Add updates up until 2025-02-25 22:04:18

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(Hawaiian Islands and
<collectingCountry id="E4163FD1C58A3AF74ECB338C0C6CB5AF" name="New Caledonia">New Caledonia</collectingCountry>
). —
<treatmentCitation id="C2A7D819299B72DA6BB47B2BF8C22818" author="Yoshino T" page="167" year="1984">
<treatmentCitation id="BEAE3411592F5C443C4029BCE6FD268E">
<bibRefCitation id="AB06055FD2AA211A29C20877200675D4" author="Yoshino T" firstAuthor="Yoshino" refId="B59" refString="Yoshino T (1984) Randallichthys filamentosus. P. 167. In: Masuda H, Amaoka K, Araga C, Uyeno T, Yoshino T (Eds.) The fishes of the Japanese Archipelago. Text. Tokai University Press, Tokyo, Japan." year="1984">Yoshino (1984)</bibRefCitation>
</treatmentCitation>
: 167
</treatmentCitation>
(southern
@ -171,9 +161,7 @@ and
<collectingCountry id="792A714A423786D8A1E3EE84802C1E12" name="New Caledonia">New Caledonia</collectingCountry>
). —
<treatmentCitation id="AC29A7E1F07E7211517DECA5B96A7480" author="Allen GR" page="156" year="1985">
<treatmentCitation id="5A59B942EABA3B49BACFE10057E29F25">
<bibRefCitation id="1045ACF8A343AA053A57400EFE292A73" author="Allen GR" firstAuthor="Allen" journalOrPublisher="FAO, Rome" refId="B3" refString="Allen GR (1985) FAO species catalogue. Vol. 6. Snappers of the world. An annotated and illustrated catalogue of lutjanid species known to date. FAO, Rome, 208 pp." title="FAO species catalogue. Vol. 6. Snappers of the world. An annotated and illustrated catalogue of lutjanid species known to date." year="1985">Allen (1985)</bibRefCitation>
</treatmentCitation>
: 156
</treatmentCitation>
(southern
@ -182,82 +170,62 @@ and
<collectingCountry id="FB5EE2E4412ADE66A29021F2CA192EAE" name="New Caledonia">New Caledonia</collectingCountry>
). —
<treatmentCitation id="00BCBE674133AA0072219D6A1FDA9196" author="Anderson WDJ &amp; Allen GR" page="2916" year="2001">
<treatmentCitation id="B5FC97B0D2C6A3F33D972432B86363BF">
<bibRefCitation id="740D3D2EA047ECF3B02CF8AE3448996C" author="Anderson WDJ &amp; Allen GR" firstAuthor="Anderson" refId="B6" refString="Anderson WDJ, Allen GR (2001) Lutjanidae. Snappers (jobfishes). Pp. 28402918. In: Carpenter KE, Niem VH (Eds.) Species identification guide for fisheries purposes. The living marine resources of the western central Pacific. Bony fishes part 3 (Menidae to Pomacentridae). FAO, Rome." year="2001">Anderson and Allen (2001)</bibRefCitation>
</treatmentCitation>
: 2916
</treatmentCitation>
(western central Pacific). —
<treatmentCitation id="29A26A8CC27EEEA78C46DB3A38B10F51" author="Shimada K" page="827" year="2002">
<treatmentCitation id="71FD1F54E69994831C133789CBCBC81D">
<bibRefCitation id="CE24629D73642E31A0F2E83B8C41B5CE" author="Shimada K" firstAuthor="Shimada" refId="B52" refString="Shimada K (2002) Lutjanidae. Labridae. Pp. 819832, 9691013, 15731579. In: Nakabo T (Ed.) Fishes of Japan with Pictorial Keys to the Species, English Edition. Tokai University Press, Tokyo, Japan." year="2002">Shimada (2002)</bibRefCitation>
</treatmentCitation>
: 827
</treatmentCitation>
(southern
<collectingCountry id="2786DB944D955D7A70D76157A5334E94" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="893C79246D1989E9F38269E6AEF4057E" author="Randall JE &amp; Bacchet P &amp; Winterbottom R &amp; Wrobel L" page="158" year="2002">
<treatmentCitation id="145664B4636220CC544D9B8E2631694B">
<bibRefCitation id="446FB0B5B153517A9BDC4AF7F077EA7C" author="Randall JE &amp; Bacchet P &amp; Winterbottom R &amp; Wrobel L" etAl="et al." firstAuthor="Randall" issue="4" journalOrPublisher="Journal of Ichthyology and Aquatic Biology" pagination="153-166" refId="B47" refString="Randall JE, Bacchet P, Winterbottom R, Wrobel L (2002) Fifty new records of shore fishes from the Society Islands and Tuamotu Archipelago. Aqua. Journal of Ichthyology and Aquatic Biology 5 (4): 153166." title="Fifty new records of shore fishes from the Society Islands and Tuamotu Archipelago. Aqua." volume="5" year="2002">Randall et al. (2002)</bibRefCitation>
</treatmentCitation>
: 158
</treatmentCitation>
(Society and Tuamotu islands). —
<treatmentCitation id="3BF93D651324D0B5BD4948387B2675E3" author="Myers RF &amp; Donaldson TJ" page="626" year="2003">
<treatmentCitation id="B5D432B5B5D96E4D0B9911A8A73D1320">
<bibRefCitation id="15FEFBE923C4EFC3B1E56E5BDF5C967D" author="Myers RF &amp; Donaldson TJ" firstAuthor="Myers" refId="B38" refString="Myers RF, Donaldson TJ (2003) The fishes of the Mariana Islands. Micronesica 3536: 598652." year="2003">Myers and Donaldson (2003)</bibRefCitation>
</treatmentCitation>
: 626
</treatmentCitation>
(
<collectingCountry id="7C211C5400C52D4FBFB6DB669450B2B4" name="Northern Mariana Islands">Mariana Islands</collectingCountry>
). —
<treatmentCitation id="D644F93E6DBBDD4124B9F8ABBE146660" author="Randall JE &amp; Williams JT &amp; Smith DG &amp; Kulbicki M &amp; Mou Tham G &amp; Labrosse P &amp; Kronen M &amp; Clua E &amp; Mann BS" page="16" year="2004">
<treatmentCitation id="37285E7DCAE9A3D5F62DEAF54599EF68">
<bibRefCitation id="5729830D6B2B8CEBFFC90E75AF70A20C" DOI="10.5479/si.00775630.502.1" author="Randall JE &amp; Williams JT &amp; Smith DG &amp; Kulbicki M &amp; Mou Tham G &amp; Labrosse P &amp; Kronen M &amp; Clua E &amp; Mann BS" etAl="et al." firstAuthor="Randall" journalOrPublisher="Atoll Research Bulletin" pagination="1-35" refId="B48" refString="Randall JE, Williams JT, Smith DG, Kulbicki M, Mou Tham G, Labrosse P, Kronen M, Clua E, Mann BS (2004) Checklist of the shore and epipelagic fishes of Tonga. Atoll Research Bulletin 502: 135. https://doi.org/10.5479/si.00775630.502.1" title="Checklist of the shore and epipelagic fishes of Tonga." volume="502" year="2004">Randall et al. (2004)</bibRefCitation>
</treatmentCitation>
: 16
</treatmentCitation>
(
<collectingCountry id="8593F2344EB38518560B7D8311317E5E" name="Tonga">Tonga</collectingCountry>
). —
<treatmentCitation id="25A2B7B33CC6041AB9053188C1452DFE" author="Leis JM" page="58" year="2005">
<treatmentCitation id="20CB6E57691E295CA07F27BF9C0B7B9B">
<bibRefCitation id="28D5E423D370D5CD86F5D6374CC0C970" DOI="10.11646/zootaxa.1008.1.7" author="Leis JM" firstAuthor="Leis" issue="1" journalOrPublisher="Zootaxa" pagination="57-64" refId="B32" refString="Leis JM (2005) A larva of the eteline lutjanid, Randallichthys filamentosus (Pisces: Perciformes), with comments on phylogenetic implications of larval morphology of basal lutjanids. Zootaxa 1008 (1): 5764. https://doi.org/10.11646/zootaxa.1008.1.7" title="A larva of the eteline lutjanid, Randallichthys filamentosus (Pisces: Perciformes), with comments on phylogenetic implications of larval morphology of basal lutjanids." volume="1008" year="2005">Leis (2005)</bibRefCitation>
</treatmentCitation>
: 58
</treatmentCitation>
(
<collectingCountry id="74FF1C34834D130236AB2C3F088936C9" name="Micronesia">Caroline Islands</collectingCountry>
);
<treatmentCitation id="E49000AB9921BD58CE9EE6D68A76E8D3" author="Mundy BC" page="385" year="2005">
<treatmentCitation id="19C7B45BC382B9D81355D3C2DA3667F8">
<bibRefCitation id="95DF1E2B5F6EB43752AEBE5A17A92B6D" author="Mundy BC" firstAuthor="Mundy" journalOrPublisher="Bishop Museum Bulletins in Zoology" pagination="1-703" refId="B37" refString="Mundy BC (2005) Checklist of the fishes of the Hawaiian Archipelago. Bishop Museum Bulletins in Zoology 6: 1703." title="Checklist of the fishes of the Hawaiian Archipelago." volume="6" year="2005">Mundy (2005)</bibRefCitation>
</treatmentCitation>
: 385
</treatmentCitation>
(Hawaiian Islands). —
<treatmentCitation id="7590C5892AFDA182C1B75BA438842806" author="Randall JE &amp; Myers RF &amp; Trevor MN &amp; Johnson SR &amp; Johnson JL &amp; Yoshi S &amp; Greene BD" page="123" year="2005">
<treatmentCitation id="93C2BD0C216B4AC3F4F83CFFD169E8AF">
<bibRefCitation id="5436152847152DDA80092C87A0135076" author="Randall JE &amp; Myers RF &amp; Trevor MN &amp; Johnson SR &amp; Johnson JL &amp; Yoshi S &amp; Greene BD" etAl="et al." firstAuthor="Randall" issue="3" journalOrPublisher="Journal of Ichthyology and Aquatic Biology" pagination="115-132" refId="B49" refString="Randall JE, Myers RF, Trevor MN, Johnson SR, Johnson JL, Yoshi S, Greene BD (2005) Ninety-one new records of fishes from the Marshall Islands. Aqua. Journal of Ichthyology and Aquatic Biology 9 (3): 115132." title="Ninety-one new records of fishes from the Marshall Islands. Aqua." volume="9" year="2005">Randall et al. (2005)</bibRefCitation>
</treatmentCitation>
: 123
</treatmentCitation>
(
<collectingCountry id="1603AEE7D961A5B2FC0372DC2AEC1E1B" name="Marshall Islands">Marshall Islands</collectingCountry>
). —
<treatmentCitation id="1BE915B0DAF0A10C5A9439185D1258F9" author="Randall JE" page="251" year="2007">
<treatmentCitation id="49DD9FE5E63C8405F67E7BA6AFF14C88">
<bibRefCitation id="583C97B05473F55B8F0D8310B6497127" author="Randall JE" firstAuthor="Randall" journalOrPublisher="Sea Grant College Program, University of Hawaii, Honolulu, HI, USA" refId="B44" refString="Randall JE (2007) Reef and shore fishes of the Hawaiian Islands. Sea Grant College Program, University of Hawaii, Honolulu, HI, USA, 546 pp." title="Reef and shore fishes of the Hawaiian Islands." year="2007">Randall (2007)</bibRefCitation>
</treatmentCitation>
: 251
</treatmentCitation>
(Hawaiian Islands). —
<treatmentCitation id="009D9199B276D5D1E0695C7F363AA593" author="Newman SJ" page="1514" year="2009">
<treatmentCitation id="7A634A611F686DDAE7D4E6C9E77B1B4A">
<bibRefCitation id="A57812CBFDBBE564F2F38BA7D12AF224" DOI="10.1111/j.1095-8649.2009.02382.x" author="Newman SJ" firstAuthor="Newman" issue="6" journalOrPublisher="Journal of Fish Biology" pagination="1513-1517" refId="B39" refString="Newman SJ (2009) First record of Randalls snapper Randallichthys filamentosus (Perciformes: Lutjanidae) from the eastern Indian Ocean (north-western Australia). Journal of Fish Biology 75 (6): 15131517. https://doi.org/10.1111/j.1095-8649.2009.02382.x" title="First record of Randalls snapper Randallichthys filamentosus (Perciformes: Lutjanidae) from the eastern Indian Ocean (north-western Australia)." volume="75" year="2009">Newman (2009)</bibRefCitation>
</treatmentCitation>
: 1514
</treatmentCitation>
(Scott Reef, northwestern
@ -274,20 +242,32 @@ and
<heading id="CD83B0F95B98B3055354AD5FFBE53F64" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="48B7CB0E668BA5DFB78E2D4D414B747B">
<abbrev content-type="institution" id="ABBRID0EGZAE" xlink_title="Research Center for Oceanography">
<collectionCode id="E4A16A91473F05C30DF15567BD1DD30F">LBRC-F</collectionCode>
</abbrev>
2425, 175.0 mm SL, off Lembeh Island, Bitung,
<collectingCountry id="C29EF1365C875675A329218FC73DDD1C" name="Indonesia">Indonesia</collectingCountry>
, hook and line, more than
<quantity id="87DC459FDEF12CDD38D8ABBA7560C697" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
depth (purchased at Girian Fish Market, Bitung, northern
<collectingRegion id="8BCF5F268518AB933A622603CD13D8CA" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
<materialsCitation id="FB9BCF7E5E8893755D20DC1821503D01" collectingDate="2011-12-20" collectingMethod="hook and line" collectionCode="LBRC-F" collectorName="T. Peristiwady" country="Indonesia" county="Bitung" elevation="100" location="off Lembeh Island" specimenCode="LBRC-F 2425" specimenCount="1">
<specimenCode id="7F00EEEA4F05E5EB88EC281258116D0D">
<abbrev content-type="institution" id="ABBRID0EGZAE" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
2425
</specimenCode>
, 175.0 mm SL,
<location id="7D312D0E4DF919FB0D4A0AA2997F2EAF" LSID="urn:lsid:plazi:treatment:2690C8BE34C95E9189515F70F238DFDD:7D312D0E4DF919FB0D4A0AA2997F2EAF" country="Indonesia" county="Bitung" name="off Lembeh Island">off Lembeh Island</location>
,
<collectionCode id="5B3EFA37E48F3FF6293CD4230F53F26F" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. Peristiwady,
<date id="7B75C2A0868FAA9C47358F02B599F1F1" value="2011-12-20">20 December 2011</date>
).
<collectingCounty id="CF9B42FD828081FAF25240DE9F36228F">Bitung</collectingCounty>
,
<collectingCountry id="C29EF1365C875675A329218FC73DDD1C" name="Indonesia">Indonesia</collectingCountry>
,
<collectingMethod id="370269B21EE63A2AEFA5804B3399FA9F">hook and line</collectingMethod>
, more than
<quantity id="87DC459FDEF12CDD38D8ABBA7560C697" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="13FF498FCA005C2C7D42A1B457402A6A" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
depth (purchased at Girian Fish Market, Bitung, northern Sulawesi,
<collectorName id="A3D71B6C319FB88658350456AA6F76C0">T. Peristiwady</collectorName>
,
<date id="7B75C2A0868FAA9C47358F02B599F1F1" value="2011-12-20">
<collectingDate id="F10E82EEE6D6A2529F11DF351C138B55" value="2011-12-20">20 December 2011</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELZAE" type="description">
@ -353,11 +333,7 @@ and
<collectingCountry id="3D30A8A58D05C603F9D25E55C1542077" name="Marshall Islands">Marshall Islands</collectingCountry>
, as well as Scott Reef (northwestern
<collectingCountry id="7CC3676A6B95A09229FE312D8FCDFA86" name="Australia">Australia</collectingCountry>
), in latitudes of 8
<geoCoordinate id="8FA309A431E8B1F7530C9CA1C32FA14F" degrees="27" direction="north" orientation="latitude" precision="55555" value="27.0">27 ° N</geoCoordinate>
and 16
<geoCoordinate id="705A310E2F27D19CD1C9A1102436BDBD" degrees="21" direction="south" orientation="latitude" precision="55555" value="-21.0">21 ° S</geoCoordinate>
(Fig.
), in latitudes of 827 ° N and 1621 ° S (Fig.
