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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e10084" ID-PMC="PMC5139134" ID-Pensoft-Pub="1314-2828-4-10084" ID-PubMed="27956851" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e10084" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Cyanobacteria of Greece: an annotated checklist" checkinTime="1478060999303" checkinUser="pensoft" docAuthor="Gkelis, Spyros, Ourailidis, Iordanis, Panou, Manthos &amp; Pappas, Nikos" docDate="2016" docId="411B18D1F8E73AC29EF1A4BB52A0A7AB" docLanguage="en" docName="BiodivDatJour 4: e10084" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e10084" docTitle="Microcystis panniformis Komarek, Komarkova-Legnerova, Sant'Anna, M. T. P. Azevedo, &amp; P. A. C. Senna 2002" docType="treatment" docVersion="5" lastPageNumber="10084" masterDocId="4C38E221FFF08B34321CFFEC8C1FFF8D" masterDocTitle="Cyanobacteria of Greece: an annotated checklist" masterLastPageNumber="10084" masterPageNumber="10084" pageNumber="10084" updateTime="1668123304914" updateUser="ExternalLinkService">
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<taxonomicName authority="Komarek, Komarkova-Legnerova, Sant'Anna, M. T. P. Azevedo, &amp; P. A. C. Senna, 2002" authorityName="Komarek, Komarkova-Legnerova, Sant'Anna, M. T. P. Azevedo, &amp; P. A. C. Senna" authorityYear="2002" class="Cyanophyceae" family="Chroococcaceae" genus="Microcystis" higherTaxonomySource="CoL" kingdom="Bacteria" lsidName="Microcystis panniformis" order="Chroococcales" pageId="0" pageNumber="10084" phylum="Cyanobacteria" rank="species" species="panniformis">
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<bibRefCitation author="Papadimitriou, Th." journalOrPublisher="Science of The Total Environment" pageId="0" pageNumber="10084" pagination="116 - 124" title="Artificially-born &quot; killer &quot; lake: Phytoplankton based water quality and microcystin affected fish in a reconstructed lake" volume="452 - 453" year="2013">Papadimitriou et al. 2013</bibRefCitation>
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<taxonomicName authority="(L.) Ker Gawl." class="Liliopsida" family="Orchidaceae" genus="Pogonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pogonia ophioglossoides" order="Asparagales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="species" species="ophioglossoides">Pogonia ophioglossoides (L.) Ker Gawl.</taxonomicName>
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<paragraph pageId="0" pageNumber="1099">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1099">Wet pine savannas (SPS-RF, VWLPS).</paragraph>
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Rare.
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. Thornhill 1286 (NCSC). Specimens seen in the vicinity: Sandy Run [Neck]: Taggart SARU 190 (WNC!). [= RAB, FNA, Weakley]
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misident.
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Siebold &amp; Zucc., Fl. Jap. 1: 110, t. 56, 57-I (1839); Ohba &amp; Akiyama in Bull. Natl. Mus. Nat. Sci., Ser. B, 39: 178 (2013).
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, Kyushu (
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Fig. 11Map 11
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CANADA: ON:Hald.-Norfolk Reg.,Cronmiller Prop., ~6km W St. Williams,
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Canada: ON; USA: GA, IN, KS (
<bibRefCitation author="Assing, V" journalOrPublisher="Beitraege zur Entomologie" pageId="42" pageNumber="161" pagination="111 - 204" title="A taxonomic and phylogenetic revision of Amarochara Thomson. I. The species of the Holarctic region (Coleoptera: Staphylinidae, Aleocharinae, Oxypodini)." volume="52" year="2002">Assing 2002</bibRefCitation>
). Native.
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This species can by distinguished from other Nearctic
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by the following combination of characters: head and pronotum with weak microsculpture; first segment of metatarsus about as long as second to fourth segments combined; punctation of abdominal tergites sparse (
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,
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). The shape of the median lobe of the aedeagus is also distinctive in lateral view.
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All specimens of this species with collection data were collected in forested reserves using flight intercept traps (
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, this study) but nothing further is known about its biology.
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:
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urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFC1FF9EFD8FFACCFE436FC6" pageId="35" pageNumber="36">948DE50E-A684- 4E57-8126-658578AF3E06</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFF08FA19FECC6CF0" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="C90D4710FFC1FF9EFF08FA19FE356C75" blockId="35.[163,779,1460,1696]" pageId="35" pageNumber="36">
<emphasis id="FBC69B02FFC1FF9EFF08FA19FE026F99" box="[163,404,1460,1481]" italics="true" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFF08FA19FEA06F99" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[163,310,1460,1481]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
bullocki
</emphasis>
misidentification, not
<taxonomicName id="0EB23C93FFC1FF9EFD25FA19FEE06FB8" ID-CoL="3P32T" authority="Collingwood, 1881" authorityName="Collingwood" authorityYear="1881" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC1FF9EFD25FA19FC9C6F99" box="[654,778,1460,1481]" italics="true" pageId="35" pageNumber="36">C. bullockii</emphasis>
<bibRefCitation id="AD233AE1FFC1FF9EFF08FA7FFEE06FB8" author="Collingwood C." box="[163,374,1490,1512]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="35" pageNumber="36" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFC1FF9EFE2AFA7FFDB26FB8" author="Johnson S." box="[385,548,1490,1512]" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/2092" refId="ref43125" refString="Johnson S. 2000. Hypselodoris bullocki from the Solomon Ids. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 2092" title="Hypselodoris bullocki from the Solomon Ids" type="url" volumeTitle="Sea slug forum" year="2000">Johnson, 2000</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC1FF9EFD99FA7FFD4A6FB8" author="Coleman N." box="[562,732,1490,1512]" journalOrPublisher="Springwood: Australia Neville Coleman's Underwater Geographic" pageId="35" pageNumber="36" refId="ref41724" refString="Coleman N. 2001. 1001 Nudibranchs: catalogue of Indo- Pacific sea slugs. Springwood: Australia Neville Coleman's Underwater Geographic." title="1001 Nudibranchs: catalogue of Indo- Pacific sea slugs" type="book" year="2001">Coleman, 2001</bibRefCitation>
: 79, lower right photograph;
<bibRefCitation id="AD233AE1FFC1FF9EFE61FA5CFD066C56" author="Gaensslen RE" box="[458,656,1521,1543]" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/20471" refId="ref42146" refString="Gaensslen RE 2007. Hypselodoris bullocki? from Papua New Guinea. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20471" title="Hypselodoris bullocki? from Papua New Guinea" type="url" volumeTitle="Sea slug forum" year="2007">Gaensslen, 2007</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC1FF9EFD34FA5CFF4C6C75" author="Rudman WB" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/hypsbull" refId="ref45077" refString="Rudman WB. 2007. Hypselodoris bullocki - feeding record. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypsbull" title="Hypselodoris bullocki - feeding record" type="url" volumeTitle="Sea slug forum" year="2007">Rudman, 2007</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC1FF9EFF4DF9BDFE0A6C75" author="Hanchard B." box="[230,412,1552,1574]" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/22275" refId="ref42817" refString="Hanchard B. 2009. Blue Hypselodoris bullocki egg-laying. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 22275" title="Blue Hypselodoris bullocki egg-laying" type="url" volumeTitle="Sea slug forum" year="2009">Hanchard, 2009</bibRefCitation>
.
</paragraph>
<paragraph id="C90D4710FFC1FF9EFF08F983FE056C33" blockId="35.[163,779,1460,1696]" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFF08F983FEAA6C13" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[163,316,1582,1603]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC1FF9EFF08F983FEAA6C13" box="[163,316,1582,1603]" italics="true" pageId="35" pageNumber="36">Hypselodoris</emphasis>
</taxonomicName>
sp. 7
<bibRefCitation id="AD233AE1FFC1FF9EFE28F983FDE26C14" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[387,628,1582,1604]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="35" pageNumber="36" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFC1FF9EFE5AF982FDBA6C13" box="[497,556,1582,1604]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2008
</bibRefCitation>
: 268, second photograph from top.
</paragraph>
<paragraph id="C90D4710FFC1FF9EFF08F9C1FECC6CF0" blockId="35.[163,779,1460,1696]" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFF08F9C1FEAF6CD1" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[163,313,1644,1665]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC1FF9EFF08F9C1FEAF6CD1" box="[163,313,1644,1665]" italics="true" pageId="35" pageNumber="36">Hypselodoris</emphasis>
</taxonomicName>
sp. 16
<bibRefCitation id="AD233AE1FFC1FF9EFE20F9C1FDE36CD2" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[395,629,1644,1666]" journalOrPublisher="Jacksonville: New World Press" pageId="35" pageNumber="36" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFC1FF9EFE5EF9C1FDB96CD1" box="[501,559,1644,1665]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2015
</bibRefCitation>
: 262, bottom left photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFF08F965FBBC6F1B" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="C90D4710FFC1FF9EFF08F965FEDA6C8F" blockId="35.[163,779,1735,1882]" box="[163,332,1735,1759]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFF08F965FEDA6C8F" box="[163,332,1735,1759]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFF08F965FEDA6C8F" box="[163,332,1735,1759]" italics="true" pageId="35" pageNumber="36">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFF08F945FEE96D09" blockId="35.[163,779,1735,1882]" pageId="35" pageNumber="36">
<materialsCitation id="79DA4D4DFFC1FF9EFF08F945FEE96D09" ID-GBIF-Occurrence="2235579121" collectingDate="2012-11-15" collectionCode="CASIZ" country="Papua New Guinea" elevation="8" latitude="-5.0985" location="Southern Sek Island" longLatPrecision="7" longitude="145.821" municipality="Lagoon" pageId="35" pageNumber="36" specimenCode="CASIZ 191246" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="1609F9B2FFC1FF9EFF08F945FE9D6CAD" box="[163,267,1768,1789]" pageId="35" pageNumber="36" type="holotype">
<emphasis id="FBC69B02FFC1FF9EFF08F945FE9D6CAD" box="[163,267,1768,1789]" italics="true" pageId="35" pageNumber="36">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFC1FF9EFEB6F945FE5D6CAE" box="[285,459,1768,1790]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFEDFF945FE5D6CAE" box="[372,459,1768,1790]" pageId="35" pageNumber="36" value="46-12-19">191246</date>
</specimenCode>
, subsampled for molecular study,
<location id="CC6D11CBFFC1FF9EFF40F8ABFE416D4C" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFF40F8ABFE416D4C" box="[235,471,1798,1820]" country="Papua New Guinea" latitude="-5.0985" longLatPrecision="7" longitude="145.821" municipality="Lagoon" name="Southern Sek Island" pageId="35" pageNumber="36" stateProvince="Madang">Southern Sek Island</location>
,
<geoCoordinate id="AC8621D7FFC1FF9EFE4AF8ABFDD16D4C" box="[481,583,1798,1820]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.0985">5.0985°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFDF9F8ABFD406D4C" box="[594,726,1798,1820]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.821">145.8210°E</geoCoordinate>
,
<elevation id="429FA023FFC1FF9EFD4AF8AAFC9D6D4C" box="[737,779,1799,1820]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">
<quantity id="0E4AEAF5FFC1FF9EFD4AF8AAFC9D6D4C" box="[737,779,1799,1820]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">8 m</quantity>
</elevation>
depth,
<collectingRegion id="0B7689F2FFC1FF9EFF58F888FEC46D6B" box="[243,338,1829,1851]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<collectingMunicipality id="2969DD6AFFC1FF9EFEF1F88BFE276D6B" box="[346,433,1830,1851]" pageId="35" pageNumber="36">Lagoon</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFC1FF9EFE16F888FD8B6D6B" box="[445,541,1829,1851]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFD82F88BFC9D6D6B" box="[553,779,1829,1851]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
<collectingDate id="AD489838FFC1FF9EFF08F8E9FEED6D09" box="[163,379,1860,1882]" pageId="35" pageNumber="36" value="2012-11-15">
<date id="BD0C61D0FFC1FF9EFF08F8E9FEED6D09" box="[163,379,1860,1882]" pageId="35" pageNumber="36" value="2012-11-15">15 November 2012</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90FF6AFBBC6F1B" blockId="35.[827,1443,198,1355]" pageId="35" pageNumber="36">
<materialsCitation id="79DA4D4DFFC1FF9EFC90FF6AFC306B68" ID-GBIF-Occurrence="2235579099" collectionCode="CASIZ" collectorName="R. Steene" elevation="9" location="Milne Bay" municipality="Milne Bay Province" pageId="35" pageNumber="36" specimenCode="CASIZ 185100" specimenCount="1" typeStatus="paratype">
<typeStatus id="1609F9B2FFC1FF9EFC90FF6AFC2D6A8C" box="[827,955,199,220]" pageId="35" pageNumber="36" type="paratype">
<emphasis id="FBC69B02FFC1FF9EFC90FF6AFC2D6A8C" box="[827,955,199,220]" italics="true" pageId="35" pageNumber="36">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFC1FF9EFC7BFF6AFB076A8C" box="[976,1169,198,220]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 185100</specimenCode>
, five specimens, one subsampled for molecular study, various locations,
<location id="CC6D11CBFFC1FF9EFC90FEA9FC576B49" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC90FEA9FC576B49" box="[827,961,260,282]" municipality="Milne Bay Province" name="Milne Bay" pageId="35" pageNumber="36">Milne Bay</location>
,
<collectingMunicipality id="2969DD6AFFC1FF9EFC79FEA9FB4E6B4A" box="[978,1240,260,282]" pageId="35" pageNumber="36">Milne Bay Province</collectingMunicipality>
,
<elevation id="429FA023FFC1FF9EFB4CFEA9FADE6B49" box="[1255,1352,260,282]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">
<quantity id="0E4AEAF5FFC1FF9EFB4CFEA9FADE6B49" box="[1255,1352,260,282]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">5–12 m</quantity>
</elevation>
depth,
<collectorName id="644722C6FFC1FF9EFC90FE8EFC306B68" box="[827,934,291,312]" pageId="35" pageNumber="36">R. Steene</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC1FF9EFC04FE8EFB916B07" ID-GBIF-Occurrence="2235579137" collectionCode="CASIZ" country="Iceland" location="Anemone Reef" municipality="Ruo" pageId="35" pageNumber="36" specimenCount="1" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC04FE8EFC6C6B68" box="[943,1018,291,312]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69754, one specimen,
<location id="CC6D11CBFFC1FF9EFB56FE8EFA096B68" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB56FE8EFA096B68" box="[1277,1439,290,312]" country="Iceland" municipality="Ruo" name="Anemone Reef" pageId="35" pageNumber="36">Anemone Reef</location>
, near
<collectingMunicipality id="2969DD6AFFC1FF9EFCD0FEEFFC3A6B07" box="[891,940,322,343]" pageId="35" pageNumber="36">Ruo</collectingMunicipality>
,
<collectingCountry id="B1A50780FFC1FF9EFC12FEECFB916B07" box="[953,1031,321,343]" name="Iceland" pageId="35" pageNumber="36">Island</collectingCountry>
</materialsCitation>
,
<materialsCitation id="79DA4D4DFFC1FF9EFBBEFEECFBF66B25" ID-GBIF-Occurrence="2235579061" collectingDate="1989-07-20" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectingRegion id="0B7689F2FFC1FF9EFBBEFEECFBE06B07" box="[1045,1142,321,343]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB2BFEEFFB4C6B07" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB2BFEEFFB4C6B07" box="[1152,1242,322,343]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB4CFEECFADC6B07" box="[1255,1354,321,343]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFAFCFEEFFC516B26" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFC79FECDFBF66B25" box="[978,1120,352,374]" pageId="35" pageNumber="36" value="1989-07-20">
<date id="BD0C61D0FFC1FF9EFC79FECDFBF66B25" box="[978,1120,352,374]" pageId="35" pageNumber="36" value="1989-07-20">20 July 1989</date>
</collectingDate>
</materialsCitation>
, M. Jebb.
<materialsCitation id="79DA4D4DFFC1FF9EFB65FECDFBF46BE2" ID-GBIF-Occurrence="2235579029" collectingDate="1989-08" collectionCode="CASIZ" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB65FECDFA8F6B25" box="[1230,1305,352,373]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69787, one specimen, dissected,
<collectingRegion id="0B7689F2FFC1FF9EFB82FED3FB116BC4" box="[1065,1159,382,404]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB25FED2FB726BC4" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB25FED2FB726BC4" box="[1166,1252,383,404]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB44FED3FAD86BC4" box="[1263,1358,382,404]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFAF1FED2FC5E6BE3" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFC79FE30FBF46BE2" box="[978,1122,413,434]" pageId="35" pageNumber="36" value="1989-08">
<date id="BD0C61D0FFC1FF9EFC79FE30FBF46BE2" box="[978,1122,413,434]" pageId="35" pageNumber="36" value="1989-08">August 1989</date>
</collectingDate>
</materialsCitation>
, M. Gosliner.
<materialsCitation id="79DA4D4DFFC1FF9EFB51FE30FBBB6BA0" ID-GBIF-Occurrence="2235579142" collectingDate="1989-08" collectionCode="CASIZ" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB51FE30FAD36BE2" box="[1274,1349,413,434]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69785, three specimens,
<collectingRegion id="0B7689F2FFC1FF9EFC50FE11FBC16B82" box="[1019,1111,444,466]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBF6FE11FB266B81" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBF6FE11FB266B81" box="[1117,1200,444,465]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB10FE11FA8E6B82" box="[1211,1304,444,466]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFA89FE11FC066BA0" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFC31FE76FBBB6BA0" box="[922,1069,475,496]" pageId="35" pageNumber="36" value="1989-08">
<date id="BD0C61D0FFC1FF9EFC31FE76FBBB6BA0" box="[922,1069,475,496]" pageId="35" pageNumber="36" value="1989-08">August 1989</date>
</collectingDate>
</materialsCitation>
, M. Gosliner.
<materialsCitation id="79DA4D4DFFC1FF9EFB63FE76FA0B687D" ID-GBIF-Occurrence="2235579072" collectingDate="1988-02-03" collectionCode="CASIZ" country="Papua New Guinea" county="Hussein Village" elevation="20" location="Hole" municipality="Wall" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB63FE76FA826BA0" box="[1224,1300,475,496]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 65356, one specimen,
<location id="CC6D11CBFFC1FF9EFC1FFE54FC7C685F" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC1FFE54FC7C685F" box="[948,1002,505,527]" country="Papua New Guinea" county="Hussein Village" municipality="Wall" name="Hole" pageId="35" pageNumber="36" stateProvince="Madang">Hole</location>
in the
<collectingMunicipality id="2969DD6AFFC1FF9EFB95FE54FBE3685F" box="[1086,1141,505,527]" pageId="35" pageNumber="36">Wall</collectingMunicipality>
, north of
<collectingCounty id="206C3F9CFFC1FF9EFB4EFE54FA09685F" box="[1253,1439,505,527]" pageId="35" pageNumber="36">Hussein Village</collectingCounty>
, north of
<collectingRegion id="0B7689F2FFC1FF9EFC06FDB5FB82687E" box="[941,1044,536,558]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<elevation id="429FA023FFC1FF9EFB88FDB5FBF5687D" box="[1059,1123,536,557]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">
<quantity id="0E4AEAF5FFC1FF9EFB88FDB5FBF5687D" box="[1059,1123,536,557]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">20 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC1FF9EFB63FDB5FA0B687D" box="[1224,1437,536,558]" pageId="35" pageNumber="36" value="1988-02-03">
<date id="BD0C61D0FFC1FF9EFB63FDB5FA0B687D" box="[1224,1437,536,558]" pageId="35" pageNumber="36" value="1988-02-03">3 February 1988</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFC1FF9EFC15FD9AFA09683A" ID-GBIF-Occurrence="2235579119" collectingDate="1988-01-24" collectionCode="CASIZ" country="Papua New Guinea" elevation="24" location="Kranket Island" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC15FD9AFB9F681C" box="[958,1033,567,588]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 71237, one specimen, patch reef off N side
<location id="CC6D11CBFFC1FF9EFC27FDF8FBA9683B" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC27FDF8FBA9683B" box="[908,1087,597,619]" country="Papua New Guinea" name="Kranket Island" pageId="35" pageNumber="36" stateProvince="Madang">Kranket Island</location>
,
<elevation id="429FA023FFC1FF9EFBE0FDF8FB12683B" box="[1099,1156,597,619]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="24.0">
<quantity id="0E4AEAF5FFC1FF9EFBE0FDF8FB12683B" box="[1099,1156,597,619]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="24.0">24 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC1FF9EFB70FDF8FA09683A" box="[1243,1439,597,619]" pageId="35" pageNumber="36" value="1988-01-24">
<date id="BD0C61D0FFC1FF9EFB70FDF8FA09683A" box="[1243,1439,597,619]" pageId="35" pageNumber="36" value="1988-01-24">24 January 1988</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFC1FF9EFC6DFDD9FAD06897" ID-GBIF-Occurrence="2235579124" collectingDate="1986-09-01" collectionCode="CASIZ" country="Solomon Islands" county="Florida Group" elevation="19" location="Takahate Bay" municipality="Big Nggela Island" pageId="35" pageNumber="36" specimenCount="1" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC6DFDD9FB8568D9" box="[966,1043,628,649]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 71474, two specimens,
<location id="CC6D11CBFFC1FF9EFA9FFDD9FCFD68F8" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFA9FFDD9FCFD68F8" country="Solomon Islands" county="Florida Group" municipality="Big Nggela Island" name="Takahate Bay" pageId="35" pageNumber="36">Takahate Bay</location>
,
<collectingMunicipality id="2969DD6AFFC1FF9EFCD1FD3FFBF268F8" box="[890,1124,658,680]" pageId="35" pageNumber="36">Big Nggela Island</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFC1FF9EFBDFFD3FFABC68F8" box="[1140,1322,658,680]" pageId="35" pageNumber="36">Florida Group</collectingCounty>
,
<collectingCountry id="B1A50780FFC1FF9EFA91FD3FFC036897" name="Solomon Islands" pageId="35" pageNumber="36">Solomon Islands</collectingCountry>
,
<elevation id="429FA023FFC1FF9EFC09FD1CFB9D6897" box="[930,1035,689,711]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="35" pageNumber="36" unit="m" value="19.0" valueMax="23.0" valueMin="15.0">
<quantity id="0E4AEAF5FFC1FF9EFC09FD1CFB9D6897" box="[930,1035,689,711]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="35" pageNumber="36" unit="m" value="19.0" valueMax="23.0" valueMin="15.0">15–23 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC1FF9EFBC3FD1CFAD06897" box="[1128,1350,689,711]" pageId="35" pageNumber="36" value="1986-09-01">
<date id="BD0C61D0FFC1FF9EFBC3FD1CFAD06897" box="[1128,1350,689,711]" pageId="35" pageNumber="36" value="1986-09-01">1 September 1986</date>
</collectingDate>
</materialsCitation>
, R. Van Syoc.
<materialsCitation id="79DA4D4DFFC1FF9EFCD7FD7DFA926973" ID-GBIF-Occurrence="2235579067" collectingDate="1922-11" collectionCode="CASIZ" collectorName="N. Tadwai Island" country="Papua New Guinea" elevation="11" latitude="-4.985" location="Lagoon" longLatPrecision="55" longitude="145.7915" pageId="35" pageNumber="36" specimenCode="CASIZ 191392" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFCD7FD7DFBB368B5" box="[892,1061,720,741]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 191392</specimenCode>
, one specimen,
<collectorName id="644722C6FFC1FF9EFB72FD7DFA0868B6" box="[1241,1438,720,742]" pageId="35" pageNumber="36">N. Tadwai Island</collectorName>
,
<geoCoordinate id="AC8621D7FFC1FF9EFC90FD43FC056954" box="[827,915,750,772]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="55" value="-4.985">4.985°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFC35FD43FBB76954" box="[926,1057,750,772]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.7915">145.7915°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFC1FF9EFB86FD43FB1C6954" box="[1069,1162,750,772]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB3AFD42FB706954" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB3AFD42FB706954" box="[1169,1254,751,772]" country="Papua New Guinea" latitude="-4.985" longLatPrecision="55" longitude="145.7915" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB5AFD43FAD96954" box="[1265,1359,750,772]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFAF0FD42FC5B6973" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFC1FF9EFC72FCA0FB856973" box="[985,1043,781,803]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="m" value="11.0">
<quantity id="0E4AEAF5FFC1FF9EFC72FCA0FB856973" box="[985,1043,781,803]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="m" value="11.0">11 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC1FF9EFBC0FCA0FA926973" box="[1131,1284,781,803]" pageId="35" pageNumber="36" value="1922-11">
<date id="BD0C61D0FFC1FF9EFBC0FCA0FA926973" box="[1131,1284,781,803]" pageId="35" pageNumber="36" value="1922-11">22 November</date>
</collectingDate>
</materialsCitation>
, 2012.
<materialsCitation id="79DA4D4DFFC1FF9EFAFDFCA0FAD96930" ID-GBIF-Occurrence="2235579035" collectingDate="2012-11-12" collectionCode="CASIZ" country="Papua New Guinea" location="Sek Island" municipality="Lagoon" pageId="35" pageNumber="36" specimenCode="CASIZ 191160" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFAFDFCA0FC056912" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFC90FC81FC056912" box="[827,915,812,834]" pageId="35" pageNumber="36" value="60-11-19">191160</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFC1FF9EFBE5FC81FB466912" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE5FC81FB466912" box="[1102,1232,812,834]" country="Papua New Guinea" municipality="Lagoon" name="Sek Island" pageId="35" pageNumber="36" stateProvince="Madang">Sek Island</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB76FC81FAAB6912" box="[1245,1341,812,834]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<collectingMunicipality id="2969DD6AFFC1FF9EFAEDFC81FA086911" box="[1350,1438,812,833]" pageId="35" pageNumber="36">Lagoon</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFC1FF9EFC90FCE7FC0E6930" box="[827,920,842,864]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFC0BFCE6FBE16930" box="[928,1143,842,864]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFB2BFCE6FAD96930" box="[1152,1359,842,864]" pageId="35" pageNumber="36" value="2012-11-12">
<date id="BD0C61D0FFC1FF9EFB2BFCE6FAD96930" box="[1152,1359,842,864]" pageId="35" pageNumber="36" value="2012-11-12">12 November 2012</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC1FF9EFAF3FCE6FB2E69EC" ID-GBIF-Occurrence="2235579100" collectingDate="2012-11-15" collectionCode="CASIZ" country="Papua New Guinea" elevation="8" latitude="-5.0985" location="Southern Sek Island" longLatPrecision="7" longitude="145.821" municipality="Lagoon" pageId="35" pageNumber="36" specimenCode="CASIZ 191227" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFAF3FCE6FC19692E" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFC90FCC4FC19692E" box="[827,911,873,894]" pageId="35" pageNumber="36" value="27-12-19">191227</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFC1FF9EFBEBFCC4FABB692F" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBEBFCC4FABB692F" box="[1088,1325,873,895]" country="Papua New Guinea" latitude="-5.0985" longLatPrecision="7" longitude="145.821" municipality="Lagoon" name="Southern Sek Island" pageId="35" pageNumber="36" stateProvince="Madang">Southern Sek Island</location>
,
<geoCoordinate id="AC8621D7FFC1FF9EFA92FCC4FA08692F" box="[1337,1438,873,895]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.0985">5.0985°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFC90FC25FC2A69CE" box="[827,956,904,926]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.821">145.8210°E</geoCoordinate>
,
<elevation id="429FA023FFC1FF9EFC6EFC25FC7B69CD" box="[965,1005,904,925]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">
<quantity id="0E4AEAF5FFC1FF9EFC6EFC25FC7B69CD" box="[965,1005,904,925]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">8 m</quantity>
</elevation>
depth,
<collectingRegion id="0B7689F2FFC1FF9EFB90FC25FB0169CE" box="[1083,1175,904,926]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<collectingMunicipality id="2969DD6AFFC1FF9EFB30FC25FB7869CD" box="[1179,1262,904,925]" pageId="35" pageNumber="36">Lagoon</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFC1FF9EFB5DFC25FAC569CE" box="[1270,1363,904,926]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFAF0FC25FC5869EC" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFC71FC0BFB2E69EC" box="[986,1208,934,956]" pageId="35" pageNumber="36" value="2012-11-15">
<date id="BD0C61D0FFC1FF9EFC71FC0BFB2E69EC" box="[986,1208,934,956]" pageId="35" pageNumber="36" value="2012-11-15">15 November 2012</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC1FF9EFB68FC0AFBFF6E48" ID-GBIF-Occurrence="2235579059" collectingDate="2012-12-12" collectionCode="CASIZ" collectorName="Cement Mixer Reef" country="Papua New Guinea" elevation="13" latitude="-5.15176" location="Lagoon" longLatPrecision="1" longitude="145.81831" pageId="35" pageNumber="36" specimenCode="CASIZ 190823" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB68FC0AFAF869EC" box="[1219,1390,935,956]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFAB3FC0AFAF869EC" box="[1304,1390,935,956]" pageId="35" pageNumber="36" value="23-08-19">190823</date>
</specimenCode>
, one specimen,
<collectorName id="644722C6FFC1FF9EFC04FC68FB1F698B" box="[943,1161,965,987]" pageId="35" pageNumber="36">Cement Mixer Reef</collectorName>
,
<geoCoordinate id="AC8621D7FFC1FF9EFB38FC68FA93698B" box="[1171,1285,965,987]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="-5.15176">5.15176°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFAA4FC68FA09698B" box="[1295,1439,965,987]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="145.81831">145.81832°E</geoCoordinate>
,
<elevation id="429FA023FFC1FF9EFC90FC49FC0669A9" box="[827,912,996,1017]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.3" metricValueMin="0.2" pageId="35" pageNumber="36" unit="m" value="12.5" valueMax="23.0" valueMin="2.0">
<quantity id="0E4AEAF5FFC1FF9EFC90FC49FC0669A9" box="[827,912,996,1017]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.3" metricValueMin="0.2" pageId="35" pageNumber="36" unit="m" value="12.5" valueMax="23.0" valueMin="2.0">2–23 m</quantity>
</elevation>
depth,
<collectingRegion id="0B7689F2FFC1FF9EFC42FC49FBDE69AA" box="[1001,1096,996,1018]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBE4FC49FB3069A9" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE4FC49FB3069A9" box="[1103,1190,996,1017]" country="Papua New Guinea" latitude="-5.15176" longLatPrecision="1" longitude="145.81831" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingRegion id="0B7689F2FFC1FF9EFB19FC49FA8469AA" box="[1202,1298,996,1018]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFAB6FC49FC186E48" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
<collectingDate id="AD489838FFC1FF9EFC3EFBAEFBFF6E48" box="[917,1129,1026,1048]" pageId="35" pageNumber="36" value="2012-12-12">
<date id="BD0C61D0FFC1FF9EFC3EFBAEFBFF6E48" box="[917,1129,1026,1048]" pageId="35" pageNumber="36" value="2012-12-12">12 December 2012</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFC1FF9EFB53FBAEFA096E05" ID-GBIF-Occurrence="2235579039" collectingDate="2012-11-19" collectionCode="CASIZ" country="Papua New Guinea" latitude="-5.0367" location="south Rempi" longLatPrecision="7" longitude="145.8066" pageId="35" pageNumber="36" specimenCode="CASIZ 191326" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB53FBAEFA096E48" box="[1272,1439,1026,1048]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 191326</specimenCode>
, one specimen,
<location id="CC6D11CBFFC1FF9EFC5BFB8CFB1A6E67" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC5BFB8CFB1A6E67" box="[1008,1164,1057,1079]" country="Papua New Guinea" latitude="-5.0367" longLatPrecision="7" longitude="145.8066" name="south Rempi" pageId="35" pageNumber="36" stateProvince="Madang">south Rempi</location>
<geoCoordinate id="AC8621D7FFC1FF9EFB3DFB8CFA956E67" box="[1174,1283,1057,1079]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.0367">5.0367°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFABAFB8CFA086E67" box="[1297,1438,1057,1079]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.8066">145.8066°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFC1FF9EFC90FBEDFC376E06" box="[827,929,1088,1110]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFC04FBEDFB326E06" box="[943,1188,1088,1110]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC1FF9EFB19FBEDFA096E05" box="[1202,1439,1088,1110]" pageId="35" pageNumber="36" value="2012-11-19">
<date id="BD0C61D0FFC1FF9EFB19FBEDFA096E05" box="[1202,1439,1088,1110]" pageId="35" pageNumber="36" value="2012-11-19">19 November 2012</date>
</collectingDate>
</materialsCitation>
, Francois Michonneau.
<materialsCitation id="79DA4D4DFFC1FF9EFBEDFBF2FB606EE1" ID-GBIF-Occurrence="2235579088" collectingDate="2012-11-12" collectionCode="CASIZ" country="Papua New Guinea" elevation="22" latitude="-5.2967" location="Bilbil Island" longLatPrecision="7" longitude="145.7816" pageId="35" pageNumber="36" specimenCode="CASIZ 191139" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFBEDFBF2FB646E24" box="[1094,1266,1119,1140]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFB37FBF2FB646E24" box="[1180,1266,1119,1140]" pageId="35" pageNumber="36" value="39-11-19">191139</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFC1FF9EFC90FBD0FC556EC3" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC90FBD0FC556EC3" box="[827,963,1149,1171]" country="Papua New Guinea" latitude="-5.2967" longLatPrecision="7" longitude="145.7816" name="Bilbil Island" pageId="35" pageNumber="36" stateProvince="Madang">Bilbil Island</location>
<geoCoordinate id="AC8621D7FFC1FF9EFC63FBD0FBBD6EC3" box="[968,1067,1149,1171]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.2967">5.2967°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFB9FFBD0FB206EC3" box="[1076,1206,1149,1171]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.7816">145.7816°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFC1FF9EFB15FBD0FA8D6EC3" box="[1214,1307,1149,1171]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFA8FFBD3FC186EE2" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFC1FF9EFC32FB31FC466EE1" box="[921,976,1180,1201]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">
<quantity id="0E4AEAF5FFC1FF9EFC32FB31FC466EE1" box="[921,976,1180,1201]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">22 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC1FF9EFB88FB31FB606EE1" box="[1059,1270,1180,1202]" pageId="35" pageNumber="36" value="2012-11-12">
<date id="BD0C61D0FFC1FF9EFB88FB31FB606EE1" box="[1059,1270,1180,1202]" pageId="35" pageNumber="36" value="2012-11-12">12 November 2012</date>
</collectingDate>
</materialsCitation>
, Heok Hui Tan.
<materialsCitation id="79DA4D4DFFC1FF9EFC90FB16FCE56F5D" ID-GBIF-Occurrence="2235579123" collectingDate="2012-11-09" collectionCode="CASIZ" collectorName="N. Siar Island" country="Papua New Guinea" latitude="-5.1967" location="Madang" longLatPrecision="7" longitude="145.8067" pageId="35" pageNumber="36" specimenCode="CASIZ 191066" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFC90FB16FC766E80" box="[827,992,1210,1232]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFC26FB17FC766E80" box="[909,992,1210,1232]" pageId="35" pageNumber="36" value="66-10-19">191066</date>
</specimenCode>
, one specimen,
<collectorName id="644722C6FFC1FF9EFB3BFB16FAB96E80" box="[1168,1327,1210,1232]" pageId="35" pageNumber="36">N. Siar Island</collectorName>
,
<geoCoordinate id="AC8621D7FFC1FF9EFA91FB17FA086E80" box="[1338,1438,1210,1232]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.1967">5.1967°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC1FF9EFC90FB74FC296EBF" box="[827,959,1241,1263]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.8067">145.8067°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFC1FF9EFC61FB74FBBE6EBF" box="[970,1064,1241,1263]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC1FF9EFB9FFB77FA876EBF" box="[1076,1297,1241,1263]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
<collectingDate id="AD489838FFC1FF9EFAB2FB74FCE56F5D" pageId="35" pageNumber="36" value="2012-11-09">
<date id="BD0C61D0FFC1FF9EFAB2FB74FCE56F5D" pageId="35" pageNumber="36" value="2012-11-09">9 November 2012</date>
</collectingDate>
</materialsCitation>
, Heok Hui Tan.
<materialsCitation id="79DA4D4DFFC1FF9EFB89FB55FBB36F1B" ID-GBIF-Occurrence="2235579079" collectionCode="CASIZ" country="Papua New Guinea" county="Marco Oliverio" location="Lagoon" municipality="November-December" pageId="35" pageNumber="36" specimenCode="CASIZ 190842" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB89FB55FB536F5D" box="[1058,1221,1272,1293]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ
<date id="BD0C61D0FFC1FF9EFBD9FB55FB536F5D" box="[1138,1221,1272,1293]" pageId="35" pageNumber="36" value="42-08-19">190842</date>
</specimenCode>
, one specimen, from orange sponge,
<collectingRegion id="0B7689F2FFC1FF9EFC41FABBFBD16F7C" box="[1002,1095,1302,1324]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBE4FABAFB326F7C" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE4FABAFB326F7C" box="[1103,1188,1303,1324]" country="Papua New Guinea" county="Marco Oliverio" municipality="November-December" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
,
<collectingMunicipality id="2969DD6AFFC1FF9EFB04FABBFA356F7C" box="[1199,1443,1302,1324]" pageId="35" pageNumber="36">November–December</collectingMunicipality>
2012,
<collectingCounty id="206C3F9CFFC1FF9EFCD6FA9BFBB36F1B" box="[893,1061,1333,1355]" pageId="35" pageNumber="36">Marco Oliverio</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFC90FADFFB276F97" pageId="35" pageNumber="36" type="distribution">
<paragraph id="C90D4710FFC1FF9EFC90FADFFBEE6FDA" blockId="35.[827,1443,1394,1479]" box="[827,1144,1394,1418]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90FADFFBEE6FDA" box="[827,1144,1394,1418]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90FADFFBEE6FDA" box="[827,1144,1394,1418]" italics="true" pageId="35" pageNumber="36">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90FA3EFB276F97" blockId="35.[827,1443,1394,1479]" pageId="35" pageNumber="36">
Known only from the
<collectingCountry id="B1A50780FFC1FF9EFBE7FA3EFAAD6FF8" box="[1100,1339,1426,1448]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
and the
<collectingCountry id="B1A50780FFC1FF9EFC90FA1CFC616F97" box="[827,1015,1457,1479]" name="Solomon Islands" pageId="35" pageNumber="36">Solomon Islands</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFC90FA43FA326C13" pageId="35" pageNumber="36" type="etymology">
<paragraph id="C90D4710FFC1FF9EFC90FA43FC2A6C56" blockId="35.[827,1444,1518,1603]" box="[827,956,1518,1542]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90FA43FC2A6C56" box="[827,956,1518,1542]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90FA43FC2A6C56" box="[827,956,1518,1542]" italics="true" pageId="35" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90F9A3FA326C13" blockId="35.[827,1444,1518,1603]" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFC90F9A3FBCE6C73" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1112,1550,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC1FF9EFC90F9A3FBCE6C73" box="[827,1112,1550,1571]" italics="true" pageId="35" pageNumber="36">Hypselodoris melanesica</emphasis>
</taxonomicName>
is named for Melanesia, the region to which this species is geographically restricted.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF98FC90F9C6FD7D6B07" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="C90D4710FFC1FF9EFC90F9C6FC516CD2" blockId="35.[827,1443,1643,1880]" box="[827,967,1643,1666]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90F9C6FC516CD2" box="[827,967,1643,1666]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90F9C6FC516CD2" box="[827,967,1643,1666]" italics="true" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF99FC90F927FD3B6F4E" blockId="35.[827,1443,1643,1880]" lastBlockId="36.[145,762,1226,1310]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="FBC69B02FFC1FF9EFC90F927FBAE6CCF" box="[827,1080,1674,1695]" italics="true" pageId="35" pageNumber="36">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFC1FF9EFABEF927FAEA6CF0" box="[1301,1404,1674,1696]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="35" pageNumber="36">Fig. 18E</figureCitation>
) of moderately size, reaching
<quantity id="0E4AEAF5FFC1FF9EFB2FF904FB4F6CEF" box="[1156,1241,1705,1727]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="mm" value="30.0">30 mm</quantity>
in length. Body translucent purple, with thin white band encircling the margin of notum and foot. Five unipinnate gill branches on notum. One large specimen (CASIZ
<date id="BD0C61D0FFC1FF9EFAA2F8A8FAC96D4B" box="[1289,1375,1797,1819]" pageId="35" pageNumber="36" value="37-07-12">071237</date>
) with seven gill branches. Gill branches light orange, with darker orange at their common base. Base of gill pocket deep violet. Bulb of rhinophores bright orange, with redder apex. Bulb with ~25 densely packed lamellae. Base of rhinophore sheath deep violet to purple.
</paragraph>
<caption id="9DCD1798FFC6FF99FF38FBF8FE4A6EF6" httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37" startId="36.[147,227,1109,1131]" targetBox="[211,1363,195,1068]" targetPageId="36">
<paragraph id="C90D4710FFC6FF99FF38FBF8FE4A6EF6" blockId="36.[147,1428,1109,1190]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF38FBF8FE9D6E3A" bold="true" box="[147,267,1109,1131]" pageId="36" pageNumber="37">Figure 20.</emphasis>
<taxonomicName id="0EB23C93FFC6FF99FEBDFBF8FC8F6E3B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[278,793,1109,1131]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="melanesica" status="sp. nov.">
<emphasis id="FBC69B02FFC6FF99FEBDFBF8FDAB6E3B" bold="true" box="[278,573,1109,1131]" italics="true" pageId="36" pageNumber="37">Hypselodoris melanesica</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC6FF99FC8BFBFBFCEF6E3B" box="[800,889,1110,1131]" pageId="36" pageNumber="37" rank="species">
<emphasis id="FBC69B02FFC6FF99FC8BFBFBFCEF6E3B" bold="true" box="[800,889,1110,1131]" pageId="36" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, paratype CASIZ 0 69787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587.
</paragraph>
</caption>
<paragraph id="C90D4710FFC6FF99FF3AFAEBFF726F2B" blockId="36.[145,761,1350,1403]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF3AFAEBFEA16F0B" box="[145,311,1350,1371]" italics="true" pageId="36" pageNumber="37">Mantle glands</emphasis>
: Subcutaneous mantle glands entirely absent.
</paragraph>
<paragraph id="C90D4710FFC6FF99FF3AFA33FB7C6F6C" blockId="36.[145,761,1438,1890]" lastBlockId="36.[809,1425,1226,1340]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF3AFA33FECD6FE4" box="[145,347,1438,1460]" italics="true" pageId="36" pageNumber="37">Buccal armature</emphasis>
: Muscular portion of buccal mass about twice the length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFC6FF99FD20FA57FD7C6C5F" box="[651,746,1530,1552]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20A</figureCitation>
). Rodlets narrow with short base and evenly curved, with single, acutely pointed apex. Radular formula of one
<typeStatus id="1609F9B2FFC6FF99FF3AF9FAFF606C3C" box="[145,246,1623,1644]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
(CASIZ 069787) 57 × 59.0.59. Rachidian row of teeth absent (
<figureCitation id="51895B95FFC6FF99FE80F9D8FE1F6CDA" box="[299,393,1653,1675]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20B</figureCitation>
). Innermost lateral teeth having one pointed denticle on inner side of bifid primary cusp, with another one or two outer denticles. Denticles not extending far beyond middle of elongate primary cusp. Next several laterals lacking inner triangular denticle but possessing three or four denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFC6FF99FDC1F880FD5D6D12" box="[618,715,1837,1859]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20C</figureCitation>
) all lacking inner denticles but having four or five rounded to triangular outer denticles and extended primary cusp. Outermost teeth having a narrower base and shorter tooth shape, with three rounded outer denticles (
<figureCitation id="51895B95FFC6FF99FC9AFA8BFC066F6B" box="[817,912,1318,1340]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20D</figureCitation>
), often larger than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFC6FF98FC82FAC8FD7D6B07" blockId="36.[809,1426,1381,1894]" lastBlockId="37.[163,779,198,343]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FC82FAC8FBB16F2B" box="[809,1063,1381,1403]" italics="true" pageId="36" pageNumber="37">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFC6FF99FC82FA29FC006FC9" box="[809,918,1412,1433]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
(CASIZ 069787) fully mature (
<figureCitation id="51895B95FFC6FF99FAB1FA29FA146FC9" box="[1306,1410,1412,1434]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="36" pageNumber="37">Fig. 13E</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick, and narrowing slightly as it transitions into muscular ejaculatory portion. Prostatic portion enveloping bursa copulatrix. Ejaculatory portion convoluted, narrow, entering short, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum seminis appressed against vagina in distal half of vaginal length. Moderately short uterine duct emerging from vagina close to base of bursa and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC7FF9BFF08FED3FDAE6B68" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="discussion">
<paragraph id="C90D4710FFC7FF98FF08FED3FE866BC6" blockId="37.[163,779,382,896]" box="[163,272,382,406]" pageId="37" pageNumber="38">
<heading id="9245F07CFFC7FF98FF08FED3FE866BC6" box="[163,272,382,406]" fontSize="10" level="4" pageId="37" pageNumber="38" reason="1">
<emphasis id="FBC69B02FFC7FF98FF08FED3FE866BC6" box="[163,272,382,406]" italics="true" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC7FF98FF08FE32FB1A683B" blockId="37.[163,779,382,896]" lastBlockId="37.[827,1443,198,895]" pageId="37" pageNumber="38">
In our phylogenetic analyses,
<taxonomicName id="0EB23C93FFC7FF98FE51FE32FD0C6BE3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[506,666,414,436]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FE51FE32FD0C6BE3" box="[506,666,414,436]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is always sister to
<taxonomicName id="0EB23C93FFC7FF98FEBBFE13FE086B82" authorityName="Collingwood" authorityYear="1881" box="[272,414,445,467]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FEBBFE13FE086B82" box="[272,414,445,467]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
but forms a distinct clade. In the ABGD analysis,
<taxonomicName id="0EB23C93FFC7FF98FE39FE71FDA06BA1" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[402,566,476,497]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FE39FE71FDA06BA1" box="[402,566,476,497]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is included in the same group as
<taxonomicName id="0EB23C93FFC7FF98FEFBFE56FE43685F" authorityName="Collingwood" authorityYear="1881" box="[336,469,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FEFBFE56FE43685F" box="[336,469,506,528]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
, thus suggesting that they should be considered as conspecific. Their uncorrected p-distances for the
<emphasis id="FBC69B02FFC7FF98FEDAFD95FE36681D" box="[369,416,568,589]" italics="true" pageId="37" pageNumber="38">COI</emphasis>
gene range between 2.0 and 2.6% different, right near the boundary for consideration of these two as distinct species. Although
<taxonomicName id="0EB23C93FFC7FF98FDC2FDDBFC9C68DA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[617,778,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FDC2FDDBFC9C68DA" box="[617,778,629,651]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
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resembles
<taxonomicName id="0EB23C93FFC7FF98FE89FD39FE3B68F9" authorityName="Collingwood" authorityYear="1881" box="[290,429,660,681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FE89FD39FE3B68F9" box="[290,429,660,681]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
in its shape and body colour, there are consistent morphological differences. In
<taxonomicName id="0EB23C93FFC7FF98FF08FD7FFEB968B6" authorityName="Collingwood" authorityYear="1881" box="[163,303,721,743]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FF08FD7FFEB968B6" box="[163,303,721,743]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
, the body colour is generally light pink to a deep purple. There may or may not be darker pigment at the base of the gill and the rhinophores. When present, the purple pigment is a wide band that is very diffuse, without distinctly defined edges. In contrast,
<taxonomicName id="0EB23C93FFC7FF98FEA7FCC6FE3C692F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[268,426,874,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FEA7FCC6FE3C692F" box="[268,426,874,896]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is always light purple in colour and always has well-defined narrow bands of darker pigment at the base of the rhinophores and gill. The bands at the base of the rhinophores of
<taxonomicName id="0EB23C93FFC7FF98FAA9FEA9FA346B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1282,1442,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAA9FEA9FA346B49" box="[1282,1442,260,281]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
always have a break in the band on the posterior side of the rhinophores. Internally,
<taxonomicName id="0EB23C93FFC7FF98FB09FEEFFAD36B06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1186,1349,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FB09FEEFFAD36B06" box="[1186,1349,321,343]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
(CASIZ 069787) has a radular formula of 65 × 68.0.68 vs. 77 × 97.0.
<quantity id="0E4AEAF5FFC7FF98FC1BFED2FC7B6BC4" box="[944,1005,382,404]" metricMagnitude="0" metricUnit="m" metricValue="2.4638" pageId="37" pageNumber="38" unit="in" value="97.0">97 in</quantity>
one specimen of
<taxonomicName id="0EB23C93FFC7FF98FB11FED2FAD26BC3" authorityName="Collingwood" authorityYear="1881" box="[1210,1348,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FB11FED2FAD26BC3" box="[1210,1348,382,404]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(CASIZ 083685). Also, the primary cusps of the radular teeth of
<taxonomicName id="0EB23C93FFC7FF98FC90FE11FC4F6B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,985,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FC90FE11FC4F6B81" box="[827,985,444,465]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
appear shorter than those of
<taxonomicName id="0EB23C93FFC7FF98FA8AFE11FA346B81" authorityName="Collingwood" authorityYear="1881" box="[1313,1442,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FA8AFE11FA346B81" box="[1313,1442,444,465]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFC7FF98FCE8FE77FC026BA0" box="[835,916,474,496]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="37" pageNumber="38">Figs 20</figureCitation>
,
<figureCitation id="51895B95FFC7FF98FC0BFE76FC2D6BA0" box="[928,955,475,496]" captionStart="Figure 21" captionStartId="37.[163,242,1850,1872]" captionTargetBox="[227,1380,938,1809]" captionTargetId="figure@37.[227,1380,938,1809]" captionTargetPageId="37" captionText="Figure 21. Hypselodoris bullockii (Collingwood, 1881), buccal mass. A, jaw rodlets, CASIZ 083685. B, inner lateral teeth, CASIZ 085905. C, middle lateral teeth, CASIZ 085905. D, outer lateral teeth, CASIZ 085905." httpUri="https://zenodo.org/record/2632820/files/figure.png" pageId="37" pageNumber="38">21</figureCitation>
). The reproductive system of
<taxonomicName id="0EB23C93FFC7FF98FAAEFE76FA356BBF" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1285,1443,474,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAAEFE76FA356BBF" box="[1285,1443,474,496]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
has a shorter penial bulb and a longer ejaculatory duct that is more convoluted than that found in
<taxonomicName id="0EB23C93FFC7FF98FA8BFDB5FA34687D" authorityName="Collingwood" authorityYear="1881" box="[1312,1442,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FA8BFDB5FA34687D" box="[1312,1442,536,557]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFC7FF98FCE8FD9BFC30681C" box="[835,934,566,588]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="37" pageNumber="38">Fig. 13F</figureCitation>
). Also,
<taxonomicName id="0EB23C93FFC7FF98FC54FD9AFB30681B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1023,1190,566,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FC54FD9AFB30681B" box="[1023,1190,566,588]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
has a longer vaginal duct than that of
<taxonomicName id="0EB23C93FFC7FF98FBA9FDFBFB13683A" authorityName="Collingwood" authorityYear="1881" box="[1026,1157,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FBA9FDFBFB13683A" box="[1026,1157,597,619]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFC7FF9BFCF8FDD9FDAE6B68" blockId="37.[827,1443,198,895]" lastBlockId="38.[145,761,198,312]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<taxonomicName id="0EB23C93FFC7FF98FCF8FDD9FBF868D9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[851,1134,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FCF8FDD9FBF868D9" box="[851,1134,628,649]" italics="true" pageId="37" pageNumber="38">Hypselodoris melanesica</emphasis>
</taxonomicName>
is known only from
<collectingCountry id="B1A50780FFC7FF98FAF1FDD9FC4C68F8" name="Papua New Guinea" pageId="37" pageNumber="38">Papua New Guinea</collectingCountry>
and the
<collectingCountry id="B1A50780FFC7FF98FBFFFD3FFABD68F8" box="[1108,1323,658,680]" name="Solomon Islands" pageId="37" pageNumber="38">Solomon Islands</collectingCountry>
, whereas
<taxonomicName id="0EB23C93FFC7FF98FC90FD1FFC5B6896" authorityName="Collingwood" authorityYear="1881" box="[827,973,689,711]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FC90FD1FFC5B6896" box="[827,973,689,711]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
is known from the western Pacific of
<collectingCountry id="B1A50780FFC7FF98FC90FD7DFC3F68B6" box="[827,937,720,742]" name="Australia" pageId="37" pageNumber="38">Australia</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FC19FD7DFBF668B6" box="[946,1120,720,742]" name="New Caledonia" pageId="37" pageNumber="38">New Caledonia</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FBC3FD7DFB4468B6" box="[1128,1234,720,742]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FB77FD7DFAD868B6" box="[1244,1358,720,742]" name="Indonesia" pageId="37" pageNumber="38">Indonesia</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FAFCFD7DFA0868B5" box="[1367,1438,720,741]" name="Japan" pageId="37" pageNumber="38">Japan</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FC90FD43FC0C6954" box="[827,922,750,772]" name="Taiwan" pageId="37" pageNumber="38">Taiwan</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FC02FD43FBAA6954" box="[937,1084,750,772]" name="Philippines" pageId="37" pageNumber="38">Philippines</collectingCountry>
,
<collectingCountry id="B1A50780FFC7FF98FBE1FD43FAB26954" box="[1098,1316,750,772]" name="Marshall Islands" pageId="37" pageNumber="38">Marshall Islands</collectingCountry>
(
<bibRefCitation id="AD233AE1FFC7FF98FA93FD43FC2D6972" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="37" pageNumber="38" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFC7FF98FC90FCA3FCE36972" box="[827,885,781,803]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
, 2008
</bibRefCitation>
) and
<collectingCountry id="B1A50780FFC7FF98FBABFCA0FBD26973" box="[1024,1092,781,803]" name="Palau" pageId="37" pageNumber="38">Palau</collectingCountry>
(present study).
<taxonomicName id="0EB23C93FFC7FF98FAA7FCA0FC2F6911" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAA7FCA0FC2F6911" italics="true" pageId="37" pageNumber="38">Hypselodoris melanesica</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC7FF98FC46FC81FBF96911" authorityName="Collingwood" authorityYear="1881" box="[1005,1135,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FC46FC81FBF96911" box="[1005,1135,812,833]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
are geographically isolated and have minor but consistent differences in their coloration and internal morphology spanning several organ systems. Based on these consistent differences, we consider
<taxonomicName id="0EB23C93FFC4FF9BFE88FF4BFE516AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,455,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC4FF9BFE88FF4BFE516AAA" box="[291,455,229,251]" italics="true" pageId="38" pageNumber="39">H. melanesica</emphasis>
</taxonomicName>
as a distinct species from
<taxonomicName id="0EB23C93FFC4FF9BFF3AFEA9FE8F6B49" authorityName="Collingwood" authorityYear="1881" box="[145,281,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC4FF9BFF3AFEA9FE8F6B49" box="[145,281,260,281]" italics="true" pageId="38" pageNumber="39">H. bullockii</emphasis>
</taxonomicName>
despite the fact that the ABGD analysis clusters these species as conspecifics.
</paragraph>
</subSubSection>
</treatment>
</document>

592
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<mods:identifier id="EA019E5A73874762357F2D3BCF2ED4BB" type="DOI">10.1093/zoolinnean/zly048</mods:identifier>
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<treatment id="411BF606FFC2FF9EFC65FED8FC9A6EC4" ID-DOI="http://doi.org/10.5281/zenodo.5980684" ID-GBIF-Taxon="156146739" ID-Zenodo-Dep="5980684" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4" httpUri="http://treatment.plazi.org/id/411BF606FFC2FF9EFC65FED8FC9A6EC4" lastPageId="35" lastPageNumber="36" pageId="32" pageNumber="33">
<subSubSection id="81A8149BFFC2FF9DFC65FED8FA856BFB" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="C90D4710FFC2FF9DFC65FED8FA856BFB" blockId="32.[934,1299,373,466]" pageId="32" pageNumber="33">
<taxonomicName id="0EB23C93FFC2FF9DFC65FED8FB3F6BFA" ID-CoL="3P34B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="35" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<heading id="9245F07CFFC2FF9DFC65FED8FB7D6BDB" allCaps="true" bold="true" box="[974,1259,373,396]" centered="true" fontSize="9" level="2" pageId="32" pageNumber="33" reason="4">
<emphasis id="FBC69B02FFC2FF9DFC65FED8FB7D6BDB" bold="true" box="[974,1259,373,396]" italics="true" pageId="32" pageNumber="33">HYPSELODORIS LACUNA</emphasis>
</heading>
<heading id="9245F07CFFC2FF9DFC0DFE3EFB3F6BFA" allCaps="true" box="[934,1193,403,427]" centered="true" fontSize="9" level="3" pageId="32" pageNumber="33" reason="4">GOSLINER &amp; JOHNSON</heading>
</taxonomicName>
<emphasis id="FBC69B02FFC2FF9DFB1BFE3AFA856BFB" bold="true" box="[1200,1299,403,427]" pageId="32" pageNumber="33">SP. NOV.</emphasis>
</paragraph>
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<subSubSection id="81A8149BFFC2FF9DFC63FE17FB676B82" box="[968,1265,442,466]" pageId="32" pageNumber="33" type="description">
<paragraph id="C90D4710FFC2FF9DFC63FE17FB676B82" blockId="32.[934,1299,373,466]" box="[968,1265,442,466]" pageId="32" pageNumber="33">
<heading id="9245F07CFFC2FF9DFC63FE17FB676B82" allCaps="true" box="[968,1265,442,466]" centered="true" fontSize="10" level="1" pageId="32" pageNumber="33" reason="1">(FIGS 2J, 13D, 18A–D, 19)</heading>
</paragraph>
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<paragraph id="C90D4710FFC2FF9DFC82FDAEFB8F6867" blockId="32.[809,1426,514,567]" pageId="32" pageNumber="33">
<emphasis id="FBC69B02FFC2FF9DFC82FDAEFCFA6848" box="[809,876,515,536]" italics="true" pageId="32" pageNumber="33">LSID</emphasis>
: urn:lsid:zoobank.org:act:CC5531BC-6A9E- 48F3-622959E3D128
</paragraph>
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<paragraph id="C90D4710FFC2FF9DFC82FDF8FC2568DA" blockId="32.[809,1425,597,711]" pageId="32" pageNumber="33">
<taxonomicName id="0EB23C93FFC2FF9DFC82FDF8FC51683A" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,967,597,618]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC2FF9DFC82FDF8FC51683A" box="[809,967,597,618]" italics="true" pageId="32" pageNumber="33">Hypselodoris</emphasis>
</taxonomicName>
sp. 11
<bibRefCitation id="AD233AE1FFC2FF9DFB88FDF8FA89683A" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1059,1311,597,619]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="32" pageNumber="33" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFC2FF9DFB3FFDFBFB45683A" box="[1172,1235,597,619]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
, 2008
</bibRefCitation>
: 270, top photograph.
</paragraph>
<paragraph id="C90D4710FFC2FF9DFC82FD3FFC646897" blockId="32.[809,1425,597,711]" pageId="32" pageNumber="33">
<taxonomicName id="0EB23C93FFC2FF9DFC82FD3FFC2A68F7" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,956,658,679]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC2FF9DFC82FD3FFC2A68F7" box="[809,956,658,679]" italics="true" pageId="32" pageNumber="33">Hypselodoris</emphasis>
</taxonomicName>
sp. 20
<bibRefCitation id="AD233AE1FFC2FF9DFBA7FD3FFB6668F8" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1036,1264,658,680]" journalOrPublisher="Jacksonville: New World Press" pageId="32" pageNumber="33" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFC2FF9DFBDFFD3EFB3A68F7" box="[1140,1196,658,680]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
, 2015
</bibRefCitation>
: 2639, bottom right photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC2FF9DFC82FD42FA046C8C" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="C90D4710FFC2FF9DFC82FD42FC456956" blockId="32.[809,1425,750,896]" box="[809,979,750,774]" pageId="32" pageNumber="33">
<heading id="9245F07CFFC2FF9DFC82FD42FC456956" box="[809,979,750,774]" fontSize="10" level="4" pageId="32" pageNumber="33" reason="1">
<emphasis id="FBC69B02FFC2FF9DFC82FD42FC456956" box="[809,979,750,774]" italics="true" pageId="32" pageNumber="33">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC2FF9DFC82FCA3FB4369D0" blockId="32.[809,1425,750,896]" pageId="32" pageNumber="33">
<materialsCitation id="79DA4D4DFFC2FF9DFC82FCA3FB4369D0" ID-GBIF-Occurrence="2235579089" collectingDate="2010-05-17" collectionCode="NMP" collectorName="T. Gosliner." country="Philippines" location="Philippines" pageId="32" pageNumber="33" specimenCode="CASIZ 182758" specimenCount="1" typeStatus="holotype">
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<emphasis id="FBC69B02FFC2FF9DFC82FCA3FC026973" box="[809,916,782,803]" italics="true" pageId="32" pageNumber="33">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFC2FF9DFC03FCA2FC716974" box="[936,999,783,804]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="32" pageNumber="33">NMP</collectionCode>
0 41280 (formerly
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), s u b s a m p l e d f o r m o l e c u l a r s t u d y, B e t h l e h e m 1 3.6 7 3 2 9°N, 1 2 0.8 4 0 9 3°E, T i n g l o y, B a t a n g a s,
<collectingCountry id="B1A50780FFC2FF9DFC82FCC7FC3A69D0" box="[809,940,874,896]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFC1CFCC6FBDD69D0" box="[951,1099,875,896]" pageId="32" pageNumber="33" value="2010-05-17">
<date id="BD0C61D0FFC2FF9DFC1CFCC6FBDD69D0" box="[951,1099,875,896]" pageId="32" pageNumber="33" value="2010-05-17">17 May 2010</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFBFEFCC6FB4369D0" box="[1109,1237,874,896]" pageId="32" pageNumber="33">T. Gosliner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFC2FF9DFC82FC07FA046C8C" blockId="32.[809,1426,937,1756]" pageId="32" pageNumber="33">
<materialsCitation id="79DA4D4DFFC2FF9DFC82FC07FC2B6E4B" ID-GBIF-Occurrence="2235579105" collectingDate="2015-03-31" collectionCode="CASIZ" collectorName="Puerto Galera &amp; P. J. Aristorenas" country="Philippines" latitude="13.51664" location="Coral Cove" longLatPrecision="1" longitude="120.99176" pageId="32" pageNumber="33" specimenCode="CASIZ 208587" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="paratype">
<typeStatus id="1609F9B2FFC2FF9DFC82FC07FC0869EF" box="[809,926,938,959]" pageId="32" pageNumber="33" type="paratype">
<emphasis id="FBC69B02FFC2FF9DFC82FC07FC0869EF" box="[809,926,938,959]" italics="true" pageId="32" pageNumber="33">Paratypes</emphasis>
</typeStatus>
:
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, one specimen, dissected,
<location id="CC6D11CBFFC2FF9DFC82FC65FC3C698D" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFC82FC65FC3C698D" box="[809,938,968,990]" country="Philippines" latitude="13.51664" longLatPrecision="1" longitude="120.99176" name="Coral Cove" pageId="32" pageNumber="33" stateProvince="Mindoro Oriental">Coral Cove</location>
,
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,
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,
<collectorName id="644722C6FFC2FF9DFB4CFC65FA1A698E" box="[1255,1420,968,990]" pageId="32" pageNumber="33">Puerto Galera</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFC82FC4BFC6A69AC" box="[809,1020,998,1020]" country="Philippines" name="Mindoro Oriental" pageId="32" pageNumber="33">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFBA2FC4BFB0469AC" box="[1033,1170,998,1020]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFB34FC4AFAC269AC" box="[1183,1364,998,1020]" pageId="32" pageNumber="33" value="2015-03-31">
<date id="BD0C61D0FFC2FF9DFB34FC4AFAC269AC" box="[1183,1364,998,1020]" pageId="32" pageNumber="33" value="2015-03-31">31 March 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFACAFC4AFC2B6E4B" pageId="32" pageNumber="33">P. J. Aristorenas</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFC61FBA8FAAD6E27" ID-GBIF-Occurrence="2235579114" collectingDate="2015-04-12" collectionCode="CASIZ" collectorName="Puerto Galera &amp; T. Gosliner" country="Philippines" latitude="13.5199" location="Channel" longLatPrecision="7" longitude="120.9604" pageId="32" pageNumber="33" specimenCode="CASIZ 208188" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFC61FBA8FB166E4A" box="[970,1152,1029,1050]" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 208188</specimenCode>
, one specimen, tissue removed for molecular study,
<collectingRegion id="0B7689F2FFC2FF9DFB09FB89FA816E69" box="[1186,1303,1060,1081]" country="Philippines" name="Batangas" pageId="32" pageNumber="33">Batangas</collectingRegion>
<location id="CC6D11CBFFC2FF9DFA89FB89FA1A6E6A" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFA89FB89FA1A6E6A" box="[1314,1420,1060,1082]" country="Philippines" latitude="13.5199" longLatPrecision="7" longitude="120.9604" name="Channel" pageId="32" pageNumber="33" stateProvince="Batangas">Channel</location>
,
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,
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,
<collectorName id="644722C6FFC2FF9DFBCBFBEEFA8F6E08" box="[1120,1305,1090,1112]" pageId="32" pageNumber="33">Puerto Galera</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFA83FBEFFC1F6E27" country="Philippines" name="Mindoro Oriental" pageId="32" pageNumber="33">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFC3FFBCCFB826E27" box="[916,1044,1121,1143]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFBB4FBCCFB2F6E27" box="[1055,1209,1121,1143]" pageId="32" pageNumber="33" value="2015-04-12">
<date id="BD0C61D0FFC2FF9DFBB4FBCCFB2F6E27" box="[1055,1209,1121,1143]" pageId="32" pageNumber="33" value="2015-04-12">12 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFB68FBCFFAAD6E27" box="[1219,1339,1121,1143]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFAEDFBCCFC316EA2" ID-GBIF-Occurrence="2235579069" collectingDate="2015-04-13" collectionCode="CASIZ" collectorName="Puerto Galera &amp; T. Gosliner" country="Philippines" latitude="13.5223" location="Channel" longLatPrecision="7" longitude="120.9485" pageId="32" pageNumber="33" specimenCode="CASIZ 208190" specimenCount="1" stateProvince="Manila" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFAEDFBCCFC166EC5" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 208190</specimenCode>
, one specimen, tissue removed for molecular study,
<collectingRegion id="0B7689F2FFC2FF9DFCDAFB33FC576EE4" box="[881,961,1182,1204]" country="Philippines" name="Manila" pageId="32" pageNumber="33">Manila</collectingRegion>
<location id="CC6D11CBFFC2FF9DFC62FB33FBBC6EE4" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFC62FB33FBBC6EE4" box="[969,1066,1182,1204]" country="Philippines" latitude="13.5223" longLatPrecision="7" longitude="120.9485" name="Channel" pageId="32" pageNumber="33" stateProvince="Manila">Channel</location>
,
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,
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,
<collectorName id="644722C6FFC2FF9DFAEEFB32FCEF6E83" pageId="32" pageNumber="33">Puerto Galera</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFC2EFB10FBC76E83" box="[901,1105,1213,1235]" country="Philippines" name="Mindoro Oriental" pageId="32" pageNumber="33">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFBF6FB10FB746E83" box="[1117,1250,1213,1235]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFB45FB10FA1A6E83" box="[1262,1420,1213,1235]" pageId="32" pageNumber="33" value="2015-04-13">
<date id="BD0C61D0FFC2FF9DFB45FB10FA1A6E83" box="[1262,1420,1213,1235]" pageId="32" pageNumber="33" value="2015-04-13">13 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFC82FB71FC316EA2" box="[809,935,1244,1266]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFC19FB71FB4C6F7F" ID-GBIF-Occurrence="2235579138" collectingDate="2016-04-18" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" location="Bethlehem" pageId="32" pageNumber="33" specimenCode="CASIZ 217345" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFC19FB71FBCB6EA2" box="[946,1117,1244,1266]" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 217345</specimenCode>
, one specimen,
<location id="CC6D11CBFFC2FF9DFAB8FB71FA076EA2" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFAB8FB71FA076EA2" box="[1299,1425,1244,1266]" country="Philippines" name="Bethlehem" pageId="32" pageNumber="33">Bethlehem</location>
1 3.6 7 3 2 9°N, 1 2 0.8 4 0 9 3°E, T i n g l o y, B a t a n g a s,
<collectingCountry id="B1A50780FFC2FF9DFC82FAB4FC3A6F7F" box="[809,940,1305,1327]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFC1CFAB4FBC26F7F" box="[951,1108,1305,1327]" pageId="32" pageNumber="33" value="2016-04-18">
<date id="BD0C61D0FFC2FF9DFC1CFAB4FBC26F7F" box="[951,1108,1305,1327]" pageId="32" pageNumber="33" value="2016-04-18">18 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFBF4FAB7FB4C6F7F" box="[1119,1242,1305,1327]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFB4EFAB4FC4A6FDB" ID-GBIF-Occurrence="2235579118" collectingDate="2008-04-17" collectionCode="CASIZ" collectorName="Tingloy &amp; T. Gosliner" country="Philippines" latitude="13.67329" location="Bethlehem" longLatPrecision="1" longitude="120.84093" pageId="32" pageNumber="33" specimenCode="CASIZ 177617" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFB4EFAB4FA1A6F7F" box="[1253,1420,1305,1327]" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 177617</specimenCode>
, one specimen, dissected,
<location id="CC6D11CBFFC2FF9DFBC2FA95FB666F1E" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFBC2FA95FB666F1E" box="[1129,1264,1336,1358]" country="Philippines" latitude="13.67329" longLatPrecision="1" longitude="120.84093" name="Bethlehem" pageId="32" pageNumber="33" stateProvince="Batangas">Bethlehem</location>
<geoCoordinate id="AC8621D7FFC2FF9DFB50FA95FA1A6F1E" box="[1275,1420,1336,1358]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="13.67329">13.67329°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC2FF9DFC82FAFAFC2A6F3C" box="[809,956,1367,1388]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="120.84093">120.84093°E</geoCoordinate>
,
<collectorName id="644722C6FFC2FF9DFC6CFAFBFB8B6F3C" box="[967,1053,1366,1388]" pageId="32" pageNumber="33">Tingloy</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFB82FAFAFB016F3C" box="[1065,1175,1367,1388]" country="Philippines" name="Batangas" pageId="32" pageNumber="33">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFB08FAFBFAB16F3C" box="[1187,1319,1366,1388]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFA98FAFAFCF76FDA" pageId="32" pageNumber="33" value="2008-04-17">
<date id="BD0C61D0FFC2FF9DFA98FAFAFCF76FDA" pageId="32" pageNumber="33" value="2008-04-17">17 April 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFCCCFADBFC4A6FDB" box="[871,988,1397,1419]" pageId="32" pageNumber="33">T.Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFC48FAD8FA1B6F98" ID-GBIF-Occurrence="2235579053" collectingDate="2015-04-26" collectionCode="CASIZ" collectorName="Puerto Galera &amp; T. M. Gosliner" country="Philippines" latitude="13.51286" location="Boulders" longLatPrecision="1" longitude="120.98309" pageId="32" pageNumber="33" specimenCode="CASIZ 208652" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFC48FAD8FB126FDB" box="[995,1156,1397,1419]" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 208652</specimenCode>
, one specimen,
<location id="CC6D11CBFFC2FF9DFA8CFAD8FA1A6FDB" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFA8CFAD8FA1A6FDB" box="[1319,1420,1397,1419]" country="Philippines" latitude="13.51286" longLatPrecision="1" longitude="120.98309" name="Boulders" pageId="32" pageNumber="33" stateProvince="Mindoro Oriental">Boulders</location>
,
<geoCoordinate id="AC8621D7FFC2FF9DFC82FA39FC216FFA" box="[809,951,1428,1450]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="13.51286">13.51286°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC2FF9DFC6EFA39FBF66FF9" box="[965,1120,1428,1449]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="120.98309">120.98309°E</geoCoordinate>
,
<collectorName id="644722C6FFC2FF9DFBC5FA39FA886FFA" box="[1134,1310,1428,1450]" pageId="32" pageNumber="33">Puerto Galera</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFA87FA39FC066F98" country="Philippines" name="Mindoro Oriental" pageId="32" pageNumber="33">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFC36FA1FFBB06F98" box="[925,1062,1458,1480]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFB98FA1FFB4E6F98" box="[1075,1240,1458,1480]" pageId="32" pageNumber="33" value="2015-04-26">
<date id="BD0C61D0FFC2FF9DFB98FA1FFB4E6F98" box="[1075,1240,1458,1480]" pageId="32" pageNumber="33" value="2015-04-26">26 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFB4FFA1EFA1B6F98" box="[1252,1421,1458,1480]" pageId="32" pageNumber="33">T. M. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFC82FA7CFB766C74" ID-GBIF-Occurrence="2235579057" collectingDate="2015-04-08" collectionCode="CASIZ" collectorName="Puerto Galera &amp; T. M. Gosliner" country="Philippines" latitude="13.52496" location="La Laguna" longLatPrecision="1" longitude="120.97114" pageId="32" pageNumber="33" specimenCode="CASIZ 208646" specimenCount="1" stateProvince="Laguna" typeStatus="paratype">
<specimenCode id="9914EF6BFFC2FF9DFC82FA7CFC596FB7" box="[809,975,1489,1511]" collectionCode="CASIZ" pageId="32" pageNumber="33">CASIZ 208646</specimenCode>
, one specimen,
<collectingRegion id="0B7689F2FFC2FF9DFBD4FA7FFB6A6FB7" box="[1151,1276,1490,1511]" country="Philippines" name="Laguna" pageId="32" pageNumber="33">La Laguna</collectingRegion>
,
<geoCoordinate id="AC8621D7FFC2FF9DFAA3FA7CFA1B6FB7" box="[1288,1421,1489,1511]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="13.52496">13.52496°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC2FF9DFC82FA5DFC5B6C55" box="[809,973,1520,1541]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="120.97114">120.97114°E</geoCoordinate>
,
<collectorName id="644722C6FFC2FF9DFC76FA5DFB0E6C56" box="[989,1176,1520,1542]" pageId="32" pageNumber="33">Puerto Galera</collectorName>
,
<collectingRegion id="0B7689F2FFC2FF9DFB03FA5DFA1A6C56" box="[1192,1420,1520,1542]" country="Philippines" name="Mindoro Oriental" pageId="32" pageNumber="33">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFC82F9A3FC3F6C74" box="[809,937,1550,1572]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFC18F9A2FBAB6C74" box="[947,1085,1550,1572]" pageId="32" pageNumber="33" value="2015-04-08">
<date id="BD0C61D0FFC2FF9DFC18F9A2FBAB6C74" box="[947,1085,1550,1572]" pageId="32" pageNumber="33" value="2015-04-08">8 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFBEDF9A2FB766C74" box="[1094,1248,1550,1572]" pageId="32" pageNumber="33">T. M. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFB42F9A2FAAF6C32" ID-GBIF-Occurrence="2235579098" collectingDate="1989-07-30" collectionCode="CASIZ" collectorName="T. Gosliner" country="Papua New Guinea" elevation="10" location="Pig" municipality="Tab" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC2FF9DFB42F9A2FAA26C74" box="[1257,1332,1551,1572]" pageId="32" pageNumber="33">CASIZ</collectionCode>
0 69756, north end of
<location id="CC6D11CBFFC2FF9DFC6DF980FC7B6C13" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFC6DF980FC7B6C13" box="[966,1005,1581,1603]" country="Papua New Guinea" municipality="Tab" name="Pig" pageId="32" pageNumber="33" stateProvince="Madang">Pig</location>
(
<collectingMunicipality id="2969DD6AFFC2FF9DFC55F980FBBA6C13" box="[1022,1068,1581,1603]" pageId="32" pageNumber="33">Tab</collectingMunicipality>
) Island,
<collectingRegion id="0B7689F2FFC2FF9DFB33F980FB6C6C13" box="[1176,1274,1581,1603]" country="Papua New Guinea" name="Madang" pageId="32" pageNumber="33">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFAA3F983FCE96C32" name="Papua New Guinea" pageId="32" pageNumber="33">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFC2FF9DFC21F9E1FC526C31" box="[906,964,1612,1633]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="m" value="10.0">
<quantity id="0E4AEAF5FFC2FF9DFC21F9E1FC526C31" box="[906,964,1612,1633]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="m" value="10.0">10 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFC2FF9DFBB0F9E1FB276C31" box="[1051,1201,1612,1634]" pageId="32" pageNumber="33" value="1989-07-30">
<date id="BD0C61D0FFC2FF9DFBB0F9E1FB276C31" box="[1051,1201,1612,1634]" pageId="32" pageNumber="33" value="1989-07-30">30 July 1989</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFB17F9E1FAAF6C32" box="[1212,1337,1612,1634]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFAEFF9E1FAAF6CCF" ID-GBIF-Occurrence="2235579066" collectingDate="1989-07-16" collectionCode="CASIZ" collectorName="T. Gosliner" country="Papua New Guinea" location="Cement Mixer Reef" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC2FF9DFAEFF9E1FA066C31" box="[1348,1424,1612,1633]" pageId="32" pageNumber="33">CASIZ</collectionCode>
0 68776, one specimen,
<location id="CC6D11CBFFC2FF9DFB92F9C6FAB66CD0" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFB92F9C6FAB66CD0" box="[1081,1312,1642,1664]" country="Papua New Guinea" name="Cement Mixer Reef" pageId="32" pageNumber="33" stateProvince="Madang">Cement Mixer Reef</location>
,
<collectingRegion id="0B7689F2FFC2FF9DFA87F9C7FA1B6CD0" box="[1324,1421,1642,1664]" country="Papua New Guinea" name="Madang" pageId="32" pageNumber="33">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFC82F927FB9A6CCF" box="[809,1036,1673,1695]" name="Papua New Guinea" pageId="32" pageNumber="33">Papua New Guinea</collectingCountry>
,
<collectingDate id="AD489838FFC2FF9DFBB3F924FB396CCE" box="[1048,1199,1673,1695]" pageId="32" pageNumber="33" value="1989-07-16">
<date id="BD0C61D0FFC2FF9DFBB3F924FB396CCE" box="[1048,1199,1673,1695]" pageId="32" pageNumber="33" value="1989-07-16">16 July 1989</date>
</collectingDate>
,
<collectorName id="644722C6FFC2FF9DFB11F927FAAF6CCF" box="[1210,1337,1673,1695]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFC2FF9DFAEFF924FA186C8C" ID-GBIF-Occurrence="2235579065" collectingDate="1990-11-23" collectionCode="CASIZ" collectorName="T. Gosliner" country="Papua New Guinea" elevation="7" location="Barracuda Point" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC2FF9DFAEFF924FA076CCE" box="[1348,1425,1673,1694]" pageId="32" pageNumber="33">CASIZ</collectionCode>
0 75842, one specimen,
<location id="CC6D11CBFFC2FF9DFB88F905FB4D6CEE" LSID="urn:lsid:plazi:treatment:411BF606FFC2FF9EFC65FED8FC9A6EC4:CC6D11CBFFC2FF9DFB88F905FB4D6CEE" box="[1059,1243,1704,1726]" country="Papua New Guinea" name="Barracuda Point" pageId="32" pageNumber="33" stateProvince="Madang">Barracuda Point</location>
,
<collectingRegion id="0B7689F2FFC2FF9DFB4FF905FAD76CEE" box="[1252,1345,1704,1726]" country="Papua New Guinea" name="Madang" pageId="32" pageNumber="33">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFC2FF9DFAE2F905FC236C8C" name="Papua New Guinea" pageId="32" pageNumber="33">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFC2FF9DFC14F96AFC7F6C8C" box="[959,1001,1735,1756]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="m" value="7.0">
<quantity id="0E4AEAF5FFC2FF9DFC14F96AFC7F6C8C" box="[959,1001,1735,1756]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="m" value="7.0">7 m</quantity>
</elevation>
depth,
<date id="BD0C61D0FFC2FF9DFB90F96AFA9A6C8C" box="[1083,1292,1734,1756]" pageId="32" pageNumber="33" value="1990-11-23">
<collectingDate id="AD489838FFC2FF9DFB90F96AFA9A6C8C" box="[1083,1292,1734,1756]" pageId="32" pageNumber="33" value="1990-11-23">23 November 1990</collectingDate>
</date>
,
<collectorName id="644722C6FFC2FF9DFABEF96AFA186C8C" box="[1301,1422,1734,1756]" pageId="32" pageNumber="33">T. Gosliner</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC2FF9DFC82F8AEFA076D27" pageId="32" pageNumber="33" type="distribution">
<paragraph id="C90D4710FFC2FF9DFC82F8AEFBF06D4B" blockId="32.[809,1425,1795,1911]" box="[809,1126,1795,1819]" pageId="32" pageNumber="33">
<heading id="9245F07CFFC2FF9DFC82F8AEFBF06D4B" box="[809,1126,1795,1819]" fontSize="10" level="4" pageId="32" pageNumber="33" reason="1">
<emphasis id="FBC69B02FFC2FF9DFC82F8AEFBF06D4B" box="[809,1126,1795,1819]" italics="true" pageId="32" pageNumber="33">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC2FF9DFC82F889FA076D27" blockId="32.[809,1425,1795,1911]" pageId="32" pageNumber="33">
Known from the western Indian Ocean of Aldabra Atoll to the western Pacific of
<collectingCountry id="B1A50780FFC2FF9DFB9AF8EEFB076D08" box="[1073,1169,1859,1880]" name="Vanuatu" pageId="32" pageNumber="33">Vanuatu</collectingCountry>
,
<collectingCountry id="B1A50780FFC2FF9DFB30F8EFFA9E6D08" box="[1179,1288,1858,1880]" name="Indonesia" pageId="32" pageNumber="33">Indonesia</collectingCountry>
,
<collectingCountry id="B1A50780FFC2FF9DFAB9F8EEFCED6D27" name="Papua New Guinea" pageId="32" pageNumber="33">Papua New Guinea</collectingCountry>
, the
<collectingCountry id="B1A50780FFC2FF9DFC06F8CCFBBF6D27" box="[941,1065,1889,1911]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
and
<collectingCountry id="B1A50780FFC2FF9DFBF6F8CFFB366D27" box="[1117,1184,1890,1911]" name="Japan" pageId="32" pageNumber="33">Japan</collectingCountry>
(
<bibRefCitation id="AD233AE1FFC2FF9DFB06F8CCFA156D26" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1197,1411,1889,1911]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="32" pageNumber="33" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFC2FF9DFABBF8CFFAD26D26" box="[1296,1348,1889,1911]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
, 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="9DCD1798FFC3FF9CFF08F984FB876D17" httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="33" pageNumber="34" startId="33.[163,244,1577,1599]" targetBox="[323,1283,196,1537]" targetPageId="33">
<paragraph id="C90D4710FFC3FF9CFF08F984FB876D17" blockId="33.[163,1444,1577,1863]" pageId="33" pageNumber="34">
<emphasis id="FBC69B02FFC3FF9CFF08F984FE8A6C6E" bold="true" box="[163,284,1577,1599]" pageId="33" pageNumber="34">Figure 18.</emphasis>
Living animals. A,
<taxonomicName id="0EB23C93FFC3FF9CFE5EF984FC5F6C6F" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[501,969,1577,1599]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFE5EF984FD7A6C6F" bold="true" box="[501,748,1577,1599]" italics="true" pageId="33" pageNumber="34">Hypselodoris lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFC7AF987FBBD6C6F" box="[977,1067,1578,1599]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFC7AF987FBBD6C6F" bold="true" box="[977,1067,1578,1599]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B,
<taxonomicName id="0EB23C93FFC3FF9CFE16F9EAFC806C0D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[445,790,1606,1629]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFE16F9EAFDA36C0C" bold="true" box="[445,565,1606,1628]" italics="true" pageId="33" pageNumber="34">H. lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFCB6F9EAFCEE6C0C" box="[797,888,1607,1628]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFCB6F9EAFCEE6C0C" bold="true" box="[797,888,1607,1628]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C,
<taxonomicName id="0EB23C93FFC3FF9CFE0EF9C9FD7B6C2A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[421,749,1636,1658]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFE0EF9C9FD816C2A" bold="true" box="[421,535,1636,1658]" italics="true" pageId="33" pageNumber="34">H. lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFD58F9C9FCDF6C29" box="[755,841,1636,1657]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFD58F9C9FCDF6C29" bold="true" box="[755,841,1636,1657]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D,
<taxonomicName id="0EB23C93FFC3FF9CFF69F92FFD8A6CC7" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[194,540,1665,1687]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFF69F92FFEAC6CC7" bold="true" box="[194,314,1665,1687]" italics="true" pageId="33" pageNumber="34">H. lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFD8FF92FFDE96CC7" box="[548,639,1666,1687]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFD8FF92FFDE96CC7" bold="true" box="[548,639,1666,1687]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E,
<taxonomicName id="0EB23C93FFC3FF9CFF6AF933FD456CE5" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[193,723,1694,1717]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="melanesica" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFF6AF933FE796CE4" bold="true" box="[193,495,1694,1716]" italics="true" pageId="33" pageNumber="34">Hypselodoris melanesica</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFD71F932FCA06CE4" box="[730,822,1695,1716]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFD71F932FCA06CE4" bold="true" box="[730,822,1695,1716]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 190823, Madang, Papua New Guinea. F,
<taxonomicName id="0EB23C93FFC3FF9CFF12F911FD066C82" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[185,656,1724,1746]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFF12F911FE286C82" bold="true" box="[185,446,1724,1746]" italics="true" pageId="33" pageNumber="34">Hypselodoris paradisa</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFD3EF911FD7C6C81" box="[661,746,1724,1745]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFD3EF911FD7C6C81" bold="true" box="[661,746,1724,1745]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G,
<taxonomicName id="0EB23C93FFC3FF9CFE0EF974FCC16CBF" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[421,855,1753,1775]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="perii" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFE0EF974FDE86CBE" bold="true" box="[421,638,1753,1775]" italics="true" pageId="33" pageNumber="34">Hypselodoris perii</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFCF5F977FC206CBF" box="[862,950,1754,1775]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFCF5F977FC206CBF" bold="true" box="[862,950,1754,1775]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, NMP 0 41281, Mabini, Batangas, Philippines. H,
<taxonomicName id="0EB23C93FFC3FF9CFF6AF95BFDC36D5D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[193,597,1782,1805]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFF6AF95BFE146D5C" bold="true" box="[193,386,1782,1804]" italics="true" pageId="33" pageNumber="34">Hypselodoris roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFDF0F95AFD276D5C" box="[603,689,1783,1804]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFDF0F95AFD276D5C" bold="true" box="[603,689,1783,1804]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I,
<taxonomicName id="0EB23C93FFC3FF9CFF13F8B9FE4F6D7A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[184,473,1812,1834]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFF13F8B9FE966D79" bold="true" box="[184,256,1812,1833]" italics="true" pageId="33" pageNumber="34">H. roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFE4BF8B9FDAE6D79" box="[480,568,1812,1833]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFE4BF8B9FDAE6D79" bold="true" box="[480,568,1812,1833]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J,
<taxonomicName id="0EB23C93FFC3FF9CFAF0F8B9FEE46D17" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFC3FF9CFAF0F8B9FA356D79" bold="true" box="[1371,1443,1812,1833]" italics="true" pageId="33" pageNumber="34">H. roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC3FF9CFED3F89FFE586D17" box="[376,462,1842,1863]" pageId="33" pageNumber="34" rank="species">
<emphasis id="FBC69B02FFC3FF9CFED3F89FFE586D17" bold="true" box="[376,462,1842,1863]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 186098, Mabini, Tingloy, Philippines.
</paragraph>
</caption>
<caption id="9DCD1798FFC0FF9FFF3AFC6AFE0F6E48" httpUri="https://zenodo.org/record/2632814/files/figure.png" pageId="34" pageNumber="35" startId="34.[145,223,967,989]" targetBox="[305,1265,197,927]" targetPageId="34">
<paragraph id="C90D4710FFC0FF9FFF3AFC6AFE0F6E48" blockId="34.[145,1426,967,1048]" pageId="34" pageNumber="35">
<emphasis id="FBC69B02FFC0FF9FFF3AFC6AFE90698C" bold="true" box="[145,262,967,989]" pageId="34" pageNumber="35">Figure 19.</emphasis>
<taxonomicName id="0EB23C93FFC0FF9FFEBAFC6AFD42698D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[273,724,967,989]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFC0FF9FFEBAFC6AFE69698D" bold="true" box="[273,511,967,989]" italics="true" pageId="34" pageNumber="35">Hypselodoris lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC0FF9FFD71FC6AFCA6698C" box="[730,816,967,988]" pageId="34" pageNumber="35" rank="species">
<emphasis id="FBC69B02FFC0FF9FFD71FC6AFCA6698C" bold="true" box="[730,816,967,988]" pageId="34" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature. A, jaw rodlets, paratype, CASIZ 208587. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587.
</paragraph>
</caption>
<subSubSection id="81A8149BFFC0FF9FFF3AFB91FE196EFF" pageId="34" pageNumber="35" type="etymology">
<paragraph id="C90D4710FFC0FF9FFF3AFB91FE846E04" blockId="34.[145,761,1084,1200]" box="[145,274,1084,1108]" pageId="34" pageNumber="35">
<heading id="9245F07CFFC0FF9FFF3AFB91FE846E04" box="[145,274,1084,1108]" fontSize="10" level="4" pageId="34" pageNumber="35" reason="1">
<emphasis id="FBC69B02FFC0FF9FFF3AFB91FE846E04" box="[145,274,1084,1108]" italics="true" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC0FF9FFF3AFBF1FE196EFF" blockId="34.[145,761,1084,1200]" pageId="34" pageNumber="35">
This species is named
<taxonomicName id="0EB23C93FFC0FF9FFE30FBF1FD1C6E21" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[411,650,1116,1137]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFC0FF9FFE30FBF1FD1C6E21" box="[411,650,1116,1137]" italics="true" pageId="34" pageNumber="35">Hypselodoris lacuna</emphasis>
</taxonomicName>
based on the translucent areas on the notum that resemble holes in the body wall.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC0FF9EFF3AFB7AFD01687E" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="C90D4710FFC0FF9FFF3AFB7AFE8B6EBE" blockId="34.[145,761,1239,1722]" box="[145,285,1239,1262]" pageId="34" pageNumber="35">
<heading id="9245F07CFFC0FF9FFF3AFB7AFE8B6EBE" box="[145,285,1239,1262]" fontSize="10" level="4" pageId="34" pageNumber="35" reason="1">
<emphasis id="FBC69B02FFC0FF9FFF3AFB7AFE8B6EBE" box="[145,285,1239,1262]" italics="true" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC0FF9FFF3AFB5AFF6F6CEA" blockId="34.[145,761,1239,1722]" pageId="34" pageNumber="35">
<emphasis id="FBC69B02FFC0FF9FFF3AFB5AFE1B6F5C" box="[145,397,1271,1292]" italics="true" pageId="34" pageNumber="35">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFC0FF9FFDC2FB5AFD2E6F5C" box="[617,696,1271,1293]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="34" pageNumber="35">Fig. 18</figureCitation>
A–D) small, reaching
<quantity id="0E4AEAF5FFC0FF9FFEE1FABBFE0F6F7B" box="[330,409,1302,1323]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="mm" value="12.0">12 mm</quantity>
in length. Body opaque white, with round translucent areas on notum that resemble holes in body wall. Two large, translucent circles situated posterior to rhinophores, and smaller circles located over surface of notum. Small black spot located in the centre of most smaller translucent circles. Ring of blue spots present along submarginal area of notum. Centre of spot darker than outer diffuse area. Blue spots also present on posterior end of foot. Gill branches white, with reddish tip. Seven unipinnate gill branches present in all specimens examined. Perfoliate rhinophores white basally and bright red apically, bearing about eight or nine densely arranged lamellae.
</paragraph>
<paragraph id="C90D4710FFC0FF9FFF3AF94EFE1F6D05" blockId="34.[145,761,1763,1877]" pageId="34" pageNumber="35">
<emphasis id="FBC69B02FFC0FF9FFF3AF94EFEAF6CA8" box="[145,313,1763,1784]" italics="true" pageId="34" pageNumber="35">Mantle glands</emphasis>
: Subcutaneous mantle glands simple, rounded in shape (
<figureCitation id="51895B95FFC0FF9FFECBF8AFFE3D6D47" box="[352,427,1794,1816]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="34" pageNumber="35">Fig. 2J</figureCitation>
). Six to twelve glands present posteriorly in seven specimens examined. Anterior and lateral glands absent.
</paragraph>
<paragraph id="C90D4710FFC0FF9FFC82FB91FB5E6C2A" blockId="34.[809,1425,1084,1658]" pageId="34" pageNumber="35">
<emphasis id="FBC69B02FFC0FF9FFC82FB91FC656E02" box="[809,1011,1084,1106]" italics="true" pageId="34" pageNumber="35">Buccal armature</emphasis>
: Muscular portion of buccal mass much smaller than oral tube. Buccal mass consisting of chitinous labial cuticle at anterior end of muscular portion of buccal mass. Jaws bearing numerous rodlets, narrow and acutely pointed, largely undivided but with an occasional bifid cusp (
<figureCitation id="51895B95FFC0FF9FFBEFFB78FB346EBB" box="[1092,1186,1237,1259]" captionStart="Figure 19" captionStartId="34.[145,223,967,989]" captionTargetBox="[305,1265,197,927]" captionTargetId="figure@34.[305,1265,197,927]" captionTargetPageId="34" captionText="Figure 19. Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, paratype, CASIZ 208587. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632814/files/figure.png" pageId="34" pageNumber="35">Fig. 19A</figureCitation>
). Radular formula of
<typeStatus id="1609F9B2FFC0FF9FFC82FB58FC196F5A" box="[809,911,1269,1290]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
(CASIZ 208587) 46 × 36.0.36. Rachidian row of teeth absent (
<figureCitation id="51895B95FFC0FF9FFC76FABEFBAC6F78" box="[989,1082,1299,1321]" captionStart="Figure 19" captionStartId="34.[145,223,967,989]" captionTargetBox="[305,1265,197,927]" captionTargetId="figure@34.[305,1265,197,927]" captionTargetPageId="34" captionText="Figure 19. Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, paratype, CASIZ 208587. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632814/files/figure.png" pageId="34" pageNumber="35">Fig. 19B</figureCitation>
). Innermost lateral teeth with one or two small, triangular denticles on inner side of bifid primary cusps and one outer denticle. Inner cusp of bifid cusp much longer than outer one. Next several laterals lacking inner and outer denticles on sides of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFC0FF9FFB44FA01FAC76F91" box="[1263,1361,1452,1474]" captionStart="Figure 19" captionStartId="34.[145,223,967,989]" captionTargetBox="[305,1265,197,927]" captionTargetId="figure@34.[305,1265,197,927]" captionTargetPageId="34" captionText="Figure 19. Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, paratype, CASIZ 208587. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632814/files/figure.png" pageId="34" pageNumber="35">Fig. 19C</figureCitation>
) also lacking inner denticles, but with a single triangular outer denticle or entirely lacking denticles. Outer cusp of bifid cusp much shorter than inner one. Outer teeth lacking inner denticles, with up to four triangular outer denticles (
<figureCitation id="51895B95FFC0FF9FFC09F9E8FB906C0B" box="[930,1030,1605,1627]" captionStart="Figure 19" captionStartId="34.[145,223,967,989]" captionTargetBox="[305,1265,197,927]" captionTargetId="figure@34.[305,1265,197,927]" captionTargetPageId="34" captionText="Figure 19. Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, paratype, CASIZ 208587. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632814/files/figure.png" pageId="34" pageNumber="35">Fig. 19D</figureCitation>
). Outermost teeth with narrower base and more elongate tooth shape.
</paragraph>
<paragraph id="C90D4710FFC0FF9EFC82F90EFD01687E" blockId="34.[809,1425,1699,1874]" lastBlockId="35.[163,780,198,558]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="FBC69B02FFC0FF9FFC82F90EFB9C6CE8" box="[809,1034,1699,1720]" italics="true" pageId="34" pageNumber="35">Reproductive system</emphasis>
: Reproductive organs fully mature (
<figureCitation id="51895B95FFC0FF9FFC9AF96CFC076C87" box="[817,913,1729,1751]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="34" pageNumber="35">Fig. 13D</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into the oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens curved and thick, narrowing gradually while transitioning into narrow, muscular ejaculatory portion. Ejaculatory portion curving into segment entering short, wider penial bulb. Penial bulb adjacent to curved, narrow vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Elongate, curved vagina leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Relatively short uterine duct emerging from vagina distal to junction of receptaculum and bursa.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFF08FDF8FC9A6EC4" pageId="35" pageNumber="36" type="discussion">
<paragraph id="C90D4710FFC1FF9EFF08FDF8FE86683D" blockId="35.[163,780,597,1172]" box="[163,272,597,621]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFF08FDF8FE86683D" box="[163,272,597,621]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFF08FDF8FE86683D" box="[163,272,597,621]" italics="true" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFF08FDD8FC9A6EC4" blockId="35.[163,780,597,1172]" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFF08FDD8FE0B68DA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,413,629,650]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFC1FF9EFF08FDD8FE0B68DA" box="[163,413,629,650]" italics="true" pageId="35" pageNumber="36">Hypselodoris lacuna</emphasis>
</taxonomicName>
is unique among members of this genus in having translucent circles and a ring of small blue spots along the submarginal area of the notum. Its colour pattern more closely resembles that of
<emphasis id="FBC69B02FFC1FF9EFF6BFD5DFE216955" box="[192,439,752,773]" italics="true" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFF6BFD5DFEC16955" authorityName="Alder &amp; Hancock" authorityYear="1855" box="[192,343,752,773]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">Chromodoris</taxonomicName>
aspersa
</emphasis>
(
<bibRefCitation id="AD233AE1FFC1FF9EFE6CFD5DFDC06956" author="Gould AA" box="[455,598,752,774]" journalOrPublisher="Mollusca and Shells" pageId="35" pageNumber="36" pagination="1 - 510" part="12" refId="ref42670" refString="Gould AA. 1852. United States exploring expedition during the years 1838 - 1842. Mollusca and Shells 12: 1 - 510." title="United States exploring expedition during the years 1838 - 1842" type="journal article" year="1852">Gould, 1852</bibRefCitation>
), but it has red rhinophoral and gill pigment rather than light orange. This species is at the base of a clade that includes
<taxonomicName id="0EB23C93FFC1FF9EFF08FCE1FF7C6931" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,234,844,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFC1FF9EFF08FCE1FF7C6931" box="[163,234,844,865]" italics="true" pageId="35" pageNumber="36">H. iba</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC1FF9EFF53FCE1FEC66931" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[248,336,844,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="reidi">
<emphasis id="FBC69B02FFC1FF9EFF53FCE1FEC66931" box="[248,336,844,865]" italics="true" pageId="35" pageNumber="36">H. reidi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC1FF9EFEF5FCE1FE5D6931" authorityName="Marcus &amp; Marcus" authorityYear="1970" box="[350,459,844,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="regina">
<emphasis id="FBC69B02FFC1FF9EFEF5FCE1FE5D6931" box="[350,459,844,865]" italics="true" pageId="35" pageNumber="36">H. regina</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC1FF9EFE73FCE1FDCF6931" authorityName="Wilson &amp; Willan" authorityYear="2007" box="[472,601,844,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="jacksoni">
<emphasis id="FBC69B02FFC1FF9EFE73FCE1FDCF6931" box="[472,601,844,865]" italics="true" pageId="35" pageNumber="36">H. jacksoni</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC1FF9EFDCCFCE1FD416931" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[615,727,844,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFC1FF9EFDCCFCE1FD416931" box="[615,727,844,865]" italics="true" pageId="35" pageNumber="36">H. cerisae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC1FF9EFF08FCC6FEA3692F" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[163,309,874,896]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFC1FF9EFF08FCC6FEA3692F" box="[163,309,874,896]" italics="true" pageId="35" pageNumber="36">H. krakatoa</emphasis>
</taxonomicName>
, although that relationship is weakly supported. All members of this clade, with the exception of
<taxonomicName id="0EB23C93FFC1FF9EFF14FC05FEB969ED" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[191,303,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFC1FF9EFF14FC05FEB969ED" box="[191,303,936,957]" italics="true" pageId="35" pageNumber="36">H. lacuna</emphasis>
</taxonomicName>
, have an elevated gill pocket and mantle glands arranged all around the perimeter of the mantle margin (
<bibRefCitation id="AD233AE1FFC1FF9EFEA3FC48FDA069AA" author="Gosliner T &amp; Johnson, R." box="[264,566,997,1019]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="35" pageNumber="36" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC1FF9EFDEFFC48FF4A6E49" author="Wilson N &amp; Willan, R." journalOrPublisher="Zootaxa" pageId="35" pageNumber="36" pagination="29 - 42" part="1549" refId="ref45538" refString="Wilson N, Willan, R. 2007. Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982. Zootaxa 1549: 29 - 42." title="Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982" type="journal article" year="2007">Wilson &amp; Willan, 2007</bibRefCitation>
; present study).
