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<mods:title id="E4358C59E7AB7A2DCAE2169BEADEBA37">A taxonomic revision of the Neotropical myrmicine ant genus Lachnomyrmex Wheeler (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="F6548DABE53CD234C81A9288163F821A">Feitosa, R. M.</mods:namePart>
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<mods:namePart id="430F59D3DA338E5FA9B88BCDE82EFCA0">Brandão, C. R. F.</mods:namePart>
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<treatment id="FCE3104571C02C962A5A11BC2EF4C715" ID-DOI="http://doi.org/10.5281/zenodo.6234231" ID-GBIF-Taxon="125134993" ID-Zenodo-Dep="6234231" LSID="urn:lsid:plazi:treatment:FCE3104571C02C962A5A11BC2EF4C715" httpUri="http://treatment.plazi.org/id/FCE3104571C02C962A5A11BC2EF4C715" lastPageNumber="19" pageNumber="17">
<subSubSection id="989FDD58C993129365F5281A54D382A7" pageNumber="17" type="nomenclature">
<paragraph id="C0C1FF7304CD8D3FF28E759E4244F416" pageNumber="17">
<treatmentCitation id="D78A974DFC99A03D3D6140B097C43CC8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" author="Smith, M. R." authority="Smith, 1944" authorityName="Smith" authorityYear="1944" genus="Lachnomyrmex" journalOrPublisher="Proceedings of the Entomological Society of Washington" lsidName-HNS="Lachnomyrmex haskinsi Smith" pageNumber="17" pagination="225 - 228" part="46" rank="species" species="haskinsi" title="The genus Lachnomyrmex, with the description of a second species" treatmentTaxon="Lachnomyrmex haskinsi" type="journal article" year="1944">
<taxonomicName id="610D68490A62161C99DC39114CF1404B" ID-CoL="3RPV9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" authority="Smith, 1944" authorityName="Smith" authorityYear="1944" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex haskinsi Smith" order="Hymenoptera" pageNumber="17" phylum="Arthropoda" rank="species" species="haskinsi">
Lachnomyrmex haskinsi
<bibRefCitation id="95F4599FD2DDBF19115B03DC73D3CC2F" author="Smith, M. R." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageNumber="49" pagination="225 - 228" part="46" title="The genus Lachnomyrmex, with the description of a second species" type="journal article" year="1944">
<authority id="805C7356A0A72B41F88742E6F39295C8" author="Smith, M. R." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageNumber="49" pagination="225 - 228" part="46" title="The genus Lachnomyrmex, with the description of a second species" type="journal article" year="1944">Smith, 1944</authority>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="27B314ED123DC91451D544A19A0E0FFD" pageNumber="17" type="description">
<paragraph id="005FE800066BA31900FC83A1679290E1" pageNumber="17">Figures 5, 19</paragraph>
</subSubSection>
<subSubSection id="D77EA411CDDC103AD339524EA3026567" pageNumber="17" type="materials_examined">
<paragraph id="AD9EFD50AF6B8A524DF0FBE628DD6812" pageNumber="17">
<treatmentCitation id="3269FAB89BC00EF3547F326AFF353CDC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" author="Smith, M. R." authority="Smith, 1944" authorityName="Smith" authorityYear="1944" genus="Lachnomyrmex" journalOrPublisher="Proceedings of the Entomological Society of Washington" lsidName-HNS="Lachnomyrmex haskinsi Smith" pageNumber="17" pagination="225 - 228" part="46" rank="species" species="haskinsi" title="The genus Lachnomyrmex, with the description of a second species" treatmentPage="227" treatmentTaxon="Lachnomyrmex haskinsi" type="journal article" year="1944">
<taxonomicName id="BDBEB7D09BE0AD73477FF302311CE70B" ID-CoL="3RPV9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" authority="Smith, 1944" authorityName="Smith" authorityYear="1944" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex haskinsi Smith" order="Hymenoptera" pageNumber="17" phylum="Arthropoda" rank="species" species="haskinsi">
Lachnomyrmex haskinsi
<bibRefCitation id="041E47BEADE32145FFCF7677F187663D" author="Smith, M. R." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageNumber="49" pagination="225 - 228" part="46" title="The genus Lachnomyrmex, with the description of a second species" type="journal article" year="1944">
<authority id="EE64D6C19B5C1DADA5E2FCA776DD1246" author="Smith, M. R." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageNumber="49" pagination="225 - 228" part="46" title="The genus Lachnomyrmex, with the description of a second species" type="journal article" year="1944">Smith, 1944</authority>
</bibRefCitation>
</taxonomicName>
: 227
</treatmentCitation>
, figs. 1-2.
