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<document id="6AFF352DCE37944D01468D06BA7E86D1" ID-CLB-Dataset="28472" ID-DOI="http://dx.doi.org/10.3897/zse.90.8705" ID-GBIF-Dataset="84f14a0e-f46f-420a-b891-854e3f88f9bd" ID-Pensoft-Pub="1860-0743-2-249" ID-ZooBank="CC6E4851C6FD4B47973E3E045390B579" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1860-0743-2-249" ModsDocOrigin="Zoosystematics and Evolution 90 (2)" ModsDocTitle="A new species of nocturnal gecko (Paroedura) from karstic limestone in northern Madagascar" checkinTime="1552154789851" checkinUser="pensoft" docAuthor="Glaw, Frank, Roesler, Herbert, Ineich, Ivan, Gehring, Philip-Sebastian, Koehler, Joern &amp; Miguel Vences," docDate="2014" docId="FCBD372AB8C1713D7EAB600063746EED" docLanguage="en" docName="ZoosystEvol 90(2): 249-259" docOrigin="Zoosystematics and Evolution 90 (2)" docSource="http://dx.doi.org/10.3897/zse.90.8705" docTitle="Paroedura hordiesi Glaw, Roesler, Ineich, Gehring, Koehler &amp; Vences, 2014, sp. n." docType="treatment" docUuid="8F5AA095-C68A-43E6-BCF8-4A02815A693F" docUuidSource="ZooBank" docVersion="7" lastPageNumber="255" masterDocId="FF8DA90E1C077258FF84FF89FFE72D64" masterDocTitle="A new species of nocturnal gecko (Paroedura) from karstic limestone in northern Madagascar" masterLastPageNumber="259" masterPageNumber="249" pageNumber="249" updateTime="1701382157146" updateUser="ExternalLinkService">
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<mods:title id="044C10CC6A4E1EBB6D43F291698A8120">A new species of nocturnal gecko (Paroedura) from karstic limestone in northern Madagascar</mods:title>
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<mods:namePart id="A66F1A0DF77DCF46AEE88436A22E1756">Glaw, Frank</mods:namePart>
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<mods:namePart id="EA63E6E4A669498934B30CA5A79F80F6">Roesler, Herbert</mods:namePart>
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<mods:namePart id="F9C17E1F4C1A191161D197EE61D8F855">Ineich, Ivan</mods:namePart>
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<mods:namePart id="415B828DE84C2082B3BF2CF60F003400">Gehring, Philip-Sebastian</mods:namePart>
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<mods:namePart id="0ABA7ADBE0C14A2704705650E6402A70">Miguel Vences,</mods:namePart>
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<mods:date id="2C312CC2C65AEB6219FC9E6C452A72AA">2014</mods:date>
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<treatment id="FCBD372AB8C1713D7EAB600063746EED" ID-GBIF-Taxon="154365922" LSID="urn:lsid:zoobank.org:act:8F5AA095-C68A-43E6-BCF8-4A02815A693F" httpUri="http://treatment.plazi.org/id/FCBD372AB8C1713D7EAB600063746EED" lastPageId="6" lastPageNumber="255" pageId="0" pageNumber="249">
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<paragraph id="89F5DAD4CD54C6BEDE0A69E0D8D2B17B" pageId="0" pageNumber="249">
<taxonomicName id="DC1D598352AD63D33C08E85068420FFC" ID-CoL="4DR4R" LSID="http://zoobank.org/8F5AA095-C68A-43E6-BCF8-4A02815A693F" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" order="Squamata" pageId="0" pageNumber="249" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
<taxonomicNameLabel id="14EC586F73B0CA8CF734F56E06D82142" pageId="0" pageNumber="249">sp. n.</taxonomicNameLabel>
Figs 2, 3, 4, Table 1
</paragraph>
</subSubSection>
<subSubSection id="50B4F614025058757D3D6ED991F5C1DF" lastPageId="1" lastPageNumber="250" pageId="0" pageNumber="249" type="remarks">
<paragraph id="06D8BD9AC0A68CB8B72E4A7A4101C5CD" pageId="0" pageNumber="249">Remarks.</paragraph>
<paragraph id="850EA118B69E0F5C682B0ABAD17ED9F6" lastPageId="1" lastPageNumber="250" pageId="0" pageNumber="249">
This species has been treated or figured under the name
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<pageBreakToken id="27E4BA291C0C4E6151467D25C6674DBE" pageId="1" pageNumber="250" start="start">Paroedura</pageBreakToken>
homalorhinus
</taxonomicName>
(
<bibRefCitation id="15A35E6B71CC2F5BBADE5BF5948D5635" author="Henkel, FW" journalOrPublisher="Ulmer Verlag, Stuttgart" pageId="7" pageNumber="256" title="Amphibien und Reptilien Madagaskars, der Maskarenen, Seychellen und Komoren." year="1995">Henkel and Schmidt 1995</bibRefCitation>
),
<taxonomicName id="5F41886CFCC768DBCCA97B7A50821030" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="karstophila">Paroedura karstophila</taxonomicName>
(
<bibRefCitation id="018C7DAC60BAD3B1515F776C4746F263" author="Glaw, F" journalOrPublisher="Spixiana" pageId="7" pageNumber="256" pagination="249 - 256" title="A new species of Paroedura Guenther from northern Madagascar (Reptilia, Squamata, Gekkonidae)." volume="24" year="2001">Glaw et al. 2001</bibRefCitation>
),
<taxonomicName id="2268B039F9788BD5D9D8C0493E46C7D3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="1" pageNumber="250" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
sp. &quot;Montagne des
<normalizedToken id="D4E12929D77E38EB1FBC1FBC0D8EEB3B" originalValue="Français”">Francais&quot;</normalizedToken>
(
<bibRefCitation id="0C1628F4895108912F38468A9C6CA1A0" author="Glaw, F" journalOrPublisher="Vences and Glaw Verlag, Cologne" pageId="7" pageNumber="256" title="A field guide to the amphibians and reptiles of Madagascar" year="2007">Glaw and Vences 2007</bibRefCitation>
),
<taxonomicName id="33E940557F0CE9C717CD81422E494609" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="1" pageNumber="250" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
sp. (
<bibRefCitation id="18A7DF21BE48C09A804B91F6A6A22E1A" author="D'Cruze, N" journalOrPublisher="Herpetological Conservation and Biology" pageId="7" pageNumber="256" pagination="87 - 99" title="The first comprehensive survey of amphibians and reptiles at Montagne des Francais, Madagascar." volume="2" year="2007">
<normalizedToken id="4D31D5E949F1408F7157CC5B6E02E10E" originalValue="DCruze">D'Cruze</normalizedToken>
et al. 2007
</bibRefCitation>
),
<taxonomicName id="D5A3B5AA236FC4525B4615CA8EF76083" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="karstophila">Paroedura aff. karstophila</taxonomicName>
(
<bibRefCitation id="9D6DA89E87CFD0940F14DD4C25AD6A4A" author="Schoenecker, P" journalOrPublisher="Edition Chimaira, Terralog" pageId="8" pageNumber="257" pagination="1 - 144" title="Geckos of Madagascar, the Seychelles, Comoros and Mascarene Islands." volume="12" year="2008">
<normalizedToken id="F0752B9E795A6AA2F5AEC2E2BEEC5E31" originalValue="Schönecker">Schoenecker</normalizedToken>
2008
</bibRefCitation>
),
<taxonomicName id="FD003263E1FE371DF1162285F0810735" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="karstophila">Paroedura cf. karstophila</taxonomicName>
(
<bibRefCitation id="FAF039C37578AA6D383B84C861AAE1B7" author="Megson, S" journalOrPublisher="Herpetology Notes" pageId="7" pageNumber="256" pagination="31 - 44" title="A comprehensive survey of amphibians and reptiles in the extreme north of Madagascar." volume="2" year="2009">Megson et al. 2009</bibRefCitation>
),
<taxonomicName id="06A9259326548E8ADEB8A588AF9E1469" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="1" pageNumber="250" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
sp. n. / sp. n. 1 (
<bibRefCitation id="309B3674A4A99D5F95D1A95024F6C16D" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. 2008</bibRefCitation>
and
<bibRefCitation id="143D3EFFC485DA1D6E63E59C7183F027" author="Nagy, ZT" journalOrPublisher="PLoS ONE" pageId="8" pageNumber="257" title="First large-scale DNA barcoding assessment of reptiles in the biodiversity hotspot of Madagascar, based on newly designed COI primers." url="10.1371/journal.pone.0034506" volume="7" year="2012">Nagy et al. 2012</bibRefCitation>
). Its karyotype (2n = 36) has been described under the names
<taxonomicName id="F5F3550472A2F5819CDF5EC442DB9EC6" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="1" pageNumber="250" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
sp. n. I 'Montagne des
<normalizedToken id="91437ADF5D564E9E9F4D1407E28BA9A8" originalValue="Français">Francais'</normalizedToken>
(
<bibRefCitation id="DE5DC0A0554A1ADD70DFA68684366FF6" author="Aprea, G" journalOrPublisher="African Zoology" pageId="7" pageNumber="256" pagination="96 - 108" title="Chromosomal rearrangements occurred repeatedly and independently during species diversification in Malagasy geckos, genus Paroedura." url="10.3377/004.048.0101" volume="48" year="2013">Aprea et al. 2013</bibRefCitation>
) and
<taxonomicName id="D9D5B46F81F883EFF648D7A851EEB9DF" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="karstophila">Paroedura karstophila</taxonomicName>
(
<bibRefCitation id="B5979A1E0EFA95489A6A67E4137C2E3C" author="Koubova, M" journalOrPublisher="Organisms, Diversity and Evolution" pageId="7" pageNumber="256" title="Chromosome Research" url="10.1007/s10577-014-9430-z" year="2014">
<normalizedToken id="0AD4052F486962B290773D5554102AE0" originalValue="Koubová">Koubova</normalizedToken>
et al. 2014
</bibRefCitation>
). DNA-sequences of this species have been published in
<bibRefCitation id="9B08EEC91BD742FC63F2011EA96CBF3C" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. (2008)</bibRefCitation>
,
<bibRefCitation id="6888F6689F725E5C82C1EEEFD2A7E647" author="Aprea, G" journalOrPublisher="African Zoology" pageId="7" pageNumber="256" pagination="96 - 108" title="Chromosomal rearrangements occurred repeatedly and independently during species diversification in Malagasy geckos, genus Paroedura." url="10.3377/004.048.0101" volume="48" year="2013">Aprea et al. (2013)</bibRefCitation>
, and
<bibRefCitation id="04AF8B39DDEAE26DCF60A58D2EFDE589" author="Koubova, M" journalOrPublisher="Organisms, Diversity and Evolution" pageId="7" pageNumber="256" title="Chromosome Research" url="10.1007/s10577-014-9430-z" year="2014">
<normalizedToken id="5A1241CCAD145BE1CE5E2173F8201A81" originalValue="Koubová">Koubova</normalizedToken>
et al. (2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1C0276E3ABEBA84AA55293F826C0522A" pageId="1" pageNumber="250" type="holotype">
<paragraph id="C8C45A69129DD8F507EDB9009259AAD8" pageId="1" pageNumber="250">Holotype.</paragraph>
<paragraph id="0C16B9960624C5E9EF08D649B2F7F794" pageId="1" pageNumber="250">
ZSM 342/2004 (field number FGZC 639), adult male with (broken) original tail and everted hemipenes, collected at Montagne des
<normalizedToken id="63E70645D4E5DE6F307154D2AFEB5F93" originalValue="Français">Francais</normalizedToken>
(
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12°19
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S
</geoCoordinate>
,
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49°20
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E
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, 334 m above sea level), Antsiranana Province, north Madagascar, on 18 February 2004 by F. Glaw, M. Puente and R. Randrianiaina. GenBank accession numbers for sequences of the holotype (
<bibRefCitation id="956A0C776C32AF19FB6FBA0BD74622DB" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. 2008</bibRefCitation>
): EF536213 (ND2), EF536239 (ND4), EF536165 (RAG1) and EF536189 (PDC).
