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<treatment ID-GBIF-Taxon="183410443" LSID="urn:lsid:plazi:treatment:1636A415D7A2210C1021C2471FF3F203" httpUri="http://treatment.plazi.org/id/1636A415D7A2210C1021C2471FF3F203" lastPageNumber="60" pageNumber="60">
<subSubSection type="nomenclature">
<paragraph pageId="29" pageNumber="60">
<treatment id="1636A415D7A2210C1021C2471FF3F203" ID-GBIF-Taxon="183410443" LSID="urn:lsid:plazi:treatment:1636A415D7A2210C1021C2471FF3F203" httpUri="http://treatment.plazi.org/id/1636A415D7A2210C1021C2471FF3F203" lastPageNumber="60" pageNumber="60">
<subSubSection id="7354D0CCE411897460290F9FEC9E9CF6" type="nomenclature">
<paragraph id="72532087082801D59175269A9BD605EC" pageId="29" pageNumber="60">
Family
<taxonomicName family="Neritidae" lsidName="" pageId="29" pageNumber="60" rank="family">Neritidae</taxonomicName>
<taxonomicName id="9D913DBE54BB331F0EBDC4922633D191" ID-CoL="7NK24" class="Gastropoda" family="Neritidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
Rafinesque, 1815
</paragraph>
</subSubSection>
<subSubSection type="reference_group">
<paragraph pageId="29" pageNumber="60">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
<subSubSection id="BD0C8BDB9D957464600CFB8173ED363A" type="reference_group">
<paragraph id="0A440D6994D79ABBADF8B98F3B52D6DD" pageId="29" pageNumber="60">
<taxonomicName id="D4B6895A4B0FBB1E24CE5FC4E568D37C" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
(Pfeiffer, 1828)
</paragraph>
<paragraph pageId="29" pageNumber="60">
<paragraph id="E223B2B01873167D82A53043AEAAEDA3" pageId="29" pageNumber="60">
*1828
<taxonomicName class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Nerita danubialis</taxonomicName>
<taxonomicName id="477C4DC905F73FE44DEED8C40959DBEE" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Nerita danubialis</taxonomicName>
Pfeiffer: 48, pl. 8, figs 17, 18.
</paragraph>
<paragraph pageId="29" pageNumber="60">
<paragraph id="A7BB0A8BFB6A025835CF3D558A526BC1" pageId="29" pageNumber="60">
2009
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
<taxonomicName id="DBF22289C0666794887C6C277FE5D57A" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
(C. Pfeiffer, 1828). -
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
<normalizedToken id="DBE8F80D7A45FAD2BABB97575964FC02" originalValue="Fehér">Feher</normalizedToken>
et al.: figs 2
<normalizedToken originalValue="a–k">a-k</normalizedToken>
<normalizedToken id="F6D1BAD69AF02BCC0998E51705499FD9" originalValue="a–k">a-k</normalizedToken>
, 4
<normalizedToken originalValue="a–c">a-c</normalizedToken>
<normalizedToken id="6EB7E93464EE1B38B08E1E6B0EC48084" originalValue="a–c">a-c</normalizedToken>
, 5
<normalizedToken originalValue="a–c">a-c</normalizedToken>
<normalizedToken id="74ED1EF15DF560D83D562C2D92CD10B9" originalValue="a–c">a-c</normalizedToken>
.
</paragraph>
<paragraph pageId="29" pageNumber="60">
<paragraph id="6F05A1644806A6824CAF635C989137EB" pageId="29" pageNumber="60">
2012
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
<taxonomicName id="05533586EC71F6172997C47A13E05E1A" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
(Pfeiffer, 1828). - Welter-Schultes: 27, unnumbered text figures.
</paragraph>
<paragraph pageId="29" pageNumber="60">
<paragraph id="221A048C7A568A5BF025C29E337C1945" pageId="29" pageNumber="60">
2016
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus (Theodoxus) danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis" subGenus="Theodoxus">Theodoxus (Theodoxus) danubialis</taxonomicName>
<taxonomicName id="D109C027630212156011F689A6F9F9CA" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus (Theodoxus) danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis" subGenus="Theodoxus">Theodoxus (Theodoxus) danubialis</taxonomicName>
(Pfeiffer, 1828). - Vinarski and Kantor: 156 [and synonyms therein].
