Add updates up until 2025-02-25 18:56:52

This commit is contained in:
ggserver 2025-02-25 19:01:55 +00:00
parent f43449b616
commit e4851425ba
3 changed files with 1665 additions and 0 deletions

View file

@ -0,0 +1,277 @@
<document id="DFEA5B831EB2D37E1CCB81A559CE4A10" ID-DOI="10.11646/zootaxa.5588.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896067" ID-ZooBank="E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1740002781111" checkinUser="plazi" docAuthor="Jacintho, Gabriela De Figueiredo, Chaul, Júlio Cezar Mário, Teixeira, Gisele Amaro, Barros, Luísa Antônia Campos, Lopes, Denilce Meneses &amp; Serrão, José Eduardo" docDate="2025" docId="03A887D2FF81693835902920FD1CF859" docLanguage="en" docName="zootaxa.5588.1.3.pdf" docOrigin="Zootaxa 5588 (1)" docSource="https://doi.org/10.11646/zootaxa.5588.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fulakora elongata" docType="treatment" docVersion="1" lastPageNumber="87" masterDocId="FF91FFAAFF8B693235072B02FFEBFFCC" masterDocTitle="A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species" masterLastPageNumber="104" masterPageNumber="77" pageNumber="87" updateTime="1740509805003" updateUser="guilherme" zenodo-license-document="CLOSED">
<mods:mods id="47E5EE796CC39E2C6DA0DAFA6C839EF6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D6302F5D523A3D8D3336E241211A1124">
<mods:title id="668D2F755F7D4D605D55F73B9C8B66E4">A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="0AE03EB715BAADA1AFCFF4DB7C7DC7CE" type="personal">
<mods:role id="D59D972DF9F9A778BD0D44C30D86C783">
<mods:roleTerm id="22060EECC8A6F066D0489BA450FBB8A5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6DB650D2E40A6DA367667ECFDA62170C">Jacintho, Gabriela De Figueiredo</mods:namePart>
<mods:affiliation id="1A163E602D6D25DF13E2B968B5E2E891">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="D7A4475CF5DEA3DE792A76089A51CC04" type="personal">
<mods:role id="104BB61698D92898A652C7CFA2B40E12">
<mods:roleTerm id="61122976A0486FD338260C28AC478440">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16F6B831068F766987D2B4B83F751E40">Chaul, Júlio Cezar Mário</mods:namePart>
<mods:affiliation id="ADB582CE48ABA5B3A9C1B7509B4A162F">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="82C8CC557F9703B5276F17C11444689E" type="personal">
<mods:role id="49EF0BCFA56C8C1F2E3ABDFF7897C564">
<mods:roleTerm id="5402CAF48001A230648510EB95467AA0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63DE90B467FFA00CB5A59CAA489CF367">Teixeira, Gisele Amaro</mods:namePart>
<mods:affiliation id="A0AD8871489CE4272C75616BC7EF4404">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="2FE7110BC18961B2639345FBDFC1E5CC" type="personal">
<mods:role id="24B5E13449BBA55129878F94FE9434E5">
<mods:roleTerm id="BDAEF8918F127377B6752EB2CC0F7A89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D00279E1419A220380826F1E7564E99">Barros, Luísa Antônia Campos</mods:namePart>
<mods:affiliation id="428892783A4F93E587B3562F289FE395">Universidade Federal do Amapá, Campus Binacional, 68980 - 000, Oiapoque, Amapá, Brazil</mods:affiliation>
</mods:name>
<mods:name id="D27534DC7CC5C6919693657B6B666049" type="personal">
<mods:role id="1E5E18149BF83BA4796DBC9577E5C2C4">
<mods:roleTerm id="79FF9DA5E12C1D8D069CE6F21B91E85D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2001FBE36CC0E871A6FEF79EE1CF3EF1">Lopes, Denilce Meneses</mods:namePart>
<mods:affiliation id="4E78CBD7B83289C4521E1F62272B5398">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="CDFC520211D25E20AD2AD70B72B2EB65" type="personal">
<mods:role id="8992FDFDDD5DB10CEB87B5C21A48930D">
<mods:roleTerm id="B8CA76FED1EE66A05F0049BC38119A80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20B06DB3431AC3489CDC0FC3E04C41C5">Serrão, José Eduardo</mods:namePart>
<mods:affiliation id="1B935611C010DA01BC76F464583674AE">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6AB11DF4F91305D47B059FB3C7CED337">text</mods:typeOfResource>
<mods:relatedItem id="504CEFBEE8B3F780FFB52AF0819DEC9A" type="host">
<mods:titleInfo id="13B51247B3954C719254C59584CE837B">
<mods:title id="DC5701D08EA55898D6163D814F3869A6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C3CFEFD04112C8B3014354516E3E0249">
<mods:date id="1A8FF1C1846B071419D84A187E81C1A8">2025</mods:date>
<mods:detail id="1DAF494D9AB1D8FE765F0C0C73E411B1" type="pubDate">
<mods:number id="C7CE5A83EF437F27A18B03DB1461DAEB">2025-02-13</mods:number>
</mods:detail>
<mods:detail id="FA4116EC7CF4BFF8466B9AA8898E91A5" type="volume">
<mods:number id="F6E3DEAD00AFA689FA292107CDF50F72">5588</mods:number>
</mods:detail>
<mods:detail id="063E1B0DB5726D8B8FB9F35E8492B13A" type="issue">
<mods:number id="30FB200B7398019220E760817A132A82">1</mods:number>
</mods:detail>
<mods:extent id="26AF34E192A4C2661FAE0E86A8B03EB4" unit="page">
<mods:start id="C787FD94DA0CAF6391BA580B18BA1970">77</mods:start>
<mods:end id="4969E1F722C7F7010F88627F613D4F56">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9C8678C40883F60C3F8F7C20DD44A8AE">
<mods:url id="86776FA32ED40F294D4BA6BBE3A8B7F3">https://doi.org/10.11646/zootaxa.5588.1.3</mods:url>
</mods:location>
<mods:classification id="E656E3552BC367B44B36126E7075E622">journal article</mods:classification>
<mods:identifier id="B7819D4C1E86830B85E9B1EBB84E0F47" type="DOI">10.11646/zootaxa.5588.1.3</mods:identifier>
<mods:identifier id="B3E836675651E9CF8E05FFFE0F5B01BD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EA1BB0A5327C86C6259C845B66A7330C" type="Zenodo-Dep">14896067</mods:identifier>
<mods:identifier id="3672B3FE927C274B4D2CA3F66403C55B" type="ZooBank">E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D</mods:identifier>
</mods:mods>
<treatment id="03A887D2FF81693835902920FD1CF859" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859" httpUri="http://treatment.plazi.org/id/03A887D2FF81693835902920FD1CF859" lastPageNumber="87" pageId="10" pageNumber="87">
<subSubSection id="C31B654FFF81693835902920FD7BFDF0" box="[151,656,546,573]" pageId="10" pageNumber="87" type="nomenclature">
<paragraph id="8BBE36C4FF81693835902920FD7BFDF0" blockId="10.[151,656,546,609]" box="[151,656,546,573]" pageId="10" pageNumber="87">
<heading id="D0F681A8FF81693835902920FD7BFDF0" bold="true" box="[151,656,546,573]" fontSize="11" level="1" pageId="10" pageNumber="87" reason="1">
<emphasis id="B975EAD6FF81693835902920FD7BFDF0" bold="true" box="[151,656,546,573]" pageId="10" pageNumber="87">
3.1.2.
<taxonomicName id="4C014D47FF81693835E62920FD7BFDF0" authority="(Santschi, 1912)" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[225,656,546,573]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF81693835E62920FE29FDF1" bold="true" box="[225,450,546,573]" italics="true" pageId="10" pageNumber="87">Fulakora elongata</emphasis>
(
<bibRefCitation id="EF904B35FF81693834D52920FD63FDF0" author="Santschi" box="[466,648,546,572]" firstAuthor="Santschi" pageId="10" pageNumber="87" pagination="519 - 534" refId="ref13214" refString="Santschi, F. (1912) Quelques fourmis de l'Amerique australe. Revue Suisse de Zoologie, 20, 519-534. https://doi.org/10.5962/bhl.part.19250" type="journal article" year="1912">Santschi, 1912</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF81693835902945FC46FBDD" pageId="10" pageNumber="87" type="description">
<paragraph id="8BBE36C4FF81693835902945FDF7FDAD" blockId="10.[151,656,546,609]" box="[151,540,583,609]" pageId="10" pageNumber="87">
(
<figureCitation id="133A2A41FF81693835982945FEE7FDAD" box="[159,268,583,609]" captionStart="FIGURE 7" captionStartId="11.[151,250,1607,1631]" captionTargetBox="[151,1435,181,1584]" captionTargetId="figure-19@11.[151,1435,181,1584]" captionTargetPageId="11" captionText="FIGURE 7. Fulakora elongata, worker of the karyotyped population (ANTWEB1047734). A, full-face view of head; B, profile view of the body; C, dorsum of mesosoma and petiole; D, anterior clypeal margin; E, genal tooth; F, profile of propodeum and petiole. Scales bars: 0.5mm (B), 0.2 mm (A, C, F), 0.05 mm (D), 0.02 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.14896082" httpUri="https://zenodo.org/record/14896082/files/figure.png" pageId="10" pageNumber="87">Figures 7</figureCitation>
;
<figureCitation id="133A2A41FF816938341F2945FEC3FDAD" box="[280,296,583,609]" captionStart="FIGURE 8" captionStartId="12.[151,250,1113,1137]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-30@12.[151,1435,181,1089]" captionTargetPageId="12" captionText="FIGURE 8. Fulakora elongata larva. A, head in profile; B, profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896084" httpUri="https://zenodo.org/record/14896084/files/figure.png" pageId="10" pageNumber="87">8</figureCitation>
;
<figureCitation id="133A2A41FF81693834322945FE73FDAD" box="[309,408,583,609]" captionStart="FIGURE 9" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[151,1436,959,1844]" captionTargetId="figure-334@13.[151,1436,959,1844]" captionTargetPageId="13" captionText="FIGURE 9. Morphological characters studied and their states. A, E, and L, Fulakora heraldoi (INPA-HYM-034964; images by Itanna Oliveira Fernandes); B and I, F. cleae (CASENT0102202; images by April Nobile) (zoom in B of F. elongata, ANTWEB1047734); C, G, and M, F.estevae(ANTWEB1047071);D, F.elongata (CASENT0915101; images by Z. Lieberman); F and K, F. elongata (ANTWEB1047734); H, F. armigera (CASENT0915647; images by Harald Bruckner); J, F. lurilabes (CASENT0172810; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896086" httpUri="https://zenodo.org/record/14896086/files/figure.png" pageId="10" pageNumber="87">9D, F, K</figureCitation>
;
<figureCitation id="133A2A41FF81693834A32945FE3CFDAD" box="[420,471,583,609]" captionStart="FIGURE 17" captionStartId="17.[151,250,780,804]" captionTargetBox="[151,1436,322,757]" captionTargetId="figure-181@17.[151,1436,322,757]" captionTargetPageId="17" captionText="FIGURE 17. A, F. elongata (CASENT0915100; images by Z. Lieberman); B, F. lurilabes (CASENT0172810; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896102" httpUri="https://zenodo.org/record/14896102/files/figure.png" pageId="10" pageNumber="87">17A</figureCitation>
;
<figureCitation id="133A2A41FF81693834E42945FDFFFDAD" box="[483,532,583,609]" captionStart="FIGURE 21" captionStartId="19.[151,250,1017,1041]" captionTargetBox="[158,1429,424,987]" captionTargetId="figure-116@19.[151,1436,417,994]" captionTargetPageId="19" captionText="FIGURE 21. A, Giemsa-stained karyotype of Fulakora elongata (2n=24; 16m + 8sm); B, the same for F. estevae (2n=24; 16m + 8sm). Scale bars: 5 μm. Abbreviations: m, metacentric; sm, submetacentric." figureDoi="http://doi.org/10.5281/zenodo.14896111" httpUri="https://zenodo.org/record/14896111/files/figure.png" pageId="10" pageNumber="87">21A</figureCitation>
)
</paragraph>
<paragraph id="8BBE36C4FF8169383590298DFC46FBDD" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
<emphasis id="B975EAD6FF8169383590298DFF0CFD65" bold="true" box="[151,231,655,681]" pageId="10" pageNumber="87">Larva.</emphasis>
TL 3.05, CW 0.19, CL 0.18, Masukos TL 2.39 (N=1). Larva with
<emphasis id="B975EAD6FF81693836F0298DFB81FD64" box="[1015,1130,655,680]" italics="true" pageId="10" pageNumber="87">meconium</emphasis>
, at advanced development stage. Small hypognathous head. In full-face view, head slightly longer than wide; occipital and lateral margins of head convex. Mandibles strongly sclerotized and slender, having less than half the lenght of head, apical teeth directed medio-posteriorly. In profile, posterior margin of mandible concave. Labrum bilobed, with lobes about the same width as the wider region of mandible in full-face view; lobes covering mandibles dorsally. Maxillary and labial palps not visualized; galea conical and slender, bearing two well-developed apical sensilla. Antennae small, round, posteriad the mid-length of the head (
<figureCitation id="133A2A41FF81693837DF2865FCC0FC4D" box="[728,811,871,897]" captionStart="FIGURE 8" captionStartId="12.[151,250,1113,1137]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-30@12.[151,1435,181,1089]" captionTargetPageId="12" captionText="FIGURE 8. Fulakora elongata larva. A, head in profile; B, profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896084" httpUri="https://zenodo.org/record/14896084/files/figure.png" pageId="10" pageNumber="87">Fig.8A</figureCitation>
). Body “pogonomyrmecoid”
<emphasis id="B975EAD6FF81693831712865FB5FFC4C" box="[1142,1204,871,896]" italics="true" pageId="10" pageNumber="87">sensu</emphasis>
<bibRefCitation id="EF904B35FF81693831BC2865FF0EFC69" author="Wheeler &amp; Wheeler" firstAuthor="Wheeler" pageId="10" pageNumber="87" pagination="1 - 108" refId="ref13844" refString="Wheeler, G. C. &amp; Wheeler, J. (1976) Ant larvae: review and synthesis. Memoirs of the Entomological Society of Washington, 7, 1-108." type="journal article" year="1976">Wheeler &amp; Wheeler (1976)</bibRefCitation>
; in profile somites slightly thickened posteriad, without any strong curve. Anus ventral (
<figureCitation id="133A2A41FF81693831D72889FACAFC69" box="[1232,1313,907,933]" captionStart="FIGURE 8" captionStartId="12.[151,250,1113,1137]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-30@12.[151,1435,181,1089]" captionTargetPageId="12" captionText="FIGURE 8. Fulakora elongata larva. A, head in profile; B, profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896084" httpUri="https://zenodo.org/record/14896084/files/figure.png" pageId="10" pageNumber="87">Fig.8B</figureCitation>
). Spiracles not visualized.
<specimenCount id="9D07FD4DFF816938344528ADFE5DFC05" box="[322,438,943,969]" count="2" pageId="10" pageNumber="87" type="generic" typeStatus="types">Two types</specimenCount>
of setae on body: (i) simple, slightly thickened, minute, stiff setae, present in all somites except the head, more abundant on dorsum; (ii) simple, thick, erect, stiff setae, about three times the size of
<typeStatus id="54BA8866FF816938305128D1FA6DFC21" box="[1366,1414,979,1005]" pageId="10" pageNumber="87">type</typeStatus>
i, densely present in all somites, sparsely distributed on the head (
<figureCitation id="133A2A41FF816938366628F5FC4BFBDD" box="[865,928,1015,1041]" captionStart="FIGURE 8" captionStartId="12.[151,250,1113,1137]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-30@12.[151,1435,181,1089]" captionTargetPageId="12" captionText="FIGURE 8. Fulakora elongata larva. A, head in profile; B, profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896084" httpUri="https://zenodo.org/record/14896084/files/figure.png" pageId="10" pageNumber="87">Fig.8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF81693835C02F18FF14FBB1" pageId="10" pageNumber="87" type="materials_examined">
<paragraph id="8BBE36C4FF81693835C02F18FF14FBB1" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
<emphasis id="B975EAD6FF81693835C02F18FE53FBF8" bold="true" box="[199,440,1050,1076]" pageId="10" pageNumber="87">Examined material.</emphasis>
<materialsCitation id="3B693C99FF81693834C42F19FF12FBB1" collectingDate="2023-03-31" collectionCode="CELC" collectorName="Jacintho G. F &amp; Chaul J." country="BRA" latitude="-20.758253" location="Mata da Biologia" longLatPrecision="1" longitude="-42.86205" municipality="Vicosa" pageId="10" pageNumber="87" specimenCode="GFJ000102" specimenCount="1" specimenCount-larva="1" stateProvince="MG">
<specimenCount id="9D07FD4DFF81693834C42F19FDDEFBF9" box="[451,565,1051,1077]" count="1" pageId="10" pageNumber="87" type="larva">One larva</specimenCount>
from the following collect event:
<collectingCountry id="F3167654FF81693836CA2F19FBE2FBF9" box="[973,1033,1051,1077]" pageId="10" pageNumber="87">BRA</collectingCountry>
,
<collectingRegion id="49C5F826FF816938311E2F19FBA2FBF9" box="[1049,1097,1051,1077]" pageId="10" pageNumber="87">MG</collectingRegion>
,
<collectingMunicipality id="6BDAACBEFF81693831502F19FB4DFBF9" box="[1111,1190,1051,1077]" pageId="10" pageNumber="87">Viçosa</collectingMunicipality>
,
<location id="8EDE601FFF81693831B12F19FA68FBF9" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693831B12F19FA68FBF9" box="[1206,1411,1051,1077]" country="BRA" latitude="-20.758253" longLatPrecision="1" longitude="-42.86205" municipality="Vicosa" name="Mata da Biologia" pageId="10" pageNumber="87" stateProvince="MG">Mata da Biologia</location>
,
<geoCoordinate id="EE355003FF81693830942F19FEE5FB95" degrees="20.758253" direction="south" orientation="latitude" pageId="10" pageNumber="87" precision="1" value="-20.758253">- 20.758253</geoCoordinate>
<geoCoordinate id="EE355003FF816938341A2F3CFE79FB95" box="[285,402,1086,1113]" degrees="42.86205" direction="west" orientation="longitude" pageId="10" pageNumber="87" precision="1" value="-42.86205">-42.86205</geoCoordinate>
,
<date id="FFBF1004FF81693834A22F3DFDCBFB95" box="[421,544,1087,1113]" pageId="10" pageNumber="87" value="2023-03-31">
<collectingDate id="EFFBE9ECFF81693834A22F3DFDCBFB95" box="[421,544,1087,1113]" pageId="10" pageNumber="87" value="2023-03-31">31.iii.2023</collectingDate>
</date>
,
<collectorName id="26F45312FF81693837332F3DFD21FB95" box="[564,714,1087,1113]" pageId="10" pageNumber="87">Jacintho G.F</collectorName>
,
<collectorName id="26F45312FF81693837DA2F3DFCAAFB95" box="[733,833,1087,1113]" pageId="10" pageNumber="87">Chaul J.</collectorName>
(ANTWEB1053725, collecting event
<specimenCode id="DBA79EBFFF81693830172F3DFA7EFB95" box="[1296,1429,1087,1113]" pageId="10" pageNumber="87">GFJ000102</specimenCode>
) [
<emphasis id="B975EAD6FF81693835A62F60FF04FBB0" bold="true" box="[161,239,1122,1148]" pageId="10" pageNumber="87">
<collectionCode id="ED10AE01FF81693835A62F60FF04FBB0" box="[161,239,1122,1148]" pageId="10" pageNumber="87">CELC</collectionCode>
</emphasis>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF81693835C02F85FB80FB6D" box="[199,1131,1159,1185]" pageId="10" pageNumber="87" type="biology_ecology">
<paragraph id="8BBE36C4FF81693835C02F85FB80FB6D" blockId="10.[151,1437,654,1941]" box="[199,1131,1159,1185]" pageId="10" pageNumber="87">
<emphasis id="B975EAD6FF81693835C02F85FEA0FB6D" bold="true" box="[199,331,1159,1185]" pageId="10" pageNumber="87">Karyology.</emphasis>
2n = 24 (16 metacentrics + 8 submetacentrics) (see 3.2 for discussion).
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF81693835C02FA8FD1CF859" pageId="10" pageNumber="87" type="discussion">
<paragraph id="8BBE36C4FF81693835C02FA8FE75FA99" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
<emphasis id="B975EAD6FF81693835C02FA8FEA4FB08" bold="true" box="[199,335,1194,1220]" pageId="10" pageNumber="87">Comments.</emphasis>
The most conspicuous difference between the larvae of both species is the
<typeStatus id="54BA8866FF81693831C22FA9FB1EFB09" box="[1221,1269,1195,1221]" pageId="10" pageNumber="87">type</typeStatus>
of pilosity:
<taxonomicName id="4C014D47FF816938308F2FA9FF02FB24" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF816938308F2FA9FF02FB24" italics="true" pageId="10" pageNumber="87">F. estevae</emphasis>
</taxonomicName>
has long, thin, and slightly curved setae (
<figureCitation id="133A2A41FF81693837C22FCDFCEFFB25" box="[709,772,1231,1257]" captionStart="FIGURE 6" captionStartId="9.[151,250,1530,1554]" captionTargetBox="[151,1435,741,1506]" captionTargetId="figure-274@9.[151,1435,741,1506]" captionTargetPageId="9" captionText="FIGURE 6. Fulakora estevae sp. nov., larva. A, head in profile; B. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="10" pageNumber="87">Fig.6</figureCitation>
), whereas
<taxonomicName id="4C014D47FF816938367A2FCDFC17FB25" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[893,1020,1231,1257]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF816938367A2FCDFC17FB25" box="[893,1020,1231,1257]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
has stiffer, shorter, and thicker setae (
<figureCitation id="133A2A41FF81693835A72FF1FF34FAC1" box="[160,223,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[151,250,1113,1137]" captionTargetBox="[151,1435,181,1089]" captionTargetId="figure-30@12.[151,1435,181,1089]" captionTargetPageId="12" captionText="FIGURE 8. Fulakora elongata larva. A, head in profile; B, profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896084" httpUri="https://zenodo.org/record/14896084/files/figure.png" pageId="10" pageNumber="87">Fig.8</figureCitation>
). It is worth noting that
<taxonomicName id="4C014D47FF81693834F52FF1FD9AFAC1" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[498,625,1267,1293]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF81693834F52FF1FD9AFAC1" box="[498,625,1267,1293]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
has overal denser pilosity on all surfaces compared to
<emphasis id="B975EAD6FF81693831E52FF1FAB2FAC0" box="[1250,1369,1267,1292]" italics="true" pageId="10" pageNumber="87">
<taxonomicName id="4C014D47FF81693831E52FF1FABEFAC0" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[1250,1365,1267,1292]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="estevae">F. estevae</taxonomicName>
.
</emphasis>
Also,
<taxonomicName id="4C014D47FF81693835902E15FEECFAFC" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[151,263,1303,1328]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF81693835902E15FEECFAFC" box="[151,263,1303,1328]" italics="true" pageId="10" pageNumber="87">F. estevae</emphasis>
</taxonomicName>
is bigger than the
<taxonomicName id="4C014D47FF81693834E32E15FD8FFAFD" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[484,612,1303,1329]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF81693834E32E15FD8FFAFD" box="[484,612,1303,1329]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
larvae, however, as we dont know exactly its developmental stage, no comparison is possible.
