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<document id="B803100FC8D5A44995E0513F295949E7" ID-DOI="10.11646/zootaxa.5005.3.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="5141647" ID-ZooBank="2D478F03-C124-436C-BE5D-1833B6BE21B3" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1627469187300" checkinUser="plazi" docAuthor="Müller, Jörg &amp; Schuh, Rudolf" docDate="2021" docId="AA3F01335A1BFFED94AB5EB73CE91DC1" docLanguage="en" docName="zootaxa.5005.3.12.pdf" docOrigin="Zootaxa 5005 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tarphius hyrcanicus Müller &amp; Schuh 2021, n. sp." docType="treatment" docVersion="1" lastPageNumber="378" masterDocId="5606794B5A1AFFEE943C5F1B39791A7A" masterDocTitle="A new species of Tarphius Erichson, 1845 (Coleoptera: Zopheridae) from Iran" masterLastPageNumber="380" masterPageNumber="375" pageNumber="376" updateTime="1732216865730" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="7A84577B9FB90A3D5199C1ECC896DDF7">A new species of Tarphius Erichson, 1845 (Coleoptera: Zopheridae) from Iran</mods:title>
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<mods:namePart id="808B1A6BAA0356FF3721A5D3E2142B4E">Müller, Jörg</mods:namePart>
<mods:affiliation id="BFF1951BF7AF91B3D853A07ECA12B5DB">Field Station Fabrikschleichach, Biocenter, University of Würzburg, Glashüttenstr. 5, 96181 Rauhenebrach, Germany; &amp; Bavarian Forest National Park, Freyunger Str. 2, 94481 Grafenau, Germany.</mods:affiliation>
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<mods:namePart id="18A9DAB5AAE89D345381335354B5CA6D">Schuh, Rudolf</mods:namePart>
<mods:affiliation id="71983AD09AE31C43B8D4FAD6A11FBD7A">Naturhistorisches Museum Wien, Zweite Zoologische Abteilung, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
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<mods:title id="BF8360A08AB0E69F50DBEF5D3AFA139E">Zootaxa</mods:title>
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<mods:date id="7CB920E273DC3C445A321204EEC594BE">2021</mods:date>
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<taxonomicName id="E596CBA65A1BFFEF94AB5EB738FF1BBC" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[151,390,428,454]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A1BFFEF94AB5EB738FF1BBC" box="[151,390,428,454]" italics="true" pageId="1" pageNumber="376">Tarphius hyrcanicus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="0BD1D14C5A1BFFEF95B15EB738B61BBC" box="[397,463,428,454]" pageId="1" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<paragraph id="2229B0255A1BFFEF94AB5ECB386B1B90" blockId="1.[151,463,428,490]" box="[151,274,464,490]" pageId="1" pageNumber="376">
(
<figureCitation id="BAADACA05A1BFFEF94A35ECB38701B90" box="[159,265,464,490]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,260,1883,1909]" captionStartId-1="2.[151,260,1495,1521]" captionStartId-2="3.[151,260,1412,1438]" captionTargetBox-0="[571,1009,995,1807]" captionTargetBox-1="[590,996,674,1466]" captionTargetBox-2="[240,1350,498,1380]" captionTargetId-0="figure-190@1.[534,1054,944,1859]" captionTargetId-1="figure-230@2.[583,1005,666,1471]" captionTargetId-2="figure-164@3.[230,1358,486,1388]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Dorsal aspect of Tarphius hyrcanicus n. sp., male holotype." captionText-1="FIGURE 2. Shape of T. hyrcanicus n. sp., dark gray patches mark the positions of gibbosities on the elytra." captionText-2="FIGURE 3. Genitalia of T. hyrcanicus n. sp. (a) male and (b, c) female." figureDoi-0="http://doi.org/10.5281/zenodo.5141649" figureDoi-1="http://doi.org/10.5281/zenodo.5141651" figureDoi-2="http://doi.org/10.5281/zenodo.5141653" httpUri-0="https://zenodo.org/record/5141649/files/figure.png" httpUri-1="https://zenodo.org/record/5141651/files/figure.png" httpUri-2="https://zenodo.org/record/5141653/files/figure.png" pageId="1" pageNumber="376">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6A8CE3AE5A1BFFEF94AB5D033B3E189C" pageId="1" pageNumber="376" type="materials_examined">
<paragraph id="2229B0255A1BFFEF94AB5D033B3E189C" blockId="1.[151,1437,536,922]" pageId="1" pageNumber="376">
<emphasis id="10E26C375A1BFFEF94AB5D03383D1848" bold="true" box="[151,324,536,562]" pageId="1" pageNumber="376">Type material:</emphasis>