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="29" pageNumber="60">Status. Accepted native species.</paragraph>
<subSubSection id="69616D5916531C3591E981CDB4AEC8C1" type="discussion">
<paragraph id="5EEBD9882013FB2F74B65BA135FC50CE" pageId="29" pageNumber="60">Status. Accepted native species.</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageId="29" pageNumber="60">Type locality. Danube River, Vienna, Austria.</paragraph>
<paragraph pageId="29" pageNumber="60">
<subSubSection id="71643852509F9F2606651A42DB5873BE" type="distribution">
<paragraph id="5B7DD101108CF7A08E24286C0B24FDB2" pageId="29" pageNumber="60">Type locality. Danube River, Vienna, Austria.</paragraph>
<paragraph id="3D33A5C82BA590A8B8F1581A2958EFD0" pageId="29" pageNumber="60">
Distribution. Danube River catchment, central to south-eastern Europe, as well as northern Italy (
<bibRefCitation author="Feher, Z" journalOrPublisher="Mollusca" pageId="74" pageNumber="105" pagination="95 - 107" title="An attempt to reveal the systematic relationship between Theodoxusprevostianus (C. Pfeiffer, 1828) and Theodoxusdanubialis (C. Pfeiffer, 1828) (Mollusca, Gastropoda, Neritidae)." volume="27" year="2009">
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
<bibRefCitation id="D41B371802968FFD1416BFE34012EFF2" author="Feher, Z" journalOrPublisher="Mollusca" pageId="74" pageNumber="105" pagination="95 - 107" title="An attempt to reveal the systematic relationship between Theodoxusprevostianus (C. Pfeiffer, 1828) and Theodoxusdanubialis (C. Pfeiffer, 1828) (Mollusca, Gastropoda, Neritidae)." volume="27" year="2009">
<normalizedToken id="B05253289E5A5F2C5E23BEFCE3A08247" originalValue="Fehér">Feher</normalizedToken>
et al. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="29" pageNumber="60">
<subSubSection id="C1DC5C942F6E36650D281055503F675F" type="discussion">
<paragraph id="998188C03EF200444A86ABD11D0BADEC" pageId="29" pageNumber="60">
Taxonomic notes. The latest phylogenetic data supports a sister relationship between
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
<taxonomicName id="EE4F19342CF012F0977DD90C29A60CE4" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus danubialis" order="Cycloneritida" pageId="29" pageNumber="60" phylum="Mollusca" rank="species" species="danubialis">Theodoxus danubialis</taxonomicName>
and the clade containing
<taxonomicName lsidName="T. fluviatilis" pageId="29" pageNumber="60" rank="species" species="fluviatilis">T. fluviatilis</taxonomicName>
<taxonomicName id="2E3F171D367DB5AF1EF758269BE6BCAF" lsidName="T. fluviatilis" pageId="29" pageNumber="60" rank="species" species="fluviatilis">T. fluviatilis</taxonomicName>
and
<taxonomicName lsidName="T. velox" pageId="29" pageNumber="60" rank="species" species="velox">T. velox</taxonomicName>
<taxonomicName id="825E345F29492B3B1B23CEA03DF8CF94" lsidName="T. velox" pageId="29" pageNumber="60" rank="species" species="velox">T. velox</taxonomicName>
(AFS, unpublished data). Some authors believe
<taxonomicName lsidName="T. danubialis" pageId="29" pageNumber="60" rank="species" species="danubialis">T. danubialis</taxonomicName>
<taxonomicName id="58C0A9B7B6C059817CC024FF7F8940FF" lsidName="T. danubialis" pageId="29" pageNumber="60" rank="species" species="danubialis">T. danubialis</taxonomicName>
and
<taxonomicName lsidName="T. prevostianus" pageId="29" pageNumber="60" rank="species" species="prevostianus">T. prevostianus</taxonomicName>
<taxonomicName id="A89AFC5D1625BED19128FCC44C08E9B4" lsidName="T. prevostianus" pageId="29" pageNumber="60" rank="species" species="prevostianus">T. prevostianus</taxonomicName>
may represent different species given some level of genetic, ecological, and morphological differentiation (
<bibRefCitation author="Feher, Z" journalOrPublisher="Mollusca" pageId="74" pageNumber="105" pagination="95 - 107" title="An attempt to reveal the systematic relationship between Theodoxusprevostianus (C. Pfeiffer, 1828) and Theodoxusdanubialis (C. Pfeiffer, 1828) (Mollusca, Gastropoda, Neritidae)." volume="27" year="2009">
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
<bibRefCitation id="99287AAD24BC49060CE62ECF53EF0D03" author="Feher, Z" journalOrPublisher="Mollusca" pageId="74" pageNumber="105" pagination="95 - 107" title="An attempt to reveal the systematic relationship between Theodoxusprevostianus (C. Pfeiffer, 1828) and Theodoxusdanubialis (C. Pfeiffer, 1828) (Mollusca, Gastropoda, Neritidae)." volume="27" year="2009">
<normalizedToken id="4505C44641F31A6139C6394F347A7763" originalValue="Fehér">Feher</normalizedToken>
et al. 2009
</bibRefCitation>
,
<bibRefCitation author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="86" pageNumber="117" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
<bibRefCitation id="F79CE39CC9226F80C73ADA252253D7DC" author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="86" pageNumber="117" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
; but see also