</paragraph>
<paragraph id="8BBE36C4FF81693835C02E5DFE8CF99D" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
We considered the specimens of our studied colonies to belong to
<taxonomicName id="4C014D47FF81693836AD2E5DFBCCFAB5" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[938,1063,1375,1401]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF81693836AD2E5DFBCCFAB5" box="[938,1063,1375,1401]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
, despite some variation observed among them and the
<typeStatus id="54BA8866FF81693834842E81FE58FA51" box="[387,435,1411,1437]" pageId="10" pageNumber="87">type</typeStatus>
species from
<collectingCountry id="F3167654FF81693837572E81FD59FA51" box="[592,690,1411,1437]" name="Uruguay" pageId="10" pageNumber="87">Uruguay</collectingCountry>
(
<bibRefCitation id="EF904B35FF81693837C72E81FC81FA51" author="Santschi" box="[704,874,1411,1437]" firstAuthor="Santschi" pageId="10" pageNumber="87" pagination="519 - 534" refId="ref13214" refString="Santschi, F. (1912) Quelques fourmis de l'Amerique australe. Revue Suisse de Zoologie, 20, 519-534. https://doi.org/10.5962/bhl.part.19250" type="journal article" year="1912">Santschi, 1912</bibRefCitation>
;
<bibRefCitation id="EF904B35FF81693836732E81FBF7FA51" accessDate="2024-05-14" author="AntWeb" box="[884,1052,1411,1437]" journalOrPublisher="California Academy of Science, San Francisco, California" pageId="10" pageNumber="87" publicationUrl="https://www.antweb.org" refId="ref11870" refString="AntWeb (2024) AntWeb. California Academy of Science, San Francisco, California. Available from: https: // www. antweb. org (accessed 14 May 2024)" title="AntWeb" type="url" year="2024">AntWeb, 2024</bibRefCitation>
). The complex taxonomic issue of this species concerning its infraspecific morphological variation requires further investigation. This is beyond the scope of this study, and involves the examination of the various
<typeStatus id="54BA8866FF81693836592EC9FC72FA29" box="[862,921,1483,1509]" pageId="10" pageNumber="87">types</typeStatus>
of subspecies currently synonymized under
<taxonomicName id="4C014D47FF816938308F2EC9FF12F9C5" baseAuthorityName="Santschi" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF816938308F2EC9FF12F9C5" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
, as well as the study of hundreds of
<typeStatus id="54BA8866FF816938379D2EEDFC91F9C5" box="[666,890,1519,1545]" isOtherTypeStatus="true" pageId="10" pageNumber="87">non-type specimens</typeStatus>
from a much broader geographical range across the Neotropics than we did. Future revisionary studies must consider the larval description is attributed to the Minas Gerais population.
</paragraph>
<paragraph id="8BBE36C4FF81693835C02D59FC70F8E5" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
<location id="8EDE601FFF81693835C02D59FF07F9B9" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693835C02D59FF07F9B9" box="[199,236,1627,1653]" country="Paraguay" county="Uruguayan" municipality="In" name="We" pageId="10" pageNumber="87">We</location>
examined, either physically at CELC or through images (
<bibRefCitation id="EF904B35FF81693836752D59FBF1F9B9" accessDate="2024-05-14" author="AntWeb" box="[882,1050,1627,1653]" journalOrPublisher="California Academy of Science, San Francisco, California" pageId="10" pageNumber="87" publicationUrl="https://www.antweb.org" refId="ref11870" refString="AntWeb (2024) AntWeb. California Academy of Science, San Francisco, California. Available from: https: // www. antweb. org (accessed 14 May 2024)" title="AntWeb" type="url" year="2024">AntWeb, 2024</bibRefCitation>
), specimens of the
<location id="8EDE601FFF81693831F72D59FABCF9B9" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693831F72D59FABCF9B9" box="[1264,1367,1627,1653]" country="Paraguay" county="Uruguayan" municipality="In" name="Brazilian" pageId="10" pageNumber="87">Brazilian</location>
states of
<location id="8EDE601FFF81693835B02D7DFEB9F955" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693835B02D7DFEB9F955" box="[183,338,1663,1689]" country="Paraguay" county="Uruguayan" municipality="In" name="Minas Gerais" pageId="10" pageNumber="87">Minas Gerais</location>
,
<location id="8EDE601FFF81693834592D7DFE46F955" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693834592D7DFE46F955" box="[350,429,1663,1689]" country="Paraguay" county="Uruguayan" municipality="In" name="Parana" pageId="10" pageNumber="87">Paraná</location>
, and
<location id="8EDE601FFF81693834EC2D7DFD7FF955" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693834EC2D7DFD7FF955" box="[491,660,1663,1689]" country="Paraguay" county="Uruguayan" municipality="In" name="Santa Catarina" pageId="10" pageNumber="87">Santa Catarina</location>
, and from the neighbor countries
<collectingCountry id="F3167654FF81693831112D7DFB94F955" box="[1046,1151,1663,1689]" name="Paraguay" pageId="10" pageNumber="87">Paraguay</collectingCountry>
,
<collectingCountry id="F3167654FF816938318D2D7DFB17F955" box="[1162,1276,1663,1689]" name="Argentina" pageId="10" pageNumber="87">Argentina</collectingCountry>
, and
<collectingCountry id="F3167654FF816938303D2D7DFA77F955" box="[1338,1436,1663,1689]" name="Uruguay" pageId="10" pageNumber="87">Uruguay</collectingCountry>
(
<bibRefCitation id="EF904B35FF81693835982DA1FEA0F971" accessDate="2024-05-14" author="AntWeb" box="[159,331,1699,1725]" journalOrPublisher="California Academy of Science, San Francisco, California" pageId="10" pageNumber="87" publicationUrl="https://www.antweb.org" refId="ref11870" refString="AntWeb (2024) AntWeb. California Academy of Science, San Francisco, California. Available from: https: // www. antweb. org (accessed 14 May 2024)" title="AntWeb" type="url" year="2024">AntWeb, 2024</bibRefCitation>
).
<location id="8EDE601FFF81693834592DA1FE69F971" LSID="urn:lsid:plazi:treatment:03A887D2FF81693835902920FD1CF859:8EDE601FFF81693834592DA1FE69F971" box="[350,386,1699,1725]" country="Paraguay" county="Uruguayan" municipality="In" name="All" pageId="10" pageNumber="87">All</location>
specimens shared the following characters: (i) small overall body size (TL 2.743.18), (ii) small genal teeth, (iii) vertex smooth, at most with piligerous punctures, (iii) presence of meso-metapleural suture/ line, and (iv) propodeal pale patches.
<collectingMunicipality id="6BDAACBEFF816938373E2DE9FDBBF8C9" box="[569,592,1771,1797]" pageId="10" pageNumber="87">In</collectingMunicipality>
comparison to the
<collectingCounty id="62DF4E48FF81693836222DE9FC4AF8C9" box="[805,929,1771,1797]" pageId="10" pageNumber="87">Uruguayan</collectingCounty>
<typeStatus id="54BA8866FF81693836AF2DE9FC30F8C9" box="[936,987,1771,1797]" pageId="10" pageNumber="87">type</typeStatus>
, the Minas Gerais population has longer mandibles, slightly longer and sharper genal teeth, and broader head.
</paragraph>
<paragraph id="8BBE36C4FF81693835C02C31FD1CF859" blockId="10.[151,1437,654,1941]" pageId="10" pageNumber="87">
The
<typeStatus id="54BA8866FF81693835FE2C31FEDFF881" box="[249,308,1843,1869]" pageId="10" pageNumber="87">types</typeStatus>
of
<taxonomicName id="4C014D47FF816938345F2C31FE3CF881" baseAuthorityName="Lattke" baseAuthorityYear="1991" box="[344,471,1843,1869]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="lurilabes">
<emphasis id="B975EAD6FF816938345F2C31FE3CF881" box="[344,471,1843,1869]" italics="true" pageId="10" pageNumber="87">F. lurilabes</emphasis>
</taxonomicName>
, a very similar species, differs from all examined populations of
<taxonomicName id="4C014D47FF81693831AE2C31FACDF881" baseAuthorityName="Santschi" baseAuthorityYear="1912" box="[1193,1318,1843,1869]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF81693831AE2C31FACDF881" box="[1193,1318,1843,1869]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
by: lateral margins of head mildly tapering posteriorly; opaque/coriaceous vertexal area; A3 cinctus narrower than propodeal spiracle; and larger total length (
<tableCitation id="C683037FFF81693837062C79FDBCF859" box="[513,599,1915,1941]" captionStart="TABLE 1" captionStartId="13.[152,240,150,174]" captionTargetPageId="13" captionText="TABLE 1. Comparison of characters, measurements and indexes among Fulakora species (see text for the description of each character state). Specimens studied: four F. estevae (ANTWEB1047071, ANTWEB1047070, ANTWEB1047069, UFV-LABECOL-000012), three F. agostii (CASENT0102201, CASENT0101451, ANTWEB1047718), one F. armigera (CASENT0915647),twoF.cleae(CASENT0102202,CASENT0101452),tenF.elongata(CASENT0915100,CASENT0915098, CASENT0102471, CASENT0178869, UFV-LABECOL-000027, CASENT0915101, UFV-LABECOL-000049, UFV- LABECOL-008530, ANTWEB1047734, ANTWEB1047061); three F. lurilabes (CASENT0102199, CASENT0172810, CASENT0172811), and one F. heraldoi (INPA-HYM-034964) (For data per specimen see Appendix 2)." pageId="10" pageNumber="87">Table 1</tableCitation>
;
<figureCitation id="133A2A41FF81693837642C79FD01F859" box="[611,746,1915,1941]" captionStart="APPENDIX 2" captionStartId="23.[152,304,978,1004]" captionTargetPageId="23" captionText="APPENDIX 2. Comparison of characters, measurements and indexes among Fulakora species. Description of each measurement is on the section 2 (Measurements and indices), characters states is on section 3.1.3." pageId="10" pageNumber="87">Appendix 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,755 @@
<document id="BCBB4B938592CA073FBB7846205BED2C" ID-DOI="10.11646/zootaxa.5588.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896067" ID-ZooBank="E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1740002781111" checkinUser="plazi" docAuthor="Jacintho, Gabriela De Figueiredo, Chaul, Júlio Cezar Mário, Teixeira, Gisele Amaro, Barros, Luísa Antônia Campos, Lopes, Denilce Meneses &amp; Serrão, José Eduardo" docDate="2025" docId="03A887D2FF85692035902A48FD06FB7C" docLanguage="en" docName="zootaxa.5588.1.3.pdf" docOrigin="Zootaxa 5588 (1)" docSource="https://doi.org/10.11646/zootaxa.5588.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fulakora Mann 1919" docType="key" docVersion="1" lastPageNumber="95" masterDocId="FF91FFAAFF8B693235072B02FFEBFFCC" masterDocTitle="A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species" masterLastPageNumber="104" masterPageNumber="77" pageNumber="91" updateTime="1740509805003" updateUser="guilherme" zenodo-license-document="CLOSED">
<mods:mods id="6DC89A2EC8C375F4C71ED113875086E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="59F0EB96CE45BE8B9EB724BD09FCCE51">
<mods:title id="ED38BBF7EA932B62DE5884CD97E42B9F">A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="6DCF2473931B603761B0A37B72910993" type="personal">
<mods:role id="2FE74AFFC22ECAC18A40E50995F904C1">
<mods:roleTerm id="DBED7A310A72788B3E978066608079E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC9A3B13465B60D6E8C0493974AE4C42">Jacintho, Gabriela De Figueiredo</mods:namePart>
<mods:affiliation id="8D802D76255A3AB1AD4703F037F90D62">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="6DFEC54A95C20BB2334BF28AA331C7DB" type="personal">
<mods:role id="6FCA85F42F97F80C1D759F1E52788ED2">
<mods:roleTerm id="736ECEBD51CC80E9C2CFC763E3CE9C5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2878CDB298A04A45CDDAB2BDBC9C390D">Chaul, Júlio Cezar Mário</mods:namePart>
<mods:affiliation id="D22EB8ACB116B5B927522A883FCFDF45">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="E4FEFE4752C51BDCF989C2B457560D0F" type="personal">
<mods:role id="0C1ABB0556793243A8451EDEF532FFA2">
<mods:roleTerm id="E9D960EA755D383B5082667DD95F23D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0CE2A4539A8D8BFC29128D77B335F2C">Teixeira, Gisele Amaro</mods:namePart>
<mods:affiliation id="C6CFA2730F0D4C53F06F87C16E157510">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="5E3649CF2E5C29BA794E9E7DFAA3B4AF" type="personal">
<mods:role id="55231C33551F80B952A1C017EE9ACB6C">
<mods:roleTerm id="759D8B96D71A4E558E385728D72010AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4B752C19CF61646E432E6E7F252DFD8">Barros, Luísa Antônia Campos</mods:namePart>
<mods:affiliation id="EA2F5E33108B7188F6CFEBD326169D5A">Universidade Federal do Amapá, Campus Binacional, 68980 - 000, Oiapoque, Amapá, Brazil</mods:affiliation>
</mods:name>
<mods:name id="3D27EB96CC87AE3F3D6C207733B2D468" type="personal">
<mods:role id="0DE7EA63226631692334F912309872B2">
<mods:roleTerm id="57D39AA12EF926FC2D54FF1206628809">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81F960DC42F43179EBB113B75D0B1806">Lopes, Denilce Meneses</mods:namePart>
<mods:affiliation id="F7E7C8B1DA4536160E758759DBB67BF6">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="2FAFB9973A1E16F339FA610E1DAE8BAA" type="personal">
<mods:role id="8549F3CA12FB291A19D928BA127343F5">
<mods:roleTerm id="6EB4056E31197820C32603C6C0E6E9F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF880A5A5B1175BDAD35FB048EF6BA57">Serrão, José Eduardo</mods:namePart>
<mods:affiliation id="9DDCB327FBA087AC6751CE7C406C0C76">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9DB5425918C80299B950D227D8A070B5">text</mods:typeOfResource>
<mods:relatedItem id="B35768E367C785070BFBECE46C47BA90" type="host">
<mods:titleInfo id="30362D4202A75164CD96F911B99F167B">
<mods:title id="896052EE61583ABD061974A30EA1442D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="198F09F9BD8AF3B975E1A4B39E0BCE33">
<mods:date id="EC5A08F4F5D213E168DA0EBEE5679FD3">2025</mods:date>
<mods:detail id="1314959359C1F357DF7C70292F53AA91" type="pubDate">
<mods:number id="AF636A9885F19171CE1A2AB88EC464F2">2025-02-13</mods:number>
</mods:detail>
<mods:detail id="B29CDBFE9C4616F0FA517308E8FECC45" type="volume">
<mods:number id="4C601475B1CF28C06D986C4DC56F3B7F">5588</mods:number>
</mods:detail>
<mods:detail id="2620C996410C2C728B51C2CBA3C5D995" type="issue">
<mods:number id="D4704E7BCC96D3E997CB44D889CA99C4">1</mods:number>
</mods:detail>
<mods:extent id="5A507E8FA355C1DDE50F706853FBC8B6" unit="page">
<mods:start id="35E294B9E4CBDD3FE980E95714D6E983">77</mods:start>
<mods:end id="3120A554792AF5943B8E5309785035E9">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1C971B80AE28AECBCD1B1612B2006D51">
<mods:url id="AD3E4CBF8EDD473BC84130E9F5DFBE60">https://doi.org/10.11646/zootaxa.5588.1.3</mods:url>
</mods:location>
<mods:classification id="F1D8EFF5DA0BAEBE1E9A50F1DCB77F41">journal article</mods:classification>
<mods:identifier id="378C0737AFA59210334517515E9718E3" type="DOI">10.11646/zootaxa.5588.1.3</mods:identifier>
<mods:identifier id="C0A19600AB7A71F8073E9B643EE501C8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="637F4FB6284B99D2F6C8F2BDDEF6E53A" type="Zenodo-Dep">14896067</mods:identifier>
<mods:identifier id="036344A1D3BECD8965F7C5026D2D1BEA" type="ZooBank">E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D</mods:identifier>
</mods:mods>
<treatment id="03A887D2FF85692035902A48FD06FB7C" LSID="urn:lsid:plazi:treatment:03A887D2FF85692035902A48FD06FB7C" httpUri="http://treatment.plazi.org/id/03A887D2FF85692035902A48FD06FB7C" lastPageId="18" lastPageNumber="95" pageId="14" pageNumber="91">
<subSubSection id="C31B654FFF85693C35902A48FC3EFEA9" box="[151,981,330,357]" pageId="14" pageNumber="91" type="nomenclature">
<paragraph id="8BBE36C4FF85693C35902A48FC3EFEA9" blockId="14.[151,981,330,357]" box="[151,981,330,357]" pageId="14" pageNumber="91">
<heading id="D0F681A8FF85693C35902A48FC3EFEA9" bold="true" box="[151,981,330,357]" fontSize="11" level="1" pageId="14" pageNumber="91" reason="1">
<emphasis id="B975EAD6FF85693C35902A48FC3EFEA9" bold="true" box="[151,981,330,357]" pageId="14" pageNumber="91">
3.1.4. Key to the Neotropical species of
<taxonomicName id="4C014D47FF85693C37782A48FD1AFEA9" authorityName="Mann" authorityYear="1919" box="[639,753,330,357]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD6FF85693C37782A48FD1AFEA9" bold="true" box="[639,753,330,357]" italics="true" pageId="14" pageNumber="91">Fulakora</emphasis>
</taxonomicName>
based on workers
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF85693C35902A91FE91FDD5" pageId="14" pageNumber="91" type="description">
<paragraph id="8BBE36C4FF85693C35902A91FA8AFE1D" blockId="14.[151,1436,403,537]" pageId="14" pageNumber="91">
In the Neotropics, any
<taxonomicName id="4C014D47FF85693C348A2A91FDAAFE61" authorityName="Forel" authorityYear="1893" box="[397,577,403,429]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="98" phylum="Arthropoda" rank="subFamily" subFamily="Amblyoponinae">Amblyoponinae</taxonomicName>
species that does not have the diagnostic characters of
<emphasis id="B975EAD6FF85693C31902A91FAE4FE61" box="[1175,1295,403,429]" italics="true" pageId="14" pageNumber="91">Prinopelta</emphasis>
, particularly the typical tridentate mandibles of that genus (
<bibRefCitation id="EF904B35FF85693C37992AB5FC46FE1D" author="Ladino &amp; Feitosa" box="[670,941,439,465]" firstAuthor="Ladino" pageId="14" pageNumber="91" pagination="201 - 249" refId="ref12650" refString="Ladino, N. &amp; Feitosa, R. M. (2020) Taxonomic revision of the genus Prionopelta Mayr, 1866 (Formicidae: Amblyoponinae) for the Neotropical region. Zootaxa, 4821 (2), 201-249. https://doi.org/10.11646/zootaxa.4821.2.1" type="journal article" year="2020">Ladino &amp; Feitosa, 2020</bibRefCitation>
), is necessarily a member of
<taxonomicName id="4C014D47FF85693C31F52AB5FAB1FE1D" authorityName="Mann" authorityYear="1919" box="[1266,1370,439,465]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAD6FF85693C31F52AB5FAB1FE1D" box="[1266,1370,439,465]" italics="true" pageId="14" pageNumber="91">Fulakora</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBE36C4FF85693C35C02AD9FE91FDD5" blockId="14.[151,1436,403,537]" pageId="14" pageNumber="91">
<taxonomicName id="4C014D47FF85693C35C02AD9FDB2FE39" authority="(Kusnezov, 1955)" baseAuthorityName="Kusnezov" baseAuthorityYear="1955" box="[199,601,474,501]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B975EAD6FF85693C35C02AD9FE60FE38" box="[199,395,475,501]" italics="true" pageId="14" pageNumber="91">Fulakora minima</emphasis>
(Kusnezov, 1955)
</taxonomicName>
and
<taxonomicName id="4C014D47FF85693C37972AD9FBC9FE39" authority="(Kusnezov, 1955)" baseAuthorityName="Kusnezov" baseAuthorityYear="1955" box="[656,1058,474,501]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="egregia">
<emphasis id="B975EAD6FF85693C37972AD9FCBFFE38" box="[656,852,475,501]" italics="true" pageId="14" pageNumber="91">Fulakora egregia</emphasis>
(Kusnezov, 1955)
</taxonomicName>
are only known by males and are not considered here.
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF85692335902941FA77F820" lastPageId="17" lastPageNumber="94" pageId="14" pageNumber="91" type="key">
<key id="FF9061F5FF85692335902941FA77F820" lastPageId="17" lastPageNumber="94" pageId="14" pageNumber="91">
<keyStep id="30F52E61FF85693C35902941FA77FD02" pageId="14" pageNumber="91">
<paragraph id="8BBE36C4FF85693C35902941FA77FD5F" blockId="14.[151,1436,579,719]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C35902941FA77FD5F" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902941FF41FD94" bold="true" box="[151,170,579,600]" pageId="14" pageNumber="91">1.</emphasis>
Frontal lobes widely separated. Mandible robust, thick along most of its length. Clypeal chaetae not rising from clypeal tubercles. Genal tooth absent. Pretarsal claws of hind legs hypertrophied (Central America up to northern
<collectingCountry id="F3167654FF85693C31DE2963FAFEFDBB" box="[1241,1301,609,631]" name="Brazil" pageId="14" pageNumber="91">Brazil</collectingCountry>
) (
<figureCitation id="133A2A41FF85693C302E2963FA91FDBB" box="[1321,1402,609,631]" captionStart="FIGURE 10" captionStartId="14.[151,250,978,1002]" captionTargetBox="[151,1436,743,954]" captionTargetId="figure-318@14.[151,1436,743,954]" captionTargetPageId="14" captionText="FIGURE 10. A, Fulakora mystriops (CASENT0173551; images by April Nobile); B, F. cleae (CASENT0101452; images by April Nobile); C, F. degenerata (CASENT0610643; images by Michele Esposito). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896088" httpUri="https://zenodo.org/record/14896088/files/figure.png" pageId="14" pageNumber="91">Fig.10A</figureCitation>
)........................................................................................
<taxonomicName id="4C014D47FF85693C31F4297CFA77FD5F" authority="Brown" authorityName="Brown" box="[1267,1436,638,659]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="mystriops">
<emphasis id="B975EAD6FF85693C31F4297CFA77FD5F" bold="true" box="[1267,1436,638,659]" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C31F4297CFAB9FD5F" bold="true" box="[1267,1362,638,659]" italics="true" pageId="14" pageNumber="91">mystriops</emphasis>
Brown
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF85693C3590299EFA77FD02" blockId="14.[151,1436,579,719]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C3590299EFA77FD02" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C3590299EFF74FD7D" bold="true" box="[151,159,668,689]" pageId="14" pageNumber="91">-</emphasis>
Frontal lobes closely approximated. Mandible delicate, becoming thin and sharp apically. Clypeal chaetae rising from clypeal tubercles. Genal tooth usually present. Pretarsal claws of hind legs not conspicuously enlarged (
<figureCitation id="133A2A41FF85693C317E29BBFB05FD02" box="[1145,1262,697,719]" captionStart="FIGURE 10" captionStartId="14.[151,250,978,1002]" captionTargetBox="[151,1436,743,954]" captionTargetId="figure-318@14.[151,1436,743,954]" captionTargetPageId="14" captionText="FIGURE 10. A, Fulakora mystriops (CASENT0173551; images by April Nobile); B, F. cleae (CASENT0101452; images by April Nobile); C, F. degenerata (CASENT0610643; images by Michele Esposito). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896088" httpUri="https://zenodo.org/record/14896088/files/figure.png" pageId="14" pageNumber="91">Figs.10B, C</figureCitation>
)............
