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<mods:title id="444296857752BFEE732D7E8BB43708EF">A bee from the middle Miocene Hindon Maar of southern New Zealand (Hymenoptera, Colletidae)</mods:title>
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<mods:title id="8D5C198C46B8F67DBB7ABA5F74609A1E">A bee from the middle Miocene Hindon Maar of southern New Zealand (Hymenoptera, Colletidae)</mods:title>
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<mods:namePart id="68935653E395BFCC693A42FA218F81F2">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="4E87568541154FD5629FDCA06CA1581C">Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, New York 10024 - 5192 (United States); mengel @ amnh. org</mods:affiliation>
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<mods:namePart id="2AFB41D2909F34DEB33F009073FC5FD3">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="C40065A7B288910B74CA37F49DADBCB0">Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, New York 10024 - 5192 (United States); mengel @ amnh. org</mods:affiliation>
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<mods:namePart id="F0FBF383AB1D638865AE44C59332D105">Kaulfuss, Uwe</mods:namePart>
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<mods:affiliation id="8C0C156E9F045A73AFCE63B64241AFC9">Georg-August-Universität GÖttingen, Johann-Friedrich-Blumenbach Institut für Zoologie und Anthropologie, Evolution und Biodiversität der Tiere, Untere Karspüle 2, 37073 GÖttingen (Germany) uwe. kaulfuss @ uni-goettingen. de</mods:affiliation>
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<mods:namePart id="9B121661399615BD0D8FAEDDB8D1640F">Kaulfuss, Uwe</mods:namePart>
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<mods:affiliation id="8D41C1AADB2B5709D3B353CE3AD87C5D">Georg-August-Universität GÖttingen, Johann-Friedrich-Blumenbach Institut für Zoologie und Anthropologie, Evolution und Biodiversität der Tiere, Untere Karspüle 2, 37073 GÖttingen (Germany) uwe. kaulfuss @ uni-goettingen. de</mods:affiliation>
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<mods:date id="232CDD55AA3F10C5936D611696847F94">2025</mods:date>
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<mods:number id="69AD59F1C795831A24F772C41158902F">2025-01-29</mods:number>
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<treatment id="03DCAE031410B13BFCF4F9C8FC43FBFB" ID-DOI="http://doi.org/10.5281/zenodo.14889262" ID-Zenodo-Dep="14889262" LSID="urn:lsid:plazi:treatment:03DCAE031410B13BFCF4F9C8FC43FBFB" httpUri="http://treatment.plazi.org/id/03DCAE031410B13BFCF4F9C8FC43FBFB" lastPageId="6" lastPageNumber="47" pageId="4" pageNumber="45" scope_family="Colletidae" scope_order="Hymenoptera" scope_subGenus="Otagocolletes">
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<subSubSection id="C36F4C9E1410B139FCF4F9C8FA92F938" box="[873,1395,1650,1677]" pageId="4" pageNumber="45" type="nomenclature">
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<paragraph id="8BCA1F151410B139FCF4F9C8FA92F938" blockId="4.[873,1395,1650,1708]" box="[873,1395,1650,1677]" pageId="4" pageNumber="45">
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@ -66,7 +69,7 @@
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<subSubSection id="C36F4C9E1410B139FBA4F928FA73F95F" pageId="4" pageNumber="45" type="description">
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<paragraph id="8BCA1F151410B139FBA4F928FB43F919" blockId="4.[873,1395,1650,1708]" box="[1081,1186,1682,1708]" pageId="4" pageNumber="45">
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(
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<figureCitation id="134E03901410B139FBDCF928FB7BF919" box="[1089,1178,1682,1708]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[280,291,1404,1422]" captionStartId-1="5.[299,309,1438,1456]" captionStartId-2="6.[150,161,833,851]" captionTargetBox-0="[132,1455,93,1361]" captionTargetBox-1="[132,1455,216,1387]" captionTargetBox-2="[174,1400,268,758]" captionTargetId-0="figure-280@4.[132,1455,215,1361]" captionTargetId-1="figure-293@5.[132,1455,216,1387]" captionTargetId-2="figure-620@6.[132,1455,216,794]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIG. 2. — Leioproctus (Otagocolletes) barrydonovani n. subgen., n. sp. holotype OU46559, habitus of part. Scale bar: 1 mm." captionText-1="FIG. 3. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, habitus of counterpart. Scale bar: 1 mm." captionText-2="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." pageId="4" pageNumber="45">Figs 2-4</figureCitation>
