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Current name:
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Allen, 1972: 72. Type locality: &quot;Brownsville [Cameron County], Texas&quot; (original citation). Holotype (♂) in MCZ [# 31957].
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)
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<emphasis id="C0B6EADF7827AB785EC3FCCFB378FCBE" bold="true" box="[151,260,785,811]" pageId="8" pageNumber="561">Remarks</emphasis>
. This species was described in the genus
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by
<bibRefCitation id="96534B3C7827AB785DF3FCCFB629FCBE" author="Zompro, O." box="[935,1109,785,811]" pageId="8" pageNumber="561" pagination="26 - 27" refId="ref5854" refString="Zompro, O. (2008) Raptophasma groehni n. sp., a new species of gladiator from Baltic amber (Insecta: Mantophasmatodea: Mantophasmatidae). Arthropoda, 16 (1), 26 - 27." type="journal article" year="2008">Zompro (2008)</bibRefCitation>
but characterized by present of numerous bristles on pro-, meso- and metatorax; therefore it is transferred to the genus
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and a new combination is proposed here.
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<emphasis id="C0B6EADF7827AB785EC3FE0DB381FE7B" bold="true" box="[151,509,467,494]" italics="true" pageId="8" pageNumber="561">Adicophasma grylloblattoides</emphasis>
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, comb. resurr.
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<emphasis id="C0B6EADF7827AB785EC3FDC4B378FDA1" bold="true" box="[151,260,538,564]" pageId="8" pageNumber="561">Remarks</emphasis>
. This species was described in the genus
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<emphasis id="C0B6EADF7827AB785CBCFDC4B1FCFDA1" box="[744,896,538,564]" italics="true" pageId="8" pageNumber="561">Adicophasma</emphasis>
(
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)
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but later transferred to
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<emphasis id="C0B6EADF7827AB785EC3FDE3B34DFDC2" box="[151,305,573,599]" italics="true" pageId="8" pageNumber="561">Raptophasma</emphasis>
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by
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. Here it returns to
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<emphasis id="C0B6EADF7827AB785C8AFDE3B10AFDC2" box="[734,886,573,599]" italics="true" pageId="8" pageNumber="561">Adicophasma</emphasis>
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based on present of distinct rows on ventral side of profemora and chaetotaxy of thoracic tergites.
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<emphasis id="C0B6EADF7827AB785EC3FF49B3CBFF24" bold="true" box="[151,439,151,178]" italics="true" pageId="8" pageNumber="561">Adicophasma spinosum</emphasis>
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, nom. resurr.
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<emphasis id="C0B6EADF7827AB785EC3FF00B378FF6D" bold="true" box="[151,260,222,248]" pageId="8" pageNumber="561">Remarks</emphasis>
. This species was described as
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<emphasis id="C0B6EADF7827AB785C3BFF03B11AFF62" box="[623,870,221,247]" italics="true" pageId="8" pageNumber="561">Adicophasma spinosa</emphasis>
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from Baltic amber (
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) but specific name was corrected as
<taxonomicName id="35C24D4E7827AB785C54FEDFB00BFE8F" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[512,631,257,283]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="8" pageNumber="561" phylum="Arthropoda" rank="species" species="spinosum">
<emphasis id="C0B6EADF7827AB785C59FEDFB00BFE8F" box="[525,631,257,282]" italics="true" pageId="8" pageNumber="561">spinosum</emphasis>
</taxonomicName>
” because ending of genus “
<emphasis id="C0B6EADF7827AB785DE3FEDFB672FE8E" box="[951,1038,257,283]" italics="true" pageId="8" pageNumber="561">phasma</emphasis>
” is gender neuter (Arillo &amp;
<bibRefCitation id="96534B3C7827AB785B00FEDFB2A8FEAB" author="Engel, M. S. &amp; Grimaldi, D. A." pageId="8" pageNumber="561" pagination="1 - 11" refId="ref4475" refString="Engel, M. S. &amp; Grimaldi, D. A. (2004) A new rock-crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates, 3431, 1 - 11. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 431 % 3 C 0001: ANRCIB % 3 E 2.0. CO; 2" type="journal article" year="2004">Engel, 2004</bibRefCitation>
). It was synonymized with
<taxonomicName id="35C24D4E7827AB785C57FEFAB161FEAB" authorityName="Zompro" authorityYear="2001" box="[515,797,292,318]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="8" pageNumber="561" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="C0B6EADF7827AB785C57FEFAB161FEAB" box="[515,797,292,318]" italics="true" pageId="8" pageNumber="561">Raptophasma kerneggeri</emphasis>
</taxonomicName>
by
<bibRefCitation id="96534B3C7827AB785D12FEFAB18DFEAB" author="Zompro, O." box="[838,1009,292,318]" pageId="8" pageNumber="561" pagination="26 - 27" refId="ref5854" refString="Zompro, O. (2008) Raptophasma groehni n. sp., a new species of gladiator from Baltic amber (Insecta: Mantophasmatodea: Mantophasmatidae). Arthropoda, 16 (1), 26 - 27." type="journal article" year="2008">Zompro (2008)</bibRefCitation>
but differs from latter, as well as from
<taxonomicName id="35C24D4E7827AB785EC3FE96B29FFEF4" authority="Perkovsky &amp; Vasilenko, 2024" authorityName="Perkovsky &amp; Vasilenko" authorityYear="2024" box="[151,227,327,353]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="8" pageNumber="561" phylum="Arthropoda" rank="species" species="neli" status="sp. nov.">
<emphasis id="C0B6EADF7827AB785EC3FE96B29FFEF4" box="[151,227,327,353]" italics="true" pageId="8" pageNumber="561">R. neli</emphasis>
</taxonomicName>
<emphasis id="C0B6EADF7827AB785EB8FE99B335FEF4" bold="true" box="[236,329,327,353]" pageId="8" pageNumber="561">
<taxonomicNameLabel id="DB8557A47827AB785EB8FE99B335FEF4" box="[236,329,327,353]" pageId="8" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present of numerous spine-like bristles on pro-, meso- and metatorax and two rows of such bristles on lower side of profemora.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:affiliation id="0D93DC3078512F83E9EBDAB2A8B1F15D">Cherepovets State University, Lunacharsky Prospect 5, Cherepovets, Vologda Region, 162600, Russia. &amp; Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia. &amp; Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 690022 Vladivostok, Russia.</mods:affiliation>
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<paragraph id="F27D36CD782BAB745EC3FDFCB113FDA8" blockId="4.[151,879,546,573]" box="[151,879,546,573]" pageId="4" pageNumber="557">
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Genus
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<emphasis id="C0B6EADF782BAB745EBAFDFCB3EFFDA8" bold="true" box="[238,403,546,573]" italics="true" pageId="4" pageNumber="557">Adicophasma</emphasis>
<bibRefCitation id="96534B3C782BAB745FCEFDFDB0B4FDA9" author="Engel, M. S. &amp; Grimaldi, D. A." box="[410,712,546,573]" pageId="4" pageNumber="557" pagination="1 - 11" refId="ref4475" refString="Engel, M. S. &amp; Grimaldi, D. A. (2004) A new rock-crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates, 3431, 1 - 11. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 431 % 3 C 0001: ANRCIB % 3 E 2.0. CO; 2" type="journal article" year="2004">Engel &amp; Grimaldi, 2004</bibRefCitation>
</taxonomicName>
, nom. resurr.
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</paragraph>
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<typeStatus id="2D79886F782BAB745EC3FDB6B2B5FD15" box="[151,201,616,640]" pageId="4" pageNumber="557">Type</typeStatus>
species:
<taxonomicName id="35C24D4E782BAB745F72FDB9B171FDEA" authority="Engel &amp; Grimaldi, 2004" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[294,781,615,639]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="C0B6EADF782BAB745F72FDB9B072FDEA" box="[294,526,615,639]" italics="true" pageId="4" pageNumber="557">Adicophasma spinosus</emphasis>
<bibRefCitation id="96534B3C782BAB745C40FDB9B171FDEA" author="Engel, M. S. &amp; Grimaldi, D. A." box="[532,781,615,639]" pageId="4" pageNumber="557" pagination="1 - 11" refId="ref4475" refString="Engel, M. S. &amp; Grimaldi, D. A. (2004) A new rock-crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates, 3431, 1 - 11. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 431 % 3 C 0001: ANRCIB % 3 E 2.0. CO; 2" type="journal article" year="2004">Engel &amp; Grimaldi, 2004</bibRefCitation>
</taxonomicName>
, by original designation.
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<emphasis id="C0B6EADF782BAB745EC3FD73B353FD52" bold="true" box="[151,303,685,711]" pageId="4" pageNumber="557">Composition</emphasis>
. Nowadays this genus consists of three species from Baltic amber known by nymphs only. A new species from Danish amber is described below.
</paragraph>
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<emphasis id="C0B6EADF782BAB745E93FD2BB346FC9A" bold="true" box="[199,314,757,783]" pageId="4" pageNumber="557">Diagnosis</emphasis>
. This genus characterized by profemora with strong spine-like bristles ventrally and by abdominal and especially thoracic tergites with numerous spine-like bristles. By these features
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<emphasis id="C0B6EADF782BAB745A4DFCC7B6CDFCA6" box="[1049,1201,793,819]" italics="true" pageId="4" pageNumber="557">Adicophasma</emphasis>
</taxonomicName>
is easy recognizable from both
<taxonomicName id="35C24D4E782BAB745F59FCE3B3DBFCC2" authorityName="Zompro" authorityYear="2001" box="[269,423,829,855]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745F59FCE3B3DBFCC2" box="[269,423,829,855]" italics="true" pageId="4" pageNumber="557">Raptophasma</emphasis>
</taxonomicName>
and
<taxonomicName id="35C24D4E782BAB745F8AFCE3B0D2FCC2" authorityName="Huang, Nel, Zompro &amp; Waller" authorityYear="2008" box="[478,686,829,855]" class="Insecta" family="Mantophasmatidae" genus="Juramantophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745F8AFCE3B0D2FCC2" box="[478,686,829,855]" italics="true" pageId="4" pageNumber="557">Juramantophasma</emphasis>
</taxonomicName>
.
</paragraph>
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<subSubSection id="BAD86546782BAB745E93FCBFB29CFC51" pageId="4" pageNumber="557" type="discussion">
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<emphasis id="C0B6EADF782BAB745E93FCBFB348FCEE" bold="true" box="[199,308,865,891]" pageId="4" pageNumber="557">Remarks</emphasis>
.
<bibRefCitation id="96534B3C782BAB745F12FCBCB38AFCEE" author="Zompro, O." box="[326,502,865,892]" pageId="4" pageNumber="557" pagination="26 - 27" refId="ref5854" refString="Zompro, O. (2008) Raptophasma groehni n. sp., a new species of gladiator from Baltic amber (Insecta: Mantophasmatodea: Mantophasmatidae). Arthropoda, 16 (1), 26 - 27." type="journal article" year="2008">Zompro (2008)</bibRefCitation>
synonymized genus
<taxonomicName id="35C24D4E782BAB745CBAFCBFB1FAFCEE" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[750,902,865,891]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745CBAFCBFB1FAFCEE" box="[750,902,865,891]" italics="true" pageId="4" pageNumber="557">Adicophasma</emphasis>
</taxonomicName>
under
<taxonomicName id="35C24D4E782BAB745D89FCBFB60BFCEE" authorityName="Zompro" authorityYear="2001" box="[989,1143,865,891]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745D89FCBFB60BFCEE" box="[989,1143,865,891]" italics="true" pageId="4" pageNumber="557">Raptophasma</emphasis>
</taxonomicName>
and
<taxonomicName id="35C24D4E782BAB745AE1FCBCB753FCEE" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[1205,1327,866,892]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="C0B6EADF782BAB745AE1FCBCB753FCEE" box="[1205,1327,866,892]" italics="true" pageId="4" pageNumber="557">A. spinosa</emphasis>
</taxonomicName>
under
<taxonomicName id="35C24D4E782BAB745BD0FCBCB36CFC0A" authorityName="Zompro" authorityYear="2001" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="C0B6EADF782BAB745BD0FCBCB36CFC0A" italics="true" pageId="4" pageNumber="557">R. kerneggeri</emphasis>
</taxonomicName>
. Here, the name
<taxonomicName id="35C24D4E782BAB745F8EFC5BB00EFC0A" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[474,626,901,927]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745F8EFC5BB00EFC0A" box="[474,626,901,927]" italics="true" pageId="4" pageNumber="557">Adicophasma</emphasis>
</taxonomicName>
is resurrected from synonymy and this taxon is considered as distinct genus.
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,159 @@
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</mods:role>
<mods:namePart id="8BBA0A8B4BB4C71D28DF502D2565EB38">Vasilenko, Dmitry V.</mods:namePart>
<mods:affiliation id="CEAF24BA3AA55ED7DF3A4BB86F68D343">Cherepovets State University, Lunacharsky Prospect 5, Cherepovets, Vologda Region, 162600, Russia. &amp; Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia. &amp; Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 690022 Vladivostok, Russia.</mods:affiliation>
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<mods:title id="9A77A912FD589C27720575AE781EEA7F">Zootaxa</mods:title>
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<mods:date id="6181FD4E96995B97DC0ED2013F4DA673">2024</mods:date>
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<subSubSection id="BAD86546782BAB745EC3FBCBB101FBBA" box="[151,893,1045,1072]" pageId="4" pageNumber="557" type="nomenclature">
<paragraph id="F27D36CD782BAB745EC3FBCBB101FBBA" blockId="4.[151,893,1045,1107]" box="[151,893,1045,1072]" pageId="4" pageNumber="557">
<heading id="A93581A1782BAB745EC3FBCBB101FBBA" bold="true" box="[151,893,1045,1072]" fontSize="11" level="1" pageId="4" pageNumber="557" reason="1">
<emphasis id="C0B6EADF782BAB745EC3FBCBB101FBBA" bold="true" box="[151,893,1045,1072]" pageId="4" pageNumber="557">
<taxonomicName id="35C24D4E782BAB745EC3FBCBB16EFBBA" authority="Perkovsky &amp; Storozhenko" authorityName="Perkovsky &amp; Storozhenko" authorityYear="2024" box="[151,786,1045,1072]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hafniensis" status="sp. nov.">
<emphasis id="C0B6EADF782BAB745EC3FBCBB3BDFBA5" bold="true" box="[151,449,1045,1072]" italics="true" pageId="4" pageNumber="557">Adicophasma hafniensis</emphasis>
Perkovsky &amp; Storozhenko
</taxonomicName>
,
<taxonomicNameLabel id="DB8557A4782BAB745D4BFBCBB101FBBA" box="[799,893,1045,1071]" pageId="4" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="BAD86546782BAB745EC3FBE7B37BFBC6" box="[151,263,1081,1107]" pageId="4" pageNumber="557" type="description">
<paragraph id="F27D36CD782BAB745EC3FBE7B37BFBC6" blockId="4.[151,893,1045,1107]" box="[151,263,1081,1107]" pageId="4" pageNumber="557">
<figureCitation id="6AF92A48782BAB745EC3FBE7B37BFBC6" box="[151,263,1081,1107]" captionStart-0="FIGURES 912" captionStart-1="FIGURES 1316" captionStart-2="FIGURES 1723" captionStartId-0="5.[152,270,1220,1244]" captionStartId-1="6.[152,270,1706,1730]" captionStartId-2="7.[152,270,1831,1855]" captionTargetBox-0="[170,1409,184,1190]" captionTargetBox-1="[247,1338,187,1676]" captionTargetBox-2="[158,1429,188,1802]" captionTargetId-0="figure-19@5.[170,1418,179,1194]" captionTargetId-1="figure-26@6.[177,1410,179,1680]" captionTargetId-2="figure-19@7.[151,1436,181,1807]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 912. Adicophasma hafniensis sp. nov. 9. Thoracic and abdominal tergites, dorsal view; 10. Fifth tarsal segment and arolium, dorsal view; 11. Anterior half of body, lateral view; 12. Head and profemur, frontal view." captionText-1="FIGURES 1316. Adicophasma hafniensis sp. nov. 13. Body, fronto-ventral view; 14. Apex of abdomen, ventral view; 15. Mesofemur from inner side, lateral view; 16. Fore leg from inner side, lateral view." captionText-2="FIGURES 1723. Adicophasma hafniensis sp. nov. 17. Head, basal segments of antennae, thoracic segments and coxae, lateral view; 18. Head, frontal view; 19. Fore leg, lateral view; 20. Metafemur, lateral view; 21. Thoracic and abdominal tergites; 22. Apex of abdomen, ventral view; 23. Fifth tarsal segment and arolium, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.11102266" figureDoi-1="http://doi.org/10.5281/zenodo.11102268" figureDoi-2="http://doi.org/10.5281/zenodo.11102270" httpUri-0="https://zenodo.org/record/11102266/files/figure.png" httpUri-1="https://zenodo.org/record/11102268/files/figure.png" httpUri-2="https://zenodo.org/record/11102270/files/figure.png" pageId="4" pageNumber="557">Figs 923</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745EC3FB5FB285FB76" pageId="4" pageNumber="557" type="etymology">
<paragraph id="F27D36CD782BAB745EC3FB5FB285FB76" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745EC3FB5FB362FB0E" bold="true" box="[151,286,1153,1179]" pageId="4" pageNumber="557">Etymology.</emphasis>
The species name is after Universitas Hafniensis (Københavns Universitet), the oldest university in
<collectingCountry id="8AD5765D782BAB745EECFB7BB35FFB2A" box="[184,291,1189,1215]" name="Denmark" pageId="4" pageNumber="557">Denmark</collectingCountry>
, where the
<typeStatus id="2D79886F782BAB745FFAFB78B3A2FB55" box="[430,478,1190,1216]" pageId="4" pageNumber="557">type</typeStatus>
was found at NHMD (the former Zoological Museum of Copenhagen University, ZMUC).