<figureCitation id="E72947273CD85B829FF51BC7C663CFA0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distributional records of Plectranthias yamakawai (squares), Aulacocephalus temminckii (dots), Iniistius geisha (triangles), and Randallichthys filamentosus (diamonds), based on previous studies. Lembeh Island, northern Sulawesi, Indonesia (the collection site of presently reported specimens) is marked by a star." figureDoi="10.3897/aiep.55.142574.figure1" httpUri="https://binary.pensoft.net/fig/1262310">1</figureCitation>
). Such a distribution has thus far supported
<bibRefCitation id="D5CA0F6D9D37682CF252D153B0E1229D" author="Randall JE" firstAuthor="Randall's" issue="3" journalOrPublisher="Pacific Science" pagination="197-209" refId="B42" refString="Randall JE (1982) Examples of antitropical and antiequatorial distribution of Indo west-Pacific fishes. Pacific Science 35 (3): 197209." title="Examples of antitropical and antiequatorial distribution of Indo - west-Pacific fishes." volume="35" year="1982">Randalls (1982)</bibRefCitation>
@ -365,9 +341,7 @@ categorization of
<taxonomicName id="0A5E0418660C50F4A600429FC0D0FC51" baseAuthorityName="Fourmanoir" baseAuthorityYear="1970" family="Lutjanidae" genus="Randallichthys" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="filamentosus">
<emphasis id="E22C2E131419DB90F2362B9F6B9F75A1" italics="true">R. filamentosus</emphasis>
</taxonomicName>
as an anti-equatorial species. However, the presently reported specimen, collected off Lembeh Island (latitude
<geoCoordinate id="E284B2D6F9A185BC05463EA83720B12B" degrees="1.4" direction="north" orientation="latitude" precision="5555" value="1.4">1.4 ° N</geoCoordinate>
), is the first record of
as an anti-equatorial species. However, the presently reported specimen, collected off Lembeh Island (latitude 1.4 ° N), is the first record of
<taxonomicName id="81396C9FCC6DB69657DE45998DC1E057" baseAuthorityName="Fourmanoir" baseAuthorityYear="1970" family="Lutjanidae" genus="Randallichthys" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="filamentosus">
<emphasis id="B164C4CE04C4BCBFB143B86AA010FBAF" italics="true">R. filamentosus</emphasis>
</taxonomicName>

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@ -0,0 +1,594 @@
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<mods:classification id="0C61347C6897C9A6F04F3E61DCA16A7D">journal article</mods:classification>
<mods:identifier id="0D47054FA65DBB2C85042E944D4BF212" type="DOI">10.3897/evolsyst.9.138575</mods:identifier>
<mods:identifier id="6CC1695792CF2A33902755C0661E9708" type="ZooBank">3B0D4E2B-B8A6-4E93-BF3A-428E4B487C40</mods:identifier>
</mods:mods>
<treatment id="565565B5F7F55D819E1CD1A9862E9E18" ID-arpha="565565B5-F7F5-5D81-9E1C-D1A9862E9E18" ID-zoobank="https://zoobank.org/092CC468-34F0-4B06-9E1E-0C93D8121C41" LSID="urn:lsid:plazi:treatment:565565B5F7F55D819E1CD1A9862E9E18" LSID-ZBK="urn:lsid:zoobank.org:act:092CC468-34F0-4B06-9E1E-0C93D8121C41" httpUri="http://treatment.plazi.org/id/565565B5F7F55D819E1CD1A9862E9E18" scope_family="Ceratopogonidae" scope_order="Diptera" scope_subGenus="Holoconops">
<subSubSection id="9052C64BF92CAAF5610A7BF7C0D11B57" type="nomenclature">
<paragraph id="DEEFA40E4D08ED2D586E2804DEAD7285">
<taxonomicName id="F8217739359055CC582338021DEC07BE" authorityName="Ghosh &amp; Pal &amp; Hazra" authorityYear="2025" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov." subGenus="Holoconops">
<emphasis id="665B4145D502ABFA7B99AEA752841A91" italics="true">Leptoconops</emphasis>
(
<emphasis id="B328A3915D1F95F00EBB6553F004BA68" italics="true">Holoconops</emphasis>
)
<emphasis id="CF8216C2A5FF3FC0B05CF546CE6C4C88" italics="true">beccus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="502A9082CA5AC186DA346108BE3619C7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUNAE" type="NCBI GenBank Accession no">
<paragraph id="D216DD3FA1ECAD72B788FED31F9599BC">
<heading id="3875664FEE6B915FBA433BDE226E6712" reason="title">NCBI GenBank Accession no.</heading>
</paragraph>
<paragraph id="D13192D53019EF2E3FAB96F67DD748CD">
<ext-link id="B5BF4769F88E8A70ECEC5AFD258919C5" ext-link-type="gen" xlink_href="PP358259" xlink_type="simple">PP 358259</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6NAE" type="type material">
<paragraph id="D291B62622099D0928189E3497F132BD">
<heading id="991C11E88140CA5F65F20C0487E0D022" reason="title">Type material.</heading>
</paragraph>
<paragraph id="4AE58C53FB3C0932F187AB1B05EA99C3">
<materialsCitation id="4C4EBD8B1C1114FB9EF4C56D78A8A0D1" collectingDate="2023-10-30" collectingMethod="Light trap" collectorName="L. Ghosh." country="India" latitude="21.621513" location="Digha" longLatPrecision="1" longitude="89.87496" municipality="Purba Medinipur" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="holotype">
<emphasis id="AD87C37C55C299609880A67E6525B29D" bold="true" italics="true">
<typeStatus id="C9782AA915C44C513152C02AC641E93B" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="E9F962BD688A4E73ED62B34398ABAB37" count="1" type="male"></specimenCount>
,
<collectingCountry id="6D37258F91BD4BC77845EA185557A784" name="India">India</collectingCountry>
<collectingRegion id="7BFEDEC2DF27C95365422EBDBD17E192" country="India" name="West Bengal">West Bengal</collectingRegion>
,
<collectingMunicipality id="EF8E3F8433D453CC231D4C8BB6658BA6">Purba Medinipur</collectingMunicipality>
,
<location id="FF3CAFBC8961236B297F0ADC86A6114B" LSID="urn:lsid:plazi:treatment:565565B5F7F55D819E1CD1A9862E9E18:FF3CAFBC8961236B297F0ADC86A6114B" country="India" latitude="21.621513" longLatPrecision="1" longitude="89.87496" municipality="Purba Medinipur" name="Digha" stateProvince="West Bengal">Digha</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[89.874962,21.621513]}">
<geoCoordinate id="DCF62A97F8B79543CBA5D94E8ED9D512" degrees="21.621513" direction="north" orientation="latitude" precision="1" value="21.621513">21.621513 ° N</geoCoordinate>
,
<geoCoordinate id="0398BBE0723B17ED468A2564C332CF1C" degrees="89.874962" direction="east" orientation="longitude" precision="1" value="89.87496">89.874962 ° E</geoCoordinate>
</named-content>
),
<date id="735642E9225E274E83592AAF43E575FD" value="2023-10-30">
<collectingDate id="4C0D412A76EB407E111445F72E922F5A" value="2023-10-30">30. X. 2023</collectingDate>
</date>
,
<collectingMethod id="FBB31C38B43935BD69B141D708906EF6">Light trap</collectingMethod>
, Col.
<collectorName id="589ABCE96F360DAE390A9D56E349422D">L. Ghosh.</collectorName>
</materialsCitation>
<materialsCitation id="2567A34826F42A69EE75C826253CA10C" collectingDate="2023-10-30" collectingMethod="Light trap" collectorName="L. Ghosh." country="India" latitude="21.621513" location="Digha" longLatPrecision="1" longitude="89.87496" municipality="Purba Medinipur" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="West Bengal" typeStatus="paratype">
<emphasis id="EC1D22FF66AB6ED66737523D31DCFB1C" bold="true" italics="true">
<typeStatus id="524AFC9AC68E15475D8B6D45BADF23DF" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="8C413E3B6A4C36819B2C340A4A3D879C" count="5" type="male">5 ♂♂</specimenCount>
and
<specimenCount id="C9319026A11A269EBC329B439ADF4C6F" count="3" type="female">3 ♀♀</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVOAE" type="diagnosis">
<paragraph id="FEC0D88A89906A1E14DA9231C44DF79A">
<heading id="AA9C6916D8A66761C6597830D8594126" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2D4740C8F6BB0A865A6699F92C8C6417">
The only species of the subgenus
<taxonomicName id="915C7FE899EB92A08111F1F12360625E" rank="subGenus" subGenus="Holoconops">
<emphasis id="D2FEFDE729B51DF13A8CCBB350A86DF7" italics="true">Holoconops</emphasis>
</taxonomicName>
from the Oriental realm that is diagnosed by the following combination of characters:
<emphasis id="EFCCAF5A23C83314E3C0C3475AB274F3" bold="true">Adult male</emphasis>
(Fig.
<figureCitation id="9BB9F9A568FB116453C05C4C6094A018" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 A I</figureCitation>
). Gonocoxite wider at base to sub-basal part and narrow at anterior part; gonostylus short and stout, curved at middle and apex with two spines; parameres jointed at middle; posterior portion roughly rectangular, apex of paramere bearing an outwardly directed small projection; aedeagus elongate, rod-shaped, little swollen at apex.
<emphasis id="5933CDBD7B31F8C92C00627FA194170E" bold="true">Adult female</emphasis>
(Fig.
<figureCitation id="B10B0C2B74B6C603E7368AE990EADA48" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 J R</figureCitation>
). Eyes bare; mandible with 1213 teeth, palpomere III with 45 well developed protruded capitate sensilla; two unequal spermathecae without any hyaline punctation.
</paragraph>
<caption id="7FF4C4A6A3D59A65BF7F9E0D1F3A6952" ID-arpha="03961768-F849-5C4D-90C4-BD3253299F7D" httpUri="https://binary.pensoft.net/fig/1262632" startId="F1">
<paragraph id="1DF90438FB21F209F58E70C28DE9B8B9">
<label id="43DC6FE0279484BE0CBA1DB7D812597C">Figure 1.</label>
</paragraph>
<paragraph id="A296ED4879EBB47D8C44E83266AE1562">
<taxonomicName id="84687DEF7829354F01CE859500C282AB" authorityName="Ghosh &amp; Pal &amp; Hazra" authorityYear="2025" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov." subGenus="Holoconops">
<emphasis id="7C1328D16A7BAAEC0EA8957095508D67" italics="true">Leptoconops</emphasis>
(
<emphasis id="B2AEDA6B22728D1210CD62D1B69021C2" italics="true">Holoconops</emphasis>
)
<emphasis id="E9D2DBDD9177734CD7FD17CA0605794E" italics="true">beccus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08F96A01859805F9351C756339535EF4" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EFCA634A59CA0867D5739BE7FE4CF55" bold="true">A I.</emphasis>
Male (♂):
<emphasis id="39FA280F8F03578FAD44EC8D02B51A3C" bold="true">A.</emphasis>
Antenna;
<emphasis id="5A8BBF011252D9893EF176E4EDE9ED80" bold="true">B.</emphasis>
Maxillary palpus;
<emphasis id="4E65996B6D0AC45D1B315EC8CAD5B0E7" bold="true">C.</emphasis>
Wing;
<emphasis id="1F4A6892588AFAFE82E4DD281A74C6F3" bold="true">D.</emphasis>
Hind, mid, fore legs (left to right);
<emphasis id="1AD8110535B6394D0695B21C00CD0687" bold="true">E.</emphasis>
Hind tibial comb;
<emphasis id="06015BF2D12A550C0BA76DD3C281B109" bold="true">F.</emphasis>
Photograph of male genitalia (dorsal view);
<emphasis id="F7044EFAD48FDA719467CDAC4B5916E1" bold="true">G.</emphasis>
Illustration of male genitalia;
<emphasis id="7D2D6DC57B6E03B79115FDB2E8C92816" bold="true">H.</emphasis>
Aedeagus;
<emphasis id="E8D88AA23204E4AEA83A675D6CD3039E" bold="true">I.</emphasis>
Parameres;
<emphasis id="C6EE96F590F2104A17747139EDECB1D5" bold="true">J R.</emphasis>
Female (♀):
<emphasis id="89696D6E984A60C409EED904CFE99216" bold="true">J.</emphasis>
Habitus (Blood engorged);
<emphasis id="B79507296FC181B16B72CBE42DE23F6E" bold="true">K.</emphasis>
Antenna;
<emphasis id="3A15DF62A4A0614BB65260C243EA7399" bold="true">L.</emphasis>
Mandibular teeth;
<emphasis id="057D3EE90A02769E9C724E4032A1C75E" bold="true">M.</emphasis>
Clypeus;
<emphasis id="16DF312D7A8A731169AA6AA29FA6FCA5" bold="true">N.</emphasis>
Maxillary palpus;
<emphasis id="4CC592E3CE8751E5D61467DA20670559" bold="true">O.</emphasis>
Wing;
<emphasis id="DB48D3F84E7D5C4112D857464B20309B" bold="true">P.</emphasis>
Hind tibial comb;
<emphasis id="A6BE610DCA4FB82CCFAEC53AC53C59BC" bold="true">Q.</emphasis>
Genital lamella;
<emphasis id="911B29155C197C0734E84BA9E3B3BE78" bold="true">R.</emphasis>
Spermathecae. Scale bars: 0.05 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ETRAE" type="description">
<paragraph id="74E4024208BC8ED3C3F720F3D5CAF91A">
<heading id="A765F39419E78FEB398A2E0E4D24D910" reason="title">Description.</heading>
</paragraph>
<paragraph id="E7C3727774AB0A6445A7DCCA17616A0E">
<emphasis id="F53DA8B7EA011A8E85B4C0DAA167796B" bold="true">Male</emphasis>
(n = 6).
<emphasis id="2FC83EF9463BA823D51D707CC22EDD4E" bold="true">
<emphasis id="35B2235E3F4D730DA6C0A64FCB63F3A7" bold="true" italics="true">Head</emphasis>
.
</emphasis>
Head dark brown, eyes bare and separated about as wide as 10 facets. All flagellomeres (I XII) small and globular except terminal flagellomere (XIII); proportion of length of flagellomeres (I XIII): 1820 (19.2): 0910 (9.66): 0811 (9.6): 0811 (9.83): 0811 (9.66): 0810 (9.83): 0710 (9.5): 0811 (9.83): 0812 (10): 1015 (11.83): 1015 (11.83): 1120 (14.66): 3645 (39.5);
<abbrev id="ABBRID0E6RAE" xlink_title="Antennal ratio">AR</abbrev>
: 0.931.05 (0.98) (Fig.
<figureCitation id="E3B527874AB137C675723A90EA4503B2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 A</figureCitation>
). Maxillary palp with four palpomeres (terminal two fused); palpomere III (Fig.
<figureCitation id="90BCDDBEFEFB5B0CA3E733C32BD0BD33" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 B</figureCitation>
) longest with a round, shallow sensory pit at apical end, with 45 protruding capitate sensilla; proportion of length of palpomeres (I IV): 1012 (11.3): 0710 (8.6): 1722 (20.5): 1620 (18);
<abbrev id="ABBRID0ELSAE" xlink_title="Palpal ratio">PR</abbrev>
3.003.33 (3.1);
<abbrev id="ABBRID0EPSAE" xlink_title="Proboscis to head ratio">P / H</abbrev>
0.620.64 (0.63).
</paragraph>
<paragraph id="23A4D98F4530C72D4B81CADE9767A6EF">
<emphasis id="CAD8250D5D2B74AF4A54346910D06032" bold="true">
<emphasis id="F1CBB56489C9740039119C0A2C40A828" bold="true" italics="true">Thorax</emphasis>
.
</emphasis>
Dark brown. Scutellum with two bristles.
</paragraph>
<paragraph id="C5209689D9ECD2E1F0D835A496AB747E">
<emphasis id="DAD24FB92A0A62283B4FBA4120215BAB" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="3CAE1B950171320071320C7D64220A87" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 C</figureCitation>
). Covered with numerous microtrichia.
<abbrev id="ABBRID0EFTAE" xlink_title="Wing length">WL</abbrev>
0.750.90 (0.82),
<abbrev id="ABBRID0EJTAE" xlink_title="Wing width">WW</abbrev>
0.300.35 (0.32),
<abbrev id="ABBRID0ENTAE" xlink_title="Costal ratio">CR</abbrev>
0.400.47 (0.45); stigma pale brown.
</paragraph>
<paragraph id="7E66949F2041E221142782F80AF4BE75">
<emphasis id="CAA643525973540BD684B92126BCA3E5" bold="true">
<emphasis id="57EFB71FCB650F43CF1D5E1F4E05D46E" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
Uniformly brown to dark brown, without any banding pattern (Fig.
<figureCitation id="7B7497ABC8F237F2C26F7DB24A26B4F9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 D</figureCitation>
).
<abbrev id="ABBRID0E3TAE" xlink_title="Hind tibial comb">HTC</abbrev>
(Fig.
<figureCitation id="DA9E95D71344128F3F1B202EE188B629" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 E</figureCitation>
) with four spines and a spur; first one smallest, near spur and second one longest; first to fourth ones starting from spur measuring 2025 (22.5), 42.545 (44.2), 3037 (34), 2532.5 (29.2) respectively. TRI 1.501.62 (1.56), TRII 1.381.51 (1.47), TRIII 1.251.40 (1.34).
</paragraph>
<paragraph id="4A8DA54679C77C87D01D97CE2A6FB3D1">
<emphasis id="6BF30183E8C16C611B2D7B8AA33B2278" bold="true">
<emphasis id="C84A911A61ED36371C8E93469A48C726" bold="true" italics="true">Abdomen</emphasis>
.
</emphasis>
Dark Brown without any scale. Sternite IX 7082 (75) long and 107112 (109.6) wide, 1.4 times wider than length. Tergite IX 130150 (140.5) long, 110122 (117) wide, almost 1.2 times longer than wide and posterior margin triangular extending up to apex of Gonocoxite; basal margin of tergite IX bearing a outwardly directed, crescent-shaped greatly sclerotised structure with a sharply pointed inwardly directed spine at middle.
</paragraph>
<paragraph id="91FB8CF0F5B7D58D35FBF5A966E78C38">
<emphasis id="B02A25093975EF59CA91A21E1546C7B8" bold="true" italics="true">Genitalia</emphasis>
(Figs
<figureCitation id="329BD97EFACECA33D279C2163CD05D8E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 F I</figureCitation>
). Apicolateral process small with 810 setae. Gonocoxite dark brown (Fig.
<figureCitation id="C256BA4DE2AB6C491FB3D2EFAF993D4E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 F</figureCitation>
); wider at base and narrower in apex, 112127 (119) long, almost twice its greatest breadth, 6267 (63) with stout setae on inner part of gonocoxite. Gonostylus brown (Fig.
<figureCitation id="E457C42F1783786DEB6EB49513F45E88" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 G</figureCitation>
), 4052 (46.5) long and 1520 (18.33) wide, usually 0.4 times gonocoxal length, apex of gonostylus with two well-sclerotised equal-sized spines. Parameres (Fig.
