Add updates up until 2025-04-15 13:22:50
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<mods:affiliation id="416E06FA6928726136BBB0BC19D924BA">Universidad de El Salvador, El Salvador, Ciudad Universitaria, San Salvador, El Salvador. C. P. 01101 & Centro de Investigación Marina y Limnológica, Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101. & Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101.</mods:affiliation>
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<mods:affiliation id="DA17C8509DCAD3DE7725626258A540C0">Universidad de El Salvador, El Salvador, Ciudad Universitaria, San Salvador, El Salvador. C. P. 01101 & Centro de Investigación Marina y Limnológica, Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101. & Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101.</mods:affiliation>
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<mods:affiliation id="E51E1849483862DBBE65605A83DF6B41">Centro de Investigación Marina y Limnológica, Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101. & Universidad Francisco Gavidia, San Salvador, El Salvador. C. P. 01101.</mods:affiliation>
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MUHNES-91-43—El Arco (
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, A. Trejo (
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.
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@ -1,63 +1,65 @@
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,
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MUHNES-91-52—El Candado (
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<elevation id="66C7D1E2FF9DFFD869EBFBDA30F40E41" box="[1220,1269,1124,1151]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="106" unit="m" value="6.0">6 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="401F5307FF9DFFD8682BFBDB31640E40" box="[1284,1381,1124,1151]" pageId="7" pageNumber="106">A. Trejo</collectorName>
|
||||
(
|
||||
<date id="99541011FF9DFFD86856FBDA34F90E9C" pageId="7" pageNumber="106" value="2018-06-19">
|
||||
<collectingDate id="8910E9F9FF9DFFD86856FBDA34F90E9C" pageId="7" pageNumber="106" value="2018-06-19">19. VI.2018</collectingDate>
|
||||
</date>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="A5F0655AFF9DFFD76DE8FB1230020A8F" lastPageId="8" lastPageNumber="107" pageId="7" pageNumber="106" type="description">
|
||||
|
|
@ -112,7 +125,7 @@ in diameter (
|
|||
<figureCitation id="75D12A54FF9DFFD86FBEFA5037260C36" box="[657,807,1517,1544]" captionStart="FIGURE 5" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[194,1393,561,1890]" captionTargetId="figure-187@8.[194,1393,561,1890]" captionTargetPageId="8" captionText="FIGURE 5. Endectyon (Endectyon) hyle. A. Collected specimen at 6 m depth. B. Ectosome with styles projecting towards the surface. C. Choanosome. D. Styles from the ectosome. E. Styles from the choanosome. F. Acanthostyles. Scale bars: A, 1 cm; B, 150 µm; C, 300 µm; D–F, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.15216246" httpUri="https://zenodo.org/record/15216246/files/figure.png" pageId="7" pageNumber="106">Figure 5D–F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="B9956659FF9DFFD86DB8F98D369B0C4E" pageId="7" pageNumber="106" startId="7.[151,239,1587,1611]" targetType="table">
|
||||
<caption id="B9956659FF9DFFD86DB8F98D369B0C4E" ID-Table-UUID="B9956659FF9DFFD86DB8F98D369B0C4E" httpUri="http://table.plazi.org/id/B9956659FF9DFFD86DB8F98D369B0C4E" pageId="7" pageNumber="106" startId="7.[151,239,1587,1611]" targetBox="[163,1420,1663,2029]" targetIsTable="true" targetPageId="7" targetType="table">
|
||||
<paragraph id="ED5536D1FF9DFFD86DB8F98D369B0C4E" blockId="7.[151,1437,1587,1648]" pageId="7" pageNumber="106">
|
||||
<emphasis id="DF9EEAC3FF9DFFD86DB8F98D350B0C72" bold="true" box="[151,266,1587,1612]" pageId="7" pageNumber="106">TABLE 5.</emphasis>
|
||||
Comparative data on spicule measurement and known distribution of
|
||||
|
|
@ -166,7 +179,7 @@ De
|
|||
<taxonomicName id="2AEA4D52FF9DFFD86D8CF8DB34E80D42" authorityName="De Laubenfels" authorityYear="1932" box="[163,233,1892,1916]" class="Demospongiae" family="Raspailiidae" genus="Endectyon" kingdom="Animalia" order="Axinellida" pageId="7" pageNumber="106" phylum="Porifera" rank="species" species="hyle">
|
||||
<emphasis id="DF9EEAC3FF9DFFD86D8CF8DB34E80D42" box="[163,233,1892,1916]" italics="true" pageId="7" pageNumber="106">E. hyle</emphasis>
|
||||
</taxonomicName>
|
||||
Aguilar- Camacho
|
||||
Aguilar-Camacho
|
||||
<emphasis id="DF9EEAC3FF9DFFD86C26F83735340D9E" box="[265,309,1928,1952]" italics="true" pageId="7" pageNumber="106">et al</emphasis>
|
||||
. 2013
|
||||
</th>
|
||||
|
|
@ -208,7 +221,7 @@ The species was described by de Laubenfels in 1930 from California. Reports have
|
|||
, the species has been recorded in Punta Amapala (
|
||||
<bibRefCitation id="897B4B20FF92FFD76871FEBD35530B7F" author="Trejo, A. & Segovia, J." pageId="8" pageNumber="107" pagination="449 - 463" refId="ref15211" refString="Trejo, A. & Segovia, J. (2024) Marine sponges of the rocky reefs of Punta Amapala, El Salvador, Eastern Tropical Pacific. Marine and Fishery Sciences, 37 (3), 449-463. https://doi.org/10.47193/mafis.37X2024010105" type="journal article" year="2024">Trejo & Segovia 2024</bibRefCitation>
|
||||
). Here we extend the species distribution in the country to Los Cóbanos (
|
||||
<tableCitation id="A068036AFF92FFD769AAFE9930EA0B7F" box="[1157,1259,295,321]" captionStart="TABLE 12" captionStartId="19.[151,239,548,572]" captionText="TABLE 12. Species of freshwater and marine sponges reported from El Salvador. *= reported in this study." pageId="8" pageNumber="107">Table 12</tableCitation>
|
||||
<tableCitation id="A068036AFF92FFD769AAFE9930EA0B7F" box="[1157,1259,295,321]" captionStart="TABLE 12" captionStartId="19.[151,239,548,572]" captionTargetPageId="19" captionText="TABLE 12. Species of freshwater and marine sponges reported from El Salvador. *= reported in this study." httpUri="http://table.plazi.org/id/B9956659FF89FFCC6DB8FD9A31120802" pageId="8" pageNumber="107" tableUuid="B9956659FF89FFCC6DB8FD9A31120802">Table 12</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
471
data/8C/4A/57/8C4A57289D60C424538DFD2DFADFF622.xml
Normal file
471
data/8C/4A/57/8C4A57289D60C424538DFD2DFADFF622.xml
Normal file
|
|
@ -0,0 +1,471 @@
|
|||
<document id="AF96867ADE354951190E34E3238BC5B6" ID-DOI="10.11646/zootaxa.5613.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="15216327" ID-ZooBank="76A3CD48-D4A3-422A-BBBC-1165F0FFEF02" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1744667614183" checkinUser="plazi" docAuthor="Bilton, David T., Mlambo, Musa C. & Balke, Michael" docDate="2025" docId="8C4A57289D60C424538DFD2DFADFF622" docLanguage="en" docName="zootaxa.5613.1.8.pdf" docOrigin="Zootaxa 5613 (1)" docSource="https://doi.org/10.11646/zootaxa.5613.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anacaena fosteri Bilton, Mlambo & Balke, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="168" masterDocId="70732F509D61C420531AFFA2FFB4F763" masterDocTitle="A new species of Anacaena Thomson, 1859 from the Cederberg Range, South Africa (Coleoptera: Hydrophilidae)" masterLastPageNumber="170" masterPageNumber="165" pageNumber="166" updateTime="1744723189982" updateUser="felipe" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="70C6F6497B72705A5DD2C4A816F11C32" xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo id="92144D5051EF27E1D0FC6B2AD2C818C3">
|
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<mods:title id="CB00CADC6DADA6C859D8C39CA09C6F7A">A new species of Anacaena Thomson, 1859 from the Cederberg Range, South Africa (Coleoptera: Hydrophilidae)</mods:title>
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</mods:titleInfo>
|
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<mods:name id="66592A5D996F2D8E8A9F63EB111E637C" type="personal">
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<mods:role id="E1038430A7AD6A0A8625C67DAF0E7973">
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<mods:namePart id="0BC31BCC9F56E02F51B0E5CAEE33B14D">Bilton, David T.</mods:namePart>
|
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<mods:affiliation id="7A4B83CF0F27B5772DC4BD587BD96F88">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK & Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa</mods:affiliation>
