,723,749]" pageId="82" pageNumber="83">trail sweep</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FB6997C0FA962EC5" box="[1223,1303,723,749]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
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<materialsCitation id="3B15930AFF8AFF81FA8097C0FAE22F38" collectingDate="1983-07-15" collectingDateMax="1983-07-23" collectingDateMin="1983-07-15" collectingMethod="flight-intercept trap" collectionCode="CNCI" collectorName="D. H. Lindeman" country="Costa Rica" elevation="1520" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FA8097C0FAF02EC5" box="[1326,1393,723,749]" count="2" pageId="82" pageNumber="83" type="male">2 ♂♂</specimenCount>
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<specimenCount id="9D7B52DEFF8AFF81FF1397E5FE802F39" box="[189,257,758,785]" count="2" pageId="82" pageNumber="83" type="female">2 ♀♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FEBF97E5FE222F38" box="[273,419,758,784]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<elevation id="00507E64FF8AFF81FE1E97E5FDCD2F38" box="[432,588,758,785]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="82" pageNumber="83" unit="m" value="1520.0">
<quantity id="4C8534B2FF8AFF81FE1E97E5FD8B2F39" box="[432,522,758,785]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="82" pageNumber="83" unit="m" value="1520.0">1520 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FDF497E5FC9C2F38" box="[602,797,758,785]" pageId="82" pageNumber="83" value="1983-07-15" valueMax="1983-07-23" valueMin="1983-07-15">
<collectingDate id="EF87467FFF8AFF81FDF497E5FC9C2F38" box="[602,797,758,785]" pageId="82" pageNumber="83" value="1983-07-15" valueMax="1983-07-23" valueMin="1983-07-15">1523 Jul. 1983</collectingDate>
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;
<collectorName id="2688FC81FF8AFF81FC8497E4FC622F38" box="[810,995,758,785]" pageId="82" pageNumber="83">D.H. Lindeman</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FB8697E5FA912F39" box="[1064,1296,758,785]" pageId="82" pageNumber="83">flight-intercept trap</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FAB397E5FAE22F38" box="[1309,1379,758,784]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="82" pageNumber="83">CNCI</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FF139609FC492F1C" box="[189,968,793,820]" collectingDate="1991-02-23" collectingDateMax="1991-02-27" collectingDateMin="1991-02-23" collectionCode="DEBU" collectorName="B. J. Sinclair" country="Costa Rica" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FF139609FF6A2F1C" box="[189,235,794,820]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7B52DEFF8AFF81FF569609FEA42F1C" box="[248,293,794,820]" count="1" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FE9D960AFE442F1B" box="[307,453,793,819]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<date id="FFC3BF97FF8AFF81FE7E9609FD182F1C" box="[464,665,793,820]" pageId="82" pageNumber="83" value="1991-02-23" valueMax="1991-02-27" valueMin="1991-02-23">
<collectingDate id="EF87467FFF8AFF81FE7E9609FD182F1C" box="[464,665,793,820]" pageId="82" pageNumber="83" value="1991-02-23" valueMax="1991-02-27" valueMin="1991-02-23">2327 Feb. 1991</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FD0B9609FCB42F1B" box="[677,821,793,820]" pageId="82" pageNumber="83">B.J. Sinclair</collectorName>
leg.;
<collectionCode id="ED6C0192FF8AFF81FCD69609FC492F1C" box="[888,968,794,820]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FC4E9609FBF92F7F" collectedFrom="treefall" collectingDate="1988-05-28" collectingDateMax="1988-06-01" collectingDateMin="1988-05-28" collectingMethod="Malaise trap" collectionCode="DEBU" collectorName="B. V. Brown" country="Costa Rica" elevation="1500" location="Finca Canada" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FC4E9609FBA42F1C" box="[992,1061,794,820]" count="2" pageId="82" pageNumber="83" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FB9C960AFB422F1B" box="[1074,1219,793,819]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
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<location id="8EA2CF8CFF8AFF81FB60960AFAF32F1B" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB60960AFAF32F1B" box="[1230,1394,793,819]" country="Costa Rica" municipality="Monteverde" name="Finca Canada" pageId="82" pageNumber="83" stateProvince="Puntarenas">Finca Canada</location>
;
<elevation id="00507E64FF8AFF81FF13962FFED42F7F" box="[189,341,828,855]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">
<quantity id="4C8534B2FF8AFF81FF13962FFE972F7F" box="[189,278,828,855]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="83" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
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;
<date id="FFC3BF97FF8AFF81FECC962EFDD72F7F" box="[354,598,829,855]" pageId="82" pageNumber="83" value="1988-05-28" valueMax="1988-06-01" valueMin="1988-05-28">
<collectingDate id="EF87467FFF8AFF81FECC962EFDD72F7F" box="[354,598,829,855]" pageId="82" pageNumber="83" value="1988-05-28" valueMax="1988-06-01" valueMin="1988-05-28">28 May1 Jun. 1988</collectingDate>
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;
<collectorName id="2688FC81FF8AFF81FDCC962EFD6D2F7F" box="[610,748,829,855]" pageId="82" pageNumber="83">B.V. Brown</collectorName>
leg.;
<collectedFrom id="3861DAAAFF8AFF81FC80962EFC052F7F" box="[814,900,829,855]" pageId="82" pageNumber="83">treefall</collectedFrom>
,
<collectingMethod id="523CE140FF8AFF81FC21962EFBA02F7F" box="[911,1057,829,855]" pageId="82" pageNumber="83">Malaise trap</collectingMethod>
<collectionCode id="ED6C0192FF8AFF81FB86962EFBF92F7F" box="[1064,1144,829,855]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
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<materialsCitation id="3B15930AFF8AFF81FB3E962EFC572F52" collectingDate="1991-02-25" collectingMethod="flight-intercept trap" collectionCode="DEBU" collectorName="H. and A. Howden" country="Costa Rica" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="3" specimenCount-male="3" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FB3E962EFB552F7F" box="[1168,1236,829,855]" count="3" pageId="82" pageNumber="83" type="male">3 ♂♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FB4F962EFAF22F7F" box="[1249,1395,829,855]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<date id="FFC3BF97FF8AFF81FF139673FEDC2F52" box="[189,349,864,891]" pageId="82" pageNumber="83" value="1991-02-25">
<collectingDate id="EF87467FFF8AFF81FF139673FEDC2F52" box="[189,349,864,891]" pageId="82" pageNumber="83" value="1991-02-25">25 Feb. 1991</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FEC59673FDCB2F52" box="[363,586,864,890]" pageId="82" pageNumber="83">H. and A. Howden</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FD3E9673FCF92F52" box="[656,888,864,890]" pageId="82" pageNumber="83">flight-intercept trap</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FC289673FC572F52" box="[902,982,864,890]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FC5C9673FCFB2FB6" collectingDate="1998-05-25" collectingMethod="sweep near Biological Station" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" location="Monteverde" municipality="Monteverde" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FC5C9673FBA32F52" box="[1010,1058,864,890]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FB9E9673FB432F52" box="[1072,1218,864,890]" pageId="82" pageNumber="83">Monteverde</collectingMunicipality>
;
<date id="FFC3BF97FF8AFF81FB619673FAF32F52" box="[1231,1394,864,891]" pageId="82" pageNumber="83" value="1998-05-25">
<collectingDate id="EF87467FFF8AFF81FB619673FAF32F52" box="[1231,1394,864,891]" pageId="82" pageNumber="83" value="1998-05-25">25 May 1998</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FF139690FEE22FB5" box="[189,355,899,925]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FE059697FC962FB5" box="[427,791,899,926]" pageId="82" pageNumber="83">sweep near Biological Station</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FC889697FCFB2FB6" box="[806,890,900,926]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF8AFF81FC0E9690FB0B2FE8" collectingDate="1995-08-09" collectingMethod="sweep along forest stream" collectionCode="MNCR" collectorName="S. A. Marshall" country="Costa Rica" elevation="2300" location="San Gerardo de Dota" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<emphasis id="B9094545FF8AFF81FC0E9690FB8E2FB5" bold="true" box="[928,1039,898,925]" pageId="82" pageNumber="83">
<collectingRegion id="49B957B5FF8AFF81FC0E9690FB8E2FB5" box="[928,1039,898,925]" country="Costa Rica" name="San Jose" pageId="82" pageNumber="83">San José</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF8AFF81FB809690FBE12FB6" box="[1070,1120,899,926]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<location id="8EA2CF8CFF8AFF81FBC19690FAF32FB6" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FBC19690FAF32FB6" box="[1135,1394,899,926]" country="Costa Rica" name="San Gerardo de Dota" pageId="82" pageNumber="83" stateProvince="San Jose">San Gerardo de Dota</location>
;
<elevation id="00507E64FF8AFF81FF1396B5FED62FE8" box="[189,343,934,961]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="82" pageNumber="83" unit="m" value="2300.0">
<quantity id="4C8534B2FF8AFF81FF1396B5FE962FE9" box="[189,279,934,961]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="82" pageNumber="83" unit="m" value="2300.0">2300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FEC896B5FE7A2FE9" box="[358,507,934,961]" pageId="82" pageNumber="83" value="1995-08-09">
<collectingDate id="EF87467FFF8AFF81FEC896B5FE7A2FE9" box="[358,507,934,961]" pageId="82" pageNumber="83" value="1995-08-09">9 Aug. 1995</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FDA696B5FD2D2FE8" box="[520,684,934,960]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<collectingMethod id="523CE140FF8AFF81FD5E96B4FBA92FE9" box="[752,1064,934,961]" pageId="82" pageNumber="83">sweep along forest stream</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FB9A96B5FB0B2FE8" box="[1076,1162,934,960]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="82" pageNumber="83">MNCR</collectionCode>
</materialsCitation>
<materialsCitation id="3B15930AFF8AFF81FB0B96B5FAE42FCC" collectingDate="1996-08-08" collectingDateMax="1996-08-09" collectingDateMin="1996-08-08" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="2500" latitude="9.55" location="San Gerardo de Dota" longLatPrecision="1299" longitude="-83.8" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FB0B96B5FB542FE9" box="[1189,1237,934,961]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<location id="8EA2CF8CFF8AFF81FB4C96B5FE9C2FCC" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB4C96B5FE9C2FCC" country="Costa Rica" latitude="9.55" longLatPrecision="1299" longitude="-83.8" name="San Gerardo de Dota" pageId="82" pageNumber="83" stateProvince="San Jose">San Gerardo de Dota</location>
;
<geoCoordinate id="EE49FF90FF8AFF81FE8696DBFE042FCC" box="[296,389,968,996]" degrees="9" direction="north" minutes="33" orientation="latitude" pageId="82" pageNumber="83" precision="925" value="9.55">9°33 N</geoCoordinate>
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<elevation id="00507E64FF8AFF81FDA096DAFD6E2FCB" box="[526,751,969,996]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="82" pageNumber="83" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">
<quantity id="4C8534B2FF8AFF81FDA096DAFD312FCC" box="[526,688,969,996]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="82" pageNumber="83" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">24002600 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FD5296D9FC2C2FCB" box="[764,941,969,996]" pageId="82" pageNumber="83" value="1996-08-08" valueMax="1996-08-09" valueMin="1996-08-08">
<collectingDate id="EF87467FFF8AFF81FD5296D9FC2C2FCB" box="[764,941,969,996]" pageId="82" pageNumber="83" value="1996-08-08" valueMax="1996-08-09" valueMin="1996-08-08">89 Aug. 1996</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FC1796D9FBDD2FCB" box="[953,1116,969,996]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<location id="8EA2CF8CFF8AFF81FB3396D9FA8B2FCB" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB3396D9FA8B2FCB" box="[1181,1290,969,996]" country="Costa Rica" latitude="9.55" longLatPrecision="1299" longitude="-83.8" name="near trail" pageId="82" pageNumber="83" stateProvince="San Jose">near trail</location>
;
<collectionCode id="ED6C0192FF8AFF81FABB96D9FAE42FCC" box="[1301,1381,970,996]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
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<materialsCitation id="3B15930AFF8AFF81FF1396FEFDFD2803" collectingDate="1995-08-09" collectingMethod="sweep" collectionCode="DEBU" collectorName="S. A. Marshall" country="Costa Rica" elevation="2000" latitude="9.55" location="near Lodge" longLatPrecision="1299" longitude="-83.8" municipality="San Gerardo de Dota" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCount id="9D7B52DEFF8AFF81FF1396FEFF6A282F" box="[189,235,1005,1031]" count="1" pageId="82" pageNumber="83" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FF5596FEFE76282F" box="[251,503,1005,1031]" pageId="82" pageNumber="83">San Gerardo de Dota</collectingMunicipality>
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<location id="8EA2CF8CFF8AFF81FDAA96FEFD0F282F" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FDAA96FEFD0F282F" box="[516,654,1005,1031]" country="Costa Rica" latitude="9.55" longLatPrecision="1299" longitude="-83.8" municipality="San Gerardo de Dota" name="near Lodge" pageId="82" pageNumber="83" stateProvince="San Jose">near Lodge</location>
;
<geoCoordinate id="EE49FF90FF8AFF81FD3496F8FD78282F" box="[666,761,1003,1031]" degrees="9" direction="north" minutes="33" orientation="latitude" pageId="82" pageNumber="83" precision="925" value="9.55">9°33 N</geoCoordinate>
,
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<elevation id="00507E64FF8AFF81FC2896FEFBA1282F" box="[902,1056,1005,1031]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="82" pageNumber="83" unit="m" value="2000.0">
<quantity id="4C8534B2FF8AFF81FC2896FEFC61282F" box="[902,992,1005,1031]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="82" pageNumber="83" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FB8196FEFB442820" box="[1071,1221,1005,1032]" pageId="82" pageNumber="83" value="1995-08-09">
<collectingDate id="EF87467FFF8AFF81FB8196FEFB442820" box="[1071,1221,1005,1032]" pageId="82" pageNumber="83" value="1995-08-09">9 Aug. 1995</collectingDate>
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;
<collectorName id="2688FC81FF8AFF81FB7C96FEFAF7282F" box="[1234,1398,1005,1031]" pageId="82" pageNumber="83">S.A. Marshall</collectorName>
leg.;
<location id="8EA2CF8CFF8AFF81FF569102FE422803" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FF569102FE422803" box="[248,451,1040,1067]" country="Costa Rica" latitude="9.55" longLatPrecision="1299" longitude="-83.8" municipality="San Gerardo de Dota" name="stream in pasture" pageId="82" pageNumber="83" stateProvince="San Jose">stream in pasture</location>
,
<collectingMethod id="523CE140FF8AFF81FE619102FD9D2803" box="[463,540,1041,1067]" pageId="82" pageNumber="83">sweep</collectingMethod>
;
<collectionCode id="ED6C0192FF8AFF81FD869102FDFD2803" box="[552,636,1041,1067]" collectionName="Canada, Ontario, Guelph, University of Guelph" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="82" pageNumber="83">DEBU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC29957FF8AFF81FF139142FE782843" blockId="82.[189,505,1105,1131]" box="[189,505,1105,1131]" pageId="82" pageNumber="83">
<heading id="D08A2E3BFF8AFF81FF139142FE782843" bold="true" box="[189,505,1105,1131]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9094545FF8AFF81FF139142FE782843" bold="true" box="[189,505,1105,1131]" pageId="82" pageNumber="83">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8AFF81FF139193FC6928C9" blockId="82.[189,1399,1150,1249]" pageId="82" pageNumber="83">
<materialsCitation id="3B15930AFF8AFF81FF139193FC6528C9" collectingDate="2015-02-12" collectingMethod="Malaise trap" collectionCode="BIOUG" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" elevation="1220" latitude="10.929167" location="Derrumbe" longLatPrecision="21" longitude="-85.464165" municipality="Area de Conservacion Guanacaste" pageId="82" pageNumber="83" specimenCode="BIOUG32728, GMCDQ410-17" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
<collectingCountry id="F36AD9C7FF8AFF81FF139193FEEB28B2" box="[189,362,1152,1178]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
<emphasis id="B9094545FF8AFF81FE3A9193FDA628B2" bold="true" box="[404,551,1152,1178]" pageId="82" pageNumber="83">
<collectingRegion id="49B957B5FF8AFF81FE3A9193FDA628B2" box="[404,551,1152,1178]" country="Costa Rica" name="Guanacaste" pageId="82" pageNumber="83">Guanacaste</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF8AFF81FDE59193FD0128B3" box="[587,640,1152,1179]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BA6032DFF8AFF81FD3F9193FBB828B2" box="[657,1081,1152,1178]" pageId="82" pageNumber="83">Area de Conservacion Guanacaste</collectingMunicipality>
,
<location id="8EA2CF8CFF8AFF81FBE49193FB4428B2" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FBE49193FB4428B2" box="[1098,1221,1152,1178]" country="Costa Rica" latitude="10.929167" longLatPrecision="21" longitude="-85.464165" municipality="Area de Conservacion Guanacaste" name="Derrumbe" pageId="82" pageNumber="83" stateProvince="Guanacaste">Derrumbe</location>
;
<geoCoordinate id="EE49FF90FF8AFF81FB78916DFAF328B3" box="[1238,1394,1150,1179]" degrees="10" direction="north" minutes="55" orientation="latitude" pageId="82" pageNumber="83" precision="15" seconds="45" value="10.929167">10°5545″ N</geoCoordinate>
,
<geoCoordinate id="EE49FF90FF8AFF81FF1391B1FEDA2896" box="[189,347,1186,1214]" degrees="85" direction="west" minutes="27" orientation="longitude" pageId="82" pageNumber="83" precision="15" seconds="51" value="-85.464165">85°2751″ W</geoCoordinate>
;
<elevation id="00507E64FF8AFF81FEC791B0FD862895" box="[361,519,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="82" pageNumber="83" unit="m" value="1220.0">
<quantity id="4C8534B2FF8AFF81FEC791B0FE442896" box="[361,453,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="82" pageNumber="83" unit="m" value="1220.0">1220 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8AFF81FDB991B0FD3B2896" box="[535,698,1187,1214]" pageId="82" pageNumber="83" value="2015-02-12">
<collectingDate id="EF87467FFF8AFF81FDB991B0FD3B2896" box="[535,698,1187,1214]" pageId="82" pageNumber="83" value="2015-02-12">12 Feb. 2015</collectingDate>
</date>
;
<collectorName id="2688FC81FF8AFF81FD6791B7FCDA2896" box="[713,859,1188,1214]" pageId="82" pageNumber="83">D.H. Janzen</collectorName>
and
<collectorName id="2688FC81FF8AFF81FC3791B7FBBE2895" box="[921,1087,1187,1214]" pageId="82" pageNumber="83">W. Hallwachs</collectorName>
leg.;
<location id="8EA2CF8CFF8AFF81FB2891B0FB4D2895" LSID="urn:lsid:plazi:treatment:03D42841FF89FF80FDF996FAFAE32851:8EA2CF8CFF8AFF81FB2891B0FB4D2895" box="[1158,1228,1187,1213]" country="Costa Rica" latitude="10.929167" longLatPrecision="21" longitude="-85.464165" municipality="Area de Conservacion Guanacaste" name="forest" pageId="82" pageNumber="83" stateProvince="Guanacaste">forest</location>
,
<collectingMethod id="523CE140FF8AFF81FB7491B0FAF32896" box="[1242,1394,1187,1214]" pageId="82" pageNumber="83">Malaise trap</collectingMethod>
;
<accessionNumber id="942E04B4FF8AFF81FF1391D4FE2828C9" box="[189,425,1223,1249]" pageId="82" pageNumber="83">BIOUG32728-A03/</accessionNumber>
GMCDQ410-17 sequenced for CO1-5;
<collectionCode id="ED6C0192FF8AFF81FC2D91D4FC6528C9" box="[899,996,1223,1249]" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="82" pageNumber="83" type="University or college">BIOUG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8AFF80FF13901BFC6C2F24" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="description">
<paragraph id="8BC29957FF8AFF81FF13901BFECF290A" blockId="82.[189,334,1288,1314]" box="[189,334,1288,1314]" pageId="82" pageNumber="83">
<heading id="D08A2E3BFF8AFF81FF13901BFECF290A" bold="true" box="[189,334,1288,1314]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9094545FF8AFF81FF13901BFECF290A" bold="true" box="[189,334,1288,1314]" pageId="82" pageNumber="83">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8AFF81FF139024FC532A42" blockId="82.[189,1398,1334,1643]" pageId="82" pageNumber="83">
<smallCapsWord id="8D240F8BFF8AFF81FF139024FF7C2967" baselines="1354,1354" box="[189,253,1335,1361]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="82" pageNumber="83">BODY</smallCapsWord>
(
<figureCitation id="134685D2FF8AFF81FEA29025FEE62978" box="[268,359,1334,1360]" captionStart="Fig" captionStartId="128.[189,232,1929,1955]" captionTargetBox="[189,1399,265,1874]" captionTargetId="figure-51@128.[189,1399,265,1874]" captionTargetPageId="128" captionText="Fig. 1. Sclerocoelus Marshall, 1995, live specimens. A. Sclerocoelus nitidistylus sp. nov. near an Eciton bivouac in Alajuela Province, Costa Rica. B. Probably Sclerocoelus copiosus sp. nov. from Bellavista Cloud Forest Reserve, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.14980856" httpUri="https://zenodo.org/record/14980856/files/figure.png" pageId="82" pageNumber="83">Figs 1A</figureCitation>
,
<figureCitation id="134685D2FF8AFF81FEDD9025FE272978" box="[371,422,1334,1360]" captionStart="Fig" captionStartId="223.[189,232,1860,1886]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure-66@223.[189,1399,265,1830]" captionTargetPageId="223" captionText="Fig. 96. Sclerocoelus nitidistylus sp. nov. A. Male paratype habitus, lateral view (debu00125692). B. Male paratype terminalia, posterior view. C. Male paratype terminalia, lateral view. Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14981080" httpUri="https://zenodo.org/record/14981080/files/figure.png" pageId="82" pageNumber="83">96A</figureCitation>
). Length
<quantity id="4C8534B2FF8AFF81FDBA9025FD232979" box="[532,674,1334,1361]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="82" pageNumber="83" unit="mm" value="2.8" valueMax="3.4" valueMin="2.2">2.23.4 mm</quantity>
. Head dark brown, lower half of frons, gena, face, and antennae orange. Frontal width 2.32.4 × frontal height. Two pairs of strong interfrontal bristles surmounting a fine third pair; anterior orbital 0.40.5 × length of posterior. Palpus yellow. Eye very large, greatest height about 5.0× shortest genal height. Thorax brown, scutum with paler lateral edges. Two pairs of dorsocentral bristles (anterior pair distinct, 0.4× length of posterior pair) separated by 78 rows of acrostichal setulae; prescutellar acrostichal setulae only slightly larger than others. Membrane around prosternum bare. Legs yellowish-brown, base of fore femur paler. Fore femur with three large ventral preapical setae. Ventral surface of male mid tibia with two rows of stout setae in apical third. Wing (
<figureCitation id="134685D2FF8AFF81FF689343FEA12A42" box="[198,288,1616,1642]" captionStart="Fig" captionStartId="135.[189,232,1850,1876]" captionTargetBox="[189,1399,277,1815]" captionTargetId="figure-86@135.[189,1399,265,1815]" captionTargetPageId="135" captionText="Fig. 8. Sclerocoelus Marshall, 1995, left wings, ventral view. A. Sclerocoelus nebulosus sp. nov. B. Sclerocoelus nitidistylus sp. nov. C. Sclerocoelus ocellatus sp. nov. D. Sclerocoelus paranebulosus sp. nov. E. Sclerocoelus pararegularis sp. nov. F. Sclerocoelus parasordipes sp. nov. G. Sclerocoelus plumiseta (Duda, 1925) H. Sclerocoelus punensis sp. nov. I. Sclerocoelus puyensis sp. nov.J. Sclerocoelus rectangularis (Malloch, 1914). Scale bars: AJ=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14980872" httpUri="https://zenodo.org/record/14980872/files/figure.png" pageId="82" pageNumber="83">Fig. 8B</figureCitation>
) slightly infuscate. CS2 0.70.8× CS3. Halter pale brown.
</paragraph>
<paragraph id="8BC29957FF8AFF80FF139381FC3F2E32" blockId="82.[189,1399,1681,2025]" lastBlockId="83.[189,1399,265,539]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<smallCapsWord id="8D240F8BFF8AFF81FF139381FF7E2A82" baselines="1701,1701" box="[189,255,1682,1708]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="82" pageNumber="83">MALE</smallCapsWord>
<smallCapsWord id="8D240F8BFF8AFF81FEAC9384FEEB2A82" baselines="1701" box="[258,362,1687,1706]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="82" pageNumber="83">ABDOMEN</smallCapsWord>
(
<figureCitation id="134685D2FF8AFF81FED99382FD822A83" box="[375,515,1681,1707]" captionStart="Fig" captionStartId="223.[189,232,1860,1886]" captionTargetBox="[189,1399,265,1788]" captionTargetId="figure-66@223.[189,1399,265,1830]" captionTargetPageId="223" captionText="Fig. 96. Sclerocoelus nitidistylus sp. nov. A. Male paratype habitus, lateral view (debu00125692). B. Male paratype terminalia, posterior view. C. Male paratype terminalia, lateral view. Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14981080" httpUri="https://zenodo.org/record/14981080/files/figure.png" pageId="82" pageNumber="83">Figs96BC</figureCitation>
,
<figureCitation id="134685D2FF8AFF81FDA59382FDAD2A83" box="[523,556,1681,1707]" captionStart="Fig" captionStartId="224.[189,232,1858,1884]" captionTargetBox="[197,1395,267,1802]" captionTargetId="figure-88@224.[189,1399,265,1830]" captionTargetPageId="224" captionText="Fig. 97. Sclerocoelus nitidistylus sp. nov., paratype. A. Aedeagus and associated structures, lateral view. B. Male S5 and genital pouch, ventral view. C. Subcercal complex, posterior view. Abbreviations: bp =basiphallus; dp =distiphallus; epi= epiphallus; hyp= hypandrium; pg =postgonite; ph =phallapodeme; sc (subcostal); se = subepandrial sclerite; ss=surstylus. Scale bars: A= 0.13 mm; B=0.25 mm; C= 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14981082" httpUri="https://zenodo.org/record/14981082/files/figure.png" pageId="82" pageNumber="83">97</figureCitation>
).Dark brown, posterior edges of tergites sometimes slightly desclerotized. T25 and S24 uniformly long-setose with large posterolateral setae. S5 rectangular, 1.8× length of S4, laterally sparsely setose with a dark, narrow, ventrally projecting patch of dense setulae flanked by pale setose areas. Anterior flange of S6+7 large, 2.0× as long as wide. Sclerite A dark, unmodified; sclerite B reduced to an indistinct filament arising from posterolateral corner of sclerite A; sclerite C broad but reduced and indistinct, arising from anterolateral corner of sclerite A; sclerite D well sclerotized, Y-shaped and bent distally; sclerite E dark, elongate, and slightly curved; sclerite F large, laterally concave, left side with an elongate apicoventral process; sclerite G small, dark, narrowly separated from sclerite F; ring sclerite small, dark. Epandrium moderate, 0.3 × length of S8, height 1.4× maximum length and 0.7× maximum width, uniformly setose, posteroventral margin deeply emarginate; anal fissure rounded; perianal pads weakly developed. Pseudocercus large, bearing three setae, and fused to posteroventral corners of epandrium; halves of subepandrial sclerite strongly arched, slightly flattened dorsally, and completely separate. Subcercus large, projecting posteriorly, elongate, apically bilobed, anteriorly with a short, dark, basal lobe. Hypandrium with Y-shaped anteromedial apodeme. Surstylus large, subtriangular, laterally concave and shining, inner surface setulose along a broad medial ridge and a dark carinate anterobasal lobe. Postgonite short, apically truncate and expanded, and posterolaterally crenulate. Phallapodeme very large, thick, apex dorsoventrally flattened; basiphallus small, tapered with a distinct, triangular epiphallus; distiphallus largely membranous.
