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<document id="C3B7F944EE14DA7669D885FA6FA4E1C1" ENCODING="UTF-8" ID-CLB-Dataset="276868" ID-DOI="https://doi.org/10.5281/zenodo.291971" ID-GBIF-Dataset="d15ba707-bee0-4d4d-a76d-52e5d488763c" ID-ISBN="978-0-9506207-7-0" ID-Zenodo-Dep="291971" checkinTime="1588973417084" checkinUser="admin" docAuthor="Jarvis, Charlie" docDate="2007" docId="7ADB46977B07820AA96F9C0A5F09F29D" docLanguage="en" docName="ch7.gg.E.xml" docOrigin="Order out of Chaos. Linnaean Plant Types and their Types, London: Linnaean Society of London in association with the Natural History Museum" docTitle="Blumea obliqua Druce 1917" docType="treatment" docVersion="9" lastPageNumber="502" masterDocId="3605B3979E7AB629CCA701EDC6B5CC30" masterDocTitle="Chapter 7: Linnaean Plant Names and their Types (part E)" masterLastPageNumber="515" masterPageNumber="490" pageNumber="502" updateTime="1699550148738" updateUser="ExternalLinkService">
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<mods:title id="62BC6B6E0B04DD2FE38CD5892D6CE43C">Chapter 7: Linnaean Plant Names and their Types (part E)</mods:title>
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<mods:namePart id="4A32D37335AB26FB47D20AF192A0E417">Jarvis, Charlie</mods:namePart>
<mods:affiliation id="851F8F1C719B08B0E526EBF1CC1B5442">Department of Botany, Natural History Museum, Cromwell Road, London, UK</mods:affiliation>
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<mods:typeOfResource id="29CE6415E39C87D1DFEFE71FFC6665AD">text</mods:typeOfResource>
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<mods:title id="FB417826F13CB52DA570923535E575D1">Order out of Chaos. Linnaean Plant Types and their Types</mods:title>
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<mods:identifier id="4507525F1E6A0AD60748E7CC12C1806E" type="DOI">https://doi.org/10.5281/zenodo.291971</mods:identifier>
<mods:identifier id="0F3CCD1C854A0C165D5068B223A4790A" type="ISBN">978-0-9506207-7-0</mods:identifier>
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<treatmentCitation id="4CC551059B13DD5A4329A535A8EA4CF6" author="Linnaeus" pageNumber="574" year="1771">
<bibCitation id="6E30FB64F0AA612D383F8643DFC5F892" author="Linnaeus" pagination="574" title="Erigeron obliquus" volumeTitle="Mantissa Plantarum Altera" year="1771">
<taxonomicName id="8613F3DB099B1692918259F4B26CC83D" ID-CoL="6H22W" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1771" class="Magnoliopsida" family="Asteraceae" genus="Erigeron" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="obliquus">
<emphasis id="C19AE67C8262D3255A756DA35C2BD99D" bold="true" pageNumber="502">Erigeron obliquus</emphasis>
Linnaeus
</taxonomicName>
,
<emphasis id="CD101C8CA2C08BD43EB9667519940253" italics="true" pageNumber="502">
<volumeTitle id="66B12F63744ACD583EB11F734FBF9496">Mantissa Plantarum Altera</volumeTitle>
</emphasis>
: 574. 1771
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.
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<paragraph id="DA20C981D1CD1178773BEB102F9ECD14" pageNumber="502">
&quot;Habitat in
<taxonomicName id="525A014CA1B6C856F5A8DC0E022E8EEA" authorityName="A.Nageswara Rao" authorityYear="1999" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="genus">India</taxonomicName>
orientali.&quot; RCN: 6240.