</paragraph>
</subSubSection>
<subSubSection id="98DCF090907DB763BA94E455DA30C2CA" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250" type="paratypes">
<paragraph id="A9521013E432E41332115631096C2988" pageId="1" pageNumber="250">Paratypes.</paragraph>
<paragraph id="2FCFBE8A3C15C9CBFE58D2D827A42A60" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250">
All paratypes were collected in the
<normalizedToken id="ADB65B3B9A400BC8B2B01EBE11CAE208" originalValue="“tsingy”">&quot;tsingy&quot;</normalizedToken>
limestone massif at Montagne des
<normalizedToken id="BF291DC1613F35DCF952A9D1659C2272" originalValue="Français">Francais</normalizedToken>
, Antsiranana province, north Madagascar. Specimens were collected on the tsingy outcrops along the way between the Hotel &quot;Kings Lodge&quot; (
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12°18
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S
</geoCoordinate>
,
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49°20
<normalizedToken id="FF5E7BF6A2D913226C2DE63E33F22192" originalValue="22,6">'22,6''</normalizedToken>
E
</geoCoordinate>
, 10 m a.s.l.) and the remains of the French Fort (
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12°19
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S
</geoCoordinate>
,
<geoCoordinate id="59499BA483BDA325C00656F8582BB2CF" direction="east" orientation="longitude" precision="15" value="49.33583">
49°20
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E
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, 334 m), except where other locality information and coordinates are given in the following: UADBA uncatalogued (FG/MV 2000-317), sex unknown, and ZSM 531/2000 (FG/MV 2000-316), adult male with everted hemipenes, both collected on 14 March 2000 by F. Glaw, K. Glaw and M. Vences; ZSM 532/2000 (no field number), adult female, collected on 21 March 2000 by F. Glaw and K. Glaw; ZSM 1108/2003 (no field number), adult female, collected on ca. 20 February 2003 by F. Glaw and R. D. Randrianiaina; ZSM 337/2004 (FGZC 634), subadult, and ZSM 338/2004 (FGZC 635), subadult, both collected at
<geoCoordinate id="DE89A62B295AC58FD79F69587120586C" direction="south" orientation="latitude" precision="15" value="-12.326111">
12°19
<normalizedToken id="2BC9C0C8A8FC359005F1A0C6A71550B1" originalValue="34“">'34&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="403C1326063D474E11EADC4E52534603" direction="east" orientation="longitude" precision="15" value="49.33583">
49°20
<normalizedToken id="C7D5FF161FCCAFCC893B1A4802D2A263" originalValue="09“">'09&quot;</normalizedToken>
E
</geoCoordinate>
, 334 m a.s.l., on 23 February 2004 by F. Glaw, M. Puente and R. D. Randrianiaina; ZSM 339/2004 (FGZC 636), adult female, ZSM 340/2004 (FGZC 637), adult female, ZSM 341/2004 (FGZC 638), adult female [original tail broken], all three with same data as holotype; ZSM 343/2004 (FGZC 640), adult male without tail, and ZSM 350/2004 (FGZC 647), adult female, both without reliable locality and collection data, but most likely with same data as holotype; UADBA uncatalogued (FGZC 612), sex unknown, collected at
<geoCoordinate id="15D37FEFECEAC977E2823965FD25BA7C" direction="south" orientation="latitude" precision="15" value="-12.326111">
12°19
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S
</geoCoordinate>
,
<geoCoordinate id="EFAE6788812C6AAC94B70C76BF8B0B1E" direction="east" orientation="longitude" precision="15" value="49.33583">
49°20
<normalizedToken id="CD26ACA564A5B72B366A5964947BCF3C" originalValue="09“">'09&quot;</normalizedToken>
E
</geoCoordinate>
, 334 m a.s.l., on 20-28 February 2004 by F. Glaw, M. Puente and R. D. Randrianiaina; ZSM 352/2004 (FGZC 649), adult, and ZSM 353/2004 (FGZC 650), adult female, both collected at
<geoCoordinate id="FF9D5C66B3F0540F539CB10D3E1523B7" direction="south" orientation="latitude" precision="15" value="-12.326111">
12°19
<normalizedToken id="4C26ADF76A36406FC594B6A1C22A59EC" originalValue="34“">'34&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="27B982AC36BC99DD8EE67ABC540D9CF4" direction="east" orientation="longitude" precision="15" value="49.33583">
49°20
<normalizedToken id="9611F0042237E00C47CFABF9818FFAA1" originalValue="09“">'09&quot;</normalizedToken>
E
</geoCoordinate>
, 334 m a.s.l.,
<pageBreakToken id="634CC15C66557A6ED7AFDF3D77970119" pageId="2" pageNumber="251" start="start">on</pageBreakToken>
18-23 February 2004 by F. Glaw, M. Puente and R. D. Randrianiaina; UADBA-R 70183 (FGZC 1109), adult male with everted hemipenes, UADBA-R 70185 (FGZC 1112), male, UADBA-R 70184 (FGZC 1114), adult female, ZSM 2106/2007 (FGZC 1099), juvenile, ZSM 2107/2007 (FGZC 1100), juvenile, ZSM 2113/2007 (FGZC 1115), adult female, all six collected around the remains of the French Fort (
<geoCoordinate id="AC8C4FDAF3B7C4C499A0DD6676A95DF2" direction="south" orientation="latitude" precision="15" value="-12.325833">
12°19
<normalizedToken id="8FC84DD761D59944A2EBDE1F7E22EE09" originalValue="33">'33''</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="0792D4F0D74398EB9FB77A798851FDFE" direction="east" orientation="longitude" precision="15" value="49.338055">
49°20
<normalizedToken id="C8E60B0692B87E7108C087F1BB1D550F" originalValue="17">'17''</normalizedToken>
E
</geoCoordinate>
), collected on 27 February 2007 by P. Bora, H. Enting, F. Glaw, A. Knoll and J.
<normalizedToken id="750607985F7E2BFB7195C47476279199" originalValue="Köhler">Koehler</normalizedToken>
; UADBA-R 70281 (FGZC 1659), sex unknown, ZSM 1530/2008 (FGZC 1660), adult female, both collected in the cave between Andavakoera and remains of French Fort, on 16 February 2008 by M. Franzen, F. Glaw, J.
<normalizedToken id="77280F9EC6FEB8FD5A643647B62FDEB1" originalValue="Köhler">Koehler</normalizedToken>
and Z. T. Nagy.
</paragraph>
</subSubSection>
<subSubSection id="03511E0AC1E010DF90CAF2A02CD9DEF7" pageId="2" pageNumber="251" type="diagnosis">
<paragraph id="30C1DF44D295758798455A55C4E5C49F" pageId="2" pageNumber="251">Diagnosis.</paragraph>
<paragraph id="DEEC4E30347B04903F03865CA0EE3863" pageId="2" pageNumber="251">
<taxonomicName id="A729DF858B10F32C886B2C0B6C3F8265" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="2" pageNumber="251" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
sp. n. is a medium-sized species (SVL up to 58 mm, tail length up to 53 mm), having moderately prominent dorsal tubercles disposed into moderately distinct, and generally regular longitudinal rows and an original tail with no spines.
</paragraph>
<paragraph id="05FB54676493F4AE053ECA870F3C6063" pageId="2" pageNumber="251">
The new species can be easily attributed to the genus
<taxonomicName id="832661E2B437DEFA67AF0F4193FC2897" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="2" pageNumber="251" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
based on its nested phylogenetic position within the genus (
<bibRefCitation id="C83EE1EA70CD9EBB7C335E1529E3DEFD" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. 2008</bibRefCitation>
) and its morphological similarity to other
<taxonomicName id="376F2C700E18F64652670DC2844E2B6F" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="2" pageNumber="251" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
species, especially concerning the ventral structure of their fingers and toes which comprise a pair of squarish terminal adhesive pads. Among Malagasy geckos this terminal toe structure is only found in
<taxonomicName id="F0605A654794A5A6438DFA5E0E657FFC" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="2" pageNumber="251" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
and the related genus
<taxonomicName id="AAA1BB1EBDBBDC29D9CDAF9A308FCEDA" class="Squamata" family="Gekkonidae" genus="Ebenavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ebenavia" pageId="2" pageNumber="251" phylum="Chordata" rank="genus">Ebenavia</taxonomicName>
(Glaw &amp; Vences, 2007). The latter genus can be easily distinguished from
<taxonomicName id="013515E35AC55651F42BD8D41A01FB3F" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="2" pageNumber="251" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
by its much narrower and strongly pointed head, its elongated body, and smaller size.