</paragraph>
<paragraph id="8BC29957FF8BFF80FF139751FC6C2F24" blockId="83.[189,1399,578,781]" pageId="83" pageNumber="84">
<smallCapsWord id="8D240F8BFF8BFF80FF139751FE952E72" baselines="597,597" box="[189,276,578,604]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="83" pageNumber="84">FEMALE</smallCapsWord>
<smallCapsWord id="8D240F8BFF8BFF80FEB59754FE022E72" baselines="597" box="[283,387,583,602]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="83" pageNumber="84">ABDOMEN</smallCapsWord>
(
<figureCitation id="134685D2FF8BFF80FE3D9751FE6B2E74" box="[403,490,578,604]" captionStart="Fig" captionStartId="225.[189,232,1927,1953]" captionTargetBox="[215,1379,266,1883]" captionTargetId="figure-49@225.[210,1381,251,1904]" captionTargetPageId="225" captionText="Fig. 98. Sclerocoelus nitidistylus sp. nov., paratype. A. Female terminalia, dorsal view. B. Female terminalia, ventral view. C. Female terminalia, lateral view. D. Spermatheca. Scale bars:AC =0.25 mm; D =0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.14981084" httpUri="https://zenodo.org/record/14981084/files/figure.png" pageId="83" pageNumber="84">Fig. 98</figureCitation>
). T7 broad, slightly emarginate posteromedially; T8 divided into a small, pale dorsal sclerite and a pair of large, dark lateral sclerites, posteroventral corners pointed. Epiproct shield-like, medially pale, apically tapered and setulose. Cercus elongate, slightly tapered with long apical and dorsal setae. S7 broad, rounded, broadly and deeply desclerotized posteromedially with four large posterior setae; S8 a single small but well-developed sclerite. Three spermathecae, bulb spherical, finely striate with a small apical invagination and a deep basal invagination.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8BFF80FF139620FE5E2F54" box="[189,479,819,892]" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BC29957FF8BFF80FF139620FED72F65" blockId="83.[189,342,819,845]" box="[189,342,819,845]" pageId="83" pageNumber="84">
<heading id="D08A2E3BFF8BFF80FF139620FED72F65" bold="true" box="[189,342,819,845]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9094545FF8BFF80FF139620FED72F65" bold="true" box="[189,342,819,845]" pageId="83" pageNumber="84">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8BFF80FF139671FE5E2F54" blockId="83.[189,479,866,892]" box="[189,479,866,892]" pageId="83" pageNumber="84">
Neotropical:
<collectingCountry id="F36AD9C7FF8BFF80FEF99671FE5A2F54" box="[343,475,866,892]" name="Costa Rica" pageId="83" pageNumber="84">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8BFF80FF1396B7FAE32851" pageId="83" pageNumber="84" type="discussion">
<paragraph id="8BC29957FF8BFF80FF1396B7FEAE2F96" blockId="83.[189,303,932,958]" box="[189,303,932,958]" pageId="83" pageNumber="84">
<heading id="D08A2E3BFF8BFF80FF1396B7FEAE2F96" bold="true" box="[189,303,932,958]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9094545FF8BFF80FF1396B7FEAE2F96" bold="true" box="[189,303,932,958]" pageId="83" pageNumber="84">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8BFF80FF1396C1FAE32851" blockId="83.[189,1399,978,1146]" pageId="83" pageNumber="84">
<taxonomicName id="4C7DE2D4FF8BFF80FF1396C1FE5D2FC4" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[189,476,978,1004]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nitidistylus" status="sp. nov.">
<emphasis id="B9094545FF8BFF80FF1396C1FE5D2FC4" box="[189,476,978,1004]" italics="true" pageId="83" pageNumber="84">Sclerocoelus nitidistylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23AF83EFF8BFF80FE4B96C0FDC02FC5" box="[485,577,979,1005]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
is similar to other members of the
<taxonomicName id="4C7DE2D4FF8BFF80FC4796C0FB1D2FC4" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[1001,1180,978,1005]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="dasysternum">
<emphasis id="B9094545FF8BFF80FC4796C0FB1D2FC4" box="[1001,1180,978,1005]" italics="true" pageId="83" pageNumber="84">S. dasysternum</emphasis>
</taxonomicName>
group, but can be readily distinguished by the very shiny outer surface of the surstylus, almost beak-shaped subcercus, and the female S7 with a deep, semicircular posterior desclerotized area.
<taxonomicName id="4C7DE2D4FF8BFF80FB97910AFAD8281B" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[1081,1369,1049,1075]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nitidistylus">
<emphasis id="B9094545FF8BFF80FB97910AFAD8281B" box="[1081,1369,1049,1075]" italics="true" pageId="83" pageNumber="84">Sclerocoelus nitidistylus</emphasis>
</taxonomicName>
is commonly collected in the cloud forests of
<collectingCountry id="F36AD9C7FF8BFF80FD65912FFCD0287E" box="[715,849,1084,1110]" name="Costa Rica" pageId="83" pageNumber="84">Costa Rica</collectingCountry>
, where it is one of four endemic Costa Rican
<taxonomicName id="4C7DE2D4FF8BFF80FF13914CFE2F2851" box="[189,430,1119,1145]" pageId="83" pageNumber="84">
<emphasis id="B9094545FF8BFF80FF13914CFED32851" box="[189,338,1119,1145]" italics="true" pageId="83" pageNumber="84">Sclerocoelus</emphasis>
species
</taxonomicName>
and one of the seven members of the
<taxonomicName id="4C7DE2D4FF8BFF80FCC59173FB9C2851" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[875,1053,1119,1146]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="dasysternum">
<emphasis id="B9094545FF8BFF80FCC59173FB9C2851" box="[875,1053,1119,1146]" italics="true" pageId="83" pageNumber="84">S. dasysternum</emphasis>
</taxonomicName>
group found in
<collectingCountry id="F36AD9C7FF8BFF80FB749173FADF2851" box="[1242,1374,1119,1146]" name="Costa Rica" pageId="83" pageNumber="84">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="188F2BF88ABF5C8A411D2438C52483EF" ID-DOI="10.5852/ejt.2025.979.2803" ID-ISSN="2118-9773" ID-Zenodo-Dep="14980852" ID-ZooBank="F40A49A4-5DCC-491E-9D0A-7A3C2EC6D186" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1741259649306" checkinUser="plazi" docAuthor="Kuwahara, Gregory K., Marshall, Stephen A. &amp; Paiero, Steven M." docDate="2025" docId="03D42841FF8BFF87FD3D91A4FAE32A87" docLanguage="en" docName="EJT.2025.979.1-277.pdf" docOrigin="European Journal of Taxonomy 979" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2803/12807" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sclerocoelus ocellatus Kuwahara, Marshall &amp; Paiero, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="85" masterDocId="FFED5039FFD8FFD3FFAE9513FF812C28" masterDocTitle="A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="84" updateTime="1743778216605" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A987DBE421C732A4DDACD75E682BFE91">A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)</mods:title>
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<mods:namePart id="273A6BA473C99C601B891C047AC0CBFD">Kuwahara, Gregory K.</mods:namePart>
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<mods:affiliation id="D24513D06F05A6B7ADAAA3D9F2AF877A">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
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<mods:namePart id="D1E5965F3E56D5E0769DDB9E0F8F4184">Marshall, Stephen A.</mods:namePart>
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<mods:affiliation id="F387C15B44E314213C338AAA6970BA5B">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
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<mods:namePart id="33A057B5E2A9CC8465981A084DE95C04">Paiero, Steven M.</mods:namePart>
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<mods:affiliation id="0DA45E68C9163B699D75BF51ECDF7310">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
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<mods:date id="31F694F0F70EA0516D4CBCC6BCE851A9">2025</mods:date>
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<paragraph id="8BC29957FF8BFF80FD3D91A4FB8028FA" blockId="83.[361,1226,1207,1304]" box="[659,1025,1207,1234]" pageId="83" pageNumber="84">
<taxonomicName id="4C7DE2D4FF8BFF80FD3D91A4FC1E28FA" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[659,927,1207,1234]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ocellatus" status="sp. nov.">
<emphasis id="B9094545FF8BFF80FD3D91A4FC1E28FA" bold="true" box="[659,927,1207,1234]" italics="true" pageId="83" pageNumber="84">Sclerocoelus ocellatus</emphasis>
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<taxonomicNameLabel id="A23AF83EFF8BFF80FC0891ABFB8028FA" box="[934,1025,1208,1234]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="134685D2FF8BFF80FD1691EDFC972930" box="[696,790,1278,1304]" captionStart="Fig" captionStartId="135.[189,232,1850,1876]" captionTargetBox="[189,1399,277,1815]" captionTargetId="figure-86@135.[189,1399,265,1815]" captionTargetPageId="135" captionText="Fig. 8. Sclerocoelus Marshall, 1995, left wings, ventral view. A. Sclerocoelus nebulosus sp. nov. B. Sclerocoelus nitidistylus sp. nov. C. Sclerocoelus ocellatus sp. nov. D. Sclerocoelus paranebulosus sp. nov. E. Sclerocoelus pararegularis sp. nov. F. Sclerocoelus parasordipes sp. nov. G. Sclerocoelus plumiseta (Duda, 1925) H. Sclerocoelus punensis sp. nov. I. Sclerocoelus puyensis sp. nov.J. Sclerocoelus rectangularis (Malloch, 1914). Scale bars: AJ=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14980872" httpUri="https://zenodo.org/record/14980872/files/figure.png" pageId="83" pageNumber="84">Figs 8C</figureCitation>
,
<figureCitation id="134685D2FF8BFF80FC8D91EDFCFA2930" box="[803,891,1278,1304]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="226.[189,232,1859,1885]" captionStartId-1="227.[189,232,1828,1854]" captionStartId-2="228.[189,232,1928,1954]" captionTargetBox-0="[189,1399,264,1819]" captionTargetBox-1="[202,1366,268,1782]" captionTargetBox-2="[213,1374,268,1892]" captionTargetId-0="figure-73@226.[189,1399,264,1829]" captionTargetId-1="figure-63@227.[189,1399,246,1811]" captionTargetId-2="figure-55@228.[208,1379,251,1904]" captionTargetPageId-0="226" captionTargetPageId-1="227" captionTargetPageId-2="228" captionText-0="Fig. 99. Sclerocoelus ocellatus sp. nov. A. Female paratype habitus, lateral view. B. Male holotype terminalia, posterior view (MNCR). C. Male holotype terminalia, lateral view (MNCR). Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." captionText-1="Fig. 100. Sclerocoelus ocellatus sp. nov., male holotype (MNCR). A. Aedeagus and associated structures, lateral view. B. Male S5 and genital pouch, ventral view.Abbreviations:bp = basiphallus; dp =distiphallus; epi= epiphallus; pg =postgonite; ph =phallapodeme. Scale bars: A=0.13 mm; B=0.25 mm." captionText-2="Fig. 101. Sclerocoelus ocellatus sp. nov., paratype. A. Female terminalia, dorsal view. B. Female terminalia, ventral view. C. Female terminalia, lateral view. D. Spermatheca. Scale bars:AC =0.25 mm; D =0.03 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14981086" figureDoi-1="http://doi.org/10.5281/zenodo.14981088" figureDoi-2="http://doi.org/10.5281/zenodo.14981092" httpUri-0="https://zenodo.org/record/14981086/files/figure.png" httpUri-1="https://zenodo.org/record/14981088/files/figure.png" httpUri-2="https://zenodo.org/record/14981092/files/figure.png" pageId="83" pageNumber="84">99101</figureCitation>
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<emphasis id="B9094545FF8BFF80FF139053FEC52972" bold="true" box="[189,324,1344,1370]" pageId="83" pageNumber="84">Etymology</emphasis>
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This name refers to the greatly reduced eye of this species, smaller than that of any other species of
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<emphasis id="B9094545FF8BFF80FF139081FED32984" box="[189,338,1426,1452]" italics="true" pageId="83" pageNumber="84">Sclerocoelus</emphasis>
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(from the Latin
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<emphasis id="B9094545FF8BFF80FDB39081FD062984" box="[541,647,1426,1452]" italics="true" pageId="83" pageNumber="84">ocellatus</emphasis>
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, meaning having small eyes).
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<emphasis id="B9094545FF8BFF80FF1390C0FE2B29C5" bold="true" box="[189,426,1491,1517]" pageId="83" pageNumber="84">Material examined</emphasis>
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<typeStatus id="54C627F5FF8BFF80FF449311FEDD2A34" box="[234,348,1538,1564]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
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;
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S of Empalme
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leg.;
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;
<elevation id="00507E64FF8BFF80FBAD93BEFB1B2AEF" box="[1027,1178,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="83" pageNumber="84" unit="m" value="3200.0">
<quantity id="4C8534B2FF8BFF80FBAD93BEFBDA2AE0" box="[1027,1115,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="83" pageNumber="84" unit="m" value="3200.0">3200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFC3BF97FF8BFF80FB0993BEFACF2AE0" box="[1191,1358,1709,1736]" pageId="83" pageNumber="84" value="1985-06-01" valueMax="1985-06-07" valueMin="1985-06-01">
<collectingDate id="EF87467FFF8BFF80FB0993BEFACF2AE0" box="[1191,1358,1709,1736]" pageId="83" pageNumber="84" value="1985-06-01" valueMax="1985-06-07" valueMin="1985-06-01">17 Jun. 1985</collectingDate>
</date>
;
<collectorName id="2688FC81FF8BFF80FAF493BDFE8C2AC3" pageId="83" pageNumber="84">H. Goulet</collectorName>
and
<collectorName id="2688FC81FF8BFF80FEE893C2FE3F2AC3" box="[326,446,1745,1771]" pageId="83" pageNumber="84">L. Masner</collectorName>
leg.;
<collectionCode id="ED6C0192FF8BFF80FDAE93C2FDD52AC3" box="[512,596,1745,1771]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="83" pageNumber="84">DEBU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B15930AFF8BFF80FDDB93C3FB4A2AC3" box="[629,1227,1744,1771]" collectingDate="1989-11" collectingDateMax="1989-12-31" collectingDateMin="1989-11" collectionCode="MNCR" collectorName="Cerro de la Muerte &amp; P. Hanson" country="Costa Rica" elevation="2800" location="San Jose" pageId="83" pageNumber="84" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<emphasis id="B9094545FF8BFF80FDDB93C3FD602AC3" bold="true" box="[629,737,1744,1771]" pageId="83" pageNumber="84">
<collectingRegion id="49B957B5FF8BFF80FDDB93C3FD602AC3" box="[629,737,1744,1771]" country="Costa Rica" name="San Jose" pageId="83" pageNumber="84">San José</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFF8BFF80FD5493C2FCA62AC3" box="[762,807,1745,1771]" count="1" pageId="83" pageNumber="84" type="female">1 ♀</specimenCount>
; same data as for holotype;
<collectionCode id="ED6C0192FF8BFF80FBDF93C2FB4A2AC3" box="[1137,1227,1745,1771]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="83" pageNumber="84">MNCR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8BFF87FF139201FC032975" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="description">
<paragraph id="8BC29957FF8BFF80FF139201FECF2B04" blockId="83.[189,334,1810,1836]" box="[189,334,1810,1836]" pageId="83" pageNumber="84">
<heading id="D08A2E3BFF8BFF80FF139201FECF2B04" bold="true" box="[189,334,1810,1836]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9094545FF8BFF80FF139201FECF2B04" bold="true" box="[189,334,1810,1836]" pageId="83" pageNumber="84">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8BFF87FF139251FE232D44" blockId="83.[189,1399,1857,2025]" lastBlockId="84.[189,1399,265,364]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
<smallCapsWord id="8D240F8BFF8BFF80FF139251FF7C2B72" baselines="1877,1877" box="[189,253,1858,1884]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="83" pageNumber="84">BODY</smallCapsWord>
(
<figureCitation id="134685D2FF8BFF80FEBF9252FE032B73" box="[273,386,1857,1883]" captionStart="Fig" captionStartId="226.[189,232,1859,1885]" captionTargetBox="[189,1399,264,1819]" captionTargetId="figure-73@226.[189,1399,264,1829]" captionTargetPageId="226" captionText="Fig. 99. Sclerocoelus ocellatus sp. nov. A. Female paratype habitus, lateral view. B. Male holotype terminalia, posterior view (MNCR). C. Male holotype terminalia, lateral view (MNCR). Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14981086" httpUri="https://zenodo.org/record/14981086/files/figure.png" pageId="83" pageNumber="84">Fig. 99A</figureCitation>
). Length 2.94.0 mm. Head dark brown, very bottom of frons orange; anterior half of gena orangish. Frontal width 2.22.3× frontal height. Three pairs of strong interfrontal bristles surmounting a fine fourth pair; anterior orbital 0.6× length of posterior. Palpus brown. Eye small, greatest height about 1.11.3 × shortest genal height. Thorax dark brown, scutum with paler lateral edges. Two pairs of dorsocentral bristles (anterior pair distinct, 0.6× length of posterior pair) separated by 67 rows of acrostichal setulae. Membrane around prosternum bare. Legs brown, mid femur slightly darker. Fore femur with four large ventral preapical setae. Wing (
<figureCitation id="134685D2FF8CFF87FC65943EFBA72D60" box="[971,1062,301,328]" captionStart="Fig" captionStartId="135.[189,232,1850,1876]" captionTargetBox="[189,1399,277,1815]" captionTargetId="figure-86@135.[189,1399,265,1815]" captionTargetPageId="135" captionText="Fig. 8. Sclerocoelus Marshall, 1995, left wings, ventral view. A. Sclerocoelus nebulosus sp. nov. B. Sclerocoelus nitidistylus sp. nov. C. Sclerocoelus ocellatus sp. nov. D. Sclerocoelus paranebulosus sp. nov. E. Sclerocoelus pararegularis sp. nov. F. Sclerocoelus parasordipes sp. nov. G. Sclerocoelus plumiseta (Duda, 1925) H. Sclerocoelus punensis sp. nov. I. Sclerocoelus puyensis sp. nov.J. Sclerocoelus rectangularis (Malloch, 1914). Scale bars: AJ=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14980872" httpUri="https://zenodo.org/record/14980872/files/figure.png" pageId="84" pageNumber="85">Fig. 8C</figureCitation>
) infuscate. CS2 subequal to CS3. Halter brown.
</paragraph>
<paragraph id="8BC29957FF8CFF87FF139486FAF62817" blockId="84.[189,1399,405,1087]" pageId="84" pageNumber="85">
<smallCapsWord id="8D240F8BFF8CFF87FF139486FF7E2D85" baselines="424,424" box="[189,255,405,431]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="84" pageNumber="85">MALE</smallCapsWord>
<smallCapsWord id="8D240F8BFF8CFF87FEA19489FEF62D85" baselines="424" box="[271,375,410,429]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="84" pageNumber="85">ABDOMEN</smallCapsWord>
(
<figureCitation id="134685D2FF8CFF87FE3F9486FDA82D87" box="[401,553,405,431]" captionStart="Fig" captionStartId="226.[189,232,1859,1885]" captionTargetBox="[189,1399,264,1819]" captionTargetId="figure-73@226.[189,1399,264,1829]" captionTargetPageId="226" captionText="Fig. 99. Sclerocoelus ocellatus sp. nov. A. Female paratype habitus, lateral view. B. Male holotype terminalia, posterior view (MNCR). C. Male holotype terminalia, lateral view (MNCR). Abbreviations: epa= epandrium; pc =pseudocercus; sc =subcercus; se =subepandrial sclerite; ss= surstylus. Scale bars: A=1.0 mm; BC =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14981086" httpUri="https://zenodo.org/record/14981086/files/figure.png" pageId="84" pageNumber="85">Figs 99BC</figureCitation>
,
<figureCitation id="134685D2FF8CFF87FD909486FDEF2D87" box="[574,622,405,431]" captionStart="Fig" captionStartId="227.[189,232,1828,1854]" captionTargetBox="[202,1366,268,1782]" captionTargetId="figure-63@227.[189,1399,246,1811]" captionTargetPageId="227" captionText="Fig. 100. Sclerocoelus ocellatus sp. nov., male holotype (MNCR). A. Aedeagus and associated structures, lateral view. B. Male S5 and genital pouch, ventral view.Abbreviations:bp = basiphallus; dp =distiphallus; epi= epiphallus; pg =postgonite; ph =phallapodeme. Scale bars: A=0.13 mm; B=0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14981088" httpUri="https://zenodo.org/record/14981088/files/figure.png" pageId="84" pageNumber="85">100</figureCitation>
). Dark brown, posterior edges of tergites sometimes slightly desclerotized. T25 and S24 uniformly long-setose with large posterolateral setae. S5 rectangular, 1.2× length of S4, sparely setose with a dark, elongate, posteromedial patch of dense setulae flanked by long-setose, pale patches. Anterior flange of S6+7 rectangular, 1.0 × as long as wide. Sclerite A pale, setulose, and weakly fused to S6+7; sclerite B dark and strongly arched into genital pouch; sclerite C apparently absent; sclerite D dark, elongate, expanded where it articulates with left side of posteromedial lobe of S5; sclerite E dark, elongate, closely associated with sclerite D, together forming a V-shaped; sclerites F and G fused, sclerite F with an elongate posteroapical process on left side, sclerite G bulbous and setulose; ring sclerite well developed but thin. Epandrium moderate, 0.6× length of S8, height 1.4× maximum length and 0.8× maximum width, uniformly long-setose; perianal pads bulging but membranous. Pseudocercus small, fused to posterolateral corner of epandrium and overlapping base of subcercus, bearing 3 setae; halves of subepandrial sclerite strongly arched with inner arch broadened. Subcercus large, setulose, flattened posteriorly with a triangular posteroventral lobe curving inwards. Hypandrium with sinuate, evenly tapered anteromedial apodeme. Surstylus large, 1.4 × as long as deep, bulbous, long-setose along inner surface with a small, triangular, inner anterobasal lobe. Postgonite short, sinuate, apically truncate with a slight constriction about midlength and a small, tooth-like, preapical anterior lobe. Phallapodeme large, slightly sinuate, with apex dorsoventrally flattened; basiphallus large, curved with an elongate, beak-like epiphallus and expanded distally along dorsal margin of distiphallus; distiphallus short, largely membranous with an elongate dorsal sclerite and a pair of short lateral sclerites.
</paragraph>
<paragraph id="8BC29957FF8CFF87FF13917BFC032975" blockId="84.[189,1399,1128,1373]" pageId="84" pageNumber="85">
<smallCapsWord id="8D240F8BFF8CFF87FF13917BFE9528A8" baselines="1147,1147" box="[189,276,1128,1154]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="84" pageNumber="85">FEMALE</smallCapsWord>
<smallCapsWord id="8D240F8BFF8CFF87FEB0917EFE0728A8" baselines="1147" box="[286,390,1133,1152]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="84" pageNumber="85">ABDOMEN</smallCapsWord>
(
<figureCitation id="134685D2FF8CFF87FE34917BFD8228AA" box="[410,515,1128,1154]" captionStart="Fig" captionStartId="228.[189,232,1928,1954]" captionTargetBox="[213,1374,268,1892]" captionTargetId="figure-55@228.[208,1379,251,1904]" captionTargetPageId="228" captionText="Fig. 101. Sclerocoelus ocellatus sp. nov., paratype. A. Female terminalia, dorsal view. B. Female terminalia, ventral view. C. Female terminalia, lateral view. D. Spermatheca. Scale bars:AC =0.25 mm; D =0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.14981092" httpUri="https://zenodo.org/record/14981092/files/figure.png" pageId="84" pageNumber="85">Fig. 101</figureCitation>
). T7 broad, simple; T8 divided into a large, pale, dorsal sclerite and two large, dark, lateral sclerites, posterolateral corners expanded and pointed. Epiproct large, subtriangular, entirely sparsely setulose with an additional posteromedial seta near posterior margin. Cercus elongate, apically rounded with long apical and dorsal setae. S7 broad, broadly desclerotized posteriorly and medially with four large posterior setae; S8 entirely membranous. Hypoproct with entirely densely setulae. Three spermathecae, bulb stout, subspherical, finely striate with shallow basal and subapical invaginations, both with a small, finger-like, central process.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8CFF87FF139096FE5E29E7" box="[189,479,1413,1487]" pageId="84" pageNumber="85" type="distribution">
<paragraph id="8BC29957FF8CFF87FF139096FED729B7" blockId="84.[189,342,1413,1439]" box="[189,342,1413,1439]" pageId="84" pageNumber="85">
<heading id="D08A2E3BFF8CFF87FF139096FED729B7" bold="true" box="[189,342,1413,1439]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
<emphasis id="B9094545FF8CFF87FF139096FED729B7" bold="true" box="[189,342,1413,1439]" pageId="84" pageNumber="85">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8CFF87FF1390A6FE5E29E7" blockId="84.[189,479,1461,1487]" box="[189,479,1461,1487]" pageId="84" pageNumber="85">
Neotropical:
<collectingCountry id="F36AD9C7FF8CFF87FEF990A6FE5A29E7" box="[343,475,1461,1487]" name="Costa Rica" pageId="84" pageNumber="85">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFF8CFF87FF1390EBFAE32A87" pageId="84" pageNumber="85" type="discussion">
<paragraph id="8BC29957FF8CFF87FF1390EBFEAE2A3A" blockId="84.[189,303,1528,1554]" box="[189,303,1528,1554]" pageId="84" pageNumber="85">
<heading id="D08A2E3BFF8CFF87FF1390EBFEAE2A3A" bold="true" box="[189,303,1528,1554]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
<emphasis id="B9094545FF8CFF87FF1390EBFEAE2A3A" bold="true" box="[189,303,1528,1554]" pageId="84" pageNumber="85">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FF8CFF87FF139334FAE32A87" blockId="84.[189,1399,1575,1711]" pageId="84" pageNumber="85">
<taxonomicName id="4C7DE2D4FF8CFF87FF139334FE422A69" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[189,451,1575,1601]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellatus" status="sp. nov.">
<emphasis id="B9094545FF8CFF87FF139334FE422A69" box="[189,451,1575,1601]" italics="true" pageId="84" pageNumber="85">Sclerocoelus ocellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23AF83EFF8CFF87FE67933BFDA22A6A" box="[457,547,1576,1602]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguished from all other species of
<taxonomicName id="4C7DE2D4FF8CFF87FBFB9334FB6B2A69" box="[1109,1258,1575,1601]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B9094545FF8CFF87FBFB9334FB6B2A69" box="[1109,1258,1575,1601]" italics="true" pageId="84" pageNumber="85">Sclerocoelus</emphasis>
</taxonomicName>
by its small eye (1.11.3× genal height).