</paragraph>
</subSubSection>
<subSubSection id="646D833CC3D997C2C933CE02EA548D9C" pageNumber="502" type="materials_examined">
<paragraph id="07C3D417CD8029ED9B1597AB34FB1C90" pageNumber="502">
<materialsCitation id="C4385094599D98B622FC707AFDCA689A" ID-GBIF-Occurrence="2611986383" collectionCode="LINN" specimenCode="994.1" typeStatus="lectotype">
<typeStatus id="DCCA9B41D451A260147D714D83E74A16">Lectotype</typeStatus>
(Randeria in
<taxonomicName id="EAFA97C750B2DF2472AC10D9037605F6" class="Magnoliopsida" family="Asteraceae" genus="Blumea" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="genus">
<emphasis id="381B3DE15D532916669402288F35603F" italics="true" pageNumber="502">Blumea</emphasis>
</taxonomicName>
10: 287. 1960): Herb. Linn. No. 994.1 (
<collectionCode id="8E02AB0347C645212D5DD91768DC61E0">LINN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2DB075525A1F0506FBF5BF4A08E1412A" pageNumber="502" type="nomenclature">
<paragraph id="4BD2278C437E3A7AE03418E1244B405A" pageNumber="502">
Current name:
<taxonomicName id="D29AFE0E5520ADBAB0ABC9F262FD080E" ID-CoL="M7JV" authority="(L.) Druce" authorityName="Druce" authorityYear="1917" baseAuthorityName="Linnaeus" baseAuthorityYear="1771" class="Magnoliopsida" family="Asteraceae" genus="Blumea" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="obliqua">
<emphasis id="C1ECA795FBF8583E602E8A02EB8DEE55" bold="true" italics="true" pageNumber="502">Blumea obliqua</emphasis>
(L.) Druce
</taxonomicName>
(
<taxonomicName id="F03E2D89BDA6FA54B56EFC7AD9E92820" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="1F45B92E606848AB52D67A89779C5D06" pageNumber="502" type="discussion">
<paragraph id="C10DAA1C5408AA6E33173A82102F6132" pageNumber="502">
<emphasis id="8FC6ECA05487FBC8F422A8ACDF482E37" italics="true" pageNumber="502">Note:</emphasis>
Specific epithet spelled
<normalizedToken id="9FB1C71C7A56DD01F3017D3880B87DDD" originalValue="“obliquum”">&quot;obliquum&quot;</normalizedToken>
in the protologue.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Revision of the Ceratocapsine Renodaeus group: Marinonicoris, Pilophoropsis, Renodaeus, and Zanchisme, with descriptions of four new genera (Heteroptera, Miridae, Orthotylinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henry, Thomas J.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<paragraph pageId="83" pageNumber="84">Taxon classification Animalia Hemiptera Miridae</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="nomenclature">
<paragraph pageId="83" pageNumber="84">
<taxonomicName authority="Distant" class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">Renodaeus Distant</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="reference_group">
<paragraph pageId="83" pageNumber="84">
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
Distant, 1893: 461 (orig. descrip.);
<bibRefCitation author="Knight, HH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="103" pageNumber="104" pagination="56 - 57" title="Description of a new Renodaeus from Texas (Hemiptera, Miridae)." volume="21" year="1926">Knight 1926</bibRefCitation>
: 57 (notes, diag., n. tribe);
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Anais da Academia Brasileira de Ciencias" pageId="101" pageNumber="102" pagination="31 - 110" title="On the major classification of the Miridae (Hemiptera). (with keys to subfamilies and tribes and a catalogue of the world genera)." volume="24" year="1952">Carvalho 1952</bibRefCitation>
: 83 (list); 1955: 80 (key);
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Orthotylinae" pageId="101" pageNumber="102" pagination="1 - 161" title="Catalogue of the Miridae of the World. Arquivos Museu Nacional, Rio de Janeiro. Part III." volume="47" year="1958">1958</bibRefCitation>
: 150 (cat.);
<bibRefCitation author="Schuh, RT" journalOrPublisher="Entomologica Americana" pageId="104" pageNumber="105" pagination="1 - 332" title="The Orthotylinae and Phylinae (Hemiptera: Miridae) of South Africa with a phylogenetic analysis of the ant-mimetic tribes of the two subfamilies for the world." volume="47" year="1974">Schuh 1974</bibRefCitation>
: 285 (note, moved to
<taxonomicName lsidName="" pageId="83" pageNumber="84" rank="tribe" tribe="Orthotylini">Orthotylini</taxonomicName>
);
<bibRefCitation author="Henry, TJ" editor="Henry, TJ" journalOrPublisher="Brill EJ, Leiden and New York" pageId="103" pageNumber="104" pagination="251 - 507" title="Family Miridae Hahn, 1833." volumeTitle="Catalog of the Heteroptera, or True Bugs, of Canada and the Continental United States" year="1988">Henry and Wheeler 1988</bibRefCitation>
: 445 (cat.);
<bibRefCitation author="Schuh, RT" journalOrPublisher="New York Entomological Society, New York" pageId="104" pageNumber="105" title="Plant Bugs of the World (Insecta: Heteroptera: Miridae). Systematic catalog, distributions, host list, and bibliography." year="1995">Schuh 1995</bibRefCitation>
: 190 (cat.). Type species:
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus ficarius" order="Hemiptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="ficarius">Renodaeus ficarius</taxonomicName>
Distant. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="diagnosis">
<paragraph pageId="83" pageNumber="84">Diagnosis.</paragraph>
<paragraph pageId="83" pageNumber="84">This genus is recognized by the constricted, punctate hemelytron with patches of golden scale-like setae and bands of silvery scale-like setae on the clavus and corium, the punctate pronotum (at least on posterior half), the heavily punctate clavus and basal half of corium, the finely punctate cuneus, and the row of file-like, tiny black spicules (stridulitrum) along the edge of the costal margin in males and females.</paragraph>
</subSubSection>
<subSubSection lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="description">
<paragraph pageId="83" pageNumber="84">Description.</paragraph>
<paragraph lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
Length 2.60-3.30 mm. Myrmecomorphic. Males and females macropterous. Head declivent, subtriangular in dorsal aspect, ventrally flattened in lateral aspect, basal margin truncate, vertex with a distinct smooth carina, basal margin of eyes level with base of vertex, frons granulate or punctate, clypeus not or hardly visible in dorsal aspect; eyes moderately large, combined widths occupying about half dorsal width of head in males, about two thirds in females, oblong oval in lateral aspect; segment I of labium extending beyond gular sulcus by about half its length, visible below buccula in lateral aspect, extending to middle coxae or beyond. Antenna generally slender, segment I shortest, II longest, slender basally, gradually enlarging to apex which is sometimes clavate, apex subequal to diameter of segments III and IV; III and IV fusiform. Pronotum strongly convex, with a narrow but distinct collar, middle of disc higher than level of head in lateral aspect; in males trapeziform, lateral margins sulcate, humeral angles flared and depressed adjacent to posterior angles, basal margin rounded; in females subquadrate, lateral margins nearly straight, humeral angles weakly flared; scutellum equilateral, base covered by posterior margin of pronotum.