<materialsCitation id="19CFD55B9A013838DD3DDB9BF3B747A4" ID-GBIF-Occurrence="1416638555" collectingDate="1994-05" collectionCode="USNM" collectorName="J. Zetek" country="Panama" location="Barro Colorado Island" stateProvince="Canal Zone" typeStatus="Holotype">
<typeStatus id="E71135848CA687FD6AA5279A667EF194">Holotype</typeStatus>
worker,
<collectingCountry id="0E2CE5D5CD644008A6DAACA8ACFBC598">Panama</collectingCountry>
:
<collectingRegion id="D89E9B9501D63FD8494C3C47DE311EF1">Canal Zone</collectingRegion>
,
<location id="8AD660EFA404A0C65617935A8E113DE7" LSID="urn:lsid:plazi:treatment:FCE3104571C02C962A5A11BC2EF4C715:8AD660EFA404A0C65617935A8E113DE7" country="Panama" name="Barro Colorado Island" stateProvince="Canal Zone">Barro Colorado Island</location>
,
<collectingDate id="5B87EC781E94E2743EB534B16598B6EC">iiiii.1994</collectingDate>
,
<collectorName id="748FEF2803BE307C1FA3C79AF25DED25">J. Zetek</collectorName>
col., no. 5121/56906 [
<collectionCode id="1F117D4C990BF531660D2390F998BFA8" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageNumber="17">USNM</collectionCode>
] (examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC241103A6FD491B5392F874E583AD6B" pageNumber="17" type="diagnosis">
<paragraph id="8ACFC273FC04278224BC3E7414379F77" pageNumber="17">Worker diagnosis. Body covered by continuous vermiculate rugae; petiolar node considerably elevated and subtriangular; dorsum of postpetiole with about six long hairs, and with a raised prominence medially; subpostpetiolar process present and well developed; first tergite of gaster devoid of long hairs.</paragraph>
</subSubSection>
<subSubSection id="53992823B4CA296A0F17E4BDA71C0B75" lastPageNumber="19" pageNumber="17" type="description">
<paragraph id="FAE84B68BBBC8466189662E769F45827" pageNumber="17">
<typeStatus id="82A5808A5676541FAFD528FE11CF322F">Holotype</typeStatus>
measurements. HL 0.60; HW 0.55; ML 0.22; SL 0.38; EL 0.09; WL 0.68; PSL 0.11; PL 0.24; PPL 0.13; GL 0.79; TL 2.65; CI 92; SI 68; OI 17.
</paragraph>
<paragraph id="6C963E24B8101A0503D1086833294833" pageNumber="17">Worker measurements (n=2). HL 0.61-0.65; HW 0.60-0.63; ML 0.21-0.24; SL 0.39; EL 0.11-0.13; WL 0.71-0.74; PSL 0.16-0.17; PL 0.25-0.30; PPL 0.13-0.14; GL 0.79-0.80; TL 2.70-2.94; CI 97-98; SI 62-66; OI 17-21.</paragraph>
<paragraph id="741CE13AFE988F40075B5F452C6BF767" pageNumber="17">Worker description. Color red ferruginous to brown, with lighter appendages. Head and mesosoma finely covered by continuous vermiculate rugae, transverse on anterior portion of pronotum, grading to longitudinal on the rest of promesonotum, mesopleura and lateral faces of propodeum; mandibles with few and short longitudinal striae restricted to basal portion; petiole and postpetiole finely and irregularly rugose. Abundant pilosity on head and dorsum of promesonotum; dorsum of petiolar node and postpetiole with about three and six long hairs, respectively; first tergite of gaster entirely devoid of long flexuous hairs.</paragraph>
<caption id="3E4A70B6A16B50D63D22D6F5C0A2EF71" pageNumber="18">
<paragraph id="0199AAACFB60A99CF6EF2C792C35190D" pageNumber="18">
<pageBreakToken id="9A0EA5C554614FC564EE449FECA68E9F" pageNumber="18" start="start">FIGURE</pageBreakToken>
5. Worker of
<taxonomicName id="25B4AC857B8F0E193B546D8A8C81954A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex haskinsi Smith" order="Hymenoptera" pageNumber="18" phylum="Arthropoda" rank="species" species="haskinsi">Lachnomyrmex haskinsi</taxonomicName>
from Heredia, Costa Rica: A, head in full face view; B, lateral view. Image by John Longino, specimen INBIOCRI001237374.