<emphasis id="B975EAD6FF85693C309729BBFA77FD02" bold="true" box="[1424,1436,697,718]" pageId="14" pageNumber="91">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF85693C359028D0FAB9FBC2" ID-DOI="http://doi.org/10.5281/zenodo.14896088" ID-Zenodo-Dep="14896088" httpUri="https://zenodo.org/record/14896088/files/figure.png" pageId="14" pageNumber="91" startId="14.[151,250,978,1002]" targetBox="[151,1436,743,954]" targetPageId="14" targetType="figure">
<paragraph id="8BBE36C4FF85693C359028D0FAB9FBC2" blockId="14.[151,1436,978,1038]" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C359028D0FED1FC26" bold="true" box="[151,314,978,1002]" pageId="14" pageNumber="91">FIGURE 10. A</emphasis>
,
<taxonomicName id="4C014D47FF85693C344028D0FDFBFC26" box="[327,528,978,1002]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="mystriops">
<emphasis id="B975EAD6FF85693C344028D0FDFBFC26" box="[327,528,978,1002]" italics="true" pageId="14" pageNumber="91">Fulakora mystriops</emphasis>
</taxonomicName>
(CASENT0173551; images by April Nobile);
<emphasis id="B975EAD6FF85693C36E928D0FC14FC26" bold="true" box="[1006,1023,978,1002]" pageId="14" pageNumber="91">B</emphasis>
,
<taxonomicName id="4C014D47FF85693C310B28D1FBB0FC26" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1036,1115,978,1002]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF85693C310B28D1FBB0FC26" box="[1036,1115,978,1002]" italics="true" pageId="14" pageNumber="91">F. cleae</emphasis>
</taxonomicName>
(CASENT0101452; images by April Nobile);
<emphasis id="B975EAD6FF85693C342928F4FEABFBC2" bold="true" box="[302,320,1014,1038]" pageId="14" pageNumber="91">C</emphasis>
,
<taxonomicName id="4C014D47FF85693C344A28F5FE33FBC2" box="[333,472,1014,1038]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF85693C344A28F5FE33FBC2" box="[333,472,1014,1038]" italics="true" pageId="14" pageNumber="91">F. degenerata</emphasis>
</taxonomicName>
(CASENT0610643; images by Michele Esposito). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF85693C35902F5FFA77FB06" pageId="14" pageNumber="91">
<paragraph id="8BBE36C4FF85693C35902F5FFA77FB5C" blockId="14.[151,1436,1117,1227]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C35902F5FFA77FB5C" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902F5FFF42FBBE" bold="true" box="[151,169,1117,1138]" pageId="14" pageNumber="91">2.</emphasis>
Clypeal chaetae thick, large, and blunt, originating from a platform. Genal tooth absent (Central Amazonia) (
<figureCitation id="133A2A41FF85693C31F12F5FFAADFBBF" box="[1270,1350,1117,1139]" captionStart="FIGURE 11" captionStartId="14.[151,250,1592,1616]" captionTargetBox="[151,1436,1252,1569]" captionTargetId="figure-571@14.[151,1436,1252,1569]" captionTargetPageId="14" captionText="FIGURE 11. A, Fulakora heraldoi (INPA-HYM-034964; images by Itanna Oliveira Fernandes); B, F. armigera (CASENT0915647; images by Harald Bruckner). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896090" httpUri="https://zenodo.org/record/14896090/files/figure.png" pageId="14" pageNumber="91">Fig.11A</figureCitation>
)....................................................................................
<taxonomicName id="4C014D47FF85693C31912F78FA77FB5C" authority="Lacau &amp; Delabie" authorityName="Lacau &amp; Delabie" box="[1174,1436,1146,1168]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="heraldoi">
<emphasis id="B975EAD6FF85693C31912F78FA77FB5C" bold="true" box="[1174,1436,1146,1168]" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C31912F78FB02FB5C" bold="true" box="[1174,1257,1146,1168]" italics="true" pageId="14" pageNumber="91">heraldoi</emphasis>
Lacau &amp; Delabie
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF85693C35902F9AFA77FB06" blockId="14.[151,1436,1117,1227]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C35902F9AFA77FB06" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902F9AFF74FB61" bold="true" box="[151,159,1176,1197]" pageId="14" pageNumber="91">-</emphasis>
Clypeal chaetae tooth-like, small, and usually sharp-tipped, originated from clypeal tubercles. Genal tooth present or absent (
<figureCitation id="133A2A41FF85693C35EE2FB7FEDCFB07" box="[233,311,1205,1227]" captionStart="FIGURE 11" captionStartId="14.[151,250,1592,1616]" captionTargetBox="[151,1436,1252,1569]" captionTargetId="figure-571@14.[151,1436,1252,1569]" captionTargetPageId="14" captionText="FIGURE 11. A, Fulakora heraldoi (INPA-HYM-034964; images by Itanna Oliveira Fernandes); B, F. armigera (CASENT0915647; images by Harald Bruckner). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896090" httpUri="https://zenodo.org/record/14896090/files/figure.png" pageId="14" pageNumber="91">Fig.11B</figureCitation>
)............................................................................................
<emphasis id="B975EAD6FF85693C30972FB7FA77FB06" bold="true" box="[1424,1436,1205,1226]" pageId="14" pageNumber="91">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF85693C35902D3AFBEFF9B9" ID-DOI="http://doi.org/10.5281/zenodo.14896090" ID-Zenodo-Dep="14896090" httpUri="https://zenodo.org/record/14896090/files/figure.png" pageId="14" pageNumber="91" startId="14.[151,250,1592,1616]" targetBox="[151,1436,1252,1569]" targetPageId="14" targetType="figure">
<paragraph id="8BBE36C4FF85693C35902D3AFBEFF9B9" blockId="14.[151,1436,1592,1653]" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902D3AFEA7F99C" bold="true" box="[151,332,1592,1616]" pageId="14" pageNumber="91">FIGURE 11. A</emphasis>
,
<taxonomicName id="4C014D47FF85693C34632D3BFDC1F99D" box="[356,554,1593,1617]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="heraldoi">
<emphasis id="B975EAD6FF85693C34632D3BFDC1F99D" box="[356,554,1593,1617]" italics="true" pageId="14" pageNumber="91">Fulakora heraldoi</emphasis>
</taxonomicName>
(INPA-HYM-034964; images by Itanna Oliveira Fernandes);
<emphasis id="B975EAD6FF85693C31F62D3BFAE9F99D" bold="true" box="[1265,1282,1593,1617]" pageId="14" pageNumber="91">B</emphasis>
,
<taxonomicName id="4C014D47FF85693C301D2D3BFA77F99C" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[1306,1436,1593,1616]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="91" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="B975EAD6FF85693C301D2D3BFA77F99C" box="[1306,1436,1593,1616]" italics="true" pageId="14" pageNumber="91">F. armigera</emphasis>
</taxonomicName>
(CASENT0915647; images by Harald Bruckner). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF85693C35902DC6FA77F8FD" pageId="14" pageNumber="91">
<paragraph id="8BBE36C4FF85693C35902DC6FA77F93A" blockId="14.[151,1437,1732,1842]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C35902DC6FA77F93A" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902DC6FF42F915" bold="true" box="[151,169,1732,1753]" pageId="14" pageNumber="91">3.</emphasis>
Chilean species. Mandibles medially thickened, as if forming a distinct basal margin. Genal tooth very small and thin. Coloration brownish or reddish, never black (
<figureCitation id="133A2A41FF85693C372D2DE3FD4AF93A" box="[554,673,1761,1783]" captionStart="FIGURE 12" captionStartId="15.[151,250,664,688]" captionTargetBox="[151,1436,181,640]" captionTargetId="figure-19@15.[151,1436,181,640]" captionTargetPageId="15" captionText="FIGURE 12. A, Fulakora monrosi (CASTYPE06893; images by April Nobile); B, F. chilensis (CASENT0915648; images by Harald Bruckner); C, F. cleae (CASENT0101452; images by April Nobile); F. degenerata (CASENT0610643; images by Michele Esposito).All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896092" httpUri="https://zenodo.org/record/14896092/files/figure.png" pageId="14" pageNumber="91">Figs.12A, B</figureCitation>
)......................................................
<emphasis id="B975EAD6FF85693C303D2DE3FAADF93A" bold="true" box="[1338,1350,1761,1782]" pageId="14" pageNumber="91">4</emphasis>
(Note 2)
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF85693C35902DFDFA77F8FD" blockId="14.[151,1437,1732,1842]" pageId="14" pageNumber="91">
<keyLead id="30F095F1FF85693C35902DFDFA77F8FD" pageId="14" pageNumber="91">
<emphasis id="B975EAD6FF85693C35902DFDFF74F8D8" bold="true" box="[151,159,1791,1812]" pageId="14" pageNumber="91">-</emphasis>
Occurring elsewhere in South America or Central America. Mandibles without a distinct basal margin. Genal tooth either small, large or absent. Coloration either yellowish, brownish, reddish or black (
<figureCitation id="133A2A41FF85693C369C2C1EFBFAF8FD" box="[923,1041,1820,1842]" captionStart="FIGURE 12" captionStartId="15.[151,250,664,688]" captionTargetBox="[151,1436,181,640]" captionTargetId="figure-19@15.[151,1436,181,640]" captionTargetPageId="15" captionText="FIGURE 12. A, Fulakora monrosi (CASTYPE06893; images by April Nobile); B, F. chilensis (CASENT0915648; images by Harald Bruckner); C, F. cleae (CASENT0101452; images by April Nobile); F. degenerata (CASENT0610643; images by Michele Esposito).All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896092" httpUri="https://zenodo.org/record/14896092/files/figure.png" pageId="14" pageNumber="91">Figs.12C, D</figureCitation>
)...............................
<emphasis id="B975EAD6FF85693C30972C1EFA77F8FD" bold="true" box="[1424,1436,1820,1841]" pageId="14" pageNumber="91">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF84693D3590299AFD27FD34" ID-DOI="http://doi.org/10.5281/zenodo.14896092" ID-Zenodo-Dep="14896092" httpUri="https://zenodo.org/record/14896092/files/figure.png" pageId="15" pageNumber="92" startId="15.[151,250,664,688]" targetBox="[151,1436,181,640]" targetPageId="15" targetType="figure">
<paragraph id="8BBE36C4FF84693D3590299AFD27FD34" blockId="15.[151,1436,664,760]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D3590299AFED7FD7C" bold="true" box="[151,316,664,688]" pageId="15" pageNumber="92">FIGURE 12. A</emphasis>
,
<taxonomicName id="4C014D47FF84693D344C299AFDEEFD7C" box="[331,517,664,688]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="monrosi">
<emphasis id="B975EAD6FF84693D344C299AFDEEFD7C" box="[331,517,664,688]" italics="true" pageId="15" pageNumber="92">Fulakora monrosi</emphasis>
</taxonomicName>
(CASTYPE06893; images by April Nobile);
<emphasis id="B975EAD6FF84693D36E3299AFC1EFD7C" bold="true" box="[996,1013,664,688]" pageId="15" pageNumber="92">B</emphasis>
,
<taxonomicName id="4C014D47FF84693D3103299BFB93FD7C" box="[1028,1144,664,688]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B975EAD6FF84693D3103299BFB93FD7C" box="[1028,1144,664,688]" italics="true" pageId="15" pageNumber="92">F. chilensis</emphasis>
</taxonomicName>
(CASENT0915648; images by Harald Bruckner);
<emphasis id="B975EAD6FF84693D347D29BEFE67FD18" bold="true" box="[378,396,700,724]" pageId="15" pageNumber="92">C</emphasis>
,
<taxonomicName id="4C014D47FF84693D349C29BFFE01FD18" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[411,490,700,724]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF84693D349C29BFFE01FD18" box="[411,490,700,724]" italics="true" pageId="15" pageNumber="92">F. cleae</emphasis>
</taxonomicName>
(CASENT0101452; images by April Nobile);
<taxonomicName id="4C014D47FF84693D36CA29BFFBB2FD18" box="[973,1113,700,724]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF84693D36CA29BFFBB2FD18" box="[973,1113,700,724]" italics="true" pageId="15" pageNumber="92">F. degenerata</emphasis>
</taxonomicName>
(CASENT0610643; images by Michele Esposito). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF84693D35902845FA77FCB5" pageId="15" pageNumber="92">
<paragraph id="8BBE36C4FF84693D35902845FA77FC90" blockId="15.[151,1436,839,890]" box="[151,1436,839,861]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902845FA77FC90" box="[151,1436,839,861]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902845FF42FC90" bold="true" box="[151,169,839,860]" pageId="15" pageNumber="92">4.</emphasis>
Sculpturing on head dorsum shallow, head mostly smooth. Metanotal suture poorly marked (
<figureCitation id="133A2A41FF84693D315C2845FB39FC90" box="[1115,1234,839,861]" captionStart="FIGURE 13" captionStartId="15.[151,250,1415,1439]" captionTargetBox="[151,1436,944,1391]" captionTargetId="figure-109@15.[151,1436,944,1391]" captionTargetPageId="15" captionText="FIGURE 13. A and B, Fulakora monrosi (CASTYPE06893; images by April Nobile), C and D, F. chilensis (CASENT0915648; images by Harald Bruckner). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896094" httpUri="https://zenodo.org/record/14896094/files/figure.png" pageId="15" pageNumber="92">Figs.13A, B</figureCitation>
)...
<taxonomicName id="4C014D47FF84693D30072845FA77FC90" authority="Brown" authorityName="Brown" box="[1280,1436,839,860]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="monrosi">
<emphasis id="B975EAD6FF84693D30072845FA77FC90" bold="true" box="[1280,1436,839,860]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D30072845FABAFC90" bold="true" box="[1280,1361,839,860]" italics="true" pageId="15" pageNumber="92">monrosi</emphasis>
Brown
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF84693D35902866FA77FCB5" blockId="15.[151,1436,839,890]" box="[151,1436,868,890]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902866FA77FCB5" box="[151,1436,868,890]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902866FF74FCB5" bold="true" box="[151,159,868,889]" pageId="15" pageNumber="92">-</emphasis>
Sculpturing on head dorsum conspicuous, head mostly opaque. Metanotal suture deep (
<figureCitation id="133A2A41FF84693D312F2866FB75FCB5" box="[1064,1182,868,890]" captionStart="FIGURE 13" captionStartId="15.[151,250,1415,1439]" captionTargetBox="[151,1436,944,1391]" captionTargetId="figure-109@15.[151,1436,944,1391]" captionTargetPageId="15" captionText="FIGURE 13. A and B, Fulakora monrosi (CASTYPE06893; images by April Nobile), C and D, F. chilensis (CASENT0915648; images by Harald Bruckner). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896094" httpUri="https://zenodo.org/record/14896094/files/figure.png" pageId="15" pageNumber="92">Figs.13C, D</figureCitation>
)........
<taxonomicName id="4C014D47FF84693D30002866FA77FCB5" authority="Mayr" authorityName="Mayr" box="[1287,1436,868,890]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B975EAD6FF84693D30002866FA77FCB5" bold="true" box="[1287,1436,868,890]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D30002866FAB5FCB6" bold="true" box="[1287,1374,868,890]" italics="true" pageId="15" pageNumber="92">chilensis</emphasis>
Mayr
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF84693D35902E85FCD8FA0F" ID-DOI="http://doi.org/10.5281/zenodo.14896094" ID-Zenodo-Dep="14896094" httpUri="https://zenodo.org/record/14896094/files/figure.png" pageId="15" pageNumber="92" startId="15.[151,250,1415,1439]" targetBox="[151,1436,944,1391]" targetPageId="15" targetType="figure">
<paragraph id="8BBE36C4FF84693D35902E85FCD8FA0F" blockId="15.[151,1437,1415,1475]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902E85FEDFFA53" bold="true" box="[151,308,1415,1439]" pageId="15" pageNumber="92">FIGURE 13. A</emphasis>
and
<emphasis id="B975EAD6FF84693D34642E85FE9FFA53" bold="true" box="[355,372,1415,1439]" pageId="15" pageNumber="92">B</emphasis>
,
<taxonomicName id="4C014D47FF84693D34782E85FDDEFA53" box="[383,565,1415,1439]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="monrosi">
<emphasis id="B975EAD6FF84693D34782E85FDDEFA53" box="[383,565,1415,1439]" italics="true" pageId="15" pageNumber="92">Fulakora monrosi</emphasis>
</taxonomicName>
(CASTYPE06893; images by April Nobile),
<emphasis id="B975EAD6FF84693D36FA2E85FBE4FA53" bold="true" box="[1021,1039,1415,1439]" pageId="15" pageNumber="92">C</emphasis>
and
<emphasis id="B975EAD6FF84693D31392E85FBBBFA53" bold="true" box="[1086,1104,1415,1439]" pageId="15" pageNumber="92">D</emphasis>
,
<taxonomicName id="4C014D47FF84693D315B2E8AFB26FA53" box="[1116,1229,1415,1439]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B975EAD6FF84693D315B2E8AFB26FA53" box="[1116,1229,1415,1439]" italics="true" pageId="15" pageNumber="92">F. chilensis</emphasis>
</taxonomicName>
(CASENT0915648; images by Harald Bruckner). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF84693D35902D11FA77F989" pageId="15" pageNumber="92">
<paragraph id="8BBE36C4FF84693D35902D11FA77F9EB" blockId="15.[151,1436,1554,1606]" box="[151,1436,1554,1576]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902D11FA77F9EB" box="[151,1436,1554,1576]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902D11FF42F9E4" bold="true" box="[151,169,1555,1576]" pageId="15" pageNumber="92">5.</emphasis>
Antenna with less than 12 antennomeres...................................................................
<emphasis id="B975EAD6FF84693D30972D10FA77F9EB" bold="true" box="[1424,1436,1554,1575]" pageId="15" pageNumber="92">6</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF84693D35902D32FA77F989" blockId="15.[151,1436,1554,1606]" box="[151,1436,1584,1606]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902D32FA77F989" box="[151,1436,1584,1606]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902D32FF74F989" bold="true" box="[151,159,1584,1605]" pageId="15" pageNumber="92">-</emphasis>
12-segmented antennae................................................................................
<emphasis id="B975EAD6FF84693D30972D32FA77F989" bold="true" box="[1424,1436,1584,1605]" pageId="15" pageNumber="92">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F52E61FF84693D35902D73FA77F912" pageId="15" pageNumber="92">
<paragraph id="8BBE36C4FF84693D35902D73FA77F968" blockId="15.[151,1436,1649,1759]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902D73FA77F968" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902D73FF42F94A" bold="true" box="[151,169,1649,1670]" pageId="15" pageNumber="92">6.</emphasis>
Antenna with 11 antennomeres. Body coloration brown to ferruginous. Genal tooth present and well-developed (Caribbean) (
<figureCitation id="133A2A41FF84693D35EE2D8CFED1F968" box="[233,314,1678,1700]" captionStart="FIGURE 14" captionStartId="16.[151,250,649,673]" captionTargetBox="[151,1436,181,625]" captionTargetId="figure-30@16.[151,1436,181,625]" captionTargetPageId="16" captionText="FIGURE 14. A, Fulakora bierigi (CASENT0915097; images by Z. Lieberman), B, F. aff. degenerata (ANTWEB1032441).All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896096" httpUri="https://zenodo.org/record/14896096/files/figure.png" pageId="15" pageNumber="92">Fig.14A</figureCitation>
)...........................................................................................
<emphasis id="B975EAD6FF84693D30972D8DFA77F968" bold="true" box="[1424,1436,1679,1700]" pageId="15" pageNumber="92">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF84693D35902DAEFA77F912" blockId="15.[151,1436,1649,1759]" pageId="15" pageNumber="92">
<keyLead id="30F095F1FF84693D35902DAEFA77F912" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D35902DAEFF74F90D" bold="true" box="[151,159,1708,1729]" pageId="15" pageNumber="92">-</emphasis>
Antenna with 69 antennomeres. Body coloration yellowish. Genal tooth absent or minute (
<figureCitation id="133A2A41FF84693D31532DAEFB48F90E" box="[1108,1187,1708,1730]" captionStart="FIGURE 14" captionStartId="16.[151,250,649,673]" captionTargetBox="[151,1436,181,625]" captionTargetId="figure-30@16.[151,1436,181,625]" captionTargetPageId="16" captionText="FIGURE 14. A, Fulakora bierigi (CASENT0915097; images by Z. Lieberman), B, F. aff. degenerata (ANTWEB1032441).All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896096" httpUri="https://zenodo.org/record/14896096/files/figure.png" pageId="15" pageNumber="92">Fig.14B</figureCitation>
)..............................................................................................
<taxonomicName id="4C014D47FF84693D31622DCBFAAEF913" authority="Borgmeier" authorityName="Borgmeier" box="[1125,1349,1737,1759]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF84693D31622DCBFAAEF913" bold="true" box="[1125,1349,1737,1759]" pageId="15" pageNumber="92">
<emphasis id="B975EAD6FF84693D31622DCBFB39F913" bold="true" box="[1125,1234,1737,1759]" italics="true" pageId="15" pageNumber="92">degenerata</emphasis>
Borgmeier
</emphasis>
</taxonomicName>
(Note 1)
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF9B69223590298BFE0AFD0A" ID-DOI="http://doi.org/10.5281/zenodo.14896096" ID-Zenodo-Dep="14896096" httpUri="https://zenodo.org/record/14896096/files/figure.png" pageId="16" pageNumber="93" startId="16.[151,250,649,673]" targetBox="[151,1436,181,625]" targetPageId="16" targetType="figure">
<paragraph id="8BBE36C4FF9B69223590298BFE0AFD0A" blockId="16.[151,1436,649,710]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B69223590298BFEDEFD6D" bold="true" box="[151,309,649,674]" pageId="16" pageNumber="93">FIGURE 14. A</emphasis>
,
<taxonomicName id="4C014D47FF9B692234462988FE02FD6E" box="[321,489,650,674]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF9B692234462988FE02FD6E" box="[321,489,650,674]" italics="true" pageId="16" pageNumber="93">Fulakora bierigi</emphasis>
</taxonomicName>
(CASENT0915097; images by Z. Lieberman),
<emphasis id="B975EAD6FF9B692236CF2988FC32FD6E" bold="true" box="[968,985,650,674]" pageId="16" pageNumber="93">B</emphasis>
,
<taxonomicName id="4C014D47FF9B692236E32988FB7DFD6E" box="[996,1174,650,674]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF9B692236E32988FC13FD6D" box="[996,1016,650,673]" italics="true" pageId="16" pageNumber="93">F.</emphasis>
aff.
