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<mods:affiliation id="DAADF7D0C821864C1190C7411AA22F66">Institute of Paleobiology, Polish Academy of Sciences, Twarda 51 / 55, 00 - 818 Warsaw, Poland & Institute of Evolutionary Biology, Biological and Chemical Research Centre, Faculty of Biology, University of Warsaw, ul. Żwirki i Wigury 101, 02 - 089 Warsaw, Poland</mods:affiliation>
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<taxonomicName id="4C2D4D25FFBFFF8FFED1FB4EF2322DBD" authority="Osi et al. 2010" authorityName="Osi" authorityYear="2010" box="[269,636,1200,1232]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="kozmai">
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@ -70,7 +73,7 @@
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<specimenCode id="DB8B9EDDFFBFFF8FFF35FB0CF1F92C7F" box="[233,439,1265,1289]" pageId="2" pageNumber="3">MTM V2009.192.1</specimenCode>
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, a proposed rostral, premaxillae, and the partial anteromedial processes of maxillae (
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<figureCitation id="13162A23FFBFFF8FFD57FAEFF2AF2C5E" box="[651,737,1297,1321]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStartId-0="3.[113,178,1823,1847]" captionStartId-1="4.[129,194,1840,1864]" captionStartId-2="5.[113,178,1411,1435]" captionStartId-3="6.[128,193,1509,1533]" captionStartId-4="7.[115,180,1219,1243]" captionStartId-5="8.[129,194,1187,1211]" captionTargetBox-0="[117,1455,147,1791]" captionTargetBox-1="[161,1441,148,1807]" captionTargetBox-2="[117,1454,147,1380]" captionTargetBox-3="[132,1469,149,1478]" captionTargetBox-4="[117,1455,147,1188]" captionTargetBox-5="[132,1470,148,1156]" captionTargetId-0="figure-50@3.[114,1458,144,1795]" captionTargetId-1="figure-46@4.[158,1444,144,1812]" captionTargetId-2="figure-264@5.[114,1458,144,1383]" captionTargetId-3="figure-243@6.[129,1473,144,1481]" captionTargetId-4="figure-421@7.[114,1458,144,1191]" captionTargetId-5="figure-403@8.[129,1473,144,1159]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." captionText-1="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." captionText-2="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." captionText-3="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." captionText-4="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." captionText-5="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="2" pageNumber="3">Figs 2–7</figureCitation>
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<figureCitation id="13162A23FFBFFF8FFD57FAEFF2AF2C5E" box="[651,737,1297,1321]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStartId-0="3.[113,178,1823,1847]" captionStartId-1="4.[129,194,1840,1864]" captionStartId-2="5.[113,178,1411,1435]" captionStartId-3="6.[128,193,1509,1533]" captionStartId-4="7.[115,180,1219,1243]" captionStartId-5="8.[129,194,1187,1211]" captionTargetBox-0="[117,1455,147,1791]" captionTargetBox-1="[161,1441,148,1807]" captionTargetBox-2="[117,1454,147,1380]" captionTargetBox-3="[132,1469,149,1478]" captionTargetBox-4="[117,1455,147,1188]" captionTargetBox-5="[132,1470,148,1156]" captionTargetId-0="figure-50@3.[114,1458,144,1795]" captionTargetId-1="figure-46@4.[158,1444,144,1812]" captionTargetId-2="figure-264@5.[114,1458,144,1383]" captionTargetId-3="figure-243@6.[129,1473,144,1481]" captionTargetId-4="figure-421@7.[114,1458,144,1191]" captionTargetId-5="figure-403@8.[129,1473,144,1159]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." captionText-1="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." captionText-2="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." captionText-3="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." captionText-4="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." captionText-5="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14893332" figureDoi-1="http://doi.org/10.5281/zenodo.14893334" figureDoi-2="http://doi.org/10.5281/zenodo.14893336" figureDoi-3="http://doi.org/10.5281/zenodo.14893338" figureDoi-4="http://doi.org/10.5281/zenodo.14893342" figureDoi-5="http://doi.org/10.5281/zenodo.14893344" httpUri-0="https://zenodo.org/record/14893332/files/figure.png" httpUri-1="https://zenodo.org/record/14893334/files/figure.png" httpUri-2="https://zenodo.org/record/14893336/files/figure.png" httpUri-3="https://zenodo.org/record/14893338/files/figure.png" httpUri-4="https://zenodo.org/record/14893342/files/figure.png" httpUri-5="https://zenodo.org/record/14893344/files/figure.png" pageId="2" pageNumber="3">Figs 2–7</figureCitation>
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).
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</paragraph>
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@ -79,28 +82,28 @@
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, an isolated predentary with the anteriormost portion of the right dentary (
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<figureCitation id="13162A23FFBFFF8FFD65FA97F2B92CF7" box="[697,759,1385,1409]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." pageId="2" pageNumber="3">Fig. 8</figureCitation>
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<figureCitation id="13162A23FFBFFF8FFD65FA97F2B92CF7" box="[697,759,1385,1409]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." figureDoi="http://doi.org/10.5281/zenodo.14893346" httpUri="https://zenodo.org/record/14893346/files/figure.png" pageId="2" pageNumber="3">Fig. 8</figureCitation>
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)
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;
|
||||
<materialsCitation id="3B453CFBFFBFFF8FFF5DFA77F1D02CD7" box="[129,414,1417,1441]" collectionCode="MTM" pageId="2" pageNumber="3" specimenCode="MTM V2009.193.1" specimenCount="1">
|
||||
<specimenCode id="DB8B9EDDFFBFFF8FFF5DFA77F1042CD7" box="[129,330,1417,1441]" pageId="2" pageNumber="3">MTM V2009.193.1</specimenCode>
|
||||
(
|
||||
<figureCitation id="13162A23FFBFFF8FFE86FA77F1DD2CD7" box="[346,403,1417,1441]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." pageId="2" pageNumber="3">Fig. 9</figureCitation>
|
||||
<figureCitation id="13162A23FFBFFF8FFE86FA77F1DD2CD7" box="[346,403,1417,1441]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." figureDoi="http://doi.org/10.5281/zenodo.14893348" httpUri="https://zenodo.org/record/14893348/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B453CFBFFBFFF8FFE75FA77F34A2CD7" box="[425,772,1417,1441]" collectionCode="MTM" pageId="2" pageNumber="3" specimenCode="MTM V2009.194.1" specimenCount="1">
|
||||
<specimenCode id="DB8B9EDDFFBFFF8FFE75FA77F23C2CD7" box="[425,626,1417,1441]" pageId="2" pageNumber="3">MTM V2009.194.1</specimenCode>
|
||||
(
|
||||
<figureCitation id="13162A23FFBFFF8FFD5FFA77F2B62CD7" box="[643,760,1417,1441]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." pageId="2" pageNumber="3">Fig. 10A–D</figureCitation>
|
||||
<figureCitation id="13162A23FFBFFF8FFD5FFA77F2B62CD7" box="[643,760,1417,1441]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." figureDoi="http://doi.org/10.5281/zenodo.14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="2" pageNumber="3">Fig. 10A–D</figureCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
, and
|
||||
<materialsCitation id="3B453CFBFFBFFF8FFF77FA56F08E2CA9" collectionCode="MTM" pageId="2" pageNumber="3" specimenCode="MTM V2009.196.1" specimenCount="1">
|
||||
<specimenCode id="DB8B9EDDFFBFFF8FFF77FA56F13D2CB6" box="[171,371,1448,1472]" pageId="2" pageNumber="3">MTM V2009.196.1</specimenCode>
|
||||
(
|
||||
<figureCitation id="13162A23FFBFFF8FFE5FFA56F1B92CB6" box="[387,503,1448,1472]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." pageId="2" pageNumber="3">Fig. 10E–H</figureCitation>
|
||||
<figureCitation id="13162A23FFBFFF8FFE5FFA56F1B92CB6" box="[387,503,1448,1472]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." figureDoi="http://doi.org/10.5281/zenodo.14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="2" pageNumber="3">Fig. 10E–H</figureCitation>
|
||||
), three isolated predentary bones
|
||||
</materialsCitation>
|
||||
.
|
||||
|
|
@ -112,7 +115,7 @@
|
|||
Iharkút, Bakony Mountains, western
|
||||
<collectingCountry id="F33A7636FFBFFF8FFF5DF9DEF0922F4E" box="[129,220,1568,1592]" name="Hungary" pageId="2" pageNumber="3">Hungary</collectingCountry>
|
||||
(
|
||||
<figureCitation id="13162A23FFBFFF8FFF2EF9DEF17F2F4E" box="[242,305,1568,1592]" captionStart="Figure 1" captionStartId="1.[114,179,656,680]" captionTargetBox="[118,758,144,628]" captionTargetId="figure-719@1.[118,758,144,628]" captionTargetPageId="1" captionText="Figure 1. European localities with purported records of ceratopsian dinosaurs: (1) Ajkaceratops kozmai from the Upper Cretaceous of Iharkút, Bakony Mountains, Hungary; (2) ceratopsian-like footprint from the Upper Cretaceous of the Młynarka Mount, Roztocze Hills, Poland; (3) Leptoceratopsidae indet. from the Upper Cretaceous of Åsen, Kristianstad Basin, Sweden; (4) Leptoceratopsidae indet. from the Upper Cretaceous of Ullstorp, Kristianstad Basin, Sweden; (5) Stenopelix valdensis from the Lower Cretaceous of Bückeburg, Lower Saxony, Germany; (6) ‘Craspedodon lonzeensis’ from the Upper Cretaceous of Lonzée, Namur Province, Belgium. The Iharkút site, from where the material redescribed here was collected, is marked by a star." pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||||
<figureCitation id="13162A23FFBFFF8FFF2EF9DEF17F2F4E" box="[242,305,1568,1592]" captionStart="Figure 1" captionStartId="1.[114,179,656,680]" captionTargetBox="[118,758,144,628]" captionTargetId="figure-719@1.[118,758,144,628]" captionTargetPageId="1" captionText="Figure 1. European localities with purported records of ceratopsian dinosaurs: (1) Ajkaceratops kozmai from the Upper Cretaceous of Iharkút, Bakony Mountains, Hungary; (2) ceratopsian-like footprint from the Upper Cretaceous of the Młynarka Mount, Roztocze Hills, Poland; (3) Leptoceratopsidae indet. from the Upper Cretaceous of Åsen, Kristianstad Basin, Sweden; (4) Leptoceratopsidae indet. from the Upper Cretaceous of Ullstorp, Kristianstad Basin, Sweden; (5) Stenopelix valdensis from the Lower Cretaceous of Bückeburg, Lower Saxony, Germany; (6) ‘Craspedodon lonzeensis’ from the Upper Cretaceous of Lonzée, Namur Province, Belgium. The Iharkút site, from where the material redescribed here was collected, is marked by a star." figureDoi="http://doi.org/10.5281/zenodo.14893330" httpUri="https://zenodo.org/record/14893330/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||||
); Csehbánya Formation (Santonian, Upper Cretaceous; see:
|
||||
<bibRefCitation id="EFBC4B57FFBFFF8FFEF1F9C1F1EE2F21" author="Szalai E" box="[301,416,1599,1623]" pageId="2" pageNumber="3" refId="ref17979" refString="Szalai E. Paleomagneses vizsgalatok Iharkuton. Unpublished manuscript, Eotvos Lorand University, Department of Environmental Geology, 2005." type="book" year="2005">Szalai 2005</bibRefCitation>
|
||||
,
|
||||
|
|
@ -249,7 +252,7 @@ spp.
