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<document id="E3F71D64F58DC992890EBA644F5DFF71" ID-DOI="10.11646/zootaxa.5492.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13211499" ID-ZooBank="35055BD1-1925-4947-91BA-5E1CC841071A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722790746796" checkinUser="plazi" docAuthor="Prena, Jens" docDate="2024" docId="326487E81924FFC2FF67ADE1FE1DFEC0" docLanguage="en" docName="zootaxa.5492.1.1.pdf" docOrigin="Zootaxa 5492 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chryasus clunis Prena 2024, new species" docType="treatment" docUuid="D7F52AB0-1155-4810-98DD-369CFDDFFB13" docUuidSource="ZooBank" docVersion="2" lastPageNumber="18" masterDocId="CE5DFF901935FFD0FFF0AC1FFFD0FFA5" masterDocTitle="Consolidation of Aniops Casey, Psiona Casey, Preglyptobaris Bondar and Prospoliata Hustache with Chryasus Champion (Coleoptera, Curculionidae, Baridinae) and descriptions of new species" masterLastPageNumber="24" masterPageNumber="1" pageNumber="18" updateTime="1722792009948" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F3DD5B59FF21D46D53210CB5A7CBF150" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E413B07A1273E51D9DA9DA8FC381443A">
<mods:title id="17325CF3B00BACF5250A27CF400DE222">Consolidation of Aniops Casey, Psiona Casey, Preglyptobaris Bondar and Prospoliata Hustache with Chryasus Champion (Coleoptera, Curculionidae, Baridinae) and descriptions of new species</mods:title>
</mods:titleInfo>
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<mods:role id="332DE8E79DA449D7F91CB1892810F6FD">
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<mods:namePart id="2EB9AB3BE37A6DC668B6CFC028E214B4">Prena, Jens</mods:namePart>
</mods:name>
<mods:typeOfResource id="11C433173D80F3C233E626FD6C98AF4F">text</mods:typeOfResource>
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<mods:title id="9A0CCEA09713B9B8A26F34200F0834AD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6FB21E73A334AB633CA69C1B3A4D31D9">
<mods:date id="16CF54A00DD25C7F7213D00DB2C42351">2024</mods:date>
<mods:detail id="F13E857D21533E0FD9CDF85212EA04F8" type="pubDate">
<mods:number id="4C6B3325924D1CB8378A3A570DE8A286">2024-08-02</mods:number>
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<mods:detail id="5629E1B7C3C64321F725004DFE95CE79" type="volume">
<mods:number id="D0E5B5F87E81EF069EC537E51D28C0BB">5492</mods:number>
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<mods:number id="6BED3B2762283E2397A8EC2CEFA7B055">1</mods:number>
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<mods:url id="C5E57602D7021D2397F8BCBC88F60089">http://dx.doi.org/10.11646/zootaxa.5492.1.1</mods:url>
</mods:location>
<mods:classification id="62CB2A8F8FA2DDA52E984E81EC11CC12">journal article</mods:classification>
<mods:identifier id="531BE97B5850BB759991EB573AA8DA36" type="DOI">10.11646/zootaxa.5492.1.1</mods:identifier>
<mods:identifier id="BE5255D7B0DD58374B13EA4258089B80" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="02C767E2FF82F58B3BE7DF0BDD709FBB" type="Zenodo-Dep">13211499</mods:identifier>
<mods:identifier id="FC9780D9AD7399C3AB211F074ABBE668" type="ZooBank">35055BD1-1925-4947-91BA-5E1CC841071A</mods:identifier>
</mods:mods>
<treatment id="326487E81924FFC2FF67ADE1FE1DFEC0" LSID="urn:lsid:zoobank.org:act:D7F52AB0-1155-4810-98DD-369CFDDFFB13" httpUri="http://treatment.plazi.org/id/326487E81924FFC2FF67ADE1FE1DFEC0" lastPageId="18" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="F2D765751924FFC1FF67ADE1FDA6FDBD" box="[151,630,510,537]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="BA7236FE1924FFC1FF67ADE1FDA6FDBD" blockId="17.[151,1187,510,609]" box="[151,630,510,537]" pageId="17" pageNumber="18">
<heading id="E13A81921924FFC1FF67ADE1FDA6FDBD" bold="true" box="[151,630,510,537]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="88B9EAEC1924FFC1FF67ADE1FDA6FDBD" bold="true" box="[151,630,510,537]" pageId="17" pageNumber="18">
16.
<taxonomicName id="7DCD4D7D1924FFC1FF33ADE1FE09FDBC" authority="Prena" authorityName="Prena" authorityYear="2024" box="[195,473,510,537]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clunis" status="sp. nov.">
<emphasis id="88B9EAEC1924FFC1FF33ADE1FE56FDBC" bold="true" box="[195,390,510,537]" italics="true" pageId="17" pageNumber="18">Chryasus clunis</emphasis>
Prena
</taxonomicName>
,
<taxonomicNameLabel id="938A57971924FFC1FE16ADE0FDA6FDBD" box="[486,630,510,537]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D765751924FFC1FF67AE3CFB73FDC4" pageId="17" pageNumber="18" type="description">
<paragraph id="BA7236FE1924FFC1FF67AE3CFE88FD98" blockId="17.[151,1187,510,609]" box="[151,344,547,573]" pageId="17" pageNumber="18">
(
<figureCitation id="22F62A7B1924FFC1FF6FAE3CFF2DFD98" box="[159,253,547,573]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="17" pageNumber="18">Figs. 21</figureCitation>
,
<figureCitation id="22F62A7B1924FFC1FEF9AE3CFE80FD98" box="[265,336,547,573]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="17" pageNumber="18">43, 44</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE1924FFC1FF67AE59FB73FDC4" blockId="17.[151,1187,510,609]" box="[151,1187,582,609]" pageId="17" pageNumber="18">
http://zoobank.org/
<uri id="CE5C3AFC1924FFC1FE82AE59FB73FDC4" box="[370,1187,582,609]" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
<uuid id="CE6B0C2B1924FFC1FD71AE59FB73FDC4" box="[641,1187,582,609]" pageId="17" pageNumber="18">D7F52AB0-1155-4810-98DD-369CFDDFFB13</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="F2D765751924FFC1FF67AE91FB48FD54" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="BA7236FE1924FFC1FF67AE91FB48FD54" blockId="17.[151,1437,654,2013]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF67AE91FEC2FD0D" bold="true" box="[151,274,654,680]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
At first glance,
<taxonomicName id="7DCD4D7D1924FFC1FE2AAE90FD93FD0C" authorityName="Prena" authorityYear="2024" box="[474,579,655,681]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1924FFC1FE2AAE90FD93FD0C" box="[474,579,655,681]" italics="true" pageId="17" pageNumber="18">C. clunis</emphasis>
</taxonomicName>
resembles other small reddish species, in particular
<taxonomicName id="7DCD4D7D1924FFC1FB46AE90FA9AFD0C" authorityName="Prena" authorityYear="2024" box="[1206,1354,655,681]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="88B9EAEC1924FFC1FB46AE90FA9AFD0C" box="[1206,1354,655,681]" italics="true" pageId="17" pageNumber="18">C. persimilis</emphasis>
</taxonomicName>
. Chief diagnostic characters are the tumid, medially depressed distal male ventrite and metaventrite, comparably long rostrum and legs and the non-confluent pronotal punctation (
<figureCitation id="22F62A7B1924FFC1FCCFAEC8FC19FD54" box="[831,969,727,753]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="17" pageNumber="18">Figs. 43, 44</figureCitation>
). See notes below.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751924FFC1FF37AEE5FEC7FBB4" pageId="17" pageNumber="18" type="description">
<paragraph id="BA7236FE1924FFC1FF37AEE5FEC7FBB4" blockId="17.[151,1437,654,2013]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37AEE5FE89FCB1" bold="true" box="[199,345,762,788]" pageId="17" pageNumber="18">Description.</emphasis>
Total length
<quantity id="7D359B1B1924FFC1FE02AEE4FDADFCB0" box="[498,637,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.8" metricValueMin="2.1" pageId="17" pageNumber="18" unit="mm" value="2.45" valueMax="2.8" valueMin="2.1">2.1–2.8 mm</quantity>
, width
<quantity id="7D359B1B1924FFC1FD24AEE4FC8DFCB0" box="[724,861,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="17" pageNumber="18" unit="mm" value="1.1" valueMax="1.3" valueMin="0.9">0.9–1.3 mm</quantity>
(n=51); integument reddish brown, body elongate ovate with moderately protruding humeri; rostrum nearly 3x as long as eye; pronotum comparatively large but not humped, with rather coarse, non-confluent punctation, interspaces not forming oblique ridges, flank with larger and more distant pits in smooth area; elytron 2.1x longer than pronotum, disc at base with transverse depression, interstriae ridged and with minute setae; metaventrite tumid in female, somewhat angularly produced and cleft by discrimen in male, male ventrite 5 tumid, 1, 2 and 5 with deep median depressions bordered by erect squamulae; profemur sublinear, 2x longer than head diameter; male genitalia as
<figureCitation id="22F62A7B1924FFC1FC78AFCCFC09FC48" box="[904,985,979,1005]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="17" pageNumber="18">Fig. 21</figureCitation>
, flagellum short, curved, with long basal appendage.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751924FFC1FF37A805FD79F8D4" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="BA7236FE1924FFC1FF37A805FE5EFA38" blockId="17.[151,1437,654,2013]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37A805FE61FB91" bold="true" box="[199,433,1050,1076]" pageId="17" pageNumber="18">Material examined.</emphasis>
<materialsCitation id="0AA53CA31924FFC1FE49A804FA49FBFC" collectingDate="1993-04-18" collectingDateMax="1993-05-19" collectingDateMin="1993-04-18" collectionCode="MNCR" collectorName="P. Rios" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="6576885C1924FFC1FE49A804FDF1FB90" box="[441,545,1051,1077]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
male, labeled “Est. Pitilla,
<quantity id="7D359B1B1924FFC1FCA6A804FC70FB90" box="[854,928,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="m" value="700.0">
<elevation id="31E0D1CD1924FFC1FCA6A804FC70FB90" box="[854,928,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<quantity id="7D359B1B1924FFC1FC5CA804FC36FB90" box="[940,998,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="km" value="9.0">9 km</quantity>
S
<location id="BF1260251924FFC1FBF4A804FB59FB90" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FBF4A804FB59FB90" box="[1028,1161,1051,1077]" country="Costa Rica" name="Sta. Cecilia" pageId="17" pageNumber="18" stateProvince="Guanacaste">Sta. Cecilia</location>
, P. N.
<collectingRegion id="7809F81C1924FFC1FB22A804FA86FB90" box="[1234,1366,1051,1077]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
, Prov. Guan.,
<collectingCountry id="C2DA766E1924FFC1FF19A820FE5FFBFC" box="[233,399,1087,1113]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
,
<date id="CE73103E1924FFC1FE6AA820FD44FBFC" box="[410,660,1087,1113]" pageId="17" pageNumber="18" value="1993-04-18" valueMax="1993-05-19" valueMin="1993-04-18">
<collectingDate id="DE37E9D61924FFC1FE6AA820FD44FBFC" box="[410,660,1087,1113]" pageId="17" pageNumber="18" value="1993-04-18" valueMax="1993-05-19" valueMin="1993-04-18">18 abr a 19 may 1993</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FD6FA820FD20FBFC" box="[671,752,1087,1113]" pageId="17" pageNumber="18">P. Ríos</collectorName>
, L-N-330200, 380200”, “INBIO CRI001 397023” (
<collectionCode id="DCDCAE3B1924FFC1FAC8A820FA41FBFC" box="[1336,1425,1087,1113]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
.
<typeStatus id="6576885C1924FFC1FF67A87CFED4FBD8" box="[151,260,1123,1149]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="ACCBFD771924FFC1FEE5A87CFEAFFBD8" box="[277,383,1123,1149]" count="32" pageId="17" pageNumber="18" type="male">32 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FE7DA87CFDDFFBD8" box="[397,527,1123,1149]" count="19" pageId="17" pageNumber="18" type="female">19 females</specimenCount>
):
<materialsCitation id="0AA53CA31924FFC1FDD3A87CFE99FB04" collectingDate="1993-04-18" collectingDateMax="1993-05-19" collectingDateMin="1993-04-18" collectionCode="MNCR" collectorName="P. Rios" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="17" pageNumber="18" specimenCount="4" specimenCount-male="4" stateProvince="Guanacaste" typeStatus="paratype">
same data as holotype, with barcodes CRI001 396769, 396773–74, 396878,
<specimenCount id="ACCBFD771924FFC1FA7EA87CFF08FB04" count="4" pageId="17" pageNumber="18" type="male">4 males</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FF18A898FE91FB04" box="[232,321,1159,1185]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31924FFC1FEA5A898FAE3FB04" box="[341,1331,1158,1185]" collectingDate="1991-11-04" collectingDateMax="1991-11-25" collectingDateMin="1991-11-04" collectionCode="MNCR" collectorName="Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
same locality as holotype,
<date id="CE73103E1924FFC1FD72A899FCF0FB04" box="[642,800,1158,1185]" pageId="17" pageNumber="18" value="1991-11-04" valueMax="1991-11-25" valueMin="1991-11-04">
<collectingDate id="DE37E9D61924FFC1FD72A899FCF0FB04" box="[642,800,1158,1185]" pageId="17" pageNumber="18" value="1991-11-04" valueMax="1991-11-25" valueMin="1991-11-04">4.–25.xi.1991</collectingDate>
</date>
, C.
<collectorName id="173853281924FFC1FCBDA898FC77FB04" box="[845,935,1159,1185]" pageId="17" pageNumber="18">Moraga</collectorName>
, CRI000 497788,
<specimenCount id="ACCBFD771924FFC1FB87A898FB13FB04" box="[1143,1219,1159,1185]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FB23A898FAFBFB04" box="[1235,1323,1159,1185]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FACFA898FCD5FB60" collectingDate="1992-08-24" collectingDateMax="1992-09-11" collectingDateMin="1992-08-24" collectionCode="MNCR" collectorName="Rios" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<date id="CE73103E1924FFC1FACFA898FEC1FB60" pageId="17" pageNumber="18" value="1992-08-24" valueMax="1992-09-11" valueMin="1992-08-24">
<collectingDate id="DE37E9D61924FFC1FACFA898FEC1FB60" pageId="17" pageNumber="18" value="1992-08-24" valueMax="1992-09-11" valueMin="1992-08-24">24.viii.– 11.ix.1992</collectingDate>
</date>
, P.
<collectorName id="173853281924FFC1FEC5A8B4FEBBFB60" box="[309,363,1195,1221]" pageId="17" pageNumber="18">Ríos</collectorName>
, CRI000 832161,
<specimenCount id="ACCBFD771924FFC1FDC6A8B4FD46FB60" box="[566,662,1195,1221]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FD54A8B4FD2CFB60" box="[676,764,1195,1221]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FCFFA8B4FAFEFB60" box="[783,1326,1195,1221]" collectingDate="1996-03-06" collectingDateMax="1996-03-12" collectingDateMin="1996-03-06" collectionCode="JPPC" collectorName="Prena" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<date id="CE73103E1924FFC1FCFFA8B4FC7EFB60" box="[783,942,1195,1221]" pageId="17" pageNumber="18" value="1996-03-06" valueMax="1996-03-12" valueMin="1996-03-06">
<collectingDate id="DE37E9D61924FFC1FCFFA8B4FC7EFB60" box="[783,942,1195,1221]" pageId="17" pageNumber="18" value="1996-03-06" valueMax="1996-03-12" valueMin="1996-03-06">6.–12.iii.1996</collectingDate>
</date>
, J.
<collectorName id="173853281924FFC1FC21A8B4FBC4FB60" box="[977,1044,1195,1221]" pageId="17" pageNumber="18">Prena</collectorName>
, 1 dissected male (
<collectionCode id="DCDCAE3B1924FFC1FB14A8B4FAF6FB60" box="[1252,1318,1195,1221]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FAC8A8B4FD65FB4C" collectingDate="1996-06" collectionCode="MNCR" collectorName="Moraga" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<date id="CE73103E1924FFC1FAC8A8B4FA48FB60" box="[1336,1432,1195,1221]" bridgedPair="/" pageId="17" pageNumber="18" value="1996-06">
<collectingDate id="DE37E9D61924FFC1FAC8A8B4FA48FB60" box="[1336,1432,1195,1221]" bridgedPair="/" pageId="17" pageNumber="18" value="1996-06">VI/1996</collectingDate>
</date>
, C.
<collectorName id="173853281924FFC1FF4CA8D0FEC6FB4C" box="[188,278,1231,1257]" pageId="17" pageNumber="18">Moraga</collectorName>
, CRI002 422519,
<specimenCount id="ACCBFD771924FFC1FE01A8D0FD90FB4C" box="[497,576,1231,1257]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FDA4A8D0FD7DFB4C" box="[596,685,1231,1257]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31924FFC1FD35A8D0FD4CFAA8" collectingDate="1990-06" collectionCode="MNCR" country="Costa Rica" location="Estac. Cacao" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
P.N.
<collectingRegion id="7809F81C1924FFC1FD0FA8D0FC53FB4C" box="[767,899,1231,1257]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
,
<location id="BF1260251924FFC1FC63A8D0FBFBFB4C" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FC63A8D0FBFBFB4C" box="[915,1067,1231,1257]" country="Costa Rica" name="Estac. Cacao" pageId="17" pageNumber="18" stateProvince="Guanacaste">Estac. Cacao</location>
, L-N-323300 375700,
<date id="CE73103E1924FFC1FAC8A8D0FA48FB4C" box="[1336,1432,1231,1257]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-06">
<collectingDate id="DE37E9D61924FFC1FAC8A8D0FA48FB4C" box="[1336,1432,1231,1257]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-06">VI/1990</collectingDate>
</date>
, CRI000 255832, 1 dissected male (
<collectionCode id="DCDCAE3B1924FFC1FDCBA8ECFD43FAA8" box="[571,659,1267,1293]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FD5BA8ECFA48FAA8" box="[683,1432,1267,1293]" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
VII–
<date id="CE73103E1924FFC1FD12A8ECFCEFFAA8" box="[738,831,1267,1293]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-03">
<collectingDate id="DE37E9D61924FFC1FD12A8ECFCEFFAA8" box="[738,831,1267,1293]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-03">III/1990</collectingDate>
</date>
,
<collectingMethod id="638C4EE91924FFC1FCBFA8ECFC0FFAA8" box="[847,991,1267,1293]" pageId="17" pageNumber="18">malaise trap</collectingMethod>
, CRI000 248710,
<specimenCount id="ACCBFD771924FFC1FB4CA8ECFAF3FAA8" box="[1212,1315,1267,1293]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FAC7A8ECFA40FAA8" box="[1335,1424,1267,1293]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31924FFC1FF67A908FAD2FA94" box="[151,1282,1303,1329]" collectingDate="1994-03" collectionCode="MNCR" collectorName="Obando" country="Costa Rica" location="Monteverde" municipality="Estac. La Casona" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="7809F81C1924FFC1FF67A908FEC5FA94" box="[151,277,1303,1329]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="18">Puntarenas</collectingRegion>
,
<collectingMunicipality id="5A16AC841924FFC1FED1A908FE38FA94" box="[289,488,1303,1329]" pageId="17" pageNumber="18">Estac. La Casona</collectingMunicipality>
,
<location id="BF1260251924FFC1FE04A908FDAEFA94" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FE04A908FDAEFA94" box="[500,638,1303,1329]" country="Costa Rica" municipality="Estac. La Casona" name="Monteverde" pageId="17" pageNumber="18" stateProvince="Puntarenas">Monteverde</location>
,
<date id="CE73103E1924FFC1FD7AA908FD39FA94" box="[650,745,1303,1329]" bridgedPair="/" pageId="17" pageNumber="18" value="1994-03">
<collectingDate id="DE37E9D61924FFC1FD7AA908FD39FA94" box="[650,745,1303,1329]" bridgedPair="/" pageId="17" pageNumber="18" value="1994-03">III/1994</collectingDate>
</date>
, N.
<collectorName id="173853281924FFC1FCE9A908FCA5FA94" box="[793,885,1303,1329]" pageId="17" pageNumber="18">Obando</collectorName>
, CRI001 780686,
<specimenCount id="ACCBFD771924FFC1FBB4A908FB40FA94" box="[1092,1168,1303,1329]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FB51A908FB2AFA94" box="[1185,1274,1303,1329]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31924FFC1FAFEA908FABCFAF0" collectingDate="2009-06-30" collectingDateMax="2009-07-09" collectingDateMin="2009-06-30" collectionCode="JPPC" collectorName="J. Prena" country="Costa Rica" elevation="1650" latitude="10.3147" location="Est. Biologica" longLatPrecision="7" longitude="-84.7969" municipality="Monteverde" pageId="17" pageNumber="18" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Puntarenas" typeStatus="paratype">
<collectingMunicipality id="5A16AC841924FFC1FAFEA908FA48FA94" box="[1294,1432,1303,1329]" pageId="17" pageNumber="18">Monteverde</collectingMunicipality>
,
<location id="BF1260251924FFC1FF67A924FEEBFAF0" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FF67A924FEEBFAF0" box="[151,315,1339,1365]" country="Costa Rica" latitude="10.3147" longLatPrecision="7" longitude="-84.7969" municipality="Monteverde" name="Est. Biologica" pageId="17" pageNumber="18" stateProvince="Puntarenas">Est. Biologica</location>
,
<geoCoordinate id="DFF950391924FFC1FEB8A924FE10FAF0" box="[328,448,1339,1365]" degrees="10.3147" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="5" value="10.3147">10.3147 N</geoCoordinate>
<geoCoordinate id="DFF950391924FFC1FE39A924FD97FAF0" box="[457,583,1339,1365]" degrees="84.7969" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="5" value="-84.7969">84.7969 W</geoCoordinate>
,
<quantity id="7D359B1B1924FFC1FDA4A925FD23FAF0" box="[596,755,1338,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="17" pageNumber="18" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<elevation id="31E0D1CD1924FFC1FDA4A925FD23FAF0" box="[596,755,1338,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="17" pageNumber="18" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1500–1800 m</elevation>
</quantity>
,
<date id="CE73103E1924FFC1FCF1A924FC13FAF0" box="[769,963,1339,1365]" pageId="17" pageNumber="18" value="2009-06-30" valueMax="2009-07-09" valueMin="2009-06-30">
<collectingDate id="DE37E9D61924FFC1FCF1A924FC13FAF0" box="[769,963,1339,1365]" pageId="17" pageNumber="18" value="2009-06-30" valueMax="2009-07-09" valueMin="2009-06-30">30.vi.–9.vii.2009</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FC20A924FBFEFAF0" box="[976,1070,1339,1365]" pageId="17" pageNumber="18">J. Prena</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FBCBA924FB46FAF0" box="[1083,1174,1339,1365]" count="8" pageId="17" pageNumber="18" type="male">8 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FB53A924FAC1FAF0" box="[1187,1297,1339,1365]" count="7" pageId="17" pageNumber="18" type="female">7 females</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FAD2A924FAB4FAF0" box="[1314,1380,1339,1365]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FA89A924FD7FFADC" collectingDate="2011-06-02" collectingDateMax="2011-06-07" collectingDateMin="2011-06-02" collectionCode="CMNC" collectorName="Prena" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<date id="CE73103E1924FFC1FA89A924FED2FADC" pageId="17" pageNumber="18" value="2011-06-02" valueMax="2011-06-07" valueMin="2011-06-02">
<collectingDate id="DE37E9D61924FFC1FA89A924FED2FADC" pageId="17" pageNumber="18" value="2011-06-02" valueMax="2011-06-07" valueMin="2011-06-02">2.– 7.vi.2011</collectingDate>
</date>
, J.
<collectorName id="173853281924FFC1FED5A940FEB8FADC" box="[293,360,1375,1401]" pageId="17" pageNumber="18">Prena</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FE82A940FE1BFADC" box="[370,459,1375,1401]" count="2" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FE25A940FD91FADC" box="[469,577,1375,1401]" count="2" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FDBFA940FD77FADC" box="[591,679,1375,1401]" pageId="17" pageNumber="18">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FD49A940FC3EFADC" box="[697,1006,1374,1401]" collectingDate="2011-06-02" collectingDateMax="2011-06-07" collectingDateMin="2011-06-02" collectionCode="JPPC" collectorName="Prena" country="Costa Rica" isTailEnumeration="true" pageId="17" pageNumber="18" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="ACCBFD771924FFC1FD49A940FCF0FADC" box="[697,800,1375,1401]" count="10" pageId="17" pageNumber="18" type="male">10 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FCDAA941FC46FADC" box="[810,918,1374,1401]" count="5" pageId="17" pageNumber="18" type="female">5 females</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FC54A940FC36FADC" box="[932,998,1375,1401]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FC09A940FAE5FADC" box="[1017,1333,1375,1401]" collectionCode="MNCR" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="ACCBFD771924FFC1FC09A940FB81FADC" box="[1017,1105,1375,1401]" count="2" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FBACA940FB17FADC" box="[1116,1223,1375,1401]" count="2" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FB25A940FAFDFADC" box="[1237,1325,1375,1401]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FAB0A940FE5BFA38" collectionCode="NHMUK" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="ACCBFD771924FFC1FAB0A940FA48FADC" box="[1344,1432,1375,1401]" count="2" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="ACCBFD771924FFC1FF67A99CFED3FA38" box="[151,259,1411,1437]" count="2" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FEE2A99CFE53FA38" box="[274,387,1411,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="18" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1924FFC1FF37A9B9FE7FF8A0" blockId="17.[151,1437,654,2013]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37A9B9FE6BFA65" bold="true" box="[199,443,1446,1472]" pageId="17" pageNumber="18">Additional material.</emphasis>
<materialsCitation id="0AA53CA31924FFC1FE34A9B8FE84FA40" collectingDate="1990-03" collectionCode="MNCR" country="Costa Rica" location="Rio San Lorencito" municipality="San Ramon" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela">
<collectingCountry id="C2DA766E1924FFC1FE34A9B8FDBFFA64" box="[452,623,1447,1473]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
.
<collectingRegion id="7809F81C1924FFC1FD8EA9B8FD31FA64" box="[638,737,1447,1473]" country="Costa Rica" name="Alajuela" pageId="17" pageNumber="18">Alajuela</collectingRegion>
:
<collectingMunicipality id="5A16AC841924FFC1FD1DA9B8FCA3FA64" box="[749,883,1447,1473]" pageId="17" pageNumber="18">San Ramón</collectingMunicipality>
,
<location id="BF1260251924FFC1FC8FA9B8FB82FA64" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FC8FA9B8FB82FA64" box="[895,1106,1447,1473]" country="Costa Rica" municipality="San Ramon" name="Rio San Lorencito" pageId="17" pageNumber="18" stateProvince="Alajuela">Río San Lorencito</location>
,
<date id="CE73103E1924FFC1FBAFA9B8FB6DFA64" box="[1119,1213,1447,1473]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-03">
<collectingDate id="DE37E9D61924FFC1FBAFA9B8FB6DFA64" box="[1119,1213,1447,1473]" bridgedPair="/" pageId="17" pageNumber="18" value="1990-03">III/1990</collectingDate>
</date>
, CRI000 157455,
<specimenCount id="ACCBFD771924FFC1FA7EA9B8FF33FA40" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FF03A9D4FE9CFA40" box="[243,332,1483,1509]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FE90A9D4FAD7FA40" box="[352,1287,1483,1509]" collectingDate="2000-04-07" collectingDateMax="2000-04-08" collectingDateMin="2000-04-07" collectionCode="JPPC" collectorName="P. N. Tapanti &amp; J. Prena" country="Costa Rica" location="Quebrada Segunda" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Cartago">
<collectingRegion id="7809F81C1924FFC1FE90A9D4FE6DFA40" box="[352,445,1483,1509]" country="Costa Rica" name="Cartago" pageId="17" pageNumber="18">Cartago</collectingRegion>
:
<collectorName id="173853281924FFC1FE39A9D4FD86FA40" box="[457,598,1483,1509]" pageId="17" pageNumber="18">P.N. Tapantí</collectorName>
,
<location id="BF1260251924FFC1FD92A9D4FCEAFA40" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FD92A9D4FCEAFA40" box="[610,826,1483,1509]" country="Costa Rica" name="Quebrada Segunda" pageId="17" pageNumber="18" stateProvince="Cartago">Quebrada Segunda</location>
,
<date id="CE73103E1924FFC1FCB6A9D4FC04FA40" box="[838,980,1483,1509]" pageId="17" pageNumber="18" value="2000-04-07" valueMax="2000-04-08" valueMin="2000-04-07">
<collectingDate id="DE37E9D61924FFC1FCB6A9D4FC04FA40" box="[838,980,1483,1509]" pageId="17" pageNumber="18" value="2000-04-07" valueMax="2000-04-08" valueMin="2000-04-07">7.–8.iv.2000</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FC11A9D4FBEEFA40" box="[993,1086,1483,1509]" pageId="17" pageNumber="18">J. Prena</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FBBAA9D4FB7CFA40" box="[1098,1196,1483,1509]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FB4CA9D4FB2FFA40" box="[1212,1279,1483,1509]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FAE3A9D4FB0CF9AC" collectingDate="2000-09-28" collectionCode="MNCR" collectorName="P. N. Hitoy Cerere &amp; W. Arana" country="Costa Rica" location="Rio Moin" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Limon">
<collectingRegion id="7809F81C1924FFC1FAE3A9D4FAB2FA40" box="[1299,1378,1483,1509]" country="Costa Rica" name="Limon" pageId="17" pageNumber="18">Limón</collectingRegion>
:
<collectorName id="173853281924FFC1FA9EA9D4FEFBF9AC" pageId="17" pageNumber="18">P.N. Hitoy Cerere</collectorName>
, sendero hacia
<location id="BF1260251924FFC1FE23A9F0FD90F9AC" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FE23A9F0FD90F9AC" box="[467,576,1519,1545]" country="Costa Rica" name="Rio Moin" pageId="17" pageNumber="18" stateProvince="Limon">Río Moín</location>
,
<date id="CE73103E1924FFC1FDBBA9F0FD15F9AC" box="[587,709,1519,1545]" pageId="17" pageNumber="18" value="2000-09-28">
<collectingDate id="DE37E9D61924FFC1FDBBA9F0FD15F9AC" box="[587,709,1519,1545]" pageId="17" pageNumber="18" value="2000-09-28">28.ix.2000</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FD3FA9F0FCE9F9AC" box="[719,825,1519,1545]" pageId="17" pageNumber="18">W. Arana</collectorName>
, INB0003337173,
<specimenCount id="ACCBFD771924FFC1FBFDA9F0FBBDF9AC" box="[1037,1133,1519,1545]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FB8BA9F0FB04F9AC" box="[1147,1236,1519,1545]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="17" pageNumber="18">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FB15A9F0FCA9F988" collectingDate="2004-05-10" collectionCode="JPPC" collectorName="P. N. Corcovado &amp; J. Prena" country="Costa Rica" location="Estac. Los Patos" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<collectingRegion id="7809F81C1924FFC1FB15A9F0FAB4F9AC" box="[1253,1380,1519,1545]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="18">Puntarenas</collectingRegion>
:
<collectorName id="173853281924FFC1FA9EA9F0FEC5F988" pageId="17" pageNumber="18">P.N. Corcovado</collectorName>
,
<location id="BF1260251924FFC1FEEFAA0CFE07F988" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FEEFAA0CFE07F988" box="[287,471,1555,1581]" country="Costa Rica" name="Estac. Los Patos" pageId="17" pageNumber="18" stateProvince="Puntarenas">Estac. Los Patos</location>
,
<date id="CE73103E1924FFC1FE11AA0CFD82F988" box="[481,594,1555,1581]" pageId="17" pageNumber="18" value="2004-05-10">
<collectingDate id="DE37E9D61924FFC1FE11AA0CFD82F988" box="[481,594,1555,1581]" pageId="17" pageNumber="18" value="2004-05-10">10.v.2004</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FDACAA0CFD67F988" box="[604,695,1555,1581]" pageId="17" pageNumber="18">J. Prena</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FD30AA0CFCF0F988" box="[704,800,1555,1581]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FCDEAA0CFCA1F988" box="[814,881,1555,1581]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FC73AA0CFCBAF9F4" collectingDate="2009-05-29" collectionCode="CMNC" country="Guatemala" location="13 km NW Machaquila" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Peten">
<collectingCountry id="C2DA766E1924FFC1FC73AA0CFBE9F988" box="[899,1081,1555,1581]" name="Guatemala" pageId="17" pageNumber="18">GUATEMALA</collectingCountry>
.