<taxonomicName id="4C7DE2D4FF8CFF87FDB4935FFCA02A4E" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[538,801,1612,1638]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellatus">
<emphasis id="B9094545FF8CFF87FDB4935FFCA02A4E" box="[538,801,1612,1638]" italics="true" pageId="84" pageNumber="85">Sclerocoelus ocellatus</emphasis>
</taxonomicName>
is known from a few specimens collected in high elevation forests in the mountains south of
<collectingRegion id="49B957B5FF8CFF87FD179362FCA32AA3" box="[697,802,1649,1675]" country="Costa Rica" name="San Jose" pageId="84" pageNumber="85">San Jose</collectingRegion>
,
<collectingCountry id="F36AD9C7FF8CFF87FC809362FC332AA2" box="[814,946,1648,1675]" name="Costa Rica" pageId="84" pageNumber="85">Costa Rica</collectingCountry>
. It is one of four endemic Costa Rican
<taxonomicName id="4C7DE2D4FF8CFF87FF139386FE2F2A87" box="[189,430,1685,1711]" pageId="84" pageNumber="85">
<emphasis id="B9094545FF8CFF87FF139386FED32A87" box="[189,338,1685,1711]" italics="true" pageId="84" pageNumber="85">Sclerocoelus</emphasis>
species
</taxonomicName>
and one of the seven members of the
<taxonomicName id="4C7DE2D4FF8CFF87FCC59386FB9C2A87" authorityName="Kuwahara &amp; Marshall &amp; Paiero" authorityYear="2025" box="[875,1053,1685,1711]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="dasysternum">
<emphasis id="B9094545FF8CFF87FCC59386FB9C2A87" box="[875,1053,1685,1711]" italics="true" pageId="84" pageNumber="85">S. dasysternum</emphasis>
</taxonomicName>
group found in
<collectingCountry id="F36AD9C7FF8CFF87FB749386FADF2A87" box="[1242,1374,1685,1711]" name="Costa Rica" pageId="84" pageNumber="85">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="37F15A31D8176E06C940382B03AE7C7B" ID-DOI="10.5852/ejt.2025.979.2803" ID-ISSN="2118-9773" ID-Zenodo-Dep="14980852" ID-ZooBank="F40A49A4-5DCC-491E-9D0A-7A3C2EC6D186" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1741259649306" checkinUser="plazi" docAuthor="Kuwahara, Gregory K., Marshall, Stephen A. &amp; Paiero, Steven M." docDate="2025" docId="03D42841FFB4FFBEFD8C9363FD6D2FF3" docLanguage="en" docName="EJT.2025.979.1-277.pdf" docOrigin="European Journal of Taxonomy 979" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2803/12807" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sclerocoelus subbrevipennis" docType="treatment" docVersion="7" lastPageNumber="110" masterDocId="FFED5039FFD8FFD3FFAE9513FF812C28" masterDocTitle="A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="109" updateTime="1743778216605" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="3F3A12A831694A6EC699E2DAB4D9450A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE2F2772CF830C475E29FE1B0660A12F">
<mods:title id="F6A10FE5A85B5DC75ED7B94B469F8BE8">A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)</mods:title>
</mods:titleInfo>
<mods:name id="D73EB79D5B1413C81C8F1350CE020590" type="personal">
<mods:role id="341387527D234DEC393437F09A875C5D">
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<mods:namePart id="61142680A6ACCB6C3BC3AE1AEBDF010B">Kuwahara, Gregory K.</mods:namePart>
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<mods:affiliation id="EEC1D0591855C353AAC90F06903986B5">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="EBA352220A828AF8CE7FA990E0E5CBDE" type="email">gkuwahar@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="63046520A646316BE5C889C786276116" type="personal">
<mods:role id="921CFACEE61485B789BADC4FEF3A58E3">
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</mods:role>
<mods:namePart id="2341983030B2AB05CFC326069EEAD6ED">Marshall, Stephen A.</mods:namePart>
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<mods:affiliation id="A4F4A284BB91D570355C518726A4A4B8">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="00655B9335F8C37364324A8224F9ED43" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
</mods:name>
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<mods:role id="4C713D4618B5A346757918003121B2BE">
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</mods:role>
<mods:namePart id="6EC597D9393D61145691576ABF792859">Paiero, Steven M.</mods:namePart>
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<mods:affiliation id="FAE8ECB27130F3A2E58C097198669847">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="E77EB5241196EDDD762DAE8139798382" type="email">paieros@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="258F5FC45BB7A3DFA15082CEC7E8B12C">European Journal of Taxonomy</mods:title>
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<mods:date id="E05A25F182A0FD92B3B93450B88F6060">2025</mods:date>
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<treatment id="03D42841FFB4FFBEFD8C9363FD6D2FF3" LSID="urn:lsid:plazi:treatment:03D42841FFB4FFBEFD8C9363FD6D2FF3" httpUri="http://treatment.plazi.org/id/03D42841FFB4FFBEFD8C9363FD6D2FF3" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
<subSubSection id="C367CADCFFB4FFBFFD8C9363FB902AA4" box="[546,1041,1648,1676]" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="8BC29957FFB4FFBFFD8C9363FB902AA4" blockId="108.[546,1041,1648,1676]" box="[546,1041,1648,1676]" pageId="108" pageNumber="109">
<heading id="D08A2E3BFFB4FFBFFD8C9363FB902AA4" box="[546,1041,1648,1676]" centered="true" fontSize="11" level="2" pageId="108" pageNumber="109" reason="2">
<taxonomicName id="4C7DE2D4FFB4FFBFFD8C9363FB902AA4" authority="(Frey, 1954)" baseAuthorityName="Frey" baseAuthorityYear="1954" box="[546,1041,1648,1676]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="subbrevipennis">
<emphasis id="B9094545FFB4FFBFFD8C9363FCFB2AA3" bold="true" box="[546,890,1648,1675]" italics="true" pageId="108" pageNumber="109">Sclerocoelus subbrevipennis</emphasis>
(
<bibRefCitation id="EFECE4A6FFB4FFBFFC279362FB882AA4" author="Frey R." box="[905,1033,1649,1676]" pageId="108" pageNumber="109" pagination="1 - 55" refId="ref114419" refString="Frey R. 1954. Diptera Brachycera and Sciaridae von Tristan da Cunha (with a contribution by J. Bequaert). Results of the Norwegian Scientific Expedition to Tristan da Cunha 1937 - 1938, Oslo 4 (26): 1-55." type="journal article" year="1954">Frey, 1954</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFFB4FFBFFF1393A0FC952B19" pageId="108" pageNumber="109" type="reference_group">
<paragraph id="8BC29957FFB4FFBFFF1393A0FD1A2AE5" blockId="108.[189,667,1714,1741]" box="[189,667,1714,1741]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="AB6DBE79FFB4FFBFFF1393A0FD1A2AE5" box="[189,667,1714,1741]" pageId="108" pageNumber="109">
<treatmentCitation id="0ADCBF46FFB4FFBFFF1393A0FD162AE5" author="Frey R." box="[189,663,1714,1741]" page="35" pageId="108" pageNumber="109" year="1954">
<taxonomicName id="4C7DE2D4FFB4FFBFFF1393A0FD162AE5" ID-CoL="8CMCB" authority="Frey, 1954: 35" authorityName="Frey" authorityPageNumber="35" authorityYear="1954" box="[189,663,1714,1741]" class="Insecta" family="Sphaeroceridae" genus="Limosina" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="subbrevipennis">
<emphasis id="B9094545FFB4FFBFFF1393A0FE632AE4" box="[189,482,1714,1740]" italics="true" pageId="108" pageNumber="109">Limosina subbrevipennis</emphasis>
<bibRefCitation id="EFECE4A6FFB4FFBFFE4793A0FD162AE5" author="Frey R." box="[489,663,1714,1741]" pageId="108" pageNumber="109" pagination="1 - 55" refId="ref114419" refString="Frey R. 1954. Diptera Brachycera and Sciaridae von Tristan da Cunha (with a contribution by J. Bequaert). Results of the Norwegian Scientific Expedition to Tristan da Cunha 1937 - 1938, Oslo 4 (26): 1-55." type="journal article" year="1954">Frey, 1954: 35</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC29957FFB4FFBFFF1393E6FCFE2B26" blockId="108.[189,895,1780,1842]" box="[189,895,1780,1806]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="AB6DBE79FFB4FFBFFF1393E6FCFE2B26" box="[189,895,1780,1806]" pageId="108" pageNumber="109">
<taxonomicName id="4C7DE2D4FFB4FFBFFF1393E6FDF52B26" box="[189,628,1780,1806]" class="Insecta" family="Sphaeroceridae" genus="Leptocera" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="subbrevipennis" subGenus="Limosina">
<emphasis id="B9094545FFB4FFBFFF1393E6FEB52B26" box="[189,308,1781,1806]" italics="true" pageId="108" pageNumber="109">Leptocera</emphasis>
(
<emphasis id="B9094545FFB4FFBFFEEB93E6FE332B26" box="[325,434,1781,1806]" italics="true" pageId="108" pageNumber="109">Limosina</emphasis>
)
<emphasis id="B9094545FFB4FFBFFE6D93E7FDF52B26" box="[451,628,1780,1806]" italics="true" pageId="108" pageNumber="109">subbrevipennis</emphasis>
</taxonomicName>
<treatmentCitation id="0ADCBF46FFB4FFBFFD3F93E7FCFA2B26" author="Richards O. W." box="[657,891,1780,1806]" page="620" pageId="108" pageNumber="109" year="1980">
<bibRefCitation id="EFECE4A6FFB4FFBFFD3F93E7FCFA2B26" author="Richards O. W." box="[657,891,1780,1806]" pageId="108" pageNumber="109" refId="ref115505" refString="Richards O. W. 1980. 57. Family Sphaeroceridae. In: Crosskey R. W. (ed.) Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London." type="book" year="1980">Richards 1980: 620</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC29957FFB4FFBFFF139204FC952B19" blockId="108.[189,895,1780,1842]" box="[189,788,1815,1842]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="AB6DBE79FFB4FFBFFF139204FC952B19" box="[189,788,1815,1842]" pageId="108" pageNumber="109">
<taxonomicName id="4C7DE2D4FFB4FFBFFF139204FD8B2B19" baseAuthorityName="Frey" baseAuthorityYear="1954" box="[189,522,1815,1841]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="subbrevipennis">
<emphasis id="B9094545FFB4FFBFFF139204FD8B2B19" box="[189,522,1815,1841]" italics="true" pageId="108" pageNumber="109">Sclerocoelus subbrevipennis</emphasis>
</taxonomicName>
<treatmentCitation id="0ADCBF46FFB4FFBFFD899204FC912B19" author="Marshall S. A." box="[551,784,1815,1841]" page="104" pageId="108" pageNumber="109" year="1997">
<bibRefCitation id="EFECE4A6FFB4FFBFFD899204FC912B19" author="Marshall S. A." box="[551,784,1815,1841]" pageId="108" pageNumber="109" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall 1997: 104</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFFB4FFBEFF13924BFD502D58" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="diagnosis">
<paragraph id="8BC29957FFB4FFBFFF13924BFEB72B5A" blockId="108.[189,310,1880,1906]" box="[189,310,1880,1906]" pageId="108" pageNumber="109">
<heading id="D08A2E3BFFB4FFBFFF13924BFEB72B5A" bold="true" box="[189,310,1880,1906]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B9094545FFB4FFBFFF13924BFEB72B5A" bold="true" box="[189,310,1880,1906]" pageId="108" pageNumber="109">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FFB4FFBEFF139294FD502D58" blockId="108.[189,1399,1927,2024]" lastBlockId="109.[189,1398,265,368]" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
Body length
<quantity id="4C8534B2FFB4FFBFFEF69294FE362B8A" box="[344,439,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="108" pageNumber="109" unit="mm" value="2.2">2.2 mm</quantity>
. Membrane around prosternum bare. Male S5 rectangular, posteromedially pale, bilobed, and covered in short, flattened setulae (cf.
<bibRefCitation id="EFECE4A6FFB4FFBFFC9892B9FC682BED" author="Marshall S. A." box="[822,1001,1962,1989]" pageId="108" pageNumber="109" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall 1997</bibRefCitation>
: figs 54, 56); perianal pads well developed and dark; surstylus subtriangular with a rectangular anteroventral lobe bearing a posteroventral patch of thickened setae (cf.
<bibRefCitation id="EFECE4A6FFB5FFBEFDB8941AFD472D0B" author="Marshall S. A." box="[534,710,265,291]" pageId="109" pageNumber="110" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall 1997</bibRefCitation>
: figs 5253); postgonite very narrow and strongly sinuate (cf.
<bibRefCitation id="EFECE4A6FFB5FFBEFF5E943CFE232D61" author="Marshall S. A." box="[240,418,303,329]" pageId="109" pageNumber="110" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall 1997</bibRefCitation>
: fig. 55). Female S8 hourglass-shaped, lateral margins curved outwards into two prominent ridges (cf.
<bibRefCitation id="EFECE4A6FFB5FFBEFE159446FDEB2D47" author="Marshall S. A." box="[443,618,341,367]" pageId="109" pageNumber="110" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall 1997</bibRefCitation>
: fig. 59).
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFFB5FFBEFF13948AFD652E23" pageId="109" pageNumber="110" type="materials_examined">
<paragraph id="8BC29957FFB5FFBEFF13948AFE2B2D9B" blockId="109.[189,426,409,435]" box="[189,426,409,435]" pageId="109" pageNumber="110">
<heading id="D08A2E3BFFB5FFBEFF13948AFE2B2D9B" bold="true" box="[189,426,409,435]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B9094545FFB5FFBEFF13948AFE2B2D9B" bold="true" box="[189,426,409,435]" pageId="109" pageNumber="110">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FFB5FFBEFF1394D8FD652E23" blockId="109.[189,1399,457,523]" pageId="109" pageNumber="110">
<materialsCitation id="3B15930AFFB5FFBEFF1394D8FD602E23" collectedFrom="albatross nest" collectingDate="1989-10-14" collectionCode="DEBU" collectorName="H. M. Barber" country="Saint Helena, Ascension and Tristan da Cunha" latitude="-37.033333" location="SW of Denstone Hill" longLatPrecision="1183" longitude="-12.2" pageId="109" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Inaccessible Island">
<collectingCountry id="F36AD9C7FFB5FFBEFF1394D8FE552DCD" box="[189,468,459,485]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="109" pageNumber="110">TRISTAN DA CUNHA</collectingCountry>
<emphasis id="B9094545FFB5FFBEFE5E94D8FD5A2DCD" bold="true" box="[496,731,459,485]" pageId="109" pageNumber="110">
<collectingRegion id="49B957B5FFB5FFBEFE5E94D8FD5A2DCD" box="[496,731,459,485]" pageId="109" pageNumber="110">Inaccessible Island</collectingRegion>
</emphasis>
<specimenCount id="9D7B52DEFFB5FFBEFD5D94D8FCA32DCD" box="[755,802,459,485]" count="1" pageId="109" pageNumber="110" type="male">1 ♂</specimenCount>
;
<location id="8EA2CF8CFFB5FFBEFC8094D8FBA72DCD" LSID="urn:lsid:plazi:treatment:03D42841FFB4FFBEFD8C9363FD6D2FF3:8EA2CF8CFFB5FFBEFC8094D8FBA72DCD" box="[814,1062,459,485]" country="Saint Helena, Ascension and Tristan da Cunha" latitude="-37.033333" longLatPrecision="1183" longitude="-12.2" name="SW of Denstone Hill" pageId="109" pageNumber="110" stateProvince="Inaccessible Island">SW of Denstone Hill</location>
;
<geoCoordinate id="EE49FF90FFB5FFBEFB9C94DAFB162DCD" box="[1074,1175,457,485]" degrees="37" direction="south" minutes="02" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-37.033333">37°02 S</geoCoordinate>
,
<geoCoordinate id="EE49FF90FFB5FFBEFB0A94DAFA942DCD" box="[1188,1301,457,485]" degrees="12" direction="west" minutes="12" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-12.2">12°12 W</geoCoordinate>
;
<date id="FFC3BF97FFB5FFBEFA8F94D8FF7A2E23" pageId="109" pageNumber="110" value="1989-10-14">
<collectingDate id="EF87467FFFB5FFBEFA8F94D8FF7A2E23" pageId="109" pageNumber="110" value="1989-10-14">14 Oct. 1989</collectingDate>
</date>
;
<collectorName id="2688FC81FFB5FFBEFEA994E2FE1D2E23" box="[263,412,497,523]" pageId="109" pageNumber="110">H.M. Barber</collectorName>
leg.;
<collectedFrom id="3861DAAAFFB5FFBEFE7094E2FD002E23" box="[478,641,497,523]" pageId="109" pageNumber="110">albatross nest</collectedFrom>
;
<collectionCode id="ED6C0192FFB5FFBEFD2394E2FD602E23" box="[653,737,497,523]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="109" pageNumber="110">DEBU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFFB5FFBEFF139726FB9E2EA9" box="[189,1055,565,641]" pageId="109" pageNumber="110" type="distribution">
<paragraph id="8BC29957FFB5FFBEFF139726FED72E67" blockId="109.[189,342,565,591]" box="[189,342,565,591]" pageId="109" pageNumber="110">
<heading id="D08A2E3BFFB5FFBEFF139726FED72E67" bold="true" box="[189,342,565,591]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B9094545FFB5FFBEFF139726FED72E67" bold="true" box="[189,342,565,591]" pageId="109" pageNumber="110">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FFB5FFBEFF139774FB9E2EA9" blockId="109.[189,1055,615,641]" box="[189,1055,615,641]" pageId="109" pageNumber="110">
South Atlantic:
<collectingCountry id="F36AD9C7FFB5FFBEFEDB9774FE462EA9" box="[373,455,615,641]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="109" pageNumber="110">Tristan</collectingCountry>
de
<collectingCountry id="F36AD9C7FFB5FFBEFE5F9774FDBF2EA9" box="[497,574,615,641]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="109" pageNumber="110">Cunha</collectingCountry>
(Inaccessible Island, Nightingale Island)
</paragraph>
</subSubSection>
<subSubSection id="C367CADCFFB5FFBEFF1397B8FD6D2FF3" pageId="109" pageNumber="110" type="discussion">
<paragraph id="8BC29957FFB5FFBEFF1397B8FEAE2EED" blockId="109.[189,303,683,709]" box="[189,303,683,709]" pageId="109" pageNumber="110">
<heading id="D08A2E3BFFB5FFBEFF1397B8FEAE2EED" bold="true" box="[189,303,683,709]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B9094545FFB5FFBEFF1397B8FEAE2EED" bold="true" box="[189,303,683,709]" pageId="109" pageNumber="110">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC29957FFB5FFBEFF1397CFFD6D2FF3" blockId="109.[189,1399,732,987]" pageId="109" pageNumber="110">
<taxonomicName id="4C7DE2D4FFB5FFBEFF1397CFFDE32EDE" authority="(Frey)" baseAuthorityName="Frey" baseAuthorityYear="1954" box="[189,610,732,758]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="subbrevipennis">
<emphasis id="B9094545FFB5FFBEFF1397CFFD8F2EDE" box="[189,526,732,758]" italics="true" pageId="109" pageNumber="110">Sclerocoelus subbrevipennis</emphasis>
(Frey)
</taxonomicName>
is the only unequivocal
<taxonomicName id="4C7DE2D4FFB5FFBEFC3997CFFB0D2EDE" box="[919,1164,732,758]" pageId="109" pageNumber="110">
<emphasis id="B9094545FFB5FFBEFC3997CFFBAD2EDE" box="[919,1068,732,758]" italics="true" pageId="109" pageNumber="110">Sclerocoelus</emphasis>
species
</taxonomicName>
with a distribution outside the New World, and one of only four species found outside the Neotropical region. The close relationship between
<taxonomicName id="4C7DE2D4FFB5FFBEFE6E963AFD132F6A" baseAuthorityName="Frey" baseAuthorityYear="1954" box="[448,658,808,835]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="subbrevipennis">
<emphasis id="B9094545FFB5FFBEFE6E963AFD132F6A" box="[448,658,808,835]" italics="true" pageId="109" pageNumber="110">S. subbrevipennis</emphasis>
</taxonomicName>
and Neotropical members of the
<taxonomicName id="4C7DE2D4FFB5FFBEFB9C963AFB6F2F6A" authorityName="Marshall" authorityYear="1997" box="[1074,1262,808,835]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="B9094545FFB5FFBEFB9C963AFB6F2F6A" box="[1074,1262,808,835]" italics="true" pageId="109" pageNumber="110">S. galapagensis</emphasis>
</taxonomicName>
group was documented by
<bibRefCitation id="EFECE4A6FFB5FFBEFED1965DFDC02F41" author="Marshall S. A." box="[383,577,846,873]" pageId="109" pageNumber="110" pagination="97 - 115" refId="ref114969" refString="Marshall S. A. 1997. A revision of the Sclerocoelus galapagensis group (Diptera: Sphaeroceridae: Limosininae). Insecta Mundi 11 (2): 97-115." type="journal article" year="1997">Marshall (1997)</bibRefCitation>
and we are confident that the species is correctly placed. Although undoubtedly a
<taxonomicName id="4C7DE2D4FFB5FFBEFEDE9666FD842FA7" box="[368,517,885,911]" class="Insecta" family="Sphaeroceridae" genus="Sclerocoelus" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B9094545FFB5FFBEFEDE9666FD842FA7" box="[368,517,885,911]" italics="true" pageId="109" pageNumber="110">Sclerocoelus</emphasis>
</taxonomicName>
, it is clearly distinct from all other species we have examined so there is no evidence that it is secondarily introduced to
<collectingCountry id="F36AD9C7FFB5FFBEFD519688FC502F9D" box="[767,977,923,949]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="109" pageNumber="110">Tristan da Cunha</collectingCountry>
. Some of the
<typeStatus id="54C627F5FFB5FFBEFBD29688FB2E2F9D" box="[1148,1199,923,949]" pageId="109" pageNumber="110">type</typeStatus>
specimens were collected in bird nests and in rotting vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>

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leg.; along stream,
leg.;
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<mods:namePart id="47687D699277FF5634E69FDF93F86784">Echternacht, Lívia</mods:namePart>
</mods:name>
<mods:name id="FA4B681B44CE6033F857C4E2B3BEA828" type="personal">
<mods:role id="66BEB21C61750A73B48EE2A200D49881">
<mods:roleTerm id="7B990FB376FAC8576022520AC9F5ED26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6DCCF83BCEE4000053E6856F5C5D31F4">Hensold, Nancy</mods:namePart>
</mods:name>
<mods:name id="FFE747CE27DE4F1113B2C02865FDCE84" type="personal">
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</mods:role>
<mods:namePart id="BCDAF47B9826B95CFD6D6C2B70100AB5">Sano, Paulo Takeo</mods:namePart>
</mods:name>
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<mods:titleInfo id="78EEA17C4CED9284D4878C6CB19A2C35">
<mods:title id="3B02BB8BB3042635B8A10FC69C875250">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="C4359CB01214E8DE54811E686066733E">
<mods:date id="8DBABF3F7B3CC16B488AB42600B7AA62">2015</mods:date>
<mods:detail id="42311A5694FD42D3FA4A4ACD40D0B0D5" type="pubDate">
<mods:number id="28DD6718E4F70CB5BCCFA6FB03B3CA6E">2015-06-09</mods:number>
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<mods:detail id="8A072DB914F3DB702D7E40D7D22430CA" type="volume">
<mods:number id="4EB74EC0A72B11BAC7A08AB308161D22">212</mods:number>
</mods:detail>
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<mods:number id="B51284518FDD3528C16E406B80CD2F40">4</mods:number>
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<mods:start id="E331A976F38D0ADE8FA8A94301A6F842">271</mods:start>
<mods:end id="25007406883AF6F3E00B49798125F2D6">282</mods:end>
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<mods:location id="4934DC210EA5CDCCF7209475DE2ECDEB">
<mods:url id="8B138B2AD61B8B028AE93DB2BE32F1D2">http://dx.doi.org/10.11646/phytotaxa.212.4.3</mods:url>
</mods:location>
<mods:classification id="31DA73D1E1EFD0888B0EB7556FCAA7D6">journal article</mods:classification>
<mods:identifier id="897775515DE4A5D39DC4CB49337566CF" type="CLB-Dataset">308979</mods:identifier>
<mods:identifier id="EFA9E06E53087B97A9B9CC115C1C97EE" type="DOI">10.11646/phytotaxa.212.4.3</mods:identifier>
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<treatment id="3E7DF728FFC1A31CA1FE35A67AA01856" ID-DOI="http://doi.org/10.5281/zenodo.15149343" ID-Zenodo-Dep="15149343" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856" httpUri="http://treatment.plazi.org/id/3E7DF728FFC1A31CA1FE35A67AA01856" lastPageId="9" lastPageNumber="280" pageId="6" pageNumber="277">
<subSubSection id="FECE15B5FFC1A313A1FE35A6780E18EE" box="[136,816,1706,1733]" pageId="6" pageNumber="277" type="nomenclature">
<paragraph id="B66B463EFFC1A313A1FE35A6780E18EE" blockId="6.[136,1451,1706,1841]" box="[136,816,1706,1733]" pageId="6" pageNumber="277">
<taxonomicName id="71D43DBDFFC1A313A1FE35A679EA18EE" ID-CoL="53WY8" authority="M.T.C.Watan. &amp; Echternacht" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[136,724,1706,1733]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="vittatus" status="sp. nov.">
<emphasis id="84A09A2CFFC1A313A1FE35A67AB618EF" bold="true" box="[136,392,1706,1733]" italics="true" pageId="6" pageNumber="277">Syngonanthus vittatus</emphasis>
M.T.C.Watan. &amp; Echternacht
</taxonomicName>
,
<emphasis id="84A09A2CFFC1A313A3AB35A0780E18EE" box="[733,816,1708,1733]" italics="true" pageId="6" pageNumber="277">
<taxonomicNameLabel id="9F932757FFC1A313A3AB35A0780E18EE" box="[733,816,1708,1733]" pageId="6" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A24235A0799F191A" pageId="6" pageNumber="277" type="materials_examined">
<paragraph id="B66B463EFFC1A313A24235A0799F191A" blockId="6.[136,1451,1706,1841]" pageId="6" pageNumber="277">
Type:—
<materialsCitation id="06BC4C63FFC1A313A2F935A07A0B191A" collectingDate="2012-07-19" collectionCode="SPF" collectorName="M. Watanabe &amp; W. Watanabe" country="Brazil" latitude="-14.106029" location="Chapada dos Veadeiros, estrada Alto Paraiso de Goias - Teresina de Goias, Fazenda Agua Fria" longLatPrecision="1" longitude="-47.509335" municipality="Alto Paraiso de Goias" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
<collectingCountry id="CEC306AEFFC1A313A2F935A078C618EE" box="[911,1016,1708,1733]" name="Brazil" pageId="6" pageNumber="277">BRAZIL</collectingCountry>
.
<collectingRegion id="741088DCFFC1A313A57735A07F7918EE" box="[1025,1095,1708,1733]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
:
<collectingMunicipality id="560FDC44FFC1A313A53B35A07E7E18EE" box="[1101,1344,1708,1733]" pageId="6" pageNumber="277">Alto Paraíso de Goiás</collectingMunicipality>
,
<location id="B30B10E5FFC1A313A43F35A07F0218C2" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856:B30B10E5FFC1A313A43F35A07F0218C2" country="Brazil" latitude="-14.106029" longLatPrecision="1" longitude="-47.509335" municipality="Alto Paraiso de Goias" name="Chapada dos Veadeiros, estrada Alto Paraiso de Goias - Teresina de Goias, Fazenda Agua Fria" pageId="6" pageNumber="277" stateProvince="Goias">Chapada dos Veadeiros, estrada Alto Paraíso de Goiás - Teresina de Goiás, Fazenda Água Fria</location>
, em campo limpo após a piscina, próximo a vereda e mata de galeria,
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14
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0621.7S
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,
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47
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3033.6W
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.
<date id="C26A60FEFFC1A313A21B35F878C21926" box="[877,1020,1780,1805]" pageId="6" pageNumber="277" value="2012-07-19">
<collectingDate id="D22E9916FFC1A313A21B35F878C21926" box="[877,1020,1780,1805]" pageId="6" pageNumber="277" value="2012-07-19">19 July 2012</collectingDate>
</date>
(fl),
<emphasis id="84A09A2CFFC1A313A54235F87E951926" box="[1076,1451,1780,1805]" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A54235F87FF91926" box="[1076,1223,1780,1805]" pageId="6" pageNumber="277">M. Watanabe</collectorName>
&amp;
<collectorName id="1B2123E8FFC1A313A59C35F87E451926" box="[1258,1403,1780,1805]" pageId="6" pageNumber="277">W. Watanabe</collectorName>
377
</emphasis>
(
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<collectionCode id="D0C5DEFBFFC1A313A18C34147A0F191A" box="[250,305,1816,1841]" pageId="6" pageNumber="277">SPF</collectionCode>
!
</materialsCitation>
,
<materialsCitation id="06BC4C63FFC1A313A03634147A8C191A" box="[320,434,1816,1841]" collectingDate="2012-07-19" collectionCode="F" collectorName="M. Watanabe &amp; W. Watanabe" country="Brazil" latitude="-14.106029" location="Chapada dos Veadeiros, estrada Alto Paraiso de Goias - Teresina de Goias, Fazenda Agua Fria" longLatPrecision="1" longitude="-47.509335" municipality="Alto Paraiso de Goias" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Goias" typeStatus="isotype">
<typeStatus id="696FF89CFFC1A313A03634147AAF191A" box="[320,401,1816,1841]" pageId="6" pageNumber="277" type="isotype">isotype</typeStatus>
<collectionCode id="D0C5DEFBFFC1A313A0EE34147A90191A" box="[408,430,1816,1841]" pageId="6" pageNumber="277">F</collectionCode>
!
</materialsCitation>
).
<figureCitation id="2EEF5ABBFFC1A313A0BF34147959191A" box="[457,615,1816,1841]" captionStart="FIGURE 2" captionStartId="4.[136,229,1920,1941]" captionTargetBox="[224,1358,207,1849]" captionTargetId="figure-18@4.[214,1373,190,1895]" captionTargetPageId="4" captionText="FIGURE 2AG. Syngonanthus incurvifolius (based on Irwin et al. 24648). A. Habit; B. Capitulum detail; C. From outermost to innermost series of involucral bracts; D. Staminate flower; E. Various forms of sepals of staminate flower; F. Pistillate flower with one sepal removed; G. Gynoecium. HN. S. vittatus (based on Watanabe &amp; Watanabe 377). H. Habit; I. Capitulum detail; J. From outermost to innermost series of involucral bracts; K. Staminate flower; L. Various forms of sepals of staminate flower; M. Pistillate flower with one sepal removed; N. Gynoecium. Illustrated by Marcelo Kubo." figureDoi="http://doi.org/10.5281/zenodo.13637542" httpUri="https://zenodo.org/record/13637542/files/figure.png" pageId="6" pageNumber="277">Figures 2HN</figureCitation>
&amp;
<figureCitation id="2EEF5ABBFFC1A313A3FA341479A3191A" box="[652,669,1816,1841]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="6" pageNumber="277">4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A1FE346E78E31994" pageId="6" pageNumber="277" type="diagnosis">
<paragraph id="B66B463EFFC1A313A1FE346E78E31994" blockId="6.[136,1451,1890,1983]" pageId="6" pageNumber="277">
The new species is similar to
<taxonomicName id="71D43DBDFFC1A313A0DA346E7874195C" box="[428,842,1890,1911]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFC1A313A0DA346E79A8195C" box="[428,662,1890,1911]" italics="true" pageId="6" pageNumber="277">Syngonanthus arenarius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A3BE346E7874195C" box="[712,842,1890,1911]" italics="true" pageId="6" pageNumber="277">heterophyllus</emphasis>
</taxonomicName>
but differs by involucral bracts narrowly lanceolate with apex markedly acute and petals glabrous in pistillate flowers.
