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<document id="E174EAECF59CE2264516628E963C6B53" ID-DOI="10.3897/zookeys.1217.129707" ID-ZooBank="541D9AC9-DD07-4435-ACA2-867C5F302D34" ID-publisher-id="129707" URI-arpha="9247C138-7213-58ED-9D52-A66F2E597D9C" URI-zoobank="http://zoobank.org/541D9AC9-DD07-4435-ACA2-867C5F302D34" article-type="research-article" checkinTime="1730829902517" checkinUser="pensoft" docAuthor="Sittichaya, Wisut, Lin, Ching-Shan, Smith, Sarah M., Pornsuriya, Chaninan &amp; Cognato, Anthony I." docDate="2024" docId="DBA136A0B71D5D539C00E4F804A32E70" docLanguage="en" docName="ZooKeys 1217: 247-262" docOrigin="ZooKeys 1217" docSource="https://zookeys.pensoft.net/article/129707/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eccoptopterus formosanus Lin, Sittichaya &amp; Smith 2024, sp. nov." docType="treatment" docUuid="DFC6F8B0-22B2-4983-B744-411097C5F087" docUuidSource="ZooBank" docVersion="1" dtd-version="3.0" lastPageNumber="262" masterDocId="9247C138721358ED9D52A66F2E597D9C" masterDocTitle="Two new species of Eccoptopterus Motschulsky, 1863 ambrosia beetle from Taiwan and Thailand (Coleoptera, Curculionidae, Scolytinae, Xyleborini)" masterLastPageNumber="262" masterPageNumber="247" pageNumber="247" updateTime="1730829973222" updateUser="pensoft">
<mods:mods id="15E55D49F3E28BD0BC5186B9EC2F62F7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E990F427F90CB6023319E6C5C8413882">
<mods:title id="5BEDC2C980513547D6A410ADE22FCC98">Two new species of Eccoptopterus Motschulsky, 1863 ambrosia beetle from Taiwan and Thailand (Coleoptera, Curculionidae, Scolytinae, Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name id="68E391A8D952D280449EE357D36984D6" type="personal">
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<mods:namePart id="CD2374B9AD3BF70E321C8BEED8EBB22C">Sittichaya, Wisut</mods:namePart>
<mods:nameIdentifier id="10BDBD36D9CFF67C3DE870CDB346074C" type="ORCID">0000-0001-6200-1285</mods:nameIdentifier>
<mods:affiliation id="F7107D06E76EE19226C05B9F7998A569">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
</mods:name>
<mods:name id="6134DFEBDDD5C89A36A0E70302E47510" type="personal">
<mods:role id="389A58B85FBBD55C2408A86022928D35">
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</mods:role>
<mods:namePart id="53927A5E76F25F6D87D506396A386317">Lin, Ching-Shan</mods:namePart>
<mods:nameIdentifier id="B72B48DD73A668AADE47F10928A86BF3" type="ORCID">https://orcid.org/0009-0006-3159-697X</mods:nameIdentifier>
<mods:affiliation id="8FA89D6E36CAC038A42C5EF3BE76D9F1">Department of Entomology, National Taiwan University, Taipei 10617, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="F489299C80D5813DF1F408B6836A2377" type="personal">
<mods:role id="552A25241F053DA877D7FCD6BA254C99">
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</mods:role>
<mods:namePart id="2E50205ACD544BD222EA06ACCC54C958">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="1DA9D02009D5010BC0044A8E1A78831D" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="FC253B6894DC7D46B40A460E96983B4F">Department of Entomology, Michigan State University, 288 Farm Lane, room 243, East Lansing, MI 48824, USA</mods:affiliation>
</mods:name>
<mods:name id="0BA8E1A7A0B491DD5C0C1CE9F4E2158E" type="personal">
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</mods:role>
<mods:namePart id="FA744B3A4AF10C17D1B26F6574730D28">Pornsuriya, Chaninan</mods:namePart>
<mods:nameIdentifier id="AB3519D1ED11DECCCA85AA57E8824191" type="ORCID">0000-0002-8233-1254</mods:nameIdentifier>
<mods:affiliation id="112EA53A3F1C18D23BE9D6CFBD7A7E71">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
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<mods:namePart id="32516626FF9F5BED502CB771A5854963">Cognato, Anthony I.</mods:namePart>
<mods:nameIdentifier id="519F1DC8DC10942B1CC1C48A39865911" type="ORCID">0000-0001-6436-2447</mods:nameIdentifier>
<mods:affiliation id="F365ED0A57D1CD887EB5AF05D1AFAC83">Department of Entomology, Michigan State University, 288 Farm Lane, room 243, East Lansing, MI 48824, USA</mods:affiliation>
</mods:name>
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<mods:title id="9D24827A390659CB2A5E4C4F6BD7B9D2">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F712FC02F41FBF4EE9613FE3D1C27202">
<mods:date id="988EE247B28754432C5DD07636163CB1">2024</mods:date>
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<mods:number id="E5E7B6C2B7DEC78A7CCF785EFACD7520">2024-11-05</mods:number>
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<subSubSection id="3B425CC17FAD21CB9F5537C687D68BAE" type="nomenclature">
<paragraph id="7904FBC212765795055272023F9D95A4">
<taxonomicName id="AC53B488465792C752C3BDD1D40F1F62" authority="Lin, Sittichaya &amp; Smith" authorityName="Lin, Sittichaya &amp; Smith" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eccoptopterus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="formosanus" status="sp. nov.">
<emphasis id="E86823D453FD79EB45769C89B576B8FA" italics="true">Eccoptopterus formosanus</emphasis>
Lin, Sittichaya &amp; Smith
</taxonomicName>
<taxonomicNameLabel id="4C6EF464B425E86B487CA50C82568286" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0365D4C49C65CA4BD0782AA1FA189667" type="description">
<paragraph id="B8DDD58B09EB9444CA7F16CF13DD114D">
<figureCitation id="94152D44142D03C620F6FA3BF8F52A39" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eccoptopterus formosanus sp. nov. A E holotype female A dorsal view B lateral view C frons D posterolateral view of abdomen E declivital face; F I paratype male F dorsal view G lateral view H frons I posterolateral view of abdomen." figureDoi="10.3897/zookeys.1217.129707.figure2" httpUri="https://binary.pensoft.net/fig/1168130">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6UBG" type="type material">
<paragraph id="89C9C3A29F48803D36BDBFC32FBED013">
<heading id="9EBC5F6F6919F24B5BE92CBFC2087F8C" reason="title">Type material.</heading>
</paragraph>
<paragraph id="3D494802FA2E971CAED0B696DB05CFEA">
<emphasis id="026A353E022E0F460B70968EF56A24D2" bold="true" italics="true">
<typeStatus id="72D92128F7C29B71EEF0CD715FC30E2D" type="holotype">Holotype</typeStatus>
</emphasis>
: • female,
<collectingCountry id="64A1D71C9F788189D09CC7F707002B67" name="Taiwan">Taiwan</collectingCountry>
,
<collectingRegion id="357AA5A329D68577E9B59086DCE5C5CE" country="Taiwan" name="Nantou">Nantou county</collectingRegion>
, Renai Township;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENVBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[121.009578,24.000102]}">
<geoCoordinate id="CC12416F030723A7371A9D299C3E70F7" degrees="24" direction="north" minutes="0" orientation="latitude" precision="1" seconds="0.3675" value="24.000103">24 ° 0 ' 0.3675 &quot; N</geoCoordinate>
,
<geoCoordinate id="DB16BC45A1FEA824B780CA009CFFC7B6" degrees="121" direction="east" minutes="0" orientation="longitude" precision="1" seconds="34.4817" value="121.009575">121 ° 0 ' 34.4817 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="2410AA8C42C506450F443B8B45BFAD8A" metricMagnitude="2" metricUnit="m" metricValue="9.69" unit="m" value="969.0">969 m</quantity>
a. s. l.; a diameter
<quantity id="7713D4554879C815FE611F54673A43C4" metricMagnitude="-2" metricUnit="m" metricValue="4.5" unit="cm" value="4.5">4.5 cm</quantity>
branch of
<taxonomicName id="0B9CA87DC63F60D236DEB7BB1E80062D" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="8314F95ABD71E372A38B68045FF395BB" italics="true">Sapium discolor</emphasis>
</taxonomicName>
(
<taxonomicName id="0F07F109C60E1B19CB150E950FD4EDAC" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
); 02. iv. 23, (C. S. Lin) (
<named-content id="6184AC283A8AC93060842A78C3AB3B08" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-science" xlink_title="National Museum of Natural Science, Taichung">
<collectionCode id="6909CB36B806967D3D1BC3C681B0A8D1" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" type="">NMNS</collectionCode>
</named-content>
).