<collectingRegion id="7809F81C1924FFC1FBB6AA0CFB59F988" box="[1094,1161,1555,1581]" country="Guatemala" name="Peten" pageId="17" pageNumber="18">Petén</collectingRegion>
:
<location id="BF1260251924FFC1FB62AA0CFA48F988" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FB62AA0CFA48F988" box="[1170,1432,1555,1581]" country="Guatemala" name="13 km NW Machaquila" pageId="17" pageNumber="18" stateProvince="Peten">
<quantity id="7D359B1B1924FFC1FB62AA0CFB08F988" box="[1170,1240,1555,1581]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="18" unit="km" value="13.0">13 km</quantity>
NW Machaquilá
</location>
,
<date id="CE73103E1924FFC1FF67AA28FED7F9F4" box="[151,263,1591,1617]" pageId="17" pageNumber="18" value="2009-05-29">
<collectingDate id="DE37E9D61924FFC1FF67AA28FED7F9F4" box="[151,263,1591,1617]" pageId="17" pageNumber="18" value="2009-05-29">29.v.2009</collectingDate>
</date>
, LLAMA #Go-B-06-3-02, 1 dissected male (
<collectionCode id="DCDCAE3B1924FFC1FCFAAA28FCB3F9F4" box="[778,867,1591,1617]" pageId="17" pageNumber="18">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FC87AA28FDC9F9D0" collectingDate="1995-06-15" collectionCode="ASUCOB" collectorName="P. N. Cusuco &amp; R. Turnbow" country="Honduras" location="Cortes" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Cortes">
<collectingCountry id="C2DA766E1924FFC1FC87AA28FBC9F9F4" box="[887,1049,1591,1617]" name="Honduras" pageId="17" pageNumber="18">HONDURAS</collectingCountry>
.
<collectingRegion id="7809F81C1924FFC1FBD9AA28FBA6F9F4" box="[1065,1142,1591,1617]" country="Honduras" name="Cortes" pageId="17" pageNumber="18">Cortés</collectingRegion>
:
<collectorName id="173853281924FFC1FB73AA28FAC1F9F4" box="[1155,1297,1591,1617]" pageId="17" pageNumber="18">P.N. Cusuco</collectorName>
,
<date id="CE73103E1924FFC1FAEDAA29FA47F9F4" box="[1309,1431,1590,1617]" pageId="17" pageNumber="18" value="1995-06-15">
<collectingDate id="DE37E9D61924FFC1FAEDAA29FA47F9F4" box="[1309,1431,1590,1617]" pageId="17" pageNumber="18" value="1995-06-15">15.vi.1995</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FF67AA44FECEF9D0" box="[151,286,1627,1653]" pageId="17" pageNumber="18">R. Turnbow</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FED9AA44FE5AF9D0" box="[297,394,1627,1653]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FE69AA44FDC0F9D0" box="[409,528,1627,1653]" pageId="17" pageNumber="18">ASUCOB</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31924FFC1FDD3AA44FBC6F9D0" box="[547,1046,1627,1653]" collectingDate="1997-03-08" collectionCode="JPPC" collectorName="Prena" country="Honduras" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Cortes">
<date id="CE73103E1924FFC1FDD3AA44FD41F9D0" box="[547,657,1627,1653]" pageId="17" pageNumber="18" value="1997-03-08">
<collectingDate id="DE37E9D61924FFC1FDD3AA44FD41F9D0" box="[547,657,1627,1653]" pageId="17" pageNumber="18" value="1997-03-08">8.iii.1997</collectingDate>
</date>
, J.
<collectorName id="173853281924FFC1FD46AA44FD29F9D0" box="[694,761,1627,1653]" pageId="17" pageNumber="18">Prena</collectorName>
, 1 dissected male (
<collectionCode id="DCDCAE3B1924FFC1FC3BAA44FBDEF9D0" box="[971,1038,1627,1653]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31924FFC1FBD2AA44FDC6F93C" collectingDate="2001-07-24" collectionCode="ASUCOB" collectorName="R. Turnbow" country="Honduras" location="Sierra del Merendon" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Cortes">
<location id="BF1260251924FFC1FBD2AA44FADBF9D0" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FBD2AA44FADBF9D0" box="[1058,1291,1627,1653]" country="Honduras" name="Sierra del Merendon" pageId="17" pageNumber="18" stateProvince="Cortes">Sierra del Merendón</location>
,
<date id="CE73103E1924FFC1FAE6AA44FA47F9D0" box="[1302,1431,1627,1653]" pageId="17" pageNumber="18" value="2001-07-24">
<collectingDate id="DE37E9D61924FFC1FAE6AA44FA47F9D0" box="[1302,1431,1627,1653]" pageId="17" pageNumber="18" value="2001-07-24">24.vii.2001</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FF67AA60FECDF93C" box="[151,285,1663,1689]" pageId="17" pageNumber="18">R. Turnbow</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FED7AA60FE58F93C" box="[295,392,1663,1689]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FE66AA60FDDEF93C" box="[406,526,1663,1689]" pageId="17" pageNumber="18">ASUCOB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FDD0AA9FFC56F93C" box="[544,902,1663,1691]" collectorName="Isla" country="Panama" location="Bocas del Toro" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Bocas del Toro">
<collectingCountry id="C2DA766E1924FFC1FDD0AA9FFD70F93E" box="[544,672,1664,1691]" name="Panama" pageId="17" pageNumber="18">PANAMÁ</collectingCountry>
.
<collectingRegion id="7809F81C1924FFC1FD59AA60FC84F93C" box="[681,852,1663,1689]" country="Panama" name="Bocas del Toro" pageId="17" pageNumber="18">Bocas del Toro</collectingRegion>
:
<collectorName id="173853281924FFC1FCAEAA60FC56F93C" box="[862,902,1663,1689]" pageId="17" pageNumber="18">Isla</collectorName>
</materialsCitation>
<materialsCitation id="0AA53CA31924FFC1FC7CAA60FED0F918" collectingDate="2004-02-26" collectionCode="CMNC" collectorName="H. Stockwell" country="Panama" location="Colon" pageId="17" pageNumber="18" specimenCount="2" stateProvince="Colon">
<collectingRegion id="7809F81C1924FFC1FC7CAA60FC03F93C" box="[908,979,1663,1689]" country="Panama" name="Colon" pageId="17" pageNumber="18">Colón</collectingRegion>
,
<date id="CE73103E1924FFC1FC2EAA60FBBBF93C" box="[990,1131,1663,1689]" pageId="17" pageNumber="18" value="2004-02-26">
<collectingDate id="DE37E9D61924FFC1FC2EAA60FBBBF93C" box="[990,1131,1663,1689]" pageId="17" pageNumber="18" value="2004-02-26">26 Feb 2004</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FB85AA60FAD9F93C" box="[1141,1289,1663,1689]" pageId="17" pageNumber="18">H. Stockwell</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FAE3AA60FA4DF93C" box="[1299,1437,1663,1689]" count="2" pageId="17" pageNumber="18" type="generic">2 specimens</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FF6FAABCFF28F918" box="[159,248,1699,1725]" pageId="17" pageNumber="18">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FEFBAABCFBEFF918" box="[267,1087,1699,1725]" collectingDate="1970-04-23" collectionCode="JPPC" collectorName="H. Hespenheide" country="Panama" location="Madden Forest" municipality="Canal Area" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Colon">
<collectingMunicipality id="5A16AC841924FFC1FEFBAABCFE5EF918" box="[267,398,1699,1725]" pageId="17" pageNumber="18">Canal Area</collectingMunicipality>
:
<location id="BF1260251924FFC1FE69AABCFD95F918" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FE69AABCFD95F918" box="[409,581,1699,1725]" country="Panama" municipality="Canal Area" name="Madden Forest" pageId="17" pageNumber="18" stateProvince="Colon">Madden Forest</location>
,
<date id="CE73103E1924FFC1FDA1AABCFD19F918" box="[593,713,1699,1725]" pageId="17" pageNumber="18" value="1970-04-23">
<collectingDate id="DE37E9D61924FFC1FDA1AABCFD19F918" box="[593,713,1699,1725]" pageId="17" pageNumber="18" value="1970-04-23">23.iv.1970</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FD26AABCFC5DF918" box="[726,909,1699,1725]" pageId="17" pageNumber="18">H. Hespenheide</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FC68AABCFC34F918" box="[920,996,1699,1725]" count="1" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FC04AABCFBE7F918" box="[1012,1079,1699,1725]" pageId="17" pageNumber="18">JPPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FBBAAABCFCA7F944" collectingDate="1995-05-26" collectingDateMax="1995-06-09" collectingDateMin="1995-05-26" collectionCode="CMNC" collectorName="F. I. T., S. Ashe &amp; R. Brooks" country="Panama" location="Reserva La Fortuna" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui">
<collectingRegion id="7809F81C1924FFC1FBBAAABCFB7CF918" box="[1098,1196,1699,1725]" country="Panama" name="Chiriqui" pageId="17" pageNumber="18">Chiriquí</collectingRegion>
:
<location id="BF1260251924FFC1FB47AABCFA48F918" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FB47AABCFA48F918" box="[1207,1432,1699,1725]" country="Panama" name="Reserva La Fortuna" pageId="17" pageNumber="18" stateProvince="Chiriqui">Reserva La Fortuna</location>
,
<date id="CE73103E1924FFC1FF67AAD9FE98F944" box="[151,328,1734,1761]" pageId="17" pageNumber="18" value="1995-05-26" valueMax="1995-06-09" valueMin="1995-05-26">
<collectingDate id="DE37E9D61924FFC1FF67AAD9FE98F944" box="[151,328,1734,1761]" pageId="17" pageNumber="18" value="1995-05-26" valueMax="1995-06-09" valueMin="1995-05-26">26.v.–9.vi.1995</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FEA5AAD8FE28F944" box="[341,504,1735,1761]" pageId="17" pageNumber="18">F.I.T., S. Ashe</collectorName>
&amp;
<collectorName id="173853281924FFC1FDEFAAD8FD45F944" box="[543,661,1735,1761]" pageId="17" pageNumber="18">R. Brooks</collectorName>
,
<specimenCount id="ACCBFD771924FFC1FD52AAD8FCD5F944" box="[674,773,1735,1761]" count="1" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1924FFC1FCE6AAD8FCBFF944" box="[790,879,1735,1761]" pageId="17" pageNumber="18">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31924FFC1FC73AAD8FE7BF8A0" collectingDate="1996-05-21" collectionCode="CMNC" collectorName="R. Turnbow" country="Panama" location="Cerro Azul" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Panama">
<collectingRegion id="7809F81C1924FFC1FC73AAD8FC30F944" box="[899,992,1735,1761]" country="Panama" name="Panama" pageId="17" pageNumber="18">Panamá</collectingRegion>
:
<location id="BF1260251924FFC1FC1DAAD8FBBDF944" LSID="urn:lsid:plazi:treatment:326487E81924FFC2FF67ADE1FE1DFEC0:BF1260251924FFC1FC1DAAD8FBBDF944" box="[1005,1133,1735,1761]" country="Panama" name="Cerro Azul" pageId="17" pageNumber="18" stateProvince="Panama">Cerro Azul</location>
,
<date id="CE73103E1924FFC1FB8AAAD8FB3AF944" box="[1146,1258,1735,1761]" pageId="17" pageNumber="18" value="1996-05-21">
<collectingDate id="DE37E9D61924FFC1FB8AAAD8FB3AF944" box="[1146,1258,1735,1761]" pageId="17" pageNumber="18" value="1996-05-21">21.v.1996</collectingDate>
</date>
,
<collectorName id="173853281924FFC1FB08AAD8FA51F944" box="[1272,1409,1735,1761]" pageId="17" pageNumber="18">R. Turnbow</collectorName>
, 1 dissected male (
<collectionCode id="DCDCAE3B1924FFC1FEBAAAF4FE73F8A0" box="[330,419,1771,1797]" pageId="17" pageNumber="18">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1924FFC1FF37AB11FD79F8D4" blockId="17.[151,1437,654,2013]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37AB11FE8AF88D" bold="true" box="[199,346,1806,1832]" pageId="17" pageNumber="18">Life history.</emphasis>
Specimens were collected on two occasions from the same two flowering
<taxonomicName id="7DCD4D7D1924FFC1FB48AB10FAC5F88D" box="[1208,1301,1807,1832]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="88B9EAEC1924FFC1FB48AB10FAC5F88D" box="[1208,1301,1807,1832]" italics="true" pageId="17" pageNumber="18">Miconia</emphasis>
</taxonomicName>
shrubs in a dense secondary forest at Monteverde. Each time, they occurred together with numerous
<taxonomicName id="7DCD4D7D1924FFC1FB60AB2CFB2EF8E9" authorityName="Prena" authorityYear="2024" box="[1168,1278,1843,1869]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC1924FFC1FB60AB2CFB2EF8E9" box="[1168,1278,1843,1869]" italics="true" pageId="17" pageNumber="18">C. montyi</emphasis>
</taxonomicName>
and, in 2009, with a single female of a still unnamed species.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751924FFC2FF37AB64FE1DFEC0" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="etymology">
<paragraph id="BA7236FE1924FFC1FF37AB64FACAF830" blockId="17.[151,1437,654,2013]" box="[199,1306,1915,1941]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37AB64FE9EF830" bold="true" box="[199,334,1915,1941]" pageId="17" pageNumber="18">Etymology.</emphasis>
The name is a Latin noun in apposition referring to the tumid fifth abdominal ventrite.
</paragraph>
<paragraph id="BA7236FE1924FFC2FF37AB80FE1DFEC0" blockId="17.[151,1437,654,2013]" lastBlockId="18.[151,1436,151,357]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37AB80FEC1F81C" bold="true" box="[199,273,1951,1977]" pageId="17" pageNumber="18">Notes.</emphasis>
<taxonomicName id="7DCD4D7D1924FFC1FEE6AB80FE19F81C" authorityName="Prena" authorityYear="2024" box="[278,457,1951,1977]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1924FFC1FEE6AB80FE19F81C" box="[278,457,1951,1977]" italics="true" pageId="17" pageNumber="18">Chryasus clunis</emphasis>
</taxonomicName>
is one of several local populations with uncertain status, with the male genitalia differing from site to site. I apply the name to Costa Rican collections from mid elevations in the Cordillera de Guanacaste (
<figureCitation id="22F62A7B1927FFC2FF6FAC88FF23FF14" box="[159,243,151,177]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="18" pageNumber="19">Fig. 43</figureCitation>
) and include also a series of slightly more slender specimens with a wider penis from the Cordillera de Tilarán (
<figureCitation id="22F62A7B1927FFC2FF08ACA4FE9FFF70" box="[248,335,187,213]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="18" pageNumber="19">Fig. 44</figureCitation>
). The few available males from
<collectingCountry id="C2DA766E1927FFC2FD32ACA4FCEBFF70" box="[706,827,187,213]" name="Guatemala" pageId="18" pageNumber="19">Guatemala</collectingCountry>
and
<collectingCountry id="C2DA766E1927FFC2FC86ACA4FC33FF70" box="[886,995,187,213]" name="Honduras" pageId="18" pageNumber="19">Honduras</collectingCountry>
(
<figureCitation id="22F62A7B1927FFC2FC04ACA4FB98FF70" box="[1012,1096,186,213]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="18" pageNumber="19">Fig. 45</figureCitation>
) and from
<collectingCountry id="C2DA766E1927FFC2FB3AACA4FAF3FF70" box="[1226,1315,187,213]" name="Panama" pageId="18" pageNumber="19">Panamá</collectingCountry>
each have the distal ventrite less tumid, the body of the penis differently shaped and a longer though basally similarly curved flagellum (
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). I have seen a photo of a male specimen from
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,
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(TAMU) that seems to belong to this complex as well. Reasonably representative collections of male
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and
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<emphasis id="88B9EAEC1927FFC2FAFAAD38FA4CFEE4" box="[1290,1436,295,321]" italics="true" pageId="18" pageNumber="19">C. persimilis</emphasis>
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exhibit much less variation.
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18.
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<emphasis id="88B9EAEC1927FFC2FF33AA0DFE03F988" bold="true" box="[195,467,1554,1581]" italics="true" pageId="18" pageNumber="19">Chryasus sculpturatus</emphasis>
(Casey)
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,
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<paragraph id="BA7236FE1927FFC2FF67AA28FD17F9F4" blockId="18.[151,797,1554,1617]" box="[151,711,1591,1617]" pageId="18" pageNumber="19">(Figs. available at https://collections.nmnh.si.edu)</paragraph>
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<emphasis id="88B9EAEC1927FFC2FF67AA64FEB1F936" box="[151,353,1659,1683]" italics="true" pageId="18" pageNumber="19">Aniops sculpturatus</emphasis>
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and 319, key and description).
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<emphasis id="88B9EAEC1927FFC2FF67AADEFEC2F97E" bold="true" box="[151,274,1729,1755]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
With a length of
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,
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is among the smallest species of the genus, all of which have conspicuous squamulae and femora with barely perceptible denticles. It is most similar to four unidentified specimens from
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in my collection (
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).
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<emphasis id="88B9EAEC1927FFC2FF37AB32FE63F8E2" bold="true" box="[199,435,1837,1863]" pageId="18" pageNumber="19">Material examined.</emphasis>
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.
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:
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, 1882–1886,
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, male
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(
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.
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<emphasis id="88B9EAEC1927FFC2FF37AB6AFEC1F82A" bold="true" box="[199,273,1909,1935]" pageId="18" pageNumber="19">Notes.</emphasis>
This specimen is another illegal exportation by H.H. Smith (see notes under 15.
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<emphasis id="88B9EAEC1927FFC2FB6AAB69FAD7F82A" box="[1178,1287,1909,1936]" italics="true" pageId="18" pageNumber="19">C. densus</emphasis>
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<treatment id="326487E81927FFC2FF67ADA9FD76FA64" LSID="urn:lsid:zoobank.org:act:CC09D1B3-9FFE-4F8C-9EB7-89E95A43BA2F" httpUri="http://treatment.plazi.org/id/326487E81927FFC2FF67ADA9FD76FA64" lastPageNumber="19" pageId="18" pageNumber="19">
<subSubSection id="F2D765751927FFC2FF67ADA9FD72FE75" box="[151,674,438,465]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="BA7236FE1927FFC2FF67ADA9FD72FE75" blockId="18.[151,1192,438,537]" box="[151,674,438,465]" pageId="18" pageNumber="19">
<heading id="E13A81921927FFC2FF67ADA9FD72FE75" bold="true" box="[151,674,438,465]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="88B9EAEC1927FFC2FF67ADA9FD72FE75" bold="true" box="[151,674,438,465]" pageId="18" pageNumber="19">
17.
<taxonomicName id="7DCD4D7D1927FFC2FF33ADA9FDD5FE74" authority="Prena" authorityName="Prena" authorityYear="2024" box="[195,517,438,465]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="persimilis" status="sp. nov.">
<emphasis id="88B9EAEC1927FFC2FF33ADA9FE62FE74" bold="true" box="[195,434,438,465]" italics="true" pageId="18" pageNumber="19">Chryasus persimilis</emphasis>
Prena
</taxonomicName>
,
<taxonomicNameLabel id="938A57971927FFC2FDE2ADA8FD72FE75" box="[530,674,438,465]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D765751927FFC2FF67ADC4FB78FDBC" pageId="18" pageNumber="19" type="description">
<paragraph id="BA7236FE1927FFC2FF67ADC4FE9AFE50" blockId="18.[151,1192,438,537]" box="[151,330,475,501]" pageId="18" pageNumber="19">
(
<figureCitation id="22F62A7B1927FFC2FF6FADC4FF3FFE50" box="[159,239,475,501]" captionStart="FIGURES 5–10" captionStartId="4.[152,270,1650,1674]" captionTargetBox="[151,1435,181,1603]" captionTargetId="figure-26@4.[151,1435,181,1626]" captionTargetPageId="4" captionText="FIGURES 5–10. Right hind legs showing gradient from clavate femur with minuscule, seta-bearing ventral denticles toward ventrally nearly straight femur with row of gradually enlarged teeth. 5, C. bicolor (type species of Preglyptobaris and subjective synonym of type species of Prosaldius); 6, C. cavernosus (type species of Chryasus); 7, C. persimilis (agreeing with Psiona); 8, 2.0 mm long species near C. carinulosus (type species of Psiona) and C. persimilis from Trinidad and Tobago; 9–10, 3.0 mm long species from São Paulo (Fig. 47) agreeing with Aniops, distal and proximal aspects. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13211505" httpUri="https://zenodo.org/record/13211505/files/figure.png" pageId="18" pageNumber="19">Figs. 7</figureCitation>
,
<figureCitation id="22F62A7B1927FFC2FF0BADC4FEC9FE50" box="[251,281,475,501]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="18" pageNumber="19">24</figureCitation>
,
<figureCitation id="22F62A7B1927FFC2FED5ADC4FE92FE50" box="[293,322,475,501]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="18" pageNumber="19">46</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE1927FFC2FF67ADE1FB78FDBC" blockId="18.[151,1192,438,537]" box="[151,1192,510,537]" pageId="18" pageNumber="19">
http://zoobank.org/
<uri id="CE5C3AFC1927FFC2FE85ADE1FB78FDBC" box="[373,1192,510,537]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="CE6B0C2B1927FFC2FD78ADE1FB78FDBC" box="[648,1192,510,537]" pageId="18" pageNumber="19">CC09D1B3-9FFE-4F8C-9EB7-89E95A43BA2F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="F2D765751927FFC2FF67AE59FF2FFD54" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="BA7236FE1927FFC2FF67AE59FF2FFD54" blockId="18.[151,1437,582,1473]" pageId="18" pageNumber="19">
<emphasis id="88B9EAEC1927FFC2FF67AE59FEC2FDC5" bold="true" box="[151,274,582,608]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="7DCD4D7D1927FFC2FD8AAE58FCCFFDC4" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[634,799,583,609]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="carinulosus">
<emphasis id="88B9EAEC1927FFC2FD8AAE58FCCFFDC4" box="[634,799,583,609]" italics="true" pageId="18" pageNumber="19">C. carinulosus</emphasis>
</taxonomicName>
,
<taxonomicName id="7DCD4D7D1927FFC2FCDEAE58FC4DFDC5" authorityName="Prena" authorityYear="2024" box="[814,925,583,609]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC1927FFC2FCDEAE58FC4DFDC5" box="[814,925,583,609]" italics="true" pageId="18" pageNumber="19">C. montyi</emphasis>
</taxonomicName>
and
<taxonomicName id="7DCD4D7D1927FFC2FC26AE58FB8CFDC5" authorityName="Prena" authorityYear="2024" box="[982,1116,583,609]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="taironius">
<emphasis id="88B9EAEC1927FFC2FC26AE58FB8CFDC5" box="[982,1116,583,609]" italics="true" pageId="18" pageNumber="19">C. taironius</emphasis>
</taxonomicName>
. They all have similar body proportions and similar male genitalia (unknown for
<taxonomicName id="7DCD4D7D1927FFC2FD2DAE74FCAFFD20" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[733,895,619,645]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="carinulosus">
<emphasis id="88B9EAEC1927FFC2FD2DAE74FCAFFD20" box="[733,895,619,645]" italics="true" pageId="18" pageNumber="19">C. carinulosus</emphasis>
</taxonomicName>
).
<taxonomicName id="7DCD4D7D1927FFC2FC64AE74FB97FD20" authorityName="Prena" authorityYear="2024" box="[916,1095,619,645]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1927FFC2FC64AE74FB97FD20" box="[916,1095,619,645]" italics="true" pageId="18" pageNumber="19">Chryasus clunis</emphasis>
</taxonomicName>
is another superficially similar species with the same color, but it is less elongate and the male has a tumid distal ventrite.
<taxonomicName id="7DCD4D7D1927FFC2FB7FAE90FABEFD0C" authorityName="Prena" authorityYear="2024" box="[1167,1390,655,681]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="88B9EAEC1927FFC2FB7FAE90FABEFD0C" box="[1167,1390,655,681]" italics="true" pageId="18" pageNumber="19">Chryasus persimilis</emphasis>
</taxonomicName>
can be distinguished from all these by its stout rostrum and femora, which measure distinctly less than twice the head diameter.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751927FFC2FF37AEE5FDBEFC48" pageId="18" pageNumber="19" type="description">
<paragraph id="BA7236FE1927FFC2FF37AEE5FDBEFC48" blockId="18.[151,1437,582,1473]" pageId="18" pageNumber="19">
<emphasis id="88B9EAEC1927FFC2FF37AEE5FE89FCB1" bold="true" box="[199,345,762,788]" pageId="18" pageNumber="19">Description.</emphasis>
Total length
<quantity id="7D359B1B1927FFC2FE1FAEE5FDA9FCB0" box="[495,633,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="18" pageNumber="19" unit="mm" value="2.2" valueMax="2.5" valueMin="1.9">1.9–2.5 mm</quantity>
, width
<quantity id="7D359B1B1927FFC2FD3EAEE4FC86FCB0" box="[718,854,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="19" unit="mm" value="0.95" valueMax="1.1" valueMin="0.8">0.8–1.1 mm</quantity>
(n=24); reddish brown, elytron somewhat brighter; body oblong ovate with protruding humeri and curved sides; rostrum 2x as long as eye; pronotum nearly as long as wide, interspaces forming indistinct oblique ridges between non-confluent punctation, flank with larger and more distant pits in smooth area; elytron 2.1x longer than pronotum, disc at base with very subtle transverse depression, interstriae ridged, metaventrite neither modified nor sexually dimorphic, male ventrites 1, 2 and 5 depressed medially and with erect squamulae; femora with 1 distinct and several minute denticles (
<figureCitation id="22F62A7B1927FFC2FBD0AFB0FBB5FC6C" box="[1056,1125,943,969]" captionStart="FIGURES 5–10" captionStartId="4.[152,270,1650,1674]" captionTargetBox="[151,1435,181,1603]" captionTargetId="figure-26@4.[151,1435,181,1626]" captionTargetPageId="4" captionText="FIGURES 5–10. Right hind legs showing gradient from clavate femur with minuscule, seta-bearing ventral denticles toward ventrally nearly straight femur with row of gradually enlarged teeth. 5, C. bicolor (type species of Preglyptobaris and subjective synonym of type species of Prosaldius); 6, C. cavernosus (type species of Chryasus); 7, C. persimilis (agreeing with Psiona); 8, 2.0 mm long species near C. carinulosus (type species of Psiona) and C. persimilis from Trinidad and Tobago; 9–10, 3.0 mm long species from São Paulo (Fig. 47) agreeing with Aniops, distal and proximal aspects. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13211505" httpUri="https://zenodo.org/record/13211505/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
); male genitalia as
<figureCitation id="22F62A7B1927FFC2FAB4AFB0FA47FC6C" box="[1348,1431,943,969]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="18" pageNumber="19">Fig. 24</figureCitation>
, endophallus with long, flexible flagellum.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751927FFC2FF37AFE9FD2AFA38" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="BA7236FE1927FFC2FF37AFE9FE49FAF0" blockId="18.[151,1437,582,1473]" pageId="18" pageNumber="19">
<emphasis id="88B9EAEC1927FFC2FF37AFE9FE62FBB5" bold="true" box="[199,434,1014,1040]" pageId="18" pageNumber="19">Material examined.</emphasis>
<materialsCitation id="0AA53CA31927FFC2FE4AAFE8FAF5FB90" collectingDate="1991-07" collectionCode="MNCR" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="6576885C1927FFC2FE4AAFE8FDF2FBB4" box="[442,546,1015,1041]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
male, labeled “Est. Pitilla,
<quantity id="7D359B1B1927FFC2FCAAAFE8FC74FBB4" box="[858,932,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="m" value="700.0">
<elevation id="31E0D1CD1927FFC2FCAAAFE8FC74FBB4" box="[858,932,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<quantity id="7D359B1B1927FFC2FC40AFE8FC3BFBB4" box="[944,1003,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="19" unit="km" value="9.0">9 km</quantity>
S
<location id="BF1260251927FFC2FBFAAFE8FB5FFBB4" LSID="urn:lsid:plazi:treatment:326487E81927FFC2FF67ADA9FD76FA64:BF1260251927FFC2FBFAAFE8FB5FFBB4" box="[1034,1167,1015,1041]" country="Costa Rica" name="Sta. Cecilia" pageId="18" pageNumber="19" stateProvince="Guanacaste">Sta. Cecilia</location>
, P.N.