<taxonomicName id="71D43DBDFFC1A313A3A0348A789719B0" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[726,937,1926,1947]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFC1A313A3A0348A789719B0" box="[726,937,1926,1947]" italics="true" pageId="6" pageNumber="277">Syngonanthus vittatus</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="71D43DBDFFC1A313A526348A7E9519B0" authorityName="Moldenke" authorityYear="1972" box="[1104,1451,1926,1947]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC1A313A526348A7E7A19B0" box="[1104,1348,1926,1947]" italics="true" pageId="6" pageNumber="277">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A404348A7E9519B0" box="[1394,1451,1926,1947]" italics="true" pageId="6" pageNumber="277">majus</emphasis>
</taxonomicName>
differing by all characteristics mentioned above as well the rosette leaves recurved.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A312A1FE34FC7E51181E" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277" type="description">
<paragraph id="B66B463EFFC1A312A1FE34FC7E51181E" blockId="6.[136,1452,2032,2093]" lastBlockId="7.[136,1452,160,2057]" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277">
Perennial herbs,
<quantity id="712CEBDBFFC1A313A03D34FC7A831622" box="[331,445,2032,2057]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="277" unit="cm" value="30.0" valueMax="40.0" valueMin="20.0">2040 cm</quantity>
tall.
<emphasis id="84A09A2CFFC1A313A08B34FC79031622" box="[509,573,2032,2057]" italics="true" pageId="6" pageNumber="277">Roots</emphasis>
<quantity id="712CEBDBFFC1A313A33E34FC79EC1622" box="[584,722,2032,2057]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="6" pageNumber="277" unit="mm" value="0.85" valueMax="1.5" valueMin="0.2">0.21.5 mm</quantity>
in diameter, white to cream-colored, spongy.
<emphasis id="84A09A2CFFC1A313A59A34FC7E721622" box="[1260,1356,2032,2057]" italics="true" pageId="6" pageNumber="277">Rhizome</emphasis>
<quantity id="712CEBDBFFC1A313A42134FC7B941606" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.3" metricValueMin="0.7" pageId="6" pageNumber="277" unit="cm" value="2.0" valueMax="3.3" valueMin="0.7">0.73.3 cm</quantity>
×
<quantity id="712CEBDBFFC1A313A1BD3B187A681606" box="[203,342,2068,2093]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="6" pageNumber="277" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
, horizontal to oblique, often branched, tomentose to villous, with white, dense and woolly curled trichomes; each branch giving rise to a rosette of leaves, each rosette bearing 13 synflorescences.
<emphasis id="84A09A2CFFC0A312A58933AC7E621E92" box="[1279,1372,160,185]" italics="true" pageId="7" pageNumber="278">Rosettes</emphasis>
<quantity id="712CEBDBFFC0A312A41333AC7B8A1EF6" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.1" metricValueMin="1.9" pageId="7" pageNumber="278" unit="mm" value="30.0" valueMax="41.0" valueMin="19.0">1941 mm</quantity>
in diameter.
<emphasis id="84A09A2CFFC0A312A03A33C87AA41EF6" box="[332,410,196,221]" italics="true" pageId="7" pageNumber="278">Leaves</emphasis>
9.022.5 × 0.31.0 mm, linear to narrowly lanceolate, flat to semi-terete, apex acute, base truncate, recurved, coriaceous, striate abaxially with 35 conspicuous veins, only the midvein visible on upper surface in young leaves; pilose on both surfaces, with arachnoid trichomes and adpressed trichomes adaxially, filamentous hairs and adpressed trichomes abaxially, mainly concentrated on midvein, ca.
<quantity id="712CEBDBFFC0A312A56E323C7F4C1F62" box="[1048,1138,304,329]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="278" unit="mm" value="0.3">0.3 mm</quantity>
long; basal sheath slightly enlarged, 0.51.0 mm wide, pale and achlorophyllous at base.
<emphasis id="84A09A2CFFC0A312A21132587F271F46" box="[871,1049,340,365]" italics="true" pageId="7" pageNumber="278">Synflorescences</emphasis>
with a primary axis bearing a lax umbel of 15 inflorescences at the apex.
<emphasis id="84A09A2CFFC0A312A3023274786D1FBA" box="[628,851,376,401]" italics="true" pageId="7" pageNumber="278">Synflorescence axis</emphasis>
5.0
<quantity id="712CEBDBFFC0A312A2E7327478DF1FBA" box="[913,993,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="278" unit="cm" value="8.5">8.5 cm</quantity>
long, emerging from the center of the rosette, erect to flexuous, yellow-reddish when young, dark reddish brown in maturity, pilose at base with filamentous trichomes, glabrescent, becoming tomentose near the apex with simple filamentous and capitate trichomes.
<emphasis id="84A09A2CFFC0A312A43332CC7A611FD6" italics="true" pageId="7" pageNumber="278">Bracts of synflorescence axis</emphasis>
usually restricted to the apex, rarely a few tiny scattered bracts along the axis ca.
<quantity id="712CEBDBFFC0A312A59532E87E1A1FD6" box="[1251,1316,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="278" unit="mm" value="3.0">3 mm</quantity>
long; apical bracts 130 × 0.31.0 mm, whorled, patent, the external series lanceolate, progressively elongating from the outer toward the inner series, the internal series linear, apex acute to rounded, erect, pubescent on both surfaces with capitate filamentous trichomes.
<emphasis id="84A09A2CFFC0A312A0E4315C790D1C42" box="[402,563,592,617]" italics="true" pageId="7" pageNumber="278">Inflorescences</emphasis>
composed of a closed spathe subtending a scape with a capitulum at the apex.
<emphasis id="84A09A2CFFC0A312A1FE31787BE11CA6" box="[136,223,628,653]" italics="true" pageId="7" pageNumber="278">Spathes</emphasis>
<quantity id="712CEBDBFFC0A312A19031787A5C1CA6" box="[230,354,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="7" pageNumber="278" unit="cm" value="4.5" valueMax="5.5" valueMin="3.5">3.55.5 cm</quantity>
long, chartaceous, cylindrical, obliquely opened, apex acute, erect, recurved or rarely involute, tip erect to uncinate, margin minutely involute, ciliate, adaxial surface glabrous to glabrescent, abaxial surface pubescent with capitate trichomes.
<emphasis id="84A09A2CFFC0A312A0ED31B07AD71CFE" box="[411,489,700,725]" italics="true" pageId="7" pageNumber="278">Scapes</emphasis>
erect,
<quantity id="712CEBDBFFC0A312A34231B079AB1CFE" box="[564,661,700,725]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="3.0" metricValueMin="0.9" pageId="7" pageNumber="278" unit="cm" value="19.5" valueMax="30.0" valueMin="9.0">930 cm</quantity>
long, greenish to straw-colored with age, 4(5)-costate, sparsely hairy to densely pilose with erect to patent glandular hairs and appressed, and simple trichomes, pubescent near the apex with a collar of patent, simple filamentous and capitate [gland-tipped] trichomes.
<emphasis id="84A09A2CFFC0A312A54930087F9F1D36" box="[1087,1185,772,797]" italics="true" pageId="7" pageNumber="278">Capitula</emphasis>
<quantity id="712CEBDBFFC0A312A5DD30087E271D36" box="[1195,1305,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="7" pageNumber="278" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">512 mm</quantity>
in diameter, obconic, later becoming depressed-hemispherical.
<emphasis id="84A09A2CFFC0A312A3BF302478B31D6A" box="[713,909,808,833]" italics="true" pageId="7" pageNumber="278">Involucral bracts</emphasis>
in 46 series, all similar in color, light to dark brown, often with a reddish tinge, with a lighter longitudinal band in center and a darker pigmentation on the margins, adaxial surface glabrous, abaxial surface strigose, ciliate, with patent, simple trichomes, rarely also capitate trichomes ca.
<quantity id="712CEBDBFFC0A312A1D830987A3D1D86" box="[174,259,916,941]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="278" unit="mm" value="0.5">0.5 mm</quantity>
long; external series 1.52.0 ×
<quantity id="712CEBDBFFC0A312A316309879D61D86" box="[608,744,916,941]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="7" pageNumber="278" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
, triangular to ovate, apex acute, bracts progressively elongating toward the internal series, middle series 45 ×
<quantity id="712CEBDBFFC0A312A3D030B479D51DFA" box="[678,747,952,977]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="278" unit="mm" value="1.0">1 mm</quantity>
, narrow elliptic, apex acute, internal series 3.05.5 × 0.71.0 mm, inner series surpassing the flowers by
<quantity id="712CEBDBFFC0A312A30630D079C71DDE" box="[624,761,988,1013]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="278" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.10.3 mm</quantity>
, narrow-lanceolate, elliptic, rarely narrow-oblanceolate, apex acute; receptacle hemispheric, pilose.
<emphasis id="84A09A2CFFC0A312A341370C79F31A32" box="[567,717,1024,1049]" italics="true" pageId="7" pageNumber="278">Floral bracts</emphasis>
absent.
<emphasis id="84A09A2CFFC0A312A25A370C78B61A32" box="[812,904,1024,1049]" italics="true" pageId="7" pageNumber="278">Flowers</emphasis>
3-merous, 60110 per capitulum, staminate and pistillate generally equal in number, with contrasting colors, staminate flowers darker and concentrated at the border of the capitulum.
<emphasis id="84A09A2CFFC0A312A024374479221A4A" box="[338,540,1096,1121]" italics="true" pageId="7" pageNumber="278">Staminate flowers</emphasis>
<quantity id="712CEBDBFFC0A312A353374479931A4A" box="[549,685,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="7" pageNumber="278" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
long, including pedicels; pedicels 0.51.0 mm long; sepals 2.0
<quantity id="712CEBDBFFC0A312A4FF37447B8A1AAE" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="278" unit="mm" value="2.5">2.5 mm</quantity>
long, coppery brown, shortly fused at base, concave, membranous, usually two sepals similar in form, elliptic to lanceolate (-oblanceolate) with acute apex, slightly curved, asymmetrical, and the third different, slightly irregular-subspathulate with round to truncate apex, but occasionally occurs a reversal of forms, or occasionally the three sepals slightly different, margins ciliate, adaxially glabrous, abaxially setose with simple filamentous trichomes ca.
<quantity id="712CEBDBFFC0A312A42337D47E921ADA" box="[1365,1452,1240,1265]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="278" unit="mm" value="0.5">0.5 mm</quantity>
long; petals ca.
<quantity id="712CEBDBFFC0A312A04937F07ABD1B3E" box="[319,387,1276,1301]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="278" unit="mm" value="2.0">2 mm</quantity>
long, hyaline, membranous, fused, tubulose, lobes triangular, obtuse, involute after anthesis, glabrous; filaments free, anthers ca.
<quantity id="712CEBDBFFC0A312A357362C79491B12" box="[545,631,1312,1337]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="278" unit="mm" value="0.6">0.6 mm</quantity>
long, white; anthophore ca.
<quantity id="712CEBDBFFC0A312A2CF362C7F311B12" box="[953,1039,1312,1337]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="278" unit="mm" value="0.3">0.3 mm</quantity>
long; pistillodes 3, ca.
<quantity id="712CEBDBFFC0A312A463362C7E521B12" box="[1301,1388,1312,1337]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="278" unit="mm" value="0.2">0.2 mm</quantity>
long, clavate, papillose at apex.
<emphasis id="84A09A2CFFC0A312A0DB364879551B76" box="[429,619,1348,1373]" italics="true" pageId="7" pageNumber="278">Pistillate flowers</emphasis>
<quantity id="712CEBDBFFC0A312A304364879C61B76" box="[626,760,1348,1373]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="7" pageNumber="278" unit="mm" value="2.95" valueMax="3.5" valueMin="2.4">2.43.5 mm</quantity>
long, including pedicels of
<quantity id="712CEBDBFFC0A312A55836487F8A1B76" box="[1070,1204,1348,1373]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="7" pageNumber="278" unit="mm" value="0.95" valueMax="1.5" valueMin="0.4">0.41.5 mm</quantity>
long; sepals ca.
<quantity id="712CEBDBFFC0A312A41D36487E921B76" box="[1387,1452,1348,1373]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="278" unit="mm" value="2.0">2 mm</quantity>
long, elliptic, concave, apex acute to obtuse, base thickened, free, cream to light castaneous, hyaline and membranous toward the apex, dark reddish brown-colored at apex, ciliate, hairy on the inside, with simple filamentous trichomes up to
<quantity id="712CEBDBFFC0A312A1B136BC7A391BE2" box="[199,263,1456,1481]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="278" unit="mm" value="1.0">1 mm</quantity>
long, glabrous outside; petals about equaling the sepals,
<quantity id="712CEBDBFFC0A312A2F736BC7F361BE2" box="[897,1032,1456,1481]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.7" metricValueMin="1.8" pageId="7" pageNumber="278" unit="mm" value="2.25" valueMax="2.7" valueMin="1.8">1.82.7 mm</quantity>
long, oblanceolate, fused at the distal third, membranous, white to hyaline, glabrous, the lobes triangular-rounded to obtuse, involute after anthesis; ovary ca.
<quantity id="712CEBDBFFC0A312A1FE36F47BE3183A" box="[136,221,1528,1553]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="278" unit="mm" value="0.6">0.6 mm</quantity>
long; style column
<quantity id="712CEBDBFFC0A312A0C136F47903183A" box="[439,573,1528,1553]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="7" pageNumber="278" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
long, cream-colored; appendages ca.
<quantity id="712CEBDBFFC0A312A2A936F47F0A183A" box="[991,1076,1528,1553]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="278" unit="mm" value="0.5">0.5 mm</quantity>
long, the glandular apex capitatecurved; stigmatic branches simple, ca.
<quantity id="712CEBDBFFC0A312A34D351079AF181E" box="[571,657,1564,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="278" unit="mm" value="1.5">1.5 mm</quantity>
long.
<emphasis id="84A09A2CFFC0A312A3A135107832181E" box="[727,780,1564,1589]" italics="true" pageId="7" pageNumber="278">Seed</emphasis>
ca. 0.40 ×
<quantity id="712CEBDBFFC0A312A2FF351078D1181E" box="[905,1007,1564,1589]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="278" unit="mm" value="0.25">0.25 mm</quantity>
, ellipsoid, reddish-brown colored.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC0A312A1CB354C7A751856" pageId="7" pageNumber="278" type="etymology">
<paragraph id="B66B463EFFC0A312A1CB354C7A751856" blockId="7.[136,1452,160,2057]" pageId="7" pageNumber="278">
<emphasis id="84A09A2CFFC0A312A1CB354C7A5A1872" bold="true" box="[189,356,1600,1625]" pageId="7" pageNumber="278">Etymology:—</emphasis>
The specific epithet refers to the well-marked longitudinal stripe evident in the center of the involucral bracts.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC0A31CA1CB358478631CA6" lastPageId="9" lastPageNumber="280" pageId="7" pageNumber="278" type="discussion">
<paragraph id="B66B463EFFC0A312A1CB35847FE8191A" blockId="7.[136,1452,160,2057]" pageId="7" pageNumber="278">
<emphasis id="84A09A2CFFC0A312A1CB35847969188A" bold="true" box="[189,599,1672,1697]" pageId="7" pageNumber="278">Notes and taxonomic affinities:—</emphasis>
In the field, the species looks superficially similar to
<taxonomicName id="71D43DBDFFC0A312A5BF35847E5D188A" box="[1225,1379,1672,1697]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC0A312A5BF35847E5D188A" box="[1225,1379,1672,1697]" italics="true" pageId="7" pageNumber="278">Paepalanthus</emphasis>
</taxonomicName>
(
<figureCitation id="2EEF5ABBFFC0A312A40035847B9018EE" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="7" pageNumber="278">FIG. 4A</figureCitation>
), because of its pilose capitula and dark involucral bracts. However, a closer examination reveals the compound inflorescences (synflorescences), with a differentiated, usually leafless primary axis, a feature that does not occur in
<taxonomicName id="71D43DBDFFC0A312A1FE35F87A1C1926" box="[136,290,1780,1805]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC0A312A1FE35F87A1C1926" box="[136,290,1780,1805]" italics="true" pageId="7" pageNumber="278">Paepalanthus</emphasis>
</taxonomicName>
. In addition, the species has petals of pistillate flowers fused in the distal part and free at the base, and staminate flowers isomerous an exclusive set of characters of
<taxonomicName id="71D43DBDFFC0A312A24B34147FEC191A" authority="(Giulietti et al. 2012)" baseAuthorityName="Giulietti" baseAuthorityYear="2012" box="[829,1234,1816,1841]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC0A312A24B341478E4191A" box="[829,986,1816,1841]" italics="true" pageId="7" pageNumber="278">Syngonanthus</emphasis>
(
<bibRefCitation id="D2453BCFFFC0A312A29E34147FF5191A" author="Giulietti, A. M. &amp; Andrade, M. J. G. de &amp; Scatena, V. L. &amp; Trovo, M. &amp; Coan, A. I. &amp; Sano, P. T. &amp; Santos, F. de &amp; Borges, R. L. B. de &amp; van den Berg, C." box="[1000,1227,1816,1841]" pageId="7" pageNumber="278" pagination="1 - 19" refId="ref7469" refString="Giulietti, A. M., Andrade, M. J. G. de, Scatena, V. L., Trovo, M., Coan, A. I., Sano, P. T., Santos, F. de A. R. dos, Borges, R. L. B. de &amp; van den Berg, C. (2012) Molecular phylogeny, morphology and their implications for the taxonomy of Eriocaulaceae. Rodriguesia 63: 1 - 19." type="journal article" year="2012">
Giulietti
<emphasis id="84A09A2CFFC0A312A53934147FBE191A" box="[1103,1152,1816,1841]" italics="true" pageId="7" pageNumber="278">et al</emphasis>
. 2012
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="B66B463EFFC0A312A1CB343079811622" blockId="7.[136,1452,160,2057]" pageId="7" pageNumber="278">
<taxonomicName id="71D43DBDFFC0A312A1CB34307A85197E" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[189,443,1852,1877]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFC0A312A1CB34307A85197E" box="[189,443,1852,1877]" italics="true" pageId="7" pageNumber="278">Syngonanthus vittatus</emphasis>
</taxonomicName>
belongs to the same morphological group as
<emphasis id="84A09A2CFFC0A312A28134307F8E197E" box="[1015,1200,1852,1877]" italics="true" pageId="7" pageNumber="278">
<taxonomicName id="71D43DBDFFC0A312A28134307F95197E" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1015,1195,1852,1877]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="species" species="incurvifolius">S. incurvifolius</taxonomicName>
,
</emphasis>
which also includes
<taxonomicName id="71D43DBDFFC0A312A1FE346C7AA21952" authorityName="Ruhland" authorityYear="1903" baseAuthorityName="Gardner" box="[136,412,1888,1913]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="species" species="arenarius">
<emphasis id="84A09A2CFFC0A312A1FE346C7AA21952" box="[136,412,1888,1913]" italics="true" pageId="7" pageNumber="278">Syngonanthus arenarius</emphasis>
</taxonomicName>
,
<taxonomicName id="71D43DBDFFC0A312A0DB346C781D1952" authorityName="Moldenke" authorityYear="1984" box="[429,803,1888,1913]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC0A312A0DB346C79781952" box="[429,582,1888,1913]" italics="true" pageId="7" pageNumber="278">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC0A312A3F2346C781D1952" box="[644,803,1888,1913]" italics="true" pageId="7" pageNumber="278">brachyphyllus</emphasis>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFC0A312A216346C7F401952" authorityName="Moldenke" authorityYear="1972" box="[864,1150,1888,1913]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC0A312A216346C78C71952" box="[864,1017,1888,1913]" italics="true" pageId="7" pageNumber="278">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC0A312A541346C7F401952" box="[1079,1150,1888,1913]" italics="true" pageId="7" pageNumber="278">majus</emphasis>
</taxonomicName>
<emphasis id="84A09A2CFFC0A312A508346C7FBC1952" box="[1150,1154,1888,1913]" italics="true" pageId="7" pageNumber="278">.</emphasis>
All these species share a similar architecture and dark and pilose involucral bracts with a contrasting pale longitudinal band.
<taxonomicName id="71D43DBDFFC0A312A47834887BE419EA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="278" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFC0A312A47834887BE419EA" italics="true" pageId="7" pageNumber="278">Syngonanthus vittatus</emphasis>
</taxonomicName>
differs from all of them by the involucral bracts narrowly lanceolate to linear, with a narrower and more conspicuous pale stripe, composing a laxer involucre (
<figureCitation id="2EEF5ABBFFC0A312A38834C0785A19CE" box="[766,868,1996,2021]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="7" pageNumber="278">FIG. 4G</figureCitation>
); the other species have large elliptical to obovate bracts, forming a denser involucre (
<figureCitation id="2EEF5ABBFFC0A312A36134FC798D1622" box="[535,691,2032,2057]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="7" pageNumber="278">Fig. 4F, 4HI</figureCitation>
).
</paragraph>
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<paragraph id="B66B463EFFCFA31DA1FE341F7F1A160F" blockId="8.[136,1452,1811,2084]" pageId="8" pageNumber="279">
<emphasis id="84A09A2CFFCFA31DA1FE341F7BC11903" bold="true" box="[136,255,1811,1832]" pageId="8" pageNumber="279">FIGURE 4.</emphasis>
<taxonomicName id="71D43DBDFFCFA31DA071341F7AE21903" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[263,476,1811,1832]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCFA31DA071341F7AE21903" box="[263,476,1811,1832]" italics="true" pageId="8" pageNumber="279">Syngonanthus vittatus</emphasis>
</taxonomicName>
and other morphologically similar
<taxonomicName id="71D43DBDFFCFA31DA241341F7F171903" box="[823,1065,1811,1832]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="84A09A2CFFCFA31DA241341F78801903" box="[823,958,1811,1832]" italics="true" pageId="8" pageNumber="279">Syngonanthus</emphasis>
species.
<emphasis id="84A09A2CFFCFA31DA56E341F7F171903" bold="true" box="[1048,1065,1811,1832]" pageId="8" pageNumber="279">A</emphasis>
</taxonomicName>
<emphasis id="84A09A2CFFCFA31DA55F341F7F7B1903" bold="true" box="[1065,1093,1811,1832]" pageId="8" pageNumber="279">E</emphasis>
.
<taxonomicName id="71D43DBDFFCFA31DA525341F7F8D1903" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1107,1203,1811,1832]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCFA31DA525341F7F8D1903" box="[1107,1203,1811,1832]" italics="true" pageId="8" pageNumber="279">S. vittatus</emphasis>
</taxonomicName>
:
<emphasis id="84A09A2CFFCFA31DA5B7341F7FE71903" bold="true" box="[1217,1241,1811,1832]" pageId="8" pageNumber="279">A.</emphasis>
Habit detail showing specimen at flowering;
<emphasis id="84A09A2CFFCFA31DA019343B7ABB1967" bold="true" box="[367,389,1847,1868]" pageId="8" pageNumber="279">B.</emphasis>
Rosette and roots;
<emphasis id="84A09A2CFFCFA31DA330343B79631967" bold="true" box="[582,605,1847,1868]" pageId="8" pageNumber="279">C.</emphasis>
Elliptic sepal of staminate flower;
<emphasis id="84A09A2CFFCFA31DA2CF343B78EE1967" bold="true" box="[953,976,1847,1868]" pageId="8" pageNumber="279">D.</emphasis>
Other forms of sepals revealing concave shapes in staminate flowers;
<emphasis id="84A09A2CFFCFA31DA01734577A49195B" bold="true" box="[353,375,1883,1904]" pageId="8" pageNumber="279">E.</emphasis>
Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals.
<emphasis id="84A09A2CFFCFA31DA4F734577E95195B" bold="true" box="[1409,1451,1883,1904]" pageId="8" pageNumber="279">FI.</emphasis>
Comparison among capitula in
<taxonomicName id="71D43DBDFFCFA31DA0C6347379BA19BF" box="[432,644,1919,1940]" pageId="8" pageNumber="279">
<emphasis id="84A09A2CFFCFA31DA0C63473790919BF" box="[432,567,1919,1940]" italics="true" pageId="8" pageNumber="279">Syngonanthus</emphasis>
species
</taxonomicName>
:
<emphasis id="84A09A2CFFCFA31DA3FD347379A019BF" bold="true" box="[651,670,1919,1940]" pageId="8" pageNumber="279">F.</emphasis>
<taxonomicName id="71D43DBDFFCFA31DA3D4347378FA19BF" box="[674,964,1919,1940]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFCFA31DA3D43473782B19BF" box="[674,789,1919,1940]" italics="true" pageId="8" pageNumber="279">S. arenarius</emphasis>
var.
<emphasis id="84A09A2CFFCFA31DA234347378FA19BF" box="[834,964,1919,1940]" italics="true" pageId="8" pageNumber="279">heterophyllus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFCFA31DA2B9347378D619BF" bold="true" box="[975,1000,1919,1940]" pageId="8" pageNumber="279">G.</emphasis>
<taxonomicName id="71D43DBDFFCFA31DA29A34737F7419BF" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1004,1098,1919,1940]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCFA31DA29A34737F7419BF" box="[1004,1098,1919,1940]" italics="true" pageId="8" pageNumber="279">S. vittatus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFCFA31DA52234737F5019BF" bold="true" box="[1108,1134,1919,1940]" pageId="8" pageNumber="279">H.</emphasis>
<taxonomicName id="71D43DBDFFCFA31DA50434737E9B19BF" authorityName="Moldenke" authorityYear="1984" box="[1138,1445,1919,1940]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFCFA31DA50434737FCE19BF" box="[1138,1264,1919,1940]" italics="true" pageId="8" pageNumber="279">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFCFA31DA46B34737E9B19BF" box="[1309,1445,1919,1940]" italics="true" pageId="8" pageNumber="279">brachyphyllus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFCFA31DA1FE34AF7BA91993" bold="true" box="[136,151,1955,1976]" pageId="8" pageNumber="279">I.</emphasis>
<taxonomicName id="71D43DBDFFCFA31DA1D634AF7AB01993" authorityName="Moldenke" authorityYear="1972" box="[160,398,1955,1976]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFCFA31DA1D634AF7A1C1993" box="[160,290,1955,1976]" italics="true" pageId="8" pageNumber="279">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFCFA31DA02334AF7AB01993" box="[341,398,1955,1976]" italics="true" pageId="8" pageNumber="279">majus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFCFA31DA0EB34AF7AF61993" bold="true" box="[413,456,1955,1976]" pageId="8" pageNumber="279">JK</emphasis>
. Pistillate flower of
<taxonomicName id="71D43DBDFFCFA31DA3E334AF79C81993" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[661,758,1955,1976]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCFA31DA3E334AF79C81993" box="[661,758,1955,1976]" italics="true" pageId="8" pageNumber="279">S. vittatus</emphasis>
</taxonomicName>
:
<emphasis id="84A09A2CFFCFA31DA27334AF78291993" bold="true" box="[773,791,1955,1976]" pageId="8" pageNumber="279">J.</emphasis>
Flower exhibiting trichomes on sepals;
<emphasis id="84A09A2CFFCFA31DA5D334AF7F801993" bold="true" box="[1189,1214,1955,1976]" pageId="8" pageNumber="279">K.</emphasis>
Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals;
<emphasis id="84A09A2CFFCFA31DA2D434CB788619F7" bold="true" box="[930,952,1991,2012]" pageId="8" pageNumber="279">L.</emphasis>
Pistillate flower of
<taxonomicName id="71D43DBDFFCFA31DA50D34CB7E9B19F7" box="[1147,1445,1991,2012]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFCFA31DA50D34CB7FCF19F7" box="[1147,1265,1991,2012]" italics="true" pageId="8" pageNumber="279">S. arenarius</emphasis>
var.