<bibRefCitation author="Bandel, K" journalOrPublisher="Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitaet Hamburg" pageId="72" pageNumber="103" pagination="65 - 164" title="The history or Theodoxus and Neritina connected with the description and systematic evaluation of related Neritimorpha (Gastropoda)." url="http://www.paleoliste.de/bandel/bandel_2001e.pdf" volume="85" year="2001">Bandel 2001</bibRefCitation>
<bibRefCitation id="CCC61CC22546542A662D0B2A3390D109" author="Bandel, K" journalOrPublisher="Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitaet Hamburg" pageId="72" pageNumber="103" pagination="65 - 164" title="The history or Theodoxus and Neritina connected with the description and systematic evaluation of related Neritimorpha (Gastropoda)." url="http://www.paleoliste.de/bandel/bandel_2001e.pdf" volume="85" year="2001">Bandel 2001</bibRefCitation>
). More recent unpublished results may suggest that the genetic level of differentiation between these species is more indicative of intraspecific diversity within a single species (AFS, unpublished data).
</paragraph>
</subSubSection>
<subSubSection type="conservation">
<paragraph pageId="29" pageNumber="60">
<subSubSection id="C44811A82EDAB31D422EF51B0BEA0D4E" type="conservation">
<paragraph id="EA1440015BB6821B045C1151A03F03DF" pageId="29" pageNumber="60">
Conservation status. Least Concern (
<bibRefCitation pageId="29" pageNumber="60">Tomovic et al. 2010</bibRefCitation>
<bibRefCitation id="886DCDD187E463C89C7E81068C3127DB" pageId="29" pageNumber="60">Tomovic et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>

213
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Dybowski, 1913
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Remarks. The distinction of the genera
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,
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, and
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follows
<bibRefCitation author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="80" pageNumber="111" pagination="43 - 103" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. (2018)</bibRefCitation>
<bibRefCitation id="CA79BB5FF6CDEF751CFA8DEED068FCA2" author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="80" pageNumber="111" pagination="43 - 103" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. (2018)</bibRefCitation>
. The three taxa are differentiated based on details of the protoconch and the expression of teleoconch sculpture.
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s. s. is characterised by a single distinct but fine spiral keel below the suture. It is usually smooth, yet within the type species some reticulate ornament can be found. Species of
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expose a distinctive, reticulate pattern on the teleoconch and a malleate protoconch with faint spiral threads. The aperture of
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often develops a distinct flat base. The discinction of the two genera is subject of current research.
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also has a malleate protoconch but with more distinct spiral threads; teleoconch sculpture is occasionally present as minute elongate nodules.
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217
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Subfamily
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Brusina, 1882
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Remarks. The genus concepts of Pontocaspian
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follow the revision of
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. Further change is expected in several of the keeled species here
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under?
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(?
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,?
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,?
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,?
<taxonomicName lsidName="T. pseudobacuana" pageId="42" pageNumber="73" rank="species" species="pseudobacuana">T. pseudobacuana</taxonomicName>
<taxonomicName id="EFDC11790208A63211EC0072B9E86B7D" lsidName="T. pseudobacuana" pageId="42" pageNumber="73" rank="species" species="pseudobacuana">T. pseudobacuana</taxonomicName>
, and?
<taxonomicName lsidName="T. pseudodimiata" pageId="42" pageNumber="73" rank="species" species="pseudodimiata">T. pseudodimiata</taxonomicName>
<taxonomicName id="222758DE61E9635ACFC0B02E68385DEB" lsidName="T. pseudodimiata" pageId="42" pageNumber="73" rank="species" species="pseudodimiata">T. pseudodimiata</taxonomicName>
) that may be grouped in their own genus for which the name
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Dybowski &amp; Grochmalicki, 1917 (type species:
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Eichwald, 1838) is available. However, such a decision will require further documentation.