<materialsCitation id="92FEBA785A1BFFEF957B5D033D5C182C" collectingDate="2017-10-05" collectorName="Jorg Muller" country="Iran" elevation="1083" latitude="37.6717" location="Gilan" longLatPrecision="1" longitude="48.75261" municipality="Shafarud" pageId="1" pageNumber="376" specimenCount="1" stateProvince="Gilan" typeStatus="holotype">
<typeStatus id="FD2D0E875A1BFFEF957B5D0338A71848" box="[327,478,536,562]" pageId="1" pageNumber="376" type="holotype">HOLOTYPE</typeStatus>
, male, deposited at CMR, labeled:
<emphasis id="10E26C375A1BFFEF976E5D033A1B1848" box="[850,866,536,562]" italics="true" pageId="1" pageNumber="376"></emphasis>
IS5_
<collectingCountry id="5A81F0B55A1BFFEF97A85D033ABA1849" box="[916,963,536,563]" name="Iran" pageId="1" pageNumber="376">Iran</collectingCountry>
Shafarud,
<collectingRegion id="E0527EC75A1BFFEF900B5D033D0C1848" box="[1079,1141,536,562]" pageId="1" pageNumber="376">Gilan</collectingRegion>
Prov.Orientbuchenurwald beweidet,
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<elevation id="A9BB57165A1BFFEF95325D273811182D" box="[270,360,572,599]" metricMagnitude="3" metricUnit="m" metricValue="1.083" pageId="1" pageNumber="376" unit="m" value="1083.0">1083 m</elevation>
</quantity>
,
<geoCoordinate id="47A2D6E25A1BFFEF95445D27389D182C" box="[376,484,572,598]" degrees="37.67170" direction="north" orientation="latitude" pageId="1" pageNumber="376" precision="1" value="37.6717">37.67170</geoCoordinate>
,
<geoCoordinate id="47A2D6E25A1BFFEF95CE5D273B27182D" box="[498,606,572,599]" degrees="48.75261" direction="east" orientation="longitude" pageId="1" pageNumber="376" precision="1" value="48.75261">48.75261</geoCoordinate>
, Gesiebe,
<date id="562896E55A1BFFEF96E55D273A35182D" box="[729,844,572,599]" pageId="1" pageNumber="376" value="2017-10-05">
<collectingDate id="466C6F0D5A1BFFEF96E55D273A35182D" box="[729,844,572,599]" pageId="1" pageNumber="376" value="2017-10-05">10.5.2017</collectingDate>
</date>
, leg.
<collectorName id="8F63D5F35A1BFFEF97B25D263D62182C" box="[910,1051,572,599]" pageId="1" pageNumber="376">Jörg Müller</collectorName>
</materialsCitation>
.
<materialsCitation id="92FEBA785A1BFFEF900F5D273A1A18E4" collectingDate="2008-05-30" collectionCode="CPE" collectorName="A. Putz." country="Iran" county="Sari County" location="Elburz Mt." municipality="Sari County Mohammadabad" pageId="1" pageNumber="376" specimenCount="1" specimenCount-female="1" stateProvince="Mazanderan" typeStatus="paratype">
<typeStatus id="FD2D0E875A1BFFEF900F5D273DA8182C" box="[1075,1233,572,598]" pageId="1" pageNumber="376" type="paratype">PARATYPES</typeStatus>
,
<specimenCount id="34907BAC5A1BFFEF90E35D273C3D182C" box="[1247,1348,572,598]" count="1" pageId="1" pageNumber="376" type="female">1 female</specimenCount>
(
<collectionCode id="448728E05A1BFFEF916B5D273CE8182C" box="[1367,1425,572,598]" collectionName="CPE" pageId="1" pageNumber="376">CPE</collectionCode>
):
<emphasis id="10E26C375A1BFFEF94AB5D7B39DE1800" box="[151,167,608,634]" italics="true" pageId="1" pageNumber="376"></emphasis>
<collectingCountry id="5A81F0B55A1BFFEF949B5D7A39AE1801" box="[167,215,609,635]" name="Iran" pageId="1" pageNumber="376">Iran</collectingCountry>
, Prov.
<collectingRegion id="E0527EC75A1BFFEF95145D7B38CB1800" box="[296,434,608,634]" country="Iran" name="Mazandaran" pageId="1" pageNumber="376">Mazanderan</collectingRegion>
IR08-06
<collectingMunicipality id="C24D2A5F5A1BFFEF961F5D7B3A0C1800" box="[547,885,608,634]" pageId="1" pageNumber="376">Sari County Mohammadabad</collectingMunicipality>
,
<location id="2749E6FE5A1BFFEF97BE5D7B3D781801" LSID="urn:lsid:plazi:treatment:AA3F01335A1BFFED94AB5EB73CE91DC1:2749E6FE5A1BFFEF97BE5D7B3D781801" box="[898,1025,608,635]" country="Iran" county="Sari County" municipality="Sari County Mohammadabad" name="Elburz Mt." pageId="1" pageNumber="376" stateProvince="Mazanderan">Elburz Mt.</location>
, N-Slope,
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<quantity id="E56E1DC05A1BFFEF90415D7B3DC11800" box="[1149,1208,608,634]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="376" unit="km" value="2.0">2 km</quantity>
SW Majid
</locationDeviation>
, 36° 09 28.453° 12 88.9 872 small stream,
<date id="562896E55A1BFFEF96755D9F3BC518E5" box="[585,700,644,671]" pageId="1" pageNumber="376" value="2008-05-30">
<collectingDate id="466C6F0D5A1BFFEF96755D9F3BC518E5" box="[585,700,644,671]" pageId="1" pageNumber="376" value="2008-05-30">30.5.2008</collectingDate>
</date>
, leg.