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745E93FB33B3ABFAE6" pageId="4" pageNumber="557" type="materials_examined">
<paragraph id="F27D36CD782BAB745E93FB33B3ABFAE6" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93FB33B308FA92" bold="true" box="[199,372,1261,1287]" pageId="4" pageNumber="557">Type material</emphasis>
.
<materialsCitation id="42AA3C90782BAB745FDCFB33B3ABFAE6" collectingDate="1969-02-01" collectionCode="V" pageId="4" pageNumber="557" specimenCount="1" typeStatus="holotype">
<typeStatus id="2D79886F782BAB745FDCFB33B38CFA92" box="[392,496,1261,1287]" pageId="4" pageNumber="557" type="holotype">Holotype</typeStatus>
ZMUC-901825, with old label Phasmoidea 1825 C.
<collectionCode id="94D3AE08782BAB745A38FB33B6FEFA92" box="[1132,1154,1261,1287]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="557" type="Museum">V</collectionCode>
. Henningsen
<date id="867C100D782BAB745B71FB33B7EFFA92" box="[1317,1427,1261,1287]" pageId="4" pageNumber="557" value="1969-02-01">
<collectingDate id="9638E9E5782BAB745B71FB33B7EFFA92" box="[1317,1427,1261,1287]" pageId="4" pageNumber="557" value="1969-02-01">1-2-1969</collectingDate>
</date>
, Danish amber, late Eocene. Nymph, probably mid instar. The sex unknown, because of in the nymphs of extant
<taxonomicName id="35C24D4E782BAB745EC3FAEBB30EFADA" authorityName="Klass, Zompro, Kristensen &amp; Adis" authorityYear="2002" box="[151,370,1333,1359]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
the apical abdominal sternites and cerci are not strongly modified such as in adult specimens. Syninclusions: stellate hairs.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745E93FAA3B620F9DE" pageId="4" pageNumber="557" type="diagnosis">
<paragraph id="F27D36CD782BAB745E93FAA3B620F9DE" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93FAA3B346FA02" bold="true" box="[199,314,1405,1431]" pageId="4" pageNumber="557">Diagnosis</emphasis>
. New species is most similar to
<taxonomicName id="35C24D4E782BAB745CF7FAA3B18FFA02" authorityName="Arillo &amp; Engel" authorityYear="2006" box="[675,1011,1405,1431]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="grylloblattoides">
<emphasis id="C0B6EADF782BAB745CF7FAA3B18FFA02" box="[675,1011,1405,1431]" italics="true" pageId="4" pageNumber="557">Adicophasma grylloblattoides</emphasis>
</taxonomicName>
but differs from the latter by enlarged pedicel and chaetotaxy of abdomen (in
<taxonomicName id="35C24D4E782BAB745C35FA7CB148FA2E" authorityName="Arillo &amp; Engel" authorityYear="2006" box="[609,820,1441,1468]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="grylloblattoides">
<emphasis id="C0B6EADF782BAB745C35FA7CB148FA2E" box="[609,820,1441,1468]" italics="true" pageId="4" pageNumber="557">A. grylloblattoides</emphasis>
</taxonomicName>
, pedicel as long as wide and sternites with scattered, simple setae). In
<taxonomicName id="35C24D4E782BAB745F08FA18B39BFA4A" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[348,487,1478,1504]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="spinosum">
<emphasis id="C0B6EADF782BAB745F08FA18B39BFA4A" box="[348,487,1478,1504]" italics="true" pageId="4" pageNumber="557">A. spinosum</emphasis>
</taxonomicName>
pedicel elongated, at least 3 times longer than wide, scapus shorter than elongated pedicel, anterior side of procoxa with 3 long bristles, and sternites with numerous sparse setae of moderated length. The features separated
<taxonomicName id="35C24D4E782BAB745FC8F9D0B06EF9B2" authorityName="Zompro" authorityYear="2008" box="[412,530,1549,1576]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="C0B6EADF782BAB745FC8F9D0B06EF9B2" box="[412,530,1549,1576]" italics="true" pageId="4" pageNumber="557">A. groehni</emphasis>
</taxonomicName>
from new species are follow: the posterior margin of pro-, meso- and metatorax with 6 long bristles and the posterior margin of sternites with a row of up to 8 bristles.
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB755E93F98BB183F81C" lastPageId="5" lastPageNumber="558" pageId="4" pageNumber="557" type="description">
<paragraph id="F27D36CD782BAB745E93F98BB602F8B6" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93F98BB32DF9FA" bold="true" box="[199,337,1621,1647]" pageId="4" pageNumber="557">Description</emphasis>
. Head hypognathous; the compound eyes large and protrude slightly above the head; vertical diameter of the eye significantly exceeds the height of the cheeks. Frontoclipeal suture broadly rounded; clipeus triangular; labrum narrow triangular with acutely rounded apex. Antennae 16-segmented; scapus cylindrical, 1.5 times as long as wide, about equal in size to pedicel; pedicel 1.5 times as long as wide; third antennomere (first flagellomere) elongated, 3.5 times as long as wide and 1.4 times as long as scapus. Maxillary palpus 5-segmented, second and fifth palpomeres longest, other palpomeres short. Labial palpus 3-segmented.
</paragraph>
<caption id="A6BD6645782AAB755ECCFB1AB604FA95" ID-DOI="http://doi.org/10.5281/zenodo.11102266" ID-Zenodo-Dep="11102266" httpUri="https://zenodo.org/record/11102266/files/figure.png" pageId="5" pageNumber="558" startId="5.[152,270,1220,1244]" targetBox="[170,1409,184,1190]" targetPageId="5" targetType="figure">
<paragraph id="F27D36CD782AAB755ECCFB1AB604FA95" blockId="5.[151,1437,1220,1280]" pageId="5" pageNumber="558">
<emphasis id="C0B6EADF782AAB755ECCFB1AB33AFB49" bold="true" box="[152,326,1220,1244]" pageId="5" pageNumber="558">FIGURES 912</emphasis>
.
<taxonomicName id="35C24D4E782AAB755F05FB1AB03BFB49" authority="Perkovsky &amp; Vasilenko, 2024" authorityName="Perkovsky &amp; Vasilenko" authorityYear="2024" box="[337,583,1220,1244]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hafniensis" status="sp. nov.">
<emphasis id="C0B6EADF782AAB755F05FB1AB03BFB49" box="[337,583,1220,1244]" italics="true" pageId="5" pageNumber="558">Adicophasma hafniensis</emphasis>
</taxonomicName>
<emphasis id="C0B6EADF782AAB755C18FB1AB0E7FB49" bold="true" box="[588,667,1220,1244]" pageId="5" pageNumber="558">
<taxonomicNameLabel id="DB8557A4782AAB755C18FB1AB0E7FB49" box="[588,667,1220,1244]" pageId="5" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9. Thoracic and abdominal tergites, dorsal view; 10. Fifth tarsal segment and arolium, dorsal view; 11. Anterior half of body, lateral view; 12. Head and profemur, frontal view.
</paragraph>
</caption>
<paragraph id="F27D36CD782AAB755E93FAF0B332FA4D" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Thorax. Pronotum distinctly shorter than head, almost as long as meso- and metanotum, with almost straight posterior margin armed by four bristles. Dorsal side of pro-, meso- and metatorax with numerous, mostly irregularly situated, spine-like bristles. Thoracic tergites distinctly marginated laterally and posteriorly; posterior margin of mesothorax straight and with a row of six bristles. Metatorax narrower than mesothorax, with a few irregularly situated bristles.</paragraph>
<paragraph id="F27D36CD782AAB755E93FA3DB6B6F924" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">
Legs. Pro- and metafemora slightly broadened and longer than more slender mesofemora. Ventral side of pro- and mesofemora with two rows of spine-like bristles. Protibia with numerous hair-like setae, inner side apically bearing 3-4 setae with cuticular thickenings at base. Mesotibia not visible. Coxae elongate, anterior side of procoxa with a long bristle. Tarsi 5-segmented, basitarsus as long as second tarsomere; third segments slightly longer than second; fourth tarsomere as long as third; third and fourth tarsomere with wide distal process in the middle; fifth segment elongate; arolium large, with typical for the
<taxonomicName id="35C24D4E782AAB755CBEF948B1B9F925" authorityName="Klass, Zompro, Kristensen &amp; Adis" authorityYear="2002" box="[746,965,1686,1712]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
preapical row of setae.
</paragraph>
<paragraph id="F27D36CD782AAB755E93F964B7F6F8D4" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Abdomen. Abdominal tergites transverse, with a few irregularly situated very long setae; anal plate trangular. Abdomem below with visible four apical sternites and subgenital plate. Hind margin of 8-th and 9-th sternites with two strong bristles; apex of subgenital plate with deep excision; paraprocts considerably extending beyond apex of subgenital plate; cerci simple. Lateral sides of sternites and cerci with very long, thin and densely arranged setae.</paragraph>
<paragraph id="F27D36CD782AAB755E93F894B183F81C" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Lengths (mm): Total: 3.2; head: 0,56; antennae: 3.35; pronotum: 0,43; mesonotum: 0,41; metanotum: 0,4; profemora: 0,75; protibiae: 0.82; protarsi: 0,65; mesofemora: 0,76; cerci 0,15.</paragraph>
</subSubSection>
<subSubSection id="BAD86546782AAB755E93F84CB6D3F839" box="[199,1199,1938,1965]" pageId="5" pageNumber="558" type="discussion">
<paragraph id="F27D36CD782AAB755E93F84CB6D3F839" blockId="5.[151,1437,1326,1965]" box="[199,1199,1938,1965]" pageId="5" pageNumber="558">
<emphasis id="C0B6EADF782AAB755E93F84CB348F839" bold="true" box="[199,308,1938,1964]" pageId="5" pageNumber="558">Remarks</emphasis>
. This is first record of the genus
<taxonomicName id="35C24D4E782AAB755CF2F84CB142F839" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[678,830,1938,1964]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782AAB755CF2F84CB142F839" box="[678,830,1938,1964]" italics="true" pageId="5" pageNumber="558">Adicophasma</emphasis>
</taxonomicName>
from late Eocene Danish amber.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0F49ECB8B68453E486963BDAAC85E5B4">New and little-known Mantophasmatidae (Insecta: Mantophasmatodea) from European amber</mods:title>
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<mods:namePart id="6DE99F281DE78DFD83A397CCADC72D66">Perkovsky, Evgeny E.</mods:namePart>
<mods:affiliation id="879B04E8A124EAA1EBE27ECF313A0986">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen 2100, Denmark.</mods:affiliation>
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<heading id="A93581A1782EAB715EC3FB10B030FB7D" bold="true" box="[151,588,1230,1257]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
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Genus
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<emphasis id="C0B6EADF782EAB715EBAFB10B3E8FB7C" bold="true" box="[238,404,1230,1257]" italics="true" pageId="1" pageNumber="554">Raptophasma</emphasis>
<bibRefCitation id="96534B3C782EAB715FC8FB11B030FB7D" author="Zompro, O." box="[412,588,1230,1257]" pageId="1" pageNumber="554" pagination="229 - 261" refId="ref5767" refString="Zompro, O. (2001) The Phasmatodea and Raptophasma n. gen., Orthoptera incertae sedis, in Baltic amber (Insecta: Orthoptera). Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitat Hamburg, 85, 229 - 261." type="journal article" year="2001">Zompro, 2001</bibRefCitation>
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<typeStatus id="2D79886F782EAB715EC3FACAB2B5FAB9" box="[151,201,1300,1324]" pageId="1" pageNumber="554">Type</typeStatus>
species:
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<emphasis id="C0B6EADF782EAB715F72FACDB05AFABE" box="[294,550,1299,1323]" italics="true" pageId="1" pageNumber="554">Raptophasma kerneggeri</emphasis>
<bibRefCitation id="96534B3C782EAB715C78FACAB0C3FABE" author="Zompro, O." box="[556,703,1299,1323]" pageId="1" pageNumber="554" pagination="229 - 261" refId="ref5767" refString="Zompro, O. (2001) The Phasmatodea and Raptophasma n. gen., Orthoptera incertae sedis, in Baltic amber (Insecta: Orthoptera). Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitat Hamburg, 85, 229 - 261." type="journal article" year="2001">Zompro, 2001</bibRefCitation>
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, by original designation.