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The following key, modified from
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genera.
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Family
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Stimpson, 1865
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<subSubSection type="discussion">
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<subSubSection id="D55231B1A269824D43CDD82612710030" type="discussion">
<paragraph id="C2B1389188391940B315E3B58126CB7E" pageId="33" pageNumber="64">
Remarks. The
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<taxonomicName id="9F178FB913E083FF7CBB512CF4B711A3" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="33" pageNumber="64" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
form the most species-rich mollusc group in the Pontocaspian region. However, in general, useful shell characters are few and highly variable (Wilke and Delicado in press). Descriptions in the past have often been very general, and illustrations of types are notably poor for several of the endemic taxa. A strong tendency of naming large numbers of species has developed throughout the 20th century (e.g.,
<bibRefCitation pageId="33" pageNumber="64">Logvinenko and Starobogatov 1969</bibRefCitation>
<bibRefCitation id="DC8411658BA3924D00CC77679D8CAABD" pageId="33" pageNumber="64">Logvinenko and Starobogatov 1969</bibRefCitation>
), but for some groups where morphological and genetic analyses could be performed (e.g.,
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<taxonomicName id="D093AFC4CA78D1DA6692D2BC0B242A91" class="Gastropoda" family="Hydrobiidae" genus="Caspiohydrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiohydrobia" order="Littorinimorpha" pageId="33" pageNumber="64" phylum="Mollusca" rank="genus">Caspiohydrobia</taxonomicName>
spp.) it has been demonstrated that actual species numbers were much lower than the number of species described (
<bibRefCitation author="Haase, M" journalOrPublisher="Journal of Molluscan Studies" pageId="76" pageNumber="107" pagination="101 - 105" title="Ecrobiagrimmi in brackish Lake Sawa, Iraq: indirect evidence for long-distance dispersal of hydrobiid gastropods (Caenogastropoda: Rissooidea) by birds." url="https://doi.org/10.1093/mollus/eyp051" volume="76" year="2010">Haase et al. 2010</bibRefCitation>
<bibRefCitation id="0EB2723EC37366F2035118C1F87242B5" author="Haase, M" journalOrPublisher="Journal of Molluscan Studies" pageId="76" pageNumber="107" pagination="101 - 105" title="Ecrobiagrimmi in brackish Lake Sawa, Iraq: indirect evidence for long-distance dispersal of hydrobiid gastropods (Caenogastropoda: Rissooidea) by birds." url="https://doi.org/10.1093/mollus/eyp051" volume="76" year="2010">Haase et al. 2010</bibRefCitation>
). For many of the endemic species, especially in the genus
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<taxonomicName id="FEED5D7E71A3BC9BBC192CA6ECE0BC58" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="33" pageNumber="64" phylum="Mollusca" rank="genus">Turricaspia</taxonomicName>
, the apparent loss of types, combined with the lack of living material makes it impossible to assess their taxonomic status. Currently, a number of taxonomic works is in progress on the endemic Pontocaspian hydrobiid groups, and some different insights on the genus-level classifications exist. Here, we adopt a conservative approach, mostly based on
<bibRefCitation author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="80" pageNumber="111" pagination="43 - 103" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. (2018)</bibRefCitation>
<bibRefCitation id="CD9774EEAD58CD1BA486A7519E178AA0" author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="80" pageNumber="111" pagination="43 - 103" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. (2018)</bibRefCitation>
.