<collectorName id="8F63D5F35A1BFFEF96C45D9F3A2D18E5" box="[760,852,644,671]" pageId="1" pageNumber="376">A. Pütz.</collectorName>
<emphasis id="10E26C375A1BFFEF976F5D9F3A1A18E4" box="[851,867,644,670]" italics="true" pageId="1" pageNumber="376"></emphasis>
</materialsCitation>
;
<materialsCitation id="92FEBA785A1BFFEF974E5D9F3B3E189C" collectingDate="2008-05-31" collectionCode="CPE, CSW" collectorName="A. Putz." country="Iran" county="Babol County" location="Elburz Mt" municipality="Babol County Mohammadabad" pageId="1" pageNumber="376" specimenCount="2" specimenCount-female="2" stateProvince="Mazanderan" typeStatus="paratype">
<specimenCount id="34907BAC5A1BFFEF974E5D9F3AA618E4" box="[882,991,644,670]" count="2" pageId="1" pageNumber="376" type="female">2 females</specimenCount>
(
<collectionCode id="448728E05A1BFFEF97D35D9F3D5E18E4" box="[1007,1063,644,670]" collectionName="CPE" pageId="1" pageNumber="376">CPE</collectionCode>
,
<collectionCode id="448728E05A1BFFEF900E5D9F3D0D18E4" box="[1074,1140,644,670]" collectionName="CSW" pageId="1" pageNumber="376">CSW</collectionCode>
): “
<collectingCountry id="5A81F0B55A1BFFEF90AE5D9F3DBD18E4" box="[1170,1220,644,670]" name="Iran" pageId="1" pageNumber="376">Iran</collectingCountry>
, Prov.
<collectingRegion id="E0527EC75A1BFFEF912E5D9F3CE518E4" box="[1298,1436,644,670]" country="Iran" name="Mazandaran" pageId="1" pageNumber="376">Mazanderan</collectingRegion>
IR08-06
<collectingMunicipality id="C24D2A5F5A1BFFEF94C75DB33B2518B8" box="[251,604,680,706]" pageId="1" pageNumber="376">Babol County Mohammadabad</collectingMunicipality>
,
<location id="2749E6FE5A1BFFEF965A5DB33BA418B9" LSID="urn:lsid:plazi:treatment:AA3F01335A1BFFED94AB5EB73CE91DC1:2749E6FE5A1BFFEF965A5DB33BA418B9" box="[614,733,680,707]" country="Iran" county="Babol County" municipality="Babol County Mohammadabad" name="Elburz Mt" pageId="1" pageNumber="376" stateProvince="Mazanderan">Elburz Mt</location>
., N-Slope,
<locationDeviation id="339BA9075A1BFFEF976B5DB33D5718B8" box="[855,1070,680,706]" location="Firuz Jah" pageId="1" pageNumber="376">
<quantity id="E56E1DC05A1BFFEF976B5DB33AF618B8" box="[855,911,680,706]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="376" unit="km" value="2.0">2 km</quantity>
SW Firuz Jah
</locationDeviation>
, 36° 66 10.0 52° 38 90.1 839 small stream,
<date id="562896E55A1BFFEF95085DD738DE189D" box="[308,423,716,743]" pageId="1" pageNumber="376" value="2008-05-31">
<collectingDate id="466C6F0D5A1BFFEF95085DD738DE189D" box="[308,423,716,743]" pageId="1" pageNumber="376" value="2008-05-31">31.5.2008</collectingDate>
</date>
, leg.
<collectorName id="8F63D5F35A1BFFEF95DC5DD73B47189C" box="[480,574,716,742]" pageId="1" pageNumber="376">A. Pütz.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6A8CE3AE5A1BFFEF94FB5DEB38591954" pageId="1" pageNumber="376" type="etymology">
<paragraph id="2229B0255A1BFFEF94FB5DEB38591954" blockId="1.[151,1437,536,922]" pageId="1" pageNumber="376">
<emphasis id="10E26C375A1BFFEF94FB5DEB38311970" bold="true" box="[199,328,752,778]" pageId="1" pageNumber="376">Etymology</emphasis>
. The species is named after its
<typeStatus id="FD2D0E875A1BFFEF96D15DEA3A641971" box="[749,797,753,779]" pageId="1" pageNumber="376">type</typeStatus>
locality, the Hyrcanian Forest, which means “wolf mountains.”
</paragraph>
</subSubSection>
<subSubSection id="6A8CE3AE5A1BFFEC94FB5C233D2B1CF8" lastPageId="2" lastPageNumber="377" pageId="1" pageNumber="376" type="description">
<paragraph id="2229B0255A1BFFEF94FB5C23382119E0" blockId="1.[151,1437,536,922]" pageId="1" pageNumber="376">
<emphasis id="10E26C375A1BFFEF94FB5C233B431928" bold="true" box="[199,570,824,850]" pageId="1" pageNumber="376">
Description (
<specimenCount id="34907BAC5A1BFFEF955D5C2338C91928" box="[353,432,824,850]" pageId="1" pageNumber="376" type="male">1 male</specimenCount>
,
<specimenCount id="34907BAC5A1BFFEF95805C233B571928" box="[444,558,824,850]" pageId="1" pageNumber="376" type="female">3 females</specimenCount>
).