<figureCitation id="AC3AF214BF34EC78428CDE372F78AF4F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 I</figureCitation>
) jointed at middle, basal part of each paramere roughly trapezoid in shape, apex of paramere with outwardly directed small projection, distal part each paramere boot-shaped. Aedeagus elongated, rod-shaped, little blunt at apex (Fig.
<figureCitation id="E06E724A1DE1DD8AA7093CA5D8B467BD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 H</figureCitation>
).
</paragraph>
<paragraph id="260C67A337EAB69AEC5E5977F06826AD">
<emphasis id="37183ACD80D33223372196B56A19F817" bold="true">Female</emphasis>
(n = 3). Overall, they have similar body colouration, and distribution of setae on body (Fig.
<figureCitation id="BFD5AFCA58CADAEFB595DAC2E6E42A6D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 J</figureCitation>
); similar shape of maxillary palp with males, hind tibial comb but some usual sexual and other differences are stated below.
</paragraph>
<paragraph id="22B919D59F1A40A420B485AF73BCB479">
<emphasis id="5B4BA6CF3293FCE2BB6940B55DD8F6BF" bold="true">
<emphasis id="979CECF3ECC61B81B1AA549C64764D64" bold="true" italics="true">Head</emphasis>
.
</emphasis>
All antennal flagellomeres rounded except terminal one (elongated) (Fig.
<figureCitation id="1B9A42FC3374FA999D5577EF3A933EF1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 K</figureCitation>
), pale brown to brown, bearing sensilla chaetica (I X); terminal flagellomere (XI) with sensilla trichodea; proportion of length of flagellomeres (I XI): 710 (8.33): 78 (7.33): 58 (6.33): 78 (7.33): 78 (7.33): 810 (8.66): 78 (7.33): 67 (6.66): 710 (8): 710 (8): 1925 (21.33);
<abbrev id="ABBRID0EZVAE" xlink_title="Antennal ratio">AR</abbrev>
0.620.67 (0.64). Mandible armed, with 1213 mandibular teeth (Fig.
<figureCitation id="D38934157E12F662948AAEFF807A938D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 L</figureCitation>
). Labrum with five serrations. Clypeus with four setae arranged in a row (Fig.
<figureCitation id="11E025DDC799A92A5E9DA83BF91397C7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 M</figureCitation>
). Proportion of length of palpomeres (Fig.
<figureCitation id="D297BDDD2C9C4099FD75773E100B801F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 N</figureCitation>
) (I III, IV + V): 810 (9): 810 (9): 1625 (21): 1019 (14.66);
<abbrev id="ABBRID0EJWAE" xlink_title="Palpal ratio">PR</abbrev>
1.72.1 (1.9) and
<abbrev id="ABBRID0ENWAE" xlink_title="Proboscis to head ratio">P / H</abbrev>
0.800.88 (0.83).
</paragraph>
<paragraph id="BB730BFB6AB4BE808E47B9354221BC4A">
<emphasis id="808D9AF54B2660B8EEBBFACDA310E4C2" bold="true">
<emphasis id="C3BB638FD4CDCF7CB921AB46B3610CDC" bold="true" italics="true">Thorax</emphasis>
.
</emphasis>
Scutum and scutellum dark brown; scutellum with four bristles.
</paragraph>
<paragraph id="74036D4F46DA298D7506A4D320F09B0F">
<emphasis id="4699E2C7D80962D36D160CA35926AE0D" bold="true" italics="true">Wings</emphasis>
(Fig.
<figureCitation id="5F7FA1FE84ECC66B5F24002BFA1A98D1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 O</figureCitation>
).
<abbrev id="ABBRID0EDXAE" xlink_title="Wing length">WL</abbrev>
0.770.90 (0.84),
<abbrev id="ABBRID0EHXAE" xlink_title="Wing width">WW</abbrev>
0.360.42 (0.38) and
<abbrev id="ABBRID0ELXAE" xlink_title="Costal ratio">CR</abbrev>
0.440.49 (0.46).
</paragraph>
<paragraph id="CCBF7C98AD06086F443E98C404FA0476">
<emphasis id="3386A728A5425FDF6F827A4C76F31141" bold="true">
<emphasis id="E99FF5EBC388ED94D92225CE430508F4" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
TRI 1.711.82 (1.77), TRII 1.331.51 (1.41), TRIII 1.401.48 (1.44);
<abbrev id="ABBRID0EWXAE" xlink_title="Hind tibial comb">HTC</abbrev>
with four spines and a spur (Fig.
<figureCitation id="42034AE45F9B5785F92438FF2EB5DA19" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 P</figureCitation>
); first to fourth spines from spur measuring 2022.5 (21), 37.542.5 (39), 3538 (36.5), 2426 (25), respectively.
</paragraph>
<paragraph id="4535D1F3174867A83E1A68FAF5D94C11">
<emphasis id="8427AB46CEFA9B77F916D317566823D0" bold="true" italics="true">Abdomen</emphasis>
(Figs
<figureCitation id="15A887B2BD16F754A81BA31FB9A485DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 Q R</figureCitation>
). Dark brown without any scales, with setae sparsely distributed. Genital lamellae pale brown (Fig.
<figureCitation id="F563DFB854B44C2F60A91D0DBA4BCCFC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 Q</figureCitation>
); little shorter, flat at base, and gradually narrow in apical part, 1.5 times longer than wide and roughly 2.5 times longer than sternite IX; sternite IX 5563 (60) long and 162175 (168) wide, well excavated. Spermathecae (Fig.
<figureCitation id="CFD320FD245CAD45FFE2BBB70FA90B89" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leptoconops (Holoconops) beccus sp. nov. A I. Male (♂): A. Antenna; B. Maxillary palpus; C. Wing; D. Hind, mid, fore legs (left to right); E. Hind tibial comb; F. Photograph of male genitalia (dorsal view); G. Illustration of male genitalia; H. Aedeagus; I. Parameres; J R. Female (♀): J. Habitus (Blood engorged); K. Antenna; L. Mandibular teeth; M. Clypeus; N. Maxillary palpus; O. Wing; P. Hind tibial comb; Q. Genital lamella; R. Spermathecae. Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/1262632">1 R</figureCitation>
) two, oval, almost equal, well sclerotised, without any short neck and hyaline punctation; measuring about 4047 (44.3) × 2530 (27.3) and 3644 (40.3) × 2532 (28.3).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERYAE" type="remarks">
<paragraph id="7088B04DEB8BD94306CF124D05739543">
<heading id="BD94F2F032ADC7706A9DBD0F2AED4DB7" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="30528CB9289236C50C5A043C9B75BE65">
The Indian species
<taxonomicName id="B534C51FF5D52197B6A70DE9E10B7BF2" authority="Mazumdar, Saha &amp; Chaudhuri, 2010" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="brevistylus">
<emphasis id="CDA8ACA7F20C0BA9CB78DC52E38B4FE5" italics="true">Leptoconops brevistylus</emphasis>
<bibRefCitation id="9A5E04067B407635DAF64DEFCA31B488" DOI="10.11646/zootaxa.2619.1.5" author="Mazumdar A &amp; Saha NC &amp; Chaudhuri PK" journalOrPublisher="Zootaxa" pagination="49-55" refId="B22" refString="Mazumdar A, Saha NC, Chaudhuri PK (2010) Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India. Zootaxa 2619: 49-55. https://doi.org/10.11646/zootaxa.2619.1.5" title="Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India." volume="2619" year="2010">Mazumdar, Saha &amp; Chaudhuri, 2010</bibRefCitation>
</taxonomicName>
differs from
<taxonomicName id="2D7C25134CC7B44A4B8FA08C76A17CF7" authorityName="Ghosh &amp; Pal &amp; Hazra" authorityYear="2025" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov.">
<emphasis id="9A74BBE472AAC719D8C578122CC1BBDE" italics="true">L. beccus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7C3201DDFE70E754224673D08C7DA768" rank="species">sp. nov.</taxonomicNameLabel>
in separate parameres, nearly U-shaped aedeagus with basal ends tapered in male, and double rows of clypeal setae,
<abbrev id="ABBRID0ENZAE" xlink_title="Hind tibial comb">HTC</abbrev>
with three spines, and two unequal, oval to sub-oval spermathecae in female. Another Indian species
<taxonomicName id="8BF065E3AA5A195F17F261033A0194AE" authority="Mazumdar, Saha &amp; Chaudhuri, 2010" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="fortipalpus">
<emphasis id="FDD62511DEBA974F36796E26575FCE96" italics="true">L. fortipalpus</emphasis>
<bibRefCitation id="D4C63F4199BF868BF0C5F7BEAAB930B2" DOI="10.11646/zootaxa.2619.1.5" author="Mazumdar A &amp; Saha NC &amp; Chaudhuri PK" journalOrPublisher="Zootaxa" pagination="49-55" refId="B22" refString="Mazumdar A, Saha NC, Chaudhuri PK (2010) Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India. Zootaxa 2619: 49-55. https://doi.org/10.11646/zootaxa.2619.1.5" title="Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India." volume="2619" year="2010">Mazumdar, Saha &amp; Chaudhuri, 2010</bibRefCitation>
</taxonomicName>
can be distinguished from the new species by double rows of clypeal setae, palpomeres I and II somewhat fused, lower
<abbrev id="ABBRID0E3ZAE" xlink_title="Palpal ratio">PR</abbrev>
(1.62) and unequal, oval spermathecae in female. Female members of
<taxonomicName id="32CD1E3CDD90008CB932DE4DF858D2C5" authority="Mazumdar, Saha &amp; Chaudhuri, 2010" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="tenebrostigmatus">
<emphasis id="C4A39E88FF509CABF6B78453B8186681" italics="true">L. tenebrostigmatus</emphasis>
<bibRefCitation id="AEB699BF146B5C3F4C538E6AB64AB1C8" DOI="10.11646/zootaxa.2619.1.5" author="Mazumdar A &amp; Saha NC &amp; Chaudhuri PK" journalOrPublisher="Zootaxa" pagination="49-55" refId="B22" refString="Mazumdar A, Saha NC, Chaudhuri PK (2010) Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India. Zootaxa 2619: 49-55. https://doi.org/10.11646/zootaxa.2619.1.5" title="Blood sucking midges of Leptoconops (Holoconops Kieffer) (Diptera: Ceratopogonidae) from India." volume="2619" year="2010">Mazumdar, Saha &amp; Chaudhuri, 2010</bibRefCitation>
</taxonomicName>
can be separated from the new species in higher antennal ratio (&gt; 1), eyes separated by six facets, double rows of clypeal setae, palpomeres I and II not well separated, and higher
<abbrev id="ABBRID0EL1AE" xlink_title="Palpal ratio">PR</abbrev>
(2.4).
</paragraph>
<paragraph id="9DCDBEB88C7179709F583B92E0830197">
The males of Nepalese species
<taxonomicName id="F4AEA5E55988DDEC1FF0C9F40FCFA4B7" authority="Chanthawanich &amp; Delfinado, 1967" authorityName="Chanthawanich &amp; Delfinado" authorityYear="1967" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="amplifemoralis">
<emphasis id="C3DE312CA6EB981F59F470B49F9F25D2" italics="true">L. amplifemoralis</emphasis>
<bibRefCitation id="7493B9A424BEA15C5906043C13D53B8B" DOI="10.1093/jmedent/4.3.294" author="Chanthawanich N &amp; Delfinado MD" journalOrPublisher="Journal of Medical Entomology" pagination="294-303" refId="B7" refString="Chanthawanich N, Delfinado MD (1967) Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae). Journal of Medical Entomology 4: 294-303. https://doi.org/10.1093/jmedent/4.3.294" title="Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae)." volume="4" year="1967">Chanthawanich &amp; Delfinado, 1967</bibRefCitation>
</taxonomicName>
differ in having the roughly rhomboid aedeagus, apex of parameres with a recurved spine and the absence of spines on the tergite band. The Chinese species
<taxonomicName id="EE652FD66E64D9B160A9E59DED5BCAA6" authority="Liu &amp; Yu, 1998" authorityName="Liu &amp; Yu" authorityYear="1998" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="menglaensis">
<emphasis id="FEA1F1EF2F20EADC05BA678BA6B9853D" italics="true">L. menglaensis</emphasis>
<bibRefCitation id="2967C3871F271A883D3B2929AD35AFD8" author="Liu ZJ &amp; Yu YX" journalOrPublisher="Journal of Medical Pest Control" pagination="24-26" refId="B21" refString="Liu ZJ, Yu YX (1998) Six new species of Leptoconops from China (Diptera: Ceratopogonidae). Journal of Medical Pest Control 14: 24-26. [in Chinese]" title="Six new species of Leptoconops from China (Diptera: Ceratopogonidae)." volume="14" year="1998">Liu &amp; Yu, 1998</bibRefCitation>
</taxonomicName>
can be segregated from the new species in separate parameres, elongated gonocoxite and club-shaped aedeagus. The Thai species
<taxonomicName id="D26660587D17F8863B6B094190E58B9E" authority="Chanthawanich &amp; Delfinado, 1967" authorityName="Chanthawanich &amp; Delfinado" authorityYear="1967" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="xuthosceles">
<emphasis id="7E67550711E5B82FA4853CFE6213D8DA" italics="true">L. xuthosceles</emphasis>
<bibRefCitation id="59E528107A5592E428096E4777C167B1" DOI="10.1093/jmedent/4.3.294" author="Chanthawanich N &amp; Delfinado MD" journalOrPublisher="Journal of Medical Entomology" pagination="294-303" refId="B7" refString="Chanthawanich N, Delfinado MD (1967) Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae). Journal of Medical Entomology 4: 294-303. https://doi.org/10.1093/jmedent/4.3.294" title="Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae)." volume="4" year="1967">Chanthawanich &amp; Delfinado, 1967</bibRefCitation>
</taxonomicName>
differs in swollen parameres in the middle and pointed recurved spine-like structure. The Chinese species
<taxonomicName id="1C6C0201AA88C857DB0F3878F1B8BEBA" authority="Yu, 1963" authorityName="Yu" authorityYear="1963" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="yunhsienensis">
<emphasis id="BE75E48827278535041083CD90FE710E" italics="true">L. yunhsienensis</emphasis>
<bibRefCitation id="25C0F215AB0DFCB861471845AFB09B1F" author="Yu YX" journalOrPublisher="Acta Zoologica Sinica" pagination="450-452" refId="B33" refString="Yu YX (1963) A new species of Leptoconops from China L. (Holoconops) yunhsienensis (Diptera: Ceratopogonidae) [in Chinese, English summary]. Acta Zoologica Sinica 15: 450-452." title="A new species of Leptoconops from China L. (Holoconops) yunhsienensis (Diptera: Ceratopogonidae) [in Chinese, English summary]." volume="15" year="1963">Yu, 1963</bibRefCitation>
</taxonomicName>
can be differentiated by separate parameres and hairy eyes. The females of
<taxonomicName id="1331AFDEC57AC53A3A92883F745F909C" authorityName="Chanthawanich &amp; Delfinado" authorityYear="1967" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="amplifemoralis">
<emphasis id="BCEDAF0D0E6736DA8F786DF7C394421C" italics="true">L. amplifemoralis</emphasis>
</taxonomicName>
differ in round, equal spermathecae, and much longer and wider wing. Females
<taxonomicName id="65C56F1D5B8955FB9864A634AF50AA45" authorityName="Yu" authorityYear="1963" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="yunhsienensis">
<emphasis id="CAC6C083DD5037CDA5E9F7A133369822" italics="true">L. yunhsienensis</emphasis>
</taxonomicName>
can be separated by hairy eyes and hyaline punctation in spermathecae. Females of
<taxonomicName id="7A99FCA01170831EC0A62E2F0338375C" authorityName="Liu &amp; Yu" authorityYear="1998" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="menglaensis">
<emphasis id="416C900596814734DAFB3A068C8E0E50" italics="true">L. menglaensis</emphasis>
</taxonomicName>
can be segregated from the new species by peach-shaped spermathecae.