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</mods:name>
|
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<mods:name id="56FB6F4354672BE793D2FB729C99C39E" type="personal">
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<mods:role id="553C4CEDF4828172E9A0D59AD4CBDE99">
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<mods:roleTerm id="503F1213FB555C1207B5664B8A7593B4">Author</mods:roleTerm>
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</mods:role>
|
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<mods:namePart id="2D8332F6879F3A8F2E71C41CFCAE9BA4">Mlambo, Musa C.</mods:namePart>
|
||||
<mods:affiliation id="00629D4C4C9498ACC356E395104B6C87">Department of Freshwater Invertebrates, Albany Museum; South African Institute for Aquatic Biodiversity and Department of Zoology and Entomology, Rhodes University, Makhanda (formerly Grahamstown) 6140, South Africa</mods:affiliation>
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</mods:name>
|
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</mods:role>
|
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<mods:namePart id="3986435926C6582783181D9A65A9A5B7">Balke, Michael</mods:namePart>
|
||||
<mods:affiliation id="7C0512D2A276B4BE5F1D4AB833F44560">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, D- 81247, Munich, Germany</mods:affiliation>
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<mods:title id="EE42151C080F12B31531F0A8A285919F">Zootaxa</mods:title>
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<mods:part id="6C03FF96CDAE1D2B6EDD10ED64825974">
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<mods:date id="0B05D3B816FA9FB8D457986F03586DEB">2025</mods:date>
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<mods:number id="FA2C38656F4304C44BA44D42BE9B36A5">2025-03-26</mods:number>
|
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|
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<mods:detail id="543E24D1E4E6FC072E94EE22FC3B6DA7" type="volume">
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|
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<mods:detail id="156FED54DAF71296B46BABB3B3AC9DDB" type="issue">
|
||||
<mods:number id="3E51757A73C8AD8642230E11BA2ABAE0">1</mods:number>
|
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</mods:detail>
|
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<mods:extent id="C9DA4DE566419ABBC2F53B485E0AB35B" unit="page">
|
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<mods:start id="EBD981B36E8D13058DA34EEF99B28909">165</mods:start>
|
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<mods:end id="5F167B867B180C67D082130FACA915AA">170</mods:end>
|
||||
</mods:extent>
|
||||
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|
||||
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|
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<mods:location id="C376AA7ECEAB500D38C1922B94D53DE4">
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<mods:url id="C510777BB5F258CDED88DB21052874A5">https://doi.org/10.11646/zootaxa.5613.1.8</mods:url>
|
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</mods:location>
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<mods:classification id="CB7644BB3636FFAF35C6FFFAAFF9488F">journal article</mods:classification>
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<mods:identifier id="F95EA221662286057C968D337A9AA321" type="DOI">10.11646/zootaxa.5613.1.8</mods:identifier>
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<mods:identifier id="2E671AD7900F48D356C89062072E9D31" type="ZooBank">76A3CD48-D4A3-422A-BBBC-1165F0FFEF02</mods:identifier>
|
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</mods:mods>
|
||||
<treatment id="8C4A57289D60C424538DFD2DFADFF622" LSID="urn:lsid:plazi:treatment:8C4A57289D60C424538DFD2DFADFF622" httpUri="http://treatment.plazi.org/id/8C4A57289D60C424538DFD2DFADFF622" lastPageId="4" lastPageNumber="168" pageId="1" pageNumber="166">
|
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<subSubSection id="4CF9B5B59D60C421538DFD2DFE7EF5CA" box="[151,458,655,681]" pageId="1" pageNumber="166" type="nomenclature">
|
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<paragraph id="045CE63E9D60C421538DFD2DFE7EF5CA" blockId="1.[151,458,655,717]" box="[151,458,655,681]" pageId="1" pageNumber="166">
|
||||
<heading id="5F1451529D60C421538DFD2DFE7EF5CA" bold="true" box="[151,458,655,681]" fontSize="11" level="1" pageId="1" pageNumber="166" reason="1">
|
||||
<emphasis id="36973A2C9D60C421538DFD2DFE7EF5CA" bold="true" box="[151,458,655,681]" pageId="1" pageNumber="166">
|
||||
<taxonomicName id="C3E39DBD9D60C421538DFD2DFED2F5CA" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[151,358,655,681]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D60C421538DFD2DFED2F5CA" bold="true" box="[151,358,655,681]" italics="true" pageId="1" pageNumber="166">Anacaena fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="2DA487579D60C4215277FD2DFE7EF5CA" box="[365,458,655,681]" pageId="1" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D60C421538DFD11FF5FF5AE" box="[151,235,691,717]" pageId="1" pageNumber="166" type="description">
|
||||
<paragraph id="045CE63E9D60C421538DFD11FF5FF5AE" blockId="1.[151,458,655,717]" box="[151,235,691,717]" pageId="1" pageNumber="166">
|
||||
(
|
||||
<figureCitation id="9CD8FABB9D60C4215385FD11FF54F5AE" box="[159,224,691,717]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D60C421538DFD59FEF3F48E" pageId="1" pageNumber="166" type="materials_examined">
|
||||
<paragraph id="045CE63E9D60C421538DFD59FC6CF45A" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
<emphasis id="36973A2C9D60C421538DFD59FE9BF476" bold="true" box="[151,303,763,789]" pageId="1" pageNumber="166">Type locality</emphasis>
|
||||
.
|
||||
<emphasis id="36973A2C9D60C4215226FD59FE66F476" bold="true" box="[316,466,763,789]" pageId="1" pageNumber="166">
|
||||
<collectingCountry id="7CF4A6AE9D60C4215226FD59FE66F476" box="[316,466,763,789]" name="South Africa" pageId="1" pageNumber="166">South Africa</collectingCountry>
|
||||
</emphasis>
|
||||
,
|
||||
<collectingRegion id="C62728DC9D60C42152C5FD5EFD51F476" box="[479,741,763,790]" country="South Africa" name="Western Cape" pageId="1" pageNumber="166">Western Cape Province</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="ED3D9EB29D60C42151EAFD59FCD3F476" box="[752,871,763,789]" pageId="1" pageNumber="166">Cederberg</collectingCounty>
|
||||
,
|
||||
<collectingMunicipality id="E4387C449D60C421506BFD59FC7BF476" box="[881,975,763,789]" pageId="1" pageNumber="166">Die Vlei</collectingMunicipality>
|
||||
along headwaters of
|
||||
<location id="013CB0E59D60C42157A0FD59FADCF476" LSID="urn:lsid:plazi:treatment:8C4A57289D60C424538DFD2DFADFF622:013CB0E59D60C42157A0FD59FADCF476" box="[1210,1384,763,789]" country="South Africa" county="Cederberg" latitude="-32.417057" longLatPrecision="1" longitude="19.024973" municipality="Die Vlei" name="Driehoek River" pageId="1" pageNumber="166" stateProvince="Western Cape">Driehoek River</location>
|
||||
,
|
||||
<quantity id="C31B4BDB9D60C4215668FD59FF1BF459" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="1" pageNumber="166" unit="m" value="943.0">
|
||||
<elevation id="8FCE010D9D60C4215668FD59FF1BF459" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="1" pageNumber="166" unit="m" value="943.0">943 m</elevation>
|
||||
</quantity>
|
||||
, waterlogged algal covered peat,
|
||||
<geoCoordinate id="61D780F99D60C421513CFCBDFD74F459" box="[550,704,799,826]" degrees="32" direction="south" minutes="25" orientation="latitude" pageId="1" pageNumber="166" precision="1" seconds="01.4" value="-32.417057">32°25’01.4”S</geoCoordinate>
|
||||
<geoCoordinate id="61D780F99D60C42151DDFCBDFCD6F45A" box="[711,866,799,825]" degrees="19" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="166" precision="1" seconds="29.9" value="19.024973">19°01’29.9”E</geoCoordinate>
|
||||
(
|
||||
<figureCitation id="9CD8FABB9D60C421506BFCBDFC7EF45A" box="[881,970,799,825]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="1" pageNumber="166">Fig. 1D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDFCE1FDC0F4C6" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
<emphasis id="36973A2C9D60C42153DDFCE1FEDBF43E" bold="true" box="[199,367,835,861]" pageId="1" pageNumber="166">Type material</emphasis>
|
||||
.