</emphasis>
TL: 3.34.0 mm (male: 3.6, females: 3.3, 3.9, 4.0). Medium size, dark brown, parts of head, legs, and antenna more red brown, body convex and ovate, setose; dorsal surface bearing large granules (
<figureCitation id="BAADACA05A1BFFEF95395C9B383E19E0" box="[261,327,896,922]" captionStart="FIGURE 1" captionStartId="1.[151,260,1883,1909]" captionTargetBox="[571,1009,995,1807]" captionTargetId="figure-190@1.[534,1054,944,1859]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal aspect of Tarphius hyrcanicus n. sp., male holotype." figureDoi="http://doi.org/10.5281/zenodo.5141649" httpUri="https://zenodo.org/record/5141649/files/figure.png" pageId="1" pageNumber="376">Fig. 1</figureCitation>
).
</paragraph>
<caption id="76E9E0AD5A1BFFEF94AB58403ABA1D0F" ID-DOI="http://doi.org/10.5281/zenodo.5141649" ID-Zenodo-Dep="5141649" box="[151,963,1883,1910]" httpUri="https://zenodo.org/record/5141649/files/figure.png" pageId="1" pageNumber="376" startId="1.[151,260,1883,1909]" targetBox="[571,1009,995,1807]" targetPageId="1">
<paragraph id="2229B0255A1BFFEF94AB58403ABA1D0F" blockId="1.[151,963,1883,1910]" box="[151,963,1883,1910]" pageId="1" pageNumber="376">
<emphasis id="10E26C375A1BFFEF94AB584038581D0F" bold="true" box="[151,289,1883,1909]" pageId="1" pageNumber="376">FIGURE 1.</emphasis>
Dorsal aspect of
<taxonomicName id="E596CBA65A1BFFEF95D858403BB11D0F" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[484,712,1883,1909]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="376" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A1BFFEF95D858403BB11D0F" box="[484,712,1883,1909]" italics="true" pageId="1" pageNumber="376">Tarphius hyrcanicus</emphasis>
</taxonomicName>
<emphasis id="10E26C375A1BFFEF96F258403A741D0F" bold="true" box="[718,781,1883,1909]" pageId="1" pageNumber="376">
<taxonomicNameLabel id="0BD1D14C5A1BFFEF96F258403A741D0F" box="[718,781,1883,1909]" pageId="1" pageNumber="376" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype.
</paragraph>
</caption>
<paragraph id="2229B0255A1BFFEC94FB58B83D031AAF" blockId="1.[151,1436,1955,2018]" lastBlockId="2.[151,1437,150,645]" lastPageId="2" lastPageNumber="377" pageId="1" pageNumber="376">
<emphasis id="10E26C375A1BFFEF94FB58B8387E1DC7" bold="true" box="[199,263,1955,1981]" pageId="1" pageNumber="376">Head</emphasis>
: frons densely granulose, granules large, separated by less than their diameters; some granules bearing a yellow, erect seta; clypeus triangular, glabrous with erect yellow setae; eyes small, round, glabrous; antenna with antennomere 2 large, globular, antennomeres 3,4 and 5 longer than wide, antennomeres 6 and 7 as long as wide, antennomeres 8 and 9 wider than long, antennomeres 10 and 11 forming a compact club.
</paragraph>
<paragraph id="2229B0255A18FFEC94FB5FC438741BAB" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB5FC438301A83" bold="true" box="[199,329,223,249]" pageId="2" pageNumber="377">Pronotum:</emphasis>
transverse (PW/PL: 1.55), maximum width in the middle; sculpture on pronotal disc consisting of large and obtuse granules larger than those on the head and on the lateral parts of the pronotum; each granule bearing a semi-erect yellow seta; median sulcus is free of granules and chagrinated; anterior margin strongly sinuate laterally, anterior angles prominent, acute; lateral margins arcuate, finely serrate with 1718 small black denticles and fringed with three irregular rows of elongate, erect setae, directed backwards in the apical half, forwards in the posterior half; posterior angles rectangular, basal margin convex, sinuate laterally; sub-basal sulcus slightly depressed.
</paragraph>
<paragraph id="2229B0255A18FFEC94FB5EC03DDD1847" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB5EC038F11B8F" bold="true" box="[199,392,475,501]" pageId="2" pageNumber="377">Elytra together:</emphasis>
EL/EW: 1.07, as wide as the pronotum; surface very rugose, with deep pits; anterior angles pointedly raised; each elytron with 5 gibbosities arranged as shown in
<figureCitation id="BAADACA05A18FFEC97955EE43A941863" box="[937,1005,511,537]" captionStart="FIGURE 2" captionStartId="2.[151,260,1495,1521]" captionTargetBox="[590,996,674,1466]" captionTargetId="figure-230@2.[583,1005,666,1471]" captionTargetPageId="2" captionText="FIGURE 2. Shape of T. hyrcanicus n. sp., dark gray patches mark the positions of gibbosities on the elytra." figureDoi="http://doi.org/10.5281/zenodo.5141651" httpUri="https://zenodo.org/record/5141651/files/figure.png" pageId="2" pageNumber="377">Fig. 2</figureCitation>
; sculpture consisting of granules, each bearing an erect yellow seta; gibbosities bearing 49 seta; apical elytral margin semicircular.
</paragraph>
<paragraph id="2229B0255A18FFEC94FB5D5C384718FF" blockId="2.[151,1437,150,645]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB5D5C3871181B" bold="true" box="[199,264,583,609]" pageId="2" pageNumber="377">Legs:</emphasis>
femora and tibiae with distinct protruding yellow setae; tarsal segment 4 slightly longer than segments 13 combined.