<emphasis id="FC7D8ECF765024CD95CBDDFF78452EEA" bold="true" italics="true">
<typeStatus id="CEC1F87E996BB37F0ECF2937C4B7EA3C" type="paratype">Paratypes</typeStatus>
</emphasis>
: • male, same as
<typeStatus id="5BDDB4540DA7C61C25500E5BFC6C4A3E" type="holotype">holotype</typeStatus>
(1,
<named-content id="D8F427BE286CB33223C425E3D3D44018" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-science" xlink_title="National Museum of Natural Science, Taichung">
<collectionCode id="C9CDCBA92F9A4D04DFE1EDC53DB7BCCE" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" type="">NMNS</collectionCode>
</named-content>
), female, (3,
<abbrev id="ABBRID0ENWBG" xlink_title="Albert J. Cook Arthropod Research Collection, Michigan State University">
<collectionCode id="35D031F80376B87D672E199A5A679E73">MSUC</collectionCode>
</abbrev>
),
<specimenCount id="FE38F468AE708B24DBBBCBA8F61A9B75" count="1" type="female">1 female</specimenCount>
,
<specimenCount id="529E77ECC04BCD8392E37DA2D6E35713" count="1" type="male">1 male</specimenCount>
(2
<abbrev id="ABBRID0ERWBG" xlink_title="National Taiwan University Insect Museum">
<collectionCode id="542DC1E43CEE9F0A33C79642DA4E30E4">NTU</collectionCode>
</abbrev>
),
<specimenCount id="CF57CB98062DBFDB33900C595D7FEEFC" count="1" type="male">1 male</specimenCount>
,
<specimenCount id="AABF6A9BFD8A1BB29936ED2CD4FF1F80" count="14" type="female">14 females</specimenCount>
(15,
<abbrev id="ABBRID0EVWBG" xlink_title="Private collection of Ching Shan Lin">
<collectionCode id="5295DC18FF045904EBF26A6175F94F57">CSL</collectionCode>
</abbrev>
) • female,
<collectingRegion id="F4D575DC92D61B1D199CD1C9F0340AE7" country="Taiwan" name="Nantou">Nantou County</collectingRegion>
(Lugu Township);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3WBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[120.816664,23.750086]}">
<geoCoordinate id="CDB2665AC9F8E36A35A0DAE218F34990" degrees="23" direction="north" minutes="45" orientation="latitude" precision="1" seconds="0.31" value="23.750086">23 ° 45 ' 0.31 &quot; N</geoCoordinate>
,
<geoCoordinate id="A126059ADD3AABA387A4B95808D6F882" degrees="120" direction="east" minutes="48" orientation="longitude" precision="1" seconds="59.99" value="120.816666">120 ° 48 ' 59.99 &quot; E</geoCoordinate>
</named-content>
;
<quantity id="A008083CC9F464AE4D3942E06F5FDE55" metricMagnitude="2" metricUnit="m" metricValue="7.2" unit="m" value="720.0">720 m</quantity>
a. s. l.; a diameter
<quantity id="FF16E3D0155A7FED939BCA46A00EFE5A" metricMagnitude="-2" metricUnit="m" metricValue="5.2" unit="cm" value="5.2">5.2 cm</quantity>
branch of
<taxonomicName id="99DBF69CF3AE15403774134B81FAF279" class="Magnoliopsida" family="Elaeocarpaceae" genus="Elaeocarpus" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="5F138EFADDFADB7035FF1A7EB01E7CA5" italics="true">Elaeocarpus sylvestris</emphasis>
</taxonomicName>
(
<taxonomicName id="D1BF6AA5924BCB0315046C0D49646A47" class="Magnoliopsida" family="Elaeocarpaceae" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="family">Elaeocarpaceae</taxonomicName>
); 022. vi. 23, (C. S. Lin),
<specimenCount id="F2A093705E78BD77004A9C2EF235FA5A" count="1" type="male">1 male</specimenCount>
,
<specimenCount id="0310D341A48D725FB3EBEC22FA319348" count="4" type="female">4 females</specimenCount>
(5,
<abbrev id="ABBRID0ERXBG" xlink_title="Private collection of Wisut Sittichaya">
<collectionCode id="4D0EBE27FF62EFAA50AD4AB7749092D0">WSTC</collectionCode>
</abbrev>
), female (1,
<named-content id="FE385968038F0E98E17185549FB51B9A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history" xlink_title="National Museum of Natural History, Smithsonian Institution">
<collectionCode id="819EE0599DA96482CCDFD42505A1BA51" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" type="Museum">NMNH</collectionCode>
</named-content>
), (1,
<named-content id="9048CC85891A71F74B4FF7F815CE3031" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="2DB85576C333C29344D70F8F660D0659" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4XBG" type="diagnosis">
<paragraph id="78E11A8ECB6669A58B6D2BFBD0AFAD58">
<heading id="EB416F9FAD7E685ABE2F4D8EB34AFDAD" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7809A443FB3BC9E737FC123021345015">
<emphasis id="A1E0BA2661F25CD80131B700E143519D" bold="true">Female</emphasis>
,
<quantity id="DC714862512C1F3F9B94F01918A67E7D" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.64" metricValueMin="2.56" unit="mm" value="2.6" valueMax="2.64" valueMin="2.56">2.56-2.64 mm</quantity>
long (mean =
<quantity id="A2DBE21EB1ED9F16A334C5F27BDC36E6" metricMagnitude="-3" metricUnit="m" metricValue="2.61" unit="mm" value="2.61">2.61 mm</quantity>
;
<emphasis id="C2666F21308160601CEEBC0E3DD0C22B" italics="true">N</emphasis>
= 4), 2.13-2.17 × as long as wide (mean = 2.14 ×;
<emphasis id="FD771D795A1CD18272651DC289A6EFFF" italics="true">N</emphasis>
= 4). Medium body size, declivital armature composed of a pair of major spines on declivital summit and 24 minor denticles unevenly spaced on each lateral margin; protibiae slender, broadest at apical 1 / 3, outer margin armed with six or seven moderated socketed denticles; scutellum broadly linguiform; elytra tapering laterally.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJYBG" type="description">
<paragraph id="653C4E39A046042833A380FF19BA0971">
<heading id="009B1C03CBB5A42997878255AF1A3B54" reason="title">Description.</heading>
</paragraph>
<paragraph id="476D5B55BC8A7ED0C448E057F66B87C0">
<emphasis id="39506971D04E122CE0C12D611E214FCE" bold="true">Female</emphasis>
(Fig.
<figureCitation id="DA9F2EB5C63A1C4D8193CDEBB24DDA58" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eccoptopterus formosanus sp. nov. A E holotype female A dorsal view B lateral view C frons D posterolateral view of abdomen E declivital face; F I paratype male F dorsal view G lateral view H frons I posterolateral view of abdomen." figureDoi="10.3897/zookeys.1217.129707.figure2" httpUri="https://binary.pensoft.net/fig/1168130">2 A E</figureCitation>
). Black brown, procoxae light brown, profemora and mesofemora paler brown, antennae, tibiae dark brown.
<emphasis id="B69D33D15D9DBDD421BA6FDD49FDA8BD" bold="true" italics="true">Head</emphasis>
: epistoma entire, transverse, with a row of hair-like setae. Frons below upper margin of eye and above epistoma flat, flattened area broadly rounded, surface subshiny, finely reticulate, sparsely punctate, punctures bearing fine, yellowish-white, hair-like setae. Eye shallowly emarginate just above antennal insertion, upper portion slightly smaller than lower portion. Submentum triangular small, moderately impressed. Antennal scape long, normal thick, slightly longer than club (12: 10). Pedicel as broad as scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club obliquely truncate, longer than wide (10: 9),
<typeStatus id="8711586B4CFD1BA79401ACD481FF13E0">type</typeStatus>
1, segment 1 corneous, occupying basal 1 / 4, margin carinate, concave, encircling anterior face, segment 2 and 3 soft, visible on anterior face only.
<emphasis id="557CE8F7203D1984230C88B3FA4BB2A5" bold="true" italics="true">Pronotum</emphasis>
: 0.97-1.00 (mean = 0.99,
<emphasis id="6D7C2539AD670F82A333BFE9F8C656F4" italics="true">N</emphasis>
= 4) × as long as wide,
<typeStatus id="B9CC43E777A79E22203CC9F0857D54A0">type</typeStatus>
<quantity id="983BCF08833ECC8BCEB5670DBA685E83" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
dorsal view, lateral sides parallel to anterior middle, broadly rounded anteriorly; anterior margin with 4-6 serrations, median pair prominent; anterior slope strongly asperate, asperities densely spaced, rugose, lower and more transverse toward the summit; disc slightly convex, finely reticulate, dull, sparsely covered with fine punctures bearing fine short hair-like setae. Base with a tuft of short hair-like setae associated with mycangium. In lateral view short and tall,
<typeStatus id="F47F080054544B286AFC8F27FA466C18">type</typeStatus>
3, summit at middle, lateral margins obliquely costate.
<emphasis id="8D0300CBD185460A0E4A40F99D7CE0D9" bold="true" italics="true">Elytra</emphasis>
: 1.14-1.17 (mean = 1.16,
<emphasis id="5F1387FF092D7654F80B035594EA168B" italics="true">N</emphasis>
= 4) × as long as wide, 1.10-1.17 (mean = 1.12,
<emphasis id="892C754236B0007CD7B297ED50BBEDE6" italics="true">N</emphasis>
= 4) × as long as pronotum. Scutellum comparatively moderately sized, narrow, linguiform, subshiny, attached on anterior slope of elytra less visible from above. Base shallowly bisinuate, with oblique edge, humeral angles rounded, lateral side tapering from humeral angle to apex. Disc short, basal area 1 / 4 of disc slightly convex, apical 3 / 4 impressed and connecting to declivital impression; disc punctate, punctures fine confused and setose, striae and interstriae hardly marked due to irregular punctures. Declivity sulcate, with a pair of major (largest) spines on declivital summit and 24 much smaller denticles on declivital margin; striae and interstriae punctate, punctures small and shallow, each bearing a short, semi-recumbent seta.