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<emphasis id="C0B6EADF782EAB715EC3FA87B376FAE6" bold="true" box="[151,266,1369,1395]" pageId="1" pageNumber="554">Diagnosis</emphasis>
. This genus characterized by profemora without strong spine-like bristles ventrally and by thoracic and abdominal tergites with numerous short hair-like setae, while the spine-like bristles absent. By these features
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is easy recognizable from
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<emphasis id="C0B6EADF782EAB715C22FA7FB172FA2E" box="[630,782,1441,1467]" italics="true" pageId="1" pageNumber="554">Adicophasma</emphasis>
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. From Jurassic
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<emphasis id="C0B6EADF782EAB715D9AFA7FB6E2FA2E" box="[974,1182,1441,1467]" italics="true" pageId="1" pageNumber="554">Juramantophasma</emphasis>
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this genus differs by smaller size, proportions of femora, anterior margin of procoxae with one strong spine, and triangular female cerci (in
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<emphasis id="C0B6EADF782EAB715EEAFA34B34DF996" box="[190,305,1514,1539]" italics="true" pageId="1" pageNumber="554">J. sinicum</emphasis>
<bibRefCitation id="96534B3C782EAB715F6DFA34B0A0F996" author="Huang, D. Y. &amp; Nel, A. &amp; Zompro, O. &amp; Waller, A." box="[313,732,1513,1540]" pageId="1" pageNumber="554" pagination="947 - 952" refId="ref4676" refString="Huang, D. Y., Nel, A., Zompro, O. &amp; Waller, A. (2008) Mantophasmatodea now in the Jurassic. Naturwissenschaften, 95, 947 - 952. https: // doi. org / 10.1007 / s 00114 - 008 - 0412 - x" type="journal article" year="2008">Huang, Nel, Zompro &amp; Waller, 2008</bibRefCitation>
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,
<emphasis id="C0B6EADF782EAB715CBCFA37B1E1F996" bold="true" box="[744,925,1513,1539]" pageId="1" pageNumber="554">nom. corrected</emphasis>
, body length
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, profemora 2.5 times as long as wide, anterior margin of coxae with a dense row of setae, and female cerci hooked).
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<emphasis id="C0B6EADF782EAB715E93F9EFB323F9DE" bold="true" box="[199,351,1585,1611]" pageId="1" pageNumber="554">Composition</emphasis>
. There are two species from Baltic amber one of which is described below.
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<emphasis id="C0B6EADF782EAB715E93F98BB348F9FA" bold="true" box="[199,308,1621,1647]" pageId="1" pageNumber="554">Remarks</emphasis>
. Recent genera of
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differ by armature of femur, by chaetotaxy of thorax and abdomen, by shape of male and female terminalia, as well as by size and coloration of body and appendages (
<bibRefCitation id="96534B3C782EAB715B09F9A7B364F922" author="Klass, K. D. &amp; Picker, M. D. &amp; Damgaard, J. &amp; Noort, Sv. &amp; Tojo, K." pageId="1" pageNumber="554" pagination="3 - 67" refId="ref4790" refString="Klass, K. D., Picker, M. D., Damgaard, J., Noort, Sv. &amp; Tojo, K. (2003) The taxonomy, genitalic morphology, and phylogenetic relationships of Southern African Mantophasmatodea (Insecta). Entomologische Abhandlungen, 61, 3 - 67." type="journal article" year="2003">
Klass
<emphasis id="C0B6EADF782EAB715EC3F940B2B4F922" box="[151,200,1693,1719]" italics="true" pageId="1" pageNumber="554">et al</emphasis>
., 2003
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;
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Wipfler
<emphasis id="C0B6EADF782EAB715FD4F940B3CEF922" box="[384,434,1693,1719]" italics="true" pageId="1" pageNumber="554">et al</emphasis>
., 2018
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). These features seem to be important for separating extinct genera also.
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Perkovsky &amp; Storozhenko
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,
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<figureCitation id="6AF92A48782EAB715D62F8D7B1E2F8B6" box="[822,926,1801,1827]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 48" captionStartId-0="2.[152,270,1318,1342]" captionStartId-1="3.[152,270,1356,1380]" captionTargetBox-0="[195,1393,179,1291]" captionTargetBox-1="[158,1429,186,1323]" captionTargetId-0="figure-26@2.[195,1393,179,1291]" captionTargetId-1="figure-19@3.[151,1436,179,1330]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 13. Raptophasma neli sp. nov. 1. Body; 2. Head, thorax, coxa, profemur and protibia; 3. Fifth tarsal segment and arolium of hind legs." captionText-1="FIGURES 48. Raptophasma neli sp. nov. 4. Body, lateral view; 5. Coxae and profemur, lateral view; 6. Head, pro- and mesonotum, lateral view; 7. Apical abdominal tergites and cerci, lateral view; 7. Fifth tarsal segment and arolium of hind legs, dorsal view; 8. Fifth tarsal segment and arolium, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.11102262" figureDoi-1="http://doi.org/10.5281/zenodo.11102264" httpUri-0="https://zenodo.org/record/11102262/files/figure.png" httpUri-1="https://zenodo.org/record/11102264/files/figure.png" pageId="1" pageNumber="554">Figs 18</figureCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782EAB715EC3F88FB32AF81A" pageId="1" pageNumber="554" type="etymology">
<paragraph id="F27D36CD782EAB715EC3F88FB32AF81A" blockId="1.[151,1437,1873,1935]" pageId="1" pageNumber="554">
<emphasis id="C0B6EADF782EAB715EC3F88FB364F8FE" bold="true" box="[151,280,1873,1899]" pageId="1" pageNumber="554">Etymology</emphasis>
. New species is named in honor of
<collectingCountry id="8AD5765D782EAB715CFFF88CB08BF8F9" box="[683,759,1874,1900]" name="France" pageId="1" pageNumber="554">France</collectingCountry>
paleoentomologist André Nel (Muséum National dHistoire Naturelle, Paris).
</paragraph>
</subSubSection>
<caption id="A6BD6645782DAB725ECCFAF8B317FAF7" ID-DOI="http://doi.org/10.5281/zenodo.11102262" ID-Zenodo-Dep="11102262" httpUri="https://zenodo.org/record/11102262/files/figure.png" pageId="2" pageNumber="555" startId="2.[152,270,1318,1342]" targetBox="[195,1393,179,1291]" targetPageId="2" targetType="figure">
<paragraph id="F27D36CD782DAB725ECCFAF8B317FAF7" blockId="2.[151,1437,1318,1378]" pageId="2" pageNumber="555">
<emphasis id="C0B6EADF782DAB725ECCFAF8B347FAAB" bold="true" box="[152,315,1318,1342]" pageId="2" pageNumber="555">FIGURES 13</emphasis>
.
<taxonomicName id="35C24D4E782DAB725F13FAF8B383FAAB" authority="Perkovsky &amp; Vasilenko, 2024" authorityName="Perkovsky &amp; Vasilenko" authorityYear="2024" box="[327,511,1318,1342]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="neli" status="sp. nov.">
<emphasis id="C0B6EADF782DAB725F13FAF8B383FAAB" box="[327,511,1318,1342]" italics="true" pageId="2" pageNumber="555">Raptophasma neli</emphasis>
</taxonomicName>
<emphasis id="C0B6EADF782DAB725C52FAF8B02BFAAB" bold="true" box="[518,599,1318,1342]" pageId="2" pageNumber="555">
<taxonomicNameLabel id="DB8557A4782DAB725C52FAF8B02BFAAB" box="[518,599,1318,1342]" pageId="2" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1. Body; 2. Head, thorax, coxa, profemur and protibia; 3. Fifth tarsal segment and arolium of hind legs.
</paragraph>
</caption>
<subSubSection id="BAD86546782DAB725E93FA6AB727F983" pageId="2" pageNumber="555" type="materials_examined">
<paragraph id="F27D36CD782DAB725E93FA6AB727F983" blockId="2.[151,1437,1460,1846]" pageId="2" pageNumber="555">
<materialsCitation id="42AA3C90782DAB725E93FA6AB72BF983" county="Yantarnyi" location="The" municipality="Baltic" pageId="2" pageNumber="555" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="C0B6EADF782DAB725E93FA6AB310FA5B" bold="true" box="[199,364,1460,1486]" pageId="2" pageNumber="555">
<typeStatus id="2D79886F782DAB725E93FA6AB37EFA5B" box="[199,258,1460,1486]" pageId="2" pageNumber="555">Type</typeStatus>
material
</emphasis>
. NHMD-614830,
<collectingCounty id="1B1C4E41782DAB725C62FA6AB0DAFA5B" box="[566,678,1460,1486]" pageId="2" pageNumber="555">Yantarnyi</collectingCounty>
,
<collectingRegion id="3006F82F782DAB725CE4FA6AB1F3FA5B" box="[688,911,1460,1486]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="555">Kaliningrad Region</collectingRegion>
,
<collectingMunicipality id="1219ACB7782DAB725DCCFA6AB1A7FA5B" box="[920,987,1460,1486]" pageId="2" pageNumber="555">Baltic</collectingMunicipality>
amber, late Eocene. Right lateral surface partly cutted, ventral and dorsal side through the thickness of amber not visible clearly.
<location id="F71D6016782DAB725A21FA06B6DDFA67" LSID="urn:lsid:plazi:treatment:7A6B87DB782EAB745EC3F8D7B0A4FE44:F71D6016782DAB725A21FA06B6DDFA67" box="[1141,1185,1496,1522]" county="Yantarnyi" municipality="Baltic" name="The" pageId="2" pageNumber="555" stateProvince="Kaliningrad">The</location>
sex male, because the enlarged cerci.
<location id="F71D6016782DAB725F16FA22B398F983" LSID="urn:lsid:plazi:treatment:7A6B87DB782EAB745EC3F8D7B0A4FE44:F71D6016782DAB725F16FA22B398F983" box="[322,484,1532,1558]" county="Yantarnyi" municipality="Baltic" name="Syninclusions" pageId="2" pageNumber="555" stateProvince="Kaliningrad">Syninclusions</location>
: worker ant
<emphasis id="C0B6EADF782DAB725C3AFA22B10EF983" box="[622,882,1532,1558]" italics="true" pageId="2" pageNumber="555">
<location id="F71D6016782DAB725C3AFA22B17CF983" LSID="urn:lsid:plazi:treatment:7A6B87DB782EAB745EC3F8D7B0A4FE44:F71D6016782DAB725C3AFA22B17CF983" box="[622,768,1532,1558]" county="Yantarnyi" municipality="Baltic" name="Ctenobetylus" pageId="2" pageNumber="555" stateProvince="Kaliningrad">Ctenobetylus</location>
goepperti
</emphasis>
(Mayr),
<taxonomicName id="35C24D4E782DAB725D83FA22B621F983" authorityName="Haliday" authorityYear="1839" box="[983,1117,1532,1558]" class="Insecta" family="Scelionidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="555" phylum="Arthropoda" rank="family">Scelionidae</taxonomicName>
, 6
<location id="F71D6016782DAB725A2FFA22B6C0F983" LSID="urn:lsid:plazi:treatment:7A6B87DB782EAB745EC3F8D7B0A4FE44:F71D6016782DAB725A2FFA22B6C0F983" box="[1147,1212,1532,1558]" county="Yantarnyi" municipality="Baltic" name="Acari" pageId="2" pageNumber="555" stateProvince="Kaliningrad">Acari</location>
, stellate hairs
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782DAB725E93F9FEB722F933" pageId="2" pageNumber="555" type="diagnosis">
<paragraph id="F27D36CD782DAB725E93F9FEB722F933" blockId="2.[151,1437,1460,1846]" pageId="2" pageNumber="555">
<emphasis id="C0B6EADF782DAB725E93F9FEB346F9AF" bold="true" box="[199,314,1568,1594]" pageId="2" pageNumber="555">Diagnosis</emphasis>
. New species similar to
<taxonomicName id="35C24D4E782DAB725C05F9FEB117F9AF" authorityName="Zompro" authorityYear="2001" box="[593,875,1568,1594]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="C0B6EADF782DAB725C05F9FEB117F9AF" box="[593,875,1568,1594]" italics="true" pageId="2" pageNumber="555">Raptophasma kerneggeri</emphasis>
</taxonomicName>
but differs from the latter by narrower profemora without two carinae dorsally and by body and legs coloration (in
<taxonomicName id="35C24D4E782DAB725DC4F99BB656F9CB" authorityName="Zompro" authorityYear="2001" box="[912,1066,1604,1630]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="2" pageNumber="555" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="C0B6EADF782DAB725DC4F99BB656F9CB" box="[912,1066,1604,1630]" italics="true" pageId="2" pageNumber="555">R. kerneggeri</emphasis>
</taxonomicName>
, profemora with two prominent carinae dorsally, 2 times as long as broad, body light brown, occiput blackish, pronotum light brown with two black, longitudinal lateral stripes, mesonotum and metanotum without black mark, and legs blackish without spots).
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782DAB745E93F96EB05EFE88" lastPageId="4" lastPageNumber="557" pageId="2" pageNumber="555" type="description">
<paragraph id="F27D36CD782DAB725E93F96EB306F8A3" blockId="2.[151,1437,1460,1846]" pageId="2" pageNumber="555">
<emphasis id="C0B6EADF782DAB725E93F96EB32DF95F" bold="true" box="[199,337,1712,1738]" pageId="2" pageNumber="555">Description</emphasis>
. Head hypognathous; compound eyes large and protrude slightly above the head; vertical diameter of the eye significantly exceeds the height of the cheeks; ommatidia small and numerous. Basal segments of antennae invisible; mid segments of flagellum elongated, 3.5-4 times as long as wide. Mouthparts typical for the
<taxonomicName id="35C24D4E782DAB725EC3F8C2B30AF8A3" authorityName="Klass, Zompro, Kristensen &amp; Adis" authorityYear="2002" box="[151,374,1820,1846]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="2" pageNumber="555" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
.
</paragraph>
<caption id="A6BD6645782CAB735ECCFA92B17AFA39" ID-DOI="http://doi.org/10.5281/zenodo.11102264" ID-Zenodo-Dep="11102264" httpUri="https://zenodo.org/record/11102264/files/figure.png" pageId="3" pageNumber="556" startId="3.[152,270,1356,1380]" targetBox="[158,1429,186,1323]" targetPageId="3" targetType="figure">
<paragraph id="F27D36CD782CAB735ECCFA92B17AFA39" blockId="3.[151,1437,1356,1453]" pageId="3" pageNumber="556">
<emphasis id="C0B6EADF782CAB735ECCFA92B341FAF1" bold="true" box="[152,317,1356,1380]" pageId="3" pageNumber="556">FIGURES 48</emphasis>
.
<taxonomicName id="35C24D4E782CAB735F19FA92B07BFAF1" authority="Perkovsky &amp; Vasilenko, 2024" authorityName="Perkovsky &amp; Vasilenko" authorityYear="2024" box="[333,519,1356,1380]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="neli" status="sp. nov.">
<emphasis id="C0B6EADF782CAB735F19FA92B07BFAF1" box="[333,519,1356,1380]" italics="true" pageId="3" pageNumber="556">Raptophasma neli</emphasis>
</taxonomicName>
<emphasis id="C0B6EADF782CAB735C44FA93B018FAF0" bold="true" box="[528,612,1357,1381]" pageId="3" pageNumber="556">
<taxonomicNameLabel id="DB8557A4782CAB735C44FA93B018FAF0" box="[528,612,1357,1381]" pageId="3" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
4. Body, lateral view; 5. Coxae and profemur, lateral view; 6. Head, pro- and mesonotum, lateral view; 7. Apical abdominal tergites and cerci, lateral view; 7. Fifth tarsal segment and arolium of hind legs, dorsal view; 8. Fifth tarsal segment and arolium, lateral view.