<emphasis id="84A09A2CFFCFA31DA45534CB7E9B19F7" box="[1315,1445,1991,2012]" italics="true" pageId="8" pageNumber="279">heterophyllus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFCFA31DA1FE34E77BFC162B" bold="true" box="[136,194,2027,2048]" pageId="8" pageNumber="279">MN.</emphasis>
Staminate flowers of
<taxonomicName id="71D43DBDFFCFA31DA0E334E77ACA162B" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[405,500,2027,2048]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCFA31DA0E334E77ACA162B" box="[405,500,2027,2048]" italics="true" pageId="8" pageNumber="279">S. vittatus</emphasis>
</taxonomicName>
:
<emphasis id="84A09A2CFFCFA31DA37634E77922162B" bold="true" box="[512,540,2027,2048]" pageId="8" pageNumber="279">M.</emphasis>
Young flower;
<emphasis id="84A09A2CFFCFA31DA3C734E779F7162B" bold="true" box="[689,713,2027,2048]" pageId="8" pageNumber="279">N.</emphasis>
Mature flower;
<emphasis id="84A09A2CFFCFA31DA21234E77843162B" bold="true" box="[868,893,2027,2048]" pageId="8" pageNumber="279">O.</emphasis>
Staminate flower of
<taxonomicName id="71D43DBDFFCFA31DA53134E77E52162B" box="[1095,1388,2027,2048]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="279" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFCFA31DA53134E77F85162B" box="[1095,1211,2027,2048]" italics="true" pageId="8" pageNumber="279">S. arenarius</emphasis>
var.
<emphasis id="84A09A2CFFCFA31DA59C34E77E52162B" box="[1258,1388,2027,2048]" italics="true" pageId="8" pageNumber="279">heterophyllus</emphasis>
</taxonomicName>
. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe).
</paragraph>
</caption>
<paragraph id="B66B463EFFCEA31CA1CB33AC7ACC1F46" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
The flowers of
<taxonomicName id="71D43DBDFFCEA31CA00533AC79521E92" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[371,620,160,185]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA00533AC79521E92" box="[371,620,160,185]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
are very similar to those of
<taxonomicName id="71D43DBDFFCEA31CA2C833AC7E231E92" box="[958,1309,160,185]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFCEA31CA2C833AC7F771E92" box="[958,1097,160,185]" italics="true" pageId="9" pageNumber="280">S. arenarius</emphasis>
var.
<emphasis id="84A09A2CFFCEA31CA5F033AC7E231E92" box="[1158,1309,160,185]" italics="true" pageId="9" pageNumber="280">heterophyllus</emphasis>
</taxonomicName>
(1903:260), except that the former has petals of pistillate flowers glabrous (
<figureCitation id="2EEF5ABBFFCEA31CA21333C878DC1EF6" box="[869,994,196,221]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4JK</figureCitation>
;
<emphasis id="84A09A2CFFCEA31CA28633C87F311EF6" box="[1008,1039,196,221]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
petals pilose,
<figureCitation id="2EEF5ABBFFCEA31CA5C133C87E2B1EF6" box="[1207,1301,196,221]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4L</figureCitation>
) and smaller flowers (up to 3.5
<emphasis id="84A09A2CFFCEA31CA01533E47ABF1F2A" box="[355,385,232,257]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
<quantity id="712CEBDBFFCEA31CA0FD33E47AD61F2A" box="[395,488,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="280" unit="mm" value="4.5">4.5 mm</quantity>
). In addition,
<taxonomicName id="71D43DBDFFCEA31CA3FB33E479C01F2A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[653,766,232,257]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA3FB33E479C01F2A" box="[653,766,232,257]" italics="true" pageId="9" pageNumber="280">S. vittatus</emphasis>
</taxonomicName>
has pistillate and staminate flowers strongly contrasting in colour, the staminate with brown sepals and the pistillate with cream sepals (
<figureCitation id="2EEF5ABBFFCEA31CA57232007E381F0E" box="[1028,1286,268,293]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4MN and 4KJ</figureCitation>
, respectively), whereas both
<typeStatus id="696FF89CFFCEA31CA052323C7A611F62" box="[292,351,304,329]" pageId="9" pageNumber="280">types</typeStatus>
of flowers in
<taxonomicName id="71D43DBDFFCEA31CA088323C78681F62" box="[510,854,304,329]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="variety" species="arenarius" variety="heterophyllus">
<emphasis id="84A09A2CFFCEA31CA088323C79B91F62" box="[510,647,304,329]" italics="true" pageId="9" pageNumber="280">S. arenarius</emphasis>
var.
<emphasis id="84A09A2CFFCEA31CA3C9323C78681F62" box="[703,854,304,329]" italics="true" pageId="9" pageNumber="280">heterophyllus</emphasis>
</taxonomicName>
have a similar pigmentation (
<figureCitation id="2EEF5ABBFFCEA31CA5D0323C7E091F62" box="[1190,1335,304,329]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4L; 4O</figureCitation>
), cream at base and reddish-brown at apex.
</paragraph>
<paragraph id="B66B463EFFCEA31CA1CB32747ADE1C6E" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
Other features differentiating
<taxonomicName id="71D43DBDFFCEA31CA364327478351FBA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[530,779,376,401]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA364327478351FBA" box="[530,779,376,401]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
from
<taxonomicName id="71D43DBDFFCEA31CA22332747F3F1FBA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[853,1025,376,401]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFCEA31CA22332747F3F1FBA" box="[853,1025,376,401]" italics="true" pageId="9" pageNumber="280">S. incurvifolius</emphasis>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFCEA31CA54B32747E6A1FBA" authorityName="Moldenke" authorityYear="1972" box="[1085,1364,376,401]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFCEA31CA54B32747FEA1FBA" box="[1085,1236,376,401]" italics="true" pageId="9" pageNumber="280">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFCEA31CA46732747E6A1FBA" box="[1297,1364,376,401]" italics="true" pageId="9" pageNumber="280">majus</emphasis>
</taxonomicName>
are the rosette leaves recurved (
<figureCitation id="2EEF5ABBFFCEA31CA0E632907AD21F9E" box="[400,492,412,437]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4B</figureCitation>
) (
<emphasis id="84A09A2CFFCEA31CA3743290791E1F9E" box="[514,544,412,437]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
incurved in the distal half or flexuous, respectively,
<figureCitation id="2EEF5ABBFFCEA31CA52332907FF31F9E" box="[1109,1229,412,437]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="9" pageNumber="280">FIG 3AB</figureCitation>
), the synflorescence axis often bearing tiny scattered bracts along its length (
<emphasis id="84A09A2CFFCEA31CA27C32CC78171FF2" box="[778,809,448,473]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
bracts restricted to the apex), and the petals of pistillate flowers glabrous (
<emphasis id="84A09A2CFFCEA31CA02232E87A4D1FD6" box="[340,371,484,509]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
pilose).
<taxonomicName id="71D43DBDFFCEA31CA0A232E879F51FD6" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[468,715,484,509]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA0A232E879F51FD6" box="[468,715,484,509]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="71D43DBDFFCEA31CA2E232E87EB61FD6" authorityName="Moldenke" authorityYear="1984" box="[916,1416,484,509]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFCEA31CA2E232E87F8F1FD6" box="[916,1201,484,509]" italics="true" pageId="9" pageNumber="280">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFCEA31CA59F32E87EB61FD6" box="[1257,1416,484,509]" italics="true" pageId="9" pageNumber="280">brachyphyllus</emphasis>
</taxonomicName>
by the narrower leaves (0.31.0 mm
<emphasis id="84A09A2CFFCEA31CA3733104791D1C0A" box="[517,547,520,545]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
2.05.0 mm), with simple and adpressed trichomes (
<emphasis id="84A09A2CFFCEA31CA50A31047FA41C0A" box="[1148,1178,520,545]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
only simple) and striate adaxial surface (
<emphasis id="84A09A2CFFCEA31CA03731207A611C6E" box="[321,351,556,581]" italics="true" pageId="9" pageNumber="280">vs.</emphasis>
reticulate).
</paragraph>
<paragraph id="B66B463EFFCEA31CA1CB315C78631CA6" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
Additionally, the evident asymmetrical sepals of staminate flowers are an important characteristic shared with
<taxonomicName id="71D43DBDFFCEA31CA4E0315C7A281CA6" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFCEA31CA4E0315C7A281CA6" italics="true" pageId="9" pageNumber="280">S. incurvifolius</emphasis>
</taxonomicName>
and the other species previously cited (
<figureCitation id="2EEF5ABBFFCEA31CA3A63178786C1CA6" box="[720,850,628,653]" captionStart="FIGURE 4" captionStartId="8.[136,229,1811,1832]" captionTargetBox="[254,1333,193,1783]" captionTargetId="figure-18@8.[251,1336,190,1786]" captionTargetPageId="8" captionText="FIGURE 4. Syngonanthus vittatus and other morphologically similar Syngonanthus species. AE. S. vittatus: A. Habit detail showing specimen at flowering; B. Rosette and roots; C. Elliptic sepal of staminate flower; D. Other forms of sepals revealing concave shapes in staminate flowers; E. Three different shapes of sepals in the same staminate flower revealing asymmetry among the sepals. FI. Comparison among capitula in Syngonanthus species:F. S. arenarius var. heterophyllus; G. S. vittatus; H. S. densifolius var. brachyphyllus; I. S. densifolius var. majus; JK. Pistillate flower of S. vittatus: J. Flower exhibiting trichomes on sepals; K. Flower with two sepals removed, the remaining sepal with hairs deciduous, showing the glabrous petals; L. Pistillate flower of S. arenarius var. heterophyllus; MN. Staminate flowers of S. vittatus: M. Young flower; N. Mature flower; O. Staminate flower of S. arenarius var. heterophyllus. Scale bars: C,M= 500 μm. DE, JL, NO= 1 mm. FI= 2 mm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637546" httpUri="https://zenodo.org/record/13637546/files/figure.png" pageId="9" pageNumber="280">FIG. 4CE</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFCEA31CA1CB31947E0E1D36" pageId="9" pageNumber="280" type="biology_ecology">
<paragraph id="B66B463EFFCEA31CA1CB31947E0E1D36" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
<emphasis id="84A09A2CFFCEA31CA1CB31947A611C9A" bold="true" box="[189,351,664,689]" pageId="9" pageNumber="280">Phenology:—</emphasis>
Flowering of
<taxonomicName id="71D43DBDFFCEA31CA08A319479CF1C9A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[508,753,664,689]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA08A319479CF1C9A" box="[508,753,664,689]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
occurs during the dry season (winter), from July to September. This period coincides with all collections reported for this species. Most
<taxonomicName id="71D43DBDFFCEA31CA2B731B07F601CFE" box="[961,1118,700,725]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFCEA31CA2B731B07F601CFE" box="[961,1118,700,725]" italics="true" pageId="9" pageNumber="280">Syngonanthus</emphasis>
</taxonomicName>
from Chapada dos Veadeiros bloom at the same time, for example,
<taxonomicName id="71D43DBDFFCEA31CA34431EC789D1CD2" authority="Moldenke (1976: 485)" authorityName="Moldenke" authorityPageNumber="485" authorityYear="1976" box="[562,931,736,761]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="decorus">
<emphasis id="84A09A2CFFCEA31CA34431EC79991CD2" box="[562,679,736,761]" italics="true" pageId="9" pageNumber="280">S. decorus</emphasis>
<bibRefCitation id="D2453BCFFFCEA31CA3D831EC789D1CD2" author="Moldenke, H. N." box="[686,931,736,761]" pageId="9" pageNumber="280" pagination="483 - 487" refId="ref7700" refString="Moldenke, H. N. (1976) Notes on new and noteworthy plants LXXXV. Phytologia 32 (6): 483 - 487." type="journal article" year="1976">Moldenke (1976:485)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFCEA31CA2AD31EC7FD51CD2" authorityName="Moldenke" authorityYear="1972" box="[987,1259,736,761]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFCEA31CA2AD31EC7F4E1CD2" box="[987,1136,736,761]" italics="true" pageId="9" pageNumber="280">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFCEA31CA5DE31EC7FD51CD2" box="[1192,1259,736,761]" italics="true" pageId="9" pageNumber="280">majus</emphasis>
</taxonomicName>
. Seeds mature at the end of the dry season, implying a dispersal period concomitant with the rains and possible hydrochory.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFCEA31CA1CB30247AA41AAE" pageId="9" pageNumber="280" type="distribution">
<paragraph id="B66B463EFFCEA31CA1CB30247AA41AAE" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
<emphasis id="84A09A2CFFCEA31CA1CB302479371D6A" bold="true" box="[189,521,808,833]" pageId="9" pageNumber="280">Habitat and distribution:—</emphasis>
All collections are known from the same population as the
<typeStatus id="696FF89CFFCEA31CA5D130247FE41D6A" box="[1191,1242,808,833]" pageId="9" pageNumber="280">type</typeStatus>
, at the farm called “Água Fria”, except by a single record from waterfall “São Bento”. These localities are about
<quantity id="712CEBDBFFCEA31CA5EA30407FDC1D4E" box="[1180,1250,844,869]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="280" unit="km" value="10.0">10 km</quantity>
apart and close to recreational and agricultural areas. The expansion and intensification of human activities in the area threatens the natural surroundings. The limited distribution of the known populations contrasts with a local abundance of individuals, and further fieldwork may reveal other populations.
<taxonomicName id="71D43DBDFFCEA31CA3EC30B478B11DFA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[666,911,952,977]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA3EC30B478B11DFA" box="[666,911,952,977]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
occurs outside conservation units, although very close to the boundaries of the PNCV. The species occurs over hydromorphic soils, in moist grasslands or wet campo, a common physiognomy on
<emphasis id="84A09A2CFFCEA31CA0BA370C79741A32" box="[460,586,1024,1049]" italics="true" pageId="9" pageNumber="280">floodplains</emphasis>
between Cerrado
<emphasis id="84A09A2CFFCEA31CA26D370C78941A32" box="[795,938,1024,1049]" italics="true" pageId="9" pageNumber="280">sensu stricto</emphasis>
and gallery forest (
<bibRefCitation id="D2453BCFFFCEA31CA5FE370C7E6A1A32" author="Ratter, J. A. &amp; Ribeiro, J. F. &amp; Bridgewater, S." box="[1160,1364,1024,1049]" pageId="9" pageNumber="280" pagination="223 - 230" refId="ref7979" refString="Ratter, J. A., Ribeiro, J. F. &amp; Bridgewater, S. (1997) The Brazilian cerrado vegetation and threats to its biodiversity. Annals of Botany 80 (3): 223 - 230." type="journal article" year="1997">
Ratter
<emphasis id="84A09A2CFFCEA31CA5A1370C7E371A32" box="[1239,1289,1024,1049]" italics="true" pageId="9" pageNumber="280">et al</emphasis>
. 1997
</bibRefCitation>
). These wet
<emphasis id="84A09A2CFFCEA31CA1CF37287A311A16" box="[185,271,1060,1085]" italics="true" pageId="9" pageNumber="280">campos</emphasis>
occur on well-drained sandy soils, which become waterlogged during summer and quite dry in winter, a variation that constrains tree growth and prevents forest formations (
<bibRefCitation id="D2453BCFFFCEA31CA2D437447F531A4A" author="Ratter, J. A. &amp; Ribeiro, J. F. &amp; Bridgewater, S." box="[930,1133,1096,1121]" pageId="9" pageNumber="280" pagination="223 - 230" refId="ref7979" refString="Ratter, J. A., Ribeiro, J. F. &amp; Bridgewater, S. (1997) The Brazilian cerrado vegetation and threats to its biodiversity. Annals of Botany 80 (3): 223 - 230." type="journal article" year="1997">
Ratter
<emphasis id="84A09A2CFFCEA31CA28637447F1C1A4A" box="[1008,1058,1096,1121]" italics="true" pageId="9" pageNumber="280">et al</emphasis>
. 1997
</bibRefCitation>
), favoring the occurrence of herbs such as
<taxonomicName id="71D43DBDFFCEA31CA05237607AAC1AAE" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[292,402,1132,1157]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA05237607AAC1AAE" box="[292,402,1132,1157]" italics="true" pageId="9" pageNumber="280">S. vittatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFCEA31CA1CB379C7AED1B12" pageId="9" pageNumber="280" type="conservation">
<paragraph id="B66B463EFFCEA31CA1CB379C7AED1B12" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
<emphasis id="84A09A2CFFCEA31CA1CB379C7AEF1A82" bold="true" box="[189,465,1168,1193]" pageId="9" pageNumber="280">Conservation status:—</emphasis>
This species is classified as critically endangered, according to criteria B1ab and B2ab of
<bibRefCitation id="D2453BCFFFCEA31CA1D137B87A091AE6" author="IUCN" box="[167,311,1204,1229]" pageId="9" pageNumber="280" refId="ref7582" refString="IUCN (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland &amp; Cambridge, UK, 70 pp." type="book" year="2001">IUCN (2001</bibRefCitation>
,
<bibRefCitation id="D2453BCFFFCEA31CA03537B87ABE1AE6" author="IUCN" box="[323,384,1204,1229]" pageId="9" pageNumber="280" refId="ref7614" refString="IUCN (2015) The IUCN red list of threatened species. Available from: www. iucnredlist. org / (accessed 1 January 2015)" type="book" year="2015">2015</bibRefCitation>
). The taxon is known only from two populations, both on privately owned land hosting human activities that may threaten the species with extinction.
<taxonomicName id="71D43DBDFFCEA31CA27037D478C01ADA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[774,1022,1240,1265]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="280" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFCEA31CA27037D478C01ADA" box="[774,1022,1240,1265]" italics="true" pageId="9" pageNumber="280">Syngonanthus vittatus</emphasis>
</taxonomicName>
has an area of occupancy of
<quantity id="712CEBDBFFCEA31CA42337D47EA21ADA" box="[1365,1436,1240,1265]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="280" unit="km" value="12.0">12 km</quantity>
<superScript id="41A1EB76FFCEA31CA4EA37DA7E9A1ACF" attach="left" box="[1436,1444,1238,1252]" fontSize="6" pageId="9" pageNumber="280">2</superScript>
. Conservation status is endangered, as evaluated by CNCFlora according to the
<bibRefCitation id="D2453BCFFFCEA31CA57437F07FAC1B3E" author="IUCN" box="[1026,1170,1276,1301]" pageId="9" pageNumber="280" refId="ref7582" refString="IUCN (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland &amp; Cambridge, UK, 70 pp." type="book" year="2001">IUCN (2001</bibRefCitation>
,
<bibRefCitation id="D2453BCFFFCEA31CA5EB37F07FE71B3E" author="IUCN" box="[1181,1241,1276,1301]" pageId="9" pageNumber="280" refId="ref7614" refString="IUCN (2015) The IUCN red list of threatened species. Available from: www. iucnredlist. org / (accessed 1 January 2015)" type="book" year="2015">2015</bibRefCitation>
), matching criteria B2ab(i,ii,iii) (Negrão 2014b).
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFCEA31CA1CB36487AA01856" pageId="9" pageNumber="280" type="materials_examined">
<paragraph id="B66B463EFFCEA31CA1CB36487AA01856" blockId="9.[136,1452,160,1661]" pageId="9" pageNumber="280">
<emphasis id="84A09A2CFFCEA31CA1CB3648795A1B76" bold="true" box="[189,612,1348,1373]" pageId="9" pageNumber="280">Additional specimens examined:—</emphasis>
<materialsCitation id="06BC4C63FFCEA31CA312364879561BAA" collectingDate="1972-08-06" collectionCode="SPF" collectorName="Rizzo" country="Brazil" location="Est. 11 - Chapada dos Veadeiros" municipality="Alto Paraiso de Goias" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
<collectingCountry id="CEC306AEFFCEA31CA312364879F31B76" box="[612,717,1348,1373]" name="Brazil" pageId="9" pageNumber="280">BRAZIL</collectingCountry>
.
<collectingRegion id="741088DCFFCEA31CA3AD3648781F1B76" box="[731,801,1348,1373]" country="Brazil" name="Goias" pageId="9" pageNumber="280">Goiás</collectingRegion>
:
<collectingMunicipality id="560FDC44FFCEA31CA25B36487F0F1B76" box="[813,1073,1348,1373]" pageId="9" pageNumber="280">Alto Paraíso de Goiás</collectingMunicipality>
.
<location id="B30B10E5FFCEA31CA54936487E991B76" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856:B30B10E5FFCEA31CA54936487E991B76" box="[1087,1447,1348,1373]" country="Brazil" municipality="Alto Paraiso de Goias" name="Est. 11 - Chapada dos Veadeiros" pageId="9" pageNumber="280" stateProvince="Goias">Est. 11-Chapada dos Veadeiros</location>
, mata,
<date id="C26A60FEFFCEA31CA1A636647AB51BAA" box="[208,395,1384,1409]" pageId="9" pageNumber="280" value="1972-08-06">
<collectingDate id="D22E9916FFCEA31CA1A636647AB51BAA" box="[208,395,1384,1409]" pageId="9" pageNumber="280" value="1972-08-06">06 August 1972</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA0EC366479271BAA" box="[410,537,1384,1409]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA0EC36647AE91BAA" box="[410,471,1384,1409]" pageId="9" pageNumber="280">Rizzo</collectorName>
8236
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA35A3664795E1BAA" box="[556,608,1384,1409]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA30E36647F2F1B8E" collectingDate="1994-09-09" collectionCode="IBGE, SPF" collectorName="Sao Bento" country="Brazil" location="Sao Jorge, cachoeira Sao Bento" municipality="Alto Paraiso" pageId="9" pageNumber="280" specimenCount="1">
<quantity id="712CEBDBFFCEA31CA30E3664798A1BAA" box="[632,692,1384,1409]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="280" unit="km" value="9.0">
<locationDeviation id="A7D95F1CFFCEA31CA30E3664798A1BAA" box="[632,692,1384,1409]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="280" unit="km" value="9.0">9 km</locationDeviation>
</quantity>
de
<collectingMunicipality id="560FDC44FFCEA31CA3943664784E1BAA" box="[738,880,1384,1409]" pageId="9" pageNumber="280">Alto Paraíso</collectingMunicipality>
/
<location id="B30B10E5FFCEA31CA2F836647E3E1BAA" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856:B30B10E5FFCEA31CA2F836647E3E1BAA" box="[910,1280,1384,1409]" country="Brazil" municipality="Alto Paraiso" name="Sao Jorge, cachoeira Sao Bento" pageId="9" pageNumber="280">São Jorge, cachoeira São Bento</location>
, campo úmido próximo a floresta de galeria,
<date id="C26A60FEFFCEA31CA0AE3680798B1B8E" box="[472,693,1420,1445]" pageId="9" pageNumber="280" value="1994-09-09">
<collectingDate id="D22E9916FFCEA31CA0AE3680798B1B8E" box="[472,693,1420,1445]" pageId="9" pageNumber="280" value="1994-09-09">09 September 1994</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA3B6368078481B8E" box="[704,886,1420,1445]" italics="true" pageId="9" pageNumber="280">
<taxonomicName id="71D43DBDFFCEA31CA3B6368079C61B8E" box="[704,760,1420,1445]" class="Aves" family="Leiothrichidae" genus="Silva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Passeriformes" pageId="9" pageNumber="280" phylum="Chordata" rank="genus">Silva</taxonomicName>
et al. 2315
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA2F3368078F51B8E" box="[901,971,1420,1445]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA2A036807F371B8E" box="[982,1033,1420,1445]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA56B36807E3C1BE2" collectingDate="1997-06-06" collectionCode="UB" collectorName="Munhoz" country="Brazil" latitude="-14.151944" location="Fazenda Agua Fria" longLatPrecision="21" longitude="-47.61889" pageId="9" pageNumber="280" specimenCount="1">
<location id="B30B10E5FFCEA31CA56B36807FCB1B8E" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856:B30B10E5FFCEA31CA56B36807FCB1B8E" box="[1053,1269,1420,1445]" country="Brazil" latitude="-14.151944" longLatPrecision="21" longitude="-47.61889" name="Fazenda Agua Fria" pageId="9" pageNumber="280">Fazenda Água Fria</location>
, próximo a sede da fazenda, borda de mata de galeria,
<geoCoordinate id="D3E020F9FFCEA31CA35136BC798C1BE2" box="[551,690,1456,1481]" degrees="14" direction="south" minutes="09" orientation="latitude" pageId="9" pageNumber="280" precision="15" seconds="07" value="-14.151944">14º0907 S</geoCoordinate>
,
<geoCoordinate id="D3E020F9FFCEA31CA3C836BC78701BE2" box="[702,846,1456,1481]" degrees="47" direction="west" minutes="37" orientation="longitude" pageId="9" pageNumber="280" precision="15" seconds="08" value="-47.61889">47º3708W</geoCoordinate>
,
<date id="C26A60FEFFCEA31CA22F36BC78D11BE2" box="[857,1007,1456,1481]" pageId="9" pageNumber="280" value="1997-06-06">
<collectingDate id="D22E9916FFCEA31CA22F36BC78D11BE2" box="[857,1007,1456,1481]" pageId="9" pageNumber="280" value="1997-06-06">06 June 1997</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA28E36BC7FB11BE2" box="[1016,1167,1456,1481]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA28E36BC7F6C1BE2" box="[1016,1106,1456,1481]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al.