<pageBreakToken pageId="84" pageNumber="85" start="start">Hemelytron</pageBreakToken>
: polished, constricted at middle of corium, widest across level of cuneus; corium and clavus with predominately golden, scale-like setae, sometimes with silvery and black scale-like setae, intermixed with long, erect, pale or black, bristle-like setae on clavus, corium, and along inner margin of cuneus; deeply punctured on clavus and middle area of corium, with very fine, scattered punctures on cuneus; costal edge in both sexes with a row of tiny black spicules, possibly serving as a stridulatory file (stridulitrum); membrane entire. Ventral surface shiny; ostiolar area pale to whitish, with a raised knob at end of scent channel; glue obscuring base of abdomen (i.e., cannot determine presence of dull or glaucous patch). Legs unmodified, except hind femur in female sometimes thickened on apical half and narrowed at apex; hind tibiae roughened and armed with two rows of tiny spicules, possibly serving as the plectrum during stridulation. Male aperture large, unarmed, genitalia as illustrated (Figs 286-291).
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="discussion">
<paragraph pageId="84" pageNumber="85">Discussion.</paragraph>
<paragraph pageId="84" pageNumber="85">
As previously noted by
<bibRefCitation author="Knight, HH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="103" pageNumber="104" pagination="56 - 57" title="Description of a new Renodaeus from Texas (Hemiptera, Miridae)." volume="21" year="1926">Knight (1926)</bibRefCitation>
, this genus was originally described by
<bibRefCitation author="Distant, WL" editor="Goodman," journalOrPublisher="London" pageId="102" pageNumber="103" pagination="1 - 88" title="Insecta. Rhynchota. Hemiptera-Heteroptera. Vol. I." volumeTitle="Biologia Centrali-Americana" year="1880 - 1893">Distant (1893)</bibRefCitation>
as an aberrant pyrrhocorid.
<bibRefCitation author="Knight, HH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="103" pageNumber="104" pagination="56 - 57" title="Description of a new Renodaeus from Texas (Hemiptera, Miridae)." volume="21" year="1926">
<normalizedToken originalValue="Knight’s">Knight's</normalizedToken>
(1926)
</bibRefCitation>
description of
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
as having
<taxonomicName class="Insecta" family="Miridae" genus="Ceratocapsus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratocapsus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Ceratocapsus</taxonomicName>
-like antennae and a head and pronotum like those of
<taxonomicName class="Insecta" family="Miridae" genus="Pilophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilophorus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Pilophorus</taxonomicName>
, convinced him that it did not belong in either
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Ceratocapsini">Ceratocapsini</taxonomicName>
or
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Pilophorini">Pilophorini</taxonomicName>
; thus, he established the new tribe
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Renodaeini">Renodaeini</taxonomicName>
.
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Anais da Academia Brasileira de Ciencias" pageId="101" pageNumber="102" pagination="31 - 110" title="On the major classification of the Miridae (Hemiptera). (with keys to subfamilies and tribes and a catalogue of the world genera)." volume="24" year="1952">Carvalho (1952</bibRefCitation>
, 1955,
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Orthotylinae" pageId="101" pageNumber="102" pagination="1 - 161" title="Catalogue of the Miridae of the World. Arquivos Museu Nacional, Rio de Janeiro. Part III." volume="47" year="1958">1958</bibRefCitation>
), however, treated
<normalizedToken originalValue="Knight’s">Knight's</normalizedToken>
tribe as a synonym of
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Pilophorini">Pilophorini</taxonomicName>
and
<bibRefCitation author="Schuh, RT" journalOrPublisher="New York Entomological Society, New York" pageId="104" pageNumber="105" title="Plant Bugs of the World (Insecta: Heteroptera: Miridae). Systematic catalog, distributions, host list, and bibliography." year="1995">Schuh (1995)</bibRefCitation>
listed it in synonymy under
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Orthotylini">Orthotylini</taxonomicName>
, along with
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Ceratocapsini">Ceratocapsini</taxonomicName>
. It is now clear that
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
belongs in
<taxonomicName lsidName="" pageId="84" pageNumber="85" rank="tribe" tribe="Ceratocapsini">Ceratocapsini</taxonomicName>
based on the male genitalia.