</paragraph>
</subSubSection>
<subSubSection id="D28DA6E7CCA2E9C36549AA67B243343A" lastPageId="3" lastPageNumber="252" pageId="2" pageNumber="251" type="comparisons">
<paragraph id="3D997DE66C7A3ECD01C31B8E2CA7D530" pageId="2" pageNumber="251">Comparisons.</paragraph>
<paragraph id="15215678C93CC5060142141C6AABA3BB" lastPageId="3" lastPageNumber="252" pageId="2" pageNumber="251">
The new species can be distinguished from the 17 other currently recognized
<taxonomicName id="23A813FA3848F119481C3C3BD4D30DC6" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="3" pageNumber="252" phylum="Chordata" rank="genus">
<pageBreakToken id="AD77D5C55366307F92ED6D83C7084818" pageId="3" pageNumber="252" start="start">Paroedura</pageBreakToken>
</taxonomicName>
species (including the three available junior synonyms in the genus) as follows: From
<taxonomicName id="0EA6E4E7FE68836897923198FE1069BD" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura androyensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="androyensis">Paroedura androyensis</taxonomicName>
,
<taxonomicName id="B7D462CDE21E11E2E5F97EE5C04454DA" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura bastardi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="bastardi">Paroedura bastardi</taxonomicName>
,
<taxonomicName id="A71BD04FD3893E21C1087030C71DA9D0" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura ibityensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="ibityensis">Paroedura ibityensis</taxonomicName>
,
<taxonomicName id="102B8C1D9D00B28CAD8A56643236B146" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura lohatsara" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="lohatsara">Paroedura lohatsara</taxonomicName>
,
<taxonomicName id="19D1E7DBCEB0205B54B79F5C08A085E2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura maingoka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="maingoka">Paroedura maingoka</taxonomicName>
,
<taxonomicName id="838DAA5037B4CA2B223C2FC3F450836D" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura picta" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="picta">Paroedura picta</taxonomicName>
, and
<taxonomicName id="485073F47B9F4226D20515CFB4874808" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vahiny" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vahiny">Paroedura vahiny</taxonomicName>
by having the nostril in contact with the rostral scale; from
<taxonomicName id="1D767CCA937B42BAACACEBC7F6340E75" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura gracilis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="gracilis">Paroedura gracilis</taxonomicName>
by absence of a raised vertebral ridge on the body and shorter forelimbs which are not extending forward beyond tip of snout; from
<taxonomicName id="ECF36291493AC8F171A48166813E6877" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura masobe" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="masobe">Paroedura masobe</taxonomicName>
by much smaller size (SVL up to 58 mm versus 107 mm), much smaller eyes with a pigmented iris (versus black iris) and absence of a dorsal row of paired spines on the tail; from the two Comoroan species
<taxonomicName id="A9ADFE4BFC9FD74E2E18A30AE9CEA044" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura sanctijohannis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="sanctijohannis">Paroedura sanctijohannis</taxonomicName>
and
<taxonomicName id="1D167814409688497F7B15C034834807" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stellata" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stellata">Paroedura stellata</taxonomicName>
by slightly smaller size (SVL up to 58 mm versus 68 mm and 62 mm, respectively) and absence of whorls with distinct spiny tubercles of the original tail; from the syntopically distributed
<taxonomicName id="8CEF104D2005575A00CC1F191CA2BE80" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
by smaller size (SVL up to 58 mm versus 70 mm) and absence of whorls with distinct spiny tubercles of the original tail; from
<taxonomicName id="58445524190AC797E1942FE0EB33AFAC" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura tanjaka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="tanjaka">Paroedura tanjaka</taxonomicName>
by much smaller size (SVL up to 58 mm versus 102 mm) and absence of whorls with distinct spiny tubercles of the original tail; from
<taxonomicName id="EA9EBB611DB7CEEBE58F88960E17D88C" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vazimba" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vazimba">Paroedura vazimba</taxonomicName>
by larger size (SVL up to 58 mm versus 49 mm) and absence of whorls with distinct spines of the original tail; from
<taxonomicName id="A463D1922939ACDD4C0FEECDDE1494F7" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura oviceps" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="oviceps">Paroedura oviceps</taxonomicName>
from its type locality Nosy Be by smaller size (SVL up to 58 mm versus 69 mm) and rather regularly arranged tubercle rows on the back (versus rather irregular rows of tubercles); from
<taxonomicName id="9770848C387613EA5E12CA5042C9FE86" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="karstophila">Paroedura karstophila</taxonomicName>
by the absence of whorls with distinct spiny tubercles of the original tail (and by a smoother regenerated tail, see
<bibRefCitation id="36E32C33B521CC76A20A412723A8A393" author="Nussbaum, RA" journalOrPublisher="Miscellaneous Publications Museum of Zoology, University of Michigan" pageId="8" pageNumber="257" pagination="1 - 26" title="Systematic revision of the genus Paroedura Guenther (Reptilia: Squamata: Gekkonidae), with description of five new species." volume="189" year="2000">Nussbaum and Raxworthy 2000</bibRefCitation>
) and by colouration (see Fig. 2 versus Fig. 5); and from its close relative
<taxonomicName id="93B2A15089F235B3E5FA3F49D2F74985" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
(
<bibRefCitation id="76E967326502AD83C71ABA5A1DA14887" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. 2008</bibRefCitation>
) by shorter limbs (finger tips reach the anterior margin of eye versus snout tip when forelimbs are adpressed along the body), slightly smaller size (SVL up to 58 mm versus 65 mm, see Table 2), distinct and generally regularly arranged tubercles rows on the back (versus less distinct and less regular rows), and a more slender habitus (see Fig. 2 versus Fig. 5). In addition,
<taxonomicName id="5F72A12006358C6D37F6419A3BAE1B59" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
can be easily distinguished from most other
<taxonomicName id="B8C6ED873990283605FC7155474CFDEC" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="3" pageNumber="252" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
species (
<taxonomicName id="7A49312E332EA6A33624B5FCB3F8DDB1" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura androyensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="androyensis">Paroedura androyensis</taxonomicName>
,
<taxonomicName id="32DBD48775AF1BAC0C141970DADA9015" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura bastardi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="bastardi">Paroedura bastardi</taxonomicName>
,
<taxonomicName id="BEE08F83E9B8D78EEC4528FA8D7E0CA4" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura gracilis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="gracilis">Paroedura gracilis</taxonomicName>
,
<taxonomicName id="D7F3F8CE982B9FD98FB1721C2D2E6AE3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura ibityensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="ibityensis">Paroedura ibityensis</taxonomicName>
,
<taxonomicName id="B1C05E2B40A1ED549BB03612C0D019A2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura lohatsara" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="lohatsara">Paroedura lohatsara</taxonomicName>
,
<taxonomicName id="912D4DFD459D9B6BCCA25E086E6C35F9" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura maingoka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="maingoka">Paroedura maingoka</taxonomicName>
,
<taxonomicName id="A2B21530172EFD2C92451A304DB2E652" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura masobe" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="masobe">Paroedura masobe</taxonomicName>
,
<taxonomicName id="6EEC28686B05BEDF5CC5029FC2680090" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura oviceps" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="oviceps">Paroedura oviceps</taxonomicName>
,
<taxonomicName id="8C61AF8DD0FD3B2C197ED108CDDA8240" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura picta" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="picta">Paroedura picta</taxonomicName>
,
<taxonomicName id="6D73AAA4731F2AD59E6D0A450630A1E6" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura sanctijohannis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="sanctijohannis">Paroedura sanctijohannis</taxonomicName>
,
<taxonomicName id="698D6707100DD60B035F2EB622C28059" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stellata" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stellata">Paroedura stellata</taxonomicName>
,
<taxonomicName id="60883BE17EF12E4B27CB48734C5BC138" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
,
<taxonomicName id="9736C50E4BD077DEE463EE58A072916B" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura tanjaka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="tanjaka">Paroedura tanjaka</taxonomicName>
,
<taxonomicName id="AC2E13463B5D69DCA75D12D89657E2C4" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vahiny" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vahiny">Paroedura vahiny</taxonomicName>
and
<taxonomicName id="C3C7889CC537EBD47A6835B23553E3A3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vazimba" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vazimba">Paroedura vazimba</taxonomicName>
) by adult colouration in life (see colour photographs in
<bibRefCitation id="4A495855D3A7D7FCB15ECF714AB9F9C4" author="Glaw, F" journalOrPublisher="Vences and Glaw Verlag, Cologne" pageId="7" pageNumber="256" title="A field guide to the amphibians and reptiles of Madagascar" year="2007">Glaw and Vences 2007</bibRefCitation>
,
<bibRefCitation id="BECA82873DE902BDAF9C818E0D8A209E" author="Schoenecker, P" journalOrPublisher="Edition Chimaira, Terralog" pageId="8" pageNumber="257" pagination="1 - 144" title="Geckos of Madagascar, the Seychelles, Comoros and Mascarene Islands." volume="12" year="2008">
<normalizedToken id="752C293CC892B9F5525A9506296AA9B0" originalValue="Schönecker">Schoenecker</normalizedToken>
2008
</bibRefCitation>
,
<bibRefCitation id="25C1EEC9299CAA5A4C11C9024FBF9EBB" author="Hawlitschek, O" journalOrPublisher="Zoologica Scripta" pageId="7" pageNumber="256" pagination="135 - 150" title="The complex colonization history of nocturnal geckos (Paroedura) on the Comoros Archipelago." url="10.1111/zsc.12001" volume="42" year="2013">Hawlitschek and Glaw 2013</bibRefCitation>
) and from
<taxonomicName id="EB4CE11057C168F8C0A20DC70EF7C70A" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura androyensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="androyensis">Paroedura androyensis</taxonomicName>
,
<taxonomicName id="9D545843810A3A3AFA3437B0E887FF6F" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura bastardi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="bastardi">Paroedura bastardi</taxonomicName>
,
<taxonomicName id="E0987E0CA78701EC3081507A3BEB1700" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura gracilis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="gracilis">Paroedura gracilis</taxonomicName>
,
<taxonomicName id="9008505837446B7A7DDBD09277890EF2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
,
<taxonomicName id="E379F4582C4FB9A6AFA98B2CCA86E8C3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura ibityensis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="ibityensis">Paroedura ibityensis</taxonomicName>
,
<taxonomicName id="82104D51EE231736145417BF35958675" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura lohatsara" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="lohatsara">Paroedura lohatsara</taxonomicName>
,
<taxonomicName id="C3E0F7BB9992BD32AAD85656E4B4FEE2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura maingoka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="maingoka">Paroedura maingoka</taxonomicName>
,
<taxonomicName id="81B39FB7081455BAF317E5AD00E0AD13" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura masobe" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="masobe">Paroedura masobe</taxonomicName>
,
<taxonomicName id="1214E9D82393426DDDD8A22C68C281C2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura picta" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="picta">Paroedura picta</taxonomicName>
,
<taxonomicName id="5283654055AEAD639B7B84B4FF485DDA" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura sanctijohannis" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="sanctijohannis">Paroedura sanctijohannis</taxonomicName>
,
<taxonomicName id="DBE13E1221668BC2486978397F347D1C" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stellata" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stellata">Paroedura stellata</taxonomicName>
,
<taxonomicName id="C10DA7323646F8DCB629FD3802A537CB" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
,
<taxonomicName id="76BE32648C0190ECD036B1FDAD8CA276" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura tanjaka" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="tanjaka">Paroedura tanjaka</taxonomicName>
, and
<taxonomicName id="C3838DB75579B25E5B8516BD2A17D2D3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vazimba" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vazimba">Paroedura vazimba</taxonomicName>
by juvenile colouration (
<bibRefCitation id="D7055D44DA49CCB10FBC2489E618D0A7" author="Glaw, F" journalOrPublisher="Vences and Glaw Verlag, Cologne" pageId="7" pageNumber="256" title="A field guide to the amphibians and reptiles of Madagascar" year="2007">Glaw and Vences 2007</bibRefCitation>
,
<bibRefCitation id="AD80EEC1C11A35ED02225CF209240762" author="Schoenecker, P" journalOrPublisher="Edition Chimaira, Terralog" pageId="8" pageNumber="257" pagination="1 - 144" title="Geckos of Madagascar, the Seychelles, Comoros and Mascarene Islands." volume="12" year="2008">
<normalizedToken id="A53584B5F69B2BAD67463C62FD72B3E4" originalValue="Schönecker">Schoenecker</normalizedToken>
2008
</bibRefCitation>
,
<bibRefCitation id="C46337ACEED4D68D3365E2961110137E" author="Hawlitschek, O" journalOrPublisher="Zoologica Scripta" pageId="7" pageNumber="256" pagination="135 - 150" title="The complex colonization history of nocturnal geckos (Paroedura) on the Comoros Archipelago." url="10.1111/zsc.12001" volume="42" year="2013">Hawlitschek and Glaw 2013</bibRefCitation>
, FG pers. obs.; juvenile colouration of the other species still unknown). Genetically,
<taxonomicName id="E76F284485B823EC49228E0B405279CF" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
can be distinguished from all other species in the genus by its molecular differentiation in mitochondrial and nuclear genes (
<bibRefCitation id="F419907BBA23AFA1D23CC12214C5E051" author="Jackman, TR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="256" pagination="74 - 81" title="Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae)." url="10.1016/j.ympev.2007.10.018" volume="46" year="2008">Jackman et al. 2008</bibRefCitation>
,
<bibRefCitation id="569A8B6A9E50E1364C936E1A5E506743" author="Nagy, ZT" journalOrPublisher="PLoS ONE" pageId="8" pageNumber="257" title="First large-scale DNA barcoding assessment of reptiles in the biodiversity hotspot of Madagascar, based on newly designed COI primers." url="10.1371/journal.pone.0034506" volume="7" year="2012">Nagy et al. 2012</bibRefCitation>
,
<bibRefCitation id="B26734839015A5ADC043DA7667C36286" author="Hawlitschek, O" journalOrPublisher="Zoologica Scripta" pageId="7" pageNumber="256" pagination="135 - 150" title="The complex colonization history of nocturnal geckos (Paroedura) on the Comoros Archipelago." url="10.1111/zsc.12001" volume="42" year="2013">Hawlitschek and Glaw 2013</bibRefCitation>
, Fig. 1) except for
<taxonomicName id="8C5B4AF237DCFC90842A004956ECE987" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura vahiny" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="vahiny">Paroedura vahiny</taxonomicName>
for which DNA sequences are not yet available.