<collectingRegion id="7809F81C1927FFC2FB22AFE8FA86FBB4" box="[1234,1366,1015,1041]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="19">Guanacaste</collectingRegion>
, Prov. Guan.,
<collectingCountry id="C2DA766E1927FFC2FF19A804FE5EFB90" box="[233,398,1051,1077]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
,
<collectorName id="173853281927FFC2FE69A804FDC3FB90" box="[409,531,1051,1077]" pageId="18" pageNumber="19">C. Moraga</collectorName>
,
<date id="CE73103E1927FFC2FDEEA804FD50FB90" box="[542,640,1051,1077]" pageId="18" pageNumber="19" value="1991-07">
<collectingDate id="DE37E9D61927FFC2FDEEA804FD50FB90" box="[542,640,1051,1077]" pageId="18" pageNumber="19" value="1991-07">Jul 1991</collectingDate>
</date>
, L-N-330200, 380200”, “INBIO CRI000 358585” (
<collectionCode id="DCDCAE3B1927FFC2FB34A804FACDFB90" box="[1220,1309,1051,1077]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FADFA804FD9BFBFC" country="Costa Rica" location="COSTA RICA" pageId="18" pageNumber="19" specimenCount="23" specimenCount-female="5" specimenCount-male="18" typeStatus="paratype">
<typeStatus id="6576885C1927FFC2FADFA804FA4CFB90" box="[1327,1436,1051,1077]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="ACCBFD771927FFC2FF50A820FEDAFBFC" box="[160,266,1087,1113]" count="18" pageId="18" pageNumber="19" type="male">18 males</specimenCount>
,
<specimenCount id="ACCBFD771927FFC2FEE8A821FE5DFBFC" box="[280,397,1086,1113]" count="5" pageId="18" pageNumber="19" type="female">5 females</specimenCount>
):
<collectingCountry id="C2DA766E1927FFC2FE52A820FD9BFBFC" box="[418,587,1087,1113]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FDA6A820FDB7FBD8" collectingDate="2011-06-20" collectingDateMax="2011-06-26" collectingDateMin="2011-06-20" collectionCode="JPPC" collectorName="J. Prena" country="Costa Rica" county="Bosque Eterno" location="Estac. Pocosol" municipality="Ninos" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="paratype">
<collectingRegion id="7809F81C1927FFC2FDA6A820FD69FBFC" box="[598,697,1087,1113]" country="Costa Rica" name="Alajuela" pageId="18" pageNumber="19">Alajuela</collectingRegion>
:
<collectingCounty id="53134E721927FFC2FD36A820FCBFFBFC" box="[710,879,1087,1113]" pageId="18" pageNumber="19">Bosque Eterno</collectingCounty>
de los
<collectingMunicipality id="5A16AC841927FFC2FC37A820FBDDFBFC" box="[967,1037,1087,1113]" pageId="18" pageNumber="19">Niños</collectingMunicipality>
,
<location id="BF1260251927FFC2FBEBA820FB14FBFC" LSID="urn:lsid:plazi:treatment:326487E81927FFC2FF67ADA9FD76FA64:BF1260251927FFC2FBEBA820FB14FBFC" box="[1051,1220,1087,1113]" country="Costa Rica" county="Bosque Eterno" municipality="Ninos" name="Estac. Pocosol" pageId="18" pageNumber="19" stateProvince="Alajuela">Estac. Pocosol</location>
,
<date id="CE73103E1927FFC2FB22A820FAACFBFC" box="[1234,1404,1087,1113]" pageId="18" pageNumber="19" value="2011-06-20" valueMax="2011-06-26" valueMin="2011-06-20">
<collectingDate id="DE37E9D61927FFC2FB22A820FAACFBFC" box="[1234,1404,1087,1113]" pageId="18" pageNumber="19" value="2011-06-20" valueMax="2011-06-26" valueMin="2011-06-20">20.–26.vi.2011</collectingDate>
</date>
,
<collectorName id="173853281927FFC2FA7AA820FF0AFBD8" pageId="18" pageNumber="19">J. Prena</collectorName>
, 1 dissected male,
<specimenCount id="ACCBFD771927FFC2FE5CA87CFDDDFBD8" box="[428,525,1123,1149]" count="1" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FDECA87CFD8FFBD8" box="[540,607,1123,1149]" pageId="18" pageNumber="19">JPPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FD82A87CFA48FBD8" box="[626,1432,1122,1149]" collectingDate="1991-07" collectionCode="MNCR" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingRegion id="7809F81C1927FFC2FD82A87CFD28FBD8" box="[626,760,1123,1149]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="19">Guanacaste</collectingRegion>
: same data as holotype, CRI000 358595,
<specimenCount id="ACCBFD771927FFC2FB38A87CFAF9FBD8" box="[1224,1321,1123,1149]" count="1" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FAC8A87CFA40FBD8" box="[1336,1424,1123,1149]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FF67A898FBCFFB04" box="[151,1055,1158,1185]" collectingDate="1991-11-04" collectingDateMax="1991-11-25" collectingDateMin="1991-11-04" collectionCode="MNCR" collectorName="C. Moraga" country="Costa Rica" elevation="700" location="Sta. Cecilia" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
same data as holotype except
<date id="CE73103E1927FFC2FE00A899FD5DFB04" box="[496,653,1158,1185]" pageId="18" pageNumber="19" value="1991-11-04" valueMax="1991-11-25" valueMin="1991-11-04">
<collectingDate id="DE37E9D61927FFC2FE00A899FD5DFB04" box="[496,653,1158,1185]" pageId="18" pageNumber="19" value="1991-11-04" valueMax="1991-11-25" valueMin="1991-11-04">4.–25.xi.1991</collectingDate>
</date>
, CRI000 497850,
<specimenCount id="ACCBFD771927FFC2FC91A898FC7EFB04" box="[865,942,1159,1185]" count="1" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FC4FA898FBC7FB04" box="[959,1047,1159,1185]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FBDDA899FDF2FB60" collectingDate="1992-07-01" collectingDateMax="1992-07-15" collectingDateMin="1992-07-01" collectionCode="MNCR" country="Costa Rica" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="CE73103E1927FFC2FBDDA899FB01FB04" box="[1069,1233,1158,1185]" pageId="18" pageNumber="19" value="1992-07-01" valueMax="1992-07-15" valueMin="1992-07-01">
<collectingDate id="DE37E9D61927FFC2FBDDA899FB01FB04" box="[1069,1233,1158,1185]" pageId="18" pageNumber="19" value="1992-07-01" valueMax="1992-07-15" valueMin="1992-07-01">1.–15.vii.1992</collectingDate>
</date>
, CRI000 690288, CRI000 690310,
<specimenCount id="ACCBFD771927FFC2FEABA8B4FE62FB60" box="[347,434,1195,1221]" count="2" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FE32A8B4FDCAFB60" box="[450,538,1195,1221]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FDDEA8B4FBD0FB60" box="[558,1024,1195,1221]" collectingDate="1996-12-03" collectionCode="CMNC" collectorName="Prena" country="Costa Rica" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
6.–
<date id="CE73103E1927FFC2FDBEA8B4FD14FB60" box="[590,708,1195,1221]" pageId="18" pageNumber="19" value="1996-12-03">
<collectingDate id="DE37E9D61927FFC2FDBEA8B4FD14FB60" box="[590,708,1195,1221]" pageId="18" pageNumber="19" value="1996-12-03">12.3.1996</collectingDate>
</date>
, J.
<collectorName id="173853281927FFC2FD1AA8B4FCFDFB60" box="[746,813,1195,1221]" pageId="18" pageNumber="19">Prena</collectorName>
,
<specimenCount id="ACCBFD771927FFC2FCC8A8B4FC5FFB60" box="[824,911,1195,1221]" count="2" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FC6FA8B4FC27FB60" box="[927,1015,1195,1221]" pageId="18" pageNumber="19">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FBFCA8B4FAE9FB60" box="[1036,1337,1195,1221]" collectingDate="1996-12-03" collectionCode="JPPC" collectorName="Prena" country="Costa Rica" isTailEnumeration="true" pageId="18" pageNumber="19" specimenCount="8" specimenCount-female="2" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="ACCBFD771927FFC2FBFCA8B4FBB5FB60" box="[1036,1125,1195,1221]" count="6" pageId="18" pageNumber="19" type="male">6 males</specimenCount>
,
<specimenCount id="ACCBFD771927FFC2FB82A8B4FB0FFB60" box="[1138,1247,1195,1221]" count="2" pageId="18" pageNumber="19" type="female">2 females</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FB1EA8B4FAE0FB60" box="[1262,1328,1195,1221]" pageId="18" pageNumber="19">JPPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FAB5A8B4FEC8FB4C" collectionCode="NHMUK" country="Costa Rica" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="ACCBFD771927FFC2FAB5A8B4FA4CFB60" box="[1349,1436,1195,1221]" count="2" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FF6FA8D0FEC0FB4C" box="[159,272,1231,1257]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="19" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FED2A8D0FF7FFAA8" collectingDate="1985-06-28" collectionCode="JPPC, MNCR" collectorName="H. Hespenheide" country="Costa Rica" location="Estac. La Selva" pageId="18" pageNumber="19" specimenCode="JPPC 1, MNCR 1" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<collectingRegion id="7809F81C1927FFC2FED2A8D0FEAFFB4C" box="[290,383,1231,1257]" country="Costa Rica" name="Heredia" pageId="18" pageNumber="19">Heredia</collectingRegion>
:
<locationDeviation id="ABC02FDC1927FFC2FE7AA8D0FDA2FB4C" box="[394,626,1231,1257]" location="Puerto Viejo" pageId="18" pageNumber="19">
<quantity id="7D359B1B1927FFC2FE7AA8D0FE13FB4C" box="[394,451,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="km" value="3.0">3 km</quantity>
S Puerto Viejo
</locationDeviation>
,
<location id="BF1260251927FFC2FD8DA8D0FCFEFB4C" LSID="urn:lsid:plazi:treatment:326487E81927FFC2FF67ADA9FD76FA64:BF1260251927FFC2FD8DA8D0FCFEFB4C" box="[637,814,1231,1257]" country="Costa Rica" name="Estac. La Selva" pageId="18" pageNumber="19" stateProvince="Heredia">Estac. La Selva</location>
,
<date id="CE73103E1927FFC2FCC9A8D1FC64FB4C" box="[825,948,1230,1257]" pageId="18" pageNumber="19" value="1985-06-28">
<collectingDate id="DE37E9D61927FFC2FCC9A8D1FC64FB4C" box="[825,948,1230,1257]" pageId="18" pageNumber="19" value="1985-06-28">28.vi.1985</collectingDate>
</date>
,
<collectorName id="173853281927FFC2FC30A8D0FBA6FB4C" box="[960,1142,1231,1257]" pageId="18" pageNumber="19">H. Hespenheide</collectorName>
,
<specimenCount id="ACCBFD771927FFC2FB71A8D0FB08FB4C" box="[1153,1240,1231,1257]" count="2" pageId="18" pageNumber="19" type="male">2 males</specimenCount>
(
<specimenCode id="EA6B9E851927FFC2FB17A8D0FAEBFB4C" box="[1255,1339,1231,1257]" collectionCode="JPPC" pageId="18" pageNumber="19">JPPC 1</specimenCode>
,
<specimenCode id="EA6B9E851927FFC2FABAA8D0FF74FAA8" collectionCode="MNCR" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR 1</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA31927FFC2FF30A8ECFCB9FAA8" box="[192,873,1267,1293]" collectingDate="1993-12-01" collectionCode="MNCR" country="Costa Rica" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="CE73103E1927FFC2FF30A8ECFEE3FAA8" box="[192,307,1267,1293]" pageId="18" pageNumber="19" value="1993-12-01">
<collectingDate id="DE37E9D61927FFC2FF30A8ECFEE3FAA8" box="[192,307,1267,1293]" pageId="18" pageNumber="19" value="1993-12-01">1.xii.1993</collectingDate>
</date>
, M/14/288, INB0003672898,
<specimenCount id="ACCBFD771927FFC2FD61A8ECFD25FAA8" box="[657,757,1267,1293]" count="1" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1927FFC2FCF8A8ECFCB1FAA8" box="[776,865,1267,1293]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FC87A8ECFD38FA94" collectingDate="1991-12" collectionCode="MNCR" collectorName="P. N. Manuel Antonio &amp; G. Varela" country="Costa Rica" location="Puntarenas" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="7809F81C1927FFC2FC87A8ECFC26FAA8" box="[887,1014,1267,1293]" country="Costa Rica" name="Puntarenas" pageId="18" pageNumber="19">Puntarenas</collectingRegion>
:
<collectorName id="173853281927FFC2FBF4A8ECFB2BFAA8" box="[1028,1275,1267,1293]" pageId="18" pageNumber="19">P.N. Manuel Antonio</collectorName>
,
<date id="CE73103E1927FFC2FAF9A8ECFAA2FAA8" box="[1289,1394,1267,1293]" bridgedPair="/" pageId="18" pageNumber="19" value="1991-12">
<collectingDate id="DE37E9D61927FFC2FAF9A8ECFAA2FAA8" box="[1289,1394,1267,1293]" bridgedPair="/" pageId="18" pageNumber="19" value="1991-12">XII/1991</collectingDate>
</date>
,
<collectorName id="173853281927FFC2FA71A8ECFF32FA94" pageId="18" pageNumber="19">G. Varela</collectorName>
, CRI000 464544, 1 dissected male (
<collectionCode id="DCDCAE3B1927FFC2FD77A908FD30FA94" box="[647,736,1303,1329]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="18" pageNumber="19">MNCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA31927FFC2FD06A907FE45FAF0" collectingDate="1973-04-28" collectionCode="JPPC" collectorName="H. Hespenheide" country="Panama" location="Cerro Campana" pageId="18" pageNumber="19" specimenCount="1" typeStatus="paratype">
<collectingCountry id="C2DA766E1927FFC2FD06A907FCA6FA96" box="[758,886,1304,1331]" name="Panama" pageId="18" pageNumber="19">PANAMÁ</collectingCountry>
.
<location id="BF1260251927FFC2FC73A908FBE9FA94" LSID="urn:lsid:plazi:treatment:326487E81927FFC2FF67ADA9FD76FA64:BF1260251927FFC2FC73A908FBE9FA94" box="[899,1081,1303,1329]" country="Panama" name="Cerro Campana" pageId="18" pageNumber="19">Cerro Campana</location>
,
<date id="CE73103E1927FFC2FBB7A908FB6AFA94" box="[1095,1210,1303,1329]" pageId="18" pageNumber="19" value="1973-04-28">
<collectingDate id="DE37E9D61927FFC2FBB7A908FB6AFA94" box="[1095,1210,1303,1329]" pageId="18" pageNumber="19" value="1973-04-28">28.4.1973</collectingDate>
</date>
,
<collectorName id="173853281927FFC2FB38A908FA51FA94" box="[1224,1409,1303,1329]" pageId="18" pageNumber="19">H. Hespenheide</collectorName>
, 1 dissected male (
<collectionCode id="DCDCAE3B1927FFC2FEBAA924FE5DFAF0" box="[330,397,1339,1365]" pageId="18" pageNumber="19">JPPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1927FFC2FF37A941FD2AFA38" blockId="18.[151,1437,582,1473]" pageId="18" pageNumber="19">
<emphasis id="88B9EAEC1927FFC2FF37A941FE89FADD" bold="true" box="[199,345,1374,1400]" pageId="18" pageNumber="19">Life history.</emphasis>
I found this species in numbers on a flowering, unidentified
<taxonomicName id="7DCD4D7D1927FFC2FBF9A940FB1BFADC" box="[1033,1227,1375,1401]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
in an open habitat at the Pitilla field station, together with
<taxonomicName id="7DCD4D7D1927FFC2FDA4A99CFD23FA39" box="[596,755,1411,1437]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC1927FFC2FDA4A99CFD23FA39" box="[596,755,1411,1437]" italics="true" pageId="18" pageNumber="19">C. cavernosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751927FFC2FF37A9B8FD76FA64" box="[199,678,1447,1473]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="BA7236FE1927FFC2FF37A9B8FD76FA64" blockId="18.[151,1437,582,1473]" box="[199,678,1447,1473]" pageId="18" pageNumber="19">
<emphasis id="88B9EAEC1927FFC2FF37A9B8FE9EFA64" bold="true" box="[199,334,1447,1473]" pageId="18" pageNumber="19">Etymology.</emphasis>
The name is a Latin adjective.
</paragraph>
</subSubSection>
</treatment>
</document>

115
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View file

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<paragraph id="BA7236FE1930FFD5FF67AA29FD50F9F5" blockId="5.[151,1177,1590,1689]" box="[151,640,1590,1617]" pageId="5" pageNumber="6">
<heading id="E13A81921930FFD5FF67AA29FD50F9F5" bold="true" box="[151,640,1590,1617]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="88B9EAEC1930FFD5FF67AA29FD50F9F5" bold="true" box="[151,640,1590,1617]" pageId="5" pageNumber="6">
2.
<taxonomicName id="7DCD4D7D1930FFD5FF45AA29FE33F9F5" authority="Prena" authorityName="Prena" authorityYear="2024" box="[181,483,1590,1617]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bicuspis" status="sp. nov.">
<emphasis id="88B9EAEC1930FFD5FF45AA29FE40F9F4" bold="true" box="[181,400,1590,1617]" italics="true" pageId="5" pageNumber="6">Chryasus bicuspis</emphasis>
Prena
</taxonomicName>
,
<taxonomicNameLabel id="938A57971930FFD5FE00AA29FD50F9F5" box="[496,640,1590,1616]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D765751930FFD5FF67AA45FB49F93C" pageId="5" pageNumber="6" type="description">
<paragraph id="BA7236FE1930FFD5FF67AA45FF29F9D1" blockId="5.[151,1177,1590,1689]" box="[151,249,1626,1652]" pageId="5" pageNumber="6">
(
<figureCitation id="22F62A7B1930FFD5FF6FAA45FF21F9D1" box="[159,241,1626,1652]" captionStart="FIGURES 26–29" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[177,1410,181,1918]" captionTargetId="figure-26@8.[177,1410,181,1918]" captionTargetPageId="8" captionText="FIGURES 26–29. Chryasus, dorsal and lateral aspects of males. 26, C. bicolor, Nova Teutônia in Santa Catarina, Brazil; 27, C. bicuspis, Shushufindi in Sucumbíos, Ecuador; 28, C. pomponella, Santa Cruz in Junín, Perú; 29, C. nimius, Estac. Magsassay, P.N. Braulio Carrillo in Heredia, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211519" httpUri="https://zenodo.org/record/13211519/files/figure.png" pageId="5" pageNumber="6">Fig. 27</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE1930FFD5FF67AA61FB49F93C" blockId="5.[151,1177,1590,1689]" box="[151,1177,1662,1689]" pageId="5" pageNumber="6">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="F2D765751930FFD5FF67AAD9FCE9F8A0" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="BA7236FE1930FFD5FF67AAD9FCE9F8A0" blockId="5.[151,1437,1734,2013]" pageId="5" pageNumber="6">
<emphasis id="88B9EAEC1930FFD5FF67AAD9FEC2F945" bold="true" box="[151,274,1734,1760]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Among the large humped species, this is the only one with dark reddish brown elytra. The species is larger and more elongate than the entirely black
<taxonomicName id="7DCD4D7D1930FFD5FD45AAF4FCE2F8A1" authorityName="Prena" authorityYear="2024" box="[693,818,1771,1797]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wappesi">
<emphasis id="88B9EAEC1930FFD5FD45AAF4FCE2F8A1" box="[693,818,1771,1797]" italics="true" pageId="5" pageNumber="6">C. wappesi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<emphasis id="88B9EAEC1930FFD5FF37AB11FE89F88D" bold="true" box="[199,345,1806,1832]" pageId="5" pageNumber="6">Description.</emphasis>
Total length 5.3, width
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(n=1); integument black, antenna, tibia, tarsus and elytron dark reddish brown; body elongate rhomboidal, sides of pronotum and elytra for most parts gradually converging, pronotum moderately vaulted, distinctly vermiculate on disc and shallowly punctate on flank, elytron with slight basal tumidity, interstriae ridged, male metaventral protuberance with anterior edge strongly produced and deeply cleft by discrimen, male ventrites 1, 2 and 5 with erect squamulae, 5 with median depression; all femora with secondary denticles but barely perceptible on fore leg; genitalia not examined.
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<paragraph id="BA7236FE1933FFD6FF37AC89FED4FF70" blockId="6.[151,1436,150,249]" pageId="6" pageNumber="7">
<emphasis id="88B9EAEC1933FFD6FF37AC89FE69FF15" bold="true" box="[199,441,150,176]" pageId="6" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="0AA53CA31933FFD6FE39AC88FED4FF70" collectionCode="CMNC" collectorName="W. Opitz" country="Ecuador" location="Sucumbios" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Sucumbios" typeStatus="holotype">
<typeStatus id="6576885C1933FFD6FE39AC88FDE1FF14" box="[457,561,151,177]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="C2DA766E1933FFD6FD09AC88FC8BFF14" box="[761,859,151,177]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
,
<collectingRegion id="7809F81C1933FFD6FC9EAC88FC3DFF14" box="[878,1005,151,177]" country="Ecuador" name="Sucumbios" pageId="6" pageNumber="7">Sucumbios</collectingRegion>
, Shushufindi, 9-VIII-98,
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” (
<collectionCode id="DCDCAE3B1933FFD6FF6FACA4FF28FF70" box="[159,248,187,213]" pageId="6" pageNumber="7">CMNC</collectionCode>
).
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<emphasis id="88B9EAEC1933FFD6FF37ACC0FE9EFF5C" bold="true" box="[199,334,223,249]" pageId="6" pageNumber="7">Etymology.</emphasis>
The name is a Latin noun referring to the cleft metaventral protuberance.
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129
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3.
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<emphasis id="88B9EAEC1933FFD6FF45AD5CFE6CFEFB" bold="true" box="[181,444,323,350]" italics="true" pageId="6" pageNumber="7">Chryasus pomponella</emphasis>
Prena
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,
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(
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)
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<paragraph id="BA7236FE1933FFD6FF67AD93FB7AFE02" blockId="6.[151,1194,323,423]" box="[151,1194,396,423]" pageId="6" pageNumber="7">
http://zoobank.org/
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<emphasis id="88B9EAEC1933FFD6FF67ADD2FEC2FE42" bold="true" box="[151,274,461,487]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
This is the largest of all known species, readily recognizable by its distinctive color pattern and the extremely humped pronotum.
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<paragraph id="BA7236FE1933FFD6FF37AE0AFE1FFCA2" blockId="6.[151,1436,461,920]" pageId="6" pageNumber="7">
<emphasis id="88B9EAEC1933FFD6FF37AE0AFE89FD8A" bold="true" box="[199,345,533,559]" pageId="6" pageNumber="7">Description.</emphasis>
Total length
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, width 3.0–
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(n=2); integument fuscous to black, antenna and legs brown, elytron yellow ochre with reddish brown base and apex; body elongate rhomboidal, sides of pronotum and elytra for most parts very gently curved, pronotum very strongly vaulted, distinctly vermiculate on disc and shallowly punctate on flank, elytron with slight basal tumidity but otherwise comparatively flat, interstriae ridged, male metaventral protuberance moderately developed and slightly channeled by discrimen, male ventrites 1, 2 and 5 shallowly depressed and with erect squamulae; all femora with secondary denticles but barely perceptible on fore leg; genitalia not examined.
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<emphasis id="88B9EAEC1933FFD6FF37AF0EFE60FC8E" bold="true" box="[199,432,785,811]" pageId="6" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="0AA53CA31933FFD6FE46AF0EFAC3FC8E" box="[438,1299,785,811]" collectingDate="2012-02" collectionCode="CMNC" country="Peru" location="Santa Cruz" municipality="Satipo" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Junin" typeStatus="holotype">
<typeStatus id="6576885C1933FFD6FE46AF0EFDCEFC8E" box="[438,542,785,811]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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,
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,
<date id="CE73103E1933FFD6FBB9AF0EFB49FC8E" box="[1097,1177,785,811]" pageId="6" pageNumber="7" value="2012-02">
<collectingDate id="DE37E9D61933FFD6FBB9AF0EFB49FC8E" box="[1097,1177,785,811]" pageId="6" pageNumber="7" value="2012-02">ii.2012</collectingDate>
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.
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<typeStatus id="6576885C1933FFD6FAEDAF0DFAAFFC89" box="[1309,1407,786,812]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
(
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): Peru, Marcapata (
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.
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<emphasis id="88B9EAEC1933FFD6FF37AF46FE9EFCD6" bold="true" box="[199,334,857,883]" pageId="6" pageNumber="7">Etymology.</emphasis>
The name is a noun in apposition, adopted from that of the last little grandniece of King Futsch of Pompolonia, a fictive character by Max Kruse.
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137
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<heading id="E13A81921933FFD6FF67AFC3FDA1FC53" bold="true" box="[151,625,987,1014]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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4.
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<emphasis id="88B9EAEC1933FFD6FF45AFC4FE51FC53" bold="true" box="[181,385,987,1014]" italics="true" pageId="6" pageNumber="7">Chryasus nimius</emphasis>
Prena
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,
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</heading>
</paragraph>
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<subSubSection id="F2D765751933FFD6FF67A81FFB58FB9A" pageId="6" pageNumber="7" type="description">
<paragraph id="BA7236FE1933FFD6FF67A81FFF29FBBF" blockId="6.[151,1160,987,1087]" box="[151,249,1024,1050]" pageId="6" pageNumber="7">
(
<figureCitation id="22F62A7B1933FFD6FF6FA81FFF21FBBF" box="[159,241,1024,1050]" captionStart="FIGURES 26–29" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[177,1410,181,1918]" captionTargetId="figure-26@8.[177,1410,181,1918]" captionTargetPageId="8" captionText="FIGURES 26–29. Chryasus, dorsal and lateral aspects of males. 26, C. bicolor, Nova Teutônia in Santa Catarina, Brazil; 27, C. bicuspis, Shushufindi in Sucumbíos, Ecuador; 28, C. pomponella, Santa Cruz in Junín, Perú; 29, C. nimius, Estac. Magsassay, P.N. Braulio Carrillo in Heredia, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211519" httpUri="https://zenodo.org/record/13211519/files/figure.png" pageId="6" pageNumber="7">Fig. 29</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE1933FFD6FF67A83BFB58FB9A" blockId="6.[151,1160,987,1087]" box="[151,1160,1060,1087]" pageId="6" pageNumber="7">
http://zoobank.org/
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<subSubSection id="F2D765751933FFD6FF67A873FE4EFA27" pageId="6" pageNumber="7" type="diagnosis">
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<emphasis id="88B9EAEC1933FFD6FF67A873FEC2FB23" bold="true" box="[151,274,1132,1158]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
<taxonomicName id="7DCD4D7D1933FFD6FEE7A873FE02FB23" authorityName="Prena" authorityYear="2024" box="[279,466,1132,1158]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nimius">
<emphasis id="88B9EAEC1933FFD6FEE7A873FE02FB23" box="[279,466,1132,1158]" italics="true" pageId="6" pageNumber="7">Chryasus nimius</emphasis>
</taxonomicName>
is the most slender among the large humped species. The interstrial ridges on the elytral disc are interrupted by two transverse depressions.