</paragraph>
<caption id="76E9E0AD5A18FFEC94AB5ACC3C361F88" ID-DOI="http://doi.org/10.5281/zenodo.5141651" ID-Zenodo-Dep="5141651" box="[151,1359,1495,1522]" httpUri="https://zenodo.org/record/5141651/files/figure.png" pageId="2" pageNumber="377" startId="2.[151,260,1495,1521]" targetBox="[590,996,674,1466]" targetPageId="2">
<paragraph id="2229B0255A18FFEC94AB5ACC3C361F88" blockId="2.[151,1359,1495,1522]" box="[151,1359,1495,1522]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94AB5ACC38591F88" bold="true" box="[151,288,1495,1522]" pageId="2" pageNumber="377">FIGURE 2.</emphasis>
Shape of
<taxonomicName id="E596CBA65A18FFEC95AD5AC23B5F1F88" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[401,550,1496,1522]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A18FFEC95AD5AC23B5F1F88" box="[401,550,1496,1522]" italics="true" pageId="2" pageNumber="377">T. hyrcanicus</emphasis>
</taxonomicName>
<emphasis id="10E26C375A18FFEC96115AC33B151F88" bold="true" box="[557,620,1496,1522]" pageId="2" pageNumber="377">
<taxonomicNameLabel id="0BD1D14C5A18FFEC96115AC33B151F88" box="[557,620,1496,1522]" pageId="2" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, dark gray patches mark the positions of gibbosities on the elytra.
</paragraph>
</caption>
<paragraph id="2229B0255A18FFEC94FB593B3C2B1C40" blockId="2.[151,1437,1567,2026]" box="[199,1362,1567,1594]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB593B380B1C43" bold="true" box="[199,370,1567,1594]" pageId="2" pageNumber="377">Male genitalia</emphasis>
(
<figureCitation id="BAADACA05A18FFEC95BD593B38AA1C40" box="[385,467,1568,1594]" captionStart="FIGURE 3" captionStartId="3.[151,260,1412,1438]" captionTargetBox="[240,1350,498,1380]" captionTargetId="figure-164@3.[230,1358,486,1388]" captionTargetPageId="3" captionText="FIGURE 3. Genitalia of T. hyrcanicus n. sp. (a) male and (b, c) female." figureDoi="http://doi.org/10.5281/zenodo.5141653" httpUri="https://zenodo.org/record/5141653/files/figure.png" pageId="2" pageNumber="377">Fig. 3a</figureCitation>
): tegmen elongate, parameres parallel-sided, slightly shorter than the basal part.
</paragraph>
<paragraph id="2229B0255A18FFEC94FB595F3D2B1CF8" blockId="2.[151,1437,1567,2026]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB595F38F31C27" bold="true" box="[199,394,1603,1630]" pageId="2" pageNumber="377">Female genitalia</emphasis>
(
<figureCitation id="BAADACA05A18FFEC95AA595F38811C25" box="[406,504,1604,1631]" captionStart="FIGURE 3" captionStartId="3.[151,260,1412,1438]" captionTargetBox="[240,1350,498,1380]" captionTargetId="figure-164@3.[230,1358,486,1388]" captionTargetPageId="3" captionText="FIGURE 3. Genitalia of T. hyrcanicus n. sp. (a) male and (b, c) female." figureDoi="http://doi.org/10.5281/zenodo.5141653" httpUri="https://zenodo.org/record/5141653/files/figure.png" pageId="2" pageNumber="377">Fig. 3b,c</figureCitation>
): gonostylus single-segmented, elongate, coxites without clear articulations between them (
<figureCitation id="BAADACA05A18FFEC94E35973384B1CF9" box="[223,306,1640,1667]" captionStart="FIGURE 3" captionStartId="3.[151,260,1412,1438]" captionTargetBox="[240,1350,498,1380]" captionTargetId="figure-164@3.[230,1358,486,1388]" captionTargetPageId="3" captionText="FIGURE 3. Genitalia of T. hyrcanicus n. sp. (a) male and (b, c) female." figureDoi="http://doi.org/10.5281/zenodo.5141653" httpUri="https://zenodo.org/record/5141653/files/figure.png" pageId="2" pageNumber="377">Fig. 3b</figureCitation>
); abdominal sternite VIII long-haired, apical margin indented (
<figureCitation id="BAADACA05A18FFEC97CF59733D3D1CF8" box="[1011,1092,1640,1666]" captionStart="FIGURE 3" captionStartId="3.[151,260,1412,1438]" captionTargetBox="[240,1350,498,1380]" captionTargetId="figure-164@3.[230,1358,486,1388]" captionTargetPageId="3" captionText="FIGURE 3. Genitalia of T. hyrcanicus n. sp. (a) male and (b, c) female." figureDoi="http://doi.org/10.5281/zenodo.5141653" httpUri="https://zenodo.org/record/5141653/files/figure.png" pageId="2" pageNumber="377">Fig. 3c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6A8CE3AE5A18FFED94FB59903AF81D08" lastPageId="3" lastPageNumber="378" pageId="2" pageNumber="377" type="diagnosis">
<paragraph id="2229B0255A18FFEC94FB59903C2A1D4C" blockId="2.[151,1437,1567,2026]" pageId="2" pageNumber="377">
<emphasis id="10E26C375A18FFEC94FB599038B41CDF" bold="true" box="[199,461,1675,1701]" pageId="2" pageNumber="377">Differential diagnosis.</emphasis>
The relationships of the species within the genus
<taxonomicName id="E596CBA65A18FFEC97CF59973D2F1CDC" authorityName="Erichson" authorityYear="1845" box="[1011,1110,1676,1702]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis id="10E26C375A18FFEC97CF59973D2F1CDC" box="[1011,1110,1676,1702]" italics="true" pageId="2" pageNumber="377">Tarphius</emphasis>
</taxonomicName>
have never been investigated and thus remain unclear.