</paragraph>
<caption id="B99A9109267D37D2BCA82AD9A8EA3560" pageId="3" pageNumber="252">
<paragraph id="F89A6735DE46398BDAF73030E1B7217B" pageId="3" pageNumber="252">
Figure 2. Paratypes of
<taxonomicName id="8ED0ACF54EAB1F04E210A1B7F425AFA2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="3" pageNumber="252" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
: (A) adult male ZSM 531/2000 (SVL 53.5 mm) in dorsolateral view; (B) adult female ZSM 2113/2007 (SVL 46.1 mm) in dorsal view; (C) subadult ZSM 2107/2007 in lateral view; (D) juvenile (ZSM 2106/2007, SVL 28.1 mm) and subadult (ZSM 2107/2007, SVL 35.2 mm) in dorsolateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="066C8247CA0FAA1D057E8BFCE1FBC8E8" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252" type="description of the holotype">
<paragraph id="058F24015A5F2DA1A38F412E9C149588" pageId="3" pageNumber="252">Description of the holotype.</paragraph>
<paragraph id="5BE980912F7C5FC90CFAFE955916E51C" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">
SVL 52.6 mm, further measurements and counts are given in Table 1. Holotype in good condition, with complete but broken original tail and everted hemipenes. Head distinctly wider than neck and wider than body. Snout angled downward to tip, slight depression between poorly developed canthal ridges. Ear opening is a vertical slit. Original tail slightly shorter than snout-vent length, nearly round in cross section in its proximal part, laterally compressed in its distal half, with sharply pointed tip; ventral pygal section with pair of postcloacal sacs. Digits moderately expanded at tips. Rostral scale rectangular, much wider than tall and wider than mental. Nostril in contact with
<pageBreakToken id="C197044DF53B73A2DF36DF048E3E3DE3" pageId="4" pageNumber="253" start="start">rostral</pageBreakToken>
, first supralabial, and five further scales. First supralabial largest, labials smooth. Snout and interorbital scales juxtaposed, some raised, some scales in front of orbits tuberculate, as are some larger lateral occipital scales. Dorsolateral neck and body scales very heterogeneous with about eight partly poorly recognizable longitudinal rows at midbody of enlarged tubercles; enlarged tubercles separated partly by small flat scales and smaller tubercles. Dorsal scales of forelimbs mostly flat. Dorsal scales of hindlimbs large and tuberculate, much smaller above knee. Ventral scales of forelimbs slightly smaller than surrounding ventral scales of the body. Dorsal pygal scales like dorsal body scales; lateroventral pygals tuberculate. Tail scales flat, tail segments without any transverse row of spiny tubercles. Mental triangular, bordered posteriorly by a pair of elongate, irregular hexagonal postmentals. Postmentals contact mental, first and second infralabial, one enlarged lateral gular, one smaller posterolateral gular, and one larger central gular. First three infralabials not significantly larger than others. Gulars small, granular. Ventrals of chest and abdomen flat. Proximal subdigitals in rows of 2-3. Pair of squarish, terminal pads. Claws curving downwards between terminal pads of digits.
</paragraph>
<caption id="049C38BBAC47223698308DDEF24B0E70" pageId="4" pageNumber="253">
<paragraph id="EC8B13861578DAEC26C021C37CE2ECD4" pageId="4" pageNumber="253">
Table 1. Morphometric and meristic variation of several type specimens of
<taxonomicName id="78EA86431EAE43A381268368E89DB0CB" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="4" pageNumber="253" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
from the type locality Montagne des
<normalizedToken id="67A8EE0B1A8F7F6BF298959C8CDB2A2B" originalValue="Français">Francais</normalizedToken>
. Abbreviations for measurements and counts (see Materials and Methods for other abbreviations): ZSM = Zoologische Staatssammlung
<normalizedToken id="4886656E04BF7A88210174BDAEED622D" originalValue="München">Muenchen</normalizedToken>
; SVL = snout-vent length; TL = tail length; HL = maximum head length (from tip of snout to posterior margin of ear); HW = maximum head width, at widest point; HH = maximum head height; AGL = axilla-groin distance; ED = maximum eye diameter; EO = maximum ear opening diameter; FOL = forelimb length, from axilla to tip of longest finger; HIL = hindlimb length, from groin to tip of longest toe; SPL = number of supralabial scales; IFL = number of infralabial scales; NAS = number of nasals in direction from rostral to labial including nasorostrals, supranasals, postnasals; IN = number of internasals; IO = number of interorbitals; PM = number of postmentals; SLM4 = number of subdigital lamellae on fourth digit of manus; SLP4 = number of subdigital lamellae on fourth toe of pes; PCT = number of postcloacal tubercles; TLT = number of tubercle rows on tail. Counts are listed left-right. All measurements in Tables 1 and 2 were taken by H.
<normalizedToken id="0D5EF2E87BEE7655AD01AFEAD809CDF6" originalValue="Rösler">Roesler</normalizedToken>
with a caliper to the nearest 0.1 mm. HT = holotype, PT = paratype, F = female, M = male, SA = subadult.
</paragraph>
</caption>
<paragraph id="7E0AA50D0DE6F0833BAA399C9CB690CE" pageId="4" pageNumber="253">
<table id="C71527270495FD15AA84973CDD99307D" pageId="4" pageNumber="253">
<tr id="5333B82395EF0BBF749001869DFCBA68" pageId="4" pageNumber="253">
<th id="B50DB60202B3666165C11284CC9C60D2" colspan="1" pageId="4" pageNumber="253" rowspan="1">ZSM</th>
<th id="26E81AE42A47A16F5F19FD53658B5906" colspan="1" pageId="4" pageNumber="253" rowspan="1">352/2004</th>
<th id="09F2FED3AE623C5277D3F4F114A163D3" colspan="1" pageId="4" pageNumber="253" rowspan="1">353/2004</th>
<th id="7189EC1E36F9ACB3F3FB5D58C3043E08" colspan="1" pageId="4" pageNumber="253" rowspan="1">340/2004</th>
<th id="61394107DF5B34FE957F6040FDE581AE" colspan="1" pageId="4" pageNumber="253" rowspan="1">350/2004</th>
<th id="C81280D361E1ADA881066BCCDA629C88" colspan="1" pageId="4" pageNumber="253" rowspan="1">339/2004</th>
<th id="364D5959CF34561553A40032C36E90B5" colspan="1" pageId="4" pageNumber="253" rowspan="1">341/2004</th>
<th id="8A4361FFDE1E0243465B7152DED57493" colspan="1" pageId="4" pageNumber="253" rowspan="1">338/2004</th>
<th id="11C5D93020FF3A237C7A8146F653E6D2" colspan="1" pageId="4" pageNumber="253" rowspan="1">337/2004</th>
<th id="DCB5E541ADC41F9ED9C4F8E211BBEBB5" colspan="1" pageId="4" pageNumber="253" rowspan="1">342/2004</th>
<th id="3E021DCC0583305EFD73A3272F38E891" colspan="1" pageId="4" pageNumber="253" rowspan="1">343/2004</th>
</tr>
<tr id="B7F46284489B383C3F0E50B3C69BB27D" pageId="4" pageNumber="253">
<td id="D0C744710477096CCA10C81504C6E52C" colspan="1" pageId="4" pageNumber="253" rowspan="1">Type status</td>
</tr>
<tr id="8CA086AC27D7CF583565C827EC1FF70E" pageId="4" pageNumber="253">
<td id="89D37B8CEAB9112F33BC814EC726BF91" colspan="1" pageId="4" pageNumber="253" rowspan="1">Sex</td>
</tr>
<tr id="FC7F433FB6D1B92D39AB2AB59A0E64A5" pageId="4" pageNumber="253">
<td id="067A30E25D8EB58F36A89879807BE46D" colspan="1" pageId="4" pageNumber="253" rowspan="1">SVL</td>
</tr>
<tr id="2899DB0A28C40842BDD5F7CE2446B024" pageId="4" pageNumber="253">
<td id="CA419CB992D65A423BA2160FBA5D0D38" colspan="1" pageId="4" pageNumber="253" rowspan="1">TL</td>
</tr>
<tr id="14558BD4B38567B01542D7CD32B5319B" pageId="4" pageNumber="253">
<td id="9C762A00AE48D998CEF2C4C8884F5040" colspan="1" pageId="4" pageNumber="253" rowspan="1">HL</td>
</tr>
<tr id="9D925500AF3D62719618CDE05E530AD6" pageId="4" pageNumber="253">
<td id="BACD3B7A43AD3246CC33B1FF573EFDAF" colspan="1" pageId="4" pageNumber="253" rowspan="1">HW</td>
</tr>
<tr id="89BD5B3B1B2417A46F9DA6EED3205AE0" pageId="4" pageNumber="253">
<td id="60F7CDE97E1AE22C7F292B2301F8C995" colspan="1" pageId="4" pageNumber="253" rowspan="1">HH</td>
</tr>
<tr id="27F169270739DBDFB58E25857718FE93" pageId="4" pageNumber="253">
<td id="253D77376C286E2339A4B81DAAA7C2F8" colspan="1" pageId="4" pageNumber="253" rowspan="1">AGL</td>
</tr>
<tr id="080D0D0162FE4772207E6AC316315E21" pageId="4" pageNumber="253">
<td id="AA19429FF13196C5DE5092B9D94C97D6" colspan="1" pageId="4" pageNumber="253" rowspan="1">ED</td>
</tr>
<tr id="46CED34EB13131B1032D1C3E1353C79B" pageId="4" pageNumber="253">
<td id="24D4E9DBDB86BA37E0961B135F35A637" colspan="1" pageId="4" pageNumber="253" rowspan="1">EO</td>
</tr>
<tr id="EEEB1E8320891946F4E2C6F31E54D67C" pageId="4" pageNumber="253">
<td id="391AEB489FAEE2F53973C9017F7D5A31" colspan="1" pageId="4" pageNumber="253" rowspan="1">FOL</td>
</tr>
<tr id="EF2F1BB389697970B0C6FF296EF06838" pageId="4" pageNumber="253">
<td id="A99AFD15389B0398D1511A78DC8158F1" colspan="1" pageId="4" pageNumber="253" rowspan="1">HIL</td>
</tr>
<tr id="25915434D0A18C56AA3DBD6204C59DBF" pageId="4" pageNumber="253">
<td id="5D995D6CACE8DFA5DEE7FEFEAA5CF8CE" colspan="1" pageId="4" pageNumber="253" rowspan="1">SVL:HIL</td>
</tr>
<tr id="FF4F5C4B2F24FA4DD642783E6C496A0A" pageId="4" pageNumber="253">
<td id="3B957F98B9102154BB09400DA6618648" colspan="1" pageId="4" pageNumber="253" rowspan="1">SPL</td>
</tr>
<tr id="2E091ED7DEC6F62129BF7B6D88CA2DA6" pageId="4" pageNumber="253">
