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<document id="D01D5423ABEAF009D2888C32B9EBAD49" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8151" ID-PMC="PMC4867695" ID-Pensoft-Pub="1314-2828-4-8151" ID-PubMed="27226759" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8151" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" checkinTime="1461288934892" checkinUser="pensoft" docAuthor="Broad, Gavin R., Shaw, Mark R. &amp; Godfray, H. Charles J." docDate="2016" docId="7AA886E7E291348120A02DB3BF21CE84" docLanguage="en" docName="BiodivDatJour 4: e8151" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8151" docTitle="Wachsmannia Szepligeti 1900" docType="treatment" docVersion="5" lastPageNumber="8151" masterDocId="EC41FFB1FFD9782CFFAEFFF87E2EFF92" masterDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" masterLastPageNumber="8151" masterPageNumber="8151" pageNumber="8151" updateTime="1668123791244" updateUser="ExternalLinkService">
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<mods:title id="309DF6EB15A3D322512BA7EE3DDCA4EB">Checklist of British and Irish Hymenoptera - Braconidae</mods:title>
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<mods:namePart id="D06230984C891CE320E6020650CD8CF3">Broad, Gavin R.</mods:namePart>
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<mods:namePart id="DDAE34D371C90B9B6AC6C0CB35C039AA">Shaw, Mark R.</mods:namePart>
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<mods:namePart id="65C76C4904DA81043682E1A1EAF261B3">Godfray, H. Charles J.</mods:namePart>
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<mods:title id="7E69CCA50831B166B6B558A27E8D9D52">Biodiversity Data Journal</mods:title>
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<mods:date id="11C01906780F6DF80FF1C96DEC786839">2016</mods:date>
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<taxonomicName id="0C262EFD87DAACC0B8C6061E0C80CCEC" ID-CoL="87D2" authority="Szepligeti, 1900" authorityName="Szepligeti" authorityYear="1900" class="Insecta" family="Braconidae" genus="Wachsmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wachsmannia" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">
Wachsmannia
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, 1900
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<paragraph id="852C85001E237402147A59D63A9591D3" pageId="0" pageNumber="8151">Notes</paragraph>
<paragraph id="EB20CA78940E1A0902EC75BF6684B29D" pageId="0" pageNumber="8151">
<bibRefCitation id="25E9F9127C0D7FF56B5AFDCB3E5B21A5" author="Belokobylskij, S. A." journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="8151" pagination="900 - 928" title="[On the classification and phylogeny of braconid wasps subfamilies Doryctinae and Exothecinae (Hymenoptera, Braconidae). I. Classification. Entomological Review 72: 109 - 137.] (in Russian)" volume="71" year="1992">Belokobylskij (1992)</bibRefCitation>
synonymised
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under
<taxonomicName id="A49EDDE26EF5E49F1D3F2BE2D13EC30C" class="Insecta" family="Braconidae" genus="Ontsira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontsira" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Ontsira</taxonomicName>
but van
<bibRefCitation id="F5C95223DC914A54D005F13CEB7E627B" author="Achterberg, C. van" journalOrPublisher="Zoologische Mededelingen, Leiden" pageId="0" pageNumber="8151" pagination="131 - 138" title="New combinations of names for Palaearctic Braconidae (Hymenoptera)" volume="69" year="1995">Achterberg (1995)</bibRefCitation>
disagreed and synonymised
<taxonomicName id="A128D230B576A3D42B0D1C11DF4A3A15" class="Insecta" family="Braconidae" genus="Wachsmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wachsmannia" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Wachsmannia</taxonomicName>
under
<taxonomicName id="19AA901006844319F98960A4895DC8AB" class="Insecta" family="Braconidae" genus="Hypodoryctes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypodoryctes" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Hypodoryctes</taxonomicName>
Kokujev, 1900. Van Achterberg (pers. comm. and in Fauna Europaea) now regards
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as a separate genus again.