</emphasis>
462 (
<collectionCode id="D0C5DEFBFFCEA31CA5BB36BC7FC61BE2" box="[1229,1272,1456,1481]" pageId="9" pageNumber="280">UB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA47A36BC7FFF183A" collectingDate="2000-06-17" collectionCode="IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Fazenda Agua Fria, cerca de 10 km em direcao a Teresina de Goias" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
<location id="B30B10E5FFCEA31CA47A36BC79F01BC6" LSID="urn:lsid:plazi:treatment:3E7DF728FFC1A31CA1FE35A67AA01856:B30B10E5FFCEA31CA47A36BC79F01BC6" country="Brazil" latitude="-14.072695" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" name="Fazenda Agua Fria, cerca de 10 km em direcao a Teresina de Goias" pageId="9" pageNumber="280" stateProvince="Goias">
Fazenda Água Fria, cerca de
<quantity id="712CEBDBFFCEA31CA05336D87A551BC6" box="[293,363,1492,1517]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="280" unit="km" value="10.0">10 km</quantity>
em direção a Teresina de Goiás
</location>
, campo limpo úmido, borda de cerrado rupestre e mata de galeria,
<geoCoordinate id="D3E020F9FFCEA31CA1FE36F47A16183A" box="[136,296,1528,1553]" degrees="14" direction="south" minutes="04" orientation="latitude" pageId="9" pageNumber="280" precision="1" seconds="21.7" value="-14.072695">14°0421.7S</geoCoordinate>
,
<geoCoordinate id="D3E020F9FFCEA31CA04236F47AE3183A" box="[308,477,1528,1553]" degrees="47" direction="west" minutes="30" orientation="longitude" pageId="9" pageNumber="280" precision="1" seconds="33.6" value="-47.509335">47°3033.6W</geoCoordinate>
, altitude
<quantity id="712CEBDBFFCEA31CA33236F479A5183A" box="[580,667,1528,1553]" metricMagnitude="3" metricUnit="m" metricValue="1.488" pageId="9" pageNumber="280" unit="m" value="1488.0">
<elevation id="3DF9A10DFFCEA31CA33236F479A5183A" box="[580,667,1528,1553]" metricMagnitude="3" metricUnit="m" metricValue="1.488" pageId="9" pageNumber="280" unit="m" value="1488.0">1488 m</elevation>
</quantity>
,
<date id="C26A60FEFFCEA31CA3D136F4787E183A" box="[679,832,1528,1553]" pageId="9" pageNumber="280" value="2000-06-17">
<collectingDate id="D22E9916FFCEA31CA3D136F4787E183A" box="[679,832,1528,1553]" pageId="9" pageNumber="280" value="2000-06-17">17 June 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA23A36F47F1B183A" box="[844,1061,1528,1553]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA23A36F47898183A" box="[844,934,1528,1553]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1651
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA54236F47F44183A" box="[1076,1146,1528,1553]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA5F336F47F86183A" box="[1157,1208,1528,1553]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA5BB36F47AC2181E" collectingDate="2000-07-01" collectionCode="IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Teresina" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
idem,
<date id="C26A60FEFFCEA31CA46536F47E99183A" box="[1299,1447,1528,1553]" pageId="9" pageNumber="280" value="2000-07-01">
<collectingDate id="D22E9916FFCEA31CA46536F47E99183A" box="[1299,1447,1528,1553]" pageId="9" pageNumber="280" value="2000-07-01">01 July 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA1FE35107A5F181E" box="[136,353,1564,1589]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA1FE35107BDC181E" box="[136,226,1564,1589]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1721
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA00635107A88181E" box="[368,438,1564,1589]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA0B735107ACA181E" box="[449,500,1564,1589]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA37E35107F5E181E" box="[520,1120,1564,1589]" collectingDate="2000-07-01" collectionCode="IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Teresina" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
idem,
<date id="C26A60FEFFCEA31CA338351079DC181E" box="[590,738,1564,1589]" pageId="9" pageNumber="280" value="2000-07-01">
<collectingDate id="D22E9916FFCEA31CA338351079DC181E" box="[590,738,1564,1589]" pageId="9" pageNumber="280" value="2000-07-01">01 July 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA39B351078FB181E" box="[749,965,1564,1589]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA39B35107879181E" box="[749,839,1564,1589]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1729
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA2A235107F24181E" box="[980,1050,1564,1589]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA55335107F66181E" box="[1061,1112,1564,1589]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA51A35107AA21872" collectingDate="2000-07-19" collectionCode="IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Teresina" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
idem,
<date id="C26A60FEFFCEA31CA5C435107E7D181E" box="[1202,1347,1564,1589]" pageId="9" pageNumber="280" value="2000-07-19">
<collectingDate id="D22E9916FFCEA31CA5C435107E7D181E" box="[1202,1347,1564,1589]" pageId="9" pageNumber="280" value="2000-07-19">19 July 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA42735107A3E1872" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA42735107E95181E" box="[1361,1451,1564,1589]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1764
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA066354C7A681872" box="[272,342,1600,1625]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA017354C7AAA1872" box="[353,404,1600,1625]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA0DE354C7F1C1872" box="[424,1058,1600,1625]" collectingDate="2000-07-19" collectionCode="F, IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Teresina" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
idem,
<date id="C26A60FEFFCEA31CA099354C79BF1872" box="[495,641,1600,1625]" pageId="9" pageNumber="280" value="2000-07-19">
<collectingDate id="D22E9916FFCEA31CA099354C79BF1872" box="[495,641,1600,1625]" pageId="9" pageNumber="280" value="2000-07-19">19 July 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA3F9354C78571872" box="[655,873,1600,1625]" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA3F9354C79D71872" box="[655,745,1600,1625]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1797
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA20F354C78B41872" box="[889,906,1600,1625]" pageId="9" pageNumber="280">F</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA2E0354C78E41872" box="[918,986,1600,1625]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA290354C7F271872" box="[998,1049,1600,1625]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFCEA31CA558354C7AA41856" collectingDate="2000-08-16" collectionCode="F, IBGE, SPF" collectorName="Munhoz" country="Brazil" elevation="1488" latitude="-14.072695" location="Teresina" longLatPrecision="1" longitude="-47.509335" municipality="Fazenda Agua Fria" pageId="9" pageNumber="280" specimenCount="1" stateProvince="Goias">
idem,
<date id="C26A60FEFFCEA31CA502354C7E141872" box="[1140,1322,1600,1625]" pageId="9" pageNumber="280" value="2000-08-16">
<collectingDate id="D22E9916FFCEA31CA502354C7E141872" box="[1140,1322,1600,1625]" pageId="9" pageNumber="280" value="2000-08-16">16 August 2000</collectingDate>
</date>
,
<emphasis id="84A09A2CFFCEA31CA443354C7BDA1856" italics="true" pageId="9" pageNumber="280">
<collectorName id="1B2123E8FFCEA31CA443354C7EB11872" box="[1333,1423,1600,1625]" pageId="9" pageNumber="280">Munhoz</collectorName>
et al. 1831
</emphasis>
(
<collectionCode id="D0C5DEFBFFCEA31CA18535687A3A1856" box="[243,260,1636,1661]" pageId="9" pageNumber="280">F</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA07935687A6D1856" box="[271,339,1636,1661]" pageId="9" pageNumber="280">IBGE</collectionCode>
,
<collectionCode id="D0C5DEFBFFCEA31CA02935687AAC1856" box="[351,402,1636,1661]" pageId="9" pageNumber="280">SPF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,835 @@
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<mods:title id="F257B4B583B5D5242AAD9040837ED740">Two new and endangered species of Syngonanthus (Eriocaulaceae) from Chapada dos Veadeiros, Goiás, Brazil</mods:title>
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<subSubSection id="FECE15B5FFC5A317A1FE375878BD1A44" box="[136,899,1108,1135]" pageId="2" pageNumber="273" type="nomenclature">
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<taxonomicName id="71D43DBDFFC5A317A1FE3758781F1A44" ID-CoL="7B8J8" authority="M.T.C.Watan. &amp; Echternacht" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[136,801,1108,1135]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="incurvifolius" status="sp. nov.">
<emphasis id="84A09A2CFFC5A317A1FE37587AF51A44" bold="true" box="[136,459,1108,1135]" italics="true" pageId="2" pageNumber="273">Syngonanthus incurvifolius</emphasis>
M.T.C.Watan. &amp; Echternacht
</taxonomicName>
,
<emphasis id="84A09A2CFFC5A317A25B375A78BD1A44" box="[813,899,1110,1135]" italics="true" pageId="2" pageNumber="273">
<taxonomicNameLabel id="9F932757FFC5A317A25B375A78BD1A44" box="[813,899,1110,1135]" pageId="2" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC5A317A2FD375A79171AF0" pageId="2" pageNumber="273" type="materials_examined">
<paragraph id="B66B463EFFC5A317A2FD375A79171AF0" blockId="2.[136,1452,1108,1243]" pageId="2" pageNumber="273">
Type:—
<materialsCitation id="06BC4C63FFC5A317A290375A7E251A9C" collectingDate="1969-03-19" collectionCode="UB" collectorName="H. S. Irwin &amp; R. Reis dos Santos &amp; R. Souza &amp; S. F. da Fonseca" country="Brazil" elevation="1000" location="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" municipality="Alto Paraiso de Goias" pageId="2" pageNumber="273" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
<collectingCountry id="CEC306AEFFC5A317A290375A7F711A44" box="[998,1103,1110,1135]" name="Brazil" pageId="2" pageNumber="273">BRAZIL</collectingCountry>
.
<collectingRegion id="741088DCFFC5A317A52D375A7F9F1A44" box="[1115,1185,1110,1135]" country="Brazil" name="Goias" pageId="2" pageNumber="273">Goiás</collectingRegion>
:
<collectingMunicipality id="560FDC44FFC5A317A5DD375A7E991A44" box="[1195,1447,1110,1135]" pageId="2" pageNumber="273">Alto Paraíso de Goiás</collectingMunicipality>
,
<location id="B30B10E5FFC5A317A1FE3776782B1AB8" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC5A317A1FE3776782B1AB8" box="[136,789,1146,1171]" country="Brazil" municipality="Alto Paraiso de Goias" name="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" pageId="2" pageNumber="273" stateProvince="Goias">
ca.
<quantity id="712CEBDBFFC5A317A1D937767BC81AB8" box="[175,246,1146,1171]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="273" unit="km" value="15.0">15 km</quantity>
south of Veadeiros, road to São João da Aliança
</location>
, cerrado on outcrops, in wet sand, adjacent campo,
<quantity id="712CEBDBFFC5A317A42137767E921AB8" box="[1367,1452,1146,1171]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="273" unit="m" value="1000.0">
<elevation id="3DF9A10DFFC5A317A42137767E921AB8" box="[1367,1452,1146,1171]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="273" unit="m" value="1000.0">1000 m</elevation>
</quantity>
alt.,
<date id="C26A60FEFFC5A317A1CF37927A561A9C" box="[185,360,1182,1207]" pageId="2" pageNumber="273" value="1969-03-19">
<collectingDate id="D22E9916FFC5A317A1CF37927A561A9C" box="[185,360,1182,1207]" pageId="2" pageNumber="273" value="1969-03-19">19 March 1969</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC5A317A00537927AD11A9C" box="[371,495,1182,1207]" italics="true" pageId="2" pageNumber="273">
<collectorName id="1B2123E8FFC5A317A00537927AD11A9C" box="[371,495,1182,1207]" pageId="2" pageNumber="273">H. S. Irwin</collectorName>
</emphasis>
,
<emphasis id="84A09A2CFFC5A317A08B379279F31A9C" box="[509,717,1182,1207]" italics="true" pageId="2" pageNumber="273">
<collectorName id="1B2123E8FFC5A317A08B379279F31A9C" box="[509,717,1182,1207]" pageId="2" pageNumber="273">R. Reis dos Santos</collectorName>
</emphasis>
,
<emphasis id="84A09A2CFFC5A317A3AD37927F561A9C" box="[731,1128,1182,1207]" italics="true" pageId="2" pageNumber="273">
<collectorName id="1B2123E8FFC5A317A3AD379278031A9C" box="[731,829,1182,1207]" pageId="2" pageNumber="273">R. Souza</collectorName>
&amp;
<collectorName id="1B2123E8FFC5A317A21737927F251A9C" box="[865,1051,1182,1207]" pageId="2" pageNumber="273">S. F. da Fonseca</collectorName>
24648
</emphasis>
(
<typeStatus id="696FF89CFFC5A317A50137927FDF1A9C" box="[1143,1249,1182,1207]" pageId="2" pageNumber="273" type="holotype">Holotype</typeStatus>
<collectionCode id="D0C5DEFBFFC5A317A59E37927E281A9C" box="[1256,1302,1182,1207]" pageId="2" pageNumber="273">UB</collectionCode>
!
</materialsCitation>
,
<materialsCitation id="06BC4C63FFC5A317A45137927E9B1A9C" box="[1319,1445,1182,1207]" collectingDate="1969-03-19" collectionCode="F" collectorName="H. S. Irwin &amp; R. Reis dos Santos &amp; R. Souza &amp; S. F. da Fonseca" country="Brazil" elevation="1000" location="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" municipality="Alto Paraiso de Goias" pageId="2" pageNumber="273" specimenCount="1" stateProvince="Goias" typeStatus="isotype">
<typeStatus id="696FF89CFFC5A317A45137927EBA1A9C" box="[1319,1412,1182,1207]" pageId="2" pageNumber="273" type="isotype">Isotypes</typeStatus>
<collectionCode id="D0C5DEFBFFC5A317A4FD37927E9F1A9C" box="[1419,1441,1182,1207]" pageId="2" pageNumber="273">F</collectionCode>
!
</materialsCitation>
,
<materialsCitation id="06BC4C63FFC5A317A1FE37CE7BFF1AF0" box="[136,193,1218,1243]" collectingDate="1969-03-19" collectionCode="MO" collectorName="H. S. Irwin &amp; R. Reis dos Santos &amp; R. Souza &amp; S. F. da Fonseca" country="Brazil" elevation="1000" location="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" municipality="Alto Paraiso de Goias" pageId="2" pageNumber="273" specimenCount="1" stateProvince="Goias" typeStatus="isotype">
<collectionCode id="D0C5DEFBFFC5A317A1FE37CE7B831AF0" box="[136,189,1218,1243]" pageId="2" pageNumber="273">MO</collectionCode>
!
</materialsCitation>
,
<materialsCitation id="06BC4C63FFC5A317A1BA37CE7BC11AF0" box="[204,255,1218,1243]" collectingDate="1969-03-19" collectionCode="NY" collectorName="H. S. Irwin &amp; R. Reis dos Santos &amp; R. Souza &amp; S. F. da Fonseca" country="Brazil" elevation="1000" location="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" municipality="Alto Paraiso de Goias" pageId="2" pageNumber="273" specimenCount="1" stateProvince="Goias" typeStatus="isotype">
<collectionCode id="D0C5DEFBFFC5A317A1BA37CE7BC41AF0" box="[204,250,1218,1243]" pageId="2" pageNumber="273">NY</collectionCode>
!
</materialsCitation>
<materialsCitation id="06BC4C63FFC5A317A07337CE7A091AF0" box="[261,311,1218,1243]" collectingDate="1969-03-19" collectionCode="NY" collectorName="H. S. Irwin &amp; R. Reis dos Santos &amp; R. Souza &amp; S. F. da Fonseca" country="Brazil" elevation="1000" location="ca. 15 km south of Veadeiros, road to Sao Joao da Alianca" municipality="Alto Paraiso de Goias" pageId="2" pageNumber="273" specimenCount="1" stateProvince="Goias" typeStatus="isotype">RB!</materialsCitation>
).
<figureCitation id="2EEF5ABBFFC5A317A03D37CE7ACE1AF0" box="[331,496,1218,1243]" captionStart="FIGURE 2" captionStartId="4.[136,229,1920,1941]" captionTargetBox="[224,1358,207,1849]" captionTargetId="figure-18@4.[214,1373,190,1895]" captionTargetPageId="4" captionText="FIGURE 2AG. Syngonanthus incurvifolius (based on Irwin et al. 24648). A. Habit; B. Capitulum detail; C. From outermost to innermost series of involucral bracts; D. Staminate flower; E. Various forms of sepals of staminate flower; F. Pistillate flower with one sepal removed; G. Gynoecium. HN. S. vittatus (based on Watanabe &amp; Watanabe 377). H. Habit; I. Capitulum detail; J. From outermost to innermost series of involucral bracts; K. Staminate flower; L. Various forms of sepals of staminate flower; M. Pistillate flower with one sepal removed; N. Gynoecium. Illustrated by Marcelo Kubo." figureDoi="http://doi.org/10.5281/zenodo.13637542" httpUri="https://zenodo.org/record/13637542/files/figure.png" pageId="2" pageNumber="273">Figures 2 AG</figureCitation>
&amp;
<figureCitation id="2EEF5ABBFFC5A317A36237CE791A1AF0" box="[532,548,1218,1243]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="2" pageNumber="273">3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC5A317A1FE36007F831BA6" pageId="2" pageNumber="273" type="diagnosis">
<paragraph id="B66B463EFFC5A317A1FE36007F831BA6" blockId="2.[136,1452,1292,1421]" pageId="2" pageNumber="273">
<taxonomicName id="71D43DBDFFC5A317A1FE36007AB01B0A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[136,398,1292,1313]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC5A317A1FE36007AB01B0A" box="[136,398,1292,1313]" italics="true" pageId="2" pageNumber="273">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
differs from all other New World species of
<taxonomicName id="71D43DBDFFC5A317A234360078F71B0A" authorityName="Ruhland" authorityYear="1900" box="[834,969,1292,1313]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC5A317A234360078F71B0A" box="[834,969,1292,1313]" italics="true" pageId="2" pageNumber="273">Syngonanthus</emphasis>
</taxonomicName>
by the rosette leaves incurved from the middle to the apex. It differs from
<taxonomicName id="71D43DBDFFC5A317A0C6363C79A41B6E" authorityName="Ruhland" authorityYear="1903" baseAuthorityName="Gardner" box="[432,666,1328,1349]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="arenarius">
<emphasis id="84A09A2CFFC5A317A0C6363C79A41B6E" box="[432,666,1328,1349]" italics="true" pageId="2" pageNumber="273">Syngonanthus arenarius</emphasis>
</taxonomicName>
, the most similar species, by pistillate flowers with cream sepals and adpressed trichomes on the leaves.
<taxonomicName id="71D43DBDFFC5A317A0C6365879891B42" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[432,695,1364,1385]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC5A317A0C6365879891B42" box="[432,695,1364,1385]" italics="true" pageId="2" pageNumber="273">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="71D43DBDFFC5A317A21136587FF91B42" authorityName="Moldenke" authorityYear="1972" box="[871,1223,1364,1385]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="273" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC5A317A21136587F621B42" box="[871,1116,1364,1385]" italics="true" pageId="2" pageNumber="273">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC5A317A5F836587FF91B42" box="[1166,1223,1364,1385]" italics="true" pageId="2" pageNumber="273">majus</emphasis>
</taxonomicName>
but differs by pistillate flowers with pilose petals, sepals cream with a light castaneous longitudinal band, and individuals smaller.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC5A316A1FE36B17F3F1DC9" lastPageId="3" lastPageNumber="274" pageId="2" pageNumber="273" type="description">
<paragraph id="B66B463EFFC5A316A1FE36B17F3F1DC9" blockId="2.[136,1452,1469,2092]" lastBlockId="3.[136,1452,160,2051]" lastPageId="3" lastPageNumber="274" pageId="2" pageNumber="273">
Perennial herbs, 7.518.0 cm tall.
<emphasis id="84A09A2CFFC5A317A36036B179681BFD" box="[534,598,1469,1494]" italics="true" pageId="2" pageNumber="273">Roots</emphasis>
<quantity id="712CEBDBFFC5A317A32936B179C91BFD" box="[607,759,1469,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.4" metricValueMin="0.75" pageId="2" pageNumber="273" unit="mm" value="1.075" valueMax="1.4" valueMin="0.75">0.751.4 mm</quantity>
in diameter, cream-colored, spongy.
<emphasis id="84A09A2CFFC5A317A5D236B17E3F1BFD" box="[1188,1281,1469,1494]" italics="true" pageId="2" pageNumber="273">Rosettes</emphasis>
<quantity id="712CEBDBFFC5A317A47A36B17EB51BFD" box="[1292,1419,1469,1494]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="3.4" metricValueMin="1.7" pageId="2" pageNumber="273" unit="cm" value="2.55" valueMax="3.4" valueMin="1.7">1.73.4 cm</quantity>
in diameter.
<emphasis id="84A09A2CFFC5A317A18F36EC7A671BD2" box="[249,345,1504,1529]" italics="true" pageId="2" pageNumber="273">Rhizome</emphasis>
1.02.5(5) cm ×
<quantity id="712CEBDBFFC5A317A34436EC79831BD2" box="[562,701,1504,1529]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.5" metricValueMin="2.5" pageId="2" pageNumber="273" unit="mm" value="5.0" valueMax="7.5" valueMin="2.5">2.57.5 mm</quantity>
, horizontal to oblique, often branched, tomentose to villous, with white, dense and interwoven trichomes ca.
<quantity id="712CEBDBFFC5A317A309350F79871837" box="[639,697,1539,1564]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="273" unit="cm" value="1.0">1 cm</quantity>
long; each branch giving rise to a rosette of leaves, each rosette bearing 15 synflorescences.
<emphasis id="84A09A2CFFC5A317A0A5352A791F1814" box="[467,545,1574,1599]" italics="true" pageId="2" pageNumber="273">Leaves</emphasis>
0.52.0 cm ×
<quantity id="712CEBDBFFC5A317A3B7352A78581814" box="[705,870,1574,1599]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="12.0" metricValueMin="7.5" pageId="2" pageNumber="273" unit="mm" value="0.975" valueMax="1.2" valueMin="0.75">0.751.20 mm</quantity>
, linear, flat to semi-terete, apex acute to round, base truncate, sinuous, recurved at base, becoming incurved from the middle to the apex, firm to rigid, striate adaxially with 3-veins prominent, or only the midvein perceptible in young leaves; densely pilose adaxially, glabrescent to pilose abaxially, with adpressed trichomes, and patent, simple trichomes ca.
<quantity id="712CEBDBFFC5A317A29E359C7F011882" box="[1000,1087,1680,1705]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="273" unit="mm" value="0.5">0.5 mm</quantity>
long; basal sheath not enlarged.
<emphasis id="84A09A2CFFC5A317A1FE35BF7A0418E7" box="[136,314,1715,1740]" italics="true" pageId="2" pageNumber="273">Synflorescences</emphasis>
with a primary axis bearing a lax umbel of 118 inflorescences at the apex.
<emphasis id="84A09A2CFFC5A317A5EC35BF7E4418E7" box="[1178,1402,1715,1740]" italics="true" pageId="2" pageNumber="273">Synflorescence axes</emphasis>
<quantity id="712CEBDBFFC5A317A4F735BF7B9418C4" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="2" pageNumber="273" unit="cm" value="5.0" valueMax="8.0" valueMin="2.0">28 cm</quantity>
long, 0.21.0 mm in diameter, emerging from the center of the rosette, erect to curved, brown, sometimes with a dark-reddish tinge, glabrescent to pilose with simple filamentous trichomes ca.
<quantity id="712CEBDBFFC5A317A57635F67F6B1938" box="[1024,1109,1786,1811]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="273" unit="mm" value="0.3">0.3 mm</quantity>
long.
<emphasis id="84A09A2CFFC5A317A5ED35F67B8B191D" italics="true" pageId="2" pageNumber="273">Bracts of synflorescence axis</emphasis>
restricted to the apex, 0.23.0 cm ×
<quantity id="712CEBDBFFC5A317A32B34117989191D" box="[605,695,1821,1846]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="273" unit="mm" value="0.5">0.5 mm</quantity>
, whorled and numerous, patent, progressively elongating from the outer toward the inner series, linear to narrowly lanceolate, apex acute, erect to uncinate, pubescent on both surfaces with adpressed trichomes, and patent, capitate and simple, filamentous trichomes, capitate trichomes mostly on abaxial surface, simple trichomes concentrated at margins.
<emphasis id="84A09A2CFFC5A317A3C8348A786119B4" box="[702,863,1926,1951]" italics="true" pageId="2" pageNumber="273">Inflorescences</emphasis>
composed of a closed spathe subtending a scape with a capitulum at the apex.
<emphasis id="84A09A2CFFC5A317A0D634A67AC919E8" box="[416,503,1962,1987]" italics="true" pageId="2" pageNumber="273">Spathes</emphasis>
<quantity id="712CEBDBFFC5A317A37634A6796A19E8" box="[512,596,1962,1987]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="2" pageNumber="273" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">14 cm</quantity>
long, chartaceous, cylindrical, lax at apex, obliquely opened, apex acute to obtuse, erect to recurved, tip erect to uncinate, margin entire, not involute, glabrous adaxially, pubescent abaxially with capitate and simple, filamentous trichomes.
<emphasis id="84A09A2CFFC5A317A30834FC79F21622" box="[638,716,2032,2057]" italics="true" pageId="2" pageNumber="273">Scapes</emphasis>
erect,
<quantity id="712CEBDBFFC5A317A26C34FC78431622" box="[794,893,2032,2057]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="2" pageNumber="273" unit="cm" value="7.5" valueMax="12.0" valueMin="3.0">312 cm</quantity>
long, green to straw-colored with age, 3-costate, pilose, pubescent near the apex with erect to patent, simple and capitate trichomes.
<emphasis id="84A09A2CFFC5A317A5263B1F7F8C1607" box="[1104,1202,2067,2092]" italics="true" pageId="2" pageNumber="273">Capitula</emphasis>
<quantity id="712CEBDBFFC5A317A5CD3B1F7E251607" box="[1211,1307,2067,2092]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="2" pageNumber="273" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">48 mm</quantity>
in diameter, radiate.
<emphasis id="84A09A2CFFC4A316A19233AC7A961E92" box="[228,424,160,185]" italics="true" pageId="3" pageNumber="274">Involucral bracts</emphasis>
in 57 series, castaneous, reddish-brown to brown, sometimes with a lighter stramineous longitudinal band in the middle and on the margins, glabrous on adaxial surface, pilose to glabrescent abaxially, ciliate, with simple filamentous trichomes, rarely also capitate trichomes, these concentrated at the apex, ca.
<quantity id="712CEBDBFFC4A316A46733EA7E571ED4" box="[1297,1385,230,255]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="274" unit="mm" value="0.3">0.3 mm</quantity>
long; external series 1.01.8 ×
<quantity id="712CEBDBFFC4A316A0D2320579131F09" box="[420,557,265,290]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="274" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
, triangular to ovate, apex acute, progressively longer toward the internal series, middle series 2.53.0 ×
<quantity id="712CEBDBFFC4A316A0EB322179161F6D" box="[413,552,301,326]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="3" pageNumber="274" unit="mm" value="1.05" valueMax="1.2" valueMin="0.9">0.91.2 mm</quantity>
, obovate, apex acute to obtuse, internal series 2.53.0 ×
<quantity id="712CEBDBFFC4A316A5C532217E001F6D" box="[1203,1342,301,326]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="3" pageNumber="274" unit="mm" value="0.5" valueMax="0.7" valueMin="0.3">0.30.7 mm</quantity>
, equaling the flower height, elliptic to slightly oblanceolate, apex acute to obtuse, sometimes membranous, lighter-colored than the external series; receptacle obconic, pilose.
<emphasis id="84A09A2CFFC4A316A3D4327F78B21FA7" box="[674,908,371,396]" italics="true" pageId="3" pageNumber="274">Floral bracts absent</emphasis>
.
<emphasis id="84A09A2CFFC4A316A2EB327F78C71FA7" box="[925,1017,371,396]" italics="true" pageId="3" pageNumber="274">Flowers</emphasis>
3-merous, 40100 per capitulum, the staminate and pistillate generally equal in number, with contrasting colors, staminate flowers darker and concentrated at the border of the capitulum or mixed.
<emphasis id="84A09A2CFFC4A316A33F32B5782F1FF9" box="[585,785,441,466]" italics="true" pageId="3" pageNumber="274">Staminate flowers</emphasis>
with pedicels 0.51.0 mm long, pilose in the bottom part of pedicel with simple trichomes ca.
<quantity id="712CEBDBFFC4A316A37232D1797B1FDD" box="[516,581,477,502]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="274" unit="mm" value="1.0">1 mm</quantity>
long; sepals 1.52.0 mm long, coppery brown, shortly fused at base, concave, membranous, the sepals slightly different from each other, or two sepals similar to each other and a third different, falcate-asymmetrical, oblanceolate, elliptic or irregular-subspathulate, apex mucronate, cuspidate, rounded or obtuse to acute, glabrous adaxially, pubescent abaxially, especially at apex and margin, with simple filamentous trichomes ca.
<quantity id="712CEBDBFFC4A316A1C631657A391CA9" box="[176,263,617,642]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="274" unit="mm" value="0.4">0.4 mm</quantity>
long; petals
<quantity id="712CEBDBFFC4A316A0EF31657AC81CA9" box="[409,502,617,642]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="274" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, hyaline, membranous, fused, tubulose, lobes obtuse to rounded, involute after anthesis, glabrous; stamens adnate to the petals at base, anthers ca.
<quantity id="712CEBDBFFC4A316A21E318178831C8D" box="[872,957,653,678]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="274" unit="mm" value="0.3">0.3 mm</quantity>
long, cream to yellowish; anthophore ca.
<quantity id="712CEBDBFFC4A316A4FF31817B8A1CE2" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="274" unit="mm" value="0.3">0.3 mm</quantity>
long; pistillodes 3, ca.
<quantity id="712CEBDBFFC4A316A0C231BC79341CE2" box="[436,522,688,713]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="274" unit="mm" value="0.3">0.3 mm</quantity>
long, clavate, papillose at apex.
<emphasis id="84A09A2CFFC4A316A20531BC7F0E1CE2" box="[883,1072,688,713]" italics="true" pageId="3" pageNumber="274">Pistillate flowers</emphasis>
<quantity id="712CEBDBFFC4A316A54131BC7FAD1CE2" box="[1079,1171,688,713]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="274" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, including pedicels; pedicels 0.41.0 mm long, with filamentous trichomes at insertion with receptacle, actinomorphic; sepals equaling the petal height, 1.22.0 mm long, oblong to widely ovate, concave, apex acute, base thickened, free, white with a light castaneous longitudinal band, adaxially glabrous, abaxially pubescent near the apex, ciliate, with simple filamentous trichomes up to
<quantity id="712CEBDBFFC4A316A04B30307AAD1D7E" box="[317,403,828,853]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="274" unit="mm" value="0.5">0.5 mm</quantity>
long; petals about equaling the sepals,
<quantity id="712CEBDBFFC4A316A23F303078981D7E" box="[841,934,828,853]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="274" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, linear to oblong, fused at the upper half, free at base and at the very top, lobe triangular, apex acute, membranous, white to hyaline when hydrated, pilose with simple filamentous trichomes up to
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; gynoecium ca.
<quantity id="712CEBDBFFC4A316A25D308F78521DB7" box="[811,876,899,924]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="274" unit="mm" value="2.0">2 mm</quantity>
long; ovary
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long; style column ca.
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long, cream-colored to castaneous; appendages ca.
<quantity id="712CEBDBFFC4A316A38B30AA786A1D94" box="[765,852,934,959]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="274" unit="mm" value="0.4">0.4 mm</quantity>
long, the glandular apex capitate; stigmatic branches simple, ca.
<quantity id="712CEBDBFFC4A316A07E30C57A601DC9" box="[264,350,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="274" unit="mm" value="1.5">1.5 mm</quantity>
long.