<emphasis id="370F593EBFA21A1CE4CEE83314778313" bold="true" italics="true">Legs</emphasis>
: procoxae contiguous; prosternal coxal piece short, conical. Protibiae slender, broadest at middle; posterior face inflated, punctate, densely covered with long hair-like setae; outer margin armed with six or seven moderate socketed denticles. Meso- and metatibiae rounded, flat, mesotibiae armed with seven or eight smaller socketed denticles, metafemora and metatibiae enlarged, metatibiae without spines.
</paragraph>
<caption id="1890BD3D7F753C59DC161E5FE0E38AE9" ID-DOI="10.3897/zookeys.1217.129707.figure2" ID-arpha="B5466BF6-7008-5F74-BF5F-3D67878B6717" httpUri="https://binary.pensoft.net/fig/1168130" startId="F2">
<paragraph id="16AF4E845D8168D68893531B42106F69">
<label id="1E640C188980D86E1AA61326DBA774C4">Figure 2.</label>
</paragraph>
<paragraph id="D5439ABED6606020C96973352CA7E3BF">
<taxonomicName id="B7877960D0B69B1153B30EDBDF53909C" authorityName="Sittichaya &amp; Lin &amp; Smith &amp; Pornsuriya &amp; Cognato" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eccoptopterus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="formosanus" status="sp. nov.">
<emphasis id="331468C2AB70C59FF2165100F438B4B2" italics="true">Eccoptopterus formosanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C75484D5C7DF8718A59C4725EFF36F7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E234BAABAB0660881823AE01D16F4D7D" bold="true">A E</emphasis>
holotype female
<emphasis id="F0E7E615DA1972FB7D9E6B48F5BB09A1" bold="true">A</emphasis>
dorsal view
<emphasis id="742E0CB42488360D4C1936FB9887A85F" bold="true">B</emphasis>
lateral view
<emphasis id="7C3137AD4CA07E7D7ADC132FAA91EBD1" bold="true">C</emphasis>
frons
<emphasis id="62B30964D4A932A771C36329D95C3D6E" bold="true">D</emphasis>
posterolateral view of abdomen
<emphasis id="D2E0FAAE5B44D4739489104EB29792AA" bold="true">E</emphasis>
declivital face;
<emphasis id="EB1F8A226A9D519A8C97E3E0D2183825" bold="true">F I</emphasis>
paratype male
<emphasis id="57D93846F33FD5720F8D5A85549DD909" bold="true">F</emphasis>
dorsal view
<emphasis id="BB85C1A833112206A0C20166ED50C10E" bold="true">G</emphasis>
lateral view
<emphasis id="EB8BD32E078C32C9D0AFA407623AE60F" bold="true">H</emphasis>
frons
<emphasis id="9B8B98DE8091F48B6FC8CB2ADF2EA1EF" bold="true">I</emphasis>
posterolateral view of abdomen.
</paragraph>
</caption>
<paragraph id="CD86F34CAB26F1579740131CBC7AC53A">
<emphasis id="D6ADBB27F1BD35839761598C957A2E44" bold="true">Male.</emphasis>
(Fig.
<figureCitation id="C954DC49247E939DE80752A7E00ECFA9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eccoptopterus formosanus sp. nov. A E holotype female A dorsal view B lateral view C frons D posterolateral view of abdomen E declivital face; F I paratype male F dorsal view G lateral view H frons I posterolateral view of abdomen." figureDoi="10.3897/zookeys.1217.129707.figure2" httpUri="https://binary.pensoft.net/fig/1168130">2 F I</figureCitation>
).
<quantity id="DC87194E756574C5E0AC8AEA47C02E95" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.3" metricValueMin="2.18" unit="mm" value="2.24" valueMax="2.3" valueMin="2.18">2.18-2.30 mm</quantity>
(mean =
<quantity id="99ABA7D1E843E8D32359926215443A58" metricMagnitude="-3" metricUnit="m" metricValue="2.24" unit="mm" value="2.24">2.24 mm</quantity>
;
<emphasis id="CFF3CF94DB24F68F5A4EC8DB470C158B" italics="true">N</emphasis>
= 4) long, 1.81-2.04 (mean 1.95;
<emphasis id="5849AB3C13623DE29B3CB7191E2569C5" italics="true">N</emphasis>
= 4) × as long as wide. Head reddish-brown, coxae light brown, femora paler brown, antennae, tibiae, pronotum and elytra dark brown in mature specimens, excepting impressed portion of pronotum is reddish-brown.
<emphasis id="1CEF3E0E66B41FFE72F0B0056D9F6A59" bold="true" italics="true">Head</emphasis>
: somewhat margined laterally and concealed under the projection of pronotum. Epistoma entire, transverse, with a row of hair-like setae. Frons below upper margin of eye and above epistoma flat, the flattened area quadrate, surface subshiny, finely reticulate, sparsely shallow punctured, each bearing a short or long hair-like seta. Eye reduced, shallowly emarginate just above antennal insertion, upper portion slightly smaller than lower portion. Submentum triangular, small, slightly impressed. Antennal scape long, normal thick, slightly longer than club. Pedicel as broad as scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club obliquely truncate, longer than wide (7.5: 5.5),
<typeStatus id="FA47FF42F9447C655058B971330CFD16">type</typeStatus>
1, segment 1 corneous, occupying basal 1 / 4, margin carinate, concave, encircling anterior face, segments 2 and 3 soft, visible on anterior face only.
<emphasis id="B23226A1AF00798BFB4DC8A07F02A6DB" bold="true" italics="true">Pronotum</emphasis>
: 0.89-1.11 (mean = 0.99;
<emphasis id="9C6E3EE8AE1D82A666B06A9FC377FF7B" italics="true">N</emphasis>
= 4) × as long as wide,
<typeStatus id="03EF73ED58AFE39426DE531F5364B62C">type</typeStatus>
<quantity id="17ED1EA5334158361CA3E17382286079" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
dorsal view, lateral sides subparallel to anterior middle, broadly rounded anteriorly; anterior margin unarmed; widely impressed on anterior slope, then gradually narrowing and slightly impressed toward disc, weakly asperate, asperities sparsely spaced, sub-rugose, becoming lower and more transverse toward the summit; slightly longitudinally impressed in middle of disc, finely reticulate, dull, sparsely covered with fine punctures bearing fine short hair-like setae. In lateral view long and tall,
<typeStatus id="BE1A0CACEABDC0A5823E35B446E85998">type</typeStatus>
9, summit at apical 2 / 3, lateral margins obliquely costate.
<emphasis id="802B47B869184C136388EE2EFD94F013" bold="true" italics="true">Elytra</emphasis>
: 1.00-1.05 (mean = 1.04;
<emphasis id="4032A345B453C83F539198F65B519640" italics="true">N</emphasis>
= 4) × as long as wide, 0.80-1.00 (mean = 0.88;
<emphasis id="3D55A450196B359B708E9BA969193CB0" italics="true">N</emphasis>
= 4) × as long as pronotum. Scutellum small, linguiform, subshiny, attachment on anterior slope of elytra less visible from above. Base procurved, with oblique edge, humeral angles rounded, subparallel-sided in basal 1 / 2, then gradually incurved to broadly rounded apex. Disc short, punctate, punctures fine confused and setose, strial setae uniseriate with long, erect hair-like setae; interstrial setae uni- or biseriate with semi-recumbent hair-like setae; basal area 1 / 4 of disc slightly convex. Declivity somewhat steeply sloping, face weakly bisulcate, declivital face much lower than declivital margin, with a pair of major spines on declivital summit and 02 much smaller minor tubercles on declivital margin; striae and interstriae punctate, punctures small and shallow, each bearing a short, semi-recumbent setae.
<emphasis id="E47F802E35F7489BF5CE06D62198FB59" bold="true" italics="true">Legs</emphasis>
: procoxae narrowly separated; prosternal coxal piece short, conical. Protibiae slender, broadest at the middle; posterior face inflated, punctate, densely covered with long hair-like setae; outer margin armed with five or six moderate socketed denticles. Meso- and metatibiae rounded, flat, mesotibiae armed with six or seven smaller socketed denticles, metafemura and metatibiae enlarged, metatibiae without apical spine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET2BG" type="etymology">
<paragraph id="FE4B7AA323A576B29B58E2E046077252">
<heading id="FB721456381F2A3A2B88EA3C81E48897" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7F18225AF915F42BE097ADA3AD14D747">
Formosa, the former name of
<collectingCountry id="3C96D13644A1A6BB95EB712B7937A1BE" name="Taiwan">Taiwan</collectingCountry>
island, in reference to the collection locality of
<typeStatus id="6E66FFBEEF67456252CE7F83CDB774B2">types</typeStatus>
. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY2BG" type="distribution">
<paragraph id="F597EC4DD4748F2DEF8DEBFF34B780E7">
<heading id="3D16DB6BBFDC950606476A2B7C0CE971" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="50BE914E5186991317016CEA2F6A51F2">
<collectingCountry id="9C16C8C474245FA6DD9266A48A349EE7" name="Taiwan">Taiwan</collectingCountry>
(
<collectingRegion id="02E33C67B894DA42F9E7A6F0D84B1EE7" country="Taiwan" name="Nantou">Nantou County</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E42BG" type="biology">
<paragraph id="7D4FE567110FC6E76AAB9EE7DCFDC165">
<heading id="7F41303BD77CCF84A951E66EB3FF00C4" reason="title">Biology.</heading>
</paragraph>
<paragraph id="D3287F37A9F24802D5BF198EC54D9545">
Bred from
<taxonomicName id="D47CB461010CB923A52C36A242EFD719" authority="(Lour.) Poir." authorityName="Poir." baseAuthorityName="Lour." class="Magnoliopsida" family="Elaeocarpaceae" genus="Elaeocarpus" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="FF6F6432DF13D73CE25BFE2F02161000" italics="true">Elaeocarpus sylvestris</emphasis>
(Lour.) Poir.