</paragraph>
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Stimpson, 1865
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<subSubSection type="discussion">
<paragraph pageId="40" pageNumber="71">
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<paragraph id="3C98FBF87A036EB122BCCCE141DBEA42" pageId="40" pageNumber="71">
Remarks. In addition to the taxa discussed below, the following species of
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have been mentioned from the Black Sea basin (updated statuses after
<bibRefCitation author="Mirzoev, GS" journalOrPublisher="International Journal of Zoological Studies" pageId="79" pageNumber="110" url="http://www.molluscabase.org" year="2018 a">MolluscaBase 2018a</bibRefCitation>
<bibRefCitation id="A03C65AFA6E57319CD821D676DED8DED" author="Mirzoev, GS" journalOrPublisher="International Journal of Zoological Studies" pageId="79" pageNumber="110" url="http://www.molluscabase.org" year="2018 a">MolluscaBase 2018a</bibRefCitation>
):
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<taxonomicName id="1A2225D3A2575612C8E7AB3FCE27AC91" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobia aciculina" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="species" species="aciculina">Hydrobia aciculina</taxonomicName>
(Bourguignat, 1876),
<taxonomicName lsidName="H. acuta" pageId="40" pageNumber="71" rank="species" species="acuta">H. acuta</taxonomicName>
<taxonomicName id="F4B2A04F97774C326D4289596F2FE143" lsidName="H. acuta" pageId="40" pageNumber="71" rank="species" species="acuta">H. acuta</taxonomicName>
(Draparnaud, 1805),
<taxonomicName lsidName="H. euryomphala" pageId="40" pageNumber="71" rank="species" species="euryomphala">H. euryomphala</taxonomicName>
<taxonomicName id="39448352DCA5B3AA4891FAE9B9EA2255" lsidName="H. euryomphala" pageId="40" pageNumber="71" rank="species" species="euryomphala">H. euryomphala</taxonomicName>
(Bourguignat, 1876),
<taxonomicName lsidName="H. mabilli" pageId="40" pageNumber="71" rank="species" species="mabilli">H. mabilli</taxonomicName>
<taxonomicName id="1FD3335174EB74057ED0BA8CA25A8068" lsidName="H. mabilli" pageId="40" pageNumber="71" rank="species" species="mabilli">H. mabilli</taxonomicName>
(Bourguignat, 1876) [currently accepted as
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Peringia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peringia mabilli" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="species" species="mabilli">Peringia mabilli</taxonomicName>
<taxonomicName id="A2E72E4E14A4993742DD0991F21BE794" class="Gastropoda" family="Hydrobiidae" genus="Peringia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peringia mabilli" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="species" species="mabilli">Peringia mabilli</taxonomicName>
],
<taxonomicName lsidName="H. macei" pageId="40" pageNumber="71" rank="species" species="macei">H. macei</taxonomicName>
<taxonomicName id="688A412094B1DC56551A68E536AC002C" lsidName="H. macei" pageId="40" pageNumber="71" rank="species" species="macei">H. macei</taxonomicName>
Paladilhe, 1867 [currently accepted as
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<taxonomicName id="58C50FD7E77B0DC1DC26659A260B3963" class="Gastropoda" family="Cochliopidae" genus="Heleobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heleobia macei" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="species" species="macei">Heleobia macei</taxonomicName>
],
<taxonomicName lsidName="H. procerula" pageId="40" pageNumber="71" rank="species" species="procerula">H. procerula</taxonomicName>
<taxonomicName id="DC9CE4443AB583A2F28ECC1D848880DE" lsidName="H. procerula" pageId="40" pageNumber="71" rank="species" species="procerula">H. procerula</taxonomicName>
(Paladilhe, 1869) [currently considered a synonym of
<taxonomicName lsidName="H. acuta" pageId="40" pageNumber="71" rank="species" species="acuta">H. acuta</taxonomicName>
<taxonomicName id="0097F5AD30E1EFDA8489D508D6E46A1D" lsidName="H. acuta" pageId="40" pageNumber="71" rank="species" species="acuta">H. acuta</taxonomicName>
] (
<bibRefCitation author="Anistratenko, VV" journalOrPublisher="Naukova dumka, Kiev" pageId="72" pageNumber="103" title="Mollyuski Azovskogo morya." url="http://ashipunov.info/shipunov/school/books/anistratenko2011_moll_azovsk_morja.pdf" year="2011">Anistratenko et al. 2011</bibRefCitation>
<bibRefCitation id="F5C49C9D5D9A404C58B31E9D6BB0C8D1" author="Anistratenko, VV" journalOrPublisher="Naukova dumka, Kiev" pageId="72" pageNumber="103" title="Mollyuski Azovskogo morya." url="http://ashipunov.info/shipunov/school/books/anistratenko2011_moll_azovsk_morja.pdf" year="2011">Anistratenko et al. 2011</bibRefCitation>
). These species were described from the Western Mediterranean and their occurrence in the Black Sea region requires re-investigation; partly the records might be misidentifications of the species of
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="genus">Ecrobia</taxonomicName>
<taxonomicName id="171CF56AD0117C10BD0BD81A05EF0775" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="genus">Ecrobia</taxonomicName>
listed below or
<taxonomicName class="Gastropoda" family="Cochliopidae" genus="Eupaludestrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupaludestrina" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="genus">Eupaludestrina</taxonomicName>
<taxonomicName id="AC4325DB6FA56C925ECF85DF86F10116" class="Gastropoda" family="Cochliopidae" genus="Eupaludestrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupaludestrina" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="genus">Eupaludestrina</taxonomicName>
(
<taxonomicName family="Cochliopidae" lsidName="" pageId="40" pageNumber="71" rank="family">Cochliopidae</taxonomicName>
<taxonomicName id="691F6D1ACB79411D1883074D7867FE45" class="Gastropoda" family="Cochliopidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="40" pageNumber="71" phylum="Mollusca" rank="family">Cochliopidae</taxonomicName>
) listed above.