|
||||
<materialsCitation id="B48BEC639D60C4215265FCE1FDC0F4C6" collectingDate="2024-09-28" collectionCode="AMG" collectorName="Bilton & Balke" country="South Africa" county="Driehoek River" elevation="943" latitude="-32.4167" location="Cederberg" longLatPrecision="1" longitude="19.12518" municipality="Die Vlei" pageId="1" pageNumber="166" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="36973A2C9D60C4215265FCE1FE58F43E" bold="true" box="[383,492,835,861]" pageId="1" pageNumber="166">
|
||||
<typeStatus id="DB58589C9D60C4215265FCE1FE58F43E" box="[383,492,835,861]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(male): “
|
||||
<date id="705DC0FE9D60C4215142FCE1FD61F43E" box="[600,725,835,861]" pageId="1" pageNumber="166" value="2024-09-28">
|
||||
<collectingDate id="601939169D60C4215142FCE1FD61F43E" box="[600,725,835,861]" pageId="1" pageNumber="166" value="2024-09-28">28/ix/2024</collectingDate>
|
||||
</date>
|
||||
<collectingCountry id="7CF4A6AE9D60C42151C7FCE1FCD9F43E" box="[733,877,835,861]" name="South Africa" pageId="1" pageNumber="166">South Africa</collectingCountry>
|
||||
WC//
|
||||
<location id="013CB0E59D60C42150A0FCE1FB85F43E" LSID="urn:lsid:plazi:treatment:8C4A57289D60C424538DFD2DFADFF622:013CB0E59D60C42150A0FCE1FB85F43E" box="[954,1073,835,861]" country="South Africa" county="Driehoek River" latitude="-32.4167" longLatPrecision="1" longitude="19.12518" municipality="Die Vlei" name="Cederberg" pageId="1" pageNumber="166">Cederberg</location>
|
||||
,
|
||||
<collectingMunicipality id="E4387C449D60C4215724FCE1FB2BF43E" box="[1086,1183,835,861]" pageId="1" pageNumber="166">Die Vlei</collectingMunicipality>
|
||||
along// headwaters of
|
||||
<collectingCounty id="ED3D9EB29D60C421538DFCC5FEF0F4E2" box="[151,324,871,897]" pageId="1" pageNumber="166">Driehoek River</collectingCounty>
|
||||
//
|
||||
<geoCoordinate id="61D780F99D60C4215247FCC5FE5DF4E2" box="[349,489,871,897]" degrees="32,41670" direction="south" orientation="latitude" pageId="1" pageNumber="166" precision="1" value="-32.4167">32,41670° S</geoCoordinate>
|
||||
<geoCoordinate id="61D780F99D60C42152EAFCC5FDCBF4E1" box="[496,639,871,898]" degrees="19,12518" direction="east" orientation="longitude" pageId="1" pageNumber="166" precision="1" value="19.12518">19,12518° E</geoCoordinate>
|
||||
,
|
||||
<quantity id="C31B4BDB9D60C4215196FCC5FD67F4E2" box="[652,723,871,897]" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="1" pageNumber="166" unit="m" value="943.0">
|
||||
<elevation id="8FCE010D9D60C4215196FCC5FD67F4E2" box="[652,723,871,897]" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="1" pageNumber="166" unit="m" value="943.0">943 m</elevation>
|
||||
</quantity>
|
||||
//
|
||||
<collectorName id="A91683E89D60C42151F1FCC5FC80F4E2" box="[747,820,871,897]" pageId="1" pageNumber="166">Bilton</collectorName>
|
||||
,
|
||||
<collectorName id="A91683E89D60C4215025FCC5FC34F4E2" box="[831,896,871,897]" pageId="1" pageNumber="166">Balke</collectorName>
|
||||
<emphasis id="36973A2C9D60C4215093FCCAFC76F4E2" box="[905,962,871,897]" italics="true" pageId="1" pageNumber="166">et al.</emphasis>
|
||||
leg.” (genitalia extracted and mounted on same card) and red holotype label (
|
||||
<collectionCode id="62F27EFB9D60C421513BFC29FDDDF4C6" box="[545,617,907,933]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="1" pageNumber="166">AMG</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDFC0DFEF3F48E" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
<materialsCitation id="B48BEC639D60C42153DDFC0DFEF0F48E" collectingDate="2024-09-28" collectionCode="AMG, CDTB, CFA, ZSM" collectorName="Bilton & Balke" country="South Africa" county="Driehoek River" elevation="943" latitude="-32.4167" location="South Africa" longLatPrecision="1" longitude="19.12518" municipality="Die Vlei" pageId="1" pageNumber="166" specimenCount="19" specimenCount-female="10" specimenCount-male="9" typeStatus="paratype">
|
||||
<emphasis id="36973A2C9D60C42153DDFC0DFE8BF4AA" bold="true" box="[199,319,943,969]" pageId="1" pageNumber="166">
|
||||
<typeStatus id="DB58589C9D60C42153DDFC0DFE8BF4AA" box="[199,319,943,969]" pageId="1" pageNumber="166" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
(19):
|
||||
<emphasis id="36973A2C9D60C4215293FC0DFD97F4AA" bold="true" box="[393,547,943,969]" pageId="1" pageNumber="166">
|
||||
<collectingCountry id="7CF4A6AE9D60C4215293FC0DFD97F4AA" box="[393,547,943,969]" name="South Africa" pageId="1" pageNumber="166">South Africa</collectingCountry>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount id="12E52DB79D60C421512FFC0DFDD0F4A9" box="[565,612,943,970]" count="9" pageId="1" pageNumber="166" type="male">9 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="12E52DB79D60C4215169FC0DFD1AF4A9" box="[627,686,943,970]" count="10" pageId="1" pageNumber="166" type="female">10 ♀</specimenCount>
|
||||
same data as holotype (
|
||||
<collectionCode id="62F27EFB9D60C42150D2FC0DFBBAF4AA" box="[968,1038,943,969]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="1" pageNumber="166">AMG</collectionCode>
|
||||
,
|
||||
<collectionCode id="62F27EFB9D60C4215706FC0DFBDDF4AA" box="[1052,1129,943,969]" pageId="1" pageNumber="166">CDTB</collectionCode>
|
||||
,
|
||||
<collectionCode id="62F27EFB9D60C4215762FC0DFB1BF4AA" box="[1144,1199,943,969]" pageId="1" pageNumber="166">CFA</collectionCode>
|
||||
,
|
||||
<collectionCode id="62F27EFB9D60C42157A7FC0DFB4FF4AA" box="[1213,1275,943,969]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="166" type="Museum">ZSM</collectionCode>
|
||||
). All with red paratype labels
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D60C42353DDFC55FAFBF592" lastPageId="3" lastPageNumber="168" pageId="1" pageNumber="166" type="description">
|
||||
<paragraph id="045CE63E9D60C42153DDFC55FAF1F355" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
<emphasis id="36973A2C9D60C42153DDFC55FEE5F372" bold="true" box="[199,337,1015,1041]" pageId="1" pageNumber="166">Description</emphasis>
|
||||
. Size:
|
||||
<typeStatus id="DB58589C9D60C42152BBFC55FDBAF372" box="[417,526,1015,1041]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
|
||||
: TL
|
||||
<quantity id="C31B4BDB9D60C421515BFC55FD1DF371" box="[577,681,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="1" pageNumber="166" unit="mm" value="2.65">2.65 mm</quantity>
|
||||
; EL
|
||||
<quantity id="C31B4BDB9D60C42151C5FC55FCF3F371" box="[735,839,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="166" unit="mm" value="1.85">1.85 mm</quantity>
|
||||
; EW
|
||||
<quantity id="C31B4BDB9D60C421509CFC55FC5AF371" box="[902,1006,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="166" unit="mm" value="1.6">1.60 mm</quantity>
|
||||
; EI 1.16.