<emphasis id="B975EAD6FF9B692231232988FB7DFD6E" box="[1060,1174,650,674]" italics="true" pageId="16" pageNumber="93">degenerata</emphasis>
</taxonomicName>
(ANTWEB1032441). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9B692235902817FA77FC4E" pageId="16" pageNumber="93">
<paragraph id="8BBE36C4FF9B692235902817FA77FC8B" blockId="16.[151,1436,789,898]" pageId="16" pageNumber="93">
<keyLead id="30F095F1FF9B692235902817FA77FC8B" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902817FF42FCE6" bold="true" box="[151,169,789,810]" pageId="16" pageNumber="93">7.</emphasis>
Foveolate piligerous punctures on head dense. Mandible shorter (ML about
<quantity id="4CF99B21FF9B692236BC2817FBEDFCE6" box="[955,1030,789,810]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="93" unit="mm" value="0.7">0.7 mm</quantity>
) (
<collectingCountry id="F3167654FF9B6922311E2817FBA5FCE7" box="[1049,1102,789,811]" name="Cuba" pageId="16" pageNumber="93">Cuba</collectingCountry>
) (
<figureCitation id="133A2A41FF9B692231652817FB58FCE7" box="[1122,1203,789,811]" captionStart="FIGURE 15" captionStartId="16.[151,250,1391,1415]" captionTargetBox="[151,1436,982,1367]" captionTargetId="figure-291@16.[151,1436,982,1367]" captionTargetPageId="16" captionText="FIGURE 15. A, Fulakora bierigi (CASENT0915097; images by Z. Lieberman), B, F. falcata (CASENT0172809; images by April Nobile). All images available at Antweb.org." figureDoi="http://doi.org/10.5281/zenodo.14896098" httpUri="https://zenodo.org/record/14896098/files/figure.png" pageId="16" pageNumber="93">Fig.15A</figureCitation>
)..................................................................................................
<taxonomicName id="4C014D47FF9B692231AE2830FAADFC84" authority="Santschi" authorityName="Santschi" box="[1193,1350,818,840]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF9B692231AE2830FAADFC84" bold="true" box="[1193,1350,818,840]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692231AE2830FB02FC84" bold="true" box="[1193,1257,818,840]" italics="true" pageId="16" pageNumber="93">bierigi</emphasis>
Santschi
</emphasis>
</taxonomicName>
(Note 2)
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF9B692235902852FA77FC4E" blockId="16.[151,1436,789,898]" pageId="16" pageNumber="93">
<keyLead id="30F095F1FF9B692235902852FA77FC4E" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902852FF74FCA9" bold="true" box="[151,159,848,869]" pageId="16" pageNumber="93">-</emphasis>
Foveolate piligerous punctures on head sparse. Mandible longer (ML about
<quantity id="4CF99B21FF9B692236BC2852FBEDFCA9" box="[955,1030,848,869]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="93" unit="mm" value="0.9">0.9 mm</quantity>
) (
<collectingCountry id="F3167654FF9B6922311F2852FB65FCA9" box="[1048,1166,847,869]" name="Puerto Rico" pageId="16" pageNumber="93">Puerto Rico</collectingCountry>
) (
<figureCitation id="133A2A41FF9B692231A6284DFB1AFCA9" box="[1185,1265,847,869]" captionStart="FIGURE 15" captionStartId="16.[151,250,1391,1415]" captionTargetBox="[151,1436,982,1367]" captionTargetId="figure-291@16.[151,1436,982,1367]" captionTargetPageId="16" captionText="FIGURE 15. A, Fulakora bierigi (CASENT0915097; images by Z. Lieberman), B, F. falcata (CASENT0172809; images by April Nobile). All images available at Antweb.org." figureDoi="http://doi.org/10.5281/zenodo.14896098" httpUri="https://zenodo.org/record/14896098/files/figure.png" pageId="16" pageNumber="93">Fig.15B</figureCitation>
).....................................................................................................
<taxonomicName id="4C014D47FF9B69223009286EFA77FC4E" authority="Lattke" authorityName="Lattke" box="[1294,1436,876,898]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="falcata">
<emphasis id="B975EAD6FF9B69223009286EFA77FC4E" bold="true" box="[1294,1436,876,898]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B69223009286EFAB9FC4E" bold="true" box="[1294,1362,876,898]" italics="true" pageId="16" pageNumber="93">falcata</emphasis>
Lattke
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF9B692235902E6DFD72FA67" ID-DOI="http://doi.org/10.5281/zenodo.14896098" ID-Zenodo-Dep="14896098" httpUri="https://zenodo.org/record/14896098/files/figure.png" pageId="16" pageNumber="93" startId="16.[151,250,1391,1415]" targetBox="[151,1436,982,1367]" targetPageId="16" targetType="figure">
<paragraph id="8BBE36C4FF9B692235902E6DFD72FA67" blockId="16.[151,1436,1391,1451]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902E6DFED2FA4B" bold="true" box="[151,313,1391,1415]" pageId="16" pageNumber="93">FIGURE 15. A</emphasis>
,
<taxonomicName id="4C014D47FF9B692234402E6DFE1AFA4B" box="[327,497,1391,1415]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF9B692234402E6DFE1AFA4B" box="[327,497,1391,1415]" italics="true" pageId="16" pageNumber="93">Fulakora bierigi</emphasis>
</taxonomicName>
(CASENT0915097; images by Z. Lieberman),
<emphasis id="B975EAD6FF9B692236DC2E6DFC07FA4B" bold="true" box="[987,1004,1391,1415]" pageId="16" pageNumber="93">B</emphasis>
,
<taxonomicName id="4C014D47FF9B692236FD2E72FBB1FA4B" box="[1018,1114,1391,1415]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="falcata">
<emphasis id="B975EAD6FF9B692236FD2E72FBB1FA4B" box="[1018,1114,1391,1415]" italics="true" pageId="16" pageNumber="93">F. falcata</emphasis>
</taxonomicName>
(CASENT0172809; images by April Nobile). All images available at Antweb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9B692235902EF8FA77F948" pageId="16" pageNumber="93">
<paragraph id="8BBE36C4FF9B692235902EF8FA77F986" blockId="16.[151,1437,1530,1670]" pageId="16" pageNumber="93">
<keyLead id="30F095F1FF9B692235902EF8FA77F986" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902EF8FF42F9C3" bold="true" box="[151,169,1530,1551]" pageId="16" pageNumber="93">8.</emphasis>
Subpetiolar process large, with a fenestra. Body light brown to yellowish. Head dorsum punctate. Genal tooth small but clearly discernible. Metanotal suture deep (
<collectingCountry id="F3167654FF9B6922373E2D1AFD86F9E1" box="[569,621,1560,1581]" name="United States of America" pageId="16" pageNumber="93">USA</collectingCountry>
,
<collectingCountry id="F3167654FF9B692237712D1AFD29F9E2" box="[630,706,1560,1582]" name="Mexico" pageId="16" pageNumber="93">Mexico</collectingCountry>
, Central America, and northern
<collectingCountry id="F3167654FF9B692236F12D1AFBBDF9E2" box="[1014,1110,1560,1582]" name="Colombia" pageId="16" pageNumber="93">Colombia</collectingCountry>
) (
<figureCitation id="133A2A41FF9B6922316D2D15FB42F9E1" box="[1130,1193,1559,1581]" captionStart="FIGURE 16" captionStartId="16.[151,250,1998,2022]" captionTargetBox="[151,1436,1694,1975]" captionTargetId="figure-492@16.[151,1436,1694,1975]" captionTargetPageId="16" captionText="FIGURE 16. A, B, and C, F. orizabana (CASENT0172807; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896100" httpUri="https://zenodo.org/record/14896100/files/figure.png" pageId="16" pageNumber="93">Fig.16</figureCitation>
).................................................................................................
<taxonomicName id="4C014D47FF9B6922319F2D36FAADF986" authority="Brown" authorityName="Brown" box="[1176,1350,1588,1610]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="orizabana">
<emphasis id="B975EAD6FF9B6922319F2D36FAADF986" bold="true" box="[1176,1350,1588,1610]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B6922319F2D36FB10F986" bold="true" box="[1176,1275,1588,1610]" italics="true" pageId="16" pageNumber="93">orizabana</emphasis>
Brown
</emphasis>
</taxonomicName>
(Note 4)
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF9B692235902D50FA77F948" blockId="16.[151,1437,1530,1670]" pageId="16" pageNumber="93">
<keyLead id="30F095F1FF9B692235902D50FA77F948" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902D50FF74F9AB" bold="true" box="[151,159,1618,1639]" pageId="16" pageNumber="93">-</emphasis>
Subpetiolar process reduced or absent and without fenestra. Body (excluding antennae, legs and mandibles) dark brown or black, rarely lighter in coloration. Head dorsum sculpturing varying. Genal tooth either small or large. Metanotal suture absent...
<emphasis id="B975EAD6FF9B692230972D6DFA77F948" bold="true" box="[1424,1436,1647,1668]" pageId="16" pageNumber="93">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF9B692235902CCCFA98F82B" ID-DOI="http://doi.org/10.5281/zenodo.14896100" ID-Zenodo-Dep="14896100" box="[151,1395,1998,2023]" httpUri="https://zenodo.org/record/14896100/files/figure.png" pageId="16" pageNumber="93" startId="16.[151,250,1998,2022]" targetBox="[151,1436,1694,1975]" targetPageId="16" targetType="figure">
<paragraph id="8BBE36C4FF9B692235902CCCFA98F82B" blockId="16.[151,1395,1998,2023]" box="[151,1395,1998,2023]" pageId="16" pageNumber="93">
<emphasis id="B975EAD6FF9B692235902CCCFEDCF82A" bold="true" box="[151,311,1998,2023]" pageId="16" pageNumber="93">FIGURE 16. A</emphasis>
,
<emphasis id="B975EAD6FF9B692234432CCDFEBEF82B" bold="true" box="[324,341,1999,2023]" pageId="16" pageNumber="93">B</emphasis>
, and
<emphasis id="B975EAD6FF9B6922348B2CCCFE75F82A" bold="true" box="[396,414,1998,2022]" pageId="16" pageNumber="93">C</emphasis>
,
<taxonomicName id="4C014D47FF9B692234AC2CCDFDC0F82B" box="[427,555,1999,2023]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="orizabana">
<emphasis id="B975EAD6FF9B692234AC2CCDFDC0F82B" box="[427,555,1999,2023]" italics="true" pageId="16" pageNumber="93">F. orizabana</emphasis>
</taxonomicName>
(CASENT0172807; images by April Nobile). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9A692335902B95FA76FF06" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A692335902B95FA77FF60" blockId="17.[151,1437,151,203]" box="[151,1436,151,173]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902B95FA77FF60" box="[151,1436,151,173]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902B95FF42FF60" bold="true" box="[151,169,151,172]" pageId="17" pageNumber="94">9.</emphasis>
Propodeum posterolateral corner with a pale patch. Small species (TL less than
<quantity id="4CF99B21FF9A692336D82B9AFBF3FF61" box="[991,1048,152,173]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="94" unit="mm" value="4.0">4 mm</quantity>
).............................
<emphasis id="B975EAD6FF9A692330832B95FA77FF60" bold="true" box="[1412,1436,151,172]" pageId="17" pageNumber="94">10</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF9A692335902BB7FA76FF06" blockId="17.[151,1437,151,203]" box="[151,1437,181,203]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902BB7FA76FF06" box="[151,1437,181,203]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902BB7FF74FF06" bold="true" box="[151,159,181,202]" pageId="17" pageNumber="94">-</emphasis>
Propodeum posterolateral corners without a pale patch. Usually larger species (TL more than
<quantity id="4CF99B21FF9A692331632BB7FB76FF06" box="[1124,1181,181,202]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="94" unit="mm" value="4.0">4 mm</quantity>
)..................
<emphasis id="B975EAD6FF9A692330812BB7FA76FF06" bold="true" box="[1414,1437,181,202]" pageId="17" pageNumber="94">11</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F52E61FF9A692335902BF4FA77FEE5" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A692335902BF4FA76FEC0" blockId="17.[151,1437,246,297]" box="[151,1437,246,268]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902BF4FA76FEC0" box="[151,1437,246,268]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902BF4FF5DFEC7" bold="true" box="[151,182,246,267]" pageId="17" pageNumber="94">10.</emphasis>
Lateral margins of head subparallel. Vertex smooth (
<figureCitation id="133A2A41FF9A692337DE2BF4FCC1FEC0" box="[729,810,246,268]" captionStart="FIGURE 17" captionStartId="17.[151,250,780,804]" captionTargetBox="[151,1436,322,757]" captionTargetId="figure-181@17.[151,1436,322,757]" captionTargetPageId="17" captionText="FIGURE 17. A, F. elongata (CASENT0915100; images by Z. Lieberman); B, F. lurilabes (CASENT0172810; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896102" httpUri="https://zenodo.org/record/14896102/files/figure.png" pageId="17" pageNumber="94">Fig.17A</figureCitation>
)....................................
<taxonomicName id="4C014D47FF9A692331ED2BF4FA76FEC0" authority="Santschi" authorityName="Santschi" box="[1258,1437,246,268]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF9A692331ED2BF4FA76FEC0" bold="true" box="[1258,1437,246,268]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692331ED2BF4FAD4FEC0" bold="true" box="[1258,1343,246,268]" italics="true" pageId="17" pageNumber="94">elongata</emphasis>
Santschi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF9A692335902A16FA77FEE5" blockId="17.[151,1437,246,297]" box="[151,1436,275,297]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902A16FA77FEE5" box="[151,1436,275,297]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902A16FF74FEE5" bold="true" box="[151,159,276,297]" pageId="17" pageNumber="94">-</emphasis>
Lateral margins of head mildly tapering posteriorly. Vertex dull or opaque (
<figureCitation id="133A2A41FF9A692336B32A11FBE8FEE5" box="[948,1027,275,297]" captionStart="FIGURE 17" captionStartId="17.[151,250,780,804]" captionTargetBox="[151,1436,322,757]" captionTargetId="figure-181@17.[151,1436,322,757]" captionTargetPageId="17" captionText="FIGURE 17. A, F. elongata (CASENT0915100; images by Z. Lieberman); B, F. lurilabes (CASENT0172810; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896102" httpUri="https://zenodo.org/record/14896102/files/figure.png" pageId="17" pageNumber="94">Fig.17B</figureCitation>
)....................
<taxonomicName id="4C014D47FF9A692331FB2A11FA77FEE5" authority="Lattke" authorityName="Lattke" box="[1276,1436,275,297]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="lurilabes">
<emphasis id="B975EAD6FF9A692331FB2A11FA77FEE5" bold="true" box="[1276,1436,275,297]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692331FB2A11FAB8FEE5" bold="true" box="[1276,1363,275,297]" italics="true" pageId="17" pageNumber="94">lurilabes</emphasis>
Lattke
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF7E664CFF9A69233590280EFD8BFC85" ID-DOI="http://doi.org/10.5281/zenodo.14896102" ID-Zenodo-Dep="14896102" httpUri="https://zenodo.org/record/14896102/files/figure.png" pageId="17" pageNumber="94" startId="17.[151,250,780,804]" targetBox="[151,1436,322,757]" targetPageId="17" targetType="figure">
<paragraph id="8BBE36C4FF9A69233590280EFD8BFC85" blockId="17.[151,1436,780,841]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A69233590280EFEDDFCE8" bold="true" box="[151,310,780,805]" pageId="17" pageNumber="94">FIGURE 17. A</emphasis>
,
<taxonomicName id="4C014D47FF9A69233445280FFE5FFCE9" box="[322,436,781,805]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF9A69233445280FFE5FFCE9" box="[322,436,781,805]" italics="true" pageId="17" pageNumber="94">F. elongata</emphasis>
</taxonomicName>
(CASENT0915100; images by Z. Lieberman);
<emphasis id="B975EAD6FF9A69233692280FFC4DFCE9" bold="true" box="[917,934,781,805]" pageId="17" pageNumber="94">B</emphasis>
,
<taxonomicName id="4C014D47FF9A692336B5280FFBCEFCE9" box="[946,1061,781,805]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="lurilabes">
<emphasis id="B975EAD6FF9A692336B5280FFBCEFCE9" box="[946,1061,781,805]" italics="true" pageId="17" pageNumber="94">F. lurilabes</emphasis>
</taxonomicName>
(CASENT0172810; images by April Nobile). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9A692335902876FA77FC0F" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A692335902876FA77FC6B" blockId="17.[151,1436,884,965]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902876FA77FC6B" pageId="17" pageNumber="94">
11. Head dorsum strongly sculptured up to the vertexal level. Medial clypeal tubercles separated (
<figureCitation id="133A2A41FF9A6923316E2876FB51FC46" box="[1129,1210,884,906]" captionStart="FIGURE 18" captionStartId="17.[151,250,1302,1326]" captionTargetBox="[151,1436,989,1279]" captionTargetId="figure-372@17.[151,1436,989,1279]" captionTargetPageId="17" captionText="FIGURE 18. A, Fulakora cleae (CASENT0102202; images by April Nobile); B, F. agostii (CASENT0101451; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896105" httpUri="https://zenodo.org/record/14896105/files/figure.png" pageId="17" pageNumber="94">Fig.18A</figureCitation>
)...................................................................................................
<taxonomicName id="4C014D47FF9A692331B12893FA77FC6B" authority="Lacau &amp; Delabie" authorityName="Lacau &amp; Delabie" box="[1206,1436,913,935]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF9A692331B12893FA77FC6B" bold="true" box="[1206,1436,913,935]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692331B12893FB02FC6B" bold="true" box="[1206,1257,913,935]" italics="true" pageId="17" pageNumber="94">cleae</emphasis>
Lacau &amp; Delabie
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<keyLead id="30F095F1FF9A6923359028ADFA77FA7B" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A6923359028ADFA77FC0F" blockId="17.[151,1436,884,965]" box="[151,1436,942,965]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A6923359028ADFF74FC08" bold="true" box="[151,159,943,964]" pageId="17" pageNumber="94">-</emphasis>
Head dorsum smooth at the vertexal level. Medial clypeal tubercles fused (
<figureCitation id="133A2A41FF9A692336B628ADFBEBFC09" box="[945,1024,943,965]" captionStart="FIGURE 18" captionStartId="17.[151,250,1302,1326]" captionTargetBox="[151,1436,989,1279]" captionTargetId="figure-372@17.[151,1436,989,1279]" captionTargetPageId="17" captionText="FIGURE 18. A, Fulakora cleae (CASENT0102202; images by April Nobile); B, F. agostii (CASENT0101451; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896105" httpUri="https://zenodo.org/record/14896105/files/figure.png" pageId="17" pageNumber="94">Fig.18B</figureCitation>
)...............................
<emphasis id="B975EAD6FF9A6923308328ACFA77FC0F" bold="true" box="[1412,1436,942,963]" pageId="17" pageNumber="94">12</emphasis>
</paragraph>
<caption id="DF7E664CFF9A692335902E14FD77FA9E" ID-DOI="http://doi.org/10.5281/zenodo.14896105" ID-Zenodo-Dep="14896105" httpUri="https://zenodo.org/record/14896105/files/figure.png" pageId="17" pageNumber="94" startId="17.[151,250,1302,1326]" targetBox="[151,1436,989,1279]" targetPageId="17" targetType="figure">
<paragraph id="8BBE36C4FF9A692335902E14FD77FA9E" blockId="17.[151,1436,1302,1362]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902E14FED7FAE2" bold="true" box="[151,316,1302,1326]" pageId="17" pageNumber="94">FIGURE 18. A</emphasis>
,
<taxonomicName id="4C014D47FF9A6923344C2E14FE03FAE2" box="[331,488,1302,1326]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF9A6923344C2E14FE03FAE2" box="[331,488,1302,1326]" italics="true" pageId="17" pageNumber="94">Fulakora cleae</emphasis>
</taxonomicName>
(CASENT0102202; images by April Nobile);
<emphasis id="B975EAD6FF9A692336D52E15FC08FAE3" bold="true" box="[978,995,1303,1327]" pageId="17" pageNumber="94">B</emphasis>
,
<taxonomicName id="4C014D47FF9A692336F52E15FBBDFAE2" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1010,1110,1303,1326]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF9A692336F52E15FBEAFAE2" box="[1010,1025,1303,1326]" italics="true" pageId="17" pageNumber="94">F</emphasis>
.
<emphasis id="B975EAD6FF9A692331162E15FBBDFAE2" box="[1041,1110,1303,1326]" italics="true" pageId="17" pageNumber="94">agostii</emphasis>
</taxonomicName>
(CASENT0101451; images by April Nobile). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9A692335902EA3FA77FA18" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A692335902EA3FA77FA7B" blockId="17.[151,1436,1441,1493]" box="[151,1436,1441,1464]" pageId="17" pageNumber="94">
12. Lateral margins of head more strongly tapering posteriorly. Meso-metapleural suture present (
<figureCitation id="133A2A41FF9A692331602EA0FB36FA7B" box="[1127,1245,1442,1464]" captionStart="FIGURE 19" captionStartId="17.[151,250,1838,1862]" captionTargetBox="[151,1436,1547,1814]" captionTargetId="figure-452@17.[151,1436,1547,1814]" captionTargetPageId="17" captionText="FIGURE 19. A, and B, Fulakora armigera (CASENT0915647; images by Harald Bruckner); C, and D, F. agostii (CASENT0101451; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896107" httpUri="https://zenodo.org/record/14896107/files/figure.png" pageId="17" pageNumber="94">Figs.19A, B</figureCitation>
)..
<taxonomicName id="4C014D47FF9A692330052EA0FA77FA7B" authority="Mayr" authorityName="Mayr" box="[1282,1436,1442,1463]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="B975EAD6FF9A692330052EA0FA77FA7B" bold="true" box="[1282,1436,1442,1463]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692330052EA0FAB6FA7B" bold="true" box="[1282,1373,1442,1463]" italics="true" pageId="17" pageNumber="94">armigera</emphasis>
Mayr
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BBE36C4FF9A692335902EBDFA77FA18" blockId="17.[151,1436,1441,1493]" box="[151,1436,1471,1493]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902EBDFA77FA18" box="[151,1436,1471,1493]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902EBDFF74FA18" bold="true" box="[151,159,1471,1492]" pageId="17" pageNumber="94">-</emphasis>
Lateral margins of head mildly tapering posteriorly. Meso-metapleural suture absent (
<figureCitation id="133A2A41FF9A6923311F2EBDFB65FA18" box="[1048,1166,1471,1493]" captionStart="FIGURE 19" captionStartId="17.[151,250,1838,1862]" captionTargetBox="[151,1436,1547,1814]" captionTargetId="figure-452@17.[151,1436,1547,1814]" captionTargetPageId="17" captionText="FIGURE 19. A, and B, Fulakora armigera (CASENT0915647; images by Harald Bruckner); C, and D, F. agostii (CASENT0101451; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896107" httpUri="https://zenodo.org/record/14896107/files/figure.png" pageId="17" pageNumber="94">Figs.19C, D</figureCitation>
)...................
<emphasis id="B975EAD6FF9A692330832EBDFA77FA18" bold="true" box="[1412,1436,1471,1492]" pageId="17" pageNumber="94">13</emphasis>
</keyLead>
</paragraph>
</keyStep>
</keyLead>
</keyStep>
<caption id="DF7E664CFF9A692335902C2CFC31F8A6" ID-DOI="http://doi.org/10.5281/zenodo.14896107" ID-Zenodo-Dep="14896107" httpUri="https://zenodo.org/record/14896107/files/figure.png" pageId="17" pageNumber="94" startId="17.[151,250,1838,1862]" targetBox="[151,1436,1547,1814]" targetPageId="17" targetType="figure">
<paragraph id="8BBE36C4FF9A692335902C2CFC31F8A6" blockId="17.[151,1436,1838,1898]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902C2CFEACF88A" bold="true" box="[151,327,1838,1862]" pageId="17" pageNumber="94">FIGURE 19. A</emphasis>
, and
<emphasis id="B975EAD6FF9A692334882C2CFE4BF88A" bold="true" box="[399,416,1838,1862]" pageId="17" pageNumber="94">B</emphasis>
,
<taxonomicName id="4C014D47FF9A692334B22C2CFD6BF88A" box="[437,640,1838,1862]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="B975EAD6FF9A692334B22C2CFD6BF88A" box="[437,640,1838,1862]" italics="true" pageId="17" pageNumber="94">Fulakora armigera</emphasis>
</taxonomicName>
(CASENT0915647; images by Harald Bruckner);
<emphasis id="B975EAD6FF9A692331B22C2CFB2CF88A" bold="true" box="[1205,1223,1838,1862]" pageId="17" pageNumber="94">C</emphasis>
, and
<emphasis id="B975EAD6FF9A692330082C2CFACAF88A" bold="true" box="[1295,1313,1838,1862]" pageId="17" pageNumber="94">D</emphasis>
,
<taxonomicName id="4C014D47FF9A692330312C2DFA77F88A" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1334,1436,1839,1862]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF9A692330312C2DFA77F88A" box="[1334,1436,1839,1862]" italics="true" pageId="17" pageNumber="94">F. agostii</emphasis>
</taxonomicName>
(CASENT0101451; images by April Nobile). All images available at AntWeb.org.
