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<mods:title>The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the &quot; hottest &quot; biodiversity hotspots</mods:title>
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<mods:namePart>Lorenz, Michelle</mods:namePart>
<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:namePart>Loria, Stephanie F.</mods:namePart>
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<mods:namePart>Harvey, Mark S.</mods:namePart>
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<taxonomicName LSID="https://zoobank.org/NomenclaturalActs/C563A8D4-7620-426E-8A7E-A257C1984516" authority="Lorenz &amp; Loria &amp; Harvey &amp; Harms, 2022" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Antsirananaella marlae" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="marlae" status="sp. nov.">Antsirananaella marlae</taxonomicName>
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<paragraph pageId="0" pageNumber="649">Type material mxamined.</paragraph>
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<paragraph id="A1ACAC2F8C53B8863F4C17D498D5F08D" pageId="0" pageNumber="649">Type material mxamined.</paragraph>
<paragraph id="E3182635913627567656F95658507105" pageId="0" pageNumber="649">
MADAGASCAR:
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Province [Sava Region]: Holotype ♂ (CAS 9072444 [BLF9974]), allotype ♀ (ZMH-A0016693), paratypes: 3 ♀, 2 ♂ (CAS 9072444 [BLF9974]), 2 ♀, 2 ♂ (ZMH-A0016694 - ZMH-A0016697),
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, 26.3km 10° NNE Daraina, 175 m,
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, 10 December 2003, B.L. Fisher.
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<paragraph pageId="0" pageNumber="649">Diagnosis.</paragraph>
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<paragraph id="96050F1FE17F91C591CA31D5855738A0" pageId="0" pageNumber="649">Diagnosis.</paragraph>
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Differs from congeners by large body size (e.g., male body length 2.17 versus 1.80-2.14, male carapace length 0.69 versus 0.55-0.66); a slender carapace (1.47 times longer than broad versus 1.36-1.40); medio-lateral mounds more pronounced than in
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<emphasis id="46826FAE30CE0348661CD863FBD8A091" italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="6D6BB5D7779A5D2F8871DF1972FE04C4" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
; a slender pedipalpal femur (1.80 (♂), 1.84 (♀) times longer than broad versus 1.70-1.79 (♂), 1.74-1.80 (♀)); pedipalpal trichobothrial pattern:
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on movable finger on same sagittal level as
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<emphasis id="AB5AE74804579A36BEA5519B30298074" italics="true" pageId="0" pageNumber="649">est</emphasis>
on fixed finger (slightly more distal in
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<emphasis italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
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<emphasis id="311F03F98BABF42AF45D9EC981DB9528" italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="BB806F0BDF7749E3AECAED8C9BCF3327" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
and in
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<emphasis italics="true" pageId="0" pageNumber="649">A. faulstichi</emphasis>
<taxonomicName id="20DD554C98659195B2D04F10EECAAD56" lsidName="A. faulstichi" pageId="0" pageNumber="649" rank="species" species="faulstichi">
<emphasis id="79DFAAC590F36F2999C1400D1FB046E3" italics="true" pageId="0" pageNumber="649">A. faulstichi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="0B894833F7B5588AE5DC88E878703CD0" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
),
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<emphasis id="C0C0DB3D6C5E57495F66FCC0B15B0333" italics="true" pageId="0" pageNumber="649">et</emphasis>
situated in the distal third between
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<emphasis id="74E8F1B619FCA2A04A5B2CF86345AB6A" italics="true" pageId="0" pageNumber="649">dt</emphasis>
and
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<emphasis id="B8471B0FD883DDF14BFA4CE54CEA70E6" italics="true" pageId="0" pageNumber="649">eb</emphasis>
on fixed finger (situated midway between
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<emphasis id="E65271D42DC802F680517A2CEC2F43DC" italics="true" pageId="0" pageNumber="649">dt</emphasis>
and
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<emphasis id="0668EDF139C8908AF7F118BEA4D0E3CE" italics="true" pageId="0" pageNumber="649">eb</emphasis>
on fixed finger in
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<emphasis italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