</paragraph>
</caption>
<paragraph id="EEA1B243743B6E2444113C5D889A4176" lastPageNumber="19" pageNumber="18">
Head longer than broad, with vertexal margin convex; frontal lobes slightly rounded laterally; eyes with six to seven facets at greatest diameter. Promesonotum evenly convex in profile; metanotal groove relatively narrow and deeply impressed; propodeal spines straight; teeth of propodeal lobes widen basally, with about
<pageBreakToken id="2FA036D592B4885409A07C254BAD8BCC" pageNumber="19" start="start">half-length</pageBreakToken>
of propodeal spines length. Petiolar node strongly elevated and subtriangular in lateral view; dorsum of postpetiole with a raised prominence medially and a well developed anteroventral projection directed anteriorly.
</paragraph>
<paragraph id="805DADF0A5E35F1507DDE2F17D3EDBA5" pageNumber="19">Gyne (first description). Differing from worker by having about 12 facets at eyes maximum diameter; dorsum of petiolar node and postpetiole with about five and 10 long hairs, respectively; first tergite of gaster with around 10 long flexuous hairs on the anterior third, near postpetiolar insertion (see picture in the &quot;Ants of Costa Rica&quot; website).</paragraph>
</subSubSection>
<subSubSection id="F62EBF3E0AC869D20FFC822420E24BEC" pageNumber="19" type="etymology">
<paragraph id="4672F426C6BA34F9CC8F26F242CA2F79" pageNumber="19">
Etymology. Dr. Marion M.
<bibRefCitation id="B92E0FD24E456982254885DB7AEE5210" author="Smith, M. R." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageNumber="49" pagination="225 - 228" part="46" title="The genus Lachnomyrmex, with the description of a second species" type="journal article" year="1944">Smith (1944)</bibRefCitation>
named this species in honor of Dr. Caryl Parker Haskins (1908- 2001), in recognition for his contribution to the study of ant biology.
</paragraph>
</subSubSection>
<subSubSection id="777FA845058B3BE9929C201B2EAE99D9" pageNumber="19" type="discussion">
<paragraph id="ED2551A3987461D7ECCF1DD36E06F18C" pageNumber="19">
Comments. This species is most similar to
<taxonomicName id="DDC6759452DDAEAB8DB820E10A9BD0BC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:245583" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex regularis Feitosa &amp; Brandão" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="regularis">Lachnomyrmex regularis</taxonomicName>
, but can be easily distinguished from the latter by the smaller size, more irregular rugulation, and presence of fine short rugae on postpetiole dorsum. Workers of
<taxonomicName id="7D03DF5361F234C89624ADF1D267B966" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:245576" class="Insecta" family="Formicidae" genus="Lachnomyrmex" genus.bestMatchDistance="4389" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex fernandezi Feitosa &amp; Brandão" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="fernandezi">L. fernandezi</taxonomicName>
and
<taxonomicName id="68FBED4AFC1544D15C86BA1A99CC61EC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:245580" class="Insecta" family="Formicidae" genus="Lachnomyrmex" genus.bestMatchDistance="3637" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex mackayi Feitosa &amp; Brandão" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="mackayi">L. mackayi</taxonomicName>
can also be confounded with
<taxonomicName id="27200A17051B545D2273059AB9249F2F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" class="Insecta" family="Formicidae" genus="Lachnomyrmex" genus.bestMatchDistance="22" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex haskinsi Smith" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="haskinsi">L. haskinsi</taxonomicName>
, but they present long flexuous hairs at least on the anterior third of the first gastral tergite.