<td id="58D696371EAA1C57A98B66805395D619" colspan="1" pageId="4" pageNumber="253" rowspan="1">IFL</td>
</tr>
<tr id="F1D845FBABEF46614CE6FB90E363DF3B" pageId="4" pageNumber="253">
<td id="FE0F7A5A18A3BDAD17179CFEFC28A43D" colspan="1" pageId="4" pageNumber="253" rowspan="1">NAS</td>
</tr>
<tr id="786D40C7EC5EBBA9038E672DFEF9411B" pageId="4" pageNumber="253">
<td id="00F36FB874772863CC080F61F7F23FFB" colspan="1" pageId="4" pageNumber="253" rowspan="1">IN</td>
</tr>
<tr id="D12565DF5ADA93EBCA8D748F3133A3C7" pageId="4" pageNumber="253">
<td id="F2D245D3D0697F6194424723B7E904F2" colspan="1" pageId="4" pageNumber="253" rowspan="1">IO</td>
</tr>
<tr id="F62AB32D57F44ABA2A0D6CB78074B57B" pageId="4" pageNumber="253">
<td id="A43256227EC35C37DC0F04C5169C211D" colspan="1" pageId="4" pageNumber="253" rowspan="1">PM</td>
</tr>
<tr id="6A84AA830FBA56933CED334406D8DEB8" pageId="4" pageNumber="253">
<td id="7D69DF3E8E0A72E2C1F7163E6DB027E6" colspan="1" pageId="4" pageNumber="253" rowspan="1">SLM4</td>
</tr>
<tr id="ED3DCB4A04785327ED168B5B5A2BABCE" pageId="4" pageNumber="253">
<td id="2C373D0A763BE29A1D9AAA5EF057F226" colspan="1" pageId="4" pageNumber="253" rowspan="1">SLP4</td>
</tr>
<tr id="C17329287E6B18A764D5199578E2C367" pageId="4" pageNumber="253">
<td id="26607DDCEB05B39AFB8CA13ECACA4B39" colspan="1" pageId="4" pageNumber="253" rowspan="1">PCT</td>
</tr>
<tr id="0B769FAF6081400A353BA2E4BC1AEA0F" pageId="4" pageNumber="253">
<td id="8147D4AA530001EA0E2D59A786C4EAFE" colspan="1" pageId="4" pageNumber="253" rowspan="1">TLT</td>
</tr>
</table>
</paragraph>
<caption id="9E86682F63ED552A93847B7D34FE3CEA" pageId="4" pageNumber="253">
<paragraph id="C2E12301E20D945CB16E08824E8C92BF" pageId="4" pageNumber="253">
Table 2. Morphometric and meristic variation of
<taxonomicName id="B9B4AD81AD88F103FF0A80D8A88B7C66" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="4" pageNumber="253" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
from the type locality Ankarana. See Table 1 for abbreviations and further details.
</paragraph>
</caption>
<paragraph id="0229093D8FAA8CAB3EC6B1FEA9D46C16" pageId="4" pageNumber="253">
<table id="AF2ABDB9872EB37C16482DB82355966B" pageId="4" pageNumber="253">
<tr id="FBAF4860C47EE884C08ECF3C5A8D0055" pageId="4" pageNumber="253">
<th id="5C5A45CE96BC19EDBFE788B3DDCD3A3C" colspan="1" pageId="4" pageNumber="253" rowspan="1">ZSM</th>
<th id="C4980A63AA787BE311367A41A11EB464" colspan="1" pageId="4" pageNumber="253" rowspan="1">345/2004</th>
<th id="71220DDC5A83552CAC8A6DA203CB3259" colspan="1" pageId="4" pageNumber="253" rowspan="1">347/2004</th>
<th id="6965814DBF278826FE64BBD3E5FEEF71" colspan="1" pageId="4" pageNumber="253" rowspan="1">348/2004</th>
<th id="6A2A11AEEA74F0B7D26377B59BAB8C78" colspan="1" pageId="4" pageNumber="253" rowspan="1">346/2004</th>
<th id="0794A3DEFD125EF18BCBCCABB696FC88" colspan="1" pageId="4" pageNumber="253" rowspan="1">349/2004</th>
<th id="836C4F1260ACC36AC8DE76239462A623" colspan="1" pageId="4" pageNumber="253" rowspan="1">344/2004</th>
<th id="14C41ABDB4F76249CE077518DDC02D72" colspan="1" pageId="4" pageNumber="253" rowspan="1">768/2003</th>
<th id="9DEC44EE3C3D220755DE688C63AC9CC4" colspan="1" pageId="4" pageNumber="253" rowspan="1">781/2003</th>
</tr>
<tr id="0DC3161DD4894AE2A2F92DE907BDE03D" pageId="4" pageNumber="253">
<td id="E94F876CCA664C05F097AE679C8DC9B4" colspan="1" pageId="4" pageNumber="253" rowspan="1">Sex</td>
</tr>
<tr id="BAB2406612275B659F1EFE56EB3111CB" pageId="4" pageNumber="253">
<td id="6275B121B4D20553B96833694AC90B3D" colspan="1" pageId="4" pageNumber="253" rowspan="1">SVL</td>
</tr>
<tr id="881B4DE47D9500BCAFEC96EB771EFD57" pageId="4" pageNumber="253">
<td id="A716313734D0DD2766B318EF935D5AAE" colspan="1" pageId="4" pageNumber="253" rowspan="1">TL</td>
</tr>
<tr id="7508A96DF5B75F5CF329297085DC4902" pageId="4" pageNumber="253">
<td id="9E59769B6B89A1C7A6B95C6919FB2F5D" colspan="1" pageId="4" pageNumber="253" rowspan="1">HL</td>
</tr>
<tr id="1623BE0C034E4D8D43477DD895143F5B" pageId="4" pageNumber="253">
<td id="766FF8225F3F81517674FAC4AB822EB4" colspan="1" pageId="4" pageNumber="253" rowspan="1">HW</td>
</tr>
<tr id="2DCC5DAD98AD3AB6F8BFC42602BA95EB" pageId="4" pageNumber="253">
<td id="772F99B61D875788ED7210D4AD79A93C" colspan="1" pageId="4" pageNumber="253" rowspan="1">HH</td>
</tr>
<tr id="5934C4C2367583A76068A3574F0540BC" pageId="4" pageNumber="253">
<td id="29ABEE808280A89BFEAA78BCD28764A6" colspan="1" pageId="4" pageNumber="253" rowspan="1">AGL</td>
</tr>
<tr id="5278BAF4F552B3E266D40551E36FD2E7" pageId="4" pageNumber="253">
<td id="54CFD810250DC8F1AE5D645D93F332D4" colspan="1" pageId="4" pageNumber="253" rowspan="1">ED</td>
</tr>
<tr id="47B950C5F4AF33E6F8A71FC8DEA6EC1C" pageId="4" pageNumber="253">
<td id="4EF80288E811B1F33CEA74BAF016D5E5" colspan="1" pageId="4" pageNumber="253" rowspan="1">EO</td>
</tr>
<tr id="C737BA43AE75E47897F33A29658E332A" pageId="4" pageNumber="253">
<td id="2491BDC7B0AB03D5C6AECE64EDDBB13D" colspan="1" pageId="4" pageNumber="253" rowspan="1">FOL</td>
</tr>
<tr id="C06DC6A8AFC526EDA692B086F9365D8D" pageId="4" pageNumber="253">
<td id="13AAB392C169B45FD8D6AAE86535C791" colspan="1" pageId="4" pageNumber="253" rowspan="1">HIL</td>
</tr>
<tr id="1A26750E36ADD275E580B37E51D6C3F4" pageId="4" pageNumber="253">
<td id="DC91FABE3A8AEABC7B42E7328C9D85EE" colspan="1" pageId="4" pageNumber="253" rowspan="1">SVL:HIL</td>
</tr>
<tr id="5D9C261034E79664A93445283EC503D0" pageId="4" pageNumber="253">
<td id="6772075C3F32883E2F570EFECB88DBC8" colspan="1" pageId="4" pageNumber="253" rowspan="1">SPL</td>
</tr>
<tr id="F690EA7C3791497E42F9F2FC0B762DD7" pageId="4" pageNumber="253">
<td id="3EEAF7DD3FA7DE2A63DF653C51E41FFE" colspan="1" pageId="4" pageNumber="253" rowspan="1">IFL</td>
</tr>
<tr id="88656EF7B3E9A63E1F4ECCC0E96F0196" pageId="4" pageNumber="253">
<td id="5FA5BF44E88B1083DDE0697FF49042E8" colspan="1" pageId="4" pageNumber="253" rowspan="1">NAS</td>
</tr>
<tr id="D83604A93902F4211936186CDF47B5F3" pageId="4" pageNumber="253">
<td id="DCF8AC4C3715A5D596765A63CED8CD61" colspan="1" pageId="4" pageNumber="253" rowspan="1">IN</td>
</tr>
<tr id="CBF646E806851DF1829D431BBC9DCEBD" pageId="4" pageNumber="253">
<td id="DA79CD8F324BC3FC8DAD34FA28EC2CCF" colspan="1" pageId="4" pageNumber="253" rowspan="1">IO</td>
</tr>
<tr id="FE86DDE28865B51061A78EEF93117F35" pageId="4" pageNumber="253">
<td id="5A9AB2B8C93C99EE125DB133235DCC2E" colspan="1" pageId="4" pageNumber="253" rowspan="1">PM</td>
</tr>
<tr id="68D0DE44379B61BE6315FE0134441996" pageId="4" pageNumber="253">
<td id="62440AA3563B768AFF03BB47A7967A1A" colspan="1" pageId="4" pageNumber="253" rowspan="1">SLM4</td>
</tr>
<tr id="9FE145E85FE240670196FA4534C90B9E" pageId="4" pageNumber="253">
<td id="F16F9E9147180C5B6B98EA211D559BD8" colspan="1" pageId="4" pageNumber="253" rowspan="1">SLP4</td>
</tr>
<tr id="2804157C4A6EABEB62253A77F428E81D" pageId="4" pageNumber="253">
<td id="50BE2A496A5B6268547072CB6528AD64" colspan="1" pageId="4" pageNumber="253" rowspan="1">PCT</td>
</tr>
<tr id="5CDA87304DA9F4E96564640529D1FD7A" pageId="4" pageNumber="253">
<td id="3418914A617CF1FAB61218D297E18E73" colspan="1" pageId="4" pageNumber="253" rowspan="1">TLT</td>
</tr>
</table>
</paragraph>
<paragraph id="6E76E25841E886066DF5E9FA771AF941" pageId="4" pageNumber="253">Colour after 10 years in alcohol (Fig. 3): head dorsally beige with almost no recognizable pattern except a whitish dorsolateral spot above each ear opening and a well-defined blackish area above each eye which represents, however, no pigmentation of the skin, but is due to the blackish eyeball shining through the skin. Dorsum greyish with four poorly defined beige spots on the back which are the remains of dorsal crossbands: one distinct spot between the forelimbs, two poorly recognizable spots on the back between forelimbs and hindlimbs, and a fourth poorly recognizable spot between the hindlimbs. Dorsal surfaces of forelimbs and hindlimbs marbled with beige and grey. Flanks with similar colour as dorsum, but without any recognizable traces of light crossbands. Tail dorsally with whitish-grey and brown alternating transverse bands which are poorly delimited in the distal portion of the tail. Throat, chest, venter, ventral parts of forelimbs and hindlimbs and ventral side of tail whitish. Iris dark grey, pupil white. Colour of holotype in life unknown (no colour photographs available).</paragraph>
<caption id="3241F0EFB81F71699E0E9BB50B171C88" pageId="4" pageNumber="253">
<paragraph id="F3756C21DB2DF53E5BD0A8E67E3818A7" pageId="4" pageNumber="253">
Figure 3. Dorsal view of the male holotype of
<taxonomicName id="11750F9B204FCBE1B0DE21511A9C99F2" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="4" pageNumber="253" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
in preservative (ZSM 342/2004).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="32658457746A8137312D25F673FB83D1" pageId="5" pageNumber="254" type="variation">
<paragraph id="17EC16BC19CED09D317C3557B8E16967" pageId="5" pageNumber="254">
<pageBreakToken id="63339E32D4343422078E7E11C02D833E" pageId="5" pageNumber="254" start="start">Variation</pageBreakToken>
.