|
|||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFBFFF8FFC88F90AF5542EDE" blockId="2.[852,1475,1654,1960]" pageId="2" pageNumber="3">(ii) **Rostral and predentary bones with flat occlusal surfaces anteriorly, deprived of the cutting lateral edges in its anterior portion, resulting in a subtriangular transverse cross-section of the bones (in contrast to all known ceratopsians, where the rostral bone is excavated anteriorly creating space for the predentary bone).</paragraph>
|
||||
<caption id="DF52662EFFBEFF8EFFADF8E1F3F92E41" box="[113,951,1823,1847]" pageId="3" pageNumber="4" startId="3.[113,178,1823,1847]" targetBox="[117,1455,147,1791]" targetPageId="3" targetType="figure">
|
||||
<caption id="DF52662EFFBEFF8EFFADF8E1F3F92E41" ID-DOI="http://doi.org/10.5281/zenodo.14893332" ID-Zenodo-Dep="14893332" box="[113,951,1823,1847]" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="3" pageNumber="4" startId="3.[113,178,1823,1847]" targetBox="[117,1455,147,1791]" targetPageId="3" targetType="figure">
|
||||
<paragraph id="8B9236A6FFBEFF8EFFADF8E1F3F92E41" blockId="3.[113,951,1823,1847]" box="[113,951,1823,1847]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B959EAB4FFBEFF8EFFADF8E1F0872E41" bold="true" box="[113,201,1823,1847]" pageId="3" pageNumber="4">Figure 2.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFBEFF8EFF12F8E1F1AE2E41" authority="MTM V" authorityName="MTM V" box="[206,480,1823,1847]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -260,7 +263,7 @@ MTM V
|
|||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B9236A6FFBEFF8EFF50F8AAF3AA2EDD" blockId="3.[140,762,1876,1963]" lastBlockId="3.[885,1458,1876,1963]" pageId="3" pageNumber="4">(iii) **Rostral and predentary bones ornamented with pits, which are absent in their anteroventral tips (in all ceratopsians the anteroventral tip is usually the most rugose part of the rostral bone, and the rugosity is mostly manifested through channels and ridges rather than pits).</paragraph>
|
||||
<caption id="DF52662EFFB9FF89FF5DF8CEF39B2E3E" box="[129,981,1840,1864]" pageId="4" pageNumber="5" startId="4.[129,194,1840,1864]" targetBox="[161,1441,148,1807]" targetPageId="4" targetType="figure">
|
||||
<caption id="DF52662EFFB9FF89FF5DF8CEF39B2E3E" ID-DOI="http://doi.org/10.5281/zenodo.14893334" ID-Zenodo-Dep="14893334" box="[129,981,1840,1864]" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="4" pageNumber="5" startId="4.[129,194,1840,1864]" targetBox="[161,1441,148,1807]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB9FF89FF5DF8CEF39B2E3E" blockId="4.[129,981,1840,1864]" box="[129,981,1840,1864]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B959EAB4FFB9FF89FF5DF8CEF0962E3E" bold="true" box="[129,216,1840,1864]" pageId="4" pageNumber="5">Figure 3.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB9FF89FF01F8CEF1A12E3E" authority="MTM V" authorityName="MTM V" box="[221,495,1840,1864]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -271,7 +274,7 @@ MTM V
|
|||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B9236A6FFB9FF89FF47F89DF3802ECF" blockId="4.[155,778,1891,1977]" lastBlockId="4.[900,1474,1891,1977]" pageId="4" pageNumber="5">(iv) **Rostral extensively covering the premaxilla in the palatal direction, reaching nearly as posteriorly as the buccal process (in all known ceratopsians the buccal process terminates significantly more posteriorly than the palatal extension of the rostral bone).</paragraph>
|
||||
<caption id="DF52662EFFB8FF88FFADFA7DF1352CC1" pageId="5" pageNumber="6" startId="5.[113,178,1411,1435]" targetBox="[117,1454,147,1380]" targetPageId="5" targetType="figure">
|
||||
<caption id="DF52662EFFB8FF88FFADFA7DF1352CC1" ID-DOI="http://doi.org/10.5281/zenodo.14893336" ID-Zenodo-Dep="14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1411,1435]" targetBox="[117,1454,147,1380]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB8FF88FFADFA7DF1352CC1" blockId="5.[113,1381,1411,1463]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B959EAB4FFB8FF88FFADFA7DF0872CED" bold="true" box="[113,201,1411,1435]" pageId="5" pageNumber="6">Figure 4.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB8FF88FF12FA7DF1AE2CED" authority="MTM V" authorityName="MTM V" box="[206,480,1411,1435]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -336,7 +339,7 @@ and other ceratopsians, the nasal seems to always contribute to the margin of th
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF52662EFFBBFF8BFF5CFA1BF3ED2C8B" box="[128,931,1509,1534]" pageId="6" pageNumber="7" startId="6.[128,193,1509,1533]" targetBox="[132,1469,149,1478]" targetPageId="6" targetType="figure">
|
||||
<caption id="DF52662EFFBBFF8BFF5CFA1BF3ED2C8B" ID-DOI="http://doi.org/10.5281/zenodo.14893338" ID-Zenodo-Dep="14893338" box="[128,931,1509,1534]" httpUri="https://zenodo.org/record/14893338/files/figure.png" pageId="6" pageNumber="7" startId="6.[128,193,1509,1533]" targetBox="[132,1469,149,1478]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8B9236A6FFBBFF8BFF5CFA1BF3ED2C8B" blockId="6.[128,931,1509,1534]" box="[128,931,1509,1534]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B959EAB4FFBBFF8BFF5CFA1BF0962C88" bold="true" box="[128,216,1509,1534]" pageId="6" pageNumber="7">Figure 5.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFBBFF8BFF01FA1BF1A12C88" authority="MTM V" authorityName="MTM V" box="[221,495,1509,1534]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -360,13 +363,13 @@ specimen of
|
|||
(MTM
|
||||
<date id="FF931066FFBBFF8BFD60F8B9F3482E29" box="[700,774,1863,1887]" pageId="6" pageNumber="7" value="2009-05">V2009</date>
|
||||
. 192.1) includes both premaxillae and the anteriormost portion of the anteromedial processes of both maxillae. The rugose anterior region of the snout has been originally interpreted as a rostral bone that would be fully and indistinguishably fused to the premaxillae (
|
||||
<figureCitation id="13162A23FFBBFF8BFB26F9B3F50E2F13" box="[1274,1344,1613,1637]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="6" pageNumber="7">Figs 2</figureCitation>
|
||||
<figureCitation id="13162A23FFBBFF8BFB26F9B3F50E2F13" box="[1274,1344,1613,1637]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFBBFF8BFA88F9B3F52F2F13" box="[1364,1377,1613,1637]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="6" pageNumber="7">3</figureCitation>
|
||||
<figureCitation id="13162A23FFBBFF8BFA88F9B3F52F2F13" box="[1364,1377,1613,1637]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFBBFF8BFAA9F9B3F5CC2F12" box="[1397,1410,1613,1636]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="6" pageNumber="7">7</figureCitation>
|
||||
<figureCitation id="13162A23FFBBFF8BFAA9F9B3F5CC2F12" box="[1397,1410,1613,1636]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="6" pageNumber="7">7</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFBBFF8BFA4AF9B3F5FF2F13" box="[1430,1457,1613,1637]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="6" pageNumber="7">11</figureCitation>
|
||||
<figureCitation id="13162A23FFBBFF8BFA4AF9B3F5FF2F13" box="[1430,1457,1613,1637]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="6" pageNumber="7">11</figureCitation>
|
||||
). The maximum length of the specimen, as measured from the anteroventral tip of the beak to the posteriormost tip of the right posterolateral process, equals
|
||||
<quantity id="4CD59B43FFBBFF8BFACDF955F5112FB4" box="[1297,1375,1707,1731]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="6" pageNumber="7" unit="mm" value="87.0">87 mm</quantity>
|
||||
. In turn, the maximum dorsoventral height, as measured from the anteroventral tip of the beak to the dorsal surface of the posterodorsal process, equals
|
||||
|
|
@ -375,12 +378,12 @@ specimen of
|
|||
</paragraph>
|
||||
<paragraph id="8B9236A6FFBBFF8AFC89F8D6F22F2C34" blockId="6.[825,1474,1581,1981]" lastBlockId="7.[113,762,1291,1346]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||||
The specimen shows a partial taphonomic deformation, with the posterior half of the left premaxilla being broken and slightly displaced posteroventrally and a bit medially (
|
||||
<figureCitation id="13162A23FFBBFF8BFAAAF899F5FF2E09" box="[1398,1457,1895,1919]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="6" pageNumber="7">Fig. 2</figureCitation>
|
||||
<figureCitation id="13162A23FFBBFF8BFAAAF899F5FF2E09" box="[1398,1457,1895,1919]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
|
||||
), and the posteroventral part of the right premaxilla being displaced posteromedially. The right side of the specimen is slightly ‘elevated’ (
|
||||
<figureCitation id="13162A23FFBAFF8AFF3CFAF5F1552C55" box="[224,283,1291,1315]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||||
<figureCitation id="13162A23FFBAFF8AFF3CFAF5F1552C55" box="[224,283,1291,1315]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
|
||||
), resulting in the posterolateral process of the right premaxilla being displaced posterodorsally.