</paragraph>
<paragraph id="B72F44FC4BD73ED3B00E8522A2C29381">
At present, only three species of the genus
<taxonomicName id="1DB9EFC6DE11AA9D10352F364726466B" authorityName="Skuse" authorityYear="1889" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="D347A10490BC42B2EB4720BBEA2F814C" italics="true">Leptoconops</emphasis>
</taxonomicName>
have publicly available COI sequences. Two of these species are categorized in the subgenus
<taxonomicName id="B2953B0D3FFE5DAD1254976859B5809D" rank="subGenus" subGenus="Leptoconops">
<emphasis id="ED26A6E36D2A60E3FCBE977FD35DA056" italics="true">Leptoconops</emphasis>
</taxonomicName>
, while the other falls under the subgenus
<taxonomicName id="E988096D3D8167E5DC48B5F870171762" authorityName="Kieffer" authorityYear="1921" class="Insecta" family="Ceratopogonidae" genus="Styloconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="A202067F03250E6D5AC5DFAB5192DF7F" italics="true">Styloconops</emphasis>
</taxonomicName>
. Our proposed new species falls in the subgenus
<taxonomicName id="ED794C5A79A66FC5E6E1623B6428AC25" rank="subGenus" subGenus="Holoconops">
<emphasis id="BF68CC0ED45BABD2E094649A39522216" italics="true">Holoconops</emphasis>
</taxonomicName>
. So at present, a molecular phylogenetic analysis may not yield fruitful results in the current scenario.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC5AE" type="etymology">
<paragraph id="A59F06AC967878849CCE7B17F97A3182">
<heading id="92099AAA892DF3EA7ED69B50EEFC26E8" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9A5F56E06B3C8FB7FC1932B119B6E448">
The specific name of the new species
<taxonomicName id="EDF485F0D0B563166C469D98AF01D984" authorityName="Ghosh &amp; Pal &amp; Hazra" authorityYear="2025" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="beccus">
<emphasis id="8CBB99E523DB37569995AA42B756F2D3" italics="true">beccus</emphasis>
</taxonomicName>
” derives from the beak-like projection of the apex of parameres.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES5AE" type="distribution">
<paragraph id="FF1B29177E969D63272EFDFAACE03C2E">
<heading id="777E4B826DB07A438891FC258F9CB72A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EC10D392F7D6FC2581118920CFDCC66B">
<collectingCountry id="C4075D0DD89BB22877F318D8A1A9F39A" name="India">India</collectingCountry>
(
<collectingRegion id="4564FD2B3C320C81E5F8199318302D45" country="India" name="West Bengal">West Bengal</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX5AE" type="Bionomics">
<paragraph id="8335F6D6B077BCBD3CBE88C613BB0DE3">
<heading id="40B1CCD7CD83D10CEF33D82BBA807858" reason="title">Bionomics.</heading>
</paragraph>
<paragraph id="4526BAB21A8269BA15423D8BC36ADD72">
<taxonomicName id="ABF5509573A273F941FFFC418D4E06E6" authorityName="Ghosh &amp; Pal &amp; Hazra" authorityYear="2025" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov." subGenus="Holoconops">
<emphasis id="B59BF74F62BC6713BA2000989D9078E2" italics="true">Leptoconops</emphasis>
(
<emphasis id="87FB85C1D304F582D57E0E68B69FEC32" italics="true">Holoconops</emphasis>
)
<emphasis id="A26276A744FC456109362315239D08E4" italics="true">beccus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4810D15F6C9C0F75B6572A1CF65FAAE" rank="species">sp. nov.</taxonomicNameLabel>
was procured from the Digha (
<quantity id="EB5743C0D5A7049DB2E3A2D8A7AAFBCB" metricMagnitude="0" metricUnit="m" metricValue="6.0" unit="m" value="6.0">6 m</quantity>
MSL), coastal plain of Purba Medinipur district, near the Bay of Bengal while the other three species [
<taxonomicName id="D7461778DDFE101C8CC0EA0B7F0667EF" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="brevistylus" subGenus="Holoconops">
<emphasis id="2A748E0E24B1281D4263453330C85B1A" italics="true">L.</emphasis>
(
<emphasis id="61055A2B4379CCEA19AE59EC8C6F8B72" italics="true">H.</emphasis>
)
<emphasis id="A212DA42BFECA9CBA90B71CB3A960080" italics="true">brevistylus</emphasis>
</taxonomicName>
;
<taxonomicName id="F64776611F4A0D7E86665BF0256E9D78" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="fortipalpus" subGenus="Holoconops">
<emphasis id="615C723CA8013B1AE168D2C33910E56D" italics="true">L.</emphasis>
(
<emphasis id="7D21F9A8C4B5FCCCFAD7A13BBFF819F3" italics="true">H.</emphasis>
)
<emphasis id="1E2205DD1170156E0B7F7AF41C95FCE6" italics="true">fortipalpus</emphasis>
</taxonomicName>
and
<taxonomicName id="6A3ABBB808FCCEE577C4B91652FBC4AE" authorityName="Mazumdar, Saha &amp; Chaudhuri" authorityYear="2010" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="tenebrostigmatus" subGenus="Holoconops">
<emphasis id="44368C340F5D9ED89A579D597EB4402F" italics="true">L.</emphasis>
(
<emphasis id="D87D4C2F97D35C3F4F9298ACBD788EF8" italics="true">H.</emphasis>
)
<emphasis id="398AD6E639C5D0FDF0678A22B61021F8" italics="true">tenebrostigmatus</emphasis>
</taxonomicName>
] were recorded mainly from the hilly area of the Chota Nagpur Plateau near the Mithon Water Reservoir (
<quantity id="6331161F75DB760C2E1031AA9D5A720D" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
MSL), Jharkhand, and Burdwan Town (
<quantity id="C0BB1E4975BD48464ABCBED6413DDD0A" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
MSL), a mature delta of the Deltaic Proper of Gangetic West Bengal. The Nepalese species
<taxonomicName id="02C9BA3E6967B95BD79A85838915285B" authority="Chanthawanich &amp; Delfinado, 1967" authorityName="Chanthawanich &amp; Delfinado" authorityYear="1967" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="amplifemoralis" subGenus="Holoconops">
<emphasis id="F9F30CD4334110838C678BA6E2D6BCA5" italics="true">L.</emphasis>
(
<emphasis id="DF04B1AEC7E29A7E8BB9BC3BB518BC1B" italics="true">H.</emphasis>
)
<emphasis id="77375132361E198532C3324A04999B3D" italics="true">amplifemoralis</emphasis>
<bibRefCitation id="1A9ADA01DCF12FDB097AAB4ED5A4A5B8" DOI="10.1093/jmedent/4.3.294" author="Chanthawanich N &amp; Delfinado MD" journalOrPublisher="Journal of Medical Entomology" pagination="294-303" refId="B7" refString="Chanthawanich N, Delfinado MD (1967) Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae). Journal of Medical Entomology 4: 294-303. https://doi.org/10.1093/jmedent/4.3.294" title="Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae)." volume="4" year="1967">Chanthawanich &amp; Delfinado, 1967</bibRefCitation>
</taxonomicName>
was reported from Kathmandu (
<quantity id="8DBA5638D6B64B4ADF35B86D606DDD81" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</quantity>
MSL), a part of Himalaya Biodiversity Hotspot. The Thai species
<taxonomicName id="D0AB5439E2108D8042ADCAFA2A58B2D7" authorityName="Chanthawanich &amp; Delfinado" authorityYear="1967" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="xuthosceles" subGenus="Holoconops">
<emphasis id="7230744FD41F07213578CDA798F62DFE" italics="true">L.</emphasis>
(
<emphasis id="74713B074A5DF902C09B9B3D4BD3820D" italics="true">H.</emphasis>
)
<emphasis id="8B39CFB080BD14330193883AD496C884" italics="true">xuthosceles</emphasis>
</taxonomicName>
was described from Chiang Mai (
<quantity id="16ACE432AB01EB3FAB8BCABE29F87B24" metricMagnitude="2" metricUnit="m" metricValue="3.1" unit="m" value="310.0">310 m</quantity>
MSL) near the Mae Ping River by
<bibRefCitation id="FA28CD4B234205DE453C12D3DA4EBDF5" DOI="10.1093/jmedent/4.3.294" author="Chanthawanich and Delfinado" firstAuthor="Chanthawanich" journalOrPublisher="Journal of Medical Entomology" pagination="294-303" refId="B7" refString="Chanthawanich N, Delfinado MD (1967) Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae). Journal of Medical Entomology 4: 294303. https://doi.org/10.1093/jmedent/4.3.294" title="Some species of Leptoconops of the Oriental and Pacific Regions (Diptera, Ceratopogonidae)." volume="4" year="1967">Chanthawanich and Delfinado 1967</bibRefCitation>
. The biting midge
<taxonomicName id="91EDA74B2E455A0B6E60D1717818265F" authority="Yu and Hui" authorityName="Yu and Hui" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="ascius" subGenus="Holoconops">
<emphasis id="E4E9EA70225FFBF2A1C839FDF26ED5E4" italics="true">L.</emphasis>
(
<emphasis id="7FB20DAF0B2E0AC0021C7AD20B0034BE" italics="true">H.</emphasis>
)
<emphasis id="52DC8853E47F948FE20E7B50E1E0DB78" italics="true">ascius</emphasis>
Yu and Hui
</taxonomicName>
, in
<bibRefCitation id="3C765D8CA42C139D0FA6DC5FC752CBF4" author="Yu" firstAuthor="Yu" refId="B34" refString="Yu YX (1988) Descriptions of new species of Ceratopogonidae (Diptera) in China [in Chinese, English summary]. Contributions to blood-sucking Diptera insects, China Public Health, Supplement 2: 127140." year="1988">Yu 1988</bibRefCitation>
was described near the Lancang River (
<quantity id="9A695FBFD270360DE36A3CE11386E9D5" metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">580 m</quantity>
MSL), Jinghong,
<collectingRegion id="C010D58AEC09A03DD743EBEE96F03FEB" country="China" name="Yunnan">Yunnan Province</collectingRegion>
of the Oriental
<collectingCountry id="7912A82AD46EC4BFF3A4BF9A09E5994C" name="China">China</collectingCountry>
. Both Oriental Chinese species,
<taxonomicName id="0D4E450EACB970B72826D0DAE976FD32" authority="Lee, 1978" authorityName="Lee" authorityYear="1978" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="menglaensis" subGenus="Holoconops">
<emphasis id="29B57D19B3722D01C6534A85B95B29A5" italics="true">L.</emphasis>
(
<emphasis id="C84EDEF5367F05148F55F77743C26AEF" italics="true">H.</emphasis>
)
<emphasis id="DE136B049F487A4C60DB2F13929F1577" italics="true">menglaensis</emphasis>
<bibRefCitation id="4229DD6A6CA668012BF6E9011BEECC8C" author="Lee TS" refId="B19" refString="Lee TS (1978) Diptera: Biting midges. Chinese Economic Entomology 13 (I - IV): 1 - 124. [in Chinese]." year="1978">Lee, 1978</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7D13833B7FA6555013743243B3FEEA5D" authority="Liu &amp; Yu, 1998" authorityName="Liu &amp; Yu" authorityYear="1998" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="yunnanensis" subGenus="Holoconops">
<emphasis id="F6A53ABEC365C8094BD1496E9BE741F4" italics="true">L.</emphasis>
(
<emphasis id="C8B9EACAA85CD808D1C5EF9AAC1492DA" italics="true">H.</emphasis>
)
<emphasis id="9FF0011C42395650FB19FC01F10C9964" italics="true">yunnanensis</emphasis>
<bibRefCitation id="C967DE8162257692ABB59B8B2B1EAD7E" author="Liu ZJ &amp; Yu YX" journalOrPublisher="Journal of Medical Pest Control" pagination="24-26" refId="B21" refString="Liu ZJ, Yu YX (1998) Six new species of Leptoconops from China (Diptera: Ceratopogonidae). Journal of Medical Pest Control 14: 24-26. [in Chinese]" title="Six new species of Leptoconops from China (Diptera: Ceratopogonidae)." volume="14" year="1998">Liu &amp; Yu, 1998</bibRefCitation>
</taxonomicName>
were reported from Mengla (
<quantity id="3FD3E791AAE17BE00C8732D126EA2A48" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
asl) of
<collectingRegion id="EBE2962C9FEF787A298C16D6505D2B17" country="China" name="Yunnan">Yunnan Province</collectingRegion>
. Another Chinese species
<taxonomicName id="CCAAEC7B9E2B03B7CEDF27BB72982A95" authorityName="Yu" authorityYear="1963" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="yunhsienensis" subGenus="Holoconops">
<emphasis id="3B1E13B8BE942C53E4D8DBDF045577FC" italics="true">L.</emphasis>
(
<emphasis id="BE81A12D356B5AD230BC64FE4525DA60" italics="true">H.</emphasis>
)
<emphasis id="DE0B2CCD634DF1469B63B2F79E342860" italics="true">yunhsienensis</emphasis>
</taxonomicName>
was described from Yunxian County (
<quantity id="C050D130C318A5A8E99B8D411DE796A9" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
MSL) of
<collectingRegion id="F91BC25C124384951BD327EF466DCDA1" country="China" name="Yunnan">Yunnan Province</collectingRegion>
by
<bibRefCitation id="47EDA645EFEE01A2D2C4A812F307181E" author="Yu YX" firstAuthor="Yu" journalOrPublisher="Acta Zoologica Sinica" pagination="450-452" refId="B33" refString="Yu YX (1963) A new species of Leptoconops from China L. (Holoconops) yunhsienensis (Diptera: Ceratopogonidae) [in Chinese, English summary]. Acta Zoologica Sinica 15: 450452." title="A new species of Leptoconops from China L. (Holoconops) yunhsienensis (Diptera: Ceratopogonidae) [in Chinese, English summary]." volume="15" year="1963">Yu (1963)</bibRefCitation>
. It was Clastier (1974) who described
<taxonomicName id="4AA66FF7020A6CF30E90FAC856BE5CA0" authorityName="Clastrier" authorityYear="1974" class="Insecta" family="Ceratopogonidae" genus="Leptoconops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="laosensis" subGenus="Holoconops">
<emphasis id="52F9238ECE2BD7FEABCA6BE315CE6079" italics="true">L.</emphasis>
(
<emphasis id="D387E7E8E9D9B174FE45109E2C87C680" italics="true">H.</emphasis>
)
<emphasis id="6D0B1ED169356624837A92284EBBC4C0" italics="true">laosensis</emphasis>
</taxonomicName>
from the Nam Khan located on the left bank of the River Mekong (
<quantity id="4E2A8B5F3024FF3B812CDABBA15BDB78" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</quantity>
MSL),
<collectingCountry id="1EB84FC74CC63D1B561FEF335A01D8AE" name="Laos">Laos</collectingCountry>
. The new species in the subgenus
<taxonomicName id="C4093031BE110D3949B54FBAE47F95D7" rank="subGenus" subGenus="Holoconops">
<emphasis id="A5FE7745017D60538DE67DB23DC0C9A7" italics="true">Holoconops</emphasis>
</taxonomicName>
is only reported from the coastal area of the Oriental region.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,80 +1,82 @@
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locality
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: 631
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: 35
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: 45
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(Amami,
@ -165,9 +155,7 @@ Islands,
<collectingCountry id="5CE806736AFA391BB12B9FCD92ACD893" name="Japan">Japan</collectingCountry>
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: 16
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(Tokara Islands,
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(purchased at Girian Fish Market, Bitung, northern
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(purchased at Girian Fish Market, Bitung, northern Sulawesi,
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,
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. Peristiwady,
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<date id="44A186B15FC7D6C835C49A7020B937FB" value="2012-04-02">
<collectingDate id="6C9CD4E2524B1D14DFE9AD8EE4545B52" value="2012-04-02">2 April 2012</collectingDate>
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@ -290,9 +287,7 @@ has been reported (based on several specimens) as having a limited Northern Hemi
<collectingCountry id="568DC35A44AE2DBF34C9BE37698412C2" name="Taiwan">Taiwan</collectingCountry>
, and the
<collectingCountry id="D9F2C8E0DF325EAE94A503F3AD6AEFC8" name="Northern Mariana Islands">Mariana Islands</collectingCountry>
, ranging from 16 to
<geoCoordinate id="A64E9480C211640497367446CD003368" degrees="30" direction="north" orientation="latitude" precision="55555" value="30.0">30 ° N</geoCoordinate>
(
, ranging from 16 to 30 ° N (
<bibRefCitation id="68CA6366E94FE8A7F962C19CDF82AAD2" author="Shimada" firstAuthor="Shimada" refId="B52" refString="Shimada K (2002) Lutjanidae. Labridae. Pp. 819832, 9691013, 15731579. In: Nakabo T (Ed.) Fishes of Japan with Pictorial Keys to the Species, English Edition. Tokai University Press, Tokyo, Japan." year="2002">Shimada 2002</bibRefCitation>
;
<bibRefCitation id="B461592F64C6552AEE993F51855150E2" author="Myers and Donaldson" firstAuthor="Myers" refId="B38" refString="Myers RF, Donaldson TJ (2003) The fishes of the Mariana Islands. Micronesica 3536: 598652." year="2003">Myers and Donaldson 2003</bibRefCitation>

View file

@ -0,0 +1,303 @@
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<taxonomicName id="E8ACC2F9CFB198303B22DFBF98E6E921" authority="Yoshino, 1972" authorityName="Yoshino" authorityYear="1972" family="Anthiadidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="yamakawai">
<emphasis id="0A0DAF75CE0161A8B668B026C8F98BD5" italics="true">Plectranthias yamakawai</emphasis>
<bibRefCitation id="F3994BF817B8C6C257C55C16FA241001" author="Yoshino T" refId="B58" refString="Yoshino T (1972) Plectranthias yamakawai, a new anthiadine fish from Ryukyu Islands, with a revision of the genus Plectranthias. Japanese Journal of Ichthyology 19 (2): 49 - 56." year="1972">Yoshino, 1972</bibRefCitation>
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<figureCitation id="4CA40172E99A821EDBB9400450036FA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of fresh specimens of (A) Plectranthias yamakawai, LBRC-F 3002, 182.5 mm SL and (B) Aulacocephalus temminckii, LBRC-F 402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia." figureDoi="10.3897/aiep.55.142574.figure2" httpUri="https://binary.pensoft.net/fig/1262311">Fig. 2 A</figureCitation>
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<emphasis id="EB35DB3B38FAB8B84497E8A7CE378744" italics="true">Plectranthias yamakawai</emphasis>
<comment id="5D268F2C52018C4C8F633234B7A17793">
<bibRefCitation id="49BE08ACF405BC76C0DD8956AABEF959" author="Yoshino T" refId="B58" refString="Yoshino T (1972) Plectranthias yamakawai, a new anthiadine fish from Ryukyu Islands, with a revision of the genus Plectranthias. Japanese Journal of Ichthyology 19 (2): 49 - 56." year="1972">Yoshino, 1972</bibRefCitation>
. —
<treatmentCitation id="93532FEA81F1436407562148A4E8BFE2" author="Yoshino T" page="53" year="1972">
<bibRefCitation id="6A937C75876D8A8E21F712BE01B97E17" author="Yoshino T" firstAuthor="Yoshino" refId="B58" refString="Yoshino T (1972) Plectranthias yamakawai, a new anthiadine fish from Ryukyu Islands, with a revision of the genus Plectranthias. Japanese Journal of Ichthyology 19 (2): 4956." year="1972">Yoshino (1972)</bibRefCitation>