<taxonomicName id="E596CBA65A18FFEC958D59AB3BEC1CB0" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[433,661,1712,1738]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A18FFEC958D59AB3BEC1CB0" box="[433,661,1712,1738]" italics="true" pageId="2" pageNumber="377">Tarphius hyrcanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BD1D14C5A18FFEC96A159AB3BA01CB0" box="[669,729,1712,1738]" pageId="2" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
shows no close relationship to any other species of that genus. In this diagnosis it is compared to species with similar habitus and the following set of external characters: pronotum ~1.5 times wider than long, lateral margin straight in the basal half, posterior angles sharp, rectangular; elytra together ~1.1 times longer than wide and with gibbosities, humeral angle sharp, rectangular, and laterally protruding.
</paragraph>
<paragraph id="2229B0255A18FFED94FB585B3B471BAB" blockId="2.[151,1437,1567,2026]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="378" pageId="2" pageNumber="377">
<taxonomicName id="E596CBA65A18FFEC94FB585B38D41D20" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[199,429,1856,1882]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A18FFEC94FB585B38D41D20" box="[199,429,1856,1882]" italics="true" pageId="2" pageNumber="377">Tarphius hyrcanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BD1D14C5A18FFEC9589585B38931D20" box="[437,490,1856,1882]" pageId="2" pageNumber="377" rank="species">n.sp.</taxonomicNameLabel>
differs from all continental species based on its larger size, the presence of elytral gibbosities, and its rectangular posterior pronotal angles. It has greater similarities to all the following species which are known.
<taxonomicName id="E596CBA65A18FFEC952758933BD41DD9" authority="Gillerfors, 1985" authorityName="Gillerfors" authorityYear="1985" box="[283,685,1928,1955]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="10E26C375A18FFEC95275893388B1DD8" box="[283,498,1928,1954]" italics="true" pageId="2" pageNumber="377">Tarphius depressus</emphasis>
Gillerfors, 1985
</taxonomicName>
and
<taxonomicName id="E596CBA65A18FFEC96DF58923D641DD9" authority="Gillerfors, 1985" authorityName="Gillerfors" authorityYear="1985" box="[739,1053,1928,1955]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="377" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="10E26C375A18FFEC96DF58923A261DD8" box="[739,863,1928,1954]" italics="true" pageId="2" pageNumber="377">T. tornvalli</emphasis>
Gillerfors, 1985
</taxonomicName>
, both of which are local endemics on particular islands in the Macaronesian Archipelago (S. Maria and S. Miguel, respectively, in the Azores) differs by the presence of 9 gibbosities on each elytron and an elytral apex distinctly produced backwards in the sutural region.
<taxonomicName id="E596CBA65A19FFED94D05F8C3BA11ACB" authority="Uyttenboogaart, 1937" authorityName="Uyttenboogaart" authorityYear="1937" box="[236,728,151,177]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="postcostatus">
<emphasis id="10E26C375A19FFED94D05F8C38A71ACA" box="[236,478,151,177]" italics="true" pageId="3" pageNumber="378">Tarphius postcostatus</emphasis>
Uyttenboogaart, 1937
</taxonomicName>
and
<taxonomicName id="E596CBA65A19FFED97305F8C3D411ACB" authority="Wollaston, 1862" authorityName="Wollaston" authorityYear="1862" box="[780,1080,151,177]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="10E26C375A19FFED97305F8C3A041ACB" box="[780,893,151,177]" italics="true" pageId="3" pageNumber="378">T. simplex</emphasis>
Wollaston, 1862
</taxonomicName>
(from Tenerife, Canary Islands) differ by the presence of 4 gibbosities on each elytron, of which the gibbosities in interval 3 near the base and apex are elongate, and the elytral apex is distinctly produced backwards in the sutural region.
<taxonomicName id="E596CBA65A19FFED909F5FC438661B67" authority="Franz, 1967" authorityName="Franz" authorityYear="1967" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="tacorontinus">
<emphasis id="10E26C375A19FFED909F5FC43CE41A82" box="[1187,1437,223,249]" italics="true" pageId="3" pageNumber="378">Tarphius tacorontinus</emphasis>
Franz, 1967
</taxonomicName>
(from Tenerife, Canary Islands) differs by the thick, almost swollen edge of its pronotal margin, the presence of 7 gibbosities on each elytron, and an elytral apex slightly produced backwards.
<taxonomicName id="E596CBA65A19FFED90BF5E3C38641B1F" authority="Franz, 1967" authorityName="Franz" authorityYear="1967" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="fernandezlopezi">
<emphasis id="10E26C375A19FFED90BF5E3C3CE51B3B" box="[1155,1436,295,321]" italics="true" pageId="3" pageNumber="378">Tarphius fernandezlopezi</emphasis>
Franz, 1967
</taxonomicName>
(from Tenerife, Canary Islands) differs in the arrangement and number of gibbosities on each elytron, an elytral apex that is strongly produced backwards, and a TL&gt;
<quantity id="E56E1DC05A19FFED97475E743AB91BF3" box="[891,960,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="378" unit="mm" value="4.0">4 mm</quantity>
.
