This commit is contained in:
maintenance 2024-06-21 12:41:11 +02:00
parent 67de862e33
commit da110f39cc
1798 changed files with 349592 additions and 0 deletions

View file

@ -0,0 +1,161 @@
<document id="7996C0D4B4EBE96CEDC7868B5B73405F" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico &amp; Vaz-De-Mello, Fernando Z." docDate="2019" docId="7AFC2C9B20ED271AE7EA69353E2474FF" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Dichotomius (Dichotomius) quinquedens Felsche 1910" docType="treatment" docVersion="7" lastPageNumber="119" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="118" updateTime="1701379963039" updateUser="ExternalLinkService">
<mods:mods id="DEC11FA72B2C1374213F409F4F0FDA27" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="78E083BF78904DF9CE946AECBDA3014E">
<mods:title id="634EA28CD31BEAA675FF97FEE1F2DE8C">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
</mods:titleInfo>
<mods:name id="7FE8EB13F47CE19B40BAA95A8F0B8A43" type="personal">
<mods:role id="F9230AC25486791E0EAAB4851CEA331E">
<mods:roleTerm id="D33E70C808456D09BDC519E66644779B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3BE4EF72AF5EE9A09FC81BBCDB68A5C3">Chamorro, William</mods:namePart>
</mods:name>
<mods:name id="040942698FCAC6A0530E1E8188410178" type="personal">
<mods:role id="B3E28FCC3E2970A33B9BBF69CDFA0721">
<mods:roleTerm id="F532F172F734D833A9A918F27AC8DDB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A460B2E61D008F29534FE09BDCD4D02C">Marin-Armijos, Diego</mods:namePart>
</mods:name>
<mods:name id="B50637DF212BF62B2714843E6753B44C" type="personal">
<mods:role id="3AE16E428327FD6CA57210AD834098D2">
<mods:roleTerm id="546A45F670218500DFB576802F4CF39D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6873E8130055DA60F2AAB53C24EFCCBB">senjo, Angelico</mods:namePart>
</mods:name>
<mods:name id="248A4E3D03C42A01540D903D52B610E2" type="personal">
<mods:role id="EE096A2CCE96FD0591B0B4109E344B19">
<mods:roleTerm id="040F44AF7B9E047731BB9E230A368284">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64CC5AB1A9D784943B942BF8901BDE8D">Vaz-De-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A4D70617CFC7AC15913E7243357A00E8">text</mods:typeOfResource>
<mods:relatedItem id="4B7BEF6ADE140478A9415F9D296ABA23" type="host">
<mods:titleInfo id="F3355CAC10CEF1BE6979B9BAFF15DACF">
<mods:title id="61C9B0D87D2AE920DB4437456D83685E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="07B196D81FB79FA33202D3D356A86A51">
<mods:date id="DAC121FBED4AF2A524D9FE903A2ED959">2019</mods:date>
<mods:detail id="4C4EA537AF26B5AFED2BEF99A286F76A" type="volume">
<mods:number id="2F2D21C4411AE241533B43238D3F7D01">826</mods:number>
</mods:detail>
<mods:extent id="BE55F4CB1EAADF4C12DBF4DE49A79514" unit="page">
<mods:start id="9902DB490A6BB79E8A3BF947CB2C965B">1</mods:start>
<mods:end id="759E7A5E012D1F77151FFDB7F713C600">343</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="80CF599F98A11AF5AF164B83077734E1">
<mods:url id="DA69A4235F8AF02C7786656AB79DF5DE">http://dx.doi.org/10.3897/zookeys.826.26488</mods:url>
</mods:location>
<mods:classification id="66321472D3B7D1027A93BA22900F80F9">journal article</mods:classification>
<mods:identifier id="32AB267E08435228B242B1D353161434" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
<mods:identifier id="0C4C0A4D24E855D18B626FB33C824874" type="Pensoft-Pub">1313-2970-826-1</mods:identifier>
<mods:identifier id="6FEED90A7D5B89F85A09DC3A81D2DDD3" type="ZooBank">B1550A3AE54744509A44BC4366D5E110</mods:identifier>
</mods:mods>
<treatment id="7AFC2C9B20ED271AE7EA69353E2474FF" ID-GBIF-Taxon="154127761" LSID="urn:lsid:plazi:treatment:7AFC2C9B20ED271AE7EA69353E2474FF" httpUri="http://treatment.plazi.org/id/7AFC2C9B20ED271AE7EA69353E2474FF" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118">
<subSubSection id="58F8AA05E3F0367408C598DC80466D2D" pageId="117" pageNumber="118" type="nomenclature">
<paragraph id="E8988D44C34206F2FE0AD1485A1D392A" pageId="117" pageNumber="118">
<taxonomicName id="15C4DA7A4785CBEE1761AF4049B9F965" ID-CoL="7ZN55" authority="Felsche, 1910" authorityName="Felsche" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens" subGenus="Dichotomius">Dichotomius (Dichotomius) quinquedens (Felsche, 1910)</taxonomicName>
Plate 27A
</paragraph>
</subSubSection>
<subSubSection id="FADE7D0C5C4C9D19D2C342FDA99F46DB" pageId="117" pageNumber="118" type="reference_group">
<paragraph id="385409ABFC7622FC6043D5701D4F4A8F" pageId="117" pageNumber="118">
<taxonomicName id="CE441F156709A4B031CFB0BAAE421003" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens">Pinotus quinquedens</taxonomicName>
Felsche, 1910: 343 (original description. Type locality: Ecuador, Los Llanos).
</paragraph>
<paragraph id="DBE1FBA2F33D3147131FA73B96DD049F" pageId="117" pageNumber="118">
<taxonomicName id="D16F53A89585924F4F62F9A530B6F20A" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens">Pinotus quinquedens</taxonomicName>
:
<bibRefCitation id="521E02BA661E97DC1D9578EA99768E41" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
: 62 (complete list of species);
<bibRefCitation id="0300CDB5A057FAD93CFD1EF59D6EC0D4" author="Luederwaldt, H" journalOrPublisher="Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 174" title="As especies brasileiras do genero Pinotus, com algumas consideractes tambem sobra outras especies." volume="16" year="1929">Luederwaldt 1929</bibRefCitation>
: 35 (characters in key);
<bibRefCitation id="2B9438F1E68B131D74AA572D67DCD58C" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 208 (list of species from Latin America).
</paragraph>
<paragraph id="CBBF6E20BF6E66E58C41AF6B93AFB5E4" pageId="117" pageNumber="118">
<taxonomicName id="DAA1C57F84F4C2E571ED36DF5F2B96FE" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens">Dichotomius quinquedens</taxonomicName>
:
<bibRefCitation id="7980555496119F2F8DF7FE8A1C41D285" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
: 138 (cited for Colombia);
<bibRefCitation id="288256FC21F008026CD88F2741CA377A" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
: 320-321 (cited for Ecuador);
<bibRefCitation id="2E4BE2C0882E1B1AD8D463FEA17D92E6" pageId="117" pageNumber="118">Krajcik 2012</bibRefCitation>
: 92 (complete list of species);
<bibRefCitation id="06BE26CB6C982CC8E76BD30685D0E102" author="Sarmiento-Garces, R" journalOrPublisher="Universidad Nacional de Colombia, Facultad de Ciencias, Instituto de Ciencias Naturales" pageId="276" pageNumber="277" title="Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia Monografia No. 4." year="2014">
<normalizedToken id="E728D6877161BF4328D1B0530C8823E2" originalValue="Sarmiento-Garcés">Sarmiento-Garces</normalizedToken>
and
<normalizedToken id="2A07C038EF7DFE719648F229D4493553" originalValue="Amat-García">Amat-Garcia</normalizedToken>
2014
</bibRefCitation>
: 53 (characters in key), 64 (diagnosis).
</paragraph>
<paragraph id="D528A042EBBC82049917E0AED4B075F8" pageId="117" pageNumber="118">
<taxonomicName id="E5978C0428D80F8C80B3AEBADA1E42BC" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens" subGenus="Dichotomius">Dichotomius (Dichotomius) quinquedens</taxonomicName>
:
<bibRefCitation id="75BB906A56477DD020C03EBA5F30D768" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
: 95 (cited for Ecuador).
</paragraph>
</subSubSection>
<subSubSection id="5CB3E50A59E37393322BF78ECD1CC473" pageId="117" pageNumber="118" type="types specimens">
<paragraph id="9A4061496FC5B70D05B451B55598D4F6" pageId="117" pageNumber="118">Types specimens.</paragraph>
<paragraph id="DACD9497D74AA404BCE20D8BE0FF8E5D" pageId="117" pageNumber="118">
<taxonomicName id="FA09BC462EEEDC56EC6FA1B745843102" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus quinquedens" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="quinquedens">Pinotus quinquedens</taxonomicName>
Felsche, 1910. Seven syntypes examined deposited at the SMTD. Lectotype to be designated in a future work on this species group.
</paragraph>
<paragraph id="B674B34224AE930C81CF1389B53DC57A" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;quinquendens- / tatus Felsche / Ecuador [hw, purpure margin]&quot;, &quot;Typus. [p, red label]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="941DA93436D3C31A1A9470E91E5FB92C" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="67DC40965B20F6D8516B2912B3D0B2AB" pageId="117" pageNumber="118">Syntype (♀): &quot;Los Lanos / Ecuador [p, red label, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="B8B2E8E6BB5879DF0CB25F6E1920C145" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="C9D7EAB06E987ED71F2FBD9A1894CA3E" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="E9548790688F41D324B03976B70058E9" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
<paragraph id="D1E84F9CDD41C3640885CB343ABCE9F4" pageId="117" pageNumber="118">Syntype (♂): &quot;Los Lanos / Ecuador [p, black margin]&quot;, &quot;Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]&quot;.</paragraph>
</subSubSection>
<subSubSection id="9D4DF1496665B16652C2F85012A7E6F1" pageId="117" pageNumber="118" type="distribution">
<paragraph id="03C708C57756527799605884CA44C706" pageId="117" pageNumber="118">Distribution.</paragraph>
<paragraph id="E60CB3BCEC95892CB22D79AE77E509DA" pageId="117" pageNumber="118">Colombia and Ecuador.</paragraph>
</subSubSection>
<subSubSection id="5E7FEEA0C3E243F08290C7A951EE01B1" pageId="117" pageNumber="118" type="records examined">
<paragraph id="D6BD005349E7EEA48A235D18582CA1C3" pageId="117" pageNumber="118">Records examined.</paragraph>
<paragraph id="DE8479554A78BAE3851FE22A3E3BC5BD" pageId="117" pageNumber="118">
BOLIVAR: Bosque Protector Filo Palanga, 970 m (1 specimen MUTPL).
<normalizedToken id="5DAD384AA97B2A5E0D7945051EB12263" originalValue="CAÑAR">CANAR</normalizedToken>
: Cochancay, 1000 m (4 specimens CEMT);
<normalizedToken id="2DA82BFF87897F9F55B2B75F73B8AC6B" originalValue="Javín">Javin</normalizedToken>
, 850-1400 m (4 specimens CEMT); Joyapal (1 specimen MQCAZ). COTOPAXI: 4 km de Guasaganda, 500 m (2 specimens MQCAZ); Bosque Integral Otonga, 1800 m (2 specimens CEMT; 7 specimens MQCAZ); Los Libres, 2015 m (1 specimen MQCAZ); San Francisco de Las Pampa, 1500 m (7 specimens CEMT). EL ORO: Salvias
<normalizedToken id="1A11EBB7B60431FB92A3BBCA04C1CFA1" originalValue="Río">Rio</normalizedToken>
Elvira, 1180 m (2 specimens MUTPL); Reserva Jocotoco, 1250 m (1 specimen MQCAZ). ESMERALDAS: El Placer, road Ibarra-San Lorenzo, 670 m (2 specimens MQCAZ);
<normalizedToken id="7A439B32C99C65F744860871327F7508" originalValue="Estación">Estacion</normalizedToken>
<normalizedToken id="02F48A57F7A608C2BF56E1FFCBC5A80D" originalValue="Biológica">Biologica</normalizedToken>
Bilsa, 500 m (1 specimen MEPN); Playa de Oro, Estero Pote, 200 m (1 specimen CEMT). IMBABURA: Lita (2 specimens CEMT); Santa Cecilia (1 specimen QCAZ).