: 53
</treatmentCitation>
, fig. 3 (type locality Ryukyu Islands, southern
<collectingCountry id="F6ADA501EA5B41896860B665389779AC" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="9832C2507A4B4DC6A7ADCF68DDCD3557" author="Randall JE" page="184" year="1980">
<bibRefCitation id="1C88548A0E661FF41BB93BE66BAFD8EC" author="Randall JE" firstAuthor="Randall" issue="1" journalOrPublisher="Micronesica" pagination="101-187" refId="B41" refString="Randall JE (1980) Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species. Micronesica 16 (1): 101187." title="Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species." volume="16" year="1980">Randall (1980)</bibRefCitation>
: 184
</treatmentCitation>
(southern
<collectingCountry id="EB505AA151B743A49B69942F426408DC" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="3A3117ACA3AED60C19DCC7473596DF05" author="Wass RC" page="12" year="1984">
<bibRefCitation id="8CF08466B340C5377EE1B5475F078DCB" author="Wass RC" firstAuthor="Wass" journalOrPublisher="National Oceanographic and Atmospheric Administration Technical Reports, National Marine Fisheries Service, Special Scientific Report, Fisheries" pagination="1-43" refId="B57" refString="Wass RC (1984) An annotated checklist of the fishes of Samoa. National Oceanographic and Atmospheric Administration Technical Reports, National Marine Fisheries Service, Special Scientific Report, Fisheries 781: 143." title="An annotated checklist of the fishes of Samoa." volume="781" year="1984">Wass (1984)</bibRefCitation>
: 12
</treatmentCitation>
(Samoan Islands). —
<treatmentCitation id="C216A6544ADD99369808357730AEF5E8" author="Chen JP &amp; Shao KT" page="64" year="2002">
<bibRefCitation id="E55AA61403F394D566D10368495A41B1" author="Chen JP &amp; Shao KT" firstAuthor="Chen" issue="1" journalOrPublisher="Zoological Studies" pagination="63-68" refId="B14" refString="Chen JP, Shao KT (2002) Plectranthias sheni, a new species and P. kamii, a new record of anthiine fishes (Perciformes: Serranidae) from Taiwan. Zoological Studies 41 (1): 6368." title="Plectranthias sheni, a new species and P. kamii, a new record of anthiine fishes (Perciformes: Serranidae) from Taiwan." volume="41" year="2002">Chen and Shao (2002)</bibRefCitation>
: 64
</treatmentCitation>
(Taiwan). —
<treatmentCitation id="0094D4591678D5A2FCEC35422BAAF35B" author="Motomura H &amp; Matsuura K (Eds.)" page="99" year="2010">
<bibRefCitation id="23207D28519D7BCE50DCDCF2D5878D64" author="Motomura H &amp; Matsuura K (Eds.)" firstAuthor="Motomura" journalOrPublisher="National Museum of Nature and Science, Tokyo, Japan" refId="B34" refString="Motomura H, Matsuura K (Eds.) (2010) Fishes of Yakushima Island — A World Heritage island in the Osumi Group, Kagoshima Prefecture, southern Japan. National Museum of Nature and Science, Tokyo, Japan, 264 pp." title="Fishes of Yakushima Island - A World Heritage island in the Osumi Group, Kagoshima Prefecture, southern Japan." year="2010">Motomura and Matsuura (2010)</bibRefCitation>
: 99
</treatmentCitation>
(southern
<collectingCountry id="3222CA684D7DFDB77F1572923F106383" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="D32EA186BCE02FE64D8379A9C760D060" author="Wada H &amp; Suzuki T &amp; Senou H &amp; Motomura H" page="305" year="2020">
<bibRefCitation id="12BBE661018C7276E35DB99BBF68D0AF" DOI="10.1007/s10228-019-00725-6" author="Wada H &amp; Suzuki T &amp; Senou H &amp; Motomura H" etAl="et al." firstAuthor="Wada" issue="2" journalOrPublisher="Ichthyological Research" pagination="294-307" refId="B56" refString="Wada H, Suzuki T, Senou H, Motomura H (2020) Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae). Ichthyological Research 67 (2): 294307. https://doi.org/10.1007/s10228-019-00725-6" title="Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae)." volume="67" year="2020">Wada et al. (2020)</bibRefCitation>
: 305
</treatmentCitation>
(southern
<collectingCountry id="71FC8E1FAF01CF08C91D5BC776012C7B" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="8BCB0BF290A563137BD4115A043D847F" author="Cabebe-Barnuevo RA &amp; Delloro Jr ESJ &amp; Penuela DFA &amp; Malay MCD &amp; Motomura H &amp; Babaran RP" page="300" year="2022">
<bibRefCitation id="BEAC5D7B0FD3BE5FEF9762F1959D3B68" DOI="10.3897/aiep.52.e96112" author="Cabebe-Barnuevo RA &amp; Delloro Jr ESJ &amp; Penuela DFA &amp; Malay MCD &amp; Motomura H &amp; Babaran RP" etAl="et al." firstAuthor="Cabebe-Barnuevo" issue="4" journalOrPublisher="Acta Ichthyologica et Piscatoria" pagination="299-307" refId="B12" refString="Cabebe-Barnuevo RA, Delloro Jr ESJ, Penuela DFA, Malay MCD, Motomura H, Babaran RP (2022) New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines. Acta Ichthyologica et Piscatoria 52 (4): 299307. https://doi.org/10.3897/aiep.52.e96112" title="New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines." volume="52" year="2022">Cabebe-Barnuevo et al. (2022)</bibRefCitation>
: 300
</treatmentCitation>
(Luzon,
<collectingCountry id="3BD8D0F12D7FD96F4E97174B1BBD763F" name="Philippines">Philippines</collectingCountry>
).
</comment>
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</paragraph>
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<paragraph id="6C782F3036255CFF9FAF5E7A2E2B423C">
<heading id="8785546EC7755A81A355DD487BE5FF77" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="75E67AB2BBBD6DE165CFA1339B491FA7">
<materialsCitation id="064076F7F3BDA92521B3D43A43A7C5F3" collectingDate="2012-12-22" collectingMethod="hook and line" collectionCode="LBRC-F" collectorName="T. Peristiwady" country="Indonesia" county="Bitung" elevation="125" location="off Lembeh Island" specimenCode="LBRC-F 3002" specimenCount="1">
<specimenCode id="0D3718F0F1E1A98CB1FB1765ACA78878">
<abbrev content-type="institution" id="ABBRID0EWLAC" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
<quantity id="1A597A9A0C283BDE6ABC4DD6857CA138" metricMagnitude="3" metricUnit="m" metricValue="3.0021825" unit="mm" value="3002182.5">3002, 182.5 mm</quantity>
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SL,
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,
<collectingCounty id="EC3AB7E3E0AC3B769D961CA8B5712072">Bitung</collectingCounty>
,
<collectingCountry id="5CFF8C9BF8F39A472B1B60C32A55DFF4" name="Indonesia">Indonesia</collectingCountry>
,
<collectingMethod id="BBB53E1CAA18A61B42C8F84E32BDC8F3">hook and line</collectingMethod>
,
<quantity id="A0C804DE7D934B445C9E183EE50EA245" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">
<elevation id="E16C2F8A9589E7967DCAEE4F923A2CDA" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100150 m</elevation>
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depth (purchased at Girian Fish Market, Bitung, northern Sulawesi by
<collectorName id="C52E81427EEACCCA4024C4F359A02DCE">T. Peristiwady</collectorName>
,
<date id="B31123DCF158BCEB7DFE7147FD155C96" value="2012-12-22">
<collectingDate id="FC38E735242CAD3276C5AB34C8B54968" value="2012-12-22">22 December 2012</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
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<paragraph id="820484A821EE1BE31AA64D4E53D007DD">
<heading id="D507CE0B1D13075EE73C0D46A9D23DB0" reason="title">Description.</heading>
</paragraph>
<paragraph id="DBF6F5FF4BF9F6DB79CC6A338AE746B9">
Dorsal fin with 10 spines and 16 soft rays; anal fin with 3 spines and 7 soft rays; pectoral fin with 13 rays, uppermost unbranched; pored lateral-line scales 32; scales above lateral line 5; scales below lateral line 16; gill rakers on upper limb of first gill arch 6, lower limb 12, total 18; body compressed, its depth
<quantity id="4793173B023D26E626B3C6FEC57A7C1A" metricMagnitude="-2" metricUnit="m" metricValue="5.842" unit="in" value="2.3">2.3 in</quantity>
SL; body width
<quantity id="147F3A5983B9B935E88DC7594FDD1EEF" metricMagnitude="-2" metricUnit="m" metricValue="5.588" unit="in" value="2.2">2.2 in</quantity>
body depth; head length
<quantity id="C8E8636689AEE376E57B30259A1ACC76" metricMagnitude="-2" metricUnit="m" metricValue="5.842" unit="in" value="2.3">2.3 in</quantity>
SL; snout length
<quantity id="36555CA533881167F3697FE0D4455D53" metricMagnitude="-2" metricUnit="m" metricValue="9.906" unit="in" value="3.9">3.9 in</quantity>
HL; orbit diameter
<quantity id="717AA9276D164C7F875FCAF5CB3CF588" metricMagnitude="-1" metricUnit="m" metricValue="1.1176" unit="in" value="4.4">4.4 in</quantity>
HL; upper jaw length
<quantity id="9552F717371979C69EA4FEF319EAC5B1" metricMagnitude="-2" metricUnit="m" metricValue="5.588" unit="in" value="2.2">2.2 in</quantity>
HL; interorbital width
<quantity id="8C43083ACD1D06936434BC2EDAA10711" metricMagnitude="-1" metricUnit="m" metricValue="1.8541999999999998" unit="in" value="7.3">7.3 in</quantity>
HL; caudal peduncle depth
<quantity id="2D77551600E34491B4184B2502642EBF" metricMagnitude="-2" metricUnit="m" metricValue="5.842" unit="in" value="2.3">2.3 in</quantity>
HL; caudal peduncle length
<quantity id="90A3F9222EB228AE9452554379A03F02" metricMagnitude="-2" metricUnit="m" metricValue="8.382" unit="in" value="3.3">3.3 in</quantity>
HL; dorsal profile of head forming angle of about 30 ° to horizontal axis of head and body, mouth terminal, oblique; vomer and palatine with teeth; posterior margin of maxilla just reaching vertical through midline of pupil; pre-opercle with two antrorse spines on ventral margin; opercle with three flat spines, uppermost blunt, middle largest; fourth dorsal-fin spine longest, fifth to last spines becoming progressively shorter; origin of first anal-fin spine about level with origin of third dorsal-fin soft ray, second anal-fin spine longest, second anal-fin soft ray longest; pectoral-fin ray long, its tip beyond base of first anal-fin spine; pelvic fin short, posterior tip not reaching anus when depressed; caudal fin weakly emarginate, some upper and lower rays somewhat filamentous.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAMAC" type="Color when fresh">
<paragraph id="DEC0B8E81E8798A4C1823FB140A0F250">
<heading id="37801AA9E025DDCB39EACD1E5E0AB4F2" reason="title">Color when fresh.</heading>
</paragraph>
<paragraph id="E61EA28C065C1BF51E6A85293C35BC56">
Body orange dorsally and rosy ventrally; dorsum of head and body, including dorsal and caudal fin bases, with numerous small distinctive, yellow-fringed dark greenish spots; large red spot just below lateral line at center of body; dorsal, anal and caudal fins yellowish; pectoral and pelvic fins reddish (Fig.
<figureCitation id="2FE68BEF5B0944B10E5E561E0D7C1DFF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of fresh specimens of (A) Plectranthias yamakawai, LBRC-F 3002, 182.5 mm SL and (B) Aulacocephalus temminckii, LBRC-F 402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia." figureDoi="10.3897/aiep.55.142574.figure2" httpUri="https://binary.pensoft.net/fig/1262311">2 A</figureCitation>
).
</paragraph>
<caption id="EE4DED334B93F533B5E7036629C36D96" ID-DOI="10.3897/aiep.55.142574.figure2" ID-arpha="98FF56E8-0D7A-5221-8E6D-4B1F2379F48D" httpUri="https://binary.pensoft.net/fig/1262311" startId="F2">
<paragraph id="6723C00B0CB4700D02DDE6453C438273">
<label id="9FC8817CFB025F78CFC82D1E91EAACB3">Figure 2.</label>
</paragraph>
<paragraph id="8C767655BC14FEA82C63B043A26D6A29">
Photographs of fresh specimens of (
<emphasis id="B49E9763653FFAB8AFD721C4275249A6" bold="true">A</emphasis>
)
<taxonomicName id="EB7306090FB0AC9F5B53816B822A6621" authorityName="Yoshino" authorityYear="1972" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="yamakawai">
<emphasis id="97921046DC1E255DCA11AC59BC39865A" italics="true">Plectranthias yamakawai</emphasis>
</taxonomicName>
,
<abbrev content-type="institution" id="ABBRID0E6MAC" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
3002, 182.5 mm SL and (
<emphasis id="9B149924FA6AD9F93C155115A5E9A12A" bold="true">B</emphasis>
)
<taxonomicName id="A410ABF75634A0CBF22A5503FC7FA8FB" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="045E9AFEF431382AF6537447244CF857" italics="true">Aulacocephalus temminckii</emphasis>
</taxonomicName>
,
<abbrev content-type="institution" id="ABBRID0ERNAC" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E2NAC" type="remarks">
<paragraph id="9A2F69703E0992FDC797EECEE12A6344">
<heading id="16203318F40798CF373659D395C9F05E" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="46E22FB759C70366E04766BA4B743AF8">
The Lembeh Island specimen agreed closely with the diagnostic characters of
<taxonomicName id="F10C0AA72D0343313DBD958A2B83C15A" authorityName="Yoshino" authorityYear="1972" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="yamakawai">
<emphasis id="64F0AE0D81782C04AA161BD27CD371F8" italics="true">P. yamakawai</emphasis>
</taxonomicName>
given by
<bibRefCitation id="4F3159A96CA5EB4D487922DD6AD3146D" author="Yoshino T" firstAuthor="Yoshino" refId="B58" refString="Yoshino T (1972) Plectranthias yamakawai, a new anthiadine fish from Ryukyu Islands, with a revision of the genus Plectranthias. Japanese Journal of Ichthyology 19 (2): 4956." year="1972">Yoshino (1972)</bibRefCitation>
,
<bibRefCitation id="7C32D63C1470D9D8411537513039407C" author="Randall" firstAuthor="Randall" issue="1" journalOrPublisher="Micronesica" pagination="101-187" refId="B41" refString="Randall JE (1980) Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species. Micronesica 16 (1): 101187." title="Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species." volume="16" year="1980">Randall (1980</bibRefCitation>
,
<bibRefCitation id="F0CE3071CA2760FDD26DC65EB43B0E19" author="Randall" firstAuthor="Randall" issue="2" journalOrPublisher="Micronesica" pagination="113-131" refId="B43" refString="Randall JE (1996) Two new anthiine fishes of the genus Plectranthias (Perciformes: Serranidae), with a key to the species. Micronesica 29 (2): 113131." title="Two new anthiine fishes of the genus Plectranthias (Perciformes: Serranidae), with a key to the species." volume="29" year="1996">1996</bibRefCitation>
),
<bibRefCitation id="629B83E602733E0F540254C9C6C65E57" DOI="10.1007/s10228-019-00725-6" author="Wada H &amp; Suzuki T &amp; Senou H &amp; Motomura H" etAl="et al." firstAuthor="Wada" issue="2" journalOrPublisher="Ichthyological Research" pagination="294-307" refId="B56" refString="Wada H, Suzuki T, Senou H, Motomura H (2020) Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae). Ichthyological Research 67 (2): 294307. https://doi.org/10.1007/s10228-019-00725-6" title="Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae)." volume="67" year="2020">Wada et al. (2020)</bibRefCitation>
, and
<bibRefCitation id="DAB1FDC689E24A2C0F4FEE9CFCC7D8A8" DOI="10.3897/aiep.52.e96112" author="Cabebe-Barnuevo RA &amp; Delloro Jr ESJ &amp; Penuela DFA &amp; Malay MCD &amp; Motomura H &amp; Babaran RP" etAl="et al." firstAuthor="Cabebe-Barnuevo" issue="4" journalOrPublisher="Acta Ichthyologica et Piscatoria" pagination="299-307" refId="B12" refString="Cabebe-Barnuevo RA, Delloro Jr ESJ, Penuela DFA, Malay MCD, Motomura H, Babaran RP (2022) New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines. Acta Ichthyologica et Piscatoria 52 (4): 299307. https://doi.org/10.3897/aiep.52.e96112" title="New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines." volume="52" year="2022">Cabebe-Barnuevo et al. (2022)</bibRefCitation>
, viz. dorsal-fin soft rays 16; pectoral fin rays 13; pored lateral-line scales 32; pre-opercle with two antrorse spines on ventral margin; fourth dorsal-fin spine longest; dorsum of head and body, including dorsal and caudal fin bases, with numerous small distinctive yellow-fringed dark greenish spots; large red spot below lateral line at center of body. In addition, the specimen also matched the figure of the
<typeStatus id="980EDDE7BA62293FE080A00343518DFE" type="holotype">holotype</typeStatus>
of the species (see
<bibRefCitation id="69B7B82019C1187C8050A3036680A998" author="Randall JE" firstAuthor="Randall" issue="1" journalOrPublisher="Micronesica" page="32" pagination="101-187" refId="B41" refString="Randall JE (1980) Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species. Micronesica 16 (1): 101187." title="Revision of the fish genus Plectranthias (Serranidae: Anthiinae) with descriptions of 13 new species." volume="16" year="1980">Randall 1980: 32</bibRefCitation>
) and the color photographs provided by
<bibRefCitation id="7D5328C42301BB13C6B9E59A04AB4A68" author="Motomura H &amp; Hagiwara K &amp; Senou H &amp; Nakae M" etAl="et al." firstAuthor="Motomura" journalOrPublisher="] Kagoshima University Museum, Kagoshima - Yokosuka City Museum, Yokosuka - Kanagawa Prefectural Museum of Natural History, Odawara - National Museum of Nature and Science, Tsukuba, Japan" page="97" refId="B36" refString="Motomura H, Hagiwara K, Senou H, Nakae M (2018) [Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan.] Kagoshima University Museum, Kagoshima Yokosuka City Museum, Yokosuka Kanagawa Prefectural Museum of Natural History, Odawara National Museum of Nature and Science, Tsukuba, Japan, 414 pp. [In Japanese]" title="[Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan." year="2018">Motomura et al. (2018: 97)</bibRefCitation>
and
<bibRefCitation id="41C3D30A192E8C0B9BFF766E1FA946D7" DOI="10.3897/aiep.52.e96112" author="Cabebe-Barnuevo" etAl="et al." firstAuthor="Cabebe-Barnuevo" issue="4" journalOrPublisher="Acta Ichthyologica et Piscatoria" pagination="299-307" refId="B12" refString="Cabebe-Barnuevo RA, Delloro Jr ESJ, Penuela DFA, Malay MCD, Motomura H, Babaran RP (2022) New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines. Acta Ichthyologica et Piscatoria 52 (4): 299307. https://doi.org/10.3897/aiep.52.e96112" title="New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines." volume="52" year="2022">Cabebe-Barnuevo et al. (2022</bibRefCitation>
: fig. 1 A).