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<figureCitation id="134E03901410B139FBDCF928FB7BF919" box="[1089,1178,1682,1708]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[280,291,1404,1422]" captionStartId-1="5.[299,309,1438,1456]" captionStartId-2="6.[150,161,833,851]" captionTargetBox-0="[132,1455,93,1361]" captionTargetBox-1="[132,1455,216,1387]" captionTargetBox-2="[174,1400,268,758]" captionTargetId-0="figure-280@4.[132,1455,215,1361]" captionTargetId-1="figure-293@5.[132,1455,216,1387]" captionTargetId-2="figure-620@6.[132,1455,216,794]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIG. 2. — Leioproctus (Otagocolletes) barrydonovani n. subgen., n. sp. holotype OU46559, habitus of part. Scale bar: 1 mm." captionText-1="FIG. 3. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, habitus of counterpart. Scale bar: 1 mm." captionText-2="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14889243" figureDoi-1="http://doi.org/10.5281/zenodo.14889245" figureDoi-2="http://doi.org/10.5281/zenodo.14889247" httpUri-0="https://zenodo.org/record/14889243/files/figure.png" httpUri-1="https://zenodo.org/record/14889245/files/figure.png" httpUri-2="https://zenodo.org/record/14889247/files/figure.png" pageId="4" pageNumber="45">Figs 2-4</figureCitation>
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)
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</paragraph>
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<paragraph id="8BCA1F151410B139FCD7F962FA73F95F" blockId="4.[842,1426,1752,1770]" box="[842,1426,1752,1770]" pageId="4" pageNumber="45">
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@ -128,7 +131,7 @@ N of Outram,
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.
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</paragraph>
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</subSubSection>
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<caption id="DF0A4F9D1411B138FEB6FA24FAEDFA05" ID-DOI="http://doi.org/10.5281/zenodo.14889245" ID-Zenodo-Dep="14889245" box="[299,1292,1437,1456]" httpUri="https://zenodo.org/record/14889245/files/figure.png" pageId="5" pageNumber="46" startId="5.[299,309,1438,1456]" targetBox="[132,1455,216,1387]" targetPageId="5" targetType="figure">
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<paragraph id="8BCA1F151411B138FEB6FA24FAEDFA05" blockId="5.[299,1292,1437,1456]" box="[299,1292,1437,1456]" pageId="5" pageNumber="46">
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. 3. —
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@ -186,7 +189,7 @@ Total length as preserved (vertex to apex of fragment of tergum II)
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<emphasis id="B901C3071411B138FCB0F8A8FC91F899" bold="true" box="[813,880,1810,1836]" pageId="5" pageNumber="46">Head.</emphasis>
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Apparently broad (incomplete and in oblique posterior position so precise dimension impossible to determine but seeming broad based on upper width as preserved); gena narrower than compound eye; malar space apparently about as long as wide (the sclerite at the lower margin of the compound eye is not the base of the mandible and indicates a malar space that is 0.5× as long as wide, or even slightly more; this seems almost certainly the case in the part but is less clear in the counterpart as preserved); mesoscutellum longer than combined lengths of metanotum and basal area of propodeum; metanotum exceedingly short (as preserved), slightly shorter than basal area of propodeum; basal area of propodeum wholly declivitous (best observed on part), almost undifferentiated from posterior surface.