</paragraph>
<paragraph pageId="84" pageNumber="85">
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
is similar to
<taxonomicName class="Insecta" family="Miridae" genus="Marinonicoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Marinonicoris" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Marinonicoris</taxonomicName>
in overall appearance, including the golden patches and silvery bands of scale-like setae on the hemelytra. It differs, however, in having erect, bristle-like setae on the hemelytra, a row of file-like spicules on the costal margin, and considerably different male genitalia. The left paramere (Figs 286, 289) in species of
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
is C-shaped, sometimes with a process arising from the middle, the right paramere (Figs 288, 291) is broad with two diverging lobes, and the phallotheca (Figs 287, 290) is relatively simple with the apex blunt or moderately pointed, whereas in
<taxonomicName class="Insecta" family="Miridae" genus="Marinonicoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Marinonicoris myrmecoides" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="myrmecoides">Marinonicoris myrmecoides</taxonomicName>
, the left paramere (Fig. 280) is slender with an apical beak-like apex and a slender, apically rounded process at the middle, the right paramere (Fig. 282) is stout and elongate, with a slender, apically bifid arm basally, and the phallotheca (Fig. 281) is broad with three, apical, comb-like processes.
</paragraph>
<paragraph pageId="84" pageNumber="85">
The lack of males precludes positive identification of
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus ficarius" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ficarius">Renodaeus ficarius</taxonomicName>
. Based on the examination of the lectototype and paralectotype females,
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus ficarius" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ficarius">Renodaeus ficarius</taxonomicName>
has a more extensive covering of scale-like setae on the hemelytra, as noted in the key.
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="key to the species of renodaeus">
<paragraph pageId="84" pageNumber="85">
Key to the species of
<taxonomicName class="Insecta" family="Miridae" genus="Renodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Renodaeus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">Renodaeus</taxonomicName>
</paragraph>
<paragraph pageId="84" pageNumber="85">
<table pageId="84" pageNumber="85">
<tr pageId="84" pageNumber="85">
<td colspan="1" pageId="84" pageNumber="85" rowspan="1">
<taxonomicName class="Insecta" family="Miridae" genus="Heteroptera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Heteroptera ficarius" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ficarius">ficarius</taxonomicName>
Distant
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<td colspan="1" pageId="84" pageNumber="85" rowspan="1">2</td>
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<td colspan="1" pageId="84" pageNumber="85" rowspan="1">
<taxonomicName class="Insecta" family="Miridae" genus="Heteroptera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Heteroptera mimeticus" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="mimeticus">mimeticus</taxonomicName>
sp. n.
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<td colspan="1" pageId="84" pageNumber="85" rowspan="1">
<taxonomicName class="Insecta" family="Miridae" genus="Heteroptera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Heteroptera gibbicollis" order="Hemiptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="gibbicollis">gibbicollis</taxonomicName>
Carvalho &amp; Becker
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159
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<paragraph pageId="31" pageNumber="116">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
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<taxonomicName LSID="http://zoobank.org/22C94495-44A0-40BD-8F4E-E6CFC415B283" authority="Klimaszewski &amp; Webster" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Dimetrota) sphagnicola" order="Coleoptera" pageId="31" pageNumber="116" phylum="Arthropoda" rank="species" species="sphagnicola" subGenus="Dimetrota">Atheta (Dimetrota) sphagnicola Klimaszewski &amp; Webster</taxonomicName>
<taxonomicNameLabel pageId="31" pageNumber="116">sp. n.</taxonomicNameLabel>
Figs 141-145
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<subSubSection pageId="32" pageNumber="117" type="holotype (male)">
<paragraph pageId="32" pageNumber="117">
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(male).
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<paragraph pageId="32" pageNumber="117">
Canada, New Brunswick, York Co., Charters Settlement,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.8267">45.8267°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.7343">66.7343°W</geoCoordinate>
, 16.IV.2005, R.P. Webster, coll. //
<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Carex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carex" order="Poales" pageId="32" pageNumber="117" phylum="Tracheophyta" rank="genus">Carex</taxonomicName>
marsh in
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hummocks (LFC). Paratypes: Canada, New Brunswick, Queens Co., Upper Gagetown, bog adjacent to Hwy 2,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.8316">45.8316°N</geoCoordinate>
,
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, 12.IV.2006, R.P. Webster, coll. // Tamarack bog, in sphagnum hummocks &amp; litter at bog margin (1 ♂, RWC). Saint John Co., Chance Harbour off Rt. 790,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.1355">45.1355°N</geoCoordinate>
,
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, 15.V.2006, R.P. Webster, coll. // Calcareous fen, in sphagnum &amp; litter in depression with
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(1 ♂, RWC).
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<subSubSection pageId="32" pageNumber="117" type="etymology">
<paragraph pageId="32" pageNumber="117">Etymology.</paragraph>
<paragraph pageId="32" pageNumber="117">
The specific name, sphagnicola, meaning &quot;living on
<taxonomicName class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="32" pageNumber="117" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
&quot;, is in reference to the
<taxonomicName class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="32" pageNumber="117" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
hummocks where the holotype was collected.