</paragraph>
</subSubSection>

223
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Family
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Lamarck, 1809
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Remarks. For the genus
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, the species status of Pontocaspian material is subject of debate where morphological and increasingly molecular arguments show the possibility of sibling species occurrences (
<bibRefCitation author="Sromek, L" journalOrPublisher="Journal of Molluscan Studies" pageId="83" pageNumber="114" pagination="515 - 524" title="Strong genetic structuring of the cockle Cerastodermaglaucum across Europe: new insights from an intronic marker and multivariate analysis." url="https://doi.org/10.1093/mollus/eyw019" volume="82" year="2016">Sromek et al. 2016</bibRefCitation>
<bibRefCitation id="35CCD6EBE516BB1657729A22F6559B6D" author="Sromek, L" journalOrPublisher="Journal of Molluscan Studies" pageId="83" pageNumber="114" pagination="515 - 524" title="Strong genetic structuring of the cockle Cerastodermaglaucum across Europe: new insights from an intronic marker and multivariate analysis." url="https://doi.org/10.1093/mollus/eyw019" volume="82" year="2016">Sromek et al. 2016</bibRefCitation>
). The genus
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is relatively well established, however much uncertainty exists over distinction between the genera
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,
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, and
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. The generic concepts have shifted through time. Only lately,
<bibRefCitation pageId="3" pageNumber="34">Kijashko in Bogutskaya et al. (2013)</bibRefCitation>
<bibRefCitation id="DB4FAAA94CC407BC35670028CC8740EE" pageId="3" pageNumber="34">Kijashko in Bogutskaya et al. (2013)</bibRefCitation>
treated
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as a subgenus of
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.
<bibRefCitation author="Bueyuekmeric, Y" journalOrPublisher="Bollettino della Societa Malacologica Italiana" pageId="73" pageNumber="104" url="https://doi.org/10.1016/j.quaint.2016.03.018" year="2018">
<normalizedToken originalValue="Büyükmeriç">Bueyuekmeric</normalizedToken>
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<normalizedToken id="171560A18BC7EA95CC00F905B0BC0AE7" originalValue="Büyükmeriç">Bueyuekmeric</normalizedToken>
and Wesselingh (2018)
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discussed the distinction between the three genera and considered
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,
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, and
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as valid.
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199
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Family
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Gray, 1840
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Remarks. Pontocaspian dreissenid taxonomy suffers from a lack of coordinated shell and DNA analyses. A large part of our considerations relies on the work of
<bibRefCitation author="Rosenberg, G" journalOrPublisher="Canadian Journal of Fisheries and Aquatic Sciences" pageId="82" pageNumber="113" pagination="1474 - 1484" title="A Nomenclatural Review of Dreissena (Bivalvia: Dreissenidae), with Identification of the Quagga Mussel as Dreissenabugensis." url="https://doi.org/10.1139/f94-147" volume="51" year="1994">Rosenberg and Ludyanskiy (1994)</bibRefCitation>
<bibRefCitation id="FEC3ED3330E790C28299C92351A7D235" author="Rosenberg, G" journalOrPublisher="Canadian Journal of Fisheries and Aquatic Sciences" pageId="82" pageNumber="113" pagination="1474 - 1484" title="A Nomenclatural Review of Dreissena (Bivalvia: Dreissenidae), with Identification of the Quagga Mussel as Dreissenabugensis." url="https://doi.org/10.1139/f94-147" volume="51" year="1994">Rosenberg and Ludyanskiy (1994)</bibRefCitation>
who examined and illustrated all type material of Pontocaspian
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.
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