</paragraph>
<paragraph id="BA7236FE1933FFD6FF37A8ABFE4EFA27" blockId="6.[151,1437,1132,1555]" pageId="6" pageNumber="7">
Description: Total length
<quantity id="7D359B1B1933FFD6FE18A8ABFDA2FB6A" box="[488,626,1204,1231]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.4" metricValueMin="5.3" pageId="6" pageNumber="7" unit="mm" value="5.35" valueMax="5.4" valueMin="5.3">5.3–5.4 mm</quantity>
, width
<quantity id="7D359B1B1933FFD6FD36A8ABFC9EFB6A" box="[710,846,1204,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="6" pageNumber="7" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
(n=2); integument black, antenna and tarsus brown; body elongate rhomboidal, sides of pronotum and elytra for most parts gradually converging, pronotum moderately vaulted, distinctly vermiculate on disc and shallowly punctate on flank, elytron with two transverse depressions, interstriae ridged, male metaventral protuberance angularly pronounced and cleft by discrimen, male ventrites 1, 2 and 5 shallowly depressed and with erect squamulae; all femora with denticles almost entirely reduced except for 1; genitalia not examined.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751933FFD6FF37A993FB70FA4B" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="BA7236FE1933FFD6FF37A993FB70FA4B" blockId="6.[151,1437,1132,1555]" pageId="6" pageNumber="7">
<emphasis id="88B9EAEC1933FFD6FF37A993FE7FFA03" bold="true" box="[199,431,1420,1446]" pageId="6" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="0AA53CA31933FFD6FE45A993FBE5FA6F" collectingDate="1991-01" collectionCode="MNCR" collectorName="M. Barrelier" country="Costa Rica" elevation="200" location="COSTA RICA" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
<typeStatus id="6576885C1933FFD6FE45A993FDCDFA03" box="[437,541,1420,1446]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
male, labeled “Est. Magsasay, P.N./ Braulio Carrillo,
<quantity id="7D359B1B1933FFD6FB9EA993FB63FA03" box="[1134,1203,1420,1446]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="200.0">
<elevation id="31E0D1CD1933FFD6FB9EA993FB63FA03" box="[1134,1203,1420,1446]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="200.0">200 m</elevation>
</quantity>
/ Prov. Here,
<collectingCountry id="C2DA766E1933FFD6FAB4A993FF09FA6F" name="Costa Rica" pageId="6" pageNumber="7">COSTA RICA</collectingCountry>
/
<collectorName id="173853281933FFD6FF1AA9AEFEA7FA6F" box="[234,375,1456,1483]" pageId="6" pageNumber="7">M. Barrelier</collectorName>
,
<date id="CE73103E1933FFD6FE73A9AEFE20FA6F" box="[387,496,1456,1483]" pageId="6" pageNumber="7" value="1991-01">
<collectingDate id="DE37E9D61933FFD6FE73A9AEFE20FA6F" box="[387,496,1456,1483]" pageId="6" pageNumber="7" value="1991-01">Ene 1991</collectingDate>
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,/ L-N-254600, 531100”, CRI000 064266 (
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)
</materialsCitation>
.
<materialsCitation id="0AA53CA31933FFD6FBB1A9AEFB4CFA4B" collectingDate="1991-04" collectionCode="MNCR" collectorName="Quepos, P. N. &amp; Manuel Antonio, R" country="Costa Rica" location="Puntarenas" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="6576885C1933FFD6FBB1A9AEFB73FA6E" box="[1089,1187,1457,1483]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
(
<specimenCount id="ACCBFD771933FFD6FB46A9AFFAD6FA6F" box="[1206,1286,1456,1482]" count="1" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
): Costa Rica,
<collectingRegion id="7809F81C1933FFD6FF67A9CAFEC5FA4A" box="[151,277,1493,1519]" country="Costa Rica" name="Puntarenas" pageId="6" pageNumber="7">Puntarenas</collectingRegion>
,
<collectorName id="173853281933FFD6FED0A9CBFD44FA4A" box="[288,660,1492,1519]" pageId="6" pageNumber="7">Quepos, P.N. Manuel Antonio, R</collectorName>
. Zúñiga,
<date id="CE73103E1933FFD6FD0AA9CBFCBCFA4B" box="[762,876,1492,1518]" pageId="6" pageNumber="7" value="1991-04">
<collectingDate id="DE37E9D61933FFD6FD0AA9CBFCBCFA4B" box="[762,876,1492,1518]" pageId="6" pageNumber="7" value="1991-04">Abr. 1991</collectingDate>
</date>
, CRI000 601953 (
<collectionCode id="DCDCAE3B1933FFD6FBCBA9CBFB44FA4B" box="[1083,1172,1492,1518]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="6" pageNumber="7">MNCR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751933FFD6FF37A9E7FC35F9B7" box="[199,997,1528,1555]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="BA7236FE1933FFD6FF37A9E7FC35F9B7" blockId="6.[151,1437,1132,1555]" box="[199,997,1528,1555]" pageId="6" pageNumber="7">
<emphasis id="88B9EAEC1933FFD6FF37A9E7FE9EF9B7" bold="true" box="[199,334,1528,1554]" pageId="6" pageNumber="7">Etymology.</emphasis>
The name is a Latin adjective meaning very big or too big.
</paragraph>
</subSubSection>
</treatment>
</document>

183
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<mods:title id="3592BAF7797A7674BE455547DFD072AA">Consolidation of Aniops Casey, Psiona Casey, Preglyptobaris Bondar and Prospoliata Hustache with Chryasus Champion (Coleoptera, Curculionidae, Baridinae) and descriptions of new species</mods:title>
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5.
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<paragraph id="BA7236FE1933FFD6FF67AA64FB40F91C" blockId="6.[151,1168,1622,1721]" pageId="6" pageNumber="7">
(
<figureCitation id="22F62A7B1933FFD6FF6FAA64FEF6F930" box="[159,294,1659,1685]" captionStart="FIGURES 11–12" captionStartId="5.[152,270,740,764]" captionTargetBox="[188,1400,181,717]" captionTargetId="figure-15@5.[188,1400,181,717]" captionTargetPageId="5" captionText="FIGURES 11–12. Externally visible apical tergites of Chryasus wappesi. 11, male with stridulatory files on tergite 7; 12, female without stridulatory files on tergite 7, tergite 8 internalized." figureDoi="http://doi.org/10.5281/zenodo.13211509" httpUri="https://zenodo.org/record/13211509/files/figure.png" pageId="6" pageNumber="7">Figs. 11, 12</figureCitation>
,
<figureCitation id="22F62A7B1933FFD6FEC2AA65FE80F930" box="[306,336,1658,1685]" captionStart="FIGURES 13–15" captionStartId="5.[152,270,1376,1400]" captionTargetBox="[189,1398,825,1352]" captionTargetId="figure-50@5.[189,1398,825,1352]" captionTargetPageId="5" captionText="FIGURES 13–15. Changes in extension and relative position of morphological parts on the prosternum associated with increased gibbosity of the pronotum and simultaneously shortened prosternum (ID = intercoxal distance; LR = length of lateral ridge of prosternal channel). 13, Chryasus montyi, with ID &lt;LR; 14, Chryasus sp. near C. bruniceps, Costa Rica (see Fig. 33), with ID = LR; 15, Chryasus wappesi, with ID&gt; LR." figureDoi="http://doi.org/10.5281/zenodo.13211511" httpUri="https://zenodo.org/record/13211511/files/figure.png" pageId="6" pageNumber="7">15</figureCitation>
,
<figureCitation id="22F62A7B1933FFD6FEACAA64FEAAF930" box="[348,378,1659,1685]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="6" pageNumber="7">16</figureCitation>
,
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<emphasis id="88B9EAEC1933FFD6FF67AAFFFEC2F95F" bold="true" box="[151,274,1760,1786]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
This is the smallest of the large humped species with male metaventral protuberance. It is less cuneate than the others and connects to the more ovate, medium-sized species, such as
<taxonomicName id="7DCD4D7D1933FFD6FBFAAB1BFB42F8BB" authorityName="Champion" authorityYear="1908" box="[1034,1170,1796,1822]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="88B9EAEC1933FFD6FBFAAB1BFB42F8BB" box="[1034,1170,1796,1822]" italics="true" pageId="6" pageNumber="7">C. plagiatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751933FFD6FF37AB37FA51F853" pageId="6" pageNumber="7" type="description">
<paragraph id="BA7236FE1933FFD6FF37AB37FA51F853" blockId="6.[151,1437,1760,2039]" pageId="6" pageNumber="7">
<emphasis id="88B9EAEC1933FFD6FF37AB37FE89F8E7" bold="true" box="[199,345,1832,1858]" pageId="6" pageNumber="7">Description.</emphasis>
Total length 4.3–4.9, width
<quantity id="7D359B1B1933FFD6FD60AB37FCC6F8E6" box="[656,790,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="6" pageNumber="7" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
(n=4); integument black, antenna, tibia, tarsus and elytron dark reddish brown; body elongate rhomboidal, sides of pronotum and elytra for most parts gradually converging, pronotum moderately vaulted, distinctly vermiculate on disc and shallowly punctate on flank, elytron with slight basal tumidity, interstriae ridged, male metaventral protuberances with anterior edge strongly produced and deeply cleft by discrimen, male ventrites 1, 2, 5 and metaventrite with erect squamulae; all femora with secondary denticles but barely perceptible on fore leg; male genitalia as
<figureCitation id="22F62A7B1933FFD6FD43ABC3FCD4F853" box="[691,772,2012,2038]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="6" pageNumber="7">Fig. 16</figureCitation>
, endophallus with moderately long, inflexible flagellum.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751932FFD7FF37AC89FAE1FEC0" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="BA7236FE1932FFD7FF37AC89FA4DFF14" blockId="7.[151,1437,150,357]" box="[199,1437,150,177]" pageId="7" pageNumber="8">
<emphasis id="88B9EAEC1932FFD7FF37AC89FE63FF15" bold="true" box="[199,435,150,176]" pageId="7" pageNumber="8">Material examined.</emphasis>
<materialsCitation id="0AA53CA31932FFD7FE4CAC88FA4DFF14" box="[444,1437,151,177]" country="French Guiana" location="Amazone Nature Lodge" municipality="Kaw Rd." pageId="7" pageNumber="8" specimenCount="1" typeStatus="holotype">
<typeStatus id="6576885C1932FFD7FE4CAC88FDF4FF14" box="[444,548,151,177]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="C2DA766E1932FFD7FD28AC88FC56FF14" box="[728,902,151,177]" name="French Guiana" pageId="7" pageNumber="8">French Guiana</collectingCountry>
,
<location id="BF1260251932FFD7FC61AC88FB75FF14" LSID="urn:lsid:plazi:treatment:326487E81933FFD7FF67AA48FAE1FEC0:BF1260251932FFD7FC61AC88FB75FF14" box="[913,1189,151,177]" country="French Guiana" municipality="Kaw Rd." name="Amazone Nature Lodge" pageId="7" pageNumber="8">Amazone Nature Lodge</location>
,
<collectingMunicipality id="5A16AC841932FFD7FB43AC88FAC9FF14" box="[1203,1305,151,177]" pageId="7" pageNumber="8">Kaw Rd.</collectingMunicipality>
6, VIII/11-
</materialsCitation>
</paragraph>
<paragraph id="BA7236FE1932FFD7FF67ACA4FA4DFF70" blockId="7.[151,1437,150,357]" box="[151,1437,186,213]" pageId="7" pageNumber="8">
23/2017, Morris &amp; Wappes”, “
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,
<geoCoordinate id="DFF950391932FFD7FD9AACA5FD34FF70" box="[618,740,186,213]" degrees="52.170310" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="52.17031">52.170310</geoCoordinate>
, beating vegetation” (
<collectionCode id="DCDCAE3B1932FFD7FC2AACA4FBE3FF70" box="[986,1075,187,213]" pageId="7" pageNumber="8">CMNC</collectionCode>
).
<typeStatus id="6576885C1932FFD7FBB6ACA4FB63FF70" box="[1094,1203,187,213]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="ACCBFD771932FFD7FB33ACA4FACDFF70" box="[1219,1309,187,213]" count="2" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
,
<specimenCount id="ACCBFD771932FFD7FAD9ACA4FA41FF70" box="[1321,1425,187,213]" count="1" pageId="7" pageNumber="8" type="female">1 female</specimenCount>
):
</paragraph>
<paragraph id="BA7236FE1932FFD7FF67ACC0FAE1FEC0" blockId="7.[151,1437,150,357]" pageId="7" pageNumber="8">
<materialsCitation id="0AA53CA31932FFD7FF67ACC0FDCCFF5C" box="[151,540,223,249]" collectorName="El Pauchi" country="Venezuela" location="Bolivar" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="C2DA766E1932FFD7FF67ACC0FEDDFF5C" box="[151,269,223,249]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
,
<collectingRegion id="7809F81C1932FFD7FEE8ACC0FEBEFF5C" box="[280,366,223,249]" country="Venezuela" name="Bolivar" pageId="7" pageNumber="8">Bolivar</collectingRegion>
,
<collectorName id="173853281932FFD7FE89ACC0FE35FF5C" box="[377,485,223,249]" pageId="7" pageNumber="8">El Pauchi</collectorName>
near
</materialsCitation>
<materialsCitation id="0AA53CA31932FFD7FDD3ACC0FE65FEB9" collectingDate="1995-01-01" collectionCode="JPPC" country="Saint Helena, Ascension and Tristan da Cunha" location="Hillert" pageId="7" pageNumber="8" specimenCount="1">
<collectingCountry id="C2DA766E1932FFD7FDD3ACC0FD7CFF5C" box="[547,684,223,249]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="8">Santa Elena</collectingCountry>
,
<date id="CE73103E1932FFD7FD46ACC1FCC4FF5C" box="[694,788,222,249]" pageId="7" pageNumber="8" value="1995-01-01">
<collectingDate id="DE37E9D61932FFD7FD46ACC1FCC4FF5C" box="[694,788,222,249]" pageId="7" pageNumber="8" value="1995-01-01">1.i.1995</collectingDate>
</date>
, G.
<location id="BF1260251932FFD7FCB2ACC0FC5BFF5C" LSID="urn:lsid:plazi:treatment:326487E81933FFD7FF67AA48FAE1FEC0:BF1260251932FFD7FCB2ACC0FC5BFF5C" box="[834,907,223,249]" country="Saint Helena, Ascension and Tristan da Cunha" name="Hillert" pageId="7" pageNumber="8">Hillert</location>
(
<collectionCode id="DCDCAE3B1932FFD7FC6AACC0FC0DFF5C" box="[922,989,223,249]" pageId="7" pageNumber="8">JPPC</collectionCode>
).
<typeStatus id="6576885C1932FFD7FF37AD1DFE65FEB9" box="[199,437,258,284]" isOtherTypeStatus="true" pageId="7" pageNumber="8">
<emphasis id="88B9EAEC1932FFD7FF37AD1DFE65FEB9" bold="true" box="[199,437,258,284]" pageId="7" pageNumber="8">Additional material</emphasis>
</typeStatus>
</materialsCitation>
<emphasis id="88B9EAEC1932FFD7FE45AD1DFE6AFEB9" bold="true" box="[437,442,258,284]" pageId="7" pageNumber="8">.</emphasis>
<materialsCitation id="0AA53CA31932FFD7FE31AD1CFB81FEE4" collectionCode="USNM" collectorName="Etymology. The &amp; Jim Wappes" country="Brazil" location="Porto Platon" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Amapa" typeStatus="holotype">
<collectingCountry id="C2DA766E1932FFD7FE31AD1CFDD8FEB8" box="[449,520,259,285]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
,
<location id="BF1260251932FFD7FDE3AD1CFD4FFEB8" LSID="urn:lsid:plazi:treatment:326487E81933FFD7FF67AA48FAE1FEC0:BF1260251932FFD7FDE3AD1CFD4FFEB8" box="[531,671,259,285]" country="Brazil" name="Porto Platon" pageId="7" pageNumber="8" stateProvince="Amapa">Pôrto Platon</location>
(
<collectingRegion id="7809F81C1932FFD7FD5DAD1CFCD4FEB8" box="[685,772,259,285]" country="Brazil" name="Amapa" pageId="7" pageNumber="8">Amapá</collectingRegion>
),
<specimenCount id="ACCBFD771932FFD7FCE7AD1CFCB2FEB8" box="[791,866,259,285]" count="1" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B1932FFD7FC81AD1CFC18FEB8" box="[881,968,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8" type="Museum">USNM</collectionCode>
).
<collectorName id="173853281932FFD7FF37AD38FE50FEE4" box="[199,384,295,321]" pageId="7" pageNumber="8">
<emphasis id="88B9EAEC1932FFD7FF37AD38FE9EFEE4" bold="true" box="[199,334,295,321]" pageId="7" pageNumber="8">Etymology.</emphasis>
The
</collectorName>
name is given for the late
<collectorName id="173853281932FFD7FD5AAD38FCE4FEE4" box="[682,820,295,321]" pageId="7" pageNumber="8">Jim Wappes</collectorName>
, collector of the
<typeStatus id="6576885C1932FFD7FC1FAD38FB81FEE4" box="[1007,1105,295,321]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
</materialsCitation>
<materialsCitation id="0AA53CA31932FFD7FBA8AD38FCC5FEC0" country="Brazil" location="Notes. Not" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
and generous friend.
<location id="BF1260251932FFD7FF37AD54FE92FEC0" LSID="urn:lsid:plazi:treatment:326487E81933FFD7FF67AA48FAE1FEC0:BF1260251932FFD7FF37AD54FE92FEC0" box="[199,322,331,357]" country="Brazil" name="Notes. Not" pageId="7" pageNumber="8" stateProvince="Amapa">
<emphasis id="88B9EAEC1932FFD7FF37AD54FEC1FEC0" bold="true" box="[199,273,331,357]" pageId="7" pageNumber="8">Notes.</emphasis>
Not
</location>
included in the type series is a male from
</materialsCitation>
<collectingCountry id="C2DA766E1932FFD7FCECAD54FCB0FEC0" box="[796,864,331,357]" name="Brazil" pageId="7" pageNumber="8">Brazil</collectingCountry>
with somewhat deviant interstrial ridges.
</paragraph>
</subSubSection>
</treatment>
</document>

154
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<paragraph id="BA7236FE1939FFDCFF67AF01FDB9FC9D" blockId="12.[151,1189,798,897]" box="[151,617,798,825]" pageId="12" pageNumber="13">
<heading id="E13A81921939FFDCFF67AF01FDB9FC9D" bold="true" box="[151,617,798,825]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="88B9EAEC1939FFDCFF67AF01FDB9FC9D" bold="true" box="[151,617,798,825]" pageId="12" pageNumber="13">
9.
<taxonomicName id="7DCD4D7D1939FFDCFF45AF01FE1CFC9C" authority="Prena" authorityName="Prena" authorityYear="2024" box="[181,460,798,825]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macer" status="sp. nov.">
<emphasis id="88B9EAEC1939FFDCFF45AF01FEA9FC9C" bold="true" box="[181,377,798,825]" italics="true" pageId="12" pageNumber="13">Chryasus macer</emphasis>
Prena
</taxonomicName>
,
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<paragraph id="BA7236FE1939FFDCFF67AF5CFEFEFCF8" blockId="12.[151,1189,798,897]" box="[151,302,835,861]" pageId="12" pageNumber="13">
(
<figureCitation id="22F62A7B1939FFDCFF6FAF5CFF2DFCF8" box="[159,253,835,861]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="12" pageNumber="13">Figs. 17</figureCitation>
,
<figureCitation id="22F62A7B1939FFDCFEF9AF5CFEF6FCF8" box="[265,294,835,861]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="12" pageNumber="13">38</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE1939FFDCFF67AF78FB75FC24" blockId="12.[151,1189,798,897]" box="[151,1189,871,897]" pageId="12" pageNumber="13">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="F2D765751939FFDCFF67AFB1FF0AFBB4" pageId="12" pageNumber="13" type="diagnosis">
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<emphasis id="88B9EAEC1939FFDCFF67AFB1FEC2FC6D" bold="true" box="[151,274,942,968]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
This is a small, slender, black species with some yellow hairs (
<figureCitation id="22F62A7B1939FFDCFC18AFB0FBEEFC6C" box="[1000,1086,943,969]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="12" pageNumber="13">Fig. 38</figureCitation>
). Other elongate species in the same mountain range are stouter and almost always with red or yellow color pattern. All of them need further study.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751939FFDCFF37A805FF20FAA8" pageId="12" pageNumber="13" type="description">
<paragraph id="BA7236FE1939FFDCFF37A805FF20FAA8" blockId="12.[151,1437,942,1509]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37A805FE89FB91" bold="true" box="[199,345,1050,1076]" pageId="12" pageNumber="13">Description.</emphasis>
Total length
<quantity id="7D359B1B1939FFDCFE03A804FDAEFB90" box="[499,638,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="12" pageNumber="13" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
, width
<quantity id="7D359B1B1939FFDCFD24A804FC8DFB90" box="[724,861,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="12" pageNumber="13" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
(n=3); integument mostly black, appendages with some red; body elongate, humeri almost flush with subconical pronotum, disc densely vermiculate, flanks smoother with coarse shallow ridges; elytron 3.0x longer than pronotum, disc at base with transverse depression, interstriae ridged, metaventrite tumid with basally deep discrimen, male ventrites 1, 2, 5 and metaventrite depressed medially, 1, 2 and 5 with erect squamulae; femora with 1 distinct and several minute denticles; male genitalia as
<figureCitation id="22F62A7B1939FFDCFAB3A8B4FA47FB60" box="[1347,1431,1195,1221]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="12" pageNumber="13">Fig. 17</figureCitation>
, endophallus without flagellum but with elongate sclerite homologous with basal appendage of flagellum of other species.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751939FFDCFF37A909FD4CFADC" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="BA7236FE1939FFDCFF37A909FD4CFADC" blockId="12.[151,1437,942,1509]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37A909FE63FA95" bold="true" box="[199,435,1302,1328]" pageId="12" pageNumber="13">Material examined.</emphasis>
<materialsCitation id="0AA53CA31939FFDCFE4CA908FBDDFAF0" collectingDate="2003-03-22" collectionCode="CMNC" collectorName="R. Anderson" country="Costa Rica" elevation="2600" latitude="9.640278" location="Pan Am Hwy." longLatPrecision="21" longitude="-83.85111" municipality="Finca Eddie Serrano" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="6576885C1939FFDCFE4CA908FDF4FA94" box="[444,548,1303,1329]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="C2DA766E1939FFDCFD29A908FC8CFA94" box="[729,860,1303,1329]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
:
<collectingRegion id="7809F81C1939FFDCFC99A908FC37FA94" box="[873,999,1303,1329]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="13">Puntarenas</collectingRegion>
,
<collectingMunicipality id="5A16AC841939FFDCFC04A908FB30FA94" box="[1012,1248,1303,1329]" pageId="12" pageNumber="13">Finca Eddie Serrano</collectingMunicipality>
, km 71
<location id="BF1260251939FFDCFAB0A908FF06FAF0" LSID="urn:lsid:plazi:treatment:326487E81939FFDCFF67AF01FE5EFA40:BF1260251939FFDCFAB0A908FF06FAF0" country="Costa Rica" latitude="9.640278" longLatPrecision="21" longitude="-83.85111" municipality="Finca Eddie Serrano" name="Pan Am Hwy." pageId="12" pageNumber="13" stateProvince="Puntarenas">Pan Am Hwy.</location>
,
<quantity id="7D359B1B1939FFDCFF10A924FEE6FAF0" box="[224,310,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="13" unit="m" value="2600.0">
<elevation id="31E0D1CD1939FFDCFF10A924FEE6FAF0" box="[224,310,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="13" unit="m" value="2600.0">2600 m</elevation>
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,
<geoCoordinate id="DFF950391939FFDCFEB1A924FE1BFAF0" box="[321,459,1339,1367]" degrees="9" direction="north" minutes="38" orientation="latitude" pageId="12" pageNumber="13" precision="15" seconds="25" value="9.640278">9° 38′ 25″ N</geoCoordinate>
<geoCoordinate id="DFF950391939FFDCFE22A924FDA0FAF0" box="[466,624,1339,1367]" degrees="83" direction="west" minutes="51" orientation="longitude" pageId="12" pageNumber="13" precision="15" seconds="04" value="-83.85111">83° 51′ 04″ W</geoCoordinate>
,
<date id="CE73103E1939FFDCFD8AA924FD2BFAF0" box="[634,763,1339,1365]" pageId="12" pageNumber="13" value="2003-03-22">
<collectingDate id="DE37E9D61939FFDCFD8AA924FD2BFAF0" box="[634,763,1339,1365]" pageId="12" pageNumber="13" value="2003-03-22">22.III.2003</collectingDate>
</date>
,
<collectorName id="173853281939FFDCFCF6A924FC44FAF0" box="[774,916,1339,1365]" pageId="12" pageNumber="13">R. Anderson</collectorName>
” (
<collectionCode id="DCDCAE3B1939FFDCFC5CA924FBD5FAF0" box="[940,1029,1339,1365]" pageId="12" pageNumber="13">CMNC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="6576885C1939FFDCFBE6A924FB53FAF0" box="[1046,1155,1339,1365]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="ACCBFD771939FFDCFB64A924FB30FAF0" box="[1172,1248,1339,1365]" count="1" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
,
<specimenCount id="ACCBFD771939FFDCFB1BA924FA81FAF0" box="[1259,1361,1339,1365]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
):
<materialsCitation id="0AA53CA31939FFDCFA93A924FD48FADC" collectingDate="2003-03-22" collectionCode="JPPC, CMNC" collectorName="R. Anderson" country="Costa Rica" elevation="2600" latitude="9.640278" location="Pan Am Hwy." longLatPrecision="21" longitude="-83.85111" municipality="Finca Eddie Serrano" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
same data, dissected male (
<collectionCode id="DCDCAE3B1939FFDCFE77A940FE19FADC" box="[391,457,1375,1401]" pageId="12" pageNumber="13">JPPC</collectionCode>
), female (
<collectionCode id="DCDCAE3B1939FFDCFDC7A940FD40FADC" box="[567,656,1375,1401]" pageId="12" pageNumber="13">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751939FFDCFF37A99CFE5EFA40" pageId="12" pageNumber="13" type="etymology">
<paragraph id="BA7236FE1939FFDCFF37A99CFE5EFA40" blockId="12.[151,1437,942,1509]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37A99CFE9EFA38" bold="true" box="[199,334,1411,1437]" pageId="12" pageNumber="13">Etymology.</emphasis>
The name is a Latin adjective meaning thin. It refers to the general appearance of the species and is a reference to the nemonychid
<taxonomicName id="7DCD4D7D1939FFDCFE04A9B8FD33FA64" authority="Kuschel" authorityName="Kuschel" authorityYear="1989" box="[500,739,1447,1473]" class="Insecta" family="Nemonychidae" genus="Atopomacer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="88B9EAEC1939FFDCFE04A9B8FDADFA64" box="[500,637,1447,1473]" italics="true" pageId="12" pageNumber="13">Atopomacer</emphasis>
Kuschel
</taxonomicName>
, the genus actually targeted at this site by Carlos Viquez, Bob Anderson and myself.
</paragraph>
</subSubSection>
</treatment>
</document>

196
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<paragraph id="BA7236FE193AFFDFFF67AA33FD0AF9E3" blockId="15.[151,730,1579,1643]" box="[151,730,1579,1606]" pageId="15" pageNumber="16">
<heading id="E13A8192193AFFDFFF67AA33FD0AF9E3" bold="true" box="[151,730,1579,1606]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="88B9EAEC193AFFDFFF67AA33FD0AF9E3" bold="true" box="[151,730,1579,1606]" pageId="15" pageNumber="16">
15.