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<paragraph pageId="32" pageNumber="117">Description.</paragraph>
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Body length 3.4 mm, narrowly elongate, subparallel; head, pronotum, and abdomen dark brown to almost black, elytra yellowish reddish brown with base and scutellar area darker (Fig. 141); integument strongly glossy with strong meshed microsculpture; forebody with fine and moderately dense punctation and pubescence; head rounded and arcuate posterolaterally, with eyes moderately large, about as long as postocular area; antennae with articles
<normalizedToken originalValue="V–X">V-X</normalizedToken>
subquadrate to moderately transverse;
<pageBreakToken pageId="33" pageNumber="118" start="start">pronotum</pageBreakToken>
transverse, rounded on sides, distinctly wider than head and narrower than elytra, pubescence directed laterad from midline of disk; elytra transverse, truncate apically, with pubescence directed posterolaterad; abdomen subparallel, narrower than
<pageBreakToken pageId="34" pageNumber="119" start="start">elytra</pageBreakToken>
and arcuate laterally. Male. Median lobe of aedeagus with bulbus moderately narrowly oval, with rounded projections apicolaterally (Fig. 142), tubus with apical part narrowly triangular and sinuate in lateral view, venter arcuate (Fig. 143); internal sac with complex structures (Figs 142, 143); tergite VIII shallowly emarginate apically, with two small lateral teeth (Fig. 144); sternite VIII elongate and rounded apically (Fig. 145). Female. Unknown.
</paragraph>
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<paragraph pageId="34" pageNumber="119">Distribution.</paragraph>
<paragraph pageId="34" pageNumber="119">Known only from NB, Canada.</paragraph>
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<paragraph pageId="34" pageNumber="119">Natural history.</paragraph>
<paragraph pageId="34" pageNumber="119">
The three known individuals of this species were collected from sphagnum in a
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marsh, a calcareous fen, and a tamarack bog. Adults were collected during April and May.
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<paragraph pageId="34" pageNumber="119">Comments.</paragraph>
<paragraph pageId="34" pageNumber="119">
This species is very distinct in its genital structures. It is superficially similar to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Dimetrota) venti" order="Coleoptera" pageId="34" pageNumber="119" phylum="Arthropoda" rank="species" species="venti" subGenus="Dimetrota">Atheta (Dimetrota) venti</taxonomicName>
(Lohse) in terms of having a broad tubus of the median lobe of aedeagus in dorsal view. For illustration of
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see
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.
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<paragraph pageId="34" pageNumber="119">
Figures 141-145.
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Klimaszewski &amp; Webster, sp. n.: 141 habitus in dorsal view 142 median lobe of aedeagus in dorsal view 143 median lobe of aedeagus in lateral view 144 male tergite VIII 145 male sternite VIII. Scale bar of habitus = 1 mm; remaining scale bars = 0.2 mm.
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<taxonomicName id="9D758C232E7685153C77D9651CD150CE" ID-CoL="4283Z" ID-ENA="1031671" LSID="7ADBE5C3-6B32-5691-8A46-E4FABF572E12" authority="Krausse, 1910" authorityName="Krausse" authorityYear="1910" class="Insecta" family="Formicidae" genus="Messor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Messor wasmanni" order="Hymenoptera" pageId="0" pageNumber="95599" phylum="Arthropoda" rank="species" species="wasmanni">Messor wasmanni Krausse, 1910</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5A9E1CFD965E616B207161E2673F6085" pageId="0" pageNumber="95599" type="notes">
<paragraph id="69AB76A8996ACD9FB15E0E1F9A58B95F" pageId="0" pageNumber="95599">Notes</paragraph>
<paragraph id="ECEBB73FC996A8DEC859422E7CE267BD" pageId="0" pageNumber="95599">
<bibRefCitation id="443010E11A604DCA3CCC55C637BA96AA" author="Atanassov, N." journalOrPublisher="Bulletin de la Societe Entomologique de Bulgarie" pageId="0" pageNumber="95599" pagination="211 - 236" refId="B8016430" refString="Atanassov, N., 1936. Zweiter Beitrag zum Studium der Ameisenfauna Bulgariens (Formicidae). Bulletin de la Societe Entomologique de Bulgarie 9: 211 - 236" title="Zweiter Beitrag zum Studium der Ameisenfauna Bulgariens (Formicidae)" volume="9" year="1936">Atanassov (1936)</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Zaika, Maxim A.</mods:namePart>
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<mods:namePart>Kilian, Norbert</mods:namePart>
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<mods:namePart>Speranskaya, Anna S.</mods:namePart>
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<treatment ID-GBIF-Taxon="161671725" LSID="urn:lsid:plazi:treatment:7ADBE6F65D695409BC6E9468436F2166" httpUri="http://treatment.plazi.org/id/7ADBE6F65D695409BC6E9468436F2166" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="7ADBE6F6-5D69-5409-BC6E-9468436F2166" authority="Nazarova, Biol. Zhurn. Armenii 43: 179. 1990." class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Takhtajaniantha Nazarova, Biol. Zhurn. Armenii 43: 179. 1990.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
=
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Fibrillosae">Scorzonera sect. Fibrillosae</taxonomicName>
Nakai, Rep. Inst. Sci. Res. Manchoukuo, ser. 1, 6: 171. 1937. Type:
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera glabra" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="glabra">Scorzonera glabra</taxonomicName>
Rupr. [=
<taxonomicName family="Asteraceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="S. austriaca" order="Asterales" pageId="0" pageNumber="1" rank="species" species="austriaca">S. austriaca</taxonomicName>
Willd.]