<taxonomicName id="E596CBA65A19FFED97F05E743C2D1BF3" authority="Gillerfors, 1991" authorityName="Gillerfors" authorityYear="1991" box="[972,1364,367,393]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="huggerti">
<emphasis id="10E26C375A19FFED97F05E743DEE1BF3" box="[972,1175,367,393]" italics="true" pageId="3" pageNumber="378">Tarphius huggerti</emphasis>
Gillerfors, 1991
</taxonomicName>
(from Gran Canaria, Canary Islands) differs in the arrangement and number of gibbosities on each elytron and the elytral apex is strongly produced backwards.
</paragraph>
<caption id="76E9E0AD5A19FFED94AB5A9F3AC71FE4" ID-DOI="http://doi.org/10.5281/zenodo.5141653" ID-Zenodo-Dep="5141653" box="[151,958,1412,1439]" httpUri="https://zenodo.org/record/5141653/files/figure.png" pageId="3" pageNumber="378" startId="3.[151,260,1412,1438]" targetBox="[240,1350,498,1380]" targetPageId="3">
<paragraph id="2229B0255A19FFED94AB5A9F3AC71FE4" blockId="3.[151,958,1412,1439]" box="[151,958,1412,1439]" pageId="3" pageNumber="378">
<emphasis id="10E26C375A19FFED94AB5A9F38591FE4" bold="true" box="[151,288,1412,1438]" pageId="3" pageNumber="378">FIGURE 3.</emphasis>
Genitalia of
<taxonomicName id="E596CBA65A19FFED958F5A9E3B301FE4" authority="Müller &amp; Schuh, 2021" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[435,585,1412,1438]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="hyrcanicus" status="sp. nov.">
<emphasis id="10E26C375A19FFED958F5A9E3B301FE4" box="[435,585,1412,1438]" italics="true" pageId="3" pageNumber="378">T. hyrcanicus</emphasis>
</taxonomicName>
<emphasis id="10E26C375A19FFED96735A9F3BF71FE4" bold="true" box="[591,654,1412,1438]" pageId="3" pageNumber="378">
<taxonomicNameLabel id="0BD1D14C5A19FFED96735A9F3BF71FE4" box="[591,654,1412,1438]" pageId="3" pageNumber="378" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(a) male and (b, c) female.
</paragraph>
</caption>
<paragraph id="2229B0255A19FFED94FB5AD73D611C54" blockId="3.[151,1437,1484,1979]" pageId="3" pageNumber="378">
<taxonomicName id="E596CBA65A19FFED94FB5AD73B1C1F9C" authority="Wollaston, 1862" authorityName="Wollaston" authorityYear="1862" box="[199,613,1484,1510]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="10E26C375A19FFED94FB5AD738DD1F9C" box="[199,420,1484,1510]" italics="true" pageId="3" pageNumber="378">Tarphius quadratus</emphasis>
Wollaston, 1862
</taxonomicName>
(from La Palma, Canary Islands) and
<taxonomicName id="E596CBA65A19FFED90225AD63C541F9C" authority="Franz, 1967" authorityName="Franz" authorityYear="1967" box="[1054,1325,1484,1511]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="gomerae">
<emphasis id="10E26C375A19FFED90225AD63DE41F9C" box="[1054,1181,1485,1510]" italics="true" pageId="3" pageNumber="378">T. gomerae</emphasis>
Franz, 1967
</taxonomicName>
(from La Gomera, Canary Islands) have 5 or 6 gibbosities of different arrangement on each elytron, a shorter elytra (EL/EW ~1.0), a depressed elytral disc, and an elytral apex produced slightly backwards.
</paragraph>
<paragraph id="2229B0255A19FFED94FB59233AF81D08" blockId="3.[151,1437,1484,1979]" pageId="3" pageNumber="378">
<taxonomicName id="E596CBA65A19FFED94FB59233B561C28" authority="Gillerfors, 1986" authorityName="Gillerfors" authorityYear="1986" box="[199,559,1592,1618]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="ericae">
<emphasis id="10E26C375A19FFED94FB5923380F1C28" box="[199,374,1592,1618]" italics="true" pageId="3" pageNumber="378">Tarphius ericae</emphasis>
Gillerfors, 1986
</taxonomicName>
(from La Gomera, Canary Islands) differs by a lateral pronotal margin slightly sinuate before the posterior angles, the presence of 4 gibbosities on each elytron, and an elytral apex slightly produced backwards.
