Add updates up until 2025-02-12 11:07:52

This commit is contained in:
ggserver 2025-02-12 11:09:57 +00:00
parent 4d0b6fcb33
commit d768b9078d
22 changed files with 4420 additions and 63 deletions

View file

@ -0,0 +1,321 @@
<document id="F0F51505F8A2CEE402B1560958EED8CC" ID-ISSN="0024-4082" ID-ZooBank="505571F-3ACC-4FB1-A0FB-05FDF18300D7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353253503" checkinUser="plazi" docAuthor="Delić, Teo, Kapla, Andrej &amp; Colla, Andrea" docDate="2020" docId="03C7373CFF8C0936FF08F91F39BBFE72" docLanguage="en" docName="zlz157.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Petraphaenops DELIC ET AL. 2020, gen. nov." docType="treatment" docVersion="1" lastPageNumber="1226" masterDocId="FFFE4F44FF89093FFFEBFFB73C20FFF9" masterDocTitle="Orogeny, sympatry and emergence of a new genus of Alpine subterranean Trechini (Coleoptera: Carabidae)" masterLastPageNumber="1231" masterPageNumber="1217" pageNumber="1222" updateTime="1739357576761" updateUser="GgImagineBatch">
<mods:mods id="1B9FFB9C660D297D4E73FD048F836D1B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48909CDAC13510DC1543BDB73C02DC03">
<mods:title id="F6BBF5C152E1279A00E4FF2867C8EE0B">Orogeny, sympatry and emergence of a new genus of Alpine subterranean Trechini (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="119307430E854ADB7EFAE0D1D8C94426" type="personal">
<mods:role id="93FC30228B9B77BB84A18EACF2C1D440">
<mods:roleTerm id="4BBA1D3732D1044C298C07C808196EAB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59CB2E686F5CD491334E38091138ED0A">Delić, Teo</mods:namePart>
</mods:name>
<mods:name id="CCCF204F1DC98CB2BC975DC5E54C49EF" type="personal">
<mods:role id="AB713C18B722A3D83B6A812A1BA5C3DD">
<mods:roleTerm id="F557F7C8398E324D1F099C5A605B3EB1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="284ADE6F93F4FD172B8AF6B782F8A48E">Kapla, Andrej</mods:namePart>
</mods:name>
<mods:name id="DF706D49AEEF023166EF0BDC63C8943A" type="personal">
<mods:role id="AE1AD51A0569F3CB7CD06DDCA5AB883A">
<mods:roleTerm id="E245AB890539E99F4001B94A72C57178">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93CAD4A56E950CB06FCCCCA1778CD43C">Colla, Andrea</mods:namePart>
</mods:name>
<mods:typeOfResource id="4D5DC127B4602336A1996A32FB41163E">text</mods:typeOfResource>
<mods:relatedItem id="BCAF043571246EECED2F58C8F429FDD1" type="host">
<mods:titleInfo id="13098DC0E5DCC044E47110E3BE5088F1">
<mods:title id="EAA813C471AA3CF5A08336BA09CF3967">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E5895EBBE6D2DD5A3796974F946CEDBA">
<mods:date id="D346F771DD6507DCA3EC7EC02A895EEC">2020</mods:date>
<mods:detail id="77BC59F502719F91996D43885A2710B9" type="volume">
<mods:number id="C216BFF24C1B09B305C84E54B108641A">189</mods:number>
</mods:detail>
<mods:extent id="A77F02CC83BD482EBDCD1D4512D5B864" unit="page">
<mods:start id="8927E8B82A78A9EFDAD77CA49AB81ABA">1217</mods:start>
<mods:end id="EC7122A3FCF925BF23B4AAFC7F0A9B57">1231</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="11C9D692730BBA12413AE3CD04BD9DFE">journal article</mods:classification>
<mods:identifier id="0D33A74D539BD17E0CC71CDDAEAA38C4" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="27EBBEC8BB64449A0BB33D4828987589" type="ZooBank">505571F-3ACC-4FB1-A0FB-05FDF18300D7</mods:identifier>
</mods:mods>
<treatment id="03C7373CFF8C0936FF08F91F39BBFE72" LSID="urn:lsid:plazi:treatment:03C7373CFF8C0936FF08F91F39BBFE72" httpUri="http://treatment.plazi.org/id/03C7373CFF8C0936FF08F91F39BBFE72" lastPageId="9" lastPageNumber="1226" pageId="5" pageNumber="1222">
<subSubSection id="C374D5A1FF8C093AFF08F91F3D87F8FD" pageId="5" pageNumber="1222" type="nomenclature">
<paragraph id="8BD1862AFF8C093AFF08F91F3EEAF946" blockId="5.[227,714,1703,1728]" box="[227,714,1703,1728]" pageId="5" pageNumber="1222">
<heading id="D0993146FF8C093AFF08F91F3EEAF946" box="[227,714,1703,1728]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="1222" reason="2">
<taxonomicName id="4C6EFDA9FF8C093AFF08F91F3E6AF939" ID-CoL="8LXLQ" authority="DELIC ET AL." authorityName="DELIC ET AL." authorityYear="2020" box="[227,586,1703,1728]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1222" phylum="Arthropoda" rank="genus" status="gen. nov.">