|
||||
</paragraph>
|
||||
<caption id="DF52662EFFBAFF8AFFAFFB3DF3FF2DAD" box="[115,945,1219,1243]" pageId="7" pageNumber="8" startId="7.[115,180,1219,1243]" targetBox="[117,1455,147,1188]" targetPageId="7" targetType="figure">
|
||||
<caption id="DF52662EFFBAFF8AFFAFFB3DF3FF2DAD" ID-DOI="http://doi.org/10.5281/zenodo.14893342" ID-Zenodo-Dep="14893342" box="[115,945,1219,1243]" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="7" pageNumber="8" startId="7.[115,180,1219,1243]" targetBox="[117,1455,147,1188]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8B9236A6FFBAFF8AFFAFFB3DF3FF2DAD" blockId="7.[115,945,1219,1243]" box="[115,945,1219,1243]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B959EAB4FFBAFF8AFFAFFB3DF0852DAD" bold="true" box="[115,203,1219,1243]" pageId="7" pageNumber="8">Figure 6.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFBAFF8AFF12FB3DF1AE2DAD" authority="MTM V" authorityName="MTM V" box="[206,480,1219,1243]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -509,13 +512,13 @@ described this element as being relatively short and curved ventrally, terminati
|
|||
<emphasis id="B959EAB4FFBAFF8AFB74F93CF53A2FAC" box="[1192,1396,1730,1754]" italics="true" pageId="7" pageNumber="8">Ajkaceratops kozmai</emphasis>
|
||||
</taxonomicName>
|
||||
snout has a relatively flat, smooth, ventral surface, lacking the developed lateral edges (
|
||||
<figureCitation id="13162A23FFBAFF8AFC27F8FEF4752E6E" box="[1019,1083,1792,1816]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="7" pageNumber="8">Figs 4</figureCitation>
|
||||
<figureCitation id="13162A23FFBAFF8AFC27F8FEF4752E6E" box="[1019,1083,1792,1816]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFBAFF8AFB96F8FEF4262E6E" box="[1098,1128,1792,1816]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="7" pageNumber="8">7C</figureCitation>
|
||||
<figureCitation id="13162A23FFBAFF8AFB96F8FEF4262E6E" box="[1098,1128,1792,1816]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="7" pageNumber="8">7C</figureCitation>
|
||||
), resulting in the subtriangular cross-section of the structure (
|
||||
<figureCitation id="13162A23FFBAFF8AFBB1F8DEF48B2E4E" box="[1133,1221,1824,1848]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="7" pageNumber="8">Fig. 12A</figureCitation>
|
||||
<figureCitation id="13162A23FFBAFF8AFBB1F8DEF48B2E4E" box="[1133,1221,1824,1848]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="7" pageNumber="8">Fig. 12A</figureCitation>
|
||||
). As such, the appearance of the bone is more robust. In all ceratopsians the ventral surface of the rostral bone is vaulted for accommodating the predentary bone, and the lateral cutting edges of the rostral are evident in the anterior portion, forming the typically V-shaped cross-section of the bone (
|
||||
<figureCitation id="13162A23FFB5FF85FE4FFAC1F2482C21" box="[403,518,1343,1367]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="8" pageNumber="9">Fig. 12F–K</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FE4FFAC1F2482C21" box="[403,518,1343,1367]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="8" pageNumber="9">Fig. 12F–K</figureCitation>
|
||||
). Such a condition is observable even in the earliest-diverging ceratopsians [e.g.
|
||||
<taxonomicName id="4C2D4D25FFB5FF85FD1DFAA0F08B2CE3" class="Reptilia" genus="Yinlong" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="downsi">
|
||||
<emphasis id="B959EAB4FFB5FF85FD1DFAA0F08B2CE3" italics="true" pageId="8" pageNumber="9">Yinlong downsi</emphasis>
|
||||
|
|
@ -527,7 +530,7 @@ spp.
|
|||
</taxonomicName>
|
||||
].
|
||||
</paragraph>
|
||||
<caption id="DF52662EFFB5FF85FF5DFB5DF3F32C79" pageId="8" pageNumber="9" startId="8.[129,194,1187,1211]" targetBox="[132,1470,148,1156]" targetPageId="8" targetType="figure">
|
||||
<caption id="DF52662EFFB5FF85FF5DFB5DF3F32C79" ID-DOI="http://doi.org/10.5281/zenodo.14893344" ID-Zenodo-Dep="14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="8" pageNumber="9" startId="8.[129,194,1187,1211]" targetBox="[132,1470,148,1156]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB5FF85FF5DFB5DF3F32C79" blockId="8.[129,1467,1187,1295]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B959EAB4FFB5FF85FF5DFB5DF0962DCD" bold="true" box="[129,216,1187,1211]" pageId="8" pageNumber="9">Figure 7.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB5FF85FF01FB5DF1A12DCD" authority="MTM V" authorityName="MTM V" box="[221,495,1187,1211]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -539,11 +542,11 @@ MTM V
|
|||
</caption>
|
||||
<paragraph id="8B9236A6FFB5FF85FF40FA63F2732E7B" blockId="8.[128,779,1343,1806]" pageId="8" pageNumber="9">
|
||||
In ventral view, the proposed rostral covers the premaxillary palate extensively (
|
||||
<figureCitation id="13162A23FFB5FF85FEABFA42F1FC2CA2" box="[375,434,1468,1492]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FEABFA42F1FC2CA2" box="[375,434,1468,1492]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||||
). The pitted surface of the bone extends far posteriorly to the point where the anteromedial premaxillary wings are not visible (
|
||||
<figureCitation id="13162A23FFB5FF85FE32FA05F2622F65" box="[494,556,1531,1555]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="8" pageNumber="9">Figs 4</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FE32FA05F2622F65" box="[494,556,1531,1555]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB5FF85FDE4FA05F2192F65" box="[568,599,1531,1555]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="8" pageNumber="9">7C</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FDE4FA05F2192F65" box="[568,599,1531,1555]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="8" pageNumber="9">7C</figureCitation>
|
||||
). The part of the vaulted premaxillary palate is not preserved in its midpoint and it is not clear how far posteriorly the proposed rostral bone develops. In any case, such condition contrasts with that seen in all ceratopsians, where the premaxilla is clearly visible in the ventral view and anteriorly wedged between the rostral buccal processes, with the medial walls of the rostral bone steeply inclined dorsomedially. No rugose surface covering the palate area, as it is in
|
||||
<taxonomicName id="4C2D4D25FFB5FF85FF46F90BF1562E7B" box="[154,280,1781,1805]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB5FF85FF46F90BF1562E7B" box="[154,280,1781,1805]" italics="true" pageId="8" pageNumber="9">Ajkaceratops</emphasis>
|
||||
|
|
@ -561,11 +564,11 @@ is rugous and covered with pits, indicating the presence of a keratinous beak du
|
|||
of pits can be distinguished. The larger ones (approximately
|
||||
<quantity id="4CD59B43FFB5FF85FCEBFAC1F33A2C20" box="[823,884,1343,1367]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="3.0">3 mm</quantity>
|
||||
in diameter) are relatively rare and shallow, concentrated mainly in the anterior region, although the anteroventral tip is deprived of any larger pits (
|
||||
<figureCitation id="13162A23FFB5FF85FB8FFA83F4DE2CE3" box="[1107,1168,1405,1429]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="8" pageNumber="9">Figs 2</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FB8FFA83F4DE2CE3" box="[1107,1168,1405,1429]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB5FF85FB47FA83F4E62CE3" box="[1179,1192,1405,1429]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="8" pageNumber="9">3</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FB47FA83F4E62CE3" box="[1179,1192,1405,1429]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB5FF85FB68FA80F49D2CE3" box="[1204,1235,1406,1429]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="8" pageNumber="9">7D</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FB68FA80F49D2CE3" box="[1204,1235,1406,1429]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="8" pageNumber="9">7D</figureCitation>
|
||||
). The smaller pits (less than
|
||||
<quantity id="4CD59B43FFB5FF85FCACFA63F3F32CC2" box="[880,957,1437,1461]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
in diameter) are present around the whole snout and are not limited only to the rostral bone.
|
||||
|
|
@ -582,7 +585,7 @@ Han
|
|||
2016
|
||||
</bibRefCitation>
|
||||
;
|
||||
<figureCitation id="13162A23FFB5FF85FA40F9E4F32F2F27" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="8" pageNumber="9">Fig. 11B</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FA40F9E4F32F2F27" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="8" pageNumber="9">Fig. 11B</figureCitation>
|
||||
). In some specimens of
|
||||
<taxonomicName id="4C2D4D25FFB5FF85FB83F9C4F5542F27" box="[1119,1306,1594,1617]" class="Reptilia" family="Psittacosauridae" genus="Psittacosaurus" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B959EAB4FFB5FF85FB83F9C4F4A42F27" box="[1119,1258,1594,1617]" italics="true" pageId="8" pageNumber="9">Psittacosaurus</emphasis>
|
||||
|
|
@ -602,7 +605,7 @@ specimens do not exhibit any longitudinal grooves (e.g.
|
|||
sp.
|
||||
</taxonomicName>
|
||||
MPC-D 100/610; see
|
||||
<figureCitation id="13162A23FFB5FF85FCE5F928F3C22F98" box="[825,908,1750,1774]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="8" pageNumber="9">Fig.11C</figureCitation>
|
||||
<figureCitation id="13162A23FFB5FF85FCE5F928F3C22F98" box="[825,908,1750,1774]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="8" pageNumber="9">Fig.11C</figureCitation>
|
||||
;
|
||||
<taxonomicName id="4C2D4D25FFB5FF85FC47F929F4E92F98" box="[923,1191,1750,1774]" class="Reptilia" family="Psittacosauridae" genus="Psittacosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="lujiatunensis">
|
||||
<emphasis id="B959EAB4FFB5FF85FC47F929F4E92F98" box="[923,1191,1750,1774]" italics="true" pageId="8" pageNumber="9">Psittacosaurus lujiatunensis</emphasis>
|
||||
|
|
@ -636,7 +639,7 @@ the pits are mostly distributed adjacent to the anteroventral tip, in contrast t
|
|||
<emphasis id="B959EAB4FFB4FF84FEFFFABAF2492C2D" box="[291,519,1347,1371]" italics="true" pageId="9" pageNumber="10">Protoceratops andrewsi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="13162A23FFB4FF84FDC0FABDF23D2C2D" box="[540,627,1347,1371]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="9" pageNumber="10">Fig. 11E</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FDC0FABDF23D2C2D" box="[540,627,1347,1371]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="9" pageNumber="10">Fig. 11E</figureCitation>
|
||||
), the rostral bone is mostly covered in longitudinal grooves, with pits being comparatively larger than those in
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FE2CFA7CF2202CEC" box="[496,622,1410,1434]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB4FF84FE2CFA7CF2202CEC" box="[496,622,1410,1434]" italics="true" pageId="9" pageNumber="10">Ajkaceratops</emphasis>
|
||||
|
|
@ -650,7 +653,7 @@ the pits are mostly distributed adjacent to the anteroventral tip, in contrast t
|
|||
<emphasis id="B959EAB4FFB4FF84FFADFA1EF13E2C8E" box="[113,368,1504,1528]" italics="true" pageId="9" pageNumber="10">Udanoceratops tschizhovi</emphasis>
|
||||
</taxonomicName>
|
||||
(PIN 3907/11), the pits are more densely distributed, and most of them fuse together dorsally, forming the longitudinal channels (
|
||||
<figureCitation id="13162A23FFB4FF84FE00F9E0F27B2F40" box="[476,565,1566,1590]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="9" pageNumber="10">Fig. 11D</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FE00F9E0F27B2F40" box="[476,565,1566,1590]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="9" pageNumber="10">Fig. 11D</figureCitation>
|
||||
). There are no longitudinal channels or ridges in
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FE64F9C0F2782F20" box="[440,566,1598,1622]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB4FF84FE64F9C0F2782F20" box="[440,566,1598,1622]" italics="true" pageId="9" pageNumber="10">Ajkaceratops</emphasis>
|
||||
|
|
@ -660,13 +663,13 @@ the pits are mostly distributed adjacent to the anteroventral tip, in contrast t
|
|||
<emphasis id="B959EAB4FFB4FF84FE5AF9A3F21D2F03" box="[390,595,1629,1653]" italics="true" pageId="9" pageNumber="10">Ajkaceratops kozmai</emphasis>
|
||||
</taxonomicName>
|
||||
are shallow and present along the posterior contact between the rugose region and the main body of the premaxilla (
|
||||
<figureCitation id="13162A23FFB4FF84FE27F962F27A2FC2" box="[507,564,1692,1716]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="9" pageNumber="10">Fig. 2</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FE27F962F27A2FC2" box="[507,564,1692,1716]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
|
||||
). If that element is indeed a rostral bone, these structures may derive from the fusion between the rostral and the premaxillae. In centrosaurine ceratopsids, such as
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FE8DF904F26B2E67" baseAuthorityName="Lambe" baseAuthorityYear="1905" box="[337,549,1786,1809]" class="Reptilia" family="Ceratopsidae" genus="Centrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="apertus">
|
||||
<emphasis id="B959EAB4FFB4FF84FE8DF904F26B2E67" box="[337,549,1786,1809]" italics="true" pageId="9" pageNumber="10">Centrosaurus apertus</emphasis>
|
||||
</taxonomicName>
|
||||
(USNM 8897; see
|
||||
<figureCitation id="13162A23FFB4FF84FFADF8E7F0862E47" box="[113,200,1817,1841]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." pageId="9" pageNumber="10">Fig. 11F</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FFADF8E7F0862E47" box="[113,200,1817,1841]" captionStart="Figure 11" captionStartId="12.[129,194,587,611]" captionTargetBox="[132,1470,147,556]" captionTargetId="figure-155@12.[129,1473,144,559]" captionTargetPageId="12" captionText="Figure 11. Simplified comparison of the ornamentation of the anterior portion of the snout of Ajkaceratops kozmai (A), and the rostral bone of the ceratopsian dinosaurs (B–F). B, Yinlong downsi (based on IVPP V18637); C, Psittacosaurus sp.(based on MPC-D 100/610); D, Udanoceratops tschizhovi (based on PIN 3907/11); E, Protoceratops andrewsi (based on AMNH FARB 6637); F, Centrosaurus apertus (based on USNM 8897). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="9" pageNumber="10">Fig. 11F</figureCitation>
|
||||
),
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FF3EF8E7F1AB2E47" box="[226,485,1817,1841]" class="Reptilia" family="Ceratopsidae" genus="Pachyrhinosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="lakustai">
|
||||
<emphasis id="B959EAB4FFB4FF84FF3EF8E7F1AB2E47" box="[226,485,1817,1841]" italics="true" pageId="9" pageNumber="10">Pachyrhinosaurus lakustai</emphasis>
|
||||
|
|
@ -697,7 +700,7 @@ spp.