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</paragraph>
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<smallCapsWord id="8D2C89C91412B13BFF0BFCFBFF4FFCE7" baselines="846,847" box="[150,174,833,851]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="47">FIG</smallCapsWord>
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||||
. 4. —
|
||||
|
|
@ -208,11 +211,11 @@ holotype OU46559, line drawings of left (
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|||
<paragraph id="8BCA1F151412B13BFF19FB2EFAD2FC5A" blockId="6.[130,777,1172,2028]" lastBlockId="6.[813,1455,917,1007]" pageId="6" pageNumber="47">
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<emphasis id="B901C3071412B13BFF19FB2EFF1BFB1B" bold="true" box="[132,250,1172,1198]" pageId="6" pageNumber="47">Forewing.</emphasis>
|
||||
All wing veins strong and tubular; marginal cell long (although incomplete as preserved the course of Rs and anterior wing margin demonstrate overall length) (
|
||||
<figureCitation id="134E03901412B13BFF10FB49FF09FABB" box="[141,232,1267,1294]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[280,291,1404,1422]" captionStartId-1="5.[299,309,1438,1456]" captionStartId-2="6.[150,161,833,851]" captionTargetBox-0="[132,1455,93,1361]" captionTargetBox-1="[132,1455,216,1387]" captionTargetBox-2="[174,1400,268,758]" captionTargetId-0="figure-280@4.[132,1455,215,1361]" captionTargetId-1="figure-293@5.[132,1455,216,1387]" captionTargetId-2="figure-620@6.[132,1455,216,794]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIG. 2. — Leioproctus (Otagocolletes) barrydonovani n. subgen., n. sp. holotype OU46559, habitus of part. Scale bar: 1 mm." captionText-1="FIG. 3. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, habitus of counterpart. Scale bar: 1 mm." captionText-2="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." pageId="6" pageNumber="47">Figs 2-4</figureCitation>
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<figureCitation id="134E03901412B13BFF10FB49FF09FABB" box="[141,232,1267,1294]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[280,291,1404,1422]" captionStartId-1="5.[299,309,1438,1456]" captionStartId-2="6.[150,161,833,851]" captionTargetBox-0="[132,1455,93,1361]" captionTargetBox-1="[132,1455,216,1387]" captionTargetBox-2="[174,1400,268,758]" captionTargetId-0="figure-280@4.[132,1455,215,1361]" captionTargetId-1="figure-293@5.[132,1455,216,1387]" captionTargetId-2="figure-620@6.[132,1455,216,794]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIG. 2. — Leioproctus (Otagocolletes) barrydonovani n. subgen., n. sp. holotype OU46559, habitus of part. Scale bar: 1 mm." captionText-1="FIG. 3. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, habitus of counterpart. Scale bar: 1 mm." captionText-2="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14889243" figureDoi-1="http://doi.org/10.5281/zenodo.14889245" figureDoi-2="http://doi.org/10.5281/zenodo.14889247" httpUri-0="https://zenodo.org/record/14889243/files/figure.png" httpUri-1="https://zenodo.org/record/14889245/files/figure.png" httpUri-2="https://zenodo.org/record/14889247/files/figure.png" pageId="6" pageNumber="47">Figs 2-4</figureCitation>
|
||||
), longer than pterostigma; prestigma about as long as wide and perhaps about as long as pterostigmal base; pterostigma slender, roughly parallel-sided, not widening apically (
|
||||
<figureCitation id="134E03901412B13BFE8EFAE9FEB6FAD8" box="[275,343,1363,1389]" captionStart="FIG" captionStartId="6.[150,161,833,851]" captionTargetBox="[174,1400,268,758]" captionTargetId="figure-620@6.[132,1455,216,794]" captionTargetPageId="6" captionText="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." pageId="6" pageNumber="47">Fig. 4</figureCitation>
|
||||
<figureCitation id="134E03901412B13BFE8EFAE9FEB6FAD8" box="[275,343,1363,1389]" captionStart="FIG" captionStartId="6.[150,161,833,851]" captionTargetBox="[174,1400,268,758]" captionTargetId="figure-620@6.[132,1455,216,794]" captionTargetPageId="6" captionText="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14889247" httpUri="https://zenodo.org/record/14889247/files/figure.png" pageId="6" pageNumber="47">Fig. 4</figureCitation>
|
||||
), tapering within marginal cell, border within marginal cell distinctly convex (
|
||||
<figureCitation id="134E03901412B13BFDA3FAC9FD62FA38" box="[574,643,1395,1421]" captionStart="FIG" captionStartId="6.[150,161,833,851]" captionTargetBox="[174,1400,268,758]" captionTargetId="figure-620@6.[132,1455,216,794]" captionTargetPageId="6" captionText="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." pageId="6" pageNumber="47">Fig. 4</figureCitation>
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||||
<figureCitation id="134E03901412B13BFDA3FAC9FD62FA38" box="[574,643,1395,1421]" captionStart="FIG" captionStartId="6.[150,161,833,851]" captionTargetBox="[174,1400,268,758]" captionTargetId="figure-620@6.[132,1455,216,794]" captionTargetPageId="6" captionText="FIG. 4. — Leioproctus (Otagocolletes) barrydonovani n. sp. holotype OU46559, line drawings of left (A) and right (B) forewing as preserved. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14889247" httpUri="https://zenodo.org/record/14889247/files/figure.png" pageId="6" pageNumber="47">Fig. 4</figureCitation>
|
||||
); basal vein (1M) faintly arched, confluent with 1cu-a; 1cu-a straight, oblique, sloping toward wing apex; 1Rs slightly longer than prestigma, oblique relative to Rs+M, much shorter than 1 M; Rs+M nearly straight; first submarginal cell longer than individual lengths of second and third submarginal cells, slightly longer than combined lengths of remaining submarginal cells; prestigmal length of first submarginal cell slightly less than 0.3× pterostigmal length of cell; 1r-rs well distad pterostigmal midlength, near pterostigmal apex, shorter than 3Rs; second submarginal cell slightly narrowed anteriorly, 2Rs longer than 3Rs, 2Rs and 1rs-m not parallel; 1m-cu entering second submarginal cell at cell midlength, basad 1rs-m by 4-5× vein width; 3Rs shorter than 4Rs; anterior border of third submarginal cell longer than anterior border of second submarginal cell; 2rs-m weakly curved distally; posterior border of third submarginal cell about as long as that of second submarginal cell, only slightly longer than anterior border of third submarginal cell; 2m-cu apparently entering third submarginal cell near apex (incomplete but given that majority of posterior border of cell present and not in contact with 2m-cu indicates it must make contact near cell’s apex); 2Cu much longer than 2cu-a.