</paragraph>
<paragraph id="6507F70E6BA0595DAA39D18B27D33D6D">
<taxonomicName id="5D92D5859B9B17FDC633D818DC4EC9B9" authorityName="Yoshino" authorityYear="1972" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="yamakawai">
<emphasis id="7C4C273E104BB812B835B4881994E8C8" italics="true">Plectranthias yamakawai</emphasis>
</taxonomicName>
has previously been reported only from southern
<collectingCountry id="3F3BBAD0DB77052E02DC240F1F32F517" name="Japan">Japan</collectingCountry>
,
<collectingCountry id="A12D5A468648131885FCE0BC80540CE9" name="Taiwan">Taiwan</collectingCountry>
, the
<collectingCountry id="4D56A20B6B04F15235AC30925268D4AE" name="Philippines">Philippines</collectingCountry>
and
<collectingCountry id="672702151ABD3AF043135279B0763608" name="Samoa">Samoa</collectingCountry>
, at latitudes of 1530 ° N and 13 ° S (
<bibRefCitation id="9FFFEDAF580DE3A69936055797649F5E" author="Wass" firstAuthor="Wass" journalOrPublisher="National Oceanographic and Atmospheric Administration Technical Reports, National Marine Fisheries Service, Special Scientific Report, Fisheries" pagination="1-43" refId="B57" refString="Wass RC (1984) An annotated checklist of the fishes of Samoa. National Oceanographic and Atmospheric Administration Technical Reports, National Marine Fisheries Service, Special Scientific Report, Fisheries 781: 143." title="An annotated checklist of the fishes of Samoa." volume="781" year="1984">Wass 1984</bibRefCitation>
;
<bibRefCitation id="96EB901903168A248882467E51DBC5E8" author="Motomura and Matsuura" firstAuthor="Motomura" journalOrPublisher="National Museum of Nature and Science, Tokyo, Japan" refId="B34" refString="Motomura H, Matsuura K (Eds.) (2010) Fishes of Yakushima Island — A World Heritage island in the Osumi Group, Kagoshima Prefecture, southern Japan. National Museum of Nature and Science, Tokyo, Japan, 264 pp." title="Fishes of Yakushima Island - A World Heritage island in the Osumi Group, Kagoshima Prefecture, southern Japan." year="2010">Motomura and Matsuura 2010</bibRefCitation>
;
<bibRefCitation id="52D3004FB8CB30D56EC4A76272AF06D0" DOI="10.1007/s10228-019-00725-6" author="Wada" etAl="et al." firstAuthor="Wada" issue="2" journalOrPublisher="Ichthyological Research" pagination="294-307" refId="B56" refString="Wada H, Suzuki T, Senou H, Motomura H (2020) Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae). Ichthyological Research 67 (2): 294307. https://doi.org/10.1007/s10228-019-00725-6" title="Plectranthias ryukyuensis, a new species of perchlet from the Ryukyu Islands, Japan, with a key to the Japanese species of Plectranthias (Serranidae: Anthiadinae)." volume="67" year="2020">Wada et al. 2020</bibRefCitation>
;
<bibRefCitation id="56C3B682955C1C9F3040E14501F0DEE7" DOI="10.3897/aiep.52.e96112" author="Cabebe-Barnuevo" etAl="et al." firstAuthor="Cabebe-Barnuevo" issue="4" journalOrPublisher="Acta Ichthyologica et Piscatoria" pagination="299-307" refId="B12" refString="Cabebe-Barnuevo RA, Delloro Jr ESJ, Penuela DFA, Malay MCD, Motomura H, Babaran RP (2022) New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines. Acta Ichthyologica et Piscatoria 52 (4): 299307. https://doi.org/10.3897/aiep.52.e96112" title="New record of anthiadine fish, Plectranthias yamakawai (Actinopterygii: Perciformes: Serranidae), from the Philippines." volume="52" year="2022">Cabebe-Barnuevo et al. 2022</bibRefCitation>
; fig. 3). The presently reported specimen, collected off Lembeh Island at a latitude of 1.4 ° N, represents the first record of
<taxonomicName id="40E9634B2AEE0B4FA2E5A1129334A33C" authorityName="Yoshino" authorityYear="1972" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="yamakawai">
<emphasis id="089CDE3BF09219A54237AB1A0937C79C" italics="true">P. yamakawai</emphasis>
</taxonomicName>
in equatorial and Indonesian waters.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D793206B87E998686B653E624412A526">Acta Ichthyologica et Piscatoria</mods:title>
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<mods:date id="5E538A416FF513D562DA871CBD744257">2025</mods:date>
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<mods:number id="A27D84326F0F06E433281EFCD6EC5C3F">2025-02-25</mods:number>
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<mods:number id="80F340F97B36252BC080C383D62F8CBD">55</mods:number>
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<mods:identifier id="0F55ABA23B5D89ED4FB750366DAFDB44" type="DOI">10.3897/aiep.55.142574</mods:identifier>
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<treatment id="DCDB7A653DAA522F8129DB5FC94D738E" ID-DOI="http://doi.org/10.5281/zenodo.14925584" ID-GBIF-Taxon="254608515" ID-Zenodo-Dep="14925584" ID-arpha="DCDB7A65-3DAA-522F-8129-DB5FC94D738E" LSID="urn:lsid:plazi:treatment:DCDB7A653DAA522F8129DB5FC94D738E" httpUri="http://treatment.plazi.org/id/DCDB7A653DAA522F8129DB5FC94D738E" scope_family="Liopropomatidae">
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<paragraph id="F03DE7CCD85F508AB07C6EF2131DCC5A">
<taxonomicName id="47568D871F60A61A99CFEE1B2B43212A" authority="Bleeker, 1855" authorityName="Bleeker" authorityYear="1855" family="Liopropomatidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="A0573AF5C4215AFB7F07EA5EF1A03B3A" italics="true">Aulacocephalus temminckii</emphasis>
<bibRefCitation id="0FA53A7B69D86A7FAA843B212D4F19B2" author="Bleeker P" issue="4" journalOrPublisher="Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen" pagination="1-132" refId="B10" refString="Bleeker P (1855) Nieuwe nalezingen op de ichthyologie van Japan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 26 (4): 1-132." title="Nieuwe nalezingen op de ichthyologie van Japan." volume="26" year="1855">Bleeker, 1855</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="FD43378091B40995ABEED4273E6361FF">
<figureCitation id="C816D8F426577A8A98281A878108063D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of fresh specimens of (A) Plectranthias yamakawai, LBRC-F 3002, 182.5 mm SL and (B) Aulacocephalus temminckii, LBRC-F 402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia." figureDoi="10.3897/aiep.55.142574.figure2" httpUri="https://binary.pensoft.net/fig/1262311">Fig. 2 B</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="7D392A35405CE110D5E1D926F54B0003">
<treatmentCitationGroup id="FFB3D836A1F43F1547E39822D4092D6F">
<treatmentCitation id="08DB14D25A3D78848D041F2308867EB3">
<taxonomicName id="568EF3FB1A6EA5F97D75D4E2B70A090B" ID-CoL="JQSB" authority="Bleeker, 1855" authorityName="Bleeker" authorityYear="1855" family="Liopropomatidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="0E55CA37A617CB4DE7979B8E9AC0DD2B" italics="true">Aulacocephalus temminckii</emphasis>
<comment id="6670C33E879BFC0D9CCC2B621B145047">
<bibRefCitation id="F4AB0B448321A3E305DBCAF19B107775" author="Bleeker P" issue="4" journalOrPublisher="Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen" pagination="1-132" refId="B10" refString="Bleeker P (1855) Nieuwe nalezingen op de ichthyologie van Japan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 26 (4): 1-132." title="Nieuwe nalezingen op de ichthyologie van Japan." volume="26" year="1855">Bleeker, 1855</bibRefCitation>
. —
<treatmentCitation id="F2866562F4725A454D3AA75AF212B6E2" author="Bleeker P" page="12" year="1855">
<bibRefCitation id="0063AE89FC8A4251280E6864D497E956" author="Bleeker P" firstAuthor="Bleeker" issue="4" journalOrPublisher="Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen" pagination="1-132" refId="B10" refString="Bleeker P (1855) Nieuwe nalezingen op de ichthyologie van Japan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 26 (4): 1132." title="Nieuwe nalezingen op de ichthyologie van Japan." volume="26" year="1855">Bleeker (1855)</bibRefCitation>
: 12
</treatmentCitation>
(type locality
<collectingRegion id="CAEA1F195047DD0314C190FD544B9646" country="Japan" name="Nagasaki">Nagasaki</collectingRegion>
,
<collectingCountry id="84B7B1CA2386B387EEBF62C8888B5ACD" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="F974737A6BDAAE8024CBFA4AA5E6CBAD" author="Randall JE &amp; Aida K &amp; Hibiya T &amp; Matsuura N &amp; Kamiya H &amp; Hashimoto Y" page="173" year="1971">
<bibRefCitation id="1D03F3AD336F60186080A0827C0BF68F" DOI="10.5134/175661" author="Randall JE &amp; Aida K &amp; Hibiya T &amp; Matsuura N &amp; Kamiya H &amp; Hashimoto Y" etAl="et al." firstAuthor="Randall" issue="2-3" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pagination="157-190" refId="B46" refString="Randall JE, Aida K, Hibiya T, Matsuura N, Kamiya H, Hashimoto Y (1971) Grammistin, the skin toxin of soapfishes and its significance in the classification of the Grammistidae. Publications of the Seto Marine Biological Laboratory 19 (23): 157190. https://doi.org/10.5134/175661" title="Grammistin, the skin toxin of soapfishes and its significance in the classification of the Grammistidae." volume="19" year="1971">Randall et al. (1971)</bibRefCitation>
: 173
</treatmentCitation>
[Natal coast (
<collectingCountry id="F1497124D5E5B07BA88C5A638BC0A1A8" name="South Africa">South Africa</collectingCountry>
),
<collectingCountry id="CB418AB2FA961E05B3D65C6FFAB720F3" name="Mauritius">Mauritius</collectingCountry>
,
<collectingCountry id="7F4B896D58011A540E80DCC828ED76E2" name="Thailand">Thailand</collectingCountry>
, northeast
<collectingCountry id="044CEF52AF7E0C7C697E357E614EB193" name="China">China</collectingCountry>
,
<collectingCountry id="D720253BCC6601467DDA81BF76EB9987" name="Taiwan">Taiwan</collectingCountry>
,
<collectingCountry id="B813FC8095C316BD0EE7EFD34431DEDE" name="South Korea">Korea</collectingCountry>
,
<collectingCountry id="3054BD24C9F80158F3127C105B3BA5F4" name="Japan">Japan</collectingCountry>
,
<collectingCountry id="891DE05E9AB623E9347526FF1037394D" name="New Zealand">New Zealand</collectingCountry>
, Kermadec Islands]. —
<treatmentCitation id="82057D3ED5373C927DE25E970E1F8D33" author="Randall JE" page="201" year="1982">
<bibRefCitation id="5DF3A7848F759D3D5A99A09E924A79AA" author="Randall JE" firstAuthor="Randall" issue="3" journalOrPublisher="Pacific Science" pagination="197-209" refId="B42" refString="Randall JE (1982) Examples of antitropical and antiequatorial distribution of Indo west-Pacific fishes. Pacific Science 35 (3): 197209." title="Examples of antitropical and antiequatorial distribution of Indo - west-Pacific fishes." volume="35" year="1982">Randall (1982)</bibRefCitation>
: 201
</treatmentCitation>
(Rapa,
<collectingCountry id="529456DEA947260C45F930FB2177B255" name="French Polynesia">French Polynesia</collectingCountry>
). —
<treatmentCitation id="0113D31C944BFD24FC977AD23FB04FC8" author="Katayama M" page="139" year="1984">
<bibRefCitation id="0634F3288F27FB226CF72BEF3C8BF194" author="Katayama M" firstAuthor="Katayama" refId="B28" refString="Katayama M (1984) Aulacocephalus temminckii. P. 139. In: Masuda H, Amaoka K, Araga C, Uyeno T, Yoshino T (Eds.) The fishes of the Japanese Archipelago. Text. Tokai University Press, Tokyo, Japan." year="1984">Katayama (1984)</bibRefCitation>
: 139
</treatmentCitation>
(southern
<collectingCountry id="14582E7638EA2A8026A2F9F1DE94977C" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="574C74A092E352B8771104B3DAC894FB" author="Francis MP" page="160" year="1993">
<bibRefCitation id="ED0823CC168A44BB4581F4D439261646" author="Francis MP" firstAuthor="Francis" issue="2" journalOrPublisher="Pacific Science" pagination="136-170" refId="B17" refString="Francis MP (1993) Checklist of the coastal fishes of Lord Howe, Norfolk, and Kermadec Island, southwest Pacific Ocean. Pacific Science 47 (2): 136170." title="Checklist of the coastal fishes of Lord Howe, Norfolk, and Kermadec Island, southwest Pacific Ocean." volume="47" year="1993">Francis (1993)</bibRefCitation>
: 160
</treatmentCitation>
(Eastern
<collectingCountry id="81680B993A899EAF73C56BD3AC69A8D4" name="Australia">Australia</collectingCountry>
,
<collectingCountry id="20A8638932D4297E5EFAA277B11141EA" name="Norfolk Island">Norfolk Island</collectingCountry>
,
<collectingCountry id="78D6CE8B484376BA6B1E146F599BB109" name="New Zealand">New Zealand</collectingCountry>
, Kermadec Islands). —
<treatmentCitation id="CFECAA1405A92236A000DE96421A0D86" author="Goren M &amp; Dor M" page="24" year="1994">
<bibRefCitation id="CD62226AE48B78F507BB4E7CE00924AB" author="Goren M &amp; Dor M" firstAuthor="Goren" journalOrPublisher="The Israel Academy of Sciences and Humanities, Jerusalem" refId="B24" refString="Goren M, Dor M (1994) An updated checklist of the fishes of the Red Sea: CLOFRES II. The Israel Academy of Sciences and Humanities, Jerusalem, 120 pp." title="An updated checklist of the fishes of the Red Sea: CLOFRES II." year="1994">Goren and Dor (1994)</bibRefCitation>
: 24
</treatmentCitation>
(Red Sea). —
<treatmentCitation id="13BB14FA57F7F0530E9FC004825CBC1F" author="Kuiter RH" page="122" year="1997">
<bibRefCitation id="652204DE1A35410B5D0600BE7025A44E" author="Kuiter RH" firstAuthor="Kuiter" journalOrPublisher="New Holland Publishers, Frenchs Forest, NSW, Australia" refId="B30" refString="Kuiter RH (1997) Guide to sea fishes of Australia. A comprehensive reference for divers and fishermen. New Holland Publishers, Frenchs Forest, NSW, Australia, 434 pp." title="Guide to sea fishes of Australia. A comprehensive reference for divers and fishermen." year="1997">Kuiter (1997)</bibRefCitation>
: 122
</treatmentCitation>
(
<collectingCountry id="02EFFE7DFC045E6C1E7B5638EA324104" name="Australia">Australia</collectingCountry>
). —
<treatmentCitation id="59BF25DA5EC78C9D07AE6332B7C37770" author="Fricke R" page="210" year="1999">
<bibRefCitation id="E3DC6EAC92B6F566688163523D287903" author="Fricke R" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Koenigstein, Germany" refId="B18" refString="Fricke R (1999) Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species. Koeltz Scientific Books, Koenigstein, Germany, 759 pp." title="Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species." year="1999">Fricke (1999)</bibRefCitation>
: 210
</treatmentCitation>
(Mascarene Islands). —
<treatmentCitation id="6AE089A9DC302C9379882981FC5EB809" author="Hutchins JB" page="30" year="2001">
<bibRefCitation id="59F94148F9C6FDB3B2A1F79D4A5EE48E" DOI="10.18195/issn.0313-122x.63.2001.009-050" author="Hutchins JB" firstAuthor="Hutchins" refId="B27" refString="Hutchins JB (2001) Checklist of the fishes of Western Australia. Records of the Western Australian Museum 63 (1, Suppl. 63): 950. https://doi.org/10.18195/issn.0313-122x.63.2001.009-050" year="2001">Hutchins (2001)</bibRefCitation>
: 30
</treatmentCitation>
(
<collectingRegion id="66BF65B96FC6BF1111DE7D646CA469E6" country="Australia" name="Western Australia">Western Australia</collectingRegion>
). —
<treatmentCitation id="033636AA267341482FAB0BAB9B94E9CA" author="Senou H" page="726" year="2002">
<bibRefCitation id="2506C1DFCAF7FBE1812715E347CD33B1" author="Senou H" firstAuthor="Senou" refId="B51" refString="Senou H (2002) Serranidae. Pp. 690731, 15321540. In: Nakabo T (Ed.) Fishes of Japan with Pictorial Keys to the Species, English Edition. Tokai University Press, Tokyo, Japan." year="2002">Senou (2002)</bibRefCitation>
: 726
</treatmentCitation>
(
<collectingCountry id="DC7F270B7B044436BF2E956B4C91757D" name="Japan">Japan</collectingCountry>
). —
<treatmentCitation id="88A131FAD73E308DF7FA6FB29ED11DFE" author="Randall JE &amp; Myers RF &amp; Trevor MN &amp; Johnson SR &amp; Johnson JL &amp; Yoshi S &amp; Greene BD" page="119" year="2005">