<smallCapsWord id="8D3710F6FF8C093AFF08F91F3D91F947" baselines="1721,1721" box="[227,433,1704,1727]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Petraphaenops" pageId="5" pageNumber="1222">
<emphasis id="B91A5A38FF8C093AFF08F91F3D91F947" bold="true" box="[227,433,1704,1727]" italics="true" pageId="5" pageNumber="1222">PETRAPHAENOPS</emphasis>
</smallCapsWord>
<smallCapsWord id="8D3710F6FF8C093AFE5CF9103DDDF947" baselines="1722,1722,1721" box="[439,509,1703,1728]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Delic" pageId="5" pageNumber="1222">DELIĆ</smallCapsWord>
<emphasis id="B91A5A38FF8C093AFDEEF91B3E64F946" box="[517,580,1707,1727]" italics="true" pageId="5" pageNumber="1222">
<smallCapsWord id="8D3710F6FF8C093AFDEEF91B3E02F947" baselines="1721" box="[517,546,1708,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="et" pageId="5" pageNumber="1222">ET</smallCapsWord>
<smallCapsWord id="8D3710F6FF8C093AFDC3F91C3E64F946" baselines="1722" box="[552,580,1707,1727]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="al" pageId="5" pageNumber="1222">AL</smallCapsWord>
</emphasis>
.
</taxonomicName>
,
<emphasis id="B91A5A38FF8C093AFDB3F9103EEAF946" bold="true" box="[600,714,1703,1727]" pageId="5" pageNumber="1222">
<taxonomicNameLabel id="A229E743FF8C093AFDB3F9103EEAF946" box="[600,714,1703,1727]" pageId="5" pageNumber="1222" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD1862AFF8C093AFF48F9673D87F8FD" blockId="5.[163,778,1744,1796]" pageId="5" pageNumber="1222">
<uri id="FFFF8A28FF8C093AFF48F9673D87F8FD" pageId="5" pageNumber="1222">
urn:lsid:zoobank.org:act:
<uuid id="FFC8BCFFFF8C093AFE3DF9673D87F8FD" pageId="5" pageNumber="1222">AC14ED47-6530-4921- 93C4-4CDEA055F5EE</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C374D5A1FF8C0936FF48F89139BBFE72" lastPageId="9" lastPageNumber="1226" pageId="5" pageNumber="1222" type="materials_examined">
<paragraph id="8BD1862AFF8C093AFF48F8913D80F8A3" blockId="5.[163,778,1829,1882]" pageId="5" pageNumber="1222">
<emphasis id="B91A5A38FF8C093AFF48F8913EFAF8C2" box="[163,730,1829,1851]" italics="true" pageId="5" pageNumber="1222">
<typeStatus id="54D53888FF8C093AFF48F8913CFBF8C2" box="[163,219,1830,1851]" pageId="5" pageNumber="1222">Type</typeStatus>
species:
<taxonomicName id="4C6EFDA9FF8C093AFEA4F8923EFAF8C2" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" box="[335,730,1829,1851]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1222" phylum="Arthropoda" rank="species" species="unguiculatus" status="sp. nov.">
<emphasis id="B91A5A38FF8C093AFEA4F8923EFAF8C2" bold="true" box="[335,730,1829,1851]" pageId="5" pageNumber="1222">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A229E743FF8C093AFD0EF8923CF6F8A0" pageId="5" pageNumber="1222" rank="species">sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
<paragraph id="8BD1862AFF8C093AFCD0FA04382BF99A" blockId="5.[827,1443,1459,1635]" pageId="5" pageNumber="1222">
<emphasis id="B91A5A38FF8C093AFCD0FA043F99FA31" box="[827,953,1459,1480]" italics="true" pageId="5" pageNumber="1222">Etymology:</emphasis>
The generic name is composed of the Greek πέτρα,
<emphasis id="B91A5A38FF8C093AFC63FA643FE3FA11" box="[904,963,1491,1512]" italics="true" pageId="5" pageNumber="1222">petra</emphasis>
rock, and
<taxonomicName id="4C6EFDA9FF8C093AFBD1FA653896FA1E" authorityName="Bonvouloir" authorityYear="1862" box="[1082,1206,1490,1511]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1222" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8C093AFBD1FA653896FA1E" box="[1082,1206,1490,1511]" italics="true" pageId="5" pageNumber="1222">Aphaenops</emphasis>
</taxonomicName>
, name of the famous Pyrenean trechine genus, which represents a classical example of troglomorphic subterranean
<taxonomicName id="4C6EFDA9FF8C093AFACCF9B83982F9DC" authorityName="LATREILLE" authorityYear="1802" box="[1319,1442,1551,1573]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1222" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
and defines the so called
<typeStatus id="54D53888FF8C093AFB89F99938B5F9BA" box="[1122,1173,1582,1603]" pageId="5" pageNumber="1222">type</typeStatus>
aphénopsien (Jeannel, 1926) morphology.