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<emphasis id="115EBEAD0AA91494370EBFF759DB1E8C" italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="2477544F678AEBDEC7DA72BF063EA01B" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
and in
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<emphasis italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
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<emphasis id="CA356817F967C8FCE38B156E6CD8A934" italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="2408B9BE639B62218561B8196C07DA82" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
);
<emphasis italics="true" pageId="0" pageNumber="649">it</emphasis>
<emphasis id="16F365D7C827FAACD38B38DF6499AA3A" italics="true" pageId="0" pageNumber="649">it</emphasis>
between
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
<emphasis id="2256C4147C4D67423D7EFA228ADB2A36" italics="true" pageId="0" pageNumber="649">eb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
<emphasis id="DEF0FE82F448530D418917889F1703C0" italics="true" pageId="0" pageNumber="649">et</emphasis>
situated closer to
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
<emphasis id="0E5A6646067CE69F69B2FBABFA0B6C51" italics="true" pageId="0" pageNumber="649">eb</emphasis>
than to
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
<emphasis id="17A742AB8DBB1352C4AC949713EB7403" italics="true" pageId="0" pageNumber="649">et</emphasis>
(equidistant in
<taxonomicName lsidName="A. leniae" pageId="0" pageNumber="649" rank="species" species="leniae">
<emphasis italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
<taxonomicName id="62D01D73E07A1CB4C6EC706AE67766B3" lsidName="A. leniae" pageId="0" pageNumber="649" rank="species" species="leniae">
<emphasis id="622EFC5D3A2826F849F486C81C73598A" italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="5B6A38AFCB9D6CFC254FFA2D256AB295" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="etymology">
<paragraph pageId="0" pageNumber="649">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="649">
<subSubSection id="CC47CAB4DAA234B8CA9B11F7BFCCA391" pageId="0" pageNumber="649" type="etymology">
<paragraph id="B32B855BDB21640A8D1F09412B2F7737" pageId="0" pageNumber="649">Etymology.</paragraph>
<paragraph id="C3E572763A97FC756649E11DD5E8040C" pageId="0" pageNumber="649">
The specific epithet is a patronym in honour of Marla Elisa Nibasumba, the senior
<normalizedToken originalValue="authors">author's</normalizedToken>
<normalizedToken id="F284BFE61DE3C5968CC620CBEE53B3A9" originalValue="authors">author's</normalizedToken>
godchild.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="description">
<paragraph pageId="0" pageNumber="649">Description.</paragraph>
<paragraph pageId="0" pageNumber="649">
<subSubSection id="82153C3A58F52B0D0FAAF082791EE3A8" pageId="0" pageNumber="649" type="description">
<paragraph id="400BDB1BF941B59750E319604D1EA2C0" pageId="0" pageNumber="649">Description.</paragraph>
<paragraph id="DA53F5F64CEA1339FB69019ED925BB3A" pageId="0" pageNumber="649">
The following description is based on holotype and allotype. -
<emphasis italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
<emphasis id="D0BA7FDACA243C3D46A51C51098D59EE" italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Drawings of Antsirananaella marlae sp. nov.: A carapace dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649">10A</figureCitation>
<figureCitation id="FAC092B9FCCB6AAC6FE5C7CBEA84C3F9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Drawings of Antsirananaella marlae sp. nov.: A carapace dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649">10A</figureCitation>
): 1.43-1.49 (♂), 1.30-1.46 (♀) times longer than broad. -
<emphasis italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
<emphasis id="1AAC42DE28A6BBB79B13054440AC09B9" italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Drawings of Antsirananaella marlae sp. nov.: A carapace dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649">10C</figureCitation>
<figureCitation id="FFC32AF3E2FEE045F23B57363C6D97DA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Drawings of Antsirananaella marlae sp. nov.: A carapace dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649">10C</figureCitation>
): Trichobothrial pattern:
<emphasis italics="true" pageId="0" pageNumber="649">sb</emphasis>
<emphasis id="BD58BCE9C0EDD10DEB4723C3AFB82122" italics="true" pageId="0" pageNumber="649">sb</emphasis>
on the movable finger and
<emphasis italics="true" pageId="0" pageNumber="649">est</emphasis>
<emphasis id="B9B5F820D6DBC91C6C7890AE818E4DC1" italics="true" pageId="0" pageNumber="649">est</emphasis>
on the fixed finger on the same sagittal level.