<taxonomicName id="7DCD4D7D193AFFDFFF33AA34FE27F9E3" authority="(Casey)" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[195,503,1579,1606]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densus" status="comb. nov.">
<emphasis id="88B9EAEC193AFFDFFF33AA34FE40F9E3" bold="true" box="[195,400,1579,1606]" italics="true" pageId="15" pageNumber="16">Chryasus densus</emphasis>
(Casey)
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,
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<paragraph id="BA7236FE193AFFDFFF67AA4FFD17F9CF" blockId="15.[151,730,1579,1643]" box="[151,711,1616,1643]" pageId="15" pageNumber="16">(Figs. available at https://collections.nmnh.si.edu)</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF67AA8AFCC4F96E" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="BA7236FE193AFFDFFF67AA8AFC0BF909" blockId="15.[151,987,1684,1739]" box="[151,987,1684,1709]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="9ADD11D0193AFFDFFF67AA8AFC0BF909" box="[151,987,1684,1709]" pageId="15" pageNumber="16">
<treatmentCitation id="3B6C10EF193AFFDFFF67AA8AFE0AF908" author="Casey, T. L." box="[151,474,1684,1709]" page="321" pageId="15" pageNumber="16" year="1922">
<taxonomicName id="7DCD4D7D193AFFDFFF67AA8AFE0AF908" ID-CoL="4PJYN" authority="Casey (1922: 321" authorityName="Casey" authorityYear="1922" box="[151,474,1684,1709]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densa">
<emphasis id="88B9EAEC193AFFDFFF67AA8AFECFF909" box="[151,287,1684,1708]" italics="true" pageId="15" pageNumber="16">Psiona densa</emphasis>
<bibRefCitation id="DE5C4B0F193AFFDFFED5AA8AFE0AF908" author="Casey, T. L." box="[293,474,1685,1709]" pageId="15" pageNumber="16" pagination="1 - 520" refId="ref14311" refString="Casey, T. L. (1922) Studies in the rhynchophorous subfamily Barinae of the Brazilian fauna. Memoirs on the Coleoptera, 10, 1 - 520." type="journal article" year="1922">Casey (1922: 321</bibRefCitation>
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, description),
<treatmentCitation id="3B6C10EF193AFFDFFD9AAA8BFCCBF909" author="Davis, S. R." box="[618,795,1684,1709]" page="153" pageId="15" pageNumber="16" year="2011">
<bibRefCitation id="DE5C4B0F193AFFDFFD9AAA8BFCCBF909" author="Davis, S. R." box="[618,795,1684,1709]" pageId="15" pageNumber="16" pagination="88 - 156" refId="ref14436" refString="Davis, S. R. (2011) Delimiting baridine weevil evolution (Coleoptera: Curculionidae: Baridinae). Zoological Journal of the Linnean Society, 161 (1), 88 - 156. https: // doi. org / 10.1111 / j. 1096 - 3642.2010.00619. x" type="journal article" year="2011">Davis (2011: 153</bibRefCitation>
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, character matrix).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BA7236FE193AFFDFFF67AAABFCC4F96E" blockId="15.[151,987,1684,1739]" box="[151,788,1715,1739]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="9ADD11D0193AFFDFFF67AAABFCC4F96E" box="[151,788,1715,1739]" pageId="15" pageNumber="16">
<treatmentCitation id="3B6C10EF193AFFDFFF67AAABFE0DF96E" author="Casey, T. L." box="[151,477,1715,1739]" page="321" pageId="15" pageNumber="16" year="1922">
<taxonomicName id="7DCD4D7D193AFFDFFF67AAABFE0DF96E" ID-CoL="4PJYQ" authority="Casey (1922: 321" authorityName="Casey" authorityYear="1922" box="[151,477,1715,1739]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="88B9EAEC193AFFDFFF67AAABFEF2F96E" box="[151,290,1716,1739]" italics="true" pageId="15" pageNumber="16">Psiona opaca</emphasis>
<bibRefCitation id="DE5C4B0F193AFFDFFED8AAACFE0DF96E" author="Casey, T. L." box="[296,477,1715,1739]" pageId="15" pageNumber="16" pagination="1 - 520" refId="ref14311" refString="Casey, T. L. (1922) Studies in the rhynchophorous subfamily Barinae of the Brazilian fauna. Memoirs on the Coleoptera, 10, 1 - 520." type="journal article" year="1922">Casey (1922: 321</bibRefCitation>
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, description);
<emphasis id="88B9EAEC193AFFDFFD9EAAACFCDEF96E" bold="true" box="[622,782,1715,1739]" pageId="15" pageNumber="16">new synonymy</emphasis>
.
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<paragraph id="BA7236FE193AFFDFFF67AAE6FC03F892" blockId="15.[151,1436,1785,1919]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF67AAE6FEC2F8B6" bold="true" box="[151,274,1785,1811]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
<taxonomicName id="7DCD4D7D193AFFDFFEE7AAE6FE02F8B6" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[279,466,1785,1811]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densus">
<emphasis id="88B9EAEC193AFFDFFEE7AAE6FE02F8B6" box="[279,466,1785,1811]" italics="true" pageId="15" pageNumber="16">Chryasus densus</emphasis>
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is an elongate,
<quantity id="7D359B1B193AFFDFFD8AAAE6FD2FF8B1" box="[634,767,1785,1812]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="15" pageNumber="16" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
long species with a coarsely, somewhat confluently punctate pronotum as in
<taxonomicName id="7DCD4D7D193AFFDFFEB6AB01FE35F892" authorityName="Champion" authorityYear="1908" box="[326,485,1822,1848]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC193AFFDFFEB6AB01FE35F892" box="[326,485,1822,1848]" italics="true" pageId="15" pageNumber="16">C. cavernosus</emphasis>
</taxonomicName>
, but lacks a deeply punctate pronotal collar.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFC1FF37AB5EFE23FE08" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="BA7236FE193AFFDFFF37AB5EFD29F8DA" blockId="15.[151,1436,1785,1919]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37AB5EFE60F8FE" bold="true" box="[199,432,1857,1883]" pageId="15" pageNumber="16">Material examined.</emphasis>
<collectingCountry id="C2DA766E193AFFDFFE46AB5DFDF0F8F9" box="[438,544,1858,1884]" name="Brazil" pageId="15" pageNumber="16">BRAZIL</collectingCountry>
.
<collectingRegion id="7809F81C193AFFDFFDDEAB5DFD13F8FE" box="[558,707,1857,1884]" country="Brazil" name="Mato Grosso" pageId="15" pageNumber="16">Mato Grosso</collectingRegion>
: Chapada dos Guimarães, 1882–1886, H.H. &amp; A. Smith,
<typeStatus id="6576885C193AFFDFFACAAB5DFA4CF8F9" box="[1338,1436,1858,1884]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="7DCD4D7D193AFFDFFF46AB79FEC2F8DA" authorityName="Casey" authorityYear="1922" box="[182,274,1893,1919]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densa">
<emphasis id="88B9EAEC193AFFDFFF46AB79FEC2F8DA" box="[182,274,1893,1919]" italics="true" pageId="15" pageNumber="16">P. densa</emphasis>
</taxonomicName>
(
<collectionCode id="DCDCAE3B193AFFDFFED1AB7AFEA4F8DA" box="[289,372,1893,1919]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16" type="Museum">USNM</collectionCode>
4),
<typeStatus id="6576885C193AFFDFFE50AB7AFDD2F8DA" box="[416,514,1893,1919]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
of
<taxonomicName id="7DCD4D7D193AFFDFFDD7AB79FD25F8DA" authority="(USNM)" baseAuthorityName="USNM" box="[551,757,1893,1919]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="88B9EAEC193AFFDFFDD7AB79FD57F8DA" box="[551,647,1894,1919]" italics="true" pageId="15" pageNumber="16">P. opaca</emphasis>
(USNM)
</taxonomicName>
.
</paragraph>
<caption id="EEB266761925FFC0FF68AB88FDE6F852" ID-DOI="http://doi.org/10.5281/zenodo.13211533" ID-Zenodo-Dep="13211533" httpUri="https://zenodo.org/record/13211533/files/figure.png" pageId="16" pageNumber="17" startId="16.[152,270,1943,1967]" targetBox="[178,1408,181,1918]" targetPageId="16" targetType="figure">
<paragraph id="BA7236FE1925FFC0FF68AB88FDE6F852" blockId="16.[151,1437,1943,2039]" pageId="16" pageNumber="17">
<emphasis id="88B9EAEC1925FFC0FF68AB88FE8AF80A" bold="true" box="[152,346,1943,1967]" pageId="16" pageNumber="17">FIGURES 42–45.</emphasis>
<taxonomicName id="7DCD4D7D1925FFC0FE90AB88FE10F80A" box="[352,448,1943,1967]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="88B9EAEC1925FFC0FE90AB88FE10F80A" box="[352,448,1943,1967]" italics="true" pageId="16" pageNumber="17">Chryasus</emphasis>
</taxonomicName>
, dorsal and lateral aspects of males.
<emphasis id="88B9EAEC1925FFC0FCC2AB88FC81F80A" bold="true" box="[818,849,1943,1967]" pageId="16" pageNumber="17">42,</emphasis>
<taxonomicName id="7DCD4D7D1925FFC0FCA7AB87FBE9F80A" authority=", Volcan" authorityName="Volcan" box="[855,1081,1943,1967]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC1925FFC0FCA7AB87FC37F80A" box="[855,999,1944,1967]" italics="true" pageId="16" pageNumber="17">C. cavernosus</emphasis>
, Volcan
</taxonomicName>
in Chiriquí, Panamá;
<emphasis id="88B9EAEC1925FFC0FAE6AB88FAE6F80A" bold="true" box="[1302,1334,1943,1967]" pageId="16" pageNumber="17">43,</emphasis>
<taxonomicName id="7DCD4D7D1925FFC0FACCAB87FECBF876" authority=", Estac. Cacao" authorityName="Estac. Cacao" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1925FFC0FACCAB87FA46F80A" box="[1340,1430,1943,1967]" italics="true" pageId="16" pageNumber="17">C. clunis</emphasis>
, Estac. Cacao
</taxonomicName>
in A.C. Guanacaste, Costa Rica;
<emphasis id="88B9EAEC1925FFC0FD85ABA4FD45F876" bold="true" box="[629,661,1979,2003]" pageId="16" pageNumber="17">44,</emphasis>
<taxonomicName id="7DCD4D7D1925FFC0FD6DABA3FC50F876" authority=", Monteverde" authorityName="Monteverde" box="[669,896,1979,2003]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1925FFC0FD6DABA3FD28F876" box="[669,760,1979,2003]" italics="true" pageId="16" pageNumber="17">C. clunis</emphasis>
, Monteverde
</taxonomicName>
in Puntarenas, Costa Rica;
<emphasis id="88B9EAEC1925FFC0FB6FABA4FB6EF876" bold="true" box="[1183,1214,1979,2003]" pageId="16" pageNumber="17">45,</emphasis>
<taxonomicName id="7DCD4D7D1925FFC0FB36ABA4FA9BF876" box="[1222,1355,1979,2003]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="88B9EAEC1925FFC0FB36ABA4FAF6F876" box="[1222,1318,1979,2003]" italics="true" pageId="16" pageNumber="17">Chryasus</emphasis>
sp.
</taxonomicName>
near
<taxonomicName id="7DCD4D7D1925FFC0FA75ABA3FE8DF852" authority=", P. N. Cusuco" authorityName="P. N. Cusuco" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC1925FFC0FA75ABA3FF04F852" italics="true" pageId="16" pageNumber="17">C. clunis</emphasis>
, P.N. Cusuco
</taxonomicName>
in Cortés, Honduras.
</paragraph>
</caption>
<paragraph id="BA7236FE1924FFC1FF37AC88FE23FE08" blockId="17.[151,1437,151,429]" pageId="17" pageNumber="18">
<emphasis id="88B9EAEC1924FFC1FF37AC88FEC1FF14" bold="true" box="[199,273,151,177]" pageId="17" pageNumber="18">Notes.</emphasis>
<taxonomicName id="7DCD4D7D1924FFC1FEEBAC88FE67FF15" box="[283,439,151,176]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="88B9EAEC1924FFC1FEEBAC88FE67FF15" box="[283,439,151,176]" italics="true" pageId="17" pageNumber="18">Psiona opaca</emphasis>
</taxonomicName>
is a small female of
<taxonomicName id="7DCD4D7D1924FFC1FD42AC88FCC1FF14" authorityName="Casey" authorityYear="1922" box="[690,785,151,177]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="densa">
<emphasis id="88B9EAEC1924FFC1FD42AC88FCC1FF14" box="[690,785,151,177]" italics="true" pageId="17" pageNumber="18">P. densa</emphasis>
</taxonomicName>
, not a distinct species. All
<specimenCount id="ACCBFD771924FFC1FBA0AC88FB2BFF14" box="[1104,1275,151,177]" count="5" pageId="17" pageNumber="18" type="generic">five specimens</specimenCount>
known to me were collected by Herbert H. Smith and his wife while on contract with the Museu Nacional, Rio de Janeiro. Smith violated the contract by illegally exporting more than 300,000 arthropod specimens to the
<collectingCountry id="C2DA766E1924FFC1FB26ACC0FABDFF5C" box="[1238,1389,223,249]" name="United States of America" pageId="17" pageNumber="18">United States</collectingCountry>
and subsequently selling them to the Carnegie Museum and numerous private collectors (
<bibRefCitation id="DE5C4B0F1924FFC1FBACAD1CFAECFEB8" author="Kunzler, J. &amp; Sequeira Fernandes, A. C. &amp; Medina da Fonseca, V. M. &amp; Jraige, S." box="[1116,1340,259,285]" pageId="17" pageNumber="18" pagination="49 - 67" refId="ref14969" refString="Kunzler, J., Sequeira Fernandes, A. C., Medina da Fonseca, V. M. &amp; Jraige, S. (2011) Herbert Huntington Smith: um naturalista injusticado? Filosofia e Historia da Biologia, 6 (1), 49 - 67." type="journal article" year="2011">
Kunzler
<emphasis id="88B9EAEC1924FFC1FB30AD1CFB29FEB8" box="[1216,1273,259,285]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2011
</bibRefCitation>
), among them Casey.
<bibRefCitation id="DE5C4B0F1924FFC1FED6AD38FE01FEE4" author="Holland, W. J." box="[294,465,295,321]" pageId="17" pageNumber="18" pagination="211 - 214" refId="ref14764" refString="Holland, W. J. (1919) Herbert Huntingdon Smith. Entomological News and Proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia, 30 (8), 211 - 214." type="journal article" year="1919">
<collectingCountry id="C2DA766E1924FFC1FED6AD38FE50FEE4" box="[294,384,295,321]" name="Netherlands" pageId="17" pageNumber="18">Holland</collectingCountry>
(1919)
</bibRefCitation>
seems to have been under the impression that he and his museum had acquired Smith’s permitted share of duplicate specimens, which was the case for everything else but not for the arthropods. None of the latter material was ever returned and, without question, this issue has contributed to the present strict export policy by Brazilian authorities.
</paragraph>
</subSubSection>
</treatment>
</document>

359
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<paragraph id="BA7236FE193AFFDFFF67AE75FD9FFD21" blockId="15.[151,591,618,681]" box="[151,591,618,645]" pageId="15" pageNumber="16">
<heading id="E13A8192193AFFDFFF67AE75FD9FFD21" bold="true" box="[151,591,618,645]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="88B9EAEC193AFFDFFF67AE75FD9FFD21" bold="true" box="[151,591,618,645]" pageId="15" pageNumber="16">
14.
<taxonomicName id="7DCD4D7D193AFFDFFF33AE75FD9FFD21" ID-CoL="5YJWK" authority="Champion" authorityName="Champion" authorityYear="1908" box="[195,591,618,645]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC193AFFDFFF33AE75FE14FD20" bold="true" box="[195,452,618,645]" italics="true" pageId="15" pageNumber="16">Chryasus cavernosus</emphasis>
Champion
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF67AE90FE9AFD0C" box="[151,330,655,681]" pageId="15" pageNumber="16" type="description">
<paragraph id="BA7236FE193AFFDFFF67AE90FE9AFD0C" blockId="15.[151,591,618,681]" box="[151,330,655,681]" pageId="15" pageNumber="16">
(
<figureCitation id="22F62A7B193AFFDFFF6FAE90FF3FFD0C" box="[159,239,655,681]" captionStart="FIGURES 5–10" captionStartId="4.[152,270,1650,1674]" captionTargetBox="[151,1435,181,1603]" captionTargetId="figure-26@4.[151,1435,181,1626]" captionTargetPageId="4" captionText="FIGURES 5–10. Right hind legs showing gradient from clavate femur with minuscule, seta-bearing ventral denticles toward ventrally nearly straight femur with row of gradually enlarged teeth. 5, C. bicolor (type species of Preglyptobaris and subjective synonym of type species of Prosaldius); 6, C. cavernosus (type species of Chryasus); 7, C. persimilis (agreeing with Psiona); 8, 2.0 mm long species near C. carinulosus (type species of Psiona) and C. persimilis from Trinidad and Tobago; 9–10, 3.0 mm long species from São Paulo (Fig. 47) agreeing with Aniops, distal and proximal aspects. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13211505" httpUri="https://zenodo.org/record/13211505/files/figure.png" pageId="15" pageNumber="16">Figs. 6</figureCitation>
,
<figureCitation id="22F62A7B193AFFDFFF0BAE90FECAFD0C" box="[251,282,655,681]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="15" pageNumber="16">20</figureCitation>
,
<figureCitation id="22F62A7B193AFFDFFED5AE90FE92FD0C" box="[293,322,655,681]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="15" pageNumber="16">42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF67AECCFCAEFCAF" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="BA7236FE193AFFDFFF67AECCFCAEFCAF" blockId="15.[151,1436,723,778]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="9ADD11D0193AFFDFFF67AECCFCAEFCAF" pageId="15" pageNumber="16">
<treatmentCitation id="3B6C10EF193AFFDFFF67AECCFDA5FD4E" author="Champion, G. C." box="[151,629,723,747]" page="337" pageId="15" pageNumber="16" year="1908">
<taxonomicName id="7DCD4D7D193AFFDFFF67AECCFDA5FD4E" ID-CoL="5YJWK" authority="Champion (1908: 337" authorityName="Champion" authorityYear="1908" box="[151,629,723,747]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC193AFFDFFF67AECCFEA7FD4E" box="[151,375,723,747]" italics="true" pageId="15" pageNumber="16">Chryasus cavernosus</emphasis>
<bibRefCitation id="DE5C4B0F193AFFDFFE75AECCFDA5FD4E" author="Champion, G. C." box="[389,629,723,747]" pageId="15" pageNumber="16" pagination="14 - 19" refId="ref14342" refString="Champion, G. C. (1908) Insecta. Coleoptera. Vol. 4. Pt. 5. Rhynchophora. Curculionidae. Curculioninae (continued). In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Porter, London, pp. 241 - 400, pls. 14 - 19." type="book chapter" year="1908">Champion (1908: 337</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, description),
<treatmentCitation id="3B6C10EF193AFFDFFCE5AECCFC3EFD4E" author="Hustache, A." box="[789,1006,723,747]" page="24" pageId="15" pageNumber="16" year="1938">
<bibRefCitation id="DE5C4B0F193AFFDFFCE5AECCFC3EFD4E" author="Hustache, A." box="[789,1006,723,747]" pageId="15" pageNumber="16" pagination="1 - 219" refId="ref14849" refString="Hustache, A. (1938) Curculionidae: Barinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, ' s-Gravenhage, 30 (163), 1 - 219." type="journal article" year="1938">Hustache (1938: 24</bibRefCitation>
</treatmentCitation>
, transfer from Centrinini to Bari[di]ni, Coelonertina),
<treatmentCitation id="3B6C10EF193AFFDFFEAFAEEDFD45FCAF" author="O'Brien, C. W. &amp; Wibmer, G. J." box="[351,661,754,778]" page="181" pageId="15" pageNumber="16" year="1982">
<bibRefCitation id="DE5C4B0F193AFFDFFEAFAEEDFD45FCAF" author="O'Brien, C. W. &amp; Wibmer, G. J." box="[351,661,754,778]" pageId="15" pageNumber="16" pagination="1 - 382" refId="ref15300" refString="O'Brien, C. W. &amp; Wibmer, G. J. (1982) Annotated checklist of the weevils (Curculionidae sensu lato) of North America, Central America, and the West Indies (Coleoptera: Curculionoidea). Memoirs of the American Entomological Institute, 34, i - ix + 1 - 382." type="journal article" year="1982">O’Brien &amp; Wibmer (1982:181</bibRefCitation>
</treatmentCitation>
, new country records).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF67AF27FBB9FC3F" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="BA7236FE193AFFDFFF67AF27FBB9FC3F" blockId="15.[151,1437,824,1498]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF67AF27FEC2FCF7" bold="true" box="[151,274,824,850]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
<taxonomicName id="7DCD4D7D193AFFDFFEE9AF27FDD9FCF7" authorityName="Champion" authorityYear="1908" box="[281,521,824,850]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="88B9EAEC193AFFDFFEE9AF27FDD9FCF7" box="[281,521,824,850]" italics="true" pageId="15" pageNumber="16">Chryasus cavernosus</emphasis>
</taxonomicName>
is
<quantity id="7D359B1B193AFFDFFDDBAF27FD63FCF6" box="[555,691,824,851]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" pageId="15" pageNumber="16" unit="mm" value="2.75" valueMax="3.2" valueMin="2.3">2.3–3.2 mm</quantity>
long and entirely brown. The anterior constriction of the coarsely punctate pronotum has deep pits separated by longitudinal ridges, a trait unique to this species. Males have the basal ventrites almost unmodified, only large ones have inconspicuous appressed squamulae.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF37AFBBFED4FA7F" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="BA7236FE193AFFDFFF37AFBBFB89FC1B" blockId="15.[151,1437,824,1498]" box="[199,1113,932,959]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37AFBBFE61FC1B" bold="true" box="[199,433,932,958]" pageId="15" pageNumber="16">Material examined.</emphasis>
<materialsCitation id="0AA53CA3193AFFDFFE48AFBBFB85FC1B" box="[440,1109,932,959]" country="Costa Rica" location="Guanacaste" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Guanacaste">
<collectingCountry id="C2DA766E193AFFDFFE48AFBBFDB2FC1B" box="[440,610,932,958]" name="Costa Rica" pageId="15" pageNumber="16">COSTA RICA</collectingCountry>
.
<collectingRegion id="7809F81C193AFFDFFD80AFBBFD26FC1B" box="[624,758,932,958]" country="Costa Rica" name="Guanacaste" pageId="15" pageNumber="16">Guanacaste</collectingRegion>
: P.N.
<collectingRegion id="7809F81C193AFFDFFCC6AFBBFC6AFC1B" box="[822,954,932,958]" country="Costa Rica" name="Guanacaste" pageId="15" pageNumber="16">Guanacaste</collectingRegion>
, Estac. Pitilla
</materialsCitation>
,
</paragraph>
<paragraph id="BA7236FE193AFFDFFF67AFD7FDF1FAA7" blockId="15.[151,1437,824,1498]" pageId="15" pageNumber="16">
<date id="CE73103E193AFFDFFF67AFD7FEDBFC47" box="[151,267,968,994]" pageId="15" pageNumber="16" value="1989-02-27">27.ii.1989</date>
–
<materialsCitation id="0AA53CA3193AFFDFFEECAFD7FD75FC47" box="[284,677,968,995]" collectingDate="1994-09-01" collectionCode="MNCR" collectorName="Moraga" pageId="15" pageNumber="16" specimenCode="MNCR 18" specimenCount="1">
<date id="CE73103E193AFFDFFEECAFD7FE53FC47" box="[284,387,968,994]" pageId="15" pageNumber="16" value="1994-09-01">
<collectingDate id="DE37E9D6193AFFDFFEECAFD7FE53FC47" box="[284,387,968,994]" pageId="15" pageNumber="16" value="1994-09-01">1.ix.1994</collectingDate>
</date>
, C.
<collectorName id="17385328193AFFDFFE46AFD6FDDDFC46" box="[438,525,969,995]" pageId="15" pageNumber="16">Moraga</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFDEFAFD7FD4CFC47" box="[543,668,968,994]" collectionCode="MNCR" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="15" pageNumber="16">MNCR 18</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFD45AFD7FD74FBA3" collectingDate="1996-03-06" collectingDateMax="1996-03-12" collectingDateMin="1996-03-06" collectionCode="CMNC, JPPC, MNCR" collectorName="J. Prena &amp; P. N. Braulio Carrillo" location="Heredia" pageId="15" pageNumber="16" specimenCode="CMNC 3, JPPC 9, MNCR 4" specimenCount="1">
<date id="CE73103E193AFFDFFD45AFD7FC84FC47" box="[693,852,968,994]" pageId="15" pageNumber="16" value="1996-03-06" valueMax="1996-03-12" valueMin="1996-03-06">
<collectingDate id="DE37E9D6193AFFDFFD45AFD7FC84FC47" box="[693,852,968,994]" pageId="15" pageNumber="16" value="1996-03-06" valueMax="1996-03-12" valueMin="1996-03-06">6.–12.iii.1996</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFC93AFD6FC6FFC46" box="[867,959,969,995]" pageId="15" pageNumber="16">J. Prena</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFC21AFD7FBEFFC47" box="[977,1087,968,994]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 3</specimenCode>
,
<specimenCode id="EA6B9E85193AFFDFFBBFAFD7FB76FC47" box="[1103,1190,968,994]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 9</specimenCode>
,
<specimenCode id="EA6B9E85193AFFDFFB46AFD7FAF4FC47" box="[1206,1316,968,994]" collectionCode="MNCR" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="15" pageNumber="16">MNCR 4</specimenCode>
).
<location id="BF126025193AFFDFFACCAFD7FA49FC47" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFACCAFD7FA49FC47" box="[1340,1433,968,994]" name="Heredia" pageId="15" pageNumber="16">Heredia</location>
:
<collectorName id="17385328193AFFDFFF67AFF2FE53FBA3" box="[151,387,1004,1031]" pageId="15" pageNumber="16">P.N. Braulio Carrillo</collectorName>
,
<locationDeviation id="ABC02FDC193AFFDFFE7FAFF3FD74FBA3" box="[399,676,1004,1030]" location="Santa Clara" pageId="15" pageNumber="16">
<quantity id="7D359B1B193AFFDFFE7FAFF3FE07FBA3" box="[399,471,1004,1030]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="16" unit="km" value="13.0">13 km</quantity>
SSW Santa Clara
</locationDeviation>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFD40AFF3FB7FFBA3" box="[688,1199,1004,1031]" collectingDate="1996-01-13" collectionCode="JPPC" collectorName="J. Prena" location="Estac. El Ceibo" pageId="15" pageNumber="16" specimenCode="JPPC 1" specimenCount="1">
<date id="CE73103E193AFFDFFD40AFF3FCCCFBA3" box="[688,796,1004,1030]" pageId="15" pageNumber="16" value="1996-01-13">
<collectingDate id="DE37E9D6193AFFDFFD40AFF3FCCCFBA3" box="[688,796,1004,1030]" pageId="15" pageNumber="16" value="1996-01-13">13.i.1996</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFCD8AFF2FC52FBA2" box="[808,898,1005,1031]" pageId="15" pageNumber="16">J. Prena</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFC61AFF3FC34FBA3" box="[913,996,1004,1030]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 1</specimenCode>
);
<location id="BF126025193AFFDFFC0EAFF2FB7FFBA3" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFC0EAFF2FB7FFBA3" box="[1022,1199,1004,1031]" name="Estac. El Ceibo" pageId="15" pageNumber="16">Estac. El Ceibo</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFB4BAFF3FCDAFB8F" collectingDate="17. - 23. ii. 2003, 11. - 16. iii. 2003" collectionCode="JPPC" collectorName="J. Prena &amp; P. N. Hitoy Cerere" location="Limon" pageId="15" pageNumber="16" specimenCode="JPPC 2" specimenCount="1">
<collectingDate id="DE37E9D6193AFFDFFB4BAFF3FEC3FB8F" isEnumeration="true" pageId="15" pageNumber="16">
<date id="CE73103E193AFFDFFB4BAFF3FAB0FBA3" box="[1211,1376,1004,1030]" pageId="15" pageNumber="16" value="2003-02-17" valueMax="2003-02-23" valueMin="2003-02-17">17.–23.ii.2003</date>
,
<date id="CE73103E193AFFDFFA9CAFF3FEC3FB8F" pageId="15" pageNumber="16" value="2003-03-11" valueMax="2003-03-16" valueMin="2003-03-11">11.– 16.iii.2003</date>
</collectingDate>
,
<collectorName id="17385328193AFFDFFEEDA80EFEA5FB8E" box="[285,373,1041,1067]" pageId="15" pageNumber="16">J. Prena</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFE72A80FFE07FB8F" box="[386,471,1040,1066]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 2</specimenCode>
).
<location id="BF126025193AFFDFFE1AA80FFDE9FB8F" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFE1AA80FFDE9FB8F" box="[490,569,1040,1066]" name="Limon" pageId="15" pageNumber="16">Limón</location>
:
<collectorName id="17385328193AFFDFFDB2A80EFCDAFB8F" box="[578,778,1040,1067]" pageId="15" pageNumber="16">P.N. Hitoy Cerere</collectorName>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFCE3A80FFE11FBEB" collectingDate="1996-01-21" collectingDateMax="1996-01-29" collectingDateMin="1996-01-21" collectionCode="JPPC" collectorName="J. Prena &amp; El Pueblito" country="Guatemala" location="Ciudad de Guatemala" pageId="15" pageNumber="16" specimenCode="JPPC 3" specimenCount="1" stateProvince="Guatemala">
<date id="CE73103E193AFFDFFCE3A80FFC61FB8F" box="[787,945,1040,1066]" pageId="15" pageNumber="16" value="1996-01-21" valueMax="1996-01-29" valueMin="1996-01-21">
<collectingDate id="DE37E9D6193AFFDFFCE3A80FFC61FB8F" box="[787,945,1040,1066]" pageId="15" pageNumber="16" value="1996-01-21" valueMax="1996-01-29" valueMin="1996-01-21">21.–29.i.1996</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFC4BA80EFBC2FB8E" box="[955,1042,1041,1067]" pageId="15" pageNumber="16">J. Prena</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFBEFA80FFBA4FB8F" box="[1055,1140,1040,1066]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 3</specimenCode>
).
<collectingCountry id="C2DA766E193AFFDFFB78A80FFAEEFB8F" box="[1160,1342,1040,1066]" name="Guatemala" pageId="15" pageNumber="16">GUATEMALA</collectingCountry>
.