</paragraph>
</subSubSection>
<subSubSection id="BD9CA63F84603CA16536D9C14297310D" pageNumber="19" type="biology_ecology">
<paragraph id="B64F479409B6B60EB6755F17AD56F657" pageNumber="19">
<taxonomicName id="186083555E1935DE49EB00B1F31D4D41" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29660" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Lachnomyrmex haskinsi Smith" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="haskinsi">Lachnomyrmex haskinsi</taxonomicName>
has been recorded from Costa Rica and Panama, in
<quantity id="7F0E99C28F20EBAD72C9C1552C804285" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" unit="m" value="500.0" valueMax="800.0" valueMin="200.0">200-800m</quantity>
elevation wet forests. Workers are usually collected in samples of sifted leaf litter. Dr. John Longino observed a few workers, one with a larva, scattered throughout the debris of a
<taxonomicName id="255DF7FA1D4692A0A53591B95D0FA53E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2407" class="Insecta" family="Formicidae" genus="Brachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Brachymyrmex Mayr" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="genus">Brachymyrmex</taxonomicName>
nest he was collecting from beneath rotten wood on the forest floor (
<bibRefCitation id="700EE21B972E5BBF06B7F1152011CF27" author="Longino, J. T." journalOrPublisher="Boletin de Entomologia Venezolana" pageNumber="49" publicationUrl="http://www.evergreen.edu/ants/AntsofCostaRica.html" title="Ants of Costa Rica" type="url" year="2007">Longino 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E6F79BBD151F14AC1EB42A317CE58752" pageNumber="19" type="materials_examined">
<paragraph id="DA49F4CD87E2D7921885EFE00F7299DF" pageNumber="19">
Additional material examined.
<materialsCitation id="F8E2C3850C906361AC263EF24D676FCB" ID-GBIF-Occurrence="1416638571" collectingDate="1990" collectionCode="BMNH" collectorName="P. Hammond" country="Costa Rica" location="Rio San Lorencito" stateProvince="Alajuela">
<collectingCountry id="F1FFDE0616569BE7FAB5F3F721D216F6">COSTA RICA</collectingCountry>
:
<collectingRegion id="815D9B657B2C671DF921580C78273A97">Alajuela</collectingRegion>
:
<location id="EA9F97B0F04758CB0F26F07448E1E864" LSID="urn:lsid:plazi:treatment:FCE3104571C02C962A5A11BC2EF4C715:EA9F97B0F04758CB0F26F07448E1E864" country="Costa Rica" name="Rio San Lorencito" stateProvince="Alajuela">Rio San Lorencito</location>
,
<collectingDate id="A0533FCCE89C57E90186FD2156EE9EB6">1990</collectingDate>
,
<collectorName id="2B57C3AE3B593D8501BEE406883E2029">P. Hammond</collectorName>
col. (1 worker) [
<collectionCode id="45BF1041AE36A17E2488407CC57764D8" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageNumber="19">BMNH</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="092DADCE3EA34EB5236367D70A12E7F1" ID-GBIF-Occurrence="1416638564" collectingDate="1986-07-15" collectionCode="INBC" collectorName="J. Longino" country="Costa Rica" latitude="10.716666" location="La Selva Biological Station" longLatPrecision="1" longitude="-84.01667" specimenCode="INBIOCRI 002280138" stateProvince="Heredia">
<collectingRegion id="AB2E35AE61AFB5B0B30E576ED5B70369">Heredia</collectingRegion>
:
<location id="87DEC1D843578CB52EF74602293E80C3" LSID="urn:lsid:plazi:treatment:FCE3104571C02C962A5A11BC2EF4C715:87DEC1D843578CB52EF74602293E80C3" country="Costa Rica" latitude="10.716666" longLatPrecision="1" longitude="-84.01667" name="La Selva Biological Station" stateProvince="Heredia">La Selva Biological Station</location>
, CCL,
<geoCoordinate id="06814CB2160F95867E7EF2052FD0DB7A" direction="north" orientation="latitude" pageNumber="19" precision="1" value="10.716666">10°43'N</geoCoordinate>
<geoCoordinate id="E2824D93BB5D65FE893D9E4E4BF3F066" direction="west" orientation="longitude" pageNumber="19" precision="1" value="-84.01667">84°01'W</geoCoordinate>
,
<date id="2B2B28702D04F20561926A0D7AA38C7D" value="1986-07-15">
<collectingDate id="D76BC86C16556CEB5BE1185534DE75B6" value="1986-07-15">15.vii.1986</collectingDate>
</date>
,
<collectorName id="A7F0B4EF6D36409B6CDBDB837A72DABF">J. Longino</collectorName>
col., no.