<normalizedToken id="86564D09A138CDC8F11C9931A264557C" originalValue="MANABÍ">MANABI</normalizedToken>
: Puerto
<normalizedToken id="75C805B888DFF4C127C7523C340131A5" originalValue="López">Lopez</normalizedToken>
, Guale, 200 m (1 specimen MUTPL); Puerto
<normalizedToken id="DA80E5F5FCE445A7C9DBAD764B080401" originalValue="López">Lopez</normalizedToken>
, San Sebastian, 480 m (1 specimen MUTPL). PICHINCHA: Choconde San Miguel de los Bancos, 1200 m (4 specimens MUTPL); Chespi, Bellavista, 1380 (4 specimens MUTPL); Chiriboga, 1500 m (1 specimen MQCAZ);
<normalizedToken id="0583286A5A1E2958DCFE23E9F6A67215" originalValue="Estación">Estacion</normalizedToken>
<normalizedToken id="73F5C954346473F07746FCE6C6E57E80" originalValue="Biológica">Biologica</normalizedToken>
Maquipucuna, 1650 (2 specimens MUTPL);
<normalizedToken id="A4B98262B189B5D0786F018379D33144" originalValue="Estación">Estacion</normalizedToken>
<normalizedToken id="DC6ECF1BE3F1286427D7531286C01E42" originalValue="Biológica">Biologica</normalizedToken>
la Hesperia, 1200 m (2 specimens CEMT); Mindo, 1500 m (1 specimen MQCAZ); Palmeras (1 specimen MQCAZ); Pampas Argentinas (1 specimen MQCAZ). SANTO DOMINGO DE LOS
<normalizedToken id="14B2FEA231B98B98945209883491BEE2" originalValue="TSÁCHILAS">TSACHILAS</normalizedToken>
:
<normalizedToken id="93C79803157388F9D2D87EDE2824D3FD" originalValue="Río">Rio</normalizedToken>
Toachi (1 specimen MQCAZ); Santo Domingo (2 specimens CEMT). UNDETERMINED PROVINCE: Los Lanos [= Los Llanos] (7 specimens SMTD).
</paragraph>
</subSubSection>
<subSubSection id="C0E58B5D1069D408C39DDE4D0409A780" pageId="118" pageNumber="119" type="temporal data">
<paragraph id="14D2BE6941EFD3DF12AB21DAE963780B" pageId="118" pageNumber="119">
<pageBreakToken id="C5FB4840E6A60D1054342CB1A99F50A1" pageId="118" pageNumber="119" start="start">Temporal</pageBreakToken>
data.
</paragraph>
<paragraph id="53028B6A2418803922BEB6CA2C52F11C" pageId="118" pageNumber="119">Collected in all months except September.</paragraph>
</subSubSection>
<subSubSection id="3E01829B7DE590C11DFCBFB91822007B" pageId="118" pageNumber="119" type="remarks">
<paragraph id="EC661D93A67BCE59BD2DCA2AF4A3C49C" pageId="118" pageNumber="119">Remarks.</paragraph>
<paragraph id="7321D4732D5F6837C45C284FC84F8634" pageId="118" pageNumber="119">Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 200-1300 m a.s.l. In the Andean region, it was registered for the evergreen lower montane forests from 1500-1800 m a.s.l. Collected manually and with pitfall traps baited with human feces.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,508 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.770.25365" ID-GBIF-Dataset="5d9d73e2-c94a-416c-b943-c3ab14a22df0" ID-PMC="PMC6041354" ID-Pensoft-Pub="1313-2970-770-43" ID-Pensoft-UUID="FFA0FF981B4D3C194F12F960FFCFFF05" ID-PubMed="30002590" ID-Zenodo-Dep="1310213" ID-ZooBank="4D984FDD93664D8B8A8E9D4B3F9B8EFB" ModsDocID="1313-2970-770-43" checkinTime="1530781150282" checkinUser="pensoft" docAuthor="Neubauer, Thomas A., Velde, Sabrina van de, Yanina, Tamara &amp; Wesselingh, Frank P." docDate="2018" docId="7AFC7930ABE592BA469B558D44CA83D8" docLanguage="en" docName="ZooKeys 770: 43-103" docOrigin="ZooKeys 770" docPubDate="2018-07-04" docSource="http://dx.doi.org/10.3897/zookeys.770.25365" docTitle="Hydrobia W. Hartmann 1821" docType="treatment" docVersion="6" id="FFA0FF981B4D3C194F12F960FFCFFF05" lastPageNumber="43" masterDocId="FFA0FF981B4D3C194F12F960FFCFFF05" masterDocTitle="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy" masterLastPageNumber="103" masterPageNumber="43" pageNumber="43" updateTime="1668165966764" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubauer, Thomas A.</mods:namePart>
<mods:affiliation>Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Velde, Sabrina van de</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yanina, Tamara</mods:namePart>
<mods:affiliation>Moscow State University, Faculty of Geography, Leninskie Gory, 1, 119991 Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesselingh, Frank P.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-07-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>770</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>103</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.770.25365</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.770.25365</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-770-43</mods:identifier>
<mods:identifier type="ZooBank">4D984FDD93664D8B8A8E9D4B3F9B8EFB</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFA0FF981B4D3C194F12F960FFCFFF05</mods:identifier>
<mods:identifier type="Zenodo-Dep">1310213</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182229637" LSID="urn:lsid:plazi:treatment:7AFC7930ABE592BA469B558D44CA83D8" httpUri="http://treatment.plazi.org/id/7AFC7930ABE592BA469B558D44CA83D8" lastPageNumber="43" pageId="0" pageNumber="43">
<subSubSection pageId="0" pageNumber="43" type="nomenclature">
<paragraph pageId="0" pageNumber="43">
Subfamily
<taxonomicName authority="Brusina, 1882" authorityName="W. Hartmann" authorityYear="1821" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae Brusina, 1882</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="reference_group">
<paragraph pageId="0" pageNumber="43">
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
1882
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
Brusina: 230.
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
1914
<taxonomicName family="Micromelaniidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="family">Micromelaniidae</taxonomicName>
B. Dybowski &amp; Grochmalicki: 276.
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
1915
<taxonomicName authorityName="B.Dybowski &amp; Grochmalicki" authorityYear="1915" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Turricaspiinae">Turricaspiinae</taxonomicName>
B. Dybowski &amp; Grochmalicki: 103.
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
2017
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
Brusina, 1882. - Bouchet et al.: 212, 346 [cum syn.].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="discussion">
<paragraph pageId="0" pageNumber="43">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="43">
The Caspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
(sensu lato) encompasses 64 species that are considered accepted in the current literature (
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). However, most of them are poorly known, documented by insufficient descriptions and drawings; for many, the
<typeStatus>type</typeStatus>
material has not been found (
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). The extreme morphological variability of several representatives, such as those detected in the material from Selitrennoye, led previous authors to introduce numerous species based on shells with only minor deviations in shape, size or whorl outline. The Caspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
therefore requires careful revision using molecular and anatomical data as far as available.
</paragraph>
<paragraph pageId="0" pageNumber="43">
In addition to the problems associated with distinguishing species, genus-level classification is poorly resolved as well. Several attempts have been made to categorize this vast variability, and genus concepts have changed tremendously (e.g., B.
<bibRefCitation author="Dybowski, B" journalOrPublisher="Petersbourg" pageId="0" pageNumber="43" refId="B32" refString="Dybowski, B, Grochmalicki, J, 1915. Ueber kaspische Schnecken aus der Abteilung &quot;Turricaspiinae&quot; subfam. nova zum Vergleich mit den Turribaikalina nobis. Petrograd, 34 pp. [numbered 103-136], 3 pls." title="Ueber kaspische Schnecken aus der Abteilung &quot; Turricaspiinae &quot; subfam. nova zum Vergleich mit den Turribaikalina nobis. Petrograd, 34 pp. [numbered 103 - 136], 3 pls." year="1915">Dybowski and Grochmalicki 1915</bibRefCitation>
,
<bibRefCitation author="Dybowski, B" journalOrPublisher="Abhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="43" pagination="1 - 55" publicationUrl="https://biodiversitylibrary.org/page/5550074" refId="B33" refString="Dybowski, B, Grochmalicki, J, 1917. Studien ueber die turmfoermigen Schnecken des Baikalsees und des Kaspimeeres (Turribaicaliinae - Turricaspiinae). Abhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 9 (3): 1 - 55, https://biodiversitylibrary.org/page/5550074" title="Studien ueber die turmfoermigen Schnecken des Baikalsees und des Kaspimeeres (Turribaicaliinae - Turricaspiinae)." url="https://biodiversitylibrary.org/page/5550074" volume="9" year="1917">1917</bibRefCitation>
,
<bibRefCitation author="Zhadin, VI" journalOrPublisher="Izdatel'stvo Akademii Nauk SSSR, Moskva, Leningrad" pageId="0" pageNumber="43" refId="B136" refString="Zhadin, VI, 1952. Mollyuski presnykh i solonovatykh vod SSSR. Izdatel'stvo Akademii Nauk SSSR, Moskva, Leningrad" title="Mollyuski presnykh i solonovatykh vod SSSR." year="1952">Zhadin 1952</bibRefCitation>
,
<bibRefCitation author="Logvinenko, BM" editor="Birshtein, YaA" journalOrPublisher="Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva" pageId="0" pageNumber="43" pagination="308 - 385" refId="B70" refString="Logvinenko, BM, Starobogatov, YaI, 1969. Mollusca. In: Birshtein, YaA, Vinogradov, LG, Kondakov, NN, Kuhn, MS, Astakhova, TV, Romanova, NN, Eds., Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva: 308 - 385" title="Mollusca." volumeTitle="Atlas bespozvonochnykh Kaspiyskogo morya." year="1969">Logvinenko and Starobogatov 1969</bibRefCitation>
,
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). Twelve genus names have been described for members of the Caspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
, based on quite different concepts of traits considered diagnostic. Currently, all species are classified within
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
</taxonomicName>
Clessin &amp; W. Dybowski, 1887,
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
</taxonomicName>
De Cristofori &amp; Jan, 1832 and
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1915 (
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). This scheme unites quite a variety of different morphologies under the same genus names, while at the same time similar species are assigned to different genera (e.g.,
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
). Unfortunately, hardly any previous study provided explanations for their genus classifications or systematic concepts in general.
</paragraph>
<paragraph pageId="0" pageNumber="43">
A thorough revision of all Caspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
is beyond the scope of this study, but we discuss and revise the concepts that have been applied to the species studied herein.
</paragraph>
<paragraph pageId="0" pageNumber="43">
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor (2016)</bibRefCitation>
listed 38 species of the genus
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
</taxonomicName>
for the Caspian Sea. The
<typeStatus>type</typeStatus>
species of
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
</taxonomicName>
De Cristofori &amp; Jan, 1832,
<taxonomicName lsidName="P. annulata" pageId="0" pageNumber="43" rank="species" species="annulata">
<emphasis italics="true" pageId="0" pageNumber="43">P. annulata</emphasis>
</taxonomicName>
(Linnaeus, 1758), lives in freshwater lakes and springs in Italy and Dalmatia (
<bibRefCitation author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="43" refId="B124" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
). Shell morphology, anatomy and protoconch characteristics are very similar to Pontocaspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
, e.g., some species of
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
(compare also discussion in
<bibRefCitation DOI="https://doi.org/10.1016/S0075-9511(01)80031-1" author="Riedel, F" journalOrPublisher="Limnologica" pageId="0" pageNumber="43" pagination="289 - 302" refId="B93" refString="Riedel, F, Healy, JM, Roepsdorf, P, Sitnikova, TYa, 2001. Ecology, Shell Morphology, Anatomy and Sperm Ultrastructure of Caenogastropod Pyrgula annulata, with a Discussion of the Relations between the ' Pyrgulidae ' and Caspian and Baikalian Rissooideans. Limnologica 31: 289 - 302, DOI: https://doi.org/10.1016/S0075-9511(01)80031-1" title="Ecology, Shell Morphology, Anatomy and Sperm Ultrastructure of Caenogastropod Pyrgula annulata, with a Discussion of the Relations between the ' Pyrgulidae ' and Caspian and Baikalian Rissooideans." url="https://doi.org/10.1016/S0075-9511(01)80031-1" volume="31" year="2001">Riedel et al. 2001</bibRefCitation>
). However, molecular evidence suggests that
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula annulata" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="annulata">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula annulata</emphasis>
</taxonomicName>
is only distantly related to the Pontocaspian species flock within the
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
, with the last common ancestor dating back to the late Miocene (
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2699.2007.01727.x" author="Wilke, T" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="43" pagination="1807 - 1821" refId="B130" refString="Wilke, T, Albrecht, C, Anistratenko, VV, Sahin, SK, Yildirim, MZ, 2007. Testing biogeographical hypotheses in space and time: faunal relationships of the putative ancient Lake Egirdir in Asia Minor. Journal of Biogeography 34: 1807 - 1821, DOI: https://doi.org/10.1111/j.1365-2699.2007.01727.x" title="Testing biogeographical hypotheses in space and time: faunal relationships of the putative ancient Lake Egirdir in Asia Minor." url="https://doi.org/10.1111/j.1365-2699.2007.01727.x" volume="34" year="2007">Wilke et al. 2007</bibRefCitation>
). Therefore, Pontocaspian species should not be attributed to
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
</taxonomicName>
, despite apparent morphological congruence, especially of some of the keeled Pontocaspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
. A separation on subfamily level as proposed by B.