</paragraph>
</caption>
<keyStep id="30F52E61FF9A692335902CBBFA77F820" pageId="17" pageNumber="94">
<paragraph id="8BBE36C4FF9A692335902CBBFA77F803" blockId="17.[151,1436,1977,2028]" box="[151,1436,1977,1999]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902CBBFA77F803" box="[151,1436,1977,1999]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902CBBFF5EF802" bold="true" box="[151,181,1977,1998]" pageId="17" pageNumber="94">13.</emphasis>
Genal tooth longer than twice clypeal tubercle plus its chaeta (Fig.20A).......................
<taxonomicName id="4C014D47FF9A692331AF2CBBFA77F803" authority="Lacau &amp; Delabie" authorityName="Lacau &amp; Delabie" box="[1192,1436,1977,1999]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF9A692331AF2CBBFA77F803" bold="true" box="[1192,1436,1977,1999]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692331AF2CBBFB02F802" bold="true" box="[1192,1257,1977,1998]" italics="true" pageId="17" pageNumber="94">agostii</emphasis>
Lacau &amp; Delabie
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36C4FF9A692335902CD5FA77F820" blockId="17.[151,1436,1977,2028]" box="[151,1436,2006,2028]" pageId="17" pageNumber="94">
<keyLead id="30F095F1FF9A692335902CD5FA77F820" box="[151,1436,2006,2028]" pageId="17" pageNumber="94">
<emphasis id="B975EAD6FF9A692335902CD5FF74F820" bold="true" box="[151,159,2007,2028]" pageId="17" pageNumber="94">-</emphasis>
Genal tooth about as long as the clypeal tubercle plus its chaeta (Fig.20B)............................
<taxonomicName id="4C014D47FF9A692330042CD5FAA1F820" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[1283,1354,2007,2028]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="94" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF9A692330042CD5FAA1F820" bold="true" box="[1283,1354,2007,2028]" italics="true" pageId="17" pageNumber="94">estevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657ADFF9A692330572CD5FA77F820" box="[1360,1436,2007,2028]" pageId="17" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<caption id="DF7E664CFF9969203590297EFE78FD74" ID-DOI="http://doi.org/10.5281/zenodo.14896109" ID-Zenodo-Dep="14896109" httpUri="https://zenodo.org/record/14896109/files/figure.png" pageId="18" pageNumber="95" startId="18.[151,250,636,660]" targetBox="[151,1436,197,612]" targetPageId="18" targetType="figure">
<paragraph id="8BBE36C4FF9969203590297EFE78FD74" blockId="18.[151,1436,636,696]" pageId="18" pageNumber="95">
<emphasis id="B975EAD6FF9969203590297EFEDDFD58" bold="true" box="[151,310,636,660]" pageId="18" pageNumber="95">FIGURE 20. A</emphasis>
,
<taxonomicName id="4C014D47FF9969203444297EFE06FD58" box="[323,493,636,660]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF9969203444297EFE06FD58" box="[323,493,636,660]" italics="true" pageId="18" pageNumber="95">Fulakora agostii</emphasis>
</taxonomicName>
(CASENT0101451; images by April Nobile);
<emphasis id="B975EAD6FF99692036C1297EFC3CFD58" bold="true" box="[966,983,636,660]" pageId="18" pageNumber="95">B</emphasis>
,
<taxonomicName id="4C014D47FF99692036E4297FFBACFD58" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[995,1095,637,660]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF99692036E4297FFBACFD58" box="[995,1095,637,660]" italics="true" pageId="18" pageNumber="95">F. estevae</emphasis>
</taxonomicName>
(ANTWEB1047071). All images available at AntWeb.org.
</paragraph>
</caption>
<subSubSection id="C31B654FFF99692035C02809FD06FB7C" pageId="18" pageNumber="95" type="discussion">
<paragraph id="8BBE36C4FF99692035C02809FAFBFC14" blockId="18.[151,1437,778,1201]" pageId="18" pageNumber="95">
<collectorName id="26F45312FF99692035C02809FF16FCE9" box="[199,253,779,805]" pageId="18" pageNumber="95">Note</collectorName>
1.
<collectorName id="26F45312FF996920341A2808FEA2FCE8" box="[285,329,778,804]" pageId="18" pageNumber="95">The</collectorName>
type locality of
<taxonomicName id="4C014D47FF99692034FD2809FD79FCE8" box="[506,658,778,804]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF99692034FD2809FD79FCE8" box="[506,658,778,804]" italics="true" pageId="18" pageNumber="95">F. degenerata</emphasis>
</taxonomicName>
is
<collectingRegion id="49C5F826FF99692037B72809FCBDFCE8" box="[688,854,778,805]" country="Brazil" name="Santa Catarina" pageId="18" pageNumber="95">Santa Catarina</collectingRegion>
,
<collectingCountry id="F3167654FF99692036582808FC4DFCE8" box="[863,934,778,804]" name="Brazil" pageId="18" pageNumber="95">Brazil</collectingCountry>
, where the
<typeStatus id="54BA8866FF99692031252808FB6FFCE8" box="[1058,1156,778,804]" pageId="18" pageNumber="95" type="holotype">holotype</typeStatus>
has seven antennomeres.
<collectorName id="26F45312FF9969203590282DFF30FC85" box="[151,219,815,841]" pageId="18" pageNumber="95">Lacau</collectorName>
&amp;
<collectorName id="26F45312FF9969203402282CFEB7FC84" box="[261,348,814,840]" pageId="18" pageNumber="95">Delabie</collectorName>
mentioned a variation from six to seven antennomeres in their key for the species.
<collectorName id="26F45312FF996920302F282DFA73FC85" box="[1320,1432,815,841]" pageId="18" pageNumber="95">Moreover</collectorName>
, they mentioned an
<collectorName id="26F45312FF99692034772850FE1AFCA0" box="[368,497,850,876]" pageId="18" pageNumber="95">Amazonian</collectorName>
specimen with 11 antennomeres similar to
<taxonomicName id="4C014D47FF99692036E52851FB96FCA0" box="[994,1149,850,876]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF99692036E52851FB96FCA0" box="[994,1149,850,876]" italics="true" pageId="18" pageNumber="95">F. degenerata</emphasis>
</taxonomicName>
, which should be further examined.
<location id="8EDE601FFF99692034142874FEBFFC5C" LSID="urn:lsid:plazi:treatment:03A887D2FF85692035902A48FD06FB7C:8EDE601FFF99692034142874FEBFFC5C" box="[275,340,886,912]" country="Brazil" name="There" pageId="18" pageNumber="95" stateProvince="Minas Gerais">There</location>
are specimens from the states of
<collectingRegion id="49C5F826FF99692037D02874FC84FC5C" box="[727,879,886,912]" country="Brazil" name="Minas Gerais" pageId="18" pageNumber="95">Minas Gerais</collectingRegion>
and
<collectingRegion id="49C5F826FF99692036AE2874FBA0FC5D" box="[937,1099,886,913]" country="Brazil" name="Espirito Santo" pageId="18" pageNumber="95">Espírito Santo</collectingRegion>
in
<collectingCountry id="F3167654FF99692031752874FB5DFC5C" box="[1138,1206,886,912]" name="Brazil" pageId="18" pageNumber="95">Brazil</collectingCountry>
and from
<collectingCountry id="F3167654FF99692030282874FA77FC5C" box="[1327,1436,886,912]" name="Honduras" pageId="18" pageNumber="95">Honduras</collectingCountry>
<materialsCitation id="3B693C99FF99692035902898FAE7FC14" country="Brazil" location="Central America" municipality="southern South America" pageId="18" pageNumber="95" specimenCount="1" typeStatus="holotype">
with nine segments, making evident that
<taxonomicName id="4C014D47FF99692037742899FCE4FC78" box="[627,783,922,948]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF99692037742899FCE4FC78" box="[627,783,922,948]" italics="true" pageId="18" pageNumber="95">F. degenerata</emphasis>
</taxonomicName>
is a complex of various cryptic species distributed from
<location id="8EDE601FFF996920359028BCFEA4FC14" LSID="urn:lsid:plazi:treatment:03A887D2FF85692035902A48FD06FB7C:8EDE601FFF996920359028BCFEA4FC14" box="[151,335,958,984]" country="Brazil" municipality="southern South America" name="Central America" pageId="18" pageNumber="95">Central America</location>
to
<collectingMunicipality id="6BDAACBEFF996920347428BCFD6EFC14" box="[371,645,958,984]" pageId="18" pageNumber="95">southern South America</collectingMunicipality>
;
<taxonomicName id="4C014D47FF996920379628BDFCC1FC14" box="[657,810,958,984]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="degenerata">
<emphasis id="B975EAD6FF996920379628BDFCC1FC14" box="[657,810,958,984]" italics="true" pageId="18" pageNumber="95">F. degenerata</emphasis>
</taxonomicName>
itself is likely restricted to southern
<collectingCountry id="F3167654FF99692031C328BCFAE7FC14" box="[1220,1292,958,984]" name="Brazil" pageId="18" pageNumber="95">Brazil</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE36C4FF99692035C028E1FD06FB7C" blockId="18.[151,1437,778,1201]" pageId="18" pageNumber="95">
Note 2. The original description of
<taxonomicName id="4C014D47FF996920375928E1FC62FC31" authority="(Santschi, 1930)" baseAuthorityName="Santschi" baseAuthorityYear="1930" box="[606,905,994,1021]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF996920375928E1FD2CFC30" box="[606,711,994,1020]" italics="true" pageId="18" pageNumber="95">F. bierigi</emphasis>
(
<bibRefCitation id="EF904B35FF99692037D028E0FC6BFC31" author="Santschi" box="[727,896,994,1021]" firstAuthor="Santschi" pageId="18" pageNumber="95" pagination="75 - 83" refId="ref13252" refString="Santschi, F. (1930) Quelques fourmis de Cuba et du Bresil. Bulletin. Societe Entomologique d'Egypte, 14, 75-83." type="journal article" year="1930">Santschi, 1930</bibRefCitation>
)
</taxonomicName>
did not mention the number of antennomeres. The species is treated as having 12-merous antennae by
<bibRefCitation id="EF904B35FF99692036122F04FC44FBED" author="Lattke" box="[789,943,1030,1057]" firstAuthor="Lattke" pageId="18" pageNumber="95" pagination="1 - 7" refId="ref12706" refString="Lattke, J. E. (1991) Studies of Neotropical Amblyopone Erichson (Hymenoptera: Formicidae). Contributions in Science, Los Angeles, 428, 1-7. https://doi.org/10.5962/p.208156" type="journal article" year="1991">Lattke (1991)</bibRefCitation>
and
<bibRefCitation id="EF904B35FF99692036EF2F05FB14FBED" author="Lacau &amp; Delabie" box="[1000,1279,1030,1057]" firstAuthor="Lacau" pageId="18" pageNumber="95" pagination="33 - 41" refId="ref12585" refString="Lacau, S. &amp; Delabie, J. H. C. (2002) Description de trois nouvelles especes d' Amblyopone avec quelques notes biogeographiques sur le genre au Bresil (Formicidae, Ponerinae). B ulletin de la Societe Entomologique de France, 107, 33-41. https://doi.org/10.3406/bsef.2002.16813" type="journal article" year="2002">Lacau &amp; Delabie (2002)</bibRefCitation>
; however, the
<typeStatus id="54BA8866FF99692035902F28FF12FB88" box="[151,249,1066,1092]" pageId="18" pageNumber="95" type="holotype">holotype</typeStatus>
images show 11 antennomeres (CASENT0915097). This condition makes
<taxonomicName id="4C014D47FF99692031502F29FB2BFB88" baseAuthorityName="Santschi" baseAuthorityYear="1930" box="[1111,1216,1066,1092]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF99692031502F29FB2BFB88" box="[1111,1216,1066,1092]" italics="true" pageId="18" pageNumber="95">F. bierigi</emphasis>
</taxonomicName>
close to
<taxonomicName id="4C014D47FF996920302D2F29FA7EFB88" box="[1322,1429,1066,1092]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="falcata">
<emphasis id="B975EAD6FF996920302D2F29FA7EFB88" box="[1322,1429,1066,1092]" italics="true" pageId="18" pageNumber="95">F. falcata</emphasis>
</taxonomicName>
, with only the characters cited in this key separating them.
<specimenCount id="9D07FD4DFF99692036362F4DFC23FBA5" box="[817,968,1103,1129]" count="1" pageId="18" pageNumber="95" type="generic" typeStatus="paratype">One paratype</specimenCount>
of
<taxonomicName id="4C014D47FF99692036F62F4DFBA5FBA4" box="[1009,1102,1102,1128]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="falcat">
<emphasis id="B975EAD6FF99692036F62F4DFBA5FBA4" box="[1009,1102,1102,1128]" italics="true" pageId="18" pageNumber="95">F. falcat</emphasis>
</taxonomicName>
a (CASENT0102200) has the mandibles considerably smaller than the
<typeStatus id="54BA8866FF99692037752F70FD3CFB40" box="[626,727,1138,1164]" pageId="18" pageNumber="95" type="holotype">holotype</typeStatus>
, and, in fact, smaller than
<taxonomicName id="4C014D47FF99692031162F71FB90FB40" baseAuthorityName="Santschi" baseAuthorityYear="1930" box="[1041,1147,1138,1164]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B975EAD6FF99692031162F71FB90FB40" box="[1041,1147,1138,1164]" italics="true" pageId="18" pageNumber="95">F. bierigi</emphasis>
</taxonomicName>
. We considered only the
<typeStatus id="54BA8866FF99692035902F94FEEFFB7C" box="[151,260,1174,1200]" pageId="18" pageNumber="95" type="holotype">holotypes</typeStatus>
of both species to separate them in the key.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,633 @@
<document id="7AD89FABE65699D231F420EB9FDEE6F1" ID-DOI="10.11646/zootaxa.5588.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896067" ID-ZooBank="E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1740002781111" checkinUser="plazi" docAuthor="Jacintho, Gabriela De Figueiredo, Chaul, Júlio Cezar Mário, Teixeira, Gisele Amaro, Barros, Luísa Antônia Campos, Lopes, Denilce Meneses &amp; Serrão, José Eduardo" docDate="2025" docId="03A887D2FF8F6938359028D0FDEFFE1D" docLanguage="en" docName="zootaxa.5588.1.3.pdf" docOrigin="Zootaxa 5588 (1)" docSource="https://doi.org/10.11646/zootaxa.5588.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fulakora estevae Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="87" masterDocId="FF91FFAAFF8B693235072B02FFEBFFCC" masterDocTitle="A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species" masterLastPageNumber="104" masterPageNumber="77" pageNumber="81" updateTime="1740509805003" updateUser="guilherme" zenodo-license-document="CLOSED">
<mods:mods id="EB4FCAB9D3F8A94D08CC833B4B3B7C5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AFE20F007DBFFD0BAE6E0ABFEAC25253">
<mods:title id="EE50C592FDA483AC665F38B91A7E15A6">A morphological and cytogenetic study of two Fulakora Mann, 1919 (Hymenoptera: Formicidae), with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="53CDB3466EC834423603215B700D8C1C" type="personal">
<mods:role id="6FCA325E244BC40F833163E11055721E">
<mods:roleTerm id="940D4D8203046BA7752B2172F1A195C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2DB3D0E01208735A5E55D89B8CC29FA5">Jacintho, Gabriela De Figueiredo</mods:namePart>
<mods:affiliation id="A11C12B6A1B4D3F5E9E54D0A15ADE775">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="1BCE1CE453105B10F8C123DB854D732A" type="personal">
<mods:role id="76D70B43A331D8B98A9C18DC7089FB84">
<mods:roleTerm id="F113C2D9E9417C754767D129C88E48F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BF9C84F4AC293C5F463D58649B2CC4B">Chaul, Júlio Cezar Mário</mods:namePart>
<mods:affiliation id="C0E8A648271B309E7F4B4D4EEF96AE73">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="58130683E0A3DA8E517AA8E93E5AB399" type="personal">
<mods:role id="65745ADDB6A851724C01891E9201353C">
<mods:roleTerm id="537FB5A5991F95062503D6A7EE5327D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E6B3AD77C09D8A5DDDF2BEDFAEF9898">Teixeira, Gisele Amaro</mods:namePart>
<mods:affiliation id="994F3BE260887DBC24E086A43BC44E1E">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="FB025D6D8D7FF515EB10897F8A47A6FE" type="personal">
<mods:role id="1824E34D53451C18CD3BB87CB9C4DAA8">
<mods:roleTerm id="EA94D254693FCE0A36831F1DEBB028C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2B4B65B937C848635FE1224611341C5">Barros, Luísa Antônia Campos</mods:namePart>
<mods:affiliation id="D58AE5C0603DC9AABF9625D57B073BF9">Universidade Federal do Amapá, Campus Binacional, 68980 - 000, Oiapoque, Amapá, Brazil</mods:affiliation>
</mods:name>
<mods:name id="A30B8FC5A7CB6EB64D3E2D505D37EEB0" type="personal">
<mods:role id="50FD1DADF1860ED323A14C47FA12EC4C">
<mods:roleTerm id="FDE67F7C399FC77A556516E6950ADCF9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A27D3BAE2554D11C19BC3C2C4E0ADC8">Lopes, Denilce Meneses</mods:namePart>
<mods:affiliation id="578361F4B1F62C5C59627A4FBC35959B">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name id="A17F37E8AF92E5CBFF2DFDB8A55C42FB" type="personal">
<mods:role id="BEC83B6D41E73C717F21C03DE5C4563F">
<mods:roleTerm id="B6F8F293A0EDFE46ED1CB0C0347830F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63ABFF03FEC0B6CA0FD50E78731C3A6E">Serrão, José Eduardo</mods:namePart>
<mods:affiliation id="F8317560ABF3677BCBC139AF7E7FB2C0">Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9E922770C35D74BBF8FF242A710D0D0B">text</mods:typeOfResource>
<mods:relatedItem id="9A60B53BD9858EC266A1D941356069DA" type="host">
<mods:titleInfo id="B1B9170DB08216D6447202CF6D793302">
<mods:title id="66A5CEB85BA041AEC31D84F3C94FF20F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9C3EA861A4972597FD3775E5CB4050A0">
<mods:date id="059B43635B3B42D13BA75294F8FC0650">2025</mods:date>
<mods:detail id="97F031B51ADBF677FF6768A680053C12" type="pubDate">
<mods:number id="F68CF1CE5B4806D8AE7BA0D10212AA1C">2025-02-13</mods:number>
</mods:detail>
<mods:detail id="95485784C4665694D1E9A7E2ED6CEDA2" type="volume">
<mods:number id="820D39A1115FEFA71CC6C6E3A31715B8">5588</mods:number>
</mods:detail>
<mods:detail id="4396D25193734A14D1946F15A9D1301D" type="issue">
<mods:number id="C711B1C620F0E85F4BF882CBD12BE201">1</mods:number>
</mods:detail>
<mods:extent id="0F0135ADCB51557DD03B1DC63CAA3341" unit="page">
<mods:start id="2A1DA3B90257E43FE629640F147728E9">77</mods:start>
<mods:end id="BCED615C1D67D8E9A3EAF8E770B5F30A">104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4995DAB5C49BC5D2182FF313C2057672">
<mods:url id="BDFECEA0B6D488C3748F40978E1A8916">https://doi.org/10.11646/zootaxa.5588.1.3</mods:url>
</mods:location>
<mods:classification id="EBC2D00917319B092807EA11986D8036">journal article</mods:classification>
<mods:identifier id="0097A528A439858BCD78ECA4AFFBD1A2" type="DOI">10.11646/zootaxa.5588.1.3</mods:identifier>
<mods:identifier id="194B579EFD3EA20EB68A23894D522F4E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="497F6BC71A5569ACF52F5DCF04E99297" type="Zenodo-Dep">14896067</mods:identifier>
<mods:identifier id="6627491ED3E261DA3519EE92451741B7" type="ZooBank">E1A536AC-FA08-4ECD-B0E3-6B8AB2BC770D</mods:identifier>
</mods:mods>
<treatment id="03A887D2FF8F6938359028D0FDEFFE1D" LSID="urn:lsid:plazi:treatment:03A887D2FF8F6938359028D0FDEFFE1D" httpUri="http://treatment.plazi.org/id/03A887D2FF8F6938359028D0FDEFFE1D" lastPageId="10" lastPageNumber="87" pageId="4" pageNumber="81">
<subSubSection id="C31B654FFF8F6936359028D0FD17FC21" box="[151,764,978,1005]" pageId="4" pageNumber="81" type="nomenclature">
<paragraph id="8BBE36C4FF8F6936359028D0FD17FC21" blockId="4.[151,764,978,1041]" box="[151,764,978,1005]" pageId="4" pageNumber="81">
<heading id="D0F681A8FF8F6936359028D0FD17FC21" bold="true" box="[151,764,978,1005]" fontSize="11" level="1" pageId="4" pageNumber="81" reason="1">
<emphasis id="B975EAD6FF8F6936359028D0FD17FC21" bold="true" box="[151,764,978,1005]" pageId="4" pageNumber="81">
3.1.1.