<bibRefCitation id="D65B22DB535AAA157662BB438FEE1065" author="Randall JE &amp; Myers RF &amp; Trevor MN &amp; Johnson SR &amp; Johnson JL &amp; Yoshi S &amp; Greene BD" etAl="et al." firstAuthor="Randall" issue="3" journalOrPublisher="Journal of Ichthyology and Aquatic Biology" pagination="115-132" refId="B49" refString="Randall JE, Myers RF, Trevor MN, Johnson SR, Johnson JL, Yoshi S, Greene BD (2005) Ninety-one new records of fishes from the Marshall Islands. Aqua. Journal of Ichthyology and Aquatic Biology 9 (3): 115132." title="Ninety-one new records of fishes from the Marshall Islands. Aqua." volume="9" year="2005">Randall et al. (2005)</bibRefCitation>
: 119
</treatmentCitation>
(
<collectingCountry id="48550814C8439BB4FED11B5C6EC27341" name="Marshall Islands">Marshall Islands</collectingCountry>
). —
<treatmentCitation id="26C096C9D5CB30DE43F123E871D74022" author="Kuiter RH &amp; Debelius H" page="313" year="2007">
<bibRefCitation id="B3C9484A250AB3F36AA79505172D7838" author="Kuiter RH &amp; Debelius H" firstAuthor="Kuiter" journalOrPublisher="IKAN-Unterwasserarchiv, Frankfurt" refId="B31" refString="Kuiter RH, Debelius H (2007) World atlas of marine fishes. 2 nd edn. IKAN-Unterwasserarchiv, Frankfurt, 728 pp." title="World atlas of marine fishes. 2 nd edn." year="2007">Kuiter and Debelius (2007)</bibRefCitation>
: 313
</treatmentCitation>
(
<collectingCountry id="88DD670985C2074E7877C858161EA8F2" name="South Africa">South Africa</collectingCountry>
). —
<treatmentCitation id="D194208812A5721CD1E099B2087A6234" author="Khalaf MA &amp; Zajonz U" page="427" year="2007">
<bibRefCitation id="EFF5CA073DF06CE62BE513DD0A3FFC28" author="Khalaf MA &amp; Zajonz U" firstAuthor="Khalaf" journalOrPublisher="Fauna of Arabia" pagination="421-433" refId="B29" refString="Khalaf MA, Zajonz U (2007) Fourteen additional fish species recorded from below 150 m depth in the Gulf of Aqaba, including Liopropoma lunulatum (Pisces: Serranidae), new record for the Red Sea. Fauna of Arabia 23: 421433." title="Fourteen additional fish species recorded from below 150 m depth in the Gulf of Aqaba, including Liopropoma lunulatum (Pisces: Serranidae), new record for the Red Sea." volume="23" year="2007">Khalaf and Zajonz (2007)</bibRefCitation>
: 427
</treatmentCitation>
(Gulf of Aqaba). —
<treatmentCitation id="D47D812A52AC6E7E271E600D3BE68B68" author="Fricke R &amp; Mulochau T &amp; Durville P &amp; Chabanet P &amp; Tessier E &amp; Letourneur Y" page="43" year="2009">
<bibRefCitation id="C64527BEC124C1E653B8282085F395E6" author="Fricke R &amp; Mulochau T &amp; Durville P &amp; Chabanet P &amp; Tessier E &amp; Letourneur Y" etAl="et al." firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde A, Neue Serie" pagination="1-168" refId="B19" refString="Fricke R, Mulochau T, Durville P, Chabanet P, Tessier E, Letourneur Y (2009) Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2: 1168." title="Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species." volume="2" year="2009">Fricke et al. (2009)</bibRefCitation>
: 43
</treatmentCitation>
(
<collectingCountry id="E75F8CA67B19C0AB825E0287ECCAD01F" name="Réunion">La Réunion</collectingCountry>
,
<collectingCountry id="E2D9B7D85C26285019E5CF9869DF699A" name="Mauritius">Mauritius</collectingCountry>
, and
<collectingCountry id="9A6FC80E36555D6D6595B5325F133220" name="Comoros">Comores</collectingCountry>
). —
<treatmentCitation id="70AAC7FEDF123EE5AE95C73AFF3688F7" author="Randall JE &amp; Schraml E" page="170" year="2010">
<bibRefCitation id="E42B805826588B6290B51B2A137F4BEB" author="Randall JE &amp; Schraml E" firstAuthor="Randall" issue="1" journalOrPublisher="International Journal of Ichthyology" pagination="169-174" refId="B45" refString="Randall JE, Schraml E (2010) Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua. International Journal of Ichthyology 16 (1): 169174." title="Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua." volume="16" year="2010">Randall and Schraml (2010)</bibRefCitation>
: 170
</treatmentCitation>
(
<collectingCountry id="4D57E51193C043533096E2BB81C7161C" name="Japan">Japan</collectingCountry>
;
<collectingCountry id="E86D943110C916C2283DAB340097B034" name="Norfolk Island">Norfolk Island</collectingCountry>
). —
<treatmentCitation id="7771529E425955763B1150D46817AEF8" author="Bos AR &amp; Gumanao GS" page="3" year="2013">
<bibRefCitation id="9FF15092CFC130C1C2F5A3ACD0F9507E" DOI="10.1017/S1755267213000614" author="Bos AR &amp; Gumanao GS" firstAuthor="Bos" journalOrPublisher="Marine Biodiversity Records" pagination="1-6" refId="B11" refString="Bos AR, Gumanao GS (2013) Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines. Marine Biodiversity Records 6: 16. https://doi.org/10.1017/S1755267213000614" title="Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines." volume="6" year="2013">Bos and Gumanao (2013)</bibRefCitation>
: 3
</treatmentCitation>
(
<collectingCountry id="CE06BF6F43EB467862F6CE8343F65E0C" name="Philippines">Philippines</collectingCountry>
). —
<treatmentCitation id="BBC9212E928AF9F7932BF6A4453BBB9A" author="Duffy CAJ &amp; Ahyong ST" page="113" year="2015">
<bibRefCitation id="7C7589D9DCE46B09280591C7901A2D3A" author="Duffy CAJ &amp; Ahyong ST" editor="Trnski T &amp; Schlumpf HA" firstAuthor="Duffy" journalOrPublisher="Bulletin of the Auckland Museum" pagination="19-124" refId="B15" refString="Duffy CAJ, Ahyong ST (2015) Annotated checklist of the marine flora and fauna of the Kermadec Islands Marine Reserve and northern Kermadec Ridge, New Zealand. In: Trnski T, Schlumpf HA (Eds.) Kermadec Biodiversity Expedition 2011. Bulletin of the Auckland Museum 60: 19124." title="Annotated checklist of the marine flora and fauna of the Kermadec Islands Marine Reserve and northern Kermadec Ridge, New Zealand." volume="60" volumeTitle="Kermadec Biodiversity Expedition 2011." year="2015">Duffy and Ahyong (2015)</bibRefCitation>
: 113
</treatmentCitation>
(
<collectingCountry id="60607F16A246F0D1C5D77B41DA8A04EF" name="Australia">Australia</collectingCountry>
,
<collectingCountry id="3FE12BDEF8B9C49783EB441D8DB06D43" name="New Zealand">New Zealand</collectingCountry>
). —
<treatmentCitation id="C8600A07F771EC59F0F12B5308F972EA" author="Stewart AL" page="1201" year="2015">
<bibRefCitation id="EBA77878ED6FDB96540F38BDE14B46F8" author="Stewart AL" firstAuthor="Stewart" refId="B54" refString="Stewart AL (2015) Aulacocephalus temminckii. Pp. 11531748. In: Roberts CD, Stewart AL, Struthers CD (Eds.) The Fishes of New Zealand (Vol. 4). Te Papa Press, Wellington, New Zealand." year="2015">Stewart (2015)</bibRefCitation>
: 1201
</treatmentCitation>
(
<collectingCountry id="74C9AE2761D925F2145A4956CE4F8FD2" name="New Zealand">New Zealand</collectingCountry>
). —
<treatmentCitation id="889297E357AEA110C4CD91E2624289E0" author="Golani D &amp; Fricke R" page="60" year="2018">
<bibRefCitation id="E656DDE3A860F113A395F4D4515EB56E" DOI="10.11646/zootaxa.4509.1.1" author="Golani D &amp; Fricke R" firstAuthor="Golani" issue="1" journalOrPublisher="Zootaxa" pagination="1-215" refId="B23" refString="Golani D, Fricke R (2018) Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants. Zootaxa 4509 (1): 1215. https://doi.org/10.11646/zootaxa.4509.1.1" title="Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants." volume="4509" year="2018">Golani and Fricke (2018)</bibRefCitation>
: 60
</treatmentCitation>
(Red Sea). —
<treatmentCitation id="D760F8B53DA6E8C109CC7DC52CDA92C5" author="Akhilesh KV &amp; Rajan PT &amp; Vineesh N &amp; Idreesbabu KK &amp; Bineesh KK &amp; Muktha M &amp; Anulekshmi C &amp; Manjebrayakath H &amp; Gladston Y &amp; Nashad M" page="49" year="2021">
<bibRefCitation id="78FAF03D503D7183D06B3E307D67B6E9" author="Akhilesh KV &amp; Rajan PT &amp; Vineesh N &amp; Idreesbabu KK &amp; Bineesh KK &amp; Muktha M &amp; Anulekshmi C &amp; Manjebrayakath H &amp; Gladston Y &amp; Nashad M" etAl="et al." firstAuthor="Akhilesh" journalOrPublisher="Journal of the Ocean Science Foundation" pagination="35-65" refId="B2" refString="Akhilesh KV, Rajan PT, Vineesh N, Idreesbabu KK, Bineesh KK, Muktha M, Anulekshmi C, Manjebrayakath H, Gladston Y, Nashad M (2021) Checklist of serranid and epinephelid fishes (Perciformes: Serranidae &amp; Epinephelidae) of India. Journal of the Ocean Science Foundation 38: 3565." title="Checklist of serranid and epinephelid fishes (Perciformes: Serranidae &amp; Epinephelidae) of India." volume="38" year="2021">Akhilesh et al. (2021)</bibRefCitation>
: 49
</treatmentCitation>
(
<collectingCountry id="065F6829B439F453CA921FDA996F93EC" name="India">India</collectingCountry>
). —
<treatmentCitation id="9584045A5B9DD6C60E93A570640E3D45" author="Heemstra PC" page="79" year="2022">
<bibRefCitation id="61E94727DD4A2334A2E3AFB6814108AF" author="Heemstra PC" firstAuthor="Heemstra" refId="B25" refString="Heemstra PC (2022) Epinephelidae: groupers, podges and soapfishes. Pp. 3687. In: Heemstra PC, Heemstra E, Ebert DA, Holleman W, Randall JE (Eds.) Coastal fishes of the western Indian Ocean (Vol. 3). South African Institute for Aquatic Biodiversity, Makhanda, South Africa." year="2022">Heemstra (2022)</bibRefCitation>
: 79
</treatmentCitation>
(western Indian Ocean).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWFAE" type="material">
<paragraph id="3F006ABAA64878F6F5774E6A79E49F16">
<heading id="D79C0DC8709DC1CEDB478AFCA1AD3F25" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="3A7ECC867E726D6140F5F4F45E80F591">
<materialsCitation id="79DCDA1FB5105413C161F2823976BE1A" collectingDate="2008-08-18" collectingDateMax="2008-12-13" collectingDateMin="2008-08-18" collectingMethod="hook and line" collectionCode="LBRC-F" collectorName="T. Peristiwady" elevation="125" location="off Lembeh Island" specimenCode="LBRC-F 367, LBRC-F 402" specimenCount="1">
<specimenCode id="0B6110EC896B10D6181B1A62AA58E499">
<abbrev content-type="institution" id="ABBRID0E3FAE" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
<quantity id="6F6CC0C501120C3AD5B05A96D2202842" metricMagnitude="2" metricUnit="m" metricValue="3.671907" unit="mm" value="367190.7">367, 190.7 mm</quantity>
</specimenCode>
SL,
<specimenCode id="A8C75D9C8EB4A4811470B8E457E5C41B">
<abbrev content-type="institution" id="ABBRID0EBGAE" xlink_title="Research Center for Oceanography">LBRC-F</abbrev>
<quantity id="30B2341E8CD5CFAEAB7DFFA5E24570DC" metricMagnitude="2" metricUnit="m" metricValue="4.021521" unit="mm" value="402152.1">402, 152.1 mm</quantity>
</specimenCode>
SL,
<location id="34366F742194A092C32A9D50F16862CA" LSID="urn:lsid:plazi:treatment:DCDB7A653DAA522F8129DB5FC94D738E:34366F742194A092C32A9D50F16862CA" name="off Lembeh Island">off Lembeh Island</location>
,
<collectingMethod id="7428ECE6110E58767EC252C399DBC772">hook and line</collectingMethod>
,
<quantity id="FE6D7A2C714E8811DF9DB26E64BEA58F" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">
<elevation id="7705BE9239A4175F969013FDD968B444" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100150 m</elevation>
</quantity>
(both purchased at Girian Fish Market, Bitung, northern Sulawesi,
<collectorName id="E5E1F1774646AEF12EABC2A67D4EEB35">T. Peristiwady</collectorName>
,
<date id="FD83352D7A93ADF41D26E0A373D0C0DB" value="2008-08-18" valueMax="2008-12-13" valueMin="2008-08-18">
<collectingDate id="8E3ABCC18DF2917F585EE8A107E97703" value="2008-08-18" valueMax="2008-12-13" valueMin="2008-08-18">18 August and 13 December 2008</collectingDate>
</date>
, respectively)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGGAE" type="description">
<paragraph id="6F02714D115F6377BB57898B4FF88B2C">
<heading id="CC9A56789446FDA5780776633F4DB996" reason="title">Description.</heading>
</paragraph>
<paragraph id="3B3D30CDAF65094116B0E58992460E7F">
Dorsal fin with 9 spines and 12 soft rays, fourth spine longest, its length
<quantity id="1B76B573EF7EBDF12D11CAAA6673A58C" metricMagnitude="-2" metricUnit="m" metricValue="8.255" metricValueMax="8.382" metricValueMin="8.128" unit="in" value="3.25" valueMax="3.3" valueMin="3.2">3.23.3 in</quantity>
HL, first and last dorsal spines subequal, their lengths about half length of fourth spine; anal fin with 3 spines and 9 soft rays, third or fourth ray longest, its length
<quantity id="2E0BD056D8B50BD1AF6F11D2E6D48C91" metricMagnitude="-2" metricUnit="m" metricValue="6.731" metricValueMax="7.111999999999999" metricValueMin="6.35" unit="in" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 in</quantity>
HL; pectoral fin with 15 rays; pored lateral-line scales 7479; scales above lateral line 13; scales below lateral line 44; gill rakers on upper limb of first gill arch 9, lower limb 18, total 27; body depth
<quantity id="4B53A65E05DCD56D129E91E97DAAB944" metricMagnitude="-2" metricUnit="m" metricValue="7.238999999999999" metricValueMax="7.366" metricValueMin="7.111999999999999" unit="in" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.82.9 in</quantity>
SL; body width behind gill opening 2.83.0 in body depth; head length
<quantity id="4001206A0CA9E98C45D40CE629749DE7" metricMagnitude="-2" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" unit="in" value="2.45" valueMax="2.5" valueMin="2.4">2.42.5 in</quantity>
SL; snout relatively long, its length
<quantity id="D2438071BB0160314B929F23FFB42873" metricMagnitude="-1" metricUnit="m" metricValue="1.7145" metricValueMax="1.7526" metricValueMin="1.6764" unit="in" value="6.75" valueMax="6.9" valueMin="6.6">6.66.9 in</quantity>
HL; mouth slightly oblique; pre-opercle coarsely serrated; subopercle and interopercle serrated; interorbital width
<quantity id="036B23D860C7C14F734A1D30DB3EBBD1" metricMagnitude="-1" metricUnit="m" metricValue="1.8034" metricValueMax="1.9304" metricValueMin="1.6764" unit="in" value="7.1" valueMax="7.6" valueMin="6.6">6.67.6 in</quantity>
HL; origin of pelvic fin just below origin of pectoral fin, pelvic fin short, its length
<quantity id="31077CE6AAC11499F88F2D2BC84BB4BC" metricMagnitude="-1" metricUnit="m" metricValue="2.4003" metricValueMax="2.4638" metricValueMin="2.3367999999999998" unit="in" value="9.45" valueMax="9.7" valueMin="9.2">9.29.7 in</quantity>
HL, posterior tip not reaching anus when depressed; caudal peduncle depth
<quantity id="6D477AD782416AD895415CEE246BA0A3" metricMagnitude="-2" metricUnit="m" metricValue="8.001" metricValueMax="8.128" metricValueMin="7.874" unit="in" value="3.15" valueMax="3.2" valueMin="3.1">3.13.2 in</quantity>
HL; caudal peduncle length
<quantity id="96B239513E8079C42891E284C2BAE9AC" metricMagnitude="-2" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" unit="in" value="2.2" valueMax="2.3" valueMin="2.1">2.12.3 in</quantity>
HL; head (except snout, interorbital, and maxillary) and body covered with ctenoid scales.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELGAE" type="Color when fresh">
<paragraph id="ECBFACF527B09D7517463601989D2CB8">
<heading id="A43D1E1F7E31FFCF30D914CC2E59EB8A" reason="title">Color when fresh.</heading>
</paragraph>
<paragraph id="D2771D638849E09517D273235D303AC8">
Body bluish grey, a bright yellow band running from near snout tip through eye, along upper part of back and dorsal fin base to upper and lower bases of caudal fin; a yellow streak along upper margin of upper jaw; soft portions of median fins with blackish rays and translucent membranes; pectoral and pelvic fin membranes translucent (Fig.