<taxonomicName id="E596CBA65A19FFED9527599B3BCE1CE1" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[283,695,1664,1691]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="10E26C375A19FFED9527599B38821CE0" box="[283,507,1664,1690]" italics="true" pageId="3" pageNumber="378">Tarphius compactus</emphasis>
Wollaston, 1854
</taxonomicName>
,
<taxonomicName id="E596CBA65A19FFED96FC599A3A8B1CE1" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[704,1010,1664,1691]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="nodosus">
<emphasis id="10E26C375A19FFED96FC599A3A4F1CE0" box="[704,822,1664,1690]" italics="true" pageId="3" pageNumber="378">T. nodosus</emphasis>
Wollaston, 1854
</taxonomicName>
,
<taxonomicName id="E596CBA65A19FFED97C7599A3C711CE1" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[1019,1288,1664,1691]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="lauri">
<emphasis id="10E26C375A19FFED97C7599A3D321CE0" box="[1019,1099,1664,1690]" italics="true" pageId="3" pageNumber="378">T. lauri</emphasis>
Wollaston, 1854
</taxonomicName>
,
<taxonomicName id="E596CBA65A19FFED912D599A38291CC5" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="10E26C375A19FFED912D599A3CE41CE0" box="[1297,1437,1664,1690]" italics="true" pageId="3" pageNumber="378">T. parallelus</emphasis>
Wollaston, 1854
</taxonomicName>
,
<taxonomicName id="E596CBA65A19FFED956559BE3BE51CC5" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[345,668,1700,1727]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="10E26C375A19FFED956559BE38A71CC4" box="[345,478,1701,1726]" italics="true" pageId="3" pageNumber="378">T. inornatus</emphasis>
Wollaston, 1854
</taxonomicName>
,
<taxonomicName id="E596CBA65A19FFED969A59BE3A8F1CC5" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[678,1014,1700,1727]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="10E26C375A19FFED969A59BE3A411CC4" box="[678,824,1700,1726]" italics="true" pageId="3" pageNumber="378">T. rotundatus</emphasis>
Wollaston, 1854
</taxonomicName>
, and
<taxonomicName id="E596CBA65A19FFED901259BE3C081CC4" authority="Wollaston, 1871" authorityName="Wollaston" authorityYear="1871" box="[1070,1393,1700,1726]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="lutulentus">
<emphasis id="10E26C375A19FFED901259BE3DCF1CC4" box="[1070,1206,1700,1726]" italics="true" pageId="3" pageNumber="378">T. lutulentus</emphasis>
Wollaston, 1871
</taxonomicName>
(all from Madeira) differ by 2 or 3 faint (sometimes absent) gibbosities on the apical half of each elytron, an elytral apex distinctly produced backwards in the sutural region, and anterior pronotal angles more distinctly pointed forwards.
<taxonomicName id="E596CBA65A19FFED94AB580B3B451D51" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" box="[151,572,1808,1835]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="testudinalis">
<emphasis id="10E26C375A19FFED94AB580B38F81D50" box="[151,385,1808,1834]" italics="true" pageId="3" pageNumber="378">Tarphius testudinalis</emphasis>
Wollaston, 1854
</taxonomicName>
and
<taxonomicName id="E596CBA65A19FFED964D580A3AA31D51" authority="Wollaston, 1857" authorityName="Wollaston" authorityYear="1857" box="[625,986,1808,1835]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="sculptipennis">
<emphasis id="10E26C375A19FFED964D580A3A661D50" box="[625,799,1808,1834]" italics="true" pageId="3" pageNumber="378">T. sculptipennis</emphasis>
Wollaston, 1857
</taxonomicName>
(from Madeira) have 5 gibbosities (with an arrangement similar to that in
<taxonomicName id="E596CBA65A19FFED9637582E3BD91D34" authorityName="Müller &amp; Schuh" authorityYear="2021" box="[523,672,1844,1870]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="378" phylum="Arthropoda" rank="species" species="hyrcanicus">
<emphasis id="10E26C375A19FFED9637582E3BD91D34" box="[523,672,1844,1870]" italics="true" pageId="3" pageNumber="378">T. hyrcanicus</emphasis>
</taxonomicName>
) on each elytron but their humeral angles protrude forwards and the elytral apex is distinctly produced backwards in the sutural region.
</paragraph>
</subSubSection>
<subSubSection id="6A8CE3AE5A19FFED94FB58673CE91DC1" pageId="3" pageNumber="378" type="biology_ecology">
<paragraph id="2229B0255A19FFED94FB58673CE91DC1" blockId="3.[151,1437,1484,1979]" pageId="3" pageNumber="378">
<emphasis id="10E26C375A19FFED94FB586738E91DEC" bold="true" box="[199,400,1916,1942]" pageId="3" pageNumber="378">Ecological notes.</emphasis>
The
<typeStatus id="FD2D0E875A19FFED95F058673B571DEC" box="[460,558,1916,1942]" pageId="3" pageNumber="378" type="holotype">holotype</typeStatus>
was extracted from leaf litter and fine woody debris in an old-growth
<taxonomicName id="E596CBA65A19FFED916A5866387B1DC0" authorityName="Lipsky" authorityYear="1898" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="378" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="10E26C375A19FFED916A5866387B1DC0" italics="true" pageId="3" pageNumber="378">Fagus orientalis</emphasis>
</taxonomicName>
forest using a Berlese funnel. The
<typeStatus id="FD2D0E875A19FFED96B658BA3B8C1DC1" box="[650,757,1953,1979]" pageId="3" pageNumber="378" type="paratype">paratypes</typeStatus>
were collected near small streams at an altitude of ~
<specimenCount id="34907BAC5A19FFED917D58BB3CE91DC1" box="[1345,1424,1952,1979]" pageId="3" pageNumber="378" type="generic">
<quantity id="E56E1DC05A19FFED917D58BB3CF51DC1" box="[1345,1420,1952,1979]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="378" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>