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.585.7503" ID-GBIF-Dataset="284e15e6-931f-4863-b037-68a83c8b53a8" ID-PMC="PMC4857038" ID-Pensoft-Pub="1313-2970-585-51" ID-PubMed="27199604" ID-ZBK="B914D8CF92A14C948EDC7CE8B0202076" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-585-51" ModsDocOrigin="ZooKeys 585" ModsDocTitle="Megastigmus seed chalcids (Hymenoptera, Torymidae) radiated much more on Angiosperms than previously considered. I- Description of 8 new species from Kenya, with a key to the females of Eastern and Southern Africa" checkinTime="1461901611862" checkinUser="pensoft" docAuthor="Roques, Alain, Copeland, Robert S., Soldati, Laurent, Denux, Olivier &amp; Auger-Rozenberg, Marie-Anne" docDate="2016" docId="7AA8E5B81F74F5F7FDEDD841A12288C1" docLanguage="en" docName="ZooKeys 585: 51-124" docOrigin="ZooKeys 585" docSource="http://dx.doi.org/10.3897/zookeys.585.7503" docTitle="Megastigmus icipeensis Roques &amp; Copeland, sp. n." docType="treatment" docUuid="4EE8D716-63AB-48F4-8B9E-E6277136C6D8" docUuidSource="ZooBank" docVersion="5" lastPageNumber="84" masterDocId="FF812555FFDD75799D3DFF99922B9D29" masterDocTitle="Megastigmus seed chalcids (Hymenoptera, Torymidae) radiated much more on Angiosperms than previously considered. I- Description of 8 new species from Kenya, with a key to the females of Eastern and Southern Africa" masterLastPageNumber="124" masterPageNumber="51" pageNumber="81" updateTime="1668163066542" updateUser="ExternalLinkService">
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<mods:namePart>Soldati, Laurent</mods:namePart>
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<mods:namePart>Denux, Olivier</mods:namePart>
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<paragraph pageId="30" pageNumber="81">Taxon classification Animalia Hymenoptera Torymidae</paragraph>
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<taxonomicName LSID="http://zoobank.org/4EE8D716-63AB-48F4-8B9E-E6277136C6D8" authority="Roques &amp; Copeland" class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus icipeensis" order="Hymenoptera" pageId="30" pageNumber="81" phylum="Arthropoda" rank="species" species="icipeensis">Megastigmus icipeensis Roques &amp; Copeland</taxonomicName>
<taxonomicNameLabel pageId="30" pageNumber="81">sp. n.</taxonomicNameLabel>
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<subSubSection pageId="30" pageNumber="81" type="type material">
<paragraph pageId="30" pageNumber="81">Type material.</paragraph>
<paragraph pageId="30" pageNumber="81">
Holotype ♀, Kenya, Nairobi Province, ICIPE Campus, Kasarani,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-1.2232">1.2232°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="36.8965">36.8965°E</geoCoordinate>
, 1600m elevation, Malaise trap, meadow in degraded shrub-/grassland, 24 Feb to 3 Mar 2014, R. Copeland leg. (NMKE)
</paragraph>
<paragraph pageId="30" pageNumber="81">Paratypes 4♀♀, the same as holotype (1♀ NMKE; 1♀ RSC; 1♀ ARC; 1♀ ICIPE).</paragraph>
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<paragraph pageId="30" pageNumber="81">Additional material.</paragraph>
<paragraph pageId="30" pageNumber="81">3♂♂, collected together with the four females (2♂♂ RSC; 1♂ ARC).</paragraph>
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<subSubSection lastPageId="33" lastPageNumber="84" pageId="30" pageNumber="81" type="description">
<paragraph pageId="30" pageNumber="81">Description.</paragraph>
<paragraph lastPageId="31" lastPageNumber="82" pageId="30" pageNumber="81">
Holotype ♀. Very small species with body length (without ovipositor) 1.2 mm; length of ovipositor sheaths 0.9mm. Body colour nearly entirely orange-
<pageBreakToken pageId="31" pageNumber="82" start="start">yellow</pageBreakToken>
(Figures 62-64). Head colour orange, except mouthparts brown (Figure 65). Pilosity on lower face pale; dorsum of head with conspicuous black hairs: a row along the internal side of each eye from parascrobal area to temple, a row along the occiput, and three isolated hairs around each of the lateral ocelli; two interocellar bristles. Antenna dark- brown except scape and pedicel yellow underneath (Figure 66). Thorax entirely orange except a small black spot on axilla at wing insertions and on lateral panel of metanotum (Figure 64). Pilosity on thorax mostly pale; sparse long hairs on pronotum, especially along the suture with mid-lobe of mesoscutum; 3 pairs of pale hairs on the lateral side of mid-lobe of mesoscutum, very close to suture with the lateral lobe; 2 pairs of pale hairs on scutellum. Legs entirely pale yellow except claws brown; Forewing stigma brown without infuscation; basal cell without setae, closed by a basal setal line with 5 setae and a costal setal line with 3 setae (Figure 67). Propodeum orange with a transverse black stripe along the suture with metanotum (Figure 64). Propodeum hairs pale; Gaster orange, with four large transverse orange-brown bands along the anterior sutures of the tergites 4-8; pilosity on gaster black. Ovipositor sheaths black, 1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than gaster, 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as body (Figure 63).