<bibRefCitation author="Dybowski, B" journalOrPublisher="Petersbourg" pageId="0" pageNumber="43" refId="B32" refString="Dybowski, B, Grochmalicki, J, 1915. Ueber kaspische Schnecken aus der Abteilung &quot;Turricaspiinae&quot; subfam. nova zum Vergleich mit den Turribaikalina nobis. Petrograd, 34 pp. [numbered 103-136], 3 pls." title="Ueber kaspische Schnecken aus der Abteilung &quot; Turricaspiinae &quot; subfam. nova zum Vergleich mit den Turribaikalina nobis. Petrograd, 34 pp. [numbered 103 - 136], 3 pls." year="1915">Dybowski and Grochmalicki (1915)</bibRefCitation>
is opposed by the latest phylogeny of rissooidean gastropods, which suggests a rather close relationship (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2012.10.025" author="Wilke, T" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="43" pagination="715 - 736" refId="B131" refString="Wilke, T, Haase, M, Hershler, R, Liu, H-P, Misof, B, Ponder, W, 2013. Pushing short DNA fragments to the limit: Phylogenetic relationships of 'hydrobioid' gastropods (Caenogastropoda: Rissooidea). Molecular Phylogenetics and Evolution 66: 715 - 736, DOI: https://doi.org/10.1016/j.ympev.2012.10.025" title="Pushing short DNA fragments to the limit: Phylogenetic relationships of ' hydrobioid' gastropods (Caenogastropoda: Rissooidea)." url="https://doi.org/10.1016/j.ympev.2012.10.025" volume="66" year="2013">Wilke et al. 2013</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1915 (
<typeStatus>type</typeStatus>
species:
<taxonomicName authorityName="Clessin &amp; W.Dybowski" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Micromelania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromelania turricula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="turricula">
<emphasis italics="true" pageId="0" pageNumber="43">Micromelania turricula</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1915) was introduced for species with turriform, elongate shells with numerous whorls. Presently, the genus includes 22 Caspian species, encompassing elongate and broad, conical and ovoid, and sculptured and smooth species (
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). Many species assigned to
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula</emphasis>
</taxonomicName>
by
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev (2006)</bibRefCitation>
and
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor (2016)</bibRefCitation>
actually resemble
<taxonomicName baseAuthorityName="Clessin &amp; W. Dybowski in W. Dybowski" baseAuthorityYear="1888" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia turricula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="turricula">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia turricula</emphasis>
</taxonomicName>
with respect to the turriform, conical shell. This similarity also regards the
<typeStatus>type</typeStatus>
species of the genera
<taxonomicName authorityName="Logvinenko &amp; Starobogatov" authorityYear="1968" class="Gastropoda" genus="Caspiopyrgula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Caspiopyrgula" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiopyrgula</emphasis>
</taxonomicName>
Logvinenko &amp; Starobogatov, 1969 (
<typeStatus>type</typeStatus>
species:
<taxonomicName authorityName="Kolesnikov" authorityYear="1947" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia nossovi" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="nossovi">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia nossovi</emphasis>
</taxonomicName>
Kolesnikov, 1947),
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Eurycaspia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eurycaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Eurycaspia</emphasis>
</taxonomicName>
Logvinenko &amp; Starobogatov, 1969 (
<taxonomicName authorityName="B.Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Micromelania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromelania pseudodimidiata" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="pseudodimidiata">
<emphasis italics="true" pageId="0" pageNumber="43">Micromelania pseudodimidiata</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1917),
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Oxypyrgula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oxypyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Oxypyrgula</emphasis>
</taxonomicName>
Logvinenko &amp; Starobogatov, 1969 (
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula pseudospica" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="pseudospica">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula pseudospica</emphasis>
</taxonomicName>
Logvinenko &amp; Starobogatov, 1969), and
<taxonomicName authorityName="B.Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Trachycaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachycaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Trachycaspia</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1917 (
<taxonomicName authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Hydrobiidae" genus="Rissoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoa dimidiata" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="dimidiata">
<emphasis italics="true" pageId="0" pageNumber="43">Rissoa dimidiata</emphasis>
</taxonomicName>
Eichwald, 1838). After examination of descriptions and illustrations of the
<typeStatus>type</typeStatus>
species (e.g.,
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
), we conclude that these genera should be considered as junior synonyms of
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="43">
Some of the species classified as
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
by
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev (2006)</bibRefCitation>
and
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor (2016)</bibRefCitation>
differ considerably from
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
s.s. in shell shape. This contains the
<typeStatus>type</typeStatus>
species of the genera
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
Thiele, 1928 (
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Rissoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoa conus" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="conus">
<emphasis italics="true" pageId="0" pageNumber="43">Rissoa conus</emphasis>
</taxonomicName>
Eichwald, 1838),
<taxonomicName authorityName="Lindholm" authorityYear="1924" class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Clessiniola</emphasis>
</taxonomicName>
Lindholm, 1924 (
<taxonomicName authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Hydrobiidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina variabilis" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="variabilis">
<emphasis italics="true" pageId="0" pageNumber="43">Paludina variabilis</emphasis>
</taxonomicName>
Eichwald, 1838), and
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
B. Dybowski &amp; Grochmalicki, 1917 (
<taxonomicName authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Hydrobiidae" genus="Rissoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoa caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="caspia">
<emphasis italics="true" pageId="0" pageNumber="43">Rissoa caspia</emphasis>
</taxonomicName>
Eichwald, 1838). In turn, some species presently attributed to the genus
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
Sitnikova &amp; Starobogatov, 1999 (
<typeStatus>type</typeStatus>
species:
<taxonomicName authorityName="Golikov &amp; Starobogatov" authorityYear="1966" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" lsidName="Pyrgula milachevitchi" order="Mesogastropoda" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="milachevitchi">
<emphasis italics="true" pageId="0" pageNumber="43">Pyrgula milachevitchi</emphasis>
</taxonomicName>
Golikov &amp; Starobogatov, 1966) closely resemble species of the
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
group (compare
<bibRefCitation author="Anistratenko, VV" journalOrPublisher="Naukova dumka, Kiev" pageId="0" pageNumber="43" refId="B9" refString="Anistratenko, VV, Khaliman, IA, Anistratenko, OY, 2011. Mollyuski Azovskogo morya. Naukova dumka, Kiev" title="Mollyuski Azovskogo morya." year="2011">Anistratenko et al. 2011</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="43">
Based on a review of the Pontocaspian species formerly attributed to these genera and illustrated in the literature (
<bibRefCitation author="Golikov, AN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="43" pagination="352 - 362" refId="B45" refString="Golikov, AN, Starobogatov, YaI, 1966. Ponto-kaspiyskiy bryukhonogiye mollyuski v Azovo-Chernomorskom basseyne. Zoologicheskiy Zhurnal 45 (3): 352 - 362" title="Ponto-kaspiyskiy bryukhonogiye mollyuski v Azovo-Chernomorskom basseyne." volume="45" year="1966">Golikov and Starobogatov 1966</bibRefCitation>
,
<bibRefCitation author="Logvinenko, BM" editor="Birshtein, YaA" journalOrPublisher="Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva" pageId="0" pageNumber="43" pagination="308 - 385" refId="B70" refString="Logvinenko, BM, Starobogatov, YaI, 1969. Mollusca. In: Birshtein, YaA, Vinogradov, LG, Kondakov, NN, Kuhn, MS, Astakhova, TV, Romanova, NN, Eds., Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva: 308 - 385" title="Mollusca." volumeTitle="Atlas bespozvonochnykh Kaspiyskogo morya." year="1969">Logvinenko and Starobogatov 1969</bibRefCitation>
,
<bibRefCitation author="Alexenko, TL" journalOrPublisher="Vestnik Zoologii" pageId="0" pageNumber="43" pagination="32 - 38" refId="B5" refString="Alexenko, TL, Starobogatov, YaI, 1987. Vidy Caspia i Turricaspia (Gastropoda, Pectinibranchia, Pyrgulidae) Azovo-Chernomorskogo basseyna. Vestnik Zoologii 21 (3): 32 - 38" title="Vidy Caspia i Turricaspia (Gastropoda, Pectinibranchia, Pyrgulidae) Azovo-Chernomorskogo basseyna." volume="21" year="1987">Alexenko and Starobogatov 1987</bibRefCitation>
,
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1002/zoos.200800001" author="Anistratenko, VV" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="43" pagination="129 - 142" refId="B7" refString="Anistratenko, VV, 2008. Evolutionary trends and relationships in hydrobiids (Mollusca, Caenogastropoda) of the Azov-Black Sea Basin in the light of their comparative morphology and paleozoogeography. Zoosystematics and Evolution 84 (2): 129 - 142, DOI: https://doi.org/10.1002/zoos.200800001" title="Evolutionary trends and relationships in hydrobiids (Mollusca, Caenogastropoda) of the Azov-Black Sea Basin in the light of their comparative morphology and paleozoogeography." url="https://doi.org/10.1002/zoos.200800001" volume="84" year="2008">Anistratenko 2008</bibRefCitation>
), we propose to distinguish the genera
<taxonomicName authorityName="Lindholm" authorityYear="1924" class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Clessiniola</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
from
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
, and to treat
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
as junior synonyms of
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName authorityName="Lindholm" authorityYear="1924" class="Gastropoda" family="Hydrobiidae" genus="Clessiniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clessiniola" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Clessiniola</emphasis>
</taxonomicName>
species can be easily distinguished from species attributed to other genera based on their broad shells with a large body whorl and aperture. The situation for the
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
is more difficult. The three
<typeStatus>type</typeStatus>
species (see above) share the ovoid shape with cyrtoconoid spire, the high whorl accretion rate, the shape, inclination, lateral sinuation and thickening of the aperture, and the extent and sculpture of the protoconch (e.g.,
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1002/zoos.200800001" author="Anistratenko, VV" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="43" pagination="129 - 142" refId="B7" refString="Anistratenko, VV, 2008. Evolutionary trends and relationships in hydrobiids (Mollusca, Caenogastropoda) of the Azov-Black Sea Basin in the light of their comparative morphology and paleozoogeography. Zoosystematics and Evolution 84 (2): 129 - 142, DOI: https://doi.org/10.1002/zoos.200800001" title="Evolutionary trends and relationships in hydrobiids (Mollusca, Caenogastropoda) of the Azov-Black Sea Basin in the light of their comparative morphology and paleozoogeography." url="https://doi.org/10.1002/zoos.200800001" volume="84" year="2008">Anistratenko 2008</bibRefCitation>
, and this study). The only differences are shell size and whorl convexity, which we do not consider sufficient to distinguish genera. The adapical thickening of the aperture resulting from downward growing of the shell in late ontogeny as stated in the diagnosis of the genus
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
by
<bibRefCitation author="Sitnikova, TYa" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="43" pagination="158 - 163" refId="B98" refString="Sitnikova, TYa, Starobogatov, YaI, 1999. Novyye rod semeystva Pyrgulidae (Gastropoda, Pectinibranchia) iz presnykh rod Azovo-Chernomorskogo basseyna (v svyazi s voprosom o Ponto-Kaspiyskikh vidakh v Azovo-Chernomorskogom basseyne). Zoologicheskiy Zhurnal 78 (2): 158 - 163" title="Novyye rod semeystva Pyrgulidae (Gastropoda, Pectinibranchia) iz presnykh rod Azovo-Chernomorskogo basseyna (v svyazi s voprosom o Ponto-Kaspiyskikh vidakh v Azovo-Chernomorskogom basseyne)." volume="78" year="1999">Sitnikova and Starobogatov (1999)</bibRefCitation>
is also shown for species that have been attributed to
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
(see below). The features of the soft-part anatomy considered diagnostic by these authors need to be rechecked and compared to live material from the Caspian Sea to reevaluate the position of
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
.