<taxonomicName id="4C014D47FF8F693635E628D0FDCFFC20" authority="Jacintho" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[225,548,978,1005]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF8F693635E628D0FE5BFC21" bold="true" box="[225,432,978,1005]" italics="true" pageId="4" pageNumber="81">Fulakora estevae</emphasis>
Jacintho
</taxonomicName>
&amp;
<collectingCountry id="F3167654FF8F6936374D28D0FD7CFC20" box="[586,663,978,1004]" name="India" pageId="4" pageNumber="81">Chaul</collectingCountry>
<taxonomicNameLabel id="A24657ADFF8F6936379928D1FD17FC21" box="[670,764,979,1005]" pageId="4" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF8F6936359028F5FDB7FBDD" box="[151,604,1014,1041]" pageId="4" pageNumber="81" type="description">
<paragraph id="8BBE36C4FF8F6936359028F5FDB7FBDD" blockId="4.[151,764,978,1041]" box="[151,604,1014,1041]" pageId="4" pageNumber="81">
(
<figureCitation id="133A2A41FF8F6936359828F5FF1BFBDD" box="[159,240,1015,1041]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Figs. 2</figureCitation>
;
<figureCitation id="133A2A41FF8F693635FB28F5FEE7FBDD" box="[252,268,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="4" pageNumber="81">3</figureCitation>
;
<figureCitation id="133A2A41FF8F6936341F28F5FEC2FBDD" box="[280,297,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,603,627]" captionTargetBox="[151,1436,181,579]" captionTargetId="figure-19@7.[151,1436,181,579]" captionTargetPageId="7" captionText="FIGURE 4. Fulakora estevae sp. nov., worker (ANTWEB1047069) dissected mouthparts. A, labrum, and B, labiomaxillary complex. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896075" httpUri="https://zenodo.org/record/14896075/files/figure.png" pageId="4" pageNumber="81">4</figureCitation>
;
<figureCitation id="133A2A41FF8F6936343228F4FEAEFBDD" box="[309,325,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="4" pageNumber="81">5</figureCitation>
;
<figureCitation id="133A2A41FF8F6936345528F5FE89FBDD" box="[338,354,1015,1041]" captionStart="FIGURE 6" captionStartId="9.[151,250,1530,1554]" captionTargetBox="[151,1435,741,1506]" captionTargetId="figure-274@9.[151,1435,741,1506]" captionTargetPageId="9" captionText="FIGURE 6. Fulakora estevae sp. nov., larva. A, head in profile; B. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="4" pageNumber="81">6</figureCitation>
;
<figureCitation id="133A2A41FF8F6936346828F5FE37FBDD" box="[367,476,1015,1041]" captionStart="FIGURE 9" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[151,1436,959,1844]" captionTargetId="figure-334@13.[151,1436,959,1844]" captionTargetPageId="13" captionText="FIGURE 9. Morphological characters studied and their states. A, E, and L, Fulakora heraldoi (INPA-HYM-034964; images by Itanna Oliveira Fernandes); B and I, F. cleae (CASENT0102202; images by April Nobile) (zoom in B of F. elongata, ANTWEB1047734); C, G, and M, F.estevae(ANTWEB1047071);D, F.elongata (CASENT0915101; images by Z. Lieberman); F and K, F. elongata (ANTWEB1047734); H, F. armigera (CASENT0915647; images by Harald Bruckner); J, F. lurilabes (CASENT0172810; images by April Nobile). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896086" httpUri="https://zenodo.org/record/14896086/files/figure.png" pageId="4" pageNumber="81">9C, G, M</figureCitation>
;
<figureCitation id="133A2A41FF8F693634E028F5FDF2FBDD" box="[487,537,1015,1041]" captionStart="FIGURE 20" captionStartId="18.[151,250,636,660]" captionTargetBox="[151,1436,197,612]" captionTargetId="figure-30@18.[151,1436,197,612]" captionTargetPageId="18" captionText="FIGURE 20. A, Fulakora agostii (CASENT0101451; images by April Nobile); B, F. estevae (ANTWEB1047071). All images available at AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.14896109" httpUri="https://zenodo.org/record/14896109/files/figure.png" pageId="4" pageNumber="81">20B</figureCitation>
;
<figureCitation id="133A2A41FF8F6936372228F5FDBFFBDD" box="[549,596,1015,1041]" captionStart="FIGURE 21" captionStartId="19.[151,250,1017,1041]" captionTargetBox="[158,1429,424,987]" captionTargetId="figure-116@19.[151,1436,417,994]" captionTargetPageId="19" captionText="FIGURE 21. A, Giemsa-stained karyotype of Fulakora elongata (2n=24; 16m + 8sm); B, the same for F. estevae (2n=24; 16m + 8sm). Scale bars: 5 μm. Abbreviations: m, metacentric; sm, submetacentric." figureDoi="http://doi.org/10.5281/zenodo.14896111" httpUri="https://zenodo.org/record/14896111/files/figure.png" pageId="4" pageNumber="81">21B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF8F693635902F3DFF14FAB5" pageId="4" pageNumber="81" type="materials_examined">
<paragraph id="8BBE36C4FF8F693635902F3DFB5FFAC1" blockId="4.[151,1437,1086,1905]" pageId="4" pageNumber="81">
<emphasis id="B975EAD6FF8F693635902F3DFEACFB94" bold="true" box="[151,327,1086,1113]" pageId="4" pageNumber="81">Type material.</emphasis>
<materialsCitation id="3B693C99FF8F693634572F3DFE64FB6D" collectingDate="2021-09-01" collectionCode="CELC" collectorName="Manual Jacintho, G. F. &amp; Chaul, J." country="BRA" latitude="-20.80183" location="M. Paraiso" longLatPrecision="1" longitude="-42.86526" municipality="Vicosa" pageId="4" pageNumber="81" specimenCode="GFJ000100" specimenCount="1" stateProvince="MG" typeStatus="holotype">
<typeStatus id="54BA8866FF8F693634572F3DFE53FB95" box="[336,440,1087,1113]" pageId="4" pageNumber="81" type="holotype">Holotype</typeStatus>
worker. Original label: “
<collectingCountry id="F3167654FF8F693637DF2F3DFCF3FB95" box="[728,792,1087,1113]" pageId="4" pageNumber="81">BRA</collectingCountry>
:
<collectingRegion id="49C5F826FF8F693636222F3DFCBEFB95" box="[805,853,1087,1113]" pageId="4" pageNumber="81">MG</collectingRegion>
,
<collectingMunicipality id="6BDAACBEFF8F693636662F3DFC5BFB95" box="[865,944,1087,1113]" pageId="4" pageNumber="81">Viçosa</collectingMunicipality>
,
<location id="8EDE601FFF8F693636B92F3DFBD2FB95" LSID="urn:lsid:plazi:treatment:03A887D2FF8F6938359028D0FDEFFE1D:8EDE601FFF8F693636B92F3DFBD2FB95" box="[958,1081,1087,1113]" country="BRA" latitude="-20.80183" longLatPrecision="1" longitude="-42.86526" municipality="Vicosa" name="M. Paraiso" pageId="4" pageNumber="81" stateProvince="MG">M. Paraíso</location>
<geoCoordinate id="EE355003FF8F693631452F3DFB53FB95" box="[1090,1208,1087,1113]" degrees="20.80183" direction="south" orientation="latitude" pageId="4" pageNumber="81" precision="1" value="-20.80183">-20.80183</geoCoordinate>
,
<geoCoordinate id="EE355003FF8F693631C22F3CFADCFB95" box="[1221,1335,1086,1113]" degrees="42.86526" direction="west" orientation="longitude" pageId="4" pageNumber="81" precision="1" value="-42.86526">-42.86526</geoCoordinate>
<date id="FFBF1004FF8F693630472F3DFA77FB95" box="[1344,1436,1087,1113]" pageId="4" pageNumber="81" value="2021-09-01">
<collectingDate id="EFFBE9ECFF8F693630472F3DFA77FB95" box="[1344,1436,1087,1113]" pageId="4" pageNumber="81" value="2021-09-01">01.ix.21</collectingDate>
</date>
<collectorName id="26F45312FF8F693635902F61FE46FBB1" box="[151,429,1123,1149]" pageId="4" pageNumber="81">Manual Jacintho, G. F.</collectorName>
;
<collectorName id="26F45312FF8F693634C72F61FDC1FBB1" box="[448,554,1123,1149]" pageId="4" pageNumber="81">Chaul, J.</collectorName>
” (pinned, unique specimen identifier ANTWEB1047071, collecting event
<specimenCode id="DBA79EBFFF8F693635902F85FEF7FB6D" box="[151,284,1159,1185]" pageId="4" pageNumber="81">GFJ000100</specimenCode>
) [
<emphasis id="B975EAD6FF8F693634312F84FE6FFB6C" bold="true" box="[310,388,1158,1184]" pageId="4" pageNumber="81">
<collectionCode id="ED10AE01FF8F693634312F84FE6FFB6C" box="[310,388,1158,1184]" collectionName="CELC" pageId="4" pageNumber="81">CELC</collectionCode>
</emphasis>
]
</materialsCitation>
.
<materialsCitation id="3B693C99FF8F693634992F85FC7CFB09" collectingDate="2021-09-01" collectionCode="DZUP" collectorName="M. Paraiso" country="India" latitude="-20.80183" location="Chaul" longLatPrecision="1" longitude="-42.86526" municipality="Vicosa" pageId="4" pageNumber="81" specimenCode="GFJ000100" specimenCount="1" specimenCount-worker="1" stateProvince="MG" typeStatus="paratype">
<typeStatus id="54BA8866FF8F693634992F85FDEFFB6D" box="[414,516,1159,1185]" pageId="4" pageNumber="81" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D07FD4DFF8F693637162F85FD79FB6D" box="[529,658,1159,1185]" count="1" pageId="4" pageNumber="81" type="worker">one worker</specimenCount>
with the same label data as the holotype (pinned, unique specimen identifier ANTWEB1047070, collecting event
<specimenCode id="DBA79EBFFF8F693637A32FA9FCC2FB09" box="[676,809,1195,1221]" pageId="4" pageNumber="81">GFJ000100</specimenCode>
) [
<emphasis id="B975EAD6FF8F693636472FA9FC67FB09" bold="true" box="[832,908,1195,1221]" pageId="4" pageNumber="81">
<collectionCode id="ED10AE01FF8F693636472FA9FC67FB09" box="[832,908,1195,1221]" collectionName="Brazil, Parana, Curitiba, Universidade Federal do Parana, Museu de Entomologia Pe. Jesus Santiago Moure" pageId="4" pageNumber="81">DZUP</collectionCode>
</emphasis>
]
</materialsCitation>
.
<materialsCitation id="3B693C99FF8F693636A42FA9FB44FAC1" collectingDate="2014-10-11" collectionCode="JCMC" collectorName="J. Chaul &amp; R. Jesus" country="BRA" latitude="-20.8" location="Mata do Paraiso" longLatPrecision="1266" longitude="-42.85" municipality="Vicosa" pageId="4" pageNumber="81" specimenCode="JCMC00652" specimenCount="1" specimenCount-worker="1" stateProvince="MG" typeStatus="paratype">
<typeStatus id="54BA8866FF8F693636A42FA9FBE2FB09" box="[931,1033,1195,1221]" pageId="4" pageNumber="81" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D07FD4DFF8F693631132FA9FB70FB09" box="[1044,1179,1195,1221]" count="1" pageId="4" pageNumber="81" type="worker">One worker</specimenCount>
. Original label: “
<collectingCountry id="F3167654FF8F6936305E2FA9FA72FB09" box="[1369,1433,1195,1221]" pageId="4" pageNumber="81">BRA</collectingCountry>
:
<collectingRegion id="49C5F826FF8F693635902FCDFF2CFB25" box="[151,199,1231,1257]" pageId="4" pageNumber="81">MG</collectingRegion>
,
<collectingMunicipality id="6BDAACBEFF8F693635D22FCDFECFFB25" box="[213,292,1231,1257]" pageId="4" pageNumber="81">Viçosa</collectingMunicipality>
,
<location id="8EDE601FFF8F693634352FCDFE06FB25" LSID="urn:lsid:plazi:treatment:03A887D2FF8F6938359028D0FDEFFE1D:8EDE601FFF8F693634352FCDFE06FB25" box="[306,493,1231,1257]" country="BRA" latitude="-20.8" longLatPrecision="1266" longitude="-42.85" municipality="Vicosa" name="Mata do Paraiso" pageId="4" pageNumber="81" stateProvince="MG">Mata do Paraíso</location>
<geoCoordinate id="EE355003FF8F693634F02FCDFDBAFB25" box="[503,593,1231,1257]" degrees="20" direction="south" minutes="48" orientation="latitude" pageId="4" pageNumber="81" precision="925" value="-20.8">20º48S</geoCoordinate>
<geoCoordinate id="EE355003FF8F6936375C2FCCFD54FB25" box="[603,703,1230,1257]" degrees="42" direction="west" minutes="51" orientation="longitude" pageId="4" pageNumber="81" precision="925" value="-42.85">42º51W</geoCoordinate>
<date id="FFBF1004FF8F693637CE2FCDFCD3FB25" box="[713,824,1231,1257]" pageId="4" pageNumber="81" value="2014-10-11">
<collectingDate id="EFFBE9ECFF8F693637CE2FCDFCD3FB25" box="[713,824,1231,1257]" pageId="4" pageNumber="81" value="2014-10-11">11.x.2014</collectingDate>
</date>
rotten log Winkler II.
<collectorName id="26F45312FF8F693631452FCDFB4AFB25" box="[1090,1185,1231,1257]" pageId="4" pageNumber="81">J. Chaul</collectorName>
&amp;
<collectorName id="26F45312FF8F693631CC2FCDFAC0FB25" box="[1227,1323,1231,1257]" pageId="4" pageNumber="81">R. Jesus</collectorName>
” (pinned, unique specimen identifier UFV-LABECOL-000012, collecting event
<specimenCode id="DBA79EBFFF8F693636AA2FF0FBD5FAC1" box="[941,1086,1266,1293]" pageId="4" pageNumber="81">JCMC00652</specimenCode>
) [
<emphasis id="B975EAD6FF8F693631512FF0FB4FFAC0" bold="true" box="[1110,1188,1266,1292]" pageId="4" pageNumber="81">MZSP</emphasis>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE36C4FF8F693635C02E15FF14FAB5" blockId="4.[151,1437,1086,1905]" pageId="4" pageNumber="81">
<emphasis id="B975EAD6FF8F693635C02E15FE42FAFC" bold="true" box="[199,425,1302,1329]" pageId="4" pageNumber="81">
<typeStatus id="54BA8866FF8F693635C02E15FE48FAFC" box="[199,419,1302,1329]" isOtherTypeStatus="true" pageId="4" pageNumber="81">Non-type material</typeStatus>
.
</emphasis>
One entirely dissected worker with the same data as the holotype (some sclerites mounted on cards and some between slides; unique specimen identifier ANTWEB1047069, collecting event GFJ000100) [
<emphasis id="B975EAD6FF8F693635A62E5CFF04FAB4" bold="true" box="[161,239,1374,1400]" pageId="4" pageNumber="81">CELC</emphasis>
].
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF8F693635C02E81FB45F99D" pageId="4" pageNumber="81" type="diagnosis">
<paragraph id="8BBE36C4FF8F693635C02E81FB45F99D" blockId="4.[151,1437,1086,1905]" pageId="4" pageNumber="81">
<emphasis id="B975EAD6FF8F693635C02E81FE48FA50" bold="true" box="[199,419,1410,1437]" pageId="4" pageNumber="81">Worker diagnosis.</emphasis>
Body mostly black, except for the orangish mandibles, antennae, and legs (
<figureCitation id="133A2A41FF8F6936300A2E81FAA0FA51" box="[1293,1355,1411,1437]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Fig.2</figureCitation>
). Head longer than wide (CI 87.3290.43), not strongly tapering posteriorly (HTI 91.92 95.48); costate on the anterior half, smooth posteriorly (
<figureCitation id="133A2A41FF8F693634B52EC9FDE6FA29" box="[434,525,1483,1509]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Figs.2B</figureCitation>
;
<figureCitation id="133A2A41FF8F6936371E2EC9FDD5FA29" box="[537,574,1483,1509]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="4" pageNumber="81">3A</figureCitation>
). Genal tooth about as long as clypeal tubercle plus its chaeta (
<figureCitation id="133A2A41FF8F693631FD2EC9FAA1FA29" box="[1274,1354,1483,1509]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Fig.2B</figureCitation>
; GentL 0.05, Ch+CltbL 0.04). Mandibles about 2/3 of head length (
<figureCitation id="133A2A41FF8F693636222EEDFC9EF9C5" box="[805,885,1519,1545]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Fig.2B</figureCitation>
; ML 0.710.74, HL 0.860.89). Mesometapleural sulcus absent (
<figureCitation id="133A2A41FF8F6936343E2D11FE7CF9E1" box="[313,407,1555,1581]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Figs.2A</figureCitation>
;
<figureCitation id="133A2A41FF8F693634A62D10FE2CF9E1" box="[417,455,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="4" pageNumber="81">5A</figureCitation>
). Propodeal corners without pale patches (
<figureCitation id="133A2A41FF8F6936369A2D11FC1BF9E1" box="[925,1008,1555,1581]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="4" pageNumber="81">Fig.2A</figureCitation>
). Petiole dorsum evidently longer than the first gaster posttergite (PtI 107.14114.41, AIIIL 0.310.34); subpetiolar process reduced.
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF8F693635C02D58FBFCF9B9" box="[199,1047,1626,1653]" pageId="4" pageNumber="81" type="distribution">
<paragraph id="8BBE36C4FF8F693635C02D58FBFCF9B9" blockId="4.[151,1437,1086,1905]" box="[199,1047,1626,1653]" pageId="4" pageNumber="81">
<emphasis id="B975EAD6FF8F693635C02D58FDEEF9B8" bold="true" box="[199,517,1626,1652]" pageId="4" pageNumber="81">Geographical distribution.</emphasis>
Viçosa,
<collectingRegion id="49C5F826FF8F693637612D59FD14F9B9" box="[614,767,1627,1653]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="81">Minas Gerais</collectingRegion>
,
<collectingCountry id="F3167654FF8F6936360D2D59FCA5F9B9" box="[778,846,1627,1653]" name="Brazil" pageId="4" pageNumber="81">Brazil</collectingCountry>
(Atlantic Forest).
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF8F693B35C02D7DFB28F90A" lastPageId="9" lastPageNumber="86" pageId="4" pageNumber="81" type="description">
<paragraph id="8BBE36C4FF8F693635C02D7DFD03F8BD" blockId="4.[151,1437,1086,1905]" pageId="4" pageNumber="81">
<emphasis id="B975EAD6FF8F693635C02D7DFD3DF955" bold="true" box="[199,726,1662,1689]" pageId="4" pageNumber="81">
Measurements (mm) and indexes.
<typeStatus id="54BA8866FF8F693637652D7DFD3AF955" box="[610,721,1663,1689]" pageId="4" pageNumber="81" type="holotype">Holotype</typeStatus>
.
</emphasis>
HL 0.89, HW 0.78, VxW 0.74, SL 0.46, ML 0.74, GentL 0.05, Ch+CltbL 0.04, WL 1.2, MsW 0.33, PpW 0.45, PtL 0.53, PtW 0.48, AIIIL 0.33, AIVL 0.40, AVL 0.15, AVIL 0.13, AVIIL 0.29, TL 4.64, CI 87.32, SI 52.11, MI 83.10, PtI 110.53, HTI 95.48, MFI 73.11.
<emphasis id="B975EAD6FF8F693631DF2DC4FF17F8C8" bold="true" pageId="4" pageNumber="81">Other examined material</emphasis>
(N=3). HL 0.860.87, HW 0.78, VxW 0.720.74, SL 0.44, ML 0.710.74, GentL 0.05, Ch+CltbL 0.04, WL 1.161.30, MsW 0.320.34, PpW 0.450.46, PtL 0.500.54, PtW 0.460.47, AIIIL 0.310.34, AIVL 0.380.41, AVL 0.150.17, AVIL 0.10, AVIIL 0.280.30 TL 4.484.69, CI 89.8690.43, SI 50.5250.72, MI 82.6185.51, PtI 107.14114.41, HTI 91.9293.88, MFI 68.9172.97.
</paragraph>
<caption id="DF7E664CFF8E693735902C2DFBF6F8A7" ID-DOI="http://doi.org/10.5281/zenodo.14896071" ID-Zenodo-Dep="14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="5" pageNumber="82" startId="5.[151,250,1839,1863]" targetBox="[151,1435,181,1815]" targetPageId="5" targetType="figure">
<paragraph id="8BBE36C4FF8E693735902C2DFBF6F8A7" blockId="5.[151,1436,1839,1899]" pageId="5" pageNumber="82">
<emphasis id="B975EAD6FF8E693735902C2DFEE6F88B" bold="true" box="[151,269,1839,1863]" pageId="5" pageNumber="82">FIGURE 2</emphasis>
.
<taxonomicName id="4C014D47FF8E6937341E2C2DFE23F88B" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[281,456,1839,1863]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF8E6937341E2C2DFE23F88B" box="[281,456,1839,1863]" italics="true" pageId="5" pageNumber="82">Fulakora estevae</emphasis>
</taxonomicName>
<emphasis id="B975EAD6FF8E693734CA2C2DFDF6F88B" bold="true" box="[461,541,1839,1863]" pageId="5" pageNumber="82">
<taxonomicNameLabel id="A24657ADFF8E693734CA2C2DFDF6F88B" box="[461,541,1839,1863]" pageId="5" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype worker (ANTWEB1047071).
<emphasis id="B975EAD6FF8E693736BF2C2DFC21F88B" bold="true" box="[952,970,1839,1863]" pageId="5" pageNumber="82">A</emphasis>
, profile view of the body;
<emphasis id="B975EAD6FF8E693731D02C2DFB03F88B" bold="true" box="[1239,1256,1839,1863]" pageId="5" pageNumber="82">B</emphasis>
, full-face view of head;
<emphasis id="B975EAD6FF8E693735D32C51FF0DF8A7" bold="true" box="[212,230,1875,1899]" pageId="5" pageNumber="82">C</emphasis>
, dorsal view of the body;
<emphasis id="B975EAD6FF8E693734F72C51FDE9F8A7" bold="true" box="[496,514,1875,1899]" pageId="5" pageNumber="82">D</emphasis>
, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B).
</paragraph>
</caption>
<paragraph id="8BBE36C4FF8D693435C02B95FAACFC91" blockId="6.[151,1437,150,861]" pageId="6" pageNumber="83">
<emphasis id="B975EAD6FF8D693435C02B95FDECFF7D" bold="true" box="[199,519,150,177]" pageId="6" pageNumber="83">Worker description. Head.</emphasis>
In full-face view, head subrectangular, longer than wide (CI 87.3290.43), wider anteriorly than posteriorly (HTI 91.9295.48); vertexal margin straight to slightly concave; lateral margins slightly convex (
<figureCitation id="133A2A41FF8D693435F02BDDFEB1FF35" box="[247,346,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="6" pageNumber="83">Figs. 2B</figureCitation>
,
<figureCitation id="133A2A41FF8D693434622BDDFE61FF35" box="[357,394,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">3A</figureCitation>
). Overlapping mandibles; mandible about 4/5 as long as head (ML 0.710.74, HL 0.860.89; MI 82.6185.51); outer margin almost straight along its basal 3/4, apically strongly curved; inner margin dentition arrangement, from apex to base: a prominent single apical tooth; a pair of rounded-tipped subapical teeth; five pairs of sharp, equally-sized, and basally fused teeth posteriorly directed, each pair with one tooth inclined dorsally and the other ventrally (
<figureCitation id="133A2A41FF8D693434792A6DFE32FE45" box="[382,473,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="6" pageNumber="83">Figs.2B</figureCitation>
;
<figureCitation id="133A2A41FF8D693434EF2A6DFDC4FE45" box="[488,559,367,393]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">3A, F</figureCitation>
); one large, broad, trapezoidal basal tooth. Anterior clypeal margin convex; medial section with five tubercles, with dentiform chaetae on their apexes, arranged as follows: medial tubercle with two chaetae diverging laterally (apparently resulted from the fusion of two tubercles); next to the medial tubercle, on each side, a thin tubercle with one chaeta inclined laterally; further laterad, a thick tubercle with two chaetae inclined laterally (apparently also resulted from the fusion of two tubercles); tubercles about two to three times their chaetae size (
<figureCitation id="133A2A41FF8D693434B52921FDE8FDF1" box="[434,515,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">Fig.3B</figureCitation>
). Genal tooth acute, relatively small, longer than its base, shorter than maximum basal width of mandible, and about as long as the thinnest clypeal tubercle plus its chaeta (
<figureCitation id="133A2A41FF8D693431B12945FAECFDAD" box="[1206,1287,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">Fig.3E</figureCitation>
). In full-face view, the distance between frontal lobes to anterior clypeal margin about half as long as lobes; frontal carinae reach about mid-length of head, with a feeble medial sulcus between them. Compound eyes with few (about four) fused, degenerate ommatidia (
<figureCitation id="133A2A41FF8D693434EF29B1FDD1FD01" box="[488,570,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">Fig.3D</figureCitation>
), located just posterior to head mid-length, barely interrupting lateral margins of head in full-face view. Twelve antennomeres; scape about half as long as remaining antennomeres combined; pedicel almost as long as antennomere 35 combined; antennomeres 312 gradually increasing; antennomeres 311 mildly constricted apically (
<figureCitation id="133A2A41FF8D693434D5281DFDCFFCF5" box="[466,548,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83">Fig.3C</figureCitation>
). Lateral margins of labrum concave; distal margin with a narrow medial notch between short, broad lobes (
<figureCitation id="133A2A41FF8D693434D42841FDCCFC91" box="[467,551,835,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,603,627]" captionTargetBox="[151,1436,181,579]" captionTargetId="figure-19@7.[151,1436,181,579]" captionTargetPageId="7" captionText="FIGURE 4. Fulakora estevae sp. nov., worker (ANTWEB1047069) dissected mouthparts. A, labrum, and B, labiomaxillary complex. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896075" httpUri="https://zenodo.org/record/14896075/files/figure.png" pageId="6" pageNumber="83">Fig.4A</figureCitation>
). Palpal formula 3, 2 (three maxillary palpomeres, two labial) (
<figureCitation id="133A2A41FF8D693431EE2841FAD1FC91" box="[1257,1338,835,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,603,627]" captionTargetBox="[151,1436,181,579]" captionTargetId="figure-19@7.[151,1436,181,579]" captionTargetPageId="7" captionText="FIGURE 4. Fulakora estevae sp. nov., worker (ANTWEB1047069) dissected mouthparts. A, labrum, and B, labiomaxillary complex. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896075" httpUri="https://zenodo.org/record/14896075/files/figure.png" pageId="6" pageNumber="83">Fig.4B</figureCitation>
).