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
<emphasis id="FD0901820CF79FAC41942EBF959B9768" italics="true" pageId="0" pageNumber="649">et</emphasis>
situated in the distal third between
<emphasis italics="true" pageId="0" pageNumber="649">dt</emphasis>
<emphasis id="35E73E953BBFF8A3F4C1F29F0194BB70" italics="true" pageId="0" pageNumber="649">dt</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
<emphasis id="5E05009C9AD9A934A77FE987354C36C4" italics="true" pageId="0" pageNumber="649">eb</emphasis>
on the fixed finger. Chelal fixed finger with 11 teeth in the OR, 21 teeth in the MR, and 12 teeth in the IR; movable finger with 11 teeth in the OR, 13 teeth in the MR, and 13 teeth in the IR. -
<taxonomicName genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="649">Dimensions</emphasis>
<taxonomicName id="9DCEEECDB13E797263063381E59B5992" genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus">
<emphasis id="C733814F4BC51C64F956D046AD9E515E" italics="true" pageId="0" pageNumber="649">Dimensions</emphasis>
</taxonomicName>
(mm):
<emphasis italics="true" pageId="0" pageNumber="649">Holotype</emphasis>
<emphasis id="B7705D03471477715C41BBABF3727AD8" italics="true" pageId="0" pageNumber="649">Holotype</emphasis>
♂: Body length 1.81; abdomen length 1.25; abdomen width 0.99 (without pleura), 1.11 (with pleura); carapace length 0.55; carapace width 0.37. Pedipalp: trochanter 0.24; femur length 0.54; femur width 0.29; patella 0.43; chela (without pedicel) 0.49; hand length (without pedicel) 0.11; width 0.13; movable finger length 0.40. Leg I: trochanter 0.10; femur 0.13; patella 0.16; tibia 0.14; tarsus 0.25. Leg IV: trochanter 0.16; femur 0.11; patella 0.29; tibia 0.28; tarsus 0.31.
<emphasis italics="true" pageId="0" pageNumber="649">Allotype</emphasis>
<emphasis id="28A805C6F1DDC153DABF92F4F4E6617A" italics="true" pageId="0" pageNumber="649">Allotype</emphasis>
♀: Body length 2.30; abdomen length 1.60; abdomen width 1.21 (without pleura), 1.44 (with pleura); carapace length 0.67; carapace width 0.46. Pedipalp: trochanter 0.31; femur length 0.68; femur width 0.39; patella length 0.54; chela (without pedicel) 0.60; hand length (without pedicel) 0.14; hand width 0.18; movable finger length 0.48. Leg I: trochanter 0.11; femur 0.24; patella 0.22; tibia 0.15; tarsus 0.25. Leg IV: trochanter 0.21; femur 0.14; patella 0.35; tibia 0.41; tarsus 0.39.