<bibRefCitation author="Sitnikova, TYa" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="43" pagination="158 - 163" refId="B98" refString="Sitnikova, TYa, Starobogatov, YaI, 1999. Novyye rod semeystva Pyrgulidae (Gastropoda, Pectinibranchia) iz presnykh rod Azovo-Chernomorskogo basseyna (v svyazi s voprosom o Ponto-Kaspiyskikh vidakh v Azovo-Chernomorskogom basseyne). Zoologicheskiy Zhurnal 78 (2): 158 - 163" title="Novyye rod semeystva Pyrgulidae (Gastropoda, Pectinibranchia) iz presnykh rod Azovo-Chernomorskogo basseyna (v svyazi s voprosom o Ponto-Kaspiyskikh vidakh v Azovo-Chernomorskogom basseyne)." volume="78" year="1999">Sitnikova and Starobogatov (1999)</bibRefCitation>
also discussed the similarities between
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
, concluding that
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
should perhaps be included in the genus
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
, possibly as a separate subgenus (which would be nomenclaturally invalid however).
</paragraph>
<paragraph pageId="0" pageNumber="43">
The ovoid shape, lateral sinuation and thickening of the aperture typical for the
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Malacostraca" family="Hydrobiidae" genus="Caspiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspiella" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspiella</emphasis>
</taxonomicName>
-
<taxonomicName authorityName="Sitnikova &amp; Starobogatov" authorityYear="1999" class="Gastropoda" family="Hydrobiidae" genus="Euxinipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euxinipyrgula" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Euxinipyrgula</emphasis>
</taxonomicName>
group are also found among species of the genus
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Prososthenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Prososthenia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Prososthenia</emphasis>
</taxonomicName>
Neumayr, 1969 from the middle Miocene of the Dinaride Lake System (e.g.,
<bibRefCitation DOI="https://doi.org/10.17738/ajes.2015.0013" author="Neubauer, TA" journalOrPublisher="Austrian Journal of Earth Sciences" pageId="0" pageNumber="43" pagination="15 - 67" refId="B78" refString="Neubauer, TA, Mandic, O, Harzhauser, M, 2016b. The freshwater mollusk fauna of the Middle Miocene Lake Drnis (Dinaride Lake System, Croatia): a taxonomic and systematic revision. Austrian Journal of Earth Sciences 108 (2): 15 - 67, DOI: https://doi.org/10.17738/ajes.2015.0013" title="The freshwater mollusk fauna of the Middle Miocene Lake Drnis (Dinaride Lake System, Croatia): a taxonomic and systematic revision." url="https://doi.org/10.17738/ajes.2015.0013" volume="108" year="2016 b">Neubauer et al. 2016b</bibRefCitation>
). These species, however, differ in the granulate protoconch making up less than one whorl.
</paragraph>
<paragraph pageId="0" pageNumber="43">
Species of
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
differ from
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1917" class="Gastropoda" family="Hydrobiidae" genus="Laevicaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laevicaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Laevicaspia</emphasis>
</taxonomicName>
in the slower, regular whorl accretion, producing a conical spire and a higher number of whorls at the same size. In addition,
<taxonomicName authorityName="B. Dybowski &amp; Grochmalicki" authorityYear="1915" class="Gastropoda" family="Hydrobiidae" genus="Turricaspia" kingdom="Animalia" lsidName="Turricaspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Turricaspia</emphasis>
</taxonomicName>
species have usually more fragile shells, thinner peristomes and often more strongly sinuate growth lines.
</paragraph>
<paragraph pageId="0" pageNumber="43">
The genus
<taxonomicName authorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" authorityYear="2018" baseAuthorityName="Neubauer &amp; Velde &amp; Yanina &amp; Wesselingh" baseAuthorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Caspia</emphasis>
</taxonomicName>
is listed among
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
in latest catalogues (
<bibRefCitation author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="43" refId="B58" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
,
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="43" refId="B121" refString="Vinarski, MV, Kantor, YuI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
), but it has been shown to be unrelated to that subfamily (
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2012.742934" author="Anistratenko, VV" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="43" pagination="51 - 64" refId="B8" refString="Anistratenko, VV, 2013. On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae). Journal of Natural History 47 (1-2): 51 - 64, DOI: https://doi.org/10.1080/00222933.2012.742934" title="On the taxonomic status of the highly endangered Ponto-Caspian gastropod genus Caspia (Gastropoda: Hydrobiidae: Caspiinae)." url="https://doi.org/10.1080/00222933.2012.742934" volume="47" year="2013">Anistratenko 2013</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.4002/040.061.0201" author="Bouchet, P" journalOrPublisher="Malacologia" pageId="0" pageNumber="43" pagination="1 - 526" refId="B15" refString="Bouchet, P, Rocroi, J-P, Hausdorf, B, Kaim, A, Kano, Y, Nuetzel, A, Parkhaev, P, Schroedl, M, Strong, EE, 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1-2): 1 - 526, DOI: https://doi.org/10.4002/040.061.0201" title="Revised classification, nomenclator and typification of gastropod and monoplacophoran families." url="https://doi.org/10.4002/040.061.0201" volume="61" year="2017">Bouchet et al. 2017</bibRefCitation>
; see discussion of the
<taxonomicName authorityName="B.Dybowski" authorityYear="1913" lsidName="" pageId="0" pageNumber="43" rank="subFamily" subFamily="Caspiinae">Caspiinae</taxonomicName>
above).
</paragraph>
<paragraph pageId="0" pageNumber="43">
Finally, several Pontocaspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
have been previously assigned to the genus
<taxonomicName authorityName="Brusina" authorityYear="1874" class="Gastropoda" family="Hydrobiidae" genus="Micromelania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromelania" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Micromelania</emphasis>
</taxonomicName>
Brusina, 1874 (e.g., W. Dybowski 1887, B.
<bibRefCitation author="Dybowski, B" journalOrPublisher="Abhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="43" pagination="1 - 55" publicationUrl="https://biodiversitylibrary.org/page/5550074" refId="B33" refString="Dybowski, B, Grochmalicki, J, 1917. Studien ueber die turmfoermigen Schnecken des Baikalsees und des Kaspimeeres (Turribaicaliinae - Turricaspiinae). Abhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 9 (3): 1 - 55, https://biodiversitylibrary.org/page/5550074" title="Studien ueber die turmfoermigen Schnecken des Baikalsees und des Kaspimeeres (Turribaicaliinae - Turricaspiinae)." url="https://biodiversitylibrary.org/page/5550074" volume="9" year="1917">Dybowski and Grochmalicki 1917</bibRefCitation>
). Its
<typeStatus>type</typeStatus>
species,
<taxonomicName authorityName="Brusina" authorityYear="1874" class="Gastropoda" family="Hydrobiidae" genus="Micromelania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micromelania cerithiopsis" order="Littorinimorpha" pageId="0" pageNumber="43" phylum="Mollusca" rank="species" species="cerithiopsis">
<emphasis italics="true" pageId="0" pageNumber="43">Micromelania cerithiopsis</emphasis>
</taxonomicName>
Brusina, 1874 (subsequent designation by
<bibRefCitation author="Dollfus, GF" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="43" pagination="179 - 270" publicationUrl="https://biodiversitylibrary.org/page/16299461" refId="B27" refString="Dollfus, GF, 1912. Recherches critiques sur quelques genre et especes d' Hydrobia vivants ou fossiles. Journal de Conchyliologie 59 (3): 179 - 270, https://biodiversitylibrary.org/page/16299461" title="Recherches critiques sur quelques genre et especes d' Hydrobia vivants ou fossiles." url="https://biodiversitylibrary.org/page/16299461" volume="59" year="1912">Dollfus 1912</bibRefCitation>
), derives from late Miocene deposits of Lake Pannon. It differs considerably from Pontocaspian
<taxonomicName baseAuthorityName="Brusina" baseAuthorityYear="1881" family="Hydrobiidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="43" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
regarding the presence of 2-4 noded keels and the small size (4.5
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.33" unit="mm" value="133.0">1.33 mm</quantity>
after
<bibRefCitation author="Brusina, S" journalOrPublisher="Actienbuchdruckerei, Agram" pageId="0" pageNumber="43" refId="B18" refString="Brusina, S, 1874. Fossile Binnen-Mollusken aus Dalmatien, Kroatien und Slavonien nebst einem Anhange. Actienbuchdruckerei, Agram" title="Fossile Binnen-Mollusken aus Dalmatien, Kroatien und Slavonien nebst einem Anhange." year="1874">Brusina 1874</bibRefCitation>
) compared to the rather high number of eight whorls.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,328 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.975.55456" ID-GBIF-Dataset="bfcb4e07-aaea-419e-8bb5-08851e9f27dd" ID-PMC="PMC7572521" ID-Pensoft-Pub="1313-2970-975-11" ID-Pensoft-UUID="FE5DCB656614590DB4C35092D89F3F3D" ID-PubMed="33117063" ID-ZooBank="785EDF225A3242178C9B6433E0AD199F" ModsDocID="1313-2970-975-11" checkinTime="1602606874639" checkinUser="pensoft" docAuthor="Hendrich, Lars, Watts, Chris H. S. &amp; Balke, Michael" docDate="2020" docId="7AFC883E9CD1562F8296A38ABA622302" docLanguage="en" docName="ZooKeys 975: 11-49" docOrigin="ZooKeys 975" docSource="http://dx.doi.org/10.3897/zookeys.975.55456" docTitle="Gibbidessus atomus Hendrich &amp; Watts &amp; Balke 2020, sp. nov." docType="treatment" docUuid="713DD48F-BA41-4581-A05F-DEECAED51519" docUuidSource="ZooBank" docVersion="4" id="FE5DCB656614590DB4C35092D89F3F3D" lastPageNumber="11" masterDocId="FE5DCB656614590DB4C35092D89F3F3D" masterDocTitle="The &quot; minute diving beetles &quot; of southern Australia - taxonomic revision of Gibbidessus Watts, 1978, with description of six new species (Coleoptera, Dytiscidae, Bidessini)" masterLastPageNumber="49" masterPageNumber="11" pageNumber="11" updateTime="1668169601453" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The &quot; minute diving beetles &quot; of southern Australia - taxonomic revision of Gibbidessus Watts, 1978, with description of six new species (Coleoptera, Dytiscidae, Bidessini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>SNSB - Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D - 81247 Muenchen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8366-0749</mods:nameIdentifier>
<mods:nameIdentifier type="email">hendrich1@aol.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watts, Chris H. S.</mods:namePart>
<mods:affiliation>South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:affiliation>SNSB - Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D - 81247 Muenchen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>975</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.975.55456</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.975.55456</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-975-11</mods:identifier>
<mods:identifier type="ZooBank">785EDF225A3242178C9B6433E0AD199F</mods:identifier>
<mods:identifier type="Pensoft-UUID">FE5DCB656614590DB4C35092D89F3F3D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168338752" LSID="urn:lsid:zoobank.org:act:713DD48F-BA41-4581-A05F-DEECAED51519" httpUri="http://treatment.plazi.org/id/7AFC883E9CD1562F8296A38ABA622302" lastPageNumber="11" pageId="0" pageNumber="11">
<subSubSection pageId="0" pageNumber="11" type="nomenclature">
<paragraph pageId="0" pageNumber="11">
<taxonomicName LSID="7AFC883E-9CD1-562F-8296-A38ABA622302" authority="Hendrich &amp; Watts &amp; Balke, 2020" authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus atomus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="atomus" status="sp. nov.">Gibbidessus atomus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="11">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">Figs 1</figureCitation>
<figureCitation captionStart="Figures 14, 15" captionStartId="F6" captionText="Figures 14, 15. 14 Gibbidessus atomus sp. nov. 15 G. chipi A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures14-15" httpUri="https://binary.pensoft.net/fig/462665" pageId="0" pageNumber="11">, 14</figureCitation>
<figureCitation captionStart="Figures 22, 23" captionStartId="F10" captionText="Figures 22, 23. Distribution of 22 Gibbidessus chipi (black dots), G. drikdrikensis sp. nov. (yellow square), G. kangarooensis sp. nov. (red square) 23 G. atomus sp. nov. (red square), G. davidi sp. nov. (yellow square)." figureDoi="10.3897/zookeys.975.55456.figures22-23" httpUri="https://binary.pensoft.net/fig/462669" pageId="0" pageNumber="11">, 23</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F13" captionText="Figure 27. Habitat of Gibbidessus atomus sp. nov. and Gibbidessus davidi sp. nov. A Seasonally flooded Melaleuca sedge swamp B small and shallow heathland pool along Windy Harbour Road, south of Northcliffe, south-western Australia." figureDoi="10.3897/zookeys.975.55456.figure27" httpUri="https://binary.pensoft.net/fig/462672" pageId="0" pageNumber="11">, 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type locality">
<paragraph pageId="0" pageNumber="11">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="11">
Western Australia, Windy Harbour Road, 11 km south of Northcliffe, small pool in sedge swamp [
<geoCoordinate degrees="34" direction="south" minutes="48" orientation="latitude" precision="15" seconds="51" value="-34.814167">34°48'51S</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="4" orientation="longitude" precision="15" seconds="8" value="116.068886">116°4'8E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type material">
<paragraph pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
,
</emphasis>
male: &quot;Australia: SW WA,
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
NP, 11 km S, Northcliffe, 77m, 4.I.2007, 34.44.048S, 116.05.354E [
<geoCoordinate degrees="34" direction="south" minutes="44" orientation="latitude" precision="15" seconds="0" value="-34.733334">34°44'0S</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="5" orientation="longitude" precision="15" seconds="13" value="116.086945">116°5'13E</geoCoordinate>
], L. &amp; E. Hendrich (WA 162)&quot;, &quot;Holotype
<taxonomicName authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus atomus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus atomus</emphasis>
</taxonomicName>
Hendrich, Watts &amp; Balke des. 2020&quot; [red printed label] (WAM).
