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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.7.85204" ID-GBIF-Dataset="69e3a7a3-d279-4b63-8070-be3b1c813968" ID-Pensoft-Pub="2535-0730-1-133" ID-Pensoft-UUID="CF6673C9B1765D47B848F9C007505762" ID-ZooBank="24DF15ADF8A04086AD8C60AD39C8A4AA" ModsDocID="2535-0730-7-1-133" checkinTime="1683131477234" checkinUser="pensoft" docAuthor="Lehmann, Ingo &amp; Dalsgaard, Thure" docDate="2023" docId="7AC207F58B7C547BB9D46BDA6FC351A9" docLanguage="en" docName="EvolutSyst 7(1): 133-182" docOrigin="Evolutionary Systematics 7 (1)" docPubDate="2023-05-03" docSource="http://dx.doi.org/10.3897/evolsyst.7.85204" docTitle="Shimbania puguensis Lehmann &amp; Dalsgaard 2023, sp. nov." docType="treatment" docUuid="244C6A80-198E-439B-A6B1-8A9F47C6C556" docUuidSource="ZooBank" docVersion="2" id="CF6673C9B1765D47B848F9C007505762" lastPageNumber="133" masterDocId="CF6673C9B1765D47B848F9C007505762" masterDocTitle="Revision of Saalmulleria Mabille, 1891 (Lepidoptera, Metarbelidae) from Madagascar with the description of three new genera and fifteen new species" masterLastPageNumber="182" masterPageNumber="133" pageNumber="133" updateTime="1683132232210" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Saalmulleria Mabille, 1891 (Lepidoptera, Metarbelidae) from Madagascar with the description of three new genera and fifteen new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lehmann, Ingo</mods:namePart>
<mods:affiliation>Leibniz-Institute for the Analysis of Biodiversity Change, ztm, Museum der Natur, Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">lehmannshimoni@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dalsgaard, Thure</mods:namePart>
<mods:affiliation>Leibniz-Institute for the Analysis of Biodiversity Change, ztm, Museum der Natur, Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">t.dalsgaard@leibniz-lib.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>133</mods:start>
<mods:end>182</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/evolsyst.7.85204</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.7.85204</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-1-133</mods:identifier>
<mods:identifier type="ZooBank">24DF15ADF8A04086AD8C60AD39C8A4AA</mods:identifier>
<mods:identifier type="Pensoft-UUID">CF6673C9B1765D47B848F9C007505762</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:244C6A80-198E-439B-A6B1-8A9F47C6C556" httpUri="http://treatment.plazi.org/id/7AC207F58B7C547BB9D46BDA6FC351A9" lastPageNumber="133" pageId="0" pageNumber="133">
<subSubSection pageId="0" pageNumber="133" type="nomenclature">
<paragraph pageId="0" pageNumber="133">
<taxonomicName LSID="https://zoobank.org/244C6A80-198E-439B-A6B1-8A9F47C6C556" authority="Lehmann &amp; Dalsgaard, 2023" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Metarbelidae" genus="Shimbania" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Shimbania puguensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="puguensis" status="sp. nov.">Shimbania puguensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a. Shimbania pwaniensis sp. nov., holotype, male, Tanzania, Pwani Region, road from Dar es-Salaam to Chalinze, near the railway crossing, ca. 2 km from Ruvu River; b. S. puguensis sp. nov., holotype, male, Tanzania, Pwani Region, Pugu Forest; c. S. kaguruensis sp. nov., holotype, male, Tanzania, Morogoro Region, Kaguru Mountains; d. S. mbarikaensis sp. nov., holotype, male, Tanzania, Morogoro Region, Mbarika Mountains, south of Mahenge Forest; e. S. kerstinhempae sp. nov., holotype, female, Tanzania, Arusha Region, Usa River, Danish Volunteer Training Centre." httpUri="https://binary.pensoft.net/fig/845632" pageId="0" pageNumber="133">Figs 2b</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a. Shimbania puguensis sp. nov., paratype, male, Tanzania, Pwani Region, Kisarawe Forest; b. S. kaguruensis sp. nov., holotype, male, Tanzania, Morogoro Region, Kaguru Mountains; c. S. mbarikaensis sp. nov., holotype, male, Tanzania, Morogoro Region, Mbarika Mountains (also called Mbaraka Mountains), south of Mahenge Forest; d. S. wanjakinuthiaae sp. nov., holotype, male, Republic of South Africa, Province KwaZulu-Natal, ca. 5 km north of Hluhluwe, probably collected on Hluhluwe Farm." httpUri="https://binary.pensoft.net/fig/845639" pageId="0" pageNumber="133">, 9a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="materials_examined">
<paragraph pageId="0" pageNumber="133">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="133">
<materialsCitation collectingDate="2004-11-29" collectorName="Ph. Darge, I. Lehmann" country="Tanzania" elevation="132" location="Pugu Forest" specimenCount="1" stateProvince="Pwani Region" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">
<typeStatus>Holotype</typeStatus>
</emphasis>
male,
<collectingCountry name="Tanzania">Tanzania</collectingCountry>
,
<collectingRegion country="Tanzania" name="Pwani">Pwani Region</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:7AC207F58B7C547BB9D46BDA6FC351A9:0A14B94389284C8365DF8CD7674B0B5E" country="Tanzania" name="Pugu Forest" stateProvince="Pwani Region">Pugu Forest</location>
[Pugu Hills Nature Forest?],
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="132.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="132.0">132 m</elevation>
</quantity>
[
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.9" metricValueMax="2.48" metricValueMin="1.3199999999999998" unit="m" value="190.0" valueMax="248.0" valueMin="132.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.9" metricValueMax="2.48" metricValueMin="1.3199999999999998" unit="m" value="190.0" valueMax="248.0" valueMin="132.0">132-248 m</elevation>
</quantity>
],
<collectingDate value="2004-11-29">29 November 2004</collectingDate>
, ex coll.