|
|||
</taxonomicName>
|
||||
there are also longitudinal channels, mostly in the posterior and the dorsalmost region of the rostral bone (e.g. LACM 45807, USNM 1201, USNM 8081).
|
||||
</paragraph>
|
||||
<caption id="DF52662EFFB4FF84FFAEFB21F2222C65" pageId="9" pageNumber="10" startId="9.[114,179,1247,1271]" targetBox="[117,1455,150,1216]" targetPageId="9" targetType="figure">
|
||||
<caption id="DF52662EFFB4FF84FFAEFB21F2222C65" ID-DOI="http://doi.org/10.5281/zenodo.14893346" ID-Zenodo-Dep="14893346" httpUri="https://zenodo.org/record/14893346/files/figure.png" pageId="9" pageNumber="10" startId="9.[114,179,1247,1271]" targetBox="[117,1455,150,1216]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB4FF84FFAEFB21F2222C65" blockId="9.[113,1379,1247,1299]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B959EAB4FFB4FF84FFAEFB21F0872D81" bold="true" box="[114,201,1247,1271]" pageId="9" pageNumber="10">Figure 8.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FF12FB21F1AE2D81" authority="MTM V" authorityName="MTM V" box="[206,480,1247,1271]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -713,9 +716,9 @@ Additionally, in
|
|||
<emphasis id="B959EAB4FFB4FF84FC31F9C0F4252F20" box="[1005,1131,1598,1622]" italics="true" pageId="9" pageNumber="10">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
the large pits cover most of the anterior portion of the snout, also in the ventral view (
|
||||
<figureCitation id="13162A23FFB4FF84FAB2F9A3F5E32F03" box="[1390,1453,1629,1653]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="9" pageNumber="10">Figs 4</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FAB2F9A3F5E32F03" box="[1390,1453,1629,1653]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="9" pageNumber="10">Figs 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB4FF84FCF6F982F3072FE2" box="[810,841,1660,1684]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="9" pageNumber="10">7C</figureCitation>
|
||||
<figureCitation id="13162A23FFB4FF84FCF6F982F3072FE2" box="[810,841,1660,1684]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="9" pageNumber="10">7C</figureCitation>
|
||||
). In ceratopsian dinosaurs, however, the grooves and pits are limited solely to the anteriormost portions of the ventral aspects of the rostrals, and their density is significantly lower than on the lateral sides [see, e.g.
|
||||
<taxonomicName id="4C2D4D25FFB4FF84FBF3F925F4952F84" box="[1071,1243,1755,1778]" class="Reptilia" family="Psittacosauridae" genus="Psittacosaurus" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B959EAB4FFB4FF84FBF3F925F4F42F84" box="[1071,1210,1755,1778]" italics="true" pageId="9" pageNumber="10">Psittacosaurus</emphasis>
|
||||
|
|
@ -738,15 +741,15 @@ sp.
|
|||
<paragraph id="8B9236A6FFB4FF87FCF6F8A6F08B2DB3" blockId="9.[810,1459,1880,1967]" lastBlockId="10.[128,779,914,1659]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||||
<emphasis id="B959EAB4FFB4FF84FCF6F8A6F3A42E06" box="[810,1002,1880,1904]" italics="true" pageId="9" pageNumber="10">The dorsal process:</emphasis>
|
||||
The proposed rostral bone apparently develops posterodorsally along the anterior surface of the premaxillary internarial bar, as indicated by the rugose texture. The dorsal process of the rostral bone would, therefore, be very long, reaching to at least the midpoint of the external nares (
|
||||
<figureCitation id="13162A23FFB7FF87FD19FC4CF34A2ABC" box="[709,772,946,970]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="10" pageNumber="11">Figs 2</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FD19FC4CF34A2ABC" box="[709,772,946,970]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="10" pageNumber="11">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB7FF87FF5DFC2FF0C02A9F" box="[129,142,977,1001]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="10" pageNumber="11">3</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FF5DFC2FF0C02A9F" box="[129,142,977,1001]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="10" pageNumber="11">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB7FF87FF45FC2FF0E82A9F" box="[153,166,977,1001]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." pageId="10" pageNumber="11">5</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FF45FC2FF0E82A9F" box="[153,166,977,1001]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14893338" httpUri="https://zenodo.org/record/14893338/files/figure.png" pageId="10" pageNumber="11">5</figureCitation>
|
||||
). It is not certain whether the proposed rostral bone would contact the nasals; however, the texture on the internarial bar indicates that it terminated just anteriorly to the purported suture with the nasals (
|
||||
<figureCitation id="13162A23FFB7FF87FE89FBD1F1DF2D31" box="[341,401,1071,1095]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." pageId="10" pageNumber="11">Figs 5</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FE89FBD1F1DF2D31" box="[341,401,1071,1095]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14893338" httpUri="https://zenodo.org/record/14893338/files/figure.png" pageId="10" pageNumber="11">Figs 5</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB7FF87FE40FBCEF1F62D31" box="[412,440,1072,1095]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="10" pageNumber="11">7B</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FE40FBCEF1F62D31" box="[412,440,1072,1095]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="10" pageNumber="11">7B</figureCitation>
|
||||
). Among ceratopsians the rostral bone typically does not contact with the nasals, with the exception of
|
||||
<taxonomicName id="4C2D4D25FFB7FF87FF11FB90F1C92DF3" box="[205,391,1134,1157]" class="Reptilia" family="Psittacosauridae" genus="Psittacosaurus" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B959EAB4FFB7FF87FF11FB90F1162DF3" box="[205,344,1134,1157]" italics="true" pageId="10" pageNumber="11">Psittacosaurus</emphasis>
|
||||
|
|
@ -757,7 +760,7 @@ spp.
|
|||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF52662EFFB7FF87FF5DFCD0F1382A14" pageId="10" pageNumber="11" startId="10.[129,194,814,838]" targetBox="[132,1470,147,784]" targetPageId="10" targetType="figure">
|
||||
<caption id="DF52662EFFB7FF87FF5DFCD0F1382A14" ID-DOI="http://doi.org/10.5281/zenodo.14893348" ID-Zenodo-Dep="14893348" httpUri="https://zenodo.org/record/14893348/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,814,838]" targetBox="[132,1470,147,784]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB7FF87FF5DFCD0F1382A14" blockId="10.[129,1444,814,867]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B959EAB4FFB7FF87FF5DFCD0F0962A30" bold="true" box="[129,216,814,838]" pageId="10" pageNumber="11">Figure 9.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB7FF87FF01FCD0F1A12A31" authority="MTM V" authorityName="MTM V" box="[221,495,814,839]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -798,7 +801,7 @@ Wolfe
|
|||
2010
|
||||
</bibRefCitation>
|
||||
;
|
||||
<figureCitation id="13162A23FFB7FF87FF5DF9DAF0812F4A" box="[129,207,1572,1596]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="10" pageNumber="11">Fig. 12J</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FF5DF9DAF0812F4A" box="[129,207,1572,1596]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="10" pageNumber="11">Fig. 12J</figureCitation>
|
||||
), and the chasmosaurine ceratopsids, where the rostral bone is slender, elongated, and deeply excavated posteriorly in lateral view (
|
||||
<bibRefCitation id="EFBC4B57FFB7FF87FEDFF99DF18C2F0D" author="Dodson P & Forster CA & Sampson SD" box="[259,450,1635,1659]" pageId="10" pageNumber="11" pagination="494 - 513" refId="ref15509" refString="Dodson P, Forster CA, Sampson SD et al. Ceratopsidae. In: Weishampel DB, Dodson P, Osmolska H (eds), The Dinosauria. Berkeley: University of California Press, 2004, 494 - 513." type="book chapter" year="2004">
|
||||
Dodson
|
||||
|
|
@ -858,9 +861,9 @@ specimen of
|
|||
<emphasis id="B959EAB4FFB7FF87FB2FFBB0F58C2D10" box="[1267,1474,1102,1126]" italics="true" pageId="10" pageNumber="11">Ajkaceratops kozmai</emphasis>
|
||||
</taxonomicName>
|
||||
(MTM V2009.192.1) are almost completely preserved. Only the posteriormost portions of the posterodorsal processes and the ventral surface of the premaxillary symphysis (just anterior to the anteromedial processes of maxillae) are not preserved (
|
||||
<figureCitation id="13162A23FFB7FF87FC98FB15F3CE2C75" box="[836,896,1259,1283]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="10" pageNumber="11">Figs 4</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FC98FB15F3CE2C75" box="[836,896,1259,1283]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="10" pageNumber="11">Figs 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB7FF87FC56FB15F3D92C75" box="[906,919,1259,1283]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="10" pageNumber="11">6</figureCitation>
|
||||
<figureCitation id="13162A23FFB7FF87FC56FB15F3D92C75" box="[906,919,1259,1283]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="10" pageNumber="11">6</figureCitation>
|
||||
). The length of the premaxilla (from the anterior tip of the subnarial fossa to the posterior tip of the posterolateral process) is
|
||||
<quantity id="4CD59B43FFB7FF87FC55FAD4F39A2C37" box="[905,980,1322,1346]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="10" pageNumber="11" unit="mm" value="67.0">67 mm</quantity>
|
||||
. The length along the ventral edge (between the anterior margin of the subnarial fossa and the posterior tip of the posteroventral process) is
|
||||
|
|
@ -958,7 +961,7 @@ Farke
|
|||
</taxonomicName>
|
||||
. It differs strikingly from the rather high premaxillae seen in protoceratopsids and leptoceratopsids.