<collectingRegion id="7809F81C193AFFDFFABBA80FFEE5FBEB" country="Guatemala" name="Guatemala" pageId="15" pageNumber="16">Ciudad de Guatemala</collectingRegion>
,
<collectorName id="17385328193AFFDFFEB0A82BFE11FBEB" box="[320,449,1076,1102]" pageId="15" pageNumber="16">El Pueblito</collectorName>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFE3CA82BFAC2FBEB" box="[460,1298,1076,1103]" collectingDate="1991-06-11" collectionCode="CMNC" collectorName="R. Anderson" country="Guatemala" location="Puenta Parada" pageId="15" pageNumber="16" specimenCode="CMNC 1" specimenCount="1" stateProvince="Guatemala">
<date id="CE73103E193AFFDFFE3CA82BFD96FBEB" box="[460,582,1076,1102]" pageId="15" pageNumber="16" value="1991-06-11">
<collectingDate id="DE37E9D6193AFFDFFE3CA82BFD96FBEB" box="[460,582,1076,1102]" pageId="15" pageNumber="16" value="1991-06-11">11.vi.1991</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFDA1A82AFD0EFBEB" box="[593,734,1076,1103]" pageId="15" pageNumber="16">R. Anderson</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFD1CA82BFC85FBEB" box="[748,853,1076,1102]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 1</specimenCode>
);
<collectingRegion id="7809F81C193AFFDFFC9EA82BFBB4FBEB" box="[878,1124,1076,1102]" country="Guatemala" name="Guatemala" pageId="15" pageNumber="16">Ciudad de Guatemala</collectingRegion>
,
<location id="BF126025193AFFDFFB9FA82AFAC2FBEB" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFB9FA82AFAC2FBEB" box="[1135,1298,1076,1103]" country="Guatemala" name="Puenta Parada" pageId="15" pageNumber="16" stateProvince="Guatemala">Puenta Parada</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFAEDA82BFDB9FB33" collectingDate="1991-06-12" collectionCode="CMNC, JPPC, NHMUK" collectorName="R. Anderson &amp; Santo Domingo &amp; T. Belt" country="Nicaragua" location="Canal Area" pageId="15" pageNumber="16" specimenCode="CMNC 2, JPPC 1, NHMUK 2" specimenCount="1" stateProvince="Chontales" typeStatus="syntype">
<date id="CE73103E193AFFDFFAEDA82BFA48FBEB" box="[1309,1432,1076,1102]" pageId="15" pageNumber="16" value="1991-06-12">
<collectingDate id="DE37E9D6193AFFDFFAEDA82BFA48FBEB" box="[1309,1432,1076,1102]" pageId="15" pageNumber="16" value="1991-06-12">12.vi.1991</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFF67A846FEF7FBD7" box="[151,295,1112,1139]" pageId="15" pageNumber="16">R. Anderson</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFEC9A847FE78FBD7" box="[313,424,1112,1138]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 2</specimenCode>
,
<specimenCode id="EA6B9E85193AFFDFFE48A847FDDCFBD7" box="[440,524,1112,1138]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 1</specimenCode>
).
<collectingCountry id="C2DA766E193AFFDFFDD9A847FD09FBD7" box="[553,729,1112,1138]" name="Nicaragua" pageId="15" pageNumber="16">NICARAGUA</collectingCountry>
.
<collectingRegion id="7809F81C193AFFDFFD1BA847FC8EFBD7" box="[747,862,1112,1138]" country="Nicaragua" name="Chontales" pageId="15" pageNumber="16">Chontales</collectingRegion>
,
<collectorName id="17385328193AFFDFFC9DA847FBF4FBD7" box="[877,1060,1112,1138]" pageId="15" pageNumber="16">Santo Domingo</collectorName>
,
<collectorName id="17385328193AFFDFFBC2A846FB54FBD7" box="[1074,1156,1112,1139]" pageId="15" pageNumber="16">T. Belt</collectorName>
,
<typeStatus id="6576885C193AFFDFFB63A846FB25FBD6" box="[1171,1269,1113,1139]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
(
<specimenCode id="EA6B9E85193AFFDFFAF7A847FA5FFBD7" box="[1287,1423,1112,1138]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="16" type="Museum">NHMUK 2</specimenCode>
).
<collectingCountry id="C2DA766E193AFFDFFF68A862FEC8FB3D" box="[152,280,1149,1176]" name="Panama" pageId="15" pageNumber="16">PANAMÁ</collectingCountry>
.
<location id="BF126025193AFFDFFED4A863FE78FB33" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFED4A863FE78FB33" box="[292,424,1148,1174]" country="Nicaragua" name="Canal Area" pageId="15" pageNumber="16" stateProvince="Chontales">Canal Area</location>
:
<locationDeviation id="ABC02FDC193AFFDFFE44A863FDB9FB33" box="[436,617,1148,1175]" location="Paraiso" pageId="15" pageNumber="16">
<quantity id="7D359B1B193AFFDFFE44A863FE39FB33" box="[436,489,1148,1174]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="15" pageNumber="16" unit="mi" value="3.0">3 mi</quantity>
W Paraíso
</locationDeviation>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFD87A863FB22FB33" box="[631,1266,1148,1175]" collectingDate="1969-12-14" collectionCode="STRI" collectorName="Stockwell" country="Nicaragua" location="5 mi NW Gamboa" pageId="15" pageNumber="16" specimenCode="STRI 1" specimenCount="1" stateProvince="Chontales" typeStatus="syntype">
<date id="CE73103E193AFFDFFD87A863FD29FB33" box="[631,761,1148,1174]" pageId="15" pageNumber="16" value="1969-12-14">
<collectingDate id="DE37E9D6193AFFDFFD87A863FD29FB33" box="[631,761,1148,1174]" pageId="15" pageNumber="16" value="1969-12-14">14.xii.1969</collectingDate>
</date>
, H.
<collectorName id="17385328193AFFDFFCDBA863FC4BFB33" box="[811,923,1148,1174]" pageId="15" pageNumber="16">Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFC5CA863FBD0FB33" box="[940,1024,1148,1174]" collectionCode="STRI" pageId="15" pageNumber="16">STRI 1</specimenCode>
);
<location id="BF126025193AFFDFFBEBA863FB22FB33" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFBEBA863FB22FB33" box="[1051,1266,1148,1175]" country="Nicaragua" name="5 mi NW Gamboa" pageId="15" pageNumber="16" stateProvince="Chontales">
<quantity id="7D359B1B193AFFDFFBEBA863FB80FB33" box="[1051,1104,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="15" pageNumber="16" unit="mi" value="5.0">5 mi</quantity>
NW Gamboa
</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFB0FA863FD36FB1F" collectingDate="1970-02-20" collectionCode="CMNC" collectorName="H. Stockwell" country="Nicaragua" location="20.5 km NW Volcan" pageId="15" pageNumber="16" specimenCode="CMNC 1" specimenCount="1" stateProvince="Chiriqui" typeStatus="syntype">
<date id="CE73103E193AFFDFFB0FA863FAA3FB33" box="[1279,1395,1148,1174]" pageId="15" pageNumber="16" value="1970-02-20">
<collectingDate id="DE37E9D6193AFFDFFB0FA863FAA3FB33" box="[1279,1395,1148,1174]" pageId="15" pageNumber="16" value="1970-02-20">20.ii.1970</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFA71A862FED7FB1F" pageId="15" pageNumber="16">H. Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFEE5A8BFFEADFB1F" box="[277,381,1184,1210]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 1</specimenCode>
).
<collectingRegion id="7809F81C193AFFDFFE64A8BFFE26FB1F" box="[404,502,1184,1210]" country="Panama" name="Chiriqui" pageId="15" pageNumber="16">Chiriquí</collectingRegion>
:
<location id="BF126025193AFFDFFDF0A8BFFD36FB1F" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFDF0A8BFFD36FB1F" box="[512,742,1184,1211]" country="Nicaragua" name="20.5 km NW Volcan" pageId="15" pageNumber="16" stateProvince="Chiriqui">
<quantity id="7D359B1B193AFFDFFDF0A8BFFD8BFB1F" box="[512,603,1184,1211]" metricMagnitude="4" metricUnit="m" metricValue="2.05" pageId="15" pageNumber="16" unit="km" value="20.5">20.5 km</quantity>
NW Volcan
</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFD00A8BFFA48FB1F" box="[752,1432,1184,1211]" collectingDate="1996-05-20" collectionCode="CMNC" collectorName="R. Turnbow" country="Nicaragua" location="Cerro Campana" pageId="15" pageNumber="16" specimenCode="CMNC 1" specimenCount="1" stateProvince="Panama" typeStatus="syntype">
<date id="CE73103E193AFFDFFD00A8BFFCB1FB1F" box="[752,865,1184,1210]" pageId="15" pageNumber="16" value="1996-05-20">
<collectingDate id="DE37E9D6193AFFDFFD00A8BFFCB1FB1F" box="[752,865,1184,1210]" pageId="15" pageNumber="16" value="1996-05-20">20.v.1996</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFC9CA8BEFC21FB1F" box="[876,1009,1184,1211]" pageId="15" pageNumber="16">R. Turnbow</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFC0FA8BFFBB7FB1F" box="[1023,1127,1184,1210]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 1</specimenCode>
).
<collectingRegion id="7809F81C193AFFDFFB8FA8BFFB0CFB1F" box="[1151,1244,1184,1210]" country="Panama" name="Panama" pageId="15" pageNumber="16">Panamá</collectingRegion>
:
<location id="BF126025193AFFDFFB15A8BFFA48FB1F" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFB15A8BFFA48FB1F" box="[1253,1432,1184,1210]" country="Nicaragua" name="Cerro Campana" pageId="15" pageNumber="16" stateProvince="Panama">Cerro Campana</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFF67A8DBFCF3FB7B" box="[151,803,1220,1247]" collectingDate="1973-04-28" collectionCode="JPPC" collectorName="H. Hespenheide" country="Nicaragua" location="Altos de Campana" pageId="15" pageNumber="16" specimenCode="JPPC 1" specimenCount="1" stateProvince="Panama" typeStatus="syntype">
<date id="CE73103E193AFFDFFF67A8DBFEC0FB7B" box="[151,272,1220,1246]" pageId="15" pageNumber="16" value="1973-04-28">
<collectingDate id="DE37E9D6193AFFDFFF67A8DBFEC0FB7B" box="[151,272,1220,1246]" pageId="15" pageNumber="16" value="1973-04-28">28.iv.1973</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFEEDA8DAFE01FB7B" box="[285,465,1220,1247]" pageId="15" pageNumber="16">H. Hespenheide</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFE11A8DBFDE5FB7B" box="[481,565,1220,1246]" collectionCode="JPPC" pageId="15" pageNumber="16">JPPC 1</specimenCode>
);
<location id="BF126025193AFFDFFDBEA8DBFCF3FB7B" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFDBEA8DBFCF3FB7B" box="[590,803,1220,1246]" country="Nicaragua" name="Altos de Campana" pageId="15" pageNumber="16" stateProvince="Panama">Altos de Campana</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFCDFA8DBFA48FB7B" box="[815,1432,1220,1247]" collectingDate="1996-05-26" collectionCode="CMNC" collectorName="R. Turnbow" country="Nicaragua" location="Cerro Jefe" municipality="Chepo" pageId="15" pageNumber="16" specimenCode="CMNC 1" specimenCount="1" stateProvince="Panama" typeStatus="syntype">
<date id="CE73103E193AFFDFFCDFA8DBFC4FFB7B" box="[815,927,1220,1246]" pageId="15" pageNumber="16" value="1996-05-26">
<collectingDate id="DE37E9D6193AFFDFFCDFA8DBFC4FFB7B" box="[815,927,1220,1246]" pageId="15" pageNumber="16" value="1996-05-26">26.v.1996</collectingDate>
</date>
,
<collectorName id="17385328193AFFDFFC5DA8DAFBE4FB7B" box="[941,1076,1220,1247]" pageId="15" pageNumber="16">R. Turnbow</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFBB4A8DBFB7EFB7B" box="[1092,1198,1220,1246]" collectionCode="CMNC" pageId="15" pageNumber="16">CMNC 1</specimenCode>
).
<collectingMunicipality id="5A16AC84193AFFDFFB37A8DBFAC3FB7B" box="[1223,1299,1220,1246]" pageId="15" pageNumber="16">Chepo</collectingMunicipality>
,
<location id="BF126025193AFFDFFAD0A8DBFA48FB7B" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFAD0A8DBFA48FB7B" box="[1312,1432,1220,1246]" country="Nicaragua" municipality="Chepo" name="Cerro Jefe" pageId="15" pageNumber="16" stateProvince="Panama">Cerro Jefe</location>
</materialsCitation>
,
<materialsCitation id="0AA53CA3193AFFDFFF67A8F7FDCCFAA7" box="[151,540,1256,1283]" collectingDate="1976-06-16" collectionCode="STRI" collectorName="Stockwell" country="Nicaragua" pageId="15" pageNumber="16" specimenCode="STRI 1" specimenCount="1" stateProvince="Panama" typeStatus="syntype">
<date id="CE73103E193AFFDFFF67A8F7FEC1FAA7" box="[151,273,1256,1282]" pageId="15" pageNumber="16" value="1976-06-16">
<collectingDate id="DE37E9D6193AFFDFFF67A8F7FEC1FAA7" box="[151,273,1256,1282]" pageId="15" pageNumber="16" value="1976-06-16">16.vi.1976</collectingDate>
</date>
, H.
<collectorName id="17385328193AFFDFFECFA8F7FE7FFAA7" box="[319,431,1256,1282]" pageId="15" pageNumber="16">Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193AFFDFFE4EA8F7FDC0FAA7" box="[446,528,1256,1282]" collectionCode="STRI" pageId="15" pageNumber="16">STRI 1</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE193AFFDFFF37A913FBF5FAEF" blockId="15.[151,1437,824,1498]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37A913FE87FA83" bold="true" box="[199,343,1292,1318]" pageId="15" pageNumber="16">Life history.</emphasis>
<materialsCitation id="0AA53CA3193AFFDFFEADA912FBF1FAEF" collectingDate="1996-03" collectorName="de Conservacion" country="Costa Rica" location="Area de Conservacion" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Guanacaste">
In
<date id="CE73103E193AFFDFFE8BA913FDD5FA83" box="[379,517,1292,1318]" pageId="15" pageNumber="16" value="1996-03">
<collectingDate id="DE37E9D6193AFFDFFE8BA913FDD5FA83" box="[379,517,1292,1318]" pageId="15" pageNumber="16" value="1996-03">March 1996</collectingDate>
</date>
, I found many specimens on a flowering, unidentified
<taxonomicName id="7DCD4D7D193AFFDFFB91A913FAF3FA83" box="[1121,1315,1292,1318]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
in an open habitat near Pitilla field station in
<location id="BF126025193AFFDFFDE4A92EFCDDFAEF" LSID="urn:lsid:plazi:treatment:326487E8193AFFDFFF67AE75FED4FA7F:BF126025193AFFDFFDE4A92EFCDDFAEF" box="[532,781,1328,1356]" country="Costa Rica" name="Area de Conservacion" pageId="15" pageNumber="16" stateProvince="Guanacaste">
Área
<collectorName id="17385328193AFFDFFDA2A92FFCDDFAEF" box="[594,781,1328,1354]" pageId="15" pageNumber="16">de Conservación</collectorName>
</location>
<collectingRegion id="7809F81C193AFFDFFCE4A92FFC48FAEF" box="[788,920,1328,1354]" country="Costa Rica" name="Guanacaste" pageId="15" pageNumber="16">Guanacaste</collectingRegion>
,
<collectingCountry id="C2DA766E193AFFDFFC53A92FFBF1FAEF" box="[931,1057,1328,1354]" name="Costa Rica" pageId="15" pageNumber="16">Costa Rica</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE193AFFDFFF37A94BFED4FA7F" blockId="15.[151,1437,824,1498]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37A94BFEC1FACB" bold="true" box="[199,273,1364,1390]" pageId="15" pageNumber="16">Notes.</emphasis>
This is the type species of
<taxonomicName id="7DCD4D7D193AFFDFFDB7A94BFD61FACB" box="[583,689,1364,1390]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="88B9EAEC193AFFDFFDB7A94BFD61FACB" box="[583,689,1364,1390]" italics="true" pageId="15" pageNumber="16">Chryasus</emphasis>
</taxonomicName>
. A
<typeStatus id="6576885C193AFFDFFD29A94BFC90FACB" box="[729,832,1364,1390]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
designation for it is not warranted at present because the
<specimenCount id="ACCBFD77193AFFDFFF33A966FE88FA36" box="[195,344,1401,1427]" count="2" pageId="15" pageNumber="16" type="generic" typeStatus="syntype">two syntypes</specimenCount>
are from the same locality and do not threaten nomenclatural stability. Specimens from higher elevations are more elongate with a smaller prothorax (often with a median carina) than specimens from the lowlands.
</paragraph>
</subSubSection>
</treatment>
</document>

188
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<subSubSection id="F2D76575193BFFDEFF67AC88FD49FF14" box="[151,665,151,178]" pageId="14" pageNumber="15" type="nomenclature">
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<emphasis id="88B9EAEC193BFFDEFF67AC88FD49FF14" bold="true" box="[151,665,151,178]" pageId="14" pageNumber="15">
11.
<taxonomicName id="7DCD4D7D193BFFDEFF32AC88FE2CFF14" authority="Prena" authorityName="Prena" authorityYear="2024" box="[194,508,151,178]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="taironius" status="sp. nov.">
<emphasis id="88B9EAEC193BFFDEFF32AC88FE78FF14" bold="true" box="[194,424,151,178]" italics="true" pageId="14" pageNumber="15">Chryasus taironius</emphasis>
Prena
</taxonomicName>
,
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF67ACA4FB46FF5F" pageId="14" pageNumber="15" type="description">
<paragraph id="BA7236FE193BFFDEFF67ACA4FEFEFF70" blockId="14.[151,1174,151,250]" box="[151,302,187,213]" pageId="14" pageNumber="15">
(
<figureCitation id="22F62A7B193BFFDEFF6FACA4FF2DFF70" box="[159,253,187,213]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="14" pageNumber="15">Figs. 18</figureCitation>
,
<figureCitation id="22F62A7B193BFFDEFEF9ACA4FEF7FF70" box="[265,295,187,213]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="14" pageNumber="15">40</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE193BFFDEFF67ACC0FB46FF5F" blockId="14.[151,1174,151,250]" box="[151,1174,223,250]" pageId="14" pageNumber="15">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="F2D76575193BFFDEFF67AD38FD38FE08" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="BA7236FE193BFFDEFF67AD38FD38FE08" blockId="14.[151,1437,295,934]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF67AD38FEC2FEE4" bold="true" box="[151,274,295,321]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
This species and the Costa Rican
<taxonomicName id="7DCD4D7D193BFFDEFD6FAD37FCDEFEE4" box="[671,782,296,322]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC193BFFDEFD6FAD37FCDEFEE4" box="[671,782,296,322]" italics="true" pageId="14" pageNumber="15">C. montyi</emphasis>
</taxonomicName>
have the same size and shape, a pronotum with elevated narrow interspaces between fine, coalescent punctation and similar male genitalia (
<figureCitation id="22F62A7B193BFFDEFBDCAD54FB62FEC0" box="[1068,1202,331,357]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="14" pageNumber="15">Figs. 18, 19</figureCitation>
). They differ in color and
<taxonomicName id="7DCD4D7D193BFFDEFF36AD6FFE9AFE2C" authorityName="Prena" authorityYear="2024" box="[198,330,368,394]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="taironius">
<emphasis id="88B9EAEC193BFFDEFF36AD6FFE9AFE2C" box="[198,330,368,394]" italics="true" pageId="14" pageNumber="15">C. taironius</emphasis>
</taxonomicName>
has the pronotal flanks more evenly punctate.
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<emphasis id="88B9EAEC193BFFDEFCA0AD70FB93FE2C" box="[848,1091,367,393]" italics="true" pageId="14" pageNumber="15">Chryasus carinulosus</emphasis>
</taxonomicName>
is another similar, plain brown species.
<taxonomicName id="7DCD4D7D193BFFDEFF06AD8CFE05FE08" authorityName="Prena" authorityYear="2024" box="[246,469,403,429]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="88B9EAEC193BFFDEFF06AD8CFE05FE08" box="[246,469,403,429]" italics="true" pageId="14" pageNumber="15">Chryasus persimilis</emphasis>
</taxonomicName>
has shorter appendages.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF37ADA8FBB8FD0C" pageId="14" pageNumber="15" type="description">
<paragraph id="BA7236FE193BFFDEFF37ADA8FBB8FD0C" blockId="14.[151,1437,295,934]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37ADA8FE89FE74" bold="true" box="[199,345,439,465]" pageId="14" pageNumber="15">Description.</emphasis>
Total length
<quantity id="7D359B1B193BFFDEFE1CADA8FDA4FE77" box="[492,628,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="14" pageNumber="15" unit="mm" value="3.15" valueMax="3.4" valueMin="2.9">2.9–3.4 mm</quantity>
, width
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(n=4); integument variously reddish brown to black, legs partially and elytron entirely fawn; body oblong ovate with protruding humeri and curved sides; rostrum sexually dimorphic, thinner and longer in female; pronotum with fine, confluent punctation, interspaces forming narrow ridges (indistinct in female), surface texture coarser and shallower on flank; elytron with transverse depression at base, interstriae ridged, metaventrite unmodified and not sexually dimorphic, male ventrites 1, 2 and 5 depressed medially and with erect squamulae; femora with denticles gradually decreasing in size (male) or minute and subequal (female); male genitalia as
<figureCitation id="22F62A7B193BFFDEFE38AE90FDC9FD0C" box="[456,537,655,681]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="14" pageNumber="15">Fig. 18</figureCitation>
, endophallus with long, basally inflexible flagellum.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF37AEACFAE1FCB0" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="BA7236FE193BFFDEFF37AEACFAE1FCB0" blockId="14.[151,1437,295,934]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37AEACFE60FD68" bold="true" box="[199,432,691,717]" pageId="14" pageNumber="15">Material examined.</emphasis>
<materialsCitation id="0AA53CA3193BFFDEFE46AEACFD0AFD54" collectingDate="1973-05-10" collectionCode="CMNC" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus id="6576885C193BFFDEFE46AEACFDCEFD68" box="[438,542,691,717]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male, labeled “COLOM[BIA]. Magd[alena]., 7000′, San Lorenzo,
<quantity id="7D359B1B193BFFDEFAF9AEACFA82FD68" box="[1289,1362,691,717]" metricMagnitude="4" metricUnit="m" metricValue="4.1" pageId="14" pageNumber="15" unit="km" value="41.0">41 km</quantity>
. S Sta. Marta,
<date id="CE73103E193BFFDEFF18AEC8FEB2FD54" box="[232,354,727,753]" pageId="14" pageNumber="15" value="1973-05-10">
<collectingDate id="DE37E9D6193BFFDEFF18AEC8FEB2FD54" box="[232,354,727,753]" pageId="14" pageNumber="15" value="1973-05-10">V-10-1973</collectingDate>
</date>
, Campbell &amp; Howden” (
<collectionCode id="DCDCAE3B193BFFDEFD89AEC8FD02FD54" box="[633,722,727,753]" pageId="14" pageNumber="15">CMNC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="6576885C193BFFDEFD15AEC7FC82FD57" box="[741,850,728,754]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="ACCBFD77193BFFDEFC91AEC8FC6BFD54" box="[865,955,727,753]" count="2" pageId="14" pageNumber="15" type="male">2 males</specimenCount>
,
<specimenCount id="ACCBFD77193BFFDEFC37AEC8FBFEFD54" box="[967,1070,727,753]" count="1" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
): same site as
<typeStatus id="6576885C193BFFDEFB3DAEC8FAE2FD54" box="[1229,1330,727,753]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
, Howden &amp; Campbell,
<materialsCitation id="0AA53CA3193BFFDEFEDFAEE4FD15FCB0" box="[303,709,763,789]" collectingDate="1973-05-01" collectionCode="CMNC" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
<date id="CE73103E193BFFDEFEDFAEE4FE41FCB0" box="[303,401,763,789]" pageId="14" pageNumber="15" value="1973-05-01">
<collectingDate id="DE37E9D6193BFFDEFEDFAEE4FE41FCB0" box="[303,401,763,789]" pageId="14" pageNumber="15" value="1973-05-01">1.v.1973</collectingDate>
</date>
, 1 dissected male (
<collectionCode id="DCDCAE3B193BFFDEFD95AEE4FD6DFCB0" box="[613,701,763,789]" pageId="14" pageNumber="15">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA3193BFFDEFD20AEE4FBDFFCB0" box="[720,1039,763,790]" collectingDate="1973-05-05" collectionCode="CMNC" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="CE73103E193BFFDEFD20AEE4FCE3FCB3" box="[720,819,763,790]" pageId="14" pageNumber="15" value="1973-05-05">
<collectingDate id="DE37E9D6193BFFDEFD20AEE4FCE3FCB3" box="[720,819,763,790]" pageId="14" pageNumber="15" value="1973-05-05">5.v.1973</collectingDate>
</date>
,
<specimenCount id="ACCBFD77193BFFDEFCCEAEE4FC4FFCB0" box="[830,927,763,789]" count="1" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B193BFFDEFC5EAEE4FBD6FCB0" box="[942,1030,763,789]" pageId="14" pageNumber="15">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="0AA53CA3193BFFDEFBEAAEE4FAFDFCB0" box="[1050,1325,763,789]" collectingDate="1973-05-07" collectionCode="JPPC" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="CE73103E193BFFDEFBEAAEE4FBACFCB0" box="[1050,1148,763,789]" pageId="14" pageNumber="15" value="1973-05-07">
<collectingDate id="DE37E9D6193BFFDEFBEAAEE4FBACFCB0" box="[1050,1148,763,789]" pageId="14" pageNumber="15" value="1973-05-07">7.v.1973</collectingDate>
</date>
,
<specimenCount id="ACCBFD77193BFFDEFB78AEE4FB03FCB0" box="[1160,1235,763,789]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
(
<collectionCode id="DCDCAE3B193BFFDEFB12AEE4FAF5FCB0" box="[1250,1317,763,789]" pageId="14" pageNumber="15">JPPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF37AF00FC4DFC00" pageId="14" pageNumber="15" type="etymology">
<paragraph id="BA7236FE193BFFDEFF37AF00FA5BFC9C" blockId="14.[151,1437,295,934]" box="[199,1419,799,826]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37AF00FE9EFC9C" bold="true" box="[199,334,799,825]" pageId="14" pageNumber="15">Etymology.</emphasis>
The name is a Latinized adjective derived from that of the native, Pre-Colombian Tairona tribes.
</paragraph>
<paragraph id="BA7236FE193BFFDEFF37AF5CFC4DFC00" blockId="14.[151,1437,295,934]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37AF5CFEC1FCF8" bold="true" box="[199,273,835,861]" pageId="14" pageNumber="15">Notes.</emphasis>
<materialsCitation id="0AA53CA3193BFFDEFEEAAF5CFC49FC00" collectionCode="CMNC" pageId="14" pageNumber="15" specimenCount="3" specimenCount-male="3">
The only available female deviates from the
<specimenCount id="ACCBFD77193BFFDEFCEBAF5CFC71FCF8" box="[795,929,835,861]" count="3" pageId="14" pageNumber="15" type="male">three males</specimenCount>
. It is somewhat larger, almost entirely black except for the fawn elytron, the pronotal ridges are fragmented and the rostrum has a pit in front of each eye. A male of a smaller species (
<figureCitation id="22F62A7B193BFFDEFE73AF94FE04FC00" box="[387,468,907,933]" captionStart="FIGURES 46–48" captionStartId="19.[152,270,1562,1586]" captionTargetBox="[151,1435,181,1538]" captionTargetId="figure-15@19.[151,1435,181,1538]" captionTargetPageId="19" captionText="FIGURES 46–48. Chryasus, dorsal and lateral aspects of males. 46, C. persimilis, male, Estac. Pitilla in A.C. Guanacaste, Costa Rica; 47, Chryasus sp. near C. sculpturatus, Município de São Paulo, Brazil; 48, Chryasus sp., San Lorenzo in Magdalena, Colombia." figureDoi="http://doi.org/10.5281/zenodo.13211535" httpUri="https://zenodo.org/record/13211535/files/figure.png" pageId="14" pageNumber="15">Fig. 48</figureCitation>
) was captured at the same site (
<collectionCode id="DCDCAE3B193BFFDEFCC8AF94FC41FC00" box="[824,913,907,933]" pageId="14" pageNumber="15">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

136
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<mods:title id="E4111A2F421F6BE52B96E07D52C2BE71">Consolidation of Aniops Casey, Psiona Casey, Preglyptobaris Bondar and Prospoliata Hustache with Chryasus Champion (Coleoptera, Curculionidae, Baridinae) and descriptions of new species</mods:title>
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<heading id="E13A8192193BFFDEFF67AFE8FCC5FBB4" bold="true" box="[151,789,1015,1042]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="88B9EAEC193BFFDEFF67AFE8FCC5FBB4" bold="true" box="[151,789,1015,1042]" pageId="14" pageNumber="15">
12.