<collectorName>Ph. Darge</collectorName>
, local collector [not named], genitalia slide number 18/012009
<collectorName>I. Lehmann</collectorName>
(ZSM)
</materialsCitation>
.
<materialsCitation collectingDate="2004-06-06" collectorName="Ph. Darge, I. Lehmann" country="Tanzania" elevation="267" latitude="-6.883333" location="Kisarawe Forest" longLatPrecision="1303" longitude="39.083332" specimenCount="1" stateProvince="Dar es-Salaam Region" typeStatus="Paratype">
<typeStatus>Paratype</typeStatus>
male,
<collectingCountry name="Tanzania">Tanzania</collectingCountry>
,
<collectingRegion country="Tanzania" name="Dar es Salaam">Dar es-Salaam Region</collectingRegion>
[incorrect
<collectingRegion country="Tanzania" name="Pwani">Pwani Region</collectingRegion>
],
<location LSID="urn:lsid:plazi:treatment:7AC207F58B7C547BB9D46BDA6FC351A9:B35D013A006602A662F9D8A37BC64E41" country="Tanzania" latitude="-6.883333" longLatPrecision="1303" longitude="39.083332" name="Kisarawe Forest" stateProvince="Dar es-Salaam Region">Kisarawe Forest</location>
,
<geoCoordinate degrees="06" direction="south" minutes="53.877" orientation="latitude" precision="1" value="-6.89795">06°53.877'S</geoCoordinate>
,
<geoCoordinate degrees="039" direction="east" minutes="05.189" orientation="longitude" precision="1" value="39.086483">039°05.189'E</geoCoordinate>
[
<geoCoordinate degrees="06" direction="south" minutes="53" orientation="latitude" precision="925" value="-6.883333">06°53'S</geoCoordinate>
,
<geoCoordinate degrees="39" direction="east" minutes="05" orientation="longitude" precision="925" value="39.083332">39°05'E</geoCoordinate>
],
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.67" unit="m" value="267.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.67" unit="m" value="267.0">267 m</elevation>
</quantity>
[incorrect, altitude is
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" unit="m" value="182.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" unit="m" value="182.0">182 m</elevation>
</quantity>
],
<collectingDate value="2004-06-06">06 June 2004</collectingDate>
,
<collectorName>Ph. Darge</collectorName>
[leg.], genitalia slide number 01/012009
<collectorName>I. Lehmann</collectorName>
(MWM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">Description.</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Head</emphasis>
: only ventrally dark chestnut, the rest is deep olive-buff (
<emphasis italics="true" pageId="0" pageNumber="133">
cf.
<taxonomicName authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Bombacaceae" genus="Shimbania" kingdom="Animalia" lsidName="Shimbania pwaniensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="pwaniensis">Shimbania pwaniensis</taxonomicName>
</emphasis>
sp. nov. that has only dark chestnut scales on the head), short scales with cream tips, glossy; eyes light brown-olive with black spots and surrounded by long hair-like scales of dark chestnut ventrally with a light lilac-vinaceous glint; a pair of pits is present on lower fronto-clypeus; pits behind labial palpi are tiny slits; antenna 0.40 length of forewing (0.35 in paratype;
<emphasis italics="true" pageId="0" pageNumber="133">cf.</emphasis>
shorter antennae in
<taxonomicName authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Bombacaceae" genus="Shimbania" kingdom="Animalia" lsidName="Shimbania pwaniensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="pwaniensis">
<emphasis italics="true" pageId="0" pageNumber="133">Shimbania pwaniensis</emphasis>
</taxonomicName>
sp. nov.), bipectinate, branches 3.0
<normalizedToken originalValue="×">x</normalizedToken>
width of shaft (2.5
<normalizedToken originalValue="×">x</normalizedToken>
in paratype), not scaled, not widely separated at base; shaft densely covered with ivory-yellow scales dorsally; labial palpi dark brown and long (longer than half of eye diameter).
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Thorax</emphasis>
: Patagia deep olive-buff, forming a collar ring, scales with light grey tips; tegulae with long hair-like dark chestnut scales with a light lilac-vinaceous glint. Metathorax with crest of deep olive-buff scales, with a glint. Hind legs deep olive-buff with fine hair-like scales with light grey tips, on lower part of tarsus dark chestnut dorsally; two pairs of long tibial spurs of unequal width and length, upper pair narrow
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
1.5 mm and 2.0 mm, lower pair of spurs slightly broader
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
1.0 mm and 1.3 mm long. Forewing length 22.5 mm and wingspan 49.0 mm in holotype; 23.0 mm and wingspan 50.0 mm in paratype. Forewing upperside dark olive-buff on inner half and along costa, olive-buff on outer half and below CuA2 with a light golden glint; below first half of 1A+2A a large dark chestnut patch with a light lilac-vinaceous glint; forewing with many narrow dark olive lines from costa to dorsum, many veins also narrowly dark olive including CuA2 that extends as a dark olive streak into the discal cell; a large and well visible dark olive subterminal patch, broadly Y-shaped, from R4 to near end of CuA1; termen without lunules; cilia short, 1.2 mm, deep olive-buff with grey tips. Underside of forewing cream-buff with weak brownish-olive lines and a golden glint. Hindwing upperside olive-buff with a light golden glint and with deep olive-buff lines; cilia as in forewing; underside as in forewing.