</paragraph>
</caption>
<paragraph id="F27D36CD782CAB735E93FA05B75AF98C" blockId="3.[151,1437,1499,2029]" pageId="3" pageNumber="556">Thorax. Pronotum distinctly shorter than head, almost as long as mesonotum and 1.5 times longer than metanotum. Thoracic tergites sharply marginated; dorsal side smooth, bristles and hair-like setae absent.</paragraph>
<paragraph id="F27D36CD782CAB735E93F9FDB0C1F964" blockId="3.[151,1437,1499,2029]" pageId="3" pageNumber="556">Legs. Profemora elongated, 2.8 times as long as broad, covered by numerous short setae. Ventral side of profemora with a dense row of short hair-like setae, long bristles absent. Protibia slightly shorter than profemora, with numerous hair-like setae; apex of tibia without thickenings at base of setae. Mesotibia not visible. All coxae elongate; anterior surface of procoxa with short spine. Tarsi 5-segmented, fifth segment elongate and slender; arolium large and broad, surpassing claws, with a row of preapical setae typical for the heelwalkers. Third tarsomere with narrow triangle distal process in the middle.</paragraph>
<paragraph id="F27D36CD782CAB735E93F925B198F8C8" blockId="3.[151,1437,1499,2029]" pageId="3" pageNumber="556">
Abdomen. First three tergites almost equal in length, each about 2 times shorter than pronotum; dorsal side smooth, bristles and hair-like setae absent. Abdominal 9-10th tergites unarmed, smooth. Cerci simple, elongated with pointed apex as in
<taxonomicName id="35C24D4E782CAB735FF6F89AB046F8C8" authorityName="Zompro" authorityYear="2001" box="[418,570,1859,1885]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="3" pageNumber="556" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="C0B6EADF782CAB735FF6F89AB046F8C8" box="[418,570,1859,1885]" italics="true" pageId="3" pageNumber="556">R. kerneggeri</emphasis>
</taxonomicName>
. Abdominal 4-8 segments not visible.
</paragraph>
<paragraph id="F27D36CD782CAB745E93F8B9B73BFF40" blockId="3.[151,1437,1499,2029]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="557" pageId="3" pageNumber="556">Coloration. Body brown with black marks. Dorsal side of head brown; occiput blackish with light spots. Compound eyes light brown; cheeks and mouthparts blackish brown. Pronotum blackish with relatively small light brown marks on lateral sides and a narrow medial longitudinal stripe. Mesonotum with black posterior part and a few black spots near the middle and at anterior portion of tergite. Metanotum brown with black posterior part. Femora brown with black marks at the base. Tibiae light brown with numerous black spots. Anterior half of the first three abdominal tergites dark brown, posterior half black. Abdominal 9-10th tergites brown; cerci blackish.</paragraph>
<paragraph id="F27D36CD782BAB745E93FF01B05EFE88" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="557">Lengths (mm): Total: 11.3; head: 2.0; pronotum: 1.7; mesonotum: 1.5; metanotum: 1.05; profemora: 2.5; protibiae: 2.0; tarsi: 1,6; cerci 0,21.</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745E93FEF9B0A4FE44" pageId="4" pageNumber="557" type="discussion">
<paragraph id="F27D36CD782BAB745E93FEF9B0A4FE44" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93FEF9B348FED4" bold="true" box="[199,308,295,321]" pageId="4" pageNumber="557">Remarks</emphasis>
. It is the first reported heelwalker finding in syninclusion with ant. It could be important because
<emphasis id="C0B6EADF782BAB745EC3FE95B3E0FEF1" box="[151,412,331,357]" italics="true" pageId="4" pageNumber="557">Ctenobetylus goepperti</emphasis>
had aphid symbionts, and in the laboratory heelwalker nymphs were fed daily with aphids (
<bibRefCitation id="96534B3C782BAB745ECBFEB1B30FFE1C" author="Conti, E. &amp; Mulder, C. &amp; Lombardo, C. F. &amp; Marletta, A. &amp; Costa, G." box="[159,371,367,393]" pageId="4" pageNumber="557" pagination="185 - 200" refId="ref4054" refString="Conti, E., Mulder, C., Lombardo, C. F., Marletta, A. &amp; Costa, G. (2019) The present is the key to the past: how living fossils in Namibia share insights on the insects of Tertiary European forests. African Entomology, 27 (1), 185 - 200. https: // doi. org / 10.4001 / 003.027.0185" type="journal article" year="2019">
Conti
<emphasis id="C0B6EADF782BAB745EBDFEB1B362FE1C" box="[233,286,367,393]" italics="true" pageId="4" pageNumber="557">et al</emphasis>
., 2019
</bibRefCitation>
). Native aphids are very rare in
<collectingCountry id="8AD5765D782BAB745CBBFEB1B1FDFE1C" box="[751,897,367,393]" name="South Africa" pageId="4" pageNumber="557">South Africa</collectingCountry>
but in European amber forest they were quite common (
<bibRefCitation id="96534B3C782BAB745F5CFE4DB020FE38" author="Perkovsky, E. E. &amp; Wegierek, P." box="[264,604,403,429]" pageId="4" pageNumber="557" pagination="297 - 310" refId="ref5224" refString="Perkovsky, E. E. &amp; Wegierek, P. (2018) Aphid- Buchnera- Ant symbiosis; or why are aphids rare in the tropics and very rare further south? Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107 (2 / 3), 297 - 310. https: // doi. org / 10.1017 / S 1755691017000147" type="journal article" year="2018">Perkovsky &amp; Wegierek, 2018</bibRefCitation>
). Therefore, we propose that nymphs and adult of
<taxonomicName id="35C24D4E782BAB745AC3FE4DB74DFE38" authorityName="Zompro" authorityYear="2001" box="[1175,1329,403,429]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782BAB745AC3FE4DB74DFE38" box="[1175,1329,403,429]" italics="true" pageId="4" pageNumber="557">Raptophasma</emphasis>
</taxonomicName>
probably fed with aphids, small insects and other arthropods.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="7A6B87EB0D43FFB69CC1FC83FCE2BCB1" ID-DOI="http://doi.org/10.5281/zenodo.6032432" ID-GBIF-Taxon="135135530" ID-Zenodo-Dep="6032432" LSID="urn:lsid:plazi:treatment:7A6B87EB0D43FFB69CC1FC83FCE2BCB1" httpUri="http://treatment.plazi.org/id/7A6B87EB0D43FFB69CC1FC83FCE2BCB1" lastPageId="4" lastPageNumber="396" pageId="1" pageNumber="393">
<subSubSection id="BAD865760D43FFB39CC1FC83FEAABDE3" pageId="1" pageNumber="393" type="nomenclature">
<paragraph id="F27D36FD0D43FFB39CC1FC83FDEBBDC1" blockId="1.[151,547,870,930]" box="[151,547,870,896]" pageId="1" pageNumber="393">
<heading id="A93581910D43FFB39CC1FC83FDEBBDC1" bold="true" box="[151,547,870,896]" fontSize="11" level="1" pageId="1" pageNumber="393" reason="1">
<taxonomicName id="35C24D7E0D43FFB39CC1FC83FDEBBDC1" ID-CoL="9257ec1a-e1f6-47ee-8f89-32bc5961595e" authority="Ramadan, 1938" authorityName="Ramadan" authorityYear="1938" box="[151,547,870,896]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D43FFB39CC1FC83FDEBBDC1" bold="true" box="[151,547,870,896]" pageId="1" pageNumber="393">
<emphasis id="C0B6EAEF0D43FFB39CC1FC83FE9FBDC1" bold="true" box="[151,343,870,896]" italics="true" pageId="1" pageNumber="393">Aristeus alcocki</emphasis>
<bibRefCitation id="96534B0C0D43FFB39D09FC83FDEBBDC1" author="Ramadan" box="[351,547,870,896]" pageId="1" pageNumber="393" refString="Ramadan, M. M. (1938) Crustacea: Penaeidae. John Murray Expedition 1933 - 1934 Scientific Reports, 5 (3), 35 - 76." type="journal article" year="1938">Ramadan, 1938</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F27D36FD0D43FFB39CC1FC6CFEAABDE3" blockId="1.[151,547,870,930]" box="[151,354,905,930]" pageId="1" pageNumber="393">
(
<figureCitation id="6AF92A780D43FFB39CC9FC6CFF24BDE3" box="[159,236,905,930]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1965]" captionTargetBox="[151,1436,956,1920]" captionTargetId="figure@2.[151,1436,956,1920]" captionTargetPageId="2" captionText="FIGURE 1. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 41.4 mm (DABFUK), atypical in having only two instead of three dorsal rostral teeth." httpUri="https://zenodo.org/record/1003054/files/figure.png" pageId="1" pageNumber="393">Figs. 1</figureCitation>
,
<figureCitation id="6AF92A780D43FFB39CACFC6FFE88BDE3" box="[250,320,906,930]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="1" pageNumber="393">2A, C</figureCitation>
,
<figureCitation id="6AF92A780D43FFB39D1DFC6FFE91BDE3" box="[331,345,906,930]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D43FFB39CC1FC37FB53BA23" pageId="1" pageNumber="393" type="reference_group">
<paragraph id="F27D36FD0D43FFB39CC1FC37FD1DBA65" blockId="1.[151,1436,977,1122]" pageId="1" pageNumber="393">
<treatmentCitationGroup id="D2D211D30D43FFB39CC1FC37FD1DBA65" pageId="1" pageNumber="393">
<taxonomicName id="35C24D7E0D43FFB39CC1FC37FDCEBDA6" ID-CoL="9257ec1a-e1f6-47ee-8f89-32bc5961595e" authority="Ramadan, 1938: 40" authorityName="Ramadan" authorityPageNumber="40" authorityYear="1938" box="[151,518,977,999]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D43FFB39CC1FC37FEF1BDA6" box="[151,313,977,999]" italics="true" pageId="1" pageNumber="393">Aristeus alcocki</emphasis>
<treatmentCitation id="736310EC0D43FFB39D69FC34FDCEBDA6" author="Ramadan" box="[319,518,977,999]" page="40" pageId="1" pageNumber="393" year="1938">
<bibRefCitation id="96534B0C0D43FFB39D69FC34FE2ABDA6" author="Ramadan" box="[319,482,977,999]" pageId="1" pageNumber="393" refString="Ramadan, M. M. (1938) Crustacea: Penaeidae. John Murray Expedition 1933 - 1934 Scientific Reports, 5 (3), 35 - 76." type="journal article" year="1938">Ramadan, 1938</bibRefCitation>
: 40
</treatmentCitation>
</taxonomicName>
, figs. 1, 2a, 3a (
<typeStatus id="2D79885F0D43FFB39EF3FC34FD1ABDA6" box="[677,722,977,999]" pageId="1" pageNumber="393">type</typeStatus>
locality: Gulf of Aden);
<treatmentCitation id="736310EC0D43FFB39F9BFC34FB4EBDA6" author="George" box="[973,1158,977,999]" page="340" pageId="1" pageNumber="393" year="1967">
<bibRefCitation id="96534B0C0D43FFB39F9BFC34FB9BBDA6" author="George" box="[973,1107,977,999]" pageId="1" pageNumber="393" refString="George, M. J. (1967) On a collection of penaeid prawns from the offshore waters of the south-west coast of India. In: Symposium on Crustacea, part 1. The Bangalore Press, Bangalore, pp. 337 - 346." type="book chapter" year="1967">George 1967</bibRefCitation>
: 340
</treatmentCitation>
; 1979: 26, fig. 2b;
<treatmentCitation id="736310EC0D43FFB39912FC34FEEBBA47" author="Crosnier" page="61" pageId="1" pageNumber="393" year="1978">
<bibRefCitation id="96534B0C0D43FFB39912FC34FF35BA47" author="Crosnier" pageId="1" pageNumber="393" refString="Crosnier, A. (1978) Crustaces Decapodes Peneides Aristeidae (Benthesicyminae, Aristeinae, Solenocerinae). Faune de Madagascar, 46, 1 - 197." type="journal article" year="1978">Crosnier 1978</bibRefCitation>
: 61
</treatmentCitation>
, fig. 25j;
<treatmentCitation id="736310EC0D43FFB39DDEFC15FDFEBA47" author="Holthuis" box="[392,566,1008,1030]" page="9" pageId="1" pageNumber="393" year="1980">
<bibRefCitation id="96534B0C0D43FFB39DDEFC15FDD5BA47" author="Holthuis" box="[392,541,1008,1030]" pageId="1" pageNumber="393" refString="Holthuis, L. B. (1980) FAO species catalogue. Vol. 1. Shrimps and prawns of the world. An annotated catalogue of species of interest to fisheries. FAO Fisheries Synopsis, 125 (1), 1 - 261." type="journal article" year="1980">Holthuis 1980</bibRefCitation>
: 9
</treatmentCitation>
; Susellan 1989: 54, fig. 1; Pérez
<treatmentCitation id="736310EC0D43FFB39FDAFC15FB73BA47" author="Farfante" box="[908,1211,1008,1030]" page="40" pageId="1" pageNumber="393" year="1997">
<bibRefCitation id="96534B0C0D43FFB39FDAFC15FB5CBA47" author="Farfante" box="[908,1172,1008,1030]" pageId="1" pageNumber="393" refString="Perez-Farfante, I. &amp; Kensley, B. (1997) Penaeoid and sergestoid shrimps and prawns of the world. Keys and diagnosis for the families and genera. Memoires du Museum National d'Histoire Naturelle, 175, 1 - 233." type="journal article" year="1997">Farfante &amp; Kensley 1997</bibRefCitation>
: 40
</treatmentCitation>
;
<treatmentCitation id="736310EC0D43FFB39890FC14FEE7BA65" author="De" page="206" pageId="1" pageNumber="393" year="2011">
<bibRefCitation id="96534B0C0D43FFB39890FC14FF34BA65" author="De" pageId="1" pageNumber="393" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, 85 (9), 195 - 588." type="journal article" year="2011">De Grave &amp; Fransen 2011</bibRefCitation>
: 206
</treatmentCitation>
;
<treatmentCitation id="736310EC0D43FFB39D6FFBEBFDA4BA65" author="Chakraborty" box="[313,620,1038,1061]" page="1719" pageId="1" pageNumber="393" year="2015">