<emphasis id="84A09A2CFFC4A316A0D230C57AE71DC9" box="[420,473,969,994]" italics="true" pageId="3" pageNumber="274">Seed</emphasis>
0.40.6 ×
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, ellipsoid, brown-colored.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC4A316A1CB30E0796E1A67" pageId="3" pageNumber="274" type="etymology">
<paragraph id="B66B463EFFC4A316A1CB30E0796E1A67" blockId="3.[136,1452,160,2051]" pageId="3" pageNumber="274">
<emphasis id="84A09A2CFFC4A316A1CB30E07A5A1A2E" bold="true" box="[189,356,1004,1029]" pageId="3" pageNumber="274">Etymology:—</emphasis>
The specific epithet refers to the orientation of the rosette leaves, which are curved inward and upward at the distal middle, a unique feature within Brazilian
<taxonomicName id="71D43DBDFFC4A316A249371C78E21A02" authorityName="Ruhland" authorityYear="1900" box="[831,988,1040,1065]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC4A316A249371C78E21A02" box="[831,988,1040,1065]" italics="true" pageId="3" pageNumber="274">Syngonanthus</emphasis>
</taxonomicName>
. Only
<taxonomicName id="71D43DBDFFC4A316A55F371C7A3A1A67" authority="Ruhland (1903: 247)" authorityName="Ruhland" authorityPageNumber="247" authorityYear="1903" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="wahlbergii">
<emphasis id="84A09A2CFFC4A316A55F371C7E7B1A02" box="[1065,1349,1040,1065]" italics="true" pageId="3" pageNumber="274">Syngonanthus wahlbergii</emphasis>
<bibRefCitation id="D2453BCFFFC4A316A43A371C7A3A1A67" author="Ruhland, W." pageId="3" pageNumber="274" pagination="1 - 294" refId="ref8150" refString="Ruhland, W. (1903) Eriocaulaceae. In: Engler, A. (Ed.) Das Pflanzenreich IV, 30 (Heft 13). W. Engelmann, Leipzig, pp. 1 - 294." type="book chapter" year="1903">Ruhland (1903:247)</bibRefCitation>
</taxonomicName>
in Africa has leaves like this.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC4A313A1CB375A78831F46" lastPageId="6" lastPageNumber="277" pageId="3" pageNumber="274" type="discussion">
<paragraph id="B66B463EFFC4A316A1CB375A7BC01B87" blockId="3.[136,1452,160,2051]" pageId="3" pageNumber="274">
<emphasis id="84A09A2CFFC4A316A1CB375A79721A44" bold="true" box="[189,588,1110,1135]" pageId="3" pageNumber="274">Notes and taxonomic affinities:—</emphasis>
Several species share a similar architecture with
<taxonomicName id="71D43DBDFFC4A316A505375A7E9B1A44" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1139,1445,1110,1135]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A505375A7E9B1A44" box="[1139,1445,1110,1135]" italics="true" pageId="3" pageNumber="274">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
, including: a rhizome with apical rosettes of leaves, bearing synflorescences with verticillate bracts at the apex and scapes arranged in lax umbels. The following taxa also share dark and pilose involucral bracts (
<figureCitation id="2EEF5ABBFFC4A316A59037907E7D1A9E" box="[1254,1347,1180,1205]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="3" pageNumber="274">Fig. 3H</figureCitation>
), usually with a lighter longitudinal band abaxially:
<taxonomicName id="71D43DBDFFC4A316A31137CC7FE71AF2" authority="(Gardner) Ruhland (1903: 260)" authorityName="Ruhland" authorityPageNumber="260" authorityYear="1903" baseAuthorityName="Gardner" box="[615,1241,1216,1241]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="arenarius">
<emphasis id="84A09A2CFFC4A316A31137CC78461AF2" box="[615,888,1216,1241]" italics="true" pageId="3" pageNumber="274">Syngonanthus arenarius</emphasis>
(Gardner)
<bibRefCitation id="D2453BCFFFC4A316A28237CC7FE71AF2" author="Ruhland, W." box="[1012,1241,1216,1241]" pageId="3" pageNumber="274" pagination="1 - 294" refId="ref8150" refString="Ruhland, W. (1903) Eriocaulaceae. In: Engler, A. (Ed.) Das Pflanzenreich IV, 30 (Heft 13). W. Engelmann, Leipzig, pp. 1 - 294." type="book chapter" year="1903">Ruhland (1903:260)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71D43DBDFFC4A316A59337CC79121AD7" authority="Moldenke (1984: 372)" authorityName="Moldenke" authorityPageNumber="372" authorityYear="1984" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC4A316A59337CC7E451AF2" box="[1253,1403,1216,1241]" italics="true" pageId="3" pageNumber="274">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A1FE37EF7A191AD7" box="[136,295,1251,1276]" italics="true" pageId="3" pageNumber="274">brachyphyllus</emphasis>
<bibRefCitation id="D2453BCFFFC4A316A05837EF79121AD7" author="Moldenke, H. N." box="[302,556,1251,1276]" pageId="3" pageNumber="274" refId="ref7727" refString="Moldenke, H. N. (1984) Notes on new and noteworthy plants CLXXVI. Phytologia 55: 372." type="journal volume" year="1984">Moldenke (1984: 372)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71D43DBDFFC4A316A34137EF7F7A1AD7" authority="Moldenke (1972: 499)" authorityName="Moldenke" authorityPageNumber="499" authorityYear="1972" box="[567,1092,1251,1276]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC4A316A34137EF79F21AD7" box="[567,716,1251,1276]" italics="true" pageId="3" pageNumber="274">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A27237EF78791AD7" box="[772,839,1251,1276]" italics="true" pageId="3" pageNumber="274">majus</emphasis>
<bibRefCitation id="D2453BCFFFC4A316A23B37EF7F7A1AD7" author="Moldenke, H. N." box="[845,1092,1251,1276]" pageId="3" pageNumber="274" pagination="498 - 499" refId="ref7679" refString="Moldenke, H. N. (1972) Four novelties from Brazil. Phytologia 24: 498 - 499." type="journal article" year="1972">Moldenke (1972:499)</bibRefCitation>
</taxonomicName>
, and
<emphasis id="84A09A2CFFC4A316A5F637EF7FCB1AD7" box="[1152,1269,1251,1276]" italics="true" pageId="3" pageNumber="274">
<taxonomicName id="71D43DBDFFC4A316A5F637EF7FCE1AD7" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1152,1264,1251,1276]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="vittatus">S. vittatus</taxonomicName>
,
</emphasis>
the new species described here below.
<taxonomicName id="71D43DBDFFC4A316A0E6360A780A1B34" authorityName="Moldenke" authorityYear="1972" box="[400,820,1286,1311]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC4A316A0E6360A798C1B34" box="[400,690,1286,1311]" italics="true" pageId="3" pageNumber="274">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A387360A780A1B34" box="[753,820,1286,1311]" italics="true" pageId="3" pageNumber="274">majus</emphasis>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFC4A316A205360A7FD51B34" authorityName="Moldenke" authorityYear="1984" box="[883,1259,1286,1311]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC4A316A205360A7F321B34" box="[883,1036,1286,1311]" italics="true" pageId="3" pageNumber="274">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A53A360A7FD51B34" box="[1100,1259,1286,1311]" italics="true" pageId="3" pageNumber="274">brachyphyllus</emphasis>
</taxonomicName>
, actually do not belong to
<taxonomicName id="71D43DBDFFC4A316A18D36257AAF1B69" box="[251,401,1321,1346]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="densifolius">
<emphasis id="84A09A2CFFC4A316A18D36257AAF1B69" box="[251,401,1321,1346]" italics="true" pageId="3" pageNumber="274">S. densifolius</emphasis>
</taxonomicName>
, which possess golden to cream, glabrous involucral bracts, and shall be elevated to species status (Echternacht
<emphasis id="84A09A2CFFC4A316A01F36407A9A1B4E" box="[361,420,1356,1381]" italics="true" pageId="3" pageNumber="274">et al.</emphasis>
in prep.). We consider these five taxa to form a cohesive morphological group due to the characters mentioned above. All of them, except
<taxonomicName id="71D43DBDFFC4A316A3C4367C78051BA2" authorityName="Ruhland" authorityYear="1903" baseAuthorityName="Gardner" box="[690,827,1392,1417]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="arenarius">
<emphasis id="84A09A2CFFC4A316A3C4367C78051BA2" box="[690,827,1392,1417]" italics="true" pageId="3" pageNumber="274">S. arenarius</emphasis>
</taxonomicName>
, occur in Chapada dos Veadeiros and they are possibly sympatric.
</paragraph>
<paragraph id="B66B463EFFC4A316A1CB36BA7ABA183E" blockId="3.[136,1452,160,2051]" pageId="3" pageNumber="274">
<taxonomicName id="71D43DBDFFC4A316A1CB36BA7AD11BE4" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[189,495,1462,1487]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A1CB36BA7AD11BE4" box="[189,495,1462,1487]" italics="true" pageId="3" pageNumber="274">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
also shares with these four taxa an unusual floral trait: sepals of staminate flowers are always clearly asymmetrical (
<figureCitation id="2EEF5ABBFFC4A316A0A436D579601BD9" box="[466,606,1497,1522]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="3" pageNumber="274">FIG. 3D; 3F</figureCitation>
). The three sepals might differ
<taxonomicName id="71D43DBDFFC4A316A2C236D57F0A1BD9" box="[948,1076,1497,1522]" class="Liliopsida" family="Eriocaulaceae" form="and" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="form" species="incurvifolius">in form and</taxonomicName>
curvature or two might be similar and the third different.
</paragraph>
<paragraph id="B66B463EFFC4A316A1CB352C7E291628" blockId="3.[136,1452,160,2051]" pageId="3" pageNumber="274">
<taxonomicName id="71D43DBDFFC4A316A1CB352C7ACA1812" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[189,500,1568,1593]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A1CB352C7ACA1812" box="[189,500,1568,1593]" italics="true" pageId="3" pageNumber="274">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
differs from the congeneric species that have synflorescences by the remarkable form of the rosette leaves, which are recurved at base and incurved at the upper half. This feature persists even after diaphanization.
<taxonomicName id="71D43DBDFFC4A316A04B356A78081854" authorityName="Moldenke" authorityYear="1984" box="[317,822,1638,1663]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC4A316A04B356A79651854" box="[317,603,1638,1663]" italics="true" pageId="3" pageNumber="274">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A3E1356A78081854" box="[663,822,1638,1663]" italics="true" pageId="3" pageNumber="274">brachyphyllus</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="71D43DBDFFC4A316A5CE356A7E5A1854" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1208,1380,1638,1663]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A5CE356A7E5A1854" box="[1208,1380,1638,1663]" italics="true" pageId="3" pageNumber="274">S. incurvifolius</emphasis>
</taxonomicName>
by its broader leaves (ca. 25
<emphasis id="84A09A2CFFC4A316A0E035857A8A1889" box="[406,436,1673,1698]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
<quantity id="712CEBDBFFC4A316A0CA3585790D1889" box="[444,563,1673,1698]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="3" pageNumber="274" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.52 mm</quantity>
), with arachnoid trichomes on the adaxial surface (
<emphasis id="84A09A2CFFC4A316A50235857FAD1889" box="[1140,1171,1673,1698]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
not arachnoid), which is reticulate (
<emphasis id="84A09A2CFFC4A316A18935A07A2018EE" box="[255,286,1708,1733]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
striate), and leaf bases achlorophyllous and pale (
<emphasis id="84A09A2CFFC4A316A23035A0785A18EE" box="[838,868,1708,1733]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
chlorophyllous and green) (
<figureCitation id="2EEF5ABBFFC4A316A5EA35A07FC518EE" box="[1180,1275,1708,1733]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="3" pageNumber="274">FIG. 3C</figureCitation>
).
<taxonomicName id="71D43DBDFFC4A316A47835A07A4018C2" authorityName="Moldenke" authorityYear="1972" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC4A316A47835A07A3F18C2" italics="true" pageId="3" pageNumber="274">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A04D35DC7A4018C2" box="[315,382,1744,1769]" italics="true" pageId="3" pageNumber="274">majus</emphasis>
</taxonomicName>
differs by pistillate flowers with glabrous petals (
<emphasis id="84A09A2CFFC4A316A2C335DC78ED18C2" box="[949,979,1744,1769]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
pilose), and darker sepals, light-brown at the upper half (
<figureCitation id="2EEF5ABBFFC4A316A04135FF7AFE1927" box="[311,448,1779,1804]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="3" pageNumber="274">FIG. 3G; 3J</figureCitation>
) (
<emphasis id="84A09A2CFFC4A316A0AD35FF7AC71927" box="[475,505,1779,1804]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
cream with a longitudinal light castaneous band); moreover, individuals are bigger than
<taxonomicName id="71D43DBDFFC4A316A1B6341A7A541904" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[192,362,1814,1839]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A1B6341A7A541904" box="[192,362,1814,1839]" italics="true" pageId="3" pageNumber="274">S. incurvifolius</emphasis>
</taxonomicName>
and the rosette leaves are erect and flexuous.
<taxonomicName id="71D43DBDFFC4A316A206341A7E1B1904" authority="var. arenarius" box="[880,1317,1814,1839]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="variety" species="arenarius" variety="arenarius">
<emphasis id="84A09A2CFFC4A316A206341A7FBF1904" box="[880,1153,1814,1839]" italics="true" pageId="3" pageNumber="274">Syngonanthus arenarius</emphasis>
var.
<emphasis id="84A09A2CFFC4A316A5CF341A7E1B1904" box="[1209,1317,1814,1839]" italics="true" pageId="3" pageNumber="274">arenarius</emphasis>
</taxonomicName>
shares with
<taxonomicName id="71D43DBDFFC4A316A1FE34357A0D1979" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[136,307,1849,1874]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A1FE34357A0D1979" box="[136,307,1849,1874]" italics="true" pageId="3" pageNumber="274">S. incurvifolius</emphasis>
</taxonomicName>
the small size and pilose petals of pistillate flowers (
<figureCitation id="2EEF5ABBFFC4A316A2F834357F2A1979" box="[910,1044,1849,1874]" captionStart="FIGURE 3" captionStartId="5.[136,229,1812,1833]" captionTargetBox="[254,1333,193,1784]" captionTargetId="figure-14@5.[251,1336,190,1787]" captionTargetPageId="5" captionText="FIGURE 3. Syngonanthus incurvifolius and other morphologically similar Syngonanthus species. AC. Comparison among leaves: A. Syngonanthus densifolius var. majus showing erect, flexuous leaves in a rosette; B. Incurved leaves of S. incurvifolius; C. Leaf of S. densifolius var. brachyphyllus, showing achlorophyllous base. DG. Comparison among sexual traits of S. incurvifolius and S. densifolius var. majus: D. Staminate flower of S. incurvifolius (trichomes removed from pedicel); E. Pistillate flower of S. incurvifolius with filamentous trichomes; F. Staminate flower of S. densifolius var. majus; G. Pistillate flower of S. densifolius var. majus exhibiting trichomes only at upper portion of the sepals; H. Capitulum detail of S. incurvifolius; IJ. Comparative vestiture of the petals in pistillate flowers: I. S. incurvifolius showing pilose petals; J. Petals glabrous in S. densifolius var. majus. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe)." figureDoi="http://doi.org/10.5281/zenodo.13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="3" pageNumber="274">FIG. 3E; 3I</figureCitation>
); it differs by the cup-shaped bracts of the synflorescence (equal size), which are fewer (
<emphasis id="84A09A2CFFC4A316A39934507833195E" box="[751,781,1884,1909]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
progressively elongating from the outer toward the inner series and more numerous), and pistillate flowers with sepals cream at base and dark reddish-brown on the upper half (
<emphasis id="84A09A2CFFC4A316A1E734AF7B8E1997" box="[145,176,1955,1980]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
cream with a longitudinal light castaneous band).
<taxonomicName id="71D43DBDFFC4A316A39834AF7F1F1997" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[750,1057,1955,1980]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC4A316A39834AF7F1F1997" box="[750,1057,1955,1980]" italics="true" pageId="3" pageNumber="274">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="71D43DBDFFC4A316A59B34AF7E621997" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1261,1372,1955,1980]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFC4A316A59B34AF7E621997" box="[1261,1372,1955,1980]" italics="true" pageId="3" pageNumber="274">S. vittatus</emphasis>
</taxonomicName>
by the synflorescence axis bearing bracts only at the apex (
<emphasis id="84A09A2CFFC4A316A39234CA783C19F4" box="[740,770,1990,2015]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
tiny bracts scattered along axis), the petals of the pistillate flowers pilose (
<emphasis id="84A09A2CFFC4A316A04034E67A6B1628" box="[310,341,2026,2051]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
glabrous), and the involucral bracts of the middle series obovate (
<emphasis id="84A09A2CFFC4A316A54C34E67F671628" box="[1082,1113,2026,2051]" italics="true" pageId="3" pageNumber="274">vs.</emphasis>
narrow-elliptic).
</paragraph>
<caption id="E2AB16B6FFC3A311A1FE348C79EF160E" ID-DOI="http://doi.org/10.5281/zenodo.13637542" ID-Zenodo-Dep="13637542" httpUri="https://zenodo.org/record/13637542/files/figure.png" pageId="4" pageNumber="275" startId="4.[136,229,1920,1941]" targetBox="[224,1358,207,1849]" targetPageId="4" targetType="figure">
<paragraph id="B66B463EFFC3A311A1FE348C79EF160E" blockId="4.[136,1452,1920,2085]" pageId="4" pageNumber="275">
<emphasis id="84A09A2CFFC3A311A1FE348C7A0E19BE" bold="true" box="[136,304,1920,1941]" pageId="4" pageNumber="275">FIGURE 2 AG.</emphasis>
<taxonomicName id="71D43DBDFFC3A311A042348C790719BE" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[308,569,1920,1941]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="275" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC3A311A042348C790719BE" box="[308,569,1920,1941]" italics="true" pageId="4" pageNumber="275">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
(based on
<emphasis id="84A09A2CFFC3A311A3E8348C787819BE" box="[670,838,1920,1941]" italics="true" pageId="4" pageNumber="275">Irwin et al. 24648</emphasis>
).
<emphasis id="84A09A2CFFC3A311A22E348C784E19BE" bold="true" box="[856,880,1920,1941]" pageId="4" pageNumber="275">A.</emphasis>
Habit;
<emphasis id="84A09A2CFFC3A311A2C3348C78F519BE" bold="true" box="[949,971,1920,1941]" pageId="4" pageNumber="275">B.</emphasis>
Capitulum detail;
<emphasis id="84A09A2CFFC3A311A50D348C7FAC19BE" bold="true" box="[1147,1170,1920,1941]" pageId="4" pageNumber="275">C.</emphasis>
From outermost to innermost series of involucral bracts;
<emphasis id="84A09A2CFFC3A311A0EC34A87A8F1992" bold="true" box="[410,433,1956,1977]" pageId="4" pageNumber="275">D.</emphasis>
Staminate flower;
<emphasis id="84A09A2CFFC3A311A30434A879B71992" bold="true" box="[626,649,1956,1977]" pageId="4" pageNumber="275">E.</emphasis>
Various forms of sepals of staminate flower;
<emphasis id="84A09A2CFFC3A311A52F34A87F551992" bold="true" box="[1113,1131,1956,1977]" pageId="4" pageNumber="275">F.</emphasis>
Pistillate flower with one sepal removed;
<emphasis id="84A09A2CFFC3A311A19C34C47A3D19F6" bold="true" box="[234,259,1992,2013]" pageId="4" pageNumber="275">G.</emphasis>
Gynoecium.
<emphasis id="84A09A2CFFC3A311A0FE34C47A8019F6" bold="true" box="[392,446,1992,2013]" pageId="4" pageNumber="275">HN.</emphasis>
<taxonomicName id="71D43DBDFFC3A311A0B334C4791B19F6" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[453,549,1992,2013]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="275" phylum="Tracheophyta" rank="species" species="vittatus">
<emphasis id="84A09A2CFFC3A311A0B334C4791B19F6" box="[453,549,1992,2013]" italics="true" pageId="4" pageNumber="275">S. vittatus</emphasis>
</taxonomicName>
(based on
<emphasis id="84A09A2CFFC3A311A3E434C478A319F6" box="[658,925,1992,2013]" italics="true" pageId="4" pageNumber="275">Watanabe &amp; Watanabe 377</emphasis>
).
<emphasis id="84A09A2CFFC3A311A2C434C478F519F6" bold="true" box="[946,971,1992,2013]" pageId="4" pageNumber="275">H.</emphasis>
Habit;
<emphasis id="84A09A2CFFC3A311A56034C47F1B19F6" bold="true" box="[1046,1061,1992,2013]" pageId="4" pageNumber="275">I.</emphasis>
Capitulum detail;
<emphasis id="84A09A2CFFC3A311A5AA34C47FD119F6" bold="true" box="[1244,1263,1992,2013]" pageId="4" pageNumber="275">J.</emphasis>
From outermost to innermost series of involucral bracts;
<emphasis id="84A09A2CFFC3A311A08234E07930162A" bold="true" box="[500,526,2028,2049]" pageId="4" pageNumber="275">K.</emphasis>
Staminate flower;
<emphasis id="84A09A2CFFC3A311A3B034E079E2162A" bold="true" box="[710,732,2028,2049]" pageId="4" pageNumber="275">L.</emphasis>
Various forms of sepals of staminate flower;
<emphasis id="84A09A2CFFC3A311A5E334E07F8F162A" bold="true" box="[1173,1201,2028,2049]" pageId="4" pageNumber="275">M.</emphasis>
Pistillate flower with one sepal removed;
<emphasis id="84A09A2CFFC3A311A0563B1C7A0F160E" bold="true" box="[288,305,2064,2085]" pageId="4" pageNumber="275">N</emphasis>
. Gynoecium. Illustrated by Marcelo Kubo.
</paragraph>
</caption>
<caption id="E2AB16B6FFC2A310A1FE3418794B160E" ID-DOI="http://doi.org/10.5281/zenodo.13637544" ID-Zenodo-Dep="13637544" httpUri="https://zenodo.org/record/13637544/files/figure.png" pageId="5" pageNumber="276" startId="5.[136,229,1812,1833]" targetBox="[254,1333,193,1784]" targetPageId="5" targetType="figure">
<paragraph id="B66B463EFFC2A310A1FE3418794B160E" blockId="5.[136,1452,1812,2085]" pageId="5" pageNumber="276">
<emphasis id="84A09A2CFFC2A310A1FE34187A3F1902" bold="true" box="[136,257,1812,1833]" pageId="5" pageNumber="276">FIGURE 3.</emphasis>
<taxonomicName id="71D43DBDFFC2A310A07D3418792A1902" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[267,532,1812,1833]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A07D3418792A1902" box="[267,532,1812,1833]" italics="true" pageId="5" pageNumber="276">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
and other morphologically similar
<taxonomicName id="71D43DBDFFC2A310A20D34187F4F1902" box="[891,1137,1812,1833]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="84A09A2CFFC2A310A20D34187F3C1902" box="[891,1026,1812,1833]" italics="true" pageId="5" pageNumber="276">Syngonanthus</emphasis>
species.
<emphasis id="84A09A2CFFC2A310A51634187F4F1902" bold="true" box="[1120,1137,1812,1833]" pageId="5" pageNumber="276">A</emphasis>
</taxonomicName>
<emphasis id="84A09A2CFFC2A310A50734187FAB1902" bold="true" box="[1137,1173,1812,1833]" pageId="5" pageNumber="276">C.</emphasis>
Comparison among leaves:
<emphasis id="84A09A2CFFC2A310A1FE34347B9E1966" bold="true" box="[136,160,1848,1869]" pageId="5" pageNumber="276">A.</emphasis>
<taxonomicName id="71D43DBDFFC2A310A1DF3434792F1966" authorityName="Moldenke" authorityYear="1972" box="[169,529,1848,1869]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC2A310A1DF34347A9F1966" box="[169,417,1848,1869]" italics="true" pageId="5" pageNumber="276">Syngonanthus densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A0AE3434792F1966" box="[472,529,1848,1869]" italics="true" pageId="5" pageNumber="276">majus</emphasis>
</taxonomicName>
showing erect, flexuous leaves in a rosette;
<emphasis id="84A09A2CFFC2A310A2A1343478D31966" bold="true" box="[983,1005,1848,1869]" pageId="5" pageNumber="276">B.</emphasis>
Incurved leaves of
<taxonomicName id="71D43DBDFFC2A310A5CF34347E701966" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1209,1358,1848,1869]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A5CF34347E701966" box="[1209,1358,1848,1869]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFC2A310A42834347E481966" bold="true" box="[1374,1398,1848,1869]" pageId="5" pageNumber="276">C.</emphasis>
Leaf of
<taxonomicName id="71D43DBDFFC2A310A1D334507ADB195A" authorityName="Moldenke" authorityYear="1984" box="[165,485,1884,1905]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC2A310A1D334507A16195A" box="[165,296,1884,1905]" italics="true" pageId="5" pageNumber="276">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A02B34507ADB195A" box="[349,485,1884,1905]" italics="true" pageId="5" pageNumber="276">brachyphyllus</emphasis>
</taxonomicName>
, showing achlorophyllous base.
<emphasis id="84A09A2CFFC2A310A25A3450785D195A" bold="true" box="[812,867,1884,1905]" pageId="5" pageNumber="276">DG.</emphasis>
Comparison among sexual traits of
<taxonomicName id="71D43DBDFFC2A310A5A634507E5A195A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1232,1380,1884,1905]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A5A634507E5A195A" box="[1232,1380,1884,1905]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFC2A310A4EF34507A6419BE" authorityName="Moldenke" authorityYear="1972" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC2A310A4EF34507BCE19BE" italics="true" pageId="5" pageNumber="276">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A057348C7A6419BE" box="[289,346,1920,1941]" italics="true" pageId="5" pageNumber="276">majus</emphasis>
</taxonomicName>
:
<emphasis id="84A09A2CFFC2A310A011348C7A4119BE" bold="true" box="[359,383,1920,1941]" pageId="5" pageNumber="276">D.</emphasis>
Staminate flower of
<taxonomicName id="71D43DBDFFC2A310A324348C79DA19BE" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[594,740,1920,1941]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A324348C79DA19BE" box="[594,740,1920,1941]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
(trichomes removed from pedicel);
<emphasis id="84A09A2CFFC2A310A534348C7F6719BE" bold="true" box="[1090,1113,1920,1941]" pageId="5" pageNumber="276">E.</emphasis>
Pistillate flower of
<taxonomicName id="71D43DBDFFC2A310A46F348C7E9519BE" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1305,1451,1920,1941]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A46F348C7E9519BE" box="[1305,1451,1920,1941]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
with filamentous trichomes;
<emphasis id="84A09A2CFFC2A310A0E934A87A8F1992" bold="true" box="[415,433,1956,1977]" pageId="5" pageNumber="276">F.</emphasis>
Staminate flower of
<taxonomicName id="71D43DBDFFC2A310A3F734A878521992" authorityName="Moldenke" authorityYear="1972" box="[641,876,1956,1977]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC2A310A3F734A8783C1992" box="[641,770,1956,1977]" italics="true" pageId="5" pageNumber="276">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A24534A878521992" box="[819,876,1956,1977]" italics="true" pageId="5" pageNumber="276">majus</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFC2A310A20D34A878AA1992" bold="true" box="[891,916,1956,1977]" pageId="5" pageNumber="276">G.</emphasis>
Pistillate flower of
<taxonomicName id="71D43DBDFFC2A310A52134A87E7C1992" authorityName="Moldenke" authorityYear="1972" box="[1111,1346,1956,1977]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC2A310A52134A87FE61992" box="[1111,1240,1956,1977]" italics="true" pageId="5" pageNumber="276">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A47F34A87E7C1992" box="[1289,1346,1956,1977]" italics="true" pageId="5" pageNumber="276">majus</emphasis>
</taxonomicName>
exhibiting trichomes only at upper portion of the sepals;
<emphasis id="84A09A2CFFC2A310A33734C4796419F6" bold="true" box="[577,602,1992,2013]" pageId="5" pageNumber="276">H.</emphasis>
Capitulum detail of
<taxonomicName id="71D43DBDFFC2A310A25634C4788E19F6" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[800,944,1992,2013]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A25634C4788E19F6" box="[800,944,1992,2013]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
;
<emphasis id="84A09A2CFFC2A310A2CB34C478DB19F6" bold="true" box="[957,997,1992,2013]" pageId="5" pageNumber="276">IJ.</emphasis>
Comparative vestiture of the petals in pistillate flowers:
<emphasis id="84A09A2CFFC2A310A1AB34E07BD2162A" bold="true" box="[221,236,2028,2049]" pageId="5" pageNumber="276">I.</emphasis>
<taxonomicName id="71D43DBDFFC2A310A18534E07ABA162A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[243,388,2028,2049]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC2A310A18534E07ABA162A" box="[243,388,2028,2049]" italics="true" pageId="5" pageNumber="276">S. incurvifolius</emphasis>
</taxonomicName>
showing pilose petals;
<emphasis id="84A09A2CFFC2A310A31134E07947162A" bold="true" box="[615,633,2028,2049]" pageId="5" pageNumber="276">J.</emphasis>
Petals glabrous in
<taxonomicName id="71D43DBDFFC2A310A24734E07F24162A" authorityName="Moldenke" authorityYear="1972" box="[817,1050,2028,2049]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="276" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC2A310A24734E0788F162A" box="[817,945,2028,2049]" italics="true" pageId="5" pageNumber="276">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC2A310A29734E07F24162A" box="[993,1050,2028,2049]" italics="true" pageId="5" pageNumber="276">majus</emphasis>
</taxonomicName>
. Scale bars: B= 3 mm. C,H= 2mm. DG, IJ: 500 μm. (Photos by Mauricio T. C. Watanabe).