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
(20 exs.)
</emphasis>
. 7 specimens with same data as holotype, three specimens with &quot;DNA M. Balke 1729&quot;, &quot;DNA M. Balke 2780&quot;, &quot;DNA M. Balke 2781&quot; [green printed labels]; 13 exs., &quot;AUSTRALIA/WA:
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
N.P., 11 km south of Northcliffe, Windy Harbour Road, 50 m, 3.1.2000,
<geoCoordinate degrees="34" direction="south" minutes="42" orientation="latitude" precision="925" value="-34.7">34°42'S</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="05" orientation="longitude" precision="925" value="116.083336">116°05'E</geoCoordinate>
[
<geoCoordinate degrees="34" direction="south" minutes="44" orientation="latitude" precision="15" seconds="0" value="-34.733334">34°44'0S</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="5" orientation="longitude" precision="15" seconds="13" value="116.086945">116°5'13E</geoCoordinate>
], Hendrich leg. (loc. WA 10c/156)&quot; (ANIC, CLH, SAMA, ZSM). All paratypes with red printed paratype labels.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="materials_examined">
<paragraph pageId="0" pageNumber="11">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="11">
1 ex., &quot;Australia, WA, RVDLE03 Riverdale Wetland [
<geoCoordinate degrees="32" direction="south" minutes="59" orientation="latitude" precision="15" seconds="22" value="-32.989445">32°59'22S</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="47" orientation="longitude" precision="15" seconds="7" value="115.78528">115°47'7E</geoCoordinate>
], 23/09/2008, South West Catchment Council Mon.&quot; (DPAW).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="diagnosis">
<paragraph pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="11">
Very small species, externally characterised by widely rounded body, with less pronounced habitus disruption between pronotum and elytron, shiny non-microreticulate dorsal surface and vague ferruginous markings on elytra. Dorsoventrally rather domed. Cervical line present (Fig.
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">1</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11" start="Figures 14" startId="F1">
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">Figures 1-4.</emphasis>
Habitus of
<emphasis bold="true" pageId="0" pageNumber="11">1</emphasis>
<taxonomicName authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus atomus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus atomus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="11">2</emphasis>
<taxonomicName lsidName="G. chipi" pageId="0" pageNumber="11" rank="species" species="chipi">
<emphasis italics="true" pageId="0" pageNumber="11">G. chipi</emphasis>
</taxonomicName>
,
<emphasis bold="true" pageId="0" pageNumber="11">3</emphasis>
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="11">4</emphasis>
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
female. Scale bar: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="description">
<paragraph pageId="0" pageNumber="11">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="11">Holotype: TL = 1.15 mm, TL-H = 1.05 mm, MW = 0.6 mm. Paratypes: TL = 1.15-1.30 mm, TL-H = 1.0-1.05 mm, MW = 0.6-0.65 mm.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Head</emphasis>
:
</emphasis>
Dark brown, around eyes almost black. Cervical line present. Strongly and coarsely punctate, rather shiny, microsculpture almost absent. Punctures weakly distributed anteriorly, strong posteriorly between eyes. Antennae relatively short, stout. Antennomeres ferruginous, darkened anteriorly.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Pronotum</emphasis>
:
</emphasis>
Ferruginous, anterior and posterior margins darker. Disc of pronotum somewhat darkened, broadest at posterior corners. Punctation very weakly, almost evenly distributed, shiny, microsculpture absent. Sides of pronotum margined and almost evenly rounded. Angle between pronotum and elytra less pronounced, basal pronotal plicae present. Striae moderately defined, on almost 1/2 length of pronotum, moderately incurved.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Elytra</emphasis>
:
</emphasis>
Dark brown with vague basal area ferruginous (Fig.
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">1</figureCitation>
). Coarsely and densely punctate, shiny, microsculpture absent. Striae weakly impressed, slightly straighter but shorter than basal pronotal striae.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Ventral side</emphasis>
:
</emphasis>
Ferruginous. Prothorax and apex of abdomen paler than other parts. Metacoxae and metaventrite covered with numerous larger punctures, surface shiny, without microreticulation. Abdominal ventrites with dense and finer punctures, shiny, microreticulation absent. Metacoxal lines almost straight, anteriorly slightly divergent. Epipleuron ferruginous, coarsely punctate, shiny, lacking microsculpture. Legs ferruginous with meta-/mesotibiae and meta-/mesotarsi set in black.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Male</emphasis>
.
</emphasis>
Dorsal surface with coarse punctures but otherwise with shiny surface (Fig.
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">1</figureCitation>
). Median lobe of aedeagus as in Fig.
<figureCitation captionStart="Figures 14, 15" captionStartId="F6" captionText="Figures 14, 15. 14 Gibbidessus atomus sp. nov. 15 G. chipi A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures14-15" httpUri="https://binary.pensoft.net/fig/462665" pageId="0" pageNumber="11">14A, B</figureCitation>
. Shape of median lobe fairly uniform, bent evenly, apex straight and pointed. Paramere, as in
<taxonomicName authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Limbodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limbodessus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Limbodessus</emphasis>
</taxonomicName>
Guignot, 1939, with hook or bent finger-like apical part with tiny setae on tip (Fig.
<figureCitation captionStart="Figures 14, 15" captionStartId="F6" captionText="Figures 14, 15. 14 Gibbidessus atomus sp. nov. 15 G. chipi A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures14-15" httpUri="https://binary.pensoft.net/fig/462665" pageId="0" pageNumber="11">14C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="affinities">
<paragraph pageId="0" pageNumber="11">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="11">
This species is similar to
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
sp. nov. but readily separated by its smaller size, the different colour pattern and the form of the median lobe and parameres (Figs
<figureCitation captionStart="Figures 14, 15" captionStartId="F6" captionText="Figures 14, 15. 14 Gibbidessus atomus sp. nov. 15 G. chipi A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures14-15" httpUri="https://binary.pensoft.net/fig/462665" pageId="0" pageNumber="11">14</figureCitation>
,
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="etymology">
<paragraph pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="11">
From Latin
<taxonomicName lsidName="atomus" pageId="0" pageNumber="11" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">atomus</emphasis>
</taxonomicName>
(smallest particle), as it is the smallest epigean diving beetle in Australia described so far.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="distribution">
<paragraph pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="11">
South-western Australia. Known only from the type locality in the
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
National Park, south of Northcliffe and the Riverdale Wetland [
<geoCoordinate degrees="32" direction="south" minutes="59" orientation="latitude" precision="15" seconds="22" value="-32.989445">32°59'22S</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="47" orientation="longitude" precision="15" seconds="7" value="115.78528">115°47'7E</geoCoordinate>
] (Fig.