<bibRefCitation id="96534B0C0D43FFB39D6FFBEBFDE5BA65" author="Chakraborty" box="[313,557,1038,1061]" pageId="1" pageNumber="393" refString="Chakraborty, R. D., Purushothaman, P., Kuberan, G., Jomon, S. &amp; Maheswarudu, G. (2015) Morphological analysis and molecular phylogeny of Aristeus alcocki Ramadan, 1938 from south-west coast of India. Indian Journal of Geo-Marine Sciences, 44 (11), 1716 - 1725." type="journal article" year="2015">
Chakraborty
<emphasis id="C0B6EAEF0D43FFB39DE8FBF5FE39BA65" box="[446,497,1038,1061]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
2015
</bibRefCitation>
: 1719
</treatmentCitation>
, figs. 45.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F27D36FD0D43FFB39CC1FBCBFB53BA23" blockId="1.[151,1436,977,1122]" pageId="1" pageNumber="393">
<treatmentCitationGroup id="D2D211D30D43FFB39CC1FBCBFB53BA23" pageId="1" pageNumber="393">
<taxonomicName id="35C24D7E0D43FFB39CC1FBCBFEBCBA02" ID-CoL="cc00a9b7-b959-48c2-b3de-6698dca654ef" authorityName="Bate" authorityYear="1888" box="[151,372,1069,1091]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D43FFB39CC1FBCBFEBCBA02" box="[151,372,1069,1091]" italics="true" pageId="1" pageNumber="393">Aristeus semidentatus</emphasis>
</taxonomicName>
<treatmentCitation id="736310EC0D43FFB39DD8FBC8FD39BA02" author="Wood-Mason" box="[398,753,1069,1091]" page="280" pageId="1" pageNumber="393" year="1891">
<bibRefCitation id="96534B0C0D43FFB39DD8FBC8FD75BA02" author="Wood-Mason" box="[398,701,1069,1091]" pageId="1" pageNumber="393" refString="Wood-Mason, J. &amp; Alcock, A. (1891) Natural history notes from H. M. Indian marine survey steamer ' Investigator', Commander R. F. Hoskyn, R. N., commanding. -- Series. II, No. 1. On the results of deep-sea dredging during the season 1890 - 91. Annals and Magazine of Natural History, 8 (6), 268 - 286, 353 - 362." type="journal article" year="1891">Wood-Mason &amp; Alcock 1891</bibRefCitation>
: 280
</treatmentCitation>
; Alcock 1901: 31; Alcock &amp; McArdle 1901: pl. 49-fig. 3;
<treatmentCitation id="736310EC0D43FFB39912FBCBFEA6BA23" author="Kemp" page="17" pageId="1" pageNumber="393" year="1912">
<bibRefCitation id="96534B0C0D43FFB39912FBCBFE80BA23" author="Kemp" pageId="1" pageNumber="393" refString="Kemp, S. &amp; Sewell, R. B. S. (1912) Notes on Decapoda in the Indian museum. III. The Species obtained by R. I. M. S. S. ' INVESTIGATOR' during the Survey Season, 1910 - 11. Records of the Indian Museum, 7, 15 - 32, pl. 1." type="journal article" year="1912">Kemp &amp; Sewell 1912</bibRefCitation>
: 17
</treatmentCitation>
;
<treatmentCitation id="736310EC0D43FFB39D2FFBA9FDFABA23" author="George" box="[377,562,1100,1122]" page="339" pageId="1" pageNumber="393" year="1967">
<bibRefCitation id="96534B0C0D43FFB39D2FFBA9FE37BA23" author="George" box="[377,511,1100,1122]" pageId="1" pageNumber="393" refString="George, M. J. (1967) On a collection of penaeid prawns from the offshore waters of the south-west coast of India. In: Symposium on Crustacea, part 1. The Bangalore Press, Bangalore, pp. 337 - 346." type="book chapter" year="1967">George 1967</bibRefCitation>
: 339
</treatmentCitation>
; 1979: 27, fig. 2c;
<treatmentCitation id="736310EC0D43FFB39EA6FBA9FC6FBA23" author="Holthuis" box="[752,935,1100,1122]" page="10" pageId="1" pageNumber="393" year="1980">
<bibRefCitation id="96534B0C0D43FFB39EA6FBA9FC4CBA23" author="Holthuis" box="[752,900,1100,1122]" pageId="1" pageNumber="393" refString="Holthuis, L. B. (1980) FAO species catalogue. Vol. 1. Shrimps and prawns of the world. An annotated catalogue of species of interest to fisheries. FAO Fisheries Synopsis, 125 (1), 1 - 261." type="journal article" year="1980">Holthuis 1980</bibRefCitation>
: 10
</treatmentCitation>
(p.p.). (
<emphasis id="C0B6EAEF0D43FFB39FAFFBA8FBD7BA23" box="[1017,1055,1101,1122]" italics="true" pageId="1" pageNumber="393">non</emphasis>
<bibRefCitation id="96534B0C0D43FFB39870FBA8FB5DBA23" author="Bate" box="[1062,1173,1100,1122]" pageId="1" pageNumber="393" refString="Bate, C. S. (1881) On the Penaeidea. Annals and Magazine of Natural History, Series 5, 8, 169 - 196. https: // doi. org / 10.1080 / 00222938109487441" type="journal article" year="1881">Bate, 1881</bibRefCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D43FFB39CC1FB6BFBACBAAE" pageId="1" pageNumber="393" type="materials_examined">
<paragraph id="F27D36FD0D43FFB39CC1FB6BFBACBAAE" blockId="1.[151,1437,1166,2019]" pageId="1" pageNumber="393">
<materialsCitation id="42AA3CA00D43FFB39CC1FB6BFE44BA8A" ID-GBIF-Occurrence="1664838850" collectingDate="2017-03-20" county="Material" location="Kollam district" municipality="Sakthikulangara" pageId="1" pageNumber="393" specimenCount="2" specimenCount-female="2" stateProvince="Kerala">
<emphasis id="C0B6EAEF0D43FFB39CC1FB6BFE48BAE6" bold="true" box="[151,384,1166,1191]" pageId="1" pageNumber="393">
<collectingCounty id="1B1C4E710D43FFB39CC1FB6BFF37BAE6" box="[151,255,1166,1191]" pageId="1" pageNumber="393">Material</collectingCounty>
examined.
</emphasis>
<collectingMunicipality id="1219AC870D43FFB39DD0FB6BFDF4BAE6" box="[390,572,1166,1191]" pageId="1" pageNumber="393">Sakthikulangara</collectingMunicipality>
fishing harbor,
<location id="F71D60260D43FFB39EBAFB6BFC5FBAE6" LSID="urn:lsid:plazi:treatment:7A6B87EB0D43FFB69CC1FC83FCE2BCB1:F71D60260D43FFB39EBAFB6BFC5FBAE6" box="[748,919,1166,1191]" county="Material" municipality="Sakthikulangara" name="Kollam district" pageId="1" pageNumber="393" stateProvince="Kerala">Kollam district</location>
,
<collectingRegion id="3006F81F0D43FFB39FF7FB6BFC26BAE6" box="[929,1006,1166,1191]" country="India" name="Kerala" pageId="1" pageNumber="393">Kerala</collectingRegion>
,
<collectingDate id="9638E9D50D43FFB39FAFFB6AFB6EBAE6" box="[1017,1190,1166,1191]" pageId="1" pageNumber="393" value="2017-03-20">
<date id="867C103D0D43FFB39FAFFB6AFB6EBAE6" box="[1017,1190,1166,1191]" pageId="1" pageNumber="393" value="2017-03-20">20 March 2017</date>
</collectingDate>
,
<specimenCount id="E4C4FD740D43FFB398E6FB6AFAD3BAE6" box="[1200,1307,1166,1191]" pageId="1" pageNumber="393" type="female">2 females</specimenCount>
cl 38.3 and
<quantity id="353A9B180D43FFB39CC1FB56FF33BA8B" box="[151,251,1203,1227]" metricMagnitude="-2" metricUnit="m" metricValue="4.14" pageId="1" pageNumber="393" unit="mm" value="41.4">41.4 mm</quantity>
(DABFUK)
</materialsCitation>
; 6 females cl
<quantity id="353A9B180D43FFB39E71FB57FD02BA8B" box="[551,714,1202,1227]" metricMagnitude="-2" metricUnit="m" metricValue="4.115" metricValueMax="5.26" metricValueMin="2.9699999999999998" pageId="1" pageNumber="393" unit="mm" value="41.15" valueMax="52.6" valueMin="29.7">29.752.6 mm</quantity>
(DABFUK), 10 females cl
<quantity id="353A9B180D43FFB39855FB57FB6EBA8B" box="[1027,1190,1202,1227]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="5.279999999999999" metricValueMin="3.12" pageId="1" pageNumber="393" unit="mm" value="42.0" valueMax="52.8" valueMin="31.2">31.252.8 mm</quantity>
(DABFUK 2029), 2 females cl 31.8 and
<quantity id="353A9B180D43FFB39D21FB32FE13BAAF" box="[375,475,1239,1263]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="1" pageNumber="393" unit="mm" value="32.9">32.9 mm</quantity>
(DABFUK), 2 females cl 30.5 and
<quantity id="353A9B180D43FFB39F3AFB33FC18BAAF" box="[876,976,1238,1263]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="1" pageNumber="393" unit="mm" value="36.3">36.3 mm</quantity>
(DABFUK).
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D43FFB39C91FB1FFC8AB9FE" pageId="1" pageNumber="393" type="diagnosis">
<paragraph id="F27D36FD0D43FFB39C91FB1FFE2AB912" blockId="1.[151,1437,1166,2019]" pageId="1" pageNumber="393">
<emphasis id="C0B6EAEF0D43FFB39C91FB1FFE89BB52" bold="true" box="[199,321,1274,1299]" pageId="1" pageNumber="393">Diagnosis.</emphasis>
Integument glabrous. Rostrum usually bearing only 3 dorsal teeth at base, very long and far overreaching scaphocerite in females but short and failed to reach distal margin of scaphocerite in males. Carapace with lower end of cervical carina located at level slightly lower than antennal spine and far from branchiostegal carina (
<figureCitation id="6AF92A780D43FFB39CBAFA83FE8EBB3E" box="[236,326,1382,1407]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="1" pageNumber="393">Fig. 2C</figureCitation>
); distance between lower end of cervical carina and branchiostegal carina similar to that between antennal and branchiostegal spines. Pleurobranchs of thoracic somites IV to VII (corresponding to pereiopods I to IV) tiny, greatly reduced to simple papillae and without pinnules (
<figureCitation id="6AF92A780D43FFB39FDAFA4BFC21BB86" box="[908,1001,1454,1479]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="1" pageNumber="393">Fig. 2A</figureCitation>
). Pereiopod I (
<figureCitation id="6AF92A780D43FFB398C5FA4BFB25BB86" box="[1171,1261,1454,1479]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">Fig. 3A</figureCitation>
) with subdistal movable spine on lateral surface of merus, chela 1.141.46 times as long as carpus, fingers 1.511.83 times longer than palm; carpus with 1424 photophores, propodus with 47 photophores, dactylus without photophores. Pereiopod II (
<figureCitation id="6AF92A780D43FFB39D65F9FFFE41B872" box="[307,393,1562,1587]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">Fig. 3B</figureCitation>
) also with sudistal movable spine on lateral surface of merus, chela 0.931.11 times as long as carpus, fingers 1.271.62 times longer than palm; carpus with 915 photophores, propodus with 36 photophores, dactylus lacking photophore. Pereiopod III (
<figureCitation id="6AF92A780D43FFB39F4DF987FCA7B83A" box="[795,879,1634,1659]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">Fig. 3C</figureCitation>
) without subdistal movable spine on merus, chela 0.700.88 time as long as carpus, fingers 1.091.38 times longer than palm; carpus bearing 613 photophores, propodus with 36 photophores and dactylus without photophore. Pereiopod IV (
<figureCitation id="6AF92A780D43FFB39860F94FFB47B882" box="[1078,1167,1706,1731]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">Fig. 3D</figureCitation>
) with carpus 0.991.10 times as long as merus; carpus with 1830 photophores, propodus with 1930 photophores while dactylus without photophore. Pereiopod V (
<figureCitation id="6AF92A780D43FFB39D89F917FDF5B94A" box="[479,573,1778,1803]" captionStart="FIGURE 3" captionStartId="4.[151,250,1682,1704]" captionTargetBox="[163,1422,823,1649]" captionTargetId="figure@4.[151,1436,807,1660]" captionTargetPageId="4" captionText="FIGURE 3. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 52.6 mm (DABFUK). A, left pereiopod I, chela to distal part of merus; B, left pereiopod II, chela to distal part of merus; C, left pereiopod III, chela to distal part of merus; D, left pereiopod IV, dactylus to distal part of merus; E, left pereiopod V, dactylus to distal part of merus. Dots representing photophores. Scale = 2 mm." httpUri="https://zenodo.org/record/1003058/files/figure.png" pageId="1" pageNumber="393">Fig. 3E</figureCitation>
) with carpus 1.081.28 times longer than merus, carpus with 1432 photophores, propodus with 2544 photophores and dactylus with 921 photophores. Abdominal somite III without posteromedian spine.
</paragraph>
<paragraph id="F27D36FD0D43FFB39C91F8BBFC8AB9FE" blockId="1.[151,1437,1166,2019]" pageId="1" pageNumber="393">
<emphasis id="C0B6EAEF0D43FFB39C91F8BBFE80B936" bold="true" box="[199,328,1886,1911]" pageId="1" pageNumber="393">Coloration</emphasis>
(
<figureCitation id="6AF92A780D43FFB39D08F8BBFE6DB936" box="[350,421,1886,1911]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1965]" captionTargetBox="[151,1436,956,1920]" captionTargetId="figure@2.[151,1436,956,1920]" captionTargetPageId="2" captionText="FIGURE 1. Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, female cl 41.4 mm (DABFUK), atypical in having only two instead of three dorsal rostral teeth." httpUri="https://zenodo.org/record/1003054/files/figure.png" pageId="1" pageNumber="393">Fig. 1</figureCitation>
). Carapace and cephalic appendages generally reddish, eyes black brown. Abdomen pinkish red and with posterior border of each somite more reddish. Pereiopods pinkish red, photophores purplish red with iridescence. Pleopods and tail fan generally reddish.