<specimenCode id="8045CB7F52CF769BD1F0841C48B494BB">INBIOCRI002280138</specimenCode>
(1 worker) [
<collectionCode id="AC18AC37F79CEBAA3BF42EC66DC2B940" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)" pageNumber="19">INBC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="D0B94D8AA718C850E8E9E3C6F2507DBB" ID-GBIF-Occurrence="1416638572" collectingDate="1991-10-23" collectionCode="INBC" collectorName="J. Longino" country="Costa Rica" latitude="10.716666" location="La Selva Biological Station" longLatPrecision="1" longitude="-84.01667" specimenCode="INBIOCRI 001237374" stateProvince="Heredia">
same locality,
<date id="CCA7AC08721B6F7CD15B8DEAE0302FBF" value="1991-10-23">
<collectingDate id="AEE64A7F664AAB2FB9A47A890ACC8673" value="1991-10-23">23.x.1991</collectingDate>
</date>
,
<collectorName id="98197FB8BE216DD3D9A156C584718C9D">J. Longino</collectorName>
col., no.
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col., no. INBIOCRI000900198 (1 worker) [
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.
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114
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<paragraph id="9A656C11BCD59D710670A900AB9EAB88" pageId="0" pageNumber="57889">Conservation status</paragraph>
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Least Concern for Italian (
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), Mediterranean (
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) and European assessments (
<bibRefCitation id="32609C534428D3F0D6D79EC4AC5A9DEA" author="Van Swaay, C." journalOrPublisher="Publications Office of the European Union, Luxembourg" pageId="0" pageNumber="57889" refId="B6009051" refString="Van Swaay, C., Cuttelod, A., Collins, S., Maes, D., Lpez Munguira, M., ai, M., Settele, J., Verovnik, R., Verstrael, T., Warren, M., Wiemers, M., Wynhof, I., 2010. European Red List of Butterflies. Publications Office of the European Union, Luxembourg" title="European Red List of Butterflies" year="2010">Van Swaay et al. 2010</bibRefCitation>
).
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It is distributed in North Africa, Europe, Turkey, Middle East and west Mongolia (
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). It can be found in mainland Italy, Sardinia and Sicily (
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).
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<paragraph id="BC01A2FBE1A737432E877DFE686A732E" pageId="0" pageNumber="57889">Notes</paragraph>
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Widespread in a wide range of habitats, often in arid and flower-rich meadows, mountain pastures and rugged fields from the plain to 2100 m a.s.l. It is also found in peatlands and wet meadows. Bivoltine species with generations in April-May, July-August, sometimes a third in October (
<bibRefCitation id="BEE0A419756B5A732E65B7A142205972" author="Paolucci, P." journalOrPublisher="WBA Handbooks" pageId="0" pageNumber="57889" pagination="1 - 480" refId="B6009042" refString="Paolucci, P., 2013. Butterflies and Burnets of the Alps and their larvae, pupae and cocoons. WBA Handbooks 4: 1 - 480" title="Butterflies and Burnets of the Alps and their larvae, pupae and cocoons" volume="4" year="2013">Paolucci 2013</bibRefCitation>
,
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,
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<bibRefCitation id="87E899B93CA37F8A420B00B650D0A45D" author="Flint Jr, O. S." journalOrPublisher="Amazoniana" pageId="0" pageNumber="1557" pagination="373 - 421" title="Studies of Neotropical Caddisflies XXII: Hydropsychidae of the Amazon Basin (Trichoptera)." volume="6" year="1978">Flint Jr 1978</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.263.4158" ID-GBIF-Dataset="5fe6c912-0997-4d09-800d-101a2a9c26e8" ID-PMC="PMC3591764" ID-Pensoft-Pub="1313-2970-263-59" ID-PubMed="23653517" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-263-59" ModsDocOrigin="ZooKeys 263" ModsDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" checkinTime="1451247727841" checkinUser="pensoft" docAuthor="Taylor, Christopher K." docDate="2013" docId="FCE33C33EACD89DA71EEDD796CBDB47F" docLanguage="en" docName="ZooKeys 263: 59-73" docOrigin="ZooKeys 263" docSource="http://dx.doi.org/10.3897/zookeys.263.4158" docTitle="Mangatangi parvum Taylor, 2013, sp. n." docType="treatment" docVersion="3" lastPageNumber="62" masterDocId="776AFFECFFB1FFC40947FF959858FFB7" masterDocTitle="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species" masterLastPageNumber="73" masterPageNumber="59" pageNumber="61" updateTime="1668155226950" updateUser="ExternalLinkService">
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<mods:title>Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species</mods:title>
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<mods:namePart>Taylor, Christopher K.</mods:namePart>
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<mods:date>2013</mods:date>
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<mods:number>263</mods:number>
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<mods:start>59</mods:start>
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<treatment ID-GBIF-Taxon="152040050" LSID="urn:lsid:plazi:treatment:FCE33C33EACD89DA71EEDD796CBDB47F" httpUri="http://treatment.plazi.org/id/FCE33C33EACD89DA71EEDD796CBDB47F" lastPageId="4" lastPageNumber="62" pageId="2" pageNumber="61">