</paragraph>
<paragraph id="1069D79BF336858BF565A2F7F9BA0D88" pageId="5" pageNumber="254">
Morphometric and meristic variation of ten specimens of the type series is summarized in Table 1, but there is also a remarkable individual variation in dorsal colouration and pattern (Fig. 2). Paratype ZSM 2106/2007, the smallest known juvenile (SVL 28.1 mm, tail length 25.2 mm), shows a moderately distinct juvenile colouration both in life (Fig. 2) and in preservative consisting of four light crossbands on the back between the insertion of forelimbs and the insertion of hindlimbs. A similarly distinct banding on body and original tail is still visible in the larger juvenile paratype ZSM 2107/2007 (Fig. 2, SVL 35.2 mm, tail length 34.7 mm). Although the juvenile colouration is distinct compared to most adults, it is less colourful and less contrasting in comparison with many other
<taxonomicName id="2C51A9DF8F1C8F0D0AABA5E2524349B3" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="5" pageNumber="254" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
species. The adult paratypes usually have a less distinct dorsal colour pattern than the juveniles, ranging from poorly recognizable (e. g. ZSM 532/2000) to distinct and well delimited (e. g. ZSM 531/2000) after 14 years in alcohol. Additional photographs of living individuals are provided in
<bibRefCitation id="139BF29D34526AE38685834A587201DE" author="Henkel, FW" journalOrPublisher="Ulmer Verlag, Stuttgart" pageId="7" pageNumber="256" title="Amphibien und Reptilien Madagaskars, der Maskarenen, Seychellen und Komoren." year="1995">Henkel and Schmidt (1995)</bibRefCitation>
,
<bibRefCitation id="202832EC05FBDF852C3EF9E596EB9696" author="Glaw, F" journalOrPublisher="Vences and Glaw Verlag, Cologne" pageId="7" pageNumber="256" title="A field guide to the amphibians and reptiles of Madagascar" year="2007">Glaw and Vences (2007)</bibRefCitation>
, and
<bibRefCitation id="AB11C67B1208A49F81928E2E74A91A57" author="Schoenecker, P" journalOrPublisher="Edition Chimaira, Terralog" pageId="8" pageNumber="257" pagination="1 - 144" title="Geckos of Madagascar, the Seychelles, Comoros and Mascarene Islands." volume="12" year="2008">
<normalizedToken id="479F9D54240D12532CA64D66E5CB383A" originalValue="Schönecker">Schoenecker</normalizedToken>
(2008)
</bibRefCitation>
. Most of the adult type specimens have a regenerated tail (without distinct transversal banding of dark and white), indicating a high pressure by predation or intraspecific aggression.
</paragraph>
</subSubSection>
<subSubSection id="4AB44E67E8EF7796CDAE7AB64BA2B195" pageId="5" pageNumber="254" type="hemipenis">
<paragraph id="35F83203D231A9922B16975E768C5128" pageId="5" pageNumber="254">Hemipenis.</paragraph>
<paragraph id="CFA0D6500BCD58D08001ADE983F78C64" pageId="5" pageNumber="254">The following description is based on the everted right organ of ZSM 343/2004 (Fig. 4). Hemipenis medium-sized (total length 6.4 mm, width at apex 4.5 mm), apex divided in two lobes. Sulcus spermaticus forms an S-shaped, narrow and deep groove, bordered by moderately distinct lips reaching the apex. On the apex, the sulcus is becoming broader and divides in two branches. A horn-like apical cone is present in the center of each lobe. Calyces on the apex are only present at the border of the lobes. The hemipenes of the holotype (ZSM 342/2004) are virtually identical in every respect.</paragraph>
<caption id="E2ECD8E0FC38F0B37E824C7E9A704C59" pageId="5" pageNumber="254">
<paragraph id="8E92133523F30583498D7B177AC3254E" pageId="5" pageNumber="254">
Figure 4. Drawings of hemipenis of
<taxonomicName id="1B036B8CDBB7E0E19BC7F8F3CD4F3F76" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
(ZSM 343/2004): (A) asulcal view; (B) lateral view; (C) detailed view on cranial lobe.
</paragraph>
</caption>
<caption id="712173DB92ECA81EA24FDF93B93B357E" pageId="5" pageNumber="254">
<paragraph id="62C76330E9EC684A6F57AAAD421ADB90" pageId="5" pageNumber="254">
Figure 5.
<taxonomicName id="3CDC5EEA0D8BDA89616D8621D81C5C5B" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="karstophila">Paroedura karstophila</taxonomicName>
from its type locality Tsingy de Namoroka (A), observed in the night of 10 September 2012 around 22:50 (not collected), and
<taxonomicName id="CFE0B6FEC83E374D27FE00695C1B95EE" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
from its type locality Ankarana (B).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F7FA9ADB9128CADCFD9A854BF6996599" pageId="5" pageNumber="254" type="habitat and habits">
<paragraph id="5ABA44537BC15EB26EAD6C80F38DA1F8" pageId="5" pageNumber="254">Habitat and habits.</paragraph>
<paragraph id="8727E578F3666B7ECDC889E52A4AEC9D" pageId="5" pageNumber="254">
<taxonomicName id="D695A5AB81C68739AA6F34323B5EB19A" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
was observed multiple times at night in karstic dry forest in the rainy season, mainly climbing on karstic rocks and the ruins of an old fort. It was found in close syntopy with
<taxonomicName id="EA4F8232D0D55FCB9B79D693A49FB077" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura lohatsara" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="lohatsara">Paroedura lohatsara</taxonomicName>
in the karstic limestone areas, whereas
<taxonomicName id="EBFBDD3A82610FA72F0A790B6C7152D8" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
was only encountered on the slope between the massif and the sea, in areas without karstic formations.
</paragraph>
</subSubSection>
<subSubSection id="92E03454BE49AD721B02974AE7211972" pageId="5" pageNumber="254" type="etymology">
<paragraph id="1D93EF6BC120FC6B51794DD93CDA1869" pageId="5" pageNumber="254">Etymology.</paragraph>
<paragraph id="8EE1AD90ADE535ABFC2BF929CF4AF83F" pageId="5" pageNumber="254">The specific name is dedicated to Freddy Hordies, in recognition of his support for biodiversity research and conservation through the BIOPAT initiative.</paragraph>
</subSubSection>
<subSubSection id="9F2577EC1C3373D32BBBB1DA8D8D9E0B" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254" type="distribution and conservation status">
<paragraph id="3265658E6A92BEE701FDDE33DF7AA5F7" pageId="5" pageNumber="254">Distribution and conservation status.</paragraph>
<paragraph id="A9CCD2E67AD2BA7989F3FC378724F39B" pageId="6" pageNumber="255">
<taxonomicName id="C97AB7B873F96579DAAF64B3266C5691" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="hordiesi">
<pageBreakToken id="9A17106C719E717973D2873A788587C6" pageId="6" pageNumber="255" start="start">Paroedura</pageBreakToken>
hordiesi
</taxonomicName>
is reliably known only from the recently established nature reserve of Montagne des
<normalizedToken id="4017C140041917890E764A4327BF7DB2" originalValue="Français">Francais</normalizedToken>
. The species possibly also occurs at Ampombofofo, ca. 25 km north of this massif (
<bibRefCitation id="7DB5F12E8C0F33936D23BC10F651ED1B" author="Megson, S" journalOrPublisher="Herpetology Notes" pageId="7" pageNumber="256" pagination="31 - 44" title="A comprehensive survey of amphibians and reptiles in the extreme north of Madagascar." volume="2" year="2009">Megson et al. 2009</bibRefCitation>
: ZSM 1531/2008), but the identity of this population needs further study as molecular data are not available thus far. Surveys in other limestone areas of northern Madagascar revealed a superficially similar
<taxonomicName id="CDF88445404BA0234EDBEE61B45014A5" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="6" pageNumber="255" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
at Nosy Hara (
<bibRefCitation id="54800F8C4C6F82EB948159A1FED908A1" author="Metcalf, CJE" journalOrPublisher="Tropical Zoology" pageId="7" pageNumber="256" pagination="151 - 161" title="Herpetofaunal assemblages on seven offshore islands of northwestern Madagascar." volume="20" year="2007">Metcalf et al. 2007</bibRefCitation>
) which is, however, strongly differentiated by colouration (see photo in
<bibRefCitation id="B1FBDE8DD9462F50DE5FC7CF7DC6D26B" author="Glaw, F" journalOrPublisher="Vences and Glaw Verlag, Cologne" pageId="7" pageNumber="256" title="A field guide to the amphibians and reptiles of Madagascar" year="2007">Glaw and Vences 2007</bibRefCitation>
) and mitochondrial DNA sequences (Fig. 1 and
<bibRefCitation id="1990141E7958466F8F3F2FB09435D033" author="Nagy, ZT" journalOrPublisher="PLoS ONE" pageId="8" pageNumber="257" title="First large-scale DNA barcoding assessment of reptiles in the biodiversity hotspot of Madagascar, based on newly designed COI primers." url="10.1371/journal.pone.0034506" volume="7" year="2012">Nagy et al. 2012</bibRefCitation>
), and is therefore considered a further undescribed candidate species. The Ankarana massif is known to harbour
<taxonomicName id="A38A93540B836E3A80666688C58F48C7" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
,
<taxonomicName id="50130FBB6A6826E3FB1B617A2B963C29" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura karstophila" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="karstophila">Paroedura karstophila</taxonomicName>
, and
<taxonomicName id="287D099F87D91A060346BE9DFDF7A2E1" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
(
<bibRefCitation id="2BE63F0633D14CDE44F967B8349D206B" author="Nussbaum, RA" journalOrPublisher="Miscellaneous Publications Museum of Zoology, University of Michigan" pageId="8" pageNumber="257" pagination="1 - 26" title="Systematic revision of the genus Paroedura Guenther (Reptilia: Squamata: Gekkonidae), with description of five new species." volume="189" year="2000">Nussbaum and Raxworthy 2000</bibRefCitation>
,
<bibRefCitation id="55745277DD7C7060E911F27F01644E23" author="Raselimanana, AP" journalOrPublisher="Malagasy Nature" pageId="8" pageNumber="257" pagination="46 - 75" title="Herpetofaune des forets seches malgaches." volume="1" year="2008">Raselimanana 2008</bibRefCitation>
). Our own surveys in Ankarana revealed only
<taxonomicName id="FEEBB1DE2A3773E340194C7CEE46B762" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura homalorhina" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="homalorhina">Paroedura homalorhina</taxonomicName>
and
<taxonomicName id="5E6F9002BF4E15A4E86666E89044EAE8" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura stumpffi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="stumpffi">Paroedura stumpffi</taxonomicName>
and we consider the occurrence of
<taxonomicName id="C684CFDE47B8C206465E72080EB5A54A" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
unlikely in this massif.