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D43FFB39C91F82FFB28B9A3" box="[199,1248,1994,2019]" pageId="1" pageNumber="393" type="distribution">
<paragraph id="F27D36FD0D43FFB39C91F82FFB28B9A3" blockId="1.[151,1437,1166,2019]" box="[199,1248,1994,2019]" pageId="1" pageNumber="393">
<emphasis id="C0B6EAEF0D43FFB39C91F82FFEA8B9A2" bold="true" box="[199,352,1994,2019]" pageId="1" pageNumber="393">Distribution.</emphasis>
Gulf of Aden, Arabian Sea,
<collectingCountry id="8AD5766D0D43FFB39EF4F82FFD14B9A2" box="[674,732,1994,2019]" name="India" pageId="1" pageNumber="393">India</collectingCountry>
and Bay of Bengal, at depths of
<quantity id="353A9B180D43FFB3981DF82FFB13B9A3" box="[1099,1243,1994,2019]" metricMagnitude="2" metricUnit="m" metricValue="6.685" metricValueMax="10.87" metricValueMin="2.5" pageId="1" pageNumber="393" unit="m" value="668.5" valueMax="1087.0" valueMin="250.0">2501087 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D40FFB69C91FF72FCE2BCB1" lastPageId="4" lastPageNumber="396" pageId="2" pageNumber="394" type="discussion">
<paragraph id="F27D36FD0D40FFB09C91FF72FCF8BDE4" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="394">
<emphasis id="C0B6EAEF0D40FFB09C91FF72FEF3BEF1" bold="true" box="[199,315,151,176]" pageId="2" pageNumber="394">Remarks.</emphasis>
Although both
<taxonomicName id="35C24D7E0D40FFB09DBCFF7DFC96BEF1" authority="Bate, 1881" authorityName="Bate" authorityYear="1881" box="[490,862,151,176]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB09DBCFF7DFD16BEF1" box="[490,734,151,176]" italics="true" pageId="2" pageNumber="394">Aristeus semidentatus</emphasis>
<bibRefCitation id="96534B0C0D40FFB09EB3FF7DFC96BEF1" author="Bate" box="[741,862,152,176]" pageId="2" pageNumber="394" refString="Bate, C. S. (1881) On the Penaeidea. Annals and Magazine of Natural History, Series 5, 8, 169 - 196. https: // doi. org / 10.1080 / 00222938109487441" type="journal article" year="1881">Bate, 1881</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="35C24D7E0D40FFB09FC2FF7DFBCCBEF1" authorityName="Ramadan" authorityYear="1938" box="[916,1028,151,176]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09FC2FF7DFBCCBEF1" box="[916,1028,151,176]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
have often been reported from
<collectingCountry id="8AD5766D0D40FFB09934FF72FA54BEF1" box="[1378,1436,151,176]" name="India" pageId="2" pageNumber="394">India</collectingCountry>
(e.g.,
<bibRefCitation id="96534B0C0D40FFB09C80FF59FEA1BE94" author="George" box="[214,361,188,213]" pageId="2" pageNumber="394" refString="George, M. J. (1967) On a collection of penaeid prawns from the offshore waters of the south-west coast of India. In: Symposium on Crustacea, part 1. The Bangalore Press, Bangalore, pp. 337 - 346." type="book chapter" year="1967">George 1967</bibRefCitation>
,
<bibRefCitation id="96534B0C0D40FFB09D22FF59FE78BE95" author="George" box="[372,432,188,212]" pageId="2" pageNumber="394" refString="George, M. J. (1979) Taxonomy of Indian prawns (Penaeidae, Crustacea, Decapoda) In: Kurian, C. V., Sharma, G. S., Mohandas, A. &amp; Antony, A. (Eds.), Contributions to Marine Sciences Dedicated to Dr. C. U. Kurian on the Occasion of his Sixtieth Birthday. Prof. Kurian's Shashtyabdhapoorthi Committee, Cochin, pp. 21 - 59." type="book chapter" year="1979">1979</bibRefCitation>
),
<bibRefCitation id="96534B0C0D40FFB09D95FF59FDB0BE95" author="Suseelan" box="[451,632,188,213]" pageId="2" pageNumber="394" refString="Suseelan, C. (1989) Taxonomic notes on a potentially commercial deep-sea prawn from the southwest coast of India. Journal of the Marine Biological Association of India, 31 (1 &amp; 2), 54 - 58." type="journal article" year="1989">Suseelan (1989)</bibRefCitation>
argued that only
<taxonomicName id="35C24D7E0D40FFB09F6FFF59FC61BE94" authorityName="Ramadan" authorityYear="1938" box="[825,937,188,213]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09F6FFF59FC61BE94" box="[825,937,188,213]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
is found in Indian waters (also see
<bibRefCitation id="96534B0C0D40FFB09964FF59FF1BBEB9" author="Ramadan" pageId="2" pageNumber="394" refString="Ramadan, M. M. (1938) Crustacea: Penaeidae. John Murray Expedition 1933 - 1934 Scientific Reports, 5 (3), 35 - 76." type="journal article" year="1938">Ramadan 1938</bibRefCitation>
). Nevertheless, recent shrimp checklists from
<collectingCountry id="8AD5766D0D40FFB09E80FF3AFCC7BEB9" box="[726,783,223,248]" name="India" pageId="2" pageNumber="394">India</collectingCountry>
still included
<taxonomicName id="35C24D7E0D40FFB09FF8FF05FB97BEB9" authorityName="Bate" authorityYear="1881" box="[942,1119,223,248]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB09FF8FF05FB97BEB9" box="[942,1119,223,248]" italics="true" pageId="2" pageNumber="394">A. semidentatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="96534B0C0D40FFB0983BFF3AFA5FBEB9" author="Radhakrishnan" box="[1133,1431,223,248]" pageId="2" pageNumber="394" refString="Radhakrishnan, E. V., Deshmukh, V. D., Maheswarudu, G., Josileen, J., Dineshababu, A. P., Philipose, K. K., Sarada, P. T., Pillai, S. L., Saleela, K. N., Chakraborty, R., Dash, G., Sajeev, C. K., Thirumilu, P., Sridhara, B., Muniuyappa, Y., Sawant, A. D., Vaidya, N. G., Johny, R. D., Verma, J. B., Baby, P. K., Unnikrishnan, C., Ramachandran, N. P., Vairamani, A., Palanichamy, A., Radhakrishnan, M. &amp; Raju, B. (2012) Prawn fauna (Crustacea: Decapoda) of India-An annotated checklist of the Penaeoid, Segestoid, Stenopodid and Caridean prawns. Journal of Marine Biological Association of India, 54 (1), 50 - 72." type="journal article" year="2012">
Radhakrishnan
<emphasis id="C0B6EAEF0D40FFB09948FF04FA9EBEB9" box="[1310,1366,223,248]" italics="true" pageId="2" pageNumber="394">et al.</emphasis>
2012
</bibRefCitation>
;
<bibRefCitation id="96534B0C0D40FFB09CC1FEE1FEB0BF5C" author="Samuel" box="[151,376,260,285]" pageId="2" pageNumber="394" refString="Samuel, V. K. D., Sreeraj, C. R., Krishnan, P., Parthiban, C., Sekar, V., Chamundeeswari, K., Immanuel, T., Shesdev, P., Purvaja, R. &amp; Ramesh, R. (2016) An updated checklist of shrimps on the Indian coast. Journal of Threatened Taxa, 8, 8977 - 8988. https: // doi. org / 10.11609 / jott. 2628.8.7.8977 - 8988" type="journal article" year="2016">
Samuel
<emphasis id="C0B6EAEF0D40FFB09CA3FEE0FEF9BF5C" box="[245,305,260,285]" italics="true" pageId="2" pageNumber="394">et al.</emphasis>
2016
</bibRefCitation>
). The abundant material of
<taxonomicName id="35C24D7E0D40FFB09EEBFEE1FCD1BF5A" authorityName="Duvernoy" authorityYear="1840" box="[701,793,260,283]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EAEF0D40FFB09EEBFEE1FCD1BF5A" box="[701,793,260,283]" italics="true" pageId="2" pageNumber="394">Aristeus</emphasis>
</taxonomicName>
observed in Sakthikulangara fishing harbor during this work contained only
<taxonomicName id="35C24D7E0D40FFB09DCEFECDFDC7BF01" authorityName="Ramadan" authorityYear="1938" box="[408,527,295,320]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09DCEFECDFDC7BF01" box="[408,527,295,320]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
. Whether
<taxonomicName id="35C24D7E0D40FFB09EC7FECDFC81BF01" authorityName="Bate" authorityYear="1881" box="[657,841,295,320]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB09EC7FECDFC81BF01" box="[657,841,295,320]" italics="true" pageId="2" pageNumber="394">A. semidentatus</emphasis>
</taxonomicName>
actually occurs in Indian water still needs to be confirmed. As mentioned above, both the relative lengths of the different segments of the pereiopods and number of photophores have been used to separate the species of
<taxonomicName id="35C24D7E0D40FFB09F72FE95FC48BFC6" authorityName="Duvernoy" authorityYear="1840" box="[804,896,368,391]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EAEF0D40FFB09F72FE95FC48BFC6" box="[804,896,368,391]" italics="true" pageId="2" pageNumber="394">Aristeus</emphasis>
</taxonomicName>
. These characters, however, are known to have large variations, including geographical variations (
<bibRefCitation id="96534B0C0D40FFB09E8AFE71FC4ABFED" author="Crosnier" box="[732,898,404,429]" pageId="2" pageNumber="394" refString="Crosnier, A. (1978) Crustaces Decapodes Peneides Aristeidae (Benthesicyminae, Aristeinae, Solenocerinae). Faune de Madagascar, 46, 1 - 197." type="journal article" year="1978">Crosnier 1978</bibRefCitation>
,
<bibRefCitation id="96534B0C0D40FFB09FD8FE71FC01BFEC" author="Crosnier" box="[910,969,404,429]" pageId="2" pageNumber="394" refString="Crosnier, A. (1985) Penaeoid shrimps (Benthesicymidae, Aristeidae, Solenoceridae, Siyoniidae) collected in Indonesia during the Corindon II and IV expeditions. Marine Research in Indonesia, 24, 19 - 47." type="journal article" year="1985">1985</bibRefCitation>
,
<bibRefCitation id="96534B0C0D40FFB09F83FE71FBD4BFED" author="Crosnier" box="[981,1052,404,428]" pageId="2" pageNumber="394" refString="Crosnier, A. (1994 a) Crustacea Decapods: Penaeoidea recoltes lors de la campagne KARUBAR en Indonesie. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 12. Memories du Museum national d'Histoire naturelle, 161, pp. 351 - 365." type="journal article" year="1994" yearSuffix="a">1994a</bibRefCitation>
,
<bibRefCitation id="96534B0C0D40FFB0987EFE71FBF1BFEC" author="Crosnier" box="[1064,1081,404,429]" pageId="2" pageNumber="394" refString="Crosnier, A. (1994 b) Crustacea Decapods: Penaeoidea, a l'exclusion des Sicyoniidae recoltes dans la zone economique des iles Wallis et Futuna, lors de la campagne MUSORSTOM 7. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 12. Memories du Museum national d'Histoire naturelle, 161, 367 - 373." type="journal article" year="1994" yearSuffix="b">b</bibRefCitation>
;
<bibRefCitation id="96534B0C0D40FFB09810FE71FB1EBFED" author="Komai" box="[1094,1238,404,429]" pageId="2" pageNumber="394" refString="Komai, T. (1993) Deep-sea shrimps of the family Aristeidae (Decapoda: Dendrobranchiata) from northern Japan, with the description of a new species of the genus Aristeus. Crustacean Research, 22, 21 - 34. https: // doi. org / 10.18353 / crustacea. 22.0 _ 21" type="journal article" year="1993">Komai 1993</bibRefCitation>
). The availability of many fresh material of
<taxonomicName id="35C24D7E0D40FFB09DEDFE5DFDE3BF91" authorityName="Ramadan" authorityYear="1938" box="[443,555,439,464]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09DEDFE5DFDE3BF91" box="[443,555,439,464]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
enables the assessment of the taxonomic values of these characters at least for distinguishing the six species in the Indo-West Pacific. Table 1 summaries the known phophophore counts and ratios of the different segments at the pereiopods in these species.
<taxonomicName id="35C24D7E0D40FFB09FC9FDE5FAECBC59" authority="(Risso, 1816)" authorityName="Risso" authorityYear="1816" baseAuthorityName="Risso" baseAuthorityYear="1816" box="[927,1316,511,536]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C0B6EAEF0D40FFB09FC9FDE5FB48BC59" box="[927,1152,512,536]" italics="true" pageId="2" pageNumber="394">Aristeus antennatus</emphasis>
(
<bibRefCitation id="96534B0C0D40FFB098C4FE1AFAD5BC59" author="Risso" box="[1170,1309,511,536]" pageId="2" pageNumber="394" refString="Risso, A. (1816) Histoire naturelles des Crustaces des environs de Nice. A la Librairie grecquelatine-allemande, Paris, 175 pp., 3 pls." type="book" year="1816">Risso, 1816</bibRefCitation>
)
</taxonomicName>
has much more numerous photophores than the other species of the genus, while
<taxonomicName id="35C24D7E0D40FFB09F8CFDC1FAF7BC7D" authority="Ramadan, 1938" authorityName="Ramadan" authorityYear="1938" box="[986,1343,548,573]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="mabahissae">
<emphasis id="C0B6EAEF0D40FFB09F8CFDC1FB49BC7C" box="[986,1153,548,573]" italics="true" pageId="2" pageNumber="394">A. mabahissae</emphasis>
<bibRefCitation id="96534B0C0D40FFB098DAFDC1FAF7BC7D" author="Ramadan" box="[1164,1343,548,573]" pageId="2" pageNumber="394" refString="Ramadan, M. M. (1938) Crustacea: Penaeidae. John Murray Expedition 1933 - 1934 Scientific Reports, 5 (3), 35 - 76." type="journal article" year="1938">Ramadan, 1938</bibRefCitation>
</taxonomicName>
has the smallest number of photophores. On the other hand,
<taxonomicName id="35C24D7E0D40FFB09EA0FDADFB8BBC21" authority="Komai, 1993" authorityName="Komai" authorityYear="1993" box="[758,1091,583,608]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="pallidicauda">
<emphasis id="C0B6EAEF0D40FFB09EA0FDADFC6EBC21" box="[758,934,583,608]" italics="true" pageId="2" pageNumber="394">A. pallidicauda</emphasis>
<bibRefCitation id="96534B0C0D40FFB09FF8FDA2FB8BBC21" author="Komai" box="[942,1091,583,608]" pageId="2" pageNumber="394" refString="Komai, T. (1993) Deep-sea shrimps of the family Aristeidae (Decapoda: Dendrobranchiata) from northern Japan, with the description of a new species of the genus Aristeus. Crustacean Research, 22, 21 - 34. https: // doi. org / 10.18353 / crustacea. 22.0 _ 21" type="journal article" year="1993">Komai, 1993</bibRefCitation>
</taxonomicName>
is unique in the carpi of both pereiopods I and II lacking photophores, thought it generally has more photophores on the posterior two pereiopods than most other species except
<taxonomicName id="35C24D7E0D40FFB09E56FD75FD55BCE9" baseAuthorityName="Risso" baseAuthorityYear="1816" box="[512,669,656,680]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C0B6EAEF0D40FFB09E56FD75FD55BCE9" box="[512,669,656,680]" italics="true" pageId="2" pageNumber="394">A. antennatus</emphasis>
</taxonomicName>
.
<taxonomicName id="35C24D7E0D40FFB09EFBFD75FC23BCE9" authority="(Bate, 1881)" authorityName="Bate" authorityYear="1881" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[685,1003,655,680]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D40FFB09EFBFD75FC9BBCE9" box="[685,851,655,680]" italics="true" pageId="2" pageNumber="394">Aristeus virilis</emphasis>
(
<bibRefCitation id="96534B0C0D40FFB09F33FD75FC2BBCE9" author="Bate" box="[869,995,656,680]" pageId="2" pageNumber="394" refString="Bate, C. S. (1881) On the Penaeidea. Annals and Magazine of Natural History, Series 5, 8, 169 - 196. https: // doi. org / 10.1080 / 00222938109487441" type="journal article" year="1881">Bate, 1881</bibRefCitation>
)
</taxonomicName>
also differs from other species in the dactyli of pereiopods II and IV sometimes bearing photophores but generally has fewer photophores than most of the other species. Nevertheless, its photophore counts always overlap with those of other species. The photophore counts of the remaining two species
<taxonomicName id="35C24D7E0D40FFB09E13FD19FD32BD54" authorityName="Bate" authorityYear="1881" box="[581,762,764,789]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB09E13FD19FD32BD54" box="[581,762,764,789]" italics="true" pageId="2" pageNumber="394">A. semidentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="35C24D7E0D40FFB09F6EFD19FC64BD54" authorityName="Ramadan" authorityYear="1938" box="[824,940,764,789]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09F6EFD19FC64BD54" box="[824,940,764,789]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
largely overlap with each other though
<taxonomicName id="35C24D7E0D40FFB099D2FD19FEE1BD79" authorityName="Bate" authorityYear="1881" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB099D2FD19FEE1BD79" italics="true" pageId="2" pageNumber="394">A. semidentatus</emphasis>
</taxonomicName>
generally has fewer photophores as compared to
<taxonomicName id="35C24D7E0D40FFB09F06FCC5FC77BD79" authorityName="Ramadan" authorityYear="1938" box="[848,959,799,824]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09F06FCC5FC77BD79" box="[848,959,799,824]" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
. At present there is a discrete difference in the number of photophores only at the propodus of pereiopod IV, and it is
<quantity id="353A9B180D40FFB09F9BFCA1FBF8BD1C" box="[973,1072,836,861]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="2" pageNumber="394" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">1518 in</quantity>
<taxonomicName id="35C24D7E0D40FFB09863FCA1FB2EBD1C" authorityName="Bate" authorityYear="1881" box="[1077,1254,836,861]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D40FFB09863FCA1FB2EBD1C" box="[1077,1254,836,861]" italics="true" pageId="2" pageNumber="394">A. semidentatus</emphasis>
</taxonomicName>
and
<quantity id="353A9B180D40FFB0994DFCA1FAB6BD1C" box="[1307,1406,836,861]" metricMagnitude="-1" metricUnit="m" metricValue="6.223" metricValueMax="7.62" metricValueMin="4.826" pageId="2" pageNumber="394" unit="in" value="24.5" valueMax="30.0" valueMin="19.0">1930 in</quantity>
<taxonomicName id="35C24D7E0D40FFB099D2FCA1FF20BDC1" authorityName="Ramadan" authorityYear="1938" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB099D2FCA1FF20BDC1" italics="true" pageId="2" pageNumber="394">A. alcocki</emphasis>
</taxonomicName>
. As the difference is extremely slight, it is highly likely that this photophore count will also be merged when more is known on the variations in these two species.
