From e0bee75c1b0ee142e007ca97f7df5625ce203374 Mon Sep 17 00:00:00 2001 From: ggserver Date: Thu, 9 Jan 2025 18:01:24 +0000 Subject: [PATCH] Add updates up until 2025-01-09 17:54:28 --- .../87/03C687DDB92A2252FF38F8D3FB01C982.xml | 227 ++-- .../87/03C687DDB93D2241FF38FD70FA25CBC9.xml | 1117 +++++++++++++++++ 2 files changed, 1234 insertions(+), 110 deletions(-) create mode 100644 data/03/C6/87/03C687DDB93D2241FF38FD70FA25CBC9.xml diff --git a/data/03/C6/87/03C687DDB92A2252FF38F8D3FB01C982.xml b/data/03/C6/87/03C687DDB92A2252FF38F8D3FB01C982.xml index bf687ff6ca2..474e150da7c 100644 --- a/data/03/C6/87/03C687DDB92A2252FF38F8D3FB01C982.xml +++ b/data/03/C6/87/03C687DDB92A2252FF38F8D3FB01C982.xml @@ -1,99 +1,99 @@ - - - -Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama + + + +Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama - - -Author + + +Author -Williams, Kevin A. +Williams, Kevin A. - - -Author + + +Author -Cambra, Roberto. A. +Cambra, Roberto. A. - - -Author + + +Author -Bartholomay, Pedro R. -Instituto Nacional da Mata Atlântica, Santa Teresa, Espírito Santo, Brazil. +Bartholomay, Pedro R. +Instituto Nacional da Mata Atlântica, Santa Teresa, Espírito Santo, Brazil. - - -Author + + +Author -Lopez, Vinicius M. -Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. +Lopez, Vinicius M. +Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. - - -Author + + +Author -Brothers, Denis J. -School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa. +Brothers, Denis J. +School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa. - - -Author + + +Author -Martins, Herbeson O. J. -Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. +Martins, Herbeson O. J. +Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. - - -Author + + +Author -Ferreira, Vinina S. -Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. +Ferreira, Vinina S. +Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. - - -Author + + +Author -Koroiva, Ricardo -Instituto de Ciências Biológicas, Universidade Federal do Pará, Belém, PA, Brazil. +Koroiva, Ricardo +Instituto de Ciências Biológicas, Universidade Federal do Pará, Belém, PA, Brazil. - - -Author + + +Author -Guillermo-Ferreira, Rhainer -Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. +Guillermo-Ferreira, Rhainer +Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-14 + +2024 + +2024-11-14 - -5538 + +5538 - -2 + +2 - -127 -152 + +127 +152 - -https://doi.org/10.11646/zootaxa.5538.2.2 + +https://doi.org/10.11646/zootaxa.5538.2.2 -journal article -10.11646/zootaxa.5538.2.2 -1175-5326 -14611598 -AA00AF28-4FA3-45A1-91CD-7C636DACAE2B +journal article +10.11646/zootaxa.5538.2.2 +1175-5326 +14611598 +AA00AF28-4FA3-45A1-91CD-7C636DACAE2B @@ -204,9 +204,9 @@ in 2010 and was almost immediately recognized by KAW as a new species. In a phyl 5♀ 3♂ ] Type material. + Holotype , - ♀ , BRAZIL @@ -215,7 +215,9 @@ in 2010 and was almost immediately recognized by KAW as a new species. In a phyl Pernambuco , -Petrolina + +Petrolina + , 09°19’44.2”S 42°33’30”W @@ -223,44 +225,55 @@ in 2010 and was almost immediately recognized by KAW as a new species. In a phyl 17.V.2019 -, coleta manual, +, +coleta manual +, H.O.J. Martins -leg, MZSPHYM0136464 ( +leg, +MZSPHYM0136464 +( MZSP ; Fig. 18 -). -Paratypes +) +. + +Paratypes : - 1♀ -, same data as holotype except MZSPHYM0145123 ( -MZSP +, same data as holotype except +MZSPHYM0145123 +( +MZSP ) ; - + 1♂ , allotype -, same data as holotype except MZSPHYM0136463 ( -MZSP +, same data as holotype except +MZSPHYM0136463 +( +MZSP ) ; - + 1♂ -same data as holotype except MZSPHYM0145122 ( -MZSP +same data as holotype except +MZSPHYM0145122 +( +MZSP ) ; - + 1♀ , same data as holotype except -Hym -–002/0051, +Hym–002/0051 +, V.2018 @@ -269,11 +282,11 @@ same data as holotype except MZSPHYM0145122 ( ) ; - + 1♂ ,same data as holotype except -Hym -–002/0048, +Hym–002/0048 +, 24–28.III.2018 @@ -282,14 +295,15 @@ same data as holotype except MZSPHYM0145122 ( ) ; - + 1♀ , Ceará -, BR116, - +, + +BR116, 5 km SW Russas @@ -307,14 +321,16 @@ SW Russas , C. Waichert col. ( -UFES -; voucher JP1779). +UFES +; voucher +JP1779 +) + +. Other material . BRAZIL - : - 1♀ , @@ -334,10 +350,7 @@ col. ( DZUP #299378 ; Fig. 19 -) - -; - +); 1♀ , Riachão @@ -349,15 +362,12 @@ col. ( , Varrediura, F. -Limeira-de- -Oliveira +Limeira-de-Oliveira leg. ( INPA ; Fig. 20 -) - -. +). @@ -397,7 +407,6 @@ species. - The sex association is based on collection of males and females at the same locality on the same date in Pernambuco @@ -423,9 +432,7 @@ corroborate the identification of a single species, nov . -Additionally, morphological similarities between both sexes and related genera further validate the sex association (see remarks for the genus above) - -. +Additionally, morphological similarities between both sexes and related genera further validate the sex association (see remarks for the genus above). The female specimens from diff --git a/data/03/C6/87/03C687DDB93D2241FF38FD70FA25CBC9.xml b/data/03/C6/87/03C687DDB93D2241FF38FD70FA25CBC9.xml new file mode 100644 index 00000000000..2122cbff1e8 --- /dev/null +++ b/data/03/C6/87/03C687DDB93D2241FF38FD70FA25CBC9.xml @@ -0,0 +1,1117 @@ + + + +Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama + + + +Author + +Williams, Kevin A. + + + +Author + +Cambra, Roberto. A. + + + +Author + +Bartholomay, Pedro R. +Instituto Nacional da Mata Atlântica, Santa Teresa, Espírito Santo, Brazil. + + + +Author + +Lopez, Vinicius M. +Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. + + + +Author + +Brothers, Denis J. +School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa. + + + +Author + +Martins, Herbeson O. J. +Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. + + + +Author + +Ferreira, Vinina S. +Universidade Federal do Vale do São Francisco, Colegiado de Ciências Biológicas. Petrolina, PE, Brazil. + + + +Author + +Koroiva, Ricardo +Instituto de Ciências Biológicas, Universidade Federal do Pará, Belém, PA, Brazil. + + + +Author + +Guillermo-Ferreira, Rhainer +Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil. + +text + + +Zootaxa + + +2024 + +2024-11-14 + + +5538 + + +2 + + +127 +152 + + + + +https://doi.org/10.11646/zootaxa.5538.2.2 + +journal article +10.11646/zootaxa.5538.2.2 +1175-5326 +14611598 +AA00AF28-4FA3-45A1-91CD-7C636DACAE2B + + + + + + +Key to American genera of +Dasymutillini + + + + + + +The genera + +Lomachaeta + +, + +Neomutilla + +, and + +Protophotopsis + +were excluded from +Dasymutillini +( + +Waldren +et al +. 2023 + +) and are not keyed here. The key is written in a way that both sexes of velvet ants from the Americas can be checked for exclusion from +Dasymutillini +or recognized as one of the 13 genera of +Dasymutillini +. + + + + +Females + + + + + + +1 +. Eye subcircular; mandible unarmed ventrally; head unarmed ventrally; mesosoma widest at pronotum or mesonotum, in dorsal view; tergal fringes composed of simple or brachyplumose setae............................................... +2 + + + + +- With one or more of the following characters: eye longitudinally ovate; mandible with ventral tooth basally; gena and/or hypostoma armed with distinct ventral tooth; propodeum and pronotum both wider than mesonotum, in dorsal view; tergal fringes composed of distinctly plumose setae........................................ +other tribes +(not treated here) + + + + + + +2 +(1). T6 convex without defined pygidial plate.................................................................. +3 + + + + +- T6 with flat pygidial plate, usually with differentiated sculpture and defined by lateral carinae........................ +5 + + + + + + +3 +(2). T1 shape narrowly petiolate, constricted with base of T2, T1 generally widest mesally............................... +4 + + + + +- T1 shape generally sessile, subsessile, or disciform, width variable, T1 generally widest at posterior margin......................................................................................... +other tribes +(not treated here) + + + + + + +4 +(3). Genal carina and scutellar scale absent; mesosoma convex dorsally; T6 without longitudinal carinae; epaulet scarcely raised ( +Panama +).............................................. +Dasyphuta + +gen +. +nov +. + +—one species: + +D. gorgupea + + +sp +. +nov +. + + + + + +- Genal carina and scutellar scale present; mesosoma flattened dorsally; T6 with two wavy longitudinal carinae mesally; epaulet usually raised onto smooth tubercle ( +Panama +).............................................................................................. + +Frigitilla +Williams + +in + +Bartholomay +et al. +2015 + +—two species, see + +Cambra +et al. +(2016) + + + + + + + +5 +(2). Pygidium with lateral carinae, with acute erect tooth at posterolateral angle (mesosomal pleura uniformly covered with dense setae; tibial spurs black)................................................................................ +6 + + + + +- Pygidium with lateral carinae but without erect tooth at posterolateral angle...................................... +7 + + + + + + +6 +(5). Second labial palpomere mesally expanded, at least twice as wide as long; metasternal process short and truncated, with medial notch of variable depth; mesosomal dorsum with one or two narrow longitudinal stripes of dense whitish setae extending from anterior margin of pronotum to posterior margin of propodeum.............................. + +Suareztilla +Casal, 1968 + + + + + +- Second labial palpomere not expanded, at least as long as wide; metasternal process extended mesally into triangular plate between metathoracic coxae; mesosomal dorsum without narrow longitudinal stripes of whitish setae................................................................................................ + +Reedomutilla +Mickel, 1964 + + + + + + + +7 +(5). Mesosoma broadly violin-shaped, propodeal sides divergent behind propodeal spiracle in dorsal view.................. +8 + + + + +- Mesosoma pear-shaped, propodeal sides convergent behind propodeal spiracle in dorsal view........................ +9 + + + + + + +8 +(7). Body uniformly pale yellow-brown; mesosoma with small scutellar scale; T2 disc with basomesal triangular patch of flattened striate setae ( +Argentina +)................ + +Gogoltilla + +Williams +et al. +2011 + + +—One species, + +G. chichikovi + +Williams +et al. +2011 + + + + + + +- With one or more of the following characters: body color partly or entirely blackish; mesosoma without scutellar scale; T2 disc without patch of striated setae.................................................... +other tribes +(not treated here) + + + + + + +9 +(7). Dorsal and posterior surfaces of vertex separated by sharp arcuate transverse carina ( +Brazil +)............................................................................ +Goncharovtilla + +gen +. +nov +. + +—one species, + +G. oblomovi + + +sp +. +nov +. + + + + + +- Posterior margin of vertex rounded or with separated posterolateral tubercles..................................... +10 + + + + + + +10 +(9). Scutellar scale bilobate; mesonotum armed with four or five dorsolateral tubercles; hypopygium apex with four spines ( +Bolivia +, +Brazil +, and +Paraguay +).................... + +Leucospilomutilla +Ashmead, 1903 + +—three species, see + +Williams +et al +. (2020) + + + + + +- Scutellar scale absent or unilobate; mesonotum usually unarmed laterally, rarely with one spine; hypopygium apex lacking spines............................................................................................. +11 + + + + + + +11 +(10). Head broad and quadrate, vertex width greater than mesosomal width (T2 with large yellow or red integumental spots, paired, with marked edges; widespread in South America).... + +Cephalomutilla +André, 1908 + +—four species, see + +Williams +et al +. (2022) + + + + + +- Head not so broad, usually rounded posteriorly, vertex width usually less than mesosomal width..................... +12 + + + + + + +12 +(11). Specimens from North and Central America or Caribbean Islands (excluding +Trinidad +)............................ +13 + + + + +- Specimens from South America (including +Trinidad +)........................................................ +14 + + + + + + +13 +(12). Body entirely black, except T2 with two or four yellow cuticular spots, these spots mostly bare, with sparse punctures; tibial spurs whitish (rarely encountered, only in Central America and Hispaniola)........................................................................... + +Traumatomutilla +André, 1901 + +, +in part— +three species, see + +Williams +et al. +(2017) + + + + + +- With one or more of the following characters: mesosoma reddish-brown; T2 disc without yellow cuticular spots or with spots having similar sculpture to surrounding T2 disc sculpture; tibial spurs black (widespread and common, in North and Central America and Caribbean Islands)................................ + +Dasymutilla +Ashmead, 1899 + +, +in part— +over 100 species, see +Manley & Pitts (2007) +, + +Cambra +et al +. (2018) + +, and + +Manley +et al +. (2020) + + + + + + + +14 +(12). T2 disc cuticle uniformly blackish, lighter markings formed by setae only (genal carina absent)..................... +15 + + + + +- T2 disc blackish and marked with yellow or reddish cuticular patches, or T2 disc cuticle lighter reddish or yellowish brown. .................................................................................................. +17 + + + + + + +15 +(14). Setal markings on T2 composed of golden circular spots (Brazilian Atlantic Forest) or T2 mostly covered with dense yellow to orange setae (countries West of Andes)................................................................. +16 + + + + +- Setal markings on T2 composed of small whitish setal stripes or spots ( +Colombia +or Brazilian Caatinga and Cerrado).... +18 + + + + + + +16 +(15). Vertex smooth, without tubercles or projections of any kind; mesonotum and T2 disc with large patches of dense yellow to orange setae ( +Chile +, +Ecuador +, and +Peru +)........................................................................................ + +Quwitilla +Williams +et al. + +in Bartholomay +et al +. 2019—three species, see + +Bartholomay +et al +. (2019b) + + + + + +- Vertex with large smooth tubercle posterolaterally; mesonotum with contrasting golden and black setae, T2 disc marked with large golden-orange spots composed of thickened bristle-like setae ( +Brazil +)...................................................... + +Atlantilla +Williams & Bartholomay + +in + +Williams +et al +. 2020 + +—one species, + +A. auriculata +( +Gerstaecker, 1874 +) + + + + + + + +17 +(14). Mesosoma pale reddish-brown; scutellar scale distinct and broad; vertex simple without tubercles or projections of any kind; genal carina present; lateral propodeal surface smooth and defined by irregular carina dorsally and posteriorly; body small, length +5 mm +or less (widespread in South America)........ + +Tobantilla +Casal, 1969 + +—nine species, see + +Williams +et al +. (2011) + + + + + +- With one or more of the following characters: mesosoma black; scutellar scale indistinct or absent; vertex armed with posterolateral tubercle; genal carina absent; lateral propodeal surface distinctly sculptured and not defined by carina dorsally and posteriorly; body length more than +5 mm +............................................................... +8 + + + + + + +18 +(15, 17). T2 disc with spots composed of setae only or with cuticular spots having similar sculpture to surrounding T2 disc sculpture (rarely encountered, in +Colombia +only, mesosoma sometimes reddish orange).................................................. + +Dasymutilla +Ashmead, 1899 + +, +in part +—four species, see + +Cambra +et al +. (2018) + +, and + +Bartholomay +et al +. (2019b) + + + + + +- T2 disc with smooth yellow or reddish cuticular spots with sparse setae (common and widespread throughout South America; in +Colombia +with mesosoma blackish)....... + +Traumatomutilla +André, 1901 + +—over 100 species, see + +Williams +et al +. (2017) + + + + + + + +Males + + + + + + +1 +. Axilla armed with posteriorly directed tooth or lobe; eye subcircular; head unarmed ventrally; T1 shape narrowly petiolate and usually constricted with base of T2; tergal fringes composed of simple or brachyplumose setae ........................ +2 + + + + +- With one or more of the following characters: axilla unarmed; eye longitudinally ovate and notched; gena and/or hypostoma armed with distinct ventral tooth; T1 broadly sessile, subsessile, or disciform; tergal fringes composed of distinctly plumose setae........................................................................ +other tribes +(not treated here) + + + + + + +2 +(1). Mandible with ventral tooth or lobe basally................................................................ +3 + + + + +- Mandible unarmed ventrally............................................................................ +4 + + + + + + +3 +(2). Mandible with bilobate lamella dorsally; paramere posteriorly upcurved with apex pointing antero-dorsally ( +Argentina +)....................................... + +Gogoltilla + +Williams +et al. +2011 + + +— +One species, + +G. chichikovi + +Williams +et al. +2011 + + + + + + +- Mandible simple dorsally; paramere not upcurved, apex pointing posteriorly.............. +other tribes +(not treated here) + + + + + + +4 +(2). Specimens from North America, Central America, or Caribbean Islands (excluding +Trinidad +)......................... +5 + + + + +- Specimens from South America (including +Trinidad +)......................................................... +7 + + + + + + +5 +(4). Nearly all erect setae on body, particularly on scape and clypeus, brachyplumose; nearly entire body covered with moderately dense whitish setae; paramere posteriorly upcurved with apex pointing dorsally ( +Panama +)................................................................................. +Dasyphuta + +gen +. +nov +. + +—one species: + +D. gorgupea + + +sp +. +nov +. + + + + +- Erect setae on body mostly simple; body setal color variable; paramere shallowly upcurved, apex pointing postero-dorsally. 6 + + + + + +6 +(5). Body cuticle entirely black, covered only with black and silvery setae; tibial spurs whitish; free paramere length with relatively sparse setae anteroventrally (rarely encountered in Central America and Hispaniola)......................................................................................... + +Traumatomutilla +André, 1901 + +, +in part— +three species + + + + +- With one or more of the following characters: at least mesosoma or T2 with cuticle yellowish or reddish-brown; head, mesosoma, or tergites densely covered with golden to reddish setae; tibial spurs blackish; free paramere length with particularly dense brush of pale yellowish setae anteroventrally (common and widespread in North and Central America and Caribbean Islands)................................................................. + +Dasymutilla +Ashmead, 1899 + +, +in part— +over 100 species +— +see +Manley & Pitts (2007) +, + +Cambra +et al +. (2018) + +, and + +Manley +et al +. (2020) + +[The male of + +Frigitilla panamensis + +Cambra +et al. +2016 + + +is not yet known, but will likely key out here with similar characters to those mentioned in couplet 9 below] + + + + + + +7 +(4). Mandible thick and somewhat vertically oriented (with distinct dorsal carina continuous to inner apical tooth); parapenial lobe longer than penis valve ( +Chile +, +Ecuador +, and +Peru +)................................................................ + +Quwitilla +Williams, Bartholomay & Cambra + +in Bartholomay +et al +. 2019—three species, see + +Bartholomay +et al +. (2019b) + + + + + +- Mandible narrowed and oblique apically (dorsal carina usually becoming obsolete before reaching inner apical tooth); parapenial lobe shorter than penis valve (widespread but rarely found West of the Andes).................................... +8 + + + + + + +8 +(7). Paramere shallowly upcurved posteriorly, apex pointing postero-dorsally (brachyplumose setae generally rare and inconspicuous on body; head usually uniformly black; clypeus usually densely punctate and bidentate antero-ventrally)............... +9 + + + + +- Paramere sharply upcurved posteriorly, apex pointing dorsally or antero-dorsally (brachyplumose setae generally abundant and conspicuous, especially on scape and clypeus; vertex often with mesal reddish cuticular spot; clypeus usually with smooth bidentate or truncate shelf antero-ventrally, antero-ventrally unidentate, or antero-ventrally rounded).................. +14 + + + + + + +9 +(8). S2 mesally convexly swollen, with asetose mesal pit (T2 and S2 with cuticle largely orange-brown; paramere virtually straight, slightly upcurved posteriorly, with especially dense ventral setal brush on basal half; +Bolivia +, +Brazil +, and +Peru +)............................................ + +Frigitilla +Williams + +in + +Bartholomay +et al. +2015 + +, +in part +— + +F. simulatrix +( +Smith, 1879 +) + + + + + +- S2 flat or simply convex, if mesal pit present, then filled with setae............................................. +10 + + + + + + +10 +(9). Axillar lobe with posterior margin flat and truncate posteriorly, additionally armed with sharp down-pointed tooth along outer margin ( +Brazil +).................................................................................................... + +Atlantilla +Williams & Bartholomay + +in + +Williams +et al +., 2020 + +—one species, + +A. auriculata +( +Gerstaecker, 1874 +) + + + + + +- Axillar lobe variable but without down-pointing tooth along outer margin....................................... +11 + + + + + + +11 +(10). Mesopleuron with lateral tooth-like tubercle; axilla posteriorly truncate and raised above level of depressed mesoscutellum; propodeum with conspicuous dorsolateral tubercle posterior to propodeal spiracle; penis valve with especially large pre-apical tooth, margin anterior to pre-apical tooth serrate ventrally ( +Bolivia +, +Brazil +, and +Paraguay +)................................................................... + +Leucospilomutilla +Ashmead, 1903 + +—three species, see + +Williams +et al +. (2020) + + + + + +- Mesopleuron usually without tooth-like tubercle (if mesopleuron armed, then axilla dentate posteriorly or not especially raised above convex mesoscutellum); propodeum without dorsolateral tubercle posterior to propodeal spiracle; penis valve with smooth margin anterior to pre-apical ventral tooth.......................................................... +12 + + + + + + +12 +(11). S2 mesally flattened, without mesal pit; paramere dorsoventrally flattened basally (widespread in South America).................................................... + +Cephalomutilla +André, 1908 + +—four species, see + +Williams +et al +. 2022 + + + + + +- S2 convex (with or without mesal pit); paramere shape variable............................................... +13 + + + + + + +13 +(12). Rarely encountered, in +Colombia +only; with metasomal cuticle mostly reddish-orange and covered with dense red setae or metasoma predominantly black except T2 entirely pale yellow or marked with pale yellow cuticular patch..... + +Dasymutilla +Ashmead, 1899 + +, +in part +—four species, see +Manley & Pitts (2007) +, + +Cambra +et al +. (2018) + +, and + +Bartholomay +et al +. (2019b) + + + + + +- Common and widespread throughout South America; Colombian species with metasomal cuticle uniformly black or mostly reddish-orange with sparse setae.................................. + +Traumatomutilla +André, 1901 + +—over 100 species + + + + + + +14 +(8). Vertex black with reddish-orange spot mesally; clypeus, frons, and gena usually with extensive black setae (if head setae predominantly whitish, then T2 marked with round yellow cuticular spots)...................................... +15 + + + + +- Vertex uniformly blackish; clypeus, frons, and gena usually with silvery setae, at most interspersed with a few erect darker setae; T2 disc entirely black, reddish-black or orange (second labial palpomere not expanded, at least as long as wide).... +16 + + + + + + +15 +(14). T2 disc with cuticle uniformly black; second labial palpomere not expanded, at least as long as wide...................................................................... + +Reedomutilla +Mickel, 1964 + +—three species, see +Casal (1968) + + + + +- T2 disc black, marked with two large circular yellow or reddish cuticular spots; second labial palpomere mesally expanded, at least twice as wide as long.................................. + +Suareztilla +Casal, 1968 + +—Nine species, see +Casal (1968) + + + + + + +16 +(14). Clypeus with longitudinal elevated crest medially, with anteromedial margin strongly projected medially ( +Fig. 8 +); body cuticle entirely black (Caatinga region of +Brazil +)................. +Goncharovtilla + +gen +. +nov +. + +—one species, + +G. oblomovi + + +sp +. +nov +. + + + + + +- Clypeus without longitudinal elevated crest medially, with anteromedial margin projected in a single raised tubercle or shallowly emarginate (see + +Figs +35–36 + +in + +Williams +et al. +2011 + +); metasomal cuticle largely reddish-orange, at least on T2 (widespread in South America, but absent from Caatinga)............... + +Tobantilla +Casal, 1969 + +—nine species, see + +Williams +et al +. (2011) + + + + + + + \ No newline at end of file