</paragraph>
<paragraph id="8BD1862AFF8C0939FCD0F93B3D6CF9BA" blockId="5.[827,1443,1675,1881]" lastBlockId="6.[145,761,1244,1603]" lastPageId="6" lastPageNumber="1223" pageId="5" pageNumber="1222">
<emphasis id="B91A5A38FF8C093AFCD0F93B3F95F958" box="[827,949,1676,1697]" italics="true" pageId="5" pageNumber="1222">Diagnosis:</emphasis>
Anophthalmous genus of trechine ground beetles, medium-sized (
<typeStatus id="54D53888FF8C093AFBB2F91D38AEF946" box="[1113,1166,1706,1727]" pageId="5" pageNumber="1222">type</typeStatus>
length
<quantity id="4C962BCFFF8C093AFB05F91D397AF946" box="[1262,1370,1706,1728]" metricMagnitude="-3" metricUnit="m" metricValue="6.11" pageId="5" pageNumber="1222" unit="mm" value="6.11">6.11 mm</quantity>
), pale brown, shiny, characterized by an aphaenopsian habitus, showing a high degree of troglomorphism. The head as wide as the pronotum, both proportionally thinner than in the closely related
<taxonomicName id="4C6EFDA9FF8C093AFB51F8923945F8C3" authorityName="Muller" authorityYear="1913" box="[1210,1381,1829,1850]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1222" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8C093AFB51F8923945F8C3" box="[1210,1381,1829,1850]" italics="true" pageId="5" pageNumber="1222">Aphaenopidius</emphasis>
</taxonomicName>
, with six supraorbital setae. Pronotum hairless, without setae in the anterior marginal position and in the posterior angles. Elytra elongated, without humeral shoulders, with all the umbilicate setigerous pores aligned in an arch along the marginal gutter, and two discal setae. Tarsi with disproportionally long claws, about twice or more as long as in
<taxonomicName id="4C6EFDA9FF8F0939FE80FAC23E35FA73" authorityName="Sturm" authorityYear="1844" box="[363,533,1397,1418]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1223" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8F0939FE80FAC23E35FA73" box="[363,533,1397,1418]" italics="true" pageId="6" pageNumber="1223">Anophthalmus</emphasis>
</taxonomicName>
or other average to large-bodied trechines. Aedeagus elongated, similar to that of
<taxonomicName id="4C6EFDA9FF8F0939FF0FFA053DB0FA3E" authorityName="Muller" authorityYear="1913" box="[228,400,1458,1479]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1223" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8F0939FF0FFA053DB0FA3E" box="[228,400,1458,1479]" italics="true" pageId="6" pageNumber="1223">Aphaenopidius</emphasis>
</taxonomicName>
, but thinner and less curved in the median lobe, with its axis forming almost a right angle with that of the basal lobe. Isotopic copulatory piece, anteriorly bilobate (
<figureCitation id="13559AAFFF8F0939FE54F9B93E3CF9DA" box="[447,540,1550,1572]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStart-3="Figure 8" captionStart-4="Figure 9" captionStartId-0="7.[163,241,1054,1076]" captionStartId-1="7.[163,241,1563,1585]" captionStartId-2="8.[146,228,1361,1383]" captionStartId-3="8.[145,225,1844,1866]" captionStartId-4="8.[809,889,953,975]" captionTargetBox-0="[165,1441,198,1013]" captionTargetBox-1="[168,1441,1208,1520]" captionTargetBox-2="[150,754,199,1320]" captionTargetBox-3="[151,755,1453,1804]" captionTargetBox-4="[816,1411,198,911]" captionTargetId-0="figure-150@7.[163,1443,195,1014]" captionTargetId-1="figure-226@7.[163,1443,1206,1523]" captionTargetId-2="figure-231@8.[149,757,195,1321]" captionTargetId-3="figure-241@8.[149,757,1451,1805]" captionTargetId-4="figure-254@8.