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Abdomen</emphasis>
: Mainly deep olive-buff mixed with ivory-yellow hair-like scales with a glint; abdominal tuft with hair-like scales of olive-buff and ivory-yellow, medium long, 1/4 length of abdomen. Genitalia with long uncus, 90% of length of whole gnathos (in holotype as well as paratype), narrow graben-like surface ventrally is present, well visible. Gnathos has gnathos arms that are small, one arm 20%-25% the size of valva (in holotype and paratype); upper part of the gnathos arm is a short band as long as 35% of basal width of valva, the lower part of the gnathal arm is small, and it does touch the other arm of broad triangular shape with a pronounced thorn-like structure and with its base 60% of the basal width of valva, with smaller thorns as well as a strongly serrate dorsal edge (in holotype and paratype); the gnathal arms are connected ventrally by a very narrow sclerotized band that is as broad as 20% of the transtilla and is not bifurcated in the middle. The Gnathos arms end well above the dorsal edge of the transtilla. The valva is triangular with a dorsal edge of 2.0
<normalizedToken originalValue="×">x</normalizedToken>
the length of uncus, ventral edge of valva slightly S-shaped with a tip that is pointed (instead of broadly rounded); sacculus pronounced, broad, sclerotized, 90% of length of ventral edge of valva; saccus absent; juxta well developed, with two broadly rounded lobes and a narrowly V-shaped emargination in between that is 20% the length of juxta, tips of lobes broadly rounded. Phallus very large, as broad as 35% of basal width of valva and 20% longer than costal width of valva, only slightly S-shaped and bent upwards at tip distally, vesica without cornuti.
</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="diagnosis">
<paragraph pageId="0" pageNumber="133">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="133">
<taxonomicName authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Bombacaceae" genus="Shimbania" kingdom="Animalia" lsidName="Shimbania puguensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">Shimbania puguensis</emphasis>
</taxonomicName>
sp. nov. can be separated from all other congeners by the long narrow uncus that has 90% of length of whole gnathos, the triangular valva with a pointed tip, the small gnathal arms of triangular shape with one thorn-like appendice in front of several smaller thorns and a strongly serrate dorsal edge. The latter character is similar in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. baginerichardi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="baginerichardi">
<emphasis italics="true" pageId="0" pageNumber="133">S. baginerichardi</emphasis>
</taxonomicName>
sp. nov. as well as in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. mbarikaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mbarikaensis">
<emphasis italics="true" pageId="0" pageNumber="133">S. mbarikaensis</emphasis>
</taxonomicName>
sp. nov., but both species have rectangular valvae with a broad and rounded end. The very narrow sclerotized band that connects the gnathal arms ventrally is only as broad as 20% of the transtilla and is not bifurcated in the middle; this is a unique character of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. puguensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">S. puguensis</emphasis>
</taxonomicName>
sp. nov. Furthermore, both males of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. puguensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">S. puguensis</emphasis>
</taxonomicName>
sp. nov. have a very contrasting forewing pattern and an olive-buff ground-colour on the outer half of forewing; the latter is short and broad in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. pwaniensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="pwaniensis">
<emphasis italics="true" pageId="0" pageNumber="133">S. pwaniensis</emphasis>
</taxonomicName>
sp. nov., its forewing is more elongated and cream-buff, and hence, more light brownish.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="distribution">
<paragraph pageId="0" pageNumber="133">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="133">
<taxonomicName authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Bombacaceae" genus="Shimbania" kingdom="Animalia" lsidName="Shimbania puguensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">Shimbania puguensis</emphasis>
</taxonomicName>
sp. nov. is known only from two lowland coastal forests that belong to the Zanzibar-Inhambane regional mosaic. A map based on forest data of the year 1970 and presented by
<bibRefCitation author="Hawthorne, WD" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="133" refId="B30" refString="Hawthorne, WD, 1984. Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1-332 [+ Appendix A-F]." title="Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1 - 332 [+ Appendix A-F]." year="1984">Hawthorne (1984)</bibRefCitation>
shows that the Pugu Hills forest assemblages, including Kisarawe forest, were once largely connected, but much of this forest area has been destroyed until 1981 for plantations and mining for Kaolin as Hawthorne noted when he did his field studies in this area. Some forest patches disappeared (
<emphasis italics="true" pageId="0" pageNumber="133">e.g.</emphasis>
, Kikoka Forest Reserve, Vikindu Forest Reserve) others are patchy, particularly north of Kisarawe town, and south as well as north of the railway line to Dar es-Salaam, the latter town is located
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
10-15 km to the East. Based on its distribution,
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="S. puguensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">S. puguensis</emphasis>
</taxonomicName>
sp. nov. can be classified as a lowland species that is endemic to the Pugu Hills forests, including Kisarawe Forest, most probably also to Kazimzumbwi Forest Reserve (
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
6-10 km southwest of Kisarawe town). These forests are located in the Usaramo floristic area
<emphasis italics="true" pageId="0" pageNumber="133">sensu</emphasis>
<bibRefCitation author="Hawthorne, WD" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="133" refId="B30" refString="Hawthorne, WD, 1984. Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1-332 [+ Appendix A-F]." title="Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1 - 332 [+ Appendix A-F]." year="1984">Hawthorne (1984)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="etymology">
<paragraph pageId="0" pageNumber="133">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="133">
<taxonomicName authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" family="Bombacaceae" genus="Shimbania" kingdom="Animalia" lsidName="Shimbania puguensis" order="Lepidoptera" pageId="0" pageNumber="133" rank="species" species="puguensis">
<emphasis italics="true" pageId="0" pageNumber="133">Shimbania puguensis</emphasis>
</taxonomicName>
is named for the Pugu Hills (Tanzania), an escarpment (altitude 93-310 m) that is built mainly of &quot;Pugu sandstone&quot; and other Neogene sediments (Tertiary), extending from south of Kisarawe town NNE for
<emphasis italics="true" pageId="0" pageNumber="133">ca.</emphasis>
10 km and comprises very different and diverse wetter and drier coastal forest types on ridge tops, slopes, valley bottoms, with swampy areas and riverine forests (
<bibRefCitation author="Hawthorne, WD" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="133" refId="B30" refString="Hawthorne, WD, 1984. Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1-332 [+ Appendix A-F]." title="Ecological and biogeographical patterns in the coastal forests of East Africa. A thesis submitted for the Doctor of Philosophy in the University of Oxford, St. Catherine's College, i-xiii + 1 - 332 [+ Appendix A-F]." year="1984">Hawthorne 1984</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="133">The gender of the new species name is feminine.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C061981257ECF464DDFCB73BDF6AA306">Flora Helvetica - Crassulaceae</mods:title>
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<mods:namePart id="DD93EA82425ABF145A21404711BDD219">Konrad Lauber</mods:namePart>
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<mods:namePart id="46245B02762865149AF100EBCC62F89E">Gerhart Wagner</mods:namePart>
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<mods:namePart id="13957B12A1D2193614193557F624FD1E">Andreas Gygax</mods:namePart>
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<mods:title id="E4093EFFB96C0A51B6468E4F2D107024">Flora Helvetica</mods:title>
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<taxonomicName id="929676B6FF256F77CA2964241E392E42" ID-CoL="79ZFN" authority="L." authorityName="L." class="Magnoliopsida" family="Crassulaceae" genus="Sedum" kingdom="Plantae" order="Saxifragales" pageNumber="194" phylum="Tracheophyta" rank="species" species="hybridum">
<emphasis id="B54974427671862ABAF93E8CC8195E4F" italics="true" pageNumber="196">Sedum hybridum</emphasis>
L.