</paragraph>
<caption id="A6BD66750D40FFB09CC1F872FC56B98D" httpUri="https://zenodo.org/record/1003054/files/figure.png" pageId="2" pageNumber="394" targetBox="[151,1436,956,1920]" targetPageId="2">
<paragraph id="F27D36FD0D40FFB09CC1F872FC56B98D" blockId="2.[151,1435,1943,1996]" pageId="2" pageNumber="394">
<emphasis id="C0B6EAEF0D40FFB09CC1F872FED1B9ED" bold="true" box="[151,281,1943,1965]" pageId="2" pageNumber="394">FIGURE 1.</emphasis>
<taxonomicName id="35C24D7E0D40FFB09D70F87DFD4DB9EC" authority="Ramadan, 1938" authorityName="Ramadan" authorityYear="1938" box="[294,645,1943,1965]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D40FFB09D70F87DFE18B9EC" box="[294,464,1943,1965]" italics="true" pageId="2" pageNumber="394">Aristeus alcocki</emphasis>
Ramadan, 1938
</taxonomicName>
, Sakthikulangara fishing harbor, southwestern India, female cl 41.4 mm (DABFUK), atypical in having only two instead of three dorsal rostral teeth.
</paragraph>
</caption>
<caption id="A6BD66750D41FFB19CC1FABCFEEBBBAB" httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="3" pageNumber="395" targetBox="[187,1412,209,1314]" targetPageId="3">
<paragraph id="F27D36FD0D41FFB19CC1FABCFEEBBBAB" blockId="3.[151,1436,1369,1514]" pageId="3" pageNumber="395">
<emphasis id="C0B6EAEF0D41FFB19CC1FABCFEDBBB2F" bold="true" box="[151,275,1369,1391]" pageId="3" pageNumber="395">FIGURE 2.</emphasis>
A, C,
<taxonomicName id="35C24D7E0D41FFB19D00FABFFD68BB2E" authority="Ramadan, 1938" authorityName="Ramadan" authorityYear="1938" box="[342,672,1369,1391]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D41FFB19D00FABFFE31BB2E" box="[342,505,1369,1391]" italics="true" pageId="3" pageNumber="395">Aristeus alcocki</emphasis>
Ramadan, 1938
</taxonomicName>
, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK)); B, D,
<taxonomicName id="35C24D7E0D41FFB19E91FA9CFC6DBBCF" authority="(Bate, 1881)" authorityName="Bate" authorityYear="1881" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[711,933,1400,1422]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D41FFB19E91FA9CFCD6BBCF" box="[711,798,1400,1422]" italics="true" pageId="3" pageNumber="395">A. virilis</emphasis>
(Bate, 1881)
</taxonomicName>
, Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU
<accessionNumber id="ED91AB1E0D41FFB19D1AFA73FE6EBBED" box="[332,422,1430,1452]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M02048" pageId="3" pageNumber="395" type="EnaNcbi">M02048</accessionNumber>
). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a= arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm.
</paragraph>
</caption>
<paragraph id="F27D36FD0D41FFB69C91F9F1FCE8BEB9" blockId="3.[151,1437,1556,2013]" lastBlockId="4.[151,1436,151,752]" lastPageId="4" lastPageNumber="396" pageId="3" pageNumber="395">
The ratios of the various segments of the pereiopods have even larger overlapping amongst the Indo-West Pacific species, though there is a tendency of longer or shorter in different species. Usually the chela and fingers of the anterior three pereiopods are shorter in
<taxonomicName id="35C24D7E0D41FFB19EC8F9B8FC8FB834" authorityName="Ramadan" authorityYear="1938" box="[670,839,1628,1653]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="mabahissae">
<emphasis id="C0B6EAEF0D41FFB19EC8F9B8FC8FB834" box="[670,839,1628,1653]" italics="true" pageId="3" pageNumber="395">A. mabahissae</emphasis>
</taxonomicName>
and
<emphasis id="C0B6EAEF0D41FFB19FDEF9B8FB8EB834" box="[904,1094,1628,1653]" italics="true" pageId="3" pageNumber="395">
<taxonomicName id="35C24D7E0D41FFB19FDEF9B8FB8AB834" authorityName="Bate" authorityYear="1881" box="[904,1090,1628,1653]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="semidentatus">A. semidentatus</taxonomicName>
.
</emphasis>
Moreover, the carpus of the pereiopod IV is also somewhat shorter for
<taxonomicName id="35C24D7E0D41FFB19EC6F964FCF0B8D8" authorityName="Ramadan" authorityYear="1938" box="[656,824,1664,1689]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="mabahissae">
<emphasis id="C0B6EAEF0D41FFB19EC6F964FCF0B8D8" box="[656,824,1664,1689]" italics="true" pageId="3" pageNumber="395">A. mabahissae</emphasis>
</taxonomicName>
. Other than the photophore counts and ratios of the segments in the pereiopods,
<taxonomicName id="35C24D7E0D41FFB19D85F940FDFBB8FC" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[467,563,1700,1725]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D41FFB19D85F940FDFBB8FC" box="[467,563,1700,1725]" italics="true" pageId="3" pageNumber="395">A. virilis</emphasis>
</taxonomicName>
and
<taxonomicName id="35C24D7E0D41FFB19E3EF940FD10B8FC" authorityName="Ramadan" authorityYear="1938" box="[616,728,1700,1725]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D41FFB19E3EF940FD10B8FC" box="[616,728,1700,1725]" italics="true" pageId="3" pageNumber="395">A. alcocki</emphasis>
</taxonomicName>
possess some unique characters amongst the Indo-West Pacific species of the genus.
<taxonomicName id="35C24D7E0D41FFB19DD5F92CFDEFB8A0" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[387,551,1736,1761]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D41FFB19DD5F92CFDEFB8A0" box="[387,551,1736,1761]" italics="true" pageId="3" pageNumber="395">Aristeus virilis</emphasis>
</taxonomicName>
is very distinctive in the pubescent body integument (versus completely naked) and the possession of a movable subdistal spine on the merus of the pereiopod III (versus lacking such a spine). In
<taxonomicName id="35C24D7E0D41FFB19CC1F8F4FECFB968" authorityName="Ramadan" authorityYear="1938" box="[151,263,1808,1833]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D41FFB19CC1F8F4FECFB968" box="[151,263,1808,1833]" italics="true" pageId="3" pageNumber="395">A. alcocki</emphasis>
</taxonomicName>
the pleurobranchs of thoracic somites IV to VII are greatly reduced to minute papillae without pinnules (
<figureCitation id="6AF92A780D41FFB19CC9F8D1FF31B90C" box="[159,249,1844,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="3" pageNumber="395">Fig. 2A</figureCitation>
, versus reduced but still obvious and feather-like with many distinct pinnules or gill filaments,
<figureCitation id="6AF92A780D41FFB19961F8D1FA47B90C" box="[1335,1423,1844,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="3" pageNumber="395">Fig. 2B</figureCitation>
). This difference in the gills, however, are difficult to observe in the field and without dissection. Nevertheless, it is found that the position of the lower end of the cervical carina relative to the branchiostegal carina as suggested by
<bibRefCitation id="96534B0C0D41FFB19CC1F845FE83B9F9" author="Crosnier" box="[151,331,1952,1977]" pageId="3" pageNumber="395" refString="Crosnier, A. (1978) Crustaces Decapodes Peneides Aristeidae (Benthesicyminae, Aristeinae, Solenocerinae). Faune de Madagascar, 46, 1 - 197." type="journal article" year="1978">Crosnier (1978)</bibRefCitation>
is a very useful character to separate
<taxonomicName id="35C24D7E0D41FFB19F54F844FCBCB9F8" authorityName="Ramadan" authorityYear="1938" box="[770,884,1952,1977]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D41FFB19F54F844FCBCB9F8" box="[770,884,1952,1977]" italics="true" pageId="3" pageNumber="395">A. alcocki</emphasis>
</taxonomicName>
from the other Indo-West Pacific species of the genus. The distance between the lower end of the cervical carina and the branchiostegal carina is nearly as long as the length between the antennal and branchiostegal spines in
<taxonomicName id="35C24D7E0D46FFB69F32FF7DFC10BEF1" authorityName="Ramadan" authorityYear="1938" box="[868,984,151,176]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D46FFB69F32FF7DFC10BEF1" box="[868,984,151,176]" italics="true" pageId="4" pageNumber="396">A. alcocki</emphasis>
</taxonomicName>
(
<figureCitation id="6AF92A780D46FFB69FBCFF72FB8CBEF1" box="[1002,1092,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="4" pageNumber="396">Fig. 2C</figureCitation>
). In all the other species, the distance between the lower end of the cervical carina and the branchiostegal carina is only about half the length between the antennal and branchiostegal spines (
<figureCitation id="6AF92A780D46FFB69EEFFF3AFCDCBEB9" box="[697,788,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1369,1391]" captionTargetBox="[187,1412,209,1314]" captionTargetId="figure@3.[151,1436,193,1348]" captionTargetPageId="3" captionText="FIGURE 2. A, C, Aristeus alcocki Ramadan, 1938, Sakthikulangara fishing harbor, southwestern India, A, female cl 41.4 mm (DABFUK), C, female cl 52.6 mm (DABFUK )); B, D, A. virilis (Bate, 1881), Dasi fishing port, northeastern Taiwan, female cl 38.8 mm (NTOU M 02048). A, B, branchial chamber of thoracic segment VI and VII (corresponding to pereiopods III and IV), a = arthobranch, e = epipods, p = pleurobranch, P = pereiopod; C, D, pterygostomian region of carapace. Scales: A, B = 2 mm, C, D = 5 mm." httpUri="https://zenodo.org/record/1003056/files/figure.png" pageId="4" pageNumber="396">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="F27D36FD0D46FFB69C91FEE1FCE2BCB1" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="396">
The GenBank only has six COI sequences of species of
<taxonomicName id="35C24D7E0D46FFB69F06FEE1FC64BF5A" authorityName="Duvernoy" authorityYear="1840" box="[848,940,260,283]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EAEF0D46FFB69F06FEE1FC64BF5A" box="[848,940,260,283]" italics="true" pageId="4" pageNumber="396">Aristeus</emphasis>
</taxonomicName>
that can be used for usual DNA barcoding analysis (
<bibRefCitation id="96534B0C0D46FFB69D53FEC2FE2EBF01" author="Hebert" box="[261,486,295,320]" pageId="4" pageNumber="396" refString="Hebert, P. D. N., Cywinska, A., Ball, S. L. &amp; deWaard, J. R. (2003) Biological identifications through DNA barcodes. Proceedings of the Royal Society of London B, 270, 313 - 321. https: // doi. org / 10.1098 / rspb. 2002.2218" type="journal article" year="2003">
Hebert
<emphasis id="C0B6EAEF0D46FFB69D36FECCFE5FBF01" box="[352,407,295,320]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2003
</bibRefCitation>
,
<bibRefCitation id="96534B0C0D46FFB69DA0FEC2FCCCBF01" author="Hajibabaei" box="[502,772,295,320]" pageId="4" pageNumber="396" refString="Hajibabaei, M., Singer, G. A. C., Hebert P. D. N. &amp; Hickey, D. A. (2007) DNA barcoding: how it complements taxonomy, molecular phylogenetics and population genetics. Trends in Genetics, 23 (4), 167 - 172. https: // doi. org / 10.1016 / j. tig. 2007.02.001" type="journal article" year="2007">
Hajibabaei
<emphasis id="C0B6EAEF0D46FFB69E2AFECCFD7BBF01" box="[636,691,295,320]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
). Amongst them four are from
<taxonomicName id="35C24D7E0D46FFB698D4FECDFB30BF01" authorityName="Ramadan" authorityYear="1938" box="[1154,1272,295,320]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D46FFB698D4FECDFB30BF01" box="[1154,1272,295,320]" italics="true" pageId="4" pageNumber="396">A. alcocki</emphasis>
</taxonomicName>
, one from
<taxonomicName id="35C24D7E0D46FFB699D2FECDFEDABF25" baseAuthorityName="Risso" baseAuthorityYear="1816" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C0B6EAEF0D46FFB699D2FECDFEDABF25" italics="true" pageId="4" pageNumber="396">A. antennatus</emphasis>
</taxonomicName>
and another from
<taxonomicName id="35C24D7E0D46FFB69DA6FEA9FD93BF24" box="[496,603,332,357]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="virilus">
<emphasis id="C0B6EAEF0D46FFB69DA6FEA9FD93BF24" box="[496,603,332,357]" italics="true" pageId="4" pageNumber="396">A. virilus</emphasis>
</taxonomicName>
. It should be noted that the sequence of
<taxonomicName id="35C24D7E0D46FFB69814FEA9FB65BF24" box="[1090,1197,332,357]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="virilus">
<emphasis id="C0B6EAEF0D46FFB69814FEA9FB65BF24" box="[1090,1197,332,357]" italics="true" pageId="4" pageNumber="396">A. virilus</emphasis>
</taxonomicName>
(
<accessionNumber id="ED91AB1E0D46FFB698E9FEA9FAFDBF24" box="[1215,1333,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ928295" pageId="4" pageNumber="396">JQ928295</accessionNumber>
) used in
<bibRefCitation id="96534B0C0D46FFB69CC1FE8AFE67BFC9" author="Chakraborty" box="[151,431,367,392]" pageId="4" pageNumber="396" refString="Chakraborty, R. D., Purushothaman, P., Kuberan, G., Jomon, S. &amp; Maheswarudu, G. (2015) Morphological analysis and molecular phylogeny of Aristeus alcocki Ramadan, 1938 from south-west coast of India. Indian Journal of Geo-Marine Sciences, 44 (11), 1716 - 1725." type="journal article" year="2015">
Chakraborty
<emphasis id="C0B6EAEF0D46FFB69D7CFE94FEABBFC9" box="[298,355,367,392]" italics="true" pageId="4" pageNumber="396">et al.</emphasis>
(2015
</bibRefCitation>
: fig. 6b) for constructing a COI parsimony tree is actually not the region commonly used for COI barcoding match and therefore excluded in the present analysis. The present work generated COI sequences (&gt; 600 bp) for all the Indo-West Pacific species except
<taxonomicName id="35C24D7E0D46FFB69F97FE5DFBAABF91" baseAuthorityName="Risso" baseAuthorityYear="1816" box="[961,1122,440,464]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C0B6EAEF0D46FFB69F97FE5DFBAABF91" box="[961,1122,440,464]" italics="true" pageId="4" pageNumber="396">A. antennatus</emphasis>
</taxonomicName>
. It reveals that there is a misidentification in the GenBank sequence for “