|
||||
</paragraph>
|
||||
<paragraph id="8BCA1F151412B13BFCB0FBAEFC83FB9B" blockId="6.[813,930,1044,1102]" box="[813,866,1044,1070]" pageId="6" pageNumber="47">
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|
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|||
|
|
@ -1,47 +1,49 @@
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|||
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<treatment id="C50387CCD8026277C697F522FEE413E8" LSID="urn:lsid:plazi:treatment:C50387CCD8026277C697F522FEE413E8" httpUri="http://treatment.plazi.org/id/C50387CCD8026277C697F522FEE413E8" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
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<treatment id="C50387CCD8026277C697F522FEE413E8" ID-DOI="http://doi.org/10.5281/zenodo.14889264" ID-Zenodo-Dep="14889264" LSID="urn:lsid:plazi:treatment:C50387CCD8026277C697F522FEE413E8" httpUri="http://treatment.plazi.org/id/C50387CCD8026277C697F522FEE413E8" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103" scope_family="Celastraceae">
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<subSubSection id="05B06551D8026276C697F522FD941336" box="[206,554,279,305]" pageId="2" pageNumber="103" type="nomenclature">
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<paragraph id="4D1536DAD8026276C697F522FD941336" blockId="2.[206,554,279,337]" box="[206,554,279,305]" pageId="2" pageNumber="103">
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|
|
@ -57,7 +59,7 @@ Jongkind
|
|||
<subSubSection id="05B06551D8026276C70DF502FE1A1356" box="[340,420,311,337]" pageId="2" pageNumber="103" type="description">
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<paragraph id="4D1536DAD8026276C70DF502FE1A1356" blockId="2.[206,554,279,337]" box="[340,420,311,337]" pageId="2" pageNumber="103">
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(
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||||
<figureCitation id="D5912A5FD8026276C705F502FE271356" box="[348,409,311,337]" captionStart="FIG" captionStartId="1.[113,124,1174,1191]" captionTargetBox="[123,1191,208,1152]" captionTargetId="figure-20@1.[133,1191,219,1137]" captionText="FIG. 1. — Salacia hallei Jongkind: A, habit; B, flower; C, calyx; D, flower bud; E, cross-section of disk and ovary.Breteler & Leal 14170. Scale bars: A, 1 cm; B-D, 2 mm; E, 1 mm." pageId="2" pageNumber="103">Fig. 1</figureCitation>
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||||
<figureCitation id="D5912A5FD8026276C705F502FE271356" box="[348,409,311,337]" captionStart="FIG" captionStartId="1.[113,124,1174,1191]" captionTargetBox="[123,1191,208,1152]" captionTargetId="figure-20@1.[133,1191,219,1137]" captionText="FIG. 1. — Salacia hallei Jongkind: A, habit; B, flower; C, calyx; D, flower bud; E, cross-section of disk and ovary.Breteler & Leal 14170. Scale bars: A, 1 cm; B-D, 2 mm; E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14889251" httpUri="https://zenodo.org/record/14889251/files/figure.png" pageId="2" pageNumber="103">Fig. 1</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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Reference in a new issue