|
||||
</paragraph>
|
||||
<caption id="DF52662EFFB6FF86FFADFBA4F1592DDD" pageId="11" pageNumber="12" startId="11.[113,178,1114,1138]" targetBox="[117,1454,147,1083]" targetPageId="11" targetType="figure">
|
||||
<caption id="DF52662EFFB6FF86FFADFBA4F1592DDD" ID-DOI="http://doi.org/10.5281/zenodo.14893350" ID-Zenodo-Dep="14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="11" pageNumber="12" startId="11.[113,178,1114,1138]" targetBox="[117,1454,147,1083]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB6FF86FFADFBA4F1592DDD" blockId="11.[113,1421,1114,1195]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B959EAB4FFB6FF86FFADFBA4F09B2D04" bold="true" box="[113,213,1114,1138]" pageId="11" pageNumber="12">Figure 10.</emphasis>
|
||||
<taxonomicName id="4C2D4D25FFB6FF86FF06FBA5F1A22D05" authority="MTM V" authorityName="MTM V" box="[218,492,1115,1139]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kozmai">
|
||||
|
|
@ -974,17 +977,17 @@ The visible parts of the premaxillae are not fused to each other in
|
|||
<emphasis id="B959EAB4FFB6FF86FF52FA07F1422F67" box="[142,268,1529,1553]" italics="true" pageId="11" pageNumber="12">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
. The posterodorsal processes preserved the suture, visible in the ventral view (
|
||||
<figureCitation id="13162A23FFB6FF86FE66F9E6F1BD2F46" box="[442,499,1560,1584]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||||
<figureCitation id="13162A23FFB6FF86FE66F9E6F1BD2F46" box="[442,499,1560,1584]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||||
), and on the anterior and dorsal margins of the external nares (
|
||||
<figureCitation id="13162A23FFB6FF86FE35F9C6F2052F39" box="[489,587,1592,1616]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="11" pageNumber="12">Fig. 7A, B</figureCitation>
|
||||
<figureCitation id="13162A23FFB6FF86FE35F9C6F2052F39" box="[489,587,1592,1616]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="11" pageNumber="12">Fig. 7A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB6FF86FFADF98FF29C2E33" blockId="11.[113,764,1649,1987]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B959EAB4FFB6FF86FFADF98FF1CC2FFF" box="[113,386,1649,1673]" italics="true" pageId="11" pageNumber="12">The posterodorsal processes:</emphasis>
|
||||
The posterodorsal processes of the premaxillae are broken just anteriorly to the contact with the nasals (
|
||||
<figureCitation id="13162A23FFB6FF86FF1EF94EF14C2FBE" box="[194,258,1712,1736]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." pageId="11" pageNumber="12">Figs 5</figureCitation>
|
||||
<figureCitation id="13162A23FFB6FF86FF1EF94EF14C2FBE" box="[194,258,1712,1736]" captionStart="Figure 5" captionStartId="6.[128,193,1509,1533]" captionTargetBox="[132,1469,149,1478]" captionTargetId="figure-243@6.[129,1473,144,1481]" captionTargetPageId="6" captionText="Figure 5. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14893338" httpUri="https://zenodo.org/record/14893338/files/figure.png" pageId="11" pageNumber="12">Figs 5</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB6FF86FECDF94EF1622FB1" box="[273,300,1712,1735]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="11" pageNumber="12">7B</figureCitation>
|
||||
<figureCitation id="13162A23FFB6FF86FECDF94EF1622FB1" box="[273,300,1712,1735]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="11" pageNumber="12">7B</figureCitation>
|
||||
). The rugose texture of the beak continues to be present on the anterior surface posterodorsally; however, in the posteriormost portion of the bar, the pits are absent and there are longitudinal striations present, suggesting that the contact with dorsally overlapping nasals was developing nearby.
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB6FF86FF51F8B2F3AB2F9B" blockId="11.[113,764,1649,1987]" lastBlockId="11.[810,1460,1216,1898]" pageId="11" pageNumber="12">
|
||||
|
|
@ -1098,7 +1101,7 @@ is placed more ventrally than that in non-ceratopsoid ceratopsians, as noted by
|
|||
</taxonomicName>
|
||||
(CMN 8887 and CMN 8880) the narial fossa is formed by a concavity on the posteroventral corner of the external nares.
|
||||
</paragraph>
|
||||
<caption id="DF52662EFFB1FF81FF5DFDB5F1C82BC1" pageId="12" pageNumber="13" startId="12.[129,194,587,611]" targetBox="[132,1470,147,556]" targetPageId="12" targetType="figure">
|
||||
<caption id="DF52662EFFB1FF81FF5DFDB5F1C82BC1" ID-DOI="http://doi.org/10.5281/zenodo.14893354" ID-Zenodo-Dep="14893354" httpUri="https://zenodo.org/record/14893354/files/figure.png" pageId="12" pageNumber="13" startId="12.[129,194,587,611]" targetBox="[132,1470,147,556]" targetPageId="12" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB1FF81FF5DFDB5F1C82BC1" blockId="12.[129,1474,587,696]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B959EAB4FFB1FF81FF5DFDB5F0AB2B15" bold="true" box="[129,229,587,611]" pageId="12" pageNumber="13">Figure 11.</emphasis>
|
||||
Simplified comparison of the ornamentation of the anterior portion of the snout of
|
||||
|
|
@ -1129,7 +1132,7 @@ sp.
|
|||
(based on USNM 8897). Not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF52662EFFB1FF81FF5DFA12F5322FDA" pageId="12" pageNumber="13" startId="12.[129,194,1516,1540]" targetBox="[132,1470,753,1485]" targetPageId="12" targetType="figure">
|
||||
<caption id="DF52662EFFB1FF81FF5DFA12F5322FDA" ID-DOI="http://doi.org/10.5281/zenodo.14893356" ID-Zenodo-Dep="14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="12" pageNumber="13" startId="12.[129,194,1516,1540]" targetBox="[132,1470,753,1485]" targetPageId="12" targetType="figure">
|
||||
<paragraph id="8B9236A6FFB1FF81FF5DFA12F5322FDA" blockId="12.[129,1442,1516,1708]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B959EAB4FFB1FF81FF5DFA12F0AB2F72" bold="true" box="[129,229,1516,1540]" pageId="12" pageNumber="13">Figure 12.</emphasis>
|
||||
Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A,
|
||||
|
|
@ -1208,17 +1211,17 @@ the fossa develops anteroventrally, reaching the highest concavity in the anteri
|
|||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FF51FE17F0BB2A0E" blockId="13.[111,764,144,1672]" pageId="13" pageNumber="14">
|
||||
Ventrally, both posterodorsal processes form a small, thin crest in the anteroventral corner of the narial fossa (
|
||||
<figureCitation id="13162A23FFB0FF80FDA6FDF6F2F02B56" box="[634,702,520,544]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="13" pageNumber="14">Fig.7A</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FDA6FDF6F2F02B56" box="[634,702,520,544]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="13" pageNumber="14">Fig.7A</figureCitation>
|
||||
). The posterodorsal processes are directly appressed to each other, but with a visible suture between the left and the right premaxilla. In early-diverging ceratopsians and
|
||||
<taxonomicName id="4C2D4D25FFB0FF80FE35FD99F2292B08" baseAuthorityName="MSM P" baseAuthorityYear="2225" box="[489,615,615,638]" genus="Zuniceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB0FF80FE35FD99F2292B08" box="[489,615,615,638]" italics="true" pageId="13" pageNumber="14">Zuniceratops</emphasis>
|
||||
</taxonomicName>
|
||||
, the two premaxillary processes meet but remain separate entities, regardless of the ontogenetic stage of the individual. The thin crest in the anteroventral corner of the narial fossa is somewhat similar to the premaxillary septum of ceratopsid dinosaurs (
|
||||
<figureCitation id="13162A23FFB0FF80FD47FD1DF2942B8D" box="[667,730,739,763]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="13" pageNumber="14">Figs 2</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FD47FD1DF2942B8D" box="[667,730,739,763]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="13" pageNumber="14">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB0FF80FD34FD1DF2BB2B8D" box="[744,757,739,763]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="13" pageNumber="14">3</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FD34FD1DF2BB2B8D" box="[744,757,739,763]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="13" pageNumber="14">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB0FF80FFADFCFDF0C12A6D" box="[113,143,771,795]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." pageId="13" pageNumber="14">7A</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FFADFCFDF0C12A6D" box="[113,143,771,795]" captionStart="Figure 7" captionStartId="8.[129,194,1187,1211]" captionTargetBox="[132,1470,148,1156]" captionTargetId="figure-403@8.[129,1473,144,1159]" captionTargetPageId="8" captionText="Figure 7. Ajkaceratops kozmai MTM V2009.192.1.A, B, close-up on the broken part of the internarial bar in left posterolateral (A) and posterior (B) views, with no visible osteological boundaries between the premaxilla and the purported rostral. C, ventral surface of the snout in the right lateroventral view.The pits cover a significant area posteriorly, with no traces of the margin between the purported rostral bone and the premaxilla. D, anterior tip of the purported rostral bone in the left anterolateral view." figureDoi="http://doi.org/10.5281/zenodo.14893344" httpUri="https://zenodo.org/record/14893344/files/figure.png" pageId="13" pageNumber="14">7A</figureCitation>
|
||||
). However, in ceratopsids the septum is much more developed and occupies the majority of the narial fossa (see:
|
||||
<bibRefCitation id="EFBC4B57FFB0FF80FD1FFCDCF0932A2F" author="Lund EK & Sampson SD & Loewen MA" pageId="13" pageNumber="14" refId="ref16665" refString="Lund EK, Sampson SD, Loewen MA. Nasutoceratops titusi (Ornithischia, Ceratopsidae), a basal centrosaurine ceratopsid from the Kaiparowits Formation, southern Utah. Journal of Vertebrate Paleontology 2016; 36: e 1054936. https: // doi. org / 10.1080 / 02724634.2015.1054 936" type="journal volume" year="2016">
|
||||
Lund
|
||||
|
|
@ -1233,14 +1236,14 @@ Lund
|
|||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FF51FC7EF20B2AA1" blockId="13.[111,764,144,1672]" pageId="13" pageNumber="14">
|
||||
In the dorsal view, just posterior to the ‘premaxillary septum’, the specimen is weathered, and it is not clear whether the posterolateral processes met medially (
|
||||
<figureCitation id="13162A23FFB0FF80FE21FC41F27B2AA1" box="[509,565,959,983]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FE21FC41F27B2AA1" box="[509,565,959,983]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FF51FC20F1292FFE" blockId="13.[111,764,144,1672]" pageId="13" pageNumber="14">
|
||||
There are at least four foramina visible on the left premaxilla,just behind the rugose part of the snout (
|
||||
<figureCitation id="13162A23FFB0FF80FE3AFC03F26F2D63" box="[486,545,1021,1045]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." pageId="13" pageNumber="14">Figs 2</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FE3AFC03F26F2D63" box="[486,545,1021,1045]" captionStart="Figure 2" captionStartId="3.[113,178,1823,1847]" captionTargetBox="[117,1455,147,1791]" captionTargetId="figure-50@3.[114,1458,144,1795]" captionTargetPageId="3" captionText="Figure 2. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893332" httpUri="https://zenodo.org/record/14893332/files/figure.png" pageId="13" pageNumber="14">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB0FF80FDF0FC03F2772D63" box="[556,569,1021,1045]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="13" pageNumber="14">3</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FDF0FC03F2772D63" box="[556,569,1021,1045]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="13" pageNumber="14">3</figureCitation>
|
||||
). The anterodorsal one is placed immediately posterior to the border of the rugose part, in the deepest part of the subnarial fossa.Posterior to it, there is a large foramen near the ‘premaxillary septum’ that is nearly twice the size of the anterodorsal foramen. The placement of these two foramina is similar to, respectively, the anterior premaxillary foramen and premaxillary foramen (
|
||||
<emphasis id="B959EAB4FFB0FF80FDC4FB44F2042DA7" box="[536,586,1210,1233]" italics="true" pageId="13" pageNumber="14">sensu</emphasis>
|
||||
<bibRefCitation id="EFBC4B57FFB0FF80FD8CFB47F2802DA7" author="Sereno PC" box="[592,718,1209,1233]" pageId="13" pageNumber="14" pagination="168 - 97" refId="ref17754" refString="Sereno PC. Lesothosaurus, ' fabrosaurids', and the early evolution of Ornithischia. Journal of Vertebrate Paleontology 1991; 11: 168 - 97. https: // doi. org / 10.1080 / 02724634.1991.10011386" type="journal article" year="1991">Sereno 1991</bibRefCitation>
|
||||
|
|
@ -1279,7 +1282,7 @@ The posterolateral processes of the premaxillae project much more posteriorly th
|
|||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FF51F89BF51D28F5" blockId="13.[113,762,1705,1949]" lastBlockId="13.[810,1461,144,1296]" pageId="13" pageNumber="14">
|
||||
The posterior tip of the posterolateral processes bears an articular surface (
|
||||
<figureCitation id="13162A23FFB0FF80FEEFF87BF13C2EEB" box="[307,370,1925,1949]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FEEFF87BF13C2EEB" box="[307,370,1925,1949]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
|
||||
). On the laterodorsal surface there are ridges that indicate a contact, most likely with the slightly overlapping nasal bone. That surface of the contact is orientated laterodorsally along the dorsal margin of the process. A similar contact surface is visible also on the medial side, covering the area just posterior to the natural dorsal border of the premaxillo-maxillary fenestra. Posteriorly, there is another ridged surface, orientated posteriorly, that may indicate contact with the posteroventral process of the nasal or with the maxilla.