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<paragraph id="C6852AFF7C1E0CB965405505E24B5D16" pageNumber="196">
Artbeschreibung:
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wie
<taxonomicName id="8401F3C831977321A88DD0C3703860F7" class="Magnoliopsida" family="Crassulaceae" genus="Sedum" kingdom="Plantae" order="Saxifragales" pageNumber="194" phylum="Tracheophyta" rank="species" species="spurium">
<emphasis id="6D5AB140937AD34BEC0CB7D358789E82" italics="true" pageNumber="196">S. spurium</emphasis>
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, aber
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<normalizedToken id="14363F4B3E2C1159667FF531FAAD9BBA" originalValue="wechselständig" pageNumber="194">wechselstaendig</normalizedToken>
.
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gelb.
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<quantity id="93DCC4E6274D97C5A4F18BD5234D90DD" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageNumber="194" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6-9 mm</quantity>
lang,
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ausgebreitet.
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<normalizedToken id="D6209B83852EC02799299AE78EFA14D5" originalValue="Blütezeit" pageNumber="194">Bluetezeit</normalizedToken>
: 5-6
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<paragraph id="1196AFF7BD2730F329BDBA1BCFE35280" pageNumber="196">Standort und Verbreitung in der Schweiz: Zierpflanze, verwildert /</paragraph>
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<paragraph id="5BCF3E8A44582CC8A422B4CD0B0A45B3" pageNumber="196">Verbreitung global: Asiatisch</paragraph>
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<paragraph id="A8A9F2EAE41BAF6EF8E4F4E2093B8373" pageNumber="196">
<normalizedToken id="32D687A1A7A69A3FCF117B3506E370AD" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="BC94734DA7CE341375B34525504571CC" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="196" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="FA876B80D06F0B5310093406F82ABA59" pageNumber="196">
<table id="A63642E7A2FC53B161D914DFCC502BD1" class="table" inLine="true" pageNumber="196">
<tr id="DA3ED6A0D971174C546301A73808C072" pageNumber="196">
<th id="CDAD163BC0853F9BC0086E256919C94C" colspan="2" pageNumber="196">
<emphasis id="2377B8B3B49AD0A5AF25E526ABA6A912" bold="true" pageNumber="196">Bodenfaktoren</emphasis>
</th>
<th id="B8356CBFC4B1DB51A5B25C4E4B92362B" colspan="2" pageNumber="196">
<emphasis id="BF021F45D86E8C822268CB54E6F91F31" bold="true" pageNumber="196">Klimafaktoren</emphasis>
</th>
<th id="A746568CA6C89E97FAB670E2C568BD7F" colspan="2" pageNumber="196">
<emphasis id="403A86A8A823819C3AECFB0D4F2EDD74" bold="true" pageNumber="196">Salztoleranz</emphasis>
</th>
</tr>
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<td id="C671CF59F46ABCF22E8B119E72F7CFE4" pageNumber="196">Feuchtezahl F</td>
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<normalizedToken id="CA3BC8C513457FA8855F4A037A37B118" originalValue="mässig">maessig</normalizedToken>
trocken; Feuchtigkeit stark wechselnd (mehr als
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2 Stufen)
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<td id="6A2D3077F03FD329639854F32A8B9576" pageNumber="196">Lichtzahl L</td>
<td id="58EE5B20AA7147072889B1F292950317" pageNumber="196" style="text-align:left;width:60px">hell</td>
<td id="A6D2DF8D394ADFEBC7ED2D68F075ABD1" pageNumber="196">Salzzeichen</td>
<td id="83CFB2DA3CD061A6006070541270CDA9" pageNumber="196" style="text-align:left;width:60px">--</td>
</tr>
<tr id="C5BF8ECB00DB5C494CB2F84250385E5D" pageNumber="196">
<td id="59B282B1FFD5BD6287C8D393866336F5" pageNumber="196">Reaktionszahl R</td>
<td id="8FE5229FA27495231BD2A3C387CBD18D" pageNumber="196" style="text-align:left">neutral bis basisch (pH 5.5-8.5)</td>
<td id="E188D5DE289AC01FE4A452A73AB96322" pageNumber="196">Temperaturzahl T</td>
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kollin (
<normalizedToken id="6332526071B6B056A46BEE6767BD67CC" originalValue="Laubmischwälder">Laubmischwaelder</normalizedToken>
mit Eichen)
</td>
</tr>
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<td id="4608923CE30D416199711EEA4712DAC7" pageNumber="196">
<normalizedToken id="D950976BD66A478EBDC84AC6DF442858" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
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<normalizedToken id="765BD582D13A3BDE6CFC573704A88FEA" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="00F8D1E8D1FC7ECBCB46F47319DD05AC" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
bis
<normalizedToken id="D36237F1EA05D360B617FEC9B013F04B" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="22B977F5286BDE44DB2E50D3E41A3E35" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
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<normalizedToken id="6B5A95C14AFF6CBE6430C416683B27CD" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="82905BF8D4E58B509AAD51EC514AF6AC" pageNumber="196" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="1A66B4DFD332AC3602B81047E8F8631D" pageNumber="196" type="vernacular_names">
<paragraph id="CB5ADD70FBE3431D688E7EE2CE424234" pageNumber="196">
Volksname Deutscher Name:
<vernacularName id="C8C4FD629B668E413CEC0800D3F4ED1F" language="deu" pageNumber="196">Bastard-Fettkraut</vernacularName>
Nom
<normalizedToken id="10DFE0E96BAC78DC50F52E1284C1A306" originalValue="français">francais</normalizedToken>
:
<vernacularName id="F0B53F6A22D0E607367C09FA67C6133C" language="fra" pageNumber="196">Orpin hybride</vernacularName>
</paragraph>
</subSubSection>
</treatment>
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112
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Tribe
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Sama, 2008: 222 [stem: Lygr-]. Type genus:
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, 1872.