</paragraph>
<paragraph pageId="0" pageNumber="1">
=
<taxonomicName authority="subsect. Egregia" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera subsect. Egregia</taxonomicName>
Kult., Tau-Sagyz Ekol. Osnovy Vvedeniya Ego
<normalizedToken originalValue="Kul’t">Kul't</normalizedToken>
.: 108. 1938.
</paragraph>
<paragraph pageId="0" pageNumber="1">
≡
<taxonomicName authority="subg. Egregia" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera subg. Egregia</taxonomicName>
(Kult.) Ovcz., Soobshch. Tadzhiksk. Fil. Akad. Nauk SSSR 20: 54. 1950.
</paragraph>
<paragraph pageId="0" pageNumber="1">
≡
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Egregia">Scorzonera sect. Egregia</taxonomicName>
(Kult.) Lipsch., Fl. URSS 29: 58. 1964. Lectotype (designated here):
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">Scorzonera tau-saghyz</taxonomicName>
Lipsch. &amp; G.G.Bosse
</paragraph>
<paragraph pageId="0" pageNumber="1">
=
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Pusillae">Scorzonera sect. Pusillae</taxonomicName>
Lipsch., Fragm. Monogr. Gen.
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera</taxonomicName>
2: 59. 1939. Type:
<taxonomicName authorityName="Pall" authorityYear="1773" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">Scorzonera pusilla</taxonomicName>
Pall.
</paragraph>
<paragraph pageId="0" pageNumber="1">
=
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Capito">Scorzonera sect. Capito</taxonomicName>
Tzvelev, Rast. Tsentral. Azii 14b: 115. 2009. Type:
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">Scorzonera capito</taxonomicName>
Maxim.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pusilla</emphasis>
</taxonomicName>
(Pall.) Nazarova
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
<paragraph pageId="0" pageNumber="1">Diagnostic features.</paragraph>
<paragraph pageId="0" pageNumber="1">Perennials; caudex often with dark brown fibrous leaf sheath residues and base of the rosulate leaves often lanate; pollen with 6 lacunae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Habit, life form, subterranean parts:</emphasis>
perennial herbs or subshrublets with a taproot or a tuber.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves:</emphasis>
rosulate or crowded in the basal part of the stem, rarely dispersed along the stem, sessile or petiolate, filiform or ovate, apically sometimes hooked (
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
), margin flat or undulate, entire or denticulate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem, synflorescence:</emphasis>
stem solitary or branched, scape-like or leafy, capitula terminal.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pollen:</emphasis>
echinolophate, tricolporate and each colpus divided into 2 lacunae, with 6 abporal lacunae (sometimes with spineless areas resembling rudimentary lacunae (
<bibRefCitation DOI="https://doi.org/10.1080/00173138209427691" author="Blackmore, S" journalOrPublisher="Grana" pageId="0" pageNumber="1" pagination="149 - 160" refId="B9" refString="Blackmore, S, 1982. Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance. Grana 21 (3): 149 - 160, DOI: https://doi.org/10.1080/00173138209427691" title="Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance." url="https://doi.org/10.1080/00173138209427691" volume="21" year="1982">Blackmore 1982</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula:</emphasis>
glabrous to arachnoid hairy or tomentose, phyllaries in several series, outer phyllaries smaller than inner ones, triangular-ovate, inner phyllaries linear to ovate, receptacle glabrous, flat, with many or rarely few (up to 12:
<taxonomicName lsidName="T. pseudodivaricata" pageId="0" pageNumber="1" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">T. pseudodivaricata</emphasis>
</taxonomicName>
) florets; florets yellow (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Images of living plants. A, B Ramaliella tortuosissima A general view B close up view of the capitula (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir C Pseudopodospermum papposum (Jordan, Nakeb, 8 May 2019). Photograph by O. Fragman-Sapir D Takhtajaniantha austriaca (Italy, South Tyrol, Schlanders, 6 May 2010). Photograph by R. Hand E Scorzonera humilis (Germany, Ammersee, 15 May 2009). Photograph by C. Niederbichler F Scorzonera meyeri (Russia, Karachaevo-Cherkessiya, Teberda, 14 July 2005). Photograph by A. S. Zernov." figureDoi="10.3897/phytokeys.137.46544.figure16" httpUri="https://binary.pensoft.net/fig/370364" pageId="0" pageNumber="1">16D</figureCitation>
) or with dark red veins, usually clearly exeeding the incolucre.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes:</emphasis>
5-15 mm, without carpopodium, straight or slightly curved, often with 10 ribs or rarely roundish, glabrous or hairy, sometimes (as in
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
) with stout conglomerations; achene wall with parenchyma of two types, with thick- and thin-walled cells, present only as subepidermal continuous layer(s) or only with thick-walled cells, air cavities absent, sclerenchyma with continuous layer with a narrow or wider hunch on either side of the principal rib or, as in
<taxonomicName lsidName="T. pseudodivaricata" pageId="0" pageNumber="1" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">T. pseudodivaricata</emphasis>
</taxonomicName>