<taxonomicName id="0EB23C93FFC1FF9EFE02FBA9FD0B6E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[425,669,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFC1FF9EFE02FBA9FD0B6E49" box="[425,669,1028,1049]" italics="true" pageId="35" pageNumber="36">Hypselodoris lacuna</emphasis>
</taxonomicName>
has only posterior mantle glands, and the gill pocket is not elevated. One feature that
<taxonomicName id="0EB23C93FFC1FF9EFE7BFBEFFDAB6E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[464,573,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFC1FF9EFE7BFBEFFDAB6E06" box="[464,573,1089,1111]" italics="true" pageId="35" pageNumber="36">H. lacuna</emphasis>
</taxonomicName>
shares with other members of this clade is the presence of radular teeth where the inner cusp is much longer than the outer one.
</paragraph>
</subSubSection>
</treatment>
</document>

292
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<materialsCitation id="79DA4D4DFFC4FF9BFF3AFD23FD526950" ID-GBIF-Occurrence="2235579030" collectingDate="1928-11" collectionCode="CASIZ" country="Papua New Guinea" county="Lagoon" elevation="39" latitude="-5.1634" location="Pig" longLatPrecision="5" longitude="145.83833" municipality="Tab Island" pageId="38" pageNumber="39" specimenCode="CASIZ 191464" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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, subsampled for molecular study, dissected, reef close,
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,
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,
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,
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,
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<date id="BD0C61D0FFC4FF9BFE4FFD46FDED6950" box="[484,635,746,768]" pageId="38" pageNumber="39" value="1928-11">28 November</date>
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, 2012.
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<emphasis id="FBC69B02FFC4FF9BFF3AFC9DFEBA6917" box="[145,300,815,839]" italics="true" pageId="38" pageNumber="39">Type locality</emphasis>
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Outer barrier reef north of Tab Island,
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Lagoon,
<collectingCountry id="B1A50780FFC4FF9BFF3AFCC2FEE769D4" box="[145,369,878,900]" name="Papua New Guinea" pageId="38" pageNumber="39">Papua New Guinea</collectingCountry>
.
</paragraph>
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<emphasis id="FBC69B02FFC4FF9BFC82FF6BFBF06A8E" box="[809,1126,198,222]" italics="true" pageId="38" pageNumber="39">Geographical distribution</emphasis>
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Presently known only from northern
<collectingCountry id="B1A50780FFC4FF9BFAA8FF4AFC166B4B" name="Papua New Guinea" pageId="38" pageNumber="39">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC4FF9BFC82FEEFFC166BE6" pageId="38" pageNumber="39" type="etymology">
<paragraph id="C90D4710FFC4FF9BFC82FEEFFC3C6B0A" blockId="38.[809,1425,322,438]" box="[809,938,322,346]" pageId="38" pageNumber="39">
<heading id="9245F07CFFC4FF9BFC82FEEFFC3C6B0A" box="[809,938,322,346]" fontSize="10" level="4" pageId="38" pageNumber="39" reason="1">
<emphasis id="FBC69B02FFC4FF9BFC82FEEFFC3C6B0A" box="[809,938,322,346]" italics="true" pageId="38" pageNumber="39">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC4FF9BFC82FECFFC166BE6" blockId="38.[809,1425,322,438]" pageId="38" pageNumber="39">
<taxonomicName id="0EB23C93FFC4FF9BFC82FECFFBA56B27" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,1075,354,375]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC4FF9BFC82FECFFBA56B27" box="[809,1075,354,375]" italics="true" pageId="38" pageNumber="39">Hypselodoris paradisa</emphasis>
</taxonomicName>
is named for the Greek word for paradise, referring to the tropical habitat of this species.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC4FF9AFC82FE70FD696CF0" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="C90D4710FFC4FF9BFC82FE70FC236BA4" blockId="38.[809,1425,477,899]" box="[809,949,477,500]" pageId="38" pageNumber="39">
<heading id="9245F07CFFC4FF9BFC82FE70FC236BA4" box="[809,949,477,500]" fontSize="10" level="4" pageId="38" pageNumber="39" reason="1">
<emphasis id="FBC69B02FFC4FF9BFC82FE70FC236BA4" box="[809,949,477,500]" italics="true" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC4FF9AFC82FE50FD5A6AAB" blockId="38.[809,1425,477,899]" lastBlockId="39.[163,778,198,251]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="FBC69B02FFC4FF9BFC82FE50FBAE6842" box="[809,1080,509,530]" italics="true" pageId="38" pageNumber="39">External morphology</emphasis>
: Living animal (
<figureCitation id="51895B95FFC4FF9BFAB0FE50FA1E6842" box="[1307,1416,509,531]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="38" pageNumber="39">Fig. 18F</figureCitation>
) moderately small, reaching
<quantity id="0E4AEAF5FFC4FF9BFBD7FDB1FB596861" box="[1148,1231,540,561]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="mm" value="10.0">10 mm</quantity>
in length. Body translucent pink, with series of rows of opaque white spots, dashes and interrupted lines on dorsal surface of notum. V-Shaped reddish mark present posterior to rhinophores. Anterior and posterior ends of notum with purple markings; also present on posterior end of foot. Pink marginal band present along notal rim. Black spots arranged in linear rows between white markings. Six unipinnate gill branches having a translucent white base and bright red–orange pigment on apical surfaces and outer margin. Bulb of perfoliate rhinophores opaque white, with three red–orange transverse bands; bearing about ten densely arranged lamellae. Base of rhinophores translucent white.
</paragraph>
<caption id="9DCD1798FFC4FF9BFF25F890FD3D6D21" httpUri="https://zenodo.org/record/2632822/files/figure.png" pageId="38" pageNumber="39" startId="38.[142,220,1853,1875]" targetBox="[206,1358,934,1813]" targetPageId="38">
<paragraph id="C90D4710FFC4FF9BFF25F890FD3D6D21" blockId="38.[142,1423,1853,1905]" pageId="38" pageNumber="39">
<emphasis id="FBC69B02FFC4FF9BFF25F890FE926D02" bold="true" box="[142,260,1853,1875]" pageId="38" pageNumber="39">Figure 22.</emphasis>
<taxonomicName id="0EB23C93FFC4FF9BFEA4F890FD7D6D03" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[271,747,1853,1875]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFC4FF9BFEA4F890FD836D03" bold="true" box="[271,533,1853,1875]" italics="true" pageId="38" pageNumber="39">Hypselodoris paradisa</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC4FF9BFD5AF893FCDE6D03" box="[753,840,1854,1875]" pageId="38" pageNumber="39" rank="species">
<emphasis id="FBC69B02FFC4FF9BFD5AF893FCDE6D03" bold="true" box="[753,840,1854,1875]" pageId="38" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, CASIZ191464. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFC5FF9AFF08FE89FD5F6BA1" blockId="39.[163,779,292,498]" pageId="39" pageNumber="40">
<emphasis id="FBC69B02FFC5FF9AFF08FE89FEDB6B69" box="[163,333,292,313]" italics="true" pageId="39" pageNumber="40">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFC5FF9AFE35FEEFFE7B6B08" box="[414,493,322,344]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="39" pageNumber="40">Fig. 2K</figureCitation>
). Glands densely situated anteriorly and posteriorly, with few glands present in the central lateral regions of body margin. About 15 glands on either side of anterior end of the body, with arc of ~20 glands situated posteriorly. Three lateral glands present on either side of lateral margin of notum.
</paragraph>
<paragraph id="C90D4710FFC5FF9AFF08FDB7FD546E4B" blockId="39.[163,779,538,1051]" pageId="39" pageNumber="40">
<emphasis id="FBC69B02FFC5FF9AFF08FDB7FEFB6860" box="[163,365,538,560]" italics="true" pageId="39" pageNumber="40">Buccal armature</emphasis>
: Muscular portion of buccal mass slightly longer than glandular portion of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFC5FF9AFE93FD38FE0A68FA" box="[312,412,661,683]" captionStart="Figure 22" captionStartId="38.[142,220,1853,1875]" captionTargetBox="[206,1358,934,1813]" captionTargetId="figure@38.[206,1358,934,1813]" captionTargetPageId="38" captionText="Figure 22. Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ191464. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632822/files/figure.png" pageId="39" pageNumber="40">Fig. 22A</figureCitation>
). Rodlets narrowly ovoid, with single, acutely pointed apex and wider base. Radular formula of
<typeStatus id="1609F9B2FFC5FF9AFEB0FD7FFEEF68B8" box="[283,377,722,744]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
46 × 55.0.55. Rachidian row of teeth absent (
<figureCitation id="51895B95FFC5FF9AFF50FD5CFECE6956" box="[251,344,753,775]" captionStart="Figure 22" captionStartId="38.[142,220,1853,1875]" captionTargetBox="[206,1358,934,1813]" captionTargetId="figure@38.[206,1358,934,1813]" captionTargetPageId="38" captionText="Figure 22. Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ191464. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632822/files/figure.png" pageId="39" pageNumber="40">Fig. 22B</figureCitation>
). Innermost lateral teeth having single triangular denticles on inner side of bifid primary cusp, with another one to two outer denticles. Next several laterals lacking inner triangular denticle but possessing two or three denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFC5FF9AFEC4FC27FE4669F0" box="[367,464,906,928]" captionStart="Figure 22" captionStartId="38.[142,220,1853,1875]" captionTargetBox="[206,1358,934,1813]" captionTargetId="figure@38.[206,1358,934,1813]" captionTargetPageId="38" captionText="Figure 22. Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ191464. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632822/files/figure.png" pageId="39" pageNumber="40">Fig. 22C</figureCitation>
) all lacking inner denticles but having four or five sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and shorter tooth shape, with four to six rounded outer denticles (
<figureCitation id="51895B95FFC5FF9AFEB8FBA8FEE66E4A" box="[275,368,1029,1051]" captionStart="Figure 22" captionStartId="38.[142,220,1853,1875]" captionTargetBox="[206,1358,934,1813]" captionTargetId="figure@38.[206,1358,934,1813]" captionTargetPageId="38" captionText="Figure 22. Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ191464. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632822/files/figure.png" pageId="39" pageNumber="40">Fig. 22D</figureCitation>
), often larger than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFC5FF9AFF08FBE9FD696CF0" blockId="39.[163,779,1092,1696]" pageId="39" pageNumber="40">
<emphasis id="FBC69B02FFC5FF9AFF08FBE9FE346E09" box="[163,418,1092,1113]" italics="true" pageId="39" pageNumber="40">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFC5FF9AFF08FBCFFE956E28" box="[163,259,1122,1144]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFC5FF9AFE0AFBCFFE696E28" box="[417,511,1122,1144]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="39" pageNumber="40">Fig. 13G</figureCitation>
). Ampulla thin, tubular, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick, and narrowing slightly as it transitions into muscular ejaculatory portion. Ejaculatory portion highly convoluted and long, widening again before entry into broad penial bulb. Penial bulb adjacent to straight, very wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively short vagina leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum adjacent to bursa at distal end of vagina. Moderately long uterine duct emerging from vagina close to base of bursa and female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC5FF9AFF08F96AFA0B685F" pageId="39" pageNumber="40" type="discussion">
<paragraph id="C90D4710FFC5FF9AFF08F96AFE866C8F" blockId="39.[163,779,1735,1912]" box="[163,272,1735,1759]" pageId="39" pageNumber="40">
<heading id="9245F07CFFC5FF9AFF08F96AFE866C8F" box="[163,272,1735,1759]" fontSize="10" level="4" pageId="39" pageNumber="40" reason="1">
<emphasis id="FBC69B02FFC5FF9AFF08F96AFE866C8F" box="[163,272,1735,1759]" italics="true" pageId="39" pageNumber="40">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC5FF9AFF08F945FA0B685F" blockId="39.[163,779,1735,1912]" lastBlockId="39.[827,1443,198,527]" pageId="39" pageNumber="40">
The phylogenetic relationship of this species to
<taxonomicName id="0EB23C93FFC5FF9AFF08F8AAFEDC6D4B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,330,1798,1820]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC5FF9AFF08F8AAFEDC6D4B" box="[163,330,1798,1820]" italics="true" pageId="39" pageNumber="40">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC5FF9AFE21F8AAFD966D4B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[394,512,1798,1820]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC5FF9AFE21F8AAFD966D4B" box="[394,512,1798,1820]" italics="true" pageId="39" pageNumber="40">H. skyleri</emphasis>
</taxonomicName>
is discussed above in the remarks section of
<taxonomicName id="0EB23C93FFC5FF9AFE6BF88BFDF16D6A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[448,615,1829,1851]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC5FF9AFE6BF88BFDF16D6A" box="[448,615,1829,1851]" italics="true" pageId="39" pageNumber="40">H. katherinae</emphasis>
</taxonomicName>
. This species has similarity in its colour pattern to the sympatric species,
<taxonomicName id="0EB23C93FFC5FF9AFEBBF8CEFE266D27" baseAuthorityName="Rudman" baseAuthorityYear="1986" box="[272,432,1890,1912]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFC5FF9AFEBBF8CEFE266D27" box="[272,432,1890,1912]" italics="true" pageId="39" pageNumber="40">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC5FF9AFE5EF8CEFD1D6D27" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[501,651,1890,1912]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFC5FF9AFE5EF8CEFD1D6D27" box="[501,651,1890,1912]" italics="true" pageId="39" pageNumber="40">H. decorata</emphasis>
</taxonomicName>
, with the presence of opaque white lines and longitudinally arranged black spots. Despite the similarity of colour pattern,
<taxonomicName id="0EB23C93FFC5FF9AFC09FEA9FBB86B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[930,1070,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC5FF9AFC09FEA9FBB86B49" box="[930,1070,260,281]" italics="true" pageId="39" pageNumber="40">H. paradisa</emphasis>
</taxonomicName>
is in a distinct clade from both
<taxonomicName id="0EB23C93FFC5FF9AFC90FE8EFC586B67" baseAuthorityName="Rudman" baseAuthorityYear="1986" box="[827,974,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFC5FF9AFC90FE8EFC586B67" box="[827,974,290,312]" italics="true" pageId="39" pageNumber="40">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC5FF9AFBA1FE8EFB056B67" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1034,1171,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFC5FF9AFBA1FE8EFB056B67" box="[1034,1171,290,312]" italics="true" pageId="39" pageNumber="40">H. decorata</emphasis>
</taxonomicName>
. Both
<taxonomicName id="0EB23C93FFC5FF9AFB49FE8EFAF96B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1250,1391,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC5FF9AFB49FE8EFAF96B67" box="[1250,1391,290,312]" italics="true" pageId="39" pageNumber="40">H. paradisa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC5FF9AFC90FEEFFC506B06" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[827,966,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFC5FF9AFC90FEEFFC506B06" box="[827,966,321,343]" italics="true" pageId="39" pageNumber="40">H. decorata</emphasis>
</taxonomicName>
have three reddish rhinophoral bands, whereas
<taxonomicName id="0EB23C93FFC5FF9AFC34FECDFBBD6B25" baseAuthorityName="Rudman" baseAuthorityYear="1986" box="[927,1067,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFC5FF9AFC34FECDFBBD6B25" box="[927,1067,352,373]" italics="true" pageId="39" pageNumber="40">H. maculosa</emphasis>
</taxonomicName>
only has two bands.
<taxonomicName id="0EB23C93FFC5FF9AFABBFECDFC376BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC5FF9AFABBFECDFC376BC3" italics="true" pageId="39" pageNumber="40">Hypselodoris paradisa</emphasis>
</taxonomicName>
has a reddish V-shaped patch on the head that is absent in both other species.
<taxonomicName id="0EB23C93FFC5FF9AFB36FE30FA0B6BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1181,1437,413,434]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC5FF9AFB36FE30FA0B6BE2" box="[1181,1437,413,434]" italics="true" pageId="39" pageNumber="40">Hypselodoris paradisa</emphasis>
</taxonomicName>
, like other members of its clade, has a broader radula than members of the
<taxonomicName id="0EB23C93FFC5FF9AFBE8FE76FB4C6BBF" baseAuthorityName="Rudman" baseAuthorityYear="1986" box="[1091,1242,474,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFC5FF9AFBE8FE76FB4C6BBF" box="[1091,1242,474,496]" italics="true" pageId="39" pageNumber="40">H. maculosa</emphasis>
</taxonomicName>
clade and has a broad penial papilla, as in other members of its clade.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="CB96EAD3129E66995C35284C10B60D88">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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<taxonomicName id="0EB23C93FFC5FF9AFC90FDE1FAAD6832" ID-CoL="3P353" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1339,587,611]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="41" phylum="Mollusca" rank="species" species="perii" status="sp. nov.">
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GOSLINER &amp; JOHNSON
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<emphasis id="FBC69B02FFC5FF9AFAE9FDE2FA356833" bold="true" box="[1346,1443,587,611]" pageId="39" pageNumber="40">SP. NOV.</emphasis>
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<emphasis id="FBC69B02FFC5FF9AFC90FCA3FC0C6973" box="[827,922,782,803]" italics="true" pageId="39" pageNumber="40">Noumea</emphasis>
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sp.
<bibRefCitation id="AD233AE1FFC5FF9AFC6DFCA0FBFB6972" author="Debelius H." box="[966,1133,781,803]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="39" pageNumber="40" refId="ref41856" refString="Debelius H. 1997. Schnecken fuhrer Indopazifik. Frankfurt: IKAN-Unterwasserarchiv." title="Schnecken fuhrer Indopazifik" type="book" year="1997">Debelius, 1997</bibRefCitation>
: 218, middle photograph.
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<emphasis id="FBC69B02FFC5FF9AFC90FC81FC0C6911" box="[827,922,812,833]" italics="true" pageId="39" pageNumber="40">Noumea</emphasis>
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sp.
<bibRefCitation id="AD233AE1FFC5FF9AFC6DFC81FBF96911" author="Coleman N." box="[966,1135,812,834]" journalOrPublisher="Springwood: Australia Neville Coleman's Underwater Geographic" pageId="39" pageNumber="40" refId="ref41724" refString="Coleman N. 2001. 1001 Nudibranchs: catalogue of Indo- Pacific sea slugs. Springwood: Australia Neville Coleman's Underwater Geographic." title="1001 Nudibranchs: catalogue of Indo- Pacific sea slugs" type="book" year="2001">Coleman, 2001</bibRefCitation>
: 84.
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<emphasis id="FBC69B02FFC5FF9AFC90FCE7FC4D690F" box="[827,987,842,863]" italics="true" pageId="39" pageNumber="40">Hypselodoris</emphasis>
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sp. 8.
<bibRefCitation id="AD233AE1FFC5FF9AFB9BFCE7FAF66930" author="Debelius H &amp; Kuiter R." box="[1072,1376,842,864]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="39" pageNumber="40" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 123, middle photograph.
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<taxonomicName id="0EB23C93FFC5FF9AFC90FC25FC4469CD" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[827,978,904,925]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC5FF9AFC90FC25FC4469CD" box="[827,978,904,925]" italics="true" pageId="39" pageNumber="40">Hypselodoris</emphasis>
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sp. 13.
<bibRefCitation id="AD233AE1FFC5FF9AFB87FC25FA8E69CE" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1068,1304,904,926]" journalOrPublisher="Jacksonville: New World Press" pageId="39" pageNumber="40" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFC5FF9AFB3CFC25FB4469CD" box="[1175,1234,904,925]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
, 2015
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: 261, upper right photograph.
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0 41281, (formerly
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), dissected,
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,
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depth,
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, Peri Paleracio.
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Mainit Bubbles, Mabini,
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,
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.
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<paragraph id="C90D4710FFC5FF9AFC90FAB4FBEE6F61" blockId="39.[827,1443,1305,1420]" box="[827,1144,1305,1329]" pageId="39" pageNumber="40">
<heading id="9245F07CFFC5FF9AFC90FAB4FBEE6F61" box="[827,1144,1305,1329]" fontSize="10" level="4" pageId="39" pageNumber="40" reason="1">
<emphasis id="FBC69B02FFC5FF9AFC90FAB4FBEE6F61" box="[827,1144,1305,1329]" italics="true" pageId="39" pageNumber="40">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC5FF9AFC90FA97FBF86FDC" blockId="39.[827,1443,1305,1420]" pageId="39" pageNumber="40">
Known only from
<collectingRegion id="0B7689F2FFC5FF9AFBADFA94FBA16F1F" box="[1030,1079,1337,1359]" country="Indonesia" name="Bali" pageId="39" pageNumber="40">Bali</collectingRegion>
,
<collectingCountry id="B1A50780FFC5FF9AFBE8FA94FB256F1F" box="[1091,1203,1337,1359]" name="Indonesia" pageId="39" pageNumber="40">Indonesia</collectingCountry>
(
<bibRefCitation id="AD233AE1FFC5FF9AFB69FA94FAFF6F1E" author="Debelius H." box="[1218,1385,1337,1359]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="39" pageNumber="40" refId="ref41856" refString="Debelius H. 1997. Schnecken fuhrer Indopazifik. Frankfurt: IKAN-Unterwasserarchiv." title="Schnecken fuhrer Indopazifik" type="book" year="1997">Debelius, 1997</bibRefCitation>
) and
<collectingRegion id="0B7689F2FFC5FF9AFC90FAF5FC3E6F3D" box="[827,936,1368,1389]" country="Philippines" name="Batangas" pageId="39" pageNumber="40">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFC5FF9AFC1FFAF5FBA06F3E" box="[948,1078,1368,1390]" name="Philippines" pageId="39" pageNumber="40">Philippines</collectingCountry>
from deep reefs (
<bibRefCitation id="AD233AE1FFC5FF9AFB57FAF5FCE46FDC" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="39" pageNumber="40" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFC5FF9AFACEFAF5FA036F3D" box="[1381,1429,1368,1389]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
., 2008
</bibRefCitation>
,
<bibRefCitation id="AD233AE1FFC5FF9AFCD5FADBFC236FDC" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[894,949,1398,1420]" journalOrPublisher="Jacksonville: New World Press" pageId="39" pageNumber="40" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">2015</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC5FF9AFC90FA1EFBA36CD3" pageId="39" pageNumber="40" type="etymology">
<paragraph id="C90D4710FFC5FF9AFC90FA1EFC2A6F9B" blockId="39.[827,1444,1459,1667]" box="[827,956,1459,1483]" pageId="39" pageNumber="40">
<heading id="9245F07CFFC5FF9AFC90FA1EFC2A6F9B" box="[827,956,1459,1483]" fontSize="10" level="4" pageId="39" pageNumber="40" reason="1">
<emphasis id="FBC69B02FFC5FF9AFC90FA1EFC2A6F9B" box="[827,956,1459,1483]" italics="true" pageId="39" pageNumber="40">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC5FF9AFC90FA79FBA36CD3" blockId="39.[827,1444,1459,1667]" pageId="39" pageNumber="40">
<taxonomicName id="0EB23C93FFC5FF9AFC90FA79FB9A6FB9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1036,1492,1513]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFC5FF9AFC90FA79FB9A6FB9" box="[827,1036,1492,1513]" italics="true" pageId="39" pageNumber="40">Hypselodoris perii</emphasis>
</taxonomicName>
is named for good friend and diver guide extraordinaire, Peri Paleracio, who found the first specimen of this species in the
<collectingCountry id="B1A50780FFC5FF9AFB4AF9BCFAFE6C77" box="[1249,1384,1553,1575]" name="Philippines" pageId="39" pageNumber="40">Philippines</collectingCountry>
. Peri has discovered many new species of marine life in the
<collectingCountry id="B1A50780FFC5FF9AFC90F9E3FC526C34" box="[827,964,1614,1636]" name="Philippines" pageId="39" pageNumber="40">Philippines</collectingCountry>
, and it is a pleasure to honor his efforts with this new species.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC5FF94FC90F906FD446B68" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40" type="description">
<paragraph id="C90D4710FFC5FF9AFC90F906FC516C92" blockId="39.[827,1443,1707,1914]" box="[827,967,1707,1730]" pageId="39" pageNumber="40">
<heading id="9245F07CFFC5FF9AFC90F906FC516C92" box="[827,967,1707,1730]" fontSize="10" level="4" pageId="39" pageNumber="40" reason="1">
<emphasis id="FBC69B02FFC5FF9AFC90F906FC516C92" box="[827,967,1707,1730]" italics="true" pageId="39" pageNumber="40">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC5FF95FC90F967FDC26F19" blockId="39.[827,1443,1707,1914]" lastBlockId="40.[145,762,1147,1353]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="FBC69B02FFC5FF9AFC90F967FBBE6C8F" box="[827,1064,1738,1759]" italics="true" pageId="39" pageNumber="40">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFC5FF9AFB5EF967FAC36CB0" box="[1269,1365,1738,1760]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="39" pageNumber="40">Fig. 18G</figureCitation>
) large, reaching
<quantity id="0E4AEAF5FFC5FF9AFC00F944FC6B6CAF" box="[939,1021,1769,1791]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="40" unit="mm" value="35.0">35 mm</quantity>
in length. Body translucent white, with wine-red medial line and scattered, elongate wine-red oval rings surrounded by diffuse purple pigment. Wide yellow to yellow–orange marginal band present along entire mantle margin. Wine-red ring encircling base of gill sheath. Translucent white foot ornamented by wine-red spots having larger areas of purple surrounding them. Eight unipinnate gill branches with translucent white base and bright redorange pigment on apical surfaces and outer margin. Perfoliate rhinophores uniformly bright red–orange, bearing ~17 densely arranged lamellae.
</paragraph>
<caption id="9DCD1798FFCAFF95FF3AFB8EFCAE6E07" httpUri="https://zenodo.org/record/2632826/files/figure.png" pageId="40" pageNumber="41" startId="40.[145,224,1059,1081]" targetBox="[209,1361,195,1018]" targetPageId="40">
<paragraph id="C90D4710FFCAFF95FF3AFB8EFCAE6E07" blockId="40.[145,1425,1059,1111]" pageId="40" pageNumber="41">
<emphasis id="FBC69B02FFCAFF95FF3AFB8EFE916E68" bold="true" box="[145,263,1059,1081]" pageId="40" pageNumber="41">Figure 23.</emphasis>
<taxonomicName id="0EB23C93FFCAFF95FEB9FB8EFD566E69" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[274,704,1059,1081]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="perii" status="sp. nov.">
<emphasis id="FBC69B02FFCAFF95FEB9FB8EFE7E6E68" bold="true" box="[274,488,1059,1081]" italics="true" pageId="40" pageNumber="41">Hypselodoris perii</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCAFF95FD6CFB89FC886E69" box="[711,798,1060,1081]" pageId="40" pageNumber="41" rank="species">
<emphasis id="FBC69B02FFCAFF95FD6CFB89FC886E69" bold="true" box="[711,798,1060,1081]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41281, buccal armature. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFCAFF95FF3AFADFFEE36C71" blockId="40.[145,761,1394,1569]" pageId="40" pageNumber="41">
<emphasis id="FBC69B02FFCAFF95FF3AFADFFEAD6FD7" box="[145,315,1394,1415]" italics="true" pageId="40" pageNumber="41">Mantle glands</emphasis>
: Subcutaneous mantle glands simple rounded in shape (
<figureCitation id="51895B95FFCAFF95FECEFA3DFE236FF5" box="[357,437,1424,1446]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="40" pageNumber="41">Fig. 2L</figureCitation>
). Glands situated anteriorly and posteriorly, with no glands present along central regions of mantle margin. Four to five glands on either side of anterior end of the body, with an arc of 13 glands situated posteriorly.
</paragraph>
<paragraph id="C90D4710FFCAFF95FF3AF9E7FB296F19" blockId="40.[145,762,1610,1908]" lastBlockId="40.[809,1425,1147,1353]" pageId="40" pageNumber="41">
<emphasis id="FBC69B02FFCAFF95FF3AF9E7FECD6C0F" box="[145,347,1610,1631]" italics="true" pageId="40" pageNumber="41">Buccal armature</emphasis>
: Muscular portion of buccal mass approximately equal in length to oral tube. Buccal mass consisting of chitinous labial cuticle at anterior end of muscular portion of buccal mass. Jaws bearing numerous rodlets (
<figureCitation id="51895B95FFCAFF95FECAF969FE2A6C89" box="[353,444,1732,1754]" captionStart="Figure 23" captionStartId="40.[145,224,1059,1081]" captionTargetBox="[209,1361,195,1018]" captionTargetId="figure@40.[209,1361,195,1018]" captionTargetPageId="40" captionText="Figure 23. Hypselodoris perii Gosliner &amp; Johnson sp. nov., holotype, NMP 041281, buccal armature. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632826/files/figure.png" pageId="40" pageNumber="41">Fig. 23A</figureCitation>
). Rodlets broadly triangular, with single acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFCAFF95FF3AF8AFFF626D48" box="[145,244,1794,1816]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
42 × 40.0.40. Rachidian row of teeth absent (
<figureCitation id="51895B95FFCAFF95FF32F88DFF6E6D65" box="[153,248,1824,1846]" captionStart="Figure 23" captionStartId="40.[145,224,1059,1081]" captionTargetBox="[209,1361,195,1018]" captionTargetId="figure@40.[209,1361,195,1018]" captionTargetPageId="40" captionText="Figure 23. Hypselodoris perii Gosliner &amp; Johnson sp. nov., holotype, NMP 041281, buccal armature. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632826/files/figure.png" pageId="40" pageNumber="41">Fig. 23B</figureCitation>
). Innermost lateral teeth have a single large, triangular denticle on inner side of bifid primary cusp and lacking outer denticles. Next several laterals lacking inner triangular denticle and lacking denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFCAFF95FA81FB37FA1E6EFF" box="[1322,1416,1178,1200]" captionStart="Figure 23" captionStartId="40.[145,224,1059,1081]" captionTargetBox="[209,1361,195,1018]" captionTargetId="figure@40.[209,1361,195,1018]" captionTargetPageId="40" captionText="Figure 23. Hypselodoris perii Gosliner &amp; Johnson sp. nov., holotype, NMP 041281, buccal armature. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632826/files/figure.png" pageId="40" pageNumber="41">Fig. 23C</figureCitation>
) also lacking inner denticles but possessing two or three triangular outer denticles. Outer teeth lacking inner denticles and having two to five triangular outer denticles (
<figureCitation id="51895B95FFCAFF95FC09FAB9FB936F79" box="[930,1029,1300,1322]" captionStart="Figure 23" captionStartId="40.[145,224,1059,1081]" captionTargetBox="[209,1361,195,1018]" captionTargetId="figure@40.[209,1361,195,1018]" captionTargetPageId="40" captionText="Figure 23. Hypselodoris perii Gosliner &amp; Johnson sp. nov., holotype, NMP 041281, buccal armature. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632826/files/figure.png" pageId="40" pageNumber="41">Fig. 23D</figureCitation>
). Outermost teeth with narrower base and more elongate tooth shape.
</paragraph>
<paragraph id="C90D4710FFCAFF94FC82FADFFD446B68" blockId="40.[809,1425,1394,1906]" lastBlockId="41.[163,779,198,312]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="FBC69B02FFCAFF95FC82FADFFB866FD7" box="[809,1040,1394,1415]" italics="true" pageId="40" pageNumber="41">Reproductive system</emphasis>
: Reproductive organs of
<typeStatus id="1609F9B2FFCAFF95FA9BFADFFA076FD8" box="[1328,1425,1394,1416]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
not fully mature, but arrangement of all major organs evident (
<figureCitation id="51895B95FFCAFF95FC27FA02FC7A6F94" box="[908,1004,1455,1477]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="40" pageNumber="41">Fig. 13H</figureCitation>
). Ampulla thick, tubular and slightly convoluted, narrowing somewhat before bifurcating into the oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens curved and thick, narrowing slightly while transitioning into muscular ejaculatory portion. Ejaculatory portion curving into segment entering short, enlarged penial bulb. Penial bulb adjacent to straight, slender vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Elongate vagina leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Short uterine duct emerging from about half of the length along duct to bursa. Uterine duct relatively short and entering female gland mass near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCBFF94FF08FEF2FA3268F8" pageId="41" pageNumber="42" type="discussion">
<paragraph id="C90D4710FFCBFF94FF08FEF2FE9B6B27" blockId="41.[163,779,351,1908]" box="[163,269,351,375]" pageId="41" pageNumber="42">
<heading id="9245F07CFFCBFF94FF08FEF2FE9B6B27" box="[163,269,351,375]" fontSize="10" level="4" pageId="41" pageNumber="42" reason="1">
<emphasis id="FBC69B02FFCBFF94FF08FEF2FE9B6B27" box="[163,269,351,375]" italics="true" pageId="41" pageNumber="42">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCBFF94FF08FE2DFDC06E81" blockId="41.[163,779,351,1908]" pageId="41" pageNumber="42">
The colour pattern of
<taxonomicName id="0EB23C93FFCBFF94FE0BFE2DFE6E6BC5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[416,504,384,405]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FE0BFE2DFE6E6BC5" box="[416,504,384,405]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
clearly differentiates it from the all other described species. It is the only species with an elongate medial longitudinal reddish line and reddish oval markings. The most similar species is
<emphasis id="FBC69B02FFCBFF94FF08FE57FE08685F" box="[163,414,506,527]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="0EB23C93FFCBFF94FF08FE57FEAA685F" authorityName="Stimpson" authorityYear="1855" box="[163,316,506,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
dollfusi
</emphasis>
(
<bibRefCitation id="AD233AE1FFCBFF94FE04FE57FDEF6841" author="Pruvot-Fol A." box="[431,633,506,529]" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="41" pageNumber="42" pagination="89 - 159" part="21" refId="ref44371" refString="Pruvot-Fol A. 1933. Mission Robert Ph. Dollfus en Egypte. Opisthobranchiata. Memoires de l'Institut d'Egypte 21: 89 - 159, pls. 1 - 4." title="Mission Robert Ph. Dollfus en Egypte. Opisthobranchiata" type="journal article" year="1933">Pruvot-Fol, 1933</bibRefCitation>
), which was re-described by
<bibRefCitation id="AD233AE1FFCBFF94FEF0FDB4FD05687F" author="Gosliner TM &amp; Behrens, DW" box="[347,659,537,559]" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="41" pageNumber="42" pagination="111 - 124" part="52" refId="ref42505" refString="Gosliner TM, Behrens, DW. 2000. Two new species of Chromodorididae (Mollusca: Nudibranchia) from the tropical Indo-Pacific, with a redescription of Hypselodoris dollfusi (Pruvot-Fol, 1933). Proceedings of the California Academy of Sciences 52: 111 - 124." title="Two new species of Chromodorididae (Mollusca: Nudibranchia) from the tropical Indo-Pacific, with a redescription of Hypselodoris dollfusi (Pruvot-Fol, 1933)" type="journal article" year="2000">Gosliner &amp; Behrens (2000)</bibRefCitation>
. Although both species are translucent white, with a yellow marginal band and wine-red and purple markings, there are numerous differences in the external colour pattern of the two species.
<emphasis id="FBC69B02FFCBFF94FE7CFD39FD5E68F9" box="[471,712,660,681]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="0EB23C93FFCBFF94FE7CFD39FDFC68F9" authorityName="Stimpson" authorityYear="1855" box="[471,618,660,681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
dollfusi
</emphasis>
lacks the medial longitudinal line that is present in
<taxonomicName id="0EB23C93FFCBFF94FD04FD1EFC936898" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[687,773,691,712]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FD04FD1EFC936898" box="[687,773,691,712]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
. In
<emphasis id="FBC69B02FFCBFF94FF6AFD7FFEA268B6" box="[193,308,721,743]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
, the wine-red markings are narrow circles, which contain purple pigment. In contrast,
<taxonomicName id="0EB23C93FFCBFF94FD1BFD5DFC9D6955" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[688,779,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FD1BFD5DFC9D6955" box="[688,779,752,773]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
has broad reddish ovals surrounded by diffuse purple pigment. In
<emphasis id="FBC69B02FFCBFF94FE8CFC83FE0C6912" box="[295,410,813,835]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
, there are red and purple rings at the base of the rhinophores and the gill pocket, whereas in
<taxonomicName id="0EB23C93FFCBFF94FF69FCC6FE8A69D0" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[194,284,875,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FF69FCC6FE8A69D0" box="[194,284,875,896]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
there is only a red ring at the base of the gill. In
<emphasis id="FBC69B02FFCBFF94FF52FC27FEE269CE" box="[249,372,905,927]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
, the gill and rhinophores are deep red, whereas they are red–orange in
<taxonomicName id="0EB23C93FFCBFF94FDE6FC05FD3269ED" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[589,676,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FDE6FC05FD3269ED" box="[589,676,936,957]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
. The red pigment on the gill of
<emphasis id="FBC69B02FFCBFF94FE3FFC6AFD91698B" box="[404,519,966,988]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
is restricted to the axis of each gill branch, whereas in
<taxonomicName id="0EB23C93FFCBFF94FE40FC4BFDAA69AB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[491,572,998,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FE40FC4BFDAA69AB" box="[491,572,998,1019]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
the entire external surface of each gill branch is red–orange. Externally, both
<emphasis id="FBC69B02FFCBFF94FF71FB8EFEDB6E67" box="[218,333,1058,1080]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
and
<taxonomicName id="0EB23C93FFCBFF94FE2FFB8EFE416E68" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[388,471,1059,1080]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FE2FFB8EFE416E68" box="[388,471,1059,1080]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
have anterior and posterior mantle glands and lack glands in the centre portion of the body. The glands are more numerous in
<emphasis id="FBC69B02FFCBFF94FD3FFBCDFC9C6E25" box="[660,778,1120,1141]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
(12–18 anterior glands per side of the body and 22 posterior glands) vs. three or four anterior glands per side and 13 posterior glands in
<taxonomicName id="0EB23C93FFCBFF94FE57FB11FDD96E81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[508,591,1212,1233]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FE57FB11FDD96E81" box="[508,591,1212,1233]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFCBFF94FF10FB77FD7C6C8B" blockId="41.[163,779,351,1908]" pageId="41" pageNumber="42">
Internally, there are also differences between the two species. The radular formula of
<emphasis id="FBC69B02FFCBFF94FD9AFB57FD306F5E" box="[561,678,1273,1295]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
contains more rows of teeth (66 vs. 42) and more teeth per row than does the radula (88 vs. 40) of
<taxonomicName id="0EB23C93FFCBFF94FD8BFA9AFDE56F1C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[544,627,1335,1356]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FD8BFA9AFDE56F1C" box="[544,627,1335,1356]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
. In the single individual of
<emphasis id="FBC69B02FFCBFF94FE9EFAFBFE3F6F3A" box="[309,425,1365,1387]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
examined, the left inner lateral teeth had a single inner denticle, whereas the right inner laterals lacked a denticle. In
<taxonomicName id="0EB23C93FFCBFF94FDEAFA3EFD0C6FF8" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[577,666,1427,1448]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FDEAFA3EFD0C6FF8" box="[577,666,1427,1448]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
, both the left and the right inner laterals bear a single denticle. In
<taxonomicName id="0EB23C93FFCBFF94FF68FA7DFE8E6FB5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[195,280,1488,1509]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FF68FA7DFE8E6FB5" box="[195,280,1488,1509]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
, the middle laterals bear two or three outer denticles, whereas the middle laterals of
<emphasis id="FBC69B02FFCBFF94FDC8FA42FD416C53" box="[611,727,1518,1540]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
lack denticles. Only the outer four to ten teeth of
<emphasis id="FBC69B02FFCBFF94FD3EF9A3FC9D6C72" box="[661,779,1549,1571]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
bear outer denticles, whereas all of the middle and outer lateral teeth of
<taxonomicName id="0EB23C93FFCBFF94FE3FF9E6FE7F6C30" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[404,489,1611,1632]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FE3FF9E6FE7F6C30" box="[404,489,1611,1632]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
have denticles. There are minor differences in the reproductive system of the two species, but given that the reproductive system of
<taxonomicName id="0EB23C93FFCBFF94FF14F90AFE806CEC" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[191,278,1703,1724]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FF14F90AFE806CEC" box="[191,278,1703,1724]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
was not fully mature, it is difficult to make detailed comparisons based on the present material.
</paragraph>
<paragraph id="C90D4710FFCBFF94FF10F949FA3268F8" blockId="41.[163,779,351,1908]" lastBlockId="41.[827,1444,198,680]" pageId="41" pageNumber="42">
The two species are geographically isolated, with
<emphasis id="FBC69B02FFCBFF94FF08F8AEFEB46D47" box="[163,290,1794,1816]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
being restricted to the Arabian Sea and
<taxonomicName id="0EB23C93FFCBFF94FF08F88FFF636D67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,245,1826,1847]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FF08F88FFF636D67" box="[163,245,1826,1847]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
being known only from Bali and the
<collectingCountry id="B1A50780FFCBFF94FD22F88CFC916D67" box="[649,775,1825,1847]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
. Unfortunately, no material suitable for molecular study is available for
<emphasis id="FBC69B02FFCBFF94FEF0F8F2FE416D23" box="[347,471,1886,1908]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
. In the present molecular analysis,
<taxonomicName id="0EB23C93FFCBFF94FC08FF6AFC616A8C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[931,1015,199,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FC08FF6AFC616A8C" box="[931,1015,199,220]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
is sister to a large clade that includes members of the
<emphasis id="FBC69B02FFCBFF94FC40FF4BFBF76AAA" box="[1003,1121,229,251]" italics="true" pageId="41" pageNumber="42">H. obscura</emphasis>
,
<emphasis id="FBC69B02FFCBFF94FBC6FF4BFB7F6AAB" box="[1133,1257,230,251]" italics="true" pageId="41" pageNumber="42">H. capensis</emphasis>
(
<bibRefCitation id="AD233AE1FFCBFF94FB5CFF48FA016AAB" author="Barnard K." box="[1271,1431,229,251]" journalOrPublisher="Annals of the South African Museum" pageId="41" pageNumber="42" pagination="171 - 215" part="25" refId="ref41182" refString="Barnard K. 1927. South African nudibranch Mollusca, with descriptions of new species, and a note on some species from Tristan d'Acunha. Annals of the South African Museum 25: 171 - 215." title="South African nudibranch Mollusca, with descriptions of new species, and a note on some species from Tristan d'Acunha" type="journal article" year="1927">Barnard, 1927</bibRefCitation>
),
<taxonomicName id="0EB23C93FFCBFF94FC90FEA9FC246B49" authorityName="Baba" authorityYear="1994" box="[827,946,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kaname">
<emphasis id="FBC69B02FFCBFF94FC90FEA9FC246B49" box="[827,946,260,281]" italics="true" pageId="41" pageNumber="42">H. kaname</emphasis>
</taxonomicName>
and
<emphasis id="FBC69B02FFCBFF94FC42FEA9FBE56B49" box="[1001,1139,260,281]" italics="true" pageId="41" pageNumber="42">H. maritima</emphasis>
clades.
<bibRefCitation id="AD233AE1FFCBFF94FB63FEA9FC176B69" author="Gosliner TM &amp; Behrens, DW" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="41" pageNumber="42" pagination="111 - 124" part="52" refId="ref42505" refString="Gosliner TM, Behrens, DW. 2000. Two new species of Chromodorididae (Mollusca: Nudibranchia) from the tropical Indo-Pacific, with a redescription of Hypselodoris dollfusi (Pruvot-Fol, 1933). Proceedings of the California Academy of Sciences 52: 111 - 124." title="Two new species of Chromodorididae (Mollusca: Nudibranchia) from the tropical Indo-Pacific, with a redescription of Hypselodoris dollfusi (Pruvot-Fol, 1933)" type="journal article" year="2000">Gosliner &amp; Behrens (2000)</bibRefCitation>
, based on morphological similarities, speculated that
<emphasis id="FBC69B02FFCBFF94FCDFFEEFFC666B06" box="[884,1008,321,343]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
was likely to be most closely related to the clade that includes
<taxonomicName id="0EB23C93FFCBFF94FBC7FECDFB636B25" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1132,1269,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="paulinae">
<emphasis id="FBC69B02FFCBFF94FBC7FECDFB636B25" box="[1132,1269,352,373]" italics="true" pageId="41" pageNumber="42">H. paulinae</emphasis>
</taxonomicName>
,
<emphasis id="FBC69B02FFCBFF94FAA8FECDFAF96B25" box="[1283,1391,352,373]" italics="true" pageId="41" pageNumber="42">H. fucata</emphasis>
and
<taxonomicName id="0EB23C93FFCBFF94FC90FED2FC216BC3" authorityName="Baba" authorityYear="1994" box="[827,951,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kaname">
<emphasis id="FBC69B02FFCBFF94FC90FED2FC216BC3" box="[827,951,382,404]" italics="true" pageId="41" pageNumber="42">H. kaname</emphasis>
</taxonomicName>
. They noted that these four species shared three apomorphies: an erect rather than spreading branchial plume, a short jaw element shaft; and a receptaculum seminis that inserts into the vagina at the base of the bursa copulatrix.
<taxonomicName id="0EB23C93FFCBFF94FB0CFE54FAE6685F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1191,1392,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FB0CFE54FAE6685F" box="[1191,1392,505,527]" italics="true" pageId="41" pageNumber="42">Hypselodoris perii</emphasis>
</taxonomicName>
also shares these characteristics. Our molecular studies place
<taxonomicName id="0EB23C93FFCBFF94FC90FD9AFC02681C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,916,567,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="perii">
<emphasis id="FBC69B02FFCBFF94FC90FD9AFC02681C" box="[827,916,567,588]" italics="true" pageId="41" pageNumber="42">H. perii</emphasis>
</taxonomicName>
in close proximity to the clade that includes
<taxonomicName id="0EB23C93FFCBFF94FC90FDFBFC5A683A" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,972,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="paulinae">
<emphasis id="FBC69B02FFCBFF94FC90FDFBFC5A683A" box="[827,972,597,619]" italics="true" pageId="41" pageNumber="42">H. paulinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFCBFF94FBA7FDFBFB07683A" authorityName="Baba" authorityYear="1994" box="[1036,1169,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kaname">
<emphasis id="FBC69B02FFCBFF94FBA7FDFBFB07683A" box="[1036,1169,597,619]" italics="true" pageId="41" pageNumber="42">H. kaname</emphasis>
</taxonomicName>
. Additional molecular studies that include
<emphasis id="FBC69B02FFCBFF94FBB7FDD9FB1768D9" box="[1052,1153,628,649]" italics="true" pageId="41" pageNumber="42">H. fucata</emphasis>
and
<emphasis id="FBC69B02FFCBFF94FB13FDD9FABA68D9" box="[1208,1324,628,649]" italics="true" pageId="41" pageNumber="42">H. dollfusi</emphasis>
need to be conducted to explore these possible relationships further.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:number id="18BE0FBAF4728672964A7588642DB527">1</mods:number>
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<mods:start id="DE74126ED1EF5CA20F898A91AA56B49F">1</mods:start>
<mods:end id="5B66E435E91F96D2AE668A7EF16551D6">74</mods:end>
</mods:extent>
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<mods:classification id="18165041FA92A6FB41D69AF955C52292">journal article</mods:classification>
<mods:identifier id="3802F0630FE4EEF946CB7A106D62AA12" type="CLB-Dataset">27363</mods:identifier>
<mods:identifier id="5D5391A30B98873D0D5A5B40BC120675" type="DOI">10.1093/zoolinnean/zly048</mods:identifier>
<mods:identifier id="07552BEAF594000B09502E0095FC73B1" type="GBIF-Dataset">49bea467-fac6-484a-9902-05483dec9a52</mods:identifier>
<mods:identifier id="F588C05A94DDF5C4FBDE2B54B20846A3" type="Zenodo-Dep">2632578</mods:identifier>
<mods:identifier id="F214DC9F7269627C9994ACE649AD7B9F" type="ZooBank">urn:lsid:zoobank.org:pub:F0065FD2-417C</mods:identifier>
</mods:mods>
<treatment id="411BF606FFC9FF93FC74FB78FAAC6BE3" ID-DOI="http://doi.org/10.5281/zenodo.5980694" ID-GBIF-Taxon="156146736" ID-Zenodo-Dep="5980694" LSID="urn:lsid:plazi:treatment:411BF606FFC9FF93FC74FB78FAAC6BE3" httpUri="http://treatment.plazi.org/id/411BF606FFC9FF93FC74FB78FAAC6BE3" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44">
<subSubSection id="81A8149BFFC9FF96FC74FB78FAB36F5B" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="C90D4710FFC9FF96FC74FB78FAB36F5B" blockId="43.[952,1317,1237,1330]" pageId="43" pageNumber="44">
<taxonomicName id="0EB23C93FFC9FF96FC74FB78FB2D6F5A" ID-CoL="3P35F" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="46" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<heading id="9245F07CFFC9FF96FC74FB78FB696EBB" allCaps="true" bold="true" box="[991,1279,1237,1260]" centered="true" fontSize="9" level="2" pageId="43" pageNumber="44" reason="4">
<emphasis id="FBC69B02FFC9FF96FC74FB78FB696EBB" bold="true" box="[991,1279,1237,1260]" italics="true" pageId="43" pageNumber="44">HYPSELODORIS ROSITOI</emphasis>
</heading>
<heading id="9245F07CFFC9FF96FC13FB5EFB2D6F5A" allCaps="true" box="[952,1211,1267,1291]" centered="true" fontSize="9" level="3" pageId="43" pageNumber="44" reason="4">GOSLINER &amp; JOHNSON</heading>
</taxonomicName>
<emphasis id="FBC69B02FFC9FF96FB69FB5AFAB36F5B" bold="true" box="[1218,1317,1267,1291]" pageId="43" pageNumber="44">SP. NOV.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC9FF96FC70FAB7FA946F62" box="[987,1282,1306,1330]" pageId="43" pageNumber="44" type="description">
<paragraph id="C90D4710FFC9FF96FC70FAB7FA946F62" blockId="43.[952,1317,1237,1330]" box="[987,1282,1306,1330]" pageId="43" pageNumber="44">
<heading id="9245F07CFFC9FF96FC70FAB7FA946F62" allCaps="true" box="[987,1282,1306,1330]" centered="true" fontSize="10" level="1" pageId="43" pageNumber="44" reason="1">(FIGS 2N, 13J, 25A, B, 26)</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC9FF96FC90FACEFBA06FC7" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="C90D4710FFC9FF96FC90FACEFBA06FC7" blockId="43.[827,1442,1378,1431]" pageId="43" pageNumber="44">
<emphasis id="FBC69B02FFC9FF96FC90FACEFCEF6F28" box="[827,889,1379,1400]" italics="true" pageId="43" pageNumber="44">LSID</emphasis>
:
<uri id="BD234B12FFC9FF96FC2FFACFFBA06FC7" pageId="43" pageNumber="44">
urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFC9FF96FB3BFACFFBA06FC7" pageId="43" pageNumber="44">0FA7479F-032D-403E- 8D72-69A1A90623DB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC9FF96FC90FA79FAD56CA0" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="C90D4710FFC9FF96FC90FA79FC726FBB" blockId="43.[827,1443,1491,1668]" box="[827,996,1491,1515]" pageId="43" pageNumber="44">
<heading id="9245F07CFFC9FF96FC90FA79FC726FBB" box="[827,996,1491,1515]" fontSize="10" level="4" pageId="43" pageNumber="44" reason="1">
<emphasis id="FBC69B02FFC9FF96FC90FA79FC726FBB" box="[827,996,1491,1515]" italics="true" pageId="43" pageNumber="44">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC9FF96FC90FA59FAB36CD4" blockId="43.[827,1443,1491,1668]" pageId="43" pageNumber="44">
<materialsCitation id="79DA4D4DFFC9FF96FC90FA59FAB36CD4" ID-GBIF-Occurrence="2235579101" collectingDate="2011-05-16" collectionCode="NMP" country="Philippines" county="Luzon" elevation="30" latitude="13.628" location="Malajibomanoc" longLatPrecision="76" longitude="120.966" municipality="Bay" pageId="43" pageNumber="44" specimenCode="CASIZ 186099" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
<typeStatus id="1609F9B2FFC9FF96FC90FA59FC316C59" box="[827,935,1524,1545]" pageId="43" pageNumber="44" type="holotype">
<emphasis id="FBC69B02FFC9FF96FC90FA59FC316C59" box="[827,935,1524,1545]" italics="true" pageId="43" pageNumber="44">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFC9FF96FC1CFA59FC606C59" box="[951,1014,1524,1545]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="43" pageNumber="44">NMP</collectionCode>
0 41283 (formerly
<specimenCode id="9914EF6BFFC9FF96FB4BFA59FA006C5A" box="[1248,1430,1524,1546]" collectionCode="CASIZ" pageId="43" pageNumber="44">CASIZ 186099</specimenCode>
), s u b s a m p l e d f o r m o l e c u l a r s t u d y, d i s s e c t e d,
<location id="CC6D11CBFFC9FF96FC90F99CFC796C17" LSID="urn:lsid:plazi:treatment:411BF606FFC9FF93FC74FB78FAAC6BE3:CC6D11CBFFC9FF96FC90F99CFC796C17" box="[827,1007,1585,1607]" country="Philippines" county="Luzon" latitude="13.628" longLatPrecision="76" longitude="120.966" municipality="Bay" name="Malajibomanoc" pageId="43" pageNumber="44" stateProvince="Batangas">Malajibomanoc</location>
(
<location id="CC6D11CBFFC9FF96FC54F99CFA8D6C17" LSID="urn:lsid:plazi:treatment:411BF606FFC9FF93FC74FB78FAAC6BE3:CC6D11CBFFC9FF96FC54F99CFA8D6C17" box="[1023,1307,1585,1607]" country="Philippines" county="Luzon" latitude="13.628" longLatPrecision="76" longitude="120.966" municipality="Bay" name="Chicken Feather Island" pageId="43" pageNumber="44" stateProvince="Batangas">Chicken Feather Island</location>
),
<geoCoordinate id="AC8621D7FFC9FF96FA9AF99CFA086C17" box="[1329,1438,1585,1607]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="13.628">13.628°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC9FF96FC90F9FDFC2B6C36" box="[827,957,1616,1638]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="120.966">120.966°E</geoCoordinate>
,
<elevation id="429FA023FFC9FF96FC67F9FDFB9D6C35" box="[972,1035,1616,1637]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="44" unit="m" value="30.0">
<quantity id="0E4AEAF5FFC9FF96FC67F9FDFB9D6C35" box="[972,1035,1616,1637]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="44" unit="m" value="30.0">30 m</quantity>
</elevation>
depth,
<collectingRegion id="0B7689F2FFC9FF96FBC4F9FDFB736C35" box="[1135,1253,1616,1637]" country="Philippines" name="Batangas" pageId="43" pageNumber="44">Batangas</collectingRegion>
<collectingMunicipality id="2969DD6AFFC9FF96FB44F9FDFA896C35" box="[1263,1311,1616,1637]" pageId="43" pageNumber="44">Bay</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFC9FF96FA85F9FDFC366CD4" country="Philippines" name="Batangas" pageId="43" pageNumber="44">Batangas Province</collectingRegion>
,
<collectingCounty id="206C3F9CFFC9FF96FC01F9C2FC656CD4" box="[938,1011,1647,1668]" pageId="43" pageNumber="44">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFC9FF96FC55F9C3FB176CD4" box="[1022,1153,1646,1668]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC9FF96FB27F9C3FAB76CD4" box="[1164,1313,1646,1668]" pageId="43" pageNumber="44" value="2011-05-16">
<date id="BD0C61D0FFC9FF96FB27F9C3FAB76CD4" box="[1164,1313,1646,1668]" pageId="43" pageNumber="44" value="2011-05-16">16 May 2011</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFC9FF96FC90F933FAD56CA0" blockId="43.[827,1443,1693,1776]" pageId="43" pageNumber="44">
<materialsCitation id="79DA4D4DFFC9FF96FC90F933FAA96CA0" ID-GBIF-Occurrence="2235579091" collectingDate="2009-05" collectionCode="CASIZ" country="Philippines" county="Company" location="Luzon" municipality="Denis Ty of Aquascapes" pageId="43" pageNumber="44" specimenCode="CASIZ 220675" specimenCount="1" stateProvince="Cavite" typeStatus="paratype">
<typeStatus id="1609F9B2FFC9FF96FC90F933FC096CE3" box="[827,927,1694,1715]" pageId="43" pageNumber="44" type="paratype">
<emphasis id="FBC69B02FFC9FF96FC90F933FC096CE3" box="[827,927,1694,1715]" italics="true" pageId="43" pageNumber="44">Paratype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFC9FF96FC0CF930FBDF6CE3" box="[935,1097,1693,1715]" collectionCode="CASIZ" pageId="43" pageNumber="44">
CASIZ
<date id="BD0C61D0FFC9FF96FC5DF930FBDF6CE3" box="[1014,1097,1693,1715]" pageId="43" pageNumber="44" value="75-06-22">220675</date>
</specimenCode>
, one specimen,
<collectingRegion id="0B7689F2FFC9FF96FB45F930FA096CE3" box="[1262,1439,1693,1715]" country="Philippines" name="Cavite" pageId="43" pageNumber="44">Cavite Province</collectingRegion>
,
<location id="CC6D11CBFFC9FF96FC90F911FC136C81" LSID="urn:lsid:plazi:treatment:411BF606FFC9FF93FC74FB78FAAC6BE3:CC6D11CBFFC9FF96FC90F911FC136C81" box="[827,901,1724,1745]" country="Philippines" county="Company" municipality="Denis Ty of Aquascapes" name="Luzon" pageId="43" pageNumber="44" stateProvince="Cavite">Luzon</location>
,
<collectingCountry id="B1A50780FFC9FF96FC3AF911FB8E6C82" box="[913,1048,1724,1746]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC9FF96FB8FF911FB0F6C81" box="[1060,1177,1724,1745]" pageId="43" pageNumber="44" value="2009-05">
<date id="BD0C61D0FFC9FF96FB8FF911FB0F6C81" box="[1060,1177,1724,1745]" pageId="43" pageNumber="44" value="2009-05">May 2009</date>
</collectingDate>
, specimen provided by
<collectingMunicipality id="2969DD6AFFC9FF96FC90F977FBD36CA0" box="[827,1093,1754,1776]" pageId="43" pageNumber="44">Denis Ty of Aquascapes</collectingMunicipality>
<collectingCountry id="B1A50780FFC9FF96FBE7F977FB5A6CA0" box="[1100,1228,1754,1776]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
<collectingCounty id="206C3F9CFFC9FF96FB78F976FAA96CA0" box="[1235,1343,1755,1776]" pageId="43" pageNumber="44">Company</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC9FF96FC90F8BAFB706D3C" pageId="43" pageNumber="44" type="distribution">
<paragraph id="C90D4710FFC9FF96FC90F8BAFBEE6D7F" blockId="43.[827,1443,1815,1900]" box="[827,1144,1815,1839]" pageId="43" pageNumber="44">
<heading id="9245F07CFFC9FF96FC90F8BAFBEE6D7F" box="[827,1144,1815,1839]" fontSize="10" level="4" pageId="43" pageNumber="44" reason="1">
<emphasis id="FBC69B02FFC9FF96FC90F8BAFBEE6D7F" box="[827,1144,1815,1839]" italics="true" pageId="43" pageNumber="44">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC9FF96FC90F895FB706D3C" blockId="43.[827,1443,1815,1900]" pageId="43" pageNumber="44">
Known only from the
<collectingRegion id="0B7689F2FFC9FF96FB93F895FB306D1D" box="[1080,1190,1848,1869]" country="Philippines" name="Batangas" pageId="43" pageNumber="44">Batangas</collectingRegion>
and
<collectingRegion id="0B7689F2FFC9FF96FB4AF895FABB6D1E" box="[1249,1325,1848,1870]" country="Philippines" name="Cavite" pageId="43" pageNumber="44">Cavite</collectingRegion>
provinces of Luzon,
<collectingCountry id="B1A50780FFC9FF96FC01F8FBFBBA6D3C" box="[938,1068,1878,1900]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<caption id="9DCD1798FFCEFF91FF3AFA16FA046CCF" httpUri="https://zenodo.org/record/2632832/files/figure.png" pageId="44" pageNumber="45" startId="44.[145,227,1467,1489]" targetBox="[305,1265,195,1426]" targetPageId="44">
<paragraph id="C90D4710FFCEFF91FF3AFA16FA046CCF" blockId="44.[145,1426,1467,1695]" pageId="44" pageNumber="45">
<emphasis id="FBC69B02FFCEFF91FF3AFA16FE9B6F80" bold="true" box="[145,269,1467,1489]" pageId="44" pageNumber="45">Figure 25.</emphasis>
Living animals. A,
<taxonomicName id="0EB23C93FFCEFF91FE40FA16FC546F81" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[491,962,1467,1489]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE40FA16FD496F80" bold="true" box="[491,735,1467,1489]" italics="true" pageId="44" pageNumber="45">Hypselodoris rositoi</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FC61FA11FBB06F81" box="[970,1062,1468,1489]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FC61FA11FBB06F81" bold="true" box="[970,1062,1468,1489]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41283, Tingloy, Batangas, Philippines. B,
<taxonomicName id="0EB23C93FFCEFF91FE1AFA74FC9E6FBF" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[433,776,1497,1519]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE1AFA74FDB26FBE" bold="true" box="[433,548,1497,1518]" italics="true" pageId="44" pageNumber="45">H. rositoi</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FCBBFA74FCFB6FBE" box="[784,877,1497,1518]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FCBBFA74FCFB6FBE" bold="true" box="[784,877,1497,1518]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, specimen from Albatross Expedition, illustrated by Kumataro Ito. C,
<taxonomicName id="0EB23C93FFCEFF91FEDEFA5BFC3B6C5C" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[373,941,1526,1548]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FEDEFA5BFD5C6C5C" bold="true" box="[373,714,1526,1548]" italics="true" pageId="44" pageNumber="45">Hypselodoris variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FC1EFA5BFB876C5B" box="[949,1041,1526,1547]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FC1EFA5BFB876C5B" bold="true" box="[949,1041,1526,1547]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 177618, Tingloy, Batangas, Philippines. D,
<taxonomicName id="0EB23C93FFCEFF91FE36F9B9FCAF6C79" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[413,825,1555,1577]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE36F9B9FDF36C79" bold="true" box="[413,613,1555,1577]" italics="true" pageId="44" pageNumber="45">H. variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FCEBF9B9FC006C79" box="[832,918,1556,1577]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FCEBF9B9FC006C79" bold="true" box="[832,918,1556,1577]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41285, Puerto Galera, Mindoro Oriental, Philippines. E,
<taxonomicName id="0EB23C93FFCEFF91FE3AF99CFCBC6C17" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[401,810,1584,1607]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE3AF99CFDCE6C16" bold="true" box="[401,600,1584,1606]" italics="true" pageId="44" pageNumber="45">H. variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FC9AF99CFC106C16" box="[817,902,1585,1606]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FC9AF99CFC106C16" bold="true" box="[817,902,1585,1606]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 208189, Puerto Galera, Mindoro Oriental, Philippines. F,
<taxonomicName id="0EB23C93FFCEFF91FE31F9E3FCD16C34" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[410,839,1614,1636]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE31F9E3FDFC6C34" bold="true" box="[410,618,1614,1636]" italics="true" pageId="44" pageNumber="45">H. variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FCE5F9E3FC3E6C33" box="[846,936,1614,1635]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FCE5F9E3FC3E6C33" bold="true" box="[846,936,1614,1635]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 212273, Mabini, Batangas, Philippines. G,
<taxonomicName id="0EB23C93FFCEFF91FE87F9C6FD606CD1" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[300,758,1643,1665]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="violacea" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE87F9C6FDB16CD1" bold="true" box="[300,551,1643,1665]" italics="true" pageId="44" pageNumber="45">Hypselodoris violacea</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FD57F9C1FCC66CD1" box="[764,848,1644,1665]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FD57F9C1FCC66CD1" bold="true" box="[764,848,1644,1665]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 0 71286, Manu Manu Island, Palawan, Batangas, Philippines. H,
<taxonomicName id="0EB23C93FFCEFF91FE31F924FD7E6CCF" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[410,744,1672,1695]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="violacea" status="sp. nov.">
<emphasis id="FBC69B02FFCEFF91FE31F924FD8C6CCE" bold="true" box="[410,538,1672,1694]" italics="true" pageId="44" pageNumber="45">H. violacea</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCEFF91FD46F924FCD76CCE" box="[749,833,1673,1694]" pageId="44" pageNumber="45" rank="species">
<emphasis id="FBC69B02FFCEFF91FD46F924FCD76CCE" bold="true" box="[749,833,1673,1694]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41286, Busuanga, Palawan, Philippines.
</paragraph>
</caption>
<subSubSection id="81A8149BFFCEFF91FF3AF96EFDD16D23" pageId="44" pageNumber="45" type="etymology">
<paragraph id="C90D4710FFCEFF91FF3AF96EFE846C8B" blockId="44.[145,761,1731,1908]" box="[145,274,1731,1755]" pageId="44" pageNumber="45">
<heading id="9245F07CFFCEFF91FF3AF96EFE846C8B" box="[145,274,1731,1755]" fontSize="10" level="4" pageId="44" pageNumber="45" reason="1">
<emphasis id="FBC69B02FFCEFF91FF3AF96EFE846C8B" box="[145,274,1731,1755]" italics="true" pageId="44" pageNumber="45">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCEFF91FF3AF94EFDD16D23" blockId="44.[145,761,1731,1908]" pageId="44" pageNumber="45">
The name
<taxonomicName id="0EB23C93FFCEFF91FEB2F949FEFC6CA9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[281,362,1764,1785]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCEFF91FEB2F949FEFC6CA9" box="[281,362,1764,1785]" italics="true" pageId="44" pageNumber="45">rositoi</emphasis>
</taxonomicName>
comes from the Latin for rose, referring to the distinctive rose-pink body colour that is predominant in this species and also honors Kumataro Ito, the artist on the 1908 Albatross Expedition who first illustrated this species (
<figureCitation id="51895B95FFCEFF91FE72F8F3FDAE6D24" box="[473,568,1886,1908]" captionStart="Figure 25" captionStartId="44.[145,227,1467,1489]" captionTargetBox="[305,1265,195,1426]" captionTargetId="figure@44.[305,1265,195,1426]" captionTargetPageId="44" captionText="Figure 25. Living animals. A, Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., holotype, NMP 041283, Tingloy, Batangas, Philippines. B, H. rositoi Gosliner &amp; Johnson sp. nov., specimen from Albatross Expedition, illustrated by Kumataro Ito. C, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177618, Tingloy, Batangas, Philippines. D, H. variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, Puerto Galera, Mindoro Oriental, Philippines. E, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208189, Puerto Galera, Mindoro Oriental, Philippines. F, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 212273, Mabini, Batangas, Philippines. G, Hypselodoris violacea Gosliner &amp; Johnson sp. nov., paratype, CASIZ 071286, Manu Manu Island, Palawan, Batangas, Philippines. H, H. violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, Busuanga, Palawan, Philippines." httpUri="https://zenodo.org/record/2632832/files/figure.png" pageId="44" pageNumber="45">Fig. 25B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCEFF93FC82F96EFDA76B68" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="C90D4710FFCEFF91FC82F96EFC236C8A" blockId="44.[809,1425,1731,1908]" box="[809,949,1731,1754]" pageId="44" pageNumber="45">
<heading id="9245F07CFFCEFF91FC82F96EFC236C8A" box="[809,949,1731,1754]" fontSize="10" level="4" pageId="44" pageNumber="45" reason="1">
<emphasis id="FBC69B02FFCEFF91FC82F96EFC236C8A" box="[809,949,1731,1754]" italics="true" pageId="44" pageNumber="45">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCEFF90FC82F94EFD986F28" blockId="44.[809,1425,1731,1908]" lastBlockId="45.[163,779,1133,1400]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="FBC69B02FFCEFF91FC82F94EFBBE6CA8" box="[809,1064,1763,1784]" italics="true" pageId="44" pageNumber="45">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFCEFF91FAA8F94EFAFC6CA9" box="[1283,1386,1763,1785]" captionStart="Figure 25" captionStartId="44.[145,227,1467,1489]" captionTargetBox="[305,1265,195,1426]" captionTargetId="figure@44.[305,1265,195,1426]" captionTargetPageId="44" captionText="Figure 25. Living animals. A, Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., holotype, NMP 041283, Tingloy, Batangas, Philippines. B, H. rositoi Gosliner &amp; Johnson sp. nov., specimen from Albatross Expedition, illustrated by Kumataro Ito. C, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177618, Tingloy, Batangas, Philippines. D, H. variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, Puerto Galera, Mindoro Oriental, Philippines. E, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208189, Puerto Galera, Mindoro Oriental, Philippines. F, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 212273, Mabini, Batangas, Philippines. G, Hypselodoris violacea Gosliner &amp; Johnson sp. nov., paratype, CASIZ 071286, Manu Manu Island, Palawan, Batangas, Philippines. H, H. violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, Busuanga, Palawan, Philippines." httpUri="https://zenodo.org/record/2632832/files/figure.png" pageId="44" pageNumber="45">Fig. 25A</figureCitation>
) of moderately large size, reaching
<quantity id="0E4AEAF5FFCEFF91FB34F8AFFA8F6D47" box="[1183,1305,1794,1816]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="44" pageNumber="45" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 mm</quantity>
in length. Entire dorsal surface deep rose pink, with thick white band encircling the margin of notum. Sides of body lighter pink, fading to white margin of foot. Five unipinnate gill branches on notum.
<typeStatus id="1609F9B2FFCFFF90FDF5FBC0FD5D6ED3" box="[606,715,1133,1155]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
with four large gill branches and five smaller ones, possibly indicating regrowth from damage.
<typeStatus id="1609F9B2FFCFFF90FD84FB06FD006E90" box="[559,662,1195,1216]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
with nine unipinnate gill branches. Gill pocket well elevated. Gill branches elongate, deep orange, with lighter pigment on inner rachis of each branch. Base of gill pocket deep pink. Bulb of rhinophores bright orange with redder apex. Bulb with ~25–32 densely packed lamellae. Base of rhinophore sheath deep pink.
</paragraph>
<caption id="9DCD1798FFCFFF90FF08FBB8FCDC6E19" httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="45" pageNumber="46" startId="45.[163,241,1045,1067]" targetBox="[227,1379,195,1004]" targetPageId="45">
<paragraph id="C90D4710FFCFFF90FF08FBB8FCDC6E19" blockId="45.[163,1443,1045,1097]" pageId="45" pageNumber="46">
<emphasis id="FBC69B02FFCFFF90FF08FBB8FE8E6E7A" bold="true" box="[163,280,1045,1067]" pageId="45" pageNumber="46">Figure 26.</emphasis>
<taxonomicName id="0EB23C93FFCFFF90FE88FBB8FD496E7B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,735,1045,1067]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<emphasis id="FBC69B02FFCFFF90FE88FBB8FD9C6E7A" bold="true" box="[291,522,1045,1067]" italics="true" pageId="45" pageNumber="46">Hypselodoris rositoi</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCFFF90FD4FFBB8FCAC6E7A" box="[740,826,1045,1066]" pageId="45" pageNumber="46" rank="species">
<emphasis id="FBC69B02FFCFFF90FD4FFBB8FCAC6E7A" bold="true" box="[740,826,1045,1066]" pageId="45" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, holotype, NMP 0 41283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFCFFF90FF08FA0CFD326FA4" blockId="45.[163,779,1441,1524]" pageId="45" pageNumber="46">
<emphasis id="FBC69B02FFCFFF90FF08FA0CFEC76FE6" box="[163,337,1441,1462]" italics="true" pageId="45" pageNumber="46">Mantle glands</emphasis>
: Small subcutaneous mantle glands present along entire mantle margin, somewhat enlarged anteriorly and posteriorly (
<figureCitation id="51895B95FFCFFF90FDEAFA73FD036FA3" box="[577,661,1502,1524]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="45" pageNumber="46">Fig. 2N</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFCFFF90FF08F9B0FB446F4C" blockId="45.[163,779,1565,1893]" lastBlockId="45.[827,1443,1133,1308]" pageId="45" pageNumber="46">
<emphasis id="FBC69B02FFCFFF90FF08F9B0FEF86C62" box="[163,366,1565,1586]" italics="true" pageId="45" pageNumber="46">Buccal armature</emphasis>
: Muscular portion of buccal mass slightly larger than oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFCFFF90FDEFF9D4FD376CDF" box="[580,673,1657,1679]" captionStart="Figure 26" captionStartId="45.[163,241,1045,1067]" captionTargetBox="[227,1379,195,1004]" captionTargetId="figure@45.[227,1379,195,1004]" captionTargetPageId="45" captionText="Figure 26. Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="45" pageNumber="46">Fig. 26A</figureCitation>
). Rodlets narrow and short with long base and evenly curved, with single, acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFCFFF90FF08F978FE946CBB" box="[163,258,1749,1771]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
(CASIZ 186099) 66 × 46.0.46. Rachidian row of teeth absent (
<figureCitation id="51895B95FFCFFF90FE96F95EFE0D6D59" box="[317,411,1779,1801]" captionStart="Figure 26" captionStartId="45.[163,241,1045,1067]" captionTargetBox="[227,1379,195,1004]" captionTargetId="figure@45.[227,1379,195,1004]" captionTargetPageId="45" captionText="Figure 26. Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="45" pageNumber="46">Fig. 26B</figureCitation>
). Innermost lateral teeth having one short denticle on inner side of bifid primary cusp, lacking outer denticles. Next several laterals lacking inner and outer denticles, possessing only primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFCFFF90FBE7FBC0FB3C6ED3" box="[1100,1194,1133,1155]" captionStart="Figure 26" captionStartId="45.[163,241,1045,1067]" captionTargetBox="[227,1379,195,1004]" captionTargetId="figure@45.[227,1379,195,1004]" captionTargetPageId="45" captionText="Figure 26. Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="45" pageNumber="46">Fig. 26C</figureCitation>
) all lacking inner and outer denticles, with exception that single outer denticle may be present on some teeth. Outermost teeth having a narrower base and lacking denticles. Three to four teeth inside outermost teeth, with one to four denticles on outer side of bifid cusp (
<figureCitation id="51895B95FFCFFF90FBCDFAABFB526F4C" box="[1126,1220,1286,1308]" captionStart="Figure 26" captionStartId="45.[163,241,1045,1067]" captionTargetBox="[227,1379,195,1004]" captionTargetId="figure@45.[227,1379,195,1004]" captionTargetPageId="45" captionText="Figure 26. Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="45" pageNumber="46">Fig. 26D</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFCFFF93FC90FAE8FDA76B68" blockId="45.[827,1443,1349,1892]" lastBlockId="46.[145,761,198,312]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="FBC69B02FFCFFF90FC90FAE8FBAD6F0A" box="[827,1083,1349,1370]" italics="true" pageId="45" pageNumber="46">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFCFFF90FC90FACEFC306F29" box="[827,934,1379,1401]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFCFFF90FBCEFACEFB5A6F29" box="[1125,1228,1379,1401]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="45" pageNumber="46">Fig. 13J</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing abruptly as it transitions into muscular ejaculatory portion. Ejaculatory portion short, curved, narrow, entering short, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled bursa copulatrix. Receptaculum seminis appressed against vagina in distal half of vaginal length. Moderately short uterine duct emerging from vagina close to receptaculum and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCCFF93FF3AFEF2FAAC6BE3" pageId="46" pageNumber="47" type="discussion">
<paragraph id="C90D4710FFCCFF93FF3AFEF2FF686B27" blockId="46.[145,762,351,1111]" box="[145,254,351,375]" pageId="46" pageNumber="47">
<heading id="9245F07CFFCCFF93FF3AFEF2FF686B27" box="[145,254,351,375]" fontSize="10" level="4" pageId="46" pageNumber="47" reason="1">
<emphasis id="FBC69B02FFCCFF93FF3AFEF2FF686B27" box="[145,254,351,375]" italics="true" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCCFF93FF3AFE2DFE4669EE" blockId="46.[145,762,351,1111]" pageId="46" pageNumber="47">
In our phylogenetic analyses,
<taxonomicName id="0EB23C93FFCCFF93FE4EFE2DFDD96BC5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[485,591,384,405]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FE4EFE2DFDD96BC5" box="[485,591,384,405]" italics="true" pageId="46" pageNumber="47">H. rositoi</emphasis>
</taxonomicName>
is sister to the rest of the
<taxonomicName id="0EB23C93FFCCFF93FEB8FE32FE0A6BE3" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[275,412,414,436]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FEB8FE32FE0A6BE3" box="[275,412,414,436]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade. In the ABGD analysis,
<taxonomicName id="0EB23C93FFCCFF93FF3AFE13FF656B83" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,243,446,467]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FF3AFE13FF656B83" box="[145,243,446,467]" italics="true" pageId="46" pageNumber="47">H.rositoi</emphasis>
</taxonomicName>
is indicated to represent a distinct species and is&gt; 13% different in its
<emphasis id="FBC69B02FFCCFF93FED0FE71FE3F6BA1" box="[379,425,476,497]" italics="true" pageId="46" pageNumber="47">COI</emphasis>
gene compared with any other members of the
<taxonomicName id="0EB23C93FFCCFF93FEECFE56FE5C685F" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[327,458,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FEECFE56FE5C685F" box="[327,458,506,528]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade.
<taxonomicName id="0EB23C93FFCCFF93FDB3FE57FD6F6840" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[536,761,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FDB3FE57FD6F6840" box="[536,761,506,528]" italics="true" pageId="46" pageNumber="47">Hypselodoris rositoi</emphasis>
</taxonomicName>
is unique among members of the
<taxonomicName id="0EB23C93FFCCFF93FDB9FDB7FD01687E" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[530,663,537,559]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FDB9FDB7FD01687E" box="[530,663,537,559]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade in having a bright rose-pink body colour. All other members of the clade are light pink to purple.
<taxonomicName id="0EB23C93FFCCFF93FDF6FDFBFF4068DB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FDF6FDFBFF4068DB" italics="true" pageId="46" pageNumber="47">Hypselodoris rositoi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FF49FDDBFECD68DA" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[226,347,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="violacea" status="sp. nov.">
<emphasis id="FBC69B02FFCCFF93FF49FDDBFECD68DA" box="[226,347,629,651]" italics="true" pageId="46" pageNumber="47">H. violacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCCFF93FEC9FDDBFE2768DB" box="[354,433,630,651]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="0EB23C93FFCCFF93FE4CFDDBFD3568DA" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[487,675,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFCCFF93FE4CFDDBFD3568DA" box="[487,675,629,651]" italics="true" pageId="46" pageNumber="47">H. variobranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCCFF93FD01FDDBFD6C68DB" box="[682,762,630,651]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
are the only members of the
<taxonomicName id="0EB23C93FFCCFF93FE74FD39FDF268F9" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[479,612,660,681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FE74FD39FDF268F9" box="[479,612,660,681]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade with a broad, solid white marginal band.
<taxonomicName id="0EB23C93FFCCFF93FDBCFD1FFD6E6898" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[535,760,690,712]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FDBCFD1FFD6E6898" box="[535,760,690,712]" italics="true" pageId="46" pageNumber="47">Hypselodoris rositoi</emphasis>
</taxonomicName>
is also the only member of the
<taxonomicName id="0EB23C93FFCCFF93FE5EFD7FFDEC68B6" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[501,634,721,743]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FE5EFD7FFDEC68B6" box="[501,634,721,743]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade with obvious mantle glands. Internally,
<taxonomicName id="0EB23C93FFCCFF93FD90FD5DFD386955" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[571,686,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FD90FD5DFD386955" box="[571,686,752,773]" italics="true" pageId="46" pageNumber="47">H. rositoi</emphasis>
</taxonomicName>
has a radula with fewer teeth per half row (46) than found in other members of the
<taxonomicName id="0EB23C93FFCCFF93FE2BFC83FE696912" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[384,511,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FE2BFC83FE696912" box="[384,511,813,835]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade. It is also unique in lacking outer denticles on the vast majority of its middle lateral teeth. All the remaining members of the
<taxonomicName id="0EB23C93FFCCFF93FF3AFC27FE8369CE" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[145,277,905,927]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FF3AFC27FE8369CE" box="[145,277,905,927]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade have numerous prominent denticles on their middle lateral teeth.
</paragraph>
<paragraph id="C90D4710FFCCFF93FF02FC6BFAAC6BE3" blockId="46.[145,762,351,1111]" lastBlockId="46.[809,1425,198,435]" pageId="46" pageNumber="47">
The reproductive system of members of the
<taxonomicName id="0EB23C93FFCCFF93FF3AFC4BFE8169AA" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[145,279,997,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FF3AFC4BFE8169AA" box="[145,279,997,1019]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
clade has slight variation. In
<taxonomicName id="0EB23C93FFCCFF93FDD8FC4BFD6F69AA" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[627,761,997,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFCCFF93FDD8FC4BFD6F69AA" box="[627,761,997,1019]" italics="true" pageId="46" pageNumber="47">H. bullockii</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFCCFF93FF32FBA9FF606E49" box="[153,246,1028,1050]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="46" pageNumber="47">Fig. 13F</figureCitation>
) and
<taxonomicName id="0EB23C93FFCCFF93FE93FBA9FE516E49" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[312,455,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFCCFF93FE93FBA9FE516E49" box="[312,455,1028,1049]" italics="true" pageId="46" pageNumber="47">H. apolegma</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFCCFF93FE7CFBA9FDBC6E49" box="[471,554,1028,1050]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="46" pageNumber="47">Fig. 4D</figureCitation>
), the vaginal duct is relatively short, whereas in
<taxonomicName id="0EB23C93FFCCFF93FE5CFB8EFDF26E68" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[503,612,1059,1080]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FE5CFB8EFDF26E68" box="[503,612,1059,1080]" italics="true" pageId="46" pageNumber="47">H. rositoi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FDD9FB8EFD646E67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[626,754,1058,1080]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFCCFF93FDD9FB8EFD646E67" box="[626,754,1058,1080]" italics="true" pageId="46" pageNumber="47">H. violacea</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FF3AFBEFFEF76E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,353,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFCCFF93FF3AFBEFFEF76E06" box="[145,353,1089,1111]" italics="true" pageId="46" pageNumber="47">H. variobranchia</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FEDAFBEFFE486E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[369,478,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFCCFF93FEDAFBEFFE486E06" box="[369,478,1089,1111]" italics="true" pageId="46" pageNumber="47">H. brycei</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFCCFF93FDB5FBEFFD5F6E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[542,713,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFCCFF93FDB5FBEFFD5F6E06" box="[542,713,1089,1111]" italics="true" pageId="46" pageNumber="47">H. melanesica</emphasis>
</taxonomicName>
the vagina is more elongate. In
<taxonomicName id="0EB23C93FFCCFF93FB2AFF6AFB656A8C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1153,1267,199,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FB2AFF6AFB656A8C" box="[1153,1267,199,220]" italics="true" pageId="46" pageNumber="47">H. rositoi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FAA8FF6AFA1D6A8B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1283,1419,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFCCFF93FAA8FF6AFA1D6A8B" box="[1283,1419,198,220]" italics="true" pageId="46" pageNumber="47">H. violacea</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FC82FF4BFC1A6AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,908,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFCCFF93FC82FF4BFC1A6AAA" box="[809,908,229,251]" italics="true" pageId="46" pageNumber="47">H. brycei</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFCCFF93FC68FF4BFBF46AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[963,1122,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFCCFF93FC68FF4BFBF46AAA" box="[963,1122,229,251]" italics="true" pageId="46" pageNumber="47">H. melanesica</emphasis>
</taxonomicName>
, the uterine duct branches from the middle of the vagina near the receptaculum seminis, whereas in
<taxonomicName id="0EB23C93FFCCFF93FBA5FE8EFB586B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1038,1230,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFCCFF93FBA5FE8EFB586B67" box="[1038,1230,290,312]" italics="true" pageId="46" pageNumber="47">H. variobranchia</emphasis>
</taxonomicName>
, the uterine duct branches near the base of the vagina. In
<taxonomicName id="0EB23C93FFCCFF93FB5DFEEFFAC96B07" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1270,1375,322,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFCCFF93FB5DFEEFFAC96B07" box="[1270,1375,322,343]" italics="true" pageId="46" pageNumber="47">H. rositoi</emphasis>
</taxonomicName>
, the ejaculatory portion of the vas deferens is relatively short, whereas it is much longer in
<taxonomicName id="0EB23C93FFCCFF93FB57FED2FA1D6BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1276,1419,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFCCFF93FB57FED2FA1D6BC3" box="[1276,1419,382,404]" italics="true" pageId="46" pageNumber="47">H. violacea</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FC82FE33FC7C6BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,1002,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFCCFF93FC82FE33FC7C6BE2" box="[809,1002,413,435]" italics="true" pageId="46" pageNumber="47">H. variobranchia</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFCCFF93FC5CFE33FBCA6BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1015,1116,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFCCFF93FC5CFE33FBCA6BE2" box="[1015,1116,413,435]" italics="true" pageId="46" pageNumber="47">H. brycei</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFCCFF93FB3FFE33FAA26BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1172,1332,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFCCFF93FB3FFE33FAA26BE2" box="[1172,1332,413,435]" italics="true" pageId="46" pageNumber="47">H. melanesica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1C56A933D1AA2553FA38EB81AA42C791">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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<mods:namePart id="E8324EB459298F81FA6A8CB1A0D4DE78">Gosliner, Terrence M.</mods:namePart>
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<mods:title id="F62319FCF804249D8EF8AE41F15C5D44">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="2C10CF07514A962E90ECA0ACA86B85A6">
<mods:date id="FF96372A4355967B57EE64FE975D06A8">2018</mods:date>
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<mods:number id="73E6B0A46C5972608269F662C612E095">20</mods:number>
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<treatment id="411BF606FFCBFF96FC97FD48FA846EC3" ID-DOI="http://doi.org/10.5281/zenodo.5980692" ID-GBIF-Taxon="156146735" ID-Zenodo-Dep="5980692" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3" httpUri="http://treatment.plazi.org/id/411BF606FFCBFF96FC97FD48FA846EC3" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
<subSubSection id="81A8149BFFCBFF94FC97FD48FA3768AD" box="[828,1441,741,765]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="C90D4710FFCBFF94FC97FD48FA3768AD" blockId="41.[828,1441,741,803]" box="[828,1441,741,765]" pageId="41" pageNumber="42">
<heading id="9245F07CFFCBFF94FC97FD48FA3768AD" allCaps="true" box="[828,1441,741,765]" centered="true" fontSize="9" level="3" pageId="41" pageNumber="42" reason="5">
<taxonomicName id="0EB23C93FFCBFF94FC97FD48FAA168AB" ID-CoL="3P35D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[828,1335,741,765]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="43" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFCBFF94FC97FD48FBB868AB" bold="true" box="[828,1070,741,764]" italics="true" pageId="41" pageNumber="42">HYPSELODORIS ROO</emphasis>
GOSLINER &amp; JOHNSON
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<emphasis id="FBC69B02FFCBFF94FA95FD45FA3768AD" bold="true" box="[1342,1441,741,765]" pageId="41" pageNumber="42">SP. NOV.</emphasis>
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(FIGS 2M,
<date id="BD0C61D0FFCBFF94FBF9FCA6FBEF6973" box="[1106,1145,779,803]" pageId="41" pageNumber="42" value="2013-01">13I</date>
, 18H–J, 24)
</heading>
</paragraph>
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<paragraph id="C90D4710FFCBFF94FC90FCF9FBE069D8" blockId="41.[827,1444,852,904]" pageId="41" pageNumber="42">
<emphasis id="FBC69B02FFCBFF94FC90FCF9FCE86939" box="[827,894,852,873]" italics="true" pageId="41" pageNumber="42">LSID</emphasis>
:
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urn:lsid:zoobank.org:act:
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<subSubSection id="81A8149BFFCBFF94FC90FC0BFB926E24" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="C90D4710FFCBFF94FC90FC0BFB19698B" blockId="41.[827,1443,934,1140]" pageId="41" pageNumber="42">
<taxonomicName id="0EB23C93FFCBFF94FC90FC0BFBB069EB" ID-CoL="3P343" authorityName="Rudman" authorityYear="1977" box="[827,1062,934,955]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFCBFF94FC90FC0BFBB069EB" box="[827,1062,934,955]" italics="true" pageId="41" pageNumber="42">Hypselodoris kanga</emphasis>
</taxonomicName>
misidentification, not
<taxonomicName id="0EB23C93FFCBFF94FA9DFC0AFC74698A" ID-CoL="3P343" authority="Rudman, 1977" authorityName="Rudman" authorityYear="1977" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFCBFF94FA9DFC0AFA3569EB" box="[1334,1443,934,956]" italics="true" pageId="41" pageNumber="42">H. kanga</emphasis>
<bibRefCitation id="AD233AE1FFCBFF94FC90FC68FC74698A" author="Rudman WB" box="[827,994,965,987]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="41" pageNumber="42" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFCBFF94FC46FC68FB1E698A" author="Krampf M." box="[1005,1160,965,987]" journalOrPublisher="Sydney: Australian Museum" pageId="41" pageNumber="42" publicationUrl="http://www.seaslugforum.net/find/20163" refId="ref43395" refString="Krampf M. 2007. Hypselodoris kanga laying eggs in Lembeh. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20163" title="Hypselodoris kanga laying eggs in Lembeh" type="url" volumeTitle="Sea slug forum" year="2007">Krampf, 2007</bibRefCitation>
.
</paragraph>
<paragraph id="C90D4710FFCBFF94FC90FC49FC646E67" blockId="41.[827,1443,934,1140]" pageId="41" pageNumber="42">
<taxonomicName id="0EB23C93FFCBFF94FC90FC49FBB069A9" ID-CoL="3P343" authorityName="Rudman" authorityYear="1977" box="[827,1062,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFCBFF94FC90FC49FBB069A9" box="[827,1062,996,1017]" italics="true" pageId="41" pageNumber="42">Hypselodoris kanga</emphasis>
</taxonomicName>
misidentification, not
<taxonomicName id="0EB23C93FFCBFF94FA9DFC49FC7C6E48" ID-CoL="3P343" authority="Rudman, 1977" authorityName="Rudman" authorityYear="1977" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFCBFF94FA9DFC49FA3569A9" box="[1334,1443,996,1017]" italics="true" pageId="41" pageNumber="42">H. kanga</emphasis>
<bibRefCitation id="AD233AE1FFCBFF94FC90FBAFFC7C6E48" author="Rudman WB" box="[827,1002,1026,1048]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="41" pageNumber="42" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFCBFF94FC5CFBAFFA8F6E48" author="Debelius H &amp; Kuiter R." box="[1015,1305,1026,1048]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="41" pageNumber="42" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 127, lower left photograph.
</paragraph>
<paragraph id="C90D4710FFCBFF94FC90FBEDFB926E24" blockId="41.[827,1443,934,1140]" pageId="41" pageNumber="42">
<taxonomicName id="0EB23C93FFCBFF94FC90FBEDFC406E05" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[827,982,1088,1109]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFCBFF94FC90FBEDFC406E05" box="[827,982,1088,1109]" italics="true" pageId="41" pageNumber="42">Hypselodoris</emphasis>
</taxonomicName>
sp. 7
<bibRefCitation id="AD233AE1FFCBFF94FBB4FBEDFA826E06" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1055,1300,1088,1110]" journalOrPublisher="Jacksonville: New World Press" pageId="41" pageNumber="42" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFCBFF94FB25FBEDFB5D6E05" box="[1166,1227,1088,1109]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
, 2015
</bibRefCitation>
: 258, upper right photograph.
</paragraph>
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<emphasis id="FBC69B02FFCBFF94FC90FB31FC726EE3" box="[827,996,1179,1203]" italics="true" pageId="41" pageNumber="42">Type material</emphasis>
</heading>
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<emphasis id="FBC69B02FFCBFF94FC90FB11FC306E81" box="[827,934,1212,1233]" italics="true" pageId="41" pageNumber="42">Holotype</emphasis>
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:
<collectionCode id="AFA3DFD5FFCBFF94FC11FB11FC6F6E81" box="[954,1017,1212,1233]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="41" pageNumber="42">NMP</collectionCode>
0 41282 (formerly
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CASIZ
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), subsampled for molecular study, dissected,
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,
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,
<collectingMunicipality id="2969DD6AFFCBFF94FA89FB54FCCF6F7D" pageId="41" pageNumber="42">Calumpan Pe</collectingMunicipality>
n i n s u l a, M a b i n i, B a t a n g a s, L u z o n I s l a n d,
<collectingCountry id="B1A50780FFCBFF94FC90FA9BFC286F1C" box="[827,958,1334,1356]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
,
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<date id="BD0C61D0FFCBFF94FC62FA9AFBCB6F1C" box="[969,1117,1335,1356]" pageId="41" pageNumber="42" value="2009-05-21">21 May 2009</date>
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,
<collectorName id="644722C6FFCBFF94FBCDFA9AFB716F1C" box="[1126,1255,1334,1356]" pageId="41" pageNumber="42">T. Gosliner.</collectorName>
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<emphasis id="FBC69B02FFCBFF94FC90FADBFC206FDB" box="[827,950,1398,1419]" italics="true" pageId="41" pageNumber="42">Paratypes</emphasis>
</typeStatus>
:
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,
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,
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,
<collectingMunicipality id="2969DD6AFFCBFF94FA8DFA39FCCF6F98" pageId="41" pageNumber="42">Calumpan Pe</collectingMunicipality>
n i n s u l a, M a b i n i, B a t a n g a s, L u z o n I s l a n d,
<collectingCountry id="B1A50780FFCBFF94FC90FA7CFC556FB7" box="[827,963,1489,1511]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
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<date id="BD0C61D0FFCBFF94FC64FA7CFBCD6FB6" box="[975,1115,1489,1511]" pageId="41" pageNumber="42" value="2011-05-04">4 May 2011</date>
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,
<collectorName id="644722C6FFCBFF94FBCDFA7FFB716FB7" box="[1126,1255,1489,1511]" pageId="41" pageNumber="42">T. Gosliner</collectorName>
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.