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<mods:namePart>Bass, Amber</mods:namePart>
<mods:affiliation>Department of Entomology, University of Manitoba, 12 Dafoe Road, Winnipeg, Manitoba, Canada &amp; Current address: Agriculture and Agri-Food Canada, Canadian National Collection of Insects, 960 Carling Avenue, Ottawa, Ontario, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morgan, Katherine</mods:namePart>
<mods:affiliation>Department of Entomology, University of Manitoba, 12 Dafoe Road, Winnipeg, Manitoba, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-18</mods:number>
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<mods:detail type="volume">
<mods:number>1089</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1089.72645</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1089-125</mods:identifier>
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<taxonomicName authority="Gibbs 2016" authorityName="Gibbs" authorityYear="2016" class="Insecta" family="Halictidae" genus="Lasioglossum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasioglossum (Dialictus) dominicense" order="Hymenoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="dominicense" subGenus="Dialictus">Lasioglossum (Dialictus) cf. dominicense Gibbs 2016</taxonomicName>
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<subSubSection pageId="0" pageNumber="125" type="description">
<paragraph pageId="0" pageNumber="125">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lasioglossum (Dialictus) cf. dominicense Gibbs, 2016, female A dorsal habitus B head, frontal view C lateral habitus." figureDoi="10.3897/zookeys.1089.72645.figure8" httpUri="https://binary.pensoft.net/fig/660574" pageId="0" pageNumber="125">Fig. 8</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="125" type="reference_group">
<paragraph pageId="0" pageNumber="125">
<taxonomicName authorityName="Gibbs" authorityYear="2016" class="Insecta" family="Halictidae" genus="Lasioglossum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasioglossum (Dialictus) dominicense" order="Hymenoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="dominicense" subGenus="Dialictus">Lasioglossum (Dialictus) dominicense</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2016.180" author="Gibbs, J" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="125" pagination="1 - 50" refId="B45" refString="Gibbs, J, 2016. Bees of the family Halictidae Thomson, 1869 from Dominica, Lesser Antilles (Hymenoptera: Apoidea). European Journal of Taxonomy 180: 1 - 50, DOI: https://doi.org/10.5852/ejt.2016.180" title="Bees of the family Halictidae Thomson, 1869 from Dominica, Lesser Antilles (Hymenoptera: Apoidea)." url="https://doi.org/10.5852/ejt.2016.180" volume="180" year="2016">Gibbs 2016</bibRefCitation>
: 6-11, 42-43.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="materials_examined">
<paragraph pageId="0" pageNumber="125">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="125">
<materialsCitation collectingDate="2009-05-08" collectingDateMax="2009-06-29" collectingDateMin="2009-05-08" collectingMethod="Louvette trap, UV light trap, Grand Anse trap" collectorName="M. L. Gimmel, C. A. Maier, R. C. Winton, E. A. Ivie" country="Saint Lucia" elevation="38" latitude="14.0052" location="Dauphin District" longLatPrecision="7" longitude="-60.8973" specimenCount="2" specimenCount-female="2" stateProvince="Dauphin District">
<emphasis bold="true" pageId="0" pageNumber="125">
<collectingCountry name="Saint Lucia">Saint Lucia</collectingCountry>
</emphasis>
•
<emphasis bold="true" pageId="0" pageNumber="125">
<collectingRegion country="Saint Lucia" name="Dauphin">Dauphin District</collectingRegion>
</emphasis>
•
<collectingMethod>Louvette trap</collectingMethod>
site,
<geoCoordinate degrees="13.9689" direction="north" orientation="latitude" precision="5" value="13.9689">13.9689</geoCoordinate>
,
<geoCoordinate degrees="60.8859" direction="west" orientation="longitude" precision="5" value="-60.8859">-60.8859</geoCoordinate>
,
<collectingDate value="2009-06-25" valueMax="2009-06-29" valueMin="2009-06-25">25-29.VI.2009</collectingDate>
, leg.
<collectorName>M.L. Gimmel</collectorName>
and
<collectorName>C.A. Maier</collectorName>
,
<collectingMethod>UV light trap</collectingMethod>
(
<specimenCount type="female">1 ♀</specimenCount>
MTEC) •
<collectingMethod>Grand Anse trap</collectingMethod>
site,
<geoCoordinate degrees="14.0052" direction="north" orientation="latitude" precision="5" value="14.0052">14.0052</geoCoordinate>
,
<geoCoordinate degrees="60.8973" direction="west" orientation="longitude" precision="5" value="-60.8973">-60.8973</geoCoordinate>
,
<specimenCount type="generic">
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.8" unit="m" value="38.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.8" unit="m" value="38.0">38 m</elevation>
</quantity>
.
</specimenCount>
<collectingDate value="2009-05-08" valueMax="2009-05-17" valueMin="2009-05-08">8-17.V.2009</collectingDate>
, leg.