<bibRefCitation id="AEDDC1969CD72A0786FA69C8380555E0" author="Rakotondravony, HA" editor="Goodman, SM" journalOrPublisher="Recherche pour le Developpement (Serie Sciences Biologiques)" pageId="8" pageNumber="257" pagination="101 - 148" title="Patterns de la diversite des reptiles et amphibiens de la region de Loky-Manambato." volume="23" volumeTitle="Inventaires de la faune et de la flore du nord de Madagascar dans la region Loky-Manambato, Analamerana et Andavakoera." year="2006 a">Rakotondravony (2006a)</bibRefCitation>
found
<taxonomicName id="91616B8124B29E7CAAB37EFE186CE548" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="6" pageNumber="255" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
sp. at Binara in the Loky-Manambato region which potentially could refer to
<taxonomicName id="CD2398BDF4BB412FBBA986A62B7FB53D" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
, but the identity of this record remains to be studied. No unidentified
<taxonomicName id="350256AE79FC0E883F616039C8046340" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura" pageId="6" pageNumber="255" phylum="Chordata" rank="genus">Paroedura</taxonomicName>
species was found at Analamera (
<bibRefCitation id="B4CCDDCFFF3920E457E64E621F307C28" author="Rakotondravony, HA" editor="Goodman, SM" journalOrPublisher="Recherche pour le Developpement, Serie Sciences Biologiques" pageId="8" pageNumber="257" pagination="149 - 173" title="Reptiles et amphibiens de la Reserve Speciale d'Analamerana et de la foret classee d'Andavakoera dans l'extreme Nord de Madagascar." volume="23" volumeTitle="Inventaires de la faune et de la flore du nord de Madagascar dans la region Loky-Manambato, Analamerana et Andavakoera." year="2006 b">Rakotondravony 2006b</bibRefCitation>
). Thus current evidence suggests that
<taxonomicName id="30784A396B0A942A26627B909CB65A22" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
is microendemic of Montagne des
<normalizedToken id="9A9DDE1E87A56754D9A20026FC14C42C" originalValue="Français">Francais</normalizedToken>
and perhaps the adjacent Ampombofofo region.
</paragraph>
<paragraph id="EF4BDBA6CFECF45BB7B2DC5B4A9C3DAE" pageId="6" pageNumber="255">
For consistency with the IUCN Red List Assessment for
<taxonomicName id="F98171BCC5883D4605DB2913FD7E8428" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura lohatsara" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="lohatsara">Paroedura lohatsara</taxonomicName>
(
<bibRefCitation id="6421C0FBC0C49B958E213FF9E7545898" author="Raxworthy, CJ" journalOrPublisher="Copeia" pageId="8" pageNumber="257" title="The IUCN Red List of Threatened species" url="http://www.iucnredlist.org" year="2011">Raxworthy et al. 2011</bibRefCitation>
) and other potential microendemic species of the Montagne des
<normalizedToken id="7A0D4DE531F887EF2514D00A3C5FD1CD" originalValue="Français">Francais</normalizedToken>
region, we suggest a classification as &quot;Critically Endangered&quot; on the basis that
<taxonomicName id="5A4309030239351B70E07FFB0A6CC168" class="Squamata" family="Gekkonidae" genus="Paroedura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroedura hordiesi" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="hordiesi">Paroedura hordiesi</taxonomicName>
has an extent of occurrence of at most 50 km², it is known from a single location, and there is a continuing decline in the extent and quality of its habitat.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.796.20736" ID-GBIF-Dataset="38274b4e-7bec-4674-964b-763328eb9f5c" ID-PMC="PMC6250776" ID-Pensoft-Pub="1313-2970-796-83" ID-PubMed="30487711" ID-ZBK="DB483AA50384461F8CF479E4939BB244" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-796-83" ModsDocOrigin="ZooKeys 796" ModsDocTitle="Miscellanea Miridologica V. Taxonomy and chorology of new or little known taxa of Continental New Guinea and neighboring islands (Insecta, Heteroptera, Miridae)" checkinTime="1542332326465" checkinUser="pensoft" docAuthor="Cerot, Frederic" docDate="2018" docId="FCBD8A3B597A30AB3724BCDEA54644A5" docLanguage="en" docName="ZooKeys 796: 83-95" docOrigin="ZooKeys 796" docSource="http://dx.doi.org/10.3897/zookeys.796.20736" docTitle="Prolygus alboscutellata Carvalho 1987" docType="treatment" docVersion="5" lastPageNumber="89" masterDocId="FFA6E751DB09FFB6FFB8FF8AB97DFF81" masterDocTitle="Miscellanea Miridologica V. Taxonomy and chorology of new or little known taxa of Continental New Guinea and neighboring islands (Insecta, Heteroptera, Miridae)" masterLastPageNumber="95" masterPageNumber="83" originalUpdateDomain="Boston" originalUpdateTime="1542332326465" originalUpdateUser="pensoft" pageNumber="88" updateTime="1668166433552" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Miscellanea Miridologica V. Taxonomy and chorology of new or little known taxa of Continental New Guinea and neighboring islands (Insecta, Heteroptera, Miridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cerot, Frederic</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>796</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>95</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.796.20736</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.796.20736</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-796-83</mods:identifier>
<mods:identifier type="ZBK">DB483AA50384461F8CF479E4939BB244</mods:identifier>
<mods:identifier type="ZooBank">DB483AA50384461F8CF479E4939BB244</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="149667290" LSID="urn:lsid:plazi:treatment:FCBD8A3B597A30AB3724BCDEA54644A5" httpUri="http://treatment.plazi.org/id/FCBD8A3B597A30AB3724BCDEA54644A5" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">
<subSubSection pageId="5" pageNumber="88" type="nomenclature">
<paragraph pageId="5" pageNumber="88">
<taxonomicName authority="Carvalho, 1987 a" class="Insecta" family="Miridae" genus="Prolygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prolygus alboscutellata" order="Heteroptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="alboscutellata">Prolygus alboscutellata Carvalho, 1987a</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88" type="materials_examined">
<paragraph pageId="5" pageNumber="88">Material examined.</paragraph>
<paragraph lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">
Papua New Guinea: 1♂, 1♀, 1?, Eastern Highlands, Kegsugl, under M(oun)t Wilhelm (coordinates provided on the label:
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-5.831111">05°49'52&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="145.0861">145°05'10&quot;E</geoCoordinate>
),
<pageBreakToken pageId="6" pageNumber="89" start="start">2780</pageBreakToken>
m, 13.v.2012, dissected for parasites, dissection numbers 623-625, negative,
<normalizedToken originalValue="Votýpka">Votypka</normalizedToken>
J. &amp;
<normalizedToken originalValue="Lukeš">Lukes</normalizedToken>
J. leg. (FC n° 6420) (NMPC).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="89" type="distribution">
<paragraph pageId="6" pageNumber="89">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="89">
Endemic to New Guinea (
<bibRefCitation author="Schuh, RT" journalOrPublisher="Systematic Catalog, Distributions, Host List, and Bibliography. Entomological Society of New York" pageId="9" pageNumber="92" url="http://research.amnh.org/pbi/catalog/" year="2002 - 2013">Schuh 2002-2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,137 @@
<document id="4FA54BC561EDE3E166EBA41C9C96C2C8" ID-DOI="10.5281/zenodo.292251" ID-ISBN="3-7643-0527-4" ID-Zenodo-Dep="292251" checkinTime="1646762458513" checkinUser="donat" docAuthor="Hess, Hans Ernst, Landolt, Elias &amp; Hirzel, Rosmarie" docDate="1976" docId="FCBDED9D89CB27AAE10D33ADAA62F920" docLanguage="de" docName="rosaceae.xml" docOrigin="Birkhaeuser Verlag" docSource="https://doi.org/10.5281/zenodo.292251" docTitle="Prunus armeniaca L." docType="treatment" docVersion="4" lastPageNumber="463" masterDocId="15FA7881DCE74474E01E9DC9A8211891" masterDocTitle="Flora der Schweiz und angrenzender Gebiete. Band 2. Nymphaceae bis Primulaceae (2 nd edition) (p. 956): Rosaceae" pageNumber="463" updateTime="1669210910520" updateUser="tatiana">
<mods:mods id="FEE0BAADD1E99F08C6C25977D03759E0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B5FF2A75744DD77A10EE15C403881EC0">Flora der Schweiz und angrenzender Gebiete. Band 2. Nymphaceae bis Primulaceae (2 nd edition) (p. 956): Rosaceae</mods:title>
</mods:titleInfo>
<mods:name id="770955BDEF81D232E292BFC1C8AB9C42" type="personal">
<mods:role id="D8C0C29ABA8E6B3C61DC0C4D5D0E2DD4">
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<mods:namePart id="E39BA9B4D33A093DC0D68A03E13A1D41">Hess, Hans Ernst</mods:namePart>
</mods:name>
<mods:name id="09E5C003A4DD4254DF9C2AB41D61D54E" type="personal">
<mods:role id="24BBCA9FCD8271E15D23639F418C73CA">