, without hunches, but discontinuous forming 5 bundles (with a gap in the principal ribs), or continuous, its fibres orientated parallel to the fruit axis, tannins absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pappus:</emphasis>
7-28 mm, ice white or dirty white, bristles scabrid in upper part.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Chromosome number:</emphasis>
x = 7 (diploids); 2n = 28 in
<taxonomicName lsidName="T. pusilla" pageId="0" pageNumber="1" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">T. pusilla</emphasis>
</taxonomicName>
(amphiploid); in
<taxonomicName lsidName="T. tau-saghyz" pageId="0" pageNumber="1" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">T. tau-saghyz</emphasis>
</taxonomicName>
polyploid cytoraces 2n = 14, 21, 28, 42 and aneuploids with chromosome numbers 15, 16, 17, 18, 22 and 24, 28 and 42 are known (
<bibRefCitation author="Krayevoy, SU" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="367 - 375" refId="B53" refString="Krayevoy, SU, 1934. K voprosu o khromosomnykh variatsiyah v Scorzonera tau-saghyz. Botanicheskii Zhurnal 19: 367 - 375" title="K voprosu o khromosomnykh variatsiyah v Scorzonera tau-saghyz." volume="19" year="1934">Krayevoy 1934</bibRefCitation>
;
<bibRefCitation author="Poddubnaya-Arnoldi, VA" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="338 - 366" refId="B80" refString="Poddubnaya-Arnoldi, VA, Steshina, N, Sosnovets, A, 1934. Razvitiye i zhiznesposobnost' pyltsy Scorzonera tau-saghyz Lipsch et Bosse. Botanicheskii Zhurnal 19: 338 - 366" title="Razvitiye i zhiznesposobnost' pyltsy Scorzonera tau-saghyz Lipsch et Bosse." volume="19" year="1934">Poddubnaya-Arnoldi et al. 1934</bibRefCitation>
;
<bibRefCitation author="Sosnovets, AP" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="1813 - 1815" refId="B91" refString="Sosnovets, AP, 1960. Tsitologiya roda Scorzonera L. Botanicheskii Zhurnal 45: 1813 - 1815" title="Tsitologiya roda Scorzonera L." volume="45" year="1960">Sosnovets 1960</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Species.</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="1">
(1)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha austriaca" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="austriaca">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha austriaca</emphasis>
</taxonomicName>
(Willd.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Images of living plants. A, B Ramaliella tortuosissima A general view B close up view of the capitula (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir C Pseudopodospermum papposum (Jordan, Nakeb, 8 May 2019). Photograph by O. Fragman-Sapir D Takhtajaniantha austriaca (Italy, South Tyrol, Schlanders, 6 May 2010). Photograph by R. Hand E Scorzonera humilis (Germany, Ammersee, 15 May 2009). Photograph by C. Niederbichler F Scorzonera meyeri (Russia, Karachaevo-Cherkessiya, Teberda, 14 July 2005). Photograph by A. S. Zernov." figureDoi="10.3897/phytokeys.137.46544.figure16" httpUri="https://binary.pensoft.net/fig/370364" pageId="0" pageNumber="1">16D</figureCitation>
) ≡
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera austriaca" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="austriaca">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera austriaca</emphasis>
</taxonomicName>
Willd., Sp. Pl. 3(3): 1498. 1803 (incl.
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. glabra" order="Asterales" pageId="0" pageNumber="1" rank="species" species="glabra">
<emphasis italics="true" pageId="0" pageNumber="1">S. glabra</emphasis>
</taxonomicName>
Rupr.). urn:lsid:ipni.org:names:77204033-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(2)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha capito</emphasis>
</taxonomicName>
(Maxim.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera capito" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="capito">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera capito</emphasis>
</taxonomicName>
Maxim., Bull. Acad. Imp. Sci.
<normalizedToken originalValue="Saint-Pétersb">Saint-Petersb</normalizedToken>
. 32: 491. 1888. urn:lsid:ipni.org:names:77204030-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(3)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha crispa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="crispa">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha crispa</emphasis>
</taxonomicName>
(M.Bieb.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Bieb" authorityYear="1808" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera crispa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="crispa">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera crispa</emphasis>
</taxonomicName>
M.Bieb., Fl. Taur.-Caucas. 2: 234. 1808. urn:lsid:ipni.org:names:77204028-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(4)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha ikonnikovii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="ikonnikovii">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha ikonnikovii</emphasis>
</taxonomicName>
(Krasch. &amp; Lipsch.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera ikonnikovii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="ikonnikovii">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera ikonnikovii</emphasis>
</taxonomicName>
Krasch. &amp; Lipsch. in Lipschitz, Fragm. Monogr. Gen.