<collectorName>R.C. Winton</collectorName>
and
<collectorName>E.A. Ivie</collectorName>
(
<specimenCount type="female">1 ♀</specimenCount>
MTEC)
</materialsCitation>
•
<materialsCitation collectingDate="2009-05-08" collectingDateMax="2009-06-06" collectingDateMin="2009-05-08" collectingMethod="pan traps" collectorName="C. M. Delphia, J. B. Runyon, R. C. Winton, Malaise" country="Saint Lucia" elevation="1" latitude="13.83242" location="Fond Bay" longLatPrecision="5" longitude="-60.8859" municipality="Escap Community Trail" specimenCount="2" specimenCount-female="2" stateProvince="Micoud District">
<emphasis bold="true" pageId="0" pageNumber="125">
<collectingRegion country="Saint Lucia" name="Micoud">Micoud District</collectingRegion>
</emphasis>
•
<collectingMunicipality>Escap Community Trail</collectingMunicipality>
to
<location LSID="urn:lsid:plazi:treatment:7AC2F013E4DCC297C9795E92A47DB91F:BE5536B80F6F86F5A2EBEA0DCCA4A8AF" country="Saint Lucia" latitude="13.83242" longLatPrecision="5" longitude="-60.8859" municipality="Escap Community Trail" name="Fond Bay" stateProvince="Micoud District">Fond Bay</location>
beach,
<geoCoordinate degrees="13.8324" direction="north" orientation="latitude" precision="5" value="13.8324">13.8324</geoCoordinate>
,
<geoCoordinate degrees="60.8986" direction="west" orientation="longitude" precision="5" value="-60.8986">-60.8986</geoCoordinate>
to
<geoCoordinate degrees="13.8316" direction="north" orientation="latitude" precision="5" value="13.8316">13.8316</geoCoordinate>
,
<geoCoordinate degrees="60.893" direction="west" orientation="longitude" precision="55" value="-60.893">-60.893</geoCoordinate>
, 46 m to
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectingDate value="2009-05-08">8.V.2009</collectingDate>
, leg.
<collectorName>C.M. Delphia</collectorName>
and
<collectorName>J.B. Runyon</collectorName>
,
<collectingMethod>pan traps</collectingMethod>
(
<specimenCount type="female">1 ♀</specimenCount>
MTEC) • Escap Community,
<geoCoordinate degrees="13.83242" direction="north" orientation="latitude" precision="1" value="13.83242">13.83242</geoCoordinate>
,
<geoCoordinate degrees="60.8859" direction="west" orientation="longitude" precision="5" value="-60.8859">-60.8859</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="4.6" unit="m" value="46.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="4.6" unit="m" value="46.0">46 m</elevation>
</quantity>
,
<collectingDate value="2009-05-22" valueMax="2009-06-06" valueMin="2009-05-22">22.V-6.VI.2009</collectingDate>
, leg.
<collectorName>R.C. Winton</collectorName>
,
<collectorName>Malaise</collectorName>
trap (
<specimenCount type="female">1 ♀</specimenCount>
WRME)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1089.72645.figure8" httpUri="https://binary.pensoft.net/fig/660574" pageId="0" pageNumber="125" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 8.</emphasis>
<taxonomicName authorityName="Gibbs" authorityYear="2016" class="Insecta" family="Halictidae" genus="Lasioglossum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasioglossum (Dialictus) dominicense" order="Hymenoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="dominicense" subGenus="Dialictus">Lasioglossum (Dialictus) cf. dominicense</taxonomicName>
Gibbs, 2016, female
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="125">C</emphasis>
lateral habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="notes">
<paragraph pageId="0" pageNumber="125">Notes.</paragraph>
<paragraph pageId="0" pageNumber="125">
We ascribe the Saint Lucia material to
<taxonomicName lsidName="L. dominicense" pageId="0" pageNumber="125" rank="species" species="dominicense">
<emphasis italics="true" pageId="0" pageNumber="125">L. dominicense</emphasis>
</taxonomicName>
without supporting evidence to the contrary. Although there seems to be some pattern of distinct species across islands in the Lesser Antilles, we are unable to confidently differentiate females of
<taxonomicName lsidName="L. dominicense" pageId="0" pageNumber="125" rank="species" species="dominicense">
<emphasis italics="true" pageId="0" pageNumber="125">L. dominicense</emphasis>
</taxonomicName>
from Saint Lucia and Dominica at this time. As a lowland species occurring near the beach, it is most consistent with a multi-island distribution. Additional comparative study including males, specimens from Martinique, and molecular data would be useful.
</paragraph>
</subSubSection>
</treatment>
</document>

173
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<taxonomicName id="7B0447CEDB7BC7183801683D1C1EE40D" ID-CoL="H2CN" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria taeniifera" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="taeniifera">Artoria taeniifera Simon, 1909</taxonomicName>
Figs 1D, 37, 38
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, 48G
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Artoria
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<taxonomicName id="7A856AC756E7F699D3C42E61B3DA13BB" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria taeniifera" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="taeniifera">Artoria taeniifera</taxonomicName>
Simon, 1909: 193-194, fig. 11. -
<bibRefCitation id="DFD098C57CE490FFDA7B01EDD34628C9" author="Rainbow, WJ" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="169" pagination="107 - 319" title="A census of Australian Araneidae." url="https://doi.org/10.3853/j.0067-1975.9.1911.928" volume="9" year="1911">Rainbow 1911</bibRefCitation>
: 275; Bonnet 1955: 751;
<bibRefCitation id="D54B0B2F2702416AD6D13381615ABFCD" author="McKay, RJ" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="169" pagination="375 - 398" title="The wolf spiders of Australia (Araneae: Lycosidae): 1. The bicolor group." volume="16" year="1973">McKay 1973</bibRefCitation>
: 380;
<bibRefCitation id="45D9963587B169B2A0E4C3CB7D1EE8D4" pageId="0" pageNumber="169">McKay 1985</bibRefCitation>
: 74;
<bibRefCitation id="804B109F58C8CBC0DC21E27D90B94521" author="Framenau, VW" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="169" pagination="209 - 235" title="Review of the wolf spider genus Artoria Thorell (Araneae: Lycosidae)." url="https://doi.org/10.1071/IT01028" volume="16" year="2002">Framenau 2002</bibRefCitation>
: 226-227, figs 21A-B, 22;
<bibRefCitation id="E612CF83C4150777C6D191DDA2CFA075" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="169" pagination="265 - 292" title="The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae)." url="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" volume="22" year="2005">Framenau 2005</bibRefCitation>
: 288-289: figs 13
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, 14.