<figureCitation captionStart="Figures 22, 23" captionStartId="F10" captionText="Figures 22, 23. Distribution of 22 Gibbidessus chipi (black dots), G. drikdrikensis sp. nov. (yellow square), G. kangarooensis sp. nov. (red square) 23 G. atomus sp. nov. (red square), G. davidi sp. nov. (yellow square)." figureDoi="10.3897/zookeys.975.55456.figures22-23" httpUri="https://binary.pensoft.net/fig/462669" pageId="0" pageNumber="11">23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="habitat">
<paragraph pageId="0" pageNumber="11">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="11">
Most specimens were obtained from an exposed, shallow and small roundish puddle without any vegetation, except some algae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F13" captionText="Figure 27. Habitat of Gibbidessus atomus sp. nov. and Gibbidessus davidi sp. nov. A Seasonally flooded Melaleuca sedge swamp B small and shallow heathland pool along Windy Harbour Road, south of Northcliffe, south-western Australia." figureDoi="10.3897/zookeys.975.55456.figure27" httpUri="https://binary.pensoft.net/fig/462672" pageId="0" pageNumber="11">27B</figureCitation>
). The remaining specimens were collected in a half-shaded pool in a
<taxonomicName class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melaleuca" order="Agaricales" pageId="0" pageNumber="11" phylum="Basidiomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Melaleuca</emphasis>
</taxonomicName>
blackwater swamp (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F13" captionText="Figure 27. Habitat of Gibbidessus atomus sp. nov. and Gibbidessus davidi sp. nov. A Seasonally flooded Melaleuca sedge swamp B small and shallow heathland pool along Windy Harbour Road, south of Northcliffe, south-western Australia." figureDoi="10.3897/zookeys.975.55456.figure27" httpUri="https://binary.pensoft.net/fig/462672" pageId="0" pageNumber="11">27A</figureCitation>
), with few clumps of
<taxonomicName class="Liliopsida" family="Juncaceae" genus="Juncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juncus" order="Poales" pageId="0" pageNumber="11" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Juncus</emphasis>
</taxonomicName>
spp. and extensive beds of macrophytes; depth up to 20 cm; bottom of sedge-filled peat (pH 5.5), twigs and rotten leaves. The heathlands south of Northcliffe are seasonally flooded, with some permanent water bodies in the summer. At Northcliffe the species is syntopic with
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
sp. nov. (
<bibRefCitation author="Hendrich, L" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="11" pagination="299 - 308" refId="B6" refString="Hendrich, L, 2001a. A new species of Antiporus Sharp, 1882 from peatland swamps of south-western Australia (Coleoptera: Dytiscidae). Linzer biologische Beitraege 33 (1): 299 - 308" title="A new species of Antiporus Sharp, 1882 from peatland swamps of south-western Australia (Coleoptera: Dytiscidae)." volume="33" year="2001 a">Hendrich 2001a</bibRefCitation>
), at the Riverdale Wetland it was collected with
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="G. rottnestensis" pageId="0" pageNumber="11" rank="species" species="rottnestensis">
<emphasis italics="true" pageId="0" pageNumber="11">G. rottnestensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph pageId="0" pageNumber="11">
Apart from the
<taxonomicName authorityName="Watts" authorityYear="1978" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus</emphasis>
</taxonomicName>
, the water beetle coenosis at Northcliffe included the following species:
<taxonomicName family="Dytiscidae" lsidName="" pageId="0" pageNumber="11" rank="family">Dytiscidae</taxonomicName>
:
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Limbodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limbodessus inornatus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis italics="true" pageId="0" pageNumber="11">Limbodessus inornatus</emphasis>
</taxonomicName>
(Sharp, 1882),
<taxonomicName authorityName="Watts" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Antiporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antiporus hollingsworthi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="hollingsworthi">
<emphasis italics="true" pageId="0" pageNumber="11">Antiporus hollingsworthi</emphasis>
</taxonomicName>
Watts, 1997,
<taxonomicName lsidName="A. mcraeae" pageId="0" pageNumber="11" rank="species" species="mcraeae">
<emphasis italics="true" pageId="0" pageNumber="11">A. mcraeae</emphasis>
</taxonomicName>
Watts &amp; Pinder, 2000,
<taxonomicName class="Insecta" family="Dytiscidae" genus="Brancuporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brancuporus gottwaldi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="gottwaldi">
<emphasis italics="true" pageId="0" pageNumber="11">Brancuporus gottwaldi</emphasis>
</taxonomicName>
(Hendrich, 2001),
<taxonomicName authorityName="Lea" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sternopriscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sternopriscus minimus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="minimus">
<emphasis italics="true" pageId="0" pageNumber="11">Sternopriscus minimus</emphasis>
</taxonomicName>
Lea, 1899,
<taxonomicName lsidName="S. eikei" pageId="0" pageNumber="11" rank="species" species="eikei">
<emphasis italics="true" pageId="0" pageNumber="11">S. eikei</emphasis>
</taxonomicName>
Hendrich &amp; Watts, 2007,
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ater" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="11">Exocelina ater</emphasis>
</taxonomicName>
(Sharp, 1882);
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Hydrophilidae" lsidName="" pageId="0" pageNumber="11" rank="family">Hydrophilidae</taxonomicName>
:
<taxonomicName baseAuthorityName="Blackburn" baseAuthorityYear="1895" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus eyrensis" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="eyrensis">
<emphasis italics="true" pageId="0" pageNumber="11">Enochrus eyrensis</emphasis>
</taxonomicName>
(Blackburn, 1895),
<taxonomicName baseAuthorityName="Broun" baseAuthorityYear="1880" class="Insecta" family="Hydrophilidae" genus="Limnoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnoxenus zealandicus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="zealandicus">
<emphasis italics="true" pageId="0" pageNumber="11">Limnoxenus zealandicus</emphasis>
</taxonomicName>
(Broun, 1880),
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Paracymus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracymus pygmaeus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis italics="true" pageId="0" pageNumber="11">Paracymus pygmaeus</emphasis>
</taxonomicName>
(Macleay, 1871).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,277 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.50.4873" ID-PMC="PMC4489085" ID-Pensoft-Pub="1314-2003-50-61" ID-Pensoft-UUID="FFB1FFBCFF9CFFDDE37D7217FF84FF94" ID-PubMed="26140021" ID-Zenodo-Dep="576278" ModsDocID="1314-2003-50-61" checkinTime="1451251517643" checkinUser="pensoft" docAuthor="Valdespino, Ivan A., Heringer, Gustavo, Salino, Alexandre, Goes-Neto, Luiz A. de Araujo &amp; Ceballos, Jorge" docDate="2015" docId="7AFCA259C6B99CC1F2F948F377232129" docLanguage="en" docName="PhytoKeys 50: 61-99" docOrigin="PhytoKeys 50" docPubDate="2015-06-16" docSource="http://dx.doi.org/10.3897/phytokeys.50.4873" docTitle="Selaginella mucugensis Valdespino 2015, sp. nov." docType="treatment" docVersion="6" id="FFB1FFBCFF9CFFDDE37D7217FF84FF94" lastPageNumber="73" masterDocId="FFB1FFBCFF9CFFDDE37D7217FF84FF94" masterDocTitle="Seven new species of Selaginella subg. Stachygynandrum (Selaginellaceae) from Brazil and new synonyms for the genus" masterLastPageNumber="99" masterPageNumber="61" pageNumber="72" updateTime="1668141371619" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of Selaginella subg. Stachygynandrum (Selaginellaceae) from Brazil and new synonyms for the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Valdespino, Ivan A.</mods:namePart>
<mods:affiliation>Departamento de Botanica, Facultad de Ciencias Naturales, Exactas y Tecnologia, Universidad de Panama, Apartado Postal 0824 - 00073, Panama</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heringer, Gustavo</mods:namePart>
<mods:affiliation>Pos-Graduacao em Botanica, Departamento de Biologia Vegetal, Universidade Federal de Vicosa, CEP 36.570 - 000 Vicosa, Minas Gerais, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salino, Alexandre</mods:namePart>
<mods:affiliation>Departamento de Botanica, Instituto de Ciencias Biologicas, Universidade Federal de Minas Gerais, CP 486, 31270 - 901, Belo Horizonte, MG, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goes-Neto, Luiz A. de Araujo</mods:namePart>
<mods:affiliation>Departamento de Botanica, Instituto de Ciencias Biologicas, Universidade Federal de Minas Gerais, CP 486, 31270 - 901, Belo Horizonte, MG, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ceballos, Jorge</mods:namePart>
<mods:affiliation>Smithsonian Tropical Research Institute, Apartado Postal 0843 - 03092, Panama</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>50</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>99</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.50.4873</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.50.4873</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-50-61</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFB1FFBCFF9CFFDDE37D7217FF84FF94</mods:identifier>
<mods:identifier type="Zenodo-Dep">576278</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152025795" LSID="urn:lsid:plazi:treatment:7AFCA259C6B99CC1F2F948F377232129" httpUri="http://treatment.plazi.org/id/7AFCA259C6B99CC1F2F948F377232129" lastPageId="12" lastPageNumber="73" pageId="11" pageNumber="72">
<subSubSection pageId="11" pageNumber="72" type="nomenclature">
<paragraph pageId="11" pageNumber="72">
<taxonomicName LSID="7AFCA259-C6B9-9CC1-F2F9-48F377232129" authority="Valdespino" authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="11" pageNumber="72" phylum="Lycopodiophyta" rank="species" species="mucugensis" status="sp. nov.">Selaginella mucugensis Valdespino</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="72">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Selaginella mucugensis Valdespino. A Section of upper surface of stem B Upper surface of median leaf C Section of lower surface of stem D Lower surface of lateral leaf A-D taken from holotype, Giulietti et al. [CFCR 1430] (NY)." figureDoi="10.3897/phytokeys.50.4873.figure10" httpUri="https://binary.pensoft.net/fig/45222" pageId="11" pageNumber="72">Figures 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Selaginella mucugensis Valdespino. A Microspore proximal face B Close-up of microspore proximal face surface C Microspore distal face D Close-up of microspore distal face surface A-D taken from holotype, Giulietti et al. [CFCR 1430] (NY)." figureDoi="10.3897/phytokeys.50.4873.figure11" httpUri="https://binary.pensoft.net/fig/45223" pageId="11" pageNumber="72">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="72" type="diagnosis">
<paragraph pageId="11" pageNumber="72">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="72">
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="11" pageNumber="72" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="11" pageNumber="72">Selaginella mucugensis</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella blepharodella" order="Selaginellales" pageId="11" pageNumber="72" phylum="Lycopodiophyta" rank="species" species="blepharodella">
<emphasis italics="true" pageId="11" pageNumber="72">Selaginella blepharodella</emphasis>
</taxonomicName>
in having median leaves distant (vs. imbricate), ovate (vs. broadly-ovate to ovate-elliptic), with margins hyaline in a band 2-5 (vs. 5-15) cells wide with cilia 30-50 (vs. 130-180)
<normalizedToken originalValue="µm">µm</normalizedToken>
long, stomata on upper surfaces on submarginal and marginal regions of the outer bases (vs. restricted to midribs), apices acute to short-acuminate (vs. long-acuminate), the acumen, if present, 0.02-0.08 (vs. acumen 0.1-0.3) mm, and lateral leaves with basiscopic margins entire along proximal
<normalizedToken originalValue="¼-½">1/4-1/2</normalizedToken>
and serrate to short-ciliate distally (vs. usually ciliate throughout).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="72" type="type">
<paragraph pageId="11" pageNumber="72">Type.</paragraph>
<paragraph pageId="11" pageNumber="72">
<emphasis bold="true" pageId="11" pageNumber="72">BRAZIL</emphasis>
. Bahia:
<normalizedToken originalValue="Mucugê">Mucuge</normalizedToken>
, campo defronte ao
<normalizedToken originalValue="cemitério">cemiterio</normalizedToken>
, [ca.
<geoCoordinate degrees="13" direction="south" minutes="00" orientation="latitude" precision="925" value="-13.0">13°00'S</geoCoordinate>
,
<geoCoordinate degrees="41" direction="west" minutes="22" orientation="longitude" precision="15" seconds="19" value="-41.371944">41°22'19&quot;W</geoCoordinate>
], [ca. 984 m], 20 Jul 1981,
<emphasis italics="true" pageId="11" pageNumber="72">A.M. Giulietti et al. [CFCR 1430]</emphasis>
(holotype: NY!; isotypes: PMA- fragment!, SPF-n.v.).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="72" type="description">
<paragraph pageId="11" pageNumber="72">Description.</paragraph>
<paragraph pageId="11" pageNumber="72">
<emphasis italics="true" pageId="11" pageNumber="72">Plants</emphasis>
terrestrial.
<emphasis italics="true" pageId="11" pageNumber="72">Stems</emphasis>
ascending to suberect, stramineous, 3-7 cm long, 0.2-0.4 mm diam., exarticulate, not flagelliform, probably shortly stoloniferous, 2- or 3-branched.
<emphasis italics="true" pageId="11" pageNumber="72">Rhizophores</emphasis>
axillary and axillary-dorsal, restricted to the bases of stems, filiform, 0.1-0.2 mm diam.
<emphasis italics="true" pageId="11" pageNumber="72">Leaves</emphasis>
heteromorphic throughout, chartaceous, both surfaces glabrous, upper surfaces green, lower surfaces silvery green or brownish (when old).
<emphasis italics="true" pageId="11" pageNumber="72">Lateral leaves</emphasis>
distant, slightly ascending, ovate to slightly ovate-oblong, 1.2-1.5
<normalizedToken originalValue="×">x</normalizedToken>
0.9-1.1 mm; bases rounded, acroscopic bases slightly to strongly overlapping stems, basiscopic bases free from stems; acroscopic margins hyaline in a band 2-8 cells wide, the cells elongate and papillate parallel to margins, papillae in 1 row over each cell lumen, long- to short-ciliate along proximal
<normalizedToken originalValue="¾">3/4</normalizedToken>
and serrate to entire along distal
<normalizedToken originalValue="¼">1/4</normalizedToken>
; basiscopic margins narrowly hyaline or greenish in a band 1 or 2 cells wide, the cells as along acroscopic margins, entire along proximal
<normalizedToken originalValue="¼-½">1/4-1/2</normalizedToken>
and serrate to short-ciliate distally, apices acute, tipped by 1-3 teeth; upper surfaces comprising rounded to quadrangular, sinuate-walled cells, some of these covered by 4-8 papillae, without idioblasts or stomata, lower surfaces comprising elongate, sinuate-walled cells, some of these papillate and idioblast-like, papillae in 2 rows over each cell lumen, with stomata in 2 or 3 rows along midribs and along proximal
<normalizedToken originalValue="½">1/2</normalizedToken>
of basiscopic margins.