</paragraph>
<caption doi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="649">
<emphasis bold="true" pageId="0" pageNumber="649">Figure 10.</emphasis>
<caption id="057CD949B686E0E5603B8347071CBAF4" doi="10.3897/asp.80.e90570.figure10" httpUri="https://binary.pensoft.net/fig/782002" pageId="0" pageNumber="649" start="Figure 10" startId="F10">
<paragraph id="DFECE8B9836F6CB96856D0DCDB923B35" pageId="0" pageNumber="649">
<emphasis id="A7ADADBB3A9E93B076BC07CBA0BCBB30" bold="true" pageId="0" pageNumber="649">Figure 10.</emphasis>
Drawings of
<taxonomicName authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella marlae" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="marlae">
<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella marlae</emphasis>
<taxonomicName id="AE16E2641D753E1EDDDBAFD8434C6BEB" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" class="Arachnida" family="Feaellidae" genus="Antsirananaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antsirananaella marlae" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="marlae">
<emphasis id="EEEE4A57AAF0127D1D205034A0DB7828" italics="true" pageId="0" pageNumber="649">Antsirananaella marlae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
<emphasis id="E9806CD46B13884481EA1F6BE796D506" bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
:
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
<emphasis id="1B3754045B3CB173E16FA245361B204A" bold="true" pageId="0" pageNumber="649">A</emphasis>
carapace dorsal;
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
<emphasis id="218107AF181298B69FC765E329913CC5" bold="true" pageId="0" pageNumber="649">B</emphasis>
leg IV;
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
<emphasis id="9CAC4803B2E7D36B1962CE92A822BD20" bold="true" pageId="0" pageNumber="649">C</emphasis>
left chela from dorsal with trichobothrial pattern;
<emphasis bold="true" pageId="0" pageNumber="649">D</emphasis>
<emphasis id="7EB610C62C6901D4616CBDBD12479465" bold="true" pageId="0" pageNumber="649">D</emphasis>
right pedipalp from dorsal. Scale bars: 0.2 mm
<emphasis bold="true" pageId="0" pageNumber="649">(A</emphasis>
<emphasis id="47EE8BD83DB730FEA58AD0CCE4892B46" bold="true" pageId="0" pageNumber="649">(A</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="649">C)</emphasis>
<emphasis id="A52BF60EC8D0D4E959D5EB6132C83FB1" bold="true" pageId="0" pageNumber="649">C)</emphasis>
; 0.4 mm
<emphasis bold="true" pageId="0" pageNumber="649">(D)</emphasis>
<emphasis id="D34D088D1B944583D281505152FB5F3D" bold="true" pageId="0" pageNumber="649">(D)</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="variation">
<paragraph pageId="0" pageNumber="649">Variation.</paragraph>
<paragraph pageId="0" pageNumber="649">Body length 1.70-1.91 (♂), 2.05-2.19 (♀); abdomen length 1.14-1.32 (♂), 1.37-1.46 (♀), width 0.98-1.07 (♂), 1.18-1.25 (♀) (without pleura), 1.02-1.18 (♂), 1.29-1.38 (♀) (with pleura), 1.17-1.28 (♂), 1.14-1.24 (♀) times longer than broad; carapace length 0.52-0.58 (♂), 0.63-0.67 (♀), width 0.36-0.39 (♂), 0.46-0.48 (♀), 1.43-1.49 (♂), 1.30-1.36 (♀) times longer than broad; Pedipalp: trochanter 0.22-0.25 (♂), 0.27-0.30 (♀); femur length 0.52-0.56 (♂), 0.70-0.71 (♀), width 0.29-0.32 (♂), 0.38-0.40 (♀), 1.73-1.86 (♂), 1.78-1.87 (♀) times longer than broad; tibia 0.39-0.46 (♂), 0.54-0.55 (♀); chela 0.45-0.51 (♂), 0.56-0.60 (♀); movable finger 0.37-0.41 (♂), 0.46-0.49 (♀); hand length (without pedicel) 0.10-0.12 (♂), 0.12-0.14 (♀), width 0.12-0.14 (♂), 0.15-0.17 (♀), 0.79-0.92 (♂), 0.75-0.88 (♀) times longer than broad.</paragraph>