</paragraph>
</caption>
<paragraph id="B66B463EFFC1A313A1CB33AC78831F46" blockId="6.[136,1452,160,1661]" pageId="6" pageNumber="277">
<taxonomicName id="71D43DBDFFC1A313A1CB33AC7AD31E92" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[189,493,160,185]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A1CB33AC7AD31E92" box="[189,493,160,185]" italics="true" pageId="6" pageNumber="277">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
has been determined as
<taxonomicName id="71D43DBDFFC1A313A38433AC7FD41E92" authority="Silveira (1928: 320)" authorityName="Silveira" authorityPageNumber="320" authorityYear="1928" box="[754,1258,160,185]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="densifolius" variety="pilosior">
<emphasis id="84A09A2CFFC1A313A38433AC78BB1E92" box="[754,901,160,185]" italics="true" pageId="6" pageNumber="277">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A2C133AC7F301E92" box="[951,1038,160,185]" italics="true" pageId="6" pageNumber="277">pilosior</emphasis>
<bibRefCitation id="D2453BCFFFC1A313A56433AC7FD41E92" author="Silveira, A." box="[1042,1258,160,185]" pageId="6" pageNumber="277" refId="ref8191" refString="Silveira, A. (1928) Floralia Montium. Imprensa Official, Belo Horizonte. 399 pp." type="book" year="1928">Silveira (1928:320)</bibRefCitation>
</taxonomicName>
in some herbaria. Actually,
<taxonomicName id="71D43DBDFFC1A313A18133C8791C1EF6" authorityName="Silveira" authorityYear="1928" box="[247,546,196,221]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="densifolius" variety="pilosior">
<emphasis id="84A09A2CFFC1A313A18133C87AB11EF6" box="[247,399,196,221]" italics="true" pageId="6" pageNumber="277">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A0BD33C8791C1EF6" box="[459,546,196,221]" italics="true" pageId="6" pageNumber="277">pilosior</emphasis>
</taxonomicName>
is probably a synonym of
<taxonomicName id="71D43DBDFFC1A313A22A33C87A521F2A" authority="(Ruhland) L. R. Parra &amp; Giulietti (1997: 228)" authorityName="L. R. Parra &amp; Giulietti" authorityPageNumber="228" authorityYear="1997" baseAuthorityName="Ruhland" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="anthemiflorus" variety="similis">
<emphasis id="84A09A2CFFC1A313A22A33C87F281EF6" box="[860,1046,196,221]" italics="true" pageId="6" pageNumber="277">S. anthemiflorus</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A52433C87FA51EF6" box="[1106,1179,196,221]" italics="true" pageId="6" pageNumber="277">similis</emphasis>
(Ruhland) L.R.
<bibRefCitation id="D2453BCFFFC1A313A43A33C87A521F2A" author="Parra, L. R. &amp; Giulietti, A. M." pageId="6" pageNumber="277" pagination="227 - 233" refId="ref7882" refString="Parra, L. R. &amp; Giulietti, A. M. (1997) Nomenclatural and taxonomic changes in Brazilian Syngonanthus (Eriocaulaceae). Willdenowia 27: 227 - 233. http: // dx. doi. org / 10.3372 / wi. 27.2723" type="journal article" year="1997">Parra &amp; Giulietti (1997:228)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="D2453BCFFFC1A313A00B33E479751F2A" author="Echternacht, L." box="[381,587,232,257]" pageId="6" pageNumber="277" refId="ref7167" refString="Echternacht, L. (2012) Sistematica de Comanthera e de Syngonanthus (Eriocaulaceae). Ph. D. Thesis, Universidade de Sao Paulo, Sao Paulo, 294 pp." type="book" year="2012">Echternacht 2012</bibRefCitation>
), which can be easily distinguished from
<taxonomicName id="71D43DBDFFC1A313A54733E47FE01F2A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1073,1246,232,257]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A54733E47FE01F2A" box="[1073,1246,232,257]" italics="true" pageId="6" pageNumber="277">S. incurvifolius</emphasis>
</taxonomicName>
by the involucral bracts surpassing the capitulum disc (
<emphasis id="84A09A2CFFC1A313A3503200797B1F0E" box="[550,581,268,293]" italics="true" pageId="6" pageNumber="277">vs.</emphasis>
not surpassing), the outer series golden to straw-colored and the internal series cream (
<emphasis id="84A09A2CFFC1A313A1A8323C7BC31F62" box="[222,253,304,329]" italics="true" pageId="6" pageNumber="277">vs.</emphasis>
all series castaneous to reddish-brown). Furthermore, the distributions of these taxa do not overlap, as
<taxonomicName id="71D43DBDFFC1A313A4E0323C7A1A1F46" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="anthemiflorus">
<emphasis id="84A09A2CFFC1A313A4E0323C7A1A1F46" italics="true" pageId="6" pageNumber="277">S. anthemiflorus</emphasis>
</taxonomicName>
is restricted to the Espinhaço Range in
<collectingRegion id="741088DCFFC1A313A397325878841F46" box="[737,954,340,365]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="277">Minas Gerais State</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A1CB32747BE41C42" pageId="6" pageNumber="277" type="biology_ecology">
<paragraph id="B66B463EFFC1A313A1CB32747BE41C42" blockId="6.[136,1452,160,1661]" pageId="6" pageNumber="277">
<emphasis id="84A09A2CFFC1A313A1CB32747A611FBA" bold="true" box="[189,351,376,401]" pageId="6" pageNumber="277">Phenology:—</emphasis>
The flowering season of
<taxonomicName id="71D43DBDFFC1A313A3E2327478F01FBA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[660,974,376,401]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A3E2327478F01FBA" box="[660,974,376,401]" italics="true" pageId="6" pageNumber="277">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
is somewhat different from congeneric sympatric species. Most species of
<taxonomicName id="71D43DBDFFC1A313A3733290799C1F9E" authorityName="Ruhland" authorityYear="1900" box="[517,674,412,437]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="genus">
<emphasis id="84A09A2CFFC1A313A3733290799C1F9E" box="[517,674,412,437]" italics="true" pageId="6" pageNumber="277">Syngonanthus</emphasis>
</taxonomicName>
bloom during the dry season but the blooming period of
<taxonomicName id="71D43DBDFFC1A313A47332907E921F9E" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1285,1452,412,437]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A47332907E921F9E" box="[1285,1452,412,437]" italics="true" pageId="6" pageNumber="277">S. incurvifolius</emphasis>
</taxonomicName>
recorded in herbaria is between February and May (the rainy season). Exceptions are
<taxonomicName id="71D43DBDFFC1A313A53832CC7E611FF2" authorityName="Moldenke" authorityYear="1972" box="[1102,1375,448,473]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="densifolius" variety="majus">
<emphasis id="84A09A2CFFC1A313A53832CC7FDD1FF2" box="[1102,1251,448,473]" italics="true" pageId="6" pageNumber="277">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A46A32CC7E611FF2" box="[1308,1375,448,473]" italics="true" pageId="6" pageNumber="277">majus</emphasis>
</taxonomicName>
and
<taxonomicName id="71D43DBDFFC1A313A4E032CC7AE91FD6" authorityName="Moldenke" authorityYear="1984" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="variety" species="densifolius" variety="brachyphyllus">
<emphasis id="84A09A2CFFC1A313A4E032CC7A3F1FD6" italics="true" pageId="6" pageNumber="277">S. densifolius</emphasis>
var.
<emphasis id="84A09A2CFFC1A313A04E32E87AE91FD6" box="[312,471,484,509]" italics="true" pageId="6" pageNumber="277">brachyphyllus</emphasis>
</taxonomicName>
, whose records also attest for a blooming in the first semester of the year. Flowers of
<taxonomicName id="71D43DBDFFC1A313A4E032E87A281C0A" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A4E032E87A281C0A" italics="true" pageId="6" pageNumber="277">S. incurvifolius</emphasis>
</taxonomicName>
are in anthesis from February to April, corresponding to the latter half of the rainy season in the region. Fruits were observed only in herbarium specimens collected in May, suggesting that seeds are dispersed during the dry season.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A1CB317878581DDE" pageId="6" pageNumber="277" type="distribution">
<paragraph id="B66B463EFFC1A313A1CB317878581DDE" blockId="6.[136,1452,160,1661]" pageId="6" pageNumber="277">
<emphasis id="84A09A2CFFC1A313A1CB317879311CA6" bold="true" box="[189,527,628,653]" pageId="6" pageNumber="277">Habitat and distribution:—</emphasis>
Records for this species have only been reported from the municipality of Alto Paraíso de
<collectingRegion id="741088DCFFC1A313A07E31947A721C9A" box="[264,332,664,689]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
, along a small stretch of GO-118 highway, that connects Alto Paraíso de
<collectingRegion id="741088DCFFC1A313A5D531947FDA1C9A" box="[1187,1252,664,689]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
and São João da Aliança, between
<quantity id="712CEBDBFFC1A313A02531B07A891CFE" box="[339,439,700,725]" metricMagnitude="4" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="6" pageNumber="277" unit="km" value="10.0" valueMax="15.0" valueMin="5.0">515 km</quantity>
south of Alto Paraíso de
<collectingRegion id="741088DCFFC1A313A3A131B078221CFE" box="[727,796,700,725]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
. According to herbarium labels,
<taxonomicName id="71D43DBDFFC1A313A5FA31B07E091CFE" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1164,1335,700,725]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A5FA31B07E091CFE" box="[1164,1335,700,725]" italics="true" pageId="6" pageNumber="277">S. incurvifolius</emphasis>
</taxonomicName>
occurs on
<emphasis id="84A09A2CFFC1A313A1FE31EC7A731CD2" box="[136,333,736,761]" italics="true" pageId="6" pageNumber="277">campos rupestres</emphasis>
within the Cerrado biome, over quartzitic sandy soils and in wet meadows, typical habitats at the border of the highway at the described locality. Field observations (
<emphasis id="84A09A2CFFC1A313A20B30087F5A1D36" box="[893,1124,772,797]" italics="true" pageId="6" pageNumber="277">King &amp; Bishop 8900</emphasis>
, MO) reported the species to be locally abundant. However, we have done fieldwork in Chapada dos Veadeiros over a large area and we did not find the reported population neither did we see any other subpopulation. This fact indicates that the populations probably occupy narrow areas. The roadside along GO-118 northwest of Alto Paraíso de
<collectingRegion id="741088DCFFC1A313A573307C7F781DA2" box="[1029,1094,880,905]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
belongs to the Parque Nacional da Chapada dos Veadeiros (PNCV); roads to the south and northeast are privately owned, used for cattle grazing and farming, fragmenting the original
<emphasis id="84A09A2CFFC1A313A37230B4798F1DFA" box="[516,689,952,977]" italics="true" pageId="6" pageNumber="277">
campo
<taxonomicName id="71D43DBDFFC1A313A32330B4798F1DFA" box="[597,689,952,977]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="rupestre">rupestre</taxonomicName>
</emphasis>
habitat.
<taxonomicName id="71D43DBDFFC1A313A26430B47F7A1DFA" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[786,1092,952,977]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A26430B47F7A1DFA" box="[786,1092,952,977]" italics="true" pageId="6" pageNumber="277">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
is registered southward, outside the PNCV and therefore might be threatened by human activities.
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A1CB370C7F6C1B76" pageId="6" pageNumber="277" type="conservation">
<paragraph id="B66B463EFFC1A313A1CB370C7F6C1B76" blockId="6.[136,1452,160,1661]" pageId="6" pageNumber="277">
<emphasis id="84A09A2CFFC1A313A1CB370C7AF11A32" bold="true" box="[189,463,1024,1049]" pageId="6" pageNumber="277">Conservation status:—</emphasis>
<taxonomicName id="71D43DBDFFC1A313A0A6370C783C1A32" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[464,770,1024,1049]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A0A6370C783C1A32" box="[464,770,1024,1049]" italics="true" pageId="6" pageNumber="277">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
is known from a stretch no longer than
<quantity id="712CEBDBFFC1A313A5C8370C7E3B1A32" box="[1214,1285,1024,1049]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="277" unit="km" value="10.0">10 km</quantity>
along GO-118 highway, where the natural habitat is severely fragmented. The species is not recorded from a protected area and the region is affected by fire, pasturage and farming, with great impact on natural habitats. As it has not been collected since 1994 and there was no success to find it in recent fieldwork, the species could be classified as extinct in the nature. However, the region is still poorly collected and a careful search might discover new populations. Because there is reasonable doubt that the last individual has died, a more thorough check of expected habitat of this species is needed to confirm it. CNCFlora (Brazilian National Center for Plant Conservation, cncflora.jbrj.gov.br) evaluated the species threatened status, following the criteria of the IUCN Red List (
<bibRefCitation id="D2453BCFFFC1A313A2EF37F07F1F1B3E" author="IUCN" box="[921,1057,1276,1301]" pageId="6" pageNumber="277" refId="ref7582" refString="IUCN (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland &amp; Cambridge, UK, 70 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
,
<bibRefCitation id="D2453BCFFFC1A313A55D37F07F561B3E" author="IUCN" box="[1067,1128,1276,1301]" pageId="6" pageNumber="277" refId="ref7614" refString="IUCN (2015) The IUCN red list of threatened species. Available from: www. iucnredlist. org / (accessed 1 January 2015)" type="book" year="2015">2015</bibRefCitation>
).
<taxonomicName id="71D43DBDFFC1A313A50C37F07E921B3E" authorityName="M. T. C. Watan. &amp; Echternacht" authorityYear="2015" box="[1146,1452,1276,1301]" class="Liliopsida" family="Eriocaulaceae" genus="Syngonanthus" kingdom="Plantae" order="Poales" pageId="6" pageNumber="277" phylum="Tracheophyta" rank="species" species="incurvifolius">
<emphasis id="84A09A2CFFC1A313A50C37F07E921B3E" box="[1146,1452,1276,1301]" italics="true" pageId="6" pageNumber="277">Syngonanthus incurvifolius</emphasis>
</taxonomicName>
has an area of occupancy of
<quantity id="712CEBDBFFC1A313A0B8362C79361B12" box="[462,520,1312,1337]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="277" unit="km" value="8.0">8 km</quantity>
<superScript id="41A1EB76FFC1A313A37E3612792E1B07" attach="left" box="[520,528,1310,1324]" fontSize="6" pageId="6" pageNumber="277">2</superScript>
, which tend to contract and deteriorate in quality. According to criteria A2ac and B2ab(i,ii,iii) this species is currently considered critically endangered (Negrão 2014a).
</paragraph>
</subSubSection>
<subSubSection id="FECE15B5FFC1A313A1CB3664780E1856" pageId="6" pageNumber="277" type="materials_examined">
<paragraph id="B66B463EFFC1A313A1CB3664780E1856" blockId="6.[136,1452,160,1661]" pageId="6" pageNumber="277">
<emphasis id="84A09A2CFFC1A313A1CB366479671BAA" bold="true" box="[189,601,1384,1409]" pageId="6" pageNumber="277">Additional specimens examined:—</emphasis>
<materialsCitation id="06BC4C63FFC1A313A32F36647A301BE2" collectingDate="1956-05-24" collectionCode="LL" collectorName="Y. Dawson" country="Brazil" county="Alto Paraiso" latitude="-14.5" location="Region of the Chapada dos Veadeiros" longLatPrecision="1288" longitude="-47.5" municipality="Alto Paraiso de Goias" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Goias">
<collectingCountry id="CEC306AEFFC1A313A32F366479FC1BAA" box="[601,706,1384,1409]" name="Brazil" pageId="6" pageNumber="277">BRAZIL</collectingCountry>
.
<collectingRegion id="741088DCFFC1A313A3BC3664782E1BAA" box="[714,784,1384,1409]" country="Brazil" name="Goias" pageId="6" pageNumber="277">Goiás</collectingRegion>
:
<collectingMunicipality id="560FDC44FFC1A313A26036647F361BAA" box="[790,1032,1384,1409]" pageId="6" pageNumber="277">Alto Paraíso de Goiás</collectingMunicipality>
,
<location id="B30B10E5FFC1A313A56736647E951BAA" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC1A313A56736647E951BAA" box="[1041,1451,1384,1409]" country="Brazil" county="Alto Paraiso" latitude="-14.5" longLatPrecision="1288" longitude="-47.5" municipality="Alto Paraiso de Goias" name="Region of the Chapada dos Veadeiros" pageId="6" pageNumber="277" stateProvince="Goias">Region of the Chapada dos Veadeiros</location>
at
<geoCoordinate id="D3E020F9FFC1A313A1D536807A3D1B8E" box="[163,259,1420,1445]" degrees="14" direction="south" minutes="30" orientation="latitude" pageId="6" pageNumber="277" precision="925" value="-14.5">14°30 S</geoCoordinate>
<geoCoordinate id="D3E020F9FFC1A313A07F36807A4B1B8E" box="[265,373,1420,1445]" degrees="47" direction="west" minutes="30" orientation="longitude" pageId="6" pageNumber="277" precision="925" value="-47.5">47°30 W</geoCoordinate>
,
<quantity id="712CEBDBFFC1A313A00936807A891B8E" box="[383,439,1420,1445]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="277" unit="km" value="7.0">7 km</quantity>
south of Veadeiros, wet meadowland below sandstone outcrop 7,
<date id="C26A60FEFFC1A313A5E236807E151B8E" box="[1172,1323,1420,1445]" pageId="6" pageNumber="277" value="1956-05-24">
<collectingDate id="D22E9916FFC1A313A5E236807E151B8E" box="[1172,1323,1420,1445]" pageId="6" pageNumber="277" value="1956-05-24">24 May 1956</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC1A313A44336807BF01BE2" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A44336807E921B8E" box="[1333,1452,1420,1445]" pageId="6" pageNumber="277">Y. Dawson</collectorName>
14639
</emphasis>
(
<collectionCode id="D0C5DEFBFFC1A313A1A836BC7A3B1BE2" box="[222,261,1456,1481]" pageId="6" pageNumber="277">LL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFC1A313A06C36BC7A371BC6" collectingDate="1975-05-26" collectionCode="C, LL, MBM" collectorName="G. Hatschbach" country="Brazil" location="Rod. GO- 12, km 5 - 10 ao Sul de Alto Paraiso" municipality="Rod." pageId="6" pageNumber="277" specimenCount="1">
<location id="B30B10E5FFC1A313A06C36BC78211BE2" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC1A313A06C36BC78211BE2" box="[282,799,1456,1481]" country="Brazil" municipality="Rod." name="Rod. GO- 12, km 5 - 10 ao Sul de Alto Paraiso" pageId="6" pageNumber="277">Rod. GO-12, km 510 ao Sul de Alto Paraíso</location>
, campo arenoso,
<date id="C26A60FEFFC1A313A29236BC7F401BE2" box="[996,1150,1456,1481]" pageId="6" pageNumber="277" value="1975-05-26">
<collectingDate id="D22E9916FFC1A313A29236BC7F401BE2" box="[996,1150,1456,1481]" pageId="6" pageNumber="277" value="1975-05-26">26 May 1975</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC1A313A5FC36BC7EBE1BE2" box="[1162,1408,1456,1481]" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A5FC36BC7E0D1BE2" box="[1162,1331,1456,1481]" pageId="6" pageNumber="277">G. Hatschbach</collectorName>
36814
</emphasis>
(
<collectionCode id="D0C5DEFBFFC1A313A4E736BC7E981BE2" box="[1425,1446,1456,1481]" pageId="6" pageNumber="277">C</collectionCode>
,
<collectionCode id="D0C5DEFBFFC1A313A1FE36D87B931BC6" box="[136,173,1492,1517]" pageId="6" pageNumber="277">LL</collectionCode>
,
<collectionCode id="D0C5DEFBFFC1A313A1CE36D87A3F1BC6" box="[184,257,1492,1517]" pageId="6" pageNumber="277">MBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFC1A313A06536D879F3183A" collectingDate="1981-02-08" collectionCode="MO, UB" collectorName="L. R. M. King &amp; L. E. Bishop" country="Brazil" elevation="1082" location="5 - 12 km S of Alto Paraiso de Goias, mostly E of road to Sao Joao da Alianca" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Goias">
<location id="B30B10E5FFC1A313A06536D87F521BC6" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC1A313A06536D87F521BC6" box="[275,1132,1492,1517]" country="Brazil" name="5 - 12 km S of Alto Paraiso de Goias, mostly E of road to Sao Joao da Alianca" pageId="6" pageNumber="277" stateProvince="Goias">
<quantity id="712CEBDBFFC1A313A06536D87A481BC6" box="[275,374,1492,1517]" metricMagnitude="3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="6" pageNumber="277" unit="km" value="8.5" valueMax="12.0" valueMin="5.0">512 km</quantity>
S of Alto Paraíso de Goiás, mostly E of road to São João da Aliança
</location>
,
<quantity id="712CEBDBFFC1A313A50136D87E161BC6" box="[1143,1320,1492,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" metricValueMax="1.15824" metricValueMin="1.0058399999999998" pageId="6" pageNumber="277" unit="ft" value="3550.0" valueMax="3800.0" valueMin="3300.0">
<elevation id="3DF9A10DFFC1A313A50136D87E161BC6" box="[1143,1320,1492,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" metricValueMax="1.15824" metricValueMin="1.0058399999999998" pageId="6" pageNumber="277" unit="ft" value="3550.0" valueMax="3800.0" valueMin="3300.0">33003800 feet</elevation>
</quantity>
,
<date id="C26A60FEFFC1A313A44436D87BFC183A" pageId="6" pageNumber="277" value="1981-02-08">
<collectingDate id="D22E9916FFC1A313A44436D87BFC183A" pageId="6" pageNumber="277" value="1981-02-08">8 February 1981</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC1A313A1B836F4796E183A" box="[206,592,1528,1553]" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A1B836F47A5A183A" box="[206,356,1528,1553]" pageId="6" pageNumber="277">L. R. M. King</collectorName>
&amp;
<collectorName id="1B2123E8FFC1A313A0FE36F4792C183A" box="[392,530,1528,1553]" pageId="6" pageNumber="277">L. E. Bishop</collectorName>
8900
</emphasis>
(
<collectionCode id="D0C5DEFBFFC1A313A32936F479AE183A" box="[607,656,1528,1553]" pageId="6" pageNumber="277">MO</collectionCode>
,
<collectionCode id="D0C5DEFBFFC1A313A3ED36F479F8183A" box="[667,710,1528,1553]" pageId="6" pageNumber="277">UB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFC1A313A3AF36F47BFE1872" collectingDate="1990-02-23" collectionCode="LL" collectorName="J. Saunders &amp; L. Breyer &amp; G. Eiten" country="Brazil" county="Brasilia" latitude="-14.5" location="10 km S of Alto Paraiso de Goias (GO- 118) heading for Brasilia, adjacent to Chapada dos Veadeiros" longLatPrecision="1288" longitude="-47.5" municipality="Chapada" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Goias">
<location id="B30B10E5FFC1A313A3AF36F47927181E" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC1A313A3AF36F47927181E" country="Brazil" county="Brasilia" latitude="-14.5" longLatPrecision="1288" longitude="-47.5" municipality="Chapada" name="10 km S of Alto Paraiso de Goias (GO- 118) heading for Brasilia, adjacent to Chapada dos Veadeiros" pageId="6" pageNumber="277" stateProvince="Goias">
<quantity id="712CEBDBFFC1A313A3AF36F4781E183A" box="[729,800,1528,1553]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="277" unit="km" value="10.0">10 km</quantity>
S of Alto Paraíso de Goiás (GO-118) heading for Brasília, adjacent to Chapada dos Veadeiros
</location>
,
<geoCoordinate id="D3E020F9FFC1A313A351351079BB181E" box="[551,645,1564,1589]" degrees="14" direction="south" minutes="30" orientation="latitude" pageId="6" pageNumber="277" precision="925" value="-14.5">14°30S</geoCoordinate>
,
<geoCoordinate id="D3E020F9FFC1A313A3E5351079C4181E" box="[659,762,1564,1589]" degrees="47" direction="west" minutes="30" orientation="longitude" pageId="6" pageNumber="277" precision="925" value="-47.5">47°30W</geoCoordinate>
,
<date id="C26A60FEFFC1A313A271351078E8181E" box="[775,982,1564,1589]" pageId="6" pageNumber="277" value="1990-02-23">
<collectingDate id="D22E9916FFC1A313A271351078E8181E" box="[775,982,1564,1589]" pageId="6" pageNumber="277" value="1990-02-23">23 February 1990</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC1A313A29535107E95181E" box="[995,1451,1564,1589]" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A29535107F54181E" box="[995,1130,1564,1589]" pageId="6" pageNumber="277">J. Saunders</collectorName>
,
<collectorName id="1B2123E8FFC1A313A50E35107FDC181E" box="[1144,1250,1564,1589]" pageId="6" pageNumber="277">L. Breyer</collectorName>
&amp;
<collectorName id="1B2123E8FFC1A313A47D35107E54181E" box="[1291,1386,1564,1589]" pageId="6" pageNumber="277">G. Eiten</collectorName>
3020
</emphasis>
(
<collectionCode id="D0C5DEFBFFC1A313A1E6354C7B891872" box="[144,183,1600,1625]" pageId="6" pageNumber="277">LL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06BC4C63FFC1A313A1B8354C78121856" collectingDate="1994-02-10" collectionCode="F, MBM" collectorName="G. Hatschbach &amp; M. Hatschbach &amp; J. M. Silva" country="Brazil" elevation="1150" location="Rod. GO- 118, 10 km S de Alto Paraiso" pageId="6" pageNumber="277" specimenCount="1">
<location id="B30B10E5FFC1A313A1B8354C79AA1872" LSID="urn:lsid:plazi:treatment:3E7DF728FFC5A313A1FE3758780E1856:B30B10E5FFC1A313A1B8354C79AA1872" box="[206,660,1600,1625]" country="Brazil" name="Rod. GO- 118, 10 km S de Alto Paraiso" pageId="6" pageNumber="277">
Rod. GO-118,
<quantity id="712CEBDBFFC1A313A001354C7AFE1872" box="[375,448,1600,1625]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="277" unit="km" value="10.0">10 km</quantity>
S de Alto Paraíso
</location>
, campo rupestre, solo arenoso úmido,
<quantity id="712CEBDBFFC1A313A539354C7F991872" box="[1103,1191,1600,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="277" unit="m" value="1150.0">
<elevation id="3DF9A10DFFC1A313A539354C7F991872" box="[1103,1191,1600,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="277" unit="m" value="1150.0">1150 m</elevation>
</quantity>
,
<date id="C26A60FEFFC1A313A5C3354C7EBD1872" box="[1205,1411,1600,1625]" pageId="6" pageNumber="277" value="1994-02-10">
<collectingDate id="D22E9916FFC1A313A5C3354C7EBD1872" box="[1205,1411,1600,1625]" pageId="6" pageNumber="277" value="1994-02-10">10 February 1994</collectingDate>
</date>
,
<emphasis id="84A09A2CFFC1A313A4E6354C79911856" italics="true" pageId="6" pageNumber="277">
<collectorName id="1B2123E8FFC1A313A4E6354C7A2F1856" pageId="6" pageNumber="277">G. Hatschbach</collectorName>
,
<collectorName id="1B2123E8FFC1A313A06A35687AF91856" box="[284,455,1636,1661]" pageId="6" pageNumber="277">M. Hatschbach</collectorName>
&amp;
<collectorName id="1B2123E8FFC1A313A09D3568795C1856" box="[491,610,1636,1661]" pageId="6" pageNumber="277">J. M. Silva</collectorName>
60290
</emphasis>
(
<collectionCode id="D0C5DEFBFFC1A313A3C9356879EE1856" box="[703,720,1636,1661]" pageId="6" pageNumber="277">F</collectionCode>
,
<collectionCode id="D0C5DEFBFFC1A313A3AD3568781A1856" box="[731,804,1636,1661]" pageId="6" pageNumber="277">MBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>