<taxonomicName id="7DCD4D7D193BFFDEFF33AFE8FDE1FBB4" authority="(Casey)" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[195,561,1015,1042]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinulosus" status="comb. nov.">
<emphasis id="88B9EAEC193BFFDEFF33AFE8FE1BFBB7" bold="true" box="[195,459,1015,1042]" italics="true" pageId="14" pageNumber="15">Chryasus carinulosus</emphasis>
(Casey)
</taxonomicName>
,
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<paragraph id="BA7236FE193BFFDEFF67A804FD17FB90" blockId="14.[151,789,1015,1078]" box="[151,711,1051,1078]" pageId="14" pageNumber="15">(Figs. available at https://collections.nmnh.si.edu)</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF67A87FFC93FBDD" box="[151,835,1120,1144]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="BA7236FE193BFFDEFF67A87FFC93FBDD" blockId="14.[151,835,1120,1144]" box="[151,835,1120,1144]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="9ADD11D0193BFFDEFF67A87FFC93FBDD" box="[151,835,1120,1144]" pageId="14" pageNumber="15">
<treatmentCitation id="3B6C10EF193BFFDEFF67A87FFDD9FBDD" author="Casey, T. L." box="[151,521,1120,1144]" page="274" pageId="14" pageNumber="15" year="1922">
<taxonomicName id="7DCD4D7D193BFFDEFF67A87FFDD9FBDD" ID-CoL="4PJYM" authority="Casey (1922: 274" authorityName="Casey" authorityYear="1922" box="[151,521,1120,1144]" class="Insecta" family="Curculionidae" genus="Psiona" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinulosa">
<emphasis id="88B9EAEC193BFFDEFF67A87FFE80FBDD" box="[151,336,1120,1144]" italics="true" pageId="14" pageNumber="15">Psiona carinulosa</emphasis>
<bibRefCitation id="DE5C4B0F193BFFDEFEA7A87FFDD9FBDD" author="Casey, T. L." box="[343,521,1120,1144]" pageId="14" pageNumber="15" pagination="1 - 520" refId="ref14311" refString="Casey, T. L. (1922) Studies in the rhynchophorous subfamily Barinae of the Brazilian fauna. Memoirs on the Coleoptera, 10, 1 - 520." type="journal article" year="1922">Casey (1922: 274</bibRefCitation>
</taxonomicName>
</treatmentCitation>
and 320, key and description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF67A8BAFB0CFB41" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="BA7236FE193BFFDEFF67A8BAFB0CFB41" blockId="14.[151,1436,1189,1432]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF67A8BAFEC2FB1A" bold="true" box="[151,274,1189,1215]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
This elongate brown species forms a morphologically uniform complex together with
<taxonomicName id="7DCD4D7D193BFFDEFAF2A8B9FAA1FB65" box="[1282,1393,1190,1216]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC193BFFDEFAF2A8B9FAA1FB65" box="[1282,1393,1190,1216]" italics="true" pageId="14" pageNumber="15">C. montyi</emphasis>
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,
<taxonomicName id="7DCD4D7D193BFFDEFA73A8B9FED5FB41" authorityName="Prena" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="88B9EAEC193BFFDEFA73A8B9FED5FB41" italics="true" pageId="14" pageNumber="15">C. persimilis</emphasis>
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and
<taxonomicName id="7DCD4D7D193BFFDEFECCA8D5FE11FB41" authorityName="Prena" authorityYear="2024" box="[316,449,1226,1252]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="taironius">
<emphasis id="88B9EAEC193BFFDEFECCA8D5FE11FB41" box="[316,449,1226,1252]" italics="true" pageId="14" pageNumber="15">C. taironius</emphasis>
</taxonomicName>
.
<taxonomicName id="7DCD4D7D193BFFDEFE3FA8D5FD5CFB41" box="[463,652,1226,1252]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC193BFFDEFE3FA8D5FD5CFB41" box="[463,652,1226,1252]" italics="true" pageId="14" pageNumber="15">Chryasus montyi</emphasis>
</taxonomicName>
is darker than
<taxonomicName id="7DCD4D7D193BFFDEFCC2A8D5FC06FB41" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[818,982,1226,1252]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="22" phylum="Arthropoda" rank="species" species="carinulosus">
C.
<emphasis id="88B9EAEC193BFFDEFCA3A8D5FC06FB41" box="[851,982,1226,1252]" italics="true" pageId="14" pageNumber="15">carinulosus</emphasis>
</taxonomicName>
, the others are reddish.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF37A8F2FBBCFA3D" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="BA7236FE193BFFDEFF37A8F2FAFCFAAD" blockId="14.[151,1436,1189,1432]" box="[199,1324,1261,1288]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37A8F2FE61FAA2" bold="true" box="[199,433,1261,1287]" pageId="14" pageNumber="15">Material examined.</emphasis>
<materialsCitation id="0AA53CA3193BFFDEFE47A8F1FAF8FAAD" box="[439,1320,1262,1288]" collectionCode="USNM" country="Brazil" location="Desbrochers des Loges" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
<collectingCountry id="C2DA766E193BFFDEFE47A8F1FE2EFAAD" box="[439,510,1262,1288]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
,
<collectingRegion id="7809F81C193BFFDEFDFAA8F1FDEFFAAD" box="[522,575,1262,1288]" country="Brazil" name="Para" pageId="14" pageNumber="15">Pará</collectingRegion>
, received from
<location id="BF126025193BFFDEFD1DA8F1FC23FAAD" LSID="urn:lsid:plazi:treatment:326487E8193BFFDEFF67AFE8FBBCFA3D:BF126025193BFFDEFD1DA8F1FC23FAAD" box="[749,1011,1262,1288]" country="Brazil" name="Desbrochers des Loges" pageId="14" pageNumber="15" stateProvince="Para">Desbrochers des Loges</location>
,
<specimenCount id="ACCBFD77193BFFDEFC0FA8F1FB9DFAAD" box="[1023,1101,1262,1288]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
,
<typeStatus id="6576885C193BFFDEFBA8A8F1FB6AFAAD" box="[1112,1210,1262,1288]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(
<collectionCode id="DCDCAE3B193BFFDEFB39A8F1FAF0FAAD" box="[1225,1312,1262,1288]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE193BFFDEFF37A90DFBBCFA3D" blockId="14.[151,1436,1189,1432]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37A90DFEC1FA89" bold="true" box="[199,273,1298,1324]" pageId="14" pageNumber="15">Notes.</emphasis>
<taxonomicName id="7DCD4D7D193BFFDEFEEDA90DFDC7FA89" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[285,535,1298,1324]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinulosus">
<emphasis id="88B9EAEC193BFFDEFEEDA90DFDC7FA89" box="[285,535,1298,1324]" italics="true" pageId="14" pageNumber="15">Chryasus carinulosus</emphasis>
</taxonomicName>
is known only by the
<typeStatus id="6576885C193BFFDEFCDFA90DFC45FA89" box="[815,917,1298,1324]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
. I dissected and documented in detail three morphologically similar species from Central America and one from
<collectingCountry id="C2DA766E193BFFDEFC4BA929FBFCFAF5" box="[955,1068,1334,1360]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
, to support future examinations of South American material. Presently available, morphologically similar specimens from the Amazon basin and
<collectingCountry id="C2DA766E193BFFDEFF67A961FE83FA3D" box="[151,339,1406,1432]" name="Trinidad Tobago" pageId="14" pageNumber="15">Trinidad Tobago</collectingCountry>
, all singletons, are insufficient to resolve the taxonomy of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

213
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<mods:title id="611A61486E2CAC1BFE67EE14EF4F97B5">Consolidation of Aniops Casey, Psiona Casey, Preglyptobaris Bondar and Prospoliata Hustache with Chryasus Champion (Coleoptera, Curculionidae, Baridinae) and descriptions of new species</mods:title>
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<paragraph id="BA7236FE193BFFDEFF67A9F5FD50F9A6" blockId="14.[151,1165,1513,1612]" box="[151,640,1513,1540]" pageId="14" pageNumber="15">
<heading id="E13A8192193BFFDEFF67A9F5FD50F9A6" bold="true" box="[151,640,1513,1540]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="88B9EAEC193BFFDEFF67A9F5FD50F9A6" bold="true" box="[151,640,1513,1540]" pageId="14" pageNumber="15">
13.
<taxonomicName id="7DCD4D7D193BFFDEFF33A9F6FE33F9A1" authority="Prena" authorityName="Prena" authorityYear="2024" box="[195,483,1513,1540]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montyi" status="sp. nov.">
<emphasis id="88B9EAEC193BFFDEFF33A9F6FE40F9A1" bold="true" box="[195,400,1513,1540]" italics="true" pageId="14" pageNumber="15">Chryasus montyi</emphasis>
Prena
</taxonomicName>
,
<taxonomicNameLabel id="938A5797193BFFDEFE00A9F5FD50F9A6" box="[496,640,1513,1540]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF67AA11FB5DF9E9" pageId="14" pageNumber="15" type="description">
<paragraph id="BA7236FE193BFFDEFF67AA11FE88F98D" blockId="14.[151,1165,1513,1612]" box="[151,344,1550,1576]" pageId="14" pageNumber="15">
(
<figureCitation id="22F62A7B193BFFDEFF6FAA11FF2DF98D" box="[159,253,1550,1576]" captionStart="FIGURES 13–15" captionStartId="5.[152,270,1376,1400]" captionTargetBox="[189,1398,825,1352]" captionTargetId="figure-50@5.[189,1398,825,1352]" captionTargetPageId="5" captionText="FIGURES 13–15. Changes in extension and relative position of morphological parts on the prosternum associated with increased gibbosity of the pronotum and simultaneously shortened prosternum (ID = intercoxal distance; LR = length of lateral ridge of prosternal channel). 13, Chryasus montyi, with ID &lt;LR; 14, Chryasus sp. near C. bruniceps, Costa Rica (see Fig. 33), with ID = LR; 15, Chryasus wappesi, with ID&gt; LR." figureDoi="http://doi.org/10.5281/zenodo.13211511" httpUri="https://zenodo.org/record/13211511/files/figure.png" pageId="14" pageNumber="15">Figs. 13</figureCitation>
,
<figureCitation id="22F62A7B193BFFDEFEF9AA11FEF7F98D" box="[265,295,1550,1576]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="14" pageNumber="15">19</figureCitation>
,
<figureCitation id="22F62A7B193BFFDEFEC3AA11FE80F98D" box="[307,336,1550,1576]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="14" pageNumber="15">41</figureCitation>
)
</paragraph>
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="F2D76575193BFFDEFF67AA6CFE16F973" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="BA7236FE193BFFDEFF67AA6CFE16F973" blockId="14.[151,1437,1651,2037]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF67AA6CFEC2F928" bold="true" box="[151,274,1651,1677]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
<taxonomicName id="7DCD4D7D193BFFDEFEE8AA6CFE05F928" authorityName="Prena" authorityYear="2024" box="[280,469,1651,1677]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montyi">
<emphasis id="88B9EAEC193BFFDEFEE8AA6CFE05F928" box="[280,469,1651,1677]" italics="true" pageId="14" pageNumber="15">Chryasus montyi</emphasis>
</taxonomicName>
is an elongate, brown species very near to but darker than
<taxonomicName id="7DCD4D7D193BFFDEFB99AA6BFADDF928" authorityName="Prena" authorityYear="2024" baseAuthorityName="Casey" baseAuthorityYear="2024" box="[1129,1293,1651,1678]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinulosus">
<emphasis id="88B9EAEC193BFFDEFB99AA6BFADDF928" box="[1129,1293,1651,1678]" italics="true" pageId="14" pageNumber="15">C. carinulosus</emphasis>
</taxonomicName>
from
<collectingCountry id="C2DA766E193BFFDEFAA1AA6CFA49F928" box="[1361,1433,1651,1677]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
.
<taxonomicName id="7DCD4D7D193BFFDEFF67AA88FEA7F914" authorityName="Prena" authorityYear="2024" box="[151,375,1687,1713]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="88B9EAEC193BFFDEFF67AA88FEA7F914" box="[151,375,1687,1713]" italics="true" pageId="14" pageNumber="15">Chryasus persimilis</emphasis>
</taxonomicName>
, the only other morphologically similar species recorded from
<collectingCountry id="C2DA766E193BFFDEFBB4AA88FB12F914" box="[1092,1218,1687,1713]" name="Costa Rica" pageId="14" pageNumber="15">Costa Rica</collectingCountry>
, is smaller, reddish and with a shorter rostrum.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193BFFDEFF37AAC0FEADF850" pageId="14" pageNumber="15" type="description">
<paragraph id="BA7236FE193BFFDEFF37AAC0FEADF850" blockId="14.[151,1437,1651,2037]" pageId="14" pageNumber="15">
<emphasis id="88B9EAEC193BFFDEFF37AAC0FE89F95C" bold="true" box="[199,345,1759,1785]" pageId="14" pageNumber="15">Description.</emphasis>
Total length
<quantity id="7D359B1B193BFFDEFE18AAC0FDA0F95F" box="[488,624,1759,1786]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="14" pageNumber="15" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.9–3.3 mm</quantity>
, width
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(n=21); integument nearly black, appendages, elytron and often pronotum brighter with purplish hue; body oblong ovate with protruding humeri and curved sides; rostrum sexually dimorphic, thinner and longer in female; pronotum with fine, confluent punctation, interspaces forming narrow ridges, surface texture coarser and shallower on flank and evanescent above coxa; elytron with transverse depression at base, interstriae ridged, metaventrite unmodified and not sexually dimorphic, male ventrites 1, 2 and 5 depressed medially and with erect squamulae; femora with denticles gradually decreasing in size, denticles occasionally scattered or in seemingly additional row; male genitalia as
<figureCitation id="22F62A7B193BFFDEFC2BABA8FBFFF874" box="[987,1071,1975,2001]" captionStart="FIGURES 16–25" captionStartId="7.[152,270,1943,1967]" captionTargetBox="[151,1435,381,1920]" captionTargetId="figure-98@7.[151,1435,381,1920]" captionTargetPageId="7" captionText="FIGURES 16–25. Chryasus, male genitalia, dorsal aspects. 16, C. wappesi; 17, C. macer; 18, C. taironius; 19, C. montyi; 20, C. cavernosus; 21, C. clunis (Cerro Cacao, Costa Rica); 22, Chryasus sp. nr. C. clunis (Machaquilá, Guatemala); 23, Chryasus sp. nr. C. clunis (Cerro Azul, Panamá); 24, C. persimilis; 25, Chryasus sp. nr. C. sculpturatus (São Paulo, Brazil)." figureDoi="http://doi.org/10.5281/zenodo.13211515" httpUri="https://zenodo.org/record/13211515/files/figure.png" pageId="14" pageNumber="15">Fig. 19</figureCitation>
, endophallus with long, basally inflexible flagellum.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF37AC89FF20FE2C" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="BA7236FE193AFFDFFF37AC89FBCEFEB8" blockId="15.[151,1437,150,537]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37AC89FE7FFF15" bold="true" box="[199,431,150,176]" pageId="15" pageNumber="16">Material examined.</emphasis>
<materialsCitation id="0AA53CA3193AFFDFFE44AC88FC59FF70" collectingDate="2009-06-30" collectingDateMax="2009-07-09" collectingDateMin="2009-06-30" collectionCode="MNCR" collectorName="J. Prena" country="Costa Rica" elevation="1650" latitude="10.3147" location="Monteverde" longLatPrecision="7" longitude="-84.7969" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="6576885C193AFFDFFE44AC88FDCCFF14" box="[436,540,151,177]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="C2DA766E193AFFDFFD37AC88FCA0FF14" box="[711,880,151,177]" name="Costa Rica" pageId="15" pageNumber="16">COSTA RICA</collectingCountry>
,
<collectingRegion id="7809F81C193AFFDFFC8CAC88FC2BFF14" box="[892,1019,151,177]" country="Costa Rica" name="Puntarenas" pageId="15" pageNumber="16">Puntarenas</collectingRegion>
:
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, Est. Biológica,
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,
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,
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(
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same data as holotype,
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(
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<specimenCount id="ACCBFD77193AFFDFFE4BAD1CFDC4FEB8" box="[443,532,259,285]" count="3" pageId="15" pageNumber="16" type="male">3 males</specimenCount>
,
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(
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(
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.
</paragraph>
<paragraph id="BA7236FE193AFFDFFF37AD39FF20FE2C" blockId="15.[151,1437,150,537]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37AD39FE88FEE5" bold="true" box="[199,344,294,320]" pageId="15" pageNumber="16">Life history.</emphasis>
All specimens were collected at two occasions from the same two flowering
<taxonomicName id="7DCD4D7D193AFFDFFB4EAD38FACBFEE5" box="[1214,1307,295,320]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="88B9EAEC193AFFDFFB4EAD38FACBFEE5" box="[1214,1307,295,320]" italics="true" pageId="15" pageNumber="16">Miconia</emphasis>
</taxonomicName>
shrubs in a dense secondary forest. They occurred together with
<taxonomicName id="7DCD4D7D193AFFDFFD1DAD54FC81FEC0" authorityName="Prena" authorityYear="2024" box="[749,849,331,357]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clunis">
<emphasis id="88B9EAEC193AFFDFFD1DAD54FC81FEC0" box="[749,849,331,357]" italics="true" pageId="15" pageNumber="16">C. clunis</emphasis>
</taxonomicName>
and in 2009 with a single female of a still unnamed species.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193AFFDFFF37AD8CFED9FDBC" pageId="15" pageNumber="16" type="etymology">
<paragraph id="BA7236FE193AFFDFFF37AD8CFED9FDBC" blockId="15.[151,1437,150,537]" pageId="15" pageNumber="16">
<emphasis id="88B9EAEC193AFFDFFF37AD8CFE9EFE08" bold="true" box="[199,334,403,429]" pageId="15" pageNumber="16">Etymology.</emphasis>
The species is named for the late Montgomery “Monty” Wood, valued friend and founder of Monteverde Biological Station. This hospitable research and education facility is not to be confused with the commercial tourist reserve of the former Quaker community, which did not cooperate with official inventory programs.
</paragraph>
</subSubSection>
</treatment>
</document>

270
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6.
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<emphasis id="88B9EAEC193CFFD9FF45AC88FE4DFF17" bold="true" box="[181,413,151,178]" italics="true" pageId="9" pageNumber="10">Chryasus plagiatus</emphasis>
Champion
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<paragraph id="BA7236FE193CFFD9FF67ACA4FF2AFF70" blockId="9.[151,552,151,213]" box="[151,250,187,213]" pageId="9" pageNumber="10">
(
<figureCitation id="22F62A7B193CFFD9FF6FACA4FF21FF70" box="[159,241,187,213]" captionStart="FIGURES 30–33" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-26@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 30–33. Chryasus, dorsal and lateral aspects of males. 30, C. wappesi, El Pauchi in Bolivar, Venezuela; 31, C. plagiatus, Puerto Viejo de Sarapiquí in Heredia, Costa Rica; 32, C. bruniceps, Agua Preta in Bahia, Brazil; 33, Chryasus sp., Finca E. Rojas north of Cariari in Limón, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211523" httpUri="https://zenodo.org/record/13211523/files/figure.png" pageId="9" pageNumber="10">Fig. 31</figureCitation>
)
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<taxonomicName id="7DCD4D7D193CFFD9FF67AD1FFDB8FEBD" ID-CoL="5YK8K" authority="Champion (1908: 338" authorityName="Champion" authorityYear="1908" box="[151,616,256,280]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="88B9EAEC193CFFD9FF67AD1FFEB5FEBD" box="[151,357,256,280]" italics="true" pageId="9" pageNumber="10">Chryasus plagiatus</emphasis>
<bibRefCitation id="DE5C4B0F193CFFD9FE85AD1FFDB8FEBD" author="Champion, G. C." box="[373,616,256,280]" pageId="9" pageNumber="10" pagination="14 - 19" refId="ref14342" refString="Champion, G. C. (1908) Insecta. Coleoptera. Vol. 4. Pt. 5. Rhynchophora. Curculionidae. Curculioninae (continued). In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Porter, London, pp. 241 - 400, pls. 14 - 19." type="book chapter" year="1908">Champion (1908: 338</bibRefCitation>
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, description),
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<bibRefCitation id="DE5C4B0F193CFFD9FCFBAD1FFC37FEBD" author="Hustache, A." box="[779,999,256,280]" pageId="9" pageNumber="10" pagination="1 - 219" refId="ref14849" refString="Hustache, A. (1938) Curculionidae: Barinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, ' s-Gravenhage, 30 (163), 1 - 219." type="journal article" year="1938">Hustache (1938: 24</bibRefCitation>
</treatmentCitation>
, transfer from Centrinini to Bari[di]ni, Coelonertina).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="BA7236FE193CFFD9FF67AD7BFE73FE63" blockId="9.[151,1436,356,707]" pageId="9" pageNumber="10">
<emphasis id="88B9EAEC193CFFD9FF67AD7BFEC2FEDB" bold="true" box="[151,274,356,382]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
This
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long species can be recognized readily by its distinctive color pattern. An unassigned, entirely brown specimen from
<collectingCountry id="C2DA766E193CFFD9FE09AD96FDA5FE07" box="[505,629,392,419]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
(
<figureCitation id="22F62A7B193CFFD9FD77AD97FD0AFE06" box="[647,730,392,419]" captionStart="FIGURES 30–33" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-26@10.[178,1408,181,1918]" captionTargetPageId="10" captionText="FIGURES 30–33. Chryasus, dorsal and lateral aspects of males. 30, C. wappesi, El Pauchi in Bolivar, Venezuela; 31, C. plagiatus, Puerto Viejo de Sarapiquí in Heredia, Costa Rica; 32, C. bruniceps, Agua Preta in Bahia, Brazil; 33, Chryasus sp., Finca E. Rojas north of Cariari in Limón, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211523" httpUri="https://zenodo.org/record/13211523/files/figure.png" pageId="9" pageNumber="10">Fig. 33</figureCitation>
) is very similar but has stouter appendages and punctate (not rugose) pronotal flanks.
</paragraph>
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<paragraph id="BA7236FE193CFFD9FF37ADCFFBD7FD66" blockId="9.[151,1436,356,707]" pageId="9" pageNumber="10">
<emphasis id="88B9EAEC193CFFD9FF37ADCFFE64FE4F" bold="true" box="[199,436,464,490]" pageId="9" pageNumber="10">Material examined.</emphasis>
<materialsCitation id="0AA53CA3193CFFD9FE4FADCEFCAFFDAA" collectingDate="1996-03-06" collectingDateMax="1996-03-12" collectingDateMin="1996-03-06" collectionCode="MNCR, JPPC" collectorName="C. Moraga &amp; J. Prena" country="Costa Rica" location="Estac. Pitilla" pageId="9" pageNumber="10" specimenCode="MNCR 3, JPPC 1" specimenCount="1" stateProvince="Guanacaste">
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.
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: P.N.
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:
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(
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<materialsCitation id="0AA53CA3193CFFD9FE18AE07FB61FD96" box="[488,1201,536,563]" collectingDate="2000-04-17" collectionCode="JPPC" collectorName="H. Hespenheide" country="Costa Rica" location="Estac. La Selva" pageId="9" pageNumber="10" specimenCode="JPPC 1" specimenCount="1" stateProvince="Heredia">
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:
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,
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</date>
,
<collectorName id="17385328193CFFD9FC61AE06FB94FD97" box="[913,1092,536,563]" pageId="9" pageNumber="10">H. Hespenheide</collectorName>
(
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)
</materialsCitation>
,
<materialsCitation id="0AA53CA3193CFFD9FB4FAE07FF2DFDF2" collectingDate="2001-04-16" collectionCode="JPPC" collectorName="Prena" country="Costa Rica" pageId="9" pageNumber="10" specimenCode="JPPC 1" specimenCount="1">
<date id="CE73103E193CFFD9FB4FAE07FAE7FD97" box="[1215,1335,536,562]" pageId="9" pageNumber="10" value="2001-04-16">
<collectingDate id="DE37E9D6193CFFD9FB4FAE07FAE7FD97" box="[1215,1335,536,562]" pageId="9" pageNumber="10" value="2001-04-16">16.iv.2001</collectingDate>
</date>
, J.
<collectorName id="17385328193CFFD9FAACAE06FA4CFD96" box="[1372,1436,537,563]" pageId="9" pageNumber="10">Prena</collectorName>
(
<specimenCode id="EA6B9E85193CFFD9FF6FAE22FF21FDF2" box="[159,241,573,599]" collectionCode="JPPC" pageId="9" pageNumber="10">JPPC 1</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA3193CFFD9FEF9AE23FBF6FDF2" box="[265,1062,572,599]" collectingDate="1992-08-12" collectionCode="MNCR" collectorName="E. Rojas" country="Costa Rica" county="Sector Cocori" location="Rojas" municipality="Fca." pageId="9" pageNumber="10" specimenCode="MNCR 1" specimenCount="1" stateProvince="Limon">
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:
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,
<collectingMunicipality id="5A16AC84193CFFD9FDFCAE22FDECFDF2" box="[524,572,573,599]" pageId="9" pageNumber="10">Fca.</collectingMunicipality>
E.
<location id="BF126025193CFFD9FD91AE23FD74FDF3" LSID="urn:lsid:plazi:treatment:326487E8193CFFD9FF67AC88FBD7FD66:BF126025193CFFD9FD91AE23FD74FDF3" box="[609,676,572,598]" country="Costa Rica" county="Sector Cocori" municipality="Fca." name="Rojas" pageId="9" pageNumber="10" stateProvince="Limon">Rojas</location>
,
<date id="CE73103E193CFFD9FD5FAE23FCE9FDF3" box="[687,825,572,598]" pageId="9" pageNumber="10" value="1992-08-12">
<collectingDate id="DE37E9D6193CFFD9FD5FAE23FCE9FDF3" box="[687,825,572,598]" pageId="9" pageNumber="10" value="1992-08-12">12.viii.1992</collectingDate>
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(
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)
</materialsCitation>
.
<materialsCitation id="0AA53CA3193CFFD9FBC2AE21FA4CFDF3" box="[1074,1436,572,601]" collectorName="Isla" country="Panama" location="Bocas del Toro" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Bocas del Toro">
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:
<collectorName id="17385328193CFFD9FA84AE23FA4CFDF3" box="[1396,1436,572,598]" pageId="9" pageNumber="10">Isla</collectorName>
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<materialsCitation id="0AA53CA3193CFFD9FF67AE7FFBDEFDDE" box="[151,1038,608,635]" collectingDate="2003-02-27" collectionCode="CMNC" collectorName="H. Stockwell" country="Panama" location="Colon" pageId="9" pageNumber="10" specimenCode="CMNC 1" specimenCount="1" stateProvince="Colon">
<collectingRegion id="7809F81C193CFFD9FF67AE7FFF0EFDDF" box="[151,222,608,634]" country="Panama" name="Colon" pageId="9" pageNumber="10">Colón</collectingRegion>
,
<date id="CE73103E193CFFD9FF19AE7FFE8DFDDF" box="[233,349,608,634]" pageId="9" pageNumber="10" value="2003-02-27">
<collectingDate id="DE37E9D6193CFFD9FF19AE7FFE8DFDDF" box="[233,349,608,634]" pageId="9" pageNumber="10" value="2003-02-27">27.ii.2003</collectingDate>
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,
<collectorName id="17385328193CFFD9FE98AE7EFE2CFDDF" box="[360,508,608,635]" pageId="9" pageNumber="10">H. Stockwell</collectorName>
, in flowers of
<taxonomicName id="7DCD4D7D193CFFD9FD6BAE7FFC5EFDDF" box="[667,910,608,634]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="micrantha">
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(
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<date id="CE73103E193CFFD9FBEBAE7FFB5EFDDF" box="[1051,1166,608,634]" pageId="9" pageNumber="10" value="2004-02-26">
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, H.
<collectorName id="17385328193CFFD9FB4BAE7FFAFBFDDF" box="[1211,1323,608,634]" pageId="9" pageNumber="10">Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193CFFD9FAC9AE7EFA5FFDDE" box="[1337,1423,609,635]" collectionCode="STRI" pageId="9" pageNumber="10">STRI 2</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA3193CFFD9FF67AE9BFC85FD3A" box="[151,853,644,671]" collectionCode="NHMUK" collectorName="Bugaba, G &amp; Champion" country="Panama" location="Chiriqui" pageId="9" pageNumber="10" specimenCode="NHMUK 1" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
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:
<collectorName id="17385328193CFFD9FEF4AE9BFE50FD3A" box="[260,384,644,671]" pageId="9" pageNumber="10">Bugaba, G</collectorName>
.