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, one specimen,
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dive site,
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,
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,
<collectingCountry id="B1A50780FFCBFF94FC90F980FC456C13" box="[827,979,1581,1603]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCBFF94FC48F980FB0B6C13" box="[995,1181,1581,1603]" pageId="41" pageNumber="42" value="2015-04-19">
<date id="BD0C61D0FFCBFF94FC48F980FB0B6C13" box="[995,1181,1581,1603]" pageId="41" pageNumber="42" value="2015-04-19">19 April 2015</date>
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,
<collectorName id="644722C6FFCBFF94FB06F983FAD66C13" box="[1197,1344,1581,1603]" pageId="41" pageNumber="42">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCBFF94FAFBF980FA366CCF" ID-GBIF-Occurrence="2235579046" collectingDate="2015-03-28" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="13.51356" location="Puerto Galera" longLatPrecision="1" longitude="120.95809" pageId="41" pageNumber="42" specimenCode="CASIZ 2" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCBFF94FAFBF980FCDF6C31" collectionCode="CASIZ" pageId="41" pageNumber="42">CASIZ 2</specimenCode>
0 8 5 4 1, o n e s p e c i m e n, G i a n t C l a m d i v e s i t e,
<geoCoordinate id="AC8621D7FFCBFF94FC90F9C7FC5C6CD0" box="[827,970,1642,1664]" direction="north" orientation="latitude" pageId="41" pageNumber="42" precision="1" value="13.51356">13.51356°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCBFF94FC7CF9C7FBE46CD0" box="[983,1138,1642,1664]" direction="east" orientation="longitude" pageId="41" pageNumber="42" precision="1" value="120.95809">120.95809°E</geoCoordinate>
,
<location id="CC6D11CBFFCBFF94FB2BF9C6FAA66CD0" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3:CC6D11CBFFCBFF94FB2BF9C6FAA66CD0" box="[1152,1328,1642,1664]" country="Philippines" latitude="13.51356" longLatPrecision="1" longitude="120.95809" name="Puerto Galera" pageId="41" pageNumber="42" stateProvince="Mindoro Oriental">Puerto Galera</location>
,
<collectingRegion id="0B7689F2FFCBFF94FA95F9C7FC306CCF" country="Philippines" name="Mindoro Oriental" pageId="41" pageNumber="42">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFCBFF94FC18F924FBD76CCF" box="[947,1089,1673,1695]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCBFF94FBE4F924FA9D6CCF" box="[1103,1291,1673,1695]" pageId="41" pageNumber="42" value="2015-03-28">
<date id="BD0C61D0FFCBFF94FBE4F924FA9D6CCF" box="[1103,1291,1673,1695]" pageId="41" pageNumber="42" value="2015-03-28">28 March 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFCBFF94FAB3F927FA366CCF" box="[1304,1440,1673,1695]" pageId="41" pageNumber="42">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCBFF94FC90F905FA366CAB" ID-GBIF-Occurrence="2235579086" collectingDate="2015-03-25" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="13.51356" location="Giant Clam" longLatPrecision="1" longitude="120.95809" municipality="Puerto Galera" pageId="41" pageNumber="42" specimenCode="CASIZ 208193" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCBFF94FC90F905FC7F6CED" box="[827,1001,1704,1725]" collectionCode="CASIZ" pageId="41" pageNumber="42">CASIZ 208193</specimenCode>
, one specimen,
<location id="CC6D11CBFFCBFF94FB0FF905FAB96CEE" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3:CC6D11CBFFCBFF94FB0FF905FAB96CEE" box="[1188,1327,1704,1726]" country="Philippines" latitude="13.51356" longLatPrecision="1" longitude="120.95809" municipality="Puerto Galera" name="Giant Clam" pageId="41" pageNumber="42" stateProvince="Mindoro Oriental">Giant Clam</location>
dive site,
<geoCoordinate id="AC8621D7FFCBFF94FC90F96BFC5C6C8C" box="[827,970,1734,1756]" direction="north" orientation="latitude" pageId="41" pageNumber="42" precision="1" value="13.51356">13.51356°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCBFF94FC7CF96BFBE46C8C" box="[983,1138,1734,1756]" direction="east" orientation="longitude" pageId="41" pageNumber="42" precision="1" value="120.95809">120.95809°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFCBFF94FB2BF96AFAA66C8C" box="[1152,1328,1734,1756]" pageId="41" pageNumber="42">Puerto Galera</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFCBFF94FA95F96BFC306CAB" country="Philippines" name="Mindoro Oriental" pageId="41" pageNumber="42">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFCBFF94FC18F948FBD76CAB" box="[947,1089,1765,1787]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCBFF94FBE4F948FA9D6CAB" box="[1103,1291,1765,1787]" pageId="41" pageNumber="42" value="2015-03-25">
<date id="BD0C61D0FFCBFF94FBE4F948FA9D6CAB" box="[1103,1291,1765,1787]" pageId="41" pageNumber="42" value="2015-03-25">25 March 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFCBFF94FAB3F94BFA366CAB" box="[1304,1440,1765,1787]" pageId="41" pageNumber="42">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCBFF94FC90F8A9FBD56D26" ID-GBIF-Occurrence="2235579049" collectingDate="2016-04-22" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Luzon Island" location="Murals" municipality="Mabini" pageId="41" pageNumber="42" specimenCode="CASIZ 217236" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFCBFF94FC90F8A9FC6E6D4A" box="[827,1016,1796,1818]" collectionCode="CASIZ" pageId="41" pageNumber="42">CASIZ 217236</specimenCode>
, one specimen,
<location id="CC6D11CBFFCBFF94FB6CF8A9FAB56D4A" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3:CC6D11CBFFCBFF94FB6CF8A9FAB56D4A" box="[1223,1315,1796,1818]" country="Philippines" county="Luzon Island" municipality="Mabini" name="Murals" pageId="41" pageNumber="42" stateProvince="Batangas">Murals</location>
dive site, 1 3.6 9 9 3°N, 1 2 0.8 8 2 4°E, C a l u m p a n
<location id="CC6D11CBFFCBFF94FAB3F88EFAA06D68" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3:CC6D11CBFFCBFF94FAB3F88EFAA06D68" box="[1304,1334,1827,1848]" country="Philippines" county="Luzon Island" municipality="Mabini" name="Pe" pageId="41" pageNumber="42" stateProvince="Batangas">Pe</location>
n i n s u l a,
<collectingMunicipality id="2969DD6AFFCBFF94FC90F8ECFC016D07" box="[827,919,1857,1879]" pageId="41" pageNumber="42">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFCBFF94FC0EF8EFFB886D07" box="[933,1054,1858,1879]" country="Philippines" name="Batangas" pageId="41" pageNumber="42">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFCBFF94FB86F8EFFB406D07" box="[1069,1238,1857,1879]" pageId="41" pageNumber="42">Luzon Island</collectingCounty>
,
<collectingCountry id="B1A50780FFCBFF94FB4EF8ECFAE16D07" box="[1253,1399,1857,1879]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCBFF94FA2DF8ECFC2E6D26" pageId="41" pageNumber="42" value="2016-04-22">
<date id="BD0C61D0FFCBFF94FA2DF8ECFC2E6D26" pageId="41" pageNumber="42" value="2016-04-22">22 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFCBFF94FC6FF8CDFBD56D26" box="[964,1091,1888,1910]" pageId="41" pageNumber="42">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCBFF97FBE5F8CDFDA16F53" ID-GBIF-Occurrence="2235579120" collectingDate="2016-04-07" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" lastPageId="42" lastPageNumber="43" latitude="9.1721" location="Pyramid" longLatPrecision="7" longitude="123.2519" municipality="Dauin" pageId="41" pageNumber="42" specimenCode="CASIZ 217308" specimenCount="1" stateProvince="Negros Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCBFF94FBE5F8CDFB6D6D25" box="[1102,1275,1888,1909]" collectionCode="CASIZ" pageId="41" pageNumber="42">CASIZ 217308</specimenCode>
,
<location id="CC6D11CBFFCBFF94FAADF8CDFAFD6D26" LSID="urn:lsid:plazi:treatment:411BF606FFCBFF96FC97FD48FA846EC3:CC6D11CBFFCBFF94FAADF8CDFAFD6D26" box="[1286,1387,1888,1910]" country="Philippines" latitude="9.1721" longLatPrecision="7" longitude="123.2519" municipality="Dauin" name="Pyramid" pageId="41" pageNumber="42" stateProvince="Negros Oriental">Pyramid</location>
dive site,
<geoCoordinate id="AC8621D7FFC8FF97FF60FB63FEAD6EB3" box="[203,315,1230,1251]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="9.1721">9.1721°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFC8FF97FEECFB63FE466EB4" box="[327,464,1230,1252]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="123.2519">123.2519°E</geoCoordinate>
<collectingMunicipality id="2969DD6AFFC8FF97FE73FB63FDB26EB4" box="[472,548,1230,1252]" pageId="42" pageNumber="43">Dauin</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFC8FF97FD9AFB62FD626EB4" box="[561,756,1230,1252]" country="Philippines" name="Negros Oriental" pageId="42" pageNumber="43">Negros Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFC8FF97FF3AFB40FE836F53" box="[145,277,1261,1283]" name="Philippines" pageId="42" pageNumber="43">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFC8FF97FE8BFB40FE396F53" box="[288,431,1261,1283]" pageId="42" pageNumber="43" value="2016-04-07">
<date id="BD0C61D0FFC8FF97FE8BFB40FE396F53" box="[288,431,1261,1283]" pageId="42" pageNumber="43" value="2016-04-07">7 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFC8FF97FE11FB40FDA16F53" box="[442,567,1261,1283]" pageId="42" pageNumber="43">T. Gosliner</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="9DCD1798FFC8FF97FF3AFBFFFECC6EF2" httpUri="https://zenodo.org/record/2632828/files/figure.png" pageId="42" pageNumber="43" startId="42.[145,225,1106,1128]" targetBox="[209,1361,197,1066]" targetPageId="42">
<paragraph id="C90D4710FFC8FF97FF3AFBFFFECC6EF2" blockId="42.[145,1425,1106,1186]" pageId="42" pageNumber="43">
<emphasis id="FBC69B02FFC8FF97FF3AFBFFFE9F6E37" bold="true" box="[145,265,1106,1128]" pageId="42" pageNumber="43">Figure 24.</emphasis>
<taxonomicName id="0EB23C93FFC8FF97FEBFFBFFFD236E38" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[276,693,1106,1128]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFC8FF97FEBFFBFFFE4C6E37" bold="true" box="[276,474,1106,1128]" italics="true" pageId="42" pageNumber="43">Hypselodoris roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC8FF97FD17FBFFFC836E37" box="[700,789,1106,1127]" pageId="42" pageNumber="43" rank="species">
<emphasis id="FBC69B02FFC8FF97FD17FBFFFC836E37" bold="true" box="[700,789,1106,1127]" pageId="42" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature. A, jaw rodlets, holotype, NMP 0 41282. B, inner lateral teeth, paratype, CASIZ 189173. C, middle lateral teeth, paratype, CASIZ 189173. D, outer lateral teeth holotype, NMP 0 41282.
</paragraph>
</caption>
<paragraph id="C90D4710FFC8FF97FF3AFA87FEBA6F12" blockId="42.[145,684,1322,1376]" box="[145,300,1322,1346]" pageId="42" pageNumber="43">
<heading id="9245F07CFFC8FF97FF3AFA87FEBA6F12" box="[145,300,1322,1346]" fontSize="10" level="4" pageId="42" pageNumber="43" reason="1">
<emphasis id="FBC69B02FFC8FF97FF3AFA87FEBA6F12" box="[145,300,1322,1346]" italics="true" pageId="42" pageNumber="43">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC8FF97FF3AFAE7FD3A6F30" blockId="42.[145,684,1322,1376]" box="[145,684,1354,1376]" pageId="42" pageNumber="43">
Mainit Bubbles, Mabini,
<collectingRegion id="0B7689F2FFC8FF97FE00FAE6FD8E6F30" box="[427,536,1355,1376]" country="Philippines" name="Batangas" pageId="42" pageNumber="43">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFC8FF97FD88FAE7FD3E6F30" box="[547,680,1354,1376]" name="Philippines" pageId="42" pageNumber="43">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC8FF97FF3AFA2AFE666F8C" pageId="42" pageNumber="43" type="distribution">
<paragraph id="C90D4710FFC8FF97FF3AFA2AFE586FCF" blockId="42.[145,761,1415,1500]" box="[145,462,1415,1439]" pageId="42" pageNumber="43">
<heading id="9245F07CFFC8FF97FF3AFA2AFE586FCF" box="[145,462,1415,1439]" fontSize="10" level="4" pageId="42" pageNumber="43" reason="1">
<emphasis id="FBC69B02FFC8FF97FF3AFA2AFE586FCF" box="[145,462,1415,1439]" italics="true" pageId="42" pageNumber="43">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC8FF97FF3AFA05FE666F8C" blockId="42.[145,761,1415,1500]" pageId="42" pageNumber="43">
Known from
<collectingCountry id="B1A50780FFC8FF97FE8CFA0AFE0F6FED" box="[295,409,1447,1469]" name="Indonesia" pageId="42" pageNumber="43">Indonesia</collectingCountry>
and the
<collectingCountry id="B1A50780FFC8FF97FDABFA0AFD126FED" box="[512,644,1447,1469]" name="Philippines" pageId="42" pageNumber="43">Philippines</collectingCountry>
(
<bibRefCitation id="AD233AE1FFC8FF97FD3FFA0AFE726F8C" author="Debelius H &amp; Kuiter R." journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="42" pageNumber="43" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC8FF97FF3AF9AEFF696CC5" pageId="42" pageNumber="43" type="etymology">
<paragraph id="C90D4710FFC8FF97FF3AF9AEFE846C4B" blockId="42.[145,761,1539,1685]" box="[145,274,1539,1563]" pageId="42" pageNumber="43">
<heading id="9245F07CFFC8FF97FF3AF9AEFE846C4B" box="[145,274,1539,1563]" fontSize="10" level="4" pageId="42" pageNumber="43" reason="1">
<emphasis id="FBC69B02FFC8FF97FF3AF9AEFE846C4B" box="[145,274,1539,1563]" italics="true" pageId="42" pageNumber="43">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC8FF97FF3AF98EFF696CC5" blockId="42.[145,761,1539,1685]" pageId="42" pageNumber="43">
<taxonomicName id="0EB23C93FFC8FF97FF3AF98EFED96C69" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,335,1571,1593]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC8FF97FF3AF98EFED96C69" box="[145,335,1571,1593]" italics="true" pageId="42" pageNumber="43">Hypselodoris roo</emphasis>
</taxonomicName>
comes from the A. A. Milne character Roo, the kangaroo whose mother is Kanga. This species is named
<taxonomicName id="0EB23C93FFC8FF97FF57F9CCFED66C26" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[252,320,1633,1654]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC8FF97FF57F9CCFED66C26" box="[252,320,1633,1654]" italics="true" pageId="42" pageNumber="43">H. roo</emphasis>
</taxonomicName>
, because it has often been mistaken for
<taxonomicName id="0EB23C93FFC8FF97FF3AF92DFF6E6CC4" authorityName="Rudman" authorityYear="1977" box="[145,248,1663,1685]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC8FF97FF3AF92DFF6E6CC4" box="[145,248,1663,1685]" italics="true" pageId="42" pageNumber="43">H. kanga</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC8FF96FF3AF96DFDCB6F5A" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="C90D4710FFC8FF97FF3AF96DFE8B6C87" blockId="42.[145,761,1728,1904]" box="[145,285,1728,1751]" pageId="42" pageNumber="43">
<heading id="9245F07CFFC8FF97FF3AF96DFE8B6C87" box="[145,285,1728,1751]" fontSize="10" level="4" pageId="42" pageNumber="43" reason="1">
<emphasis id="FBC69B02FFC8FF97FF3AF96DFE8B6C87" box="[145,285,1728,1751]" italics="true" pageId="42" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC8FF97FF3AF972FB7C6C65" blockId="42.[145,761,1728,1904]" lastBlockId="42.[809,1426,1230,1589]" pageId="42" pageNumber="43">
<emphasis id="FBC69B02FFC8FF97FF3AF972FE186CA4" box="[145,398,1759,1780]" italics="true" pageId="42" pageNumber="43">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFC8FF97FDC0F972FD2B6CA5" box="[619,701,1759,1781]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="42" pageNumber="43">Fig. 18</figureCitation>
H–J) moderately large, reaching
<quantity id="0E4AEAF5FFC8FF97FE10F953FD906D44" box="[443,518,1790,1812]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="45.0">45 mm</quantity>
in length. Body colour whitish to grey–blue. Sides of body high, with narrow mantle margin tapering posteriorly into rounded lobe. Notum ornamented with small to large yellow spots and smaller dark blue to black spots scattered over the surface.Areas of blue are present on the notum and sides of body. Additional spots of same colour found on sides of body and foot. Gill pocket slightly elevated from notum. Nine to eleven narrow, thin, unipinnate gill branches held erectly from gill pocket. Gill branches with red lines along edges of inner and outer surface. Apex of gill branch red–orange. Central portion of middle of outer face of gill branches with several opaque white spots. Base of rhinophores red, with opaque white spot on posterior face. Upper half of rhinophore club bright red. Rhinophores with 13–14 small lamellae.
</paragraph>
<paragraph id="C90D4710FFC8FF97FC82F9F3FAE76D5D" blockId="42.[809,1425,1630,1805]" pageId="42" pageNumber="43">
<emphasis id="FBC69B02FFC8FF97FC82F9F3FC506C23" box="[809,966,1630,1651]" italics="true" pageId="42" pageNumber="43">Mantle glands</emphasis>
: Subcutaneous mantle glands variable in distribution (
<figureCitation id="51895B95FFC8FF97FC17F9D0FB876CC2" box="[956,1041,1661,1683]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="42" pageNumber="43">Fig. 2M</figureCitation>
). Posterior glands always present; lateral glands always absent; anterior glands present or absent. This arrangement was based on seven specimens examined, four of which had both anterior and posterior mantle glands and three that lacked anterior glands.
</paragraph>
<paragraph id="C90D4710FFC8FF96FC82F89BFD33683B" blockId="42.[809,1425,1846,1899]" lastBlockId="43.[163,779,198,619]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="FBC69B02FFC8FF97FC82F89BFC656D1C" box="[809,1011,1846,1868]" italics="true" pageId="42" pageNumber="43">Buccal armature</emphasis>
: Muscular portion of buccal mass somewhat larger than length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFC9FF96FEA2FEA9FEF96B49" box="[265,367,260,282]" captionStart="Figure 24" captionStartId="42.[145,225,1106,1128]" captionTargetBox="[209,1361,197,1066]" captionTargetId="figure@42.[209,1361,197,1066]" captionTargetPageId="42" captionText="Figure 24. Hypselodoris roo Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, holotype, NMP 041282. B, inner lateral teeth, paratype, CASIZ 189173. C, middle lateral teeth, paratype, CASIZ 189173. D, outer lateral teeth holotype, NMP 041282." httpUri="https://zenodo.org/record/2632828/files/figure.png" pageId="43" pageNumber="44">Fig. 24A</figureCitation>
). Rodlets narrow with short base and evenly curved, with single, acutely pointed apex. Radula broad, nearly as wide as long. Radular formula of
<typeStatus id="1609F9B2FFC9FF96FF6BFECDFEB36B26" box="[192,293,352,374]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(CASIZ
<date id="BD0C61D0FFC9FF96FE27FECDFE746B26" box="[396,482,352,374]" pageId="43" pageNumber="44" value="70-10-19">191070</date>
) 71 × 79.0.79. Rachidian row of teeth absent (
<figureCitation id="51895B95FFC9FF96FE27FED3FE7C6BC4" box="[396,490,382,404]" captionStart="Figure 24" captionStartId="42.[145,225,1106,1128]" captionTargetBox="[209,1361,197,1066]" captionTargetId="figure@42.[209,1361,197,1066]" captionTargetPageId="42" captionText="Figure 24. Hypselodoris roo Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, holotype, NMP 041282. B, inner lateral teeth, paratype, CASIZ 189173. C, middle lateral teeth, paratype, CASIZ 189173. D, outer lateral teeth holotype, NMP 041282." httpUri="https://zenodo.org/record/2632828/files/figure.png" pageId="43" pageNumber="44">Fig. 24B</figureCitation>
). Innermost lateral teeth having irregular triangular denticle on inner side of bifid primary cusp. Denticles absent from outer side of tooth. Next several laterals and middle lateral teeth (
<figureCitation id="51895B95FFC9FF96FF46FE54FEDA685E" box="[237,332,505,527]" captionStart="Figure 24" captionStartId="42.[145,225,1106,1128]" captionTargetBox="[209,1361,197,1066]" captionTargetId="figure@42.[209,1361,197,1066]" captionTargetPageId="42" captionText="Figure 24. Hypselodoris roo Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, holotype, NMP 041282. B, inner lateral teeth, paratype, CASIZ 189173. C, middle lateral teeth, paratype, CASIZ 189173. D, outer lateral teeth holotype, NMP 041282." httpUri="https://zenodo.org/record/2632828/files/figure.png" pageId="43" pageNumber="44">Fig. 24C</figureCitation>
) with bifid cusp, lacking inner or outer denticles. Two outermost teeth having a narrower base and shorter tooth shape, with one or two rounded outer denticles (
<figureCitation id="51895B95FFC9FF96FEBCFDF8FEE1683A" box="[279,375,597,619]" captionStart="Figure 24" captionStartId="42.[145,225,1106,1128]" captionTargetBox="[209,1361,197,1066]" captionTargetId="figure@42.[209,1361,197,1066]" captionTargetPageId="42" captionText="Figure 24. Hypselodoris roo Gosliner &amp; Johnson sp. nov., buccal armature. A, jaw rodlets, holotype, NMP 041282. B, inner lateral teeth, paratype, CASIZ 189173. C, middle lateral teeth, paratype, CASIZ 189173. D, outer lateral teeth holotype, NMP 041282." httpUri="https://zenodo.org/record/2632828/files/figure.png" pageId="43" pageNumber="44">Fig. 24D</figureCitation>
), smaller than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFC9FF96FF08FD23FDCB6F5A" blockId="43.[163,779,654,1290]" pageId="43" pageNumber="44">
<emphasis id="FBC69B02FFC9FF96FF08FD23FEE968F4" box="[163,383,654,676]" italics="true" pageId="43" pageNumber="44">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFC9FF96FD06FD23FC9C68F4" box="[685,778,654,676]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFC9FF96FE93FD00FE1D6892" box="[312,395,685,707]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="43" pageNumber="44">
Fig.
<date id="BD0C61D0FFC9FF96FECCFD00FE1D6892" box="[359,395,685,706]" pageId="43" pageNumber="44" value="2013-01">13I</date>
</figureCitation>
). Ampulla thick, short, tubular and curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Proximal prostatic portion of vas deferens relatively long, convoluted, curved and thick and narrowing slightly as it transitions into muscular ejaculatory portion. Ejaculatory portion relatively long, convoluted and narrow, entering elongate, wider penial bulb. Penial bulb adjacent to straight, wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively moderately long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum seminis curving upwards towards base of bursa. Moderately long uterine duct emerging from vagina immediately below receptaculum, entering female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC9FF96FF08FA86FA846EC3" pageId="43" pageNumber="44" type="discussion">
<paragraph id="C90D4710FFC9FF96FF08FA86FE866F13" blockId="43.[163,779,1323,1899]" box="[163,272,1323,1347]" pageId="43" pageNumber="44">
<heading id="9245F07CFFC9FF96FF08FA86FE866F13" box="[163,272,1323,1347]" fontSize="10" level="4" pageId="43" pageNumber="44" reason="1">
<emphasis id="FBC69B02FFC9FF96FF08FA86FE866F13" box="[163,272,1323,1347]" italics="true" pageId="43" pageNumber="44">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC9FF96FF08FAE1FD726C07" blockId="43.[163,779,1323,1899]" pageId="43" pageNumber="44">
<taxonomicName id="0EB23C93FFC9FF96FF08FAE1FEF36F31" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,357,1356,1377]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FF08FAE1FEF36F31" box="[163,357,1356,1377]" italics="true" pageId="43" pageNumber="44">Hypselodoris roo</emphasis>
</taxonomicName>
, together with
<taxonomicName id="0EB23C93FFC9FF96FDB2FAE1FD076F31" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[537,657,1356,1377]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FDB2FAE1FD076F31" box="[537,657,1356,1377]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
, has often been misidentified as
<taxonomicName id="0EB23C93FFC9FF96FE10FAC6FDB86F2F" authorityName="Rudman" authorityYear="1977" box="[443,558,1386,1408]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FE10FAC6FDB86F2F" box="[443,558,1386,1408]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFC9FF96FDEAFAC7FC936FD0" author="Rudman WB" box="[577,773,1386,1408]" journalOrPublisher="Sydney: Australian Museum" pageId="43" pageNumber="44" publicationUrl="http://www.seaslugforum.net/find/hypskang" refId="ref44958" refString="Rudman WB. 1999 b. Hypselodoris kanga Rudman, 1977. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypskang" title="Hypselodoris kanga Rudman, 1977" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999b</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC9FF96FF08FA24FE216FCE" author="Debelius H &amp; Kuiter R." box="[163,439,1417,1439]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="43" pageNumber="44" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
). These two species also have some external similarity to
<taxonomicName id="0EB23C93FFC9FF96FE5CFA05FD166FED" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[503,640,1448,1469]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FE5CFA05FD166FED" box="[503,640,1448,1469]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
. A detailed comparison of
<taxonomicName id="0EB23C93FFC9FF96FEE9FA6AFE246F8B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[322,434,1478,1500]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FEE9FA6AFE246F8B" box="[322,434,1478,1500]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FE4CFA6AFDDA6F8B" authorityName="Rudman" authorityYear="1977" box="[487,588,1478,1500]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FE4CFA6AFDDA6F8B" box="[487,588,1478,1500]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
is found above in the remarks for
<taxonomicName id="0EB23C93FFC9FF96FEF8FA4BFE526FAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[339,452,1509,1531]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FEF8FA4BFE526FAA" box="[339,452,1509,1531]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
. In our phylogenetic analysis,
<taxonomicName id="0EB23C93FFC9FF96FF08F9A9FF716C49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,231,1540,1561]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FF08F9A9FF716C49" box="[163,231,1540,1561]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
is sister to
<taxonomicName id="0EB23C93FFC9FF96FEC1F9A9FC9C6C49" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[362,778,1540,1562]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FEC1F9A9FE756C49" box="[362,483,1540,1561]" italics="true" pageId="43" pageNumber="44">H. zephyra</emphasis>
<bibRefCitation id="AD233AE1FFC9FF96FE42F9A9FC9C6C49" author="Gosliner T &amp; Johnson, R." box="[489,778,1540,1562]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="43" pageNumber="44" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
and both are sister to
<taxonomicName id="0EB23C93FFC9FF96FE3BF98EFD966C67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[400,512,1570,1592]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FE3BF98EFD966C67" box="[400,512,1570,1592]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
. In contrast,
<taxonomicName id="0EB23C93FFC9FF96FD24F98EFC9D6C67" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[655,779,1570,1592]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FD24F98EFC9D6C67" box="[655,779,1570,1592]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
is a well-separated clade and is sister to
<emphasis id="FBC69B02FFC9FF96FDCDF9EFFD4A6C06" box="[614,732,1601,1623]" italics="true" pageId="43" pageNumber="44">H. obscura</emphasis>
.
</paragraph>
<paragraph id="C90D4710FFC9FF96FF10F9CDFDF36D3B" blockId="43.[163,779,1323,1899]" pageId="43" pageNumber="44">
<taxonomicName id="0EB23C93FFC9FF96FF10F9CDFE296C25" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[187,447,1632,1653]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FF10F9CDFE296C25" box="[187,447,1632,1653]" italics="true" pageId="43" pageNumber="44">Hypselodoris zephyra</emphasis>
</taxonomicName>
has intersecting blue lines on the notum, whereas
<taxonomicName id="0EB23C93FFC9FF96FE6FF9D2FD996CC4" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[452,527,1663,1684]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FE6FF9D2FD996CC4" box="[452,527,1663,1684]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
has bluish pigment, but the blue never forms lines.
<taxonomicName id="0EB23C93FFC9FF96FD93F930FC9D6CE3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[568,779,1693,1715]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FD93F930FC9D6CE3" box="[568,779,1693,1715]" italics="true" pageId="43" pageNumber="44">Hypselodoris roo</emphasis>
</taxonomicName>
also has yellow spots, compared with the irregular yellow pustules found in
<taxonomicName id="0EB23C93FFC9FF96FE71F976FDC86CBF" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[474,606,1754,1776]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FE71F976FDC86CBF" box="[474,606,1754,1776]" italics="true" pageId="43" pageNumber="44">H. zephyra</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFC9FF96FDC5F977FF5E6D5F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FDC5F977FF5E6D5F" italics="true" pageId="43" pageNumber="44">Hypselodoris roo</emphasis>
</taxonomicName>
has black spots, which are absent in
<taxonomicName id="0EB23C93FFC9FF96FD2EF957FC926D5E" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[645,772,1785,1807]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FD2EF957FC926D5E" box="[645,772,1785,1807]" italics="true" pageId="43" pageNumber="44">H. zephyra</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFC9FF96FF08F8B5FE3C6D7D" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[163,426,1816,1837]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FF08F8B5FE3C6D7D" box="[163,426,1816,1837]" italics="true" pageId="43" pageNumber="44">Hypselodoris zephyra</emphasis>
</taxonomicName>
lacks the opaque white spot on the basal portion of the posterior part of the rhinophores that are present in
<taxonomicName id="0EB23C93FFC9FF96FDB3F8FBFDC86D3B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[536,606,1878,1899]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FDB3F8FBFDC86D3B" box="[536,606,1878,1899]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFC9FF96FCF8FF6BFA1A68F8" blockId="43.[827,1443,198,1171]" pageId="43" pageNumber="44">
There are also clear anatomical differences between
<taxonomicName id="0EB23C93FFC9FF96FC90FF4BFC176AAB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,897,230,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FC90FF4BFC176AAB" box="[827,897,230,251]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC9FF96FC24FF4BFB916AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[911,1031,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FC24FF4BFB916AAA" box="[911,1031,229,251]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC9FF96FBBEFF4BFBE96AAA" authorityName="Rudman" authorityYear="1977" box="[1045,1151,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FBBEFF4BFBE96AAA" box="[1045,1151,229,251]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FB11FF4BFAAA6AAA" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[1210,1340,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FB11FF4BFAAA6AAA" box="[1210,1340,229,251]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
. All four species have bluish pigment and yellow spots. In
<taxonomicName id="0EB23C93FFC9FF96FAF4FEA9FA356B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1375,1443,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FAF4FEA9FA356B49" box="[1375,1443,260,281]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FCDBFE8EFC6F6B67" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[880,1017,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FCDBFE8EFC6F6B67" box="[880,1017,290,312]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
, the markings on the gill branches are red, whereas they are deep blue in
<taxonomicName id="0EB23C93FFC9FF96FAAEFEEFFAE66B06" authorityName="Rudman" authorityYear="1977" box="[1285,1392,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FAAEFEEFFAE66B06" box="[1285,1392,321,343]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FC90FECDFC396B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,943,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FC90FECDFC396B25" box="[827,943,352,373]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFC9FF96FC77FECDFBCA6B25" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[988,1116,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FC77FECDFBCA6B25" box="[988,1116,352,373]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
, the red pigment on the gills is found on the gill rachis and pinnae, whereas in
<taxonomicName id="0EB23C93FFC9FF96FC90FE33FCE96BE3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,895,414,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FC90FE33FCE96BE3" box="[827,895,414,435]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
there is red pigment on the inner gill rachis and in two lines on the outer edge, with red and opaque white markings in between the two outer lines. The rhinophores of
<taxonomicName id="0EB23C93FFC9FF96FC44FE57FBAE685F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1007,1080,506,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FC44FE57FBAE685F" box="[1007,1080,506,527]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
have an opaque white spot on the inner side of the base that is absent in
<taxonomicName id="0EB23C93FFC9FF96FAB6FDB5FA0B687D" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[1309,1437,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FAB6FDB5FA0B687D" box="[1309,1437,536,557]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
. The sides of the body of
<taxonomicName id="0EB23C93FFC9FF96FBC0FD9AFB20681C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1131,1206,567,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FBC0FD9AFB20681C" box="[1131,1206,567,588]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
are higher than in
<taxonomicName id="0EB23C93FFC9FF96FC90FDFBFC2A683A" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[827,956,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FC90FDFBFC2A683A" box="[827,956,597,619]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
, and the mantle margin is narrower. Also,
<taxonomicName id="0EB23C93FFC9FF96FC90FDD9FC5468D9" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[827,962,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="infucata">
<emphasis id="FBC69B02FFC9FF96FC90FDD9FC5468D9" box="[827,962,628,649]" italics="true" pageId="43" pageNumber="44">H. infucata</emphasis>
</taxonomicName>
has a broad posterior end of the notum rather than a tapered posterior lobe found in
<taxonomicName id="0EB23C93FFC9FF96FAEBFD3EFA1068F8" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1344,1414,659,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FAEBFD3EFA1068F8" box="[1344,1414,659,680]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFC9FF96FCF8FD1CFAFA698B" blockId="43.[827,1443,198,1171]" pageId="43" pageNumber="44">
There are differences in the arrangement of mantle glands in the species within this clade. In
<emphasis id="FBC69B02FFC9FF96FA8CFD7DFC2A6953" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="0EB23C93FFC9FF96FA8CFD7DFAF868B5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1319,1390,720,741]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">H. roo</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FC90FD42FC2A6953" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,956,750,772]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">H. zephyra</taxonomicName>
</emphasis>
there are usually anterior and posterior mantle glands, but anterior glands may also be lacking in some specimens of
<emphasis id="FBC69B02FFC9FF96FB99FC81FAE66911" box="[1074,1392,812,833]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="0EB23C93FFC9FF96FB99FC81FBEF6911" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1074,1145,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">H. roo</taxonomicName>
.
<taxonomicName id="0EB23C93FFC9FF96FB28FC81FAE66911" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1155,1392,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">Hypselodoris confetti</taxonomicName>
</emphasis>
and
<taxonomicName id="0EB23C93FFC9FF96FC90FCE6FC676930" authorityName="Eliot" authorityYear="1904" box="[827,1009,843,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="nigrostriata">
<emphasis id="FBC69B02FFC9FF96FC90FCE6FC676930" box="[827,1009,843,864]" italics="true" pageId="43" pageNumber="44">H. nigrostriata</emphasis>
</taxonomicName>
have mantle glands all around the mantle margin, whereas
<taxonomicName id="0EB23C93FFC9FF96FBF4FCC7FB5C692E" authorityName="Rudman" authorityYear="1977" box="[1119,1226,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FBF4FCC7FB5C692E" box="[1119,1226,873,895]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
has only posterior glands. The shape of the jaw rodlets and radular teeth is also similar in
<taxonomicName id="0EB23C93FFC9FF96FBA7FC0AFBC369EC" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1036,1109,935,956]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FBA7FC0AFBC369EC" box="[1036,1109,935,956]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC9FF96FBCFFC0AFB7369EB" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1124,1253,934,956]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FBCFFC0AFB7369EB" box="[1124,1253,934,956]" italics="true" pageId="43" pageNumber="44">H. zephyra</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC9FF96FB58FC0AFAF869EB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1267,1390,934,956]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FB58FC0AFAF869EB" box="[1267,1390,934,956]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FC90FC6BFC34698A" authorityName="Rudman" authorityYear="1977" box="[827,930,965,987]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFC9FF96FC90FC6BFC34698A" box="[827,930,965,987]" italics="true" pageId="43" pageNumber="44">H. kanga</emphasis>
</taxonomicName>
, but the number of teeth varies slightly.
</paragraph>
<paragraph id="C90D4710FFC9FF96FCF8FC49FA846EC3" blockId="43.[827,1443,198,1171]" pageId="43" pageNumber="44">
As noted above, in the reproductive system of
<taxonomicName id="0EB23C93FFC9FF96FC90FBAEFC296E47" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,959,1026,1048]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFC9FF96FC90FBAEFC296E47" box="[827,959,1026,1048]" italics="true" pageId="43" pageNumber="44">H. confetti</emphasis>
</taxonomicName>
the receptaculum seminis is situated immediately adjacent to the bursa copulatrix, whereas in
<taxonomicName id="0EB23C93FFC9FF96FCF3FBEDFC086E05" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[856,926,1088,1109]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFC9FF96FCF3FBEDFC086E05" box="[856,926,1088,1109]" italics="true" pageId="43" pageNumber="44">H. roo</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC9FF96FC01FBEDFBB26E05" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[938,1060,1088,1109]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFC9FF96FC01FBEDFBB26E05" box="[938,1060,1088,1109]" italics="true" pageId="43" pageNumber="44">H. zephyra</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC9FF96FBF6FBEDFA906E05" authorityName="Eliot" authorityYear="1904" box="[1117,1286,1088,1109]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="nigrostriata">
<emphasis id="FBC69B02FFC9FF96FBF6FBEDFA906E05" box="[1117,1286,1088,1109]" italics="true" pageId="43" pageNumber="44">H. nigrostriata</emphasis>
</taxonomicName>
, it is situated more proximally on the vagina (
<bibRefCitation id="AD233AE1FFC9FF96FB48FBF3FA0B6E24" author="Rudman WB" box="[1251,1437,1118,1140]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="43" pageNumber="44" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFC9FF96FC90FBD0FBCC6EC2" author="Gosliner T &amp; Johnson, R." box="[827,1114,1149,1171]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="43" pageNumber="44" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson 1999</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:number id="2605C5A7A8C940AD17572600AE58658A">1</mods:number>
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<mods:identifier id="261D9FF9C494DEFCC3F1D86A99787054" type="DOI">10.1093/zoolinnean/zly048</mods:identifier>
<mods:identifier id="86A55D8BAB93F9B8D016AA68313A17EF" type="GBIF-Dataset">49bea467-fac6-484a-9902-05483dec9a52</mods:identifier>
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<treatment id="411BF606FFCCFF8DFC67FE48FAFB6E67" ID-DOI="http://doi.org/10.5281/zenodo.5980696" ID-GBIF-Taxon="156146734" ID-Zenodo-Dep="5980696" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67" httpUri="http://treatment.plazi.org/id/411BF606FFCCFF8DFC67FE48FAFB6E67" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="81A8149BFFCCFF93FC67FE48FA85684B" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="C90D4710FFCCFF93FC67FE48FA85684B" blockId="46.[934,1299,485,578]" pageId="46" pageNumber="47">
<taxonomicName id="0EB23C93FFCCFF93FC67FE48FB3F684A" ID-CoL="3P35P" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="48" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<heading id="9245F07CFFCCFF93FC67FE48FB7B6BAB" allCaps="true" bold="true" box="[972,1261,485,508]" centered="true" fontSize="9" level="2" pageId="46" pageNumber="47" reason="4">
<emphasis id="FBC69B02FFCCFF93FC67FE48FB7B6BAB" bold="true" box="[972,1261,485,508]" italics="true" pageId="46" pageNumber="47">HYPSELODORIS SKYLERI</emphasis>
</heading>
<heading id="9245F07CFFCCFF93FC0DFDAEFB3F684A" allCaps="true" box="[934,1193,515,539]" centered="true" fontSize="9" level="3" pageId="46" pageNumber="47" reason="4">GOSLINER &amp; JOHNSON</heading>
</taxonomicName>
<emphasis id="FBC69B02FFCCFF93FB1BFDAAFA85684B" bold="true" box="[1200,1299,515,539]" pageId="46" pageNumber="47">SP. NOV.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCCFF93FC75FD87FB4D6812" box="[990,1243,554,578]" pageId="46" pageNumber="47" type="description">
<paragraph id="C90D4710FFCCFF93FC75FD87FB4D6812" blockId="46.[934,1299,485,578]" box="[990,1243,554,578]" pageId="46" pageNumber="47">
<heading id="9245F07CFFCCFF93FC75FD87FB4D6812" allCaps="true" box="[990,1243,554,578]" centered="true" fontSize="10" level="1" pageId="46" pageNumber="47" reason="1">(FIGS 2O, 13K, 27, 28)</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCCFF93FC82FDDEFBCA68F7" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="C90D4710FFCCFF93FC82FDDEFBCA68F7" blockId="46.[809,1426,626,679]" pageId="46" pageNumber="47">
<emphasis id="FBC69B02FFCCFF93FC82FDDEFCFB68D8" box="[809,877,627,648]" italics="true" pageId="46" pageNumber="47">LSID</emphasis>
:
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urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFCCFF93FB1BFDDFFBCA68F7" pageId="46" pageNumber="47">8944E0DB-1780- 41E4-B926-82328F762D68</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCCFF93FC82FD68FC646906" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="C90D4710FFCCFF93FC82FD68FB7B688B" blockId="46.[809,1425,709,854]" box="[809,1261,709,731]" pageId="46" pageNumber="47">
<taxonomicName id="0EB23C93FFCCFF93FC82FD68FB70688A" ID-CoL="3P34H" authority="? Rudman, 2003" authorityName="? Rudman" authorityYear="2003" box="[809,1254,709,731]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFCCFF93FC82FD68FBAF688A" box="[809,1081,709,730]" italics="true" pageId="46" pageNumber="47">Hypselodoris maculosa?</emphasis>
<bibRefCitation id="AD233AE1FFCCFF93FBEBFD68FB70688A" author="Rudman WB" box="[1088,1254,709,731]" journalOrPublisher="Australian Museum, Sydney" pageId="46" pageNumber="47" publicationUrl="http://www.seaslugforum.net/find/11083" refId="ref45035" refString="Rudman WB. 2003. Comment on Hypselodoris maculosa? from Philippines by Yukari Sato. In: Sea slug forum. Australian Museum, Sydney. Available at: http: // www. seaslugforum. net / find / 11083" title="Comment on Hypselodoris maculosa? from Philippines by Yukari Sato" type="url" volumeTitle="Sea slug forum" year="2003">Rudman, 2003</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFCCFF93FC82FD49FC256948" blockId="46.[809,1425,709,854]" pageId="46" pageNumber="47">
<taxonomicName id="0EB23C93FFCCFF93FC82FD49FC5068A9" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,966,740,761]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFCCFF93FC82FD49FC5068A9" box="[809,966,740,761]" italics="true" pageId="46" pageNumber="47">Hypselodoris</emphasis>
</taxonomicName>
sp. 5
<bibRefCitation id="AD233AE1FFCCFF93FBBAFD49FA9F68A9" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1041,1289,740,762]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="46" pageNumber="47" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFCCFF93FB2AFD49FB2968A9" box="[1153,1215,740,761]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 2008
</bibRefCitation>
: 265, third photograph.
</paragraph>
<paragraph id="C90D4710FFCCFF93FC82FC8CFC646906" blockId="46.[809,1425,709,854]" pageId="46" pageNumber="47">
<taxonomicName id="0EB23C93FFCCFF93FC82FC8CFC296966" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,959,801,822]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFCCFF93FC82FC8CFC296966" box="[809,959,801,822]" italics="true" pageId="46" pageNumber="47">Hypselodoris</emphasis>
</taxonomicName>
sp. 10
<bibRefCitation id="AD233AE1FFCCFF93FBB9FC8CFB6A6967" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1042,1276,801,823]" journalOrPublisher="Jacksonville: New World Press" pageId="46" pageNumber="47" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFCCFF93FBD7FC8FFB206966" box="[1148,1206,801,823]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 2015
</bibRefCitation>
: 259, middle right photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCCFF92FC82FCDEFEBD6CBD" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="C90D4710FFCCFF93FC82FCDEFC4569DA" blockId="46.[809,1424,882,1028]" box="[809,979,882,906]" pageId="46" pageNumber="47">
<heading id="9245F07CFFCCFF93FC82FCDEFC4569DA" box="[809,979,882,906]" fontSize="10" level="4" pageId="46" pageNumber="47" reason="1">
<emphasis id="FBC69B02FFCCFF93FC82FCDEFC4569DA" box="[809,979,882,906]" italics="true" pageId="46" pageNumber="47">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCCFF93FC82FC3FFABE6E54" blockId="46.[809,1424,882,1028]" pageId="46" pageNumber="47">
<materialsCitation id="79DA4D4DFFCCFF93FC82FC3FFABE6E54" ID-GBIF-Occurrence="2235579131" collectingDate="2010-05-18" collectionCode="NMP" collectorName="T. Gosliner." country="Philippines" county="Luzon" latitude="13.68736" location="Sea Pen" longLatPrecision="1" longitude="120.83283" municipality="Tingloy" pageId="46" pageNumber="47" specimenCode="CASIZ 182780" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
<typeStatus id="1609F9B2FFCCFF93FC82FC3FFC0369F7" box="[809,917,914,935]" pageId="46" pageNumber="47" type="holotype">
<emphasis id="FBC69B02FFCCFF93FC82FC3FFC0369F7" box="[809,917,914,935]" italics="true" pageId="46" pageNumber="47">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFCCFF93FC0DFC3EFC7369F8" box="[934,997,915,936]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="46" pageNumber="47">NMP</collectionCode>
0 41284 (formerly
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), subsampled for molecular study, dissected,
<location id="CC6D11CBFFCCFF93FA9AFC1CFA066997" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCCFF93FA9AFC1CFA066997" box="[1329,1424,945,967]" country="Philippines" county="Luzon" latitude="13.68736" longLatPrecision="1" longitude="120.83283" municipality="Tingloy" name="Sea Pen" pageId="46" pageNumber="47" stateProvince="Batangas">Sea Pen</location>
dive site,
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,
<geoCoordinate id="AC8621D7FFCCFF93FB8FFC7DFB2069B5" box="[1060,1206,976,997]" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="120.83283">120.83283°E</geoCoordinate>
,
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,
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,
<collectingCounty id="206C3F9CFFCCFF93FC82FC42FCE46E54" box="[809,882,1007,1028]" pageId="46" pageNumber="47">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFCCFF93FCD7FC43FC696E54" box="[892,1023,1006,1028]" name="Philippines" pageId="46" pageNumber="47">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCCFF93FBA1FC42FB086E54" box="[1034,1182,1007,1028]" pageId="46" pageNumber="47" value="2010-05-18">
<date id="BD0C61D0FFCCFF93FBA1FC42FB086E54" box="[1034,1182,1007,1028]" pageId="46" pageNumber="47" value="2010-05-18">18 May 2010</date>
</collectingDate>
,
<collectorName id="644722C6FFCCFF93FB03FC42FABE6E54" box="[1192,1320,1006,1028]" pageId="46" pageNumber="47">T. Gosliner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFCCFF92FC82FB8BFDE06C21" blockId="46.[809,1425,1061,1114]" lastBlockId="47.[163,780,1198,1649]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<materialsCitation id="79DA4D4DFFCCFF92FC82FB8BFDCF6E94" ID-GBIF-Occurrence="2235579129" collectingDate="2008-03-17" collectionCode="CASIZ" collectorName="Tingloy &amp; T. Gosliner" country="Philippines" lastPageId="47" lastPageNumber="48" latitude="13.68806" location="Sepok Wall" longLatPrecision="1" longitude="120.82678" municipality="Luzon" pageId="46" pageNumber="47" specimenCode="CASIZ 177305" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<typeStatus id="1609F9B2FFCCFF93FC82FB8BFC056E6B" box="[809,915,1062,1083]" pageId="46" pageNumber="47" type="paratype">
<emphasis id="FBC69B02FFCCFF93FC82FB8BFC056E6B" box="[809,915,1062,1083]" italics="true" pageId="46" pageNumber="47">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFCCFF93FC35FB88FBAA6E6B" box="[926,1084,1061,1083]" collectionCode="CASIZ" pageId="46" pageNumber="47">CASIZ 177305</specimenCode>
, one specimen, dissected,
<location id="CC6D11CBFFCCFF93FAE5FB88FCCA6E0A" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCCFF93FAE5FB88FCCA6E0A" country="Philippines" latitude="13.68806" longLatPrecision="1" longitude="120.82678" municipality="Luzon" name="Sepok Wall" pageId="46" pageNumber="47" stateProvince="Batangas">Sepok Wall</location>
,
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,
<geoCoordinate id="AC8621D7FFCCFF93FC40FBE9FBE06E0A" box="[1003,1142,1092,1114]" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="120.82678">120.82678°E</geoCoordinate>
,
<collectorName id="644722C6FFCCFF93FBD5FBE9FB586E0A" box="[1150,1230,1092,1114]" pageId="46" pageNumber="47">Tingloy</collectorName>
,
<collectingRegion id="0B7689F2FFCCFF93FB7DFBE9FAA86E09" box="[1238,1342,1092,1113]" country="Philippines" name="Batangas" pageId="46" pageNumber="47">Batangas</collectingRegion>
,
<collectingMunicipality id="2969DD6AFFCCFF93FAECFBE9FA1B6E09" box="[1351,1421,1092,1113]" pageId="46" pageNumber="47">Luzon</collectingMunicipality>
,
<collectingCountry id="B1A50780FFCDFF92FF08FB03FEB56E94" box="[163,291,1198,1220]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FE86FB03FE406E93" box="[301,470,1198,1220]" pageId="47" pageNumber="48" value="2008-03-17">
<date id="BD0C61D0FFCDFF92FE86FB03FE406E93" box="[301,470,1198,1220]" pageId="47" pageNumber="48" value="2008-03-17">17 March 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE4BFB03FDCF6E94" box="[480,601,1198,1220]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FDC8FB03FE356F70" ID-GBIF-Occurrence="2235579040" collectingDate="2014-05-04" collectionCode="CASIZ" collectorName="Tingloy &amp; T. Gosliner" country="Philippines" county="Luzon" latitude="13.62978" location="Bonito" longLatPrecision="1" longitude="120.94738" municipality="Culebra" pageId="47" pageNumber="48" specimenCode="CASIZ 200649" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FDC8FB03FC916E94" box="[611,775,1198,1220]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FD1FFB03FC916E94" box="[692,775,1198,1220]" pageId="47" pageNumber="48" value="49-06-20">200649</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FEF9FB61FE366EB2" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FEF9FB61FE366EB2" box="[338,416,1228,1250]" country="Philippines" county="Luzon" latitude="13.62978" longLatPrecision="1" longitude="120.94738" municipality="Culebra" name="Bonito" pageId="47" pageNumber="48" stateProvince="Batangas">Bonito</location>
(
<collectingMunicipality id="2969DD6AFFCDFF92FE1AFB61FD876EB2" box="[433,529,1228,1250]" pageId="47" pageNumber="48">Culebra</collectingMunicipality>
) Island,
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,
<geoCoordinate id="AC8621D7FFCDFF92FF08FB46FEA26F50" box="[163,308,1259,1280]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="1" value="120.94738">120.94738°E</geoCoordinate>
,
<collectorName id="644722C6FFCDFF92FEEBFB46FE036F51" box="[320,405,1259,1281]" pageId="47" pageNumber="48">Tingloy</collectorName>
,
<collectingRegion id="0B7689F2FFCDFF92FE0BFB41FD9B6F51" box="[416,525,1260,1281]" country="Philippines" name="Batangas" pageId="47" pageNumber="48">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFCDFF92FDB1FB41FDF56F51" box="[538,611,1260,1281]" pageId="47" pageNumber="48">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFCDFF92FDC5FB46FD676F51" box="[622,753,1259,1281]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FD56FB46FE8E6F4F" pageId="47" pageNumber="48" value="2014-05-04">
<date id="BD0C61D0FFCDFF92FD56FB46FE8E6F4F" pageId="47" pageNumber="48" value="2014-05-04">4 May 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE8FFAA7FE356F70" box="[292,419,1290,1312]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FE04FAA7FC916F0D" ID-GBIF-Occurrence="2235579097" collectingDate="2014-05-29" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="13.77931" location="Binacas" longLatPrecision="1" longitude="120.10959" municipality="Lubang Island" pageId="47" pageNumber="48" specimenCode="CASIZ 200552" specimenCount="1" stateProvince="Mindoro Occidental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FE04FAA7FDCD6F70" box="[431,603,1290,1312]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FDAEFAA7FDCD6F70" box="[517,603,1290,1312]" pageId="47" pageNumber="48" value="52-05-20">200552</date>
</specimenCode>
, one specimen, off
<location id="CC6D11CBFFCDFF92FF6CFA85FEB46F6E" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FF6CFA85FEB46F6E" box="[199,290,1320,1342]" country="Philippines" latitude="13.77931" longLatPrecision="1" longitude="120.10959" municipality="Lubang Island" name="Binacas" pageId="47" pageNumber="48" stateProvince="Mindoro Occidental">Binacas</location>
,
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,
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,
<collectingMunicipality id="2969DD6AFFCDFF92FDF6FA85FC906F6E" box="[605,774,1320,1342]" pageId="47" pageNumber="48">Lubang Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFCDFF92FF08FAEAFEEE6F0D" box="[163,376,1351,1373]" country="Philippines" name="Mindoro Occidental" pageId="47" pageNumber="48">Mindoro Occidental</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FE2BFAEAFE6A6F0D" box="[384,508,1351,1373]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FDAFFAEAFD1B6F0C" box="[516,653,1351,1373]" pageId="47" pageNumber="48" value="2014-05-29">
<date id="BD0C61D0FFCDFF92FDAFFAEAFD1B6F0C" box="[516,653,1351,1373]" pageId="47" pageNumber="48" value="2014-05-29">29 May 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FD3EFAE5FC916F0D" box="[661,775,1351,1373]" pageId="47" pageNumber="48">T.Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FF08FACBFEB06FE9" ID-GBIF-Occurrence="2235579076" collectingDate="2016-04-10" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="9.0502" location="Tubod" longLatPrecision="7" longitude="123.5144" municipality="Island" pageId="47" pageNumber="48" specimenCode="CASIZ 217258" specimenCount="1" stateProvince="Siquijor" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FF08FACBFED66F2C" box="[163,320,1382,1404]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 217258</specimenCode>
, one specimen, south side
<location id="CC6D11CBFFCDFF92FDF8FACBFD0F6F2C" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FDF8FACBFD0F6F2C" box="[595,665,1382,1404]" country="Philippines" latitude="9.0502" longLatPrecision="7" longitude="123.5144" municipality="Island" name="Tubod" pageId="47" pageNumber="48" stateProvince="Siquijor">Tubod</location>
,
<geoCoordinate id="AC8621D7FFCDFF92FD09FACBFC916F2C" box="[674,775,1382,1404]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="9.0502">9.0502°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FF08FA29FEB26FCA" box="[163,292,1412,1434]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="123.5144">123.5144°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFCDFF92FE84FA29FE116FCA" box="[303,391,1412,1434]" country="Philippines" name="Siquijor" pageId="47" pageNumber="48">Siquijor</collectingRegion>
<collectingMunicipality id="2969DD6AFFCDFF92FE24FA29FE416FCA" box="[399,471,1412,1434]" pageId="47" pageNumber="48">Island</collectingMunicipality>
,
<collectingCountry id="B1A50780FFCDFF92FE49FA29FDF46FCA" box="[482,610,1412,1434]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FDC6FA28FC916FCA" box="[621,775,1412,1434]" pageId="47" pageNumber="48" value="2016-04-10">
<date id="BD0C61D0FFCDFF92FDC6FA28FC916FCA" box="[621,775,1412,1434]" pageId="47" pageNumber="48" value="2016-04-10">10 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FF08FA09FEB06FE9" box="[163,294,1443,1465]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FE98FA0EFDB96FA6" ID-GBIF-Occurrence="2235579036" collectingDate="2016-04-05" collectionCode="CASIZ" collectorName="Siaton &amp; T. Gosliner" country="Philippines" latitude="9.1374" location="Tambobo Bay" longLatPrecision="7" longitude="123.1149" pageId="47" pageNumber="48" specimenCode="CASIZ 217424" specimenCount="1" stateProvince="Negros Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FE98FA0EFE756FE8" box="[307,483,1443,1464]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 217424</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FD0BFA0EFF466F87" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FD0BFA0EFF466F87" country="Philippines" latitude="9.1374" longLatPrecision="7" longitude="123.1149" name="Tambobo Bay" pageId="47" pageNumber="48" stateProvince="Negros Oriental">Tambobo Bay</location>
,
<geoCoordinate id="AC8621D7FFCDFF92FF77FA6FFEDC6F87" box="[220,330,1474,1495]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="9.1374">9.1374°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FEFDFA6FFE486F87" box="[342,478,1474,1495]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="123.1149">123.1149°E</geoCoordinate>
,
<collectorName id="644722C6FFCDFF92FE40FA6FFDAF6F88" box="[491,569,1474,1496]" pageId="47" pageNumber="48">Siaton</collectorName>
,
<collectingRegion id="0B7689F2FFCDFF92FDEDFA6FFC916F88" box="[582,775,1474,1496]" country="Philippines" name="Negros Oriental" pageId="47" pageNumber="48">Negros Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08FA4DFE896FA6" box="[163,287,1504,1526]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FE81FA4DFE276FA6" box="[298,433,1504,1526]" pageId="47" pageNumber="48" value="2016-04-05">
<date id="BD0C61D0FFCDFF92FE81FA4DFE276FA6" box="[298,433,1504,1526]" pageId="47" pageNumber="48" value="2016-04-05">5 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE11FA4CFDB96FA6" box="[442,559,1504,1526]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FD92FA4CFDB06C64" ID-GBIF-Occurrence="2235579095" collectingDate="1983-04-21" collectionCode="CASIZ" collectorName="S. Johnson" country="Marshall Islands" location="Munjor Pinnacle" pageId="47" pageNumber="48" specimenCode="CASIZ 120921" specimenCount="1" stateProvince="Enewetak Atoll" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FD92FA4CFD4F6FA6" box="[569,729,1505,1526]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FD22FA4CFD4F6FA6" box="[649,729,1505,1526]" pageId="47" pageNumber="48" value="21-12-09">120921</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FEB1FA52FE4F6C45" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FEB1FA52FE4F6C45" box="[282,473,1535,1557]" country="Marshall Islands" name="Munjor Pinnacle" pageId="47" pageNumber="48" stateProvince="Enewetak Atoll">Munjor Pinnacle</location>
,
<collectingRegion id="0B7689F2FFCDFF92FE4EFA52FD016C45" box="[485,663,1535,1557]" inferredBySuffix="true" pageId="47" pageNumber="48">Enewetak Atoll</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FD08FA52FF616C64" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FEA9F9B3FE086C63" box="[258,414,1566,1588]" pageId="47" pageNumber="48" value="1983-04-21">
<date id="BD0C61D0FFCDFF92FEA9F9B3FE086C63" box="[258,414,1566,1588]" pageId="47" pageNumber="48" value="1983-04-21">21 April 1983</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE01F9B3FDB06C64" box="[426,550,1566,1588]" pageId="47" pageNumber="48">S. Johnson</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FD9AF9B3FDE46C21" ID-GBIF-Occurrence="2235579050" collectingDate="2013-07-22" collectionCode="CASIZ" collectorName="S. Johnson" country="Marshall Islands" location="Pinnacle" pageId="47" pageNumber="48" specimenCode="CASIZ 194004" specimenCount="1" stateProvince="Kwajalein" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FD9AF9B3FD416C63" box="[561,727,1566,1587]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 194004</specimenCode>
, one specimen, under rock, K9
<location id="CC6D11CBFFCDFF92FE73F991FDA96C02" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FE73F991FDA96C02" box="[472,575,1596,1618]" country="Marshall Islands" name="Pinnacle" pageId="47" pageNumber="48" stateProvince="Kwajalein">Pinnacle</location>
,
<collectingRegion id="0B7689F2FFCDFF92FDE6F991FC906C02" box="[589,774,1596,1618]" country="Marshall Islands" name="Kwajalein" pageId="47" pageNumber="48">Kwajalein Atoll</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08F9F6FECA6C21" box="[163,348,1627,1649]" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FECDF9F6FE676C20" box="[358,497,1627,1649]" pageId="47" pageNumber="48" value="2013-07-22">
<date id="BD0C61D0FFCDFF92FECDF9F6FE676C20" box="[358,497,1627,1649]" pageId="47" pageNumber="48" value="2013-07-22">22 July 2013</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE50F9F6FDE46C21" box="[507,626,1627,1649]" pageId="47" pageNumber="48">S. Johnson</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="9DCD1798FFCCFF93FF3AF8B7FDBA6D3B" httpUri="https://zenodo.org/record/2632840/files/figure.png" pageId="46" pageNumber="47" startId="46.[145,223,1818,1840]" targetBox="[145,1425,1158,1778]" targetPageId="46">
<paragraph id="C90D4710FFCCFF93FF3AF8B7FDBA6D3B" blockId="46.[145,1425,1818,1899]" pageId="46" pageNumber="47">
<emphasis id="FBC69B02FFCCFF93FF3AF8B7FE936D7F" bold="true" box="[145,261,1818,1840]" pageId="46" pageNumber="47">Figure 27.</emphasis>
<taxonomicName id="0EB23C93FFCCFF93FEBBF8B7FD5C6D60" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[272,714,1818,1840]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFCCFF93FEBBF8B7FE6F6D60" bold="true" box="[272,505,1818,1840]" italics="true" pageId="46" pageNumber="47">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCCFF93FD64F8B6FCB26D60" box="[719,804,1819,1840]" pageId="46" pageNumber="47" rank="species">
<emphasis id="FBC69B02FFCCFF93FD64F8B6FCB26D60" bold="true" box="[719,804,1819,1840]" pageId="46" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
, living animals. A, paratype, CASIZ 217258, Tubod, Siquijor, Philippines. B, paratype, CASIZ 217424, Siaton, Negros Oriental, Philippines. C, holotype, NMP 0 41284. D, paratype, CASIZ 177305, Tingloy, Batangas, Philippines.
</paragraph>
</caption>
<caption id="9DCD1798FFCDFF92FF08FBFBFCF36ED9" httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="47" pageNumber="48" startId="47.[163,241,1110,1132]" targetBox="[227,1379,195,1069]" targetPageId="47">
<paragraph id="C90D4710FFCDFF92FF08FBFBFCF36ED9" blockId="47.[163,1444,1110,1161]" pageId="47" pageNumber="48">
<emphasis id="FBC69B02FFCDFF92FF08FBFBFE8E6E3B" bold="true" box="[163,280,1110,1132]" pageId="47" pageNumber="48">Figure 28.</emphasis>
<taxonomicName id="0EB23C93FFCDFF92FE88FBFBFD756E3C" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,739,1110,1132]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFCDFF92FE88FBFBFD986E3C" bold="true" box="[291,526,1110,1132]" italics="true" pageId="47" pageNumber="48">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCDFF92FD42FBFBFCA96E3B" box="[745,831,1110,1131]" pageId="47" pageNumber="48" rank="species">
<emphasis id="FBC69B02FFCDFF92FD42FBFBFCA96E3B" bold="true" box="[745,831,1110,1131]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFCDFF92FF08F934FEAB6CE0" blockId="47.[163,779,1688,1773]" box="[163,317,1688,1712]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FF08F934FEAB6CE0" box="[163,317,1688,1712]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FF08F934FEAB6CE0" box="[163,317,1688,1712]" italics="true" pageId="47" pageNumber="48">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FF08F914FEBD6CBD" blockId="47.[163,779,1688,1773]" pageId="47" pageNumber="48">
Sea Pen, Tingloy, Maricaban Island,
<collectingRegion id="0B7689F2FFCDFF92FD27F914FC906C9E" box="[652,774,1721,1742]" country="Philippines" name="Batangas" pageId="47" pageNumber="48">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08F97AFEBE6CBD" box="[163,296,1751,1773]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF92FF08F8B9FDB66D39" pageId="47" pageNumber="48" type="distribution">
<paragraph id="C90D4710FFCDFF92FF08F8B9FE766D7C" blockId="47.[163,779,1812,1897]" box="[163,480,1812,1836]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FF08F8B9FE766D7C" box="[163,480,1812,1836]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FF08F8B9FE766D7C" box="[163,480,1812,1836]" italics="true" pageId="47" pageNumber="48">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FF08F898FDB66D39" blockId="47.[163,779,1812,1897]" pageId="47" pageNumber="48">
Known from the
<collectingCountry id="B1A50780FFCDFF92FED4F899FD846D1A" box="[383,530,1844,1866]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingCountry id="B1A50780FFCDFF92FD89F899FD096D1A" box="[546,671,1844,1866]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
and the
<collectingCountry id="B1A50780FFCDFF92FF08F8FEFEF36D39" box="[163,357,1875,1897]" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF92FC90FB00FA176F2D" pageId="47" pageNumber="48" type="etymology">
<paragraph id="C90D4710FFCDFF92FC90FB00FC2A6E95" blockId="47.[827,1443,1197,1405]" box="[827,956,1197,1221]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FC90FB00FC2A6E95" box="[827,956,1197,1221]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FC90FB00FC2A6E95" box="[827,956,1197,1221]" italics="true" pageId="47" pageNumber="48">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FC90FB63FA176F2D" blockId="47.[827,1443,1197,1405]" pageId="47" pageNumber="48">This species is named for author Rebecca Johnson’s son, Skyler (Sky) Rodgers. The beautiful bright white spots on the mantle of this species are reminiscent of bright stars in the sky on a clear night. Sky, thanks for your patience, kindness and thoughtfulness. May you always stay curious about the stars and our place in the universe.</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF8DFC90FA08FF416C8E" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="C90D4710FFCDFF92FC90FA08FC516FEC" blockId="47.[827,1444,1445,1897]" box="[827,967,1445,1468]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FC90FA08FC516FEC" box="[827,967,1445,1468]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FC90FA08FC516FEC" box="[827,967,1445,1468]" italics="true" pageId="47" pageNumber="48">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF8DFC90FA69FEF46AAB" blockId="47.[827,1444,1445,1897]" lastBlockId="48.[145,760,198,251]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="FBC69B02FFCDFF92FC90FA69FBAF6F89" box="[827,1081,1476,1497]" italics="true" pageId="47" pageNumber="48">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFCDFF92FAB9FA69FAF56F8A" box="[1298,1379,1476,1498]" captionStart="Figure 27" captionStartId="46.[145,223,1818,1840]" captionTargetBox="[145,1425,1158,1778]" captionTargetId="figure@46.[145,1425,1158,1778]" captionTargetPageId="46" captionText="Figure 27. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., living animals. A, paratype, CASIZ 217258, Tubod, Siquijor, Philippines. B, paratype, CASIZ 217424, Siaton, Negros Oriental, Philippines. C, holotype, NMP 041284. D, paratype, CASIZ 177305, Tingloy, Batangas, Philippines." httpUri="https://zenodo.org/record/2632840/files/figure.png" pageId="47" pageNumber="48">Fig. 27</figureCitation>
A–D) small, reaching
<quantity id="0E4AEAF5FFCDFF92FC5DFA4EFBD16FA9" box="[1014,1095,1507,1529]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="47" pageNumber="48" unit="mm" value="15.0">15 mm</quantity>
in length. Body long, slender, translucent pink to peach. Fine parallel or intersecting brown longitudinal lines on notum sides of body and foot. Small, scattered opaque white spot situated on small tubercles on notum sides of body and foot. Scattered blue–purple spots found around margin of notum and foot; pigment more dense in centre of spot. Seven to eight unipinnate gill branches having a translucent white base and inner margin and bright red–orange pigment on outer surface of rachis and some gill filaments. Inner surface with opaque white. Bulb of perfoliate rhinophores opaque white with three red transverse bands and bearing about seven to ten densely arranged lamellae. Base of rhinophores translucent white.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFE89FECC6BE4" blockId="48.[145,761,292,436]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFE89FEAA6B69" box="[145,316,292,313]" italics="true" pageId="48" pageNumber="49">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFD2FF8DFE12FEEFFD806B07" box="[441,534,322,344]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="48" pageNumber="49">Fig. 2O</figureCitation>
). Glands situated around entire margin of body in six specimens
<quantity id="0E4AEAF5FFD2FF8DFD35FECCFD6F6B26" box="[670,761,353,374]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="48" pageNumber="49" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
preserved length. Anterior and posterior glands larger than lateral ones.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFE70FEE36E4A" blockId="48.[145,761,477,1051]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFE70FECA6BA3" box="[145,348,477,499]" italics="true" pageId="48" pageNumber="49">Buccal armature</emphasis>
: Muscular portion of buccal mass about equal in length to oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFD2FF8DFD20FD94FD7D681E" box="[651,747,569,591]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28A</figureCitation>
). Rodlets sharply angled, acutely pointed with one to three apices and posterior lateral extensions. Radular formula of
<typeStatus id="1609F9B2FFD2FF8DFEB2FD38FE1068FA" box="[281,390,661,682]" pageId="48" pageNumber="49" type="paratype">paratype</typeStatus>
(CASIZ 177305) 53 × 38.0.38. Rachidian row of teeth absent (
<figureCitation id="51895B95FFD2FF8DFDAEFD19FDFE6899" box="[517,616,692,714]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28B</figureCitation>
). Innermost lateral teeth having two or three large, triangular denticles on inner side of bifid primary cusp, with another two or three outer denticles. Next several laterals lacking inner triangular denticle but possessing two or three denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFD2FF8DFDCFFCC1FD5F69D1" box="[612,713,876,898]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
) all lacking inner denticles but having three or four sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and somewhat shorter tooth shape, with five or six rounded to pointed outer denticles (
<figureCitation id="51895B95FFD2FF8DFEAEFBA8FEF36E4A" box="[261,357,1029,1051]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28D</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFBE9FF416C8E" blockId="48.[145,761,1092,1758]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFBE9FEE26E09" box="[145,372,1092,1113]" italics="true" pageId="48" pageNumber="49">Reproductive system</emphasis>
: Reproductive organs fully mature in one specimen examined (CASIZ 177305;
<figureCitation id="51895B95FFD2FF8DFD23FBCFFD7D6E27" box="[648,747,1122,1144]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="48" pageNumber="49">Fig. 13K</figureCitation>
). Ampulla thick, convoluted, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing slightly as it transitions into long, convoluted, muscular ejaculatory portion. Ejaculatory portion widening into much enlarged penial bulb. Penial papilla distinctly curved with broad base, devoid of penial hooks. Penial bulb adjacent to straight, slightly narrow, widening into penial sac. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina widening proximally, leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum situated near base of bursa. Moderately short, narrow uterine duct emerging from vagina close to base of bursa and female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD2FF8DFF3AF8A9FAFB6E67" pageId="48" pageNumber="49" type="discussion">
<paragraph id="C90D4710FFD2FF8DFF3AF8A9FF686D4C" blockId="48.[145,761,1796,1882]" box="[145,254,1796,1820]" pageId="48" pageNumber="49">
<heading id="9245F07CFFD2FF8DFF3AF8A9FF686D4C" box="[145,254,1796,1820]" fontSize="10" level="4" pageId="48" pageNumber="49" reason="1">
<emphasis id="FBC69B02FFD2FF8DFF3AF8A9FF686D4C" box="[145,254,1796,1820]" italics="true" pageId="48" pageNumber="49">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AF888FA1C68D9" blockId="48.[145,761,1796,1882]" lastBlockId="48.[809,1426,198,1079]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFF3AF888FE1D6D6A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,395,1829,1850]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFF3AF888FE1D6D6A" box="[145,395,1829,1850]" italics="true" pageId="48" pageNumber="49">Hypselodoris skyleri</emphasis>
</taxonomicName>
is a member of a large clade that includes
<taxonomicName id="0EB23C93FFD2FF8DFE9EF8E9FE406D09" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[309,470,1860,1881]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFD2FF8DFE9EF8E9FE406D09" box="[309,470,1860,1881]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD2FF8DFE74F8E9FDAE6D09" box="[479,568,1860,1881]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="0EB23C93FFD2FF8DFDEFF8E9FD466D09" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[580,720,1860,1881]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFD2FF8DFDEFF8E9FD466D09" box="[580,720,1860,1881]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD2FF8DFD72F8E9FCCC6A8C" box="[729,858,199,1881]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="FBC69B02FFD2FF8DFCCDFF6AFC6F6A8C" box="[870,1017,199,220]" italics="true" pageId="48" pageNumber="49">H. maritima</emphasis>
,
<taxonomicName id="0EB23C93FFD2FF8DFBA3FF6AFB026A8B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1032,1172,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="rudmani">
<emphasis id="FBC69B02FFD2FF8DFBA3FF6AFB026A8B" box="[1032,1172,198,220]" italics="true" pageId="48" pageNumber="49">H. rudmani</emphasis>
</taxonomicName>
and
<emphasis id="FBC69B02FFD2FF8DFB7BFF6AFACF6A8B" box="[1232,1369,198,220]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFB7BFF6AFAC36A8B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1232,1365,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="bertschi">H. bertschi</taxonomicName>
.
</emphasis>
It is sister to
<taxonomicName id="0EB23C93FFD2FF8DFC38FF4BFBA16AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[915,1079,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFC38FF4BFBA16AAA" box="[915,1079,229,251]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBDFFF4BFA946AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1140,1282,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBDFFF4BFA946AAA" box="[1140,1282,229,251]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. The ABGD analysis indicates that
<taxonomicName id="0EB23C93FFD2FF8DFBEFFEA9FB2E6B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1092,1208,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFBEFFEA9FB2E6B49" box="[1092,1208,260,281]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
is clearly distinct from
<taxonomicName id="0EB23C93FFD2FF8DFCC5FE8EFB886B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[878,1054,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFCC5FE8EFB886B67" box="[878,1054,290,312]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBC8FE8EFB6A6B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1123,1276,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBC8FE8EFB6A6B67" box="[1123,1276,290,312]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. A detailed comparison of these species is found in the remarks section of
<taxonomicName id="0EB23C93FFD2FF8DFC0BFECDFBD76B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[928,1089,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFC0BFECDFBD76B25" box="[928,1089,352,373]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBD7FECDFA9E6B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1148,1288,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBD7FECDFA9E6B25" box="[1148,1288,352,373]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. The colour pattern of
<taxonomicName id="0EB23C93FFD2FF8DFC08FED2FB856BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[931,1043,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC08FED2FB856BC3" box="[931,1043,382,404]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0EB23C93FFD2FF8DFAAAFED2FA076BC3" authorityName="? Rudman" authorityYear="2003" box="[1281,1425,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFAAAFED2FA076BC3" box="[1281,1425,382,404]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFCF5FE33FC7E6BE2" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[862,1000,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCF5FE33FC7E6BE2" box="[862,1000,413,435]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
but differs in having brown rather than opaque white lines. In
<taxonomicName id="0EB23C93FFD2FF8DFB2EFE11FB6D6B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1157,1275,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB2EFE11FB6D6B81" box="[1157,1275,444,465]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
, the opaque white spots are scattered over the surface of the body and elevated on small tubercles. In
<taxonomicName id="0EB23C93FFD2FF8DFB67FE57FACB685E" authorityName="? Rudman" authorityYear="2003" box="[1228,1373,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB67FE57FACB685E" box="[1228,1373,505,527]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC82FDB5FC3C687D" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,938,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FDB5FC3C687D" box="[809,938,536,557]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
, the spots are found only on the margins of the body and are flush with the body surface. In both
<taxonomicName id="0EB23C93FFD2FF8DFC82FDFBFC03683A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,917,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC82FDFBFC03683A" box="[809,917,597,619]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC65FDFBFBC6683A" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[974,1104,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC65FDFBFBC6683A" box="[974,1104,597,619]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
there are three rhinophoral red rings, whereas there are only two in
<taxonomicName id="0EB23C93FFD2FF8DFB5DFDD9FA1568D9" authorityName="? Rudman" authorityYear="2003" box="[1270,1411,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB5DFDD9FA1568D9" box="[1270,1411,628,649]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFCEAFD3FFAFB6E67" blockId="48.[809,1426,198,1079]" pageId="48" pageNumber="49">
Both
<taxonomicName id="0EB23C93FFD2FF8DFCD6FD3EFC6B68F7" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[893,1021,658,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCD6FD3EFC6B68F7" box="[893,1021,658,680]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFBA0FD3FFBCD68F7" box="[1035,1115,658,680]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="48" pageNumber="49">Fig. 2D</figureCitation>
) and
<taxonomicName id="0EB23C93FFD2FF8DFB33FD3EFAB568F7" authorityName="? Rudman" authorityYear="2003" box="[1176,1315,658,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB33FD3EFAB568F7" box="[1176,1315,658,680]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFD2FF8DFA84FD3FFBDA6896" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="48" pageNumber="49" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999: fig. 29D</bibRefCitation>
) have mantle glands only at the anterior and posterior ends of the body, whereas
<taxonomicName id="0EB23C93FFD2FF8DFC82FD42FC356953" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,931,750,772]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC82FD42FC356953" box="[809,931,750,772]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
also has many lateral glands. The jaw rodlets of
<taxonomicName id="0EB23C93FFD2FF8DFC37FCA3FB9A6972" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[924,1036,781,803]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC37FCA3FB9A6972" box="[924,1036,781,803]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
have one to three cusps, whereas
<taxonomicName id="0EB23C93FFD2FF8DFC82FC81FC3D6911" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,939,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FC81FC3D6911" box="[809,939,812,833]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFC11FC81FB936911" box="[954,1029,812,834]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="48" pageNumber="49">Fig. 10</figureCitation>
) and
<taxonomicName id="0EB23C93FFD2FF8DFBEDFC81FB456911" authorityName="? Rudman" authorityYear="2003" box="[1094,1235,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFBEDFC81FB456911" box="[1094,1235,812,833]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFB4AFC81FABB6911" box="[1249,1325,812,834]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="48" pageNumber="49">Fig. 11</figureCitation>
) possess a single cusp. The radulae of
<taxonomicName id="0EB23C93FFD2FF8DFB2AFCE6FB64690F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1153,1266,842,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB2AFCE6FB64690F" box="[1153,1266,842,864]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD2FF8DFAABFCE6FA04690F" authorityName="? Rudman" authorityYear="2003" box="[1280,1426,842,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFAABFCE6FA04690F" box="[1280,1426,842,864]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFCF7FCC7FC74692E" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[860,994,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCF7FCC7FC74692E" box="[860,994,873,895]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
all have a similar formula and teeth of a similar shape. One notable difference is that the cusps and adjacent denticles on the inner and middle lateral teeth are more elongate in
<taxonomicName id="0EB23C93FFD2FF8DFB6AFC6BFAA4698A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1217,1330,965,987]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB6AFC6BFAA4698A" box="[1217,1330,965,987]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
than in
<taxonomicName id="0EB23C93FFD2FF8DFC82FC49FC2F69A9" authorityName="? Rudman" authorityYear="2003" box="[809,953,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFC82FC49FC2F69A9" box="[809,953,996,1017]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC5FFC49FBEF69A9" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1012,1145,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC5FFC49FBEF69A9" box="[1012,1145,996,1017]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFD2FF8DFB03FC49FA8E69A9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1192,1304,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB03FC49FA8E69A9" box="[1192,1304,996,1017]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
, the penis is proportionately wider than in
<taxonomicName id="0EB23C93FFD2FF8DFB69FBAEFACC6E47" authorityName="? Rudman" authorityYear="2003" box="[1218,1370,1026,1048]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB69FBAEFACC6E47" box="[1218,1370,1026,1048]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC82FB8FFC3D6E66" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,939,1057,1079]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FB8FFC3D6E66" box="[809,939,1057,1079]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
and possesses a distinct penial papilla.
</paragraph>
</subSubSection>
</treatment>
</document>

286
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<heading id="9245F07CFFD1FF8EFC7EFDE1FA9F6832" allCaps="true" bold="true" box="[981,1289,588,611]" centered="true" fontSize="9" level="2" pageId="51" pageNumber="52" reason="4">
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<heading id="9245F07CFFD1FF8EFC13FDC7FB2D68D0" allCaps="true" box="[952,1211,618,642]" centered="true" fontSize="9" level="3" pageId="51" pageNumber="52" reason="4">GOSLINER &amp; JOHNSON</heading>
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<emphasis id="FBC69B02FFD1FF8EFB69FDC0FAB368D2" bold="true" box="[1218,1317,618,642]" pageId="51" pageNumber="52">SP. NOV.</emphasis>
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<emphasis id="FBC69B02FFD1FF8EFC90FC81FBF46911" box="[827,1122,812,833]" italics="true" pageId="51" pageNumber="52">H y p s e l o d o r i s b u l l o ck i</emphasis>
, m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFD1FF8EFC90FCE6FB706931" ID-CoL="3P32T" authority="(Collingwood, 1881)" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[827,1254,842,865]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFD1FF8EFC90FCE6FC40690F" box="[827,982,842,864]" italics="true" pageId="51" pageNumber="52">H. bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFD1FF8EFC42FCE7FB4B6931" author="Collingwood C." box="[1001,1245,842,865]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="51" pageNumber="52" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFD1FF8EFB5CFCE7FA0B6930" author="Kohler E." box="[1271,1437,842,864]" journalOrPublisher="Sydney: Australian Museum" pageId="51" pageNumber="52" publicationUrl="http://www.seaslugforum.net/find/2073" refId="ref43316" refString="Kohler E. 2000. (Mar 13) Hypselodoris cf. bullocki. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 2073" title="(Mar 13) Hypselodoris cf. bullocki" type="url" volumeTitle="Sea slug forum" year="2000">Köhler, 2000</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD1FF8EFC90FCC4FC48692E" author="Sullivan K." box="[827,990,873,895]" journalOrPublisher="Sydney: Australian Museum" pageId="51" pageNumber="52" publicationUrl="http://www.seaslugforum.net/find/2274" refId="ref45218" refString="Sullivan K. 2000. Blue Hypselodoris cf. bullocki. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 2274" title="Blue Hypselodoris cf. bullocki" type="url" volumeTitle="Sea slug forum" year="2000">Sullivan, 2000</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD1FF8EFC40FCC7FBCA692F" author="Lau A." box="[1003,1116,873,895]" journalOrPublisher="Sydney: Australian Museum" pageId="51" pageNumber="52" publicationUrl="http://www.seaslugforum.net/find/15410" refId="ref43519" refString="Lau A. 2005. Hypselodoris bullocki? from Malaysia. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 15410" title="Hypselodoris bullocki? from Malaysia" type="url" volumeTitle="Sea slug forum" year="2005">Lau, 2005</bibRefCitation>
.
</paragraph>
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<caption id="9DCD1798FFD1FF8EFF08F897FCF36D3D" httpUri="https://zenodo.org/record/2632854/files/figure.png" pageId="51" pageNumber="52" startId="51.[163,242,1850,1872]" targetBox="[227,1379,932,1809]" targetPageId="51">
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<emphasis id="FBC69B02FFD1FF8EFF08F897FE8E6D1F" bold="true" box="[163,280,1850,1872]" pageId="51" pageNumber="52">Figure 31.</emphasis>
<taxonomicName id="0EB23C93FFD1FF8EFE88F894FD6E6D00" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,760,1849,1872]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea" status="sp. nov.">
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Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD1FF8EFD54F897FCC36D1F" box="[767,853,1850,1871]" pageId="51" pageNumber="52" rank="species">
<emphasis id="FBC69B02FFD1FF8EFD54F897FCC36D1F" bold="true" box="[767,853,1850,1871]" pageId="51" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, holotype NMP 0 41286. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<subSubSection id="81A8149BFFD6FF89FF3AFF6AFDCA6BB2" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="C90D4710FFD6FF89FF3AFF6AFEAD6A8E" blockId="52.[145,761,198,375]" box="[145,315,198,222]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FF3AFF6AFEAD6A8E" box="[145,315,198,222]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FF3AFF6AFEAD6A8E" box="[145,315,198,222]" italics="true" pageId="52" pageNumber="53">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFF4BFE596B27" blockId="52.[145,761,198,375]" pageId="52" pageNumber="53">
<materialsCitation id="79DA4D4DFFD6FF89FF3AFF4BFE596B27" ID-GBIF-Occurrence="2235579136" collectingDate="2010-02-24" collectionCode="NMP" collectorName="G. Williams." country="Philippines" county="Busuanga Island" elevation="10" latitude="11.99" location="Magic Reef" longLatPrecision="776" longitude="120.08" municipality="Coron" pageId="52" pageNumber="53" specimenCode="CASIZ 182307" specimenCount="1" stateProvince="Palawan" typeStatus="holotype">
<typeStatus id="1609F9B2FFD6FF89FF3AFF4BFF6B6AAB" box="[145,253,230,251]" pageId="52" pageNumber="53" type="holotype">
<emphasis id="FBC69B02FFD6FF89FF3AFF4BFF6B6AAB" box="[145,253,230,251]" italics="true" pageId="52" pageNumber="53">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFD6FF89FEA5FF4AFEDB6AAC" box="[270,333,231,252]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="52" pageNumber="53">NMP</collectionCode>
0 41286 (formerly
<specimenCode id="9914EF6BFFD6FF89FD9DFF4AFD7A6AAD" box="[566,748,231,253]" collectionCode="CASIZ" pageId="52" pageNumber="53">CASIZ 182307</specimenCode>
), subsampled for molecular study, dissected,
<location id="CC6D11CBFFD6FF89FDDDFEA8FD636B4B" LSID="urn:lsid:plazi:treatment:411BF606FFD1FF89FC7EFDE1FBF569E8:CC6D11CBFFD6FF89FDDDFEA8FD636B4B" box="[630,757,261,283]" country="Philippines" county="Busuanga Island" latitude="11.99" longLatPrecision="776" longitude="120.08" municipality="Coron" name="Magic Reef" pageId="52" pageNumber="53" stateProvince="Palawan">Magic Reef</location>
,
<geoCoordinate id="AC8621D7FFD6FF89FF3AFE89FF646B69" box="[145,242,292,313]" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="555" value="11.99">11.99°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD6FF89FF54FE89FEF96B69" box="[255,367,292,313]" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="555" value="120.08">120.08°E</geoCoordinate>
,
<elevation id="429FA023FFD6FF89FED7FE89FE2E6B69" box="[380,440,292,313]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="52" pageNumber="53" unit="m" value="10.0">
<quantity id="0E4AEAF5FFD6FF89FED7FE89FE2E6B69" box="[380,440,292,313]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="52" pageNumber="53" unit="m" value="10.0">10 m</quantity>
</elevation>
depth,
<location id="CC6D11CBFFD6FF89FDBEFE89FD626B69" LSID="urn:lsid:plazi:treatment:411BF606FFD1FF89FC7EFDE1FBF569E8:CC6D11CBFFD6FF89FDBEFE89FD626B69" box="[533,756,292,314]" country="Philippines" county="Busuanga Island" latitude="11.99" longLatPrecision="776" longitude="120.08" municipality="Coron" name="Bintuan Barangay" pageId="52" pageNumber="53" stateProvince="Palawan">Bintuan Barangay</location>
,
<collectingMunicipality id="2969DD6AFFD6FF89FF3AFEEEFF4A6B08" box="[145,220,323,344]" pageId="52" pageNumber="53">Coron</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFD6FF89FF41FEEEFE2A6B09" box="[234,444,323,345]" pageId="52" pageNumber="53">Busuanga Island</collectingCounty>
,
<collectingRegion id="0B7689F2FFD6FF89FE61FEEEFDA26B09" box="[458,564,323,345]" country="Philippines" name="Palawan" pageId="52" pageNumber="53">Palawan</collectingRegion>
,
<collectingCountry id="B1A50780FFD6FF89FDE9FEEEFD596B09" box="[578,719,323,345]" name="Philippines" pageId="52" pageNumber="53">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD6FF89FD76FEEEFEAC6B26" pageId="52" pageNumber="53" value="2010-02-24">
<date id="BD0C61D0FFD6FF89FD76FEEEFEAC6B26" pageId="52" pageNumber="53" value="2010-02-24">24 February 2010</date>
</collectingDate>
,
<collectorName id="644722C6FFD6FF89FEEFFECCFE596B27" box="[324,463,353,375]" pageId="52" pageNumber="53">G. Williams.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFE3CFDCA6BB2" blockId="52.[145,761,400,483]" pageId="52" pageNumber="53">
<materialsCitation id="79DA4D4DFFD6FF89FF3AFE3CFDCE6BB2" ID-GBIF-Occurrence="2235579122" collectingDate="1988-06-09" collectionCode="CASIZ" collectorName="R. Van Syoc" country="Philippines" elevation="12" location="Manu Manu Island" municipality="North Verde Island" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Palawan" typeStatus="paratype">
<typeStatus id="1609F9B2FFD6FF89FF3AFE3CFF6C6BF6" box="[145,250,401,422]" pageId="52" pageNumber="53" type="paratype">
<emphasis id="FBC69B02FFD6FF89FF3AFE3CFF6C6BF6" box="[145,250,401,422]" italics="true" pageId="52" pageNumber="53">Paratype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFD6FF89FEA3FE3DFEC06BF5" box="[264,342,400,421]" pageId="52" pageNumber="53">CASIZ</collectionCode>
0 71286,
<location id="CC6D11CBFFD6FF89FE68FE3DFD3D6BF6" LSID="urn:lsid:plazi:treatment:411BF606FFD1FF89FC7EFDE1FBF569E8:CC6D11CBFFD6FF89FE68FE3DFD3D6BF6" box="[451,683,400,422]" country="Philippines" municipality="North Verde Island" name="Manu Manu Island" pageId="52" pageNumber="53" stateProvince="Palawan">Manu Manu Island</location>
, north end of
<collectingMunicipality id="2969DD6AFFD6FF89FF43FE02FE4B6B95" box="[232,477,431,453]" pageId="52" pageNumber="53">North Verde Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD6FF89FE40FE02FDC16B95" box="[491,599,431,453]" country="Philippines" name="Palawan" pageId="52" pageNumber="53">Palawan</collectingRegion>
,
<collectingCountry id="B1A50780FFD6FF89FDCDFE02FD636B95" box="[614,757,431,453]" name="Philippines" pageId="52" pageNumber="53">Philippines</collectingCountry>
,
<elevation id="429FA023FFD6FF89FF3AFE60FF726BB3" box="[145,228,461,483]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="52" pageNumber="53" unit="m" value="12.0" valueMax="17.0" valueMin="7.0">
<quantity id="0E4AEAF5FFD6FF89FF3AFE60FF726BB3" box="[145,228,461,483]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="52" pageNumber="53" unit="m" value="12.0" valueMax="17.0" valueMin="7.0">7–17 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFD6FF89FE92FE60FE536BB2" box="[313,453,461,483]" pageId="52" pageNumber="53" value="1988-06-09">
<date id="BD0C61D0FFD6FF89FE92FE60FE536BB2" box="[313,453,461,483]" pageId="52" pageNumber="53" value="1988-06-09">9 June 1988</date>
</collectingDate>
,
<collectorName id="644722C6FFD6FF89FE7BFE63FDCE6BB2" box="[464,600,461,483]" pageId="52" pageNumber="53">R. Van Syoc</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FF3AFDA7FD87680F" pageId="52" pageNumber="53" type="distribution">
<paragraph id="C90D4710FFD6FF89FF3AFDA7FE586872" blockId="52.[145,761,522,607]" box="[145,462,522,546]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FF3AFDA7FE586872" box="[145,462,522,546]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FF3AFDA7FE586872" box="[145,462,522,546]" italics="true" pageId="52" pageNumber="53">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFD86FD87680F" blockId="52.[145,761,522,607]" pageId="52" pageNumber="53">
Known only from
<collectingRegion id="0B7689F2FFD6FF89FECAFD86FE516811" box="[353,455,555,577]" country="Philippines" name="Palawan" pageId="52" pageNumber="53">Palawan</collectingRegion>
,
<collectingCountry id="B1A50780FFD6FF89FE78FD86FDC16811" box="[467,599,555,577]" name="Philippines" pageId="52" pageNumber="53">Philippines</collectingCountry>
and northern Borneo,
<collectingCountry id="B1A50780FFD6FF89FF45FDE4FEC0680F" box="[238,342,585,607]" name="Malaysia" pageId="52" pageNumber="53">Malaysia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FF3AFD2BFDB268AA" pageId="52" pageNumber="53" type="etymology">
<paragraph id="C90D4710FFD6FF89FF3AFD2BFE8468CE" blockId="52.[145,760,646,762]" box="[145,274,646,670]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FF3AFD2BFE8468CE" box="[145,274,646,670]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FF3AFD2BFE8468CE" box="[145,274,646,670]" italics="true" pageId="52" pageNumber="53">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFD0AFDB268AA" blockId="52.[145,760,646,762]" pageId="52" pageNumber="53">
The name
<taxonomicName id="0EB23C93FFD6FF89FEA7FD0BFEFF68EB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[268,361,678,699]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD6FF89FEA7FD0BFEFF68EB" box="[268,361,678,699]" italics="true" pageId="52" pageNumber="53">violacea</emphasis>
</taxonomicName>
is Latin for violet, referring to the intense purple body colour that ornaments the body, gill and rhinophores of this species.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FF3AFC8CFA126973" pageId="52" pageNumber="53" type="description">
<paragraph id="C90D4710FFD6FF89FF3AFC8CFE8B6968" blockId="52.[145,762,801,1162]" box="[145,285,801,824]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FF3AFC8CFE8B6968" box="[145,285,801,824]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FF3AFC8CFE8B6968" box="[145,285,801,824]" italics="true" pageId="52" pageNumber="53">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFCECFD286EDA" blockId="52.[145,762,801,1162]" pageId="52" pageNumber="53">
<emphasis id="FBC69B02FFD6FF89FF3AFCECFE1D6906" box="[145,395,833,854]" italics="true" pageId="52" pageNumber="53">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFD6FF89FDCAFCECFD666907" box="[609,752,833,855]" captionStart="Figure 25" captionStartId="44.[145,227,1467,1489]" captionTargetBox="[305,1265,195,1426]" captionTargetId="figure@44.[305,1265,195,1426]" captionTargetPageId="44" captionText="Figure 25. Living animals. A, Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., holotype, NMP 041283, Tingloy, Batangas, Philippines. B, H. rositoi Gosliner &amp; Johnson sp. nov., specimen from Albatross Expedition, illustrated by Kumataro Ito. C, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177618, Tingloy, Batangas, Philippines. D, H. variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, Puerto Galera, Mindoro Oriental, Philippines. E, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208189, Puerto Galera, Mindoro Oriental, Philippines. F, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 212273, Mabini, Batangas, Philippines. G, Hypselodoris violacea Gosliner &amp; Johnson sp. nov., paratype, CASIZ 071286, Manu Manu Island, Palawan, Batangas, Philippines. H, H. violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, Busuanga, Palawan, Philippines." httpUri="https://zenodo.org/record/2632832/files/figure.png" pageId="52" pageNumber="53">Figs 25G, H</figureCitation>
) of moderately large size, reaching
<quantity id="0E4AEAF5FFD6FF89FD87FCCDFDE86925" box="[556,638,864,886]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="53" unit="mm" value="50.0">50 mm</quantity>
in length. Entire dorsal surface deep, dark purple with thick, solid opaque white band encircling the margin of notum. Sides of body and margin of foot same colour as notum. Ten to eleven unipinnate gill branches on notum that may be divided further basally or apically. Gill branches deep purple. Base of gill pocket well elevated, deep purple. Bulb and base of rhinophores deep purple throughout. Bulb with ~31 densely packed lamellae. Base of rhinophore sheath deep purple.
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFB0FFED46E87" blockId="52.[145,760,1186,1239]" pageId="52" pageNumber="53">
<emphasis id="FBC69B02FFD6FF89FF3AFB0FFEAB6EE7" box="[145,317,1186,1207]" italics="true" pageId="52" pageNumber="53">Mantle glands</emphasis>
: Mantle glands entirely absent from mantle margin.
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AFAADFD826D51" blockId="52.[145,761,1280,1793]" pageId="52" pageNumber="53">
<emphasis id="FBC69B02FFD6FF89FF3AFAADFEDD6F46" box="[145,331,1280,1302]" italics="true" pageId="52" pageNumber="53">Buccal armature</emphasis>
: Muscular portion of buccal mass about equal in size with oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFD6FF89FD99FAF1FD196F21" box="[562,655,1372,1394]" captionStart="Figure 31" captionStartId="51.[163,242,1850,1872]" captionTargetBox="[227,1379,932,1809]" captionTargetId="figure@51.[227,1379,932,1809]" captionTargetPageId="51" captionText="Figure 31. Hypselodoris violacea Gosliner &amp; Johnson sp. nov., buccal armature, holotype NMP 041286. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632854/files/figure.png" pageId="52" pageNumber="53">Fig. 31A</figureCitation>
). Rodlets narrow and short with long base and evenly curved, with single, acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFD6FF89FF3AFA15FF606F9E" box="[145,246,1464,1486]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
(CASIZ 182307) 66 × 103.0.103. Rachidian row of teeth absent (
<figureCitation id="51895B95FFD6FF89FED1FA7AFE4F6FBC" box="[378,473,1495,1517]" captionStart="Figure 31" captionStartId="51.[163,242,1850,1872]" captionTargetBox="[227,1379,932,1809]" captionTargetId="figure@51.[227,1379,932,1809]" captionTargetPageId="51" captionText="Figure 31. Hypselodoris violacea Gosliner &amp; Johnson sp. nov., buccal armature, holotype NMP 041286. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632854/files/figure.png" pageId="52" pageNumber="53">Fig. 31B</figureCitation>
). Innermost lateral teeth having one to three short denticles on inner of bifid primary cusp and one denticle on the outer side of the bifid cusp. Outer cusp of bifid cusps much shorter than inner one. Next several laterals lacking inner denticles possessing primary bifid cusps and zero or one outer denticle. Midlateral teeth (
<figureCitation id="51895B95FFD6FF89FE63F922FDBF6CF4" box="[456,553,1679,1701]" captionStart="Figure 31" captionStartId="51.[163,242,1850,1872]" captionTargetBox="[227,1379,932,1809]" captionTargetId="figure@51.[227,1379,932,1809]" captionTargetPageId="51" captionText="Figure 31. Hypselodoris violacea Gosliner &amp; Johnson sp. nov., buccal armature, holotype NMP 041286. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632854/files/figure.png" pageId="52" pageNumber="53">Fig. 31C</figureCitation>
) all lacking inner denticles but with five or six prominent outer denticles. Outermost teeth (
<figureCitation id="51895B95FFD6FF89FEFCF961FE236CB1" box="[343,437,1740,1762]" captionStart="Figure 31" captionStartId="51.[163,242,1850,1872]" captionTargetBox="[227,1379,932,1809]" captionTargetId="figure@51.[227,1379,932,1809]" captionTargetPageId="51" captionText="Figure 31. Hypselodoris violacea Gosliner &amp; Johnson sp. nov., buccal armature, holotype NMP 041286. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632854/files/figure.png" pageId="52" pageNumber="53">Fig. 31D</figureCitation>
) having a narrower base and having four or five outer denticles.
</paragraph>
<paragraph id="C90D4710FFD6FF89FF3AF884FA126973" blockId="52.[145,761,1833,1886]" lastBlockId="52.[809,1425,198,803]" pageId="52" pageNumber="53">
<emphasis id="FBC69B02FFD6FF89FF3AF884FE076D6F" box="[145,401,1833,1855]" italics="true" pageId="52" pageNumber="53">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFD6FF89FF3AF8E5FF6A6D0E" box="[145,252,1864,1886]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFD6FF89FE12F8E5FDB26D0D" box="[441,548,1864,1886]" captionStart="Figure 30" captionStartId="50.[146,229,834,856]" captionTargetBox="[306,1266,197,792]" captionTargetId="figure@50.[305,1268,196,794]" captionTargetPageId="50" captionText="Figure 30. Reproductive systems. A, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, scale bar: 1.5 mm. B, Hypseldoris violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, scale bar: 2.0 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632852/files/figure.png" pageId="52" pageNumber="53">Fig. 30B</figureCitation>
). Ampulla thick, tubular and curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens long, convoluted, curved and thick and narrowing abruptly as it transitions into muscular ejaculatory portion. Ejaculatory portion moderately short curved, narrow, entering moderately long, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled bursa copulatrix. Receptaculum seminis appressed against vagina in distal half of vaginal length. Moderately short uterine duct emerging from vagina adjacent to receptaculum and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FC82FCE8FBF569E8" pageId="52" pageNumber="53" type="discussion">
<paragraph id="C90D4710FFD6FF89FC82FCE8FC00690D" blockId="52.[809,1425,837,952]" box="[809,918,837,861]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FC82FCE8FC00690D" box="[809,918,837,861]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FC82FCE8FC00690D" box="[809,918,837,861]" italics="true" pageId="52" pageNumber="53">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82FCC8FBF569E8" blockId="52.[809,1425,837,952]" pageId="52" pageNumber="53">
The differences between
<taxonomicName id="0EB23C93FFD6FF89FBECFCCBFB50692A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1095,1222,869,891]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD6FF89FBECFCCBFB50692A" box="[1095,1222,869,891]" italics="true" pageId="52" pageNumber="53">H. violacea</emphasis>
</taxonomicName>
and other closely related species are discussed above in the remarks section of
<taxonomicName id="0EB23C93FFD6FF89FC31FC0EFBCA69E7" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[922,1116,930,952]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD6FF89FC31FC0EFBCA69E7" box="[922,1116,930,952]" italics="true" pageId="52" pageNumber="53">H. variobranchia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DC13F1FFF05EA188A1989EBFDC0C5119">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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<taxonomicName id="0EB23C93FFD2FF8DFC35FBD1FB3F6EE0" ID-CoL="3P35V" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="51" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<heading id="9245F07CFFD2FF8DFC35FBD1FA8A6EC2" allCaps="true" bold="true" box="[926,1308,1148,1171]" centered="true" fontSize="9" level="2" pageId="48" pageNumber="49" reason="4">
<emphasis id="FBC69B02FFD2FF8DFC35FBD1FA8A6EC2" bold="true" box="[926,1308,1148,1171]" italics="true" pageId="48" pageNumber="49">HYPSELODORIS VARIOBRANCHIA</emphasis>
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<heading id="9245F07CFFD2FF8DFC0DFB37FB3F6EE0" allCaps="true" box="[934,1193,1178,1202]" centered="true" fontSize="9" level="3" pageId="48" pageNumber="49" reason="4">GOSLINER &amp; JOHNSON</heading>
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<emphasis id="FBC69B02FFD2FF8DFB1BFB30FA856EE2" bold="true" box="[1200,1299,1178,1202]" pageId="48" pageNumber="49">SP. NOV.</emphasis>
</paragraph>
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<heading id="9245F07CFFD2FF8DFC75FB6CFB4D6E89" allCaps="true" box="[990,1243,1216,1241]" centered="true" fontSize="10" level="1" pageId="48" pageNumber="49" reason="1">(FIGS 25C–F, 29, 30A)</heading>
</paragraph>
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<subSubSection id="81A8149BFFD2FF8DFC82FAA4FBC06F6D" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="C90D4710FFD2FF8DFC82FAA4FBC06F6D" blockId="48.[809,1426,1289,1341]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFC82FAA4FCFA6F4E" box="[809,876,1289,1310]" italics="true" pageId="48" pageNumber="49">LSID</emphasis>
:
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urn:lsid:zoobank.org:act:
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<paragraph id="C90D4710FFD2FF8DFC82FAF1FAC76FBC" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFC82FAF1FBC66F21" box="[809,1104,1372,1393]" italics="true" pageId="48" pageNumber="49">H y p s e l o d o r i s b u l l o ck i</emphasis>
, m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFD2FF8DFC82FAD6FB2D6FC0" ID-CoL="3P32T" authority="(Collingwood, 1881)" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[809,1211,1402,1424]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFD2FF8DFC82FAD6FC2C6FDF" box="[809,954,1402,1424]" italics="true" pageId="48" pageNumber="49">H. bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFD2FF8DFC60FAD7FB276FC0" author="Collingwood C." box="[971,1201,1402,1424]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="48" pageNumber="49" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFD2FF8DFB62FAD7FA1D6FDF" author="Rudman WB" box="[1225,1419,1402,1424]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/hypsbull" refId="ref44915" refString="Rudman WB. 1999 a. Hypselodoris bullocki (Collingwood, 1881) - Page 1. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypsbull" title="Hypselodoris bullocki (Collingwood, 1881) - Page 1" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999a</bibRefCitation>
: photograph E;
<bibRefCitation id="AD233AE1FFD2FF8DFC77FA34FBFC6FFE" author="Izumi R." box="[988,1130,1433,1455]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/10353" refId="ref43007" refString="Izumi R. 2003. Hypselodoris from Anilao. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 10353" title="Hypselodoris from Anilao" type="url" volumeTitle="Sea slug forum" year="2003">Izumi, 2003</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD2FF8DFBD2FA34FA856FFE" author="Adams MJ" box="[1145,1299,1433,1455]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/11904" refId="ref41035" refString="Adams MJ. 2004. Blue Hypselodoris bullocki mating. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 11904" title="Blue Hypselodoris bullocki mating" type="url" volumeTitle="Sea slug forum" year="2004">Adams, 2004</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD2FF8DFA88FA34FCF56F9D" author="Brauchli F." journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/11986" refId="ref41511" refString="Brauchli F. 2004. Hypselodoris bullocki mating. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 11986" title="Hypselodoris bullocki mating" type="url" volumeTitle="Sea slug forum" year="2004">Brauchli, 2004</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD2FF8DFCD8FA15FC7B6F9E" author="Lau A." box="[883,1005,1464,1486]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/18974" refId="ref43556" refString="Lau A. 2006. Re: Hypselodoris bullocki? from Malaysia. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 18974" title="Re: Hypselodoris bullocki? from Malaysia" type="url" volumeTitle="Sea slug forum" year="2006">Lau, 2006</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD2FF8DFC56FA15FB0C6F9E" author="Turker K." box="[1021,1178,1464,1486]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/16653" refId="ref45319" refString="Turker K. 2006. Hypselodoris bullocki? - feeding? In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 16653" title="Hypselodoris bullocki? - feeding?" type="url" volumeTitle="Sea slug forum" year="2006">Turker, 2006</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD2FF8DFB00FA15FCF66FBB" author="Debelius H &amp; Kuiter R." journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="48" pageNumber="49" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 116: middle right photograph;
<bibRefCitation id="AD233AE1FFD2FF8DFB6EFA7BFADC6FBB" author="Tanke MA" box="[1221,1354,1494,1516]" journalOrPublisher="Sydney: Australian Museum" pageId="48" pageNumber="49" publicationUrl="http://www.seaslugforum.net/find/21737" refId="ref45281" refString="Tanke MA. 2008. Re: Hypselodoris bullocki? again. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 21737" title="Re: Hypselodoris bullocki? again" type="url" volumeTitle="Sea slug forum" year="2008">Tanke 2008</bibRefCitation>
.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82FA58FC306C79" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82FA58FC2A6C5A" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,956,1525,1546]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82FA58FC2A6C5A" box="[809,956,1525,1546]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="AD233AE1FFD2FF8DFC49FA58FB1E6C5A" author="Coleman N." box="[994,1160,1525,1547]" journalOrPublisher="Springwood: Australia Neville Coleman's Underwater Geographic" pageId="48" pageNumber="49" refId="ref41724" refString="Coleman N. 2001. 1001 Nudibranchs: catalogue of Indo- Pacific sea slugs. Springwood: Australia Neville Coleman's Underwater Geographic." title="1001 Nudibranchs: catalogue of Indo- Pacific sea slugs" type="book" year="2001">Coleman, 2001</bibRefCitation>
: 82, middle photograph, fourth row.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82F99FFC256C37" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82F99FFC5A6C17" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,972,1586,1607]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82F99FFC5A6C17" box="[809,972,1586,1607]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="AD233AE1FFD2FF8DFBAFF99FFB576C17" author="Coleman N." box="[1028,1217,1586,1608]" journalOrPublisher="Springwood: Australia Neville Coleman's Underwater Geographic" pageId="48" pageNumber="49" refId="ref41724" refString="Coleman N. 2001. 1001 Nudibranchs: catalogue of Indo- Pacific sea slugs. Springwood: Australia Neville Coleman's Underwater Geographic." title="1001 Nudibranchs: catalogue of Indo- Pacific sea slugs" type="book" year="2001">Coleman, 2001</bibRefCitation>
: 82, lower right photograph.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82F9DDFB576CF4" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82F9DDFC556CD5" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,963,1648,1669]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82F9DDFC556CD5" box="[809,963,1648,1669]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
cf.