|
||||
<typeStatus id="DB58589C9D60C4215747FC5AFB7AF371" box="[1117,1230,1016,1042]" pageId="1" pageNumber="166" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
<specimenCount id="12E52DB79D60C42157C0FC5AFB44F371" box="[1242,1264,1016,1042]" count="1" pageId="1" pageNumber="166" type="male">♂</specimenCount>
|
||||
s BL
|
||||
<quantity id="C31B4BDB9D60C4215636FC55FF72F355" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.55" metricValueMin="2.35" pageId="1" pageNumber="166" unit="mm" value="2.45" valueMax="2.55" valueMin="2.35">2.35–2.55 mm</quantity>
|
||||
; EL
|
||||
<quantity id="C31B4BDB9D60C42153E0FBB9FE14F355" box="[250,416,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.7" metricValueMin="1.55" pageId="1" pageNumber="166" unit="mm" value="1.625" valueMax="1.7" valueMin="1.55">1.55–1.70 mm</quantity>
|
||||
; EW
|
||||
<quantity id="C31B4BDB9D60C42152C7FBB9FD37F355" box="[477,643,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.5" metricValueMin="1.45" pageId="1" pageNumber="166" unit="mm" value="1.475" valueMax="1.5" valueMin="1.45">1.45–1.50 mm</quantity>
|
||||
.
|
||||
<specimenCount id="12E52DB79D60C4215194FBBEFD15F355" box="[654,673,1052,1078]" count="1" pageId="1" pageNumber="166" type="female">♀</specimenCount>
|
||||
s BL
|
||||
<quantity id="C31B4BDB9D60C42151C4FBB9FC30F355" box="[734,900,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="2.775" metricValueMax="2.95" metricValueMin="2.6" pageId="1" pageNumber="166" unit="mm" value="2.775" valueMax="2.95" valueMin="2.6">2.60–2.95 mm</quantity>
|
||||
; EL
|
||||
<quantity id="C31B4BDB9D60C42150A2FBB9FBEAF355" box="[952,1118,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.95" metricValueMin="1.75" pageId="1" pageNumber="166" unit="mm" value="1.85" valueMax="1.95" valueMin="1.75">1.75–1.95 mm</quantity>
|
||||
; EW
|
||||
<quantity id="C31B4BDB9D60C4215781FBB9FAF5F355" box="[1179,1345,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.65" metricValueMin="1.45" pageId="1" pageNumber="166" unit="mm" value="1.5499999999999998" valueMax="1.65" valueMin="1.45">1.45–1.65 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDFB9DFE48F38A" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
Colour: Dorsum (
|
||||
<figureCitation id="9CD8FABB9D60C4215288FB9DFE5FF33A" box="[402,491,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="1" pageNumber="166">Fig. 1A</figureCitation>
|
||||
) with head black, apex of labrum paler, reddish brown; pronotum and elytra dark brown to black, posterolateral angles and lateral margins of pronotum paler, yellowish brown. Maxillary palpi with palpomeres 2–3 dark yellow; palpomere 4 infuscated. Legs reddish brown, distal portions of femora somewhat lighter. Underside of head black. Pronotal and abdominal ventrites dark brown to black; pronotal hypomera and elytral epipleurs reddish brown.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDFB51FEADF169" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
Head: Clypeus large, with bluntly rounded anterolateral angles and distinctly concave anterior margin; lateral margin longer than longitudinal diameter of compound eyes. Clypeus and frons shining, without microreticulation and with shallow, irregular, medium punctures, spaced on average 2–3 puncture-widths apart. Frontoclypeal suture very weak and shallow, not visible on all specimens. Compound eyes moderate, very slightly protruding, emarginate anteriorly; ventral portion smaller than dorsal (approximately ¾ size). Antennae with eight antennomeres. Maxillary palpi moderately slender, half as long as maximum width of clypeus; palpomere 2 distinctly inflated. Palpomeres 2: 3: 4 =
|
||||
<quantity id="C31B4BDB9D60C42153F8FA69FEFFF285" box="[226,331,1483,1510]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="166" unit="mm" value="0.08">0.08 mm</quantity>
|
||||
:
|
||||
<quantity id="C31B4BDB9D60C421524CFA69FE0AF285" box="[342,446,1483,1510]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="166" unit="mm" value="0.07">0.07 mm</quantity>
|
||||
:
|
||||
<quantity id="C31B4BDB9D60C42152D3FA69FD85F285" box="[457,561,1483,1510]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="166" unit="mm" value="0.14">0.14 mm</quantity>
|
||||
. Labial palpi moderately slender, elongate, slightly longer than lateral margin of mentum.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDF9B6FD07F1DE" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">Pronotum: Transverse, highly arched, broadest at posterior angles. Anterior and posterior angles broadly rounded; anterior angles slightly protruding. Anterior margin arcuate over central 0.5; posterior margin almost straight over central third, arcuate laterally. All margins with narrow bead. Lateral margins without evident setae. Surface shining, without microreticulation; punctures spaced and sized as on head; somewhat shallower in some specimens, particularly posteriorly and laterally.</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDF965FB9DF02E" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">Elytra: Oval, broadest behind middle; arcuate over anterior third, then evenly rounded to apex; weakly emarginate around suture. Lateral margins without setae. Humeral region without evident callosity. Sutural stria present on posterior two thirds of elytra. Upper surface shining, without microreticulation. Punctures spaced as on head and pronotum, larger and deeper; distinct series and impressed striae absent.</paragraph>
|
||||
<paragraph id="045CE63E9D60C42153DDF8FAFB47F0BE" blockId="1.[151,1437,763,2014]" pageId="1" pageNumber="166">
|
||||
Venter:
|
||||
<collectingRegion id="C62728DC9D60C421523EF8FAFE37F011" box="[292,387,1880,1906]" country="Myanmar" name="Chin" pageId="1" pageNumber="166">Mentum</collectingRegion>
|
||||
flat with convex lateral margins; distinctly depressed anteriorly, anterior margin slightly emarginated; lateral margins with dense brushes of long, flattened setae. Surface shining, without microreticulation and with sparse setiferous punctures. Prosternum slightly tectiform, without median carina. Mesoventrite flat, without evident process. Abdominal ventrites entirely pubescent; ventrite 5 not emarginate apically.