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<paragraph id="35E3ED454C60248231BEA44C8214D011" pageId="0" pageNumber="169">
<taxonomicName id="159D84E334DC81CBE988649B49DE95CF" class="Arachnida" family="Lycosidae" genus="Artoriella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Artoriella taeniifera" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="taeniifera">Artoriella taeniifera</taxonomicName>
(Simon).-
<bibRefCitation id="06FDA8825AE7C085E5FE8608B16E821B" pageId="0" pageNumber="169">Roewer 1955</bibRefCitation>
: 233; Rower 1960: 563.
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<paragraph id="0A6EF97F876F179BC9CB9F66A911CF7F" pageId="0" pageNumber="169">Material examined.</paragraph>
<paragraph id="AD940E5C36B59E9BDC004948F8F14036" pageId="0" pageNumber="169">
Holotype female, Bunbury ('Station 142') (
<geoCoordinate id="C15AD4E1FB685F402AABD23E8FB34337" direction="south" orientation="latitude" precision="925" value="-33.333332">33°20'S</geoCoordinate>
<geoCoordinate id="E85A208302A0C11F70204AAA852E3744" direction="east" orientation="longitude" precision="925" value="115.65">115°39'E</geoCoordinate>
, Western Australia, AUSTRALIA], collected during the 'Hamburger südwest-australische Forschungsreise 1905' (ZMB 10549) (examined).
</paragraph>
</subSubSection>
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<paragraph id="9AF0AAFF0801244D027985DEA053B5E4" pageId="0" pageNumber="169">Other material examined.</paragraph>
<paragraph id="A6368D8955F5D7819BE3BC3132C1FB67" pageId="0" pageNumber="169">
AUSTRALIA: New South Wales: 1 female, Round Hill, Euabalong,
<geoCoordinate id="CE9E105EDA84F540F1DACD82466CFDEA" direction="south" orientation="latitude" precision="925" value="-32.95">32°57'S</geoCoordinate>
,
<geoCoordinate id="28B8924D4FF5DED7CA88FF3231CF8814" direction="east" orientation="longitude" precision="925" value="146.15">146°09'E</geoCoordinate>
(AM KS50283).
</paragraph>
</subSubSection>
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<paragraph id="DA3000F5AF9463A3427BB28B1C9E1804" pageId="0" pageNumber="169">Diagnosis.</paragraph>
<paragraph id="820A6EC4403B5A53937029DF55A81770" pageId="0" pageNumber="169">
The male of
<taxonomicName id="D789F523BC52A32106537C72F480DC5A" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
is most similar to that of
<taxonomicName id="47250EB866186F62D338EE7312CC3F97" lsidName="A. belfordensis" pageId="0" pageNumber="169" rank="species" species="belfordensis">A. belfordensis</taxonomicName>
sp. n. based on the structure of the basoembolic apophysis, which is basally drawn out into a tip or sharp edge and apically truncated. However, these species differ in the shape of the tegular apophysis, which is two-pronged in
<taxonomicName id="C9BE1A28D1F744367E31CB388BE90446" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
and three lobed in
<taxonomicName id="DE8B35BB7234FAA839903576874063F7" lsidName="A. belfordensis" pageId="0" pageNumber="169" rank="species" species="belfordensis">A. belfordensis</taxonomicName>
sp. n. The epigyne of female
<taxonomicName id="179382250249057E07D7A1187A758EAB" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
is distinctive within
<taxonomicName id="32AE77B7280E77ABD902F86305C0C320" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">Artoria</taxonomicName>
, with its atrium semicircular along its anterior border and strongly sclerotised posterior edges that point medially.
</paragraph>
</subSubSection>
<subSubSection id="15634D22F330019C57DBB8C35896DF8A" pageId="0" pageNumber="169" type="description">
<paragraph id="6C9AD5B95C3F9437906BEDDA2E252B18" pageId="0" pageNumber="169">Description.</paragraph>
<paragraph id="6CBF6020D2EBCC477F5C02B20788940E" pageId="0" pageNumber="169">
The female of
<taxonomicName id="723A88F23EAAA1E5938505B79E979E98" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
has been described in detail (
<bibRefCitation id="27AEF6319AE3AAB6F1BA27A9255B1094" author="Framenau, VW" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="169" pagination="209 - 235" title="Review of the wolf spider genus Artoria Thorell (Araneae: Lycosidae)." url="https://doi.org/10.1071/IT01028" volume="16" year="2002">Framenau 2002</bibRefCitation>
;
<bibRefCitation id="AB6D36B94FC7A275F7ACC291482C10F0" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="169" pagination="265 - 292" title="The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae)." url="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" volume="22" year="2005">2005</bibRefCitation>
). A diagnosis and diagnostic images of the single female found in NSW (Figs 1D, 38C, D, G, H) are provided here to facilitate identification. The male is described for the first time based on a specimen collected in Nedlands (
<geoCoordinate id="90D480D89D5FEC246D1DF9C168AB756C" direction="south" orientation="latitude" precision="925" value="-31.983334">31°59'S</geoCoordinate>
,
<geoCoordinate id="8B324084DA870B97C0411AA58BB378A2" direction="east" orientation="longitude" precision="925" value="115.8">115°48'E</geoCoordinate>
, Western Australia).