</taxonomicName>
(
<taxonomicName id="2CCD06A586E80C8CDA08347F12C9C81A" class="Magnoliopsida" family="Elaeocarpaceae" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="family">Elaeocarpaceae</taxonomicName>
),
<taxonomicName id="5F654D3BB4DBD1ADC4ABE6252BAEF6EC" authority="(Benth.) Rehder" authorityName="Rehder" baseAuthorityName="Benth." class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="hancei">
<emphasis id="99E9BCE9E4C4A01BFD5964FE3628C8A6" italics="true">Lithocarpus hancei</emphasis>
(Benth.) Rehder
</taxonomicName>
,
<taxonomicName id="2B2AF078793DE66F03DB9787645BE938" authority="Thunb. ex Murray" authorityName="Murray" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="glauca">
<emphasis id="234D9617EC110AB630D2A7EBD4DA85C8" italics="true">Quercus glauca</emphasis>
Thunb. ex Murray
</taxonomicName>
(
<taxonomicName id="C9EA79FB1ADE8295DF22EF2D68EEFDF0" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
),
<taxonomicName id="7A76B21EAF8460124FA4E9202CD1A52E" authority="Muell." authorityName="Muell." class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="9B6C7DED1C8D1BECD17C2C278385F54A" italics="true">Sapium discolor</emphasis>
Muell.
</taxonomicName>
- Arg. (
<taxonomicName id="87AF36E50E15E17279F14232AFAAD47E" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
),
<taxonomicName id="0E9F24A13985E6223E881CB5933DA4EC" class="Magnoliopsida" family="Cannabaceae" genus="Trema" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="orientale">
<emphasis id="723CBB3102924DD7412373FA607173DA" italics="true">Trema orientale</emphasis>
</taxonomicName>
(L.) Blume (
<taxonomicName id="66E18852CACF31CE0EAD843DEC6762F6" class="Magnoliopsida" family="Cannabaceae" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="family">Cannabaceae</taxonomicName>
) with a diameter of about
<quantity id="9290379E3F8AD84A20C31B01DFEDE81B" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.2" metricValueMin="4.8" unit="cm" value="5.5" valueMax="6.2" valueMin="4.8">4.86.2 cm</quantity>
in
<collectingCountry id="C6F10F48966D571C83E31E3D14BB73BC" name="Taiwan">Taiwan</collectingCountry>
. The radial entrance gallery leads to several branches in various planes without enlarged brood chambers (C. S. Lin pers. obs.).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,174 @@
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<mods:title id="AC96210ADB3317F3BF658BF7E77B76F1">On the Brazilian Exocnophila stick insects (Insecta: Phasmatodea), with the description of a new species</mods:title>
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Genus
<taxonomicName id="91A50DFAE16AFF85FF37FC76FDF8FC2E" ID-CoL="8NRQQ" authority="Zompro, 2001" authorityName="Zompro" authorityYear="2001" box="[238,576,979,1006]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="64D1AA6BE16AFF85FF37FC76FE31FC2D" bold="true" box="[238,393,979,1006]" italics="true" pageId="2" pageNumber="61">Exocnophila</emphasis>
<bibRefCitation id="32340B88E16AFF85FE48FC76FDF8FC2E" author="Zompro, O." box="[401,576,979,1005]" pageId="2" pageNumber="61" pagination="189 - 255" refId="ref16401" refString="Zompro, O. (2001) A generic revision of the insect order Phasmatodea: The New World genera of the stick insect subfamily Diapheromeridae: Diapheromerinae = Heteronemiidae: Heteronemiinae sensu Bradley &amp; Galil, 1977. Revue Suisse de Zoologie, 108, 189 - 255. https: // doi. org / 10.5962 / bhl. part. 79626" type="journal article" year="2001">Zompro, 2001</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1EBF25F2E16AFF85FF4EFBBEFAF6FB9A" pageId="2" pageNumber="61" type="distribution">
<paragraph id="561A7679E16AFF85FF4EFBBEFAF6FB9A" blockId="2.[151,1436,1051,1114]" pageId="2" pageNumber="61">
<emphasis id="64D1AA6BE16AFF85FF4EFBBEFE91FBF6" bold="true" box="[151,297,1051,1077]" pageId="2" pageNumber="61">Distribution</emphasis>
(
<figureCitation id="CE9E6AFCE16AFF85FEE3FBBEFE34FBF6" box="[314,396,1051,1077]" captionStart="FIGURE 31" captionStartId="36.[151,250,1852,1876]" captionTargetBox="[151,1435,921,1829]" captionTargetId="figure-366@36.[151,1435,921,1829]" captionTargetPageId="36" captionText="FIGURE 31. Map of the known distribution for Exocnophila species, in southeast Brazil. Precise localities are known only for Rio de Janeiro state; the inlet map shows Brazilian states with records for the genus highlighted in red, map colours indicating biomes (dark green: Amazon, green: Atlantic Forest; orange: Cerrado; beige: Caatinga)." figureDoi="http://doi.org/10.5281/zenodo.14024131" httpUri="https://zenodo.org/record/14024131/files/figure.png" pageId="2" pageNumber="61">Fig. 31</figureCitation>
). The genus is endemic to
<collectingCountry id="2EB236E9E16AFF85FD66FBBEFCBBFBF6" box="[703,771,1051,1077]" name="Brazil" pageId="2" pageNumber="61">Brazil</collectingCountry>
and is recorded only from the Atlantic Forest in Southeast and Northeast region of the country, in the states of
<collectingRegion id="9461B89BE16AFF85FD05FB9AFC98FB9A" box="[732,800,1087,1113]" country="Brazil" name="Bahia" pageId="2" pageNumber="61">Bahia</collectingRegion>
,
<collectingRegion id="9461B89BE16AFF85FCF2FB9AFC76FB9A" box="[811,974,1087,1113]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="61">Espírito Santo</collectingRegion>
,
<collectingRegion id="9461B89BE16AFF85FC00FB9AFBC2FB9A" box="[985,1146,1087,1113]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="61">Rio de Janeiro</collectingRegion>
and
<collectingRegion id="9461B89BE16AFF85FB68FB9AFAF3FB9A" box="[1201,1355,1087,1113]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="61">Minas Gerais</collectingRegion>
.