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FC99FE75F3A02AC1" blockId="13.[810,1461,144,1296]" pageId="13" pageNumber="14">
|
||||
|
|
@ -1308,7 +1311,7 @@ and
|
|||
<emphasis id="B959EAB4FFB0FF80FBA3FCDDF4B32A4C" box="[1151,1277,803,826]" italics="true" pageId="13" pageNumber="14">Zuniceratops</emphasis>
|
||||
</taxonomicName>
|
||||
the posterolateral process is deeply bifurcated and its dorsal tip forms the anterior margin of the premaxillo-maxillary fenestra, while the posterior tip forms the anteroventral one, resulting in contact with the maxilla (
|
||||
<figureCitation id="13162A23FFB0FF80FC59FC61F3932AC1" box="[901,989,927,951]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="13" pageNumber="14">Fig. 12H</figureCitation>
|
||||
<figureCitation id="13162A23FFB0FF80FC59FC61F3932AC1" box="[901,989,927,951]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="13" pageNumber="14">Fig. 12H</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB0FF80FC99FC41F3E62C79" blockId="13.[810,1461,144,1296]" pageId="13" pageNumber="14">
|
||||
|
|
@ -1402,15 +1405,15 @@ The posterolateral processes of the premaxillary bone expand laterally, resultin
|
|||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB3FF83FF40FF6EF21F2812" blockId="14.[128,777,144,356]" pageId="14" pageNumber="15">
|
||||
On the lateral surface of the base of the posterolateral processes, just anterior to the premaxillo-maxillary fenestra, there is a foramen present in a shallow pocket. Such a foramen has not been traced in any ceratopsian with the premaxillo-maxillary fenestration. The additional foramen is present on the anteromedial wall of the premaxillo-maxillary fenestra, at the base of the posterolateral process of premaxilla (
|
||||
<figureCitation id="13162A23FFB3FF83FE24FEB2F20E2812" box="[504,576,332,356]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="14" pageNumber="15">Fig. 6A</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FE24FEB2F20E2812" box="[504,576,332,356]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB3FF83FF5DFE7BF3462A22" blockId="14.[128,779,389,1291]" pageId="14" pageNumber="15">
|
||||
<emphasis id="B959EAB4FFB3FF83FF5DFE7BF13028EB" box="[129,382,389,413]" italics="true" pageId="14" pageNumber="15">The premaxillary palate:</emphasis>
|
||||
The premaxillary palate is ventrally vaulted to a high degree and strongly arched. However, the arching is present much more posteriorly (posterior to the external nares;
|
||||
<figureCitation id="13162A23FFB3FF83FED7FE1DF12B288D" box="[267,357,483,507]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="14" pageNumber="15">Fig. 12A</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FED7FE1DF12B288D" box="[267,357,483,507]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="14" pageNumber="15">Fig. 12A</figureCitation>
|
||||
) than in ceratopsian dinosaurs, making the anteriormost portion of the snout solid in the cross-section. The dorsalmost part of the premaxillary palate is not preserved and it is not certain how deeply concave it was; however, due to the rugose surface of the proposed rostral bone preserved on the medial walls of the palate, it can be concluded that it reached up to 75% of the subnarial premaxillary height. In ceratopsian dinosaurs, the premaxillary palate is usually vaulted to the level of the lower quarter or third of the premaxillary body (
|
||||
<figureCitation id="13162A23FFB3FF83FD4BFD20F2932B80" box="[663,733,734,758]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." pageId="14" pageNumber="15">Fig. 12</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FD4BFD20F2932B80" box="[663,733,734,758]" captionStart="Figure 12" captionStartId="12.[129,194,1516,1540]" captionTargetBox="[132,1470,753,1485]" captionTargetId="figure-223@12.[129,1473,750,1488]" captionTargetPageId="12" captionText="Figure 12. Comparison of the snouts of selected ornithischian dinosaurs, with emphasis on the shape of the vaulted premaxillary palate and the cross-section of the anterior part (near the posterior border of the rostral bone, as indicated by the dashed line). A, Ajkaceratops kozmai MTM V2009.192.1; B, Stegouros elengassen CPAP-3165; C, Lesothosaurus diagnosticus NHMUK PV RU B17; D, Thescelosaurus neglectus NCSM 15728; E, Yinlong downsi IVPP V14530; F, Psittacosaurus sp. MPC-D 100/610; G, Leptoceratops gracilis CMN 8889; H, Bagaceratops rozhdestvenskyi (based on of ZPAL MgD-I/129 and MPC-D 100/535); I, Protoceratops andrewsi AMNH FARB 6433; J, Zuniceratops christopheri (based on MSM P2225, MSM P3197 and MSM P4185); K, Chasmosaurus sp. CMN 8801.The anterior part of the premaxillary palate is convex even in the early-diverging ceratopsians, while in Ajkaceratops the palatal convexity is displaced posteriorly.Not to scale." figureDoi="http://doi.org/10.5281/zenodo.14893356" httpUri="https://zenodo.org/record/14893356/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
|
||||
). In
|
||||
<taxonomicName id="4C2D4D25FFB3FF83FF5DFD03F0B12A63" box="[129,255,765,789]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB3FF83FF5DFD03F0B12A63" box="[129,255,765,789]" italics="true" pageId="14" pageNumber="15">Ajkaceratops</emphasis>
|
||||
|
|
@ -1503,11 +1506,11 @@ further shows a foramen that is located at the bottom of a shallow fossa, just a
|
|||
Only the anteriormost portions of the anteromedial processes of both maxillae are preserved, measuring
|
||||
<quantity id="4CD59B43FFB3FF83FB2CFEB2F5782815" box="[1264,1334,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="15" unit="mm" value="14.0">14 mm</quantity>
|
||||
each, and are wedged between the posteroventral processes of the premaxillae (
|
||||
<figureCitation id="13162A23FFB3FF83FC98FE75F3C628D5" box="[836,904,395,419]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="14" pageNumber="15">Figs 3</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FC98FE75F3C628D5" box="[836,904,395,419]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="14" pageNumber="15">Figs 3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB3FF83FC46FE75F3E928D5" box="[922,935,395,419]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." pageId="14" pageNumber="15">4</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FC46FE75F3E928D5" box="[922,935,395,419]" captionStart="Figure 4" captionStartId="5.[113,178,1411,1435]" captionTargetBox="[117,1454,147,1380]" captionTargetId="figure-264@5.[114,1458,144,1383]" captionTargetPageId="5" captionText="Figure 4. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in ventral view. The posterodorsal processes of premaxillae were digitally removed for clarity." figureDoi="http://doi.org/10.5281/zenodo.14893336" httpUri="https://zenodo.org/record/14893336/files/figure.png" pageId="14" pageNumber="15">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="13162A23FFB3FF83FC66FE75F38928D5" box="[954,967,395,419]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="14" pageNumber="15">6</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FC66FE75F38928D5" box="[954,967,395,419]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="14" pageNumber="15">6</figureCitation>
|
||||
). Because of the taphonomic deformation of the specimen, they are slightly displaced. Specifically, the anteromedial process of the right maxilla overlaps the left one dorsally. The intermaxillary and premaxillo-maxillary sutures are clearly visible in the ventral view. They seem to be almost completely preserved anteriorly, although the dorsal surface of the vaulted palate is eroded just in front of them. On the ventral surfaces of both maxillae there are small foramina present medially. Such foramina are sometimes present on the anteroventral processes of maxillae in some ceratopsians, as in
|
||||
<taxonomicName id="4C2D4D25FFB3FF83FA9CFD5BF39D2BAA" authorityName="Maryanska & Osmólska" authorityYear="1975" class="Reptilia" family="Protoceratopsidae" genus="Bagaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rozhdestvenskyi">
|
||||
<emphasis id="B959EAB4FFB3FF83FA9CFD5BF39D2BAA" italics="true" pageId="14" pageNumber="15">Bagaceratops rozhdestvenskyi</emphasis>
|
||||
|
|
@ -1544,7 +1547,7 @@ In
|
|||
<emphasis id="B959EAB4FFB3FF83FCAEFC20F3A02A80" box="[882,1006,990,1014]" italics="true" pageId="14" pageNumber="15">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
the preserved portions of the maxillae direct anterodorsally, with an angle of nearly 45° in relation to the longer axis of the ventral margin of the premaxilla. They are placed more ventrally than the posteroventral processes of the premaxillae, resulting in being almost visible in the lateral view (
|
||||
<figureCitation id="13162A23FFB3FF83FAE0FBA5F5382D05" box="[1340,1398,1115,1139]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FAE0FBA5F5382D05" box="[1340,1398,1115,1139]" captionStart="Figure 3" captionStartId="4.[129,194,1840,1864]" captionTargetBox="[161,1441,148,1807]" captionTargetId="figure-46@4.[158,1444,144,1812]" captionTargetPageId="4" captionText="Figure 3. Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14893334" httpUri="https://zenodo.org/record/14893334/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
). In all ceratopsian dinosaurs the anteroventral processes of the maxillae are placed much more dorsally, nearly in the same plane as the premaxillary palate, and are deeply hidden in the lateral view by the lateroventral edge of the premaxillae and maxillae. In the
|
||||
<typeStatus id="54968804FFB3FF83FCE7FB06F3D82C66" box="[827,918,1272,1296]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
|
|
@ -1561,7 +1564,7 @@ In
|
|||
<emphasis id="B959EAB4FFB3FF83FCB3FA8BF3A42CFB" box="[879,1002,1397,1421]" italics="true" pageId="14" pageNumber="15">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
the contact between the maxilla and premaxilla seems to be limited to the posterior surface of the posteroventral process of the premaxilla only, with no lateral overlapping of the premaxilla by the anterolateral maxillary process (
|
||||
<figureCitation id="13162A23FFB3FF83FAEFFA2DF5252C9D" box="[1331,1387,1491,1515]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." pageId="14" pageNumber="15">Fig. 6</figureCitation>
|
||||