<emphasis italics="true" pageId="11" pageNumber="72">Median leaves</emphasis>
distant, ascending, ovate, 0.8-1.3
<normalizedToken originalValue="×">x</normalizedToken>
0.5-0.7 mm; bases oblique, inner bases plane in profile, outer bases ventricose (i.e., swollen); margins hyaline in a band 2-5 cells wide, the cells elongate and papillate parallel to margins, papillae in 1 row over each cell lumen, shortly ciliate throughout; apices acute to short-acuminate, each acumen 0.02-0.08 mm, occasionally with 1 or 2 hairs on upper surfaces, tipped by 1-3 teeth; both surfaces without idioblasts, upper surfaces comprising rounded to quadrangular, sinuate-walled cells covered by 4-8 papillae, with stomata along midribs and some on submarginal and marginal regions of the outer bases, lower surfaces comprising elongate, sinuate-walled cells, without stomata.
<emphasis italics="true" pageId="11" pageNumber="72">Axillary leaves</emphasis>
similar to lateral leaves.
<emphasis italics="true" pageId="11" pageNumber="72">Strobili</emphasis>
terminal on branch tips, compact, quadrangular, 2.0-7.0 mm.
<emphasis italics="true" pageId="11" pageNumber="72">Sporophylls</emphasis>
monomorphic, without a laminar flap, ovate, 0.8-1
<normalizedToken originalValue="×">x</normalizedToken>
0.4-0.5 mm, each with a well-developed, frequently puberulous keel along the midribs; bases rounded; margins hyaline, short-ciliate to serrate; apices acute, tipped by 1 or 2 teeth;
<emphasis italics="true" pageId="11" pageNumber="72">dorsal sporophylls</emphasis>
with upper surfaces green and cells as in median leaves, except for the half that overlaps the ventral sporophylls, there hyaline with elongate, papillate, and slightly sinuate-walled cells, lower surfaces silvery green and comprising elongate, sinuate-walled cells;
<emphasis italics="true" pageId="11" pageNumber="72">ventral sporophylls</emphasis>
with both surfaces hyaline, comprising elongate, sinuate-walled cells.
<emphasis italics="true" pageId="11" pageNumber="72">Megasporangia</emphasis>
frequently proximal in 2 ventral rows or the proximal megasporangia abortive and a few intermixed with microsporangia;
<emphasis italics="true" pageId="11" pageNumber="72">megaspores</emphasis>
lemon-yellow, mostly immature or absent, proximal faces not observed, reticulate on distal faces, 275-285
<normalizedToken originalValue="µm">µm</normalizedToken>
.
<emphasis italics="true" pageId="11" pageNumber="72">Microsporangia</emphasis>
in 2 dorsal rows and, in distal portion, also in 2 ventral rows;
<emphasis italics="true" pageId="11" pageNumber="72">microspores</emphasis>
orange, gemmate-rugulate or broadly baculate-rugulate with psilate to echinulate microstructure on proximal faces, vermiculate with echinulate microstructure on distal faces, 30-40
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="72" type="habitat and distribution">
<paragraph pageId="11" pageNumber="72">Habitat and distribution.</paragraph>
<paragraph pageId="11" pageNumber="72">
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="11" pageNumber="72" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="11" pageNumber="72">Selaginella mucugensis</emphasis>
</taxonomicName>
is known only from the type collection from
<normalizedToken originalValue="Mucugê">Mucuge</normalizedToken>
, Serra do
<normalizedToken originalValue="Sincorá">Sincora</normalizedToken>
, in the Chapada Diamantina region of the
<normalizedToken originalValue="Espinhaço">Espinhaco</normalizedToken>
Mountain Range, where it is probably a local endemic. It grows terrestrially on damp soil in Campos Rupestres vegetation at ca. 984 m.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="etymology">
<paragraph pageId="12" pageNumber="73">
<pageBreakToken pageId="12" pageNumber="73" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="12" pageNumber="73">This species is named for the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="conservation status">
<paragraph pageId="12" pageNumber="73">Conservation status.</paragraph>
<paragraph pageId="12" pageNumber="73">
At present, there is limited information available to allow a conclusive determination of the conservation status of
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella mucugensis</emphasis>
</taxonomicName>
. Nevertheless, according to
<bibRefCitation author="IUCN" journalOrPublisher="IUCN, Gland, Switzerland and Cambridge, United Kingdom" pageId="23" pageNumber="84" refId="B10" refString="IUCN, 2012. IUCN Red List Categories and Criteria, Version 3.1. Second edition. Prepared by the IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, United Kingdom" title="IUCN Red List Categories and Criteria, Version 3.1. Second edition. Prepared by the IUCN Species Survival Commission" year="2012">IUCN (2012)</bibRefCitation>
categories and criteria, we tentatively considered this species to be vulnerable (VU) on account that it is so far known from a single locality in the
<normalizedToken originalValue="Espinhaço">Espinhaco</normalizedToken>
Mountain Range, which is threatened by human activities (
<bibRefCitation author="Rapini, A" journalOrPublisher="Megadiversidade" pageId="24" pageNumber="85" pagination="16 - 23" publicationUrl="http://www.conservation.org.br/publicacoes/files_mega4/02_a_flora_dos_campos_rupestres_da_cadeia_do_espinhaco.pdf" refId="B20" refString="Rapini, A, Ribeiro, PL, Lambert, S, Pirani, JR, 2008. A flora dos campos rupestres da Cadeia do Espinhaco. Megadiversidade 4 (1-2): 16 - 23, http://www.conservation.org.br/publicacoes/files_mega4/02_a_flora_dos_campos_rupestres_da_cadeia_do_espinhaco.pdf" title="A flora dos campos rupestres da Cadeia do Espinhaco." url="http://www.conservation.org.br/publicacoes/files_mega4/02_a_flora_dos_campos_rupestres_da_cadeia_do_espinhaco.pdf" volume="4" year="2008">Rapini et al. 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="discussion">
<paragraph pageId="12" pageNumber="73">Discussion.</paragraph>
<paragraph pageId="12" pageNumber="73">
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella mucugensis</emphasis>
</taxonomicName>
is a member of subg.
<taxonomicName class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="(Stachygynandrum)" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="subGenus" subGenus="Stachygynandrum">
<emphasis italics="true" pageId="12" pageNumber="73">Stachygynandrum</emphasis>
</taxonomicName>
and may be confused with
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella blepharodella" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="blepharodella">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella blepharodella</emphasis>
</taxonomicName>
because they have similar leaf margins and indument on the upper surfaces in the distal region of median leaves and sporophylls (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Selaginella crinita Valdespino. A Megaspore proximal face B Close-up of megaspore proximal face surface C Megaspore distal face D Close-up of megaspore distal face surface E Microspore proximal face F Microspore distal face A-F taken from holotype, Harley &amp; Taylor 27048 (NY)." figureDoi="10.3897/phytokeys.50.4873.figure7" httpUri="https://binary.pensoft.net/fig/45219" pageId="12" pageNumber="73">7B</figureCitation>
). In fact, these two species may prove to be sympatric in the Serra do
<normalizedToken originalValue="Sincorá">Sincora</normalizedToken>
, where both were collected. According to
<bibRefCitation author="Harley, RM" journalOrPublisher="Royal Botanic Gardens, Kew" pageId="23" pageNumber="84" pagination="1 - 228" refId="B5" refString="Harley, RM, Simmons, NA, 1986. Florula of Mucuge, Chapada Diamantina, Bahia, Brazil: A descriptive check-list of a campo rupestre area. Royal Botanic Gardens, Kew: 1 - 228" title="Florula of Mucuge, Chapada Diamantina, Bahia, Brazil: A descriptive check-list of a campo rupestre area" year="1986">Harley and Simmons (1986)</bibRefCitation>
, this area is an important center of diversity of the Brazilian montane flora.
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella mucugensis</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella blepharodella" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="blepharodella">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella blepharodella</emphasis>
</taxonomicName>
by the characters discussed under the diagnosis.
</paragraph>
<caption ID-Zenodo-Dep="963553" doi="10.3897/phytokeys.50.4873.figure10" httpUri="https://binary.pensoft.net/fig/45222" pageId="12" pageNumber="73" start="Figure 10" startId="F10">
<paragraph pageId="12" pageNumber="73">
<emphasis bold="true" pageId="12" pageNumber="73">Figure 10.</emphasis>
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella mucugensis</emphasis>
</taxonomicName>
Valdespino.
<emphasis bold="true" pageId="12" pageNumber="73">A</emphasis>
Section of upper surface of stem
<emphasis bold="true" pageId="12" pageNumber="73">B</emphasis>
Upper surface of median leaf
<emphasis bold="true" pageId="12" pageNumber="73">C</emphasis>
Section of lower surface of stem
<emphasis bold="true" pageId="12" pageNumber="73">D</emphasis>
Lower surface of lateral leaf
<emphasis bold="true" pageId="12" pageNumber="73">A-D</emphasis>
taken from holotype,
<emphasis italics="true" pageId="12" pageNumber="73">Giulietti et al. [CFCR 1430]</emphasis>
(NY).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="963555" doi="10.3897/phytokeys.50.4873.figure11" httpUri="https://binary.pensoft.net/fig/45223" pageId="12" pageNumber="73" start="Figure 11" startId="F11">
<paragraph pageId="12" pageNumber="73">
<emphasis bold="true" pageId="12" pageNumber="73">Figure 11.</emphasis>
<taxonomicName authorityName="Valdespino" authorityYear="2015" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Selaginella mucugensis" order="Selaginellales" pageId="12" pageNumber="73" phylum="Lycopodiophyta" rank="species" species="mucugensis">
<emphasis italics="true" pageId="12" pageNumber="73">Selaginella mucugensis</emphasis>
</taxonomicName>
Valdespino.