</paragraph>
<paragraph id="A6C6D1B3B4FDFED064349B12B3F80E60" pageId="0" pageNumber="169">Male (based on WAM 98/2179).</paragraph>
<paragraph id="092F205C3E598A0AE836EE9DF5245B26" pageId="0" pageNumber="169">Total length 4.8.</paragraph>
<paragraph id="DAF69B17ACAFD3977EAB71A3F7A5DD64" pageId="0" pageNumber="169">Prosoma. Length 2.6, width 2.0; carapace brown with darker radial pattern; light brown elongated V-shaped median band and indistinct light brown submarginal bands (Fig. 38A); sternum glabrous light brown (Fig. 38B).</paragraph>
<paragraph id="46A200289D0B3818DF5F25D55258FA25" pageId="0" pageNumber="169">Eyes. Diameter of AME: 0.09; ALE: 0.09; PME: 0.25; PLE: 0.21.</paragraph>
<paragraph id="642FC5E34E2759B698379F11AB11F5BE" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, evenly spaced.</paragraph>
<paragraph id="AB4312171DB4752811E3D2CF33FB5779" pageId="0" pageNumber="169">Chelicerae. Dark brown.</paragraph>
<paragraph id="9F9D98C32F1B3F486143E5668CF4D457" pageId="0" pageNumber="169">Labium. Brown, with lighter anterior rim (Fig. 38B)</paragraph>
<paragraph id="ACB1DF2A80AF98407A7F1228EDDAADEA" pageId="0" pageNumber="169">Pedipalp coxae. Brown, with lighter anterior rim (Fig. 38B).</paragraph>
<paragraph id="6578A030F455A14D9AD20B49BECCB237" pageId="0" pageNumber="169">Legs. Yellow brown with darker annulations; tibiae, metatarsi and tarsi darker, less annulated (Fig. 38A).</paragraph>
<paragraph id="9D7B4F17109F020DFE096EA5035F49BD" pageId="0" pageNumber="169">Opisthosoma. Length 2.1, width 1.6; olive-brown with irregular darker pattern, centrally darker in particular in posterior half, light yellow-brown anterior cardiac mark (Fig. 38A). Venter pale yellow with darker central pattern (Fig. 38B); spinnerets dark grey.</paragraph>
<paragraph id="1008F5867F5A6A6A34BE2C523DC8407E" pageId="0" pageNumber="169">Pedipalps. Tibia slightly longer than broad; cymbium apically with ca. 10 stronger setae (Fig. 38E, F); dorsal scopula patch absent; tegular apophysis distally two-pronged, (Fig. 38F); palea about twice as long as wide; basoembolic apophysis about long than broad and apically truncated; embolus apically with wide tip; terminal apophysis apically widely triangular and strongly sclerotised (Fig. 48G).</paragraph>
<caption id="8AA6E9E88E969E18B037FC09AEF7372B" pageId="0" pageNumber="169">
<paragraph id="CFB11B21391896A1590A3E6518AEE95D" pageId="0" pageNumber="169">
Figure 38.
<taxonomicName id="A637EE937B478908456AE6F3667FB5E0" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria taeniifera" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="taeniifera">Artoria taeniifera</taxonomicName>
Simon, 1909, male (WAM 98/2179), female (AM KS50283): A, male habitus, dorsal view; B, male habitus, ventral view; C, female habitus, dorsal view; D, female habitus, ventral view; E, male pedipalp, ventral view; F, male pedipalp, retrolateral view; G, epigyne, ventral view; H, epigyne, dorsal view. Scale bars: habitus 1.0 mm; pedipalp, epigyne 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="91A729310C713885CA4CE3E7E3D5F9ED" pageId="0" pageNumber="169" type="life history and habitat preferences">
<paragraph id="43E71E83A39D9C01F37D9632FF717ECD" pageId="0" pageNumber="169">Life history and habitat preferences.</paragraph>
<paragraph id="4DA20644D2E5E48BE9A13CCF84DF622B" pageId="0" pageNumber="169">
The habitat of
<taxonomicName id="57F1CBBA472CD0DC0E93A4B473781ED3" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
in NSW is unknown. In Western Australia, from where most records of this species have been reported,
<taxonomicName id="7ABB22667EECA509C782246A2A67F9D9" lsidName="A. taeniifera" pageId="0" pageNumber="169" rank="species" species="taeniifera">A. taeniifera</taxonomicName>
prefers open habitats such as coastal dunes, gardens and open woodlands. The single female from NSW was found in March suggesting the species to be autumn-mature. In south-western Western Australia, mature spiders have been found in winter and early spring.
</paragraph>
</subSubSection>
<subSubSection id="97AD2CE86D6A58EC26ADD341A5460925" pageId="0" pageNumber="169" type="distribution">
<paragraph id="75EBD46B54BA5679C1B1D1BBE47E6986" pageId="0" pageNumber="169">Distribution.</paragraph>
<paragraph id="7039EF2A2E2CCBFCF1AD7CB749B2981F" pageId="0" pageNumber="169">
<taxonomicName id="9673BA40B51AF49B28D032E52A79D16B" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria taeniifera" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="taeniifera">Artoria taeniifera</taxonomicName>
has only been found once in NSW, centrally in the Cobar Plain (COP) IBRA region (Fig. 37). There is a second isolated record of the species in South Australia (Framenau, unpublished data). It is unknown if these represent persistent populations. The species occurs commonly in south-western Western Australia, from where it was initially described (
<bibRefCitation id="ABDBB1BAF5EE4430C8389279D522229C" author="Framenau, VW" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="169" pagination="209 - 235" title="Review of the wolf spider genus Artoria Thorell (Araneae: Lycosidae)." url="https://doi.org/10.1071/IT01028" volume="16" year="2002">Framenau 2002</bibRefCitation>
;
<bibRefCitation id="0A16EE28BBA46E23D442CFD2F9CCC858" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="169" pagination="265 - 292" title="The wolf spider genus Artoria Thorell in Australia: new synonymies and generic transfers (Araneae, Lycosidae)." url="https://doi.org/10.18195/issn.0312-3162.22(4).2005.265-292" volume="22" year="2005">2005</bibRefCitation>
; unpublished data).
</paragraph>
</subSubSection>
</treatment>
</document>