|
||||
</paragraph>
|
||||
<caption id="509CB6B69D63C4225382F8A6FC8CF0EA" ID-DOI="http://doi.org/10.5281/zenodo.15216329" ID-Zenodo-Dep="15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="2" pageNumber="167" startId="2.[152,255,1796,1820]" targetBox="[175,1421,182,1772]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="045CE63E9D63C4225382F8A6FC8CF0EA" blockId="2.[151,1437,1796,1929]" pageId="2" pageNumber="167">
|
||||
<emphasis id="36973A2C9D63C4225382F8A6FEACF07E" bold="true" box="[152,280,1796,1821]" pageId="2" pageNumber="167">FIGURE 1.</emphasis>
|
||||
<taxonomicName id="C3E39DBD9D63C4225204F8A7FE7EF07E" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[286,458,1797,1821]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D63C4225204F8A7FE7EF07E" box="[286,458,1797,1821]" italics="true" pageId="2" pageNumber="167">Anacaena fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="36973A2C9D63C42252CAF8A7FD94F07E" bold="true" box="[464,544,1797,1821]" pageId="2" pageNumber="167">
|
||||
<taxonomicNameLabel id="2DA487579D63C42252CAF8A7FD94F07E" box="[464,544,1797,1821]" pageId="2" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="045CE63E9D62C42353DDFF35FD4FF606" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="168">
|
||||
Legs: Procoxae without spine-like setae. Profemora (
|
||||
<figureCitation id="9CD8FABB9D62C4235007FF35FCC7F7D2" box="[797,883,151,177]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1B</figureCitation>
|
||||
) pubescent on proximal half of ventral face with slightly oblique hairline, broadest approximately one third from anterior margin; mesofemora (
|
||||
<figureCitation id="9CD8FABB9D62C42357DFFF19FAAFF7B6" box="[1221,1307,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1B</figureCitation>
|
||||
) pubescent on proximal half along anterior margin, obliquely reducing to ¼ of posterior margin; metafemoral pubescence (
|
||||
<figureCitation id="9CD8FABB9D62C4235675FF7DFF0CF67E" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1B</figureCitation>
|
||||
) reduced to a narrow strip over mesal half of anterior margin and on proximal portion, adjacent to metatrochanter. Metatibia with strong spines on lateral face; distal spurs slender, longer mesal spur reaching midway along tarsomere 2 (
|
||||
<figureCitation id="9CD8FABB9D62C42353AEFEE9FEBAF606" box="[180,270,331,357]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1A</figureCitation>
|
||||
). Metatarsus slightly shorter than metatibia.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D62C42353DDFECDFBF0F5CA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="168">
|
||||
Aedeagus: Phallobase shorter than parameres (
|
||||
<figureCitation id="9CD8FABB9D62C42351EAFECDFCF8F6EA" box="[752,844,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1C</figureCitation>
|
||||
), slightly longer than wide, evenly converging to the distinct, broad knob-like manubrium. Ventral face of phallobase largely unpigmented, borderline between unpigmented and pigmented part of ventral face reaching manubrium. Parameres with undulated mesal margins and straight to weakly arcuate lateral margins, distinctly inflated and angled internally at their apices. Ventral bases of parameres fused. Dorsal bases curved, just reaching into phallobase. Median lobe wide, short, widening from the base to approximately halfway along its length, then almost parallel to approximately 1/5 from apex, then narrowing abruptly to a distinct point. Apex of median lobe not reaching apex of parameres, ending close to start of apical expansion of parameres; corona in apical position. Basal apophyses as long as main piece of median lobe, with distinct extension into phallobase, pointing mesad. Aedeagal length
|
||||
<quantity id="C31B4BDB9D62C423508BFD2DFBF4F5CA" box="[913,1088,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="3" pageNumber="168" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40− 0.45 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D62C42353DDFD11FB03F5AE" blockId="3.[151,1437,151,2013]" box="[199,1207,691,717]" pageId="3" pageNumber="168">Female: Not significantly different from males, other than slightly larger size—see above.</paragraph>
|
||||
<paragraph id="045CE63E9D62C42353DDFD75FAFBF592" blockId="3.[151,1437,151,2013]" box="[199,1359,727,753]" pageId="3" pageNumber="168">Variation: Some variation in punctation and impression of frontoclypeal suture and size as noted above.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D62C42353DDFD59FD31F45A" pageId="3" pageNumber="168" type="etymology">
|
||||
<paragraph id="045CE63E9D62C42353DDFD59FD31F45A" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="168">
|
||||
<emphasis id="36973A2C9D62C42353DDFD59FEFCF476" bold="true" box="[199,328,763,789]" pageId="3" pageNumber="168">Etymology</emphasis>
|
||||
. Named after Professor Garth Foster, Ayr,
|
||||
<collectingCountry id="7CF4A6AE9D62C4235033FD59FC3AF476" box="[809,910,763,789]" name="United Kingdom" pageId="3" pageNumber="168">Scotland</collectingCountry>
|
||||
, water beetler extraordinaire and mentor to the first author, on occasion of his 80
|
||||
<superScript id="F3964B769D62C4235114FCBCFDAFF44F" attach="left" box="[526,539,798,812]" fontSize="6" pageId="3" pageNumber="168">th</superScript>
|
||||
birthday.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D62C42353DDFCE0FF47F066" pageId="3" pageNumber="168" type="distribution">
|
||||
<paragraph id="045CE63E9D62C42353DDFCE0FE89F2A2" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="168">
|
||||
<emphasis id="36973A2C9D62C42353DDFCE0FE59F43E" bold="true" box="[199,493,834,861]" pageId="3" pageNumber="168">Distribution and ecology</emphasis>
|
||||
.
|
||||
<taxonomicName id="C3E39DBD9D62C42352E0FCE1FDD5F43E" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[506,609,835,861]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D62C42352E0FCE1FDD5F43E" box="[506,609,835,861]" italics="true" pageId="3" pageNumber="168">A. fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="36973A2C9D62C423517DFCE1FD74F43E" bold="true" box="[615,704,835,861]" pageId="3" pageNumber="168">
|
||||
<taxonomicNameLabel id="2DA487579D62C423517DFCE1FD74F43E" box="[615,704,835,861]" pageId="3" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is known only from the type series, taken at an altitude of ca.
|
||||
<quantity id="C31B4BDB9D62C4235668FCE1FF19F4E2" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="3" pageNumber="168" unit="m" value="943.0">943 m</quantity>
|
||||
in a plateau seepage bog (
|
||||
<figureCitation id="9CD8FABB9D62C42352CCFCC5FD98F4E2" box="[470,556,871,897]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="3" pageNumber="168">Fig. 1D</figureCitation>
|
||||
) surrounded by montane fynbos below Uitkyk Pass in the Cederberg,
|
||||
<collectingRegion id="C62728DC9D62C423565BFCC5FE88F4C6" country="South Africa" name="Western Cape" pageId="3" pageNumber="168">Western Cape Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="7CF4A6AE9D62C423525DFC29FE63F4C6" box="[327,471,907,933]" name="South Africa" pageId="3" pageNumber="168">South Africa</collectingCountry>
|
||||
. Beetles were netted from algal encrusted wet peat amongst grasses, sedges and restios, the saturated substrate having to be stamped down in order to reveal nettable water. DTB has visited this area a number of times since 2009 and every time this algal-covered peat has been damp, but without accessible water. It thus seems highly likely that the species is semi-terrestrial, living much of the time on damp ground out of liquid water. Such a lifestyle is known in other
|
||||
<taxonomicName id="C3E39DBD9D62C423516FFBB9FD51F356" authorityName="Thomson" authorityYear="1859" box="[629,741,1051,1077]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="36973A2C9D62C423516FFBB9FD51F356" box="[629,741,1051,1077]" italics="true" pageId="3" pageNumber="168">Anacaena</emphasis>
|
||||
</taxonomicName>
|
||||
in the region including
|
||||
<taxonomicName id="C3E39DBD9D62C42350E5FBB9FAFCF356" authority="(Hansen, 1999)" baseAuthorityName="Hansen" baseAuthorityYear="1999" box="[1023,1352,1051,1077]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="endroedyi">
|
||||
<emphasis id="36973A2C9D62C42350E5FBB9FB3BF356" box="[1023,1167,1051,1077]" italics="true" pageId="3" pageNumber="168">A. endroedyi</emphasis>
|
||||
(
|
||||
<bibRefCitation id="60729BCF9D62C42357BBFBB9FAF4F356" author="Hansen, M." box="[1185,1344,1051,1077]" pageId="3" pageNumber="168" pagination="121 - 172" refId="ref2928" refString="Hansen, M. (1999) Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family. Entomologica Scandinavica, 30, 121-172. https://doi.org/10.1163/187631200X00228" type="journal article" year="1999">Hansen, 1999</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="C3E39DBD9D62C423569EFBB9FE21F33A" authority="(Hansen, 1999)" baseAuthorityName="Hansen" baseAuthorityYear="1999" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="striata">
|
||||
<emphasis id="36973A2C9D62C423569EFBB9FF54F33B" italics="true" pageId="3" pageNumber="168">A. striata</emphasis>
|
||||
(
|
||||
<bibRefCitation id="60729BCF9D62C42353F4FB9DFE39F33A" author="Hansen, M." box="[238,397,1087,1113]" pageId="3" pageNumber="168" pagination="121 - 172" refId="ref2928" refString="Hansen, M. (1999) Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family. Entomologica Scandinavica, 30, 121-172. https://doi.org/10.1163/187631200X00228" type="journal article" year="1999">Hansen, 1999</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="C3E39DBD9D62C4235285FB9DFCB0F33A" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[415,772,1087,1113]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="glabriventris">
|
||||
<emphasis id="36973A2C9D62C4235285FB9DFDFAF33A" box="[415,590,1087,1113]" italics="true" pageId="3" pageNumber="168">A. glabriventris</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C423514EFB9DFCB0F33A" author="Komarek, A." box="[596,772,1087,1113]" pageId="3" pageNumber="168" pagination="303 - 349" refId="ref3006" refString="Komarek, A. (2004) Taxonomic revision of Anacaena Thomson, 1859. I. Afrotropical species (Coleoptera: Hydrophilidae). Koleopterologische Rundschau, 74, 303-349." type="journal article" year="2004">Komarek, 2004</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, whilst often living in water, can also frequently be found in damp terrestrial litter beside seepages and madicolous habitats in
|
||||
<collectingCountry id="7CF4A6AE9D62C423506CFBC1FBB1F31E" box="[886,1029,1123,1149]" name="South Africa" pageId="3" pageNumber="168">South Africa</collectingCountry>
|
||||
(DTB,
|
||||
<emphasis id="36973A2C9D62C4235741FBC1FB73F31E" box="[1115,1223,1123,1149]" italics="true" pageId="3" pageNumber="168">pers. obs.</emphasis>
|
||||
). Elsewhere, some species of the genus frequently occur in similar habitats, including the widespread Palaearctic
|
||||
<taxonomicName id="C3E39DBD9D62C42357ABFB25FF6DF3A6" authority="(Paykull, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="globulus">
|
||||
<emphasis id="36973A2C9D62C42357ABFB25FA9BF3C2" box="[1201,1327,1159,1185]" italics="true" pageId="3" pageNumber="168">A. globulus</emphasis>
|
||||
(Paykull, 1798)
|
||||
</taxonomicName>
|
||||
(e.g.