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<paragraph pageId="2" pageNumber="61">
<taxonomicName LSID="urn:lsid:zoobank.org:act:F3982336-8FDC-49EF-B17D-E899B32F312D" class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">
<pageBreakToken pageId="2" pageNumber="61" start="start">Mangatangi</pageBreakToken>
parvum
</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="61">sp. n.</taxonomicNameLabel>
Figure 1
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="61" type="holotype">
<paragraph pageId="2" pageNumber="61">Holotype.</paragraph>
<paragraph pageId="2" pageNumber="61">
♂. New Zealand, AK. Mangatangi, Hunua Ra., 8
<normalizedToken originalValue="Feb–">Feb-</normalizedToken>
8 Mar 1977, I. Barton, ARA Kauri Seed Project, pit trap.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="61" type="paratypes">
<paragraph pageId="2" pageNumber="61">Paratypes.</paragraph>
<paragraph pageId="2" pageNumber="61">1 ♀, as for holotype; 1 ♂, Cuvier Is, July, R. Forster.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="61" type="etymology">
<paragraph pageId="2" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="61">
From the Latin parvus, small, in reference to its small size compared to other New Zealand
<taxonomicName lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Enantiobuninae">Enantiobuninae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="61" type="diagnosis">
<paragraph pageId="2" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="61">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
can be distinguished from all other New Zealand
<taxonomicName lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Enantiobuninae">Enantiobuninae</taxonomicName>
by the presence of a well developed tooth comb on the pedipalpal tarsal claw. It can be distinguished from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Monoscutum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monoscutum titirangiense" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="titirangiense">Monoscutum titirangiense</taxonomicName>
,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Acihasta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acihasta salebrosa" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="salebrosa">Acihasta salebrosa</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Templar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Templar incongruens" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="incongruens">Templar incongruens</taxonomicName>
by its relatively long legs and unsclerotised dorsum. It differs from
<normalizedToken originalValue="‘Megalopsalis’">'Megalopsalis'</normalizedToken>
triascuta, all
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
and most
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
species in the absence of either a mediodistal apophysis or hypersetose region on the pedipalpal patella, from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
species by its relatively bowed cheliceral fingers, and from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
species by the absence of denticles on the medial side of the pedipalp coxa.
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
can also be distinguished from all other New Zealand species, so far as is known, by its genital morphology: all other New Zealand species investigated to date have a relatively long glans that is either narrow in lateral view (most species) or possesses a distinct dorsal keel (
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
). The deep and short glans of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
is also distinct from that of
<taxonomicName lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Enantiobuninae">Enantiobuninae</taxonomicName>
elsewhere:
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Neopantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neopantopsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Neopantopsalis</taxonomicName>
species have a very elongate and relatively flat glans, and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Spinicrus</taxonomicName>
species have a short but also distally flattened glans. The only other
<taxonomicName lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Enantiobuninae">Enantiobuninae</taxonomicName>
in Australasia to possess comparatively deep glans are
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Australiscutum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Australiscutum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Australiscutum</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Tercentenarium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tercentenarium linnaei" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="linnaei">Tercentenarium linnaei</taxonomicName>
;
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
differs from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Australiscutum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Australiscutum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Australiscutum</taxonomicName>
in possessing relatively long legs and retaining an anterior grill of spines over the spiracle, and from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Tercentenarium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tercentenarium linnaei" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="linnaei">Tercentenarium linnaei</taxonomicName>