<subSubSection id="2902CD1B8FEEB72981AFE4DB20987F61" pageId="0" pageNumber="649" type="variation">
<paragraph id="8D08601B1AEF3DBD89585A017CFFF096" pageId="0" pageNumber="649">Variation.</paragraph>
<paragraph id="6C6A071D0BDA8783B26FE00114A2ABBA" pageId="0" pageNumber="649">Body length 1.70-1.91 (♂), 2.05-2.19 (♀); abdomen length 1.14-1.32 (♂), 1.37-1.46 (♀), width 0.98-1.07 (♂), 1.18-1.25 (♀) (without pleura), 1.02-1.18 (♂), 1.29-1.38 (♀) (with pleura), 1.17-1.28 (♂), 1.14-1.24 (♀) times longer than broad; carapace length 0.52-0.58 (♂), 0.63-0.67 (♀), width 0.36-0.39 (♂), 0.46-0.48 (♀), 1.43-1.49 (♂), 1.30-1.36 (♀) times longer than broad; Pedipalp: trochanter 0.22-0.25 (♂), 0.27-0.30 (♀); femur length 0.52-0.56 (♂), 0.70-0.71 (♀), width 0.29-0.32 (♂), 0.38-0.40 (♀), 1.73-1.86 (♂), 1.78-1.87 (♀) times longer than broad; tibia 0.39-0.46 (♂), 0.54-0.55 (♀); chela 0.45-0.51 (♂), 0.56-0.60 (♀); movable finger 0.37-0.41 (♂), 0.46-0.49 (♀); hand length (without pedicel) 0.10-0.12 (♂), 0.12-0.14 (♀), width 0.12-0.14 (♂), 0.15-0.17 (♀), 0.79-0.92 (♂), 0.75-0.88 (♀) times longer than broad.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="genbank code">
<paragraph pageId="0" pageNumber="649">GenBank Code.</paragraph>
<paragraph pageId="0" pageNumber="649">OP589965. The species differs from all congeners by more than 4.8% in the H3 dataset.</paragraph>
<subSubSection id="F96C2F43937AA35C2377B85CFBC7C365" pageId="0" pageNumber="649" type="genbank code">
<paragraph id="19235ADCAB44540F624B72D9691B950E" pageId="0" pageNumber="649">GenBank Code.</paragraph>
<paragraph id="6AFAC1F39B0C28A445DBE276AF6F3014" pageId="0" pageNumber="649">OP589965. The species differs from all congeners by more than 4.8% in the H3 dataset.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="habitat">
<paragraph pageId="0" pageNumber="649">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="649">Isolated tropical dry forest at a moderate elevation (100-500 m) located northwest of Ambarilao. The forest leads up to the mountain ranges of the Central Highlands. The specimens were found in sifted litter (leaf mold, rotten wood).</paragraph>
<subSubSection id="42906DC1AB5FD0277DF9DF479B5C4B27" pageId="0" pageNumber="649" type="habitat">
<paragraph id="41B1E32D89C4E53240A4BA9C84AD0A9B" pageId="0" pageNumber="649">Habitat.</paragraph>
<paragraph id="7D6804DA8377BC227D4AE2FDB6B2CC91" pageId="0" pageNumber="649">Isolated tropical dry forest at a moderate elevation (100-500 m) located northwest of Ambarilao. The forest leads up to the mountain ranges of the Central Highlands. The specimens were found in sifted litter (leaf mold, rotten wood).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="distribution">
<paragraph pageId="0" pageNumber="649">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="649">
<subSubSection id="BE5CB169025B94E7AFFB95CE4DFD2A1E" pageId="0" pageNumber="649" type="distribution">
<paragraph id="79F4C6E6A73EEFAF0F94A4B04D195EC6" pageId="0" pageNumber="649">Distribution.</paragraph>
<paragraph id="9701A08CF3A8A9FB5B4344F0753A51F2" pageId="0" pageNumber="649">
Only known from the type locality (BLF9974) in Sava Region (formerly eastern
<taxonomicName genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
<taxonomicName id="EDE2D865B4CCF7D7FCF8979162FA9047" genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
Province) of northern Madagascar.
</paragraph>
</subSubSection>