<emphasis id="FBC69B02FFD2FF8DFC46F9DDFBDD6CD5" box="[1005,1099,1648,1669]" italics="true" pageId="48" pageNumber="49">bullocki</emphasis>
-1
<bibRefCitation id="AD233AE1FFD2FF8DFBC1F9DDFA1D6CD5" author="Debelius H &amp; Kuiter R." box="[1130,1419,1648,1670]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="48" pageNumber="49" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 116: lower two rows of photographs.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82F900FC256CB2" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82F900FC5C6C92" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,970,1709,1730]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82F900FC5C6C92" box="[809,970,1709,1730]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
sp. 6
<bibRefCitation id="AD233AE1FFD2FF8DFBB2F900FA8A6C92" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1049,1308,1709,1731]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="48" pageNumber="49" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFD2FF8DFB25F903FB586C92" box="[1166,1230,1709,1731]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2008
</bibRefCitation>
: 268, top photograph.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82F947FBAA6D4F" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82F947FC536CAF" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,965,1770,1791]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82F947FC536CAF" box="[809,965,1770,1791]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
sp. 1
<bibRefCitation id="AD233AE1FFD2FF8DFBA5F946FAC66CAF" author="Humann P &amp; DeLoach N." box="[1038,1360,1770,1792]" journalOrPublisher="Jacksonville: New World Publications" pageId="48" pageNumber="49" refId="ref42986" refString="Humann P, DeLoach N. 2010. Reef creature identification, tropical Pacific. Jacksonville: New World Publications." title="Reef creature identification, tropical Pacific" type="book" year="2010">Humann &amp; DeLoach, 2010</bibRefCitation>
: 339, upper right photograph.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFC82F885FC766D0C" blockId="48.[809,1425,1372,1884]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFC82F885FC296D6D" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,959,1832,1853]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD2FF8DFC82F885FC296D6D" box="[809,959,1832,1853]" italics="true" pageId="48" pageNumber="49">Hypselodoris</emphasis>
</taxonomicName>
sp. 15
<bibRefCitation id="AD233AE1FFD2FF8DFBB9F885FB6A6D6D" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1042,1276,1831,1854]" journalOrPublisher="Jacksonville: New World Press" pageId="48" pageNumber="49" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFD2FF8DFBD7F885FB206D6D" box="[1148,1206,1832,1853]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2015
</bibRefCitation>
: 262, middle left photograph.
</paragraph>
</subSubSection>
<caption id="9DCD1798FFD3FF8CFF09FBFEFC2E6ED7" httpUri="https://zenodo.org/record/2632848/files/figure.png" pageId="49" pageNumber="50" startId="49.[162,241,1107,1129]" targetBox="[226,1378,196,1069]" targetPageId="49">
<paragraph id="C90D4710FFD3FF8CFF09FBFEFC2E6ED7" blockId="49.[162,1443,1107,1159]" pageId="49" pageNumber="50">
<emphasis id="FBC69B02FFD3FF8CFF09FBFEFE8F6E39" bold="true" box="[162,281,1107,1129]" pageId="49" pageNumber="50">Figure 29.</emphasis>
<taxonomicName id="0EB23C93FFD3FF8CFE8FFBFEFCD46E3A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[292,834,1107,1130]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFD3FF8CFE8FFBFEFDFD6E39" bold="true" box="[292,619,1107,1129]" italics="true" pageId="49" pageNumber="50">Hypselodoris variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD3FF8CFCE2FBF9FC376E39" box="[841,929,1108,1129]" pageId="49" pageNumber="50" rank="species">
<emphasis id="FBC69B02FFD3FF8CFCE2FBF9FC376E39" bold="true" box="[841,929,1108,1129]" pageId="49" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, holotype, NMP 0 41285. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<subSubSection id="81A8149BFFD3FF8CFF08FB01FBE46FE7" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="C90D4710FFD3FF8CFF08FB01FEDA6E93" blockId="49.[163,779,1195,1372]" box="[163,332,1195,1219]" pageId="49" pageNumber="50">
<heading id="9245F07CFFD3FF8CFF08FB01FEDA6E93" box="[163,332,1195,1219]" fontSize="10" level="4" pageId="49" pageNumber="50" reason="1">
<emphasis id="FBC69B02FFD3FF8CFF08FB01FEDA6E93" box="[163,332,1195,1219]" italics="true" pageId="49" pageNumber="50">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD3FF8CFF08FB66FE716F0C" blockId="49.[163,779,1195,1372]" pageId="49" pageNumber="50">
<materialsCitation id="79DA4D4DFFD3FF8CFF08FB66FE716F0C" ID-GBIF-Occurrence="2235579031" collectingDate="2008-05-17" collectionCode="NMP" country="Philippines" county="Luzon" elevation="30" latitude="13.6586" location="Aphol's Rock" longLatPrecision="5" longitude="120.90129" municipality="Tingloy" pageId="49" pageNumber="50" specimenCode="CASIZ 177455" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
<typeStatus id="1609F9B2FFD3FF8CFF08FB66FE996EB0" box="[163,271,1227,1248]" pageId="49" pageNumber="50" type="holotype">
<emphasis id="FBC69B02FFD3FF8CFF08FB66FE996EB0" box="[163,271,1227,1248]" italics="true" pageId="49" pageNumber="50">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFD3FF8CFEB4FB61FEC86EB1" box="[287,350,1228,1249]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="49" pageNumber="50">NMP</collectionCode>
0 41285 (formerly
<specimenCode id="9914EF6BFFD3FF8CFDE3FB61FD686EB1" box="[584,766,1227,1249]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 177455</specimenCode>
), subsampled for molecular study, dissected,
<location id="CC6D11CBFFD3FF8CFD18FB47FF4B6F4F" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFD18FB47FF4B6F4F" country="Philippines" county="Luzon" latitude="13.6586" longLatPrecision="5" longitude="120.90129" municipality="Tingloy" name="Aphol's Rock" pageId="49" pageNumber="50" stateProvince="Batangas">Aphol’s Rock</location>
,
<geoCoordinate id="AC8621D7FFD3FF8CFF4DFAA4FEC86F4F" box="[230,350,1289,1311]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="5" value="13.6586">13.6586°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFEC3FAA4FE6C6F4E" box="[360,506,1289,1310]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.90129">120.90129°E</geoCoordinate>
,
<elevation id="429FA023FFD3FF8CFDAEFAA4FDAA6F4E" box="[517,572,1289,1310]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="m" value="30.0">
<quantity id="0E4AEAF5FFD3FF8CFDAEFAA4FDAA6F4E" box="[517,572,1289,1310]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="m" value="30.0">30 m</quantity>
</elevation>
depth,
<location id="CC6D11CBFFD3FF8CFD3BFAA4FF7B6F6D" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFD3BFAA4FF7B6F6D" country="Philippines" county="Luzon" latitude="13.6586" longLatPrecision="5" longitude="120.90129" municipality="Tingloy" name="Maricaban Island" pageId="49" pageNumber="50" stateProvince="Batangas">Maricaban Island</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFF5EFA8AFEDC6F6D" box="[245,330,1319,1341]" pageId="49" pageNumber="50">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFEF8FA85FDBF6F6D" box="[339,553,1319,1341]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas Province</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFD99FA85FDED6F6D" box="[562,635,1320,1341]" pageId="49" pageNumber="50">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFD2FFA8AFC916F6D" box="[644,775,1319,1341]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<date id="BD0C61D0FFD3FF8CFF08FAEBFEA16F0B" box="[163,311,1350,1372]" pageId="49" pageNumber="50" value="2008-05-17">
<collectingDate id="AD489838FFD3FF8CFF08FAEBFEA16F0B" box="[163,311,1350,1372]" pageId="49" pageNumber="50" value="2008-05-17">17 May 2008</collectingDate>
</date>
, Peri Paleracio.
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFD3FF8CFF08FA28FBE46FE7" blockId="49.[163,780,1413,1895]" lastBlockId="49.[827,1443,1195,1463]" pageId="49" pageNumber="50">
<materialsCitation id="79DA4D4DFFD3FF8CFF08FA28FE276F88" ID-GBIF-Occurrence="2235579117" collectingDate="1993-03-29" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Mindanao" location="Liuay Rock" municipality="Dakak" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Zamboanga del Norte" typeStatus="paratype">
<typeStatus id="1609F9B2FFD3FF8CFF08FA28FE826FCA" box="[163,276,1413,1434]" pageId="49" pageNumber="50" type="paratype">
<emphasis id="FBC69B02FFD3FF8CFF08FA28FE826FCA" box="[163,276,1413,1434]" italics="true" pageId="49" pageNumber="50">Paratypes</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFD3FF8CFE89FA28FEF96FCA" box="[290,367,1413,1434]" pageId="49" pageNumber="50">CASIZ</collectionCode>
0 85901, one specimen,
<location id="CC6D11CBFFD3FF8CFD2AFA28FC906FCB" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFD2AFA28FC906FCB" box="[641,774,1413,1435]" country="Philippines" county="Mindanao" municipality="Dakak" name="Liuay Rock" pageId="49" pageNumber="50" stateProvince="Zamboanga del Norte">Liuay Rock</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFF08FA0EFF786FE9" box="[163,238,1443,1465]" pageId="49" pageNumber="50">Dakak</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFF53FA0EFE726FE9" box="[248,484,1443,1465]" country="Philippines" name="Zamboanga del Norte" pageId="49" pageNumber="50">Zamboanga del Norte</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFE45FA0EFDC86FE9" box="[494,606,1443,1465]" pageId="49" pageNumber="50">Mindanao</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFDC2FA0EFD706FE9" box="[617,742,1443,1465]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFD5BFA09FEBD6F87" pageId="49" pageNumber="50" value="1993-03-29">
<date id="BD0C61D0FFD3FF8CFD5BFA09FEBD6F87" pageId="49" pageNumber="50" value="1993-03-29">29 March 1993</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFE9DFA6EFE276F88" box="[310,433,1474,1496]" pageId="49" pageNumber="50">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFE10FA6FFEAC6C63" ID-GBIF-Occurrence="2235579125" collectingDate="1994-03-14" collectionCode="CASIZ" collectorName="Mike Severns" country="Philippines" county="Luzon" latitude="13.68806" location="Sepok Point" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFD3FF8CFE10FA6FFD906F87" box="[443,518,1474,1495]" pageId="49" pageNumber="50">CASIZ</collectionCode>
0 96279, one specimen,
<location id="CC6D11CBFFD3FF8CFF08FA4CFEB56FA7" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFF08FA4CFEB56FA7" box="[163,291,1505,1527]" country="Philippines" county="Luzon" latitude="13.68806" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" name="Sepok Point" pageId="49" pageNumber="50" stateProvince="Batangas">Sepok Point</location>
,
<geoCoordinate id="AC8621D7FFD3FF8CFE80FA4CFE3D6FA7" box="[299,427,1505,1527]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="1" value="13.68806">13.68806°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFE19FA4CFDA96FA7" box="[434,575,1505,1527]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.82678">120.82678°E</geoCoordinate>
,
<location id="CC6D11CBFFD3FF8CFDEDFA4CFC906FA7" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFDEDFA4CFC906FA7" box="[582,774,1505,1527]" country="Philippines" county="Luzon" latitude="13.68806" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" name="Maricaban Island" pageId="49" pageNumber="50" stateProvince="Batangas">Maricaban Island</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFF08FA52FF616C45" box="[163,247,1535,1557]" pageId="49" pageNumber="50">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFEA9F9ADFEF86C45" box="[258,366,1536,1557]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFED1F9ADFE546C45" box="[378,450,1536,1557]" pageId="49" pageNumber="50">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFE65FA52FDC66C45" box="[462,592,1535,1557]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFDF0F9ADFC906C45" box="[603,774,1535,1557]" pageId="49" pageNumber="50" value="1994-03-14">
<date id="BD0C61D0FFD3FF8CFDF0F9ADFC906C45" box="[603,774,1535,1557]" pageId="49" pageNumber="50" value="1994-03-14">14 March 1994</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFF08F9B3FEAC6C63" box="[163,314,1566,1588]" pageId="49" pageNumber="50">Mike Severns</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFEEFF9B3FDC86C21" ID-GBIF-Occurrence="2235579093" collectingDate="2008-04-17" collectionCode="CASIZ" collectorName="Peri Paleracio" country="Philippines" elevation="30" latitude="13.6586" location="Aphol's Rock" longLatPrecision="5" longitude="120.90129" pageId="49" pageNumber="50" specimenCode="CASIZ 177618" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFEEFF9B3FE736C64" box="[324,485,1566,1588]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 177618</specimenCode>
, subsampled for molecular study, dissected,
<location id="CC6D11CBFFD3FF8CFEF7F990FE7A6C03" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFEF7F990FE7A6C03" box="[348,492,1597,1619]" country="Philippines" latitude="13.6586" longLatPrecision="5" longitude="120.90129" name="Aphol's Rock" pageId="49" pageNumber="50" stateProvince="Batangas">Aphol’s Rock</location>
,
<geoCoordinate id="AC8621D7FFD3FF8CFE5CF990FDFA6C03" box="[503,620,1597,1619]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="5" value="13.6586">13.6586°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFDDCF990FC906C02" box="[631,774,1597,1618]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.90129">120.90129°E</geoCoordinate>
,
<elevation id="429FA023FFD3FF8CFF08F9F1FF4E6C21" box="[163,216,1628,1649]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="m" value="30.0">
<quantity id="0E4AEAF5FFD3FF8CFF08F9F1FF4E6C21" box="[163,216,1628,1649]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="m" value="30.0">30 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFD3FF8CFE83F9F1FE2D6C21" box="[296,443,1627,1649]" pageId="49" pageNumber="50" value="2008-04-17">
<date id="BD0C61D0FFD3FF8CFE83F9F1FE2D6C21" box="[296,443,1627,1649]" pageId="49" pageNumber="50" value="2008-04-17">17 April 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFE6EF9F6FDC86C21" box="[453,606,1627,1649]" pageId="49" pageNumber="50">Peri Paleracio</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFDCDF9F1FDC46C9D" ID-GBIF-Occurrence="2235579064" collectingDate="2010-05-21" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Luzon" latitude="13.65083" location="Devil's Point" longLatPrecision="1" longitude="120.84127" municipality="Maricaban Island" pageId="49" pageNumber="50" specimenCode="CASIZ 182841" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFDCDF9F1FC906C21" box="[614,774,1628,1649]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 182841</specimenCode>
, one specimen,
<location id="CC6D11CBFFD3FF8CFEE3F9D7FE4C6CC0" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFEE3F9D7FE4C6CC0" box="[328,474,1658,1680]" country="Philippines" county="Luzon" latitude="13.65083" longLatPrecision="1" longitude="120.84127" municipality="Maricaban Island" name="Devil's Point" pageId="49" pageNumber="50" stateProvince="Batangas">Devil’s Point</location>
,
<geoCoordinate id="AC8621D7FFD3FF8CFE4EF9D7FDFF6CC0" box="[485,617,1658,1680]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="1" value="13.65083">13.65083°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFDDEF9D7FC906CDF" box="[629,774,1658,1679]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.84127">120.84127°E</geoCoordinate>
,
<location id="CC6D11CBFFD3FF8CFF08F934FF6E6CFF" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFF08F934FF6E6CFF" box="[163,248,1689,1711]" country="Philippines" county="Luzon" latitude="13.65083" longLatPrecision="1" longitude="120.84127" municipality="Maricaban Island" name="Tingloy" pageId="49" pageNumber="50" stateProvince="Batangas">Tingloy</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFEA8F934FE596CFF" box="[259,463,1689,1711]" pageId="49" pageNumber="50">Maricaban Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFE70F934FD246CFF" box="[475,690,1689,1711]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas Province</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFD15F934FC916CFE" box="[702,775,1689,1710]" pageId="49" pageNumber="50">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFF08F91AFEB16C9D" box="[163,295,1719,1741]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFE98F915FE5C6C9D" box="[307,458,1720,1741]" pageId="49" pageNumber="50" value="2010-05-21">
<date id="BD0C61D0FFD3FF8CFE98F915FE5C6C9D" box="[307,458,1720,1741]" pageId="49" pageNumber="50" value="2010-05-21">21 May 2010</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFE7EF915FDC46C9D" box="[469,594,1719,1741]" pageId="49" pageNumber="50">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFDF5F915FC916D79" ID-GBIF-Occurrence="2235579126" collectingDate="2015-04-26" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="13.525953" location="Puerto Galera" longLatPrecision="1" longitude="120.97016" pageId="49" pageNumber="50" specimenCode="CASIZ 208189" specimenCount="1" stateProvince="Laguna" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFDF5F915FC916C9D" box="[606,775,1720,1741]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 208189</specimenCode>
, one specimen, subsampled for molecular study,
<collectingRegion id="0B7689F2FFD3FF8CFD47F97AFF686D5A" country="Philippines" name="Laguna" pageId="49" pageNumber="50">La Laguna</collectingRegion>
,
<geoCoordinate id="AC8621D7FFD3FF8CFEA1F958FE376D5B" box="[266,417,1781,1803]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="1" value="13.525953">13.525953°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFE05F958FDC76D5B" box="[430,593,1781,1803]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.97016">120.970160°E</geoCoordinate>
,
<location id="CC6D11CBFFD3FF8CFDF5F958FC906D5B" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFDF5F958FC906D5B" box="[606,774,1781,1803]" country="Philippines" latitude="13.525953" longLatPrecision="1" longitude="120.97016" name="Puerto Galera" pageId="49" pageNumber="50" stateProvince="Laguna">Puerto Galera</location>
,
<collectingRegion id="0B7689F2FFD3FF8CFF08F8BEFEF56D79" box="[163,355,1811,1833]" country="Philippines" name="Mindoro Oriental" pageId="49" pageNumber="50">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFD3FF8CFEC6F8BEFE7C6D79" box="[365,490,1811,1833]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFE5EF8BEFD1E6D79" box="[501,648,1811,1833]" pageId="49" pageNumber="50" value="2015-04-26">
<date id="BD0C61D0FFD3FF8CFE5EF8BEFD1E6D79" box="[501,648,1811,1833]" pageId="49" pageNumber="50" value="2015-04-26">26 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFD39F8B9FC916D79" box="[658,775,1811,1833]" pageId="49" pageNumber="50">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFF08F89FFBE26E90" ID-GBIF-Occurrence="2235579032" collectingDate="2016-04-21" collectionCode="CASIZ" collectorName="Brenna Green" country="Philippines" county="Luzon" latitude="13.6305" location="Bonito Island" longLatPrecision="5" longitude="120.94763" municipality="Tingloy" pageId="49" pageNumber="50" specimenCode="CASIZ 217246" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFF08F89FFED16D18" box="[163,327,1842,1864]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 217246</specimenCode>
,
<location id="CC6D11CBFFD3FF8CFEF9F89FFE7D6D18" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFEF9F89FFE7D6D18" box="[338,491,1842,1864]" country="Philippines" county="Luzon" latitude="13.6305" longLatPrecision="5" longitude="120.94763" municipality="Tingloy" name="Bonito Island" pageId="49" pageNumber="50" stateProvince="Batangas">Bonito Island</location>
,
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,
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,
<location id="CC6D11CBFFD3FF8CFF08F8FCFEF26D37" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFF08F8FCFEF26D37" box="[163,356,1873,1895]" country="Philippines" county="Luzon" latitude="13.6305" longLatPrecision="5" longitude="120.94763" municipality="Tingloy" name="Maricaban Island" pageId="49" pageNumber="50" stateProvince="Batangas">Maricaban Island</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFEC6F8FCFE286D37" box="[365,446,1873,1895]" pageId="49" pageNumber="50">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFE63F8FCFDA76D36" box="[456,561,1873,1894]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFD91F8FCFD166D36" box="[570,640,1873,1894]" pageId="49" pageNumber="50">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFD22F8FCFC916D37" box="[649,775,1873,1895]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFC90FB06FC5B6E91" box="[827,973,1195,1217]" pageId="49" pageNumber="50" value="2016-04-21">
<date id="BD0C61D0FFD3FF8CFC90FB06FC5B6E91" box="[827,973,1195,1217]" pageId="49" pageNumber="50" value="2016-04-21">21 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFC7DFB01FBE26E90" box="[982,1140,1195,1217]" pageId="49" pageNumber="50">Brenna Green</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFBD7FB06FC126F4C" ID-GBIF-Occurrence="2235579060" collectingDate="2016-04-15" collectionCode="CASIZ" collectorName="Brenna Green" country="Philippines" county="Luzon" latitude="13.68806" location="Sepok Wall" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" pageId="49" pageNumber="50" specimenCode="CASIZ 217389" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFBD7FB06FA8D6E90" box="[1148,1307,1195,1216]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 217389</specimenCode>
,
<location id="CC6D11CBFFD3FF8CFA8FFB06FA096E91" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFA8FFB06FA096E91" box="[1316,1439,1195,1217]" country="Philippines" county="Luzon" latitude="13.68806" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" name="Sepok Wall" pageId="49" pageNumber="50" stateProvince="Batangas">Sepok Wall</location>
,
<geoCoordinate id="AC8621D7FFD3FF8CFC90FB67FC556EB0" box="[827,963,1226,1248]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="1" value="13.68806">13.68806°N</geoCoordinate>
,
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<location id="CC6D11CBFFD3FF8CFBC0FB67FAAB6EB0" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFBC0FB67FAAB6EB0" box="[1131,1341,1226,1248]" country="Philippines" county="Luzon" latitude="13.68806" longLatPrecision="1" longitude="120.82678" municipality="Tingloy" name="Maricaban Island" pageId="49" pageNumber="50" stateProvince="Batangas">Maricaban Island</location>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFAE3FB67FA096EB0" box="[1352,1439,1226,1248]" pageId="49" pageNumber="50">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFD3FF8CFC90FB44FC3C6EAE" box="[827,938,1257,1278]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFD3FF8CFC1CFB44FB976EAE" box="[951,1025,1257,1278]" pageId="49" pageNumber="50">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFD3FF8CFBA6FB44FB056EAF" box="[1037,1171,1257,1279]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFD3FF8CFB34FB45FAD66EAE" box="[1183,1344,1256,1279]" pageId="49" pageNumber="50" value="2016-04-15">
<date id="BD0C61D0FFD3FF8CFB34FB45FAD66EAE" box="[1183,1344,1256,1279]" pageId="49" pageNumber="50" value="2016-04-15">15 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFAE7FB44FC126F4C" pageId="49" pageNumber="50">Brenna Green</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFC24FAAAFADB6F0B" ID-GBIF-Occurrence="2235579055" collectingDate="2016-04-22" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Bubbles" latitude="13.688027" location="Calumpan Peninsula" longLatPrecision="1" longitude="120.89719" municipality="Luzon Island" pageId="49" pageNumber="50" specimenCode="CASIZ 217273" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFC24FAAAFBAE6F4C" box="[911,1080,1287,1308]" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 217273</specimenCode>
,
<geoCoordinate id="AC8621D7FFD3FF8CFBE8FAAAFB726F4D" box="[1091,1252,1287,1309]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="1" value="13.688027">13.6880278°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD3FF8CFB5BFAAAFA006F4C" box="[1264,1430,1287,1308]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="120.89719">120.8971833E</geoCoordinate>
°,
<location id="CC6D11CBFFD3FF8CFC90FA8BFBBC6F6C" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFC90FA8BFBBC6F6C" box="[827,1066,1318,1340]" country="Philippines" county="Bubbles" latitude="13.688027" longLatPrecision="1" longitude="120.89719" municipality="Luzon Island" name="Calumpan Peninsula" pageId="49" pageNumber="50" stateProvince="Batangas">Calumpan Peninsula</location>
,
<location id="CC6D11CBFFD3FF8CFB9FFA8BFB106F6C" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFB9FFA8BFB106F6C" box="[1076,1158,1318,1340]" country="Philippines" county="Bubbles" latitude="13.688027" longLatPrecision="1" longitude="120.89719" municipality="Luzon Island" name="Mabini" pageId="49" pageNumber="50" stateProvince="Batangas">Mabini</location>
,
<collectingRegion id="0B7689F2FFD3FF8CFB39FA8BFB686F6B" box="[1170,1278,1318,1339]" country="Philippines" name="Batangas" pageId="49" pageNumber="50">Batangas</collectingRegion>
,
<collectingMunicipality id="2969DD6AFFD3FF8CFAA2FA8BFA096F6C" box="[1289,1439,1318,1340]" pageId="49" pageNumber="50">Luzon Island</collectingMunicipality>
,
<collectingCountry id="B1A50780FFD3FF8CFC90FAE8FC2A6F0B" box="[827,956,1349,1371]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
,
<collectingCounty id="206C3F9CFFD3FF8CFC6CFAE8FBB56F0B" box="[967,1059,1349,1371]" pageId="49" pageNumber="50">Bubbles</collectingCounty>
,
<collectingDate id="AD489838FFD3FF8CFB85FAE8FB5F6F0A" box="[1070,1225,1348,1371]" pageId="49" pageNumber="50" value="2016-04-22">
<date id="BD0C61D0FFD3FF8CFB85FAE8FB5F6F0A" box="[1070,1225,1348,1371]" pageId="49" pageNumber="50" value="2016-04-22">22 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFD3FF8CFB78FAE8FADB6F0B" box="[1235,1357,1349,1371]" pageId="49" pageNumber="50">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD3FF8CFAF3FAE8FBE46FE7" ID-GBIF-Occurrence="2235579087" collectionCode="CASIZ" collectorName="R. Bolland." country="Japan" elevation="69" latitude="26.5" location="Horseshoe Cliffs" longLatPrecision="5" longitude="127.85431" pageId="49" pageNumber="50" specimenCode="CASIZ 104704" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCode id="9914EF6BFFD3FF8CFAF3FAE8FC076F28" collectionCode="CASIZ" pageId="49" pageNumber="50">CASIZ 104704</specimenCode>
, one specimen,
<elevation id="429FA023FFD3FF8CFBE2FACEFB156F29" box="[1097,1155,1379,1401]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="49" pageNumber="50" unit="m" value="69.0">
<quantity id="0E4AEAF5FFD3FF8CFBE2FACEFB156F29" box="[1097,1155,1379,1401]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="49" pageNumber="50" unit="m" value="69.0">69 m</quantity>
</elevation>
depth,
<location id="CC6D11CBFFD3FF8CFB76FACEFA096F29" LSID="urn:lsid:plazi:treatment:411BF606FFD2FF8EFC35FBD1FB68685F:CC6D11CBFFD3FF8CFB76FACEFA096F29" box="[1245,1439,1379,1401]" country="Japan" latitude="26.5" longLatPrecision="5" longitude="127.85431" name="Horseshoe Cliffs" pageId="49" pageNumber="50" stateProvince="Okinawa">Horseshoe Cliffs</location>
, (
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,
<geoCoordinate id="AC8621D7FFD3FF8CFC69FA2FFBC86FC8" box="[962,1118,1410,1432]" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="1" value="127.85431">127.854307°E</geoCoordinate>
,
<locationDeviation id="D8BF5E32FFD3FF8CFBC2FA2FFA096FC8" box="[1129,1439,1410,1432]" location="Onna Village" pageId="49" pageNumber="50">
<quantity id="0E4AEAF5FFD3FF8CFBC2FA2FFB376FC8" box="[1129,1185,1410,1432]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="km" value="1.0">1 km</quantity>
WNW of Onna Village
</locationDeviation>
,
<collectingRegion id="0B7689F2FFD3FF8CFC90FA0CFC366FE7" box="[827,928,1441,1463]" country="Japan" name="Okinawa" pageId="49" pageNumber="50">Okinawa</collectingRegion>
,
<collectingCountry id="B1A50780FFD3FF8CFC01FA0CFC786FE6" box="[938,1006,1441,1462]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
,
<collectorName id="644722C6FFD3FF8CFC52FA0CFBE46FE7" box="[1017,1138,1441,1463]" pageId="49" pageNumber="50">R. Bolland.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD3FF8CFC90FA70FB356C01" pageId="49" pageNumber="50" type="distribution">
<paragraph id="C90D4710FFD3FF8CFC90FA70FBEE6FA5" blockId="49.[827,1443,1501,1617]" box="[827,1144,1501,1525]" pageId="49" pageNumber="50">
<heading id="9245F07CFFD3FF8CFC90FA70FBEE6FA5" box="[827,1144,1501,1525]" fontSize="10" level="4" pageId="49" pageNumber="50" reason="1">
<emphasis id="FBC69B02FFD3FF8CFC90FA70FBEE6FA5" box="[827,1144,1501,1525]" italics="true" pageId="49" pageNumber="50">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD3FF8CFC90FA53FB356C01" blockId="49.[827,1443,1501,1617]" pageId="49" pageNumber="50">
Known from
<collectingRegion id="0B7689F2FFD3FF8CFC7BFA53FBCC6C44" box="[976,1114,1534,1556]" country="Australia" name="Queensland" pageId="49" pageNumber="50">Queensland</collectingRegion>
,
<collectingCountry id="B1A50780FFD3FF8CFBCEFA53FB476C44" box="[1125,1233,1534,1556]" name="Australia" pageId="49" pageNumber="50">Australia</collectingCountry>
(
<bibRefCitation id="AD233AE1FFD3FF8CFB4BFA53FA026C43" author="Rudman WB" box="[1248,1428,1534,1556]" journalOrPublisher="Sydney: Australian Museum" pageId="49" pageNumber="50" publicationUrl="http://www.seaslugforum.net/find/hypsbull" refId="ref44915" refString="Rudman WB. 1999 a. Hypselodoris bullocki (Collingwood, 1881) - Page 1. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypsbull" title="Hypselodoris bullocki (Collingwood, 1881) - Page 1" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999a</bibRefCitation>
),
<collectingCountry id="B1A50780FFD3FF8CFC90F9B0FC326C63" box="[827,932,1565,1587]" name="Malaysia" pageId="49" pageNumber="50">Malaysia</collectingCountry>
(
<bibRefCitation id="AD233AE1FFD3FF8CFC18F9B0FBB26C62" author="Lau A." box="[947,1060,1564,1586]" journalOrPublisher="Sydney: Australian Museum" pageId="49" pageNumber="50" publicationUrl="http://www.seaslugforum.net/find/18974" refId="ref43556" refString="Lau A. 2006. Re: Hypselodoris bullocki? from Malaysia. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 18974" title="Re: Hypselodoris bullocki? from Malaysia" type="url" volumeTitle="Sea slug forum" year="2006">Lau, 2006</bibRefCitation>
),
<collectingCountry id="B1A50780FFD3FF8CFB92F9B0FB3D6C63" box="[1081,1195,1565,1587]" name="Indonesia" pageId="49" pageNumber="50">Indonesia</collectingCountry>
,
<collectingRegion id="0B7689F2FFD3FF8CFB1CF9B0FAB66C63" box="[1207,1312,1565,1587]" country="Japan" name="Okinawa" pageId="49" pageNumber="50">Okinawa</collectingRegion>
,
<collectingCountry id="B1A50780FFD3FF8CFA80F9B0FAE76C62" box="[1323,1393,1565,1586]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
and the
<collectingCountry id="B1A50780FFD3FF8CFCCCF996FC7F6C01" box="[871,1001,1595,1617]" name="Philippines" pageId="49" pageNumber="50">Philippines</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD3FF8CFC90F9D5FC616CBC" pageId="49" pageNumber="50" type="etymology">
<paragraph id="C90D4710FFD3FF8CFC90F9D5FC2A6CC0" blockId="49.[827,1443,1656,1772]" box="[827,956,1656,1680]" pageId="49" pageNumber="50">
<heading id="9245F07CFFD3FF8CFC90F9D5FC2A6CC0" box="[827,956,1656,1680]" fontSize="10" level="4" pageId="49" pageNumber="50" reason="1">
<emphasis id="FBC69B02FFD3FF8CFC90F9D5FC2A6CC0" box="[827,956,1656,1680]" italics="true" pageId="49" pageNumber="50">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD3FF8CFC90F934FC616CBC" blockId="49.[827,1443,1656,1772]" pageId="49" pageNumber="50">
The name
<taxonomicName id="0EB23C93FFD3FF8CFC1AF934FBC76CFE" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[945,1105,1689,1710]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD3FF8CFC1AF934FBC76CFE" box="[945,1105,1689,1710]" italics="true" pageId="49" pageNumber="50">variobranchia</emphasis>
</taxonomicName>
comes from Latin for variable gills, referring to the gill, which may be either orange or bright purple.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD3FF8FFC90F8B9FA126C3E" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
<paragraph id="C90D4710FFD3FF8CFC90F8B9FC516D7B" blockId="49.[827,1443,1812,1896]" box="[827,967,1812,1835]" pageId="49" pageNumber="50">
<heading id="9245F07CFFD3FF8CFC90F8B9FC516D7B" box="[827,967,1812,1835]" fontSize="10" level="4" pageId="49" pageNumber="50" reason="1">
<emphasis id="FBC69B02FFD3FF8CFC90F8B9FC516D7B" box="[827,967,1812,1835]" italics="true" pageId="49" pageNumber="50">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD3FF8FFC90F89EFD6C6EB0" blockId="49.[827,1443,1812,1896]" lastBlockId="50.[145,762,980,1248]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="FBC69B02FFD3FF8CFC90F89EFBB86D18" box="[827,1070,1843,1864]" italics="true" pageId="49" pageNumber="50">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFD3FF8CFB56F89EFADA6D19" box="[1277,1356,1843,1865]" captionStart="Figure 25" captionStartId="44.[145,227,1467,1489]" captionTargetBox="[305,1265,195,1426]" captionTargetId="figure@44.[305,1265,195,1426]" captionTargetPageId="44" captionText="Figure 25. Living animals. A, Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., holotype, NMP 041283, Tingloy, Batangas, Philippines. B, H. rositoi Gosliner &amp; Johnson sp. nov., specimen from Albatross Expedition, illustrated by Kumataro Ito. C, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177618, Tingloy, Batangas, Philippines. D, H. variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, Puerto Galera, Mindoro Oriental, Philippines. E, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208189, Puerto Galera, Mindoro Oriental, Philippines. F, H. variobranchia Gosliner &amp; Johnson sp. nov., paratype, CASIZ 212273, Mabini, Batangas, Philippines. G, Hypselodoris violacea Gosliner &amp; Johnson sp. nov., paratype, CASIZ 071286, Manu Manu Island, Palawan, Batangas, Philippines. H, H. violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, Busuanga, Palawan, Philippines." httpUri="https://zenodo.org/record/2632832/files/figure.png" pageId="49" pageNumber="50">Fig. 25</figureCitation>
C–F) of moderately large size, reaching
<quantity id="0E4AEAF5FFD3FF8CFB32F8FFFB736D37" box="[1177,1253,1874,1896]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="49" pageNumber="50" unit="mm" value="50.0">50 mm</quantity>
in length. Entire dorsal surface deep purple, with thick, solid opaque white band encircling the margin of notum. Sides of body and margin of foot same colour as notum. Five to nine unipinnate gill branches on notum. Gill branches deep orange or deep purple. In specimens with orange gill branches, common base often with purple. Base of gill pocket well elevated, deep purple. Bulb of rhinophores bright orange throughout. Bulb with 19–23 densely packed lamellae. Base of rhinophore sheath deep purple.
</paragraph>
<caption id="9DCD1798FFD0FF8FFF39FCEFFAD269E0" httpUri="https://zenodo.org/record/2632852/files/figure.png" pageId="50" pageNumber="51" startId="50.[146,229,834,856]" targetBox="[306,1266,197,792]" targetPageId="50">
<paragraph id="C90D4710FFD0FF8FFF39FCEFFAD269E0" blockId="50.[146,1427,834,944]" pageId="50" pageNumber="51">
<emphasis id="FBC69B02FFD0FF8FFF39FCEFFE866907" bold="true" box="[146,272,834,856]" pageId="50" pageNumber="51">Figure 30.</emphasis>
Reproductive systems. A,
<taxonomicName id="0EB23C93FFD0FF8FFD97FCEFFBEC6908" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[572,1146,834,856]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFD0FF8FFD97FCEFFC036908" bold="true" box="[572,917,834,856]" italics="true" pageId="50" pageNumber="51">Hypselodoris variobranchia</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD0FF8FFB29FCEFFB496907" box="[1154,1247,834,855]" pageId="50" pageNumber="51" rank="species">
<emphasis id="FBC69B02FFD0FF8FFB29FCEFFB496907" bold="true" box="[1154,1247,834,855]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41285, scale bar: 1.5 mm. B,
<taxonomicName id="0EB23C93FFD0FF8FFE71FCF2FC236925" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[474,949,863,885]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="violacea" status="sp. nov.">
<emphasis id="FBC69B02FFD0FF8FFE71FCF2FD406925" bold="true" box="[474,726,863,885]" italics="true" pageId="50" pageNumber="51">Hypseldoris violacea</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD0FF8FFC16FCCDFB8E6925" box="[957,1048,864,885]" pageId="50" pageNumber="51" rank="species">
<emphasis id="FBC69B02FFD0FF8FFC16FCCDFB8E6925" bold="true" box="[957,1048,864,885]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, NMP 0 41286, scale bar: 2.0 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland.
</paragraph>
</caption>
<paragraph id="C90D4710FFD0FF8FFF3AFAA5FED46F6D" blockId="50.[145,760,1288,1341]" pageId="50" pageNumber="51">
<emphasis id="FBC69B02FFD0FF8FFF3AFAA5FEAB6F4D" box="[145,317,1288,1309]" italics="true" pageId="50" pageNumber="51">Mantle glands</emphasis>
: Mantle glands entirely absent from mantle margin.
</paragraph>
<paragraph id="C90D4710FFD0FF8FFF3AFACBFDD56D36" blockId="50.[145,761,1382,1894]" pageId="50" pageNumber="51">
<emphasis id="FBC69B02FFD0FF8FFF3AFACBFEDC6F2B" box="[145,330,1382,1403]" italics="true" pageId="50" pageNumber="51">Buccal armature</emphasis>
: Muscular portion of buccal mass much shorter than oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFD0FF8FFE48FA6FFDA86F87" box="[483,574,1474,1496]" captionStart="Figure 29" captionStartId="49.[162,241,1107,1129]" captionTargetBox="[226,1378,196,1069]" captionTargetId="figure@49.[226,1378,196,1069]" captionTargetPageId="49" captionText="Figure 29. Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041285. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632848/files/figure.png" pageId="50" pageNumber="51">Fig. 29A</figureCitation>
). Rodlets narrow and short with long base and evenly curved, with single, acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFD0FF8FFD38FA52FD6F6C45" box="[659,761,1535,1557]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
(CASIZ 177455) 71 × 98.0.98, and 68 × 106.0.106 (CASIZ 177618) in one
<typeStatus id="1609F9B2FFD0FF8FFE3BF990FE606C02" box="[400,502,1597,1618]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
. Rachidian row of teeth absent (
<figureCitation id="51895B95FFD0FF8FFF46F9F6FEDB6C20" box="[237,333,1627,1649]" captionStart="Figure 29" captionStartId="49.[162,241,1107,1129]" captionTargetBox="[226,1378,196,1069]" captionTargetId="figure@49.[226,1378,196,1069]" captionTargetPageId="49" captionText="Figure 29. Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041285. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632848/files/figure.png" pageId="50" pageNumber="51">Fig. 29B</figureCitation>
). Innermost lateral teeth having one short denticle on inner and outer sides of bifid primary cusps. Outer cusp of bifid cusps much shorter than inner one. Next several laterals lacking inner denticles possessing primary bifid cusps and two or three outer denticles. Midlateral teeth (
<figureCitation id="51895B95FFD0FF8FFE65F959FDBB6D59" box="[462,557,1780,1802]" captionStart="Figure 29" captionStartId="49.[162,241,1107,1129]" captionTargetBox="[226,1378,196,1069]" captionTargetId="figure@49.[226,1378,196,1069]" captionTargetPageId="49" captionText="Figure 29. Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041285. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632848/files/figure.png" pageId="50" pageNumber="51">Fig. 29C</figureCitation>
) all lacking inner denticles, but with five or six prominent outer denticles. Outermost teeth having a narrower base and having three or four outer denticles (
<figureCitation id="51895B95FFD0FF8FFE7CF8FDFDA36D35" box="[471,565,1872,1894]" captionStart="Figure 29" captionStartId="49.[162,241,1107,1129]" captionTargetBox="[226,1378,196,1069]" captionTargetId="figure@49.[226,1378,196,1069]" captionTargetPageId="49" captionText="Figure 29. Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041285. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632848/files/figure.png" pageId="50" pageNumber="51">Fig. 29D</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFD0FF8FFC82FC79FA126C3E" blockId="50.[809,1425,980,1646]" pageId="50" pageNumber="51">
<emphasis id="FBC69B02FFD0FF8FFC82FC79FBBF69BA" box="[809,1065,980,1002]" italics="true" pageId="50" pageNumber="51">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFD0FF8FFC82FC5EFC026E59" box="[809,916,1011,1033]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFD0FF8FFBFAFC5EFB2A6E58" box="[1105,1212,1011,1033]" captionStart="Figure 30" captionStartId="50.[146,229,834,856]" captionTargetBox="[306,1266,197,792]" captionTargetId="figure@50.[305,1268,196,794]" captionTargetPageId="50" captionText="Figure 30. Reproductive systems. A, Hypselodoris variobranchia Gosliner &amp; Johnson sp. nov., holotype, NMP 041285, scale bar: 1.5 mm. B, Hypseldoris violacea Gosliner &amp; Johnson sp. nov., holotype, NMP 041286, scale bar: 2.0 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632852/files/figure.png" pageId="50" pageNumber="51">Fig. 30A</figureCitation>
). Ampulla thick, tubular and straight, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens long, convoluted, curved and thick and narrowing abruptly as it transitions into muscular ejaculatory portion. Ejaculatory portion elongate, convoluted, narrow, entering moderately long, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled bursa copulatrix. Receptaculum seminis appressed against vagina in distal half of vaginal length. Moderately short uterine duct emerging from vagina proximally to receptaculum and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD0FF8EFC82F938FB68685F" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="discussion">
<paragraph id="C90D4710FFD0FF8FFC82F938FC006CFD" blockId="50.[809,1425,1685,1893]" box="[809,918,1685,1709]" pageId="50" pageNumber="51">
<heading id="9245F07CFFD0FF8FFC82F938FC006CFD" box="[809,918,1685,1709]" fontSize="10" level="4" pageId="50" pageNumber="51" reason="1">
<emphasis id="FBC69B02FFD0FF8FFC82F938FC006CFD" box="[809,918,1685,1709]" italics="true" pageId="50" pageNumber="51">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD0FF8EFC82F91BFE6C6BC4" blockId="50.[809,1425,1685,1893]" lastBlockId="51.[163,779,198,895]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
In our phylogenetic analyses,
<taxonomicName id="0EB23C93FFD0FF8FFBDFF91BFAA26C9B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1140,1332,1718,1739]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD0FF8FFBDFF91BFAA26C9B" box="[1140,1332,1718,1739]" italics="true" pageId="50" pageNumber="51">H. variobranchia</emphasis>
</taxonomicName>
is sister to the clade containing
<taxonomicName id="0EB23C93FFD0FF8FFBECF978FB456CB9" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[1095,1235,1748,1770]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFD0FF8FFBECF978FB456CB9" box="[1095,1235,1748,1770]" italics="true" pageId="50" pageNumber="51">H. bullockii</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD0FF8FFB49F978FA1D6CB9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1250,1419,1748,1770]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFD0FF8FFB49F978FA1D6CB9" box="[1250,1419,1748,1770]" italics="true" pageId="50" pageNumber="51">H. melanesica</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD0FF8FFC82F959FC0F6D58" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,921,1779,1801]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFD0FF8FFC82F959FC0F6D58" box="[809,921,1779,1801]" italics="true" pageId="50" pageNumber="51">H. brycei</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD0FF8FFC70F959FBE16D58" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[987,1143,1779,1801]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFD0FF8FFC70F959FBE16D58" box="[987,1143,1779,1801]" italics="true" pageId="50" pageNumber="51">H. apolegma</emphasis>
</taxonomicName>
. Externally, it is most similar to
<taxonomicName id="0EB23C93FFD0FF8FFC06F8BFFBB46D77" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[941,1058,1810,1831]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFD0FF8FFC06F8BFFBB46D77" box="[941,1058,1810,1831]" italics="true" pageId="50" pageNumber="51">H. rositoi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD0FF8FFB98F8BFFB2B6D77" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1075,1213,1810,1831]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD0FF8FFB98F8BFFB2B6D77" box="[1075,1213,1810,1831]" italics="true" pageId="50" pageNumber="51">H. violacea</emphasis>
</taxonomicName>
and some colour morphs of
<taxonomicName id="0EB23C93FFD0FF8FFC37F89CFC496D15" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[924,991,1840,1862]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD0FF8FFC37F89CFC496D15" box="[924,991,1840,1862]" italics="true" pageId="50" pageNumber="51">H. iba</emphasis>
</taxonomicName>
, which is a member of a separate clade. All of these species have a wide, solid opaque white marginal band. Externally,
<taxonomicName id="0EB23C93FFD1FF8EFE73FF6AFDD76A8C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[472,577,199,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFD1FF8EFE73FF6AFDD76A8C" box="[472,577,199,220]" italics="true" pageId="51" pageNumber="52">H. rositoi</emphasis>
</taxonomicName>
has a pink rather than purple body colour, and
<taxonomicName id="0EB23C93FFD1FF8EFDABFF4BFD126AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[512,644,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFDABFF4BFD126AAA" box="[512,644,229,251]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
has purple rhinophores in contrast to the orange rhinophores of
<taxonomicName id="0EB23C93FFD1FF8EFF08FE8EFF716B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,231,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD1FF8EFF08FE8EFF716B67" box="[163,231,290,312]" italics="true" pageId="51" pageNumber="52">H. iba</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFEB5FE8EFE486B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[286,478,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFEB5FE8EFE486B67" box="[286,478,290,312]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
. Both
<taxonomicName id="0EB23C93FFD1FF8EFD8CFE8EFDFA6B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[551,620,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD1FF8EFD8CFE8EFDFA6B67" box="[551,620,290,312]" italics="true" pageId="51" pageNumber="52">H. iba</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFD08FE8EFC9D6B68" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[675,779,291,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFD1FF8EFD08FE8EFC9D6B68" box="[675,779,291,312]" italics="true" pageId="51" pageNumber="52">H. rositoi</emphasis>
</taxonomicName>
have mantle glands that are absent in
<taxonomicName id="0EB23C93FFD1FF8EFDE0FEEFFC9C6B06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[587,778,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFDE0FEEFFC9C6B06" box="[587,778,321,343]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFF7EFECDFEC26B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[213,340,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFF7EFECDFEC26B25" box="[213,340,352,373]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
. Also,
<taxonomicName id="0EB23C93FFD1FF8EFE34FECDFE736B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[415,485,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD1FF8EFE34FECDFE736B25" box="[415,485,352,373]" italics="true" pageId="51" pageNumber="52">H. iba</emphasis>
</taxonomicName>
has a higher body profile than that of
<taxonomicName id="0EB23C93FFD1FF8EFE9AFED2FE656BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[305,499,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFE9AFED2FE656BC3" box="[305,499,382,404]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFD1FF8EFF10FE30FB68685F" blockId="51.[163,779,198,895]" lastBlockId="51.[827,1443,198,527]" pageId="51" pageNumber="52">
The radula formula of
<taxonomicName id="0EB23C93FFD1FF8EFE48FE33FDA56BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[483,563,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD1FF8EFE48FE33FDA56BE2" box="[483,563,413,435]" italics="true" pageId="51" pageNumber="52">H. iba</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD1FF8EFDEEFE33FD446BE2" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[581,722,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFDEEFE33FD446BE2" box="[581,722,413,435]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFF08FE11FEF36B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,357,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFF08FE11FEF36B81" box="[163,357,444,465]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
is similar, with almost 100 teeth per half row, whereas
<taxonomicName id="0EB23C93FFD1FF8EFED8FE76FE4A6BA0" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[371,476,475,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFD1FF8EFED8FE76FE4A6BA0" box="[371,476,475,496]" italics="true" pageId="51" pageNumber="52">H. rositoi</emphasis>
</taxonomicName>
has only 46 teeth per half row. In
<taxonomicName id="0EB23C93FFD1FF8EFF52FE57FED7685E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[249,321,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFD1FF8EFF52FE57FED7685E" box="[249,321,505,527]" italics="true" pageId="51" pageNumber="52">H. iba</emphasis>
</taxonomicName>
, only the innermost radular teeth have denticles other than the two primary cusps, whereas in
<taxonomicName id="0EB23C93FFD1FF8EFF08FD9AFEF2681B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,356,566,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFF08FD9AFEF2681B" box="[163,356,566,588]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFE36FD9AFD8C681B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[413,538,566,588]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFE36FD9AFD8C681B" box="[413,538,566,588]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
the majority of teeth have numerous outer denticles. In
<taxonomicName id="0EB23C93FFD1FF8EFD9FFDFBFD36683B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[564,672,598,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="rositoi">
<emphasis id="FBC69B02FFD1FF8EFD9FFDFBFD36683B" box="[564,672,598,619]" italics="true" pageId="51" pageNumber="52">H. rositoi</emphasis>
</taxonomicName>
, only the outer teeth have prominent outer denticles, with the exception of some of the middle lateral teeth, which may have a single denticles (
<figureCitation id="51895B95FFD1FF8EFE5DFD1CFDCE6897" box="[502,600,689,711]" captionStart="Figure 26" captionStartId="45.[163,241,1045,1067]" captionTargetBox="[227,1379,195,1004]" captionTargetId="figure@45.[227,1379,195,1004]" captionTargetPageId="45" captionText="Figure 26. Hypselodoris rositoi Gosliner &amp; Johnson sp. nov., buccal armature, holotype, NMP 041283. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632834/files/figure.png" pageId="51" pageNumber="52">Fig. 26C</figureCitation>
). The radula is very similar in
<taxonomicName id="0EB23C93FFD1FF8EFEFDFD7DFD8D68B5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[342,539,720,741]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFEFDFD7DFD8D68B5" box="[342,539,720,741]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFDFDFD7DFD4168B5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[598,727,720,741]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFDFDFD7DFD4168B5" box="[598,727,720,741]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
, but in
<taxonomicName id="0EB23C93FFD1FF8EFF6BFD42FEA86953" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[192,318,750,772]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFF6BFD42FEA86953" box="[192,318,750,772]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
the innermost teeth have a longer, more acutely pointed inner cusp of the bifid cusps. This difference in inner cusp length and sharpness is also found in the middle lateral teeth. The reproductive systems of
<taxonomicName id="0EB23C93FFD1FF8EFE80FCC7FE6C692E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[299,506,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFE80FCC7FE6C692E" box="[299,506,873,895]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFD92FCC7FD56692E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[569,704,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFD92FCC7FD56692E" box="[569,704,873,895]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
differ in a couple of key regards. In
<taxonomicName id="0EB23C93FFD1FF8EFB09FF6AFAF86A8B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1186,1390,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFB09FF6AFAF86A8B" box="[1186,1390,198,220]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
, the ejaculatory portion of the vas deferens is far more elongate than in
<taxonomicName id="0EB23C93FFD1FF8EFC54FEA9FBE86B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1023,1150,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFC54FEA9FBE86B49" box="[1023,1150,260,281]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFD1FF8EFB07FEA9FAE66B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1196,1392,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFB07FEA9FAE66B49" box="[1196,1392,260,281]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
, the uterine duct emerges from the proximal portion of the vagina, whereas in
<taxonomicName id="0EB23C93FFD1FF8EFBA4FEEFFB1C6B06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1039,1162,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFBA4FEEFFB1C6B06" box="[1039,1162,321,343]" italics="true" pageId="51" pageNumber="52">H.violacea</emphasis>
</taxonomicName>
the uterine duct emerges from the distal third of the vagina. In the ABGD analysis,
<taxonomicName id="0EB23C93FFD1FF8EFC0DFED2FBFF6BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[934,1129,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFC0DFED2FBFF6BC3" box="[934,1129,382,404]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD1FF8EFB08FED2FAB76BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1187,1313,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFB08FED2FAB76BC3" box="[1187,1313,382,404]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
are clearly indicated as distinct species. The three specimens of
<taxonomicName id="0EB23C93FFD1FF8EFC90FE11FC6A6B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1020,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="variobranchia">
<emphasis id="FBC69B02FFD1FF8EFC90FE11FC6A6B81" box="[827,1020,444,465]" italics="true" pageId="51" pageNumber="52">H. variobranchia</emphasis>
</taxonomicName>
are only 0.15–0.3% different in their
<emphasis id="FBC69B02FFD1FF8EFC90FE76FCFC6BA0" box="[827,874,475,496]" italics="true" pageId="51" pageNumber="52">COI</emphasis>
gene from each other, whereas all three specimens are 8.8–9.0% different from
<taxonomicName id="0EB23C93FFD1FF8EFBD1FE57FB61685E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1146,1271,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFD1FF8EFBD1FE57FB61685E" box="[1146,1271,505,527]" italics="true" pageId="51" pageNumber="52">H. violacea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

474
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<paragraph id="C90D4710FFD6FF89FC73FBADFA856E66" blockId="52.[934,1299,1024,1117]" pageId="52" pageNumber="53">
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<heading id="9245F07CFFD6FF89FC73FBADFB746E46" allCaps="true" bold="true" box="[984,1250,1024,1047]" centered="true" fontSize="9" level="2" pageId="52" pageNumber="53" reason="4">
<emphasis id="FBC69B02FFD6FF89FC73FBADFB746E46" bold="true" box="[984,1250,1024,1047]" italics="true" pageId="52" pageNumber="53">HYPSELODORIS YARAE</emphasis>
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<heading id="9245F07CFFD6FF89FC0DFBB3FB3F6E64" allCaps="true" box="[934,1193,1054,1078]" centered="true" fontSize="9" level="3" pageId="52" pageNumber="53" reason="4">GOSLINER &amp; JOHNSON</heading>
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<emphasis id="FBC69B02FFD6FF89FB1BFB8CFA856E66" bold="true" box="[1200,1299,1054,1078]" pageId="52" pageNumber="53">SP. NOV.</emphasis>
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</subSubSection>
<subSubSection id="81A8149BFFD6FF89FBB8FBE8FB306E0D" box="[1043,1190,1093,1117]" pageId="52" pageNumber="53" type="description">
<paragraph id="C90D4710FFD6FF89FBB8FBE8FB306E0D" blockId="52.[934,1299,1024,1117]" box="[1043,1190,1093,1117]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FBB8FBE8FB306E0D" allCaps="true" box="[1043,1190,1093,1117]" centered="true" fontSize="10" level="1" pageId="52" pageNumber="53" reason="1">
(
<figureCitation id="51895B95FFD6FF89FBB0FBE8FBE26E0D" box="[1051,1140,1093,1117]" pageId="52" pageNumber="53">FIGS 32</figureCitation>
–34)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FC82FB20FBF76E92" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="C90D4710FFD6FF89FC82FB20FBF76E92" blockId="52.[809,1426,1165,1218]" pageId="52" pageNumber="53">
<emphasis id="FBC69B02FFD6FF89FC82FB20FCFA6EF2" box="[809,876,1165,1186]" italics="true" pageId="52" pageNumber="53">LSID</emphasis>
:
<uri id="BD234B12FFD6FF89FC2BFB20FBF76E92" pageId="52" pageNumber="53">
urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFD6FF89FB02FB20FBF76E92" pageId="52" pageNumber="53">5F96599D-CBC6- 4073-BF7C-6A059710F8B8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF89FC82FB4DFC7D6C78" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="C90D4710FFD6FF89FC82FB4DFB406F01" blockId="52.[809,1425,1248,1576]" pageId="52" pageNumber="53">
<taxonomicName id="0EB23C93FFD6FF89FC82FB4DFBB96EA5" ID-CoL="3P34H" authorityName="? Rudman" authorityYear="2003" box="[809,1071,1248,1269]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD6FF89FC82FB4DFBB96EA5" box="[809,1071,1248,1269]" italics="true" pageId="52" pageNumber="53">Hypselodoris maculosa</emphasis>
</taxonomicName>
not
<taxonomicName id="0EB23C93FFD6FF89FBCAFB4DFA1B6EA6" ID-CoL="3P34H" authority="(Pease, 1871)" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1121,1421,1248,1270]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD6FF89FBCAFB4DFB796EA5" box="[1121,1263,1248,1269]" italics="true" pageId="52" pageNumber="53">H. maculosa</emphasis>
(
<bibRefCitation id="AD233AE1FFD6FF89FB56FB4DFA136EA6" author="Pease WH" box="[1277,1413,1248,1270]" journalOrPublisher="American Journal of Conchology" pageId="52" pageNumber="53" pagination="11 - 19" part="7" refId="ref44179" refString="Pease WH. 1871. Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2. American Journal of Conchology, 7: 11 - 19." title="Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2." type="journal article" year="1871">Pease, 1871</bibRefCitation>
)
</taxonomicName>
: misidentification in part,
<bibRefCitation id="AD233AE1FFD6FF89FBF1FB53FA1D6F44" author="Gosliner T &amp; Johnson, R." box="[1114,1419,1278,1300]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="52" pageNumber="53" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
: specimen CASIZ 073533;
<bibRefCitation id="AD233AE1FFD6FF89FBE1FAB3FB406F62" author="Fraser V." box="[1098,1238,1309,1331]" journalOrPublisher="Sydney: Australian Museum" pageId="52" pageNumber="53" publicationUrl="http://www.seaslugforum.net/find/1628" refId="ref42054" refString="Fraser V. 1999. Hypselodoris from South Africa. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 1628" title="Hypselodoris from South Africa" type="url" volumeTitle="Sea slug forum" year="1999">Fraser, 1999</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD6FF89FB48FAB0FCF66F01" author="Poddubetskaia M." journalOrPublisher="Sydney: Australian Museum" pageId="52" pageNumber="53" publicationUrl="http://www.seaslugforum.net/find/9266" refId="ref44244" refString="Poddubetskaia M. 2003. Hypselodoris maculosa from Egypt. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 9266" title="Hypselodoris maculosa from Egypt" type="url" volumeTitle="Sea slug forum" year="2003">Poddubetskaia, 2003</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD6FF89FCC7FA91FBB36F02" author="Lederman O." box="[876,1061,1340,1362]" journalOrPublisher="Sydney: Australian Museum" pageId="52" pageNumber="53" publicationUrl="http://www.seaslugforum.net/find/13209" refId="ref43595" refString="Lederman O. 2005. Hypselodoris maculosa from the Red Sea. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 13209" title="Hypselodoris maculosa from the Red Sea" type="url" volumeTitle="Sea slug forum" year="2005">Lederman, 2005</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD6FF89FB99FA91FB596F01" author="Houben A." box="[1074,1231,1340,1362]" journalOrPublisher="Sydney: Australian Museum" pageId="52" pageNumber="53" publicationUrl="http://www.seaslugforum.net/find/20856" refId="ref42947" refString="Houben A. 2007. Hypselodoris cf. maculosa from Sri Lanka. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20856" title="Hypselodoris cf. maculosa from Sri Lanka" type="url" volumeTitle="Sea slug forum" year="2007">Houben, 2007</bibRefCitation>
.
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82FAF7FC256FDF" blockId="52.[809,1425,1248,1576]" pageId="52" pageNumber="53">
<taxonomicName id="0EB23C93FFD6FF89FC82FAF7FC546F3F" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,962,1370,1391]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD6FF89FC82FAF7FC546F3F" box="[809,962,1370,1391]" italics="true" pageId="52" pageNumber="53">Hypselodoris</emphasis>
</taxonomicName>
sp. 1
<bibRefCitation id="AD233AE1FFD6FF89FBA2FAF7FB6F6F20" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1033,1273,1370,1392]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="52" pageNumber="53" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFD6FF89FBDDFAF6FB276F3F" box="[1142,1201,1370,1392]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
, 2008
</bibRefCitation>
: 259, middle photograph.
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82FA35FAE86FFD" blockId="52.[809,1425,1248,1576]" box="[809,1406,1432,1454]" pageId="52" pageNumber="53">
<taxonomicName id="0EB23C93FFD6FF89FC82FA35FAE16FFD" ID-CoL="3P34H" authority="Johnson &amp; Gosliner, 2012" authorityName="Johnson &amp; Gosliner" authorityYear="2012" box="[809,1399,1432,1454]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD6FF89FC82FA35FC2A6FFD" box="[809,956,1432,1453]" italics="true" pageId="52" pageNumber="53">Hypselodoris</emphasis>
cf.
<emphasis id="FBC69B02FFD6FF89FC49FA35FBD86FFD" box="[994,1102,1432,1453]" italics="true" pageId="52" pageNumber="53">maculosa</emphasis>
<bibRefCitation id="AD233AE1FFD6FF89FBFFFA35FAE16FFD" author="Johnson R &amp; Gosliner T." box="[1108,1399,1432,1454]" journalOrPublisher="PLoS One" pageId="52" pageNumber="53" pagination="33479" part="7" refId="ref43042" refString="Johnson R, Gosliner T. 2012. Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs. PLoS One 7: e 33479." title="Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs" type="journal article" year="2012">Johnson &amp; Gosliner, 2012</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82FA1BFC256FBB" blockId="52.[809,1425,1248,1576]" pageId="52" pageNumber="53">
<taxonomicName id="0EB23C93FFD6FF89FC82FA1BFC2A6F9B" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,956,1462,1483]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD6FF89FC82FA1BFC2A6F9B" box="[809,956,1462,1483]" italics="true" pageId="52" pageNumber="53">Hypselodoris</emphasis>
</taxonomicName>
sp. 4
<bibRefCitation id="AD233AE1FFD6FF89FC56FA1BFB496F9C" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1021,1247,1462,1484]" journalOrPublisher="Jacksonville: New World Press" pageId="52" pageNumber="53" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFD6FF89FBCFFA1AFB0A6F9B" box="[1124,1180,1462,1484]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
, 2015
</bibRefCitation>
: 255, upper left photograph.
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82FA59FC7D6C78" blockId="52.[809,1425,1248,1576]" pageId="52" pageNumber="53">
<taxonomicName id="0EB23C93FFD6FF89FC82FA59FC526C59" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[809,964,1524,1545]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFD6FF89FC82FA59FC526C59" box="[809,964,1524,1545]" italics="true" pageId="52" pageNumber="53">Hypselodoris</emphasis>
</taxonomicName>
sp. 1
<bibRefCitation id="AD233AE1FFD6FF89FBA6FA59FA946C5A" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1037,1282,1524,1546]" journalOrPublisher="Jacksonville: New World Press" pageId="52" pageNumber="53" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFD6FF89FBD7FA59FB2F6C59" box="[1148,1209,1524,1545]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
, 2015
</bibRefCitation>
: 253, upper right photograph
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD6FF8BFC82F9FDFEF96C21" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="C90D4710FFD6FF89FC82F9FDFC456C37" blockId="52.[809,1425,1615,1762]" box="[809,979,1615,1639]" pageId="52" pageNumber="53">
<heading id="9245F07CFFD6FF89FC82F9FDFC456C37" box="[809,979,1615,1639]" fontSize="10" level="4" pageId="52" pageNumber="53" reason="1">
<emphasis id="FBC69B02FFD6FF89FC82F9FDFC456C37" box="[809,979,1615,1639]" italics="true" pageId="52" pageNumber="53">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD6FF89FC82F9DDFB666CB2" blockId="52.[809,1425,1615,1762]" pageId="52" pageNumber="53">
<materialsCitation id="79DA4D4DFFD6FF89FC82F9DDFB666CB2" ID-GBIF-Occurrence="2235579045" collectingDate="2014-05-11" collectionCode="CASIZ" country="Mozambique" elevation="16" latitude="-26.8389" location="Doodles" municipality="Ponta do Ouro" pageId="52" pageNumber="53" specimenCode="CASIZ 223316" specimenCount="1" typeStatus="holotype">
<typeStatus id="1609F9B2FFD6FF89FC82F9DDFC0E6CD5" box="[809,920,1648,1669]" pageId="52" pageNumber="53" type="holotype">
<emphasis id="FBC69B02FFD6FF89FC82F9DDFC0E6CD5" box="[809,920,1648,1669]" italics="true" pageId="52" pageNumber="53">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFD6FF89FC01F9DDFBF16CD6" box="[938,1127,1648,1670]" collectionCode="CASIZ" pageId="52" pageNumber="53">CASIZ 223316</specimenCode>
, one specimen,
<quantity id="0E4AEAF5FFD6FF89FA98F9DDFA1A6CD5" box="[1331,1420,1648,1669]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="52" pageNumber="53" unit="mm" value="17.0">17 mm</quantity>
, subsampled for molecular study,
<location id="CC6D11CBFFD6FF89FB6BF923FAB66CF4" LSID="urn:lsid:plazi:treatment:411BF606FFD6FF8AFC73FBADFC0F69CF:CC6D11CBFFD6FF89FB6BF923FAB66CF4" box="[1216,1312,1678,1700]" country="Mozambique" latitude="-26.8389" municipality="Ponta do Ouro" name="Doodles" pageId="52" pageNumber="53">Doodles</location>
dive site (
<geoCoordinate id="AC8621D7FFD6FF89FC9AF900FC3F6C93" box="[817,937,1709,1731]" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="5" value="-26.8389">26.8389°S</geoCoordinate>
328917°E),
<collectingMunicipality id="2969DD6AFFD6FF89FB91F903FB716C92" box="[1082,1255,1709,1731]" pageId="52" pageNumber="53">Ponta do Ouro</collectingMunicipality>
,
<collectingCountry id="B1A50780FFD6FF89FB5FF900FA1A6C93" box="[1268,1420,1709,1731]" name="Mozambique" pageId="52" pageNumber="53">Mozambique</collectingCountry>
,
<elevation id="429FA023FFD6FF89FC82F961FCF76CB1" box="[809,865,1740,1762]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="52" pageNumber="53" unit="m" value="16.0">
<quantity id="0E4AEAF5FFD6FF89FC82F961FCF76CB1" box="[809,865,1740,1762]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="52" pageNumber="53" unit="m" value="16.0">16 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFD6FF89FC1DF961FBDC6CB1" box="[950,1098,1740,1761]" pageId="52" pageNumber="53" value="2014-05-11">
<date id="BD0C61D0FFD6FF89FC1DF961FBDC6CB1" box="[950,1098,1740,1761]" pageId="52" pageNumber="53" value="2014-05-11">11 May 2014</date>
</collectingDate>
, Yara Tibiriçá.
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFD6FF8BFC82F8A6FEF96C21" blockId="52.[809,1425,1802,1886]" lastBlockId="54.[145,761,1535,1649]" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
<materialsCitation id="79DA4D4DFFD6FF89FC82F8A6FB3A6D0E" ID-GBIF-Occurrence="2235579112" collectingDate="1990-03-27" collectionCode="CASIZ" collectorName="Ifaty, SW &amp; T. Gosliner" country="Madagascar" elevation="2" location="Mora Mora Resort" pageId="52" pageNumber="53" specimenCount="1" typeStatus="paratype">
<typeStatus id="1609F9B2FFD6FF89FC82F8A6FC096D70" box="[809,927,1803,1824]" pageId="52" pageNumber="53" type="paratype">
<emphasis id="FBC69B02FFD6FF89FC82F8A6FC096D70" box="[809,927,1803,1824]" italics="true" pageId="52" pageNumber="53">Paratypes</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFD6FF89FC04F8A6FC696D70" box="[943,1023,1803,1824]" pageId="52" pageNumber="53">CASIZ</collectionCode>
0 73533, one specimen,
<collectorName id="644722C6FFD6FF89FA8AF8A7FA076D70" box="[1313,1425,1802,1824]" pageId="52" pageNumber="53">Ifaty, SW</collectorName>
<collectingCountry id="B1A50780FFD6FF89FC82F884FC2B6D6F" box="[809,957,1833,1855]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
,
<quantity id="0E4AEAF5FFD6FF89FC61F884FB9E6D6F" box="[970,1032,1833,1855]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="52" pageNumber="53" unit="km" value="1.0">1 km</quantity>
west of
<location id="CC6D11CBFFD6FF89FBD9F887FAC36D6F" LSID="urn:lsid:plazi:treatment:411BF606FFD6FF8AFC73FBADFC0F69CF:CC6D11CBFFD6FF89FBD9F887FAC36D6F" box="[1138,1365,1834,1855]" country="Madagascar" name="Mora Mora Resort" pageId="52" pageNumber="53">Mora Mora Resort</location>
,
<elevation id="429FA023FFD6FF89FAC8F884FA076D6F" box="[1379,1425,1833,1855]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="53" unit="m" value="2.0">
<quantity id="0E4AEAF5FFD6FF89FAC8F884FA076D6F" box="[1379,1425,1833,1855]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="53" unit="m" value="2.0">2 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFD6FF89FCD3F8E5FBB36D0D" box="[888,1061,1864,1886]" pageId="52" pageNumber="53" value="1990-03-27">
<date id="BD0C61D0FFD6FF89FCD3F8E5FBB36D0D" box="[888,1061,1864,1886]" pageId="52" pageNumber="53" value="1990-03-27">27 March 1990</date>
</collectingDate>
,
<collectorName id="644722C6FFD6FF89FB84F8E5FB3A6D0E" box="[1071,1196,1864,1886]" pageId="52" pageNumber="53">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFD6FF8BFB1DF8E5FEFA6C21" ID-GBIF-Occurrence="2235579133" collectingDate="2014-06-09" collectionCode="CASIZ" collectorName="Yara Tibirica" country="Mozambique" elevation="12" lastPageId="54" lastPageNumber="55" latitude="-14.490278" location="Kwalala Lodge" longLatPrecision="21" longitude="40.74167" municipality="Nacala" pageId="52" pageNumber="53" specimenCode="CASIZ 223317" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFD6FF89FB1DF8E5FAC86D0D" box="[1206,1374,1864,1885]" collectionCode="CASIZ" pageId="52" pageNumber="53">CASIZ 223317</specimenCode>
, one specimen,
<quantity id="0E4AEAF5FFD4FF8BFEACFA52FEC06C45" box="[263,342,1535,1557]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="54" pageNumber="55" unit="mm" value="26.0">26 mm</quantity>
, dissected, subsampled for molecular study, in front of
<location id="CC6D11CBFFD4FF8BFE29F9B3FDDD6C64" LSID="urn:lsid:plazi:treatment:411BF606FFD6FF8AFC73FBADFC0F69CF:CC6D11CBFFD4FF8BFE29F9B3FDDD6C64" box="[386,587,1566,1588]" country="Mozambique" latitude="-14.490278" longLatPrecision="21" longitude="40.74167" municipality="Nacala" name="Kwalala Lodge" pageId="54" pageNumber="55">Kwalala Lodge</location>
, (
<geoCoordinate id="AC8621D7FFD4FF8BFDCFF9B3FD656C63" box="[612,755,1565,1588]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="15" value="-14.490278">
14°29
<emphasis id="FBC69B02FFD4FF8BFD04F9B0FD236C64" box="[687,693,1565,1588]" italics="true" pageId="54" pageNumber="55">′</emphasis>
25
<emphasis id="FBC69B02FFD4FF8BFD7CF9B0FD776C64" box="[727,737,1565,1588]" italics="true" pageId="54" pageNumber="55">″</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFD4FF8BFF3AF990FE866C02" box="[145,272,1595,1618]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="15" value="40.74167">
40°44
<emphasis id="FBC69B02FFD4FF8BFF7AF996FF416C02" box="[209,215,1595,1618]" italics="true" pageId="54" pageNumber="55">′</emphasis>
30
<emphasis id="FBC69B02FFD4FF8BFF58F996FF6B6C02" box="[243,253,1595,1618]" italics="true" pageId="54" pageNumber="55">″</emphasis>
E
</geoCoordinate>
),
<collectingMunicipality id="2969DD6AFFD4FF8BFE8FF991FEE06C02" box="[292,374,1596,1618]" pageId="54" pageNumber="55">Nacala</collectingMunicipality>
,
<collectingCountry id="B1A50780FFD4FF8BFE2AF991FD826C02" box="[385,532,1596,1618]" name="Mozambique" pageId="54" pageNumber="55">Mozambique</collectingCountry>
,
<elevation id="429FA023FFD4FF8BFDB4F990FDC16C02" box="[543,599,1597,1618]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="54" pageNumber="55" unit="m" value="12.0">
<quantity id="0E4AEAF5FFD4FF8BFDB4F990FDC16C02" box="[543,599,1597,1618]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="54" pageNumber="55" unit="m" value="12.0">12 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFD4FF8BFD06F990FF5C6C20" pageId="54" pageNumber="55" value="2014-06-09">
<date id="BD0C61D0FFD4FF8BFD06F990FF5C6C20" pageId="54" pageNumber="55" value="2014-06-09">9 June 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFD4FF8BFF78F9F1FEFA6C21" box="[211,364,1627,1649]" pageId="54" pageNumber="55">Yara Tibiriçá</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="9DCD1798FFD7FF88FF08FC73FAD06E7E" httpUri="https://zenodo.org/record/2632858/files/figure.png" pageId="53" pageNumber="54" startId="53.[163,244,990,1012]" targetBox="[163,1443,195,966]" targetPageId="53">
<paragraph id="C90D4710FFD7FF88FF08FC73FAD06E7E" blockId="53.[163,1443,989,1070]" pageId="53" pageNumber="54">
<emphasis id="FBC69B02FFD7FF88FF08FC73FE8A69A3" bold="true" box="[163,284,990,1012]" pageId="53" pageNumber="54">Figure 32.</emphasis>
<taxonomicName id="0EB23C93FFD7FF88FE8CFC70FD7E69A4" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[295,744,989,1012]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="yarae" status="sp. nov.">
<emphasis id="FBC69B02FFD7FF88FE8CFC70FD9D69A3" bold="true" box="[295,523,989,1011]" italics="true" pageId="53" pageNumber="54">Hypselodoris yarae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD7FF88FD44FC73FCDF69A3" box="[751,841,990,1011]" pageId="53" pageNumber="54" rank="species">
<emphasis id="FBC69B02FFD7FF88FD44FC73FCDF69A3" bold="true" box="[751,841,990,1011]" pageId="53" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
, living animals. A, photograph of specimen from Îles de Radama, October 2005. B, holotype, CASIZ 223316, Ponto de Ouro, Mozambique, photograph by Yara Tibiriçá. C, paratype, CASIZ 223317, Nacala, Mozambique, photograph by Yara Tibiriçá. D, paratype, CASIZ 0 73533, Ifaty, Madagascar.
</paragraph>
</caption>
<caption id="9DCD1798FFD7FF88FF08F95EFB0B6D14" httpUri="https://zenodo.org/record/2632860/files/figure.png" pageId="53" pageNumber="54" startId="53.[163,241,1779,1801]" targetBox="[322,1282,1106,1736]" targetPageId="53">
<paragraph id="C90D4710FFD7FF88FF08F95EFB0B6D14" blockId="53.[163,1444,1779,1860]" pageId="53" pageNumber="54">
<emphasis id="FBC69B02FFD7FF88FF08F95EFE8E6D58" bold="true" box="[163,280,1779,1801]" pageId="53" pageNumber="54">Figure 33.</emphasis>
<taxonomicName id="0EB23C93FFD7FF88FE89F95EFD456D59" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[290,723,1779,1801]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="yarae" status="sp. nov.">
<emphasis id="FBC69B02FFD7FF88FE89F95EFE696D58" bold="true" box="[290,511,1779,1801]" italics="true" pageId="53" pageNumber="54">Hypselodoris yarae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD7FF88FD72F95EFCB96D58" box="[729,815,1779,1800]" pageId="53" pageNumber="54" rank="species">
<emphasis id="FBC69B02FFD7FF88FD72F95EFCB96D58" bold="true" box="[729,815,1779,1800]" pageId="53" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
A, mantle glands. B, reproductive system, scale bar: 0.5 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland.
</paragraph>
</caption>
<caption id="9DCD1798FFD4FF8BFF3AFA0AFB166F8B" httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="54" pageNumber="55" startId="54.[145,223,1447,1469]" targetBox="[209,1361,197,1407]" targetPageId="54">
<paragraph id="C90D4710FFD4FF8BFF3AFA0AFB166F8B" blockId="54.[145,1425,1447,1499]" pageId="54" pageNumber="55">
<emphasis id="FBC69B02FFD4FF8BFF3AFA0AFE906FEC" bold="true" box="[145,262,1447,1469]" pageId="54" pageNumber="55">Figure 34.</emphasis>
<taxonomicName id="0EB23C93FFD4FF8BFEBAFA0AFD536FEE" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[273,709,1447,1470]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="yarae" status="sp. nov.">
<emphasis id="FBC69B02FFD4FF8BFEBAFA0AFE796FED" bold="true" box="[273,495,1447,1469]" italics="true" pageId="54" pageNumber="55">Hypselodoris yarae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD4FF8BFD60FA05FCB46FED" box="[715,802,1448,1469]" pageId="54" pageNumber="55" rank="species">
<emphasis id="FBC69B02FFD4FF8BFD60FA05FCB46FED" bold="true" box="[715,802,1448,1469]" pageId="54" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth.
</paragraph>
</caption>
<subSubSection id="81A8149BFFD4FF8BFF3AF935FE0B6D19" pageId="54" pageNumber="55" type="distribution">
<paragraph id="C90D4710FFD4FF8BFF3AF935FE586CE0" blockId="54.[145,761,1688,1865]" box="[145,462,1688,1712]" pageId="54" pageNumber="55">
<heading id="9245F07CFFD4FF8BFF3AF935FE586CE0" box="[145,462,1688,1712]" fontSize="10" level="4" pageId="54" pageNumber="55" reason="1">
<emphasis id="FBC69B02FFD4FF8BFF3AF935FE586CE0" box="[145,462,1688,1712]" italics="true" pageId="54" pageNumber="55">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD4FF8BFF3AF914FE0B6D19" blockId="54.[145,761,1688,1865]" pageId="54" pageNumber="55">
Known only from
<collectingCountry id="B1A50780FFD4FF8BFECFF915FE656C9E" box="[356,499,1720,1742]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
and
<collectingCountry id="B1A50780FFD4FF8BFD84F915FD536C9E" box="[559,709,1720,1742]" name="Mozambique" pageId="54" pageNumber="55">Mozambique</collectingCountry>
and possibly from
<collectingCountry id="B1A50780FFD4FF8BFE99F975FEEA6CBD" box="[306,380,1752,1773]" name="Kenya" pageId="54" pageNumber="55">Kenya</collectingCountry>
(
<bibRefCitation id="AD233AE1FFD4FF8BFE27F97AFDE76CBD" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[396,625,1751,1773]" journalOrPublisher="Jacksonville: New World Press" pageId="54" pageNumber="55" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFD4FF8BFE5FF975FDBB6CBC" box="[500,557,1751,1773]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
, 2015
</bibRefCitation>
: 252, upper right photograph),
<collectingCountry id="B1A50780FFD4FF8BFED2F95BFD856D5C" box="[377,531,1782,1804]" name="South Africa" pageId="54" pageNumber="55">South Africa</collectingCountry>
(
<bibRefCitation id="AD233AE1FFD4FF8BFD8EF95BFD2A6D5B" author="Fraser V." box="[549,700,1782,1803]" journalOrPublisher="Sydney: Australian Museum" pageId="54" pageNumber="55" publicationUrl="http://www.seaslugforum.net/find/1628" refId="ref42054" refString="Fraser V. 1999. Hypselodoris from South Africa. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 1628" title="Hypselodoris from South Africa" type="url" volumeTitle="Sea slug forum" year="1999">Fraser, 1999</bibRefCitation>
),
<collectingCountry id="B1A50780FFD4FF8BFD7EF95BFF4D6D7A" name="Sri Lanka" pageId="54" pageNumber="55">Sri Lanka</collectingCountry>
(
<bibRefCitation id="AD233AE1FFD4FF8BFF4CF8B9FE176D7A" author="Houben A." box="[231,385,1812,1834]" journalOrPublisher="Sydney: Australian Museum" pageId="54" pageNumber="55" publicationUrl="http://www.seaslugforum.net/find/20856" refId="ref42947" refString="Houben A. 2007. Hypselodoris cf. maculosa from Sri Lanka. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20856" title="Hypselodoris cf. maculosa from Sri Lanka" type="url" volumeTitle="Sea slug forum" year="2007">Houben, 2007</bibRefCitation>
) and the Red Sea (
<bibRefCitation id="AD233AE1FFD4FF8BFDE0F8B9FF5E6D18" author="Poddubetskaia M." journalOrPublisher="Sydney: Australian Museum" pageId="54" pageNumber="55" publicationUrl="http://www.seaslugforum.net/find/9266" refId="ref44244" refString="Poddubetskaia M. 2003. Hypselodoris maculosa from Egypt. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 9266" title="Hypselodoris maculosa from Egypt" type="url" volumeTitle="Sea slug forum" year="2003">Poddubetskaia, 2003</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFD4FF8BFF7FF89EFE1B6D19" author="Lederman O." box="[212,397,1843,1865]" journalOrPublisher="Sydney: Australian Museum" pageId="54" pageNumber="55" publicationUrl="http://www.seaslugforum.net/find/13209" refId="ref43595" refString="Lederman O. 2005. Hypselodoris maculosa from the Red Sea. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 13209" title="Hypselodoris maculosa from the Red Sea" type="url" volumeTitle="Sea slug forum" year="2005">Lederman, 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD4FF8BFC82FA52FC166CC1" pageId="54" pageNumber="55" type="etymology">
<paragraph id="C90D4710FFD4FF8BFC82FA52FC3C6C47" blockId="54.[809,1425,1535,1681]" box="[809,938,1535,1559]" pageId="54" pageNumber="55">
<heading id="9245F07CFFD4FF8BFC82FA52FC3C6C47" box="[809,938,1535,1559]" fontSize="10" level="4" pageId="54" pageNumber="55" reason="1">
<emphasis id="FBC69B02FFD4FF8BFC82FA52FC3C6C47" box="[809,938,1535,1559]" italics="true" pageId="54" pageNumber="55">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD4FF8BFC82F9B2FC166CC1" blockId="54.[809,1425,1535,1681]" pageId="54" pageNumber="55">
The name
<taxonomicName id="0EB23C93FFD4FF8BFC1EF98DFC6A6C65" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[949,1020,1568,1589]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFD4FF8BFC1EF98DFC6A6C65" box="[949,1020,1568,1589]" italics="true" pageId="54" pageNumber="55">yarae</emphasis>
</taxonomicName>
honors Yara Tibiriçá, who has advanced our knowledge of western Indian Ocean nudibranchs and collected several specimens of this species.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD4FF8AFC82F914FAB96A8C" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph id="C90D4710FFD4FF8BFC82F914FC236C80" blockId="54.[809,1425,1721,1867]" box="[809,949,1721,1744]" pageId="54" pageNumber="55">
<heading id="9245F07CFFD4FF8BFC82F914FC236C80" box="[809,949,1721,1744]" fontSize="10" level="4" pageId="54" pageNumber="55" reason="1">
<emphasis id="FBC69B02FFD4FF8BFC82F914FC236C80" box="[809,949,1721,1744]" italics="true" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD4FF8AFC82F974FD076BA0" blockId="54.[809,1425,1721,1867]" lastBlockId="55.[163,779,198,496]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis id="FBC69B02FFD4FF8BFC82F974FBB86CBE" box="[809,1070,1753,1774]" italics="true" pageId="54" pageNumber="55">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFD4FF8BFABAF974FAFE6CBE" box="[1297,1384,1753,1775]" captionStart="Figure 32" captionStartId="53.[163,244,990,1012]" captionTargetBox="[163,1443,195,966]" captionTargetId="figure@53.[163,1443,195,966]" captionTargetPageId="53" captionText="Figure 32. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., living animals. A, photograph of specimen from Îles de Radama, October 2005. B, holotype, CASIZ 223316, Ponto de Ouro, Mozambique, photograph by Yara Tibiriçá. C, paratype, CASIZ 223317, Nacala, Mozambique, photograph by Yara Tibiriçá. D, paratype, CASIZ 073533, Ifaty, Madagascar." httpUri="https://zenodo.org/record/2632858/files/figure.png" pageId="54" pageNumber="55">Fig. 32</figureCitation>
) of moderate size,
<quantity id="0E4AEAF5FFD4FF8BFC48F95AFBF26D5D" box="[995,1124,1783,1805]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="54" pageNumber="55" unit="mm" value="27.5" valueMax="35.0" valueMin="20.0">20–35 mm</quantity>
in length. Entire dorsal surface light pink to tan, with thick orange band encircling the margin of notum. Marginal band often with undulating margins on inner side. Notum with series of narrow, opaque white longitudinal lines. Large plum to brown spots located between lines. Sides of body and margin of foot same colour as notum. Gill branches orange basally, with opaque white apices. Seven to nine unipinnate, well-elevated gill branches present on notum. Bulb of rhinophores opaque white, with two prominent orange bands and traces of third band basally. Base of rhinophores translucent white. Bulb with ~12–13 densely packed lamellae.
</paragraph>
<paragraph id="C90D4710FFD5FF8AFF08FDA4FE8F680C" blockId="55.[163,778,521,604]" pageId="55" pageNumber="56">
<emphasis id="FBC69B02FFD5FF8AFF08FDA4FEC7684E" box="[163,337,521,542]" italics="true" pageId="55" pageNumber="56">Mantle glands</emphasis>
: Mantle glands present along entire mantle margin, larger posteriorly and anteriorly (
<figureCitation id="51895B95FFD5FF8AFF00FDEBFE9C680C" box="[171,266,582,604]" captionStart="Figure 33" captionStartId="53.[163,241,1779,1801]" captionTargetBox="[322,1282,1106,1736]" captionTargetId="figure@53.[321,1284,1105,1738]" captionTargetPageId="53" captionText="Figure 33. Hypselodoris yarae Gosliner &amp; Johnson sp. nov. A, mantle glands. B, reproductive system, scale bar: 0.5 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632860/files/figure.png" pageId="55" pageNumber="56">Fig. 33A</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFD5FF8AFF08FDD8FD4D6EE3" blockId="55.[163,779,629,1203]" pageId="55" pageNumber="56">
<emphasis id="FBC69B02FFD5FF8AFF08FDD8FEF868DB" box="[163,366,629,651]" italics="true" pageId="55" pageNumber="56">Buccal armature</emphasis>
: Muscular portion of buccal mass smaller in size than oral tube. Chitinous labial cuticle elongate (
<figureCitation id="51895B95FFD5FF8AFEBCFD1FFEEC6898" box="[279,378,690,712]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34A</figureCitation>
), narrow, found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFD5FF8AFE9CFD5DFE0D6955" box="[311,411,752,774]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34B</figureCitation>
). Rodlets straight, narrow and long, with long base. Rodlets evenly curved, with single, acutely pointed apex with lateral lobes. Radular formula of
<typeStatus id="1609F9B2FFD5FF8AFE86FCE1FE0D6931" box="[301,411,844,865]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
(CASIZ 073533) 45 × 31.0.31 (
<figureCitation id="51895B95FFD5FF8AFF00FCC7FE9B69D0" box="[171,269,874,896]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34C</figureCitation>
). Rachidian row of teeth absent (
<figureCitation id="51895B95FFD5FF8AFD33FCC7FD6A69D0" box="[664,764,874,896]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34D</figureCitation>
). Innermost lateral teeth having zero to two short denticles on inner side of bifid primary cusp and one or two denticles on the outer side of the bifid cusp. Outer cusp of bifid cusps about same size as inner one. Second lateral tooth and next several laterals lacking inner denticles, possessing primary bifid cusps and one ot three outer denticles. Midlateral teeth (
<figureCitation id="51895B95FFD5FF8AFD34FBECFC946E06" box="[671,770,1089,1111]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34E</figureCitation>
) all lacking inner denticles but with three or four prominent outer denticles. Outermost teeth (
<figureCitation id="51895B95FFD5FF8AFD0FFBD3FC946EC4" box="[676,770,1150,1172]" captionStart="Figure 34" captionStartId="54.[145,223,1447,1469]" captionTargetBox="[209,1361,197,1407]" captionTargetId="figure@54.[209,1361,197,1407]" captionTargetPageId="54" captionText="Figure 34. Hypselodoris yarae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 223316. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632864/files/figure.png" pageId="55" pageNumber="56">Fig. 34F</figureCitation>
) having a narrower base and three outer denticles.
</paragraph>
<paragraph id="C90D4710FFD5FF8AFF08FB71FAB96A8C" blockId="55.[163,779,1244,1879]" lastBlockId="55.[827,1327,198,220]" pageId="55" pageNumber="56">
<emphasis id="FBC69B02FFD5FF8AFF08FB71FE356EA1" box="[163,419,1244,1265]" italics="true" pageId="55" pageNumber="56">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFD5FF8AFF08FB56FE9C6F40" box="[163,266,1275,1296]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
(CASIZ 223317) are fully mature (
<figureCitation id="51895B95FFD5FF8AFD37FB57FD6A6F40" box="[668,764,1274,1296]" captionStart="Figure 33" captionStartId="53.[163,241,1779,1801]" captionTargetBox="[322,1282,1106,1736]" captionTargetId="figure@53.[321,1284,1105,1738]" captionTargetPageId="53" captionText="Figure 33. Hypselodoris yarae Gosliner &amp; Johnson sp. nov. A, mantle glands. B, reproductive system, scale bar: 0.5 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632860/files/figure.png" pageId="55" pageNumber="56">Fig. 33B</figureCitation>
). Ampulla think, tubular and curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens long, convoluted, curved and thick and narrowing abruptly as it transitions into muscular ejaculatory portion. Ejaculatory portion moderately short, straight, narrow, entering moderately short, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, curved receptaculum seminis and larger spherical, thin-walled bursa copulatrix. Receptaculum seminis located in distal half of vaginal length. Long uterine duct, wide at base of vagina, narrowing and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD5FF8AFC90FEAEFC0F69CF" pageId="55" pageNumber="56" type="discussion">
<paragraph id="C90D4710FFD5FF8AFC90FEAEFC3E6B4B" blockId="55.[827,1444,259,927]" box="[827,936,259,283]" pageId="55" pageNumber="56">
<heading id="9245F07CFFD5FF8AFC90FEAEFC3E6B4B" box="[827,936,259,283]" fontSize="10" level="4" pageId="55" pageNumber="56" reason="1">
<emphasis id="FBC69B02FFD5FF8AFC90FEAEFC3E6B4B" box="[827,936,259,283]" italics="true" pageId="55" pageNumber="56">Remarks</emphasis>
</heading>
</paragraph>
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This species was initially recognized as genetically distinct from
<taxonomicName id="0EB23C93FFD5FF8AFC4EFEEEFBE86B07" authorityName="Johnson &amp; Gosliner" authorityYear="2012" box="[997,1150,322,344]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD5FF8AFC4EFEEEFBE86B07" box="[997,1150,322,344]" italics="true" pageId="55" pageNumber="56">H. maculosa</emphasis>
</taxonomicName>
by
<bibRefCitation id="AD233AE1FFD5FF8AFB05FEEFFC126B27" author="Johnson R &amp; Gosliner T." journalOrPublisher="PLoS One" pageId="55" pageNumber="56" pagination="33479" part="7" refId="ref43042" refString="Johnson R, Gosliner T. 2012. Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs. PLoS One 7: e 33479." title="Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs" type="journal article" year="2012">Johnson &amp; Gosliner (2012)</bibRefCitation>
, as
<emphasis id="FBC69B02FFD5FF8AFC18FECCFBA96B26" box="[947,1087,353,374]" italics="true" pageId="55" pageNumber="56">Hypselodori</emphasis>
s sp.
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cf.
<emphasis id="FBC69B02FFD5FF8AFB30FECCFA9F6B26" box="[1179,1289,353,374]" italics="true" pageId="55" pageNumber="56">maculosa</emphasis>
</taxonomicName>
. The present study also indicates a basal position as being sister to the remaining members of its large clade. Its distinctness is also strongly supported by the ABGD analysis. Externally, there are differences between
<taxonomicName id="0EB23C93FFD5FF8AFC90FE56FC316840" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,935,507,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFD5FF8AFC90FE56FC316840" box="[827,935,507,528]" italics="true" pageId="55" pageNumber="56">H. yarae</emphasis>
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and other members of the
<taxonomicName id="0EB23C93FFD5FF8AFAACFE56FA32685F" authorityName="Johnson &amp; Gosliner" authorityYear="2012" box="[1287,1444,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD5FF8AFAACFE56FA32685F" box="[1287,1444,506,528]" italics="true" pageId="55" pageNumber="56">H. maculosa</emphasis>
</taxonomicName>
clade. This species has a darker body colour than
<taxonomicName id="0EB23C93FFD5FF8AFC90FD95FC7D681D" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1003,568,589]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFD5FF8AFC90FD95FC7D681D" box="[827,1003,568,589]" italics="true" pageId="55" pageNumber="56">H. alburtuqali</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD5FF8AFC51FD95FB11681D" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1018,1159,568,589]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD5FF8AFC51FD95FB11681D" box="[1018,1159,568,589]" italics="true" pageId="55" pageNumber="56">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD5FF8AFB6DFD95FACB681D" authorityName="Johnson &amp; Gosliner" authorityYear="2012" box="[1222,1373,568,589]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD5FF8AFB6DFD95FACB681D" box="[1222,1373,568,589]" italics="true" pageId="55" pageNumber="56">H. maculosa</emphasis>
</taxonomicName>
, with fewer, larger brown to plum spots. It also differs from
<taxonomicName id="0EB23C93FFD5FF8AFCD2FDDBFBB468DA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[889,1058,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFD5FF8AFCD2FDDBFBB468DA" box="[889,1058,629,651]" italics="true" pageId="55" pageNumber="56">H. alburtuqali</emphasis>
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in having a wide orange margin that usually has undulations along its inner margin. The jaws of
<taxonomicName id="0EB23C93FFD5FF8AFC64FD1EFBA06898" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[975,1078,691,712]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFD5FF8AFC64FD1EFBA06898" box="[975,1078,691,712]" italics="true" pageId="55" pageNumber="56">H. yarae</emphasis>
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are much narrower and more elongate than are those of
<taxonomicName id="0EB23C93FFD5FF8AFBD4FD7FFABA68B6" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1151,1324,721,743]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFD5FF8AFBD4FD7FFABA68B6" box="[1151,1324,721,743]" italics="true" pageId="55" pageNumber="56">H. alburtuqali</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD5FF8AFA95FD7CFA0268B6" box="[1342,1428,721,743]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="55" pageNumber="56">Fig. 3A</figureCitation>
),
<taxonomicName id="0EB23C93FFD5FF8AFC90FD5DFC506955" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[827,966,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD5FF8AFC90FD5DFC506955" box="[827,966,752,773]" italics="true" pageId="55" pageNumber="56">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD5FF8AFC73FD5DFBA86955" box="[984,1086,752,774]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="55" pageNumber="56">Fig. 10A</figureCitation>
) and
<taxonomicName id="0EB23C93FFD5FF8AFB2DFD5DFA8B6955" authorityName="Johnson &amp; Gosliner" authorityYear="2012" box="[1158,1309,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD5FF8AFB2DFD5DFA8B6955" box="[1158,1309,752,773]" italics="true" pageId="55" pageNumber="56">H. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD5FF8AFA85FD5DFA026955" box="[1326,1428,752,774]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="55" pageNumber="56">Fig. 11A</figureCitation>
). The most distinctive anatomical features of
<taxonomicName id="0EB23C93FFD5FF8AFAEBFCA2FA356974" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1344,1443,783,804]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFD5FF8AFAEBFCA2FA356974" box="[1344,1443,783,804]" italics="true" pageId="55" pageNumber="56">H. yarae</emphasis>
</taxonomicName>
from the above three species are the basal insertion of the uterine duct into the vagina that has an expanded base, and the more basal position of the receptaculum seminis.