|
||||
<bibRefCitation id="60729BCF9D62C4235206FB09FDACF3A6" author="Balfour-Browne, W. A. F." box="[284,536,1194,1221]" pageId="3" pageNumber="168" refId="ref2612" refString="Balfour-Browne, W. A. F. (1958) British Water Beetles. Vol. 3. Ray Society, London, 210 pp." type="book" year="1958">Balfour-Browne 1958</bibRefCitation>
|
||||
). The only other water beetle found in association with
|
||||
<taxonomicName id="C3E39DBD9D62C4235781FB09FAB0F3A6" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[1179,1284,1195,1221]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D62C4235781FB09FAB0F3A6" box="[1179,1284,1195,1221]" italics="true" pageId="3" pageNumber="168">A. fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="36973A2C9D62C4235616FB09FADCF3A6" bold="true" box="[1292,1384,1195,1221]" pageId="3" pageNumber="168">
|
||||
<taxonomicNameLabel id="2DA487579D62C4235616FB09FADCF3A6" box="[1292,1384,1195,1221]" pageId="3" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
was
|
||||
<taxonomicName id="C3E39DBD9D62C423538DFB6DFB9CF38A" authority="Seidel, Minoshima, Arriaga-Varela & Fikacek, 2018" authorityName="Seidel, Minoshima, Arriaga-Varela & Fikacek" authorityYear="2018" box="[151,1064,1231,1257]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="trevornoahi">
|
||||
<emphasis id="36973A2C9D62C423538DFB6DFE71F38A" box="[151,453,1231,1257]" italics="true" pageId="3" pageNumber="168">Relictorygmus trevornoahi</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C42352D5FB6DFB9CF38A" author="Seidel, M. & Minoshima, Y. N. & Arriaga-Varela, E. & Fikacek, M." box="[463,1064,1231,1257]" pageId="3" pageNumber="168" pagination="375 - 402" refId="ref3561" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. & Fikacek, M. (2018) Breaking a disjunct distribution: A review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375-402. https://doi.org/10.3161/00034541ANZ2018.68.2.011" type="journal article" year="2018">Seidel, Minoshima, Arriaga-Varela & Fikáček, 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. This beetle was described from wetlands in the southern
|
||||
<collectingRegion id="C62728DC9D62C42352A3FB51FDE3F26E" box="[441,599,1267,1293]" country="South Africa" name="Western Cape" pageId="3" pageNumber="168">Western Cape</collectingRegion>
|
||||
(
|
||||
<bibRefCitation id="60729BCF9D62C4235172FB51FC8DF26E" author="Seidel, M. & Minoshima, Y. N. & Arriaga-Varela, E. & Fikacek, M." box="[616,825,1267,1293]" pageId="3" pageNumber="168" pagination="375 - 402" refId="ref3561" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. & Fikacek, M. (2018) Breaking a disjunct distribution: A review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375-402. https://doi.org/10.3161/00034541ANZ2018.68.2.011" type="journal article" year="2018">
|
||||
Seidel
|
||||
<emphasis id="36973A2C9D62C42351A3FB51FD40F26E" box="[697,756,1267,1293]" italics="true" pageId="3" pageNumber="168">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
), at low elevation and its discovery in the Cederberg is rather unexpected. Interestingly, the third species of
|
||||
<taxonomicName id="C3E39DBD9D62C4235008FAB5FC00F252" authorityName="Seidel, Minoshima, Arriaga-Varela & Fikacek" authorityYear="2018" box="[786,948,1303,1329]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="36973A2C9D62C4235008FAB5FC00F252" box="[786,948,1303,1329]" italics="true" pageId="3" pageNumber="168">Relictorygmus</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<collectingCountry id="7CF4A6AE9D62C42350C7FAB5FBC4F252" box="[989,1136,1303,1329]" name="South Africa" pageId="3" pageNumber="168">South Africa</collectingCountry>
|
||||
, recently discovered from high-altitude (
|
||||
<quantity id="C31B4BDB9D62C423522FFA99FE3FF236" box="[309,395,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.167" pageId="3" pageNumber="168" unit="m" value="1167.0">1167 m</quantity>
|
||||
) in the Kamiesberg,
|
||||
<collectingRegion id="C62728DC9D62C423516EFA99FC33F236" box="[628,903,1339,1365]" country="South Africa" name="Northern Cape" pageId="3" pageNumber="168">Northern Cape Province</collectingRegion>
|
||||
, also co-occurred with an
|
||||
<taxonomicName id="C3E39DBD9D62C42357B6FA99FAC0F236" box="[1196,1396,1339,1365]" pageId="3" pageNumber="168">
|
||||
<emphasis id="36973A2C9D62C42357B6FA99FAA8F236" box="[1196,1308,1339,1365]" italics="true" pageId="3" pageNumber="168">Anacaena</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="C3E39DBD9D62C423569EFA99FE9CF21A" authorityName="Komarek" authorityYear="2004" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="glabriventris">
|
||||
<emphasis id="36973A2C9D62C423569EFA99FE9CF21A" italics="true" pageId="3" pageNumber="168">A. glabriventris</emphasis>
|
||||
</taxonomicName>
|
||||
) (Bilton & Mlambo 2014) and
|
||||
<taxonomicName id="C3E39DBD9D62C4235197FAFDFC84F21A" authorityName="Seidel, Minoshima, Arriaga-Varela & Fikacek" authorityYear="2018" box="[653,816,1375,1401]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="trevornoahi">
|
||||
<emphasis id="36973A2C9D62C4235197FAFDFC84F21A" box="[653,816,1375,1401]" italics="true" pageId="3" pageNumber="168">R. trevornoahi</emphasis>
|
||||
</taxonomicName>
|
||||
co-occurred with
|
||||
<taxonomicName id="C3E39DBD9D62C42350E4FAFDFA9DF21A" authority="Hebauer, 1999" authorityName="Hebauer" authorityYear="1999" box="[1022,1321,1375,1401]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="capensis">
|
||||
<emphasis id="36973A2C9D62C42350E4FAFDFBCAF21B" box="[1022,1150,1375,1401]" italics="true" pageId="3" pageNumber="168">A. capensis</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C423579FFAFDFA9DF21A" author="Hebauer, F." box="[1157,1321,1375,1401]" pageId="3" pageNumber="168" pagination="7 - 16" refId="ref2975" refString="Hebauer, F. (1999) Neue und wenig bekannte Hydrophiloidea aus dem sudlichen Afrika (Coleoptera, Hydrophilidae). Acta Coleopterologica, 15, 7-16." type="journal article" year="1999">Hebauer, 1999</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="60729BCF9D62C4235623FAFDFF4EF2FE" author="Seidel, M. & Minoshima, Y. N. & Arriaga-Varela, E. & Fikacek, M." pageId="3" pageNumber="168" pagination="375 - 402" refId="ref3561" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. & Fikacek, M. (2018) Breaking a disjunct distribution: A review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375-402. https://doi.org/10.3161/00034541ANZ2018.68.2.011" type="journal article" year="2018">
|
||||
Seidel
|
||||
<emphasis id="36973A2C9D62C4235692FAFDFF00F2FE" italics="true" pageId="3" pageNumber="168">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
). Specimens collected in 2024 closely match an examined
|
||||
<typeStatus id="DB58589C9D62C4235084FA21FC4AF2FE" box="[926,1022,1411,1437]" pageId="3" pageNumber="168" type="paratype">paratype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="C3E39DBD9D62C4235732FA21FB78F2FE" authorityName="Seidel, Minoshima, Arriaga-Varela & Fikacek" authorityYear="2018" box="[1064,1228,1411,1437]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="trevornoahi">
|
||||
<emphasis id="36973A2C9D62C4235732FA21FB78F2FE" box="[1064,1228,1411,1437]" italics="true" pageId="3" pageNumber="168">R. trevornoahi</emphasis>
|
||||
</taxonomicName>
|
||||
externally and on male genitalia.