</paragraph>
</subSubSection>
<subSection id="3F2A6F92E16AFF85FF4EFB0EFAE8FA02" pageId="2" pageNumber="61" type="multiple">
<subSubSection id="1EBF25F2E16AFF85FF4EFB0EFE46FA5E" pageId="2" pageNumber="61" type="taxon_list">
<paragraph id="561A7679E16AFF85FF4EFB0EFE8FFB06" blockId="2.[151,311,1195,1221]" box="[151,311,1195,1221]" pageId="2" pageNumber="61">
<heading id="0D52C115E16AFF85FF4EFB0EFE8FFB06" bold="true" box="[151,311,1195,1221]" fontSize="11" level="1" pageId="2" pageNumber="61" reason="1">
<emphasis id="64D1AA6BE16AFF85FF4EFB0EFE8FFB06" bold="true" box="[151,311,1195,1221]" pageId="2" pageNumber="61">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="561A7679E16AFF85FF4EFB56FC1EFACE" blockId="2.[151,1360,1267,1474]" box="[151,934,1267,1294]" pageId="2" pageNumber="61">
<taxonomicName id="91A50DFAE16AFF85FF4EFB56FCA7FACE" authority="(Brunner von Wattenwyl, 1907)" authorityName="Ghirotto &amp; Conle &amp; Hennemann &amp; Valero &amp; Cancello" authorityYear="2024" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1907" box="[151,799,1267,1294]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="brevitarsata" status="comb. nov.">
<emphasis id="64D1AA6BE16AFF85FF4EFB56FE0CFACE" box="[151,436,1267,1293]" italics="true" pageId="2" pageNumber="61">Exocnophila brevitarsata</emphasis>
(
<bibRefCitation id="32340B88E16AFF85FE1BFB56FCAFFACE" author="Brunner von Wattenwyl, K." box="[450,791,1267,1294]" pageId="2" pageNumber="61" pagination="181 - 338" refId="ref15134" refString="Brunner von Wattenwyl, K. (1907) IX. Tribus Clitumnini, X. Tribus Lonchodini, XI. Tribus Bacunculini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 181 - 338, tafel VII - XV." type="book chapter" year="1907">Brunner von Wattenwyl, 1907</bibRefCitation>
)
</taxonomicName>
<emphasis id="64D1AA6BE16AFF85FCFFFB56FC1EFACE" bold="true" box="[806,934,1267,1293]" pageId="2" pageNumber="61">
<taxonomicNameLabel id="7FE21710E16AFF85FCFFFB56FC1EFACE" box="[806,934,1267,1293]" pageId="2" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="561A7679E16AFF85FF4EFAB2FD59FAF2" blockId="2.[151,1360,1267,1474]" box="[151,737,1303,1329]" pageId="2" pageNumber="61">
<taxonomicName id="91A50DFAE16AFF85FF4EFAB2FDE1FAF2" authority="(Burmeister, 1838)" authorityName="Ghirotto &amp; Conle &amp; Hennemann &amp; Valero &amp; Cancello" authorityYear="2024" baseAuthorityName="Burmeister" baseAuthorityYear="1838" box="[151,601,1303,1329]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="hastata" status="comb. nov.">
<emphasis id="64D1AA6BE16AFF85FF4EFAB2FEC6FAF2" box="[151,382,1303,1329]" italics="true" pageId="2" pageNumber="61">Exocnophila hastata</emphasis>
(
<bibRefCitation id="32340B88E16AFF85FE54FAB2FDE9FAF2" author="Burmeister, H." box="[397,593,1303,1329]" pageId="2" pageNumber="61" pagination="397 - 756" refId="ref15201" refString="Burmeister, H. (1838) Handbuch der Entomologie. Zweiter Band. Theod. Chr. Friedr. Enslin., Berlin, pp. 397 - 756." type="book chapter" year="1838">Burmeister, 1838</bibRefCitation>
)
</taxonomicName>
<emphasis id="64D1AA6BE16AFF85FDB9FAB2FD59FAF2" bold="true" box="[608,737,1303,1329]" pageId="2" pageNumber="61">
<taxonomicNameLabel id="7FE21710E16AFF85FDB9FAB2FD59FAF2" box="[608,737,1303,1329]" pageId="2" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="561A7679E16AFF85FF4EFA9EFCAFFA96" blockId="2.[151,1360,1267,1474]" box="[151,791,1339,1366]" pageId="2" pageNumber="61">
<taxonomicName id="91A50DFAE16AFF85FF4EFA9EFCAFFA96" authority="(Brunner von Wattenwyl, 1907)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1907" box="[151,791,1339,1366]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="64D1AA6BE16AFF85FF4EFA9EFE14FA96" box="[151,428,1339,1365]" italics="true" pageId="2" pageNumber="61">Exocnophila tuberculata</emphasis>
(
<bibRefCitation id="32340B88E16AFF85FE63FA9EFCB7FA96" author="Brunner von Wattenwyl, K." box="[442,783,1339,1366]" pageId="2" pageNumber="61" pagination="181 - 338" refId="ref15134" refString="Brunner von Wattenwyl, K. (1907) IX. Tribus Clitumnini, X. Tribus Lonchodini, XI. Tribus Bacunculini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 181 - 338, tafel VII - XV." type="book chapter" year="1907">Brunner von Wattenwyl, 1907</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="561A7679E16AFF85FF4EFAFAFD56FABA" blockId="2.[151,1360,1267,1474]" box="[151,750,1375,1402]" pageId="2" pageNumber="61">
<taxonomicName id="91A50DFAE16AFF85FF4EFAFAFD56FABA" authority="(Brunner von Wattenwyl, 1907)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1907" box="[151,750,1375,1402]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="64D1AA6BE16AFF85FF4EFAFAFE3BFABA" box="[151,387,1375,1401]" italics="true" pageId="2" pageNumber="61">Exocnophila cornuta</emphasis>
(
<bibRefCitation id="32340B88E16AFF85FE48FAFAFD5EFABA" author="Brunner von Wattenwyl, K." box="[401,742,1375,1402]" pageId="2" pageNumber="61" pagination="181 - 338" refId="ref15134" refString="Brunner von Wattenwyl, K. (1907) IX. Tribus Clitumnini, X. Tribus Lonchodini, XI. Tribus Bacunculini. In: Brunner von Wattenwyl, K. &amp; Redtenbacher, J. (Eds.), 1906 - 1908, Die Insektenfamilie der Phasmiden. Wilhelm Engelmann, Leipzig, pp. 181 - 338, tafel VII - XV." type="book chapter" year="1907">Brunner von Wattenwyl, 1907</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="561A7679E16AFF85FF4EFA26FE46FA5E" blockId="2.[151,1360,1267,1474]" box="[151,510,1411,1437]" pageId="2" pageNumber="61">
<taxonomicName id="91A50DFAE16AFF85FF4EFA26FE26FA5E" authorityName="Ghirotto &amp; Conle &amp; Hennemann &amp; Valero &amp; Cancello" authorityYear="2024" box="[151,414,1411,1437]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="ovifuscum" status="sp. nov.">
<emphasis id="64D1AA6BE16AFF85FF4EFA26FE26FA5E" box="[151,414,1411,1437]" italics="true" pageId="2" pageNumber="61">Exocnophila ovifuscum</emphasis>
</taxonomicName>
<emphasis id="64D1AA6BE16AFF85FE7DFA26FE46FA5E" bold="true" box="[420,510,1411,1437]" pageId="2" pageNumber="61">
<taxonomicNameLabel id="7FE21710E16AFF85FE7DFA26FE46FA5E" box="[420,510,1411,1437]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="1EBF25F2E16AFF85FF4EFA0DFAE8FA02" box="[151,1360,1447,1474]" pageId="2" pageNumber="61" type="type_taxon">
<paragraph id="561A7679E16AFF85FF4EFA0DFAE8FA02" blockId="2.[151,1360,1267,1474]" box="[151,1360,1447,1474]" pageId="2" pageNumber="61">
<typeStatus id="891EC8DBE16AFF85FF4EFA0DFF77FA01" box="[151,207,1448,1474]" pageId="2" pageNumber="61">Type</typeStatus>
species:
<taxonomicName id="91A50DFAE16AFF85FEECFA02FD8CFA02" authorityName="Zompro" authorityYear="2001" box="[309,564,1447,1473]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="exintegra">
<emphasis id="64D1AA6BE16AFF85FEECFA02FD8CFA02" box="[309,564,1447,1473]" italics="true" pageId="2" pageNumber="61">Exocnophila exintegra</emphasis>
</taxonomicName>
(junior synonym of
<taxonomicName id="91A50DFAE16AFF85FCC3FA02FB8EFA02" authorityName="Ghirotto &amp; Conle &amp; Hennemann &amp; Valero &amp; Cancello" authorityYear="2024" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1907" box="[794,1078,1447,1473]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="brevitarsata">
<emphasis id="64D1AA6BE16AFF85FCC3FA02FB8EFA02" box="[794,1078,1447,1473]" italics="true" pageId="2" pageNumber="61">Exocnophila brevitarsata</emphasis>
</taxonomicName>
) by original designation.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>

View file

@ -1,105 +0,0 @@
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<paragraph id="561A7679E16AFF85FF4EFC76FDF8FC2E" blockId="2.[151,576,979,1006]" box="[151,576,979,1006]" pageId="2" pageNumber="61">
<heading id="0D52C115E16AFF85FF4EFC76FDF8FC2E" bold="true" box="[151,576,979,1006]" fontSize="11" level="1" pageId="2" pageNumber="61" reason="1">
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Genus
<taxonomicName id="91A50DFAE16AFF85FF37FC76FDF8FC2E" ID-CoL="8NRQQ" authority="Zompro, 2001" authorityName="Zompro" authorityYear="2001" box="[238,576,979,1006]" class="Insecta" family="Diapheromeridae" genus="Exocnophila" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="64D1AA6BE16AFF85FF37FC76FE31FC2D" bold="true" box="[238,393,979,1006]" italics="true" pageId="2" pageNumber="61">Exocnophila</emphasis>
<bibRefCitation id="32340B88E16AFF85FE48FC76FDF8FC2E" author="Zompro, O." box="[401,576,979,1005]" pageId="2" pageNumber="61" pagination="189 - 255" refId="ref16401" refString="Zompro, O. (2001) A generic revision of the insect order Phasmatodea: The New World genera of the stick insect subfamily Diapheromeridae: Diapheromerinae = Heteronemiidae: Heteronemiinae sensu Bradley &amp; Galil, 1977. Revue Suisse de Zoologie, 108, 189 - 255. https: // doi. org / 10.5962 / bhl. part. 79626" type="journal article" year="2001">Zompro, 2001</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1EBF25F2E16AFF85FF4EFBBEFAF6FB9A" pageId="2" pageNumber="61" type="distribution">
<paragraph id="561A7679E16AFF85FF4EFBBEFAF6FB9A" blockId="2.[151,1436,1051,1114]" pageId="2" pageNumber="61">
<emphasis id="64D1AA6BE16AFF85FF4EFBBEFE91FBF6" bold="true" box="[151,297,1051,1077]" pageId="2" pageNumber="61">Distribution</emphasis>
(
<figureCitation id="CE9E6AFCE16AFF85FEE3FBBEFE34FBF6" box="[314,396,1051,1077]" captionStart="FIGURE 31" captionStartId="36.[151,250,1852,1876]" captionTargetBox="[151,1435,921,1829]" captionTargetId="figure-366@36.[151,1435,921,1829]" captionTargetPageId="36" captionText="FIGURE 31. Map of the known distribution for Exocnophila species, in southeast Brazil. Precise localities are known only for Rio de Janeiro state; the inlet map shows Brazilian states with records for the genus highlighted in red, map colours indicating biomes (dark green: Amazon, green: Atlantic Forest; orange: Cerrado; beige: Caatinga)." figureDoi="http://doi.org/10.5281/zenodo.14024131" httpUri="https://zenodo.org/record/14024131/files/figure.png" pageId="2" pageNumber="61">Fig. 31</figureCitation>
). The genus is endemic to
<collectingCountry id="2EB236E9E16AFF85FD66FBBEFCBBFBF6" box="[703,771,1051,1077]" name="Brazil" pageId="2" pageNumber="61">Brazil</collectingCountry>