[813,1421,195,913]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="8" captionText-0="Figure 5. Morphological comparison of the four closely related and syntopically occurring species from Devettisoča jama: A, Petraphaenops unguiculatus; B, Aphaenopidius sp.; C, Anophthalmus erebus; and D, Anophthalmus pretneri. Note the differences in the general body plan, size of the head, shape of the mandibles, relative length of the legs and size and shape of claws. Differences in chaetotaxy are presented on the left part of the body as white dots." captionText-1="Figure 6. Comparison of the aedeagi and endophallus copulatory piece of the four closely related and syntopically occurring species from Devettisoča jama: A, Petraphaenops unguiculatus; B, Aphaenopidius sp.; C, Anophthalmus erebus; and D, Anophthalmus pretneri. All aedeagi are presented in the same scale." captionText-2="Figure 7. Petraphaenops unguiculatus, holotype male, habitus." captionText-3="Figure 8. Aedeagus of a male holotype of Petraphaenops unguiculatus." captionText-4="Figure 9. Comparison of shape and length of tarsomeres and claws in the three closely related and syntopically occurring trechines: A, Petraphaenops unguiculatus; B, Aphaenopidius spp.; and C, Anophthalmus erebus. All photos in the same scale." pageId="6" pageNumber="1223">Figs 5-9</figureCitation>
). Known only from the
<typeStatus id="54D53888FF8F0939FF56F99A3CCEF9BB" box="[189,238,1581,1602]" pageId="6" pageNumber="1223">type</typeStatus>
species.
</paragraph>
<caption id="DF11D6A2FF8F0939FF79FBB13CDDFB56" pageId="6" pageNumber="1223" startId="6.[146,224,1030,1052]" targetBox="[146,1409,197,987]" targetPageId="6" targetType="figure">
<paragraph id="8BD1862AFF8F0939FF79FBB13CDDFB56" blockId="6.[145,1425,1030,1199]" pageId="6" pageNumber="1223">
<emphasis id="B91A5A38FF8F0939FF79FBB13CD9FBE5" bold="true" box="[146,249,1030,1052]" pageId="6" pageNumber="1223">Figure 4.</emphasis>
Time-calibrated phylogeny of the selected European subterranean
<taxonomicName id="4C6EFDA9FF8F0939FC51FBB03835FBE4" box="[954,1045,1031,1053]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1223" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
. Calibrated time-tree corresponds to the Bayesian relaxed-clock analysis based on the mean clock-rate values published in
<bibRefCitation id="EFFFFBDBFF8F0939FBECFB9338E0FBC3" author="Faille A &amp; Casale A &amp; Balke M &amp; Ribera I" box="[1031,1216,1060,1082]" pageId="6" pageNumber="1223" refId="ref8817" refString="Faille A, Casale A, Balke M, Ribera I. 2013. A molecular phylogeny of Alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology 13: 248." type="journal volume" year="2013">
Faille
<emphasis id="B91A5A38FF8F0939FBACFB92385AFBC3" box="[1095,1146,1060,1082]" italics="true" pageId="6" pageNumber="1223">et al.</emphasis>
(2013)
</bibRefCitation>
and 95% confidence intervals corresponding to values from
<bibRefCitation id="EFFFFBDBFF8F0939FDDAFBF63F2CFBAE" author="Andujar C &amp; Serrano J &amp; Gomez-Zurita J" box="[561,780,1089,1111]" pageId="6" pageNumber="1223" refId="ref7990" refString="Andujar C, Serrano J, Gomez-Zurita J. 2012. Winding up the molecular clock in the genus Carabus (Coleoptera: Carabidae): assessment of methodological decisions on rate and node age estimation. BMC Evolutionary Biology 12: 40." type="journal volume" year="2012">
Andújar
<emphasis id="B91A5A38FF8F0939FD64FBF53E9DFBAE" box="[655,701,1089,1111]" italics="true" pageId="6" pageNumber="1223">et al</emphasis>
. (2012)
</bibRefCitation>
. Onset of the Raduha orogenesis (southern Calcareous Alps) is noted with a red line and corresponds to the estimated age of split between
<taxonomicName id="4C6EFDA9FF8F0939FC31FBE83856FB8C" box="[986,1142,1119,1141]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1223" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8F0939FC31FBE83856FB8C" box="[986,1142,1119,1141]" italics="true" pageId="6" pageNumber="1223">Petraphaenops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6EFDA9FF8F0939FB46FBE8396DFB8C" authorityName="Muller" authorityYear="1913" box="[1197,1357,1119,1141]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1223" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF8F0939FB46FBE8396DFB8C" box="[1197,1357,1119,1141]" italics="true" pageId="6" pageNumber="1223">Aphaenopidius</emphasis>
</taxonomicName>
. Node supports are denoted as white&gt; 0.95, grey&gt; 0.99 and black = 1 circles and the 95 % confidence intervals are shown as blue node bars.