<figureCitation id="13162A23FFB3FF83FAEFFA2DF5252C9D" box="[1331,1387,1491,1515]" captionStart="Figure 6" captionStartId="7.[115,180,1219,1243]" captionTargetBox="[117,1455,147,1188]" captionTargetId="figure-421@7.[114,1458,144,1191]" captionTargetPageId="7" captionText="Figure 6.Ajkaceratops kozmai MTM V2009.192.1, holotype specimen in posterior view." figureDoi="http://doi.org/10.5281/zenodo.14893342" httpUri="https://zenodo.org/record/14893342/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
|
||||
). It suggests that the anterolateral process of the maxilla was not well-developed in
|
||||
<taxonomicName id="4C2D4D25FFB3FF83FC63F9ECF4772F5C" box="[959,1081,1554,1578]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB3FF83FC63F9ECF4772F5C" box="[959,1081,1554,1578]" italics="true" pageId="14" pageNumber="15">Ajkaceratops</emphasis>
|
||||
|
|
@ -1624,7 +1627,7 @@ to
|
|||
<emphasis id="B959EAB4FFB3FF83FBE4F857F5462EB7" box="[1080,1288,1961,1985]" italics="true" pageId="14" pageNumber="15">Ajkaceratops kozmai</emphasis>
|
||||
</taxonomicName>
|
||||
. Only the largest specimen, MTM V2009.195.1, preserves the anterodorsal tip of the bone (
|
||||
<figureCitation id="13162A23FFB2FF82FF02FF51F15929B1" box="[222,279,175,199]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FF02FF51F15929B1" box="[222,279,175,199]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." figureDoi="http://doi.org/10.5281/zenodo.14893346" httpUri="https://zenodo.org/record/14893346/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||||
). The anterior end of the predentary is sharply pointed in dorsal view, and it curves anterodorsally in lateral view, as in all ceratopsians (with the exception of
|
||||
<taxonomicName id="4C2D4D25FFB2FF82FDADFF11F0D12853" class="Reptilia" family="Psittacosauridae" genus="Psittacosaurus" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="undetermined">
|
||||
<emphasis id="B959EAB4FFB2FF82FDADFF11F2B22870" box="[625,764,239,262]" italics="true" pageId="15" pageNumber="16">Psittacosaurus</emphasis>
|
||||
|
|
@ -1679,7 +1682,7 @@ Riguetti
|
|||
)
|
||||
</taxonomicName>
|
||||
. Similarly to the proposed rostral bone of the type specimen (MTM V2009.192.1) and other referred predentaries (MTM V2009.193.1;
|
||||
<figureCitation id="13162A23FFB2FF82FED9FE17F1712B77" box="[261,319,489,513]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FED9FE17F1712B77" box="[261,319,489,513]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." figureDoi="http://doi.org/10.5281/zenodo.14893348" httpUri="https://zenodo.org/record/14893348/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||||
), the anterior surface is unkeeled, and the tip of the predentary is subtriangular in transverse cross-section. The pattern of ornamentation accurately matches that of the
|
||||
<typeStatus id="54968804FFB2FF82FFADFDB9F0812B29" box="[113,207,583,607]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||||
, such as the tip being deprived of any rugosities, with the two ventralmost pits placed in the longitudinal grooves some distance from the anterodorsal end of the bone. On the dorsal surface of the predentary MTM V2009.195.1, the pits are less common and there are three longitudinal grooves visible, running from just behind the anterior tip of the bone.
|
||||
|
|
@ -1691,9 +1694,9 @@ The smaller specimen referred to
|
|||
, MTM V
|
||||
</taxonomicName>
|
||||
2009.194.1, exhibits distinct morphology, with relatively densely distributed, large, and deep pits (
|
||||
<figureCitation id="13162A23FFB2FF82FDF7FCBFF2EB2A2F" box="[555,677,833,857]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." pageId="15" pageNumber="16">Fig. 10A–D</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FDF7FCBFF2EB2A2F" box="[555,677,833,857]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." figureDoi="http://doi.org/10.5281/zenodo.14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="15" pageNumber="16">Fig. 10A–D</figureCitation>
|
||||
). There is a quite deep medial longitudinal groove on the dorsal surface of MTM V2009.194.1, and in the smallest specimen, MTM V2009.196.1 (
|
||||
<figureCitation id="13162A23FFB2FF82FED0FC61F1C62AC1" box="[268,392,927,951]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." pageId="15" pageNumber="16">Fig. 10E–H</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FED0FC61F1C62AC1" box="[268,392,927,951]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." figureDoi="http://doi.org/10.5281/zenodo.14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="15" pageNumber="16">Fig. 10E–H</figureCitation>
|
||||
). The posterolateral processes are not well-developed in these
|
||||
<specimenCount id="9D2BFD2FFFB2FF82FE7CFC41F2732AA1" box="[416,573,959,983]" count="2" pageId="15" pageNumber="16" type="generic">two specimens</specimenCount>
|
||||
. As noted by
|
||||
|
|
@ -1724,7 +1727,7 @@ There is no visible slit into the posterolateral process that would enclose the
|
|||
. A clear articulation surface is visible on the right posterolateral process of MTM V2009.194.1, matching the Morphotype I by
|
||||
<bibRefCitation id="EFBC4B57FFB2FF82FFADF9B5F24F2F15" author="Nabavizadeh A & Weishampel DB" box="[113,513,1611,1635]" pageId="15" pageNumber="16" pagination="1358 - 88" refId="ref17235" refString="Nabavizadeh A, Weishampel DB. The predentary bone and its significance in the evolution of feeding mechanisms in ornithischian dinosaurs. Anatomical Record (Hoboken, N. J.: 2007) 2016; 299: 1358 - 88. https: // doi. org / 10.1002 / ar. 23455" type="journal article" year="2016">Nabavizadeh and Weishampel (2016)</bibRefCitation>
|
||||
. However, in specimen MTM V2009.193.1, the articular surface for the dentaries on the posterior surface of the predentary seems to be relatively deeply excavated (
|
||||
<figureCitation id="13162A23FFB2FF82FF38F957F1522FB7" box="[228,284,1705,1729]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FF38F957F1522FB7" box="[228,284,1705,1729]" captionStart="Figure 9" captionStartId="10.[129,194,814,838]" captionTargetBox="[132,1470,147,784]" captionTargetId="figure-660@10.[129,1473,144,787]" captionTargetPageId="10" captionText="Figure 9. Ajkaceratops kozmai MTM V2009.193.1, referred predentary in right lateral (A), anterior (B), dorsal (C), left lateral (D), posterior (E) and ventral (F) views." figureDoi="http://doi.org/10.5281/zenodo.14893348" httpUri="https://zenodo.org/record/14893348/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B9236A6FFB2FF82FFADF91DF5D528B4" blockId="15.[113,763,1763,1975]" lastBlockId="15.[809,1461,144,1077]" pageId="15" pageNumber="16">
|
||||
|
|
@ -1736,7 +1739,7 @@ A posteroventral process is completely preserved in MTM V2009.194.1 and partiall
|
|||
(2010
|
||||
</bibRefCitation>
|
||||
:SI), but it is no longer preserved in that specimen (
|
||||
<figureCitation id="13162A23FFB2FF82FE0FF89EF2062E0E" box="[467,584,1888,1912]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." pageId="15" pageNumber="16">Fig. 10E–H</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FE0FF89EF2062E0E" box="[467,584,1888,1912]" captionStart="Figure 10" captionStartId="11.[113,178,1114,1138]" captionTargetBox="[117,1454,147,1083]" captionTargetId="figure-469@11.[114,1458,144,1087]" captionTargetPageId="11" captionText="Figure 10. Ajkaceratops kozmai MTM V2009.194.1 (A–D) and MTM V2009.196.1 (E–H), referred predentaries.MTM V2009.194.1 in dorsal (A), ventral (B), left lateral (C), and right lateral (D) views. MTM V2009.196.1 in posterior (E), left lateral (F), right lateral (G), and dorsal (H) views." figureDoi="http://doi.org/10.5281/zenodo.14893350" httpUri="https://zenodo.org/record/14893350/files/figure.png" pageId="15" pageNumber="16">Fig. 10E–H</figureCitation>
|
||||
). In
|
||||
<taxonomicName id="4C2D4D25FFB2FF82FDAAF89EF2BA2E0E" box="[630,756,1888,1912]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFB2FF82FDAAF89EF2BA2E0E" box="[630,756,1888,1912]" italics="true" pageId="15" pageNumber="16">Ajkaceratops</emphasis>
|
||||
|
|
@ -1902,7 +1905,7 @@ One of the specimens referred to
|
|||
<emphasis id="B959EAB4FFB2FF82FB52FB70F52F2DD0" box="[1166,1377,1166,1190]" italics="true" pageId="15" pageNumber="16">Ajkaceratops kozmai</emphasis>
|
||||
</taxonomicName>
|
||||
(MTM V2009.195.1) preserves a fused, 17 mm-long anteriormost portion of the right dentary bone (
|
||||
<figureCitation id="13162A23FFB2FF82FB68FB33F4BD2D93" box="[1204,1267,1229,1253]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||||
<figureCitation id="13162A23FFB2FF82FB68FB33F4BD2D93" box="[1204,1267,1229,1253]" captionStart="Figure 8" captionStartId="9.[114,179,1247,1271]" captionTargetBox="[117,1455,150,1216]" captionTargetId="figure-380@9.[114,1458,144,1219]" captionTargetPageId="9" captionText="Figure 8. Ajkaceratops kozmai MTM V2009.195.1, referred predentary and a dentary fragment in left lateral (A, B), right lateral (C, D), anterior (E, F), dorsal (G, H), and ventral (I, J) views." figureDoi="http://doi.org/10.5281/zenodo.14893346" httpUri="https://zenodo.org/record/14893346/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||||
). The merging of the predentary with the dentary is rarely reported for ornithischians, although the bones fuse late in ontogeny in
|
||||
<emphasis id="B959EAB4FFB2FF82FCF6FAD5F3E32C34" box="[810,941,1323,1346]" italics="true" pageId="15" pageNumber="16">Psittacosurus</emphasis>
|
||||
(
|
||||
|
|
@ -2124,7 +2127,7 @@ As could be expected, OTUs 1 and 2 were reconstructed as part of Ceratopsia usin
|
|||
<emphasis id="B959EAB4FFADFF9DFB7AFCFDF56F2A6D" box="[1190,1313,771,795]" italics="true" pageId="16" pageNumber="17">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
within the clade is well-supported in both these runs (see
|
||||
<figureCitation id="13162A23FFADFF9DFB3CFCDCF56A2A4C" box="[1248,1316,802,826]" captionStart="Figure 13" captionStartId="17.[113,178,1138,1162]" captionTargetBox="[117,1449,147,1107]" captionTargetId="figure-355@17.[114,1458,144,1110]" captionTargetPageId="17" captionText="Figure 13. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A, D, G) Han et al. (2018), (B, E, H) Herne et al. (2019), and (C, F, I) Dieudonné et al.(2021). The placements are indicated on segments of the phylogenetic trees and reflect alternative interpretations of the osteological features of Ajkaceratops kozmai: A–C, operational taxonomic unit (OTU) scored using the information from the holotype as well as the referred material; with the anterior section interpreted as the rostral bone (‘OTU 1’; for explanation see the main text); D–F, OTU scored using the information from the holotype only; with the anterior section interpreted as the rostral bone (‘OTU 2’); G–I, OTU scored using the information from the holotype and the referred material; with the anterior section interpreted as the premaxillary bone (‘OTU 3’). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||||