and is recorded only from the Atlantic Forest in Southeast and Northeast region of the country, in the states of
<collectingRegion id="9461B89BE16AFF85FD05FB9AFC98FB9A" box="[732,800,1087,1113]" country="Brazil" name="Bahia" pageId="2" pageNumber="61">Bahia</collectingRegion>
,
<collectingRegion id="9461B89BE16AFF85FCF2FB9AFC76FB9A" box="[811,974,1087,1113]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="61">Espírito Santo</collectingRegion>
,
<collectingRegion id="9461B89BE16AFF85FC00FB9AFBC2FB9A" box="[985,1146,1087,1113]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="61">Rio de Janeiro</collectingRegion>
and
<collectingRegion id="9461B89BE16AFF85FB68FB9AFAF3FB9A" box="[1201,1355,1087,1113]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="61">Minas Gerais</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,339 @@
<document id="47F8EA6146F41E6D2FD8CE9BDAE8B059" ID-DOI="10.3897/zookeys.1217.129707" ID-ZooBank="541D9AC9-DD07-4435-ACA2-867C5F302D34" ID-publisher-id="129707" URI-arpha="9247C138-7213-58ED-9D52-A66F2E597D9C" URI-zoobank="http://zoobank.org/541D9AC9-DD07-4435-ACA2-867C5F302D34" article-type="research-article" checkinTime="1730829902517" checkinUser="pensoft" docAuthor="Sittichaya, Wisut, Lin, Ching-Shan, Smith, Sarah M., Pornsuriya, Chaninan &amp; Cognato, Anthony I." docDate="2024" docId="F2E0A6A18C5F5F878C6CBB465C1AB33D" docLanguage="en" docName="ZooKeys 1217: 247-262" docOrigin="ZooKeys 1217" docSource="https://zookeys.pensoft.net/article/129707/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eccoptopterus intermedius Sittichaya, Lin &amp; Smith 2024, sp. nov." docType="treatment" docUuid="BCEE85CB-6BE4-412A-B1B7-E3096D103600" docUuidSource="ZooBank" docVersion="1" dtd-version="3.0" lastPageNumber="262" masterDocId="9247C138721358ED9D52A66F2E597D9C" masterDocTitle="Two new species of Eccoptopterus Motschulsky, 1863 ambrosia beetle from Taiwan and Thailand (Coleoptera, Curculionidae, Scolytinae, Xyleborini)" masterLastPageNumber="262" masterPageNumber="247" pageNumber="247" updateTime="1730829973222" updateUser="pensoft">
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<mods:title id="17FC0F249BA1A68D56A0CD9E1251A328">Two new species of Eccoptopterus Motschulsky, 1863 ambrosia beetle from Taiwan and Thailand (Coleoptera, Curculionidae, Scolytinae, Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name id="2B7A7E8F85208C67496C5AF7F46DB091" type="personal">
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<mods:namePart id="76737C0F88966D340B1CEE889483B2C9">Sittichaya, Wisut</mods:namePart>
<mods:nameIdentifier id="593A547542C50C9F43FAB4BD65DE3E22" type="ORCID">0000-0001-6200-1285</mods:nameIdentifier>
<mods:affiliation id="2254D5557411389295C4F3A6B399680A">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="0524CCD8B333E457FBC7E8247B50DDF4">Lin, Ching-Shan</mods:namePart>
<mods:nameIdentifier id="4F713697064643DBE79AB7055A4BF5EC" type="ORCID">https://orcid.org/0009-0006-3159-697X</mods:nameIdentifier>
<mods:affiliation id="34D9BC2EE5FEDAEB134E83020133E873">Department of Entomology, National Taiwan University, Taipei 10617, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="0C4CD49924196C04AE38869D6AAF5E84" type="personal">
<mods:role id="AF60F27D6558396878332D3D37D8E7FE">
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</mods:role>
<mods:namePart id="0FE8EF4921D4CC1A7EF0810EE90C093F">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="A10A38ECA6901E4445E4F0FFEBDF3826" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="54E8384897D29AC199BB3C44ECCF2FB6">Department of Entomology, Michigan State University, 288 Farm Lane, room 243, East Lansing, MI 48824, USA</mods:affiliation>
</mods:name>
<mods:name id="3C033EE859DFF0E7E79189F36DB4AC32" type="personal">
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</mods:role>
<mods:namePart id="12CC9BB46A7757922986D24C909732FB">Pornsuriya, Chaninan</mods:namePart>
<mods:nameIdentifier id="2C169057DEE5D079D9CE70F277D89622" type="ORCID">0000-0002-8233-1254</mods:nameIdentifier>
<mods:affiliation id="20DE090887484B60084C63E06558C874">Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand</mods:affiliation>
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<mods:namePart id="4ECD8727C71FBD783E60C86A7F8EB84F">Cognato, Anthony I.</mods:namePart>
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<mods:affiliation id="4A4169EA79E89E5176D6A0618987EF1C">Department of Entomology, Michigan State University, 288 Farm Lane, room 243, East Lansing, MI 48824, USA</mods:affiliation>
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<mods:date id="7D51162FDA1B6E78AA5AED40FC44A3AC">2024</mods:date>
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<mods:number id="45C2BBE0439A342CE7A4435A8005A3E4">2024-11-05</mods:number>
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<mods:classification id="EB9DE1AE340C7EFD6946B74973364647">journal article</mods:classification>
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<treatment id="F2E0A6A18C5F5F878C6CBB465C1AB33D" ID-arpha="F2E0A6A1-8C5F-5F87-8C6C-BB465C1AB33D" ID-zoobank="https://zoobank.org/BCEE85CB-6BE4-412A-B1B7-E3096D103600" LSID="urn:lsid:zoobank.org:act:BCEE85CB-6BE4-412A-B1B7-E3096D103600" LSID-ZBK="urn:lsid:zoobank.org:act:BCEE85CB-6BE4-412A-B1B7-E3096D103600" httpUri="http://treatment.plazi.org/id/F2E0A6A18C5F5F878C6CBB465C1AB33D">
<subSubSection id="32557F75AE70DB0FCC02EA24793B2EC7" type="nomenclature">
<paragraph id="2E1267ECE93FCF773F35560259FA8DEC">
<taxonomicName id="BF37F720070CF2125F9D1784B336FC5F" authority="Sittichaya, Lin &amp; Smith" authorityName="Sittichaya, Lin &amp; Smith" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eccoptopterus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="3F5FAA629A44AC50B05F802A4A2998A5" italics="true">Eccoptopterus intermedius</emphasis>
Sittichaya, Lin &amp; Smith
</taxonomicName>
<taxonomicNameLabel id="26CDAA57C31A5A1B0549025ADB1C7A47" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B6E0E2C00D0017047346572DF64CF067" type="description">
<paragraph id="62DDE5B2985A25D3E3BD52315745581E">
<figureCitation id="343B4A656B81F65F61B04F65681E68A7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eccoptopterus intermedius sp. nov. Holotype, female, A dorsal view B lateral view C posterolateral view D frons E antenna." figureDoi="10.3897/zookeys.1217.129707.figure3" httpUri="https://binary.pensoft.net/fig/1168131">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E36BG" type="type material">
<paragraph id="3D27A25C96663FE1B6632F557CEAE1CC">
<heading id="0DA9449B3E13598113224D9CBEC08675" reason="title">Type material.</heading>
</paragraph>
<paragraph id="2BAC38F6C5681134981341454A922DE0">
<materialsCitation id="3B01FC7F3F3B9414DDF587929C94DB10" collectionCode="NHMW" collectorName="W. Sittichaya" country="Thailand" latitude="17.671583" location="Tak Province" longLatPrecision="1" longitude="97.851166" specimenCount="1" stateProvince="Tak" typeStatus="holotype">
<emphasis id="D445530006D32B80FCC8D8FC8D587D1D" bold="true" italics="true">
<typeStatus id="6E1FFB26D22FDCEA6C1BC2BAE11E5EDC" type="holotype">Holotype</typeStatus>
</emphasis>
, • female,
<collectingCountry id="10C4C4C6535E413E63F94BF7CB12B105" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="4FA66EC4FBB8BAD4913F8A34E2A832AA" country="Thailand" name="Tak">Tak Province</collectingRegion>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELAAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[97.851167,17.671583]}">
<geoCoordinate id="80167FA20A12088A0577022CA027D91B" degrees="17" direction="north" minutes="40" orientation="latitude" precision="1" seconds="17.7" value="17.671583">17 ° 40 ' 17.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="C270ED599947EDBE4421BD9F02C9E029" degrees="97" direction="east" minutes="51" orientation="longitude" precision="1" seconds="04.2" value="97.851166">97 ° 51 ' 04.2 &quot; E</geoCoordinate>
</named-content>
; 600 msl; semiagricultural area, ex. small branch of unknown tree; 08. ix. 19, (
<collectorName id="766CB84B1D8A001E27ADACA448C9A661">W. Sittichaya</collectorName>
) (
<named-content id="5FF20C3AF3193FEFDE75DFCE5F280CA9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="5FB23B13069D3882B9F4709CB1C3A2C5" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
</named-content>
)
</materialsCitation>
;
<materialsCitation id="1ED86227EC5F48825778D4C5909EAAA6" collectingDate="2019-05-01" collectorName="W. Sittichaya" country="Thailand" elevation="420" latitude="15.62275" location="Pha Taem National Park" longLatPrecision="1" longitude="105.609634" specimenCount="1" stateProvince="Ubon Ratchathani" typeStatus="paratype">
<emphasis id="ED1B803CEE491013EFF5F0F5E8DE4B33" bold="true" italics="true">
<typeStatus id="A982A9E8EFB8FE6C8F63F5FA556D2913" type="paratype">Paratypes</typeStatus>
</emphasis>
: • females,
<collectingRegion id="978AF7954F6E290754E2EF97B4CEAD35" country="Thailand" name="Ubon Ratchathani">Ubon Ratchathani Province</collectingRegion>
,
<location id="77645B4973A8A041802CB9011019523F" LSID="urn:lsid:plazi:treatment:F2E0A6A18C5F5F878C6CBB465C1AB33D:77645B4973A8A041802CB9011019523F" country="Thailand" latitude="15.62275" longLatPrecision="1" longitude="105.609634" name="Pha Taem National Park" stateProvince="Ubon Ratchathani">Pha Taem National Park</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1AAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.609639,15.622750]}">
<geoCoordinate id="C4D798749A3C33BE896B9F0B1FFD307D" degrees="15" direction="north" minutes="37" orientation="latitude" precision="1" seconds="21.9" value="15.62275">15 ° 37 ' 21.9 &quot; N</geoCoordinate>
,
<geoCoordinate id="3C53731DBEE802455FB34DA3D73E69F0" degrees="105" direction="east" minutes="36" orientation="longitude" precision="1" seconds="34.7" value="105.609634">105 ° 36 ' 34.7 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="BA8B2897F74B79E5CAA7AB8FAE1F2503" metricMagnitude="2" metricUnit="m" metricValue="4.2" unit="m" value="420.0">
<quantity id="FFBA546FA3C6323ABB8D199AEF393D6B" metricMagnitude="2" metricUnit="m" metricValue="4.2" unit="m" value="420.0">420 m</quantity>
a. s. l.