</paragraph>
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GOSLINER &amp; JOHNSON
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<emphasis id="FBC69B02FFE4FFBBFDB3FA7EFDED6FB7" bold="true" box="[536,635,1487,1511]" pageId="6" pageNumber="7">SP. NOV.</emphasis>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE4FFBBFF3AF93CFEDE6C96" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="C90D4710FFE4FFBBFF3AF93CFEDE6C96" blockId="6.[145,731,1681,1734]" pageId="6" pageNumber="7">
<taxonomicName id="0EB23C93FFE4FFBBFF3AF93CFEB26CF6" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,292,1681,1702]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFE4FFBBFF3AF93CFEB26CF6" box="[145,292,1681,1702]" italics="true" pageId="6" pageNumber="7">Hypselodoris</emphasis>
</taxonomicName>
sp. 2
<bibRefCitation id="AD233AE1FFE4FFBBFECEF93CFDDE6CF7" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[357,584,1681,1703]" journalOrPublisher="Jacksonville: New World Press" pageId="6" pageNumber="7" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFE4FFBBFE66F93FFD936CF6" box="[461,517,1681,1703]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2015
</bibRefCitation>
: 254, bottom left photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE4FFBBFF3AF940FA046AAC" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="C90D4710FFE4FFBBFF3AF940FEAD6D55" blockId="6.[145,761,1773,1919]" box="[145,315,1773,1797]" pageId="6" pageNumber="7">
<heading id="9245F07CFFE4FFBBFF3AF940FEAD6D55" box="[145,315,1773,1797]" fontSize="10" level="4" pageId="6" pageNumber="7" reason="1">
<emphasis id="FBC69B02FFE4FFBBFF3AF940FEAD6D55" box="[145,315,1773,1797]" italics="true" pageId="6" pageNumber="7">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFF3AF8A0FF616D2F" blockId="6.[145,761,1773,1919]" pageId="6" pageNumber="7">
<materialsCitation id="79DA4D4DFFE4FFBBFF3AF8A0FF616D2F" ID-GBIF-Occurrence="2235579096" collectingDate="2013-03-10" collectionCode="CASIZ" collectorName="T. M. Gosliner." country="Saudi Arabia" county="Red Sea" elevation="8" location="Abulad Island" municipality="Farasan Islands" pageId="6" pageNumber="7" specimenCode="CASIZ 192295" specimenCount="1" typeStatus="holotype">
<typeStatus id="1609F9B2FFE4FFBBFF3AF8A0FF6F6D72" box="[145,249,1805,1826]" pageId="6" pageNumber="7" type="holotype">
<emphasis id="FBC69B02FFE4FFBBFF3AF8A0FF6F6D72" box="[145,249,1805,1826]" italics="true" pageId="6" pageNumber="7">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFE4FFBBFEA7F8A0FE2F6D73" box="[268,441,1805,1827]" collectionCode="CASIZ" pageId="6" pageNumber="7">CASIZ 192295</specimenCode>
, subsampled for molecular study, dissected,
<location id="CC6D11CBFFE4FFBBFEF9F881FE6E6D12" LSID="urn:lsid:plazi:treatment:411BF606FFE4FFB6FEB8FA1CFA326F3A:CC6D11CBFFE4FFBBFEF9F881FE6E6D12" box="[338,504,1836,1858]" country="Saudi Arabia" county="Red Sea" municipality="Farasan Islands" name="Abulad Island" pageId="6" pageNumber="7">Abulad Island</location>
,
<collectingMunicipality id="2969DD6AFFE4FFBBFDAFF881FD566D12" box="[516,704,1836,1858]" pageId="6" pageNumber="7">Farasan Islands</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFE4FFBBFD67F881FF286D30" pageId="6" pageNumber="7">Red Sea</collectingCounty>
,
<collectingCountry id="B1A50780FFE4FFBBFF60F8E7FEFC6D30" box="[203,362,1866,1888]" name="Saudi Arabia" pageId="6" pageNumber="7">Saudi Arabia</collectingCountry>
,
<elevation id="429FA023FFE4FFBBFEDDF8E6FE356D30" box="[374,419,1867,1888]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="m" value="8.0">
<quantity id="0E4AEAF5FFE4FFBBFEDDF8E6FE356D30" box="[374,419,1867,1888]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="m" value="8.0">8 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFE4FFBBFE55F8E6FD256D30" box="[510,691,1866,1888]" pageId="6" pageNumber="7" value="2013-03-10">
<date id="BD0C61D0FFE4FFBBFE55F8E6FD256D30" box="[510,691,1866,1888]" pageId="6" pageNumber="7" value="2013-03-10">10 March 2013</date>
</collectingDate>
,
<collectorName id="644722C6FFE4FFBBFD15F8E6FF616D2F" pageId="6" pageNumber="7">T. M. Gosliner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FF6AFC526A8E" blockId="6.[809,1426,198,253]" box="[809,964,198,222]" pageId="6" pageNumber="7">
<heading id="9245F07CFFE4FFBBFC82FF6AFC526A8E" box="[809,964,198,222]" fontSize="10" level="4" pageId="6" pageNumber="7" reason="1">
<emphasis id="FBC69B02FFE4FFBBFC82FF6AFC526A8E" box="[809,964,198,222]" italics="true" pageId="6" pageNumber="7">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FF4AFA046AAC" blockId="6.[809,1426,198,253]" box="[809,1426,231,253]" pageId="6" pageNumber="7">
Abulad Island, Farasan Islands,
<collectingCountry id="B1A50780FFE4FFBBFB39FF4AFAB16AAD" box="[1170,1319,231,253]" name="Saudi Arabia" pageId="6" pageNumber="7">Saudi Arabia</collectingCountry>
, Red Sea.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE4FFBBFC82FE8EFBAF6B09" pageId="6" pageNumber="7" type="distribution">
<paragraph id="C90D4710FFE4FFBBFC82FE8EFBF06B6B" blockId="6.[809,1126,291,346]" box="[809,1126,291,315]" pageId="6" pageNumber="7">
<heading id="9245F07CFFE4FFBBFC82FE8EFBF06B6B" box="[809,1126,291,315]" fontSize="10" level="4" pageId="6" pageNumber="7" reason="1">
<emphasis id="FBC69B02FFE4FFBBFC82FE8EFBF06B6B" box="[809,1126,291,315]" italics="true" pageId="6" pageNumber="7">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FEE9FBAF6B09" blockId="6.[809,1126,291,346]" box="[809,1081,324,346]" pageId="6" pageNumber="7">
Saudi Arabian
<collectingRegion id="0B7689F2FFE4FFBBFC7EFEE9FBA36B09" box="[981,1077,324,346]" country="Egypt" name="Al Bahr al Ahmar" pageId="6" pageNumber="7">Red Sea</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE4FFBBFC82FE2CFBD06BA5" pageId="6" pageNumber="7" type="etymology">
<paragraph id="C90D4710FFE4FFBBFC82FE2CFC3C6BC9" blockId="6.[809,1425,385,501]" box="[809,938,385,409]" pageId="6" pageNumber="7">
<heading id="9245F07CFFE4FFBBFC82FE2CFC3C6BC9" box="[809,938,385,409]" fontSize="10" level="4" pageId="6" pageNumber="7" reason="1">
<emphasis id="FBC69B02FFE4FFBBFC82FE2CFC3C6BC9" box="[809,938,385,409]" italics="true" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FE0CFBD06BA5" blockId="6.[809,1425,385,501]" pageId="6" pageNumber="7">
<taxonomicName id="0EB23C93FFE4FFBBFC82FE0CFBD26BE6" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,1092,417,438]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE4FFBBFC82FE0CFBD26BE6" box="[809,1092,417,438]" italics="true" pageId="6" pageNumber="7">Hypselodoris alburtuqali</emphasis>
</taxonomicName>
is named for the Arabic word for ‘the orange one’, al burtuqali, owing to the orange coloration of this species.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE4FFB6FC82FDB1FDCF68F8" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7" type="description">
<paragraph id="C90D4710FFE4FFBBFC82FDB1FC236863" blockId="6.[809,1425,540,1054]" box="[809,949,540,563]" pageId="6" pageNumber="7">
<heading id="9245F07CFFE4FFBBFC82FDB1FC236863" box="[809,949,540,563]" fontSize="10" level="4" pageId="6" pageNumber="7" reason="1">
<emphasis id="FBC69B02FFE4FFBBFC82FDB1FC236863" box="[809,949,540,563]" italics="true" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FD91FAD66E4E" blockId="6.[809,1425,540,1054]" pageId="6" pageNumber="7">
<emphasis id="FBC69B02FFE4FFBBFC82FD91FBAE6801" box="[809,1080,572,593]" italics="true" pageId="6" pageNumber="7">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFE4FFBBFA82FD91FA1F6801" box="[1321,1417,572,594]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
) moderately large, reaching
<quantity id="0E4AEAF5FFE4FFBBFBD2FDF6FB5A6820" box="[1145,1228,603,624]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="30.0">30 mm</quantity>
in length. Body translucent pink, with about eight continuous or interrupted longitudinal lines on dorsal surface of notum. Anterior and posterior ends of the animal more opaque white than pink. Additional opaque white lines located on sides of animal and on posterior portion of foot. Longitudinal rows of dark brown spots situated between opaque white lines of notum and foot. Wide red–orange marginal band present along entire mantle and foot margins. Eight unipinnate gill branches having a translucent white base and bright red–orange pigment on apical surfaces and outer margin. Bulb of perfoliate rhinophores opaque white with two red–orange transverse bands and bearing about ten densely arranged lamellae. Base of rhinophores translucent white.
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FBEBFB486EA6" blockId="6.[809,1425,1094,1270]" pageId="6" pageNumber="7">
<emphasis id="FBC69B02FFE4FFBBFC82FBEBFC456E0B" box="[809,979,1094,1115]" italics="true" pageId="6" pageNumber="7">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFE4FFBBFBF9FBC8FB386E2A" box="[1106,1198,1125,1147]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
). Glands situated anteriorly and posteriorly, with no glands present in the central lateral regions of body margin. Nine to ten glands on either side of anterior end of the body, with arc of nine glands situated posteriorly.
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82FAB3FAA86D4F" blockId="6.[809,1425,1310,1823]" pageId="6" pageNumber="7">
<emphasis id="FBC69B02FFE4FFBBFC82FAB3FC656F64" box="[809,1011,1310,1332]" italics="true" pageId="6" pageNumber="7">Buccal armature</emphasis>
: Muscular portion of buccal mass about twice length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass (
<figureCitation id="51895B95FFE4FFBBFCD9FAD6FC546FC0" box="[882,962,1403,1425]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
) bearing numerous jaw rodlets (
<figureCitation id="51895B95FFE4FFBBFA99FAD6FA156FC0" box="[1330,1411,1403,1425]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3B</figureCitation>
). Rodlets narrowly ovoid, with single, acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFE4FFBBFB94FA15FB086F9E" box="[1087,1182,1464,1486]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
44 × 26.0.26 (
<figureCitation id="51895B95FFE4FFBBFA9EFA15FA156F9D" box="[1333,1411,1464,1486]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
). Rachidian row of teeth absent (
<figureCitation id="51895B95FFE4FFBBFB0DFA7AFB686FBC" box="[1190,1278,1495,1517]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3D</figureCitation>
). Innermost lateral teeth having one or two large, triangular denticles on inner side of bifid primary cusp, with another one to two outer denticles. Next several laterals lacking inner triangular denticle but possessing two or three denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFE4FFBBFC47F922FBAD6CF4" box="[1004,1083,1679,1701]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
) all lacking inner denticles but having three or four sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and shorter tooth shape, with three or four rounded outer denticles (
<figureCitation id="51895B95FFE4FFBBFC31F8A4FC7E6D4E" box="[922,1000,1801,1823]" captionStart="Figure 3" captionStartId="9.[159,237,1815,1837]" captionTargetBox="[223,1375,459,1775]" captionTargetId="figure@9.[223,1375,459,1775]" captionTargetPageId="9" captionText="Figure 3. Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth." httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="6" pageNumber="7">Fig. 3F</figureCitation>
), often larger than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFE4FFBBFC82F8E5FA066D2D" blockId="6.[809,1425,1864,1917]" pageId="6" pageNumber="7">
<emphasis id="FBC69B02FFE4FFBBFC82F8E5FB946D0E" box="[809,1026,1864,1886]" italics="true" pageId="6" pageNumber="7">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFE4FFBBFA9FF8E5FA076D0E" box="[1332,1425,1864,1886]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFE4FFBBFC15F8CAFB9D6D2C" box="[958,1035,1895,1917]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
). Ampulla thick, tubular, narrowing
</paragraph>
<caption id="9DCD1798FFE5FFBAFF0AF9FAFB016D07" httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="7" pageNumber="8" startId="7.[161,241,1623,1645]" targetBox="[225,1377,195,1582]" targetPageId="7">
<paragraph id="C90D4710FFE5FFBAFF0AF9FAFB016D07" blockId="7.[161,1443,1622,1879]" pageId="7" pageNumber="8">
<emphasis id="FBC69B02FFE5FFBAFF0AF9FAFE9A6C3C" bold="true" box="[161,268,1623,1645]" pageId="7" pageNumber="8">Figure 1.</emphasis>
Living animals. A,
<taxonomicName id="0EB23C93FFE5FFBAFE49F9FBFC7E6C3D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[482,1000,1622,1645]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="alburtuqali" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFE49F9FBFC9B6C3C" bold="true" box="[482,781,1622,1644]" italics="true" pageId="7" pageNumber="8">Hypselodoris alburtuqali</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFC44F9FAFBDE6C3C" bold="true" box="[1007,1096,1623,1644]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFC44F9FAFBDE6C3C" box="[1007,1096,1623,1644]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 192295, holotype, Saudi Arabia. B,
<taxonomicName id="0EB23C93FFE5FFBAFEB3F9D9FD7F6CDA" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[280,745,1652,1674]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="brycei" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFEB3F9D9FD906CDA" bold="true" box="[280,518,1652,1674]" italics="true" pageId="7" pageNumber="8">Hypselodoris brycei</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFD5AF9D9FCDB6CD9" bold="true" box="[753,845,1652,1673]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFD5AF9D9FCDB6CD9" box="[753,845,1652,1673]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, WAM
<accessionNumber id="D6E1DAF3FFE5FFBAFC35F9D9FC676CDA" box="[926,1009,1652,1674]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S96082" pageId="7" pageNumber="8" type="EnaNcbi">S96082</accessionNumber>
, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C,
<taxonomicName id="0EB23C93FFE5FFBAFDDEF93CFBD46CF7" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[629,1090,1681,1703]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="cerisae" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFDDEF93CFCF16CF6" bold="true" box="[629,871,1681,1703]" italics="true" pageId="7" pageNumber="8">Hypselodoris cerisae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFBE2F93FFB376CF7" bold="true" box="[1097,1185,1682,1703]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFBE2F93FFB376CF7" box="[1097,1185,1682,1703]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, specimen from Hachijo Island, Japan, Nishina Masayoshi. D,
<taxonomicName id="0EB23C93FFE5FFBAFD91F903FB846C95" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[570,1042,1710,1733]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="confetti" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFD91F903FCA06C94" bold="true" box="[570,822,1710,1732]" italics="true" pageId="7" pageNumber="8">Hypselodoris confetti</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFBB2F902FBE46C94" bold="true" box="[1049,1138,1711,1732]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFBB2F902FBE46C94" box="[1049,1138,1711,1732]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E,
<taxonomicName id="0EB23C93FFE5FFBAFDE1F961FBC96CB1" authority="(Gohar &amp; Abu-Ela, 1957)" authorityName="Gohar &amp; Abu-Ela" authorityYear="1957" baseAuthorityName="Gohar &amp; Abu-Ela" baseAuthorityYear="1957" box="[586,1119,1739,1762]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFE5FFBAFDE1F961FCC66CB1" box="[586,848,1740,1761]" italics="true" pageId="7" pageNumber="8">Hypseldoris ghardaqana</emphasis>
(Gohar &amp; Abu-Ela, 1957)
</taxonomicName>
, CASIZ 192282, Saudi Arabia. F,
<taxonomicName id="0EB23C93FFE5FFBAFF11F944FD096CAF" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[186,671,1769,1791]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="juniperae" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFF11F944FE5F6CAE" bold="true" box="[186,457,1769,1791]" italics="true" pageId="7" pageNumber="8">Hypselodoris juniperae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFD0DF947FD6B6CAF" bold="true" box="[678,765,1770,1791]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFD0DF947FD6B6CAF" box="[678,765,1770,1791]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 177550, Radama Islands, Madagascar. G,
<taxonomicName id="0EB23C93FFE5FFBAFAA0F944FD956D4D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFAA0F944FEB36D4C" bold="true" italics="true" pageId="7" pageNumber="8">Hypselodoris katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFDA1F8AAFDF56D4C" bold="true" box="[522,611,1799,1820]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFDA1F8AAFDF56D4C" box="[522,611,1799,1820]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 176773, eastern Peninsular Malaysia. H,
<taxonomicName id="0EB23C93FFE5FFBAFBD5F8AAFF6C6D6A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFBD5F8AAFAB26D4C" bold="true" box="[1150,1316,1798,1820]" italics="true" pageId="7" pageNumber="8">H. katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFEAAF889FECC6D69" bold="true" box="[257,346,1828,1849]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFEAAF889FECC6D69" box="[257,346,1828,1849]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 177532, Batangas, Luzon, Philippines. I,
<taxonomicName id="0EB23C93FFE5FFBAFCC6F889FB786D6A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[877,1262,1828,1850]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFE5FFBAFCC6F889FB846D6A" bold="true" box="[877,1042,1828,1850]" italics="true" pageId="7" pageNumber="8">H. katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE5FFBAFB5EF889FAD86D69" bold="true" box="[1269,1358,1828,1849]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="E0F52679FFE5FFBAFB5EF889FAD86D69" box="[1269,1358,1828,1849]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated.
</paragraph>
</caption>
<caption id="9DCD1798FFEAFFB5FF3BF95AFA076D34" httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="8" pageNumber="9" startId="8.[144,228,1783,1805]" targetBox="[146,1425,197,1740]" targetPageId="8">
<paragraph id="C90D4710FFEAFFB5FF3BF95AFA076D34" blockId="8.[144,1425,1783,1893]" pageId="8" pageNumber="9">
<emphasis id="FBC69B02FFEAFFB5FF3BF95AFE946D5C" bold="true" box="[144,258,1783,1805]" pageId="8" pageNumber="9">Figure 2.</emphasis>
Mantle glands. A,
<taxonomicName id="0EB23C93FFEAFFB5FE75F95AFB926D5D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[478,1028,1783,1805]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="alburtuqali" status="sp. nov.">
<emphasis id="FBC69B02FFEAFFB5FE75F95AFC8C6D5D" bold="true" box="[478,794,1783,1805]" italics="true" pageId="8" pageNumber="9">Hypselodoris alburtuqali</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFEAFFB5FBA7F95AFBFC6D5C" bold="true" box="[1036,1130,1783,1804]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="E0F52679FFEAFFB5FBA7F95AFBFC6D5C" box="[1036,1130,1783,1804]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 192295, holotype. B,
<taxonomicName id="0EB23C93FFEAFFB5FF05F8B9FD1E6D7A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[174,648,1812,1834]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="cerisae" status="sp. nov.">
<emphasis id="FBC69B02FFEAFFB5FF05F8B9FE316D79" bold="true" box="[174,423,1812,1834]" italics="true" pageId="8" pageNumber="9">Hypselodoris cerisae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFEAFFB5FD24F8B9FD7C6D79" bold="true" box="[655,746,1812,1833]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="E0F52679FFEAFFB5FD24F8B9FD7C6D79" box="[655,746,1812,1833]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, CASIZ 178350. C,
<taxonomicName id="0EB23C93FFEAFFB5FB80F8B9FE976D18" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="confetti" status="sp. nov.">
<emphasis id="FBC69B02FFEAFFB5FB80F8B9FABA6D7A" bold="true" box="[1067,1324,1812,1834]" italics="true" pageId="8" pageNumber="9">Hypselodoris confetti</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFEAFFB5FEA3F89FFEC96D17" bold="true" box="[264,351,1842,1863]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="E0F52679FFEAFFB5FEA3F89FFEC96D17" box="[264,351,1842,1863]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 191070. D,
<taxonomicName id="0EB23C93FFEAFFB5FD85F89FFC246D18" authority="(Risbec, 1928)" authorityName="Risbec" authorityYear="1928" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[558,946,1842,1864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFEAFFB5FD85F89FFC816D17" box="[558,791,1842,1863]" italics="true" pageId="8" pageNumber="9">Hypselodoris decorata</emphasis>
(Risbec, 1928)
</taxonomicName>
, CASIZ 184316. E,
<taxonomicName id="0EB23C93FFEAFFB5FB2BF89FFE326D34" authority="(Gohar &amp; Abu-Ela, 1957)" authorityName="Gohar &amp; Abu-Ela" authorityYear="1957" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFEAFFB5FB2BF89FFA076D17" box="[1152,1425,1842,1863]" italics="true" pageId="8" pageNumber="9">Hypselodoris ghardaqana</emphasis>
(Gohar &amp; Abu-Ela, 1957)
</taxonomicName>
, CASIZ 192282. F,
<taxonomicName id="0EB23C93FFEAFFB5FDDEF8E2FBB76D35" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[629,1057,1871,1893]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFEAFFB5FDDEF8E2FCD66D35" bold="true" box="[629,832,1871,1893]" italics="true" pageId="8" pageNumber="9">Hypselodoris iba</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFEAFFB5FB82F8E2FB126D34" bold="true" box="[1065,1156,1871,1892]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="E0F52679FFEAFFB5FB82F8E2FB126D34" box="[1065,1156,1871,1892]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NMP 0 41279.
</paragraph>
</caption>
<paragraph id="C90D4710FFEBFFB4FF08FF68FDFA6BDB" blockId="9.[163,1444,196,395]" pageId="9" pageNumber="10">
G,
<taxonomicName id="0EB23C93FFEBFFB4FF16FF69FD0F6A8B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[189,665,196,219]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="juniperae" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FF16FF69FE5F6A8A" bold="true" box="[189,457,196,218]" italics="true" pageId="9" pageNumber="10">Hypselodoris juniperae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FD36FF68FD676A8A" box="[669,753,197,218]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FD36FF68FD676A8A" bold="true" box="[669,753,197,218]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ 177550.
<materialsCitation id="79DA4D4DFFEBFFB4FC30FF68FE356AA8" ID-GBIF-Occurrence="2235579082" collectionCode="CASIZ" pageId="9" pageNumber="10" specimenCode="CASIZ 177532" specimenCount="1">
H,
<taxonomicName id="0EB23C93FFEBFFB4FC13FF69FA356A8B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[952,1443,196,219]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FC13FF69FB456A8A" bold="true" box="[952,1235,196,218]" italics="true" pageId="9" pageNumber="10">Hypselodoris katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FF08FF4FFF6C6AA7" box="[163,250,226,247]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FF08FF4FFF6C6AA7" bold="true" box="[163,250,226,247]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<specimenCode id="9914EF6BFFEBFFB4FEACFF4FFE356AA8" box="[263,419,226,248]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 177532</specimenCode>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEBFFB4FE06FF4FFB196AA8" ID-GBIF-Occurrence="2235579071" box="[429,1167,226,248]" collectionCode="CASIZ" pageId="9" pageNumber="10" specimenCode="CASIZ 206801" specimenCount="1">
I,
<taxonomicName id="0EB23C93FFEBFFB4FE69FF4FFC7F6AA7" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[450,1001,226,248]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEBFFB4FE69FF4FFD596AA8" bold="true" box="[450,719,226,248]" italics="true" pageId="9" pageNumber="10">Hypselodoris krakatoa</emphasis>
<bibRefCitation id="AD233AE1FFEBFFB4FD7DFF4FFC7F6AA7" author="Gosliner T &amp; Johnson, R." box="[726,1001,226,248]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="9" pageNumber="10" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
,
<specimenCode id="9914EF6BFFEBFFB4FC58FF4FFB196AA8" box="[1011,1167,226,248]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 206801</specimenCode>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEBFFB4FB32FF4FFD796B45" ID-GBIF-Occurrence="2235579141" collectionCode="CASIZ" pageId="9" pageNumber="10" specimenCode="CASIZ 208652" specimenCount="1" typeStatus="paratype">
J,
<taxonomicName id="0EB23C93FFEBFFB4FB1AFF4FFEEE6B45" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FB1AFF4FFA356AA8" bold="true" box="[1201,1443,226,248]" italics="true" pageId="9" pageNumber="10">Hypselodoris lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FED4FEADFE4E6B45" box="[383,472,256,277]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FED4FEADFE4E6B45" bold="true" box="[383,472,256,277]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFEBFFB4FE4FFEADFDD36B45" box="[484,581,256,277]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
,
<specimenCode id="9914EF6BFFEBFFB4FDFAFF52FD796B45" box="[593,751,255,277]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 208652</specimenCode>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEBFFB4FD52FEADFEE06B63" ID-GBIF-Occurrence="2235579127" collectionCode="CASIZ" pageId="9" pageNumber="10" specimenCode="CASIZ 191464" specimenCount="1" typeStatus="holotype">
K,
<taxonomicName id="0EB23C93FFEBFFB4FCB3FF52FA946B45" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[792,1282,255,277]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FCB3FF52FBB06B45" bold="true" box="[792,1062,255,277]" italics="true" pageId="9" pageNumber="10">Hypselodoris paradisa</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FAA2FEADFAF46B45" box="[1289,1378,256,277]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FAA2FEADFAF46B45" bold="true" box="[1289,1378,256,277]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFEBFFB4FAC5FF52FF446B62" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
,
<specimenCode id="9914EF6BFFEBFFB4FF77FEB0FEE06B63" box="[220,374,285,307]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 191464</specimenCode>
</materialsCitation>
. L,
<taxonomicName id="0EB23C93FFEBFFB4FE30FEB1FCD16B63" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[411,839,284,307]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="perii" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FE30FEB1FDE66B62" bold="true" box="[411,624,284,306]" italics="true" pageId="9" pageNumber="10">Hypselodoris perii</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FCE6FEB0FC326B62" box="[845,932,285,306]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FCE6FEB0FC326B62" bold="true" box="[845,932,285,306]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, NMP
<materialsCitation id="79DA4D4DFFEBFFB4FC45FEB0FDF36B1F" ID-GBIF-Occurrence="2235579132" collectionCode="CASIZ" pageId="9" pageNumber="10" specimenCode="CASIZ 204801" specimenCount="1" typeStatus="paratype">
0 41281. M,
<taxonomicName id="0EB23C93FFEBFFB4FBCCFEB1FF6C6B00" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FBCCFEB1FABC6B62" bold="true" box="[1127,1322,284,306]" italics="true" pageId="9" pageNumber="10">Hypselodoris roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FEABFE97FEC06B1F" box="[256,342,314,335]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FEABFE97FEC06B1F" bold="true" box="[256,342,314,335]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFEBFFB4FEC9FE97FE576B1F" box="[354,449,314,335]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
,
<specimenCode id="9914EF6BFFEBFFB4FE60FE97FDF36B1F" box="[459,613,314,335]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 204801</specimenCode>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEBFFB4FDC5FE97FADD6B1F" ID-GBIF-Occurrence="2235579111" box="[622,1355,314,336]" collectionCode="NMP" pageId="9" pageNumber="10" specimenCount="1" typeStatus="holotype">
N,
<taxonomicName id="0EB23C93FFEBFFB4FD20FE97FBDE6B00" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[651,1096,314,336]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FD20FE97FCE46B1F" bold="true" box="[651,882,314,336]" italics="true" pageId="9" pageNumber="10">Hypselodoris rositoi</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FBE5FE97FB326B1F" box="[1102,1188,314,335]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FBE5FE97FB326B1F" bold="true" box="[1102,1188,314,335]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFEBFFB4FB1BFE97FA9C6B00" box="[1200,1290,314,336]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
,
<collectionCode id="AFA3DFD5FFEBFFB4FABFFE97FADD6B1F" box="[1300,1355,314,335]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="9" pageNumber="10">NMP</collectionCode>
</materialsCitation>
<materialsCitation id="79DA4D4DFFEBFFB4FAF9FE97FDFE6BDB" ID-GBIF-Occurrence="2235579102" collectionCode="NMP" pageId="9" pageNumber="10" specimenCode="CASIZ 200649, CASIZ 177305, CASIZ 200552, CASIZ 217424" specimenCount="1" typeStatus="holotype">
0 41283. O,
<taxonomicName id="0EB23C93FFEBFFB4FF6BFEFAFD136B3D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[192,645,343,365]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FF6BFEFAFE3B6B3D" bold="true" box="[192,429,343,365]" italics="true" pageId="9" pageNumber="10">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEBFFB4FD20FEF5FD756B3D" box="[651,739,344,365]" pageId="9" pageNumber="10" rank="species">
<emphasis id="FBC69B02FFEBFFB4FD20FEF5FD756B3D" bold="true" box="[651,739,344,365]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, composite of five specimens (
<specimenCode id="9914EF6BFFEBFFB4FB8BFEFAFB2A6B3D" box="[1056,1212,343,365]" collectionCode="CASIZ" pageId="9" pageNumber="10">
CASIZ
<date id="BD0C61D0FFEBFFB4FBC5FEFAFB2A6B3D" box="[1134,1212,343,365]" pageId="9" pageNumber="10" value="49-06-20">200649</date>
</specimenCode>
,
<specimenCode id="9914EF6BFFEBFFB4FB6DFEFAFAF76B3D" box="[1222,1377,343,365]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 177305</specimenCode>
,
<collectionCode id="AFA3DFD5FFEBFFB4FAC7FEF5FA356B3D" box="[1388,1443,344,365]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="9" pageNumber="10">NMP</collectionCode>
0 41284,
<specimenCode id="9914EF6BFFEBFFB4FF51FED8FE046BDA" box="[250,402,372,394]" collectionCode="CASIZ" pageId="9" pageNumber="10">
CASIZ
<date id="BD0C61D0FFEBFFB4FEEDFED9FE046BDA" box="[326,402,372,394]" pageId="9" pageNumber="10" value="52-05-20">200552</date>
</specimenCode>
and
<specimenCode id="9914EF6BFFEBFFB4FE6DFED8FDF46BDB" box="[454,610,373,395]" collectionCode="CASIZ" pageId="9" pageNumber="10">CASIZ 217424</specimenCode>
)
</materialsCitation>
.
</paragraph>
<caption id="9DCD1798FFEBFFB4FF34F8BAFB286D1B" httpUri="https://zenodo.org/record/2632777/files/figure.png" pageId="9" pageNumber="10" startId="9.[159,237,1815,1837]" targetBox="[223,1375,459,1775]" targetPageId="9">
<paragraph id="C90D4710FFEBFFB4FF34F8BAFB286D1B" blockId="9.[159,1440,1815,1867]" pageId="9" pageNumber="10">
<emphasis id="FBC69B02FFEBFFB4FF34F8BAFE916D7D" bold="true" box="[159,263,1815,1837]" pageId="9" pageNumber="10">Figure 3.</emphasis>
<taxonomicName id="0EB23C93FFEBFFB4FEB8F8BAFC986D7E" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[275,782,1815,1838]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="alburtuqali" status="sp. nov.">
<emphasis id="FBC69B02FFEBFFB4FEB8F8BAFDAE6D7D" bold="true" box="[275,568,1815,1837]" italics="true" pageId="9" pageNumber="10">Hypselodoris alburtuqali</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFEBFFB4FCBEF8B5FCFD6D7D" bold="true" box="[789,875,1816,1837]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="E0F52679FFEBFFB4FCBEF8B5FCFD6D7D" box="[789,875,1816,1837]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, CASIZ 192295, holotype, buccal armature. A, portion of jaw. B, jaw rodlets. C, entire radula. D, innermost teeth. E, middle lateral teeth. F, outermost teeth.
</paragraph>
</caption>
<caption id="9DCD1798FFE8FFB7FF39F9F0FA056D10" httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="10" pageNumber="11" startId="10.[146,224,1629,1651]" targetBox="[244,1328,198,1587]" targetPageId="10">
<paragraph id="C90D4710FFE8FFB7FF39F9F0FA056D10" blockId="10.[146,1427,1629,1856]" pageId="10" pageNumber="11">
<emphasis id="FBC69B02FFE8FFB7FF39F9F0FF6C6C22" bold="true" box="[146,250,1629,1651]" pageId="10" pageNumber="11">Figure 4.</emphasis>
Reproductive systems A,
<taxonomicName id="0EB23C93FFE8FFB7FDA1F9F0FB956C23" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[522,1027,1629,1651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="alburtuqali" status="sp. nov.">
<emphasis id="FBC69B02FFE8FFB7FDA1F9F0FCB86C23" bold="true" box="[522,814,1629,1651]" italics="true" pageId="10" pageNumber="11">Hypselodoris alburtuqali</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE8FFB7FBA2F9F0FBC96C22" bold="true" box="[1033,1119,1629,1650]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0F52679FFE8FFB7FBA2F9F0FBC96C22" box="[1033,1119,1629,1650]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, CASIZ19229, scale bar: 0.5 mm. B,
<taxonomicName id="0EB23C93FFE8FFB7FE9CF9D7FD216CC1" authority="(Pease, 1871)" authorityName="Pease" authorityYear="1871" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[311,695,1658,1681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE8FFB7FE9CF9D7FDB16CDF" box="[311,551,1658,1679]" italics="true" pageId="10" pageNumber="11">Hypselodoris maculosa</emphasis>
(Pease, 1871)
</taxonomicName>
, CASIZ 139595, scale bar: 0.75 mm. C,
<taxonomicName id="0EB23C93FFE8FFB7FBFAF9D7FE926CFE" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="brycei" status="sp. nov.">
<emphasis id="FBC69B02FFE8FFB7FBFAF9D7FAA46CC0" bold="true" box="[1105,1330,1658,1680]" italics="true" pageId="10" pageNumber="11">Hypselodoris brycei</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE8FFB7FEA1F935FEF46CFD" bold="true" box="[266,354,1688,1709]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0F52679FFE8FFB7FEA1F935FEF46CFD" box="[266,354,1688,1709]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, WAM
<accessionNumber id="D6E1DAF3FFE8FFB7FDB0F935FDFC6CFE" box="[539,618,1688,1710]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S12628" pageId="10" pageNumber="11" type="EnaNcbi">S12628</accessionNumber>
, scale bar: 3.0 mm. D,
<taxonomicName id="0EB23C93FFE8FFB7FCF1F935FB7E6CFE" authority="(Yonow, 2001)" authorityName="Yonow" authorityYear="2001" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[858,1256,1688,1710]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFE8FFB7FCF1F935FBD96CFD" box="[858,1103,1688,1709]" italics="true" pageId="10" pageNumber="11">Hypselodoris apolegma</emphasis>
(Yonow, 2001)
</taxonomicName>
, CASIZ 0 83743, scale bar: 2.0 mm. E,
<taxonomicName id="0EB23C93FFE8FFB7FED1F918FCDD6C9B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[378,843,1717,1739]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cerisae" status="sp. nov.">
<emphasis id="FBC69B02FFE8FFB7FED1F918FDF96C9A" bold="true" box="[378,623,1717,1739]" italics="true" pageId="10" pageNumber="11">Hypselodoris cerisae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE8FFB7FCF9F918FC3D6C9A" bold="true" box="[850,939,1717,1738]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0F52679FFE8FFB7FCF9F918FC3D6C9A" box="[850,939,1717,1738]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype CASIZ 178350, scale bar: 0.85 mm. F,
<taxonomicName id="0EB23C93FFE8FFB7FF02F97FFD086CB8" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[169,670,1746,1768]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFE8FFB7FF02F97FFE076CB7" box="[169,401,1746,1767]" italics="true" pageId="10" pageNumber="11">Hypselodoris krakatoa</emphasis>
Gosliner &amp; Johnson, 1999
</taxonomicName>
, CASIZ 206801, Philippines, scale bar: 1.0 mm. G,
<taxonomicName id="0EB23C93FFE8FFB7FB0BF97FFEF46D56" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="confetti" status="sp. nov.">
<emphasis id="FBC69B02FFE8FFB7FB0BF97FFA046CB8" bold="true" box="[1184,1426,1746,1768]" italics="true" pageId="10" pageNumber="11">Hypselodoris confetti</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<emphasis id="FBC69B02FFE8FFB7FEC3F95DFE296D55" bold="true" box="[360,447,1776,1797]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0F52679FFE8FFB7FEC3F95DFE296D55" box="[360,447,1776,1797]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, CASIZ 191070, scale bar: 0.60 mm. H,
<taxonomicName id="0EB23C93FFE8FFB7FC69F95DFAD46D56" authority="(Risbec, 1928)" authorityName="Risbec" authorityYear="1928" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[962,1346,1776,1798]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE8FFB7FC69F95DFB3E6D55" box="[962,1192,1776,1797]" italics="true" pageId="10" pageNumber="11">Hypselodoris decorata</emphasis>
(Risbec, 1928)
</taxonomicName>
, CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland.
</paragraph>
</caption>
<paragraph id="C90D4710FFE9FFB6FF08FF6BFDCF68F8" blockId="11.[163,779,198,680]" pageId="11" pageNumber="12">somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing slightly as it transitions into muscular, ejaculatory portion. Ejaculatory portion widening again before entry into wider penial bulb. Penial bulb adjacent to straight, very wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively short vagina leading to minute receptaculum seminis and larger spherical, thinwalled receptaculum seminis. Moderately long uterine duct emerging from vagina close to base of bursa and female gland mass, near albumen gland.</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFE9FFB6FF08FD62FA326F3A" pageId="11" pageNumber="12" type="discussion">
<paragraph id="C90D4710FFE9FFB6FF08FD62FE8668B7" blockId="11.[163,779,719,1878]" box="[163,272,719,743]" pageId="11" pageNumber="12">
<heading id="9245F07CFFE9FFB6FF08FD62FE8668B7" box="[163,272,719,743]" fontSize="10" level="4" pageId="11" pageNumber="12" reason="1">
<emphasis id="FBC69B02FFE9FFB6FF08FD62FE8668B7" box="[163,272,719,743]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFE9FFB6FF08FD5DFE2A6D05" blockId="11.[163,779,719,1878]" pageId="11" pageNumber="12">
The colour pattern of
<taxonomicName id="0EB23C93FFE9FFB6FE1CFD5DFDFD6955" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[439,619,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FE1CFD5DFDFD6955" box="[439,619,752,773]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
most closely resembles that of
<taxonomicName id="0EB23C93FFE9FFB6FE2EFCA3FDA86973" authorityName="Pease" authorityYear="1866" box="[389,574,782,803]" class="Gastropoda" family="Chromodorididae" genus="Goniobranchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFE9FFB6FE2EFCA3FDA86973" box="[389,574,782,803]" italics="true" pageId="11" pageNumber="12">Goniobranchus</emphasis>
</taxonomicName>
sp. 29 (
<bibRefCitation id="AD233AE1FFE9FFB6FD08FCA3FEB66913" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" journalOrPublisher="Jacksonville: New World Press" pageId="11" pageNumber="12" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFE9FFB6FF08FC83FF4A6912" box="[163,220,813,835]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
, 2015
</bibRefCitation>
: 227 upper left photograph), with which it is sympatric. Our molecular phylogeny (
<figureCitation id="51895B95FFE9FFB6FDCFFCE1FD266932" box="[612,688,844,866]" captionStart="Figure 35" captionStartId="56.[146,226,1818,1840]" captionTargetBox="[210,1362,196,1775]" captionTargetId="figure@56.[209,1364,195,1777]" captionTargetPageId="56" captionText="Figure 35. Bayesian inference and maximum likelihood tree of Hypselodoris, indicating support values for each node. Bootstrap values are on the left and Bayesian posterior probablilities on the right." httpUri="https://zenodo.org/record/2632866/files/figure.png" pageId="11" pageNumber="12">Fig. 35</figureCitation>
) clearly indicates that this species is in the
<taxonomicName id="0EB23C93FFE9FFB6FD88FCC6FD38692F" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[547,686,874,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FD88FCC6FD38692F" box="[547,686,874,896]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
clade. It is sister to the clade that includes
<taxonomicName id="0EB23C93FFE9FFB6FD82FC27FF7569EE" authority="(Risbec, 1928)" baseAuthorityName="Risbec" baseAuthorityYear="1928" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FD82FC27FD3A69CE" box="[553,684,905,927]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
(
<bibRefCitation id="AD233AE1FFE9FFB6FD11FC24FF4A69EE" author="Risbec J." journalOrPublisher="Faune des Colonies Francaises" pageId="11" pageNumber="12" pagination="1 - 328" part="2" refId="ref44543" refString="Risbec J. 1928. Contribution a l'etude des nudibranches Neo- Caledoniens. Faune des Colonies Francaises 2: 1 - 328." title="Contribution a l'etude des nudibranches Neo- Caledoniens" type="journal article" year="1928">Risbec, 1928</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FF44FC05FEED69ED" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[239,379,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="juniperae" status="sp. nov.">
<emphasis id="FBC69B02FFE9FFB6FF44FC05FEED69ED" box="[239,379,936,957]" italics="true" pageId="11" pageNumber="12">H. juniperae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFE9FFB6FE29FC05FE4169ED" box="[386,471,936,957]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="0EB23C93FFE9FFB6FE49FC05FC9D69EE" authority="(Pease, 1871)" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[482,779,936,958]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FE49FC05FDF969ED" box="[482,623,936,957]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
(
<bibRefCitation id="AD233AE1FFE9FFB6FDD6FC05FC9569EE" author="Pease WH" box="[637,771,936,958]" journalOrPublisher="American Journal of Conchology" pageId="11" pageNumber="12" pagination="11 - 19" part="7" refId="ref44179" refString="Pease WH. 1871. Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2. American Journal of Conchology, 7: 11 - 19." title="Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2." type="journal article" year="1871">Pease, 1871</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FF79FC6AFEB9698C" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[210,303,967,988]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae" status="sp. nov.">
<emphasis id="FBC69B02FFE9FFB6FF79FC6AFEB9698C" box="[210,303,967,988]" italics="true" pageId="11" pageNumber="12">H.yarae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFE9FFB6FE99FC6AFE14698C" box="[306,386,967,988]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
and is shown to represent a distinct species in our ABGD analysis. The colour pattern most closely resembles that of
<taxonomicName id="0EB23C93FFE9FFB6FE14FBA9FDD76E49" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[447,577,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FE14FBA9FDD76E49" box="[447,577,1028,1049]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FDD2FBA9FD4E6E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[633,728,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FDD2FBA9FD4E6E49" box="[633,728,1028,1049]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
, but
<taxonomicName id="0EB23C93FFE9FFB6FF08FB8EFEC76E67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,337,1058,1080]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FF08FB8EFEC76E67" box="[163,337,1058,1080]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
has a narrower red–orange margin, more longitudinal opaque white lines and more, smaller brown spots compared with
<taxonomicName id="0EB23C93FFE9FFB6FE4DFBCDFDFC6E25" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[486,618,1120,1141]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FE4DFBCDFDFC6E25" box="[486,618,1120,1141]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FD0FFBCDFC936E25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[676,773,1120,1141]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FD0FFBCDFC936E25" box="[676,773,1120,1141]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName id="0EB23C93FFE9FFB6FE96FBD2FE546EC3" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[317,450,1150,1172]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FE96FBD2FE546EC3" box="[317,450,1150,1172]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
has opaque white spots and purple markings on its anterior and posterior margins of the notum that are absent in
<taxonomicName id="0EB23C93FFE9FFB6FD82FB11FD406E81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[553,726,1212,1233]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FD82FB11FD406E81" box="[553,726,1212,1233]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FF08FB76FE916EA0" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[163,263,1243,1264]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FF08FB76FE916EA0" box="[163,263,1243,1264]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
. All of the members of this clade, including those species that were studied by
<bibRefCitation id="AD233AE1FFE9FFB6FD80FB54FF7A6F7E" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson (1999)</bibRefCitation>
and are not included in the present molecular phylogeny, have distinctive colour patterns on their rhinophores.
<taxonomicName id="0EB23C93FFE9FFB6FE95FAF8FDF36F3A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[318,613,1365,1386]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FE95FAF8FDF36F3A" box="[318,613,1365,1386]" italics="true" pageId="11" pageNumber="12">Hypselodoris alburtuqali</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FDD8FAFBFC936F3A" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[627,773,1365,1387]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FDD8FAFBFC936F3A" box="[627,773,1365,1387]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
,
<emphasis id="FBC69B02FFE9FFB6FF08FAD9FE9D6FD9" box="[163,267,1396,1417]" italics="true" pageId="11" pageNumber="12">H. peasei</emphasis>
,
<taxonomicName id="0EB23C93FFE9FFB6FEB3FAD9FF4A6FF8" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="insulana">
<emphasis id="FBC69B02FFE9FFB6FEB3FAD9FD8C6FD9" box="[280,538,1396,1417]" italics="true" pageId="11" pageNumber="12">Hypselodoris insulana</emphasis>
<bibRefCitation id="AD233AE1FFE9FFB6FD8AFAD9FF4A6FF8" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FF4EFA3FFF4D6F96" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alboterminata">
<emphasis id="FBC69B02FFE9FFB6FF4EFA3FFD896FF7" box="[229,543,1426,1447]" italics="true" pageId="11" pageNumber="12">Hypselodoris alboterminata</emphasis>
<bibRefCitation id="AD233AE1FFE9FFB6FD8DFA3FFF4D6F96" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FEBDFA1FFE316F97" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[278,423,1458,1479]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFE9FFB6FEBDFA1FFE316F97" box="[278,423,1458,1479]" italics="true" pageId="11" pageNumber="12">H. juniperae</emphasis>
</taxonomicName>
have two reddish rhinophoral rings. In contrast, the rhinophores of
<taxonomicName id="0EB23C93FFE9FFB6FDDBFA7DFD3A6C54" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="violabranchia">
<emphasis id="FBC69B02FFE9FFB6FDDBFA7DFEC56C53" italics="true" pageId="11" pageNumber="12">Hypselodoris violabranchia</emphasis>
<bibRefCitation id="AD233AE1FFE9FFB6FEF5FA43FD3A6C54" author="Gosliner T &amp; Johnson, R." box="[350,684,1518,1540]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
have
<bibRefCitation id="AD233AE1FFE9FFB6FD56FA42FC9D6C54" author="Gosliner T &amp; Johnson, R." box="[765,779,1519,1540]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">a</bibRefCitation>
reddish base and a violet apex. The rhinophores of
<taxonomicName id="0EB23C93FFE9FFB6FF08F981FEBF6C11" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[163,297,1580,1601]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FF08F981FEBF6C11" box="[163,297,1580,1601]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FECFF981FE516C11" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[356,455,1580,1601]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FECFF981FE516C11" box="[356,455,1580,1601]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
have three red rhinophoral rings, and those of
<taxonomicName id="0EB23C93FFE9FFB6FEDDF9E7FF4B6C2E" authority="Rudman, 1977" authorityName="Rudman" authorityYear="1977" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maridadilus">
<emphasis id="FBC69B02FFE9FFB6FEDDF9E7FD0B6C0F" box="[374,669,1610,1631]" italics="true" pageId="11" pageNumber="12">Hypselodoris maridadilus</emphasis>
<bibRefCitation id="AD233AE1FFE9FFB6FD08F9E7FF4B6C2E" author="Rudman WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FF43F9C7FDFE6C2F" authority="(Adams &amp; Reeve, 1850)" baseAuthorityName="Adams &amp; Reeve" baseAuthorityYear="1850" box="[232,616,1641,1663]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="whitei">
<emphasis id="FBC69B02FFE9FFB6FF43F9C7FEC46C2E" box="[232,338,1641,1663]" italics="true" pageId="11" pageNumber="12">H. whitei</emphasis>
(
<bibRefCitation id="AD233AE1FFE9FFB6FECAF9C4FDF66C2F" author="Adams H &amp; Reeve L." box="[353,608,1641,1663]" journalOrPublisher="London: Reeve, Benham and Reeve" pageId="11" pageNumber="12" refId="ref40984" refString="Adams H, Reeve L. 1850. Mollusca, part 3. The zoology of the voyage of the H. M. S. Samarang under the command of Cap. Sir Edward Belcher during the years 1843 - 46. London: Reeve, Benham and Reeve." title="Mollusca, part 3. The zoology of the voyage of the H. M. S. Samarang under the command of Cap. Sir Edward Belcher during the years 1843 - 46" type="book" year="1850">Adams &amp; Reeve, 1850</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FD08F9C7FC9D6C2F" authorityName="Rudman" authorityYear="1977" box="[675,779,1642,1663]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="emma">
<emphasis id="FBC69B02FFE9FFB6FD08F9C7FC9D6C2F" box="[675,779,1642,1663]" italics="true" pageId="11" pageNumber="12">H. emma</emphasis>
</taxonomicName>
all have uniformly red rhinophores with a white apex. Of those species with two red rhinophoral rings,
<taxonomicName id="0EB23C93FFE9FFB6FF08F96BFEB96C8A" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[163,303,1733,1755]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="insulana">
<emphasis id="FBC69B02FFE9FFB6FF08F96BFEB96C8A" box="[163,303,1733,1755]" italics="true" pageId="11" pageNumber="12">H. insulana</emphasis>
</taxonomicName>
and
<emphasis id="FBC69B02FFE9FFB6FEC7F96BFE416C8B" box="[364,471,1734,1755]" italics="true" pageId="11" pageNumber="12">H. peasei</emphasis>
lack dark brownish spots on the notum and have more numerous opaque white lines on the notum, and
<taxonomicName id="0EB23C93FFE9FFB6FE19F8AEFDAB6D48" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[434,573,1795,1816]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFE9FFB6FE19F8AEFDAB6D48" box="[434,573,1795,1816]" italics="true" pageId="11" pageNumber="12">H. juniperae</emphasis>
</taxonomicName>
has purple rather than dark brown spots and opaque white spots rather than lines on the notum.
</paragraph>
<paragraph id="C90D4710FFE9FFB6FCF8FF6BFAC168F8" blockId="11.[827,1444,198,1386]" pageId="11" pageNumber="12">
<taxonomicName id="0EB23C93FFE9FFB6FCF8FF6BFBFD6A8B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[851,1131,198,219]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FCF8FF6BFBFD6A8B" box="[851,1131,198,219]" italics="true" pageId="11" pageNumber="12">Hypselodoris alburtuqali</emphasis>
</taxonomicName>
has mantle glands that are distributed completely around the anterior and posterior margins of the mantle, with no glands present along most of the lateral notal margins. Of the other members of this clade, only
<taxonomicName id="0EB23C93FFE9FFB6FC57FEEFFB156B06" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1020,1155,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FC57FEEFFB156B06" box="[1020,1155,321,343]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FB25FEEFFA9D6B06" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1166,1291,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FB25FEEFFA9D6B06" box="[1166,1291,321,343]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FABDFEEFFA0B6B07" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1302,1437,322,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFE9FFB6FABDFEEFFA0B6B07" box="[1302,1437,322,343]" italics="true" pageId="11" pageNumber="12">H. juniperae</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FC90FECDFC636B25" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,1013,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alboterminata">
<emphasis id="FBC69B02FFE9FFB6FC90FECDFC636B25" box="[827,1013,352,373]" italics="true" pageId="11" pageNumber="12">H. alboterminata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FB82FECDFB1D6B25" baseAuthorityName="Adams &amp; Reeve" baseAuthorityYear="1850" box="[1065,1163,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="whitei">
<emphasis id="FBC69B02FFE9FFB6FB82FECDFB1D6B25" box="[1065,1163,352,373]" italics="true" pageId="11" pageNumber="12">H. whitei</emphasis>
</taxonomicName>
have this arrangement of mantle glands (
<bibRefCitation id="AD233AE1FFE9FFB6FC4EFED3FCE26BE2" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">
Gosliner &amp; Johnson, 1999; present study,
<figureCitation id="51895B95FFE9FFB6FC90FE30FCE26BE2" box="[827,884,413,435]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
</bibRefCitation>
).
<emphasis id="FBC69B02FFE9FFB6FC2DFE30FBCC6BE3" box="[902,1114,413,435]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0EB23C93FFE9FFB6FC2DFE30FB856BE2" authorityName="Stimpson" authorityYear="1855" box="[902,1043,413,434]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
peasei
</emphasis>
,
<taxonomicName id="0EB23C93FFE9FFB6FBCFFE33FB756BE2" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1124,1251,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="insulana">
<emphasis id="FBC69B02FFE9FFB6FBCFFE33FB756BE2" box="[1124,1251,413,435]" italics="true" pageId="11" pageNumber="12">H. insulana</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FB46FE33FA346BE2" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1261,1442,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="violabranchia">
<emphasis id="FBC69B02FFE9FFB6FB46FE33FA346BE2" box="[1261,1442,413,435]" italics="true" pageId="11" pageNumber="12">H. violabranchia</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FCC4FE11FC4E6B81" authorityName="Rudman" authorityYear="1977" box="[879,984,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="emma">
<emphasis id="FBC69B02FFE9FFB6FCC4FE11FC4E6B81" box="[879,984,444,465]" italics="true" pageId="11" pageNumber="12">H. emma</emphasis>
</taxonomicName>
have a similar arrangement of mantle glands, but lack glands along the anterior edge and have anterior glands only along the anterolateral margins (
<bibRefCitation id="AD233AE1FFE9FFB6FCE8FDB5FBEC687D" author="Gosliner T &amp; Johnson, R." box="[835,1146,536,558]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFE9FFB6FB1EFDB5FAFB687D" authorityName="Rudman" authorityYear="1977" box="[1205,1389,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maridadilus">
<emphasis id="FBC69B02FFE9FFB6FB1EFDB5FAFB687D" box="[1205,1389,536,557]" italics="true" pageId="11" pageNumber="12">H. maridadilus</emphasis>
</taxonomicName>
, the mantle glands are distributed along the entire margin of the mantle with the exception of the anterior margin, which is devoid of glands, whereas in
<taxonomicName id="0EB23C93FFE9FFB6FB63FDD9FAB468D9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1224,1314,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FB63FDD9FAB468D9" box="[1224,1314,628,649]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
, the mantle glands are present all around the mantle margin.
</paragraph>
<paragraph id="C90D4710FFE9FFB6FCF8FD1CFA1A69CE" blockId="11.[827,1444,198,1386]" pageId="11" pageNumber="12">
The radulae of most members of this clade are very similar in the shape of their teeth and in the number of denticles on the teeth.
<taxonomicName id="0EB23C93FFE9FFB6FBF3FD43FAE26953" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1112,1396,750,771]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FBF3FD43FAE26953" box="[1112,1396,750,771]" italics="true" pageId="11" pageNumber="12">Hypselodoris alburtuqali</emphasis>
</taxonomicName>
has three or four denticles on the middle lateral teeth, as do
<taxonomicName id="0EB23C93FFE9FFB6FCF0FC81FB8A6911" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[859,1052,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alboterminata">
<emphasis id="FBC69B02FFE9FFB6FCF0FC81FB8A6911" box="[859,1052,812,833]" italics="true" pageId="11" pageNumber="12">H. alboterminata</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FB8CFC81FB3E6911" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1063,1192,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FB8CFC81FB3E6911" box="[1063,1192,812,833]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FB1FFC81FAAF6911" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1204,1337,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="insulana">
<emphasis id="FBC69B02FFE9FFB6FB1FFC81FAAF6911" box="[1204,1337,812,833]" italics="true" pageId="11" pageNumber="12">H. insulana</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FAEFFC81FA356911" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1348,1443,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FAEFFC81FA356911" box="[1348,1443,812,833]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FCC5FCE6FC6B6930" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[878,1021,843,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFE9FFB6FCC5FCE6FC6B6930" box="[878,1021,843,864]" italics="true" pageId="11" pageNumber="12">H. juniperae</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFE9FFB6FB87FCE6FB026930" authorityName="Rudman" authorityYear="1977" box="[1068,1172,843,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="emma">
<emphasis id="FBC69B02FFE9FFB6FB87FCE6FB026930" box="[1068,1172,843,864]" italics="true" pageId="11" pageNumber="12">H. emma</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FB65FCE6FAC9690F" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1230,1375,842,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FB65FCE6FAC9690F" box="[1230,1375,842,864]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
there are four or five denticles, and
<taxonomicName id="0EB23C93FFE9FFB6FB2CFCC7FA95692E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1159,1283,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="violacea">
<emphasis id="FBC69B02FFE9FFB6FB2CFCC7FA95692E" box="[1159,1283,873,895]" italics="true" pageId="11" pageNumber="12">H. violacea</emphasis>
</taxonomicName>
has five or six denticles (
<bibRefCitation id="AD233AE1FFE9FFB6FC04FC25FA1669CE" author="Gosliner T &amp; Johnson, R." box="[943,1408,904,926]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="11" pageNumber="12" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999; present study</bibRefCitation>
).
</paragraph>
<paragraph id="C90D4710FFE9FFB6FCF8FC0BFA326F3A" blockId="11.[827,1444,198,1386]" pageId="11" pageNumber="12">
The reproductive system of
<taxonomicName id="0EB23C93FFE9FFB6FB24FC0AFAA469EB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1167,1330,934,956]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FB24FC0AFAA469EB" box="[1167,1330,934,956]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
is unique among members of the
<taxonomicName id="0EB23C93FFE9FFB6FBE3FC6BFB40698A" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1096,1238,965,987]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FBE3FC6BFB40698A" box="[1096,1238,965,987]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
clade in having a much wider vagina (
<figureCitation id="51895B95FFE9FFB6FB8EFC49FBE069A9" box="[1061,1142,996,1018]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
). In
<taxonomicName id="0EB23C93FFE9FFB6FB07FC49FAAF69A9" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1196,1337,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FB07FC49FAAF69A9" box="[1196,1337,996,1017]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFE9FFB6FAE2FC49FA0C69A9" box="[1353,1434,996,1018]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="11" pageNumber="12">Fig. 4B</figureCitation>
)
<taxonomicName id="0EB23C93FFE9FFB6FC90FBAEFC2E6E47" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[827,952,1026,1048]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FC90FBAEFC2E6E47" box="[827,952,1026,1048]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFE9FFB6FC6DFBAFFB816E48" box="[966,1047,1026,1048]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="11" pageNumber="12">Fig. 4H</figureCitation>
) and
<taxonomicName id="0EB23C93FFE9FFB6FBFEFBAEFB276E48" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1109,1201,1027,1048]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FBFEFBAEFB276E48" box="[1109,1201,1027,1048]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFE9FFB6FB6BFBAFFA8C6E48" box="[1216,1306,1026,1048]" captionStart="Figure 33" captionStartId="53.[163,241,1779,1801]" captionTargetBox="[322,1282,1106,1736]" captionTargetId="figure@53.[321,1284,1105,1738]" captionTargetPageId="53" captionText="Figure 33. Hypselodoris yarae Gosliner &amp; Johnson sp. nov. A, mantle glands. B, reproductive system, scale bar: 0.5 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632860/files/figure.png" pageId="11" pageNumber="12">Fig. 33B</figureCitation>
), the vagina is much narrower. In
<taxonomicName id="0EB23C93FFE9FFB6FB9DFB8FFB4A6E66" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1078,1244,1057,1079]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FB9DFB8FFB4A6E66" box="[1078,1244,1057,1079]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFE9FFB6FB41FB8FFAF96E66" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1258,1391,1057,1079]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FB41FB8FFAF96E66" box="[1258,1391,1057,1079]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FC90FBEDFC5F6E05" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[827,969,1088,1109]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FC90FBEDFC5F6E05" box="[827,969,1088,1109]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
, the receptaculum seminis is situated at the base of the bursa copulatrix or immediately below it, whereas in
<taxonomicName id="0EB23C93FFE9FFB6FC12FBD3FB836EC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[953,1045,1150,1171]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FC12FBD3FB836EC3" box="[953,1045,1150,1171]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
it is much more distally situated. In
<taxonomicName id="0EB23C93FFE9FFB6FC90FB31FC4A6EE1" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,988,1180,1201]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFE9FFB6FC90FB31FC4A6EE1" box="[827,988,1180,1201]" italics="true" pageId="11" pageNumber="12">H. alburtuqali</emphasis>
</taxonomicName>
, the uterine duct connects to the vagina immediately below the bursa and receptaculum, whereas in
<taxonomicName id="0EB23C93FFE9FFB6FCFDFB77FC466EBE" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[854,976,1241,1263]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFE9FFB6FCFDFB77FC466EBE" box="[854,976,1241,1263]" italics="true" pageId="11" pageNumber="12">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFE9FFB6FBAEFB77FB1C6EBE" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1029,1162,1241,1263]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFE9FFB6FBAEFB77FB1C6EBE" box="[1029,1162,1241,1263]" italics="true" pageId="11" pageNumber="12">H. maculosa</emphasis>
</taxonomicName>
the uterine duct branches from the vagina well below the base of the bursa and receptaculum. In
<taxonomicName id="0EB23C93FFE9FFB6FC5DFABAFBC76F7C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1014,1105,1303,1324]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="yarae">
<emphasis id="FBC69B02FFE9FFB6FC5DFABAFBC76F7C" box="[1014,1105,1303,1324]" italics="true" pageId="11" pageNumber="12">H. yarae</emphasis>
</taxonomicName>
, the uterine duct emerges from the base of the vagina, near the genital opening, and has an expanded base adjacent to its juncture with the vagina.
</paragraph>
</subSubSection>
</treatment>
</document>

494
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<taxonomicName id="0EB23C93FFE9FFB6FC4FFA6BFB2D6FAB" ID-CoL="3P32S" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="13" phylum="Mollusca" rank="species" species="brycei" status="sp. nov.">
<heading id="9245F07CFFE9FFB6FC4FFA6BFB6F6F8C" allCaps="true" bold="true" box="[996,1273,1478,1501]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="FBC69B02FFE9FFB6FC4FFA6BFB6F6F8C" bold="true" box="[996,1273,1478,1501]" italics="true" pageId="11" pageNumber="12">HYPSELODORIS BRYCEI</emphasis>
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<heading id="9245F07CFFE9FFB6FC13FA48FB2D6FAB" allCaps="true" box="[952,1211,1509,1533]" centered="true" fontSize="9" level="3" pageId="11" pageNumber="12" reason="4">GOSLINER &amp; JOHNSON</heading>
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<emphasis id="FBC69B02FFE9FFB6FB69FA45FAB36FAD" bold="true" box="[1218,1317,1509,1533]" pageId="11" pageNumber="12">SP. NOV.</emphasis>
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urn:lsid:zoobank.org:act:
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<emphasis id="FBC69B02FFE9FFB6FC90F90AFBF36CEC" box="[827,1125,1702,1724]" italics="true" pageId="11" pageNumber="12">C h r o m o d o r i s b u l l o c k i</emphasis>
m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFE9FFB6FC90F968FB046C8A" ID-CoL="3P32T" authority="Collingwood, 1881" authorityName="Collingwood" authorityYear="1881" box="[827,1170,1733,1755]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFE9FFB6FC90F968FC2C6C8A" box="[827,954,1733,1754]" italics="true" pageId="11" pageNumber="12">C. bullockii</emphasis>
<bibRefCitation id="AD233AE1FFE9FFB6FC6AF968FB046C8A" author="Collingwood C." box="[961,1170,1733,1755]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="11" pageNumber="12" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="AD233AE1FFE9FFB6FB36F968FA146C8A" author="Wells FE &amp; Bryce CW" box="[1181,1410,1733,1755]" journalOrPublisher="Perth: Western Australian Museum" pageId="11" pageNumber="12" refId="ref45476" refString="Wells FE, Bryce CW. 1993. Sea slugs of Western Australia. Perth: Western Australian Museum." title="Sea slugs of Western Australia" type="book" year="1993">Wells &amp; Bryce, 1993</bibRefCitation>
: 8, upper figure.
</paragraph>
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<emphasis id="FBC69B02FFE9FFB6FC90F8AEFBF36D48" box="[827,1125,1794,1816]" italics="true" pageId="11" pageNumber="12">H y p s e l o d o r i s b u l l o c k i</emphasis>
m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFE9FFB6FC90F88FFB246D67" ID-CoL="3P32T" authority="(Collingwood, 1881)" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[827,1202,1825,1847]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFE9FFB6FC90F88FFC556D66" box="[827,963,1825,1847]" italics="true" pageId="11" pageNumber="12">H. bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFE9FFB6FC79F88CFB3C6D67" author="Collingwood C." box="[978,1194,1825,1847]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="11" pageNumber="12" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
)
</taxonomicName>
,
<bibRefCitation id="AD233AE1FFE9FFB6FB14F88CFAFD6D66" author="Debelius H." box="[1215,1387,1825,1847]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="11" pageNumber="12" refId="ref41856" refString="Debelius H. 1997. Schnecken fuhrer Indopazifik. Frankfurt: IKAN-Unterwasserarchiv." title="Schnecken fuhrer Indopazifik" type="book" year="1997">Debelius, 1997</bibRefCitation>
: 233 uppermost photograph.
</paragraph>
<caption id="9DCD1798FFEEFFB1FF3BFBF4FDFD6EDC" httpUri="https://zenodo.org/record/2632781/files/figure.png" pageId="12" pageNumber="13" startId="12.[144,222,1113,1135]" targetBox="[208,1360,195,1072]" targetPageId="12">
<paragraph id="C90D4710FFEEFFB1FF3BFBF4FDFD6EDC" blockId="12.[144,1425,1113,1164]" pageId="12" pageNumber="13">
<emphasis id="FBC69B02FFEEFFB1FF3BFBF4FF616E3E" bold="true" box="[144,247,1113,1135]" pageId="12" pageNumber="13">Figure 5.</emphasis>
<taxonomicName id="0EB23C93FFEEFFB1FEA8FBF4FD206E3F" ID-CoL="3P32S" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[259,694,1113,1135]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="brycei" status="sp. nov.">
<emphasis id="FBC69B02FFEEFFB1FEA8FBF4FE726E3F" bold="true" box="[259,484,1113,1135]" italics="true" pageId="12" pageNumber="13">Hypselodoris brycei</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
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<emphasis id="FBC69B02FFEEFFB1FD10FBF4FC876E3E" bold="true" box="[699,785,1113,1134]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, WAM
<accessionNumber id="D6E1DAF3FFEEFFB1FC68FBF4FB856E3F" box="[963,1043,1113,1135]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S12628" pageId="12" pageNumber="13" type="EnaNcbi">S12628</accessionNumber>
. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFEEFFB1FF3AFB1CFE336EB5" blockId="12.[145,761,1201,1376]" pageId="12" pageNumber="13">
<taxonomicName id="0EB23C93FFEEFFB1FF3AFB1CFEBD6E96" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,299,1201,1222]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEEFFB1FF3AFB1CFEBD6E96" box="[145,299,1201,1222]" italics="true" pageId="12" pageNumber="13">Hypselodoris</emphasis>
</taxonomicName>
cf.
<emphasis id="FBC69B02FFEEFFB1FEFEFB1CFE256E96" box="[341,435,1201,1222]" italics="true" pageId="12" pageNumber="13">bullocki</emphasis>
2
<bibRefCitation id="AD233AE1FFEEFFB1FE79FB1CFD656E96" author="Debelius H &amp; Kuiter R." box="[466,755,1201,1223]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="12" pageNumber="13" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 117, upper photographs.
</paragraph>
<paragraph id="C90D4710FFEEFFB1FF3AFB43FE8D6F73" blockId="12.[145,761,1201,1376]" pageId="12" pageNumber="13">
<taxonomicName id="0EB23C93FFEEFFB1FF3AFB43FEBC6F53" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,298,1262,1283]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEEFFB1FF3AFB43FEBC6F53" box="[145,298,1262,1283]" italics="true" pageId="12" pageNumber="13">Hypselodoris</emphasis>
</taxonomicName>
sp. 9
<bibRefCitation id="AD233AE1FFEEFFB1FEDBFB43FDF66F53" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[368,608,1262,1284]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="12" pageNumber="13" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFEEFFB1FE76FB42FD8E6F53" box="[477,536,1262,1284]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
, 2008
</bibRefCitation>
: 268, bottom photograph.
</paragraph>
<paragraph id="C90D4710FFEEFFB1FF3AFA86FEDE6F30" blockId="12.[145,761,1201,1376]" pageId="12" pageNumber="13">
<taxonomicName id="0EB23C93FFEEFFB1FF3AFA86FEBF6F10" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,297,1323,1344]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEEFFB1FF3AFA86FEBF6F10" box="[145,297,1323,1344]" italics="true" pageId="12" pageNumber="13">Hypselodoris</emphasis>
</taxonomicName>
sp. 18
<bibRefCitation id="AD233AE1FFEEFFB1FED5FA86FDFB6F11" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[382,621,1323,1345]" journalOrPublisher="Jacksonville: New World Press" pageId="12" pageNumber="13" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFEEFFB1FE40FA81FDB06F10" box="[491,550,1323,1345]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
, 2015
</bibRefCitation>
: 263, upper left photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEEFFB1FF3AFA25FB2D6EB7" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="C90D4710FFEEFFB1FF3AFA25FEAD6FCF" blockId="12.[145,761,1415,1561]" box="[145,315,1415,1439]" pageId="12" pageNumber="13">
<heading id="9245F07CFFEEFFB1FF3AFA25FEAD6FCF" box="[145,315,1415,1439]" fontSize="10" level="4" pageId="12" pageNumber="13" reason="1">
<emphasis id="FBC69B02FFEEFFB1FF3AFA25FEAD6FCF" box="[145,315,1415,1439]" italics="true" pageId="12" pageNumber="13">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEEFFB1FF3AFA0AFEF66C49" blockId="12.[145,761,1415,1561]" pageId="12" pageNumber="13">
<materialsCitation id="79DA4D4DFFEEFFB1FF3AFA0AFEF66C49" ID-GBIF-Occurrence="2235579128" collectingDate="2015-04-15" collectionCode="WAM" collectorName="S. Morrison." elevation="7" latitude="-20.4513" location="Montebello Islands" longLatPrecision="7" longitude="115.5827" municipality="Epsilon Island" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="1609F9B2FFEEFFB1FF3AFA0AFF6C6FEC" box="[145,250,1447,1468]" pageId="12" pageNumber="13" type="holotype">
<emphasis id="FBC69B02FFEEFFB1FF3AFA0AFF6C6FEC" box="[145,250,1447,1468]" italics="true" pageId="12" pageNumber="13">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFEEFFB1FEA6FA05FED96FED" box="[269,335,1448,1469]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="13" type="Museum">WAM</collectionCode>
<accessionNumber id="D6E1DAF3FFEEFFB1FEF3FA0AFE256FED" box="[344,435,1447,1469]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S96082" pageId="12" pageNumber="13" type="EnaNcbi">S96082</accessionNumber>
, subsampled for molecular study,
<collectingMunicipality id="2969DD6AFFEEFFB1FF4BFA6BFE026F8C" box="[224,404,1478,1500]" pageId="12" pageNumber="13">Epsilon Island</collectingMunicipality>
,
<location id="CC6D11CBFFEEFFB1FE0AFA6BFD1F6F8C" LSID="urn:lsid:plazi:treatment:411BF606FFE9FFB0FC4FFA6BFA3269EC:CC6D11CBFFEEFFB1FE0AFA6BFD1F6F8C" box="[417,649,1478,1500]" latitude="-20.4513" longLatPrecision="7" longitude="115.5827" municipality="Epsilon Island" name="Montebello Islands" pageId="12" pageNumber="13" stateProvince="Western Australia">Montebello Islands</location>
,
<collectingRegion id="0B7689F2FFEEFFB1FD3DFA6AFF696FAB" country="Australia" name="Western Australia" pageId="12" pageNumber="13">Western Australia</collectingRegion>
,
<geoCoordinate id="AC8621D7FFEEFFB1FEA0FA48FEE86FAB" box="[267,382,1509,1531]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="-20.4513">20.4513°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFEEFFB1FE21FA48FD996FAB" box="[394,527,1509,1531]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="115.5827">115.5827°E</geoCoordinate>
,
<elevation id="429FA023FFEEFFB1FDB0FA48FDD06FAA" box="[539,582,1509,1530]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="7.0">
<quantity id="0E4AEAF5FFEEFFB1FDB0FA48FDD06FAA" box="[539,582,1509,1530]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="7.0">7 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEEFFB1FD37FA48FF5C6C49" pageId="12" pageNumber="13" value="2015-04-15">
<date id="BD0C61D0FFEEFFB1FD37FA48FF5C6C49" pageId="12" pageNumber="13" value="2015-04-15">15 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFEEFFB1FF7FF9A9FEF66C49" box="[212,352,1539,1561]" pageId="12" pageNumber="13">S. Morrison.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFEEFFB1FF3AF9EDFD4E6D1A" blockId="12.[145,761,1599,1867]" pageId="12" pageNumber="13">
<materialsCitation id="79DA4D4DFFEEFFB1FF3AF9EDFD266CC3" ID-GBIF-Occurrence="2235579048" collectingDate="2015-04-15" collectionCode="WAM" collectorName="A. Hara &amp; A. Hosie" elevation="7" latitude="-20.4513" location="Western Australia" longLatPrecision="7" longitude="115.5827" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<typeStatus id="1609F9B2FFEEFFB1FF3AF9EDFE936C05" box="[145,261,1600,1621]" pageId="12" pageNumber="13" type="paratype">
<emphasis id="FBC69B02FFEEFFB1FF3AF9EDFE936C05" box="[145,261,1600,1621]" italics="true" pageId="12" pageNumber="13">Paratypes</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFEEFFB1FEBCF9EDFECF6C05" box="[279,345,1600,1621]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="13" type="Museum">WAM</collectionCode>
<accessionNumber id="D6E1DAF3FFEEFFB1FECAF992FE2C6C05" box="[353,442,1599,1621]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S96083" pageId="12" pageNumber="13" type="EnaNcbi">S96083</accessionNumber>
, subsampled for molecular study,
<collectingRegion id="0B7689F2FFEEFFB1FF49F9F2FE5C6C24" box="[226,458,1630,1652]" country="Australia" name="Western Australia" pageId="12" pageNumber="13">Western Australia</collectingRegion>
,
<geoCoordinate id="AC8621D7FFEEFFB1FE73F9F3FDC16C24" box="[472,599,1630,1652]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="-20.4513">20.4513°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFEEFFB1FDCEF9F3FD626C24" box="[613,756,1630,1652]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="115.5827">115.5827°E</geoCoordinate>
,
<elevation id="429FA023FFEEFFB1FF3AF9D0FF2D6CC2" box="[145,187,1661,1682]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="7.0">
<quantity id="0E4AEAF5FFEEFFB1FF3AF9D0FF2D6CC2" box="[145,187,1661,1682]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="7.0">7 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEEFFB1FEBAF9D0FE396CC3" box="[273,431,1661,1683]" pageId="12" pageNumber="13" value="2015-04-15">
<date id="BD0C61D0FFEEFFB1FEBAF9D0FE396CC3" box="[273,431,1661,1683]" pageId="12" pageNumber="13" value="2015-04-15">15 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFEEFFB1FE12F9D0FD826CC2" box="[441,532,1661,1682]" pageId="12" pageNumber="13">A. Hara</collectorName>
and
<collectorName id="644722C6FFEEFFB1FDE7F9D0FD266CC3" box="[588,688,1661,1683]" pageId="12" pageNumber="13">A. Hosie</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEEFFB1FD13F9D0FD436D1A" ID-GBIF-Occurrence="2235579054" collectingDate="2000-07-31" collectingDateMax="2015-04-16" collectingDateMin="2000-07-31" collectionCode="WAM" collectorName="A. Hara. &amp; St. &amp; Dampier Archipelago, NW &amp; C. Bryce" elevation="15" latitude="-20.4992" location="Legendre Island" longLatPrecision="7" longitude="115.5899" municipality="Montebello Islands" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFEEFFB1FD13F9D0FD6F6CC2" box="[696,761,1661,1682]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="13" type="Museum">WAM</collectionCode>
<accessionNumber id="D6E1DAF3FFEEFFB1FF3AF936FF7C6CE1" box="[145,234,1691,1713]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S96157" pageId="12" pageNumber="13" type="EnaNcbi">S96157</accessionNumber>
, subsampled for molecular study,
<locationDeviation id="D8BF5E32FFEEFFB1FDD1F936FE106C80" location="Ah Chong Island" pageId="12" pageNumber="13">
<quantity id="0E4AEAF5FFEEFFB1FDD1F936FD5C6CE1" box="[634,714,1691,1713]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="13" unit="km" value="5.2">5.2 km</quantity>
NE of Ah Chong Island
</locationDeviation>
,
<collectingMunicipality id="2969DD6AFFEEFFB1FE3EF917FD136C80" box="[405,645,1722,1744]" pageId="12" pageNumber="13">Montebello Islands</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFEEFFB1FD38F916FE9F6CBF" country="Australia" name="Western Australia" pageId="12" pageNumber="13">Western Australia</collectingRegion>
,
<geoCoordinate id="AC8621D7FFEEFFB1FEBDF974FE026CBE" box="[278,404,1753,1774]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="-20.4992">20.4992°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFEEFFB1FE0AF974FDA76CBF" box="[417,561,1753,1775]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="115.5899">115.5899°E</geoCoordinate>
,
<elevation id="429FA023FFEEFFB1FD94F974FDEA6CBE" box="[575,636,1753,1775]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="m" value="15.0">
<quantity id="0E4AEAF5FFEEFFB1FD94F974FDEA6CBE" box="[575,636,1753,1775]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="m" value="15.0">15 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEEFFB1FD77F974FE986D5D" pageId="12" pageNumber="13" value="2015-04-16">
<date id="BD0C61D0FFEEFFB1FD77F974FE986D5D" pageId="12" pageNumber="13" value="2015-04-16">16 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFEEFFB1FEB3F955FEEC6D5D" box="[280,378,1784,1805]" pageId="12" pageNumber="13">A. Hara.</collectorName>
<collectionCode id="AFA3DFD5FFEEFFB1FED4F955FE566D5D" box="[383,448,1784,1805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="13" type="Museum">WAM</collectionCode>
12628, five specimens, one dissected,
<collectorName id="644722C6FFEEFFB1FEA3F8BBFEBE6D7B" box="[264,296,1814,1835]" pageId="12" pageNumber="13">St.</collectorName>
16,
<collectorName id="644722C6FFEEFFB1FEFDF8BBFD106D7C" box="[342,646,1814,1836]" pageId="12" pageNumber="13">Dampier Archipelago, NW</collectorName>
<location id="CC6D11CBFFEEFFB1FD20F8BBFF4B6D1B" LSID="urn:lsid:plazi:treatment:411BF606FFE9FFB0FC4FFA6BFA3269EC:CC6D11CBFFEEFFB1FD20F8BBFF4B6D1B" latitude="-20.4992" longLatPrecision="7" longitude="115.5899" municipality="Montebello Islands" name="Legendre Island" pageId="12" pageNumber="13" stateProvince="Western Australia">Legendre Island</location>
,
<collectingRegion id="0B7689F2FFEEFFB1FF43F898FE566D1B" box="[232,448,1845,1867]" country="Australia" name="Western Australia" pageId="12" pageNumber="13">Western Australia</collectingRegion>
,
<collectingDate id="AD489838FFEEFFB1FE67F898FDF56D1A" box="[460,611,1845,1867]" pageId="12" pageNumber="13" value="2000-07-31">
<date id="BD0C61D0FFEEFFB1FE67F898FDF56D1A" box="[460,611,1845,1867]" pageId="12" pageNumber="13" value="2000-07-31">31 July 2000</date>
</collectingDate>
,
<collectorName id="644722C6FFEEFFB1FDC4F898FD436D1A" box="[623,725,1845,1866]" pageId="12" pageNumber="13">C. Bryce</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFEEFFB1FC82FB1CFC526E98" blockId="12.[809,1211,1200,1255]" box="[809,964,1200,1224]" pageId="12" pageNumber="13">
<heading id="9245F07CFFEEFFB1FC82FB1CFC526E98" box="[809,964,1200,1224]" fontSize="10" level="4" pageId="12" pageNumber="13" reason="1">
<emphasis id="FBC69B02FFEEFFB1FC82FB1CFC526E98" box="[809,964,1200,1224]" italics="true" pageId="12" pageNumber="13">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEEFFB1FC82FB7CFB2D6EB7" blockId="12.[809,1211,1200,1255]" box="[809,1211,1233,1255]" pageId="12" pageNumber="13">Epsilon Island, Montebello Islands.</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEEFFB1FC82FAA3FADD6FD1" pageId="12" pageNumber="13" type="distribution">
<paragraph id="C90D4710FFEEFFB1FC82FAA3FBF06F76" blockId="12.[809,1425,1294,1410]" box="[809,1126,1294,1318]" pageId="12" pageNumber="13">
<heading id="9245F07CFFEEFFB1FC82FAA3FBF06F76" box="[809,1126,1294,1318]" fontSize="10" level="4" pageId="12" pageNumber="13" reason="1">
<emphasis id="FBC69B02FFEEFFB1FC82FAA3FBF06F76" box="[809,1126,1294,1318]" italics="true" pageId="12" pageNumber="13">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEEFFB1FC82FA82FADD6FD1" blockId="12.[809,1425,1294,1410]" pageId="12" pageNumber="13">
Known only from the Houtman Abrolhos Islands to the Exmouth Region and Dampier Archipelago,
<collectingRegion id="0B7689F2FFEEFFB1FA9BFAE0FC036FD2" country="Australia" name="Western Australia" pageId="12" pageNumber="13">Western Australia</collectingRegion>
(
<bibRefCitation id="AD233AE1FFEEFFB1FC0FFAC1FB1E6FD1" author="Wells FE &amp; Bryce CW" box="[932,1160,1388,1410]" journalOrPublisher="Perth: Western Australian Museum" pageId="12" pageNumber="13" refId="ref45476" refString="Wells FE, Bryce CW. 1993. Sea slugs of Western Australia. Perth: Western Australian Museum." title="Sea slugs of Western Australia" type="book" year="1993">Wells &amp; Bryce, 1993</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFEEFFB1FB3EFAC1FAAA6FD1" author="Debelius H." box="[1173,1340,1388,1410]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="12" pageNumber="13" refId="ref41856" refString="Debelius H. 1997. Schnecken fuhrer Indopazifik. Frankfurt: IKAN-Unterwasserarchiv." title="Schnecken fuhrer Indopazifik" type="book" year="1997">Debelius, 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEEFFB1FC82FA05FB6C6C0A" pageId="12" pageNumber="13" type="etymology">
<paragraph id="C90D4710FFEEFFB1FC82FA05FC3C6F90" blockId="12.[809,1426,1448,1626]" box="[809,938,1448,1472]" pageId="12" pageNumber="13">
<heading id="9245F07CFFEEFFB1FC82FA05FC3C6F90" box="[809,938,1448,1472]" fontSize="10" level="4" pageId="12" pageNumber="13" reason="1">
<emphasis id="FBC69B02FFEEFFB1FC82FA05FC3C6F90" box="[809,938,1448,1472]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEEFFB1FC82FA64FB6C6C0A" blockId="12.[809,1426,1448,1626]" pageId="12" pageNumber="13">
<taxonomicName id="0EB23C93FFEEFFB1FC82FA64FC6E6F8E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,1016,1481,1502]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEEFFB1FC82FA64FC6E6F8E" box="[809,1016,1481,1502]" italics="true" pageId="12" pageNumber="13">Hypselodoris brycei</emphasis>
</taxonomicName>
is named for our colleague, Clay Bryce of the Western Australian Museum, who has collected and photographed very many new species of nudibranchs. He was the first to document this new species and has collected most of the material studied here.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEEFFB0FC82F9D4FE626CF0" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="C90D4710FFEEFFB1FC82F9D4FC236CC0" blockId="12.[809,1425,1657,1864]" box="[809,949,1657,1680]" pageId="12" pageNumber="13">
<heading id="9245F07CFFEEFFB1FC82F9D4FC236CC0" box="[809,949,1657,1680]" fontSize="10" level="4" pageId="12" pageNumber="13" reason="1">
<emphasis id="FBC69B02FFEEFFB1FC82F9D4FC236CC0" box="[809,949,1657,1680]" italics="true" pageId="12" pageNumber="13">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEEFFB0FC82F934FD316B68" blockId="12.[809,1425,1657,1864]" lastBlockId="13.[163,779,198,312]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="FBC69B02FFEEFFB1FC82F934FB9E6CFE" box="[809,1032,1689,1710]" italics="true" pageId="12" pageNumber="13">Externalmorphology</emphasis>
: Livinganimals(
<figureCitation id="51895B95FFEEFFB1FB68F934FA9D6CFE" box="[1219,1291,1689,1711]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="12" pageNumber="13">Fig.1B</figureCitation>
)moderately large, reaching
<quantity id="0E4AEAF5FFEEFFB1FC64F91AFB8D6C9D" box="[975,1051,1719,1741]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="50.0">50 mm</quantity>
in length. Body translucent white, with a deep violet marginal band encircling the margin of notum. More purple submarginal pigment found inside marginal band. Nine to 11 thin, unipinnate gill branches on notum. Gill branches with purple base and red–orange outer two-thirds. Base of gill pocket purple. Bulb and base of rhinophores bright red orange, with ~41 small lamellae. Base of rhinophore sheath deep violet to purple. Edge of foot with purple marginal band.
</paragraph>
<paragraph id="C90D4710FFEFFFB0FF08FECCFF606BC6" blockId="13.[163,779,353,406]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FF08FECCFEDF6B26" box="[163,329,353,374]" italics="true" pageId="13" pageNumber="14">Mantle glands</emphasis>
: Subcutaneous mantle glands entirely absent.
</paragraph>
<paragraph id="C90D4710FFEFFFB0FF08FE13FC9C69AC" blockId="13.[163,779,446,1020]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FF08FE13FEFB6B84" box="[163,365,446,468]" italics="true" pageId="13" pageNumber="14">Buccal armature</emphasis>
: Muscular portion of buccal mass about twice length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFEFFFB0FDE6FDB7FD096860" box="[589,671,538,560]" captionStart="Figure 5" captionStartId="12.[144,222,1113,1135]" captionTargetBox="[208,1360,195,1072]" captionTargetId="figure@12.[208,1360,195,1072]" captionTargetPageId="12" captionText="Figure 5. Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632781/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
). Rodlets narrow, with short base and evenly curved, with single, acutely pointed apex. Radular formula of one
<typeStatus id="1609F9B2FFEFFFB0FD34FDF5FC90683D" box="[671,774,600,621]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
, WAM 12628, ~70 × 92.0.92. Rachidian row of teeth absent (
<figureCitation id="51895B95FFEFFFB0FEAAFD38FEC068FB" box="[257,342,661,683]" captionStart="Figure 5" captionStartId="12.[144,222,1113,1135]" captionTargetBox="[208,1360,195,1072]" captionTargetId="figure@12.[208,1360,195,1072]" captionTargetPageId="12" captionText="Figure 5. Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632781/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
). Innermost lateral teeth having one rounded denticle on inner side of bifid primary cusp with another three to four outer denticles. Denticles not extending far beyond middle of elongate primary cusp. Next several laterals lacking inner triangular denticle but possessing four to six denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFEFFFB0FD1BFCE0FC946933" box="[688,770,845,867]" captionStart="Figure 5" captionStartId="12.[144,222,1113,1135]" captionTargetBox="[208,1360,195,1072]" captionTargetId="figure@12.[208,1360,195,1072]" captionTargetPageId="12" captionText="Figure 5. Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632781/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
) all lacking inner denticles, but having five to nine rounded, triangular outer denticles and extended primary cusp. Outermost teeth having a narrower base and shorter tooth shape, with five or six rounded outer denticles (
<figureCitation id="51895B95FFEFFFB0FEF1FC4BFE3A69AC" box="[346,428,998,1020]" captionStart="Figure 5" captionStartId="12.[144,222,1113,1135]" captionTargetBox="[208,1360,195,1072]" captionTargetId="figure@12.[208,1360,195,1072]" captionTargetPageId="12" captionText="Figure 5. Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632781/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
), often larger than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFEFFFB0FF08FB88FE626CF0" blockId="13.[163,780,1061,1696]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FF08FB88FEEB6E6B" box="[163,381,1061,1083]" italics="true" pageId="13" pageNumber="14">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFEFFFB0FD06FB88FC9C6E6B" box="[685,778,1061,1083]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFEFFFB0FE9EFBE9FE166E09" box="[309,384,1092,1114]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="13" pageNumber="14">Fig.4C</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing slightly as it transitions into muscular ejaculatory portion. Prostatic portion enveloping bursa copulatrix. Ejaculatory portion convoluted, narrow, entering short, wider penial bulb. Penial bulb adjacent to slightly curved, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum seminis appressed against vagina. Moderately long uterine duct emerging from vagina close to base of bursa and female gland mass, near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FF08F917FA3269EC" pageId="13" pageNumber="14" type="discussion">
<paragraph id="C90D4710FFEFFFB0FF08F917FE986C82" blockId="13.[163,779,1722,1899]" box="[163,270,1722,1746]" pageId="13" pageNumber="14">
<heading id="9245F07CFFEFFFB0FF08F917FE986C82" box="[163,270,1722,1746]" fontSize="10" level="4" pageId="13" pageNumber="14" reason="1">
<emphasis id="FBC69B02FFEFFFB0FF08F917FE986C82" box="[163,270,1722,1746]" italics="true" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEFFFB0FF08F977FA3269EC" blockId="13.[163,779,1722,1899]" lastBlockId="13.[827,1444,198,956]" pageId="13" pageNumber="14">
The colour pattern of
<taxonomicName id="0EB23C93FFEFFFB0FE09F976FD9C6CBF" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[418,522,1754,1776]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FE09F976FD9C6CBF" box="[418,522,1754,1776]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
is distinctive from all other members of the genus. It most closely resembles
<taxonomicName id="0EB23C93FFEFFFB0FF08F8B5FD026D7E" authority="(Kay &amp; Young, 1969)" baseAuthorityName="Kay &amp; Young" baseAuthorityYear="1969" box="[163,660,1816,1838]" class="Gastropoda" family="Chromodorididae" genus="Thorunna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="daniellae">
<emphasis id="FBC69B02FFEFFFB0FF08F8B5FE046D7D" box="[163,402,1816,1837]" italics="true" pageId="13" pageNumber="14">Thorunna daniellae</emphasis>
(
<bibRefCitation id="AD233AE1FFEFFFB0FE09F8B5FD1D6D7E" author="Kay EA &amp; Young, DK" box="[418,651,1816,1838]" journalOrPublisher="Pacific Science" pageId="13" pageNumber="14" pagination="18 - 37" part="23" refId="ref43288" refString="Kay EA, Young, DK. 1969. The Doridacea (Opisthobranchia Mollusca) of the Hawaiian islands. Pacific Science 23: 18 - 37." title="The Doridacea (Opisthobranchia Mollusca) of the Hawaiian islands" type="journal article" year="1969">Kay &amp; Young, 1969</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFEFFFB0FD0EF8B5FE8B6D1C" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" journalOrPublisher="Jacksonville: New World Press" pageId="13" pageNumber="14" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFEFFFB0FF08F89AFF4C6D1B" box="[163,218,1846,1868]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2015
</bibRefCitation>
: 250 upper right photograph), but
<taxonomicName id="0EB23C93FFEFFFB0FD0CF89AFC9D6D1B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[679,779,1846,1868]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FD0CF89AFC9D6D1B" box="[679,779,1846,1868]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
is a larger species, reaching
<quantity id="0E4AEAF5FFEFFFB0FE48F8F8FDA76D3B" box="[483,561,1877,1899]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="50.0">50 mm</quantity>
in length, whereas
<taxonomicName id="0EB23C93FFEFFFB0FC90FF6AFC5B6A8B" baseAuthorityName="Kay &amp; Young" baseAuthorityYear="1969" box="[827,973,198,220]" class="Gastropoda" family="Chromodorididae" genus="Thorunna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="daniellae">
<emphasis id="FBC69B02FFEFFFB0FC90FF6AFC5B6A8B" box="[827,973,198,220]" italics="true" pageId="13" pageNumber="14">T. daniellae</emphasis>
</taxonomicName>
rarely exceeds
<quantity id="0E4AEAF5FFEFFFB0FB39FF6AFB7E6A8C" box="[1170,1256,199,220]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="20.0">20 mm</quantity>
in length. Our molecular phylogeny (
<figureCitation id="51895B95FFEFFFB0FBE9FF48FB056AAB" box="[1090,1171,229,251]" captionStart="Figure 35" captionStartId="56.[146,226,1818,1840]" captionTargetBox="[210,1362,196,1775]" captionTargetId="figure@56.[209,1364,195,1777]" captionTargetPageId="56" captionText="Figure 35. Bayesian inference and maximum likelihood tree of Hypselodoris, indicating support values for each node. Bootstrap values are on the left and Bayesian posterior probablilities on the right." httpUri="https://zenodo.org/record/2632866/files/figure.png" pageId="13" pageNumber="14">Fig. 35</figureCitation>
) clearly indicates that
<taxonomicName id="0EB23C93FFEFFFB0FC90FEA9FC326B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,932,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FC90FEA9FC326B49" box="[827,932,260,281]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
is in the
<taxonomicName id="0EB23C93FFEFFFB0FBB3FEA9FB346B49" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[1048,1186,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFEFFFB0FBB3FEA9FB346B49" box="[1048,1186,260,281]" italics="true" pageId="13" pageNumber="14">H. bullockii</emphasis>
</taxonomicName>
clade, where it is the sister species to
<taxonomicName id="0EB23C93FFEFFFB0FBAFFE8EFADD6B69" authority="(Yonow, 2001)" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[1028,1355,290,313]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FBAFFE8EFB0D6B67" box="[1028,1179,290,312]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
(
<bibRefCitation id="AD233AE1FFEFFFB0FB00FE8EFAD56B69" author="Yonow N." box="[1195,1347,291,313]" journalOrPublisher="Zoologische Mededelingen" pageId="13" pageNumber="14" pagination="1 - 50" part="75" refId="ref45664" refString="Yonow N. 2001. Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material. Zoologische Mededelingen 75: 1 - 50." title="Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material" type="journal article" year="2001">Yonow, 2001</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="AD233AE1FFEFFFB0FAFDFE8EFC146B07" author="Yonow N." journalOrPublisher="Zoologische Mededelingen" pageId="13" pageNumber="14" pagination="1 - 50" part="75" refId="ref45664" refString="Yonow N. 2001. Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material. Zoologische Mededelingen 75: 1 - 50." title="Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material" type="journal article" year="2001">Yonow (2001)</bibRefCitation>
described
<taxonomicName id="0EB23C93FFEFFFB0FC56FEEFFB1A6B06" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[1021,1164,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FC56FEEFFB1A6B06" box="[1021,1164,321,343]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
</taxonomicName>
as a species of
<taxonomicName id="0EB23C93FFEFFFB0FA96FEECFA0B6B06" authorityName="Odhner" authorityYear="1934" box="[1341,1437,321,342]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEFFFB0FA96FEECFA0B6B06" box="[1341,1437,321,342]" italics="true" pageId="13" pageNumber="14">Risbecia</emphasis>
</taxonomicName>
, but
<bibRefCitation id="AD233AE1FFEFFFB0FCC1FECDFB326B26" author="Johnson R &amp; Gosliner T." box="[874,1188,352,374]" journalOrPublisher="PLoS One" pageId="13" pageNumber="14" pagination="33479" part="7" refId="ref43042" refString="Johnson R, Gosliner T. 2012. Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs. PLoS One 7: e 33479." title="Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs" type="journal article" year="2012">Johnson &amp; Gosliner (2012)</bibRefCitation>
clearly demonstrated that
<taxonomicName id="0EB23C93FFEFFFB0FCDFFED3FC436BC3" authorityName="Odhner" authorityYear="1934" box="[884,981,382,403]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEFFFB0FCDFFED3FC436BC3" box="[884,981,382,403]" italics="true" pageId="13" pageNumber="14">Risbecia</emphasis>
</taxonomicName>
is nested within
<taxonomicName id="0EB23C93FFEFFFB0FB0AFED3FAA16BC3" authorityName="Stimpson" authorityYear="1855" box="[1185,1335,382,403]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEFFFB0FB0AFED3FAA16BC3" box="[1185,1335,382,403]" italics="true" pageId="13" pageNumber="14">Hypselodoris</emphasis>
</taxonomicName>
and that
<taxonomicName id="0EB23C93FFEFFFB0FC90FE33FC466BE2" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[827,976,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FC90FE33FC466BE2" box="[827,976,413,435]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
</taxonomicName>
is not in the same clade as the species formerly included in
<taxonomicName id="0EB23C93FFEFFFB0FB91FE11FB086B81" authorityName="Odhner" authorityYear="1934" box="[1082,1182,444,465]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFEFFFB0FB91FE11FB086B81" box="[1082,1182,444,465]" italics="true" pageId="13" pageNumber="14">Risbecia</emphasis>
</taxonomicName>
but is closely related to
<taxonomicName id="0EB23C93FFEFFFB0FCF0FE76FC7F6BBF" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[859,1001,474,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFEFFFB0FCF0FE76FC7F6BBF" box="[859,1001,474,496]" italics="true" pageId="13" pageNumber="14">H. bullockii</emphasis>
</taxonomicName>
. Of the members of the
<taxonomicName id="0EB23C93FFEFFFB0FABDFE76FA326BBF" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[1302,1444,474,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFEFFFB0FABDFE76FA326BBF" box="[1302,1444,474,496]" italics="true" pageId="13" pageNumber="14">H. bullockii</emphasis>
</taxonomicName>
clade,
<taxonomicName id="0EB23C93FFEFFFB0FC2FFE57FC7A685E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[900,1004,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FC2FFE57FC7A685E" box="[900,1004,505,527]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
is the only species with a white body colour and a violet marginal band. All members of the
<taxonomicName id="0EB23C93FFEFFFB0FC90FD9AFC57681B" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[827,961,566,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFEFFFB0FC90FD9AFC57681B" box="[827,961,566,588]" italics="true" pageId="13" pageNumber="14">H. bullockii</emphasis>
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clade lack mantle glands and are unique among
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species in this regard. The radula of
<taxonomicName id="0EB23C93FFEFFFB0FCF2FDD9FC6668D9" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[857,1008,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FCF2FDD9FC6668D9" box="[857,1008,628,649]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
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appears to contain more teeth than that of
<taxonomicName id="0EB23C93FFEFFFB0FC3BFD3EFC6168F7" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[912,1015,658,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FC3BFD3EFC6168F7" box="[912,1015,658,680]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFEFFFB0FBACFD3EFAD368F8" author="Yonow N." box="[1031,1349,658,680]" journalOrPublisher="Zoologische Mededelingen" pageId="13" pageNumber="14" pagination="1 - 50" part="75" refId="ref45664" refString="Yonow N. 2001. Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material. Zoologische Mededelingen 75: 1 - 50." title="Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material" type="journal article" year="2001">Yonow, 2001; present study</bibRefCitation>
).