<emphasis bold="true" pageId="12" pageNumber="73">A</emphasis>
Microspore proximal face
<emphasis bold="true" pageId="12" pageNumber="73">B</emphasis>
Close-up of microspore proximal face surface
<emphasis bold="true" pageId="12" pageNumber="73">C</emphasis>
Microspore distal face
<emphasis bold="true" pageId="12" pageNumber="73">D</emphasis>
Close-up of microspore distal face surface
<emphasis bold="true" pageId="12" pageNumber="73">A-D</emphasis>
taken from holotype,
<emphasis italics="true" pageId="12" pageNumber="73">Giulietti et al. [CFCR 1430]</emphasis>
(NY).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,170 @@
<document ID-CLB-Dataset="297056" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="57b8b4bd-5d41-457f-9201-f4a0de0367c7" ID-ISBN="0-8018-8221-4" checkinTime="1670905223902" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="7AFCCD90A722B2A1C44BDE2B7DB940B5" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Phyllostomidae.395-426.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Desmodus rotundus" docType="treatment" docVersion="4" id="0DE5F2F887EEE913B0D59311D5EEEDFD" lastPageNumber="395" masterDocId="0DE5F2F887EEE913B0D59311D5EEEDFD" masterDocTitle="Order Chiroptera - Family Phyllostomidae" masterLastPageNumber="426" masterPageNumber="395" pageNumber="395" updateTime="1716773540486" updateUser="ExternalLinkService">
<mods:mods id="6DB3E0FAC4CCE23422D7866034E6C39E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="488CAD157BB6E10562E54CD393D4587F">
<mods:title id="6D95D956F0DBE23DD6701FC9BA93DE1C">Order Chiroptera - Family Phyllostomidae</mods:title>
</mods:titleInfo>
<mods:name id="E2AFB3747E9279595C12359BFE0A1A2A" type="personal">
<mods:role id="DBF13564DFD40F546D74E0C5E2ECE5AB">
<mods:roleTerm id="415A1976B83B79C2738D6E8E059E20F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="48FBD775AC391E6021CD151443070F33">Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name id="A45B3523A136594D578E9F473B99C0D4" type="personal">
<mods:role id="2AEA3EBC3A8CB053E63B138A5B370C3C">
<mods:roleTerm id="7EBFAE5F8DAD79F92210A80789D134CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3F5037A54DA03505270F5F589C4C82C">Reeder, DeeAnn</mods:namePart>
</mods:name>
<mods:typeOfResource id="666ACDA59D29EBACEE1F834DD26F6A2A">text</mods:typeOfResource>
<mods:relatedItem id="74B8F88909E261C00EE468B5667D3FD5" type="host">
<mods:originInfo id="4A9938AD4C5929BCAF680FA1BECE6A91">
<mods:dateIssued id="E01B135B5CCDD87E1553C2EF641C8C3E">2005</mods:dateIssued>
<mods:publisher id="965B5DF155FC7BED40F412783AA59DCB">The Johns Hopkins University Press</mods:publisher>
<mods:place id="859E7B4B9277FCFC5AFDD866D48379B6">
<mods:placeTerm id="2400101ACFD2E403E01D0541588C5C5C">Baltimore</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo id="F723615DA952CC6AB98A35FFACC44425">
<mods:title id="2679124CB0A8C0C788B37D979CD38BD6">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1</mods:title>
</mods:titleInfo>
<mods:part id="0F2B8160C8D8ECEE2BB3B9DAA60251A9">
<mods:extent id="6DAF965D2C728FA0E7534191EBAC3DF4" unit="page">
<mods:start id="4C856D72B594D9CFB28FB821982A0470">395</mods:start>
<mods:end id="AAC8EDE9938275A3D6FD593E97AA8B7A">426</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B86D04E996BBC3B3789E1D6B2B78FCD9">book chapter</mods:classification>
<mods:identifier id="996BEF4EA1CD79309ED0AE414D9723D9" type="ISBN">0-8018-8221-4</mods:identifier>
<mods:identifier id="DC2AB79F9A612DD807873FD03DD464D9" type="DOI">10.5281/zenodo.7316519</mods:identifier>
</mods:mods>
<treatment id="7AFCCD90A722B2A1C44BDE2B7DB940B5" ID-DOI="http://doi.org/10.5281/zenodo.11326170" ID-GBIF-Taxon="231521085" ID-Zenodo-Dep="11326170" LSID="urn:lsid:plazi:treatment:7AFCCD90A722B2A1C44BDE2B7DB940B5" httpUri="http://treatment.plazi.org/id/7AFCCD90A722B2A1C44BDE2B7DB940B5" lastPageNumber="395" pageNumber="395" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13801094">
<subSubSection id="81E14BD8DCB8FFA029FA4ED640034339" pageNumber="395" type="nomenclature">
<paragraph id="3253B090E979FD87B8006E40DD77C5E9" pageNumber="395">
<taxonomicName id="FA8F4612D6E6528C261328C79D54CA9C" ID-CoL="353LY" authority="(E. Geoffroy 1810)" baseAuthorityName="E. Geoffroy" baseAuthorityYear="1810" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="rotundus">
<emphasis id="4C269E36A4FA53DDCD4DD21301514C5A" italics="true" pageNumber="395">Desmodus rotundus</emphasis>
(E. Geoffroy 1810)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="669EE3FEB6FB2669E6913AC6438B3C2F" pageNumber="395" type="reference_group">
<paragraph id="D23184BE16BBD03B9068830FE2CC3E35" pageNumber="395">
<treatmentCitation id="EDD1D475796A64D9598E0FCD2D2011D8" author="E. Geoffroy" authority="E. Geoffroy 1810" authorityName="E. Geoffroy" authorityYear="1810" class="Mammalia" family="Phyllostomidae" genus="Phyllostoma" journal="Ann. Mus. Natn. Hist. Nat. Paris" kingdom="Animalia" order="Chiroptera" page="181" pageNumber="395" phylum="Chordata" rank="species" species="rotundum" title="Phyllostoma rotundum" volume="15" year="1810">
<taxonomicName id="200DBCF170BC28439948CCC83D421C47" ID-CoL="8546H" authority="E. Geoffroy 1810" authorityName="E. Geoffroy" authorityYear="1810" class="Mammalia" family="Phyllostomidae" genus="Phyllostoma" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="rotundum">
<emphasis id="E1C472CEC36F936E4B722E31637B2591" italics="true" pageNumber="395">[Phyllostoma] rotundum</emphasis>
E. Geoffroy 1810
</taxonomicName>
,
<bibCitation id="46503716F9A928D146AD6A2D199E9B87" author="E. Geoffroy" journal="Ann. Mus. Natn. Hist. Nat. Paris" pageNumber="395" pagination="181" title="Phyllostoma rotundum" volume="15" year="1810">Ann. Mus. Natn. Hist. Nat. Paris, 15: 181</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AD258710AC33E7A05C847B8513803F94" pageNumber="395" type="materials_examined">
<paragraph id="644552474388B4D69954DB3852D7A748" pageNumber="395">
<emphasis id="15855C436CA8A3BCC4A76B3F5F5D192E" inLineHeading="true" pageNumber="395">Type Locality:</emphasis>
<materialsCitation id="939B859C88679650779B914D48963471" ID-GBIF-Occurrence="4867283383" country="Paraguay" pageNumber="395" stateProvince="Asuncion">
<collectingCountry id="710153491AEB71D0CD5A409870ADF786" name="Paraguay">Paraguay</collectingCountry>
,
<collectingRegion id="F557B083FCF747030EFB19582A466756" country="Paraguay" name="Asuncion">Asunción</collectingRegion>
(restricted by
<bibRefCitation id="F0B03751666C9403CE8327DF23525C7D" author="Cabrera, A." refId="ref48744" refString="Cabrera, A. 1958. Dos felidos argentinos ineditos (Mammalia, Carnivora). Neotropica, 3 (12): 70 - 72." year="1958">Cabrera, 1958</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="512D5F07CAFFF73FE430AF5CF418D327" pageNumber="395" type="vernacular_names">
<paragraph id="7CB37DC51FFEABEEE35E3E0EAC9A7367" pageNumber="395">
<emphasis id="843BCC1F88FBF24EEC3984246C4F1C7F" inLineHeading="true" pageNumber="395">Vernacular Names:</emphasis>
<vernacularName id="545EFBE157A8DB75EA8645996B4F46DB" language="eng" pageNumber="395">Common Vampire Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="FE0353EA562F79368779CC32110DC687" pageNumber="395" type="synonym_list">
<paragraph id="8EB94D57AAF4A22F5B8D5A81861C195F" pageNumber="395">
<emphasis id="CFB5D7D8071E650BACA771ACC9CBC341" inLineHeading="true" pageNumber="395">Synonyms:</emphasis>
<taxonomicName id="FF5AAC10BE634331F7A403ECFF292714" authority="D'Orbigny 1834" authorityName="D'Orbigny" authorityYear="1834" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="cinerea">
<emphasis id="9543A6CC12393E500354C1A74C9ACB1E" italics="true" pageNumber="395">Desmodus cinerea</emphasis>
D'Orbigny 1834
</taxonomicName>
;
<taxonomicName id="F7E81B8ADB721161C6D5CF820A151D7C" authority="Waterhouse 1838" authorityName="Waterhouse" authorityYear="1838" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="dorbignyi">
<emphasis id="898EDD7C93CB3EBF8457B6CADF62270D" italics="true" pageNumber="395">Desmodus dorbignyi</emphasis>
<bibRefCitation id="9D8179590AC18854DE03660511FE2ED0" author="Waterhouse, G. R." refId="ref383195" refString="Waterhouse, G. R. 1838 [1839]. On the skull and dentition of the American badger (Meles labradoria). Proceedings of the Zoological Society of London, 1838: 153 - 154." year="1838">Waterhouse 1838</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2822295F7BB6F73EF32DD76A3400D526" authority="Schinz 1821" authorityName="Schinz" authorityYear="1821" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="ecaudatus">
<emphasis id="3EDACFB459C3871A3EB4681F3B565BDE" italics="true" pageNumber="395">Desmodus ecaudatus</emphasis>
Schinz 1821
</taxonomicName>
;
<taxonomicName id="B1031D942D0299BFC3872AA2ACEB3C49" authority="Burmeister 1854" authorityName="Burmeister" authorityYear="1854" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="947D9FA1B861B6156A0711196380B3F7" italics="true" pageNumber="395">Desmodus fuscus</emphasis>
Burmeister 1854
</taxonomicName>
;
<taxonomicName id="2AF2ABA3BF581AF0AE1A9AB723703231" authority="Burmeister 1879" authorityName="Burmeister" authorityYear="1879" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="mordax">
<emphasis id="C2DC680128CD4D5A2F7C67D17F76E461" italics="true" pageNumber="395">Desmodus mordax</emphasis>
Burmeister 1879
</taxonomicName>
;
<taxonomicName id="6444682D012B42BF19A5FDA488822282" authority="Wagner 1840" authorityName="Wagner" authorityYear="1840" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="murinus">
<emphasis id="96A69E4DF2899191D23E36700B324A25" italics="true" pageNumber="395">Desmodus murinus</emphasis>
Wagner 1840
</taxonomicName>
;
<taxonomicName id="79E7AD164F88134D52B0C8E5CE665C20" authority="Wied-Neuwied 1824" authorityName="Wied-Neuwied" authorityYear="1824" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageNumber="395" phylum="Chordata" rank="species" species="rufus">
<emphasis id="345491D860CFD28E20756CD3ED0D7CFF" italics="true" pageNumber="395">Desmodus rufus</emphasis>
Wied-Neuwied 1824
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9497100D0A104117CAAA380DF071BE54" pageNumber="395" type="distribution">
<paragraph id="FCEE2CDF44D2488855C11F6342E23466" pageNumber="395">
<emphasis id="7B819658C2DBF87F17B82661DF6393D7" inLineHeading="true" pageNumber="395">Distribution:</emphasis>
<collectingCountry id="3FC3826065A3B9C3BD81C97263F57812" name="Uruguay">Uruguay</collectingCountry>
, N
<collectingCountry id="F56F2DEF0706B2029CDA7E676D4116B5" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="59C3A019F66765AC35D9CBA711E37524" name="Paraguay">Paraguay</collectingCountry>
,
<collectingCountry id="289112549E3D026B3173DEC04FC2AE83" name="Bolivia">Bolivia</collectingCountry>
, and N
<collectingCountry id="E69184FA331ED9C6C213D7781F2FCE16" name="Chile">Chile</collectingCountry>
north to
<collectingRegion id="BCCD68DF28E9F57D9A399CEA3CE89B97" country="Mexico" name="Sonora">Sonora</collectingRegion>
,
<collectingRegion id="11C902360D8919094E72D6B35F6527D8" country="Mexico" name="Nuevo Leon">Nuevo León</collectingRegion>
and
<collectingRegion id="9CCB6658B9ABA0B69E8421A53F9876D9" country="Mexico" name="Tamaulipas">Tamaulipas</collectingRegion>
(
<collectingCountry id="DD5EB44C61EBFD11ECB8A7882561C840" name="Mexico">Mexico</collectingCountry>
); Margarita Isl (
<collectingCountry id="FB97D7949587E9BFAC602E7E3BFACD3B" name="Venezuela">Venezuela</collectingCountry>
);
<collectingCountry id="26970250FECEF8FDC03FBE59C0749E13" name="Trinidad and Tobago">Trinidad</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6D145C8EC5DBF365D83C281DF74EBBE7" pageNumber="395" type="conservation">
<paragraph id="749749AE0E249AFC5407EADF7B835281" pageNumber="395">
<emphasis id="0F4BC4FAC059291E79015ED3701D73D2" inLineHeading="true" pageNumber="395">Conservation:</emphasis>
<collectionCode id="3AD42A75CD01F081DD42D0AC26D6C778">IUCN</collectionCode>
2003 and
<collectionCode id="F3D42BEE5EDB239985E18DC2791157C5">IUCN</collectionCode>
/
<collectionCode id="B2803EA4595A5E3C6677F2359A30364E" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="FECA54B1F8B503B997753055495B7A10" pageNumber="395" type="discussion">
<paragraph id="DA8BA60E34C1C03521349D9AE6D11B3E" pageNumber="395">
<emphasis id="FAADFDBA36A3D7BE336DEEB8A00F3C16" inLineHeading="true" pageNumber="395">Discussion:</emphasis>
See
<bibRefCitation id="C75973451564A683214E71C0484605E0" author="Greenhall, A. M. &amp; G. Joermann &amp; U. Schmidt &amp; M. R. Seidel" refId="ref129368" refString="Greenhall, A. M., G. Joermann, U. Schmidt, and M. R. Seidel. 1983. Desmodus rotundus. Mammalian Species, 202: 1 - 6." year="1983">Greenhall et al. (1983)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>