</paragraph>
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Head about 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long in front view (Figure 65). POL large, POL: OOL: 2.7. Scape 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as combined length of pedicel, anellus, and F1 (Figure 66); scape 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as combined length of pedicel, anellus, F1 and F2; pedicel elongate, 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than F1; F2 subquadrate, small, 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as F1; following funicular segments subquadrate from F2. Pronotum, mid-lobe of mesoscutum, axilla and scutellum with fine transverse striae. Mid-lobe of mesoscutum conspicuously short, only 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum length; scutellum 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; frenum roughly 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum length, with the same sculpture as the remainder of scutellum. Propodeum with a zig-zag median carina. Forewing stigma oval, without infuscation, about 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (Figure 67); upper part of stigmal vein short, 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as stigma length; uncus as long as upper part of stigmal vein; marginal vein 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as postmarginal vein.
</paragraph>
<paragraph pageId="31" pageNumber="82">
Variation. Body ranges 1.2-1.3 mm with ovipositor up to 1.2 mm. Ovipositor length varies from 0.7 to 0.9
<normalizedToken originalValue="×">x</normalizedToken>
the body length, and from 1.6 to 2.0
<normalizedToken originalValue="×">x</normalizedToken>
the gaster length. No variation in colour and pilosity.
</paragraph>
<paragraph pageId="31" pageNumber="82">
Male. In the absence of molecular analyses, the males caught in Malaise traps along with the four female
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus icipeensis" order="Hymenoptera" pageId="31" pageNumber="82" phylum="Arthropoda" rank="species" species="icipeensis">Megastigmus icipeensis</taxonomicName>
could not be attributed with certainty to this species but noticeable convergences in morphology make it probable. Body length 1mm. Same body colour as in female of
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus icipeensis" order="Hymenoptera" pageId="31" pageNumber="82" phylum="Arthropoda" rank="species" species="icipeensis">Megastigmus icipeensis</taxonomicName>
(Figures 68-70) except a paler yellow pronotum and some additional black patterns on head and thorax: an irregular spot surrounding the three ocelli, an annelation at insertion of pronotum, a darkening of the lateral sutures of mid-lobe of mesoscutum, a narrow transverse band on lateral panel of metanotum, and a transverse band covering the base of propodeum and extending in its middle into a large longitudinal spot in shape of inverted Y (Figures 62, 64). Gaster petiolate, mostly brown- black on dorsum: T3 black, T4 brown black with a large transverse yellow band distally, T5 similar but the yellow band narrower, T6 brown-black, T7 yellow.
</paragraph>
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yellow with the two last tarsal segments and claws black. Pilosity pale on head and thorax, showing the same setal pattern as in female
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus icipeensis" order="Hymenoptera" pageId="32" pageNumber="83" phylum="Arthropoda" rank="species" species="icipeensis">Megastigmus icipeensis</taxonomicName>
, including the interocellar brisles and 2 pairs of pale hairs on scutellum (Figure 70). Forewing stigma
<pageBreakToken pageId="33" pageNumber="84" start="start">without</pageBreakToken>
infuscation (Figure 73); basal cell closed, but without setae on disc; basal setal line with 6 setae, costal setal line with 3 long setae; costal cell with only 3 small setae in the apical 1/2 to 2/3.