<bibRefCitation id="AD233AE1FFEFFFB0FAF3FD3EFC146897" author="Yonow N." journalOrPublisher="Zoologische Mededelingen" pageId="13" pageNumber="14" pagination="1 - 50" part="75" refId="ref45664" refString="Yonow N. 2001. Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material. Zoologische Mededelingen 75: 1 - 50." title="Results of the Rumphius Biohistorical Expedition to Ambon (1990). Part 11. Doridacea of the families Chromodorididae and Hexabranchidae (Mollusca, Gastropoda, Opisthobranchia, Nudibranchia), including additional Moluccan material" type="journal article" year="2001">Yonow (2001)</bibRefCitation>
described the radula formula of
<taxonomicName id="0EB23C93FFEFFFB0FB5EFD1FFA156896" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[1269,1411,689,711]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FB5EFD1FFA156896" box="[1269,1411,689,711]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
</taxonomicName>
as 96 × 120.0.120, whereas the specimen of
<taxonomicName id="0EB23C93FFEFFFB0FAA7FD7DFAE768B5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1292,1393,720,741]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FAA7FD7DFAE768B5" box="[1292,1393,720,741]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
had a formula of 70 × 92.0.92. The denticles of the radular teeth of
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<emphasis id="FBC69B02FFEFFFB0FC38FCA3FC636972" box="[915,1013,781,803]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
</taxonomicName>
do not extend much beyond the middle of the primary cusp, whereas in
<taxonomicName id="0EB23C93FFEFFFB0FB3FFC81FA8B6911" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[1172,1309,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FB3FFC81FA8B6911" box="[1172,1309,812,833]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
</taxonomicName>
they extend almost to the end of the cusp. The reproductive system of
<taxonomicName id="0EB23C93FFEFFFB0FCFDFCC7FC2F692E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[854,953,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="brycei">
<emphasis id="FBC69B02FFEFFFB0FCFDFCC7FC2F692E" box="[854,953,873,895]" italics="true" pageId="13" pageNumber="14">H. brycei</emphasis>
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has a much shorter penial papilla and long ejaculatory duct, whereas
<taxonomicName id="0EB23C93FFEFFFB0FBCDFC25FB6269CD" baseAuthorityName="Yonow" baseAuthorityYear="2001" box="[1126,1268,904,925]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolegma">
<emphasis id="FBC69B02FFEFFFB0FBCDFC25FB6269CD" box="[1126,1268,904,925]" italics="true" pageId="13" pageNumber="14">H. apolegma</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFEFFFB0FAAFFC25FAC069CD" box="[1284,1366,904,926]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="13" pageNumber="14">Fig. 4D</figureCitation>
) has a longer, wider penial bulb and a shorter ejaculatory duct.
</paragraph>
</subSubSection>
</treatment>
</document>

528
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misidentification, not
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<bibRefCitation id="AD233AE1FFEDFFB2FC90F8AEFC746D48" author="Rudman WB" box="[827,994,1795,1817]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="15" pageNumber="16" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
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;
<bibRefCitation id="AD233AE1FFEDFFB2FC46F8AEFB366D49" author="Rudman WB" box="[1005,1184,1795,1817]" journalOrPublisher="Sydney: Australian Museum" pageId="15" pageNumber="16" publicationUrl="http://www.seaslugforum.net/find/hypskang" refId="ref44958" refString="Rudman WB. 1999 b. Hypselodoris kanga Rudman, 1977. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypskang" title="Hypselodoris kanga Rudman, 1977" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999b</bibRefCitation>
: lower photograph.
</paragraph>
<paragraph id="C90D4710FFEDFFB2FC90F88CFC466D25" blockId="15.[827,1443,1764,1909]" pageId="15" pageNumber="16">
<taxonomicName id="0EB23C93FFEDFFB2FC90F88CFBB06D66" ID-CoL="3P343" authorityName="Rudman" authorityYear="1977" box="[827,1062,1825,1846]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFEDFFB2FC90F88CFBB06D66" box="[827,1062,1825,1846]" italics="true" pageId="15" pageNumber="16">Hypselodoris kanga</emphasis>
</taxonomicName>
misidentification, not
<taxonomicName id="0EB23C93FFEDFFB2FA9DF88FFC746D05" ID-CoL="3P343" authority="Rudman, 1977" authorityName="Rudman" authorityYear="1977" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFEDFFB2FA9DF88FFA356D66" box="[1334,1443,1825,1847]" italics="true" pageId="15" pageNumber="16">H. kanga</emphasis>
<bibRefCitation id="AD233AE1FFEDFFB2FC90F8EDFC746D05" author="Rudman WB" box="[827,994,1856,1878]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="15" pageNumber="16" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="AD233AE1FFEDFFB2FC40F8EDFB6D6D05" author="Debelius H &amp; Kuiter R." box="[1003,1275,1856,1878]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="15" pageNumber="16" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 126, top three photographs.
</paragraph>
<caption id="9DCD1798FFF2FFADFF39F9BBFCA06C1A" httpUri="https://zenodo.org/record/2632785/files/figure.png" pageId="16" pageNumber="17" startId="16.[146,224,1558,1580]" targetBox="[210,1362,200,1518]" targetPageId="16">
<paragraph id="C90D4710FFF2FFADFF39F9BBFCA06C1A" blockId="16.[146,1426,1558,1610]" pageId="16" pageNumber="17">
<emphasis id="FBC69B02FFF2FFADFF39F9BBFF6C6C7C" bold="true" box="[146,250,1558,1580]" pageId="16" pageNumber="17">Figure 7.</emphasis>
<taxonomicName id="0EB23C93FFF2FFADFEAEF9BBFD6F6C7B" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[261,761,1558,1580]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFF2FFADFEAEF9BBFE736C7B" box="[261,485,1558,1579]" italics="true" pageId="16" pageNumber="17">Hypseldoris krakatoa</emphasis>
Gosliner &amp; Johnson, 1999
</taxonomicName>
, CASIZ 206801. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFF2FFADFF3AF9C3FE8D6CF3" blockId="16.[145,761,1646,1760]" pageId="16" pageNumber="17">
<taxonomicName id="0EB23C93FFF2FFADFF3AF9C3FEBC6CD3" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,298,1646,1667]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFF2FFADFF3AF9C3FEBC6CD3" box="[145,298,1646,1667]" italics="true" pageId="16" pageNumber="17">Hypselodoris</emphasis>
</taxonomicName>
sp. 2
<bibRefCitation id="AD233AE1FFF2FFADFEDBF9C3FDF66CD3" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[368,608,1646,1668]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="16" pageNumber="17" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFF2FFADFE76F9C2FD8E6CD3" box="[477,536,1646,1668]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, 2008
</bibRefCitation>
: 264: bottom photograph.
</paragraph>
<paragraph id="C90D4710FFF2FFADFF3AF906FE396CB0" blockId="16.[145,761,1646,1760]" pageId="16" pageNumber="17">
<taxonomicName id="0EB23C93FFF2FFADFF3AF906FEBB6C90" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,301,1707,1728]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFF2FFADFF3AF906FEBB6C90" box="[145,301,1707,1728]" italics="true" pageId="16" pageNumber="17">Hypselodoris</emphasis>
</taxonomicName>
sp. 2
<bibRefCitation id="AD233AE1FFF2FFADFEDDF901FD2E6C90" author="Humann P &amp; DeLoach N." box="[374,696,1707,1729]" journalOrPublisher="Jacksonville: New World Publications" pageId="16" pageNumber="17" refId="ref42986" refString="Humann P, DeLoach N. 2010. Reef creature identification, tropical Pacific. Jacksonville: New World Publications." title="Reef creature identification, tropical Pacific" type="book" year="2010">Humann &amp; DeLoach, 2010</bibRefCitation>
: 339: middle right photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF2FFACFF3AF8A5FDE16CF3" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="C90D4710FFF2FFADFF3AF8A5FEAD6D4F" blockId="16.[145,761,1799,1884]" box="[145,315,1799,1823]" pageId="16" pageNumber="17">
<heading id="9245F07CFFF2FFADFF3AF8A5FEAD6D4F" box="[145,315,1799,1823]" fontSize="10" level="4" pageId="16" pageNumber="17" reason="1">
<emphasis id="FBC69B02FFF2FFADFF3AF8A5FEAD6D4F" box="[145,315,1799,1823]" italics="true" pageId="16" pageNumber="17">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF2FFADFF3AF88AFD6F6D0C" blockId="16.[145,761,1799,1884]" pageId="16" pageNumber="17">
<materialsCitation id="79DA4D4DFFF2FFADFF3AF88AFD6F6D0C" ID-GBIF-Occurrence="2235579047" collectionCode="CASIZ" latitude="-5.183333" location="Siar Island" longLatPrecision="1" longitude="145.80667" pageId="16" pageNumber="17" specimenCode="CASIZ 191070" specimenCount="1" typeStatus="holotype">
<typeStatus id="1609F9B2FFF2FFADFF3AF88AFF6F6D6C" box="[145,249,1831,1852]" pageId="16" pageNumber="17" type="holotype">
<emphasis id="FBC69B02FFF2FFADFF3AF88AFF6F6D6C" box="[145,249,1831,1852]" italics="true" pageId="16" pageNumber="17">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFF2FFADFEA7F88AFE2F6D6C" box="[268,441,1831,1852]" collectionCode="CASIZ" pageId="16" pageNumber="17">
CASIZ
<date id="BD0C61D0FFF2FFADFEC9F88AFE2F6D6C" box="[354,441,1831,1852]" pageId="16" pageNumber="17" value="70-10-19">191070</date>
</specimenCode>
, subsampled for molecular study, dissected,
<location id="CC6D11CBFFF2FFADFEE3F8EBFE536D0C" LSID="urn:lsid:plazi:treatment:411BF606FFEDFFAFFC7FF9A9FB906D24:CC6D11CBFFF2FFADFEE3F8EBFE536D0C" box="[328,453,1862,1884]" latitude="-5.183333" longLatPrecision="1" longitude="145.80667" name="Siar Island" pageId="16" pageNumber="17">Siar Island</location>
,
<geoCoordinate id="AC8621D7FFF2FFADFE67F8EBFDDB6D0C" box="[460,589,1862,1884]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-5.183333">5.183333°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF2FFADFDFEF8EBFD636D0C" box="[597,757,1862,1884]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="145.80667">145.806667°E</geoCoordinate>
,
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFF2FFADFC82F9C3FB826CF2" blockId="16.[809,1425,1646,1699]" pageId="16" pageNumber="17">
<materialsCitation id="79DA4D4DFFF2FFADFC82F9C3FB996CF2" ID-GBIF-Occurrence="2235579041" collectingDate="2012-11-09" country="Papua New Guinea" elevation="10" location="Madang" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Madang">
<collectingRegion id="0B7689F2FFF2FFADFC82F9C3FC116CD4" box="[809,903,1646,1668]" country="Papua New Guinea" name="Madang" pageId="16" pageNumber="17">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFF2FFADFC39F9C3FBE66CD4" box="[914,1136,1646,1668]" name="Papua New Guinea" pageId="16" pageNumber="17">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFF2FFADFBD0F9C3FB256CD3" box="[1147,1203,1646,1667]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="10.0">
<quantity id="0E4AEAF5FFF2FFADFBD0F9C3FB256CD3" box="[1147,1203,1646,1667]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="10.0">10 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFF2FFADFAACF9C3FCF46CF2" pageId="16" pageNumber="17" value="2012-11-09">
<date id="BD0C61D0FFF2FFADFAACF9C3FCF46CF2" pageId="16" pageNumber="17" value="2012-11-09">9 November 2012</date>
</collectingDate>
, Heok Hui Tan
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFF2FFADFC82F961FB006D0B" blockId="16.[809,1425,1739,1884]" pageId="16" pageNumber="17">
<materialsCitation id="79DA4D4DFFF2FFADFC82F961FB7A6D50" ID-GBIF-Occurrence="2235579056" collectionCode="CASIZ" country="Philippines" location="Panglao Channel" municipality="Tagbilaran" pageId="16" pageNumber="17" specimenCode="CASIZ 176067" specimenCount="1" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="1609F9B2FFF2FFADFC82F961FC376CB1" box="[809,929,1740,1761]" pageId="16" pageNumber="17" type="paratype">
<emphasis id="FBC69B02FFF2FFADFC82F961FC376CB1" box="[809,929,1740,1761]" italics="true" pageId="16" pageNumber="17">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFF2FFADFC1EF966FBFD6CB1" box="[949,1131,1739,1761]" collectionCode="CASIZ" pageId="16" pageNumber="17">CASIZ 176067</specimenCode>
, one specimen,
<location id="CC6D11CBFFF2FFADFA9BF966FC1A6D50" LSID="urn:lsid:plazi:treatment:411BF606FFEDFFAFFC7FF9A9FB906D24:CC6D11CBFFF2FFADFA9BF966FC1A6D50" country="Philippines" municipality="Tagbilaran" name="Panglao Channel" pageId="16" pageNumber="17" stateProvince="Bohol">Panglao Channel</location>
,
<collectingMunicipality id="2969DD6AFFF2FFADFC3FF947FB856D50" box="[916,1043,1770,1792]" pageId="16" pageNumber="17">Tagbilaran</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF2FFADFBB7F947FBC96D50" box="[1052,1119,1770,1792]" country="Philippines" name="Bohol" pageId="16" pageNumber="17">Bohol</collectingRegion>
,
<collectingCountry id="B1A50780FFF2FFADFBC2F947FB7A6D50" box="[1129,1260,1770,1792]" name="Philippines" pageId="16" pageNumber="17">Philippines</collectingCountry>
</materialsCitation>
,
<date id="BD0C61D0FFF2FFADFB5DF947FA1B6CAF" box="[1270,1421,1770,1791]" pageId="16" pageNumber="17" value="2004-06-28">28 June 2004</date>
, Gosliner, 2004 Panglao Expedition.
<materialsCitation id="79DA4D4DFFF2FFADFB12F8A4FBCC6D6D" ID-GBIF-Occurrence="2235579090" collectionCode="CASIZ" country="Indonesia" location="Bali" pageId="16" pageNumber="17" specimenCode="CASIZ 190503" specimenCount="1" stateProvince="Bali" typeStatus="paratype">
<specimenCode id="9914EF6BFFF2FFADFB12F8A4FAC96D4E" box="[1209,1375,1800,1822]" collectionCode="CASIZ" pageId="16" pageNumber="17">
CASIZ
<date id="BD0C61D0FFF2FFADFAA0F8A5FAC96D4E" box="[1291,1375,1800,1822]" pageId="16" pageNumber="17" value="03-05-19">190503</date>
</specimenCode>
, one specimen,
<collectingRegion id="0B7689F2FFF2FFADFC0EF88AFC4E6D6D" box="[933,984,1831,1853]" country="Indonesia" name="Bali" pageId="16" pageNumber="17">Bali</collectingRegion>
,
<collectingCountry id="B1A50780FFF2FFADFC4FF88AFBCC6D6D" box="[996,1114,1831,1853]" name="Indonesia" pageId="16" pageNumber="17">Indonesia</collectingCountry>
</materialsCitation>
, 1998.
<materialsCitation id="79DA4D4DFFF2FFADFB05F88AFBDB6D0C" ID-GBIF-Occurrence="2235579043" collectionCode="CASIZ" country="Indonesia" location="Bali" pageId="16" pageNumber="17" specimenCode="CASIZ 190504" specimenCount="1" stateProvince="Bali" typeStatus="paratype">
<specimenCode id="9914EF6BFFF2FFADFB05F88AFACA6D6D" box="[1198,1372,1831,1853]" collectionCode="CASIZ" pageId="16" pageNumber="17">
CASIZ
<date id="BD0C61D0FFF2FFADFAAEF88AFACA6D6D" box="[1285,1372,1831,1853]" pageId="16" pageNumber="17" value="04-05-19">190504</date>
</specimenCode>
, one specimen,
<collectingRegion id="0B7689F2FFF2FFADFC0BF8EBFC476D0C" box="[928,977,1862,1884]" country="Indonesia" name="Bali" pageId="16" pageNumber="17">Bali</collectingRegion>
,
<collectingCountry id="B1A50780FFF2FFADFC70F8EBFBDB6D0C" box="[987,1101,1862,1884]" name="Indonesia" pageId="16" pageNumber="17">Indonesia</collectingCountry>
</materialsCitation>
, 1998.
</paragraph>
<caption id="9DCD1798FFF3FFACFF0FF9B8FC6C6C19" httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="17" pageNumber="18" startId="17.[164,243,1557,1579]" targetBox="[228,1380,195,1517]" targetPageId="17">
<paragraph id="C90D4710FFF3FFACFF0FF9B8FC6C6C19" blockId="17.[164,1444,1557,1609]" pageId="17" pageNumber="18">
<emphasis id="FBC69B02FFF3FFACFF0FF9B8FE986C7B" bold="true" box="[164,270,1557,1579]" pageId="17" pageNumber="18">Figure 8.</emphasis>
<taxonomicName id="0EB23C93FFF3FFACFEB2F9B8FD7D6C7C" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[281,747,1557,1580]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="confetti" status="sp. nov.">
Hypselodoris
<emphasis id="FBC69B02FFF3FFACFE13F9B8FD846C7B" bold="true" box="[440,530,1557,1579]" italics="true" pageId="17" pageNumber="18">confetti</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFF3FFACFD59F9BBFCDF6C7B" box="[754,841,1558,1579]" pageId="17" pageNumber="18" rank="species">
<emphasis id="FBC69B02FFF3FFACFD59F9BBFCDF6C7B" bold="true" box="[754,841,1558,1579]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFF3FFACFF08F9C3FEAB6CD5" blockId="17.[163,631,1645,1699]" box="[163,317,1645,1669]" pageId="17" pageNumber="18">
<heading id="9245F07CFFF3FFACFF08F9C3FEAB6CD5" box="[163,317,1645,1669]" fontSize="10" level="4" pageId="17" pageNumber="18" reason="1">
<emphasis id="FBC69B02FFF3FFACFF08F9C3FEAB6CD5" box="[163,317,1645,1669]" italics="true" pageId="17" pageNumber="18">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF3FFACFF08F920FDE16CF3" blockId="17.[163,631,1645,1699]" box="[163,631,1677,1699]" pageId="17" pageNumber="18">
Siar Island,
<collectingRegion id="0B7689F2FFF3FFACFE85F920FE1D6CF3" box="[302,395,1677,1699]" country="Papua New Guinea" name="Madang" pageId="17" pageNumber="18">Madang</collectingRegion>
<collectingCountry id="B1A50780FFF3FFACFE38F923FDE56CF3" box="[403,627,1677,1699]" name="Papua New Guinea" pageId="17" pageNumber="18">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF3FFACFF08F945FDD56D0B" pageId="17" pageNumber="18" type="distribution">
<paragraph id="C90D4710FFF3FFACFF08F945FE766D50" blockId="17.[163,779,1768,1883]" box="[163,480,1768,1792]" pageId="17" pageNumber="18">
<heading id="9245F07CFFF3FFACFF08F945FE766D50" box="[163,480,1768,1792]" fontSize="10" level="4" pageId="17" pageNumber="18" reason="1">
<emphasis id="FBC69B02FFF3FFACFF08F945FE766D50" box="[163,480,1768,1792]" italics="true" pageId="17" pageNumber="18">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF3FFACFF08F8A4FDD56D0B" blockId="17.[163,779,1768,1883]" pageId="17" pageNumber="18">
Known from
<collectingCountry id="B1A50780FFF3FFACFE90F8A4FD8B6D4E" box="[315,541,1800,1822]" name="Papua New Guinea" pageId="17" pageNumber="18">Papua New Guinea</collectingCountry>
and the
<collectingCountry id="B1A50780FFF3FFACFD2DF8A5FC9D6D4E" box="[646,779,1800,1822]" name="Philippines" pageId="17" pageNumber="18">Philippines</collectingCountry>
(present study) and probably also
<collectingCountry id="B1A50780FFF3FFACFDE0F88AFD436D6C" box="[587,725,1831,1852]" name="Hong Kong" pageId="17" pageNumber="18">Hong Kong</collectingCountry>
and
<collectingCountry id="B1A50780FFF3FFACFF08F8E8FE856D0B" box="[163,275,1861,1883]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
(
<bibRefCitation id="AD233AE1FFF3FFACFE89F8E8FDA56D0B" author="Debelius H &amp; Kuiter R." box="[290,563,1861,1883]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="17" pageNumber="18" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF3FFACFC90F9C0FAF76D0B" pageId="17" pageNumber="18" type="etymology">
<paragraph id="C90D4710FFF3FFACFC90F9C0FC2A6CD5" blockId="17.[827,1443,1645,1883]" box="[827,956,1645,1669]" pageId="17" pageNumber="18">
<heading id="9245F07CFFF3FFACFC90F9C0FC2A6CD5" box="[827,956,1645,1669]" fontSize="10" level="4" pageId="17" pageNumber="18" reason="1">
<emphasis id="FBC69B02FFF3FFACFC90F9C0FC2A6CD5" box="[827,956,1645,1669]" italics="true" pageId="17" pageNumber="18">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF3FFACFC90F923FAF76D0B" blockId="17.[827,1443,1645,1883]" pageId="17" pageNumber="18">
<emphasis id="FBC69B02FFF3FFACFC90F923FBCE6CF3" box="[827,1112,1677,1699]" italics="true" pageId="17" pageNumber="18">H y p s e l o d o r i s c o n f e t t i</emphasis>
c o m e s f r o m t h e I t a l i a n word
<taxonomicName id="0EB23C93FFF3FFACFC2AF901FC496C92" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[897,991,1708,1730]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="confetti">confetti</taxonomicName>
that means sweets, referring to the multicoloured sweets that were thrown to people at Italian carnivals. In the 19th century it was used to refer to brightly coloured pieces of paper tossed out during parades. This species is marked with bright blue, yellow and black spots resembling
<taxonomicName id="0EB23C93FFF3FFACFAAFF8E8FACB6D0B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1284,1373,1861,1883]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="confetti">confetti</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF0FFAFFF3AFF6AFB7A6B68" pageId="18" pageNumber="19" type="description">
<paragraph id="C90D4710FFF0FFAFFF3AFF6AFE8B6A8E" blockId="18.[145,761,199,682]" box="[145,285,199,222]" pageId="18" pageNumber="19">
<heading id="9245F07CFFF0FFAFFF3AFF6AFE8B6A8E" box="[145,285,199,222]" fontSize="10" level="4" pageId="18" pageNumber="19" reason="1">
<emphasis id="FBC69B02FFF0FFAFFF3AFF6AFE8B6A8E" box="[145,285,199,222]" italics="true" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF0FFAFFF3AFF4BFDC568FA" blockId="18.[145,761,199,682]" pageId="18" pageNumber="19">
<emphasis id="FBC69B02FFF0FFAFFF3AFF4BFE366AAB" box="[145,416,230,251]" italics="true" pageId="18" pageNumber="19">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFF0FFAFFD3BFF4AFD666AAC" box="[656,752,231,253]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="18" pageNumber="19">Fig. 1D</figureCitation>
) moderately large, reaching
<quantity id="0E4AEAF5FFF0FFAFFE4AFEA8FDA26B4B" box="[481,564,261,283]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="mm" value="35.0">35 mm</quantity>
in length. Body colour whitish to grey–blue. Notum ornamented with large yellow spots and smaller dark blue to black spots scattered over the surface. Large blue areas found near the mantle margin. Additional spots of same colour found on sides of body and foot. Gill pocket slightly elevated from notum. Seven to nine narrow, thin, unipinnate gill branches on notum. Gill branches with purple lines along edges of inner and outer surface.Apex of gill branch red–orange. Central portion of middle of outer face of gill branches with three to five yellow spots. Base of rhinophores deep purple, extending onto basal half of club. Upper half of rhinophore club bright red rhinophores with 19 small lamellae.
</paragraph>
<paragraph id="C90D4710FFF0FFAFFF3AFD7DFE446973" blockId="18.[145,761,720,803]" pageId="18" pageNumber="19">
<emphasis id="FBC69B02FFF0FFAFFF3AFD7DFEA368B5" box="[145,309,720,741]" italics="true" pageId="18" pageNumber="19">Mantle glands</emphasis>
: Subcutaneous mantle glands (
<figureCitation id="51895B95FFF0FFAFFD0BFD7DFD6668B5" box="[672,752,720,742]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="18" pageNumber="19">Fig. 2C</figureCitation>
) uniformly distributed along entire margin in the specimen from the
<typeStatus id="1609F9B2FFF0FFAFFEC1FCA0FE586973" box="[362,462,781,803]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="C90D4710FFF0FFAFFF3AFCE1FD816F7D" blockId="18.[145,761,844,1326]" pageId="18" pageNumber="19">
<emphasis id="FBC69B02FFF0FFAFFF3AFCE1FECD6932" box="[145,347,844,866]" italics="true" pageId="18" pageNumber="19">Buccal armature</emphasis>
: Muscular portion of buccal mass much larger than length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass (
<figureCitation id="51895B95FFF0FFAFFE9AFC05FE1069ED" box="[305,390,936,958]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8A</figureCitation>
) bearing numerous jaw rodlets (
<figureCitation id="51895B95FFF0FFAFFF32FC6AFF7A698C" box="[153,236,967,989]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8B</figureCitation>
). Rodlets narrow with short base and evenly curved, with single, acutely pointed apex. Radula broad, nearly as wide as long (
<figureCitation id="51895B95FFF0FFAFFE77FBA9FDBC6E49" box="[476,554,1028,1050]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8C</figureCitation>
). Radular formula of
<typeStatus id="1609F9B2FFF0FFAFFF05FB8EFE856E69" box="[174,275,1059,1081]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(CASIZ
<date id="BD0C61D0FFF0FFAFFED0FB8EFE476E69" box="[379,465,1059,1081]" pageId="18" pageNumber="19" value="70-10-19">191070</date>
) 66 × 75.0.75. Rachidian row of teeth absent (
<figureCitation id="51895B95FFF0FFAFFED4FBECFE456E06" box="[383,467,1089,1111]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8D</figureCitation>
). Innermost lateral teeth having three triangular denticles on inner side of bifid primary cusp. Denticles absent from outer side of tooth. Next several laterals and middle lateral teeth (
<figureCitation id="51895B95FFF0FFAFFD0AFB30FD676EE2" box="[673,753,1181,1203]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8E</figureCitation>
) with bifid cusp, lacking inner or outer denticles. Two outermost teeth having a narrower base and shorter tooth shape, with one to five rounded outer denticles (
<figureCitation id="51895B95FFF0FFAFFF32FAB5FF7E6F7D" box="[153,232,1304,1326]" captionStart="Figure 8" captionStartId="17.[164,243,1557,1579]" captionTargetBox="[228,1380,195,1517]" captionTargetId="figure@17.[228,1380,195,1517]" captionTargetPageId="17" captionText="Figure 8. Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632787/files/figure.png" pageId="18" pageNumber="19">Fig. 8F</figureCitation>
), smaller than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFF0FFAFFF3AFAFBFB7A6B68" blockId="18.[145,761,1366,1910]" lastBlockId="18.[809,1425,198,312]" pageId="18" pageNumber="19">
<emphasis id="FBC69B02FFF0FFAFFF3AFAFBFE066F3C" box="[145,400,1366,1388]" italics="true" pageId="18" pageNumber="19">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFF0FFAFFF3AFAD8FF656FDB" box="[145,243,1397,1419]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
fully mature (
<figureCitation id="51895B95FFF0FFAFFE30FAD8FE796FDA" box="[411,495,1397,1419]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="18" pageNumber="19">Fig. 4G</figureCitation>
). Ampulla thick, short, tubular and straight, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Proximal prostatic portion of vas deferens relatively long, convoluted, curved and thick and narrowing slightly as it transitions into muscular, ejaculatory portion. Ejaculatory portion relatively long, slightly curved and narrow, entering elongate, wider penial bulb. Penial bulb adjacent to straight, wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Moderately long vagina narrowing and leading to small, straight receptaculum seminis and larger spherical, thin-walled bursa copulatrix. Receptaculum seminis appressed against vagina, at base of bursa. Moderately short uterine duct emerging from vagina immediately below receptaculum, entering female gland mass near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF0FFAFFC82FEF2FB906D24" pageId="18" pageNumber="19" type="discussion">
<paragraph id="C90D4710FFF0FFAFFC82FEF2FC006B27" blockId="18.[809,1426,351,1908]" box="[809,918,351,375]" pageId="18" pageNumber="19">
<heading id="9245F07CFFF0FFAFFC82FEF2FC006B27" box="[809,918,351,375]" fontSize="10" level="4" pageId="18" pageNumber="19" reason="1">
<emphasis id="FBC69B02FFF0FFAFFC82FEF2FC006B27" box="[809,918,351,375]" italics="true" pageId="18" pageNumber="19">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF0FFAFFC82FE2DFB5169EE" blockId="18.[809,1426,351,1908]" pageId="18" pageNumber="19">
<taxonomicName id="0EB23C93FFF0FFAFFC82FE2DFB9C6BC5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,1034,384,405]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC82FE2DFB9C6BC5" box="[809,1034,384,405]" italics="true" pageId="18" pageNumber="19">Hypselodoris confetti</emphasis>
</taxonomicName>
, together with
<taxonomicName id="0EB23C93FFF0FFAFFB00FE2DFB7D6BC5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1195,1259,384,405]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFF0FFAFFB00FE2DFB7D6BC5" box="[1195,1259,384,405]" italics="true" pageId="18" pageNumber="19">H. roo</emphasis>
</taxonomicName>
, has often been misidentified as
<taxonomicName id="0EB23C93FFF0FFAFFC49FE32FBDE6BE3" authorityName="Rudman" authorityYear="1977" box="[994,1096,414,436]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC49FE32FBDE6BE3" box="[994,1096,414,436]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFF0FFAFFBFCFE33FA9E6BE4" author="Rudman WB" box="[1111,1288,414,436]" journalOrPublisher="Sydney: Australian Museum" pageId="18" pageNumber="19" publicationUrl="http://www.seaslugforum.net/find/hypskang" refId="ref44958" refString="Rudman WB. 1999 b. Hypselodoris kanga Rudman, 1977. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypskang" title="Hypselodoris kanga Rudman, 1977" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999b</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFF0FFAFFABFFE33FC216B82" author="Debelius H &amp; Kuiter R." journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="18" pageNumber="19" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
). The distinguishing features separating
<taxonomicName id="0EB23C93FFF0FFAFFC82FE71FC326BA1" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,932,476,497]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC82FE71FC326BA1" box="[809,932,476,497]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFF0FFAFFC49FE71FBBA6BA1" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[994,1068,476,497]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="roo">
<emphasis id="FBC69B02FFF0FFAFFC49FE71FBBA6BA1" box="[994,1068,476,497]" italics="true" pageId="18" pageNumber="19">H. roo</emphasis>
</taxonomicName>
are discussed in the remarks after the description of the latter species.
<taxonomicName id="0EB23C93FFF0FFAFFB54FE57FCEE687E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFB54FE57FCEE687E" italics="true" pageId="18" pageNumber="19">Hypselodoris confetti</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFF0FFAFFC05FDB7FB84687E" authorityName="Rudman" authorityYear="1977" box="[942,1042,537,559]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC05FDB7FB84687E" box="[942,1042,537,559]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
are geographically separated, with
<taxonomicName id="0EB23C93FFF0FFAFFC82FD95FC1A681D" authorityName="Rudman" authorityYear="1977" box="[809,908,568,589]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC82FD95FC1A681D" box="[809,908,568,589]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
being found in the Indian Ocean from
<collectingCountry id="B1A50780FFF0FFAFFA81FD95FA07681E" box="[1322,1425,568,590]" name="Tanzania" pageId="18" pageNumber="19">Tanzania</collectingCountry>
to
<collectingCountry id="B1A50780FFF0FFAFFCEFFDFBFC3C683C" box="[836,938,598,620]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
(
<bibRefCitation id="AD233AE1FFF0FFAFFC12FDFBFB0F683C" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[953,1177,598,620]" journalOrPublisher="Jacksonville: New World Press" pageId="18" pageNumber="19" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFF0FFAFFB8BFDFAFBC1683B" box="[1056,1111,598,620]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
, 2015
</bibRefCitation>
) and
<taxonomicName id="0EB23C93FFF0FFAFFB71FDFAFADA683B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1242,1356,598,620]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFB71FDFAFADA683B" box="[1242,1356,598,620]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
being restricted to the western Pacific (present study). Several consistent differences permit the separation of
<taxonomicName id="0EB23C93FFF0FFAFFA89FD39FA0668F9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1314,1424,660,681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFA89FD39FA0668F9" box="[1314,1424,660,681]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
and
<emphasis id="FBC69B02FFF0FFAFFCF6FD1EFB2D6897" box="[861,1211,690,712]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="0EB23C93FFF0FFAFFCF6FD1EFC5C6897" authorityName="Rudman" authorityYear="1977" box="[861,970,690,712]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">H. kanga</taxonomicName>
.
<taxonomicName id="0EB23C93FFF0FFAFFC7DFD1FFB2D6897" authorityName="Rudman" authorityYear="1977" box="[982,1211,690,711]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">Hypselodoris kanga</taxonomicName>
</emphasis>
has bluish purple lines on the notum and sides of body that are absent in
<taxonomicName id="0EB23C93FFF0FFAFFC82FD5DFC0F6955" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,921,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC82FD5DFC0F6955" box="[809,921,752,773]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
, and
<taxonomicName id="0EB23C93FFF0FFAFFC7CFD5DFBD16955" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[983,1095,752,773]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC7CFD5DFBD16955" box="[983,1095,752,773]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
has additional black spots on the notum and marginal blue spots that are absent in
<taxonomicName id="0EB23C93FFF0FFAFFC82FC83FC1B6912" authorityName="Rudman" authorityYear="1977" box="[809,909,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC82FC83FC1B6912" box="[809,909,813,835]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
. In our molecular analysis (
<figureCitation id="51895B95FFF0FFAFFB69FC80FA9A6913" box="[1218,1292,813,835]" captionStart="Figure 35" captionStartId="56.[146,226,1818,1840]" captionTargetBox="[210,1362,196,1775]" captionTargetId="figure@56.[209,1364,195,1777]" captionTargetPageId="56" captionText="Figure 35. Bayesian inference and maximum likelihood tree of Hypselodoris, indicating support values for each node. Bootstrap values are on the left and Bayesian posterior probablilities on the right." httpUri="https://zenodo.org/record/2632866/files/figure.png" pageId="18" pageNumber="19">Fig. 35</figureCitation>
),
<taxonomicName id="0EB23C93FFF0FFAFFA8BFC83FA076912" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1312,1425,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFA8BFC83FA076912" box="[1312,1425,813,835]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
is sister to a clade that includes
<taxonomicName id="0EB23C93FFF0FFAFFB3DFCE1FC5D69D0" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFF0FFAFFB3DFCE1FA996931" box="[1174,1295,844,865]" italics="true" pageId="18" pageNumber="19">H. zephyra</emphasis>
<bibRefCitation id="AD233AE1FFF0FFAFFABDFCE1FC5D69D0" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFF0FFAFFBAEFCC6FBDC69D0" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1029,1098,875,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFF0FFAFFBAEFCC6FBDC69D0" box="[1029,1098,875,896]" italics="true" pageId="18" pageNumber="19">H. roo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFF0FFAFFBF9FCC6FB3169D0" box="[1106,1191,875,896]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, and
<emphasis id="FBC69B02FFF0FFAFFB4FFCC6FA07692F" box="[1252,1425,874,896]" italics="true" pageId="18" pageNumber="19">H. nigrolineata</emphasis>
is sister to these three species. Molecular samples of
<taxonomicName id="0EB23C93FFF0FFAFFC82FC05FC1B69ED" authorityName="Rudman" authorityYear="1977" box="[809,909,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC82FC05FC1B69ED" box="[809,909,936,957]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
were not available for study.
</paragraph>
<paragraph id="C90D4710FFF0FFAFFCEAFC6BFA976C30" blockId="18.[809,1426,351,1908]" pageId="18" pageNumber="19">
The shape of the jaw rodlets and radular teeth is also similar in
<taxonomicName id="0EB23C93FFF0FFAFFC34FC4BFB8269AA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[927,1044,997,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC34FC4BFB8269AA" box="[927,1044,997,1019]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFF0FFAFFBE6FC4BFB2369AA" authorityName="Rudman" authorityYear="1977" box="[1101,1205,997,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFBE6FC4BFB2369AA" box="[1101,1205,997,1019]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
, but the number of teeth differs markedly.In
<taxonomicName id="0EB23C93FFF0FFAFFB94FBA9FB386E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1087,1198,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFB94FBA9FB386E49" box="[1087,1198,1028,1049]" italics="true" pageId="18" pageNumber="19">H.confetti</emphasis>
</taxonomicName>
, the radular formula is 66 × 75.0.75, whereas in
<taxonomicName id="0EB23C93FFF0FFAFFBF9FB8EFB2E6E67" authorityName="Rudman" authorityYear="1977" box="[1106,1208,1058,1080]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFBF9FB8EFB2E6E67" box="[1106,1208,1058,1080]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
the radula formula was 107 × 93.0.93 (
<bibRefCitation id="AD233AE1FFF0FFAFFBA3FBECFB266E06" author="Rudman WB" box="[1032,1200,1089,1111]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFF0FFAFFB4CFBEFFACB6E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1255,1373,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFB4CFBEFFACB6E06" box="[1255,1373,1089,1111]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
, the innermost lateral tooth has three denticles on the inner side of the two primary cusps, whereas in
<taxonomicName id="0EB23C93FFF0FFAFFA8FFBD2FA076EC3" authorityName="Rudman" authorityYear="1977" box="[1316,1425,1150,1172]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFA8FFBD2FA076EC3" box="[1316,1425,1150,1172]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
there is only a single inner denticle (
<bibRefCitation id="AD233AE1FFF0FFAFFB6BFB30FAF56EE2" author="Rudman WB" box="[1216,1379,1181,1203]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFF0FFAFFC82FB11FC196E81" authorityName="Rudman" authorityYear="1977" box="[809,911,1212,1233]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC82FB11FC196E81" box="[809,911,1212,1233]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
, the remaining teeth except for the outermost two teeth all lack denticles other than the two primary cusps. In
<taxonomicName id="0EB23C93FFF0FFAFFC0AFB57FB876F5E" authorityName="Rudman" authorityYear="1977" box="[929,1041,1273,1295]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFC0AFB57FB876F5E" box="[929,1041,1273,1295]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
, the second to fifth teeth have three to five denticles on their outer margin. From the sixth to the 82nd tooth there are no denticles, and the two outermost teeth have a single denticle. In the radula of
<emphasis id="FBC69B02FFF0FFAFFC37FAD9FBD96FD9" box="[924,1103,1396,1417]" italics="true" pageId="18" pageNumber="19">H. nigrolineata</emphasis>
, the innermost tooth has a single inner denticle and the remaining teeth are all devoid of denticles (
<bibRefCitation id="AD233AE1FFF0FFAFFBA1FA1CFB396F96" author="Rudman WB" box="[1034,1199,1457,1479]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFF0FFAFFB4FFA1FFAC96F96" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1252,1375,1457,1479]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="zephyra">
<emphasis id="FBC69B02FFF0FFAFFB4FFA1FFAC96F96" box="[1252,1375,1457,1479]" italics="true" pageId="18" pageNumber="19">H. zephyra</emphasis>
</taxonomicName>
, the innermost tooth has a single inner denticle and most of the remaining teeth entirely lack denticles other than the two primary cusps (
<bibRefCitation id="AD233AE1FFF0FFAFFB99F9A0FAC46C72" author="Gosliner T &amp; Johnson, R." box="[1074,1362,1549,1571]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
). The outermost teeth have three to six denticles, and the two adjacent teeth have only a single denticle.
</paragraph>
<paragraph id="C90D4710FFF0FFAFFCEAF9C4FB906D24" blockId="18.[809,1426,351,1908]" pageId="18" pageNumber="19">
The reproductive system of
<taxonomicName id="0EB23C93FFF0FFAFFB2CF9C7FA966C2E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1159,1280,1641,1663]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFB2CF9C7FA966C2E" box="[1159,1280,1641,1663]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="0EB23C93FFF0FFAFFC25F925FB8A6CCD" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[910,1052,1672,1693]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFF0FFAFFC25F925FB8A6CCD" box="[910,1052,1672,1693]" italics="true" pageId="18" pageNumber="19">H. krakatoa</emphasis>
</taxonomicName>
,
<emphasis id="FBC69B02FFF0FFAFFB80F925FB776CCD" box="[1067,1249,1672,1693]" italics="true" pageId="18" pageNumber="19">H. nigrolineata</emphasis>
and
<taxonomicName id="0EB23C93FFF0FFAFFAB5F925FA1D6CCD" authorityName="Rudman" authorityYear="1977" box="[1310,1419,1672,1693]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kanga">
<emphasis id="FBC69B02FFF0FFAFFAB5F925FA1D6CCD" box="[1310,1419,1672,1693]" italics="true" pageId="18" pageNumber="19">H. kanga</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFF0FFAFFCE0F90AFC516CEB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[843,967,1702,1724]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFCE0F90AFC516CEB" box="[843,967,1702,1724]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
, the receptaculum seminis is situated immediately adjacent to the bursa copulatrix, whereas in the other three species it is situated more proximally on the vagina (
<bibRefCitation id="AD233AE1FFF0FFAFFC49F8AFFB076D48" author="Rudman WB" box="[994,1169,1794,1816]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="351 - 397" part="61" refId="ref44619" refString="Rudman WB. 1977. Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific. Zoological Journal of the Linnean Society 61: 351 - 397." title="Chromodorid opisthobranch Mollusca from East Africa and the tropical West Pacific" type="journal article" year="1977">Rudman, 1977</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFF0FFAFFB0AF8AFFCF66D66" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="18" pageNumber="19" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson 1999</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFF0FFAFFC3EF88FFB9C6D66" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[917,1034,1825,1847]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="confetti">
<emphasis id="FBC69B02FFF0FFAFFC3EF88FFB9C6D66" box="[917,1034,1825,1847]" italics="true" pageId="18" pageNumber="19">H. confetti</emphasis>
</taxonomicName>
, the ejaculatory portion of the vas deferens is shorter and less convoluted than in the other three species.
</paragraph>
</subSubSection>
</treatment>
</document>

458
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<mods:number id="DE587E9FA198DCD39662AB934C67FC88">2018-09-14</mods:number>
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<mods:classification id="B5101DE69EECA9B7D8708ADED5F4707F">journal article</mods:classification>
<mods:identifier id="88C2213B5CBCC48DD431AF6DE3CEAFFF" type="CLB-Dataset">27363</mods:identifier>
<mods:identifier id="C6E5E7E9283DB57547C89567C4300906" type="DOI">10.1093/zoolinnean/zly048</mods:identifier>
<mods:identifier id="BEECF9D1D6F1EA6F1951B1F714487DFD" type="GBIF-Dataset">49bea467-fac6-484a-9902-05483dec9a52</mods:identifier>
<mods:identifier id="AD1CF7AD81B68B2A2CBCDA827E0044F1" type="Zenodo-Dep">2632578</mods:identifier>
<mods:identifier id="8A1D8EB10199E498241931224654A474" type="ZooBank">urn:lsid:zoobank.org:pub:F0065FD2-417C</mods:identifier>
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<treatment id="411BF606FFEFFFB2FC76FC51FABE6F97" ID-DOI="http://doi.org/10.5281/zenodo.5980670" ID-GBIF-Taxon="156146726" ID-Zenodo-Dep="5980670" LSID="urn:lsid:plazi:treatment:411BF606FFEFFFB2FC76FC51FABE6F97" httpUri="http://treatment.plazi.org/id/411BF606FFEFFFB2FC76FC51FABE6F97" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="81A8149BFFEFFFB0FC76FC51FAB36E62" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="C90D4710FFEFFFB0FC76FC51FAB36E62" blockId="13.[952,1317,1020,1113]" pageId="13" pageNumber="14">
<taxonomicName id="0EB23C93FFEFFFB0FC76FC51FB2D6E60" ID-CoL="3P32Z" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="15" phylum="Mollusca" rank="species" species="cerisae" status="sp. nov.">
<heading id="9245F07CFFEFFFB0FC76FC51FA966E42" allCaps="true" bold="true" box="[989,1280,1020,1043]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="FBC69B02FFEFFFB0FC76FC51FA966E42" bold="true" box="[989,1280,1020,1043]" italics="true" pageId="13" pageNumber="14">HYPSELODORIS CERISAE</emphasis>
</heading>
<heading id="9245F07CFFEFFFB0FC13FBB7FB2D6E60" allCaps="true" box="[952,1211,1050,1074]" centered="true" fontSize="9" level="3" pageId="13" pageNumber="14" reason="4">GOSLINER &amp; JOHNSON</heading>
</taxonomicName>
<emphasis id="FBC69B02FFEFFFB0FB69FBB0FAB36E62" bold="true" box="[1218,1317,1050,1074]" pageId="13" pageNumber="14">SP. NOV.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FC56FBECFB496E09" box="[1021,1247,1089,1113]" pageId="13" pageNumber="14" type="description">
<paragraph id="C90D4710FFEFFFB0FC56FBECFB496E09" blockId="13.[952,1317,1020,1113]" box="[1021,1247,1089,1113]" pageId="13" pageNumber="14">
<heading id="9245F07CFFEFFFB0FC56FBECFB496E09" allCaps="true" box="[1021,1247,1089,1113]" centered="true" fontSize="10" level="1" pageId="13" pageNumber="14" reason="1">(FIGS 1C, 2B, 4E, 6)</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FC90FBD2FBFB6EE3" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="C90D4710FFEFFFB0FC90FBD2FBFB6EE3" blockId="13.[827,1444,1150,1203]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FC90FBD2FCEB6EC4" box="[827,893,1151,1172]" italics="true" pageId="13" pageNumber="14">LSID</emphasis>
:
<uri id="BD234B12FFEFFFB0FC3AFBD3FBFB6EE3" pageId="13" pageNumber="14">
urn:LSID:zoobank.org:act:
<uuid id="BD147DC5FFEFFFB0FB67FBD3FBFB6EE3" pageId="13" pageNumber="14">A53FA948-8E58- 47A8-9BA3-681900B19537</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FC90FB7CFBD56F74" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="C90D4710FFEFFFB0FC90FB7CFBD56F74" blockId="13.[827,1442,1233,1316]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FC90FB7CFBE36EB6" box="[827,1141,1233,1254]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="0EB23C93FFEFFFB0FC90FB7CFBFF6EB6" ID-CoL="3P348" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,1129,1233,1254]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="krakatoa">Hypselodoris krakatoa</taxonomicName>
?
</emphasis>
misidentification, not
<taxonomicName id="0EB23C93FFEFFFB0FC90FB5DFA9E6F55" ID-CoL="3P348" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,1288,1264,1286]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEFFFB0FC90FB5DFC5F6F55" box="[827,969,1264,1285]" italics="true" pageId="13" pageNumber="14">H. krakatoa</emphasis>
<bibRefCitation id="AD233AE1FFEFFFB0FC7AFB5DFA9E6F55" author="Gosliner T &amp; Johnson, R." box="[977,1288,1264,1286]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="13" pageNumber="14" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFEFFFB0FABEFB5DFCE46F74" author="Garthwaite T." journalOrPublisher="Sydney: Australian Museum" pageId="13" pageNumber="14" publicationUrl="http://www.seaslugforum.net/find/7246" refId="ref42208" refString="Garthwaite T. 2002. Hypselodoris krakatoa? from Taiwan. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 7246" title="Hypselodoris krakatoa? from Taiwan" type="url" volumeTitle="Sea slug forum" year="2002">Garthwaite, 2002</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFEFFFB0FCD5FAA3FBAA6F74" author="Masayoshi N." box="[894,1084,1294,1316]" journalOrPublisher="Sydney: Australian Museum" pageId="13" pageNumber="14" publicationUrl="http://www.seaslugforum.net/find/7370" refId="ref43725" refString="Masayoshi N. 2002. Re: Hypselodoris krakatoa? from Taiwan. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 7370" title="Re: Hypselodoris krakatoa? from Taiwan" type="url" volumeTitle="Sea slug forum" year="2002">Masayoshi, 2002</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FC90FAECFB516D48" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="C90D4710FFEFFFB0FC90FAECFC726F09" blockId="13.[827,1443,1345,1460]" box="[827,996,1345,1369]" pageId="13" pageNumber="14">
<heading id="9245F07CFFEFFFB0FC90FAECFC726F09" box="[827,996,1345,1369]" fontSize="10" level="4" pageId="13" pageNumber="14" reason="1">
<emphasis id="FBC69B02FFEFFFB0FC90FAECFC726F09" box="[827,996,1345,1369]" italics="true" pageId="13" pageNumber="14">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90FACCFBDA6FE4" blockId="13.[827,1443,1345,1460]" pageId="13" pageNumber="14">
<materialsCitation id="79DA4D4DFFEFFFB0FC90FACCFBDA6FE4" ID-GBIF-Occurrence="2235579074" collectingDate="2008-08-09" collectionCode="CASIZ" collectorName="C. Chen." country="Taiwan" elevation="10" location="Long Dong" municipality="County" pageId="13" pageNumber="14" specimenCode="CASIZ 178350" specimenCount="1" stateProvince="Taipei" typeStatus="holotype">
<typeStatus id="1609F9B2FFEFFFB0FC90FACCFC356F26" box="[827,931,1377,1398]" pageId="13" pageNumber="14" type="holotype">
<emphasis id="FBC69B02FFEFFFB0FC90FACCFC356F26" box="[827,931,1377,1398]" italics="true" pageId="13" pageNumber="14">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFEFFFB0FC1EFACCFBF56F27" box="[949,1123,1377,1399]" collectionCode="CASIZ" pageId="13" pageNumber="14">CASIZ 178350</specimenCode>
, subsampled for molecular study,
<location id="CC6D11CBFFEFFFB0FC2EFA2DFB956FC5" LSID="urn:lsid:plazi:treatment:411BF606FFEFFFB2FC76FC51FABE6F97:CC6D11CBFFEFFFB0FC2EFA2DFB956FC5" box="[901,1027,1408,1429]" country="Taiwan" municipality="County" name="Long Dong" pageId="13" pageNumber="14" stateProvince="Taipei">Long Dong</location>
,
<collectingRegion id="0B7689F2FFEFFFB0FBA6FA2DFBC26FC6" box="[1037,1108,1408,1430]" country="Taiwan" name="Taipei" pageId="13" pageNumber="14">Taipei</collectingRegion>
<collectingMunicipality id="2969DD6AFFEFFFB0FBF7FA2DFB396FC5" box="[1116,1199,1408,1429]" pageId="13" pageNumber="14">County</collectingMunicipality>
,
<collectingCountry id="B1A50780FFEFFFB0FB12FA2DFA996FC6" box="[1209,1295,1408,1430]" name="Taiwan" pageId="13" pageNumber="14">Taiwan</collectingCountry>
,
<elevation id="429FA023FFEFFFB0FAB0FA2DFAC26FC5" box="[1307,1364,1408,1429]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="10.0">
<quantity id="0E4AEAF5FFEFFFB0FAB0FA2DFAC26FC5" box="[1307,1364,1408,1429]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="10.0">10 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEFFFB0FC90FA32FC766FE4" box="[827,992,1439,1460]" pageId="13" pageNumber="14" value="2008-08-09">
<date id="BD0C61D0FFEFFFB0FC90FA32FC766FE4" box="[827,992,1439,1460]" pageId="13" pageNumber="14" value="2008-08-09">9 August 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFEFFFB0FC40FA32FBDA6FE4" box="[1003,1100,1438,1460]" pageId="13" pageNumber="14">C. Chen.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90FA7EFBCB6C33" blockId="13.[827,1443,1490,1635]" pageId="13" pageNumber="14">
<materialsCitation id="79DA4D4DFFEFFFB0FC90FA7EFA366C57" ID-GBIF-Occurrence="2235579080" collectingDate="2001-07-01" collectionCode="CASIZ" collectorName="Nishina Masayoshi" country="Japan" elevation="7" location="Sokoda" municipality="Hachijo Island" pageId="13" pageNumber="14" specimenCode="CASIZ 219744" specimenCount="1" typeStatus="paratype">
<typeStatus id="1609F9B2FFEFFFB0FC90FA7EFC276FB8" box="[827,945,1491,1512]" pageId="13" pageNumber="14" type="paratype">
<emphasis id="FBC69B02FFEFFFB0FC90FA7EFC276FB8" box="[827,945,1491,1512]" italics="true" pageId="13" pageNumber="14">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFEFFFB0FC6EFA7EFBE06FB8" box="[965,1142,1491,1512]" collectionCode="CASIZ" pageId="13" pageNumber="14">CASIZ 219744</specimenCode>
,
<location id="CC6D11CBFFEFFFB0FB2FFA7FFB486FB8" LSID="urn:lsid:plazi:treatment:411BF606FFEFFFB2FC76FC51FABE6F97:CC6D11CBFFEFFFB0FB2FFA7FFB486FB8" box="[1156,1246,1490,1512]" country="Japan" municipality="Hachijo Island" name="Sokoda" pageId="13" pageNumber="14">Sokoda</location>
,
<collectingMunicipality id="2969DD6AFFEFFFB0FB40FA7FFA086FB8" box="[1259,1438,1490,1512]" pageId="13" pageNumber="14">Hachijo Island</collectingMunicipality>
,
<collectingCountry id="B1A50780FFEFFFB0FC90FA5FFC106C57" box="[827,902,1522,1543]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
,
<elevation id="429FA023FFEFFFB0FC39FA5FFC566C57" box="[914,960,1522,1543]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">
<quantity id="0E4AEAF5FFEFFFB0FC39FA5FFC566C57" box="[914,960,1522,1543]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="m" value="7.0">7 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEFFFB0FBB0FA5CFB3F6C56" box="[1051,1193,1521,1543]" pageId="13" pageNumber="14" value="2001-07-01">
<date id="BD0C61D0FFEFFFB0FBB0FA5CFB3F6C56" box="[1051,1193,1521,1543]" pageId="13" pageNumber="14" value="2001-07-01">1 July 2001</date>
</collectingDate>
,
<collectorName id="644722C6FFEFFFB0FB1DFA5CFA366C57" box="[1206,1440,1521,1543]" pageId="13" pageNumber="14">Nishina Masayoshi</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFEFFFB0FC90F9BDFBCC6C33" ID-GBIF-Occurrence="2235579044" collectingDate="2007-10-03" collectionCode="CASIZ" collectorName="T. Gosliner" country="Malaysia" elevation="20" location="Rayner's Rock" municipality="Pulau Aur" pageId="13" pageNumber="14" specimenCode="CASIZ 175726" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFEFFFB0FC90F9BDFC636C76" box="[827,1013,1552,1574]" collectionCode="CASIZ" pageId="13" pageNumber="14">CASIZ 175726</specimenCode>
, subsampled for molecular study,
<location id="CC6D11CBFFEFFFB0FC90F982FC496C14" LSID="urn:lsid:plazi:treatment:411BF606FFEFFFB2FC76FC51FABE6F97:CC6D11CBFFEFFFB0FC90F982FC496C14" box="[827,991,1582,1604]" country="Malaysia" municipality="Pulau Aur" name="Rayner's Rock" pageId="13" pageNumber="14">Rayner’s Rock</location>
, off
<collectingMunicipality id="2969DD6AFFEFFFB0FBA5F983FB136C14" box="[1038,1157,1582,1604]" pageId="13" pageNumber="14">Pulau Aur</collectingMunicipality>
,
<collectingCountry id="B1A50780FFEFFFB0FB3BF983FB6D6C14" box="[1168,1275,1582,1604]" name="Malaysia" pageId="13" pageNumber="14">Malaysia</collectingCountry>
,
<elevation id="429FA023FFEFFFB0FAADF982FAA96C14" box="[1286,1343,1583,1604]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="20.0">
<quantity id="0E4AEAF5FFEFFFB0FAADF982FAA96C14" box="[1286,1343,1583,1604]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="20.0">20 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFEFFFB0FA3EF982FC426C32" pageId="13" pageNumber="14" value="2007-10-03">
<date id="BD0C61D0FFEFFFB0FA3EF982FC426C32" pageId="13" pageNumber="14" value="2007-10-03">3 October 2007</date>
</collectingDate>
,
<collectorName id="644722C6FFEFFFB0FC76F9E3FBCC6C33" box="[989,1114,1613,1635]" pageId="13" pageNumber="14">T. Gosliner</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90F9D1FC696CE0" blockId="13.[827,1443,1660,1712]" pageId="13" pageNumber="14">
<emphasis id="FBC69B02FFEFFFB0FC90F9D1FB3A6CC1" box="[827,1196,1660,1681]" italics="true" pageId="13" pageNumber="14">Comparative material examined:</emphasis>
<taxonomicName id="0EB23C93FFEFFFB0FB1CF9D1FC6D6CE0" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEFFFB0FB1CF9D1FAAB6CC1" box="[1207,1341,1660,1681]" italics="true" pageId="13" pageNumber="14">H. krakatoa</emphasis>
<bibRefCitation id="AD233AE1FFEFFFB0FAE9F9D1FC6D6CE0" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="13" pageNumber="14" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90F94EFC436CAA" blockId="13.[827,1223,1762,1816]" box="[827,981,1762,1786]" pageId="13" pageNumber="14">
<heading id="9245F07CFFEFFFB0FC90F94EFC436CAA" box="[827,981,1762,1786]" fontSize="10" level="4" pageId="13" pageNumber="14" reason="1">
<emphasis id="FBC69B02FFEFFFB0FC90F94EFC436CAA" box="[827,981,1762,1786]" italics="true" pageId="13" pageNumber="14">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90F8AEFB516D48" blockId="13.[827,1223,1762,1816]" box="[827,1223,1794,1816]" pageId="13" pageNumber="14">
Long Dong,
<collectingRegion id="0B7689F2FFEFFFB0FC69F8AFFB9F6D48" box="[962,1033,1794,1816]" country="Taiwan" name="Taipei" pageId="13" pageNumber="14">Taipei</collectingRegion>
County,
<collectingCountry id="B1A50780FFEFFFB0FBC0F8AFFB556D48" box="[1131,1219,1794,1816]" name="Taiwan" pageId="13" pageNumber="14">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEFFFB0FC90F898FA8B6D3B" pageId="13" pageNumber="14" type="distribution">
<paragraph id="C90D4710FFEFFFB0FC90F898FBEE6D1D" blockId="13.[827,1309,1845,1899]" box="[827,1144,1845,1869]" pageId="13" pageNumber="14">
<heading id="9245F07CFFEFFFB0FC90F898FBEE6D1D" box="[827,1144,1845,1869]" fontSize="10" level="4" pageId="13" pageNumber="14" reason="1">
<emphasis id="FBC69B02FFEFFFB0FC90F898FBEE6D1D" box="[827,1144,1845,1869]" italics="true" pageId="13" pageNumber="14">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEFFFB0FC90F8FBFA8B6D3B" blockId="13.[827,1309,1845,1899]" box="[827,1309,1877,1899]" pageId="13" pageNumber="14">
Known from
<collectingCountry id="B1A50780FFEFFFB0FC64F8FBFB806D3B" box="[975,1046,1878,1899]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
,
<collectingCountry id="B1A50780FFEFFFB0FB8BF8F8FBE26D3B" box="[1056,1140,1877,1899]" name="Taiwan" pageId="13" pageNumber="14">Taiwan</collectingCountry>
and
<collectingCountry id="B1A50780FFEFFFB0FB06F8F8FA8F6D3B" box="[1197,1305,1877,1899]" name="Malaysia" pageId="13" pageNumber="14">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="9DCD1798FFECFFB3FF24F9A3FC3B6C0F" httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="14" pageNumber="15" startId="14.[143,221,1550,1572]" targetBox="[207,1359,195,1510]" targetPageId="14">
<paragraph id="C90D4710FFECFFB3FF24F9A3FC3B6C0F" blockId="14.[143,1424,1550,1631]" pageId="14" pageNumber="15">
<emphasis id="FBC69B02FFECFFB3FF24F9A3FF616C73" bold="true" box="[143,247,1550,1572]" pageId="14" pageNumber="15">Figure 6.</emphasis>
<taxonomicName id="0EB23C93FFECFFB3FEA8F9A3FD506C74" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[259,710,1550,1572]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="cerisae" status="sp. nov.">
<emphasis id="FBC69B02FFECFFB3FEA8F9A3FE676C73" bold="true" box="[259,497,1550,1572]" italics="true" pageId="14" pageNumber="15">Hypselodoris cerisae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFECFFB3FD67F9A2FCB56C74" box="[716,803,1551,1572]" pageId="14" pageNumber="15" rank="species">
<emphasis id="FBC69B02FFECFFB3FD67F9A2FCB56C74" bold="true" box="[716,803,1551,1572]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726.
</paragraph>
</caption>
<subSubSection id="81A8149BFFECFFB3FF3AF93DFEE26D12" pageId="14" pageNumber="15" type="etymology">
<paragraph id="C90D4710FFECFFB3FF3AF93DFE846CF8" blockId="14.[145,761,1680,1858]" box="[145,274,1680,1704]" pageId="14" pageNumber="15">
<heading id="9245F07CFFECFFB3FF3AF93DFE846CF8" box="[145,274,1680,1704]" fontSize="10" level="4" pageId="14" pageNumber="15" reason="1">
<emphasis id="FBC69B02FFECFFB3FF3AF93DFE846CF8" box="[145,274,1680,1704]" italics="true" pageId="14" pageNumber="15">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFECFFB3FF3AF91CFEE26D12" blockId="14.[145,761,1680,1858]" pageId="14" pageNumber="15">
<taxonomicName id="0EB23C93FFECFFB3FF3AF91CFE156C97" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,387,1713,1735]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFECFFB3FF3AF91CFE156C97" box="[145,387,1713,1735]" italics="true" pageId="14" pageNumber="15">Hypselodoris cerisae</emphasis>
</taxonomicName>
is named for Cerise Chen, who first found this species in
<collectingCountry id="B1A50780FFECFFB3FE67F97DFDB26CB6" box="[460,548,1744,1766]" name="Taiwan" pageId="14" pageNumber="15">Taiwan</collectingCountry>
and collected the
<typeStatus id="1609F9B2FFECFFB3FF3AF943FF6D6D54" box="[145,251,1774,1796]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
specimen. Cerise has a keen interest in octocoral biology but also a more general interest in marine biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFECFFB2FC82F93CFE196D27" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="C90D4710FFECFFB3FC82F93CFC236CF8" blockId="14.[809,1425,1681,1858]" box="[809,949,1681,1704]" pageId="14" pageNumber="15">
<heading id="9245F07CFFECFFB3FC82F93CFC236CF8" box="[809,949,1681,1704]" fontSize="10" level="4" pageId="14" pageNumber="15" reason="1">
<emphasis id="FBC69B02FFECFFB3FC82F93CFC236CF8" box="[809,949,1681,1704]" italics="true" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFECFFB2FC82F91CFD566BE3" blockId="14.[809,1425,1681,1858]" lastBlockId="15.[163,779,198,435]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="FBC69B02FFECFFB3FC82F91CFBAE6C96" box="[809,1080,1713,1734]" italics="true" pageId="14" pageNumber="15">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFECFFB3FA82F91CFA1F6C96" box="[1321,1417,1713,1735]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="14" pageNumber="15">Fig. 1C</figureCitation>
) moderately large, reaching
<quantity id="0E4AEAF5FFECFFB3FBF2F962FB586CB5" box="[1113,1230,1743,1765]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="14" pageNumber="15" unit="mm" value="27.5" valueMax="35.0" valueMin="20.0">20–35 mm</quantity>
in length. Colour pattern complex with golden honey ground colour. Notum ornamented with purple on anterior, lateral and posterior margins. Irregular dark brown to black longitudinal lines present on notum, with few opaque white spots along sides of dark lines. Gill pocket well elevated from notum. Seven to nine narrow, thin, unipinnate gill branches on notum. Gill branches red–orange on upper outer surface, opaque white internally and externally at base. Rhinophores uniformly bright red–orange except at opaque white apex. Rhinophores with ~21 small lamellae, Edge of foot with purple marginal band.
</paragraph>
<paragraph id="C90D4710FFEDFFB2FF08FE71FDF6683C" blockId="15.[163,779,476,620]" pageId="15" pageNumber="16">
<emphasis id="FBC69B02FFEDFFB2FF08FE71FEDE6BA1" box="[163,328,476,497]" italics="true" pageId="15" pageNumber="16">Mantle glands</emphasis>
: Subcutaneous mantle glands (
<figureCitation id="51895B95FFEDFFB2FD1AFE71FC956BA1" box="[689,771,476,498]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="15" pageNumber="16">Fig. 2B</figureCitation>
) uniformly distributed along entire margin in the specimens from
<collectingCountry id="B1A50780FFEDFFB2FF70FDB4FEBD687F" box="[219,299,537,559]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
(CASIZ 178350) and
<collectingCountry id="B1A50780FFEDFFB2FDBAFDB7FDC5687F" box="[529,595,538,559]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
(CASIZ 219744), but absent from some portions of the lateral edges in the specimen from
<collectingCountry id="B1A50780FFEDFFB2FEEEFDFBFE31683C" box="[325,423,598,620]" name="Malaysia" pageId="15" pageNumber="16">Malaysia</collectingCountry>
(CASIZ 175726).
</paragraph>
<paragraph id="C90D4710FFEDFFB2FF08FD38FDEF6E27" blockId="15.[163,780,661,1143]" pageId="15" pageNumber="16">
<emphasis id="FBC69B02FFEDFFB2FF08FD38FEC068FB" box="[163,342,661,683]" italics="true" pageId="15" pageNumber="16">Buccal armature</emphasis>
: Muscular portion of buccal mass slightly larger than length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass (
<figureCitation id="51895B95FFEDFFB2FF46FD5CFEA86957" box="[237,318,753,775]" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6A</figureCitation>
) bearing numerous jaw rodlets (
<figureCitation id="51895B95FFEDFFB2FD1FFD5CFF206975" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6B, C</figureCitation>
. Rodlets narrow with short base and evenly curved, with single, acutely pointed apex and occasional lateral flange. Radular formula of
<typeStatus id="1609F9B2FFEDFFB2FEDEFCE0FE476933" box="[373,465,845,867]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(CASIZ 178350) 59 × 36.0.36 and
<typeStatus id="1609F9B2FFEDFFB2FF7BFCC1FEA769D1" box="[208,305,876,897]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
(CAS 175726) 52 × 56.0.56. Rachidian row of teeth absent (
<figureCitation id="51895B95FFEDFFB2FE9EFC27FE1469F0" box="[309,386,906,928]" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6D</figureCitation>
). Innermost lateral teeth having one to two triangular denticles on inner side of bifid primary cusp. Denticles absent from outer side of tooth (
<figureCitation id="51895B95FFEDFFB2FD06FC65FD6B698E" box="[685,765,968,990]" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6D</figureCitation>
). Next several laterals and middle lateral teeth (
<figureCitation id="51895B95FFEDFFB2FD1FFC4BFF236E4B" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6D, E</figureCitation>
) with bifid cusp, lacking inner or outer denticles. Outermost one to three teeth having a narrower base and shorter tooth shape, with one to five rounded outer denticles (
<figureCitation id="51895B95FFEDFFB2FEBBFBCCFECD6E27" box="[272,347,1121,1143]" captionStart="Figure 6" captionStartId="14.[143,221,1550,1572]" captionTargetBox="[207,1359,195,1510]" captionTargetId="figure@14.[207,1359,195,1510]" captionTargetPageId="14" captionText="Figure 6. Hypselodoris cerisae Gosliner &amp; Johnson sp. nov. A, entire jaw, holotype, CASIZ 178350. B, jaw rodlets, paratype, CAS 175726. C, jaw rodlets, holotype, CASIZ 178350. D, inner lateral teeth, holotype, CAS 178350. E, middle lateral teeth, holotype, CASIZ, 178350. F, outer lateral teeth, paratype, CAS175726." httpUri="https://zenodo.org/record/2632783/files/figure.png" pageId="15" pageNumber="16">Fig. 6F</figureCitation>
), smaller than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFEDFFB2FF08FB0DFE196D27" blockId="15.[163,779,1184,1911]" pageId="15" pageNumber="16">
<emphasis id="FBC69B02FFEDFFB2FF08FB0DFE346EE5" box="[163,418,1184,1205]" italics="true" pageId="15" pageNumber="16">Reproductive system</emphasis>
: Reproductive organs of the
<typeStatus id="1609F9B2FFEDFFB2FF08FB13FE926E84" box="[163,260,1214,1236]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(
<figureCitation id="51895B95FFEDFFB2FEA5FB13FECD6E84" box="[270,347,1214,1236]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="15" pageNumber="16">Fig.4E</figureCitation>
) and one
<typeStatus id="1609F9B2FFEDFFB2FE17FB12FDB46E84" box="[444,546,1215,1236]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
(CASIZ175726) fully mature and virtually identical anatomically. Ampulla thick, short, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens relatively short, convoluted, curved and thick and narrowing slightly as it transitions into muscular, ejaculatory portion. Ejaculatory portion relatively short, convoluted, narrow, entering short, wider penial bulb. Penial bulb adjacent to slightly curved, wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thinwalled receptaculum seminis. Receptaculum seminis appressed against vagina, near middle of vagina. Moderately short uterine duct emerging from vagina opposite receptaculum, entering female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFEDFFB2FC90FF6BFABE6F97" pageId="15" pageNumber="16" type="discussion">
<paragraph id="C90D4710FFEDFFB2FC90FF6BFC3E6A8E" blockId="15.[827,1444,198,1479]" box="[827,936,198,222]" pageId="15" pageNumber="16">
<heading id="9245F07CFFEDFFB2FC90FF6BFC3E6A8E" box="[827,936,198,222]" fontSize="10" level="4" pageId="15" pageNumber="16" reason="1">
<emphasis id="FBC69B02FFEDFFB2FC90FF6BFC3E6A8E" box="[827,936,198,222]" italics="true" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFEDFFB2FC90FF4AFA3269CF" blockId="15.[827,1444,198,1479]" pageId="15" pageNumber="16">
The colour pattern of
<taxonomicName id="0EB23C93FFEDFFB2FB94FF4AFB256AAC" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1087,1203,231,252]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FB94FF4AFB256AAC" box="[1087,1203,231,252]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0EB23C93FFEDFFB2FC90FEABFB6B6B4A" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,1277,261,283]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FC90FEABFC506B4A" box="[827,966,261,283]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
<bibRefCitation id="AD233AE1FFEDFFB2FC65FEA8FB6B6B4A" author="Gosliner T &amp; Johnson, R." box="[974,1277,261,283]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="15" pageNumber="16" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
, but differs in several consistent respects. In
<taxonomicName id="0EB23C93FFEDFFB2FB1FFE88FABD6B69" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1204,1323,292,314]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FB1FFE88FABD6B69" box="[1204,1323,292,314]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
, the body has much more pink to purple pigment, whereas in
<taxonomicName id="0EB23C93FFEDFFB2FC90FECFFC2B6B26" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,957,353,375]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FC90FECFFC2B6B26" box="[827,957,353,375]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
the notum is suffused with patches of burnt orange. Both species have prominent purple pigment on the anterior margin of the mantle, but in
<taxonomicName id="0EB23C93FFEDFFB2FAA0FE32FAE16BE4" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1291,1399,415,436]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FAA0FE32FAE16BE4" box="[1291,1399,415,436]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
the posterior end is also purple, whereas it is burnt orange in
<taxonomicName id="0EB23C93FFEDFFB2FCF1FE70FC726BA1" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[858,996,476,498]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FCF1FE70FC726BA1" box="[858,996,476,498]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
. The body appears lower and wider in
<taxonomicName id="0EB23C93FFEDFFB2FC90FE56FC3C6840" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,938,507,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FC90FE56FC3C6840" box="[827,938,507,528]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
than in
<taxonomicName id="0EB23C93FFEDFFB2FBA7FE56FB03685F" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1036,1173,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FBA7FE56FB03685F" box="[1036,1173,506,528]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
, but in
<taxonomicName id="0EB23C93FFEDFFB2FB45FE56FAE0685F" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1262,1398,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FB45FE56FAE0685F" box="[1262,1398,506,528]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
the mantle margin is more extensive laterally, overhanging the lateral edges of the body. There are seven to nine gill branches in
<taxonomicName id="0EB23C93FFEDFFB2FBAAFDFAFBE1683C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1025,1143,599,620]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FBAAFDFAFBE1683C" box="[1025,1143,599,620]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
and only six or seven in
<taxonomicName id="0EB23C93FFEDFFB2FC90FDDBFC5168DA" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,967,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FC90FDDBFC5168DA" box="[827,967,629,651]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
. The mantle glands are distributed in a similar manner in both species, where they are largely uniformly spaced around the entire mantle margin. In our molecular analysis (
<figureCitation id="51895B95FFEDFFB2FBC4FD7CFB2A68B7" box="[1135,1212,721,743]" captionStart="Figure 35" captionStartId="56.[146,226,1818,1840]" captionTargetBox="[210,1362,196,1775]" captionTargetId="figure@56.[209,1364,195,1777]" captionTargetPageId="56" captionText="Figure 35. Bayesian inference and maximum likelihood tree of Hypselodoris, indicating support values for each node. Bootstrap values are on the left and Bayesian posterior probablilities on the right." httpUri="https://zenodo.org/record/2632866/files/figure.png" pageId="15" pageNumber="16">Fig. 35</figureCitation>
),
<taxonomicName id="0EB23C93FFEDFFB2FB79FD7FFAD768B7" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1234,1345,722,743]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FB79FD7FFAD768B7" box="[1234,1345,722,743]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
is sister to a clade that includes
<taxonomicName id="0EB23C93FFEDFFB2FBFFFD5CFCE56974" authority="Wilson &amp; Willan, 2007" authorityName="Wilson &amp; Willan" authorityYear="2007" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="jacksoni">
<emphasis id="FBC69B02FFEDFFB2FBFFFD5CFB436955" box="[1108,1237,752,774]" italics="true" pageId="15" pageNumber="16">H. jacksoni</emphasis>
<bibRefCitation id="AD233AE1FFEDFFB2FB76FD5DFCE56974" author="Wilson N &amp; Willan, R." journalOrPublisher="Zootaxa" pageId="15" pageNumber="16" pagination="29 - 42" part="1549" refId="ref45538" refString="Wilson N, Willan, R. 2007. Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982. Zootaxa 1549: 29 - 42." title="Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982" type="journal article" year="2007">Wilson &amp; Willan, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFEDFFB2FCD5FCA2FB686974" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[894,1278,782,805]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="reidi">
<emphasis id="FBC69B02FFEDFFB2FCD5FCA2FC426973" box="[894,980,782,804]" italics="true" pageId="15" pageNumber="16">H. reidi</emphasis>
<bibRefCitation id="AD233AE1FFEDFFB2FC70FCA2FB686974" author="Gosliner T &amp; Johnson, R." box="[987,1278,783,805]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="15" pageNumber="16" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFEDFFB2FA93FCA2FAEB6973" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1336,1405,782,804]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFEDFFB2FA93FCA2FAEB6973" box="[1336,1405,782,804]" italics="true" pageId="15" pageNumber="16">H. iba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFEDFFB2FA2FFCA2FCFE6913" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
Additionally,
<taxonomicName id="0EB23C93FFEDFFB2FBA3FC83FB046912" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1032,1170,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FBA3FC83FB046912" box="[1032,1170,813,835]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFEDFFB2FB66FC83FC796931" authority="Marcus &amp; Marcus, 1970" authorityName="Marcus &amp; Marcus" authorityYear="1970" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="reginae">
<emphasis id="FBC69B02FFEDFFB2FB66FC83FAD36913" box="[1229,1349,814,835]" italics="true" pageId="15" pageNumber="16">H. reginae</emphasis>
<bibRefCitation id="AD233AE1FFEDFFB2FAE7FC83FC796931" author="Marcus EV &amp; Marcus ER" journalOrPublisher="Malacologia" pageId="15" pageNumber="16" pagination="181 - 223" part="10" refId="ref43702" refString="Marcus EV, Marcus ER. 1970. Some gastropods from Madagascar and West Mexico. Malacologia 10: 181 - 223." title="Some gastropods from Madagascar and West Mexico" type="journal article" year="1970">Marcus &amp; Marcus, 1970</bibRefCitation>
</taxonomicName>
form a trichotomy with the clade that includes
<taxonomicName id="0EB23C93FFEDFFB2FC08FCC6FB8269D0" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[931,1044,875,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FC08FCC6FB8269D0" box="[931,1044,875,896]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
. All members of this largest clade have a gill peduncle that is well elevated from the notum.
</paragraph>
<paragraph id="C90D4710FFEDFFB2FCF8FC05FADC6EA0" blockId="15.[827,1444,198,1479]" pageId="15" pageNumber="16">
The shape of the jaw rodlets and radular teeth is also similar in
<taxonomicName id="0EB23C93FFEDFFB2FC19FC6AFBB6698C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[946,1056,967,988]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FC19FC6AFBB6698C" box="[946,1056,967,988]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFEDFFB2FBF1FC6AFB74698B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1114,1250,966,988]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FBF1FC6AFB74698B" box="[1114,1250,966,988]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
, but the number of teeth per radular row differs. In the two specimens of
<taxonomicName id="0EB23C93FFEDFFB2FCF3FBA8FC5D6E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[856,971,1028,1050]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FCF3FBA8FC5D6E49" box="[856,971,1028,1050]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
(CASIZ 178350 and 175726) the radula formula was 59 × 36.0.36 and 52 × 56.0.56, respectively. In four specimens of
<taxonomicName id="0EB23C93FFEDFFB2FB9AFBEFFB2A6E06" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1073,1212,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FB9AFBEFFB2A6E06" box="[1073,1212,1089,1111]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
the radula formula was 57 × 74.0.74 (CASIZ 206801;
<figureCitation id="51895B95FFEDFFB2FB6BFBCDFB696E25" box="[1216,1279,1120,1142]" captionStart="Figure 7" captionStartId="16.[146,224,1558,1580]" captionTargetBox="[210,1362,200,1518]" captionTargetId="figure@16.[210,1362,200,1518]" captionTargetPageId="16" captionText="Figure 7. Hypseldoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632785/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
), 57 × 72.0.72 (CASIZ 177371), 51 × 66.0.66, 46 × 60.0.60 and 30 × 35.0.35 (last three from
<bibRefCitation id="AD233AE1FFEDFFB2FB02FB30FCE26E81" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="15" pageNumber="16" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
). Generally, the radula of
<taxonomicName id="0EB23C93FFEDFFB2FB1AFB10FAB06E81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1201,1318,1212,1234]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FB1AFB10FAB06E81" box="[1201,1318,1212,1234]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
has fewer lateral teeth per row than that of
<taxonomicName id="0EB23C93FFEDFFB2FB10FB76FAD56EBF" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1211,1347,1242,1264]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FB10FB76FAD56EBF" box="[1211,1347,1242,1264]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFEDFFB2FCF8FB54FABE6F97" blockId="15.[827,1444,198,1479]" pageId="15" pageNumber="16">
The reproductive system of
<taxonomicName id="0EB23C93FFEDFFB2FB0BFB57FA856F5F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1184,1299,1274,1295]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FB0BFB57FA856F5F" box="[1184,1299,1274,1295]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
has several consistent differences from that of
<taxonomicName id="0EB23C93FFEDFFB2FB69FAB4FADF6F7D" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1218,1353,1304,1326]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FB69FAB4FADF6F7D" box="[1218,1353,1304,1326]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
(CASIZ 206801;
<figureCitation id="51895B95FFEDFFB2FC3CFA9AFC726F1C" box="[919,996,1335,1357]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="15" pageNumber="16">Fig. 4F</figureCitation>
). In
<taxonomicName id="0EB23C93FFEDFFB2FBBDFA9AFB176F1C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1046,1153,1335,1356]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFEDFFB2FBBDFA9AFB176F1C" box="[1046,1153,1335,1356]" italics="true" pageId="15" pageNumber="16">H. cerisae</emphasis>
</taxonomicName>
, the vagina is shorter and wider than that of
<taxonomicName id="0EB23C93FFEDFFB2FBADFAFBFB1F6F3A" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1030,1161,1365,1387]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FBADFAFBFB1F6F3A" box="[1030,1161,1365,1387]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
, and the uterine duct and receptaculum seminis are situated more distally than in
<taxonomicName id="0EB23C93FFEDFFB2FC90FA3EFC576FF7" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[827,961,1426,1448]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFEDFFB2FC90FA3EFC576FF7" box="[827,961,1426,1448]" italics="true" pageId="15" pageNumber="16">H. krakatoa</emphasis>
</taxonomicName>
, where the uterine duct and receptaculum are immediately below the bursa copulatrix.
</paragraph>
</subSubSection>
</treatment>
</document>

549
data/41/1B/F6/411BF606FFF1FFABFF42FF6BFC456BE3.xml Normal file
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@ -0,0 +1,549 @@
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<mods:title id="3283A7679BCACACA634992EF760B8C15">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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</mods:role>
<mods:namePart id="D0C8092B609115CC1A705B344D6DC9DC">Lopez, Alessandra</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="E1E3AC7348FAF0B3A5121FD6DCA905F3">Gosliner, Terrence M.</mods:namePart>
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<mods:titleInfo id="D33D09F428B6BFE17D572814735866BB">
<mods:title id="28961A0AD812FB39E6FFD71CF909DFA9">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="4302B0851780766E700A6D33B8F2ED00">
<mods:date id="F2B32E73AA9CBEE9A88F647834A78A56">2018</mods:date>
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<mods:number id="24380D812D41DF2833021A7A671EEA04">20</mods:number>
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<mods:start id="1817320D2D91967C9C5CE152CEE33CAB">1</mods:start>
<mods:end id="59D26086662C25EF2F18384F6BEF32D4">74</mods:end>
</mods:extent>
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<mods:identifier id="27441BA8383AC89A4F70A5DD36CF2720" type="CLB-Dataset">27363</mods:identifier>
<mods:identifier id="88A67CCCB5F4AD5EB2A079744A60C035" type="DOI">10.1093/zoolinnean/zly048</mods:identifier>
<mods:identifier id="F395C9DAB71FC0538367996809D84A5A" type="GBIF-Dataset">49bea467-fac6-484a-9902-05483dec9a52</mods:identifier>
<mods:identifier id="F9288104CC23F6236571A9AAA0EB215A" type="Zenodo-Dep">2632578</mods:identifier>
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<treatment id="411BF606FFF1FFABFF42FF6BFC456BE3" ID-DOI="http://doi.org/10.5281/zenodo.5980674" ID-GBIF-Taxon="156146727" ID-Zenodo-Dep="5980674" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3" httpUri="http://treatment.plazi.org/id/411BF606FFF1FFABFF42FF6BFC456BE3" lastPageId="22" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="81A8149BFFF1FFAEFF42FF6BFD526A8D" box="[233,708,197,221]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="C90D4710FFF1FFAEFF42FF6BFD526A8D" blockId="19.[163,779,197,421]" box="[233,708,197,221]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFF42FF6BFD526A8D" allCaps="true" box="[233,708,197,221]" centered="true" fontSize="9" level="3" pageId="19" pageNumber="20" reason="5">
<taxonomicName id="0EB23C93FFF1FFAEFF42FF6BFD526A8D" ID-CoL="3P337" authority="(RISBEC, 1928)" baseAuthorityName="RISBEC" baseAuthorityYear="1928" box="[233,708,197,221]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF1FFAEFF42FF6BFD826A8C" box="[233,532,198,221]" italics="true" pageId="19" pageNumber="20">HYPSELODORIS DECORATA</emphasis>
(
<bibRefCitation id="AD233AE1FFF1FFAEFD88FF6BFD2A6A8D" author="Risbec J." box="[547,700,197,221]" journalOrPublisher="Faune des Colonies Francaises" pageId="19" pageNumber="20" pagination="1 - 328" part="2" refId="ref44543" refString="Risbec J. 1928. Contribution a l'etude des nudibranches Neo- Caledoniens. Faune des Colonies Francaises 2: 1 - 328." title="Contribution a l'etude des nudibranches Neo- Caledoniens" type="journal article" year="1928">RISBEC, 1928</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF1FFAEFECEFF41FDDE6B54" box="[357,584,236,260]" pageId="19" pageNumber="20" type="description">
<paragraph id="C90D4710FFF1FFAEFECEFF41FDDE6B54" blockId="19.[163,779,197,421]" box="[357,584,236,260]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFECEFF41FDDE6B54" allCaps="true" box="[357,584,236,260]" centered="true" fontSize="10" level="1" pageId="19" pageNumber="20" reason="1">(FIGS 2D, 4H, 9, 10)</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF1FFAEFF08FEB9FDA06BF5" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="C90D4710FFF1FFAEFF08FEB9FEA36B18" blockId="19.[163,779,197,421]" pageId="19" pageNumber="20">
<taxonomicName id="0EB23C93FFF1FFAEFF08FEB9FDE16B7A" ID-CoL="5YHKH" authority="Risbec, 1928: 152" authorityName="Risbec" authorityPageNumber="152" authorityYear="1928" box="[163,631,276,298]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF1FFAEFF08FEB9FE306B79" box="[163,422,276,297]" italics="true" pageId="19" pageNumber="20">Chromodoris decorata</emphasis>
<bibRefCitation id="AD233AE1FFF1FFAEFE06FEB9FDD46B79" author="Risbec J." box="[429,578,276,298]" journalOrPublisher="Faune des Colonies Francaises" pageId="19" pageNumber="20" pagination="1 - 328" part="2" refId="ref44543" refString="Risbec J. 1928. Contribution a l'etude des nudibranches Neo- Caledoniens. Faune des Colonies Francaises 2: 1 - 328." title="Contribution a l'etude des nudibranches Neo- Caledoniens" type="journal article" year="1928">Risbec, 1928</bibRefCitation>
: 152
</taxonomicName>
–154, fig. 43, pl. VII, fig. 4.