</elevation>
; dry dipterocarp rainforest,
<collectingMethod id="ABA34AB8CE6D07008682CD3618F1F011">ethanol baited trap</collectingMethod>
;
<date id="261EE0680E9A266B25679B70B251054C" value="2019-05-01">
<collectingDate id="5B94549E5DF4B9E5EC83B2EE2996193D" value="2019-05-01">01. v. 2019</collectingDate>
</date>
, (
<collectorName id="5A2D78DB9BCCB8C9A14076F29E1443EF">W. Sittichaya</collectorName>
)
</materialsCitation>
<materialsCitation id="5030B88608D3D28E8ADAF5E12862F44B" collectionCode="THNHM" collectorName="Small &amp; W. Sittichaya" country="Thailand" elevation="600" latitude="17.671583" location="Tak Province" longLatPrecision="1" longitude="97.851166" specimenCount="1" stateProvince="Tak" typeStatus="paratype">
<collectingRegion id="AD7B74D0215D9B1D8183BDF1CCE9D9F3" country="Thailand" name="Tak">Tak Province</collectingRegion>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECBAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[97.851167,17.671583]}">
<geoCoordinate id="E65768A1D298A84F780A027D6508D540" degrees="17" direction="north" minutes="40" orientation="latitude" precision="1" seconds="17.7" value="17.671583">17 ° 40 ' 17.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="5EBB813C5FB3F3D13BAB5AA7CFCCC7B3" degrees="97" direction="east" minutes="51" orientation="longitude" precision="1" seconds="04.2" value="97.851166">97 ° 51 ' 04.2 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="D4B48B6FF14303B36AC37E131CBF7FE4" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity id="B4CE10570BF70D4B7F146394C5BAD6F9" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a. s. l.
</elevation>
; semiagricultural area, ex.
<collectorName id="F9791E5700C219C5895F490E8279FB7D">Small</collectorName>
branch of unknown tree; 08. ix. 19 (1), (
<collectorName id="CB02497CD8E6B91630AD88DF3BB5B292">W. Sittichaya</collectorName>
), (1
<abbrev id="ABBRID0EHBAI" xlink_title="Natural History Museum of the National Science Museum">
<collectionCode id="DFBF15911941C976426CC9D7017B4864">THNHM</collectionCode>
</abbrev>
)
</materialsCitation>
<materialsCitation id="A9C4F5C13218F2FC1772443B6D89DBE2" collectionCode="WSTC" collectorName="W. Sittichaya" country="Thailand" elevation="600" latitude="17.558222" location="Dry Dipterocarp forest" longLatPrecision="1" longitude="98.87945" municipality="Maeping National Park" specimenCount="1" stateProvince="Lamphun" typeStatus="paratype">
<collectingRegion id="31B1DA4927852210606A62AA7457D936" country="Thailand" name="Lamphun">Lamphun Province</collectingRegion>
,
<collectingMunicipality id="96F7E929119CDDD8F8974E276B46C9A2">Maeping National Park</collectingMunicipality>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOBAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[98.879444,17.558222]}">
<geoCoordinate id="B9B4736B99FA57B7A61C8121339D211A" degrees="17" direction="north" minutes="33" orientation="latitude" precision="1" seconds="29.6" value="17.558222">17 ° 33 ' 29.6 &quot; N</geoCoordinate>
,
<geoCoordinate id="2AD2A062E9824CA32BF36FBB0D67D81D" degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="46.0" value="98.87945">98 ° 52 ' 46.0 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="863FB5F76C91F904D045281A0A8C2034" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity id="67C940B1B9AE08590901F3852EC8E0A8" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a. s. l.
</elevation>
;
<location id="92DFE4800B9F5887A1A74D8A094FE632" LSID="urn:lsid:plazi:treatment:F2E0A6A18C5F5F878C6CBB465C1AB33D:92DFE4800B9F5887A1A74D8A094FE632" country="Thailand" latitude="17.558222" longLatPrecision="1" longitude="98.87945" municipality="Maeping National Park" name="Dry Dipterocarp forest" stateProvince="Lamphun">Dry Dipterocarp forest</location>
,
<collectingMethod id="CCF469C0200C7530AE0618AD1007D62D">ethanol baited trap</collectingMethod>
; 01. ii. 19 (1), 01. iii. 19 (1), 01. v. 19 (1) (all
<collectorName id="4978FCA02EA8B4B34C17B94463EB2ADF">W. Sittichaya</collectorName>
) (3
<abbrev id="ABBRID0ETBAI" xlink_title="Private collection of Wisut Sittichaya">
<collectionCode id="9E8DE447E684FB579B7D54719A362B69">WSTC</collectionCode>
</abbrev>
)
</materialsCitation>
<materialsCitation id="8D888D1DC736EB61E47FB5540EB11B4B" collectionCode="MSUC" collectorName="W. Sittichaya" country="Thailand" elevation="600" latitude="17.558222" location="Chiang Dao Wildlife Sanctuary" longLatPrecision="1" longitude="98.87945" specimenCount="1" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingRegion id="A0FBF2C3C8783AEEC2D9CF341EE70F80" country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
,
<location id="9D3851E6B7845F18F586FB98D83335CC" LSID="urn:lsid:plazi:treatment:F2E0A6A18C5F5F878C6CBB465C1AB33D:9D3851E6B7845F18F586FB98D83335CC" country="Thailand" latitude="17.558222" longLatPrecision="1" longitude="98.87945" name="Chiang Dao Wildlife Sanctuary" stateProvince="Chiang Mai">Chiang Dao Wildlife Sanctuary</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1BAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[98.879444,17.558222]}">
<geoCoordinate id="5BD69A93F0395D73EA9F4503F263F29D" degrees="17" direction="north" minutes="33" orientation="latitude" precision="1" seconds="29.6" value="17.558222">17 ° 33 ' 29.6 &quot; N</geoCoordinate>
,
<geoCoordinate id="D336F98768D92C2C1715E43B47C9CF21" degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="46.0" value="98.87945">98 ° 52 ' 46.0 &quot; E</geoCoordinate>
</named-content>
;
<elevation id="BEFD8CE6951ABAFBBCEDA6148F9F8572" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity id="C870140E437DBBA14AEEE9184E70B1D4" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a. s. l.
</elevation>
; mixed deciduous forest,
<collectingMethod id="AFE254ABE7B74623559F8192FFA6C93D">ethanol baited trap</collectingMethod>
; 01. vi. 19 (
<collectorName id="49DE8385FBAA59F2F4DFC9F85E975724">W. Sittichaya</collectorName>
) (1
<abbrev id="ABBRID0E6BAI" xlink_title="Albert J. Cook Arthropod Research Collection, Michigan State University">
<collectionCode id="3C3A7028AB9FC0FF0E26DE4AC2FD57F9">MSUC</collectionCode>
</abbrev>
)
</materialsCitation>
.