in lacking a large dorsolateral flange at the junction between shaft and glans. The glans of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Thrasychirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thrasychirus" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Thrasychirus</taxonomicName>
has never been illustrated in lateral view, but
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
is clearly distinguished from that genus by possessing paired bristle groups at the junction between shaft and glans rather than single bristles.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="description">
<paragraph pageId="2" pageNumber="61">Description.</paragraph>
<paragraph lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
Male (Figs 1
<normalizedToken originalValue="a–b">a-b</normalizedToken>
,
<normalizedToken originalValue="d–e">d-e</normalizedToken>
,
<normalizedToken originalValue="g–i">g-i</normalizedToken>
, l): Total body length 2.06-2.74 (larger value in all measurements represents holotype), prosoma length 0.97-1.19, prosoma width 1.76-2.01. Dorsal prosomal plate mostly light orange-yellow, unarmed except short, spinose black setae scattered over entire body; anterior propeltidium lighter yellow-cream, supracheliceral groove extending roughly halfway between anterior margin of carapace and ocularium; median propeltidium with diffuse purple stripes along border with anterior propeltidium with diffuse silver-white markings behind purple stripes, dark brown markings on lateral edge of dorsal prosomal plate; ocularium silver with black stripes margining eyes, unarmed; postocularium not distinguished from remainder of posterior propeltidium. Mesopeltidium forming raised ridge, medially pale yellow, laterally dark brown. Ozopores on raised
<pageBreakToken pageId="3" pageNumber="62" start="start">lateral</pageBreakToken>
lobes, anterior lobes of prosoma and ozopore lobes dark brown, posterior of ozopore lobes silver-white, remainder of lateral shelves mostly yellow with dark brown lateral margins broadening to diffuse dark brown patch at about three-quarters of distance from front of prosoma. Metapeltidium and dorsum of opisthosoma with background colour of purple broken by pale yellow mottling, particularly along segment boundaries, longitudinal mediolateral broken stripes of silver-white present as well as longitudinal medial rows of silver-white spots, sides of opisthosoma with purple background heavily broken by pale yellow punctations. Mouthparts white; coxae proximally pale yellow; coxae I and II distally with purple mottling, coxae III and IV with dark yellow-brown mottling laterally; genital operculum pale yellow; venter of opisthosoma mottled light purple with pale yellow stripes along segment boundaries.
</paragraph>
<paragraph pageId="3" pageNumber="62">Chelicerae: Segment I 2.85-3.51, segment II 3.82-4.62. Segment I ventrally cream, dorsally orange-yellow, sparsely denticulate dorsally; segment II inflated, orange-yellow, densely dorsally and sparsely ventrally denticulate. Cheliceral fingers (Fig. 1d) long, bowed, movable finger with setae close to median tooth.</paragraph>
<paragraph pageId="3" pageNumber="62">Pedipalps: Femur 1.53-2.13, patella 0.65-0.77, tibia 0.71-1.03, tarsus 1.80-2.47. Coxae unarmed. Femur to tarsus long, slender, unarmed, femur to tibia cream with paler distal ends to each segment, tarsus off-white with yellow-brown shading at distal end. Patella and tibia (Fig. 1e) straight, patella without distal prolateral apophysis or hypersetose region. Plumose setae absent. Microtrichia on distal half of tarsus only. Claw with ventral tooth-comb.</paragraph>
<paragraph pageId="3" pageNumber="62">Legs: Leg I femur 2.99-3.80, patella 0.71-0.91, tibia 2.93-3.87; leg II not preserved; leg III 2.56-3.38, patella 0.77-0.93, tibia 2.70-3.45; leg IV femur 4.05-5.01, patella and tibia not preserved. All segments unarmed. Trochanters pale yellow, trochanters III and IV with dark yellow-brown mottling laterally. Femora to tarsi pale yellow, patellae and distal ends of femora and tibiae darkening to orange-yellow. Leg II not preserved; tibia IV with three pseudosegments.</paragraph>
<paragraph pageId="3" pageNumber="62">
Penis (Figs 1
<normalizedToken originalValue="g–i">g-i</normalizedToken>
): Glans noticeably short and deep, sides parabolic in ventral view. Bristle groups of medium length. Tendon short, not extending far behind bristle groups.
</paragraph>
<paragraph pageId="3" pageNumber="62">Spiracle (Fig. 1l): Curtain of distally anastomosing spines extending over entire spiracle; shortening to cluster of tubercles (possibly lace tubercles) at medial corner.</paragraph>
<paragraph pageId="3" pageNumber="62">
Female (Figs 1c, f,
<normalizedToken originalValue="j–k">j-k</normalizedToken>
): Coloration similar to that of male. Other features as for male except for following: Chelicerae not enlarged, unarmed, segment I without ventral spine. Pedipalp (Fig. 1f) with microtrichia over entire patella, tibia and tarsus except glabrous dorsal line on patella and tibia. Ovipositor (Figs 1
<normalizedToken originalValue="j–k">j-k</normalizedToken>
) with single pair of seminal receptacles.
</paragraph>
<caption pageId="3" pageNumber="62">
<paragraph pageId="3" pageNumber="62">
Figure 1.