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08FEFFFDA06BF5" blockId="19.[163,779,197,421]" pageId="19" pageNumber="20">
<emphasis id="FBC69B02FFF1FFAEFF08FEFFFE416B37" box="[163,471,337,359]" italics="true" pageId="19" pageNumber="20">H y p s e l o d o r i s m a c u l o s a</emphasis>
m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFF1FFAEFF08FEDCFE5B6BD5" ID-CoL="3P34H" authority="Pease, 1871" authorityName="Pease" authorityYear="1871" box="[163,461,368,390]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF1FFAEFF08FEDCFEA16BD5" box="[163,311,368,390]" italics="true" pageId="19" pageNumber="20">H. maculosa</emphasis>
<bibRefCitation id="AD233AE1FFF1FFAEFE94FEDCFE5B6BD5" author="Pease WH" box="[319,461,368,390]" journalOrPublisher="American Journal of Conchology" pageId="19" pageNumber="20" pagination="11 - 19" part="7" refId="ref44179" refString="Pease WH. 1871. Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2. American Journal of Conchology, 7: 11 - 19." title="Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2." type="journal article" year="1871">Pease, 1871</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFF1FFAEFE71FEDDFD5D6BD6" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[474,715,368,390]" journalOrPublisher="Jacksonville: New World Press" pageId="19" pageNumber="20" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFF1FFAEFDECFEDCFD156BD5" box="[583,643,368,390]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2015
</bibRefCitation>
: 254, second photograph from top on left.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF1FFAEFF08FE61FEC16F10" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="C90D4710FFF1FFAEFF08FE61FE1C6BB4" blockId="19.[163,779,460,1250]" box="[163,394,460,484]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFF08FE61FE1C6BB4" box="[163,394,460,484]" fontSize="10" level="4" pageId="19" pageNumber="20" reason="1">
<emphasis id="FBC69B02FFF1FFAEFF08FE61FE1C6BB4" box="[163,394,460,484]" italics="true" pageId="19" pageNumber="20">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08FE41FE5D6EB1" blockId="19.[163,779,460,1250]" pageId="19" pageNumber="20">
<materialsCitation id="79DA4D4DFFF1FFAEFF08FE41FE8A680E" ID-GBIF-Occurrence="2235579108" collectingDate="2012-11-18" collectionCode="CASIZ" collectorName="V. Knutson" country="Papua New Guinea" elevation="20" latitude="-5.16" location="Mizegwadan Reef" longLatPrecision="1" longitude="145.82333" pageId="19" pageNumber="20" specimenCode="CASIZ 191315" specimenCount="1" stateProvince="Madang">
<specimenCode id="9914EF6BFFF1FFAEFF08FE41FECB6852" box="[163,349,492,514]" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 191315</specimenCode>
, subsampled for molecular study,
<location id="CC6D11CBFFF1FFAEFF08FDA6FEE26871" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFF08FDA6FEE26871" box="[163,372,523,545]" country="Papua New Guinea" latitude="-5.16" longLatPrecision="1" longitude="145.82333" name="Mizegwadan Reef" pageId="19" pageNumber="20" stateProvince="Madang">Mizegwadan Reef</location>
,
<geoCoordinate id="AC8621D7FFF1FFAEFE2BFDA6FE616871" box="[384,503,523,545]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="-5.16">5.16000°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFDA8FDA6FD0C6871" box="[515,666,523,545]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="145.82333">145.82333°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFF1FFAEFD0DFDA6FC906871" box="[678,774,523,545]" country="Papua New Guinea" name="Madang" pageId="19" pageNumber="20">Madang</collectingRegion>
,
<collectingCountry id="B1A50780FFF1FFAEFF08FD87FE1E6810" box="[163,392,554,576]" name="Papua New Guinea" pageId="19" pageNumber="20">Papua New Guinea</collectingCountry>
,
<elevation id="429FA023FFF1FFAEFE3EFD87FE59686F" box="[405,463,554,575]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="20.0">
<quantity id="0E4AEAF5FFF1FFAEFE3EFD87FE59686F" box="[405,463,554,575]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="20.0">20 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFF1FFAEFD82FD87FC91686F" box="[553,775,554,576]" pageId="19" pageNumber="20" value="2012-11-18">
<date id="BD0C61D0FFF1FFAEFD82FD87FC91686F" box="[553,775,554,576]" pageId="19" pageNumber="20" value="2012-11-18">18 November 2012</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFF08FDE4FE8A680E" box="[163,284,585,606]" pageId="19" pageNumber="20">V.Knutson</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFE8FFDE5FD3F68CC" ID-GBIF-Occurrence="2235579052" collectingDate="2012-11-14" collectionCode="CASIZ" collectorName="Anthony Berberian" country="Papua New Guinea" latitude="-5.196667" location="Pana Tibun" longLatPrecision="1" longitude="145.80667" pageId="19" pageNumber="20" specimenCode="CASIZ 191214" specimenCount="1" stateProvince="Madang">
<specimenCode id="9914EF6BFFF1FFAEFE8FFDE5FE52680D" box="[292,452,584,605]" collectionCode="CASIZ" pageId="19" pageNumber="20">
CASIZ
<date id="BD0C61D0FFF1FFAEFED9FDE5FE52680D" box="[370,452,584,605]" pageId="19" pageNumber="20" value="14-12-19">191214</date>
</specimenCode>
, two specimens, one dissected, south point,
<location id="CC6D11CBFFF1FFAEFE83FDCAFE31682D" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFE83FDCAFE31682D" box="[296,423,615,637]" country="Papua New Guinea" latitude="-5.196667" longLatPrecision="1" longitude="145.80667" name="Pana Tibun" pageId="19" pageNumber="20" stateProvince="Madang">Pana Tibun</location>
islet,
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,
<geoCoordinate id="AC8621D7FFF1FFAEFDC1FDCAFC91682D" box="[618,775,615,637]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="145.80667">145.806667°E</geoCoordinate>
,
<collectingDate id="AD489838FFF1FFAEFF08FD2BFE0D68CB" box="[163,411,646,668]" pageId="19" pageNumber="20" value="2012-11-14">
<date id="BD0C61D0FFF1FFAEFF08FD2BFE0D68CB" box="[163,411,646,668]" pageId="19" pageNumber="20" value="2012-11-14">14 November 2012</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFE01FD2BFD3F68CC" box="[426,681,646,668]" pageId="19" pageNumber="20">Anthony Berberian</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFD13FD2BFD2668A8" ID-GBIF-Occurrence="2235579037" collectingDate="2012-11-17" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Luzon" latitude="13.67" location="Bethlehem" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" pageId="19" pageNumber="20" specimenCode="CASIZ 190661" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFD13FD2BFF6168EA" collectionCode="CASIZ" pageId="19" pageNumber="20">
CASIZ
<date id="BD0C61D0FFF1FFAEFF08FD09FF6168EA" box="[163,247,676,698]" pageId="19" pageNumber="20" value="61-06-19">190661</date>
</specimenCode>
, one specimen, dissected,
<location id="CC6D11CBFFF1FFAEFD8BFD09FD0968EA" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFD8BFD09FD0968EA" box="[544,671,676,698]" country="Philippines" county="Luzon" latitude="13.67" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" name="Bethlehem" pageId="19" pageNumber="20" stateProvince="Batangas">Bethlehem</location>
,
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,
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,
<location id="CC6D11CBFFF1FFAEFEBFFD6EFEFE6889" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFEBFFD6EFEFE6889" box="[276,360,707,729]" country="Philippines" county="Luzon" latitude="13.67" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" name="Tingloy" pageId="19" pageNumber="20" stateProvince="Batangas">Tingloy</location>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFED8FD6EFDAB6889" box="[371,573,707,729]" pageId="19" pageNumber="20">Maricaban Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF1FFAEFDE3FD6EFD226888" box="[584,692,707,728]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFF1FFAEFD14FD6EFC916888" box="[703,775,707,728]" pageId="19" pageNumber="20">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFF1FFAEFF08FD4FFEBB68A8" box="[163,301,738,760]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFE91FD4FFD8968A7" box="[314,543,738,760]" pageId="19" pageNumber="20" value="2012-11-17">
<date id="BD0C61D0FFF1FFAEFE91FD4FFD8968A7" box="[314,543,738,760]" pageId="19" pageNumber="20" value="2012-11-17">17 November 2012</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFD80FD4FFD2668A8" box="[555,688,738,760]" pageId="19" pageNumber="20">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFD17FD4FFDCF6904" ID-GBIF-Occurrence="2235579081" collectingDate="2010-10-02" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Luzon" latitude="13.67" location="Bethlehem" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" pageId="19" pageNumber="20" specimenCode="CASIZ 184316" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFD17FD4FFF616946" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 184316</specimenCode>
, one specimen, dissected,
<location id="CC6D11CBFFF1FFAEFD8BFCADFD096946" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFD8BFCADFD096946" box="[544,671,768,790]" country="Philippines" county="Luzon" latitude="13.67" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" name="Bethlehem" pageId="19" pageNumber="20" stateProvince="Batangas">Bethlehem</location>
,
<geoCoordinate id="AC8621D7FFF1FFAEFD01FCADFC906946" box="[682,774,768,790]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="13.67">13.67°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFF08FCB2FE9C6964" box="[163,266,799,820]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="120.84">120.84°E</geoCoordinate>
,
<location id="CC6D11CBFFF1FFAEFEBFFCB2FEFE6965" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFEBFFCB2FEFE6965" box="[276,360,799,821]" country="Philippines" county="Luzon" latitude="13.67" longLatPrecision="774" longitude="120.84" municipality="Maricaban Island" name="Tingloy" pageId="19" pageNumber="20" stateProvince="Batangas">Tingloy</location>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFED8FCB2FDAB6965" box="[371,573,799,821]" pageId="19" pageNumber="20">Maricaban Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF1FFAEFDE3FCB2FD226964" box="[584,692,799,820]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFF1FFAEFD14FCB2FC916964" box="[703,775,799,820]" pageId="19" pageNumber="20">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFF1FFAEFF08FC93FEB56904" box="[163,291,830,852]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFE86FC93FE416903" box="[301,471,830,852]" pageId="19" pageNumber="20" value="2010-10-02">
<date id="BD0C61D0FFF1FFAEFE86FC93FE416903" box="[301,471,830,852]" pageId="19" pageNumber="20" value="2010-10-02">2 October 2010</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFE4AFC93FDCF6904" box="[481,601,830,852]" pageId="19" pageNumber="20">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFDC8FC93FDC269E0" ID-GBIF-Occurrence="2235579130" collectingDate="2015-04-11" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Puerto Galera" latitude="13.52" location="Bamboo Beach" longLatPrecision="774" longitude="120.96" municipality="Channel" pageId="19" pageNumber="20" specimenCode="CASIZ 208606" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFDC8FC93FC916903" box="[611,775,829,851]" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 208606</specimenCode>
, one specimen, off
<location id="CC6D11CBFFF1FFAEFEDCFCF1FDB06922" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFEDCFCF1FDB06922" box="[375,550,860,882]" country="Philippines" county="Puerto Galera" latitude="13.52" longLatPrecision="774" longitude="120.96" municipality="Channel" name="Bamboo Beach" pageId="19" pageNumber="20" stateProvince="Batangas">Bamboo Beach</location>
<geoCoordinate id="AC8621D7FFF1FFAEFD85FCF1FD1B6922" box="[558,653,860,882]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="13.52">13.52°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFD31FCF1FC906922" box="[666,774,860,882]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="120.96">120.96°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFF1FFAEFF08FCD6FE8769C0" box="[163,273,891,912]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
<collectingMunicipality id="2969DD6AFFF1FFAEFEB3FCD6FEEA69C1" box="[280,380,891,913]" pageId="19" pageNumber="20">Channel</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFF1FFAEFE23FCD6FDB869C1" box="[392,558,891,913]" pageId="19" pageNumber="20">Puerto Galera</collectingCounty>
,
<collectingRegion id="0B7689F2FFF1FFAEFD91FCD6FC9069C1" box="[570,774,891,913]" country="Philippines" name="Mindoro Oriental" pageId="19" pageNumber="20">Mindoro Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFF1FFAEFF08FC37FEB069E0" box="[163,294,922,944]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFE9AFC37FE5869FF" box="[305,462,921,944]" pageId="19" pageNumber="20" value="2015-04-11">
<date id="BD0C61D0FFF1FFAEFE9AFC37FE5869FF" box="[305,462,921,944]" pageId="19" pageNumber="20" value="2015-04-11">11 April 2015</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFE72FC37FDC269E0" box="[473,596,922,944]" pageId="19" pageNumber="20">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFDF4FC37FC9169BD" ID-GBIF-Occurrence="2235579063" collectingDate="2007-09-29" collectionCode="CASIZ" collectorName="T. Gosliner" country="China" county="Sea" location="West Beach" municipality="South" pageId="19" pageNumber="20" specimenCode="CASIZ 176776" specimenCount="1">
<specimenCode id="9914EF6BFFF1FFAEFDF4FC37FC9069FF" box="[607,774,921,943]" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 176776</specimenCode>
, two specimens,
<location id="CC6D11CBFFF1FFAEFECFFC14FE63699E" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFECFFC14FE63699E" box="[356,501,952,974]" country="China" county="Sea" municipality="South" name="West Beach" pageId="19" pageNumber="20">West Beach</location>
,
<location id="CC6D11CBFFF1FFAEFDA8FC15FD20699E" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFDA8FC15FD20699E" box="[515,694,952,974]" country="China" county="Sea" municipality="South" name="Pulau Tenggol" pageId="19" pageNumber="20">Pulau Tenggol</location>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFD6FFC15FC9D699E" box="[708,779,952,974]" pageId="19" pageNumber="20">South</collectingMunicipality>
<collectingCountry id="B1A50780FFF1FFAEFF08FC7AFF7F69BD" box="[163,233,983,1005]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
<collectingCounty id="206C3F9CFFF1FFAEFF5BFC7AFE8A69BC" box="[240,284,983,1004]" pageId="19" pageNumber="20">Sea</collectingCounty>
,
<collectingCountry id="B1A50780FFF1FFAEFE83FC7AFE0569BD" box="[296,403,983,1005]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFE34FC7AFDE969BC" box="[415,639,983,1005]" pageId="19" pageNumber="20" value="2007-09-29">
<date id="BD0C61D0FFF1FFAEFE34FC7AFDE969BC" box="[415,639,983,1005]" pageId="19" pageNumber="20" value="2007-09-29">29 September 2007</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFD21FC7AFC9169BD" box="[650,775,983,1005]" pageId="19" pageNumber="20">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFF08FC5BFED06E19" ID-GBIF-Occurrence="2235579075" collectingDate="2014-05-19" collectionCode="CASIZ" collectorName="C. Piotrowski" country="Philippines" latitude="13.91" location="Calatagan" longLatPrecision="774" longitude="120.6" municipality="Luzon" pageId="19" pageNumber="20" specimenCode="CASIZ 197309" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFF08FC5BFEC16E5B" box="[163,343,1014,1035]" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 197309</specimenCode>
, one specimen,
<location id="CC6D11CBFFF1FFAEFDB7FC5BFD0D6E5C" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFDB7FC5BFD0D6E5C" box="[540,667,1014,1036]" country="Philippines" latitude="13.91" longLatPrecision="774" longitude="120.6" municipality="Luzon" name="Calatagan" pageId="19" pageNumber="20" stateProvince="Batangas">Calatagan</location>
<geoCoordinate id="AC8621D7FFF1FFAEFD0FFC5BFC916E5B" box="[676,775,1014,1035]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="13.91">13.91°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFF08FBB9FE9A6E7A" box="[163,268,1044,1066]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="120.6">120.60°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFF1FFAEFEBCFBB8FE126E7A" box="[279,388,1045,1066]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFE3BFBB8FE4F6E7A" box="[400,473,1045,1066]" pageId="19" pageNumber="20">Luzon</collectingMunicipality>
,
<collectingCountry id="B1A50780FFF1FFAEFE4FFBB9FDF16E7A" box="[484,615,1044,1066]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFDD9FBB9FC916E79" box="[626,775,1044,1066]" pageId="19" pageNumber="20" value="2014-05-19">
<date id="BD0C61D0FFF1FFAEFDD9FBB9FC916E79" box="[626,775,1044,1066]" pageId="19" pageNumber="20" value="2014-05-19">19 May 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFF08FB9EFED06E19" box="[163,326,1075,1097]" pageId="19" pageNumber="20">C. Piotrowski</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFEF9FB9EFD256ED6" ID-GBIF-Occurrence="2235579038" collectingDate="2016-04-21" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" county="Luzon" latitude="13.63" location="Bonito" longLatPrecision="774" longitude="120.95" municipality="Tingloy" pageId="19" pageNumber="20" specimenCode="CASIZ 217267" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFEF9FB9EFD976E19" box="[338,513,1075,1097]" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 217267</specimenCode>
, one specimen,
<location id="CC6D11CBFFF1FFAEFD16FB9EFC9D6E19" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFD16FB9EFC9D6E19" box="[701,779,1075,1097]" country="Philippines" county="Luzon" latitude="13.63" longLatPrecision="774" longitude="120.95" municipality="Tingloy" name="Bonito" pageId="19" pageNumber="20" stateProvince="Batangas">Bonito</location>
(
<location id="CC6D11CBFFF1FFAEFF01FBFFFE906E38" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFF01FBFFFE906E38" box="[170,262,1106,1128]" country="Philippines" county="Luzon" latitude="13.63" longLatPrecision="774" longitude="120.95" municipality="Tingloy" name="Culebra" pageId="19" pageNumber="20" stateProvince="Batangas">Culebra</location>
) Island
<geoCoordinate id="AC8621D7FFF1FFAEFEC8FBFCFE286E37" box="[355,446,1105,1127]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="13.63">13.63°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFE62FBFCFDA76E37" box="[457,561,1105,1127]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="120.95">120.95°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFD90FBFFFD196E38" box="[571,655,1106,1128]" pageId="19" pageNumber="20">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF1FFAEFD31FBFFFC906E37" box="[666,774,1106,1127]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFF1FFAEFF08FBDCFF7B6ED6" box="[163,237,1137,1158]" pageId="19" pageNumber="20">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFF1FFAEFF53FBDDFEEB6ED6" box="[248,381,1136,1158]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFE21FBDDFDBC6ED6" box="[394,554,1136,1158]" pageId="19" pageNumber="20" value="2016-04-21">
<date id="BD0C61D0FFF1FFAEFE21FBDDFDBC6ED6" box="[394,554,1136,1158]" pageId="19" pageNumber="20" value="2016-04-21">21 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFD9FFBDCFD256ED6" box="[564,691,1136,1158]" pageId="19" pageNumber="20">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFF1FFAEFD15FBDDFE516EB1" ID-GBIF-Occurrence="2235579094" collectingDate="2016-04-17" collectionCode="CASIZ" collectorName="Brenna Green" country="Philippines" county="Luzon" latitude="13.8" location="Dive" longLatPrecision="774" longitude="120.91" municipality="Bauan" pageId="19" pageNumber="20" specimenCode="CASIZ 217358" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF1FFAEFD15FBDDFF6F6EF5" collectionCode="CASIZ" pageId="19" pageNumber="20">CASIZ 217358</specimenCode>
, two specimens,
<location id="CC6D11CBFFF1FFAEFE16FB22FE656EF5" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFE16FB22FE656EF5" box="[445,499,1167,1189]" country="Philippines" county="Luzon" latitude="13.8" longLatPrecision="774" longitude="120.91" municipality="Bauan" name="Dive" pageId="19" pageNumber="20" stateProvince="Batangas">Dive</location>
and
<location id="CC6D11CBFFF1FFAEFD85FB22FDF06EF5" LSID="urn:lsid:plazi:treatment:411BF606FFF1FFABFF42FF6BFC456BE3:CC6D11CBFFF1FFAEFD85FB22FDF06EF5" box="[558,614,1167,1189]" country="Philippines" county="Luzon" latitude="13.8" longLatPrecision="774" longitude="120.91" municipality="Bauan" name="Trek" pageId="19" pageNumber="20" stateProvince="Batangas">Trek</location>
reef,
<geoCoordinate id="AC8621D7FFF1FFAEFD03FB22FC906EF4" box="[680,774,1167,1188]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="13.8">13.80°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF1FFAEFF08FB03FE876E93" box="[163,273,1198,1219]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="120.91">120.91°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFF1FFAEFEB5FB03FEF96E93" box="[286,367,1198,1219]" pageId="19" pageNumber="20">Bauan</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF1FFAEFED6FB03FE666E93" box="[381,496,1198,1219]" country="Philippines" name="Batangas" pageId="19" pageNumber="20">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFF1FFAEFE56FB03FDDF6E93" box="[509,585,1198,1219]" pageId="19" pageNumber="20">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFF1FFAEFDFCFB03FD776E94" box="[599,737,1198,1220]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFF1FFAEFD45FB03FE8D6EB2" pageId="19" pageNumber="20" value="2016-04-17">
<date id="BD0C61D0FFF1FFAEFD45FB03FE8D6EB2" pageId="19" pageNumber="20" value="2016-04-17">17 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFF1FFAEFE8EFB60FE516EB1" box="[293,455,1228,1250]" pageId="19" pageNumber="20">Brenna Green</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08FAA7FEAB6F71" blockId="19.[163,343,1289,1344]" box="[163,317,1289,1313]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFF08FAA7FEAB6F71" box="[163,317,1289,1313]" fontSize="10" level="4" pageId="19" pageNumber="20" reason="1">
<emphasis id="FBC69B02FFF1FFAEFF08FAA7FEAB6F71" box="[163,317,1289,1313]" italics="true" pageId="19" pageNumber="20">
<typeStatus id="1609F9B2FFF1FFAEFF08FAA7FF4A6F71" box="[163,220,1290,1313]" pageId="19" pageNumber="20">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08FA87FEC16F10" blockId="19.[163,343,1289,1344]" box="[163,343,1322,1344]" pageId="19" pageNumber="20">
<collectingCountry id="B1A50780FFF1FFAEFF08FA87FEC56F10" box="[163,339,1322,1344]" name="New Caledonia" pageId="19" pageNumber="20">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF1FFAEFF08FACBFD166F8A" pageId="19" pageNumber="20" type="distribution">
<paragraph id="C90D4710FFF1FFAEFF08FACBFE766F2E" blockId="19.[163,779,1382,1498]" box="[163,480,1382,1406]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFF08FACBFE766F2E" box="[163,480,1382,1406]" fontSize="10" level="4" pageId="19" pageNumber="20" reason="1">
<emphasis id="FBC69B02FFF1FFAEFF08FACBFE766F2E" box="[163,480,1382,1406]" italics="true" pageId="19" pageNumber="20">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08FA2AFD166F8A" blockId="19.[163,779,1382,1498]" pageId="19" pageNumber="20">
Known from the
<collectingCountry id="B1A50780FFF1FFAEFEDCFA2AFDDB6FCD" box="[375,589,1415,1437]" name="Marshall Islands" pageId="19" pageNumber="20">Marshall Islands</collectingCountry>
,
<collectingCountry id="B1A50780FFF1FFAEFDF0FA2AFD516FCC" box="[603,711,1415,1436]" name="Vanuatu" pageId="19" pageNumber="20">Vanuatu</collectingCountry>
,
<collectingCountry id="B1A50780FFF1FFAEFD7EFA2AFEBF6FEC" name="New Caledonia" pageId="19" pageNumber="20">New Caledonia</collectingCountry>
,
<collectingCountry id="B1A50780FFF1FFAEFE91FA0BFE286FEC" box="[314,446,1446,1468]" name="Indonesia" pageId="19" pageNumber="20">Indonesia</collectingCountry>
,
<collectingCountry id="B1A50780FFF1FFAEFE65FA0BFD476FEC" box="[462,721,1446,1468]" name="Papua New Guinea" pageId="19" pageNumber="20">Papua New Guinea</collectingCountry>
, the
<collectingCountry id="B1A50780FFF1FFAEFF08FA69FEB36F8A" box="[163,293,1476,1498]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
and
<collectingCountry id="B1A50780FFF1FFAEFEF6FA69FE536F8A" box="[349,453,1476,1498]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF1FFAEFF08F9AFFA9E6D3A" pageId="19" pageNumber="20" type="description">
<paragraph id="C90D4710FFF1FFAEFF08F9AFFEB96C49" blockId="19.[163,779,1538,1898]" box="[163,303,1538,1561]" pageId="19" pageNumber="20">
<heading id="9245F07CFFF1FFAEFF08F9AFFEB96C49" box="[163,303,1538,1561]" fontSize="10" level="4" pageId="19" pageNumber="20" reason="1">
<emphasis id="FBC69B02FFF1FFAEFF08F9AFFEB96C49" box="[163,303,1538,1561]" italics="true" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF1FFAEFF08F98CFACE6BC4" blockId="19.[163,779,1538,1898]" lastBlockId="19.[827,1443,198,404]" pageId="19" pageNumber="20">
<emphasis id="FBC69B02FFF1FFAEFF08F98CFE106C66" box="[163,390,1569,1590]" italics="true" pageId="19" pageNumber="20">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFF1FFAEFDEEF98FFD166C67" box="[581,640,1570,1592]" captionStart="Figure 9" captionStartId="20.[145,223,1589,1611]" captionTargetBox="[273,1297,195,1548]" captionTargetId="figure@20.[273,1297,195,1548]" captionTargetPageId="20" captionText="Figure 9. Hypselodoris decorata (Risbec, 1928), living animals. A, CASIZ 191315, Madang, Papua New Guinea, photograph by Vanessa Knutson. B, CASIZ 217614, Mabini, Batangas, Philippines. C, CASIZ 184316, Tingloy, Batangas, Philippines. D, CASIZ 190661, Tingloy, Batangas, Philippines. E, CASIZ 217358, Bauan, Batangas, Philippines. F, CASIZ 191214, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, living animal from Risbec (1928), New Caledonia." httpUri="https://zenodo.org/record/2632789/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
) moderately large, reaching
<quantity id="0E4AEAF5FFF1FFAEFEF1F9EDFE3D6C06" box="[346,427,1600,1622]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="mm" value="25.0">25 mm</quantity>
in length. Body long, slender, translucent pink, with four thin, continuous opaque white longitudinal lines on dorsal surface of notum. Broad orange band found along lateral margins of notum, often with interior undulations and opaque white spots.Anterior and posterior ends of the animal with purple to orange areas with opaque white spots. Rows of reddish to purple spots found on notum, with more diffuse purple pigment found around periphery situated between opaque white lines. Similar pattern of lines and spots found on lateral margins of the body and foot. Seven to eight unipinnate gill branches having a translucent white base and inner margin and bright red–orange pigment on apical surfaces and outer margin. Bulb of perfoliate rhinophores opaque white with two red transverse bands and bearing ~14–16 densely arranged lamellae. Base of rhinophores translucent white with a third red transverse ring.
</paragraph>
<paragraph id="C90D4710FFF1FFAEFC90FE10FA3268DB" blockId="19.[827,1444,445,651]" pageId="19" pageNumber="20">
<emphasis id="FBC69B02FFF1FFAEFC90FE10FC736B82" box="[827,997,445,466]" italics="true" pageId="19" pageNumber="20">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFF1FFAEFBC8FE71FB566BA1" box="[1123,1216,476,498]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="19" pageNumber="20">Fig. 2D</figureCitation>
). Glands situated anteriorly and posteriorly, with no glands present in the central lateral regions of body margin. Eight to ten glands on either side of anterior end of the body, with arc of 15–20 glands situated posteriorly. Their arrangement was based upon four specimens examined.
</paragraph>
<paragraph id="C90D4710FFF1FFAEFC90FD19FAAC6E83" blockId="19.[827,1443,692,1235]" pageId="19" pageNumber="20">
<emphasis id="FBC69B02FFF1FFAEFC90FD19FB936899" box="[827,1029,692,713]" italics="true" pageId="19" pageNumber="20">Buccal armature</emphasis>
: Muscular portion of buccal mass slightly longer than oral tube. Chitinous labial cuticle (
<figureCitation id="51895B95FFF1FFAEFCE8FD5CFC376956" box="[835,929,753,775]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig. 10A</figureCitation>
) found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFF1FFAEFA90FCBDFA036975" box="[1339,1429,784,806]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig.10B</figureCitation>
). Rodlets sharply angled with single, acutely pointed apex and posterolateral extensions. Radular formula of two specimens 55 × 26.0.26 (CASIZ
<date id="BD0C61D0FFF1FFAEFAA4FCC1FAF069D2" box="[1295,1382,876,898]" pageId="19" pageNumber="20" value="61-06-19">190661</date>
) and 48 × 26.0.26 (CASIZ
<date id="BD0C61D0FFF1FFAEFB91FC26FB0769F1" box="[1082,1169,907,929]" pageId="19" pageNumber="20" value="14-12-19">191214</date>
) (
<figureCitation id="51895B95FFF1FFAEFB00FC27FA8669F0" box="[1195,1296,906,928]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig. 10C</figureCitation>
). Rachidian row of teeth absent (
<figureCitation id="51895B95FFF1FFAEFB8FFC04FB1569EE" box="[1060,1155,937,959]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig. 10D</figureCitation>
). Innermost lateral teeth having one or two large, triangular denticles on inner side of bifid primary cusp, with another one to two outer denticles. Next several laterals lacking inner triangular denticle but possessing two denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFF1FFAEFAA6FBEFFAE46E08" box="[1293,1394,1090,1112]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig. 10E</figureCitation>
) all lacking inner denticles but having three to four sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and shorter tooth shape, with two to four rounded outer denticles (
<figureCitation id="51895B95FFF1FFAEFB7BFB10FABD6E82" box="[1232,1323,1213,1235]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="19" pageNumber="20">Fig. 10F</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFF1FFAEFC90FB43FA9E6D3A" blockId="19.[827,1443,1262,1898]" pageId="19" pageNumber="20">
<emphasis id="FBC69B02FFF1FFAEFC90FB43FBBA6F54" box="[827,1068,1262,1284]" italics="true" pageId="19" pageNumber="20">Reproductive system</emphasis>
: Reproductive organs (
<figureCitation id="51895B95FFF1FFAEFAE8FB43FA0C6F54" box="[1347,1434,1262,1284]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="19" pageNumber="20">Fig. 4H</figureCitation>
) fully mature in two specimens examined (CASIZ 1908661, CASIZ 184316). Ampulla thick, tubular, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing slightly as it transitions into short, narrow, muscular ejaculatory portion. Ejaculatory portion widening into penial bulb. Penial bulb adjacent to curved, slightly wider vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum situated near base of bursa. Moderately long uterine duct emerging from vagina close to base of bursa and female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<caption id="9DCD1798FFF6FFA9FF3AF998FB7F6CF3" httpUri="https://zenodo.org/record/2632789/files/figure.png" pageId="20" pageNumber="21" startId="20.[145,223,1589,1611]" targetBox="[273,1297,195,1548]" targetPageId="20">
<paragraph id="C90D4710FFF6FFA9FF3AF998FB7F6CF3" blockId="20.[145,1425,1589,1699]" pageId="20" pageNumber="21">
<emphasis id="FBC69B02FFF6FFA9FF3AF998FF6E6C1A" bold="true" box="[145,248,1589,1611]" pageId="20" pageNumber="21">Figure 9.</emphasis>
<taxonomicName id="0EB23C93FFF6FFA9FEA8F998FD166C1B" authority="(Risbec, 1928)" authorityName="Risbec" authorityYear="1928" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[259,640,1589,1611]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF6FFA9FEA8F998FE7F6C1A" box="[259,489,1589,1610]" italics="true" pageId="20" pageNumber="21">Hypselodoris decorata</emphasis>
(Risbec, 1928)
</taxonomicName>
, living animals. A, CASIZ 191315, Madang, Papua New Guinea, photograph by Vanessa Knutson. B, CASIZ 217614, Mabini, Batangas, Philippines. C, CASIZ 184316, Tingloy, Batangas, Philippines. D, CASIZ 190661, Tingloy, Batangas, Philippines. E, CASIZ 217358, Bauan, Batangas, Philippines. F, CASIZ 191214, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, living animal from Risbec (1928), New Caledonia.
</paragraph>
</caption>
<subSubSection id="81A8149BFFF6FFABFF3AF96AFC456BE3" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="discussion">
<paragraph id="C90D4710FFF6FFA9FF3AF96AFF686C8F" blockId="20.[145,761,1735,1881]" box="[145,254,1735,1759]" pageId="20" pageNumber="21">
<heading id="9245F07CFFF6FFA9FF3AF96AFF686C8F" box="[145,254,1735,1759]" fontSize="10" level="4" pageId="20" pageNumber="21" reason="1">
<emphasis id="FBC69B02FFF6FFA9FF3AF96AFF686C8F" box="[145,254,1735,1759]" italics="true" pageId="20" pageNumber="21">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF6FFABFF3AF94AFC456BE3" blockId="20.[145,761,1735,1881]" lastBlockId="22.[809,1425,198,435]" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<taxonomicName id="0EB23C93FFF6FFA9FF3AF94AFE046CAC" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[145,402,1767,1788]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF6FFA9FF3AF94AFE046CAC" box="[145,402,1767,1788]" italics="true" pageId="20" pageNumber="21">Hypselodoris decorata</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="0EB23C93FFF6FFA9FDC3F945FEBC6D4C" authority="(Pease, 1871)" baseAuthorityName="Pease" baseAuthorityYear="1871" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF6FFA9FDC3F945FD6F6CAC" box="[616,761,1767,1789]" italics="true" pageId="20" pageNumber="21">H. maculosa</emphasis>
(
<bibRefCitation id="AD233AE1FFF6FFA9FF32F8ABFEB76D4C" author="Pease WH" box="[153,289,1798,1820]" journalOrPublisher="American Journal of Conchology" pageId="20" pageNumber="21" pagination="11 - 19" part="7" refId="ref44179" refString="Pease WH. 1871. Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2. American Journal of Conchology, 7: 11 - 19." title="Descriptions of new species of nudibranchiate Mollusca inhabiting Polynesia. No. 2." type="journal article" year="1871">Pease, 1871</bibRefCitation>
)
</taxonomicName>
and was regarded as a synonym of this species by
<bibRefCitation id="AD233AE1FFF6FFA9FEA6F888FE576D6B" author="Rudman WB" box="[269,449,1829,1851]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="20" pageNumber="21" pagination="309 - 353" part="86" refId="ref44765" refString="Rudman WB. 1986. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups. Zoological Journal of the Linnean Society 86: 309 - 353." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups" type="journal article" year="1986">Rudman (1986)</bibRefCitation>
.
<bibRefCitation id="AD233AE1FFF6FFA9FE65F888FDE96D6B" author="Johnson S." box="[462,639,1829,1851]" journalOrPublisher="Sydney: Australian Museum" pageId="20" pageNumber="21" publicationUrl="http://www.underwaterkwaj.com/nudi/chromodorids/e017.htm" refId="ref43163" refString="Johnson S. 2005. Available at: http: // www. underwaterkwaj. com / nudi / chromodorids / e 017. htm" type="url" year="2005">Johnson (2005)</bibRefCitation>
suggested that
<taxonomicName id="0EB23C93FFF6FFA9FF63F8E9FEDD6D08" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[200,331,1859,1881]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF6FFA9FF63F8E9FEDD6D08" box="[200,331,1859,1881]" italics="true" pageId="20" pageNumber="21">H. decorata</emphasis>
</taxonomicName>
has several distinguishing attributes of its colour pattern that separate it from
<taxonomicName id="0EB23C93FFF6FFA9FB55F965FA1D6C8C" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1278,1419,1735,1757]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF6FFA9FB55F965FA1D6C8C" box="[1278,1419,1735,1757]" italics="true" pageId="20" pageNumber="21">H. maculosa</emphasis>
</taxonomicName>
, including the presence of three rather than two red rhinophoral rings. Owing to this confusion, we have provided a complete description of
<taxonomicName id="0EB23C93FFF6FFA9FB72F889FAF06D68" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1241,1382,1827,1849]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF6FFA9FB72F889FAF06D68" box="[1241,1382,1827,1849]" italics="true" pageId="20" pageNumber="21">H. decorata</emphasis>
</taxonomicName>
for purposes of comparison with that described for
<taxonomicName id="0EB23C93FFF7FFA8FF08F90AFEAD6CEB" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[163,315,1702,1724]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF7FFA8FF08F90AFEAD6CEB" box="[163,315,1702,1724]" italics="true" pageId="21" pageNumber="22">H. maculosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFF7FFA8FEE5F90BFD966CEC" author="Rudman WB" box="[334,512,1702,1724]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="21" pageNumber="22" pagination="309 - 353" part="86" refId="ref44765" refString="Rudman WB. 1986. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups. Zoological Journal of the Linnean Society 86: 309 - 353." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups" type="journal article" year="1986">Rudman, 1986</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFF7FFA8FDBBF90BFF486C8A" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="21" pageNumber="22" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
). The fact that specimens identified here as
<taxonomicName id="0EB23C93FFF7FFA8FF08F949FEA86CA9" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[163,318,1764,1785]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF7FFA8FF08F949FEA86CA9" box="[163,318,1764,1785]" italics="true" pageId="21" pageNumber="22">H. maculosa</emphasis>
</taxonomicName>
, with two rhinophoral rings, cluster together with species separate from
<taxonomicName id="0EB23C93FFF7FFA8FDEFF8AEFD5E6D47" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[580,712,1794,1816]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF7FFA8FDEFF8AEFD5E6D47" box="[580,712,1794,1816]" italics="true" pageId="21" pageNumber="22">H. decorata</emphasis>
</taxonomicName>
(with three rhinophoral rings) in our molecular phylogeny gives credence to the observation by
<bibRefCitation id="AD233AE1FFF7FFA8FDF3F8EDFC9D6D06" author="Johnson S." box="[600,779,1856,1878]" journalOrPublisher="Sydney: Australian Museum" pageId="21" pageNumber="22" publicationUrl="http://www.underwaterkwaj.com/nudi/chromodorids/e017.htm" refId="ref43163" refString="Johnson S. 2005. Available at: http: // www. underwaterkwaj. com / nudi / chromodorids / e 017. htm" type="url" year="2005">Johnson (2005)</bibRefCitation>
that these represent distinct species.
<taxonomicName id="0EB23C93FFF7FFA8FDC4F8F3FC376CEB" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[623,929,1702,1907]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF7FFA8FDC4F8F3FC376CEB" box="[623,929,1702,1907]" italics="true" pageId="21" pageNumber="22">Hypselodoris decorata</emphasis>
</taxonomicName>
is sister to
<taxonomicName id="0EB23C93FFF7FFA8FB9BF90AFB526CEC" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1072,1220,1703,1724]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="juniperae" status="sp. nov.">
<emphasis id="FBC69B02FFF7FFA8FB9BF90AFB526CEC" box="[1072,1220,1703,1724]" italics="true" pageId="21" pageNumber="22">H. juniperae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFF7FFA8FB67F90AFAB06CEC" box="[1228,1318,1703,1724]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, described later in this study, and
<taxonomicName id="0EB23C93FFF7FFA8FBF2F96BFB786C8A" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1113,1262,1733,1755]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF7FFA8FBF2F96BFB786C8A" box="[1113,1262,1733,1755]" italics="true" pageId="21" pageNumber="22">H. maculosa</emphasis>
</taxonomicName>
. In addition to the colour differences noted by
<bibRefCitation id="AD233AE1FFF7FFA8FB0CF949FACE6CAA" author="Johnson S." box="[1191,1368,1764,1786]" journalOrPublisher="Sydney: Australian Museum" pageId="21" pageNumber="22" publicationUrl="http://www.underwaterkwaj.com/nudi/chromodorids/e017.htm" refId="ref43163" refString="Johnson S. 2005. Available at: http: // www. underwaterkwaj. com / nudi / chromodorids / e 017. htm" type="url" year="2005">Johnson (2005)</bibRefCitation>
, there appear to be distinct anatomical differences between the two species. In
<taxonomicName id="0EB23C93FFF7FFA8FBB4F88FFB316D66" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1055,1191,1825,1847]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF7FFA8FBB4F88FFB316D66" box="[1055,1191,1825,1847]" italics="true" pageId="21" pageNumber="22">H. decorata</emphasis>
</taxonomicName>
, posterior glands are present, but there are no anterolateral mantle glands, and anterior mantle glands may be present or absent. In
<taxonomicName id="0EB23C93FFF4FFABFF04FF6AFEAC6A8B" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[175,314,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF4FFABFF04FF6AFEAC6A8B" box="[175,314,198,220]" italics="true" pageId="22" pageNumber="23">H. maculosa</emphasis>
</taxonomicName>
, posterior mantle glands are present, as are anterolateral and anterior glands. In
<taxonomicName id="0EB23C93FFF4FFABFDC5FF4BFD646AAA" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[622,754,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF4FFABFDC5FF4BFD646AAA" box="[622,754,229,251]" italics="true" pageId="22" pageNumber="23">H. decorata</emphasis>
</taxonomicName>
, the middle lateral teeth possess fewer denticles (three or four), whereas those of
<taxonomicName id="0EB23C93FFF4FFABFE60FE8EFDC96B67" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[459,607,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF4FFABFE60FE8EFDC96B67" box="[459,607,290,312]" italics="true" pageId="22" pageNumber="23">H. maculosa</emphasis>
</taxonomicName>
have four or five denticles (
<figureCitation id="51895B95FFF4FFABFEEFFEECFE016B06" box="[324,407,321,343]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
). The reproductive system of
<taxonomicName id="0EB23C93FFF4FFABFF3AFECDFE846B25" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[145,274,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF4FFABFF3AFECDFE846B25" box="[145,274,352,373]" italics="true" pageId="22" pageNumber="23">H. decorata</emphasis>
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has a very short ejaculatory segment of the vas deferens, whereas that of
<taxonomicName id="0EB23C93FFF4FFABFE48FED2FDE66BC3" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[483,624,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF4FFABFE48FED2FDE66BC3" box="[483,624,382,404]" italics="true" pageId="22" pageNumber="23">H. maculosa</emphasis>
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has a much more elongate ejaculatory segment (
<figureCitation id="51895B95FFF4FFABFDE4FE30FD306BE2" box="[591,678,413,435]" captionStart="Figure 4" captionStartId="10.[146,224,1629,1651]" captionTargetBox="[244,1328,198,1587]" captionTargetId="figure@10.[242,1330,196,1589]" captionTargetPageId="10" captionText="Figure 4. Reproductive systems A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., holotype, CASIZ19229, scale bar: 0.5 mm. B, Hypselodoris maculosa (Pease, 1871), CASIZ 139595, scale bar: 0.75 mm. C, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., paratype, WAM S12628, scale bar: 3.0 mm. D, Hypselodoris apolegma (Yonow, 2001), CASIZ 083743, scale bar: 2.0 mm. E, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype CASIZ 178350, scale bar: 0.85 mm. F, Hypselodoris krakatoa Gosliner &amp; Johnson, 1999, CASIZ 206801, Philippines, scale bar: 1.0 mm. G, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191070, scale bar: 0.60 mm. H, Hypselodoris decorata (Risbec, 1928), CASIZ 184316, scale bar: 0.60 mm. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland." httpUri="https://zenodo.org/record/2632779/files/figure.png" pageId="22" pageNumber="23">Fig. 4B</figureCitation>
). Also, the receptaculum seminis of
<taxonomicName id="0EB23C93FFF4FFABFB29FF6AFA9D6A8B" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1154,1291,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFF4FFABFB29FF6AFA9D6A8B" box="[1154,1291,198,220]" italics="true" pageId="22" pageNumber="23">H. decorata</emphasis>
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is situated immediately below the bursa, whereas it is much closer to the uterine duct in
<taxonomicName id="0EB23C93FFF4FFABFBC2FEA9FB606B49" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[1129,1270,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF4FFABFBC2FEA9FB606B49" box="[1129,1270,260,281]" italics="true" pageId="22" pageNumber="23">H. maculosa</emphasis>
</taxonomicName>
. Both species appear to be sympatric in the
<collectingCountry id="B1A50780FFF4FFABFBD0FE8FFAA86B68" box="[1147,1342,290,312]" name="Marshall Islands" pageId="22" pageNumber="23">Marshall Islands</collectingCountry>
,
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and the
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. Differences between this species and
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<emphasis id="FBC69B02FFF4FFABFC59FECDFB096B25" box="[1010,1183,352,373]" italics="true" pageId="22" pageNumber="23">H. alburtuqali</emphasis>
</taxonomicName>
and other members of
<taxonomicName id="0EB23C93FFF4FFABFCEEFED2FC406BC3" baseAuthorityName="Pease" baseAuthorityYear="1871" box="[837,982,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFF4FFABFCEEFED2FC406BC3" box="[837,982,382,404]" italics="true" pageId="22" pageNumber="23">H. maculosa</emphasis>
</taxonomicName>
are detailed in the remarks section of
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<emphasis id="FBC69B02FFF4FFABFC82FE33FC5A6BE2" box="[809,972,413,435]" italics="true" pageId="22" pageNumber="23">H. alburtuqali</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

367
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(GOHAR &amp; ABOUL-
<bibRefCitation id="AD233AE1FFF5FFAAFDAAFF49FDE16AAC" author="Gohar HAF &amp; Aboul-Ela IA" box="[513,631,228,252]" journalOrPublisher="Al-Ghardaqa, Egypt: Publications of the Marine Biological Station" pageId="23" pageNumber="24" pagination="203 - 228" refId="ref42289" refString="Gohar HAF, Aboul-Ela IA. 1957. The development of three chromodorids (with the description of a new species), Vol. 9. Al-Ghardaqa, Egypt: Publications of the Marine Biological Station, 203 - 228." title="The development of three chromodorids (with the description of a new species), Vol. 9" type="book chapter" year="1957">ELA, 1957</bibRefCitation>
)
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<emphasis id="FBC69B02FFF5FFAAFF08FE9EFE616B18" box="[163,503,307,328]" italics="true" pageId="23" pageNumber="24">Chromodoris ghardaqana</emphasis>
<bibRefCitation id="AD233AE1FFF5FFAAFDA8FE9EFF496B37" author="Gohar HAF &amp; Aboul-Ela IA" journalOrPublisher="Al-Ghardaqa, Egypt: Publications of the Marine Biological Station" pageId="23" pageNumber="24" pagination="203 - 228" refId="ref42289" refString="Gohar HAF, Aboul-Ela IA. 1957. The development of three chromodorids (with the description of a new species), Vol. 9. Al-Ghardaqa, Egypt: Publications of the Marine Biological Station, 203 - 228." title="The development of three chromodorids (with the description of a new species), Vol. 9" type="book chapter" year="1957">Gohar &amp; Aboul-Ela, 1957</bibRefCitation>
: 220
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–222, pl. 1, fig. 3, pl. 2, figs. 4, 5;
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<emphasis id="FBC69B02FFF5FFAAFD0CFEFCFEB96BD5" italics="true" pageId="23" pageNumber="24">Risbecia ghardaqana</emphasis>
<bibRefCitation id="AD233AE1FFF5FFAAFE9EFEDDFDC36BD6" author="Gohar HAF &amp; Aboul-Ela IA" box="[309,597,368,390]" journalOrPublisher="Al-Ghardaqa, Egypt: Publications of the Marine Biological Station" pageId="23" pageNumber="24" pagination="203 - 228" refId="ref42289" refString="Gohar HAF, Aboul-Ela IA. 1957. The development of three chromodorids (with the description of a new species), Vol. 9. Al-Ghardaqa, Egypt: Publications of the Marine Biological Station, 203 - 228." title="The development of three chromodorids (with the description of a new species), Vol. 9" type="book chapter" year="1957">Gohar &amp; Aboul-Ela, 1957</bibRefCitation>
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;
<bibRefCitation id="AD233AE1FFF5FFAAFDF4FEDDFC936BD5" author="Rudman WB" box="[607,773,368,390]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="23" pageNumber="24" pagination="305 - 407" part="90" refId="ref44802" refString="Rudman WB. 1987. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups. Zoological Journal of the Linnean Society 90: 305 - 407." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups" type="journal article" year="1987">Rudman, 1987</bibRefCitation>
: 374, figs 37b, 39b, 40–41.
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</collectingDate>
,
<collectorName id="644722C6FFF5FFAAFD8FFD88FD36686A" box="[548,672,548,570]" pageId="23" pageNumber="24">T. Gosliner</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF5FFAAFF08FDFBFF6468FC" pageId="23" pageNumber="24" type="distribution">
<paragraph id="C90D4710FFF5FFAAFF08FDFBFE76683E" blockId="23.[163,779,598,684]" box="[163,480,598,622]" pageId="23" pageNumber="24">
<heading id="9245F07CFFF5FFAAFF08FDFBFE76683E" box="[163,480,598,622]" fontSize="10" level="4" pageId="23" pageNumber="24" reason="1">
<emphasis id="FBC69B02FFF5FFAAFF08FDFBFE76683E" box="[163,480,598,622]" italics="true" pageId="23" pageNumber="24">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF5FFAAFF08FDDAFF6468FC" blockId="23.[163,779,598,684]" pageId="23" pageNumber="24">
Known only from the
<collectingRegion id="0B7689F2FFF5FFAAFE33FDDAFE6368DC" box="[408,501,631,653]" country="Egypt" name="Al Bahr al Ahmar" pageId="23" pageNumber="24">Red Sea</collectingRegion>
(
<bibRefCitation id="AD233AE1FFF5FFAAFDA8FDDAFF7068FC" author="Rudman WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="23" pageNumber="24" pagination="305 - 407" part="90" refId="ref44802" refString="Rudman WB. 1987. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups. Zoological Journal of the Linnean Society 90: 305 - 407." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups" type="journal article" year="1987">Rudman, 1987; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF5FFA4FF08FD63FE1369FE" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="C90D4710FFF5FFAAFF08FD63FEB968B5" blockId="23.[163,779,718,894]" box="[163,303,718,741]" pageId="23" pageNumber="24">
<heading id="9245F07CFFF5FFAAFF08FD63FEB968B5" box="[163,303,718,741]" fontSize="10" level="4" pageId="23" pageNumber="24" reason="1">
<emphasis id="FBC69B02FFF5FFAAFF08FD63FEB968B5" box="[163,303,718,741]" italics="true" pageId="23" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF5FFAAFF08FD40FC9C692E" blockId="23.[163,779,718,894]" pageId="23" pageNumber="24">
<emphasis id="FBC69B02FFF5FFAAFF08FD40FE036952" box="[163,405,749,770]" italics="true" pageId="23" pageNumber="24">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFF5FFAAFDCCFD43FD2D6953" box="[615,699,750,772]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="23" pageNumber="24">Fig. 1E</figureCitation>
) large, reaching
<quantity id="0E4AEAF5FFF5FFAAFEB8FCA1FEF36972" box="[275,357,780,802]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="24" unit="mm" value="75.0">75 mm</quantity>
in length. Body translucent white, with irregular large yellowish spots and a continuous dark blue marginal band. Translucent white foot ornamented by dark blue marginal band. Gill white,
</paragraph>
<paragraph id="C90D4710FFF5FFAAFC90FF6BFA346A8C" blockId="23.[827,1443,199,312]" box="[827,1442,198,220]" pageId="23" pageNumber="24">with dark blue line on interior and outer edge or gill</paragraph>
<paragraph id="C90D4710FFF5FFAAFC90FF48FB686B68" blockId="23.[827,1443,199,312]" pageId="23" pageNumber="24">rachis. Thirteen unipinnate gill branches. Perfoliate rhinophores white basally and dark blue apically, bearing ~18 densely arranged lamellae.</paragraph>
<paragraph id="C90D4710FFF5FFAAFC90FECCFAAE6BE4" blockId="23.[827,1443,353,436]" pageId="23" pageNumber="24">
<emphasis id="FBC69B02FFF5FFAAFC90FECCFC736B26" box="[827,997,353,374]" italics="true" pageId="23" pageNumber="24">Mantle glands</emphasis>
: Subcutaneous mantle glands simple rounded in shape (
<figureCitation id="51895B95FFF5FFAAFBBBFE2DFBF76BC5" box="[1040,1121,384,406]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="23" pageNumber="24">Fig. 2E</figureCitation>
). Glands dense, found along entire mantle margin except at anterior end.
</paragraph>
<paragraph id="C90D4710FFF5FFA4FC90FE70FD766B07" blockId="23.[827,1443,477,898]" lastBlockId="25.[163,779,198,343]" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<emphasis id="FBC69B02FFF5FFAAFC90FE70FB936BA3" box="[827,1029,477,499]" italics="true" pageId="23" pageNumber="24">Buccal armature</emphasis>
: Muscular portion of buccal mass approximately equal in length to oral tube. Buccal mass consisting of chitinous labial cuticle at anterior end of muscular portion of buccal mass. Jaws bearing numerous rodlets (
<figureCitation id="51895B95FFF5FFAAFB9DFDF5FB33683D" box="[1078,1189,600,622]" captionStart="Figure 12" captionStartId="23.[163,242,1850,1872]" captionTargetBox="[227,1379,935,1809]" captionTargetId="figure@23.[227,1379,935,1809]" captionTargetPageId="23" captionText="Figure 12. Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632795/files/figure.png" pageId="23" pageNumber="24">Fig. 12A</figureCitation>
). Rodlets narrowly triangular, with single, acutely pointed apex. Radular formula of
<typeStatus id="1609F9B2FFF5FFAAFC19FD38FB8768FB" box="[946,1041,661,683]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
63 × 55.0.55. Rachidian row of teeth absent (
<figureCitation id="51895B95FFF5FFAAFC3FFD19FC666899" box="[916,1008,692,714]" captionStart="Figure 12" captionStartId="23.[163,242,1850,1872]" captionTargetBox="[227,1379,935,1809]" captionTargetId="figure@23.[227,1379,935,1809]" captionTargetPageId="23" captionText="Figure 12. Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632795/files/figure.png" pageId="23" pageNumber="24">Fig. 12B</figureCitation>
). Innermost lateral teeth have a single small triangular denticle on inner side of bifid primary cusp and lacking outer denticles. Next several laterals lacking inner triangular denticle and also lacking denticles on outer side of primary bifid cusps. Outer lateral cusp much shorter than inner one. Midlateral teeth (
<figureCitation id="51895B95FFF5FFAAFC3BFCC1FC6C69D1" box="[912,1018,876,898]" captionStart="Figure 12" captionStartId="23.[163,242,1850,1872]" captionTargetBox="[227,1379,935,1809]" captionTargetId="figure@23.[227,1379,935,1809]" captionTargetPageId="23" captionText="Figure 12. Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632795/files/figure.png" pageId="23" pageNumber="24">Fig. 12C</figureCitation>
) also lacking inner denticles but possessing five to ten triangular outer denticles. Outer cusp of bifid cusp much shorter than inner one. Outer teeth lacking inner denticles and having six to eight triangular outer denticles (
<figureCitation id="51895B95FFFBFFA4FE73FE8FFDAE6B68" box="[472,568,290,312]" captionStart="Figure 12" captionStartId="23.[163,242,1850,1872]" captionTargetBox="[227,1379,935,1809]" captionTargetId="figure@23.[227,1379,935,1809]" captionTargetPageId="23" captionText="Figure 12. Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632795/files/figure.png" pageId="25" pageNumber="26">Fig. 12D</figureCitation>
). Outermost teeth with narrower base and more elongate tooth shape.
</paragraph>
<caption id="9DCD1798FFF5FFAAFF08F897FDF46D3D" httpUri="https://zenodo.org/record/2632795/files/figure.png" pageId="23" pageNumber="24" startId="23.[163,242,1850,1872]" targetBox="[227,1379,935,1809]" targetPageId="23">
<paragraph id="C90D4710FFF5FFAAFF08F897FDF46D3D" blockId="23.[163,1443,1849,1901]" pageId="23" pageNumber="24">
<emphasis id="FBC69B02FFF5FFAAFF08F897FE8F6D1F" bold="true" box="[163,281,1850,1872]" pageId="23" pageNumber="24">Figure 12.</emphasis>
<taxonomicName id="0EB23C93FFF5FFAAFE8FF897FCC06D1F" authority="(Gohar &amp; Aboul-Ela, 1957)" authorityName="Gohar &amp; Aboul-Ela" authorityYear="1957" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[292,854,1849,1872]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFF5FFAAFE8FF897FDA26D1F" box="[292,564,1850,1871]" italics="true" pageId="23" pageNumber="24">Hypselodoris ghardaqana</emphasis>
(Gohar &amp; Aboul-Ela, 1957)
</taxonomicName>
, CASIZ 192282. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<caption id="9DCD1798FFFAFFA5FF24F88FFA196D22" httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="24" pageNumber="25" startId="24.[143,224,1826,1848]" targetBox="[207,1359,196,1783]" targetPageId="24">
<paragraph id="C90D4710FFFAFFA5FF24F88FFA196D22" blockId="24.[143,1424,1825,1907]" pageId="24" pageNumber="25">
<emphasis id="FBC69B02FFFAFFA5FF24F88FFE9F6D67" bold="true" box="[143,265,1826,1848]" pageId="24" pageNumber="25">Figure 13.</emphasis>
Reproductive systems. A,
<taxonomicName id="0EB23C93FFFAFFA5FD86F88FFBE46D67" authority="(Gohar &amp; Aboul-Ela, 1957)" authorityName="Gohar &amp; Aboul-Ela" authorityYear="1957" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[557,1138,1825,1848]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFAFFA5FD86F88FFCD06D67" box="[557,838,1826,1847]" italics="true" pageId="24" pageNumber="25">Hypselodoris ghardaqana</emphasis>
(Gohar &amp; Aboul-Ela, 1957)
</taxonomicName>
, CASIZ 192282, scale bar: 3.5 mm. B,
<taxonomicName id="0EB23C93FFFAFFA5FEAAF892FD026D05" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[257,660,1855,1877]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFFAFFA5FEAAF892FE016D05" bold="true" box="[257,407,1855,1877]" italics="true" pageId="24" pageNumber="25">Hypselodoris</emphasis>
<emphasis id="FBC69B02FFFAFFA5FE37F892FE546D05" bold="true" box="[412,450,1855,1877]" italics="true" pageId="24" pageNumber="25">iba</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFAFFA5FD32F8EDFD796D05" box="[665,751,1856,1877]" pageId="24" pageNumber="25" rank="species">
<emphasis id="FBC69B02FFFAFFA5FD32F8EDFD796D05" bold="true" box="[665,751,1856,1877]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ 177777, scale bar: 3.0 mm. C,
<taxonomicName id="0EB23C93FFFAFFA5FBD8F892FEF56D23" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFFAFFA5FBD8F892FA9F6D05" bold="true" box="[1139,1289,1855,1877]" italics="true" pageId="24" pageNumber="25">Hypselodoris</emphasis>
<emphasis id="FBC69B02FFFAFFA5FAA5F892FA196D05" bold="true" box="[1294,1423,1855,1877]" italics="true" pageId="24" pageNumber="25">katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFAFFA5FEC1F8F0FE546D22" box="[362,450,1885,1906]" pageId="24" pageNumber="25" rank="species">
<emphasis id="FBC69B02FFFAFFA5FEC1F8F0FE546D22" bold="true" box="[362,450,1885,1906]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ 181257, scale bar: 1.0 mm. D,
<taxonomicName id="0EB23C93FFFAFFA5FCF7F8F1FABC6D23" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[860,1322,1884,1907]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="lacuna" status="sp. nov.">
<emphasis id="FBC69B02FFFAFFA5FCF7F8F1FC636D22" bold="true" box="[860,1013,1884,1906]" italics="true" pageId="24" pageNumber="25">Hypselodoris</emphasis>
<emphasis id="FBC69B02FFFAFFA5FC57F8F1FBC66D22" bold="true" box="[1020,1104,1884,1906]" italics="true" pageId="24" pageNumber="25">lacuna</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFAFFA5FA9AF8F0FA1C6D22" box="[1329,1418,1885,1906]" pageId="24" pageNumber="25" rank="species">
<emphasis id="FBC69B02FFFAFFA5FA9AF8F0FA1C6D22" bold="true" box="[1329,1418,1885,1906]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
,
</paragraph>
</caption>
<paragraph id="C90D4710FFFBFFA4FF08FEDDFE1369FE" blockId="25.[163,779,368,942]" pageId="25" pageNumber="26">
<emphasis id="FBC69B02FFFBFFA4FF08FEDDFE3F6BD5" box="[163,425,368,389]" italics="true" pageId="25" pageNumber="26">Reproductive system</emphasis>
: Reproductive organs fully mature (
<figureCitation id="51895B95FFFBFFA4FEAEFE23FEF26BF4" box="[261,356,398,420]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="25" pageNumber="26">Fig. 13A</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into the oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens curved and thick, narrowing slightly while transitioning into muscular, ejaculatory portion. Ejaculatory portion curving into segment entering elongate, slightly widened penial bulb. Penial bulb adjacent to curved, wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Elongate, curved vagina leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Relatively short uterine duct emerging from about half of the length along duct to bursa.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFBFFA4FF08FC78FC5A685E" pageId="25" pageNumber="26" type="discussion">
<paragraph id="C90D4710FFFBFFA4FF08FC78FE8669BD" blockId="25.[163,779,981,1618]" box="[163,272,981,1005]" pageId="25" pageNumber="26">
<heading id="9245F07CFFFBFFA4FF08FC78FE8669BD" box="[163,272,981,1005]" fontSize="10" level="4" pageId="25" pageNumber="26" reason="1">
<emphasis id="FBC69B02FFFBFFA4FF08FC78FE8669BD" box="[163,272,981,1005]" italics="true" pageId="25" pageNumber="26">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFBFFA4FF08FC58FC5A685E" blockId="25.[163,779,981,1618]" lastBlockId="25.[827,1443,198,527]" pageId="25" pageNumber="26">
This species was originally described as a species of
<taxonomicName id="0EB23C93FFFBFFA4FF08FBB9FEA16E79" authorityName="Alder &amp; Hancock" authorityYear="1855" box="[163,311,1044,1065]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FF08FBB9FEA16E79" box="[163,311,1044,1065]" italics="true" pageId="25" pageNumber="26">Chromodoris</emphasis>
</taxonomicName>
distinct from both
<taxonomicName id="0EB23C93FFFBFFA4FDA3FBB9FEC36E19" authority="(Ruppell &amp; Leuckart, 1828)" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="pulchella">
<emphasis id="FBC69B02FFFBFFA4FDA3FBB9FD1A6E79" box="[520,652,1044,1065]" italics="true" pageId="25" pageNumber="26">C. pulchella</emphasis>
(Rüppell &amp; Leuckart, 1828)
</taxonomicName>
and
<taxonomicName id="0EB23C93FFFBFFA4FE21FB9FFC9D6E18" authority="Eliot, 1904" authorityName="Eliot" authorityYear="1904" box="[394,779,1074,1096]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="annulata">
<emphasis id="FBC69B02FFFBFFA4FE21FB9FFD1A6E17" box="[394,652,1074,1095]" italics="true" pageId="25" pageNumber="26">Chromodoris annulata</emphasis>
<bibRefCitation id="AD233AE1FFFBFFA4FD3AFB9FFC9D6E18" author="Eliot C." box="[657,779,1074,1096]" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="25" pageNumber="26" pagination="380 - 406" part="1904" refId="ref41933" refString="Eliot C. 1904. On some nudibranchs from East Africa and Zanzibar. Part IV. Proceedings of the Zoological Society of London 1904: 380 - 406." title="On some nudibranchs from East Africa and Zanzibar. Part IV" type="journal article" year="1904">Eliot, 1904</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFFBFFA4FF00FBFCFE2E6E37" author="Gohar HAF &amp; Aboul-Ela IA" box="[171,440,1105,1127]" journalOrPublisher="Al-Ghardaqa, Egypt: Publications of the Marine Biological Station" pageId="25" pageNumber="26" pagination="203 - 228" refId="ref42289" refString="Gohar HAF, Aboul-Ela IA. 1957. The development of three chromodorids (with the description of a new species), Vol. 9. Al-Ghardaqa, Egypt: Publications of the Marine Biological Station, 203 - 228." title="The development of three chromodorids (with the description of a new species), Vol. 9" type="book chapter" year="1957">Gohar &amp; Abu-Ela, 1957</bibRefCitation>
).
<bibRefCitation id="AD233AE1FFFBFFA4FE65FBFCFDE96E37" author="Rudman WB" box="[462,639,1105,1127]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="25" pageNumber="26" pagination="305 - 407" part="90" refId="ref44802" refString="Rudman WB. 1987. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups. Zoological Journal of the Linnean Society 90: 305 - 407." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Chromodoris epicuria, C. aureopurpurea, C. annulata, C. coi and Risbecia tryoni colour groups" type="journal article" year="1987">Rudman (1987)</bibRefCitation>
maintained the distinctness of the three species but transferred
<taxonomicName id="0EB23C93FFFBFFA4FF08FB22FEBE6EF3" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[163,296,1166,1188]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="pulchella">
<emphasis id="FBC69B02FFFBFFA4FF08FB22FEBE6EF3" box="[163,296,1166,1188]" italics="true" pageId="25" pageNumber="26">C. pulchella</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFFBFFA4FEF5FB22FD906EF3" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[350,518,1166,1188]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FEF5FB22FD906EF3" box="[350,518,1166,1188]" italics="true" pageId="25" pageNumber="26">C. ghardaqana</emphasis>
</taxonomicName>
to
<taxonomicName id="0EB23C93FFFBFFA4FD8CFB23FD106EF3" authorityName="Odhner" authorityYear="1934" box="[551,646,1166,1187]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FD8CFB23FD106EF3" box="[551,646,1166,1187]" italics="true" pageId="25" pageNumber="26">Risbecia</emphasis>
</taxonomicName>
.
<bibRefCitation id="AD233AE1FFFBFFA4FD39FB23FEC66E93" author="Johnson R &amp; Gosliner T." journalOrPublisher="PLoS One" pageId="25" pageNumber="26" pagination="33479" part="7" refId="ref43042" refString="Johnson R, Gosliner T. 2012. Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs. PLoS One 7: e 33479." title="Traditional taxonomic groupings mask evolutionary history: a molecular phylogeny and new classification of the chromodorid nudibranchs" type="journal article" year="2012">Johnson &amp; Gosliner (2012)</bibRefCitation>
, based on molecular phylogeny, showed that species of
<taxonomicName id="0EB23C93FFFBFFA4FEE2FB61FE3E6EB1" authorityName="Odhner" authorityYear="1934" box="[329,424,1228,1249]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FEE2FB61FE3E6EB1" box="[329,424,1228,1249]" italics="true" pageId="25" pageNumber="26">Risbecia</emphasis>
</taxonomicName>
are nested in
<taxonomicName id="0EB23C93FFFBFFA4FDECFB61FD4C6EB1" authorityName="Stimpson" authorityYear="1855" box="[583,730,1228,1249]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FDECFB61FD4C6EB1" box="[583,730,1228,1249]" italics="true" pageId="25" pageNumber="26">Hypselodoris</emphasis>
</taxonomicName>
and that maintenance of
<taxonomicName id="0EB23C93FFFBFFA4FE31FB47FE6B6EAF" authorityName="Odhner" authorityYear="1934" box="[410,509,1258,1279]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FE31FB47FE6B6EAF" box="[410,509,1258,1279]" italics="true" pageId="25" pageNumber="26">Risbecia</emphasis>
</taxonomicName>
creates a paraphyletic
<taxonomicName id="0EB23C93FFFBFFA4FF08FAA4FEA06F4E" authorityName="Stimpson" authorityYear="1855" box="[163,310,1289,1310]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FF08FAA4FEA06F4E" box="[163,310,1289,1310]" italics="true" pageId="25" pageNumber="26">Hypselodoris</emphasis>
</taxonomicName>
. They also demonstrated that
<taxonomicName id="0EB23C93FFFBFFA4FD2CFAA4FC9D6F4E" authorityName="Eliot" authorityYear="1904" box="[647,779,1289,1310]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="annulata">
<emphasis id="FBC69B02FFFBFFA4FD2CFAA4FC9D6F4E" box="[647,779,1289,1310]" italics="true" pageId="25" pageNumber="26">C. annulata</emphasis>
</taxonomicName>
should be considered as a species of
<taxonomicName id="0EB23C93FFFBFFA4FDF8FA85FC936F6D" authorityName="Pease" authorityYear="1866" box="[595,773,1320,1341]" class="Gastropoda" family="Chromodorididae" genus="Goniobranchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FDF8FA85FC936F6D" box="[595,773,1320,1341]" italics="true" pageId="25" pageNumber="26">Goniobranchus</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFFBFFA4FF08FAEBFE406F0B" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[163,470,1350,1371]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FF08FAEBFE406F0B" box="[163,470,1350,1371]" italics="true" pageId="25" pageNumber="26">Hypselodoris ghardaqana</emphasis>
</taxonomicName>
was not included in their analysis but was hypothesized to be a member of the
<taxonomicName id="0EB23C93FFFBFFA4FF08FA29FE9F6FC9" authorityName="Odhner" authorityYear="1934" box="[163,265,1412,1433]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FF08FA29FE9F6FC9" box="[163,265,1412,1433]" italics="true" pageId="25" pageNumber="26">Risbecia</emphasis>
</taxonomicName>
clade of
<emphasis id="FBC69B02FFFBFFA4FEDCFA29FD8D6FC9" box="[375,539,1412,1433]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FEDCFA29FD806FC9" authorityName="Stimpson" authorityYear="1855" box="[375,534,1412,1433]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
.
</emphasis>
A surprising result of the present study is that
<taxonomicName id="0EB23C93FFFBFFA4FE5FFA0EFD3F6FE7" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[500,681,1442,1464]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FE5FFA0EFD3F6FE7" box="[500,681,1442,1464]" italics="true" pageId="25" pageNumber="26">H. ghardaqana</emphasis>
</taxonomicName>
is not a close relative of the members of the
<taxonomicName id="0EB23C93FFFBFFA4FDF0FA6CFD296F86" authorityName="Odhner" authorityYear="1934" box="[603,703,1473,1494]" class="Gastropoda" family="Chromodorididae" genus="Risbecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FDF0FA6CFD296F86" box="[603,703,1473,1494]" italics="true" pageId="25" pageNumber="26">Risbecia</emphasis>
</taxonomicName>
clade, but rather is sister to
<taxonomicName id="0EB23C93FFFBFFA4FE6CFA4DFDC16FA5" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[455,599,1504,1525]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bollandi">
<emphasis id="FBC69B02FFFBFFA4FE6CFA4DFDC16FA5" box="[455,599,1504,1525]" italics="true" pageId="25" pageNumber="26">H. bollandi</emphasis>
</taxonomicName>
.
<emphasis id="FBC69B02FFFBFFA4FDC2FA4DFEAD6C43" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FDC2FA4DFC9C6FA5" authorityName="Stimpson" authorityYear="1855" box="[617,778,1504,1525]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">Hypselodoris</taxonomicName>
nigrolineata
</emphasis>
is sister to both
<taxonomicName id="0EB23C93FFFBFFA4FDBCFA52FD426C43" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[535,724,1534,1556]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FDBCFA52FD426C43" box="[535,724,1534,1556]" italics="true" pageId="25" pageNumber="26">H. ghardaqana</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFFBFFA4FF08F9B3FEB46C62" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[163,290,1565,1587]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bollandi">
<emphasis id="FBC69B02FFFBFFA4FF08F9B3FEB46C62" box="[163,290,1565,1587]" italics="true" pageId="25" pageNumber="26">H. bollandi</emphasis>
</taxonomicName>
. The colour pattern of
<taxonomicName id="0EB23C93FFFBFFA4FDB6F9B3FD0D6C62" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[541,667,1565,1587]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bollandi">
<emphasis id="FBC69B02FFFBFFA4FDB6F9B3FD0D6C62" box="[541,667,1565,1587]" italics="true" pageId="25" pageNumber="26">H. bollandi</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0EB23C93FFFBFFA4FEBCF991FE5E6C01" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[279,456,1596,1617]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FEBCF991FE5E6C01" box="[279,456,1596,1617]" italics="true" pageId="25" pageNumber="26">H. ghardaqana</emphasis>
</taxonomicName>
in that both species have a white body colour with yellow spots. The rhinophores and gill of
<taxonomicName id="0EB23C93FFFBFFA4FC1CFF4BFBFE6AAA" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[951,1128,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FC1CFF4BFBFE6AAA" box="[951,1128,229,251]" italics="true" pageId="25" pageNumber="26">H. ghardaqana</emphasis>
</taxonomicName>
are ornamented with dark blue pigment, whereas they are ornamented with red in
<taxonomicName id="0EB23C93FFFBFFA4FCF0FE8EFC756B67" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[859,995,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bollandi">
<emphasis id="FBC69B02FFFBFFA4FCF0FE8EFC756B67" box="[859,995,290,312]" italics="true" pageId="25" pageNumber="26">H. bollandi</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFFBFFA4FC59FE8FFB6F6B67" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1010,1273,290,311]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bollandi">
<emphasis id="FBC69B02FFFBFFA4FC59FE8FFB6F6B67" box="[1010,1273,290,311]" italics="true" pageId="25" pageNumber="26">Hypselodoris bollandi</emphasis>
</taxonomicName>
also has dark brown pigment on the notum. In all three species, the middle lateral teeth bifid cusps have a characteristic shape, with an elongated inner cusp and a short outer cusp, with numerous denticles below the outer cusp. The colour pattern of
<taxonomicName id="0EB23C93FFFBFFA4FBEBFE11FB636B81" baseAuthorityName="Gohar &amp; Aboul-Ela" baseAuthorityYear="1957" box="[1088,1269,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ghardaqana">
<emphasis id="FBC69B02FFFBFFA4FBEBFE11FB636B81" box="[1088,1269,444,465]" italics="true" pageId="25" pageNumber="26">H. ghardaqana</emphasis>
</taxonomicName>
is remarkably convergent with that of its sympatric congener,
<emphasis id="FBC69B02FFFBFFA4FC90FE57FC5A685E" box="[827,972,505,527]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FC90FE57FC5E685E" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1828" box="[827,968,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="pulchella">H. pulchella</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

658
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<mods:identifier id="BF7EE4D304403808C532F56777860E5D" type="ZooBank">urn:lsid:zoobank.org:pub:F0065FD2-417C</mods:identifier>
</mods:mods>
<treatment id="411BF606FFFBFFA1FCEAFDE1FB626BA0" ID-DOI="http://doi.org/10.5281/zenodo.5980678" ID-GBIF-Taxon="156146725" ID-Zenodo-Dep="5980678" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0" httpUri="http://treatment.plazi.org/id/411BF606FFFBFFA1FCEAFDE1FB626BA0" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="81A8149BFFFBFFA4FCEAFDE1FB1A68BF" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="C90D4710FFFBFFA4FCEAFDE1FB7C68DA" blockId="25.[833,1436,587,650]" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FCEAFDE1FAA46832" ID-CoL="3P33T" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[833,1330,587,611]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="28" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FCEAFDE1FBBE6832" bold="true" box="[833,1064,588,611]" italics="true" pageId="25" pageNumber="26">HYPSELODORIS IBA</emphasis>
GOSLINER &amp; JOHNSON
</taxonomicName>
<emphasis id="FBC69B02FFFBFFA4FA92FDE2FA0A6833" bold="true" box="[1337,1436,587,611]" pageId="25" pageNumber="26">SP. NOV.</emphasis>
(FIGS 2F, 13B, 14, 15)
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FD16FB1A68BF" blockId="25.[827,1444,698,751]" pageId="25" pageNumber="26">
<emphasis id="FBC69B02FFFBFFA4FC90FD16FCE86880" box="[827,894,699,720]" italics="true" pageId="25" pageNumber="26">LSID</emphasis>
:
<uri id="BD234B12FFFBFFA4FC39FD17FB1A68BF" pageId="25" pageNumber="26">
urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFFBFFA4FB14FD17FB1A68BF" pageId="25" pageNumber="26">A5DF9265-3D8B- 4BB8-B54D-DC66C89C7BFC</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFBFFA4FC90FCA3FB966E48" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="C90D4710FFFBFFA4FC90FCA3FC46692F" blockId="25.[827,1443,781,1048]" pageId="25" pageNumber="26">
<emphasis id="FBC69B02FFFBFFA4FC90FCA3FBF46973" box="[827,1122,781,803]" italics="true" pageId="25" pageNumber="26">H y p s e l o d o r i s b u l l o ck i</emphasis>
, m i s i d e n t i f i c a t i o n, n o t
<taxonomicName id="0EB23C93FFFBFFA4FC90FC81FAB96912" ID-CoL="5YHJT" authority="(Collingwood, 1881)" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[827,1327,812,834]" class="Gastropoda" family="Chromodorididae" genus="Chromodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFFBFFA4FC90FC81FBA96911" box="[827,1087,812,833]" italics="true" pageId="25" pageNumber="26">Chromodoris bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFFBFFA4FBE5FC81FAB16912" author="Collingwood C." box="[1102,1319,812,834]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="25" pageNumber="26" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="AD233AE1FFFBFFA4FA97FC81FB9A6930" author="Humann P &amp; DeLoach N." journalOrPublisher="Jacksonville: New World Publications" pageId="25" pageNumber="26" refId="ref42986" refString="Humann P, DeLoach N. 2010. Reef creature identification, tropical Pacific. Jacksonville: New World Publications." title="Reef creature identification, tropical Pacific" type="book" year="2010">Humann &amp; DeLoach, 2010</bibRefCitation>
: 338, middle right and lower left photographs.
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FC25FAC869CD" blockId="25.[827,1443,781,1048]" box="[827,1374,904,926]" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FC90FC25FC5869CD" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[827,974,904,925]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FC90FC25FC5869CD" box="[827,974,904,925]" italics="true" pageId="25" pageNumber="26">Hypselodoris</emphasis>
</taxonomicName>
sp. 5
<bibRefCitation id="AD233AE1FFFBFFA4FBA4FC25FAB769CD" author="Debelius H &amp; Kuiter R." box="[1039,1313,904,926]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="25" pageNumber="26" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
: 118.
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FC0BFC53698B" blockId="25.[827,1443,781,1048]" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FC90FC0BFC4F69EB" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[827,985,934,955]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FC90FC0BFC4F69EB" box="[827,985,934,955]" italics="true" pageId="25" pageNumber="26">Hypselodoris</emphasis>
</taxonomicName>
sp. 10
<bibRefCitation id="AD233AE1FFFBFFA4FB9FFC0BFAA669EC" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[1076,1328,934,956]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="25" pageNumber="26" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFFBFFA4FB0DFC0AFB7369EB" box="[1190,1253,934,956]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
, 2008
</bibRefCitation>
: 269, top photograph.
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FC49FB966E48" blockId="25.[827,1443,781,1048]" pageId="25" pageNumber="26">
<taxonomicName id="0EB23C93FFFBFFA4FC90FC49FC4769A9" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[827,977,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFBFFA4FC90FC49FC4769A9" box="[827,977,996,1017]" italics="true" pageId="25" pageNumber="26">Hypselodoris</emphasis>
</taxonomicName>
sp. 14
<bibRefCitation id="AD233AE1FFFBFFA4FB88FC49FA9B69AA" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[1059,1293,996,1018]" journalOrPublisher="Jacksonville: New World Press" pageId="25" pageNumber="26" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFFBFFA4FB26FC49FB5169A9" box="[1165,1223,996,1017]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
, 2015
</bibRefCitation>
: 261, bottom two photographs.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFBFFA7FC90FBEDFE8F6EA1" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="C90D4710FFFBFFA4FC90FBEDFC726E07" blockId="25.[827,1443,1087,1264]" box="[827,996,1087,1111]" pageId="25" pageNumber="26">
<heading id="9245F07CFFFBFFA4FC90FBEDFC726E07" box="[827,996,1087,1111]" fontSize="10" level="4" pageId="25" pageNumber="26" reason="1">
<emphasis id="FBC69B02FFFBFFA4FC90FBEDFC726E07" box="[827,996,1087,1111]" italics="true" pageId="25" pageNumber="26">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FBCDFC536EA0" blockId="25.[827,1443,1087,1264]" pageId="25" pageNumber="26">
<materialsCitation id="79DA4D4DFFFBFFA4FC90FBCDFC536EA0" ID-GBIF-Occurrence="2235579106" collectingDate="2008-03-21" collectionCode="NMP" collectorName="P. Paleracio." country="Philippines" county="Maricaban Island" elevation="35" latitude="13.65835" location="Aphol's Reef" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" pageId="25" pageNumber="26" specimenCode="CASIZ 177511" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
<typeStatus id="1609F9B2FFFBFFA4FC90FBCDFC306E25" box="[827,934,1120,1141]" pageId="25" pageNumber="26" type="holotype">
<emphasis id="FBC69B02FFFBFFA4FC90FBCDFC306E25" box="[827,934,1120,1141]" italics="true" pageId="25" pageNumber="26">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="AFA3DFD5FFFBFFA4FC11FBCDFC6F6E25" box="[954,1017,1120,1141]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="25" pageNumber="26">NMP</collectionCode>
0 41279 (formerly
<specimenCode id="9914EF6BFFFBFFA4FB4AFBCDFA006E26" box="[1249,1430,1120,1142]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 177511</specimenCode>
), subsampled for molecular study,
<location id="CC6D11CBFFFBFFA4FB53FBD3FA096EC4" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFFBFFA4FB53FBD3FA096EC4" box="[1272,1439,1150,1172]" country="Philippines" county="Maricaban Island" latitude="13.65835" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" name="Aphol's Reef" pageId="25" pageNumber="26" stateProvince="Batangas">Aphol’s Reef</location>
,
<geoCoordinate id="AC8621D7FFFBFFA4FC90FB30FC556EE3" box="[827,963,1181,1203]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="13.65835">13.65835°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFBFFA4FC65FB30FBF46EE2" box="[974,1122,1181,1202]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="120.90144">120.90144°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFFBFFA4FBC6FB30FB526EE3" box="[1133,1220,1181,1203]" pageId="25" pageNumber="26">Tingloy</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFFBFFA4FB64FB30FA096EE3" box="[1231,1439,1181,1203]" pageId="25" pageNumber="26">Maricaban Island</collectingCounty>
,
<collectingRegion id="0B7689F2FFFBFFA4FC90FB11FC386E81" box="[827,942,1212,1233]" country="Philippines" name="Batangas" pageId="25" pageNumber="26">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFFBFFA4FC11FB11FBD26E82" box="[954,1092,1212,1234]" name="Philippines" pageId="25" pageNumber="26">Philippines</collectingCountry>
,
<elevation id="429FA023FFFBFFA4FBFAFB11FB1B6E81" box="[1105,1165,1212,1234]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">
<quantity id="0E4AEAF5FFFBFFA4FBFAFB11FB1B6E81" box="[1105,1165,1212,1234]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">35 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFFBFFA4FB42FB11FA086E81" box="[1257,1438,1212,1234]" pageId="25" pageNumber="26" value="2008-03-21">
<date id="BD0C61D0FFFBFFA4FB42FB11FA086E81" box="[1257,1438,1212,1234]" pageId="25" pageNumber="26" value="2008-03-21">21 March 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFFBFFA4FC90FB76FC536EA0" box="[827,965,1242,1264]" pageId="25" pageNumber="26">P. Paleracio.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFFBFFA4FC90FAA7FA356C02" blockId="25.[827,1443,1289,1618]" pageId="25" pageNumber="26">
<materialsCitation id="79DA4D4DFFFBFFA4FC90FAA7FA096F2B" ID-GBIF-Occurrence="2235579107" collectingDate="2008-04-23" collectionCode="CASIZ" collectorName="P. Paleracio" country="Philippines" county="Maricaban Island" elevation="35" latitude="13.65835" location="Aphol's Reef" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" pageId="25" pageNumber="26" specimenCode="CASIZ 177777" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<typeStatus id="1609F9B2FFFBFFA4FC90FAA7FC236F4F" box="[827,949,1290,1311]" pageId="25" pageNumber="26" type="paratype">
<emphasis id="FBC69B02FFFBFFA4FC90FAA7FC236F4F" box="[827,949,1290,1311]" italics="true" pageId="25" pageNumber="26">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFFBFFA4FC6DFAA4FBE86F4E" box="[966,1150,1289,1310]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 177777</specimenCode>
, dissected, subsampled for molecular study,
<location id="CC6D11CBFFFBFFA4FBFBFA85FB616F6E" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFFBFFA4FBFBFA85FB616F6E" box="[1104,1271,1320,1342]" country="Philippines" county="Maricaban Island" latitude="13.65835" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" name="Aphol's Reef" pageId="25" pageNumber="26" stateProvince="Batangas">Aphol’s Reef</location>
,
<geoCoordinate id="AC8621D7FFFBFFA4FAACFA85FA086F6E" box="[1287,1438,1320,1342]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="13.65835">13.65835°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFBFFA4FC90FAEAFC426F0C" box="[827,980,1351,1372]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="120.90144">120.90144°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFFBFFA4FC4AFAEBFBAD6F0C" box="[993,1083,1350,1372]" pageId="25" pageNumber="26">Tingloy</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFFBFFA4FBECFAEBFA896F0C" box="[1095,1311,1350,1372]" pageId="25" pageNumber="26">Maricaban Island</collectingCounty>
,
<collectingRegion id="0B7689F2FFFBFFA4FA87FAEAFA086F0C" box="[1324,1438,1351,1372]" country="Philippines" name="Batangas" pageId="25" pageNumber="26">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFFBFFA4FC90FAC8FC556F2B" box="[827,963,1381,1403]" name="Philippines" pageId="25" pageNumber="26">Philippines</collectingCountry>
,
<elevation id="429FA023FFFBFFA4FC64FAC8FB9C6F2B" box="[975,1034,1381,1403]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">
<quantity id="0E4AEAF5FFFBFFA4FC64FAC8FB9C6F2B" box="[975,1034,1381,1403]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">35 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFFBFFA4FBCFFAC8FA9E6F2A" box="[1124,1288,1381,1403]" pageId="25" pageNumber="26" value="2008-04-23">
<date id="BD0C61D0FFFBFFA4FBCFFAC8FA9E6F2A" box="[1124,1288,1381,1403]" pageId="25" pageNumber="26" value="2008-04-23">23 April 2008</date>
</collectingDate>
,
<collectorName id="644722C6FFFBFFA4FABFFACBFA096F2B" box="[1300,1439,1381,1403]" pageId="25" pageNumber="26">P. Paleracio</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFFBFFA4FC90FA29FB4D6FA6" ID-GBIF-Occurrence="2235579140" collectingDate="2009-05-17" collectionCode="CASIZ" collectorName="P. Paleracio" country="Philippines" county="Maricaban Island" elevation="35" latitude="13.65835" location="Aphol's Reef" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" pageId="25" pageNumber="26" specimenCode="CASIZ 180419" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFBFFA4FC90FA29FC7E6FC9" box="[827,1000,1412,1433]" collectionCode="CASIZ" pageId="25" pageNumber="26">
CASIZ
<date id="BD0C61D0FFFBFFA4FC3AFA29FC7E6FC9" box="[913,1000,1412,1433]" pageId="25" pageNumber="26" value="19-04-18">180419</date>
</specimenCode>
, dissected, subsampled for molecular study,
<location id="CC6D11CBFFFBFFA4FC28FA0FFB846FE8" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFFBFFA4FC28FA0FFB846FE8" box="[899,1042,1442,1464]" country="Philippines" county="Maricaban Island" latitude="13.65835" longLatPrecision="1" longitude="120.90144" municipality="Tingloy" name="Aphol's Reef" pageId="25" pageNumber="26" stateProvince="Batangas">Aphol’s Reef</location>
,
<geoCoordinate id="AC8621D7FFFBFFA4FBB7FA0FFB376FE8" box="[1052,1185,1442,1464]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="13.65835">13.65835°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFBFFA4FB06FA0EFAA96FE8" box="[1197,1343,1443,1464]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="120.90144">120.90144°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFFBFFA4FAE1FA0FFA096FE8" box="[1354,1439,1442,1464]" pageId="25" pageNumber="26">Tingloy</collectingMunicipality>
,
<collectingCounty id="206C3F9CFFFBFFA4FC90FA6CFBB56F87" box="[827,1059,1473,1495]" pageId="25" pageNumber="26">Maricaban Island</collectingCounty>
,
<collectingRegion id="0B7689F2FFFBFFA4FB98FA6FFB386F87" box="[1075,1198,1474,1495]" country="Philippines" name="Batangas" pageId="25" pageNumber="26">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFFBFFA4FB15FA6CFAC56F87" box="[1214,1363,1473,1495]" name="Philippines" pageId="25" pageNumber="26">Philippines</collectingCountry>
,
<elevation id="429FA023FFFBFFA4FAC8FA6CFA356F87" box="[1379,1443,1473,1495]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">
<quantity id="0E4AEAF5FFFBFFA4FAC8FA6CFA356F87" box="[1379,1443,1473,1495]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="26" unit="m" value="35.0">35 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFFBFFA4FC39FA4DFBA16FA5" box="[914,1079,1504,1525]" pageId="25" pageNumber="26" value="2009-05-17">
<date id="BD0C61D0FFFBFFA4FC39FA4DFBA16FA5" box="[914,1079,1504,1525]" pageId="25" pageNumber="26" value="2009-05-17">17 May 2009</date>
</collectingDate>
,
<collectorName id="644722C6FFFBFFA4FBEEFA4DFB4D6FA6" box="[1093,1243,1504,1526]" pageId="25" pageNumber="26">P. Paleracio</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFFBFFA4FB43FA4DFA096C02" ID-GBIF-Occurrence="2235579062" collectingDate="2011-05-16" collectionCode="CASIZ" collectorName="P. Paleracio" country="Philippines" elevation="40" latitude="13.62772" location="Malajibomanoc" longLatPrecision="1" longitude="120.96592" municipality="Tingloy" pageId="25" pageNumber="26" specimenCode="CASIZ 186103" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFBFFA4FB43FA4DFA086FA6" box="[1256,1438,1504,1526]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 186103</specimenCode>
, two specimens,
<location id="CC6D11CBFFFBFFA4FC54FA53FB2C6C44" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFFBFFA4FC54FA53FB2C6C44" box="[1023,1210,1534,1556]" country="Philippines" latitude="13.62772" longLatPrecision="1" longitude="120.96592" municipality="Tingloy" name="Malajibomanoc" pageId="25" pageNumber="26" stateProvince="Batangas">Malajibomanoc</location>
(Chicken Feather) Island,
<geoCoordinate id="AC8621D7FFFBFFA4FC38F9B0FB8B6C63" box="[915,1053,1565,1587]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="13.62772">13.62772°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFBFFA4FB82F9B0FB296C63" box="[1065,1215,1565,1587]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="120.96592">120.96592°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFFBFFA4FB61F9B0FAB46C63" box="[1226,1314,1565,1587]" pageId="25" pageNumber="26">Tingloy</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFFBFFA4FA85F9B3FA086C63" box="[1326,1438,1566,1587]" country="Philippines" name="Batangas" pageId="25" pageNumber="26">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFFBFFA4FC90F991FC536C02" box="[827,965,1596,1618]" name="Philippines" pageId="25" pageNumber="26">Philippines</collectingCountry>
,
<elevation id="429FA023FFFBFFA4FC7AF991FB9B6C01" box="[977,1037,1596,1617]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="m" value="40.0">
<quantity id="0E4AEAF5FFFBFFA4FC7AF991FB9B6C01" box="[977,1037,1596,1617]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="m" value="40.0">40 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFFBFFA4FBC2F991FA936C01" box="[1129,1285,1596,1618]" pageId="25" pageNumber="26" value="2011-05-16">
<date id="BD0C61D0FFFBFFA4FBC2F991FA936C01" box="[1129,1285,1596,1618]" pageId="25" pageNumber="26" value="2011-05-16">16 May 2011</date>
</collectingDate>
,
<collectorName id="644722C6FFFBFFA4FAB9F991FA096C02" box="[1298,1439,1596,1618]" pageId="25" pageNumber="26">P. Paleracio</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFFBFFA4FF08F927FC326D3D" blockId="25.[163,1444,1673,1901]" pageId="25" pageNumber="26">
<materialsCitation id="79DA4D4DFFFBFFA4FF08F927FB106CCF" ID-GBIF-Occurrence="2235579073" box="[163,1158,1673,1696]" collectionCode="CASIZ" pageId="25" pageNumber="26" specimenCode="CASIZ 208587" specimenCount="1">
<specimenCode id="9914EF6BFFFBFFA4FF08F927FED66CF0" box="[163,320,1674,1696]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 208587</specimenCode>
, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FE1EF927FD916CCF" box="[437,519,1674,1696]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
. E,
<taxonomicName id="0EB23C93FFFBFFA4FD85F924FBBB6CF0" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[558,1069,1673,1696]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="melanesica" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FD85F924FCC56CCF" bold="true" box="[558,851,1673,1695]" italics="true" pageId="25" pageNumber="26">Hypselodoris melanesica</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FB98F927FB106CCF" box="[1075,1158,1674,1695]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FB98F927FB106CCF" bold="true" box="[1075,1158,1674,1695]" pageId="25" pageNumber="26">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="FBC69B02FFFBFFA4FB2CF927FB1A6CCF" bold="true" box="[1159,1164,1674,1695]" pageId="25" pageNumber="26">.</emphasis>
,
<collectionCode id="AFA3DFD5FFFBFFA4FB33F927FB496CCF" box="[1176,1247,1674,1695]" pageId="25" pageNumber="26">CASIZ</collectionCode>
0 69787, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FF08F90AFF616CED" box="[163,247,1703,1725]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="mm" value="0.7">0.7 mm</quantity>
. F,
<taxonomicName id="0EB23C93FFFBFFA4FEB7F90AFD796CED" authority="(Collingwood, 1881)" baseAuthorityName="Collingwood" baseAuthorityYear="1881" box="[284,751,1703,1725]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFFBFFA4FEB7F90AFD986CEC" box="[284,526,1703,1724]" italics="true" pageId="25" pageNumber="26">Hypselodoris bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFFBFFA4FDB6F90AFD716CED" author="Collingwood C." box="[541,743,1703,1725]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="25" pageNumber="26" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
)
</taxonomicName>
,
<materialsCitation id="79DA4D4DFFFBFFA4FD51F90AFDD16C8A" ID-GBIF-Occurrence="2235579116" collectionCode="CASIZ" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<collectionCode id="AFA3DFD5FFFBFFA4FD51F90AFCD46CEC" box="[762,834,1703,1724]" pageId="25" pageNumber="26">CASIZ</collectionCode>
0 85905, scale bar: 1.0 mm. G,
<taxonomicName id="0EB23C93FFFBFFA4FB39F90AFEEC6C8A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FB39F90AFA356CED" bold="true" box="[1170,1443,1703,1725]" italics="true" pageId="25" pageNumber="26">Hypselodoris paradisa</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FE29F968FE4A6C8A" box="[386,476,1733,1754]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FE29F968FE4A6C8A" bold="true" box="[386,476,1733,1754]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFFBFFA4FE42F969FDD16C8A" box="[489,583,1732,1754]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="79DA4D4DFFFBFFA4FDF9F968FEF16CA8" ID-GBIF-Occurrence="2235579034" collectionCode="CASIZ" pageId="25" pageNumber="26" specimenCode="CASIZ 191464" specimenCount="1" typeStatus="holotype">
<specimenCode id="9914EF6BFFFBFFA4FDF9F968FD676C8A" box="[594,753,1732,1754]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 191464</specimenCode>
, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FCC1F969FC286C8A" box="[874,958,1732,1754]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
. H,
<taxonomicName id="0EB23C93FFFBFFA4FC42F969FA356C8A" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1001,1443,1732,1754]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="perii" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FC42F969FB506C89" bold="true" box="[1001,1222,1732,1754]" italics="true" pageId="25" pageNumber="26">Hypselodoris perii</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FF08F94FFF6B6CA7" box="[163,253,1762,1783]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FF08F94FFF6B6CA7" bold="true" box="[163,253,1762,1783]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFFBFFA4FEA1F94FFEF16CA8" box="[266,359,1762,1784]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="79DA4D4DFFFBFFA4FED9F94FFAF66CA8" ID-GBIF-Occurrence="2235579113" box="[370,1376,1761,1784]" collectionCode="NMP" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<collectionCode id="AFA3DFD5FFFBFFA4FED9F94FFE3C6CA7" box="[370,426,1762,1783]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="25" pageNumber="26">NMP</collectionCode>
0 41281, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FDD0F94FFD596CA7" box="[635,719,1762,1783]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="26" unit="mm" value="0.3">0.3 mm</quantity>
. I,
<taxonomicName id="0EB23C93FFFBFFA4FD44F94CFB036CA8" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[751,1173,1761,1784]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="roo" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FD44F94CFC2E6CA7" bold="true" box="[751,952,1761,1783]" italics="true" pageId="25" pageNumber="26">Hypselodoris roo</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FB37F94FFB606CA7" box="[1180,1270,1762,1783]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FB37F94FFB606CA7" bold="true" box="[1180,1270,1762,1783]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFFBFFA4FAA8F94FFAF66CA8" box="[1283,1376,1762,1784]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="79DA4D4DFFFBFFA4FAC7F94FFB076D45" ID-GBIF-Occurrence="2235579068" collectionCode="NMP" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<collectionCode id="AFA3DFD5FFFBFFA4FAC7F94FFA326CA7" box="[1388,1444,1762,1783]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="25" pageNumber="26">NMP</collectionCode>
0 41282, scale bar: 1.0 mm. J,
<taxonomicName id="0EB23C93FFFBFFA4FE41F952FC546D45" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[490,962,1791,1813]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="rositoi" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FE41F952FD486D44" bold="true" box="[490,734,1791,1813]" italics="true" pageId="25" pageNumber="26">Hypselodoris rositoi</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FC62F952FBB36D44" box="[969,1061,1791,1812]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FC62F952FBB36D44" bold="true" box="[969,1061,1791,1812]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="1609F9B2FFFBFFA4FB99F952FB076D45" box="[1074,1169,1791,1813]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<collectionCode id="AFA3DFD5FFFBFFA4FB36F8ADFB406D45" box="[1181,1238,1792,1813]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="25" pageNumber="26">NMP</collectionCode>
0 41283, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FF08F8B0FF636D62" box="[163,245,1821,1842]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="mm" value="1.2">1.2 mm</quantity>
. K,
<taxonomicName id="0EB23C93FFFBFFA4FEB7F8B1FD496D62" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[284,735,1820,1842]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFFBFFA4FEB7F8B1FD9E6D62" bold="true" box="[284,520,1820,1842]" italics="true" pageId="25" pageNumber="26">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFBFFA4FD4DF8B0FCA86D62" box="[742,830,1821,1842]" pageId="25" pageNumber="26" rank="species">
<emphasis id="FBC69B02FFFBFFA4FD4DF8B0FCA86D62" bold="true" box="[742,830,1821,1842]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<materialsCitation id="79DA4D4DFFFBFFA4FCE1F8B0FC366D3D" ID-GBIF-Occurrence="2235579077" collectionCode="CASIZ" pageId="25" pageNumber="26" specimenCode="CASIZ 177305" specimenCount="1" typeStatus="holotype">
<specimenCode id="9914EF6BFFFBFFA4FCE1F8B0FC726D62" box="[842,996,1820,1842]" collectionCode="CASIZ" pageId="25" pageNumber="26">CASIZ 177305</specimenCode>
, scale bar:
<quantity id="0E4AEAF5FFFBFFA4FBF2F8B1FB3C6D62" box="[1113,1194,1820,1842]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
. am, ampulla; bc, bursa copulatrix; ej, ejaculatory portion of the vas deferens; fgm, female gland mass; p, penis; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina; vg, vestibular gland
</materialsCitation>
.
</paragraph>
<caption id="9DCD1798FFF8FFA7FF3AFC23FAB6698F" httpUri="https://zenodo.org/record/2632799/files/figure.png" pageId="26" pageNumber="27" startId="26.[145,223,910,932]" targetBox="[209,1361,195,880]" targetPageId="26">
<paragraph id="C90D4710FFF8FFA7FF3AFC23FAB6698F" blockId="26.[145,1425,910,991]" pageId="26" pageNumber="27">
<emphasis id="FBC69B02FFF8FFA7FF3AFC23FE9369F4" bold="true" box="[145,261,910,932]" pageId="26" pageNumber="27">Figure 14.</emphasis>
<taxonomicName id="0EB23C93FFF8FFA7FEBBFC23FD3069F5" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[272,678,910,933]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFF8FFA7FEBBFC23FE4469F4" bold="true" box="[272,466,910,932]" italics="true" pageId="26" pageNumber="27">Hypselodoris iba</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFF8FFA7FD00FC22FC9769F4" box="[683,769,911,932]" pageId="26" pageNumber="27" rank="species">
<emphasis id="FBC69B02FFF8FFA7FD00FC22FC9769F4" bold="true" box="[683,769,911,932]" pageId="26" pageNumber="27">sp. nov.</emphasis>
</taxonomicNameLabel>
, living animals. A, paratype, CASIZ 180419, Tingloy, Batangas, Philippines. B, holotype, NMP 0 41279, Tingloy, Batangas, Philippines. C, two specimens of different colour morphs mating, southwestern Sangeang, Indonesia, David Cowdery. D, paratype, CASIZ 177777, Tingloy, Batangas, Philippines.
</paragraph>
</caption>
<paragraph id="C90D4710FFF8FFA7FF3AFBA9FE366E25" blockId="26.[145,761,1027,1141]" pageId="26" pageNumber="27">
<materialsCitation id="79DA4D4DFFF8FFA7FF3AFBA9FE0A6E25" ID-GBIF-Occurrence="2235579033" collectingDate="2016-04-16" collectionCode="CASIZ" collectorName="P. Paleracio" country="Philippines" county="Luzon" elevation="30" latitude="13.694934" location="Dead Palm" longLatPrecision="1" longitude="120.88469" municipality="Mabini" pageId="26" pageNumber="27" specimenCode="CASIZ 217217" specimenCount="1" stateProvince="Batangas">
<specimenCode id="9914EF6BFFF8FFA7FF3AFBA9FED96E49" box="[145,335,1028,1049]" collectionCode="CASIZ" pageId="26" pageNumber="27">CASIZ 217217</specimenCode>
, MAB 325,
<location id="CC6D11CBFFF8FFA7FE41FBAEFDE16E49" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFF8FFA7FE41FBAEFDE16E49" box="[490,631,1027,1049]" country="Philippines" county="Luzon" latitude="13.694934" longLatPrecision="1" longitude="120.88469" municipality="Mabini" name="Dead Palm" pageId="26" pageNumber="27" stateProvince="Batangas">Dead Palm</location>
dive site,
<geoCoordinate id="AC8621D7FFF8FFA7FF3AFB8FFED56E68" box="[145,323,1058,1080]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="13.694934">13.69493372°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFF8FFA7FEE4FB8FFE646E68" box="[335,498,1058,1080]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="120.88469">120.884687°E</geoCoordinate>
,
<location id="CC6D11CBFFF8FFA7FE55FB8FFD636E68" LSID="urn:lsid:plazi:treatment:411BF606FFFBFFA1FCEAFDE1FB626BA0:CC6D11CBFFF8FFA7FE55FB8FFD636E68" box="[510,757,1058,1080]" country="Philippines" county="Luzon" latitude="13.694934" longLatPrecision="1" longitude="120.88469" municipality="Mabini" name="Calumpan Peninsula" pageId="26" pageNumber="27" stateProvince="Batangas">Calumpan Peninsula</location>
,
<collectingMunicipality id="2969DD6AFFF8FFA7FF3AFBECFF736E07" box="[145,229,1089,1111]" pageId="26" pageNumber="27">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFF8FFA7FF5BFBECFEC86E06" box="[240,350,1089,1110]" country="Philippines" name="Batangas" pageId="26" pageNumber="27">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFF8FFA7FEC1FBECFE256E06" box="[362,435,1089,1110]" pageId="26" pageNumber="27">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFF8FFA7FE15FBECFDD46E07" box="[446,578,1089,1111]" name="Philippines" pageId="26" pageNumber="27">Philippines</collectingCountry>
,
<elevation id="429FA023FFF8FFA7FDE5FBECFD116E06" box="[590,647,1089,1110]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="m" value="30.0">
<quantity id="0E4AEAF5FFF8FFA7FDE5FBECFD116E06" box="[590,647,1089,1110]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="m" value="30.0">30 m</quantity>
</elevation>
depth,
<collectingDate id="AD489838FFF8FFA7FD76FBECFE9D6E25" pageId="26" pageNumber="27" value="2016-04-16">
<date id="BD0C61D0FFF8FFA7FD76FBECFE9D6E25" pageId="26" pageNumber="27" value="2016-04-16">16 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFF8FFA7FEBDFBCDFE0A6E25" box="[278,412,1119,1141]" pageId="26" pageNumber="27">P. Paleracio</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="C90D4710FFF8FFA7FF3AFB30FEBA6EE4" blockId="26.[145,761,1180,1265]" box="[145,300,1180,1204]" pageId="26" pageNumber="27">
<heading id="9245F07CFFF8FFA7FF3AFB30FEBA6EE4" box="[145,300,1180,1204]" fontSize="10" level="4" pageId="26" pageNumber="27" reason="1">
<emphasis id="FBC69B02FFF8FFA7FF3AFB30FEBA6EE4" box="[145,300,1180,1204]" italics="true" pageId="26" pageNumber="27">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF8FFA7FF3AFB10FE8F6EA1" blockId="26.[145,761,1180,1265]" pageId="26" pageNumber="27">
Aphol’s Reef, Tingloy, Maricaban Island,
<collectingRegion id="0B7689F2FFF8FFA7FD29FB10FD636E82" box="[642,757,1213,1234]" country="Philippines" name="Batangas" pageId="26" pageNumber="27">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFF8FFA7FF3AFB76FE806EA1" box="[145,278,1243,1265]" name="Philippines" pageId="26" pageNumber="27">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF8FFA7FF3AFAB5FE8A6F3D" pageId="26" pageNumber="27" type="distribution">
<paragraph id="C90D4710FFF8FFA7FF3AFAB5FE586F60" blockId="26.[145,761,1304,1389]" box="[145,462,1304,1328]" pageId="26" pageNumber="27">
<heading id="9245F07CFFF8FFA7FF3AFAB5FE586F60" box="[145,462,1304,1328]" fontSize="10" level="4" pageId="26" pageNumber="27" reason="1">
<emphasis id="FBC69B02FFF8FFA7FF3AFAB5FE586F60" box="[145,462,1304,1328]" italics="true" pageId="26" pageNumber="27">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF8FFA7FF3AFA94FE8A6F3D" blockId="26.[145,761,1304,1389]" pageId="26" pageNumber="27">
Known from
<collectingCountry id="B1A50780FFF8FFA7FE83FA94FE0C6F1F" box="[296,410,1337,1359]" name="Indonesia" pageId="26" pageNumber="27">Indonesia</collectingCountry>
and the
<collectingCountry id="B1A50780FFF8FFA7FDA9FA94FD106F1F" box="[514,646,1337,1359]" name="Philippines" pageId="26" pageNumber="27">Philippines</collectingCountry>
(
<bibRefCitation id="AD233AE1FFF8FFA7FD3DFA94FE9A6F3D" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="26" pageNumber="27" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFF8FFA7FF3AFAF5FF5F6F3C" box="[145,201,1367,1389]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
, 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF8FFA7FF3AFA3BFDE76F9B" pageId="26" pageNumber="27" type="etymology">
<paragraph id="C90D4710FFF8FFA7FF3AFA3BFDE76F9B" blockId="26.[145,761,1430,1483]" pageId="26" pageNumber="27">
<emphasis id="FBC69B02FFF8FFA7FF3AFA3BFE4E6FFB" box="[145,472,1430,1451]" italics="true" pageId="26" pageNumber="27">
Etymology:
<taxonomicName id="0EB23C93FFF8FFA7FEB1FA3BFE4E6FFB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[282,472,1430,1451]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="iba">Hypselodoris iba</taxonomicName>
</emphasis>
is named for the Tagalog word
<taxonomicName id="0EB23C93FFF8FFA7FF7BFA18FF636F9B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[208,245,1461,1483]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="iba">iba</taxonomicName>
, meaning different or distinctive.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFF8FFA1FF3AFA5FFD6C68F8" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="C90D4710FFF8FFA7FF3AFA5FFE8B6C59" blockId="26.[145,761,1522,1883]" box="[145,285,1522,1545]" pageId="26" pageNumber="27">
<heading id="9245F07CFFF8FFA7FF3AFA5FFE8B6C59" box="[145,285,1522,1545]" fontSize="10" level="4" pageId="26" pageNumber="27" reason="1">
<emphasis id="FBC69B02FFF8FFA7FF3AFA5FFE8B6C59" box="[145,285,1522,1545]" italics="true" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFF8FFA7FF3AF9BFFCE06F1C" blockId="26.[145,761,1522,1883]" lastBlockId="26.[809,1425,1027,1356]" pageId="26" pageNumber="27">
<emphasis id="FBC69B02FFF8FFA7FF3AF9BFFE136C77" box="[145,389,1554,1575]" italics="true" pageId="26" pageNumber="27">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFF8FFA7FDF3F9BFFD3E6C77" box="[600,680,1554,1576]" captionStart="Figure 14" captionStartId="26.[145,223,910,932]" captionTargetBox="[209,1361,195,880]" captionTargetId="figure@26.[209,1361,195,880]" captionTargetPageId="26" captionText="Figure 14. Hypselodoris iba Gosliner &amp; Johnson sp. nov., living animals. A, paratype, CASIZ 180419, Tingloy, Batangas, Philippines. B, holotype, NMP 041279, Tingloy, Batangas, Philippines. C, two specimens of different colour morphs mating, southwestern Sangeang, Indonesia, David Cowdery. D, paratype, CASIZ 177777, Tingloy, Batangas, Philippines." httpUri="https://zenodo.org/record/2632799/files/figure.png" pageId="26" pageNumber="27">Fig. 14</figureCitation>
) large, reaching
<quantity id="0E4AEAF5FFF8FFA7FF5DF99CFED66C16" box="[246,320,1585,1606]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="27" unit="mm" value="70.0">70 mm</quantity>
in length.Body wide, tapering to narrow, rounded posterior end of notum. Foot wide, elongate posteriorly. Gill with six to eight unipinnate branches. Gill branches with white pigment at base, becoming orange in outer portion. Gill pocket well elevated from notum. Rhinophores with 20–24 lamellae. Colour pattern occurs in two distinct morphs. In first morph, body generally uniformly purple, with uniformly wide opaque white marginal band or occasionally orange band. Rhinophores uniformly orange, with orange or purple pigment on base of rhinophore sheaths. Second morph generally off-white to beige, with thin white marginal band, transitioning to orange anteriorly and posteriorly. Submarginal blood red band may be present, transitioning to purple anteriorly. Blood red blotches present on central region of notum, on base of gill pocket and base of rhinophore sheaths. purple band present near opening of rhinophore sheaths. Blood red blotches may also be present on lateral faces of body. Wide purple band present along lateral margin of foot.