</paragraph>
<caption id="DF7E664CFF8D693435902C1BFAEEF851" ID-DOI="http://doi.org/10.5281/zenodo.14896073" ID-Zenodo-Dep="14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="6" pageNumber="83" startId="6.[151,250,1817,1841]" targetBox="[151,1436,885,1793]" targetPageId="6" targetType="figure">
<paragraph id="8BBE36C4FF8D693435902C1BFAEEF851" blockId="6.[151,1436,1817,1950]" pageId="6" pageNumber="83">
<emphasis id="B975EAD6FF8D693435902C1BFEE4F8FD" bold="true" box="[151,271,1817,1841]" pageId="6" pageNumber="83">FIGURE 3</emphasis>
.
<taxonomicName id="4C014D47FF8D6934341A2C1BFE25F8FD" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[285,462,1817,1841]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF8D6934341A2C1BFE25F8FD" box="[285,462,1817,1841]" italics="true" pageId="6" pageNumber="83">Fulakora estevae</emphasis>
</taxonomicName>
<emphasis id="B975EAD6FF8D693434D22C1BFDC3F8FD" bold="true" box="[469,552,1817,1841]" pageId="6" pageNumber="83">
<taxonomicNameLabel id="A24657ADFF8D693434D22C1BFDC3F8FD" box="[469,552,1817,1841]" pageId="6" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype worker (ANTWEB1047070).
<emphasis id="B975EAD6FF8D693436CD2C1BFC37F8FD" bold="true" box="[970,988,1817,1841]" pageId="6" pageNumber="83">A</emphasis>
, full-face view of head;
<emphasis id="B975EAD6FF8D693431DA2C1BFB05F8FD" bold="true" box="[1245,1262,1817,1841]" pageId="6" pageNumber="83">B</emphasis>
, anterior clypeal margin;
<emphasis id="B975EAD6FF8D693435EB2C3FFF15F899" bold="true" box="[236,254,1853,1877]" pageId="6" pageNumber="83">C</emphasis>
, antennal funiculus;
<emphasis id="B975EAD6FF8D693434D32C3FFE0DF899" bold="true" box="[468,486,1853,1877]" pageId="6" pageNumber="83">D</emphasis>
, compound eye, arrows indicate fused and degenerate ommatidia;
<emphasis id="B975EAD6FF8D693431882C3FFB4BF899" bold="true" box="[1167,1184,1853,1877]" pageId="6" pageNumber="83">E</emphasis>
, genal tooth;
<emphasis id="B975EAD6FF8D6934302B2C3FFAD0F899" bold="true" box="[1324,1339,1853,1877]" pageId="6" pageNumber="83">F</emphasis>
, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations:
<emphasis id="B975EAD6FF8D693430252C63FAD0F8B5" bold="true" box="[1314,1339,1889,1913]" pageId="6" pageNumber="83">ch</emphasis>
, chaetae;
<emphasis id="B975EAD6FF8D693435902C87FF2BF851" bold="true" box="[151,192,1925,1949]" pageId="6" pageNumber="83">cltb</emphasis>
, clypeal tubercles;
<emphasis id="B975EAD6FF8D693434822C87FE4AF851" bold="true" box="[389,417,1925,1949]" pageId="6" pageNumber="83">cls</emphasis>
, median pair of clypeal setae;
<emphasis id="B975EAD6FF8D693437D02C87FD06F851" bold="true" box="[727,749,1925,1949]" pageId="6" pageNumber="83">ce</emphasis>
, compound eye;
<emphasis id="B975EAD6FF8D6934369B2C87FC5AF851" bold="true" box="[924,945,1925,1949]" pageId="6" pageNumber="83">gt</emphasis>
, genal tooth;
<emphasis id="B975EAD6FF8D6934313A2C87FBB4F851" bold="true" box="[1085,1119,1925,1949]" pageId="6" pageNumber="83">sps</emphasis>
, spatulate setae.
</paragraph>
</caption>
<caption id="DF7E664CFF8C693535902959FE50FD5B" ID-DOI="http://doi.org/10.5281/zenodo.14896075" ID-Zenodo-Dep="14896075" httpUri="https://zenodo.org/record/14896075/files/figure.png" pageId="7" pageNumber="84" startId="7.[151,250,603,627]" targetBox="[151,1436,181,579]" targetPageId="7" targetType="figure">
<paragraph id="8BBE36C4FF8C693535902959FE50FD5B" blockId="7.[151,1436,603,663]" pageId="7" pageNumber="84">
<emphasis id="B975EAD6FF8C693535902959FEFFFDBF" bold="true" box="[151,276,603,627]" pageId="7" pageNumber="84">FIGURE 4.</emphasis>
<taxonomicName id="4C014D47FF8C6935341B2959FE26FDBF" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[284,461,603,627]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF8C6935341B2959FE26FDBF" box="[284,461,603,627]" italics="true" pageId="7" pageNumber="84">Fulakora estevae</emphasis>
</taxonomicName>
<emphasis id="B975EAD6FF8C693534D22959FDCCFDBF" bold="true" box="[469,551,603,627]" pageId="7" pageNumber="84">
<taxonomicNameLabel id="A24657ADFF8C693534D22959FDCCFDBF" box="[469,551,603,627]" pageId="7" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, worker (ANTWEB1047069) dissected mouthparts.
<emphasis id="B975EAD6FF8C693531402959FBB2FDBF" bold="true" box="[1095,1113,603,627]" pageId="7" pageNumber="84">A</emphasis>
, labrum, and
<emphasis id="B975EAD6FF8C693531EE2959FB11FDBF" bold="true" box="[1257,1274,603,627]" pageId="7" pageNumber="84">B</emphasis>
, labiomaxillary complex. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BBE36C4FF8C693535C029E8FCBDFB5C" blockId="7.[151,1437,745,1996]" pageId="7" pageNumber="84">
<emphasis id="B975EAD6FF8C693535C029E8FEA7FCC8" bold="true" box="[199,332,746,772]" pageId="7" pageNumber="84">Mesosoma.</emphasis>
In dorsal view, anterior margin of pronotum round (excluding pronotal neck), lateral margins slightly converging posteriorly; pronotum separated from the posterior mesosoma by a constriction, the promesonotal articulation (
<figureCitation id="133A2A41FF8C693534212833FE93FC87" box="[294,376,817,843]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Fig.2C</figureCitation>
). Posterior to the pronotum, all mesosoma sclerites (mesothorax, metathorax, and propodeum) fused, block-like, without sutures, lines, or carinae delimiting them. In dorsal view, area corresponding to the mesonotum, metanotum, and dorsum of propodeum trapezoidal, as long as pronotum (excluding the mesothorax articulatory anterior edge); pronotum as wide as propodeum posterior (maximum) width. Articulatory anterior edge (mesothrorax presclerite) thick, sometimes visible (
<figureCitation id="133A2A41FF8C6935362328C3FC93FC10" box="[804,888,961,988]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5A</figureCitation>
). In profile, dorsal outline of mesosoma almost straight, except for the oblique anterior pronotum and posterior propodeum faces. Latero-posterior margins of propodeum well-defined and conspicuously angled in relation to posterior surface; dorsal surface softly curving into posterior surface (
<figureCitation id="133A2A41FF8C693534662F2FFE54FB8B" box="[353,447,1069,1095]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Figs.2A</figureCitation>
;
<figureCitation id="133A2A41FF8C693534CD2F2FFE1BFB84" box="[458,496,1069,1096]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">5A</figureCitation>
). Metathoracic spiracle tiny, twice as distant from the propodeal spiracle as from the mesothoracic spiracle; propodeal spiracle round, just below mid-height of propodeum. Metapleural gland opening a curved slit, overlapped ventrally by a cuticular flap (
<figureCitation id="133A2A41FF8C693537F12F77FCA1FB5C" box="[758,842,1141,1168]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5A</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C4FF8C693535C02F98FDD2FA18" blockId="7.[151,1437,745,1996]" pageId="7" pageNumber="84">
<emphasis id="B975EAD6FF8C693535C02F98FEEDFB78" bold="true" box="[199,262,1178,1204]" pageId="7" pageNumber="84">Legs.</emphasis>
Fore- and hindlegs more robust than midleg; with the legs in postero-lateral view, pro- and metafemora about 1.3x thicker than mesofemur; mesotibia shorter and thinner than pro- and metatibiae (
<figureCitation id="133A2A41FF8C6935318A2FBFFB0AFB1B" box="[1165,1249,1213,1239]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Fig.2A</figureCitation>
). Protibial calcar fully pectinate, without velum (
<figureCitation id="133A2A41FF8C693534FA2FE3FDBBFB30" box="[509,592,1249,1276]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5D</figureCitation>
). Mesotibia with short posterior spur, which is basally glabrous with some apical microtrichia; anterior spur not observed. Anterior metatibial spur slightly longer than mesotibial posterior spur, almost glabrous, with a few inconspicuous microtrichia; posterior spur resembling the calcar, sinuous, fully pectinate with lanceolate microtrichia. Posterior surface of probasitarsal notch armed with a single stout seta; inner portion of probasitarsus without a basal projection. Basitarsal sulcus gland present on mesobasitarsus (
<bibRefCitation id="EF904B35FF8C693530172E73FF38FA7C" author="Billen" etAl="et al." firstAuthor="Billen" pageId="7" pageNumber="84" pagination="1 - 10" refId="ref11904" refString="Billen, J., Khalife, A., Ito, F., Anh, N. D. &amp; Esteves, F. A. (2021) The basitarsal sulcus gland, a novel exocrine structure in ants. Arthropod Structure &amp; Development, 61 (101041), 1-10. https://doi.org/10.1016/j.asd.2021.101041" type="journal article" year="2021">
Billen
<emphasis id="B975EAD6FF8C6935305A2E70FA65FA47" box="[1373,1422,1393,1419]" italics="true" pageId="7" pageNumber="84">et al</emphasis>
., 2021
</bibRefCitation>
); absent on metabasitarsus. Pretarsal claws subequal on all legs, about the size of the fourth tarsomere, without teeth on their inner margins (
<figureCitation id="133A2A41FF8C693534DB2EBBFDC7FA18" box="[476,556,1465,1492]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5E</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C4FF8C693535C02EDCFDCFF800" blockId="7.[151,1437,745,1996]" pageId="7" pageNumber="84">
<emphasis id="B975EAD6FF8C693535C02EDCFEA0FA34" bold="true" box="[199,331,1502,1528]" pageId="7" pageNumber="84">Metasoma.</emphasis>
Petiole sessile. In dorsal view, petiole longer than wide, subrectangular (
<figureCitation id="133A2A41FF8C6935318B2EDFFB35FA3B" box="[1164,1246,1501,1527]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Fig.2C</figureCitation>
); in lateral view, petiole longer than high, trapezoidal (
<figureCitation id="133A2A41FF8C693537412D03FD4FF9D7" box="[582,676,1537,1563]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Figs.2A</figureCitation>
;
<figureCitation id="133A2A41FF8C693537B62D03FD3EF9D0" box="[689,725,1537,1564]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">5B</figureCitation>
). In lateral view, anterior margin of petiole straight to slightly concave, perpendicular to the slightly convex dorsal margin; paired protuberances just anterior to the petiolar spiracle small. In lateral view, petiolar sternite visible at the anterior two thirds of the ventral margin of the petiole, with constant width along its anterior visible half, obtusely angled at midpoint and becoming progressively narrower along its posterior visible half; laterotergite triangular, visible in lateral view at the posterior third of the ventral margin of the petiole. Petiolar sternite anterior process not surpassing anteroventral “pinch-like&quot; fusion area of tergite; its anterior edge about half as tall as the anteroventral “pinch-like&quot; fusion area of tergite (
<bibRefCitation id="EF904B35FF8C6935300A2DDBFA7CF938" author="Ward" box="[1293,1431,1753,1780]" firstAuthor="Ward" pageId="7" pageNumber="84" pagination="159 - 175" refId="ref13590" refString="Ward, P. S. (1994) Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology, 19, 159-175. https://doi.org/10.1111/j.1365-3113.1994.tb00585.x" type="journal article" year="1994">Ward, 1994</bibRefCitation>
;
<bibRefCitation id="EF904B35FF8C693535902DFFFECCF8D4" author="Keller" box="[151,295,1789,1816]" firstAuthor="Keller" pageId="7" pageNumber="84" pagination="1 - 90" refId="ref12495" refString="Keller, R. A. (2011) A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the poneromorph subfamilies. Bulletin of the American Museum of Natural History, 355, 1-90. https://doi.org/10.1206/355.1" type="journal article" year="2011">Keller, 2011</bibRefCitation>
), that lies dorsad it. Prora well-developed as a pair of swellings ventrally carinated. Helcium supraaxial (
<figureCitation id="133A2A41FF8C693535DB2C23FEC5F8F0" box="[220,302,1825,1852]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5B</figureCitation>
). Helcium and presclerites of abdominal segment IV separated from postsclerites by scrobiculate sulci (cinctus); cinctus of AIII about 0.80x as thick as cinctus of AIV, in profile. Posttergite of AIV slightly longer than that of AIII; length of posttergites V and VI subequal and the shortest in metasoma; posttergite VII (pygidium) about as long as posttergite III (
<figureCitation id="133A2A41FF8C693534BD2C8FFDE6F86B" box="[442,525,1933,1959]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="7" pageNumber="84">Fig.2A</figureCitation>
). Poststernite VII (hypopygium) with a row of 10 stout setae on each side. Sting apparatus well-developed (
<figureCitation id="133A2A41FF8C693534C12CB3FDF3F800" box="[454,536,1969,1996]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="7" pageNumber="84">Fig.5C</figureCitation>
).
</paragraph>
<caption id="DF7E664CFF83693A35902F0FFDAFFB15" ID-DOI="http://doi.org/10.5281/zenodo.14896077" ID-Zenodo-Dep="14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="8" pageNumber="85" startId="8.[151,250,1037,1061]" targetBox="[151,1436,181,1013]" targetPageId="8" targetType="figure">
<paragraph id="8BBE36C4FF83693A35902F0FFDAFFB15" blockId="8.[151,1437,1037,1241]" pageId="8" pageNumber="85">
<emphasis id="B975EAD6FF83693A35902F0FFEE4FBE9" bold="true" box="[151,271,1037,1061]" pageId="8" pageNumber="85">FIGURE 5</emphasis>
.
<taxonomicName id="4C014D47FF83693A341A2F0FFE25FBE9" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[285,462,1037,1061]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF83693A341A2F0FFE25FBE9" box="[285,462,1037,1061]" italics="true" pageId="8" pageNumber="85">Fulakora estevae</emphasis>
</taxonomicName>
<emphasis id="B975EAD6FF83693A34D12F0FFDC3FBE9" bold="true" box="[470,552,1037,1061]" pageId="8" pageNumber="85">
<taxonomicNameLabel id="A24657ADFF83693A34D12F0FFDC3FBE9" box="[470,552,1037,1061]" pageId="8" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype worker (ANTWEB1047070).
<emphasis id="B975EAD6FF83693A36CC2F0FFC36FBE9" bold="true" box="[971,989,1037,1061]" pageId="8" pageNumber="85">A</emphasis>
, profile of mesosoma;
<emphasis id="B975EAD6FF83693A31C82F0FFB0BFBE9" bold="true" box="[1231,1248,1037,1061]" pageId="8" pageNumber="85">B</emphasis>
, profile of petiole and AIII;
<emphasis id="B975EAD6FF83693A35F92F33FEFBFB85" bold="true" box="[254,272,1073,1097]" pageId="8" pageNumber="85">C</emphasis>
, zoom of poststernites VII (pygidium and hypopygium);
<emphasis id="B975EAD6FF83693A36622F33FC9CFB85" bold="true" box="[869,887,1073,1097]" pageId="8" pageNumber="85">D</emphasis>
, zoom of apex of protibia and base of probasitarsus;
<emphasis id="B975EAD6FF83693A35902F57FF43FBA1" bold="true" box="[151,168,1109,1133]" pageId="8" pageNumber="85">E</emphasis>
, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations:
<emphasis id="B975EAD6FF83693A316C2F57FB68FBA1" bold="true" box="[1131,1155,1109,1133]" pageId="8" pageNumber="85">ca</emphasis>
, calcar;
<emphasis id="B975EAD6FF83693A31D92F57FB1CFBA1" bold="true" box="[1246,1271,1109,1133]" pageId="8" pageNumber="85">he</emphasis>
, helcium;
<emphasis id="B975EAD6FF83693A30602F57FA7EFBA1" bold="true" box="[1383,1429,1109,1133]" pageId="8" pageNumber="85">hyss</emphasis>
, hypopygium spiniform setae;
<emphasis id="B975EAD6FF83693A34D42F7BFDE3FB5D" bold="true" box="[467,520,1145,1169]" pageId="8" pageNumber="85">mtsp</emphasis>
, metapleural spiracle;
<emphasis id="B975EAD6FF83693A37FB2F7BFCC7FB5D" bold="true" box="[764,812,1145,1169]" pageId="8" pageNumber="85">pbss</emphasis>
, probasitarsal spiniform seta;
<emphasis id="B975EAD6FF83693A31692F7BFB75FB5D" bold="true" box="[1134,1182,1145,1169]" pageId="8" pageNumber="85">petp</emphasis>
, petiolar protuberances;
<emphasis id="B975EAD6FF83693A35902F9FFF24FB79" bold="true" box="[151,207,1181,1205]" pageId="8" pageNumber="85">pettb</emphasis>
, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion;
<emphasis id="B975EAD6FF83693A36A42F9FFC03FB79" bold="true" box="[931,1000,1181,1205]" pageId="8" pageNumber="85">prmsa</emphasis>
, promesontal articulation;
<emphasis id="B975EAD6FF83693A30062F9FFACCFB79" bold="true" box="[1281,1319,1181,1205]" pageId="8" pageNumber="85">psp</emphasis>
, propodeal spiracle;
<emphasis id="B975EAD6FF83693A35F42FC3FEF9FB15" bold="true" box="[243,274,1217,1241]" pageId="8" pageNumber="85">scs</emphasis>
, scrobiculate sulcus;
<emphasis id="B975EAD6FF83693A34EB2FC3FE15FB15" bold="true" box="[492,510,1217,1241]" pageId="8" pageNumber="85">st</emphasis>
, sting.
</paragraph>
</caption>
<paragraph id="8BBE36C4FF83693A35C02E29FF2DF91E" blockId="8.[151,1437,1323,2034]" pageId="8" pageNumber="85">
<emphasis id="B975EAD6FF83693A35C02E29FEBEFA89" bold="true" box="[199,341,1323,1349]" pageId="8" pageNumber="85">Sculpturing</emphasis>
. Major area of dorsum of head irregularly costate anteriorly until mid-height, smooth posteriorly and laterally except for foveolate piligerous punctures; frontal lobes and area between the frontal carinae mostly smooth (
<figureCitation id="133A2A41FF83693A35FF2E71FEBFFA41" box="[248,340,1395,1421]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="8" pageNumber="85">Figs.2B</figureCitation>
;
<figureCitation id="133A2A41FF83693A34582E71FE6EFA41" box="[351,389,1395,1421]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">3A</figureCitation>
); clypeus smooth, except for clypeal tubercles; tubercles with tiny, sharp and thick, dentiform, chaetae that are spirally costulate (
<figureCitation id="133A2A41FF83693A371D2E95FD87FA7D" box="[538,620,1431,1457]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">Fig.3B</figureCitation>
). Mandible dorsal surface irregularly costate; apical, preapical, and basal teeth smooth (
<figureCitation id="133A2A41FF83693A343B2EB9FE73FA19" box="[316,408,1467,1493]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="8" pageNumber="85">Figs.2B</figureCitation>
;
<figureCitation id="133A2A41FF83693A34A02EB9FE27FA19" box="[423,460,1467,1493]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">3A</figureCitation>
); remaining teeth irregularly sculptured (
<figureCitation id="133A2A41FF83693A36A12EB9FC1EFA19" box="[934,1013,1467,1493]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">Fig.3F</figureCitation>
). Antennal scape irregularly strigate; remaining antennomeres smooth but for piligerous punctures (
<figureCitation id="133A2A41FF83693A36602EDDFC52FA35" box="[871,953,1503,1529]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">Fig.3C</figureCitation>
). Mesosoma smooth, except for foveolate piligerous punctures on the pronotum and dorsum of the posterior mesosoma. Lower metapleuron costate. Presclerite of the mesonotum imbricate (
<figureCitation id="133A2A41FF83693A34EF2D25FDD0F98E" box="[488,571,1575,1602]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="8" pageNumber="85">Fig.5A</figureCitation>
). Legs smooth except for the foveolate piligerous punctures. Petiolar tergite anterior and anterolateral surfaces smooth, except for the imbricate area ventrad the spiracle and posterolateral foveolate piligerous punctures. Petiolar sternite imbricate. Helcium and presclerites of abdominal segment IV mostly imbricate (
<figureCitation id="133A2A41FF83693A34162D91FE88F962" box="[273,355,1683,1710]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="8" pageNumber="85">Fig.5B</figureCitation>
). Piligerous foveolate punctures present on the petiolar dorsum and abdominal postsclerites III VII.