<figureCitation id="4E8F6443548FECD485F417C6F02EA96D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of fresh specimens of (A) Plectranthias yamakawai, LBRC-F 3002, 182.5 mm SL and (B) Aulacocephalus temminckii, LBRC-F 402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia." figureDoi="10.3897/aiep.55.142574.figure2" httpUri="https://binary.pensoft.net/fig/1262311">2 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVGAE" type="remarks">
<paragraph id="11EF01BE0103216C776E771BDD6B4F12">
<heading id="07C73E41F64E4C2EB269FE2DD11EE023" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A96E066EA540985E994594C34564325B">
The following diagnostic features of the presently reported specimens closely matched those of
<taxonomicName id="BBC7C8CCAC00D6284B276E13234F9750" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="D7641024B2E39024E481551BD5BEAC46" italics="true">A. temminckii</emphasis>
</taxonomicName>
given by
<bibRefCitation id="159037C71CE3DFDC6FFDAF4A5305E513" author="Heemstra PC &amp; Randall JE" firstAuthor="Heemstra" refId="B26" refString="Heemstra PC, Randall JE (1999) Serranidae. Pp. 24422548. In: Carpenter KE, Niem VH (Eds.) Species identification guide for fisheries purposes. The living marine resources of the western central Pacific. Bony fishes part 2 (Mugilidae to Carangidae). FAO, Rome." year="1999">Heemstra and Randall (1999)</bibRefCitation>
: for example, dorsal fin single, with 9 spines and 12 soft rays, last two spines subequal; anal fin with 3 spines and 9 soft rays; body depth less than head length
<quantity id="73A353586A15543ABD66275C60260F2C" metricMagnitude="-2" metricUnit="m" metricValue="7.238999999999999" metricValueMax="7.366" metricValueMin="7.111999999999999" unit="in" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.82.9 in</quantity>
SL; caudal fin rounded; pelvic fin short, its posterior tip not reaching anus when depressed; posterior margin of pre-opercle, subopercle and interorpercle serrated; head and body scales ctenoid; body bluish-grey, with a bright yellow band from near front of upper jaw through eye, along upper part of back and dorsal fin base to caudal fin base.
</paragraph>
<paragraph id="5AEB8CF77F31073887293A532AD36797">
There appears to be variation in the coloration of the posterior end of the yellow band running along the upper part of the body. Specimens of
<taxonomicName id="9BAFF4F9DE71A4CE3BF5229318454109" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="5EA3B6158D629F8FF05AAAE1ABF5A51F" italics="true">A. temminckii</emphasis>
</taxonomicName>
collected from the Red Sea and Japanese waters exhibit the band terminating at the upper caudal fin base, accompanied by a yellow blotch on the lower caudal fin base (see
<bibRefCitation id="BB3D2B5E62F1C0BEE71C5E9695CA7453" author="Khalaf MA &amp; Zajonz U" firstAuthor="Khalaf" journalOrPublisher="Fauna of Arabia" page="427" pagination="421-433" refId="B29" refString="Khalaf MA, Zajonz U (2007) Fourteen additional fish species recorded from below 150 m depth in the Gulf of Aqaba, including Liopropoma lunulatum (Pisces: Serranidae), new record for the Red Sea. Fauna of Arabia 23: 421433." title="Fourteen additional fish species recorded from below 150 m depth in the Gulf of Aqaba, including Liopropoma lunulatum (Pisces: Serranidae), new record for the Red Sea." volume="23" year="2007">Khalaf and Zajonz 2007: 427</bibRefCitation>
;
<bibRefCitation id="70B3ABBD3127DAB56685B4007FF1C041" author="Randall and Schraml" firstAuthor="Randall" issue="1" journalOrPublisher="International Journal of Ichthyology" pagination="169-174" refId="B45" refString="Randall JE, Schraml E (2010) Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua. International Journal of Ichthyology 16 (1): 169174." title="Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua." volume="16" year="2010">Randall and Schraml 2010</bibRefCitation>
: fig. 4). In contrast, specimens from
<collectingCountry id="D751540ED041756747B5B4B4D5D0FB27" name="Norfolk Island">Norfolk Island</collectingCountry>
show the band ending at the upper caudal fin base without a blotch on the lower base (see
<bibRefCitation id="34085A2E248D9C8F08F57D513A8BF833" author="Randall and Schraml" firstAuthor="Randall" issue="1" journalOrPublisher="International Journal of Ichthyology" pagination="169-174" refId="B45" refString="Randall JE, Schraml E (2010) Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua. International Journal of Ichthyology 16 (1): 169174." title="Notes on soapfishes (Serranidae: Grammistinae), with a pictorial review of the genus Pogonoperca. Aqua." volume="16" year="2010">Randall and Schraml 2010</bibRefCitation>
: fig. 5). Meanwhile, specimens from the
<collectingCountry id="9346EAB9DBA6814B1436C827EC83FC75" name="Philippines">Philippines</collectingCountry>
and Lembeh Island waters show the band extending posteriorly to form a vertical band across both the upper and lower bases of the caudal fin (see
<bibRefCitation id="5F7397E0474BE797BE5D672CAFF91D4D" DOI="10.1017/S1755267213000614" author="Bos and Gumanao" firstAuthor="Bos" journalOrPublisher="Marine Biodiversity Records" pagination="1-6" refId="B11" refString="Bos AR, Gumanao GS (2013) Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines. Marine Biodiversity Records 6: 16. https://doi.org/10.1017/S1755267213000614" title="Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines." volume="6" year="2013">Bos and Gumanao 2013</bibRefCitation>
: fig. 4 A; this study: Fig.
<figureCitation id="15F54E03A94A5DE7CA7AA51D34260765" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of fresh specimens of (A) Plectranthias yamakawai, LBRC-F 3002, 182.5 mm SL and (B) Aulacocephalus temminckii, LBRC-F 402, 152.1 SL, collected off Lembeh Island, northern Sulawesi, Indonesia." figureDoi="10.3897/aiep.55.142574.figure2" httpUri="https://binary.pensoft.net/fig/1262311">2 B</figureCitation>
).
</paragraph>
<paragraph id="5286D013AD2CEBCC5646021EF6DE74A5">
Although
<taxonomicName id="17F917503FCB93A93252BC37EB449517" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="087BECD5740ED4E4EBB632A91066D30C" italics="true">A. temminckii</emphasis>
</taxonomicName>
is widely distributed in the Red Sea and Indo west Pacific, except for the Hawaiian Islands (Fig.
<figureCitation id="867DC3DDFB2D536F6EDEF9BF6C381B86" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distributional records of Plectranthias yamakawai (squares), Aulacocephalus temminckii (dots), Iniistius geisha (triangles), and Randallichthys filamentosus (diamonds), based on previous studies. Lembeh Island, northern Sulawesi, Indonesia (the collection site of presently reported specimens) is marked by a star." figureDoi="10.3897/aiep.55.142574.figure1" httpUri="https://binary.pensoft.net/fig/1262310">1</figureCitation>
); it is relatively rare in collections (
<bibRefCitation id="7C77C8B3D83D3BB8E5B09A887AD7A2DC" DOI="10.1007/BF02725185" author="Baldwin and Smith" firstAuthor="Baldwin" issue="4" journalOrPublisher="Ichthyological Research" pagination="325-339" refId="B9" refString="Baldwin CC, Smith WL (1998) Belonoperca pylei, a new species of seabass (Teleostei: Serranidae: Epinephelinae: Diploprionini) from the Cook Islands with comments on relationships among diploprionins. Ichthyological Research 45 (4): 325339. https://doi.org/10.1007/BF02725185" title="Belonoperca pylei, a new species of seabass (Teleostei: Serranidae: Epinephelinae: Diploprionini) from the Cook Islands with comments on relationships among diploprionins." volume="45" year="1998">Baldwin and Smith 1998</bibRefCitation>
). The distribution of
<taxonomicName id="18E4F2DE38748F0C8C93BFDB6B3BD2F3" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="F4234315724CE8CEA7065581BB0612E3" italics="true">A. temminckii</emphasis>
</taxonomicName>
in Southeast Asia was previously reported as limited to the
<collectingCountry id="0A58D070D7E09D62CA1C470C82346849" name="Philippines">Philippines</collectingCountry>
(
<bibRefCitation id="3B241D4E3CF6B50262CD5F33D0C72F25" DOI="10.1017/S1755267213000614" author="Bos and Gumanao" firstAuthor="Bos" journalOrPublisher="Marine Biodiversity Records" pagination="1-6" refId="B11" refString="Bos AR, Gumanao GS (2013) Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines. Marine Biodiversity Records 6: 16. https://doi.org/10.1017/S1755267213000614" title="Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines." volume="6" year="2013">Bos and Gumanao 2013</bibRefCitation>
), the presently reported specimens being the first recorded from
<collectingCountry id="913854E618FC7305653CA14CB7251BD2" name="Indonesia">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="CCECE906CCE253A3BD44356C03B5B848">
<taxonomicName id="4663E669441D93C4895175CADCC2EDC5" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="97B0A64BDD037D5A2762F3D9CC8992F6" italics="true">Aulacocephalus temminckii</emphasis>
</taxonomicName>
is common in subtropical waters between 20 and 36 ° N or S (
<bibRefCitation id="7DFF17493FF79F693A4AF82FF758AD69" DOI="10.5134/175661" author="Randall" etAl="et al." firstAuthor="Randall" issue="2-3" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pagination="157-190" refId="B46" refString="Randall JE, Aida K, Hibiya T, Matsuura N, Kamiya H, Hashimoto Y (1971) Grammistin, the skin toxin of soapfishes and its significance in the classification of the Grammistidae. Publications of the Seto Marine Biological Laboratory 19 (23): 157190. https://doi.org/10.5134/175661" title="Grammistin, the skin toxin of soapfishes and its significance in the classification of the Grammistidae." volume="19" year="1971">Randall et al. 1971</bibRefCitation>
) but has not previously been reported in equatorial waters (
<bibRefCitation id="AFD305817E538CCD649BD1337404C31E" author="Randall" firstAuthor="Randall" issue="3" journalOrPublisher="Pacific Science" pagination="197-209" refId="B42" refString="Randall JE (1982) Examples of antitropical and antiequatorial distribution of Indo west-Pacific fishes. Pacific Science 35 (3): 197209." title="Examples of antitropical and antiequatorial distribution of Indo - west-Pacific fishes." volume="35" year="1982">Randall 1982</bibRefCitation>
). Prior to the presently reported account, the lowest latitudes reported for
<taxonomicName id="C31A26D8C7ACB67C9DD934C80BAB6F15" authorityName="Bleeker" authorityYear="1855" family="Serranidae" genus="Aulacocephalus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="temminckii">
<emphasis id="CAB37EAC19EB66013203382BFAC8E0FB" italics="true">A. temminckii</emphasis>
</taxonomicName>
were 11 ° N (Port Blair, Andaman Islands) (
<bibRefCitation id="0ECE2AAD4357432AF493408A55B7B4FA" author="Rangarajan" firstAuthor="Rangarajan" issue="2" journalOrPublisher="Journal of the Marine Biological Association of India" pagination="442-444" refId="B50" refString="Rangarajan K (1969) Aulacocephalus temminckii Bleeker (Pisces: Serranidae), a new record from the Andaman Sea. Journal of the Marine Biological Association of India 9 (2): 442444." title="Aulacocephalus temminckii Bleeker (Pisces: Serranidae), a new record from the Andaman Sea." volume="9" year="1969">Rangarajan 1969</bibRefCitation>
) and 11 ° S (
<collectingCountry id="0A503691E6989B395245020D55314E7C" name="Comoros">Comoros</collectingCountry>
, eastern Africa) (
<bibRefCitation id="2DA36C5A15CE5B433EFAE9C3530DD57D" author="Fricke" etAl="et al." firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde A, Neue Serie" pagination="1-168" refId="B19" refString="Fricke R, Mulochau T, Durville P, Chabanet P, Tessier E, Letourneur Y (2009) Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2: 1168." title="Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species." volume="2" year="2009">Fricke et al. 2009</bibRefCitation>
), considered as evidence of an anti-equatorial distribution (
<bibRefCitation id="3C40FB520352E4AA2E872EA83DC9A6D2" author="Randall" firstAuthor="Randall" issue="3" journalOrPublisher="Pacific Science" pagination="197-209" refId="B42" refString="Randall JE (1982) Examples of antitropical and antiequatorial distribution of Indo west-Pacific fishes. Pacific Science 35 (3): 197209." title="Examples of antitropical and antiequatorial distribution of Indo - west-Pacific fishes." volume="35" year="1982">Randall 1982</bibRefCitation>
;
<bibRefCitation id="99BB4F81F4321BC60FFD6CCAC487705B" DOI="10.5479/si.00810282.367" author="Springer" firstAuthor="Springer" issue="367" journalOrPublisher="Smithsonian Contributions to Zoology" pagination="1-182" refId="B53" refString="Springer V (1982) Pacific plate biogeography with special reference to shorefishes. Smithsonian Contributions to Zoology 367 (367): 1182. https://doi.org/10.5479/si.00810282.367" title="Pacific plate biogeography with special reference to shorefishes." volume="367" year="1982">Springer 1982</bibRefCitation>
;
<bibRefCitation id="D41D358C66DCDE7834B1C2807EE56EA2" author="Fricke" etAl="et al." firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde A, Neue Serie" pagination="1-168" refId="B19" refString="Fricke R, Mulochau T, Durville P, Chabanet P, Tessier E, Letourneur Y (2009) Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2: 1168." title="Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species." volume="2" year="2009">Fricke et al. 2009</bibRefCitation>
). However,
<bibRefCitation id="CBEDD60F091E5D8DAEF0D09EE52ABAD5" DOI="10.1017/S1755267213000614" author="Bos AR &amp; Gumanao GS" firstAuthor="Bos" journalOrPublisher="Marine Biodiversity Records" pagination="1-6" refId="B11" refString="Bos AR, Gumanao GS (2013) Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines. Marine Biodiversity Records 6: 16. https://doi.org/10.1017/S1755267213000614" title="Seven new records of fish (Teleostei: Perciformes) from coral reefs and pelagic habitats in southern Mindanao, the Philippines." volume="6" year="2013">Bos and Gumanao (2013)</bibRefCitation>
reported the first record of the species from the
<collectingCountry id="D22A1958A4FE8CED444D64FB3D9C9637" name="Philippines">Philippines</collectingCountry>
, based on a specimen collected at Peñaplata Public Market, southern Mindanao (caught latitude ca. 7 ° N in Samal Island,
<collectingRegion id="F1BF12EAF0AF0D21618A594C9FE493F2" country="Philippines" name="Davao">Davao</collectingRegion>
Gulf). The presently reported specimen, collected off Lembeh Island (latitude 1.4 ° N) indicates that the anti-equatorial distribution, recognized by
<bibRefCitation id="F2C97E1AD392BE62E1EF1B6F0B6E38DA" author="Randall JE" firstAuthor="Randall" issue="3" journalOrPublisher="Pacific Science" pagination="197-209" refId="B42" refString="Randall JE (1982) Examples of antitropical and antiequatorial distribution of Indo west-Pacific fishes. Pacific Science 35 (3): 197209." title="Examples of antitropical and antiequatorial distribution of Indo - west-Pacific fishes." volume="35" year="1982">Randall (1982)</bibRefCitation>
,
<bibRefCitation id="9F4C063E81196FE41831B815FB7C9D4C" DOI="10.5479/si.00810282.367" author="Springer V" firstAuthor="Springer" issue="367" journalOrPublisher="Smithsonian Contributions to Zoology" pagination="1-182" refId="B53" refString="Springer V (1982) Pacific plate biogeography with special reference to shorefishes. Smithsonian Contributions to Zoology 367 (367): 1182. https://doi.org/10.5479/si.00810282.367" title="Pacific plate biogeography with special reference to shorefishes." volume="367" year="1982">Springer (1982)</bibRefCitation>
, and
<bibRefCitation id="B21D1719C4EE5987D7FB55B6D1DE7364" author="Fricke R &amp; Mulochau T &amp; Durville P &amp; Chabanet P &amp; Tessier E &amp; Letourneur Y" etAl="et al." firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde A, Neue Serie" pagination="1-168" refId="B19" refString="Fricke R, Mulochau T, Durville P, Chabanet P, Tessier E, Letourneur Y (2009) Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2: 1168." title="Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species." volume="2" year="2009">Fricke et al. (2009)</bibRefCitation>
, is no longer valid.
</paragraph>
</subSubSection>
</treatment>
</document>