|
||||
</paragraph>
|
||||
<paragraph id="045CE63E9D62C42353DDFA69FF47F066" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="168">
|
||||
The area from which the new species was collected forms the source of the Driehoek River, being labelled Die Vlei or Driehoek Vlei on maps and is a major river system draining central Cederberg before joining the Doring River (
|
||||
<bibRefCitation id="60729BCF9D62C42353F2F9B1FE4CF14E" author="Quick, L. J. & Eckardt, F. D." box="[232,504,1554,1581]" pageId="3" pageNumber="168" pagination="85 - 93" refId="ref3337" refString="Quick, L. J. & Eckardt, F. D. (2015) The Cederberg: a rugged sandstone topography. In: Grab, S. & Knight, J. (Eds.), Landscapes and Landforms of South Africa. World Geomorphological Landscapes. Springer, Cham., pp. 85-93. https://doi.org/10.1007/978-3-319-03560-4_10" type="book chapter" year="2015">Quick & Eckardt 2015</bibRefCitation>
|
||||
). It is one of the sites from which pollen cores have been extracted to examine Late Pleistocene to Holocene vegetation changes in the region (
|
||||
<bibRefCitation id="60729BCF9D62C423506BF995FB14F132" author="Meadows, M. E. & Sugden J. M." box="[881,1184,1591,1617]" pageId="3" pageNumber="168" pagination="34 - 43" refId="ref3276" refString="Meadows, M. E. & Sugden J. M. (1991) A vegetation history of the last 14 000 years on the Cederberg, south-western Cape Province. South African Journal of Science, 87, 34-43. [https://hdl.handle.net/10520/AJA00382353_9382]" type="journal article" year="1991">Meadows & Sugden 1991</bibRefCitation>
|
||||
), peat deposition here apparently dating to at least 14,600 bp. This area of the high Cederberg supports a number of localised endemics, including water beetles (
|
||||
<bibRefCitation id="60729BCF9D62C42352B0F9DDFD80F1FA" author="Bilton, D. T." box="[426,564,1663,1689]" pageId="3" pageNumber="168" pagination="438 - 462" refId="ref2639" refString="Bilton, D. T. (2014) New species and new records of Pterosthetops: eumadicolous water beetles of the South African Cape (Coleoptera, Hydraenidae). Zootaxa, 3811 (4), 438-462. https://doi.org/10.11646/zootaxa.3811.4.2" type="journal article" year="2014">Bilton 2014</bibRefCitation>
|
||||
) and mesic terrestrial invertebrates such as Onychophora (likely
|
||||
<taxonomicName id="C3E39DBD9D62C4235615F9DDFD5EF1DE" authority="Daniels, McDonald & Picker, 2013" authorityName="Daniels" authorityYear="2013" family="Peripatopsidae" genus="Peripatopsis" kingdom="Animalia" pageId="3" pageNumber="168" phylum="Onychophora" rank="species" species="cederbergiensis">
|
||||
<emphasis id="36973A2C9D62C4235615F9DDFEF3F1DE" italics="true" pageId="3" pageNumber="168">Peripatopsis cederbergiensis</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C423524BF901FD5EF1DE" author="Daniels, S. R. & McDonald, D. E. & Picker, M. D." box="[337,746,1699,1725]" pageId="3" pageNumber="168" pagination="656 - 674" refId="ref2810" refString="Daniels, S. R., McDonald, D. E. & Picker, M. D. (2013) Evolutionary insight into the Peripatopsis balfouri sensu lato species complex (Onychophora: Peripatopsidae) reveals novel lineages and zoogeographic patterning. Zoologica Scripta, 42, 656-674. https://doi.org/10.1111/zsc.12025" type="journal article" year="2013">Daniels, McDonald & Picker, 2013</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; DTB
|
||||
<emphasis id="36973A2C9D62C4235021F901FC1EF1DE" box="[827,938,1699,1725]" italics="true" pageId="3" pageNumber="168">pers. obs.</emphasis>
|
||||
).