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
. A Holotype, dorsal view B holotype, lateral view C female, dorsal view D holotype, cheliceral fingers, anterior view E holotype, patella and tibia of left pedipalp, dorsal view F female, patella and tibia of right pedipalp, dorsal view G penis, right lateral view H glans, ventral view I glans, left lateral view J ovipositor K close-up of seminal receptacles L left spiracle of female.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="phylogeny">
<paragraph pageId="3" pageNumber="62">Phylogeny.</paragraph>
<paragraph lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
is probably related to the clade formed by
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
, with which it shares the presence of sharp papillae on the glans, and of setae close to the major tooth of the mobile finger of the chelicera (this last feature is also present in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Neopantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neopantopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Neopantopsalis</taxonomicName>
). The retention in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
of a plesiomorphic tooth-comb on the pedipalpal tarsal claw, together with
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
's distinctly short glans, could suggest a sister relationship between
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
and the
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="4" pageNumber="63" start="start">Pantopsalis</pageBreakToken>
</taxonomicName>
+
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
clade, but this should perhaps be treated with caution.
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis rennelli" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="rennelli">Pantopsalis rennelli</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis cheliferoides" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="cheliferoides">Pantopsalis cheliferoides</taxonomicName>
each retain reduced teeth arrays (a single tooth in the latter species) on the tarsal claw, and that of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis albipalpis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="albipalpis">Pantopsalis albipalpis</taxonomicName>
has a ventral rugose area that may correspond to the remains of the tooth-row. The loss of the tooth-row in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
has therefore happened at least partially in parallel. As regards the short glans of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi parvum" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="parvum">Mangatangi parvum</taxonomicName>
compared to the long glans of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
, our understanding of enantiobunine phylogeny is not yet robustly resolved (
<bibRefCitation pageId="4" pageNumber="63">Taylor 2011</bibRefCitation>
) and it is questionable which state is plesiomorphic for the clade.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ENCODING="UTF-8" ID-DOI="https://doi.org/10.5962/bhl.title.542" ID-GBIF-Dataset="57ea6fdb-2533-44e5-b60b-736758a88d19" ID-Zenodo-Dep="3922206" ID-ZooBank="2C6327E1-5560-4DB4-B9CA-76A0FA03D975" checkinTime="1593218702860" checkinUser="admin" docAuthor="Linnaeus, Carolus" docDate="1758" docId="FCE3C4F4E3542715E13DEBA4D4AFFD00" docLanguage="de" docName="Linnaeus1758Treatments.xml" docOrigin="Stockholm: Laurentius Salvius" docSource="https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf" docTitle="Holothuria Linnaeus, 1758, gen. nov." docType="treatment" docUuid="4F57143A-CCE4-45E0-B9C3-CFA62CB07EED" docUuidSource="ZooBank" docVersion="13" lastPageNumber="657" masterDocId="3F896AFC21C459639D0D71D9EDB8875C" masterDocTitle="Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" pageNumber="657" updateTime="1643600433100" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Linnaeus, Carolus</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:dateIssued>1758</mods:dateIssued>
<mods:publisher>Laurentius Salvius</mods:publisher>
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<mods:placeTerm>Stockholm</mods:placeTerm>
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<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3917852" ID-GBIF-Taxon="164807582" ID-Zenodo-Dep="3917852" LSID="urn:lsid:zoobank.org:act:4F57143A-CCE4-45E0-B9C3-CFA62CB07EED" httpUri="http://treatment.plazi.org/id/FCE3C4F4E3542715E13DEBA4D4AFFD00" lastPageNumber="657" pageNumber="657" sourcePageUrl="https://www.biodiversitylibrary.org/page/727568">
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<paragraph pageNumber="657">
<taxonomicName LSID="urn:lsid:zoobank.org:act:4F57143A-CCE4-45E0-B9C3-CFA62CB07EED" authorityName="Linnaeus" authorityYear="1758" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" phylum="Echinodermata" rank="genus" status="gen. nov." zbkClass="Vermes" zbkKingdom="Animalia" zbkOrder="Mollusca">Holothuria</taxonomicName>
[
<taxonomicNameLabel rank="genus">gen. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="657">
<emphasis italics="true">
<pageBreakToken pageNumber="657">Corpus</pageBreakToken>
</emphasis>
gibbum, nudum, ovale, natans.
</paragraph>
<paragraph pageNumber="657">
<emphasis italics="true">Tentacula</emphasis>
saepius ad alteram extremitatem, inaequalia numero &amp; figura.
</paragraph>
</subSubSection>
</treatment>
</document>