<figureCitation id="13162A23FFADFF9DFB3CFCDCF56A2A4C" box="[1248,1316,802,826]" captionStart="Figure 13" captionStartId="17.[113,178,1138,1162]" captionTargetBox="[117,1449,147,1107]" captionTargetId="figure-355@17.[114,1458,144,1110]" captionTargetPageId="17" captionText="Figure 13. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A, D, G) Han et al. (2018), (B, E, H) Herne et al. (2019), and (C, F, I) Dieudonné et al.(2021). The placements are indicated on segments of the phylogenetic trees and reflect alternative interpretations of the osteological features of Ajkaceratops kozmai: A–C, operational taxonomic unit (OTU) scored using the information from the holotype as well as the referred material; with the anterior section interpreted as the rostral bone (‘OTU 1’; for explanation see the main text); D–F, OTU scored using the information from the holotype only; with the anterior section interpreted as the rostral bone (‘OTU 2’); G–I, OTU scored using the information from the holotype and the referred material; with the anterior section interpreted as the premaxillary bone (‘OTU 3’). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." figureDoi="http://doi.org/10.5281/zenodo.14893358" httpUri="https://zenodo.org/record/14893358/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||||
). In HA18, the dataset that was originally constructed to focus especially on early-diverging ceratopsians and that comprises the highest number of ceratopsian taxa from among the ornithischian-wide datasets used in the present study, OTUs 1 and 2 are nested within Euceratopsia, as the sister-taxon to
|
||||
<taxonomicName id="4C2D4D25FFADFF9DFBC9FC41F4C32AA0" baseAuthorityName="MSM P" baseAuthorityYear="2225" box="[1045,1165,959,982]" genus="Zuniceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFADFF9DFBC9FC41F4C32AA0" box="[1045,1165,959,982]" italics="true" pageId="16" pageNumber="17">Zuniceratops</emphasis>
|
||||
|
|
@ -2134,7 +2137,7 @@ within the clade is well-supported in both these runs (see
|
|||
<emphasis id="B959EAB4FFADFF9DFCE5FBA5F3FA2D05" box="[825,948,1115,1139]" italics="true" pageId="16" pageNumber="17">Ajkaceratops</emphasis>
|
||||
</taxonomicName>
|
||||
as a ceratopsian, though in D21 the support for such placement was poorer (
|
||||
<figureCitation id="13162A23FFADFF9DFBF1FB85F43D2DE5" box="[1069,1139,1147,1171]" captionStart="Figure 13" captionStartId="17.[113,178,1138,1162]" captionTargetBox="[117,1449,147,1107]" captionTargetId="figure-355@17.[114,1458,144,1110]" captionTargetPageId="17" captionText="Figure 13. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A, D, G) Han et al. (2018), (B, E, H) Herne et al. (2019), and (C, F, I) Dieudonné et al.(2021). The placements are indicated on segments of the phylogenetic trees and reflect alternative interpretations of the osteological features of Ajkaceratops kozmai: A–C, operational taxonomic unit (OTU) scored using the information from the holotype as well as the referred material; with the anterior section interpreted as the rostral bone (‘OTU 1’; for explanation see the main text); D–F, OTU scored using the information from the holotype only; with the anterior section interpreted as the rostral bone (‘OTU 2’); G–I, OTU scored using the information from the holotype and the referred material; with the anterior section interpreted as the premaxillary bone (‘OTU 3’). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||||
<figureCitation id="13162A23FFADFF9DFBF1FB85F43D2DE5" box="[1069,1139,1147,1171]" captionStart="Figure 13" captionStartId="17.[113,178,1138,1162]" captionTargetBox="[117,1449,147,1107]" captionTargetId="figure-355@17.[114,1458,144,1110]" captionTargetPageId="17" captionText="Figure 13. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A, D, G) Han et al. (2018), (B, E, H) Herne et al. (2019), and (C, F, I) Dieudonné et al.(2021). The placements are indicated on segments of the phylogenetic trees and reflect alternative interpretations of the osteological features of Ajkaceratops kozmai: A–C, operational taxonomic unit (OTU) scored using the information from the holotype as well as the referred material; with the anterior section interpreted as the rostral bone (‘OTU 1’; for explanation see the main text); D–F, OTU scored using the information from the holotype only; with the anterior section interpreted as the rostral bone (‘OTU 2’); G–I, OTU scored using the information from the holotype and the referred material; with the anterior section interpreted as the premaxillary bone (‘OTU 3’). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." figureDoi="http://doi.org/10.5281/zenodo.14893358" httpUri="https://zenodo.org/record/14893358/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||||
). The analysis of HE19 inferred
|
||||
<taxonomicName id="4C2D4D25FFADFF9DFCE5FB64F3FA2DC4" box="[825,948,1178,1202]" genus="Ajkaceratops" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B959EAB4FFADFF9DFCE5FB64F3FA2DC4" box="[825,948,1178,1202]" italics="true" pageId="16" pageNumber="17">Ajkaceratops</emphasis>
|
||||
|
|
@ -2152,11 +2155,11 @@ was inferred as the sister-taxon to
|
|||
and
|
||||
<taxonomicName id="4C2D4D25FFADFF9DFB0DFAC9F5262C39" baseAuthorityName="Cooper" baseAuthorityYear="1985" box="[1233,1384,1335,1359]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neornithischia" pageId="16" pageNumber="17" phylum="Chordata" rank="order">Neornithischia</taxonomicName>
|
||||
, respectively (
|
||||
<figureCitation id="13162A23FFADFF9DFCA4FAA8F3F22C18" box="[888,956,1366,1390]" captionStart="Figure 14" captionStartId="18.[128,193,1548,1572]" captionTargetBox="[132,1470,147,1518]" captionTargetId="figure-173@18.[129,1473,144,1521]" captionTargetPageId="18" captionText="Figure 14. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A) Han et al. (2018), (B) Herne et al. (2019), and (C) Dieudonné et al. (2021). The placements are indicated on segments of the phylogenetic trees.In all three analyses, the operational taxonomic unit (OTU) was scored using the information from the holotype only; with the anterior section interpreted as the premaxillary bone (‘OTU 4’; for explanation see the main text). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||||
<figureCitation id="13162A23FFADFF9DFCA4FAA8F3F22C18" box="[888,956,1366,1390]" captionStart="Figure 14" captionStartId="18.[128,193,1548,1572]" captionTargetBox="[132,1470,147,1518]" captionTargetId="figure-173@18.[129,1473,144,1521]" captionTargetPageId="18" captionText="Figure 14. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A) Han et al. (2018), (B) Herne et al. (2019), and (C) Dieudonné et al. (2021). The placements are indicated on segments of the phylogenetic trees.In all three analyses, the operational taxonomic unit (OTU) was scored using the information from the holotype only; with the anterior section interpreted as the premaxillary bone (‘OTU 4’; for explanation see the main text). Numbers at nodes indicate Bremer support values. For full tree topologies see Supporting Information, S1." figureDoi="http://doi.org/10.5281/zenodo.14893360" httpUri="https://zenodo.org/record/14893360/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||||
). When assessing the majority rule consensus trees (that, however, should be treated with considerably greater caution), HE19 tends to place OTU 4 at an early-diverging position among ornithopods, while D21 infers it at the basal polytomy of
|
||||
<taxonomicName id="4C2D4D25FFADFF9DFCE6FA2DF3812C9D" baseAuthorityName="Cooper" baseAuthorityYear="1985" box="[826,975,1491,1515]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neornithischia" pageId="16" pageNumber="17" phylum="Chordata" rank="order">Neornithischia</taxonomicName>
|
||||
(Supplementary Material 1: Figs S18 and S28, respectively). Finally, when assessing the referred predentaries only (OTU 5), the analyses place the material either in a large polytomy with other ornithischians (HA18), with other ceratopsians (D21) or possibly within Neoceratopsia (HE19) (
|
||||
<figureCitation id="13162A23FFADFF9DFB3AF9AFF5662F1E" box="[1254,1320,1616,1641]" captionStart="Figure 15" captionStartId="19.[113,178,1876,1900]" captionTargetBox="[146,1355,152,1845]" captionTargetId="figure-6@19.[143,1429,149,1848]" captionTargetPageId="19" captionText="Figure 15. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A) Han et al. (2018), (B) Herne et al. (2019), and (C) Dieudonné et al. (2021). The placements are indicated on segments of the phylogenetic trees.In all three analyses, the operational taxonomic unit (OTU) was scored using the information from the referred predentaries only (‘OTU 5’; for explanation see the main text). Numbers at nodes indicate Bremer support values.For full tree topologies see Supporting Information, S1." pageId="16" pageNumber="17">Fig. 15</figureCitation>
|
||||
<figureCitation id="13162A23FFADFF9DFB3AF9AFF5662F1E" box="[1254,1320,1616,1641]" captionStart="Figure 15" captionStartId="19.[113,178,1876,1900]" captionTargetBox="[146,1355,152,1845]" captionTargetId="figure-6@19.[143,1429,149,1848]" captionTargetPageId="19" captionText="Figure 15. Phylogenetic placement of Ajkaceratops kozmai among ornithischian dinosaurs reconstructed using the matrices of (A) Han et al. (2018), (B) Herne et al. (2019), and (C) Dieudonné et al. (2021). The placements are indicated on segments of the phylogenetic trees.In all three analyses, the operational taxonomic unit (OTU) was scored using the information from the referred predentaries only (‘OTU 5’; for explanation see the main text). Numbers at nodes indicate Bremer support values.For full tree topologies see Supporting Information, S1." figureDoi="http://doi.org/10.5281/zenodo.14893362" httpUri="https://zenodo.org/record/14893362/files/figure.png" pageId="16" pageNumber="17">Fig. 15</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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Reference in a new issue