<taxonomicName id="35C24D7E0D46FFB69E92FE39FCEEBFB4" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[708,806,476,501]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D46FFB69E92FE39FCEEBFB4" box="[708,806,476,501]" italics="true" pageId="4" pageNumber="396">A. virilis</emphasis>
</taxonomicName>
” (
<accessionNumber id="ED91AB1E0D46FFB69F12FE39FC0DBFB5" box="[836,965,476,500]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX381807" pageId="4" pageNumber="396">KX381807</accessionNumber>
, from
<collectingCountry id="8AD5766D0D46FFB69845FE39FB99BFB4" box="[1043,1105,476,501]" name="India" pageId="4" pageNumber="396">India</collectingCountry>
), which has 99.7% sequence similarity with the present material of
<taxonomicName id="35C24D7E0D46FFB69E08FDE5FD1BBC59" authorityName="Ramadan" authorityYear="1938" box="[606,723,511,536]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D46FFB69E08FDE5FD1BBC59" box="[606,723,511,536]" italics="true" pageId="4" pageNumber="396">A. alcocki</emphasis>
</taxonomicName>
. With the consideration of this misidentification, the present molecular genetic analysis confirmed that the six currently recognized taxa of
<taxonomicName id="35C24D7E0D46FFB69854FDC1FB96BC7A" authorityName="Duvernoy" authorityYear="1840" box="[1026,1118,548,571]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EAEF0D46FFB69854FDC1FB96BC7A" box="[1026,1118,548,571]" italics="true" pageId="4" pageNumber="396">Aristeus</emphasis>
</taxonomicName>
in the Indo-West Pacific are all distinct species with more than 6.9% sequence divergence (between
<taxonomicName id="35C24D7E0D46FFB69FE8FDADFBAABC21" authorityName="Ramadan" authorityYear="1938" box="[958,1122,583,608]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="mabahissae">
<emphasis id="C0B6EAEF0D46FFB69FE8FDADFBAABC21" box="[958,1122,583,608]" italics="true" pageId="4" pageNumber="396">A. mabahissae</emphasis>
</taxonomicName>
and
<taxonomicName id="35C24D7E0D46FFB698CCFDADFA83BC21" authorityName="Bate" authorityYear="1881" box="[1178,1355,583,608]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D46FFB698CCFDADFA83BC21" box="[1178,1355,583,608]" italics="true" pageId="4" pageNumber="396">A. semidentatus</emphasis>
</taxonomicName>
) in the barcoding gene COI (see
<bibRefCitation id="96534B0C0D46FFB69DEBFD89FD9DBCC5" author="Darling" box="[445,597,620,645]" pageId="4" pageNumber="396" refString="Darling, J. A. (2011) Interspecific hybridization and mitochondrial introgression in invasive Carcinus shore crabs. PLoS ONE, 6 (3), e 17828. https: // doi. org / 10.1371 / journal. pone. 0017828" type="journal article" year="2011">Darling 2011</bibRefCitation>
,
<bibRefCitation id="96534B0C0D46FFB69E34FD89FC94BCC5" author="Vergamini" box="[610,860,620,645]" pageId="4" pageNumber="396" refString="Vergamini, F. G., Pileggi, L. G. &amp; Mantelatto, F. L. (2011) Genetic variability of the Amazon river prawn Macrobrachium amazonicum (Decapoda, Caridea, Palaemonidae). Contributions to Zoology, 80 (1), 67 - 83." type="journal article" year="2011">
Vergamini
<emphasis id="C0B6EAEF0D46FFB69E89FD88FCD2BCC4" box="[735,794,620,645]" italics="true" pageId="4" pageNumber="396">et al.</emphasis>
2011
</bibRefCitation>
,
<bibRefCitation id="96534B0C0D46FFB69F3FFD88FBE4BCC4" author="Yang" box="[873,1068,620,645]" pageId="4" pageNumber="396" refString="Yang, C. H., Tsuchida, S., Fujikura, K., Fujiwara, Y., Kawato, M. &amp; Chan, T. Y. (2016) Connectivity of the squat lobsters Shinkaia crosnieri (Crustacea: Decapoda: Galatheidae) between cold seep and hydrothermal vent habitats. Bulletin of Marine Science, 92 (1), 17 - 31. https: // doi. org / 10.5343 / bms. 2015.1031" type="journal article" year="2016">
Yang
<emphasis id="C0B6EAEF0D46FFB69FFAFD88FC2EBCC4" box="[940,998,620,645]" italics="true" pageId="4" pageNumber="396">et al.</emphasis>
2016
</bibRefCitation>
). On the other hand, the highest intraspecific sequence divergence noticed is 3.8% in
<taxonomicName id="35C24D7E0D46FFB69EA2FD75FCADBCE9" authorityName="Ramadan" authorityYear="1938" box="[756,869,655,680]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D46FFB69EA2FD75FCADBCE9" box="[756,869,655,680]" italics="true" pageId="4" pageNumber="396">A. alcocki</emphasis>
</taxonomicName>
amongst the Indian material. Thus, the following revised key is constructed for the Indo-West Pacific species of
<taxonomicName id="35C24D7E0D46FFB69F33FD51FC09BC8A" authorityName="Duvernoy" authorityYear="1840" box="[869,961,692,715]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EAEF0D46FFB69F33FD51FC09BC8A" box="[869,961,692,715]" italics="true" pageId="4" pageNumber="396">Aristeus</emphasis>
</taxonomicName>
, with the consideration of long preserved material might have the photophores difficult to recognize.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,137 @@
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<paragraph id="F27D36FD0D46FFB69CC1F8AAFD2CB928" blockId="4.[151,740,1871,1897]" box="[151,740,1871,1897]" pageId="4" pageNumber="396">
<heading id="A93581910D46FFB69CC1F8AAFD2CB928" bold="true" box="[151,740,1871,1897]" fontSize="11" level="1" pageId="4" pageNumber="396" reason="1">
<emphasis id="C0B6EAEF0D46FFB69CC1F8AAFD2CB928" bold="true" box="[151,740,1871,1897]" pageId="4" pageNumber="396">
Key to the Indo-West Pacific species of
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<emphasis id="C0B6EAEF0D46FFB69ED7F8AAFD2CB928" bold="true" box="[641,740,1871,1897]" italics="true" pageId="4" pageNumber="396">Aristeus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BAD865760D46FFB79CC1F870FA53BC87" lastPageId="5" lastPageNumber="397" pageId="4" pageNumber="396" type="key">
<keyStep id="49362E580D46FFB69CC1F870FA54B989" pageId="4" pageNumber="396">
<paragraph id="F27D36FD0D46FFB69CC1F870FA54B9EB" blockId="4.[151,1436,1941,2022]" box="[151,1436,1941,1963]" pageId="4" pageNumber="396">
<keyLead id="493395C80D46FFB69CC1F870FA54B9EB" box="[151,1436,1941,1963]" pageId="4" pageNumber="396">
1 Body integument distinctly pubescent; lateral surface of merus of pereiopod III with subdistal movable spine.......
<taxonomicName id="35C24D7E0D46FFB6991FF870FA54B9EB" ID-CoL="c58e3dde-229a-4d5e-b88e-e720abeeae88" baseAuthorityName="Bate" baseAuthorityYear="1881" box="[1353,1436,1941,1962]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="virilis">
<emphasis id="C0B6EAEF0D46FFB6991FF870FA54B9EB" box="[1353,1436,1941,1962]" italics="true" pageId="4" pageNumber="396">A. virilis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F27D36FD0D46FFB69CC1F856FA54B989" blockId="4.[151,1436,1941,2022]" box="[151,1436,1970,1992]" pageId="4" pageNumber="396">
<keyLead id="493395C80D46FFB69CC1F856FA54B989" box="[151,1436,1970,1992]" pageId="4" pageNumber="396">-. Body integument naked; lateral surface of merus of pereiopod III without subdistal spine.............................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="49362E580D46FFB79CC1F835FA54BEA6" lastPageId="5" lastPageNumber="397" pageId="4" pageNumber="396">
<paragraph id="F27D36FD0D46FFB69CC1F835FA54B9A7" blockId="4.[151,1436,1941,2022]" box="[151,1436,2000,2022]" pageId="4" pageNumber="396">
<keyLead id="493395C80D46FFB69CC1F835FA54B9A7" box="[151,1436,2000,2022]" pageId="4" pageNumber="396">2. Distance between lower end of cervical carina and branchiostegal carina nearly as long as distance between antennal and bran-</keyLead>
</paragraph>
<paragraph id="F27D36FD0D47FFB79CB7FF72FA54BEA6" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
chiostegal spines; pleurobranchs of thoractic somites IV to VII greatly reduced to minute papillae without pinnules.
<taxonomicName id="35C24D7E0D47FFB7996AFF72FA53BEED" ID-CoL="9257ec1a-e1f6-47ee-8f89-32bc5961595e" authorityName="Ramadan" authorityYear="1938" box="[1340,1435,151,172]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="C0B6EAEF0D47FFB7996AFF72FA53BEED" box="[1340,1435,151,172]" italics="true" pageId="5" pageNumber="397">A. alcocki</emphasis>
</taxonomicName>
<keyLead id="493395C80D47FFB79CC1FF50FA54BEA6" pageId="5" pageNumber="397">-. Distance between lower end of cervical carina and branchiostegal carina about half distance between antennal and branchiostegal spines; pleurobranchs of thoractic somites IV to VII small but distinct, feather-like and with distinct branches.........3</keyLead>
</paragraph>
</keyStep>
<keyStep id="49362E580D47FFB79CC1FF0AFA53BFDB" pageId="5" pageNumber="397">
<paragraph id="F27D36FD0D47FFB79CC1FF0AFA54BF7E" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FF0AFA54BF7E" pageId="5" pageNumber="397">3. Pereiopods I and II with chela more or less as long as carpus (&lt;1.3 times in pereiopod I and &lt;1.01 times in pereiopod II); pereiopod IV with carpus and propodus bearing less than 33 and 19 photophores, respectively; pereiopod V with carpus and dactylus bearing less than 34 and 17 photophores, respectively..................................................4</keyLead>
</paragraph>
<paragraph id="F27D36FD0D47FFB79CC1FEAEFA53BFDB" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FEAEFA53BFDB" pageId="5" pageNumber="397">
-. Pereiopods I and II with chela always longer than carpus (
<emphasis id="C0B6EAEF0D47FFB79F5DFEAFFCEBBF21" box="[779,803,330,352]" italics="true" pageId="5" pageNumber="397"></emphasis>
1.3 times in pereiopod I and
<emphasis id="C0B6EAEF0D47FFB79862FEAFFB84BF21" box="[1076,1100,330,352]" italics="true" pageId="5" pageNumber="397"></emphasis>
1.04 times in pereiopod II); pereiopod IV with carpus and propodus bearing more than 32 and 19 photophores, respectively; pereiopod V with carpus and dactylus bearing more than 35 and 21 photophores, respectively.....................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="49362E580D47FFB79CC1FE40FA53BC57" pageId="5" pageNumber="397">
<paragraph id="F27D36FD0D47FFB79CC1FE40FA53BF99" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FE40FA53BF99" pageId="5" pageNumber="397">
4. Pereiopod IV with carpus as long as or shorter than (
<emphasis id="C0B6EAEF0D47FFB79E89FE40FD3FBFFA" box="[735,759,421,443]" italics="true" pageId="5" pageNumber="397">Ĺ</emphasis>
1.00 times) merus; carpi of pereiopods I and IV with less than 10 photophores..................................................................................
<taxonomicName id="35C24D7E0D47FFB79959FE26FA53BF99" ID-CoL="3a80a356-dc28-4abd-938b-1b39691538fc" authorityName="Ramadan" authorityYear="1938" box="[1295,1435,451,472]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="mabahissae">
<emphasis id="C0B6EAEF0D47FFB79959FE26FA53BF99" box="[1295,1435,451,472]" italics="true" pageId="5" pageNumber="397">A. mabahissae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F27D36FD0D47FFB79CC1FE01FA53BC57" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FE01FA53BC57" pageId="5" pageNumber="397">
-. Pereiopod IV with carpus longer than (
<emphasis id="C0B6EAEF0D47FFB79E0BFE06FDBDBFB8" box="[605,629,483,505]" italics="true" pageId="5" pageNumber="397"></emphasis>
1.05 times) merus; carpi of pereiopods I and IV with more than 10 photophores...........................................................................................
<taxonomicName id="35C24D7E0D47FFB79955FDE4FA53BC57" ID-CoL="cc00a9b7-b959-48c2-b3de-6698dca654ef" authorityName="Bate" authorityYear="1881" box="[1283,1435,513,534]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="semidentatus">
<emphasis id="C0B6EAEF0D47FFB79955FDE4FA53BC57" box="[1283,1435,513,534]" italics="true" pageId="5" pageNumber="397">A. semidentatus</emphasis>
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<paragraph id="F27D36FD0D47FFB79CC1FDFBFA53BC2F" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FDFBFA53BC2F" pageId="5" pageNumber="397">
5. Pereiopod II with fingers 1.5 times longer than palm; pereiopod III with fingers 1.35 times longer than palm; carpi of pereiopods II and III bearing 3 or more photophores, pereiopods IV and V with more than 55 photophores each on carpi and propodi..........................................................................................
<taxonomicName id="35C24D7E0D47FFB79947FDBCFA53BC2F" ID-CoL="cc256e90-9046-4bd2-8dec-26da591c0f70" baseAuthorityName="Risso" baseAuthorityYear="1816" box="[1297,1435,601,622]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C0B6EAEF0D47FFB79947FDBCFA53BC2F" box="[1297,1435,601,622]" italics="true" pageId="5" pageNumber="397">A. antennatus</emphasis>
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<paragraph id="F27D36FD0D47FFB79CC1FD92FA53BC87" blockId="5.[151,1436,151,710]" pageId="5" pageNumber="397">
<keyLead id="493395C80D47FFB79CC1FD92FA53BC87" pageId="5" pageNumber="397">
-. Pereiopod II with fingers less than 1.5 times as long as palm; pereiopod III with fingers less than 1.30 times as long as palm; carpi of pereiopods II and III without photophore, pereiopods IV and V with less than 40 photophores each on carpi and propodi..................................................................................
<taxonomicName id="35C24D7E0D47FFB79953FD54FA53BC87" ID-CoL="13a36a79-a4ff-4d38-80e0-8c00cb1d5fcd" authorityName="Komai" authorityYear="1993" box="[1285,1435,689,710]" class="Malacostraca" family="Aristeidae" genus="Aristeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="397" phylum="Arthropoda" rank="species" species="pallidicauda">
<emphasis id="C0B6EAEF0D47FFB79953FD54FA53BC87" box="[1285,1435,689,710]" italics="true" pageId="5" pageNumber="397">A. pallidicauda</emphasis>
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