|
||||
<taxonomicName id="C3E39DBD9D62C42350D9F901FB99F1DE" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[963,1069,1699,1725]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D62C42350D9F901FB99F1DE" box="[963,1069,1699,1725]" italics="true" pageId="3" pageNumber="168">A. fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="36973A2C9D62C423572DF901FB27F1DE" bold="true" box="[1079,1171,1699,1725]" pageId="3" pageNumber="168">
|
||||
<taxonomicNameLabel id="2DA487579D62C423572DF901FB27F1DE" box="[1079,1171,1699,1725]" pageId="3" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
occurs relatively close geographically to
|
||||
<taxonomicName id="C3E39DBD9D62C4235279F965FCA4F182" authority="Bilton & Komarek, 2016" authorityName="Bilton & Komarek" authorityYear="2016" box="[355,784,1735,1761]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="namaqua">
|
||||
<emphasis id="36973A2C9D62C4235279F965FE5FF183" box="[355,491,1735,1761]" italics="true" pageId="3" pageNumber="168">A. namaqua</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C42352E9F965FCA4F182" author="Bilton, D. T. & Komarek, A." box="[499,784,1735,1761]" pageId="3" pageNumber="168" pagination="593 - 599" refId="ref2693" refString="Bilton, D. T. & Komarek, A. (2016) A new species of Anacaena Thomson, 1859 from South Africa (Coleoptera: Hydrophilidae). Zootaxa, 4139 (4), 593-599. https://doi.org/10.11646/zootaxa.4139.4.11" type="journal article" year="2016">Bilton & Komarek, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, but in quite a different microhabitat apparently, at higher altitude.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4CF9B5B59D62C42453DDF8ACFADFF622" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168" type="diagnosis">
|
||||
<paragraph id="045CE63E9D62C42453DDF8ACFADFF622" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,150,321]" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168">
|
||||
<emphasis id="36973A2C9D62C42353DDF8ACFE70F04B" bold="true" box="[199,452,1806,1832]" pageId="3" pageNumber="168">Differential diagnosis</emphasis>
|
||||
. The new species is most similar structurally to
|
||||
<taxonomicName id="C3E39DBD9D62C42350D7F8ADFBFEF04B" authorityName="Hebauer" authorityYear="1999" box="[973,1098,1807,1833]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="capensis">
|
||||
<emphasis id="36973A2C9D62C42350D7F8ADFBFEF04B" box="[973,1098,1807,1833]" italics="true" pageId="3" pageNumber="168">A. capensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C3E39DBD9D62C423574DF8ADFAB1F04A" authorityName="Komarek" authorityYear="2004" box="[1111,1285,1807,1833]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="glabriventris">
|
||||
<emphasis id="36973A2C9D62C423574DF8ADFAB1F04A" box="[1111,1285,1807,1833]" italics="true" pageId="3" pageNumber="168">A. glabriventris</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C3E39DBD9D62C423560AF8ADFA21F04B" authorityName="Bilton & Komarek" authorityYear="2016" box="[1296,1429,1807,1833]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="namaqua">
|
||||
<emphasis id="36973A2C9D62C423560AF8ADFA21F04B" box="[1296,1429,1807,1833]" italics="true" pageId="3" pageNumber="168">A. namaqua</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="C3E39DBD9D62C423538DF891FE70F02E" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[151,452,1843,1869]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="reducta">
|
||||
<emphasis id="36973A2C9D62C423538DF891FEB9F02E" box="[151,269,1843,1869]" italics="true" pageId="3" pageNumber="168">A. reducta</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C423520FF891FE70F02E" author="Komarek, A." box="[277,452,1843,1869]" pageId="3" pageNumber="168" pagination="303 - 349" refId="ref3006" refString="Komarek, A. (2004) Taxonomic revision of Anacaena Thomson, 1859. I. Afrotropical species (Coleoptera: Hydrophilidae). Koleopterologische Rundschau, 74, 303-349." type="journal article" year="2004">Komarek, 2004</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="C3E39DBD9D62C42352E7F891FCAEF02E" authority="Hebauer, 1999" authorityName="Hebauer" authorityYear="1999" box="[509,794,1843,1869]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="tenella">
|
||||
<emphasis id="36973A2C9D62C42352E7F891FDDEF02E" box="[509,618,1843,1869]" italics="true" pageId="3" pageNumber="168">A. tenella</emphasis>
|
||||
<bibRefCitation id="60729BCF9D62C4235168F891FCAEF02E" author="Hebauer, F." box="[626,794,1843,1869]" pageId="3" pageNumber="168" pagination="7 - 16" refId="ref2975" refString="Hebauer, F. (1999) Neue und wenig bekannte Hydrophiloidea aus dem sudlichen Afrika (Coleoptera, Hydrophilidae). Acta Coleopterologica, 15, 7-16." type="journal article" year="1999">Hebauer, 1999</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, sharing the following features with these South African species: eight antennomeres, no preocular patches on clypeus, flat mesoventrite, colouration of the pronotum (dark brown to black with yellow lateral margins), absence of serial elytral punctures and reduced metafemoral pubescence. The new species can be readily separated from
|
||||
<taxonomicName id="C3E39DBD9D62C423505EF83DFC79F0DB" authorityName="Bilton & Komarek" authorityYear="2016" box="[836,973,1951,1977]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="namaqua">
|
||||
<emphasis id="36973A2C9D62C423505EF83DFC79F0DB" box="[836,973,1951,1977]" italics="true" pageId="3" pageNumber="168">A. namaqua</emphasis>
|
||||
</taxonomicName>
|
||||
by its uniformly dark coloration and the form of the aedeagus.
|
||||
<taxonomicName id="C3E39DBD9D62C4235294F861FDF8F0BE" authorityName="Bilton & Mlambo & Balke" authorityYear="2025" box="[398,588,1987,2013]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="fosteri" status="sp. nov.">
|
||||
<emphasis id="36973A2C9D62C4235294F861FDF8F0BE" box="[398,588,1987,2013]" italics="true" pageId="3" pageNumber="168">Anacaena fosteri</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="36973A2C9D62C4235148F861FD1FF0BE" bold="true" box="[594,683,1987,2013]" pageId="3" pageNumber="168">
|
||||
<taxonomicNameLabel id="2DA487579D62C4235148F861FD1FF0BE" box="[594,683,1987,2013]" pageId="3" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be distinguished from
|
||||
<taxonomicName id="C3E39DBD9D62C42350C2F861FBFFF0BE" authorityName="Komarek" authorityYear="2004" box="[984,1099,1987,2013]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="reducta">
|
||||
<emphasis id="36973A2C9D62C42350C2F861FBFFF0BE" box="[984,1099,1987,2013]" italics="true" pageId="3" pageNumber="168">A. reducta</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="C3E39DBD9D62C423579AF861FB5EF0BE" authorityName="Hebauer" authorityYear="1999" box="[1152,1258,1987,2013]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="tenella">
|
||||
<emphasis id="36973A2C9D62C423579AF861FB5EF0BE" box="[1152,1258,1987,2013]" italics="true" pageId="3" pageNumber="168">A. tenella</emphasis>
|
||||
</taxonomicName>
|
||||
on its generally larger body size (2.35–2.95 vs. <
|
||||
<quantity id="C31B4BDB9D65C4245114FF34FDDCF7D2" box="[526,616,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="169" unit="mm" value="2.5">2.5 mm</quantity>
|
||||
), from
|
||||
<taxonomicName id="C3E39DBD9D65C42451A3FF35FCDDF7D2" authorityName="Komarek" authorityYear="2004" box="[697,873,151,177]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="glabriventris">
|
||||
<emphasis id="36973A2C9D65C42451A3FF35FCDDF7D2" box="[697,873,151,177]" italics="true" pageId="4" pageNumber="169">A. glabriventris</emphasis>
|
||||
</taxonomicName>
|
||||
on the entirely pubescent abdominal ventrites and the very different aedeagus and from
|
||||
<taxonomicName id="C3E39DBD9D65C424512CFF19FD01F7B7" authorityName="Hebauer" authorityYear="1999" box="[566,693,187,213]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="capensis">
|
||||
<emphasis id="36973A2C9D65C424512CFF19FD01F7B7" box="[566,693,187,213]" italics="true" pageId="4" pageNumber="169">A. capensis</emphasis>
|
||||
</taxonomicName>
|
||||
on its distinctly different aedeagus. It is further distinguished from
|
||||
<taxonomicName id="C3E39DBD9D65C424538DFF7DFEA2F79B" authorityName="Hebauer" authorityYear="1999" box="[151,278,223,249]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="capensis">
|
||||
<emphasis id="36973A2C9D65C424538DFF7DFEA2F79B" box="[151,278,223,249]" italics="true" pageId="4" pageNumber="169">A. capensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="C3E39DBD9D65C4245256FF7DFE48F79A" authorityName="Komarek" authorityYear="2004" box="[332,508,223,249]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="glabriventris">
|
||||
<emphasis id="36973A2C9D65C4245256FF7DFE48F79A" box="[332,508,223,249]" italics="true" pageId="4" pageNumber="169">A. glabriventris</emphasis>
|
||||
</taxonomicName>
|
||||
by the less elongate habitus, the less impressed frontoclypeal suture and the darker dorsal pigmentation, with pale pronotal margins narrower and much less evident. The aedeagus, with the paramere apices greatly surpassing the tip of the median lobe (
|
||||
<figureCitation id="9CD8FABB9D65C42451FEFE85FC8DF622" box="[740,825,295,321]" captionStart="FIGURE 1" captionStartId="2.[152,255,1796,1820]" captionTargetBox="[175,1421,182,1772]" captionTargetId="figure-22@2.[165,1421,181,1773]" captionTargetPageId="2" captionText="FIGURE 1. Anacaena fosteri sp. nov. A) Holotype male, dorsal habitus; B) Paratype male, ventral habitus; C) Holotype male, aedeagus; D) Type locality, South Africa, Western Cape Province, Cederberg, Die Vlei along headwaters of Driehoek River, 943 m. Red arrow indicates microhabitat from which specimens were netted (after trampling), yellow arrow indicates approximate position of the river channel. Scale bar A–B = 1 mm; C = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15216329" httpUri="https://zenodo.org/record/15216329/files/figure.png" pageId="4" pageNumber="169">Fig. 1C</figureCitation>
|
||||
) is unique within the known South African fauna.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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