</paragraph>
<caption id="8FB48488AFD587531BA4EA88A965C38E" ID-DOI="10.3897/zookeys.1217.129707.figure3" ID-arpha="15CE74E5-874E-59A5-94A1-873E00B64A9C" httpUri="https://binary.pensoft.net/fig/1168131" startId="F3">
<paragraph id="EDF1929AB27B60E18CBE9153A16B0661">
<label id="DFBC21883734EAC8E9C2D51D082C3056">Figure 3.</label>
</paragraph>
<paragraph id="8F3AA74FCDCF55B17C476EDDDB3F4C8F">
<taxonomicName id="F630264F3F778D613D1C3537CAD11253" authorityName="Sittichaya &amp; Lin &amp; Smith &amp; Pornsuriya &amp; Cognato" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eccoptopterus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="8FC371E201DC927B8D01AB3BAA8CA8C3" italics="true">Eccoptopterus intermedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC7196294C28354F7D42D7E4570E2176" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, female,
<emphasis id="B08721C1E7E20B1C32AE561755DF8754" bold="true">A</emphasis>
dorsal view
<emphasis id="0D0F6DF869F54486C612812B988E1850" bold="true">B</emphasis>
lateral view
<emphasis id="B4CDA6362E17E2386E40E9F1DD4518BB" bold="true">C</emphasis>
posterolateral view
<emphasis id="733C4B502AABC23557715D9769AA0285" bold="true">D</emphasis>
frons
<emphasis id="58041D67EDFDFC5870A2E4C41F2C4790" bold="true">E</emphasis>
antenna.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EFDAI" type="diagnosis">
<paragraph id="F3181007713F09A3A4F5BF4C1C44F1CC">
<heading id="E0298FE77F591155176A8E83A7F84BC3" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D4DC5F2C0C6550335E0687A5E9756586">
<emphasis id="C3EDBAC60FFECB7DA11B176E34AD09C1" bold="true">Female</emphasis>
,
<quantity id="EAB574DAE908EE555417BC9E8E71E636" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.70-1.90 mm</quantity>
long (mean =
<quantity id="0641A9BBA034703CD2ADBEAB73DC1DF8" metricMagnitude="-3" metricUnit="m" metricValue="1.8" unit="mm" value="1.8">1.80 mm</quantity>
;
<emphasis id="037D36310E43269113E5C88E917EDEFF" italics="true">N</emphasis>
= 6), 2.03-2.38 × as long as wide (mean = 2.13 ×;
<emphasis id="C94A7EB7D43A3766783C7F1BAD5EFEE2" italics="true">N</emphasis>
= 6). Small body size, declivital armature composed of a pair of major spines at interstriae 3 on declivital summit and four minor spines unevenly spaced on each lateral margin, declivity covered with flattened scale-like setae; protibiae slender, broadest at apical 1 / 3, outer margin armed with four or five moderated socketed denticles, elytra tapering laterally.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERDAI" type="description">
<paragraph id="FA26DA90809D0BF5395A3D07F0F5EFF2">
<heading id="A24F250A89FB1C846D59F922EFEAD083" reason="title">Description.</heading>
</paragraph>
<paragraph id="35256DFEDFC002C95D0B11611E8B0D52">
<emphasis id="C0B13E1C93B6CFD5D68D8B07EC4718A1" bold="true">Female</emphasis>
. Body brown, dark brown to black, impressed portion of elytral disc and declivital face paler and bearing grayish-brown scale-like setae; antennae, prolegs, middle legs and associated coxae paler brown, hind legs dark brown to black.
<emphasis id="B080F89D3340EFDC94ACDE30738C8438" bold="true" italics="true">Head</emphasis>
: epistoma complete, margin bisinuated, with a row of hair-like setae. Frons below upper margin of eye and above epistoma impressed, without raised median line, surface reticulate, subshiny, sparsely covered with fine long setae, setal insertion shallowly punctate. Frons below upper portion of the eye slightly convex. Eye shallowly emarginate just above antennal insertion, upper portion slightly smaller than lower part. Submentum triangular small, shallowly impressed. Antennal scape long, normal thick, slightly longer than club (9: 8). Pedicel as broad as scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club obliquely truncate, longer than wide (8: 6.5),
<typeStatus id="B6160567B330E074D6514CA76EB724AB">type</typeStatus>
1, segment 1 corneous, occupying basal 1 / 4, margin carinate, concave, encircling in anterior face, segment 2 and 3 soft, visible on anterior face only.
<emphasis id="C85B66996A4CE5ED15F18EDFFE805D6B" bold="true" italics="true">Pronotum</emphasis>
: 0.93-0.97 (mean = 0.95,
<emphasis id="835F03A87B7BAF26EEDAABBF3C10CE31" italics="true">N</emphasis>
= 6) × as long as wide, round shorter than long,
<typeStatus id="2996C80E43F76EF12A16FACF86AB9F33">type</typeStatus>
<quantity id="D740D9EB88C7D673C9BF22F27298F2F3" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
dorsal view, lateral sides parallel to anterior middle, broadly rounded anteriorly; anterior margin with 2-4 serrations, median pair prominent; anterior slope strongly asperate, asperities densely spaced, rugose, becoming lower and more transverse toward the summit; disc slightly convex, finely alutaceous, dull, sparsely covered with fine punctures bearing fine short hair-like setae. Base with a tuft of short hair-like setae associated with mycangium. In lateral view short and tall,
<typeStatus id="02B2457C1473521D259D8FA6AA06BE1C">type</typeStatus>
3, summit at middle, lateral margins obliquely costate.
<emphasis id="E3DA33B3E9077A1227570CBBCED5EE71" bold="true" italics="true">Elytra</emphasis>
: 1.12-1.18 (mean = 1.15,
<emphasis id="A8469F6D92EA321C8D5B2E076563BE65" italics="true">N</emphasis>
= 6) × as long as wide, 1.07-1.19 (mean = 1.15,
<emphasis id="604C6468AE3283B0C9BB8D12872E384C" italics="true">N</emphasis>
= 6) × as long as pronotum. Scutellum comparatively moderately sized, narrow, linguiform, finely punctate, subshiny, attachment on anterior slope of elytra less visible from above. Base shallowly bisinuate, with oblique edge, humeral angles rounded, lateral side tapering from humeral angle to apex. Disc short, basal area ¼ of disc slightly convex, apical 3 / 4 impressed and connecting to declivital impression; disc punctate, punctures fine confused and setose, strial setae uniseriate with long, erect hair-like setae; interstrial setae bi- or triseriate with semi-recumbent hair-like setae. Impressed portion of disc covered with leaf-like setae. Declivity sulcate, with a pair of major (largest) spines on declivital summit and four much smaller minor spines on declivital margin, first minor spine located far from the major spine; striae and interstriae punctate, punctures small and shallow; striae 1 shallowly impressed, 2-3 flattened; interstriae 1 slightly convex, 2 and 3 flattened. Striae and interstriae with flattened bristle-like setae, setae semi-recumbent, near median suture in vertical rows (34 rows on each side), apically pointed, near lateral margins (four or five rows per side) pointed inwardly to elytral suture, at apical margin without upwardly setae.
<emphasis id="B0AABF4BAC71F0245C9BD98DBD526D2F" bold="true" italics="true">Legs</emphasis>
: procoxae contiguous; prosternal coxal piece short, inconspicuous. Protibiae slender, broadest at middle; posterior face inflated, punctate, densely covered with long hair-like setae; outer margin armed with four or five moderate socketed denticles. Meso- and metatibiae rounded, flat, mesotibiae armed with three or four smaller socketed denticles, metafemora and metatibiae enlarged, the latter without spine.
</paragraph>
<paragraph id="1CF9CD1622E65FD425F6AA7E79697981">
<emphasis id="157F2F7101A84616DBA97FCEB26AE5F7" bold="true">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPEAI" type="etymology">
<paragraph id="85D893920B35AC543DDAD62A45EE0166">
<heading id="494E20026587DAD396360ED7ABA27778" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3B86D9A0369828F6DE210D5BF9C30336">
L.
<emphasis id="E94E62EA63A1E80286EB650E07A35BA9" italics="true">inter</emphasis>
+
<emphasis id="E6B81A3F6B26C87681E2197406CD2BD2" italics="true">medius</emphasis>
= in the middle. The name refers to the morphological characters of the species which lie between those of
<taxonomicName id="448D4DA130881AAF0DE792FFD0548B30" class="Insecta" family="Curculionidae" genus="Eccoptopterus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="limbus">
<emphasis id="7F063779277DFAFFC6705E09121137D2" italics="true">E. limbus</emphasis>
</taxonomicName>
and
<taxonomicName id="95A9BFB3C4673AECB053BA73B214A713" authorityName="Olivier" authorityYear="1800" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="7493B9583A4716845687E9C5F7F5B4CB" italics="true">E. spinosus</emphasis>
</taxonomicName>
. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPFAI" type="distribution">
<paragraph id="E3B8C959EAB3A77D58515F57D45569D2">
<heading id="1F7864CE079CEA7F0011E646928F620A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B92B2359167B7D8694ADE9DA4DA92D4B">
<collectingCountry id="D6FA50FFFC794608BB15057BE81A08E4" name="Thailand">Thailand</collectingCountry>
(
<collectingRegion id="8D58F565F4E0902EDAE5AB4100DE1B72" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
,
<collectingRegion id="DED3070FEF577E6D4F2F3F6180087557" country="Thailand" name="Lamphun">Lamphun</collectingRegion>
,
<collectingRegion id="6199899F1F9972C500398F01D9B54FD2" country="Thailand" name="Tak">Tak</collectingRegion>
,
<collectingRegion id="8C79CD43B2855444352EA22A0046E866" country="Thailand" name="Ubon Ratchathani">Ubon Ratchathani</collectingRegion>
provinces).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUFAI" type="host">
<paragraph id="8E59B4C900A6307E3F8531FAF087C440">
<heading id="7940ADCB4173906E79DD9B06223A4F89" reason="title">Host plants.</heading>
</paragraph>
<paragraph id="F6575D2D879E61C0FF25FD7D0C058735">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>