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</mods:role>
<mods:namePart id="3583AB59B8426BC37EF1C6CEBC624926">Landolt, Elias</mods:namePart>
</mods:name>
<mods:name id="8E5022843452BF9AA63011B9ACD3DDF4" type="personal">
<mods:role id="2400FFFBCD8BA4A4C261F4BBCD8BE1C8">
<mods:roleTerm id="C1DC5824A339909F0ADBE6BEB7436300">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D3126C5D3ACF3946DB6E2DA689D0B15">Hirzel, Rosmarie</mods:namePart>
</mods:name>
<mods:typeOfResource id="C3C046980536E0746D12EC01032E7F64">text</mods:typeOfResource>
<mods:originInfo id="293EFA331CCAB14AF3A8FF1FB6103D1A">
<mods:dateIssued id="CFE65CC0902EE2F3ACDC77B9F11CA2F3">1976</mods:dateIssued>
<mods:publisher id="8A8FA63BCDF01BB5F2E094738C1CC6AC">Birkhaeuser Verlag</mods:publisher>
</mods:originInfo>
<mods:location id="00EB2C3F8996AF3B73A45A06E57BC756">
<mods:url id="10C56D8856ACF9BC43908C8F1C5B88FE">https://doi.org/10.5281/zenodo.292251</mods:url>
</mods:location>
<mods:classification id="E16B43AEE1B69B653350D84C7208293B">book</mods:classification>
<mods:identifier id="509C8BDA422AF437E4358C1C8F3207AE" type="Zenodo-Dep">292251</mods:identifier>
<mods:identifier id="2AAB345C54E63FFC44E97964E174F07C" type="DOI">10.5281/zenodo.292251</mods:identifier>
<mods:identifier id="71A8A272CAD95914D679A63185F9D246" type="ISBN">3-7643-0527-4</mods:identifier>
</mods:mods>
<treatment id="FCBDED9D89CB27AAE10D33ADAA62F920" LSID="urn:lsid:plazi:treatment:FCBDED9D89CB27AAE10D33ADAA62F920" httpUri="http://treatment.plazi.org/id/FCBDED9D89CB27AAE10D33ADAA62F920" lastPageNumber="463" pageId="null" pageNumber="463" sec="rosaceae.s303">
<title id="C843CBAE2BAF2D1E7A978C83B973E80D" pageId="null" pageNumber="463">
<subSubSection id="D4FE50102D518A37D68FEA4F18D12AED" pageId="null" pageNumber="463" type="nomenclature">
<paragraph id="D9E1E136372D00EE3EE14245A7B20644" pageId="null" pageNumber="463">
<taxonomicName id="16B174C03538706CBC6FEFB3F6377E5A" authorityName="L." class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="null" pageNumber="463" phylum="Tracheophyta" rank="species" species="armeniaca">
<pageBreakToken id="7F0CBA3F610500CE6050EA1EC1C2BED1" pageId="null" pageNumber="463" start="start">Prunus</pageBreakToken>
</taxonomicName>
<normalizedToken id="9470D63DE4EAB26D9319DECE93433C6D" originalValue="Armeníaca" pageId="null" pageNumber="463">Armeniaca</normalizedToken>
<authorityName id="354AAB75C7079052871C8AC4730708B2" pageId="null" pageNumber="463">L.</authorityName>
</paragraph>
</subSubSection>
<subSubSection id="CAF79697BADDA12AB7CF47CE2EBD1124" pageId="null" pageNumber="463" type="reference_group">
<paragraph id="A2286C5C044E0F596A82E08B6FB21BBD" pageId="null" pageNumber="463">
(
<taxonomicName id="4EA5470C836DD6FD547FA7100A302E03" authority="Lam." authorityName="Lam." class="Magnoliopsida" family="Rosaceae" genus="Armeniaca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="null" pageNumber="463" phylum="Tracheophyta" rank="species" species="vulgaris">
<emphasis id="DF5BFE54A075C8BC7AF3851D7E319645" italics="true" pageId="null" pageNumber="463">Armeniaca vulgaris</emphasis>
Lam.
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="16614E03480D99D7D205A8F45CF5638C" pageId="null" pageNumber="463" type="vernacular_names">
<paragraph id="6354FAAD3556CA56963FAE15EE987243" pageId="null" pageNumber="463">Aprikose</paragraph>
</subSubSection>
</title>
<subSubSection id="01BB45B63C7C969CF28D0752FBF823EA" pageId="null" pageNumber="463" type="description">
<paragraph id="685B0C4C25C2DDC6C44EDE1012989070" pageId="null" pageNumber="463">
Bis 4 m hoher Strauch. Junge Triebe kahl.
<emphasis id="29A0FF49EE45E289365590AFF1407FF0" italics="true" pageId="null" pageNumber="463">
<normalizedToken id="80977C0BF0EF10847AD678510D92C402" originalValue="Blätter" pageId="null" pageNumber="463">Blaetter</normalizedToken>
rundlich bis
<normalizedToken id="0901B0BB8F527C322392A4984BCE97D3" originalValue="herzförmig" pageId="null" pageNumber="463">herzfoermig</normalizedToken>
, 5-10 cm lang, 1-1
</emphasis>
<normalizedToken id="8C96E0856F6EA7F976C4CA09138086AB" originalValue="½" pageId="null" pageNumber="463">1/2</normalizedToken>
<emphasis id="6272CEC7488D34F8A69DB61CE4D2F8AD" italics="true" pageId="null" pageNumber="463">
-mal so lang wie breit, spitz, am Grunde wenig ausgerandet und in den Stiel
<normalizedToken id="9943923E86BF8A98005A3A4673B6EDDD" originalValue="verschmälert" pageId="null" pageNumber="463">verschmaelert</normalizedToken>
</emphasis>
, beiderseits
<normalizedToken id="07BC53A0C1A02C05FF3FA56A17309FC8" originalValue="dunkelgrün" pageId="null" pageNumber="463">dunkelgruen</normalizedToken>
, unterseits in den innern Nervenwinkeln
<normalizedToken id="6A52C53D20726701AEFB91870C6BBA55" originalValue="bärtig" pageId="null" pageNumber="463">baertig</normalizedToken>
behaart, sonst kahl, am Rande
<emphasis id="E86173964ABC43D0F0430199A65B9CA2" italics="true" pageId="null" pageNumber="463">
oft doppelt
<normalizedToken id="B61F68F971E2222101B289F104239AE6" originalValue="gezähnt" pageId="null" pageNumber="463">gezaehnt</normalizedToken>
, in der Knospenlage eingerollt;
</emphasis>
am Blattstiel mit einigen
<normalizedToken id="0C16009955813B252630838E51828AC3" originalValue="Drüsen" pageId="null" pageNumber="463">Druesen</normalizedToken>
.
<normalizedToken id="57FF2FB4E17B555C616C1928E5EAFB8A" originalValue="Blüten" pageId="null" pageNumber="463">Blueten</normalizedToken>
einzeln oder in Paaren.
<normalizedToken id="5E30FDAB29F152A80A3C5307D01A5C49" originalValue="Blütenstiele" pageId="null" pageNumber="463">Bluetenstiele</normalizedToken>
flaumig behaart,
<normalizedToken id="FE0B5E3A16A3E5B861DF24CD7D5AF9B5" originalValue="kürzer" pageId="null" pageNumber="463">kuerzer</normalizedToken>
als 5 mm.
<normalizedToken id="6F0723260479A7E77674E6CE34A92E40" originalValue="Kelchblätter" pageId="null" pageNumber="463">Kelchblaetter</normalizedToken>
ganzrandig, flaumig behaart.
<normalizedToken id="175845030124FD0C563092A40F8156F0" originalValue="Kronblätter" pageId="null" pageNumber="463">Kronblaetter</normalizedToken>
rundlich bis oval, 1-1,5 cm lang, rosa bis
<normalizedToken id="904754F90A11AEEB0016910CF2FE1FC1" originalValue="weiß" pageId="null" pageNumber="463">weiss</normalizedToken>
.
<normalizedToken id="07AB83B2E539EE358E45E552CD9582B5" originalValue="Staubblätter" pageId="null" pageNumber="463">Staubblaetter</normalizedToken>
etwa 20; Staubbeutel gelb. Fruchtknoten und Frucht weich behaart; Frucht fast kugelig, im
<normalizedToken id="F13E4BB8BF6C734FCD1D08BE06A464CB" originalValue="größten" pageId="null" pageNumber="463">groessten</normalizedToken>
Durchmesser 4-8 cm, mit einseitiger
<normalizedToken id="E5977806B92CCD4CD1CBD6D19E7B148E" originalValue="Längsfurche" pageId="null" pageNumber="463">Laengsfurche</normalizedToken>
, gelb bis rot, mit sich leicht vom Stein
<normalizedToken id="A017F197F03F89D42A058155F6BC0DE9" originalValue="lösendem" pageId="null" pageNumber="463">loesendem</normalizedToken>
Fruchtfleisch und etwas rauhem, scharfkantigem Stein. -
<normalizedToken id="481679206180C4E1EDF7A1769C254F7B" originalValue="Blüte" pageId="null" pageNumber="463">Bluete</normalizedToken>
:
<normalizedToken id="950C344F3399DB8654C12174B4233D97" originalValue="Frühling" pageId="null" pageNumber="463">Fruehling</normalizedToken>
.
</paragraph>
<paragraph id="E897C03A98B7713D6AF3E8A7E1C5879C" pageId="null" pageNumber="463">
<emphasis id="2C8B4CDA7A54B35931E8E9D78EEBC46C" italics="true" pageId="null" pageNumber="463">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="1E791FB78FAFCAA4BE02723187CC8318" italics="true" pageId="null" pageNumber="463">16:</emphasis>
3 Kultursorten aus der Schweiz untersucht, keine
<normalizedToken id="B54FBE215494B74EE10E5D55C02A5D0B" originalValue="Störungen" pageId="null" pageNumber="463">Stoerungen</normalizedToken>
in der Pollenmeiose, normale
<normalizedToken id="7902E9A19384679D8D5B2140EACA2F54" originalValue="Pollenkörner" pageId="null" pageNumber="463">Pollenkoerner</normalizedToken>
(Kobel 1927).
</paragraph>
<paragraph id="F76D8B7FDB5DB61F65597B13451C00D3" pageId="null" pageNumber="463">
<emphasis id="FCC5A6804618203CDAB4E3FF9A144365" italics="true" pageId="null" pageNumber="463">Standort.</emphasis>
Angepflanzt in Weinbaugebieten; im Gebiet kaum verwildert.
</paragraph>
<paragraph id="D95868386B3F726CD74ADBCF089F0E58" pageId="null" pageNumber="463">
<emphasis id="C97B7DA236245B75297F84DA8B4AA8C7" italics="true" pageId="null" pageNumber="463">Verbreitung.</emphasis>
<normalizedToken id="87D2E0C89D11D46DD4B3035797F63A9B" originalValue="Ursprünglich" pageId="null" pageNumber="463">Urspruenglich</normalizedToken>
wahrscheinlich
<emphasis id="2D586FBAE3E08F6CF5E690616FAAC35B" italics="true" pageId="null" pageNumber="463">ostasiatische Pflanze</emphasis>
(China). Seit der Antike im Gebiet angepflanzt.
</paragraph>
</subSubSection>
</treatment>
</document>