</paragraph>
<paragraph pageId="33" pageNumber="84">
Head transverse, about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long in front view (Figure 71). OOL short, POL: OOL: 2.8. Scape 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as combined length of pedicel, anellus, and F1 (Figure 72); scape as long as combined length of pedicel, anellus, F1 and F2; pedicel elongate, 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than F1; F2 subquadrate, small, 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as F1; following funicular segments also subquadrate. Pronotum, mid-lobe of mesoscutum, axilla and scutellum finely transversely striated. Mid-lobe of mesoscutum conspicuously short, only as long as scutellum length; scutellum 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; frenum roughly 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum length, with the same sculpture as the rest of scutellum. Propodeum with an irregular median carina. Forewing stigma nearly rounded, only 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (Figure 74); upper part of stigmal vein very short, only 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as stigma length; uncus 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than upper part of stigmal vein; marginal vein 1.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than postmarginal vein. Genital digitus with only 2 teeth (Figure 73).
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="84" type="host plants">
<paragraph pageId="33" pageNumber="84">Host plants.</paragraph>
<paragraph pageId="33" pageNumber="84">
Unknown. In the molecular study,
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus icipeensis" order="Hymenoptera" pageId="33" pageNumber="84" phylum="Arthropoda" rank="species" species="icipeensis">Megastigmus icipeensis</taxonomicName>
did not group with other taxa, but the phylogenetic position (Figure 14) and the genetic distances between this species and the other groups led to suggest that it could be a seed-feeder associated with another plant family than those identified in this paper.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="84" type="distribution">
<paragraph pageId="33" pageNumber="84">Distribution.</paragraph>
<paragraph pageId="33" pageNumber="84">Only known from the collection site in a degraded meadow near a small stream.</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="84" type="etymology">
<paragraph pageId="33" pageNumber="84">Etymology.</paragraph>
<paragraph pageId="33" pageNumber="84">Named after the institution on whose grounds the species was collected, the campus of the International Centre of Insect Physiology and Ecology (ICIPE), Nairobi.</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="84" type="diagnosis">
<paragraph pageId="33" pageNumber="84">Diagnosis.</paragraph>
<paragraph pageId="33" pageNumber="84">
The species can be easily distinguished from other tiny
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus" order="Hymenoptera" pageId="33" pageNumber="84" phylum="Arthropoda" rank="genus">Megastigmus</taxonomicName>
spp. known from the Afrotropical region by the pale bristles on the thoracic dorsum whereas they are mostly black in
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus zebrinus" order="Hymenoptera" pageId="33" pageNumber="84" phylum="Arthropoda" rank="species" species="zebrinus">Megastigmus zebrinus</taxonomicName>
(Grissell, 2006), and by its orange body whereas
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus pretorianensis" order="Hymenoptera" pageId="33" pageNumber="84" phylum="Arthropoda" rank="species" species="pretorianensis">Megastigmus pretorianensis</taxonomicName>
is mostly black with brown areas (
<bibRefCitation author="Doǧanlar, M" journalOrPublisher="Entomofauna- Zeitschrift fuer Entomologie" pageId="58" pageNumber="109" pagination="561 - 580" title="Diagnosis of Megastigmus spp. (Hymenoptera: Torymidae) reared from galls of Leptocybeinvasa Fisher &amp; Lasalle, 2004, (Hymenoptera: Eulophidae) on Eucalyptus spp. (Myrtaceae), with description of a new species from South Africa." volume="36" year="2015">Doǧanlar 2015</bibRefCitation>
). In contrast to most other species, the male genital digitus bears only two teeth such as in
<taxonomicName class="Insecta" family="Torymidae" genus="Megastigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megastigmus helinae" order="Hymenoptera" pageId="33" pageNumber="84" phylum="Arthropoda" rank="species" species="helinae">Megastigmus helinae</taxonomicName>
but the phallobase of the latter species is more elongate (Figure 29).
</paragraph>
</subSubSection>
</treatment>
</document>