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera</taxonomicName>
1: 109. 1935. urn:lsid:ipni.org:names:77204024-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(5)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha mongolica" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mongolica">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha mongolica</emphasis>
</taxonomicName>
(Maxim.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Maxim" authorityYear="1888" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera mongolica" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="mongolica">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera mongolica</emphasis>
</taxonomicName>
Maxim., Bull. Acad. Imp. Sci.
<normalizedToken originalValue="Saint-Pétersb">Saint-Petersb</normalizedToken>
. 32: 492. 1888. urn:lsid:ipni.org:names:77204020-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(6)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pseudodivaricata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pseudodivaricata</emphasis>
</taxonomicName>
(Lipsch.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Lipsch" authorityYear="1933" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera pseudodivaricata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pseudodivaricata">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera pseudodivaricata</emphasis>
</taxonomicName>
Lipsch., Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 42: 158. 1933. urn:lsid:ipni.org:names:77204019-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(7)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha pusilla" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pusilla">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha pusilla</emphasis>
</taxonomicName>
(Pall.) Nazarova
</paragraph>
<paragraph pageId="0" pageNumber="1">
(8)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha subacaulis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="subacaulis">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha subacaulis</emphasis>
</taxonomicName>
(Regel) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Regel" authorityYear="1880" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera austriaca var. subacaulis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="austriaca" variety="subacaulis">Scorzonera austriaca var. subacaulis</taxonomicName>
Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 323. 1880. urn:lsid:ipni.org:names:77204015-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(9)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha tau-saghyz</emphasis>
</taxonomicName>
(Lipsch. &amp; G.G.Bosse) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera tau-saghyz" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera tau-saghyz</emphasis>
</taxonomicName>
Lipsch. &amp; G.G.Bosse, Trudy Nauchno-Issl. Lab. Tresta &quot;Kauchukonos&quot; 4: 27. 1930. urn:lsid:ipni.org:names:77204010-1
</paragraph>
<paragraph pageId="0" pageNumber="1">
(10)
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha veresczaginii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="veresczaginii">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha veresczaginii</emphasis>
</taxonomicName>
(Kamelin &amp; S.V.Smirn.) Zaika, Sukhor. &amp; N.Kilian, comb. nov. ≡
<taxonomicName authorityName="Kamelin &amp; S. V. Smirn" authorityYear="2002" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera veresczaginii" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="veresczaginii">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera veresczaginii</emphasis>
</taxonomicName>
Kamelin &amp; S.V.Smirn., Turczaninowia 5(1): 17. 2002. urn:lsid:ipni.org:names:77204008-1
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution area.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">N Africa</emphasis>
: EG.
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Temperate</emphasis>
: AF; AR; AZ; CN; IL; IR; JO; KG; KP; KR; KZ; MN; RU (Asiatic part); SA; TJ; TM; UZ.
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Tropical</emphasis>
: PK.
<emphasis bold="true" pageId="0" pageNumber="1">Europe</emphasis>
: AL; AT; BA; BG; BY; CH; CZ; DE; FR; GR; HR; HU; IT; MD; ME; RO; RS; RU; SI; SK; UA.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The following species may belong to
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Takhtajaniantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Takhtajaniantha" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Takhtajaniantha</emphasis>
</taxonomicName>
, based on their morphological characters (they were included neither in the carpological nor in molecular investigations):
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="N. Kilian" authorityYear="2011" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera aniana" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="aniana">
<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera aniana</emphasis>
</taxonomicName>
N.Kilian,
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. grubovii" order="Asterales" pageId="0" pageNumber="1" rank="species" species="grubovii">
<emphasis italics="true" pageId="0" pageNumber="1">S. grubovii</emphasis>
</taxonomicName>
Lipsch.,
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. karataviensis" order="Asterales" pageId="0" pageNumber="1" rank="species" species="karataviensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. karataviensis</emphasis>
</taxonomicName>
Kult. (possibly a synonym of
<taxonomicName lsidName="T. tau-saghyz" pageId="0" pageNumber="1" rank="species" species="tau-saghyz">
<emphasis italics="true" pageId="0" pageNumber="1">T. tau-saghyz</emphasis>
</taxonomicName>
),
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. manshurica" order="Asterales" pageId="0" pageNumber="1" rank="species" species="manshurica">
<emphasis italics="true" pageId="0" pageNumber="1">S. manshurica</emphasis>
</taxonomicName>
Nakai,
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. pamirica" order="Asterales" pageId="0" pageNumber="1" rank="species" species="pamirica">
<emphasis italics="true" pageId="0" pageNumber="1">S. pamirica</emphasis>
</taxonomicName>
C.Shih,
<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. sinensis" order="Asterales" pageId="0" pageNumber="1" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. sinensis</emphasis>
</taxonomicName>
(Lipsch. &amp; Krasch.) Nakai
</paragraph>
</subSubSection>
</treatment>
</document>