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, 1881,
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(
<specimenCode id="EA6B9E85193CFFD9FD38AE9AFC99FD3A" box="[712,841,645,671]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 1</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="0AA53CA3193CFFD9FC93AE9AFF2DFD66" collectingDate="1973-05-13" collectionCode="STRI" collectorName="H. Stockwell" country="Panama" location="Cerro Campana" pageId="9" pageNumber="10" specimenCode="STRI 1" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
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,
<date id="CE73103E193CFFD9FB7CAE9AFB2CFD3A" box="[1164,1276,645,671]" pageId="9" pageNumber="10" value="1973-05-13">
<collectingDate id="DE37E9D6193CFFD9FB7CAE9AFB2CFD3A" box="[1164,1276,645,671]" pageId="9" pageNumber="10" value="1973-05-13">13.v.1973</collectingDate>
</date>
,
<collectorName id="17385328193CFFD9FAF9AE9AFA4CFD3B" box="[1289,1436,644,671]" pageId="9" pageNumber="10">H. Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193CFFD9FF6FAEB6FF21FD66" box="[159,241,681,707]" collectionCode="STRI" pageId="9" pageNumber="10">STRI 1</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA3193CFFD9FEFAAEB7FBD2FD66" box="[266,1026,680,707]" collectingDate="1995-06-04" collectionCode="STRI" collectorName="El Llano-Carti &amp; H. Stockwell" country="Panama" location="Chepo" pageId="9" pageNumber="10" specimenCode="STRI 1" specimenCount="1" typeStatus="holotype">
<location id="BF126025193CFFD9FEFAAEB7FE86FD67" LSID="urn:lsid:plazi:treatment:326487E8193CFFD9FF67AC88FBD7FD66:BF126025193CFFD9FEFAAEB7FE86FD67" box="[266,342,680,706]" country="Panama" name="Chepo" pageId="9" pageNumber="10">Chepo</location>
,
<collectorName id="17385328193CFFD9FE91AEB7FDD4FD67" box="[353,516,680,706]" pageId="9" pageNumber="10">El Llano-Cartí</collectorName>
road km 8,
<date id="CE73103E193CFFD9FD7AAEB7FD26FD66" box="[650,758,680,707]" pageId="9" pageNumber="10" value="1995-06-04">
<collectingDate id="DE37E9D6193CFFD9FD7AAEB7FD26FD66" box="[650,758,680,707]" pageId="9" pageNumber="10" value="1995-06-04">4.vi.1995</collectingDate>
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,
<collectorName id="17385328193CFFD9FCF2AEB6FC44FD67" box="[770,916,680,707]" pageId="9" pageNumber="10">H. Stockwell</collectorName>
(
<specimenCode id="EA6B9E85193CFFD9FC53AEB6FC25FD66" box="[931,1013,681,707]" collectionCode="STRI" pageId="9" pageNumber="10">STRI 1</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

312
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<mods:identifier id="4133F84F0C387CF56B645390124EC38A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B26C8F060E5C3CC8950325CD5586CE52" type="Zenodo-Dep">13211499</mods:identifier>
<mods:identifier id="6924B2F0572109108EE4352969AD7109" type="ZooBank">35055BD1-1925-4947-91BA-5E1CC841071A</mods:identifier>
</mods:mods>
<treatment id="326487E8193CFFDCFF67A9FCFEC8FD68" LSID="urn:lsid:zoobank.org:act:1C8C00B9-A002-420B-979E-7E3541199E61" httpUri="http://treatment.plazi.org/id/326487E8193CFFDCFF67A9FCFEC8FD68" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="F2D76575193CFFD9FF67A9FCFD4DFA58" box="[151,669,1507,1534]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="BA7236FE193CFFD9FF67A9FCFD4DFA58" blockId="9.[151,1163,1507,1606]" box="[151,669,1507,1534]" pageId="9" pageNumber="10">
<heading id="E13A8192193CFFD9FF67A9FCFD4DFA58" bold="true" box="[151,669,1507,1534]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="88B9EAEC193CFFD9FF67A9FCFD4DFA58" bold="true" box="[151,669,1507,1534]" pageId="9" pageNumber="10">
8.
<taxonomicName id="7DCD4D7D193CFFD9FF45A9FCFDD0FA58" authority="Prena" authorityName="Prena" authorityYear="2024" box="[181,512,1507,1534]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="talamanca" status="sp. nov.">
<emphasis id="88B9EAEC193CFFD9FF45A9FCFE7DFA5B" bold="true" box="[181,429,1507,1534]" italics="true" pageId="9" pageNumber="10">Chryasus talamanca</emphasis>
Prena
</taxonomicName>
,
<taxonomicNameLabel id="938A5797193CFFD9FDFEA9FCFD4DFA58" box="[526,669,1507,1533]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193CFFD9FF67AA18FB5BF9E3" pageId="9" pageNumber="10" type="description">
<paragraph id="BA7236FE193CFFD9FF67AA18FEFEF987" blockId="9.[151,1163,1507,1606]" box="[151,302,1543,1570]" pageId="9" pageNumber="10">
(
<figureCitation id="22F62A7B193CFFD9FF6FAA18FEF6F987" box="[159,294,1543,1570]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="9" pageNumber="10">Figs. 34, 35</figureCitation>
)
</paragraph>
<paragraph id="BA7236FE193CFFD9FF67AA34FB5BF9E3" blockId="9.[151,1163,1507,1606]" box="[151,1163,1579,1606]" pageId="9" pageNumber="10">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="CE6B0C2B193CFFD9FD75AA34FB5BF9E3" box="[645,1163,1579,1606]" pageId="9" pageNumber="10">1C8C00B9-A002-420B-979E-7E3541199E61</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193CFFD9FF67AA6CFA56F95C" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="BA7236FE193CFFD9FF67AA6CFA56F95C" blockId="9.[151,1437,1651,2038]" pageId="9" pageNumber="10">
<emphasis id="88B9EAEC193CFFD9FF67AA6CFEC2F928" bold="true" box="[151,274,1651,1677]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
<taxonomicName id="7DCD4D7D193CFFD9FEEBAA6CFDD4F928" authorityName="Prena" authorityYear="2024" box="[283,516,1651,1677]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis id="88B9EAEC193CFFD9FEEBAA6CFDD4F928" box="[283,516,1651,1677]" italics="true" pageId="9" pageNumber="10">Chryasus talamanca</emphasis>
</taxonomicName>
is one of several elongate species with a subconical prothorax. The
<typeStatus id="6576885C193CFFD9FAEEAA6CFA50F928" box="[1310,1408,1651,1677]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is an almost completely black male with a distinctive yellow elytral fascia (
<figureCitation id="22F62A7B193CFFD9FC3FAA88FBF3F914" box="[975,1059,1687,1713]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="9" pageNumber="10">Fig. 34</figureCitation>
), the smaller female
<typeStatus id="6576885C193CFFD9FAFFAA87FABFF917" box="[1295,1391,1688,1714]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
has dorsum and appendages mostly reddish instead of black (
<figureCitation id="22F62A7B193CFFD9FCFDAAA4FC8FF973" box="[781,863,1723,1750]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="9" pageNumber="10">Fig. 35</figureCitation>
). A lack of representative collections does not allow at present the distinction between local populations and color varieties. See notes below for further explanations.
</paragraph>
</subSubSection>
<subSubSection id="F2D76575193CFFD9FF37AB1CFD91F850" pageId="9" pageNumber="10" type="description">
<paragraph id="BA7236FE193CFFD9FF37AB1CFD91F850" blockId="9.[151,1437,1651,2038]" pageId="9" pageNumber="10">
<emphasis id="88B9EAEC193CFFD9FF37AB1CFE89F8B8" bold="true" box="[199,345,1795,1821]" pageId="9" pageNumber="10">Description.</emphasis>
Total length 2.7–3.0 mm, width
<quantity id="7D359B1B193CFFD9FD1BAB1CFCA8F8BB" box="[747,888,1795,1822]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="9" pageNumber="10" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
(n=2); integument black and red (particularly appendages), elytron with irregular yellow ochre fascia between interstriae 2–8; body elongate cuneate with slightly protruding humeri and gently curved sides; pronotum subconical, disc densely vermiculate, flanks smoother with coarse shallow ridges; elytron 2.6x longer than pronotum, disc at base with very subtle transverse depression, interstriae ridged, metaventrite tumid with basally deep discrimen, male ventrites 1, 2, 5 and metaventrite depressed medially, 1, 2 and 5 with erect squamulae; femora with 1 distinct and several minute denticles (barely perceptible on profemur); genitalia not examined.
</paragraph>
</subSubSection>
<caption id="EEB26676193FFFDAFF68AB88FD66F852" ID-DOI="http://doi.org/10.5281/zenodo.13211523" ID-Zenodo-Dep="13211523" httpUri="https://zenodo.org/record/13211523/files/figure.png" pageId="10" pageNumber="11" startId="10.[152,270,1943,1967]" targetBox="[178,1408,181,1918]" targetPageId="10" targetType="figure">
<paragraph id="BA7236FE193FFFDAFF68AB88FD66F852" blockId="10.[151,1437,1943,2039]" pageId="10" pageNumber="11">
<emphasis id="88B9EAEC193FFFDAFF68AB88FE8EF80A" bold="true" box="[152,350,1943,1967]" pageId="10" pageNumber="11">FIGURES 30–33.</emphasis>
<taxonomicName id="7DCD4D7D193FFFDAFE97AB88FE17F80A" box="[359,455,1943,1967]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="88B9EAEC193FFFDAFE97AB88FE17F80A" box="[359,455,1943,1967]" italics="true" pageId="10" pageNumber="11">Chryasus</emphasis>
</taxonomicName>
, dorsal and lateral aspects of males.
<emphasis id="88B9EAEC193FFFDAFCA1AB88FCA0F80A" bold="true" box="[849,880,1943,1967]" pageId="10" pageNumber="11">30,</emphasis>
<taxonomicName id="7DCD4D7D193FFFDAFC8AAB87FBB2F80A" authority=", El Pauchi" authorityName="El Pauchi" box="[890,1122,1943,1967]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="wappesi">
<emphasis id="88B9EAEC193FFFDAFC8AAB87FC3DF80A" box="[890,1005,1944,1967]" italics="true" pageId="10" pageNumber="11">C. wappesi</emphasis>
, El Pauchi
</taxonomicName>
in Bolivar, Venezuela;
<emphasis id="88B9EAEC193FFFDAFAACAB88FAABF80A" bold="true" box="[1372,1403,1943,1967]" pageId="10" pageNumber="11">31,</emphasis>
<taxonomicName id="7DCD4D7D193FFFDAFA75AB87FDDBF876" authority=", Puerto Viejo de Sarapiqui" authorityName="Puerto Viejo de Sarapiqui" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="88B9EAEC193FFFDAFA75AB87FF25F876" italics="true" pageId="10" pageNumber="11">C. plagiatus</emphasis>
, Puerto Viejo de Sarapiquí
</taxonomicName>
in Heredia, Costa Rica;
<emphasis id="88B9EAEC193FFFDAFCFAABA4FCFAF876" bold="true" box="[778,810,1979,2003]" pageId="10" pageNumber="11">32,</emphasis>
<taxonomicName id="7DCD4D7D193FFFDAFCC1ABA3FBE3F876" authority=", Agua Preta" authorityName="Agua Preta" box="[817,1075,1979,2003]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bruniceps">
<emphasis id="88B9EAEC193FFFDAFCC1ABA3FC64F876" box="[817,948,1979,2003]" italics="true" pageId="10" pageNumber="11">C. bruniceps</emphasis>
, Agua Preta
</taxonomicName>
in Bahia, Brazil;
<emphasis id="88B9EAEC193FFFDAFB1BABA4FADAF876" bold="true" box="[1259,1290,1979,2003]" pageId="10" pageNumber="11">33,</emphasis>
<taxonomicName id="7DCD4D7D193FFFDAFAE2ABA4FA48F876" box="[1298,1432,1979,2003]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="88B9EAEC193FFFDAFAE2ABA4FAA2F876" box="[1298,1394,1979,2003]" italics="true" pageId="10" pageNumber="11">Chryasus</emphasis>
sp.
</taxonomicName>
, Finca E. Rojas north of Cariari in Limón, Costa Rica.
</paragraph>
</caption>
<caption id="EEB26676193EFFDBFF68AB88FD2FF852" ID-DOI="http://doi.org/10.5281/zenodo.13211525" ID-Zenodo-Dep="13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="11" pageNumber="12" startId="11.[152,270,1943,1967]" targetBox="[178,1408,181,1918]" targetPageId="11" targetType="figure">
<paragraph id="BA7236FE193EFFDBFF68AB88FD2FF852" blockId="11.[151,1437,1943,2039]" pageId="11" pageNumber="12">
<emphasis id="88B9EAEC193EFFDBFF68AB88FE8BF80A" bold="true" box="[152,347,1943,1967]" pageId="11" pageNumber="12">FIGURES 34–47.</emphasis>
<taxonomicName id="7DCD4D7D193EFFDBFE91AB88FDE2F80A" authorityName="Prena" authorityYear="2024" box="[353,562,1943,1967]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis id="88B9EAEC193EFFDBFE91AB88FDE2F80A" box="[353,562,1943,1967]" italics="true" pageId="11" pageNumber="12">Chryasus talamanca</emphasis>
</taxonomicName>
complex, dorsal and lateral aspects.
<emphasis id="88B9EAEC193EFFDBFC58AB88FC18F80A" bold="true" box="[936,968,1943,1967]" pageId="11" pageNumber="12">34,</emphasis>
Male holotype, Buquete in Chiriquí, Panamá;
<emphasis id="88B9EAEC193EFFDBFF67ABA4FF67F876" bold="true" box="[151,183,1979,2003]" pageId="11" pageNumber="12">35,</emphasis>
Female paratype, Buquete in Chiriquí, Panamá;
<emphasis id="88B9EAEC193EFFDBFD52ABA4FD12F876" bold="true" box="[674,706,1979,2003]" pageId="11" pageNumber="12">36,</emphasis>
Teneral female from gall of
<taxonomicName id="7DCD4D7D193EFFDBFC11ABA3FAD7F876" authority=", Monteverde" authorityName="Monteverde" box="[993,1287,1979,2003]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="pittieri">
<emphasis id="88B9EAEC193EFFDBFC11ABA3FB51F876" box="[993,1153,1980,2003]" italics="true" pageId="11" pageNumber="12">Miconia pittieri</emphasis>
, Monteverde
</taxonomicName>
in Puntarenas, Costa Rica;
<emphasis id="88B9EAEC193EFFDBFEE3ABC0FEE3F852" bold="true" box="[275,307,2015,2039]" pageId="11" pageNumber="12">37,</emphasis>
Female, P.N. Tapantí in Cartago, Costa Rica.
</paragraph>
</caption>
<subSubSection id="F2D765751939FFDCFF37AC89FCDDFD20" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="BA7236FE1939FFDCFF37AC89FC59FF5C" blockId="12.[151,1437,150,717]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37AC89FE64FF15" bold="true" box="[199,436,150,176]" pageId="12" pageNumber="13">Material examined.</emphasis>
<materialsCitation id="0AA53CA31939FFDCFE4FAC88FE17FF70" collectingDate="1981-06-25" collectingDateMax="1981-06-26" collectingDateMin="1981-06-25" collectionCode="CMNC" collectorName="B. Gill" country="Panama" elevation="1700" location="Boquete" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<typeStatus id="6576885C1939FFDCFE4FAC88FDF7FF14" box="[447,551,151,177]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="C2DA766E1939FFDCFD12AC87FCB3FF16" box="[738,867,152,179]" name="Panama" pageId="12" pageNumber="13">PANAMÁ</collectingCountry>
:
<collectingRegion id="7809F81C1939FFDCFC81AC88FC01FF14" box="[881,977,151,177]" country="Panama" name="Chiriqui" pageId="12" pageNumber="13">Chiriquí</collectingRegion>
,
<quantity id="7D359B1B1939FFDCFC10AC88FBCCFF14" box="[992,1052,151,177]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="km" value="8.0">8 km</quantity>
NW
<location id="BF1260251939FFDCFB90AC88FB10FF14" LSID="urn:lsid:plazi:treatment:326487E8193CFFDCFF67A9FCFEC8FD68:BF1260251939FFDCFB90AC88FB10FF14" box="[1120,1216,151,177]" country="Panama" name="Boquete" pageId="12" pageNumber="13" stateProvince="Chiriqui">Boquete</location>
,
<date id="CE73103E1939FFDCFB3FAC89FA47FF14" box="[1231,1431,150,177]" pageId="12" pageNumber="13" value="1981-06-25" valueMax="1981-06-26" valueMin="1981-06-25">
<collectingDate id="DE37E9D61939FFDCFB3FAC89FA47FF14" box="[1231,1431,150,177]" pageId="12" pageNumber="13" value="1981-06-25" valueMax="1981-06-26" valueMin="1981-06-25">25–26 June 1981</collectingDate>
</date>
,
<collectorName id="173853281939FFDCFF67ACA4FF37FF70" box="[151,231,187,213]" pageId="12" pageNumber="13">B. Gill</collectorName>
,
<quantity id="7D359B1B1939FFDCFF04ACA4FE9CFF70" box="[244,332,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="m" value="1700.0">
<elevation id="31E0D1CD1939FFDCFF04ACA4FE9CFF70" box="[244,332,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="m" value="1700.0">1700 m</elevation>
</quantity>
” (
<collectionCode id="DCDCAE3B1939FFDCFE96ACA4FE6FFF70" box="[358,447,187,213]" pageId="12" pageNumber="13">CMNC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="6576885C1939FFDCFE22ACA4FDE4FF70" box="[466,564,187,213]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
(
<specimenCount id="ACCBFD771939FFDCFDB7ACA4FD7EFF70" box="[583,686,187,213]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
):
<materialsCitation id="0AA53CA31939FFDCFD31ACA4FC55FF5C" collectingDate="1996-06-15" collectionCode="CMNC" collectorName="R. Anderson" country="Panama" elevation="1800" location="La Culebra Trail" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Panama, Chiriqui" typeStatus="paratype">
<collectingRegion id="7809F81C1939FFDCFD31ACA4FC59FF70" box="[705,905,187,213]" isEnumeration="true" pageId="12" pageNumber="13">Panamá, Chiriquí</collectingRegion>
,
<locationDeviation id="ABC02FDC1939FFDCFC65ACA5FBB9FF70" box="[917,1129,186,213]" location="Boquete" pageId="12" pageNumber="13">
<quantity id="7D359B1B1939FFDCFC65ACA5FC34FF70" box="[917,996,186,213]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="13" unit="km" value="5.6">5.6 km</quantity>
N Boquete
</locationDeviation>
,
<location id="BF1260251939FFDCFB85ACA4FAE3FF70" LSID="urn:lsid:plazi:treatment:326487E8193CFFDCFF67A9FCFEC8FD68:BF1260251939FFDCFB85ACA4FAE3FF70" box="[1141,1331,187,213]" country="Panama" name="La Culebra Trail" pageId="12" pageNumber="13" stateProvince="Panama, Chiriqui">La Culebra Trail</location>
,
<quantity id="7D359B1B1939FFDCFACFACA4FA47FF70" box="[1343,1431,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">
<elevation id="31E0D1CD1939FFDCFACFACA4FA47FF70" box="[1343,1431,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="CE73103E1939FFDCFF67ACC1FEC2FF5C" box="[151,274,222,249]" pageId="12" pageNumber="13" value="1996-06-15">
<collectingDate id="DE37E9D61939FFDCFF67ACC1FEC2FF5C" box="[151,274,222,249]" pageId="12" pageNumber="13" value="1996-06-15">15.vi.1996</collectingDate>
</date>
,
<collectorName id="173853281939FFDCFEEDACC0FE7CFF5C" box="[285,428,223,249]" pageId="12" pageNumber="13">R. Anderson</collectorName>
, 96-131E, wet cloud forest litter (
<collectionCode id="DCDCAE3B1939FFDCFCD5ACC0FCAEFF5C" box="[805,894,223,249]" pageId="12" pageNumber="13">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1939FFDCFF37AD1DFDFBFEC0" blockId="12.[151,1437,150,717]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37AD1DFE6DFEB9" bold="true" box="[199,445,258,284]" pageId="12" pageNumber="13">Additional material.</emphasis>
<materialsCitation id="0AA53CA31939FFDCFE37AD1CFA48FEB8" box="[455,1432,259,285]" collectingDate="2003-03-27" collectionCode="JPPC" collectorName="P. N. Tapanti" country="Costa Rica" location="Estac. La Esperanza" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Cartago">
<collectingCountry id="C2DA766E1939FFDCFE37AD1CFD94FEB8" box="[455,580,259,285]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
,
<collectingRegion id="7809F81C1939FFDCFDA5AD1CFD60FEB8" box="[597,688,259,285]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="13">Cartago</collectingRegion>
,
<collectorName id="173853281939FFDCFD4FAD1CFC9EFEB8" box="[703,846,259,285]" pageId="12" pageNumber="13">P.N. Tapantí</collectorName>
,
<location id="BF1260251939FFDCFCACAD1CFB97FEB8" LSID="urn:lsid:plazi:treatment:326487E8193CFFDCFF67A9FCFEC8FD68:BF1260251939FFDCFCACAD1CFB97FEB8" box="[860,1095,259,285]" country="Costa Rica" name="Estac. La Esperanza" pageId="12" pageNumber="13" stateProvince="Cartago">Estac. La Esperanza</location>
,
<date id="CE73103E1939FFDCFBA6AD1CFB19FEB8" box="[1110,1225,259,285]" pageId="12" pageNumber="13" value="2003-03-27">
<collectingDate id="DE37E9D61939FFDCFBA6AD1CFB19FEB8" box="[1110,1225,259,285]" pageId="12" pageNumber="13" value="2003-03-27">27.3.2003</collectingDate>
</date>
,
<specimenCount id="ACCBFD771939FFDCFB27AD1CFAEBFEB8" box="[1239,1339,259,285]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectionCode id="DCDCAE3B1939FFDCFABDAD1CFA40FEB8" box="[1357,1424,259,285]" pageId="12" pageNumber="13">JPPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AA53CA31939FFDCFF67AD38FDF6FEC0" collectingDate="2002-03-20" collectionCode="JPPC" collectorName="P. N. Braulio Carrillo &amp; K. Nishida" country="Costa Rica" elevation="2000" location="Vara Blanca" pageId="12" pageNumber="13" specimenCode="JPPC 1" specimenCount="1" specimenCount-female="1" stateProvince="Heredia">
<collectingCountry id="C2DA766E1939FFDCFF67AD38FEC3FEE4" box="[151,275,295,321]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
,
<collectingRegion id="7809F81C1939FFDCFEEEAD38FEAAFEE4" box="[286,378,295,321]" country="Costa Rica" name="Heredia" pageId="12" pageNumber="13">Heredia</collectingRegion>
,
<collectorName id="173853281939FFDCFE74AD38FDBDFEE4" box="[388,621,295,321]" pageId="12" pageNumber="13">P.N. Braulio Carrillo</collectorName>
,
<quantity id="7D359B1B1939FFDCFD87AD38FD60FEE4" box="[631,688,295,321]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="km" value="6.0">6 km</quantity>
ENE
<location id="BF1260251939FFDCFD02AD38FCACFEE4" LSID="urn:lsid:plazi:treatment:326487E8193CFFDCFF67A9FCFEC8FD68:BF1260251939FFDCFD02AD38FCACFEE4" box="[754,892,295,321]" country="Costa Rica" name="Vara Blanca" pageId="12" pageNumber="13" stateProvince="Heredia">Vara Blanca</location>
,
<quantity id="7D359B1B1939FFDCFC76AD38FC0DFEE4" box="[902,989,295,321]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="2000.0">
<elevation id="31E0D1CD1939FFDCFC76AD38FC0DFEE4" box="[902,989,295,321]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="CE73103E1939FFDCFC18AD38FB8AFEE4" box="[1000,1114,295,321]" pageId="12" pageNumber="13" value="2002-03-20">
<collectingDate id="DE37E9D61939FFDCFC18AD38FB8AFEE4" box="[1000,1114,295,321]" pageId="12" pageNumber="13" value="2002-03-20">20.3.2002</collectingDate>
</date>
,
<collectorName id="173853281939FFDCFB95AD38FB30FEE4" box="[1125,1248,295,321]" pageId="12" pageNumber="13">K. Nishida</collectorName>
,
<specimenCount id="ACCBFD771939FFDCFB1BAD38FA9BFEE4" box="[1259,1355,295,321]" count="1" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
ex leaf petiole of
<taxonomicName id="7DCD4D7D1939FFDCFEF9AD54FE69FEC1" box="[265,441,331,356]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="pittieri">
<emphasis id="88B9EAEC1939FFDCFEF9AD54FE69FEC1" box="[265,441,331,356]" italics="true" pageId="12" pageNumber="13">Miconia pittieri</emphasis>
</taxonomicName>
(
<specimenCode id="EA6B9E851939FFDCFE38AD54FDCAFEC0" box="[456,538,331,357]" collectionCode="JPPC" pageId="12" pageNumber="13">JPPC 1</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1939FFDCFF37AD70FDBEFDBC" blockId="12.[151,1437,150,717]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37AD70FEC1FE2C" bold="true" box="[199,273,367,393]" pageId="12" pageNumber="13">Notes.</emphasis>
<materialsCitation id="0AA53CA31939FFDCFEEAAD70FDBAFDBC" collectionCode="JPPC" country="Costa Rica" location="The" municipality="Monteverde" pageId="12" pageNumber="13" specimenCount="3" typeStatus="holotype">
I consider a small reddish female (
<figureCitation id="22F62A7B1939FFDCFD5AAD70FD2DFE2C" box="[682,765,366,393]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="12" pageNumber="13">Fig. 35</figureCitation>
) from the type locality, with the same elytral fascia as the black
<typeStatus id="6576885C1939FFDCFF2AAD8CFEEFFE08" box="[218,319,403,429]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, as conspecific and include it as a
<typeStatus id="6576885C1939FFDCFD46AD8CFCCAFE08" box="[694,794,403,429]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
. Three morphologically similar females (
<collectionCode id="DCDCAE3B1939FFDCFB11AD8CFAF1FE08" box="[1249,1313,403,429]" pageId="12" pageNumber="13">JPPC</collectionCode>
) from high elevations in
<collectingCountry id="C2DA766E1939FFDCFEC0ADA8FE7EFE74" box="[304,430,439,465]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
, all singleton from different places, seem to represent different species.
<location id="BF1260251939FFDCFB13ADA8FADFFE74" LSID="urn:lsid:plazi:treatment:326487E8193CFFDCFF67A9FCFEC8FD68:BF1260251939FFDCFB13ADA8FADFFE74" box="[1251,1295,439,465]" country="Costa Rica" municipality="Monteverde" name="The" pageId="12" pageNumber="13">The</location>
two reddish ones (
<figureCitation id="22F62A7B1939FFDCFF2CADC4FEB7FE50" box="[220,359,475,501]" captionStart="FIGURES 34–47" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[178,1408,181,1918]" captionTargetId="figure-15@11.[178,1408,181,1918]" captionTargetPageId="11" captionText="FIGURES 34–47. Chryasus talamanca complex, dorsal and lateral aspects. 34, Male holotype, Buquete in Chiriquí, Panamá; 35, Female paratype, Buquete in Chiriquí, Panamá; 36, Teneral female from gall of Miconia pittieri, Monteverde in Puntarenas, Costa Rica; 37, Female, P.N. Tapantí in Cartago, Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13211525" httpUri="https://zenodo.org/record/13211525/files/figure.png" pageId="12" pageNumber="13">Figs. 36, 37</figureCitation>
) are very similar to
<taxonomicName id="7DCD4D7D1939FFDCFDA5ADC4FD3EFE50" authorityName="Prena" authorityYear="2024" box="[597,750,475,501]" class="Insecta" family="Curculionidae" genus="Chryasus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis id="88B9EAEC1939FFDCFDA5ADC4FD3EFE50" box="[597,750,475,501]" italics="true" pageId="12" pageNumber="13">C. talamanca</emphasis>
</taxonomicName>
and therefore included under
<typeStatus id="6576885C1939FFDCFBBAADC4FAF2FE50" box="[1098,1314,475,501]" isOtherTypeStatus="true" pageId="12" pageNumber="13">additional material</typeStatus>
. A smaller black specimen was taken in
<collectingMunicipality id="5A16AC841939FFDCFE2FADE0FDBAFDBC" box="[479,618,511,537]" pageId="12" pageNumber="13">Monteverde</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="BA7236FE1939FFDCFF37AE3DFCDDFD20" blockId="12.[151,1437,150,717]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37AE3DFE8CFD99" bold="true" box="[199,348,546,572]" pageId="12" pageNumber="13">Life history.</emphasis>
Kenji Nishida discovered swollen leaf petioles on
<taxonomicName id="7DCD4D7D1939FFDCFC45AE3CFB68FD98" authority="Cogn." authorityName="Cogn." box="[949,1208,547,573]" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="pittieri">
<emphasis id="88B9EAEC1939FFDCFC45AE3CFBB9FD99" box="[949,1129,547,572]" italics="true" pageId="12" pageNumber="13">Miconia pittieri</emphasis>
Cogn.
</taxonomicName>
(
<figureCitation id="22F62A7B1939FFDCFB3AAE3CFAC2FD98" box="[1226,1298,547,573]" captionStart="FIGURES 1–4" captionStartId="1.[152,270,1403,1427]" captionTargetBox="[151,1435,417,1379]" captionTargetId="figure-119@1.[151,1435,417,1379]" captionTargetPageId="1" captionText="FIGURES 1–4. Development of specimens near Chryasus talamanca in leaf petioles of Miconia pittieri. 1, slender, elongate galls; 2, short, more swollen galls; 3, dissected gall with pupa; 4, teneral female emerging from dissected gall. Photos by Kenji Nishida taken in Parque Nacional Braulio Carrillo at 2000 m elevation on 11.iv.2002." figureDoi="http://doi.org/10.5281/zenodo.13211503" httpUri="https://zenodo.org/record/13211503/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
) at
<quantity id="7D359B1B1939FFDCFAB4AE3CFA4CFD98" box="[1348,1436,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="2000.0">2000 m</quantity>
elevation in Braulio Carrillo National Park. When he opened these galls, most were parasitized. He obtained one teneral female and took the images used for figures 1–4.
</paragraph>
</subSubSection>
<subSubSection id="F2D765751939FFDCFF37AE90FEC8FD68" pageId="12" pageNumber="13" type="etymology">
<paragraph id="BA7236FE1939FFDCFF37AE90FEC8FD68" blockId="12.[151,1437,150,717]" pageId="12" pageNumber="13">
<emphasis id="88B9EAEC1939FFDCFF37AE90FE9EFD0C" bold="true" box="[199,334,655,681]" pageId="12" pageNumber="13">Etymology.</emphasis>
The name is a noun in apposition referring to the known range of this species, the Cordillera de Talamanca.
</paragraph>
</subSubSection>
</treatment>
</document>