</paragraph>
<paragraph id="C90D4710FFF8FFA7FC82FAD8FB4B6FB6" blockId="26.[809,1425,1397,1511]" pageId="26" pageNumber="27">
<emphasis id="FBC69B02FFF8FFA7FC82FAD8FC416FDA" box="[809,983,1397,1418]" italics="true" pageId="26" pageNumber="27">Mantle glands</emphasis>
: Subcutaneous mantle glands large and concentrated posteriorly, with 12–16 glands. Smaller, irregularly distributed glands present along lateral and anterior margins (
<figureCitation id="51895B95FFF8FFA7FBD5FA7CFB5B6FB6" box="[1150,1229,1489,1511]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="26" pageNumber="27">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFF8FFA6FC82F9A2FEFD6D0A" blockId="26.[809,1425,1551,1880]" lastBlockId="27.[163,779,1676,1882]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="FBC69B02FFF8FFA7FC82F9A2FC656C75" box="[809,1011,1551,1573]" italics="true" pageId="26" pageNumber="27">Buccal armature</emphasis>
: Muscular portion of buccal mass about the same size as the length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFF8FFA7FC9AF927FC076CF0" box="[817,913,1674,1696]" captionStart="Figure 15" captionStartId="27.[163,241,1559,1581]" captionTargetBox="[227,1379,195,1518]" captionTargetId="figure@27.[227,1379,195,1518]" captionTargetPageId="27" captionText="Figure 15. Hypselodoris iba Gosliner &amp; Johnson sp. nov., buccal armature.A, jaw rodlets, CASIZ 177777. B, inner lateral teeth, CASIZ 177777. C, inner lateral teeth, CASIZ 180419. D, middle lateral teeth, CASIZ 177777. E, middle lateral teeth, CASIZ 180419. F, outer lateral teeth, CASIZ 180419." httpUri="https://zenodo.org/record/2632803/files/figure.png" pageId="26" pageNumber="27">Fig. 15A</figureCitation>
). Rodlets narrow, triangular with short base and evenly curved, with single, acutely pointed apex. Radular formula of two
<typeStatus id="1609F9B2FFF8FFA7FB9DF965FB316C8D" box="[1078,1191,1736,1757]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
63 × 98.0.98 (CASIZ 177777) and 66 × 88.0.88 (CASIZ
<date id="BD0C61D0FFF8FFA7FB1AF94BFA9E6CAC" box="[1201,1288,1766,1788]" pageId="26" pageNumber="27" value="19-04-18">180419</date>
). Rachidian row of teeth absent (
<figureCitation id="51895B95FFF8FFA7FB89F8A8FB336D4A" box="[1058,1189,1797,1819]" captionStart="Figure 15" captionStartId="27.[163,241,1559,1581]" captionTargetBox="[227,1379,195,1518]" captionTargetId="figure@27.[227,1379,195,1518]" captionTargetPageId="27" captionText="Figure 15. Hypselodoris iba Gosliner &amp; Johnson sp. nov., buccal armature.A, jaw rodlets, CASIZ 177777. B, inner lateral teeth, CASIZ 177777. C, inner lateral teeth, CASIZ 180419. D, middle lateral teeth, CASIZ 177777. E, middle lateral teeth, CASIZ 180419. F, outer lateral teeth, CASIZ 180419." httpUri="https://zenodo.org/record/2632803/files/figure.png" pageId="26" pageNumber="27">Fig. 15B, C</figureCitation>
). Innermost lateral teeth lacking inner denticles or having single acutely pointed denticle on inner side of bifid primary cusp. Outer surface of inner lateral without outer lateral denticles. Next several laterals (
<figureCitation id="51895B95FFF9FFA6FDA1F906FD136C90" box="[522,645,1707,1729]" captionStart="Figure 15" captionStartId="27.[163,241,1559,1581]" captionTargetBox="[227,1379,195,1518]" captionTargetId="figure@27.[227,1379,195,1518]" captionTargetPageId="27" captionText="Figure 15. Hypselodoris iba Gosliner &amp; Johnson sp. nov., buccal armature.A, jaw rodlets, CASIZ 177777. B, inner lateral teeth, CASIZ 177777. C, inner lateral teeth, CASIZ 180419. D, middle lateral teeth, CASIZ 177777. E, middle lateral teeth, CASIZ 180419. F, outer lateral teeth, CASIZ 180419." httpUri="https://zenodo.org/record/2632803/files/figure.png" pageId="27" pageNumber="28">Fig. 15D, E</figureCitation>
) and all but outermost lateral teeth with only a bifid primary cusp. Outer cusp much shorter than inner cusp. Outermost teeth having a narrower base and shorter tooth shape, with zero to two rounded outer denticles (
<figureCitation id="51895B95FFF9FFA6FD34F88BFC946D6B" box="[671,770,1829,1852]" captionStart="Figure 15" captionStartId="27.[163,241,1559,1581]" captionTargetBox="[227,1379,195,1518]" captionTargetId="figure@27.[227,1379,195,1518]" captionTargetPageId="27" captionText="Figure 15. Hypselodoris iba Gosliner &amp; Johnson sp. nov., buccal armature.A, jaw rodlets, CASIZ 177777. B, inner lateral teeth, CASIZ 177777. C, inner lateral teeth, CASIZ 180419. D, middle lateral teeth, CASIZ 177777. E, middle lateral teeth, CASIZ 180419. F, outer lateral teeth, CASIZ 180419." httpUri="https://zenodo.org/record/2632803/files/figure.png" pageId="27" pageNumber="28">Fig. 15F</figureCitation>
) below bifid cusps.
</paragraph>
<caption id="9DCD1798FFF9FFA6FF08F9BAFD5E6C37" httpUri="https://zenodo.org/record/2632803/files/figure.png" pageId="27" pageNumber="28" startId="27.[163,241,1559,1581]" targetBox="[227,1379,195,1518]" targetPageId="27">
<paragraph id="C90D4710FFF9FFA6FF08F9BAFD5E6C37" blockId="27.[163,1443,1559,1640]" pageId="27" pageNumber="28">
<emphasis id="FBC69B02FFF9FFA6FF08F9BAFE816C7C" bold="true" box="[163,279,1559,1581]" pageId="27" pageNumber="28">Figure 15.</emphasis>
<taxonomicName id="0EB23C93FFF9FFA6FE89F9BAFD236C7D" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[290,693,1559,1581]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="iba" status="sp. nov.">
<emphasis id="FBC69B02FFF9FFA6FE89F9BAFE756C7D" bold="true" box="[290,483,1559,1581]" italics="true" pageId="27" pageNumber="28">Hypselodoris iba</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFF9FFA6FD12F9B5FC996C7D" box="[697,783,1560,1581]" pageId="27" pageNumber="28" rank="species">
<emphasis id="FBC69B02FFF9FFA6FD12F9B5FC996C7D" bold="true" box="[697,783,1560,1581]" pageId="27" pageNumber="28">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature. A, jaw rodlets, CASIZ 177777. B, inner lateral teeth, CASIZ 177777. C, inner lateral teeth, CASIZ 180419. D, middle lateral teeth, CASIZ 177777. E, middle lateral teeth, CASIZ 180419. F, outer lateral teeth, CASIZ 180419.
</paragraph>
</caption>
<paragraph id="C90D4710FFF9FFA1FC90F921FD6C68F8" blockId="27.[827,1443,1676,1882]" lastBlockId="28.[145,762,198,680]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="FBC69B02FFF9FFA6FC90F921FB896CF2" box="[827,1055,1676,1698]" italics="true" pageId="27" pageNumber="28">Reproductive system</emphasis>
: Reproductive organs (
<figureCitation id="51895B95FFF9FFA6FA8DF921FA146CF1" box="[1318,1410,1676,1698]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="27" pageNumber="28">Fig.13B</figureCitation>
) of two
<typeStatus id="1609F9B2FFF9FFA6FCC1F906FC4D6C90" box="[874,987,1707,1728]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
fully mature (CASIZ 177777 and CASIZ
<date id="BD0C61D0FFF9FFA6FC90F967FC056CB0" box="[827,915,1738,1760]" pageId="27" pageNumber="28" value="19-04-18">180419</date>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens curved and thick and narrowing slightly as it transitions into muscular, ejaculatory portion. Prostatic portion enveloping bursa copulatrix. Ejaculatory portion moderately to highly convoluted, narrow, entering short, wider penial bulb. Penial bulb adjacent to slightly curved, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long, wide vagina leading to small, curved receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum seminis appressed against vagina immediately below bursa copulatrix. Moderately long uterine duct emerging from vagina close to base of bursa and female gland mass, near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA1FF3AFD78FB626BA0" pageId="28" pageNumber="29" type="discussion">
<paragraph id="C90D4710FFFEFFA1FF3AFD78FF6868BD" blockId="28.[145,761,725,1914]" box="[145,254,725,749]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FF3AFD78FF6868BD" box="[145,254,725,749]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FF3AFD78FF6868BD" box="[145,254,725,749]" italics="true" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FF3AFD58FD506FFE" blockId="28.[145,761,725,1914]" pageId="28" pageNumber="29">
This species has been considered as a synonym of
<taxonomicName id="0EB23C93FFFEFFA1FF3AFCB9FD1F6979" authority="(Humann &amp; DeLoach, 2010)" baseAuthorityName="Humann &amp; DeLoach" baseAuthorityYear="2010" box="[145,649,788,810]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFFEFFA1FF3AFCB9FEB46979" box="[145,290,788,809]" italics="true" pageId="28" pageNumber="29">H. bullockii</emphasis>
(
<bibRefCitation id="AD233AE1FFFEFFA1FE9EFCB9FD166979" author="Humann P &amp; DeLoach N." box="[309,640,788,810]" journalOrPublisher="Jacksonville: New World Publications" pageId="28" pageNumber="29" refId="ref42986" refString="Humann P, DeLoach N. 2010. Reef creature identification, tropical Pacific. Jacksonville: New World Publications." title="Reef creature identification, tropical Pacific" type="book" year="2010">Humann &amp; DeLoach, 2010</bibRefCitation>
)
</taxonomicName>
or as an undescribed species (
<bibRefCitation id="AD233AE1FFFEFFA1FED5FC9FFD1A6918" author="Debelius H &amp; Kuiter R." box="[382,652,818,840]" journalOrPublisher="Frankfurt: IKAN-Unterwasserarchiv" pageId="28" pageNumber="29" refId="ref41869" refString="Debelius H, Kuiter R. 2007. Nudibranchs of the world. Frankfurt: IKAN-Unterwasserarchiv." title="Nudibranchs of the world" type="book" year="2007">Debelius &amp; Kuiter, 2007</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFFEFFA1FD32FC9FFE866936" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="28" pageNumber="29" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFFEFFA1FF3AFCFFFF556936" box="[145,195,849,871]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
., 2008
</bibRefCitation>
,
<bibRefCitation id="AD233AE1FFFEFFA1FEB5FCFCFEC06937" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[286,342,849,871]" journalOrPublisher="Jacksonville: New World Press" pageId="28" pageNumber="29" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">2015</bibRefCitation>
). This has been complicated further by the fact that
<taxonomicName id="0EB23C93FFFEFFA1FEE3FCDDFE1B69D5" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[328,397,880,901]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FEE3FCDDFE1B69D5" box="[328,397,880,901]" italics="true" pageId="28" pageNumber="29">H. iba</emphasis>
</taxonomicName>
has two distinct colour morphs, and one of them is also similar to
<taxonomicName id="0EB23C93FFFEFFA1FD85FC22FD6F69F3" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[558,761,910,932]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFFEFFA1FD85FC22FD6F69F3" box="[558,761,910,932]" italics="true" pageId="28" pageNumber="29">H. variobranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFEFFA1FF3AFC03FF726993" box="[145,228,942,963]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
described here. For a detailed comparison, see the remarks on
<taxonomicName id="0EB23C93FFFEFFA1FEF8FC61FDB669B1" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[339,544,972,993]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="variobranchia" status="sp. nov.">
<emphasis id="FBC69B02FFFEFFA1FEF8FC61FDB669B1" box="[339,544,972,993]" italics="true" pageId="28" pageNumber="29">H. variobranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFEFFA1FD82FC61FD1469B1" box="[553,642,972,993]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
The most reliable way to distinguish
<taxonomicName id="0EB23C93FFFEFFA1FE68FC46FD9E69AF" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[451,520,1002,1024]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FE68FC46FD9E69AF" box="[451,520,1002,1024]" italics="true" pageId="28" pageNumber="29">H. iba</emphasis>
</taxonomicName>
from members of the
<taxonomicName id="0EB23C93FFFEFFA1FF3AFBA7FE876E4E" baseAuthorityName="Humann &amp; DeLoach" baseAuthorityYear="2010" box="[145,273,1033,1055]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFFEFFA1FF3AFBA7FE876E4E" box="[145,273,1033,1055]" italics="true" pageId="28" pageNumber="29">H. bullockii</emphasis>
</taxonomicName>
clade is by the presence of mantle glands in
<taxonomicName id="0EB23C93FFFEFFA1FF3AFB85FF446E6D" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,210,1064,1085]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FF3AFB85FF446E6D" box="[145,210,1064,1085]" italics="true" pageId="28" pageNumber="29">H. iba</emphasis>
</taxonomicName>
. No members of the
<taxonomicName id="0EB23C93FFFEFFA1FE04FB85FDBB6E6D" baseAuthorityName="Humann &amp; DeLoach" baseAuthorityYear="2010" box="[431,557,1064,1085]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFFEFFA1FE04FB85FDBB6E6D" box="[431,557,1064,1085]" italics="true" pageId="28" pageNumber="29">H. bullockii</emphasis>
</taxonomicName>
clade have mantle glands (present study). The two colour morphs have been observed copulating with each other (
<figureCitation id="51895B95FFFEFFA1FD3BFBC8FD656E2A" box="[656,755,1125,1147]" captionStart="Figure 14" captionStartId="26.[145,223,910,932]" captionTargetBox="[209,1361,195,880]" captionTargetId="figure@26.[209,1361,195,880]" captionTargetPageId="26" captionText="Figure 14. Hypselodoris iba Gosliner &amp; Johnson sp. nov., living animals. A, paratype, CASIZ 180419, Tingloy, Batangas, Philippines. B, holotype, NMP 041279, Tingloy, Batangas, Philippines. C, two specimens of different colour morphs mating, southwestern Sangeang, Indonesia, David Cowdery. D, paratype, CASIZ 177777, Tingloy, Batangas, Philippines." httpUri="https://zenodo.org/record/2632799/files/figure.png" pageId="28" pageNumber="29">Fig. 14C</figureCitation>
;
<bibRefCitation id="AD233AE1FFFEFFA1FF3AFB29FEBD6EC9" author="Kohler E." box="[145,299,1156,1178]" journalOrPublisher="Sydney: Australian Museum" pageId="28" pageNumber="29" publicationUrl="http://www.seaslugforum.net/find/6816" refId="ref43356" refString="Kohler E. 2002. H. bullocki ' colour forms' mating. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 6816" title="H. bullocki ' colour forms' mating" type="url" volumeTitle="Sea slug forum" year="2002">Köhler, 2002</bibRefCitation>
). Not only are the two colour morphs observed copulating, but they show very little genetic divergence. The
<typeStatus id="1609F9B2FFFEFFA1FEE8FB6CFE346E87" box="[323,418,1217,1239]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(CASIZ 177511), a light morph specimen, is only 0.3% different in its
<emphasis id="FBC69B02FFFEFFA1FD95FB4DFDFB6EA5" box="[574,621,1248,1269]" italics="true" pageId="28" pageNumber="29">COI</emphasis>
gene from a purple morph specimen from the same locality (CASIZ
<date id="BD0C61D0FFFEFFA1FF3AFAB0FF716F63" box="[145,231,1309,1331]" pageId="28" pageNumber="29" value="19-04-18">180419</date>
), and a third specimen (CASIZ 177777, also a light morph) from the same locality is 0.6% different from the other two. The subsequent ABGD analysis confirms that these three specimens are conspecific with each other and distinct from all other species.
</paragraph>
<paragraph id="C90D4710FFFEFFA1FF02FA1BFE206CAF" blockId="28.[145,761,725,1914]" pageId="28" pageNumber="29">
<taxonomicName id="0EB23C93FFFEFFA1FF02FA1BFEE56F9B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[169,371,1462,1483]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FF02FA1BFEE56F9B" box="[169,371,1462,1483]" italics="true" pageId="28" pageNumber="29">Hypselodoris iba</emphasis>
</taxonomicName>
is also similar in appearance to
<emphasis id="FBC69B02FFFEFFA1FF3AFA78FEF26FBB" box="[145,356,1493,1515]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="0EB23C93FFFEFFA1FF3AFA78FE926FBA" authorityName="Bergh" authorityYear="1878" box="[145,260,1493,1514]" class="Gastropoda" family="Chromodorididae" genus="Thorunna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">Thorunna</taxonomicName>
punicea
</emphasis>
(
<bibRefCitation id="AD233AE1FFFEFFA1FED8FA78FD8B6FBB" author="Rudman WB" box="[371,541,1493,1515]" journalOrPublisher="Molluscan Research" pageId="28" pageNumber="29" pagination="1 - 43" part="16" refId="ref44878" refString="Rudman WB. 1995. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: further species from New Caledonia and the Noumea romeri colour group. Molluscan Research 16: 1 - 43." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: further species from New Caledonia and the Noumea romeri colour group" type="journal article" year="1995">Rudman, 1995</bibRefCitation>
). In
<emphasis id="FBC69B02FFFEFFA1FDFAFA7BFD506FBB" box="[593,710,1494,1515]" italics="true" pageId="28" pageNumber="29">T. punicea</emphasis>
, the gills vibrate when the animal is active, the reproductive system has a large receptaculum seminis and the foot has an opaque white line along its margin. In
<taxonomicName id="0EB23C93FFFEFFA1FD00F99FFD656C16" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[683,755,1585,1607]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FD00F99FFD656C16" box="[683,755,1585,1607]" italics="true" pageId="28" pageNumber="29">H. iba</emphasis>
</taxonomicName>
, the gills do not vibrate and the foot is lined with purple, and it has a small receptaculum seminis and denticulate outer lateral teeth rather than simply bifid ones. In our phylogenetic analysis,
<emphasis id="FBC69B02FFFEFFA1FD80F901FD356C91" box="[555,675,1708,1729]" italics="true" pageId="28" pageNumber="29">T. punicea</emphasis>
clearly nests with other species of
<taxonomicName id="0EB23C93FFFEFFA1FE69F967FDA26C8F" authorityName="Bergh" authorityYear="1878" box="[450,564,1738,1759]" class="Gastropoda" family="Chromodorididae" genus="Thorunna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFEFFA1FE69F967FDA26C8F" box="[450,564,1738,1759]" italics="true" pageId="28" pageNumber="29">Thorunna</emphasis>
</taxonomicName>
rather than with members of
<taxonomicName id="0EB23C93FFFEFFA1FEB7F944FE396CAE" authorityName="Stimpson" authorityYear="1855" box="[284,431,1769,1790]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFEFFA1FEB7F944FE396CAE" box="[284,431,1769,1790]" italics="true" pageId="28" pageNumber="29">Hypselodoris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFFEFFA1FF02F8A5FB626BA0" blockId="28.[145,761,725,1914]" lastBlockId="28.[809,1425,198,496]" pageId="28" pageNumber="29">
<taxonomicName id="0EB23C93FFFEFFA1FF02F8A5FEE56D4D" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[169,371,1800,1821]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FF02F8A5FEE56D4D" box="[169,371,1800,1821]" italics="true" pageId="28" pageNumber="29">Hypselodoris iba</emphasis>
</taxonomicName>
is a member of large clade that includes
<taxonomicName id="0EB23C93FFFEFFA1FF53F88AFEC66D6B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[248,336,1830,1852]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="reidi">
<emphasis id="FBC69B02FFFEFFA1FF53F88AFEC66D6B" box="[248,336,1830,1852]" italics="true" pageId="28" pageNumber="29">H. reidi</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFFEFFA1FEF5F88AFE486D6B" authorityName="Wilson &amp; Willan" authorityYear="2007" box="[350,478,1830,1852]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="jacksoni">
<emphasis id="FBC69B02FFFEFFA1FEF5F88AFE486D6B" box="[350,478,1830,1852]" italics="true" pageId="28" pageNumber="29">H. jacksoni</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFFEFFA1FE40F88AFDCA6D6C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[491,604,1831,1852]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="cerisae">
<emphasis id="FBC69B02FFFEFFA1FE40F88AFDCA6D6C" box="[491,604,1831,1852]" italics="true" pageId="28" pageNumber="29">H. cerisae</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFFEFFA1FDC2F88AFD656D6B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[617,755,1830,1852]" class="Gastropoda" family="Chromodorididae" genus="Hypseldoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="krakatoa">
<emphasis id="FBC69B02FFFEFFA1FDC2F88AFD656D6B" box="[617,755,1830,1852]" italics="true" pageId="28" pageNumber="29">H. krakatoa</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFFEFFA1FF3AF8EBFE976D0B" authorityName="Marcus &amp; Marcus" authorityYear="1970" box="[145,257,1862,1883]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="regina">
<emphasis id="FBC69B02FFFEFFA1FF3AF8EBFE976D0B" box="[145,257,1862,1883]" italics="true" pageId="28" pageNumber="29">H. regina</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFFEFFA1FE94F8EBFE236D0A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[319,437,1861,1883]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFFEFFA1FE94F8EBFE236D0A" box="[319,437,1861,1883]" italics="true" pageId="28" pageNumber="29">H. lacuna</emphasis>
</taxonomicName>
. None of these species has a colour pattern similar to either colour morph of
<taxonomicName id="0EB23C93FFFEFFA1FC82FF6AFCE76A8B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,881,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="iba">
<emphasis id="FBC69B02FFFEFFA1FC82FF6AFCE76A8B" box="[809,881,198,220]" italics="true" pageId="28" pageNumber="29">H. iba</emphasis>
</taxonomicName>
. All members of this clade, with the exception of
<taxonomicName id="0EB23C93FFFEFFA1FCEEFF4BFC206AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[837,950,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="lacuna">
<emphasis id="FBC69B02FFFEFFA1FCEEFF4BFC206AAA" box="[837,950,229,251]" italics="true" pageId="28" pageNumber="29">H. lacuna</emphasis>
</taxonomicName>
, have an elevated gill pocket and mantle glands arranged all around the perimeter of the mantle margin (
<bibRefCitation id="AD233AE1FFFEFFA1FC25FE8FFB2A6B68" author="Gosliner T &amp; Johnson, R." box="[910,1212,290,312]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="28" pageNumber="29" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFFEFFA1FB61FE8FFCF66B06" author="Wilson N &amp; Willan, R." journalOrPublisher="Zootaxa" pageId="28" pageNumber="29" pagination="29 - 42" part="1549" refId="ref45538" refString="Wilson N, Willan, R. 2007. Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982. Zootaxa 1549: 29 - 42." title="Hypselodoris jacksoni, a new species from the south-western Pacific Ocean (Nudibranchia: Chromodorididae), with a discussion on intraspecific variation in mantle glands in Chromodoris willani Rudman, 1982" type="journal article" year="2007">Wilson &amp; Willan, 2007</bibRefCitation>
; present study). Members of this clade also have an inner lateral tooth with a single inner denticle and no denticles on the outer side of the bifid cusps. All members of this clade have the majority of teeth with no inner or outer denticles and the outer cusp of the tooth much smaller than the inner cusp.
</paragraph>
</subSubSection>
</treatment>
</document>

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</mods:mods>
<treatment id="411BF606FFFCFF9DFEB1FD45FAE66B68" ID-DOI="http://doi.org/10.5281/zenodo.5980682" ID-GBIF-Taxon="156146722" ID-Zenodo-Dep="5980682" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68" httpUri="http://treatment.plazi.org/id/411BF606FFFCFF9DFEB1FD45FAE66B68" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="81A8149BFFFCFFA3FEB1FD45FDED694E" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="C90D4710FFFCFFA3FEB1FD45FDED694E" blockId="30.[270,635,744,837]" pageId="30" pageNumber="31">
<taxonomicName id="0EB23C93FFFCFFA3FEB1FD45FD87694C" ID-CoL="3P345" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="32" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<heading id="9245F07CFFFCFFA3FEB1FD45FDE668AE" allCaps="true" bold="true" box="[282,624,744,767]" centered="true" fontSize="9" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="FBC69B02FFFCFFA3FEB1FD45FDE668AE" bold="true" box="[282,624,744,767]" italics="true" pageId="30" pageNumber="31">HYPSELODORIS KATHERINAE</emphasis>
</heading>
<heading id="9245F07CFFFCFFA3FEA5FCABFD87694C" allCaps="true" box="[270,529,774,798]" centered="true" fontSize="9" level="3" pageId="30" pageNumber="31" reason="4">GOSLINER &amp; JOHNSON</heading>
</taxonomicName>
<emphasis id="FBC69B02FFFCFFA3FDB3FCA4FDED694E" bold="true" box="[536,635,774,798]" pageId="30" pageNumber="31">SP. NOV.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FE9CFC80FDC46915" box="[311,594,813,837]" pageId="30" pageNumber="31" type="description">
<paragraph id="C90D4710FFFCFFA3FE9CFC80FDC46915" blockId="30.[270,635,744,837]" box="[311,594,813,837]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FE9CFC80FDC46915" allCaps="true" box="[311,594,813,837]" centered="true" fontSize="10" level="1" pageId="30" pageNumber="31" reason="1">(FIGS 1G–I, 2H, 13C, 17)</heading>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FF3AFCD8FE2169FA" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="C90D4710FFFCFFA3FF3AFCD8FE2169FA" blockId="30.[145,762,885,938]" pageId="30" pageNumber="31">
<emphasis id="FBC69B02FFFCFFA3FF3AFCD8FF4269DA" box="[145,212,885,906]" italics="true" pageId="30" pageNumber="31">LSID</emphasis>
:
<uri id="BD234B12FFFCFFA3FF42FCD8FE2169FA" pageId="30" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid id="BD147DC5FFFCFFA3FDB8FCD8FE2169FA" pageId="30" pageNumber="31">F501859D-DC12- 4156-A465-307918472727</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FF3AFC65FEDE6EE4" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="C90D4710FFFCFFA3FF3AFC65FE8D69AC" blockId="30.[145,761,968,1204]" pageId="30" pageNumber="31">
<taxonomicName id="0EB23C93FFFCFFA3FF3AFC65FEA4698D" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,306,968,989]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFCFFA3FF3AFC65FEA4698D" box="[145,306,968,989]" italics="true" pageId="30" pageNumber="31">Hypselodoris</emphasis>
</taxonomicName>
sp. 3
<bibRefCitation id="AD233AE1FFFCFFA3FE2AFC65FD12698D" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[385,644,968,990]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="30" pageNumber="31" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFFCFFA3FE5DFC65FDA0698D" box="[502,566,968,989]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2008
</bibRefCitation>
: 265, top photograph.
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF3AFBA8FE8D6E6A" blockId="30.[145,761,968,1204]" pageId="30" pageNumber="31">
<taxonomicName id="0EB23C93FFFCFFA3FF3AFBA8FEBC6E4A" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,298,1029,1050]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFCFFA3FF3AFBA8FEBC6E4A" box="[145,298,1029,1050]" italics="true" pageId="30" pageNumber="31">Hypselodoris</emphasis>
</taxonomicName>
sp. 4
<bibRefCitation id="AD233AE1FFFCFFA3FEDAFBA8FDF46E4A" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[369,610,1029,1051]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="30" pageNumber="31" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFFCFFA3FE74FBABFD8C6E4A" box="[479,538,1029,1051]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2008
</bibRefCitation>
: 265, second photograph.
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF3AFBEFFE8D6E27" blockId="30.[145,761,968,1204]" pageId="30" pageNumber="31">
<taxonomicName id="0EB23C93FFFCFFA3FF3AFBEFFEB36E07" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,293,1090,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFCFFA3FF3AFBEFFEB36E07" box="[145,293,1090,1111]" italics="true" pageId="30" pageNumber="31">Hypselodoris</emphasis>
</taxonomicName>
sp. 8
<bibRefCitation id="AD233AE1FFFCFFA3FEC2FBEFFDD96E08" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[361,591,1090,1112]" journalOrPublisher="Jacksonville: New World Press" pageId="30" pageNumber="31" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFFCFFA3FE79FBEEFD9D6E07" box="[466,523,1090,1112]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2015
</bibRefCitation>
: 259, top right photograph.
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF3AFB2DFEDE6EE4" blockId="30.[145,761,968,1204]" pageId="30" pageNumber="31">
<taxonomicName id="0EB23C93FFFCFFA3FF3AFB2DFEBC6EC5" ID-CoL="7P5LJ" authorityName="Stimpson" authorityYear="1855" box="[145,298,1152,1173]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFFCFFA3FF3AFB2DFEBC6EC5" box="[145,298,1152,1173]" italics="true" pageId="30" pageNumber="31">Hypselodoris</emphasis>
</taxonomicName>
sp. 9
<bibRefCitation id="AD233AE1FFFCFFA3FEDAFB2DFDF76EC6" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[369,609,1152,1174]" journalOrPublisher="Jacksonville: New World Press" pageId="30" pageNumber="31" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner
<emphasis id="FBC69B02FFFCFFA3FE75FB2DFD8F6EC5" box="[478,537,1152,1173]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2015
</bibRefCitation>
: 259, middle left photograph.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FF3AFB71FC0F687F" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="C90D4710FFFCFFA3FF3AFB71FEAD6EA3" blockId="30.[145,761,1243,1359]" box="[145,315,1243,1267]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FF3AFB71FEAD6EA3" box="[145,315,1243,1267]" fontSize="10" level="4" pageId="30" pageNumber="31" reason="1">
<emphasis id="FBC69B02FFFCFFA3FF3AFB71FEAD6EA3" box="[145,315,1243,1267]" italics="true" pageId="30" pageNumber="31">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF3AFB51FD796F1F" blockId="30.[145,761,1243,1359]" pageId="30" pageNumber="31">
<materialsCitation id="79DA4D4DFFFCFFA3FF3AFB51FD796F1F" ID-GBIF-Occurrence="2235579110" collectingDate="2007-10-05" collectionCode="CASIZ" collectorName="T. Gosliner." country="Malaysia" location="Pulau Chebeh" municipality="Tioman Island" pageId="30" pageNumber="31" specimenCode="CASIZ 176771" specimenCount="1" typeStatus="holotype">
<typeStatus id="1609F9B2FFFCFFA3FF3AFB51FF6F6F41" box="[145,249,1276,1297]" pageId="30" pageNumber="31" type="holotype">
<emphasis id="FBC69B02FFFCFFA3FF3AFB51FF6F6F41" box="[145,249,1276,1297]" italics="true" pageId="30" pageNumber="31">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFFCFFA3FEA7FB51FE2F6F42" box="[268,441,1276,1298]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 176771</specimenCode>
, subsampled for molecular study, dissected,
<location id="CC6D11CBFFFCFFA3FEF8FAB7FE6C6F60" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FEF8FAB7FE6C6F60" box="[339,506,1306,1328]" country="Malaysia" municipality="Tioman Island" name="Pulau Chebeh" pageId="30" pageNumber="31">Pulau Chebeh</location>
, off NW tip of
<collectingMunicipality id="2969DD6AFFFCFFA3FD0BFAB7FF4D6F1F" pageId="30" pageNumber="31">Tioman Island</collectingMunicipality>
, eastern
<collectingCountry id="B1A50780FFFCFFA3FEE9FA94FE3A6F1F" box="[322,428,1337,1359]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FE1CFA94FDF36F1E" box="[439,613,1337,1359]" pageId="30" pageNumber="31" value="2007-10-05">
<date id="BD0C61D0FFFCFFA3FE1CFA94FDF36F1E" box="[439,613,1337,1359]" pageId="30" pageNumber="31" value="2007-10-05">5 October 2007</date>
</collectingDate>
,
<collectorName id="644722C6FFFCFFA3FDC5FA97FD796F1F" box="[622,751,1337,1359]" pageId="30" pageNumber="31">T. Gosliner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF3AFAD5FA196BE3" blockId="30.[145,761,1400,1912]" lastBlockId="30.[809,1425,198,435]" pageId="30" pageNumber="31">
<materialsCitation id="79DA4D4DFFFCFFA3FF3AFAD5FD636F9A" ID-GBIF-Occurrence="2235579085" collectingDate="2007-10-04" collectionCode="CASIZ" collectorName="Batu Sepoi" country="Malaysia" location="Tioman Island" pageId="30" pageNumber="31" specimenCode="CASIZ 176772" specimenCount="1" typeStatus="paratype">
<typeStatus id="1609F9B2FFFCFFA3FF3AFAD5FE946FDD" box="[145,258,1400,1421]" pageId="30" pageNumber="31" type="paratype">
<emphasis id="FBC69B02FFFCFFA3FF3AFAD5FE946FDD" box="[145,258,1400,1421]" italics="true" pageId="30" pageNumber="31">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFFCFFA3FEBFFAD5FE2B6FDE" box="[276,445,1400,1422]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 176772</specimenCode>
, one specimen, subsampled for molecular study,
<collectorName id="644722C6FFFCFFA3FE31FA3AFDBF6FFC" box="[410,553,1430,1452]" pageId="30" pageNumber="31">Batu Sepoi</collectorName>
, off SW coast of
<location id="CC6D11CBFFFCFFA3FF3AFA18FED16F9B" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FF3AFA18FED16F9B" box="[145,327,1461,1483]" country="Malaysia" name="Tioman Island" pageId="30" pageNumber="31">Tioman Island</location>
, eastern
<collectingCountry id="B1A50780FFFCFFA3FE12FA18FDBD6F9B" box="[441,555,1461,1483]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FD93FA18FD636F9A" box="[568,757,1461,1483]" pageId="30" pageNumber="31" value="2007-10-04">
<date id="BD0C61D0FFFCFFA3FD93FA18FD636F9A" box="[568,757,1461,1483]" pageId="30" pageNumber="31" value="2007-10-04">4 October 2007</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="79DA4D4DFFFCFFA3FF3AFA79FDA86FBA" ID-GBIF-Occurrence="2235579083" box="[145,574,1492,1514]" collectionCode="CASIZ" country="Malaysia" pageId="30" pageNumber="31" specimenCode="CASIZ 177532" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FF3AFA79FED66FBA" box="[145,320,1492,1514]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 177532</specimenCode>
, molecular sample of
</materialsCitation>
<materialsCitation id="79DA4D4DFFFCFFA3FDEDFA79FE056C58" ID-GBIF-Occurrence="2235579134" collectionCode="CASIZ" country="Malaysia" pageId="30" pageNumber="31" specimenCode="CASIZ 176772" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FDEDFA79FD626FBA" box="[582,756,1492,1514]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 176772</specimenCode>
, same locality and date
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFFCFFA3FE36FA5EFE6B6C15" ID-GBIF-Occurrence="2235579084" collectingDate="2007-10-06" collectionCode="CASIZ" collectorName="Nichi Asu Maru" country="Malaysia" location="Kuantan" pageId="30" pageNumber="31" specimenCode="CASIZ 176773" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FE36FA5EFDD56C58" box="[413,579,1522,1544]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 176773</specimenCode>
, two specimens,
<collectorName id="644722C6FFFCFFA3FF3AF9BCFEF76C77" box="[145,353,1553,1575]" pageId="30" pageNumber="31">Nichi Asu Maru</collectorName>
wreck, off
<location id="CC6D11CBFFFCFFA3FE5DF9BFFDFF6C77" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FE5DF9BFFDFF6C77" box="[502,617,1554,1575]" country="Malaysia" name="Kuantan" pageId="30" pageNumber="31">Kuantan</location>
, east coast of
<collectingCountry id="B1A50780FFFCFFA3FF18F99DFEBF6C16" box="[179,297,1584,1606]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FE93F99DFE6B6C15" box="[312,509,1584,1606]" pageId="30" pageNumber="31" value="2007-10-06">
<date id="BD0C61D0FFFCFFA3FE93F99DFE6B6C15" box="[312,509,1584,1606]" pageId="30" pageNumber="31" value="2007-10-06">6 October 2007</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FD0CF99DFD8E6CD2" ID-GBIF-Occurrence="2235579109" collectingDate="2007-10-02" collectionCode="CASIZ" collectorName="Tiger Point" country="Malaysia" location="Pulau Tioman" pageId="30" pageNumber="31" specimenCode="CASIZ 176774" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FD0CF99DFF7D6C34" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 176774</specimenCode>
, one specimen,
<collectorName id="644722C6FFFCFFA3FE00F9E3FDAE6C34" box="[427,568,1614,1636]" pageId="30" pageNumber="31">Tiger Point</collectorName>
,
<location id="CC6D11CBFFFCFFA3FDEDF9E3FD626C34" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FDEDF9E3FD626C34" box="[582,756,1614,1636]" country="Malaysia" name="Pulau Tioman" pageId="30" pageNumber="31">Pulau Tioman</location>
, eastern
<collectingCountry id="B1A50780FFFCFFA3FF44F9C0FECD6CD3" box="[239,347,1645,1667]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FECCF9C0FD8E6CD2" box="[359,536,1645,1667]" pageId="30" pageNumber="31" value="2007-10-02">
<date id="BD0C61D0FFFCFFA3FECCF9C0FD8E6CD2" box="[359,536,1645,1667]" pageId="30" pageNumber="31" value="2007-10-02">2 October 2007</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FD07F9C0FD636C90" ID-GBIF-Occurrence="2235579058" collectingDate="2007-10-06" collectionCode="CASIZ" collectorName="Nichi Asu Maru" country="Malaysia" location="Kuantan" pageId="30" pageNumber="31" specimenCode="CASIZ 175728" specimenCount="1" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FD07F9C0FF7C6CF2" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 175728</specimenCode>
, two specimens,
<collectorName id="644722C6FFFCFFA3FE1CF921FDEC6CF1" box="[439,634,1676,1698]" pageId="30" pageNumber="31">Nichi Asu Maru</collectorName>
wreck, off
<location id="CC6D11CBFFFCFFA3FF3AF906FE966C90" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FF3AF906FE966C90" box="[145,256,1707,1728]" country="Malaysia" name="Kuantan" pageId="30" pageNumber="31">Kuantan</location>
, east coast of
<collectingCountry id="B1A50780FFFCFFA3FE1FF907FDB16C90" box="[436,551,1706,1728]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FD9EF907FD636C90" box="[565,757,1706,1728]" pageId="30" pageNumber="31" value="2007-10-06">
<date id="BD0C61D0FFFCFFA3FD9EF907FD636C90" box="[565,757,1706,1728]" pageId="30" pageNumber="31" value="2007-10-06">6 October 2007</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FEBCF964FDFC6D6A" ID-GBIF-Occurrence="2235579139" collectingDate="2009-05-20" collectionCode="CASIZ" collectorName="Mainit Bubbles" country="Philippines" county="Luzon Island" latitude="13.688027" location="Calumpan Peninsula" longLatPrecision="1" longitude="120.95809" municipality="Mabini" pageId="30" pageNumber="31" specimenCode="CASIZ 181300" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FEBCF964FE286C8E" box="[279,446,1737,1758]" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 181300</specimenCode>
, one specimen, subsampled for molecular study,
<collectorName id="644722C6FFFCFFA3FE2DF945FDD66CAE" box="[390,576,1768,1790]" pageId="30" pageNumber="31">Mainit Bubbles</collectorName>
,
<geoCoordinate id="AC8621D7FFFCFFA3FDE6F945FD636CAE" box="[589,757,1768,1790]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="13.688027">13.6880278°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFCFFA3FF3AF8ABFEB56D4C" box="[145,291,1798,1820]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="120.95809">120.95809°E</geoCoordinate>
,
<location id="CC6D11CBFFFCFFA3FE86F8ABFD886D4C" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FE86F8ABFD886D4C" box="[301,542,1798,1820]" country="Philippines" county="Luzon Island" latitude="13.688027" longLatPrecision="1" longitude="120.95809" municipality="Mabini" name="Calumpan Peninsula" pageId="30" pageNumber="31" stateProvince="Batangas">Calumpan Peninsula</location>
,
<collectingMunicipality id="2969DD6AFFFCFFA3FD82F8ABFDEA6D4C" box="[553,636,1798,1820]" pageId="30" pageNumber="31">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFFCFFA3FD2CF8AAFD626D4C" box="[647,756,1799,1820]" country="Philippines" name="Batangas" pageId="30" pageNumber="31">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFFCFFA3FF3AF88BFEB96D6B" box="[145,303,1829,1851]" pageId="30" pageNumber="31">Luzon Island</collectingCounty>
,
<collectingCountry id="B1A50780FFFCFFA3FE90F888FE556D6B" box="[315,451,1829,1851]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FE64F888FDFC6D6A" box="[463,618,1829,1851]" pageId="30" pageNumber="31" value="2009-05-20">
<date id="BD0C61D0FFFCFFA3FE64F888FDFC6D6A" box="[463,618,1829,1851]" pageId="30" pageNumber="31" value="2009-05-20">20 May 2009</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FF3AF8E9FC086AAA" ID-GBIF-Occurrence="2235579042" collectingDate="2009-05-21" collectionCode="CASIZ" collectorName="Mainit Bubbles" country="Philippines" county="Luzon Island" latitude="13.688027" location="Calumpan Peninsula" longLatPrecision="1" longitude="120.95809" municipality="Mabini" pageId="30" pageNumber="31" specimenCode="CASIZ 181251" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FF3AF8E9FEDB6D0A" box="[145,333,1860,1882]" collectionCode="CASIZ" pageId="30" pageNumber="31">
CASIZ
<date id="BD0C61D0FFFCFFA3FF44F8E9FEDB6D0A" box="[239,333,1860,1882]" pageId="30" pageNumber="31" value="51-12-18">181251</date>
</specimenCode>
, two specimens,
<collectorName id="644722C6FFFCFFA3FD80F8E9FD626D0A" box="[555,756,1860,1882]" pageId="30" pageNumber="31">Mainit Bubbles</collectorName>
,
<geoCoordinate id="AC8621D7FFFCFFA3FF3AF8CFFEAA6D28" box="[145,316,1890,1912]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="13.688027">13.6880278°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFCFFA3FEE1F8CFFE736D28" box="[330,485,1890,1912]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="120.95809">120.95809°E</geoCoordinate>
,
<location id="CC6D11CBFFFCFFA3FE58F8CFFD636D28" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FE58F8CFFD636D28" box="[499,757,1890,1912]" country="Philippines" county="Luzon Island" latitude="13.688027" longLatPrecision="1" longitude="120.95809" municipality="Mabini" name="Calumpan Peninsula" pageId="30" pageNumber="31" stateProvince="Batangas">Calumpan Peninsula</location>
,
<collectingMunicipality id="2969DD6AFFFCFFA3FC82FF6BFC136A8C" box="[809,901,198,220]" pageId="30" pageNumber="31">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFFCFFA3FC38FF6AFB9D6A8C" box="[915,1035,199,220]" country="Philippines" name="Batangas" pageId="30" pageNumber="31">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFFCFFA3FBB0FF6AFB526A8C" box="[1051,1220,198,220]" pageId="30" pageNumber="31">Luzon Island</collectingCounty>
,
<collectingCountry id="B1A50780FFFCFFA3FB78FF6BFAF36A8C" box="[1235,1381,198,220]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FADFFF6AFC086AAA" pageId="30" pageNumber="31" value="2009-05-21">
<date id="BD0C61D0FFFCFFA3FADFFF6AFC086AAA" pageId="30" pageNumber="31" value="2009-05-21">21 May 2009</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FB9EFF48FB3B6B06" ID-GBIF-Occurrence="2235579135" collectingDate="2009-05-21" collectionCode="CASIZ" country="Philippines" county="Luzon Island" location="Calumpan Peninsula" municipality="Mabini" pageId="30" pageNumber="31" specimenCode="CASIZ 181227" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FB9EFF48FB776AAA" box="[1077,1249,229,250]" collectionCode="CASIZ" pageId="30" pageNumber="31">
CASIZ
<date id="BD0C61D0FFFCFFA3FB21FF48FB776AAA" box="[1162,1249,229,250]" pageId="30" pageNumber="31" value="27-12-18">181227</date>
</specimenCode>
, one specimen, M a i n i t B u b b l e s, 1 3.6 8 8 0 2 7 8° N, 1 2 0.9 5 8 0 9°E,
<location id="CC6D11CBFFFCFFA3FC82FE8FFBAD6B68" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FC82FE8FFBAD6B68" box="[809,1083,290,312]" country="Philippines" county="Luzon Island" municipality="Mabini" name="Calumpan Peninsula" pageId="30" pageNumber="31" stateProvince="Batangas">Calumpan Peninsula</location>
,
<collectingMunicipality id="2969DD6AFFFCFFA3FBE0FE8FFB3F6B68" box="[1099,1193,290,312]" pageId="30" pageNumber="31">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFFCFFA3FB13FE8EFAA56B68" box="[1208,1331,291,312]" country="Philippines" name="Batangas" pageId="30" pageNumber="31">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFFCFFA3FAEFFE8EFCE36B07" pageId="30" pageNumber="31">Luzon Island</collectingCounty>
,
<collectingCountry id="B1A50780FFFCFFA3FC2AFEECFB9E6B07" box="[897,1032,321,343]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FBBFFEECFB3B6B06" box="[1044,1197,321,343]" pageId="30" pageNumber="31" value="2009-05-21">
<date id="BD0C61D0FFFCFFA3FBBFFEECFB3B6B06" box="[1044,1197,321,343]" pageId="30" pageNumber="31" value="2009-05-21">21 May 2009</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
<materialsCitation id="79DA4D4DFFFCFFA3FAE8FEECFA936BE2" ID-GBIF-Occurrence="2235579070" collectingDate="2008-03-21" collectionCode="CASIZ" collectorName="Mainit Bubbles" country="Philippines" county="Luzon Island" latitude="13.688027" location="Calumpan Peninsula" longLatPrecision="1" longitude="120.95809" municipality="Mabini" pageId="30" pageNumber="31" specimenCode="CASIZ 177532" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFFCFFA3FAE8FEECFCEB6B26" collectionCode="CASIZ" pageId="30" pageNumber="31">CASIZ 177532</specimenCode>
, one specimen,
<collectorName id="644722C6FFFCFFA3FB84FECDFB766B26" box="[1071,1248,352,374]" pageId="30" pageNumber="31">Mainit Bubbles</collectorName>
,
<geoCoordinate id="AC8621D7FFFCFFA3FB47FECDFA1B6B26" box="[1260,1421,352,374]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="13.688027">13.6880278°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFCFFA3FC82FED3FC2D6BC4" box="[809,955,382,404]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="120.95809">120.95809°E</geoCoordinate>
,
<location id="CC6D11CBFFFCFFA3FC6EFED3FB206BC4" LSID="urn:lsid:plazi:treatment:411BF606FFFCFF9DFEB1FD45FAE66B68:CC6D11CBFFFCFFA3FC6EFED3FB206BC4" box="[965,1206,382,404]" country="Philippines" county="Luzon Island" latitude="13.688027" longLatPrecision="1" longitude="120.95809" municipality="Mabini" name="Calumpan Peninsula" pageId="30" pageNumber="31" stateProvince="Batangas">Calumpan Peninsula</location>
,
<collectingMunicipality id="2969DD6AFFFCFFA3FB6AFED3FA826BC4" box="[1217,1300,382,404]" pageId="30" pageNumber="31">Mabini</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFFCFFA3FAB4FED2FA1A6BC4" box="[1311,1420,383,404]" country="Philippines" name="Batangas" pageId="30" pageNumber="31">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFFCFFA3FC82FE33FC566BE3" box="[809,960,413,435]" pageId="30" pageNumber="31">Luzon Island</collectingCounty>
,
<collectingCountry id="B1A50780FFFCFFA3FC60FE30FBD86BE3" box="[971,1102,413,435]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFFCFFA3FBF2FE30FA936BE2" box="[1113,1285,413,435]" pageId="30" pageNumber="31" value="2008-03-21">
<date id="BD0C61D0FFFCFFA3FBF2FE30FA936BE2" box="[1113,1285,413,435]" pageId="30" pageNumber="31" value="2008-03-21">21 March 2008</date>
</collectingDate>
</materialsCitation>
, T. Gosliner.
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82FE76FC526BA2" blockId="30.[809,1425,474,559]" box="[809,964,474,498]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FC82FE76FC526BA2" box="[809,964,474,498]" fontSize="10" level="4" pageId="30" pageNumber="31" reason="1">
<emphasis id="FBC69B02FFFCFFA3FC82FE76FC526BA2" box="[809,964,474,498]" italics="true" pageId="30" pageNumber="31">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82FE57FC0F687F" blockId="30.[809,1425,474,559]" pageId="30" pageNumber="31">
Pulau Chebeh, off NW tip of Tioman Island, eastern
<collectingCountry id="B1A50780FFFCFFA3FC82FDB4FC03687F" box="[809,917,537,559]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FC82FDFBFB0C68FB" pageId="30" pageNumber="31" type="distribution">
<paragraph id="C90D4710FFFCFFA3FC82FDFBFBF0683E" blockId="30.[809,1424,598,683]" box="[809,1126,598,622]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FC82FDFBFBF0683E" box="[809,1126,598,622]" fontSize="10" level="4" pageId="30" pageNumber="31" reason="1">
<emphasis id="FBC69B02FFFCFFA3FC82FDFBFBF0683E" box="[809,1126,598,622]" italics="true" pageId="30" pageNumber="31">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82FDDAFB0C68FB" blockId="30.[809,1424,598,683]" pageId="30" pageNumber="31">
Known from the
<collectingCountry id="B1A50780FFFCFFA3FC5EFDDBFBE968DC" box="[1013,1151,630,652]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
, eastern
<collectingCountry id="B1A50780FFFCFFA3FB45FDDBFACD68DC" box="[1262,1371,630,652]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
and
<collectingCountry id="B1A50780FFFCFFA3FC82FD38FC0F68FB" box="[809,921,661,683]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
(
<bibRefCitation id="AD233AE1FFFCFFA3FC03FD38FB1D68FA" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[936,1163,661,683]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="30" pageNumber="31" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner
<emphasis id="FBC69B02FFFCFFA3FBA4FD3BFBD168FA" box="[1039,1095,661,683]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFFA3FC82FD7FFB8C6990" pageId="30" pageNumber="31" type="etymology">
<paragraph id="C90D4710FFFCFFA3FC82FD7FFC3C68BA" blockId="30.[809,1425,722,960]" box="[809,938,722,746]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FC82FD7FFC3C68BA" box="[809,938,722,746]" fontSize="10" level="4" pageId="30" pageNumber="31" reason="1">
<emphasis id="FBC69B02FFFCFFA3FC82FD7FFC3C68BA" box="[809,938,722,746]" italics="true" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82FD5FFB8C6990" blockId="30.[809,1425,722,960]" pageId="30" pageNumber="31">
This species is named for Katherine Piatek, Senior Program Manager of the Institute of Biodiversity Science and Sustainability at the California Academy of Sciences, who has provided immense support for the research undertaken in the
<collectingCountry id="B1A50780FFFCFFA3FBF4FCC0FB7569D3" box="[1119,1251,877,899]" name="Philippines" pageId="30" pageNumber="31">Philippines</collectingCountry>
. Her assistance has been crucial to the successful completion of many research expeditions.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFCFF9DFC82FC45FE6A6897" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="C90D4710FFFCFFA3FC82FC45FC2369AF" blockId="30.[809,1425,1000,1514]" box="[809,949,1000,1023]" pageId="30" pageNumber="31">
<heading id="9245F07CFFFCFFA3FC82FC45FC2369AF" box="[809,949,1000,1023]" fontSize="10" level="4" pageId="30" pageNumber="31" reason="1">
<emphasis id="FBC69B02FFFCFFA3FC82FC45FC2369AF" box="[809,949,1000,1023]" italics="true" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82FBA5FB2E6FBA" blockId="30.[809,1425,1000,1514]" pageId="30" pageNumber="31">
<emphasis id="FBC69B02FFFCFFA3FC82FBA5FBB86E4D" box="[809,1070,1032,1053]" italics="true" pageId="30" pageNumber="31">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFFCFFA3FAB9FBA5FAC06E4D" box="[1298,1366,1032,1054]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="30" pageNumber="31">Fig. 1</figureCitation>
G–I) moderately large,
<quantity id="0E4AEAF5FFFCFFA3FC41FB8AFBCD6E6C" box="[1002,1115,1063,1084]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.2" metricValueMin="2.0" pageId="30" pageNumber="31" unit="mm" value="26.0" valueMax="32.0" valueMin="20.0">20–32 mm</quantity>
in length. Body long, slender, translucent pink to peach. Fine parallel or intersecting red longitudinal lines or interrupted dashes on notum sides of body and foot. Large, scattered opaque white tubercles on notum of larger individuals. Scattered bluepurple spots found around margin of notum and foot, forming continuous line in some specimens. Pigment more dense in centre of spot. Seven to nine unipinnate gill branches having a translucent white base and inner margin and bright red–orange pigment on inner and outer surfaces of rachis and some gill filaments. Bulb of perfoliate rhinophores mostly red–orange, with some translucent white on anterior face. Bulb with eight to 16 congested lamellae. Base of rhinophores red anteriorly, with opaque white on posterior face.
</paragraph>
<paragraph id="C90D4710FFFCFFA3FC82F9BFFA046C91" blockId="30.[809,1426,1554,1730]" pageId="30" pageNumber="31">
<emphasis id="FBC69B02FFFCFFA3FC82F9BFFC456C77" box="[809,979,1554,1575]" italics="true" pageId="30" pageNumber="31">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFFCFFA3FBFBF99CFB386C16" box="[1104,1198,1585,1607]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="30" pageNumber="31">Fig. 2H</figureCitation>
). Glands situated around entire margin of anterior and posterior ends in two specimens examined (CASIZ 177532, CASIZ 176771). Few scattered lateral glands also present. Anterior and posterior glands larger than lateral ones.
</paragraph>
<paragraph id="C90D4710FFFCFFA2FC82F947FA356C86" blockId="30.[809,1425,1770,1915]" lastBlockId="31.[827,1443,1637,1751]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="FBC69B02FFFCFFA3FC82F947FC486D50" box="[809,990,1770,1792]" italics="true" pageId="30" pageNumber="31">Buccal armature</emphasis>
: Muscular portion of buccal mass about equal in length to oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFFCFFA3FB61F8EBFAB16D0C" box="[1226,1319,1862,1884]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="30" pageNumber="31">Fig. 17A</figureCitation>
). Rodlets narrow and acutely pointed anteriorly, with single apex (
<figureCitation id="51895B95FFFDFFA2FF41F9C8FEDA6C2A" box="[234,332,1637,1659]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32">Fig. 17B</figureCitation>
). Radular formula of
<typeStatus id="1609F9B2FFFDFFA2FDE0F9C8FD3B6C2B" box="[587,685,1637,1659]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
(CASIZ 176771) 55 × 53.0.53, and 61 × 67.0.
<quantity id="0E4AEAF5FFFDFFA2FD93F929FDE76CCA" box="[568,625,1668,1690]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" pageId="31" pageNumber="32" unit="in" value="67.0">67 in</quantity>
one
<typeStatus id="1609F9B2FFFDFFA2FD0CF929FC9D6CC9" box="[679,779,1668,1689]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
(CASIZ
<date id="BD0C61D0FFFDFFA2FF51F90FFEDF6CE8" box="[250,329,1698,1720]" pageId="31" pageNumber="32" value="51-12-18">181251</date>
;
<figureCitation id="51895B95FFFDFFA2FEFEF90FFE396CE7" box="[341,431,1698,1720]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32">Fig. 17C</figureCitation>
). Rachidian row of teeth absent (
<figureCitation id="51895B95FFFDFFA2FF00F96CFE9C6C86" box="[171,266,1729,1751]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32">Fig. 17D</figureCitation>
). Innermost lateral teeth having one to three triangular denticles on inner side of bifid primary cusp, with another one or two denticles on the outside. Next several laterals lacking inner triangular denticles but possessing one to three denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFFDFFA2FE42F8F7FDD36D3F" box="[489,581,1882,1904]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32">Fig. 17E</figureCitation>
) all lacking inner denticles but having three to five sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and somewhat shorter tooth shape, with four to seven rounded to pointed outer denticles (
<figureCitation id="51895B95FFFDFFA2FA95F96CFA036C86" box="[1342,1429,1729,1751]" captionStart="Figure 17" captionStartId="31.[162,241,1549,1571]" captionTargetBox="[226,1378,195,1508]" captionTargetId="figure@31.[226,1378,195,1508]" captionTargetPageId="31" captionText="Figure 17. Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32">Fig. 17F</figureCitation>
).
</paragraph>
<caption id="9DCD1798FFFDFFA2FF09F9A0FB4C6C11" httpUri="https://zenodo.org/record/2632808/files/figure.png" pageId="31" pageNumber="32" startId="31.[162,241,1549,1571]" targetBox="[226,1378,195,1508]" targetPageId="31">
<paragraph id="C90D4710FFFDFFA2FF09F9A0FB4C6C11" blockId="31.[162,1443,1549,1601]" pageId="31" pageNumber="32">
<emphasis id="FBC69B02FFFDFFA2FF09F9A0FE8E6C72" bold="true" box="[162,280,1549,1571]" pageId="31" pageNumber="32">Figure 17.</emphasis>
<taxonomicName id="0EB23C93FFFDFFA2FE88F9A0FC8E6C73" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,792,1549,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFFDFFA2FE88F9A0FDD76C73" bold="true" box="[291,577,1549,1571]" italics="true" pageId="31" pageNumber="32">Hypselodoris katherinae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFDFFA2FCB4F9A3FCE06C73" box="[799,886,1550,1571]" pageId="31" pageNumber="32" rank="species">
<emphasis id="FBC69B02FFFDFFA2FCB4F9A3FCE06C73" bold="true" box="[799,886,1550,1571]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, CASIZ 181251, buccal armature. A, entire jaw. B, jaw rodlets. C, whole radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFFDFF9DFC90F950FE6A6897" blockId="31.[827,1443,1789,1903]" lastBlockId="32.[145,762,198,711]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="FBC69B02FFFDFFA2FC90F950FB8A6D43" box="[827,1052,1789,1811]" italics="true" pageId="31" pageNumber="32">Reproductive system</emphasis>
: Reproductive organs fully mature in one specimen examined (CASIZ 176771;
<figureCitation id="51895B95FFFDFFA2FA98F8B1FA026D61" box="[1331,1428,1820,1842]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="31" pageNumber="32">Fig. 13C</figureCitation>
). Ampulla thick, tubular, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick, covering bursa copulatrix. Vas deferens narrowing slightly as it transitions into long, convoluted, muscular ejaculatory portion. Ejaculatory portion widening into enlarged penial bulb. Penial papilla distinctly curved with broad base, devoid of penial hooks. Penial bulb adjacent to thick, slightly narrower vaginal duct at common gonopore. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Vagina thick, curved, leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum situated near base of bursa. Moderately short, narrow uterine duct emerging from vagina distal to base of bursa and female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC2FF9DFF3AFD43FAE66B68" pageId="32" pageNumber="33" type="discussion">
<paragraph id="C90D4710FFC2FF9DFF3AFD43FF686956" blockId="32.[145,762,750,1908]" box="[145,254,750,774]" pageId="32" pageNumber="33">
<heading id="9245F07CFFC2FF9DFF3AFD43FF686956" box="[145,254,750,774]" fontSize="10" level="4" pageId="32" pageNumber="33" reason="1">
<emphasis id="FBC69B02FFC2FF9DFF3AFD43FF686956" box="[145,254,750,774]" italics="true" pageId="32" pageNumber="33">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC2FF9DFF3AFCA3FD906FF8" blockId="32.[145,762,750,1908]" pageId="32" pageNumber="33">
<taxonomicName id="0EB23C93FFC2FF9DFF3AFCA3FE3E6973" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,424,782,803]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFF3AFCA3FE3E6973" box="[145,424,782,803]" italics="true" pageId="32" pageNumber="33">Hypselodoris katherinae</emphasis>
</taxonomicName>
is a member of a large clade that includes
<taxonomicName id="0EB23C93FFC2FF9DFE9AFC83FE376912" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[305,417,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFC2FF9DFE9AFC83FE376912" box="[305,417,813,835]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC2FF9DFE02FC83FD966913" box="[425,512,814,835]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="0EB23C93FFC2FF9DFDA7FC83FD006912" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[524,662,813,835]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFC2FF9DFDA7FC83FD006912" box="[524,662,813,835]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC2FF9DFD35FC83FD636913" box="[670,757,814,835]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="FBC69B02FFC2FF9DFF3AFCE1FEB26931" box="[145,292,844,865]" italics="true" pageId="32" pageNumber="33">H. maritima</emphasis>
,
<taxonomicName id="0EB23C93FFC2FF9DFE98FCE1FD6F6931" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[307,761,844,866]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="rudmani">
<emphasis id="FBC69B02FFC2FF9DFE98FCE1FE286931" box="[307,446,844,865]" italics="true" pageId="32" pageNumber="33">H. rudmani</emphasis>
<bibRefCitation id="AD233AE1FFC2FF9DFE6CFCE1FD6F6931" author="Gosliner T &amp; Johnson, R." box="[455,761,844,866]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="32" pageNumber="33" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFF69FCC6FDF769D0" authority="Gosliner &amp; Johnson, 1999" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[194,609,874,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="bertschi">
<emphasis id="FBC69B02FFC2FF9DFF69FCC6FEAF692F" box="[194,313,874,896]" italics="true" pageId="32" pageNumber="33">H. bertschi</emphasis>
<bibRefCitation id="AD233AE1FFC2FF9DFEEBFCC7FDF769D0" author="Gosliner T &amp; Johnson, R." box="[320,609,874,896]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="32" pageNumber="33" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999</bibRefCitation>
</taxonomicName>
. It is sister to
<taxonomicName id="0EB23C93FFC2FF9DFF3AFC27FE8569CE" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,275,905,927]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFF3AFC27FE8569CE" box="[145,275,905,927]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFEECFC27FE3869CE" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[327,430,905,927]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFEECFC27FE3869CE" box="[327,430,905,927]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
. The ABGD analysis indicates that
<taxonomicName id="0EB23C93FFC2FF9DFF61FC05FEFD69ED" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[202,363,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFF61FC05FEFD69ED" box="[202,363,936,957]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
is clearly distinct from
<taxonomicName id="0EB23C93FFC2FF9DFD23FC05FD6F69ED" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[648,761,936,957]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFD23FC05FD6F69ED" box="[648,761,936,957]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFF6EFC6AFEC6698B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[197,336,966,988]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFF6EFC6AFEC6698B" box="[197,336,966,988]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
. All three species have longitudinal lines, but they are opaque white in
<taxonomicName id="0EB23C93FFC2FF9DFD93FC4BFD5369AA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[568,709,997,1019]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFD93FC4BFD5369AA" box="[568,709,997,1019]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
and brown in
<taxonomicName id="0EB23C93FFC2FF9DFF56FBA9FEFF6E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[253,361,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFF56FBA9FEFF6E49" box="[253,361,1028,1049]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFC2FF9DFEDDFBA9FD1F6E49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[374,649,1028,1049]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFEDDFBA9FD1F6E49" box="[374,649,1028,1049]" italics="true" pageId="32" pageNumber="33">Hypselodoris katherinae</emphasis>
</taxonomicName>
is unique among described members of
<taxonomicName id="0EB23C93FFC2FF9DFE47FB8FFD146E67" authorityName="Stimpson" authorityYear="1855" box="[492,642,1058,1079]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC2FF9DFE47FB8FFD146E67" box="[492,642,1058,1079]" italics="true" pageId="32" pageNumber="33">Hypselodoris</emphasis>
</taxonomicName>
in having longitudinal red lines or dashes. In
<taxonomicName id="0EB23C93FFC2FF9DFD92FBEFFD536E06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[569,709,1089,1111]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFD92FBEFFD536E06" box="[569,709,1089,1111]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFF3AFBCDFF6A6E25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,252,1120,1141]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFF3AFBCDFF6A6E25" box="[145,252,1120,1141]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
there are additional opaque white spots that are absent in
<taxonomicName id="0EB23C93FFC2FF9DFE92FBD2FE4B6EC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[313,477,1150,1172]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFE92FBD2FE4B6EC3" box="[313,477,1150,1172]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
.
<taxonomicName id="0EB23C93FFC2FF9DFE46FBD3FD6F6EC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[493,761,1150,1171]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFE46FBD3FD6F6EC3" box="[493,761,1150,1171]" italics="true" pageId="32" pageNumber="33">Hypselodoris paradisa</emphasis>
</taxonomicName>
also has minute black spots that are absent in the other two species. The three species also differ in the ornamentation of the rhinophores. All three have red pigment on the rhinophores, but in
<taxonomicName id="0EB23C93FFC2FF9DFDB5FB57FD206F5E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[542,694,1273,1295]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFDB5FB57FD206F5E" box="[542,694,1273,1295]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
there are no distinct rings, whereas there are two rings in
<taxonomicName id="0EB23C93FFC2FF9DFF3AFA9AFE876F1B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,273,1334,1356]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFF3AFA9AFE876F1B" box="[145,273,1334,1356]" italics="true" pageId="32" pageNumber="33">H.paradisa</emphasis>
</taxonomicName>
and three rings in
<taxonomicName id="0EB23C93FFC2FF9DFE73FA9AFDA96F1B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[472,575,1334,1356]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFE73FA9AFDA96F1B" box="[472,575,1334,1356]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFC2FF9DFDCFFA9AFD6E6F1B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[612,760,1334,1356]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFDCFFA9AFD6E6F1B" box="[612,760,1334,1356]" italics="true" pageId="32" pageNumber="33">H.katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFF6EFAFBFEC46F3A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[197,338,1365,1387]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFF6EFAFBFEC46F3A" box="[197,338,1365,1387]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
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there are only a few lateral mantle glands, whereas they are largely continuous in the many specimens of
<taxonomicName id="0EB23C93FFC2FF9DFE8FFA3EFE1B6FF7" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[292,397,1426,1448]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFE8FFA3EFE1B6FF7" box="[292,397,1426,1448]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
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examined.
</paragraph>
<paragraph id="C90D4710FFC2FF9DFF02FA1CFD6C6C12" blockId="32.[145,762,750,1908]" pageId="32" pageNumber="33">
The jaw rodlets of
<taxonomicName id="0EB23C93FFC2FF9DFE2AFA1FFDB66F96" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[385,544,1457,1479]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFE2AFA1FFDB66F96" box="[385,544,1457,1479]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
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have only a single cusp, whereas there may be one or two cusps in
<taxonomicName id="0EB23C93FFC2FF9DFF3AFA42FE8D6C53" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,283,1518,1540]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFF3AFA42FE8D6C53" box="[145,283,1518,1540]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
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and one to three cusps in
<taxonomicName id="0EB23C93FFC2FF9DFDFBFA42FD296C53" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[592,703,1518,1540]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFDFBFA42FD296C53" box="[592,703,1518,1540]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
. The radulae of
<taxonomicName id="0EB23C93FFC2FF9DFEBBF9A3FE246C72" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[272,434,1549,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFEBBF9A3FE246C72" box="[272,434,1549,1571]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFC2FF9DFE6BF9A3FDDA6C72" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[448,588,1549,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFE6BF9A3FDDA6C72" box="[448,588,1549,1571]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFD23F9A3FD6F6C72" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[648,761,1549,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFD23F9A3FD6F6C72" box="[648,761,1549,1571]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
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all have a similar formula and teeth of a similar shape.
</paragraph>
<paragraph id="C90D4710FFC2FF9DFF02F9E7FAE66B68" blockId="32.[145,762,750,1908]" lastBlockId="32.[809,1425,198,312]" pageId="32" pageNumber="33">
The reproductive system is similar in all three species. In
<taxonomicName id="0EB23C93FFC2FF9DFEB8F9C7FE226C2E" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[275,436,1641,1663]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFEB8F9C7FE226C2E" box="[275,436,1641,1663]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
, the vagina is elongate and uniformly wide. In
<taxonomicName id="0EB23C93FFC2FF9DFECDF925FE7A6CCD" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[358,492,1672,1693]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFECDF925FE7A6CCD" box="[358,492,1672,1693]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
, the vagina is short and uniformly wide, and
<taxonomicName id="0EB23C93FFC2FF9DFE2CF90AFE6E6CEB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[391,504,1702,1724]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFE2CF90AFE6E6CEB" box="[391,504,1702,1724]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
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has a relatively thin, elongate vagina, where the portion below the bursa copulatrix is much wider. All three species, together with the sister clade including
<emphasis id="FBC69B02FFC2FF9DFE42F8AEFDE26D48" box="[489,628,1795,1816]" italics="true" pageId="32" pageNumber="33">H. maritima</emphasis>
,
<taxonomicName id="0EB23C93FFC2FF9DFD2BF8AEFD6F6D47" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[640,761,1794,1816]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="bertschi">
<emphasis id="FBC69B02FFC2FF9DFD2BF8AEFD6F6D47" box="[640,761,1794,1816]" italics="true" pageId="32" pageNumber="33">H. bertschi</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFC2FF9DFF68F88FFEDC6D66" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[195,330,1825,1847]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="rudmani">
<emphasis id="FBC69B02FFC2FF9DFF68F88FFEDC6D66" box="[195,330,1825,1847]" italics="true" pageId="32" pageNumber="33">H. rudmani</emphasis>
</taxonomicName>
, have a much wider penial bulb than is found in other species of
<taxonomicName id="0EB23C93FFC2FF9DFE79F8EDFDFF6D05" authorityName="Stimpson" authorityYear="1855" box="[466,617,1856,1877]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="FBC69B02FFC2FF9DFE79F8EDFDFF6D05" box="[466,617,1856,1877]" italics="true" pageId="32" pageNumber="33">Hypselodoris</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFC2FF9DFDD1F8EDFE7A6D24" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="32" pageNumber="33" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999; present study</bibRefCitation>
). In
<taxonomicName id="0EB23C93FFC2FF9DFD88F8F2FD526D23" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[547,708,1886,1908]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFC2FF9DFD88F8F2FD526D23" box="[547,708,1886,1908]" italics="true" pageId="32" pageNumber="33">H. katherinae</emphasis>
</taxonomicName>
, the penial papilla is curved with a broad base, whereas it is blunt and discoidal in
<taxonomicName id="0EB23C93FFC2FF9DFBFFFF4BFB746AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1108,1250,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFC2FF9DFBFFFF4BFB746AAA" box="[1108,1250,229,251]" italics="true" pageId="32" pageNumber="33">H. paradisa</emphasis>
</taxonomicName>
and conical in
<taxonomicName id="0EB23C93FFC2FF9DFC82FEA9FC026B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,916,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFC2FF9DFC82FEA9FC026B49" box="[809,916,260,281]" italics="true" pageId="32" pageNumber="33">H. skyleri</emphasis>
</taxonomicName>
. The penis of
<taxonomicName id="0EB23C93FFC2FF9DFB85FEA9FB246B49" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1070,1202,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="rudmani">
<emphasis id="FBC69B02FFC2FF9DFB85FEA9FB246B49" box="[1070,1202,260,281]" italics="true" pageId="32" pageNumber="33">H. rudmani</emphasis>
</taxonomicName>
is unique in having a ring of muscular plates at the apex of the papilla.
</paragraph>
</subSubSection>
</treatment>
</document>

347
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<paragraph id="C90D4710FFFEFFA1FC10FD80FA856833" blockId="28.[934,1299,557,650]" pageId="28" pageNumber="29">
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<emphasis id="FBC69B02FFFEFFA1FC10FD80FB696813" bold="true" box="[955,1279,557,580]" italics="true" pageId="28" pageNumber="29">HYPSELODORIS JUNIPERAE</emphasis>
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<heading id="9245F07CFFFEFFA1FC0DFDE6FB3F6832" allCaps="true" box="[934,1193,587,611]" centered="true" fontSize="9" level="3" pageId="28" pageNumber="29" reason="4">GOSLINER &amp; JOHNSON</heading>
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<emphasis id="FBC69B02FFFEFFA1FB1BFDE2FA856833" bold="true" box="[1200,1299,587,611]" pageId="28" pageNumber="29">SP. NOV.</emphasis>
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<heading id="9245F07CFFFEFFA1FC57FDDFFB2B68DA" allCaps="true" box="[1020,1213,626,650]" centered="true" fontSize="10" level="1" pageId="28" pageNumber="29" reason="1">(FIGS 1F, 2G, 16)</heading>
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<emphasis id="FBC69B02FFFEFFA1FC82FD16FCEC6880" box="[809,890,699,720]" italics="true" pageId="28" pageNumber="29">L S I D:</emphasis>
u r n: l s i d: z o o b a n k. o r g:a c t:
<uuid id="BD147DC5FFFEFFA1FB59FD16FB2768BF" pageId="28" pageNumber="29">3 E5 9 F D 3 C - 9CDD-4663-A867-EF74877AB7A8</uuid>
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA1FC82FCA0FAFF6973" box="[809,1385,781,803]" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="C90D4710FFFEFFA1FC82FCA0FAFF6973" blockId="28.[809,1385,781,803]" box="[809,1385,781,803]" pageId="28" pageNumber="29">
<taxonomicName id="0EB23C93FFFEFFA1FC82FCA0FA9F6973" ID-CoL="3P34H" authority="Rudman, 1999 b" authorityName="Rudman" authorityYear="1999" box="[809,1289,781,803]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFEFFA1FC82FCA0FC2A6972" box="[809,956,781,802]" italics="true" pageId="28" pageNumber="29">Hypselodoris</emphasis>
cf.
<emphasis id="FBC69B02FFFEFFA1FC49FCA0FBD86972" box="[994,1102,781,802]" italics="true" pageId="28" pageNumber="29">maculosa</emphasis>
<bibRefCitation id="AD233AE1FFFEFFA1FBFFFCA0FA9F6973" author="Rudman WB" box="[1108,1289,781,803]" journalOrPublisher="Sydney: Australian Museum" pageId="28" pageNumber="29" publicationUrl="http://www.seaslugforum.net/find/hypskang" refId="ref44958" refString="Rudman WB. 1999 b. Hypselodoris kanga Rudman, 1977. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypskang" title="Hypselodoris kanga Rudman, 1977" type="url" volumeTitle="Sea slug forum" year="1999">Rudman, 1999b</bibRefCitation>
</taxonomicName>
, in part.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA1FC82FCE6FAD36E6A" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="C90D4710FFFEFFA1FC82FCE6FC456932" blockId="28.[809,1426,842,988]" box="[809,979,842,866]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FC82FCE6FC456932" box="[809,979,842,866]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FC82FCE6FC456932" box="[809,979,842,866]" italics="true" pageId="28" pageNumber="29">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FC82FCC7FA04698C" blockId="28.[809,1426,842,988]" pageId="28" pageNumber="29">
<materialsCitation id="79DA4D4DFFFEFFA1FC82FCC7FA04698C" ID-GBIF-Occurrence="2235579078" collectingDate="2005-10-15" collectionCode="CASIZ" collectorName="T. Gosliner &amp; S. Fahey." country="Madagascar" latitude="-14.157833" location="Nosy Valiha" longLatPrecision="5" longitude="47.6485" municipality="Iles Radama" pageId="28" pageNumber="29" specimenCode="CASIZ 175550" specimenCount="1" typeStatus="holotype">
<typeStatus id="1609F9B2FFFEFFA1FC82FCC7FC07692F" box="[809,913,874,895]" pageId="28" pageNumber="29" type="holotype">
<emphasis id="FBC69B02FFFEFFA1FC82FCC7FC07692F" box="[809,913,874,895]" italics="true" pageId="28" pageNumber="29">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="9914EF6BFFFEFFA1FC0FFCC6FBC769D0" box="[932,1105,874,896]" collectionCode="CASIZ" pageId="28" pageNumber="29">CASIZ 175550</specimenCode>
, subsampled for molecular study, dissected, wall at south end of reef,
<geoCoordinate id="AC8621D7FFFEFFA1FB55FC24FA1A69CF" box="[1278,1420,905,927]" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="-14.157833">14.157833°S</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFFEFFA1FC82FC05FC2469EE" box="[809,946,936,958]" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="47.6485">47.6485°E</geoCoordinate>
, west of
<location id="CC6D11CBFFFEFFA1FB84FC05FB4469EE" LSID="urn:lsid:plazi:treatment:411BF606FFFEFFA3FC10FD80FD5168F8:CC6D11CBFFFEFFA1FB84FC05FB4469EE" box="[1071,1234,936,958]" country="Madagascar" latitude="-14.157833" longLatPrecision="5" longitude="47.6485" municipality="Iles Radama" name="Nosy Valiha" pageId="28" pageNumber="29">Nosy Valiha</location>
,
<collectingMunicipality id="2969DD6AFFFEFFA1FB48FC08FA1A69EE" box="[1251,1420,933,958]" pageId="28" pageNumber="29">Îles Radama</collectingMunicipality>
,
<collectingCountry id="B1A50780FFFEFFA1FC82FC6BFC25698C" box="[809,947,966,988]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
,
<collectingDate id="AD489838FFFEFFA1FC16FC6BFBE0698C" box="[957,1142,966,988]" pageId="28" pageNumber="29" value="2005-10-15">
<date id="BD0C61D0FFFEFFA1FC16FC6BFBE0698C" box="[957,1142,966,988]" pageId="28" pageNumber="29" value="2005-10-15">15 October 2005</date>
</collectingDate>
,
<collectorName id="644722C6FFFEFFA1FBD5FC6AFB6E698C" box="[1150,1272,966,988]" pageId="28" pageNumber="29">T. Gosliner</collectorName>
and
<collectorName id="644722C6FFFEFFA1FA85FC6AFA04698C" box="[1326,1426,966,988]" pageId="28" pageNumber="29">S. Fahey.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FC82FBA9FC526E4B" blockId="28.[809,1349,1027,1082]" box="[809,964,1027,1051]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FC82FBA9FC526E4B" box="[809,964,1027,1051]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FC82FBA9FC526E4B" box="[809,964,1027,1051]" italics="true" pageId="28" pageNumber="29">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FC82FB89FAD36E6A" blockId="28.[809,1349,1027,1082]" box="[809,1349,1057,1082]" pageId="28" pageNumber="29">
West of Nosy Valiha, Îles Radama,
<collectingCountry id="B1A50780FFFEFFA1FB1EFB89FAD76E6A" box="[1205,1345,1060,1082]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA1FC82FBCCFB466E84" pageId="28" pageNumber="29" type="distribution">
<paragraph id="C90D4710FFFEFFA1FC82FBCCFBF06E29" blockId="28.[809,1425,1121,1236]" box="[809,1126,1121,1145]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FC82FBCCFBF06E29" box="[809,1126,1121,1145]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FC82FBCCFBF06E29" box="[809,1126,1121,1145]" italics="true" pageId="28" pageNumber="29">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FC82FB2FFB466E84" blockId="28.[809,1425,1121,1236]" pageId="28" pageNumber="29">
Known from
<collectingCountry id="B1A50780FFFEFFA1FC16FB2CFBDE6EC7" box="[957,1096,1153,1175]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
(present study) and possibly
<collectingCountry id="B1A50780FFFEFFA1FC82FB0DFC0A6EE6" box="[809,924,1184,1206]" name="Sri Lanka" pageId="28" pageNumber="29">Sri Lanka</collectingCountry>
(
<bibRefCitation id="AD233AE1FFFEFFA1FC01FB0DFBDE6EE5" author="Houben A." box="[938,1096,1184,1206]" journalOrPublisher="Sydney: Australian Museum" pageId="28" pageNumber="29" publicationUrl="http://www.seaslugforum.net/find/20856" refId="ref42947" refString="Houben A. 2007. Hypselodoris cf. maculosa from Sri Lanka. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20856" title="Hypselodoris cf. maculosa from Sri Lanka" type="url" volumeTitle="Sea slug forum" year="2007">Houben, 2007</bibRefCitation>
),
<collectingCountry id="B1A50780FFFEFFA1FBF6FB0DFB7D6EE6" box="[1117,1259,1184,1206]" name="South Africa" pageId="28" pageNumber="29">South Africa</collectingCountry>
(
<bibRefCitation id="AD233AE1FFFEFFA1FB50FB0DFA1F6EE6" author="Ogden CM" box="[1275,1417,1184,1206]" journalOrPublisher="Sydney: Australian Museum" pageId="28" pageNumber="29" publicationUrl="http://www.seaslugforum.net/find/1485" refId="ref43821" refString="Ogden CM. 2005. Hypselodoris from South Africa. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 1485" title="Hypselodoris from South Africa" type="url" volumeTitle="Sea slug forum" year="2005">Ogden, 2005</bibRefCitation>
) and
<collectingCountry id="B1A50780FFFEFFA1FCF0FB13FC2C6E84" box="[859,954,1214,1236]" name="Réunion" pageId="28" pageNumber="29">Reunion</collectingCountry>
Island (
<bibRefCitation id="AD233AE1FFFEFFA1FBB3FB13FB566E84" author="Bidgrain P." box="[1048,1216,1214,1236]" journalOrPublisher="Sydney: Australian Museum" pageId="28" pageNumber="29" publicationUrl="http://www.seaslugforum.net/find/14854" refId="ref41470" refString="Bidgrain P. 2005. Variations in Hypselodoris cf. maculosa from Reunion Island. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 14854" title="Variations in Hypselodoris cf. maculosa from Reunion Island" type="url" volumeTitle="Sea slug forum" year="2005">Bidgrain, 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA1FC82FB56FAFE6FFC" pageId="28" pageNumber="29" type="etymology">
<paragraph id="C90D4710FFFEFFA1FC82FB56FC3C6F43" blockId="28.[809,1425,1275,1452]" box="[809,938,1275,1299]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FC82FB56FC3C6F43" box="[809,938,1275,1299]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FC82FB56FC3C6F43" box="[809,938,1275,1299]" italics="true" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA1FC82FAB1FAFE6FFC" blockId="28.[809,1425,1275,1452]" pageId="28" pageNumber="29">This small but striking species is named for author Rebecca Johnson’s daughter, Juniper Rodgers. Juniper, also one of a kind, loves bright colours and patterns, and when she was only 4 years old said, ‘Slugs are just like land nudibranchs’. This nudibranch is for you.</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFEFFA0FC82FA79FA0B6FEE" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="C90D4710FFFEFFA1FC82FA79FC236FBB" blockId="28.[809,1425,1492,1914]" box="[809,949,1492,1515]" pageId="28" pageNumber="29">
<heading id="9245F07CFFFEFFA1FC82FA79FC236FBB" box="[809,949,1492,1515]" fontSize="10" level="4" pageId="28" pageNumber="29" reason="1">
<emphasis id="FBC69B02FFFEFFA1FC82FA79FC236FBB" box="[809,949,1492,1515]" italics="true" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFEFFA0FC82FA59FD506F12" blockId="28.[809,1425,1492,1914]" lastBlockId="29.[163,779,1202,1346]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="FBC69B02FFFEFFA1FC82FA59FBAC6C59" box="[809,1082,1524,1545]" italics="true" pageId="28" pageNumber="29">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFFEFFA1FA81FA59FA1F6C59" box="[1322,1417,1524,1546]" captionStart="Figure 1" captionStartId="7.[161,241,1623,1645]" captionTargetBox="[225,1377,195,1582]" captionTargetId="figure@7.[225,1377,195,1582]" captionTargetPageId="7" captionText="Figure 1. Living animals. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype, Saudi Arabia. B, Hypselodoris brycei Gosliner &amp; Johnson sp. nov., WAM S96082, holotype, Montebello Islands, Western Australia, photograph by Nerida Wilson. C, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., specimen from Hachijo Island, Japan, Nishina Masayoshi. D, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070, Papua New Guinea, photograph by J. Goodheart. E, Hypseldoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282, Saudi Arabia. F, Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., CASIZ 177550, Radama Islands, Madagascar. G, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 176773, eastern Peninsular Malaysia. H, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 177532, Batangas, Luzon, Philippines. I, H. katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181300, Batangas, Luzon, Philippines. All photographs by T. Gosliner unless otherwise indicated." httpUri="https://zenodo.org/record/2632773/files/figure.png" pageId="28" pageNumber="29">Fig. 1F</figureCitation>
) small, reaching
<quantity id="0E4AEAF5FFFEFFA1FC42F9BEFBBA6C78" box="[1001,1068,1555,1576]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="mm" value="9.0">9 mm</quantity>
in length. Body long, slender, translucent pink, with four groups of small white tubercles arranged longitudinally in linear fashion. Two additional rows of submarginal spots. Deep purple spots found between areas of white spots, arranged in longitudinal rows. Centre of spots darker purple than outer portions. Broad orange band along lateral margins of notum, with deeper orange anteriorly and posteriorly in region of mantle glands. Anterior and posterior ends of the animal with purple areas. Similar pattern of lines and spots found on lateral margins of the body and foot. Five to six unipinnate gill branches having a translucent white base and inner margin and bright red–orange pigment on outer surface margin. Bulb of perfoliate rhinophores opaque white, with two red transverse bands and bearing ~11 densely arranged lamellae. Base of rhinophores translucent white.
</paragraph>
<caption id="9DCD1798FFFFFFA0FF08FBF7FC006EDD" httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="29" pageNumber="30" startId="29.[163,242,1114,1136]" targetBox="[227,1379,195,1073]" targetPageId="29">
<paragraph id="C90D4710FFFFFFA0FF08FBF7FC006EDD" blockId="29.[163,1442,1114,1165]" pageId="29" pageNumber="30">
<emphasis id="FBC69B02FFFFFFA0FF08FBF7FE8E6E3F" bold="true" box="[163,280,1114,1136]" pageId="29" pageNumber="30">Figure 16.</emphasis>
<taxonomicName id="0EB23C93FFFFFFA0FE88FBF7FC9E6E20" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,776,1114,1136]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="juniperae" status="sp. nov.">
<emphasis id="FBC69B02FFFFFFA0FE88FBF7FDA76E3F" bold="true" box="[291,561,1114,1136]" italics="true" pageId="29" pageNumber="30">Hypselodoris juniperae</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFFFFFA0FCA5FBF7FCF36E3F" box="[782,869,1114,1135]" pageId="29" pageNumber="30" rank="species">
<emphasis id="FBC69B02FFFFFFA0FCA5FBF7FCF36E3F" bold="true" box="[782,869,1114,1135]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFFFFFA0FF08FAC6FD936C4A" blockId="29.[163,779,1387,1562]" pageId="29" pageNumber="30">
<emphasis id="FBC69B02FFFFFFA0FF08FAC6FEDB6FD0" box="[163,333,1387,1408]" italics="true" pageId="29" pageNumber="30">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFFFFFA0FE60FA27FDB16FCF" box="[459,551,1418,1440]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="29" pageNumber="30">Fig. 2G</figureCitation>
). Glands situated anteriorly and posteriorly, with no glands present in the central lateral regions of body margin. Ten glands on either side of anterior end of the body, with arc of ten glands situated posteriorly.
</paragraph>
<paragraph id="C90D4710FFFFFFA0FF08F9EEFB616F2F" blockId="29.[163,779,1603,1901]" lastBlockId="29.[827,1443,1202,1408]" pageId="29" pageNumber="30">
<emphasis id="FBC69B02FFFFFFA0FF08F9EEFEFB6C09" box="[163,365,1603,1625]" italics="true" pageId="29" pageNumber="30">Buccal armature</emphasis>
: Muscular portion of buccal mass longer than oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFFFFFA0FD87F932FD036CE5" box="[556,661,1695,1717]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="29" pageNumber="30">Fig. 16A</figureCitation>
). Rodlets sharply angled with single, acutely pointed apex and posterolateral extensions. Radular formula of
<typeStatus id="1609F9B2FFFFFFA0FD01F971FC9D6CA2" box="[682,779,1756,1778]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
(CASIZ 175550) 33 × 19.0.19. Rachidian row of teeth absent (
<figureCitation id="51895B95FFFFFFA0FEACF8B7FEF96D7F" box="[263,367,1817,1840]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="29" pageNumber="30">Fig. 16B</figureCitation>
). Innermost lateral teeth having one large triangular denticle on inner side of bifid primary cusp, with another two outer denticles. Next several laterals lacking inner triangular denticle but possessing two denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFFFFFA0FBCFFB42FB5F6F55" box="[1124,1225,1263,1285]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="29" pageNumber="30">Fig. 16C</figureCitation>
) all lacking inner denticles but having two or three sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and shorter tooth shape, with three to five rounded outer denticles (
<figureCitation id="51895B95FFFFFFA0FB2CFAC7FB716F2F" box="[1159,1255,1385,1408]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="29" pageNumber="30">Fig. 16D</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFFFFFA0FC90FA05FA0B6FEE" blockId="29.[827,1437,1448,1470]" box="[827,1437,1448,1470]" pageId="29" pageNumber="30">
<emphasis id="FBC69B02FFFFFFA0FC90FA05FBB46FEE" box="[827,1058,1448,1470]" italics="true" pageId="29" pageNumber="30">Reproductive system</emphasis>
: Reproductive organs immature.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFFFFFA3FC90FA48FD5168F8" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="discussion">
<paragraph id="C90D4710FFFFFFA0FC90FA48FC3E6FAD" blockId="29.[827,1444,1509,1901]" box="[827,936,1509,1533]" pageId="29" pageNumber="30">
<heading id="9245F07CFFFFFFA0FC90FA48FC3E6FAD" box="[827,936,1509,1533]" fontSize="10" level="4" pageId="29" pageNumber="30" reason="1">
<emphasis id="FBC69B02FFFFFFA0FC90FA48FC3E6FAD" box="[827,936,1509,1533]" italics="true" pageId="29" pageNumber="30">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFFFFFA0FC90F9A8FB426D3D" blockId="29.[827,1444,1509,1901]" pageId="29" pageNumber="30">
<taxonomicName id="0EB23C93FFFFFFA0FC90F9A8FBD96C4B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1103,1541,1563]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFFFFA0FC90F9A8FBD96C4B" box="[827,1103,1541,1563]" italics="true" pageId="29" pageNumber="30">Hypselodoris juniperae</emphasis>
</taxonomicName>
is a member of a clade that includes
<taxonomicName id="0EB23C93FFFFFFA0FC09F988FBDE6C69" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[930,1096,1572,1594]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="alburtuqali">
<emphasis id="FBC69B02FFFFFFA0FC09F988FBDE6C69" box="[930,1096,1572,1594]" italics="true" pageId="29" pageNumber="30">H. alburtuqali</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFFFFFA0FBFEF988FB4F6C69" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1109,1241,1572,1594]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFFFFFA0FBFEF988FB4F6C69" box="[1109,1241,1572,1594]" italics="true" pageId="29" pageNumber="30">H. decorata</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFFFFFA0FABFF988FA356C69" authorityName="Rudman" authorityYear="1999" box="[1300,1443,1572,1594]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFFFFA0FABFF988FA356C69" box="[1300,1443,1572,1594]" italics="true" pageId="29" pageNumber="30">H. maculosa</emphasis>
</taxonomicName>
and is sister to
<emphasis id="FBC69B02FFFFFFA0FBA8F9E9FB336C08" box="[1027,1189,1603,1625]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="0EB23C93FFFFFFA0FBA8F9E9FB346C08" authorityName="Rudman" authorityYear="1999" box="[1027,1186,1603,1625]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="maculosa">H. maculosa</taxonomicName>
.
</emphasis>
The ABGD analysis indicates that
<taxonomicName id="0EB23C93FFFFFFA0FC4CF9CFFBEF6C27" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[999,1145,1634,1655]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFFFFA0FC4CF9CFFBEF6C27" box="[999,1145,1634,1655]" italics="true" pageId="29" pageNumber="30">H. juniperae</emphasis>
</taxonomicName>
from the western Indian Ocean is distinct from
<taxonomicName id="0EB23C93FFFFFFA0FBE8F92CFB426CC5" authorityName="Rudman" authorityYear="1999" box="[1091,1236,1664,1686]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFFFFA0FBE8F92CFB426CC5" box="[1091,1236,1664,1686]" italics="true" pageId="29" pageNumber="30">H. maculosa</emphasis>
</taxonomicName>
from the western Pacific and they are 6.5–7.2% divergent in their
<emphasis id="FBC69B02FFFFFFA0FAD8F932FA356CE4" box="[1395,1443,1695,1716]" italics="true" pageId="29" pageNumber="30">COI</emphasis>
gene, whereas the two western Pacific
<taxonomicName id="0EB23C93FFFFFFA0FAA5F913FA326C82" authorityName="Rudman" authorityYear="1999" box="[1294,1444,1725,1747]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFFFFA0FAA5F913FA326C82" box="[1294,1444,1725,1747]" italics="true" pageId="29" pageNumber="30">H. maculosa</emphasis>
</taxonomicName>
are 2.7% divergent from each other. The colour pattern of
<taxonomicName id="0EB23C93FFFFFFA0FCF3F951FC7F6D40" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[856,1001,1787,1809]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFFFFA0FCF3F951FC7F6D40" box="[856,1001,1787,1809]" italics="true" pageId="29" pageNumber="30">H. juniperae</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0EB23C93FFFFFFA0FB77F951FAFB6D40" authorityName="Rudman" authorityYear="1999" box="[1244,1389,1787,1809]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFFFFA0FB77F951FAFB6D40" box="[1244,1389,1787,1809]" italics="true" pageId="29" pageNumber="30">H. maculosa</emphasis>
</taxonomicName>
, but differs in having white spots arranged in lines rather than thin continuous lines (
<bibRefCitation id="AD233AE1FFFFFFA0FB26F895FAD76D1E" author="Bidgrain P." box="[1165,1345,1848,1870]" journalOrPublisher="Sydney: Australian Museum" pageId="29" pageNumber="30" publicationUrl="http://www.seaslugforum.net/find/14854" refId="ref41470" refString="Bidgrain P. 2005. Variations in Hypselodoris cf. maculosa from Reunion Island. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 14854" title="Variations in Hypselodoris cf. maculosa from Reunion Island" type="url" volumeTitle="Sea slug forum" year="2005">Bidgrain, 2005</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFFFFFA0FAE4F895FCE46D3D" author="Ogden CM" journalOrPublisher="Sydney: Australian Museum" pageId="29" pageNumber="30" publicationUrl="http://www.seaslugforum.net/find/1485" refId="ref43821" refString="Ogden CM. 2005. Hypselodoris from South Africa. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 1485" title="Hypselodoris from South Africa" type="url" volumeTitle="Sea slug forum" year="2005">Ogden, 2005</bibRefCitation>
;
<bibRefCitation id="AD233AE1FFFFFFA0FCD5F8FAFB8D6D3C" author="Houben A." box="[894,1051,1879,1901]" journalOrPublisher="Sydney: Australian Museum" pageId="29" pageNumber="30" publicationUrl="http://www.seaslugforum.net/find/20856" refId="ref42947" refString="Houben A. 2007. Hypselodoris cf. maculosa from Sri Lanka. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20856" title="Hypselodoris cf. maculosa from Sri Lanka" type="url" volumeTitle="Sea slug forum" year="2007">Houben, 2007</bibRefCitation>
; present study).
</paragraph>
<paragraph id="C90D4710FFFCFFA3FF02FF6BFD5168F8" blockId="30.[145,761,198,680]" pageId="30" pageNumber="31">
There appear to be fewer mantle glands in
<taxonomicName id="0EB23C93FFFCFFA3FF3AFF4BFE8D6AAB" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,283,230,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFCFFA3FF3AFF4BFE8D6AAB" box="[145,283,230,251]" italics="true" pageId="30" pageNumber="31">H. juniperae</emphasis>
</taxonomicName>
than in
<taxonomicName id="0EB23C93FFFCFFA3FED0FF4BFD906AAA" authorityName="Rudman" authorityYear="1999" box="[379,518,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFCFFA3FED0FF4BFD906AAA" box="[379,518,229,251]" italics="true" pageId="30" pageNumber="31">H. maculosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFFCFFA3FDBEFF48FE696B4A" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="30" pageNumber="31" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999: fig. 29D; present study</bibRefCitation>
). The jaw rodlets of
<taxonomicName id="0EB23C93FFFCFFA3FF3AFE8EFEBF6B68" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,297,291,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFCFFA3FF3AFE8EFEBF6B68" box="[145,297,291,312]" italics="true" pageId="30" pageNumber="31">H. juniperae</emphasis>
</taxonomicName>
have a shorter cusp than do those of
<taxonomicName id="0EB23C93FFFCFFA3FF3AFEEFFEB66B06" authorityName="Rudman" authorityYear="1999" box="[145,288,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFCFFA3FF3AFEEFFEB66B06" box="[145,288,321,343]" italics="true" pageId="30" pageNumber="31">H. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFFCFFA3FE84FEECFE036B06" box="[303,405,321,343]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="30" pageNumber="31">Figs 11B</figureCitation>
,
<figureCitation id="51895B95FFFCFFA3FE09FEECFE466B07" box="[418,464,321,343]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="30" pageNumber="31">16A</figureCitation>
;
<bibRefCitation id="AD233AE1FFFCFFA3FE76FEECFD126B07" author="Rudman WB" box="[477,644,321,343]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="30" pageNumber="31" pagination="309 - 353" part="86" refId="ref44765" refString="Rudman WB. 1986. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups. Zoological Journal of the Linnean Society 86: 309 - 353." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups" type="journal article" year="1986">Rudman, 1986</bibRefCitation>
: fig. 16E). The radula of
<taxonomicName id="0EB23C93FFFCFFA3FEEBFECDFE416B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[320,471,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFCFFA3FEEBFECDFE416B25" box="[320,471,352,373]" italics="true" pageId="30" pageNumber="31">H. juniperae</emphasis>
</taxonomicName>
is much narrower than that of
<taxonomicName id="0EB23C93FFFCFFA3FF49FED2FEF96BC3" authorityName="Rudman" authorityYear="1999" box="[226,367,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFCFFA3FF49FED2FEF96BC3" box="[226,367,382,404]" italics="true" pageId="30" pageNumber="31">H. maculosa</emphasis>
</taxonomicName>
, with only 19 teeth per half row as compared with 28–51 teeth per half row in
<taxonomicName id="0EB23C93FFFCFFA3FDC5FE33FED06B81" authority="(Rudman, 1986)" baseAuthorityName="Rudman" baseAuthorityYear="1986" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFCFFA3FDC5FE33FD6F6BE2" box="[622,761,413,435]" italics="true" pageId="30" pageNumber="31">H. maculosa</emphasis>
(
<bibRefCitation id="AD233AE1FFFCFFA3FF32FE11FEAA6B82" author="Rudman WB" box="[153,316,444,466]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="30" pageNumber="31" pagination="309 - 353" part="86" refId="ref44765" refString="Rudman WB. 1986. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups. Zoological Journal of the Linnean Society 86: 309 - 353." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups" type="journal article" year="1986">Rudman, 1986</bibRefCitation>
)
</taxonomicName>
. The inner lateral teeth of
<taxonomicName id="0EB23C93FFFCFFA3FDC5FE11FD6F6B81" baseAuthorityName="Rudman" baseAuthorityYear="1986" box="[622,761,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFFCFFA3FDC5FE11FD6F6B81" box="[622,761,444,465]" italics="true" pageId="30" pageNumber="31">H. maculosa</emphasis>
</taxonomicName>
have two or three denticles on the inner side of the bifid cusps and an additional two denticles on the outer side of the cusps (
<figureCitation id="51895B95FFFCFFA3FEF2FDB5FE2E687D" box="[345,440,536,558]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="30" pageNumber="31">Fig. 11D</figureCitation>
;
<bibRefCitation id="AD233AE1FFFCFFA3FE6EFDB5FDFF687E" author="Rudman WB" box="[453,617,536,558]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="30" pageNumber="31" pagination="309 - 353" part="86" refId="ref44765" refString="Rudman WB. 1986. The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups. Zoological Journal of the Linnean Society 86: 309 - 353." title="The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: Noumea purpurea and Chromodoris decora colour groups" type="journal article" year="1986">Rudman, 1986</bibRefCitation>
: fig. 16A, B), whereas in
<taxonomicName id="0EB23C93FFFCFFA3FEB0FD9AFE39681C" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[283,431,567,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFCFFA3FEB0FD9AFE39681C" box="[283,431,567,588]" italics="true" pageId="30" pageNumber="31">H. juniperae</emphasis>
</taxonomicName>
there is only a single inner denticle and one or two outer ones (
<figureCitation id="51895B95FFFCFFA3FDE6FDF8FD25683B" box="[589,691,597,619]" captionStart="Figure 16" captionStartId="29.[163,242,1114,1136]" captionTargetBox="[227,1379,195,1073]" captionTargetId="figure@29.[227,1379,195,1073]" captionTargetPageId="29" captionText="Figure 16. Hypselodoris juniperae Gosliner &amp; Johnson sp. nov., buccal armature, holotype, CASIZ 175550. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632806/files/figure.png" pageId="30" pageNumber="31">Fig. 16B</figureCitation>
). The reproductive systems cannot be compared owing to the immaturity of the single specimen of
<taxonomicName id="0EB23C93FFFCFFA3FD9EFD3EFD5668F8" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[565,704,659,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="juniperae">
<emphasis id="FBC69B02FFFCFFA3FD9EFD3EFD5668F8" box="[565,704,659,680]" italics="true" pageId="30" pageNumber="31">H. juniperae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>