</paragraph>
<paragraph id="8BBE36C4FF83693B35C02DD9FC2DFF35" blockId="8.[151,1437,1323,2034]" lastBlockId="9.[151,1437,151,717]" lastPageId="9" lastPageNumber="86" pageId="8" pageNumber="85">
<emphasis id="B975EAD6FF83693A35C02DD9FE74F939" bold="true" box="[199,415,1755,1781]" pageId="8" pageNumber="85">Pilosity and color.</emphasis>
Simple, suberect to subdecumbent pilosity all over the body. Mandible with various setae oriented medially and arranged contralaterally along its inner margin, with filiform setae on the dorsal surface and spatulate setae on the ventral surface (
<figureCitation id="133A2A41FF83693A37732C21FD1BF8F1" box="[628,752,1827,1853]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">Figs.3A, F</figureCitation>
). Clypeal anterior margin with a pair of longer filiform setae enclosing the medial tubercle laterally (
<figureCitation id="133A2A41FF83693A37642C45FD5FF8AD" box="[611,692,1863,1889]" captionStart="FIGURE 3" captionStartId="6.[151,250,1817,1841]" captionTargetBox="[151,1436,885,1793]" captionTargetId="figure-358@6.[151,1436,885,1793]" captionTargetPageId="6" captionText="FIGURE 3. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, full-face view of head; B, anterior clypeal margin; C, antennal funiculus; D, compound eye, arrows indicate fused and degenerate ommatidia; E, genal tooth; F, zoom of mandible inner (masticatory) margin. Scale bars: 0.2 mm (A, C), 0.1 mm (B), 0.02 mm (D, E, F). Abbreviations: ch, chaetae; cltb, clypeal tubercles; cls, median pair of clypeal setae; ce, compound eye; gt, genal tooth; sps, spatulate setae." figureDoi="http://doi.org/10.5281/zenodo.14896073" httpUri="https://zenodo.org/record/14896073/files/figure.png" pageId="8" pageNumber="85">Fig.3B</figureCitation>
). Labrum with simple, minute setae (
<figureCitation id="133A2A41FF83693A316E2C45FB56F8AD" box="[1129,1213,1863,1889]" captionStart="FIGURE 4" captionStartId="7.[151,250,603,627]" captionTargetBox="[151,1436,181,579]" captionTargetId="figure-19@7.[151,1436,181,579]" captionTargetPageId="7" captionText="FIGURE 4. Fulakora estevae sp. nov., worker (ANTWEB1047069) dissected mouthparts. A, labrum, and B, labiomaxillary complex. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896075" httpUri="https://zenodo.org/record/14896075/files/figure.png" pageId="8" pageNumber="85">Fig.4A</figureCitation>
). Antennae densely pilose. Mesopleuron, metapleuron, and lateral surface of the propodeum glabrous (
<figureCitation id="133A2A41FF83693A312E2C69FB96F84A" box="[1065,1149,1899,1926]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="8" pageNumber="85">Fig.5A</figureCitation>
). Legs with short pilosity; tibia and tarsi with denser pilosity than the femur. Petiole sternite with long erect setae (
<figureCitation id="133A2A41FF83693A314A2C8DFB74F866" box="[1101,1183,1935,1962]" captionStart="FIGURE 5" captionStartId="8.[151,250,1037,1061]" captionTargetBox="[151,1436,181,1013]" captionTargetId="figure-30@8.[151,1436,181,1013]" captionTargetPageId="8" captionText="FIGURE 5. Fulakora estevae sp. nov., paratype worker (ANTWEB1047070). A, profile of mesosoma; B, profile of petiole and AIII; C, zoom of poststernites VII (pygidium and hypopygium); D, zoom of apex of protibia and base of probasitarsus; E, pretarsal claws. Scale bars: 0.2 mm (A, B), 0.05 mm (C, D), 0.02 mm (E). Abbreviations: ca, calcar; he, helcium; hyss, hypopygium spiniform setae; mtsp, metapleural spiracle; pbss, probasitarsal spiniform seta; petp, petiolar protuberances; pettb, petiolar tergite anteroventral bridge, or &quot;pinch-like&quot; area of fusion; prmsa, promesontal articulation; psp, propodeal spiracle; scs, scrobiculate sulcus; st, sting." figureDoi="http://doi.org/10.5281/zenodo.14896077" httpUri="https://zenodo.org/record/14896077/files/figure.png" pageId="8" pageNumber="85">Fig.5B</figureCitation>
). Pilosity on metasoma slightly longer than on the mesosoma; AVVII with longer setae; AVII with the longest and more densely distributed pilosity. Abdominal presclerites glabrous. Head and body mostly shiny black. Mandible reddish with orangish apex. Antennae orangish. Genal teeth, clypeal anterior portion, and frontal lobes brownish to reddish (
<figureCitation id="133A2A41FF82693B31CE2B95FAF1FF7D" box="[1225,1306,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="9" pageNumber="86">Fig.2B</figureCitation>
). Pronotum and metasomal sclerites with a narrow, lighter-colored area at their posterior limits. Coxae basally black, gradually lightening distally, blending with the orangish color of the legs (
<figureCitation id="133A2A41FF82693B36622BDDFC52FF35" box="[869,953,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1863]" captionTargetBox="[151,1435,181,1815]" captionTargetId="figure-19@5.[151,1435,181,1815]" captionTargetPageId="5" captionText="FIGURE 2. Fulakora estevae sp. nov., holotype worker (ANTWEB1047071). A, profile view of the body; B, full-face view of head; C, dorsal view of the body; D, labels. Scales bars: 0.5 mm (A, C), and 0.2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.14896071" httpUri="https://zenodo.org/record/14896071/files/figure.png" pageId="9" pageNumber="86">Fig.2A</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C4FF82693B35C02A01FB9EFD65" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C02A01FE4FFED0" bold="true" box="[199,420,258,285]" pageId="9" pageNumber="86">Larva description.</emphasis>
TL 4.79, CW 0.21, CL 0.18, Masukos TL 3.68 (N=1). Larva with meconium, at advanced development stage. Small hypognathous head. In full-face view, head slightly longer than wide; occipital margin and lateral margins of head convex. Mandibles strongly sclerotized, slender, half as long as head; in full-face view, outer margins with a notch at mid-length; in profile, posterior margins conspicuously concave; apical teeth curved medially and posteriorly. Labrum bilobed; lobes with round apex. Maxillary and labial palps not visualized; galea finger-shaped, slender and bearing two minute apical sensilla.Antennae small, round, located just posteriad the mid-length of head (
<figureCitation id="133A2A41FF82693B344E2AD9FE76FE39" box="[329,413,475,501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1530,1554]" captionTargetBox="[151,1435,741,1506]" captionTargetId="figure-274@9.[151,1435,741,1506]" captionTargetPageId="9" captionText="FIGURE 6. Fulakora estevae sp. nov., larva. A, head in profile; B. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="9" pageNumber="86">Fig.6A</figureCitation>
). Body “pogonomyrmecoid”
<emphasis id="B975EAD6FF82693B37EE2AD9FCCCFE38" box="[745,807,475,500]" italics="true" pageId="9" pageNumber="86">sensu</emphasis>
<bibRefCitation id="EF904B35FF82693B36292AD9FB8EFE39" author="Wheeler &amp; Wheeler" box="[814,1125,475,501]" firstAuthor="Wheeler" pageId="9" pageNumber="86" pagination="1 - 108" refId="ref13844" refString="Wheeler, G. C. &amp; Wheeler, J. (1976) Ant larvae: review and synthesis. Memoirs of the Entomological Society of Washington, 7, 1-108." type="journal article" year="1976">Wheeler &amp; Wheeler (1976)</bibRefCitation>
, in profile somites up to AI or AII thin, abruptly thickened posteriad AIII, also strongly curving at this point. Anus ventral (
<figureCitation id="133A2A41FF82693B31D72AFDFAC9FDD5" box="[1232,1314,511,537]" captionStart="FIGURE 6" captionStartId="9.[151,250,1530,1554]" captionTargetBox="[151,1435,741,1506]" captionTargetId="figure-274@9.[151,1435,741,1506]" captionTargetPageId="9" captionText="FIGURE 6. Fulakora estevae sp. nov., larva. A, head in profile; B. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="9" pageNumber="86">Fig.6B</figureCitation>
). Spiracles not visualized.
<specimenCount id="9D07FD4DFF82693B34472921FE28FDF1" box="[320,451,547,573]" count="3" pageId="9" pageNumber="86" type="generic" typeStatus="types">Three types</specimenCount>
of seta on body: (i) minute, stiff, slightly thickened, simple, present in all somites, more abundant on dorsum; (ii) simple, slender, flexible, slightly angled setae, about three times the size of
<typeStatus id="54BA8866FF82693B31FC2945FAC0FDAD" box="[1275,1323,583,609]" pageId="9" pageNumber="86">type</typeStatus>
i, densely present in all somites, including some sparse setae on the head; (iii) same morphology of
<typeStatus id="54BA8866FF82693B31992969FB25FD49" box="[1182,1230,619,645]" pageId="9" pageNumber="86">type</typeStatus>
ii, but longer and sparsely distributed dorsally on the thoracic somites and ventrally on all somites (
<figureCitation id="133A2A41FF82693B312E298DFB83FD65" box="[1065,1128,655,681]" captionStart="FIGURE 6" captionStartId="9.[151,250,1530,1554]" captionTargetBox="[151,1435,741,1506]" captionTargetId="figure-274@9.[151,1435,741,1506]" captionTargetPageId="9" captionText="FIGURE 6. Fulakora estevae sp. nov., larva. A, head in profile; B. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations: ant, antenna; ga, galea; hi, hair type i; hii, hair type ii; hiii, hair type iii; lb, labrum; md, mandible." figureDoi="http://doi.org/10.5281/zenodo.14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="9" pageNumber="86">Fig.6</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C4FF82693B35C029B0FC32FD01" blockId="9.[151,1437,151,717]" box="[199,985,690,717]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C029B0FE5FFD00" bold="true" box="[199,436,690,716]" pageId="9" pageNumber="86">Examined material.</emphasis>
<specimenCount id="9D07FD4DFF82693B34BC29B1FDC3FD01" box="[443,552,691,717]" count="1" pageId="9" pageNumber="86" type="larva">One larva</specimenCount>
from the
<typeStatus id="54BA8866FF82693B379129B1FD2DFD01" box="[662,710,691,717]" pageId="9" pageNumber="86">type</typeStatus>
series' colony fragment.
</paragraph>
<caption id="DF7E664CFF82693B35902EF8FACAF9FA" ID-DOI="http://doi.org/10.5281/zenodo.14896080" ID-Zenodo-Dep="14896080" httpUri="https://zenodo.org/record/14896080/files/figure.png" pageId="9" pageNumber="86" startId="9.[151,250,1530,1554]" targetBox="[151,1435,741,1506]" targetPageId="9" targetType="figure">
<paragraph id="8BBE36C4FF82693B35902EF8FACAF9FA" blockId="9.[151,1436,1530,1590]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35902EF8FEFAF9DE" bold="true" box="[151,273,1530,1554]" pageId="9" pageNumber="86">FIGURE 6</emphasis>
.
<taxonomicName id="4C014D47FF82693B34252EF8FE3DF9DE" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[290,470,1530,1554]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="estevae" status="sp. nov.">
<emphasis id="B975EAD6FF82693B34252EF8FE3DF9DE" box="[290,470,1530,1554]" italics="true" pageId="9" pageNumber="86">Fulakora estevae</emphasis>
</taxonomicName>
<emphasis id="B975EAD6FF82693B34E72EF8FDDFF9DE" bold="true" box="[480,564,1530,1554]" pageId="9" pageNumber="86">
<taxonomicNameLabel id="A24657ADFF82693B34E72EF8FDDFF9DE" box="[480,564,1530,1554]" pageId="9" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, larva.
<emphasis id="B975EAD6FF82693B378E2EF8FD70F9DE" bold="true" box="[649,667,1530,1554]" pageId="9" pageNumber="86">A</emphasis>
, head in profile;
<emphasis id="B975EAD6FF82693B365D2EF8FC80F9DE" bold="true" box="[858,875,1530,1554]" pageId="9" pageNumber="86">B</emphasis>
. profile view. Scale bars: 0.1 mm (A), 0.5 mm (B). Abbreviations:
<emphasis id="B975EAD6FF82693B34332D1CFEBCF9FA" bold="true" box="[308,343,1566,1590]" pageId="9" pageNumber="86">ant</emphasis>
, antenna;
<emphasis id="B975EAD6FF82693B34C72D1CFE32F9FA" bold="true" box="[448,473,1566,1590]" pageId="9" pageNumber="86">ga</emphasis>
, galea;
<emphasis id="B975EAD6FF82693B372E2D1CFDD5F9FA" bold="true" box="[553,574,1566,1590]" pageId="9" pageNumber="86">hi</emphasis>
, hair type i;
<emphasis id="B975EAD6FF82693B37B82D1CFD30F9FA" bold="true" box="[703,731,1566,1590]" pageId="9" pageNumber="86">hii</emphasis>
, hair type ii;
<emphasis id="B975EAD6FF82693B36642D1CFC6DF9FA" bold="true" box="[867,902,1566,1590]" pageId="9" pageNumber="86">hiii</emphasis>
, hair type iii;
<emphasis id="B975EAD6FF82693B31122D1CFBC1F9FA" bold="true" box="[1045,1066,1566,1590]" pageId="9" pageNumber="86">lb</emphasis>
, labrum;
<emphasis id="B975EAD6FF82693B318B2D1CFB44F9FA" bold="true" box="[1164,1199,1566,1590]" pageId="9" pageNumber="86">md</emphasis>
, mandible.
</paragraph>
</caption>
<paragraph id="8BBE36C4FF82693B35C02D8AFE0AF96E" blockId="9.[151,1437,1672,2023]" box="[199,481,1672,1699]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C02D8AFEB6F96E" bold="true" box="[199,349,1672,1698]" pageId="9" pageNumber="86">Queen, male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BBE36C4FF82693B35C02DAEFB28F90A" blockId="9.[151,1437,1672,2023]" box="[199,1219,1708,1735]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C02DAEFEA0F90A" bold="true" box="[199,331,1708,1734]" pageId="9" pageNumber="86">Karyology.</emphasis>
2n = 24 (16 metacentrics + 8 submetacentrics) (see session 3.2 for discussion).
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF82693B35C02DD2FAF0F89A" pageId="9" pageNumber="86" type="etymology">
<paragraph id="8BBE36C4FF82693B35C02DD2FAF0F89A" blockId="9.[151,1437,1672,2023]" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C02DD2FEA5F926" bold="true" box="[199,334,1744,1770]" pageId="9" pageNumber="86">Etymology.</emphasis>
The species is named in honor of the Brazilian myrmecologist Flávia de Araújo Esteves, an amblyoponine specialist. The name is formed directly from a modern personal name “Esteves”. It was formed by adding -ae to the stem of her last name, “Estev”, as it reffers to a woman, according to the ICZN code's article 31.1.2. Eponym orthography is unchangeable and does not depend on the generic name for which it is applied.
</paragraph>
</subSubSection>
<subSubSection id="C31B654FFF82693835C02C62FDEFFE1D" lastPageId="10" lastPageNumber="87" pageId="9" pageNumber="86" type="discussion">
<paragraph id="8BBE36C4FF82693835C02C62FAC4FEA9" blockId="9.[151,1437,1672,2023]" lastBlockId="10.[151,1437,151,465]" lastPageId="10" lastPageNumber="87" pageId="9" pageNumber="86">
<emphasis id="B975EAD6FF82693B35C02C62FEA4F8B6" bold="true" box="[199,335,1888,1914]" pageId="9" pageNumber="86">Comments.</emphasis>
In comparison to other species of the genus,
<taxonomicName id="4C014D47FF82693B36532C63FC28F8B6" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[852,963,1889,1914]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF82693B36532C63FC28F8B6" box="[852,963,1889,1914]" italics="true" pageId="9" pageNumber="86">F. estevae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C014D47FF82693B319F2C63FAEAF8B6" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1176,1281,1889,1914]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF82693B319F2C63FAEAF8B6" box="[1176,1281,1889,1914]" italics="true" pageId="9" pageNumber="86">F. agostii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014D47FF82693B30172C63FA7EF8B6" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[1296,1429,1889,1914]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="B975EAD6FF82693B30172C63FA7EF8B6" box="[1296,1429,1889,1914]" italics="true" pageId="9" pageNumber="86">F. armigera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014D47FF82693B35902C87FF05F852" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[151,238,1924,1950]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF82693B35902C87FF05F852" box="[151,238,1924,1950]" italics="true" pageId="9" pageNumber="86">F. cleae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014D47FF82693B35FA2C87FE90F852" box="[253,379,1924,1950]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF82693B35FA2C87FE90F852" box="[253,379,1924,1950]" italics="true" pageId="9" pageNumber="86">F. elongata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014D47FF82693B348D2C87FDE1F852" baseAuthorityName="Lattke" baseAuthorityYear="1991" box="[394,522,1924,1950]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="lurilabes">
<emphasis id="B975EAD6FF82693B348D2C87FDE1F852" box="[394,522,1924,1950]" italics="true" pageId="9" pageNumber="86">F. lurilabes</emphasis>
</taxonomicName>
, and to certain extent, even to the strange
<taxonomicName id="4C014D47FF82693B36ED2C87FB8EF852" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1002,1125,1924,1950]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="heraldoi">
<emphasis id="B975EAD6FF82693B36ED2C87FB8EF852" box="[1002,1125,1924,1950]" italics="true" pageId="9" pageNumber="86">F. heraldoi</emphasis>
</taxonomicName>
, as all share a mostly darkcolored body, clypeal chaetae arising from distinct clypeal tubercles (or platform), a relatively conserved mandibular dentition pattern, and poorly developed subpetiolar anterior process. Despite the odd appearance of
<taxonomicName id="4C014D47FF82693B31E22CCFFAB5F82A" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1253,1374,1996,2022]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="heraldoi">
<emphasis id="B975EAD6FF82693B31E22CCFFAB5F82A" box="[1253,1374,1996,2022]" italics="true" pageId="9" pageNumber="86">F. heraldoi</emphasis>
</taxonomicName>
, with crescent-shaped mandibles, tubercles on clypeus fused into a platform, and absence of genal tooth, it is similar to these species in the other respects.
<taxonomicName id="4C014D47FF816938372A2BB9FD1BFF18" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[557,752,187,213]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF816938372A2BB9FD1BFF18" box="[557,752,187,213]" italics="true" pageId="10" pageNumber="87">Fulakora estevae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C014D47FF816938368B2BB9FBE7FF19" box="[908,1036,187,213]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B975EAD6FF816938368B2BB9FBE7FF19" box="[908,1036,187,213]" italics="true" pageId="10" pageNumber="87">F. elongata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C014D47FF81693831402BB9FB22FF19" baseAuthorityName="Lattke" baseAuthorityYear="1991" box="[1095,1225,187,213]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="lurilabes">
<emphasis id="B975EAD6FF81693831402BB9FB22FF19" box="[1095,1225,187,213]" italics="true" pageId="10" pageNumber="87">F. lurilabes</emphasis>
</taxonomicName>
by the absence of propodeal pale patches, absence of mesometanotal suture, and, from the latter, by the smooth rather than opaque vertex. The genal tooth in
<taxonomicName id="4C014D47FF81693834C22A01FDDFFED0" authorityName="Jacintho &amp; Chaul &amp; Teixeira &amp; Barros &amp; Lopes &amp; Serrão" authorityYear="2025" box="[453,564,259,284]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="estevae">
<emphasis id="B975EAD6FF81693834C22A01FDDFFED0" box="[453,564,259,284]" italics="true" pageId="10" pageNumber="87">F. estevae</emphasis>
</taxonomicName>
is much smaller than that of
<taxonomicName id="4C014D47FF81693836842A01FC07FED0" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[899,1004,259,284]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="agostii">
<emphasis id="B975EAD6FF81693836842A01FC07FED0" box="[899,1004,259,284]" italics="true" pageId="10" pageNumber="87">F. agostii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C014D47FF81693836FB2A01FB6BFED0" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[1020,1152,259,284]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="B975EAD6FF81693836FB2A01FB6BFED0" box="[1020,1152,259,284]" italics="true" pageId="10" pageNumber="87">F. armigera</emphasis>
</taxonomicName>
, and
<emphasis id="B975EAD6FF81693831C62A01FACBFED1" box="[1217,1312,259,285]" italics="true" pageId="10" pageNumber="87">
<taxonomicName id="4C014D47FF81693831C62A01FAF6FED1" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[1217,1309,259,285]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="cleae">F. cleae</taxonomicName>
.
</emphasis>
Moreover, it differs from
<taxonomicName id="4C014D47FF81693834472A25FE73FE8D" baseAuthorityName="Lacau &amp; Delabie" baseAuthorityYear="2002" box="[320,408,295,321]" class="Insecta" family="Formicidae" genus="Fulakora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="cleae">
<emphasis id="B975EAD6FF81693834472A25FE73FE8D" box="[320,408,295,321]" italics="true" pageId="10" pageNumber="87">F. cleae</emphasis>
</taxonomicName>
by the subrectangular head, rather than the subquadrate head. A comprehensive list of the differences and similarities between these species is presented in section 3.1.3 (see
<tableCitation id="C683037FFF816938313C2A49FB64FEA9" box="[1083,1167,331,357]" captionStart="TABLE 1" captionStartId="13.[152,240,150,174]" captionTargetPageId="13" captionText="TABLE 1. Comparison of characters, measurements and indexes among Fulakora species (see text for the description of each character state). Specimens studied: four F. estevae (ANTWEB1047071, ANTWEB1047070, ANTWEB1047069, UFV-LABECOL-000012), three F. agostii (CASENT0102201, CASENT0101451, ANTWEB1047718), one F. armigera (CASENT0915647),twoF.cleae(CASENT0102202,CASENT0101452),tenF.elongata(CASENT0915100,CASENT0915098, CASENT0102471, CASENT0178869, UFV-LABECOL-000027, CASENT0915101, UFV-LABECOL-000049, UFV- LABECOL-008530, ANTWEB1047734, ANTWEB1047061); three F. lurilabes (CASENT0102199, CASENT0172810, CASENT0172811), and one F. heraldoi (INPA-HYM-034964) (For data per specimen see Appendix 2)." pageId="10" pageNumber="87">Table 1</tableCitation>
;
<figureCitation id="133A2A41FF816938319C2A49FAC9FEA9" box="[1179,1314,331,357]" captionStart="APPENDIX 2" captionStartId="23.[152,304,978,1004]" captionTargetPageId="23" captionText="APPENDIX 2. Comparison of characters, measurements and indexes among Fulakora species. Description of each measurement is on the section 2 (Measurements and indices), characters states is on section 3.1.3." pageId="10" pageNumber="87">Appendix 2</figureCitation>
).
</paragraph>
<paragraph id="8BBE36C4FF81693835C02A6DFDEFFE1D" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="87">
Out of the
<specimenCount id="9D07FD4DFF81693834432A6DFE29FE45" box="[324,450,367,393]" count="10" pageId="10" pageNumber="87" type="worker">10 workers</specimenCount>
originally found in the colony fragment, only three became part of the
<typeStatus id="54BA8866FF81693831F42A6DFAC8FE45" box="[1267,1315,367,393]" pageId="10" pageNumber="87">type</typeStatus>
series due to an accident during the period the colony was maintained for karyological studies, where
<specimenCount id="9D07FD4DFF816938319C2A91FAD7FE61" box="[1179,1340,403,429]" count="7" pageId="10" pageNumber="87" type="worker">seven workers</specimenCount>
escaped from the lab colony and got lost.
</paragraph>
</subSubSection>
</treatment>
</document>