</paragraph>
</caption>
<paragraph id="8BD1862AFF8F0936FF7AF9DB3D33FB68" blockId="6.[145,760,1643,1880]" lastBlockId="9.[163,779,197,1170]" lastPageId="9" lastPageNumber="1226" pageId="6" pageNumber="1223">
<emphasis id="B91A5A38FF8F0939FF7AF9DB3D04F978" box="[145,292,1644,1665]" italics="true" pageId="6" pageNumber="1223">Description:</emphasis>
Medium-sized genus of subterranean trechines, blind, with a high level of troglomorphism; body elongated, glabrous, with head as wide as the prothorax, the latter short, elytra without humeral angles; legs extremely long, especially the femurs, tarsi with claws indicating specialization to a specific ecological niche within the cave environment presumably cave hygropetric; posterior wings absent; depigmented, colour pale brown, almost amber, shiny. Head as wide as the pronotum, elongated but robust, about twice as long as wide with closed mandibles included, with the maximum width at level of a section between the anterior and intermediate supraorbital setae, progressively narrowing to its posterior part and neck, without prominent temples (the cuticle between the ocular region and neck); neck robust and wide but elongated, with a large articular surface between the neck constriction and the pronotal anterior margin (with the head in the same axis as the body); frontal furrows well marked, but incomplete, parallel, suddenly stopping at the level of the intermediate supraorbital setae; head surface glabrous, with six supraorbital setae (three on each side), the front pair at the level of half the length of the frontal furrows, the middle pair equidistant from the clypeofrontal suture and the neck constriction; the hind pair of setae at level of the temples; eyes totally absent; vertex frontally convex; clypeus with the anterior margin weakly convex, with a glabrous central area between the setae, that are two or three on each side (character asymmetrical in the
<typeStatus id="54D53888FF8E0938FF48F96E3D2EF916" box="[163,270,1753,1775]" pageId="7" pageNumber="1224" type="holotype">holotype</typeStatus>
); labrum concave in the middle, bilobate, normally sexsetose; mandibles long and robust, gently curved in the middle, then arcuate at the acute apex, bidentate in the molar region; maxillae almost as long as the mandibles; maxillary palps long, thin, the penultimate and ultimate articles each as long as half of the third antennomer, the ultimate in the shape of a narrow bowling pin; antennae reaching about 84% of the total body length, with the scapus more than twice as long as wide, the second article almost four, the third, fourth and the fifth article about ten, the sixth and seventh article more than eight, the eighth and ninth more than six, and the tenth and 11
<superScript id="7C1B2B62FF810937FAD8FBD83961FB82" attach="left" box="[1331,1345,1135,1147]" fontSize="5" pageId="8" pageNumber="1225">th</superScript>
article almost six times longer than wide. Prothorax small, thinner than a single elytron, less than a third of the maximum body width at elytral level, about as long as wide, round-ovate, with shallow and narrow marginal gutters; regularly arched lateral margins, barely sinuated in correspondence to the small anterior and posterior lobes, the later with rounded hind angles; front angles more acute anteriorly, as a consequence of the concavity of the pronotal front margin, resembling a widely open V; hind margin shorter than the front one, only slightly concave; transverse sulcus present; disc convex, shiny, glabrous, without any visible setae or setigerous pores, apart from three asymmetrically inserted setae near the anterior margin. Elytra elongated, slightly longer than half of the total body length, pair of elytra approximately 1.8 times as long as wide, maximum width slightly behind middle, at the level of the third discal setigerous pore; surface flat in the first third, moderately convex in the central and apical part, with shallow but still recognizable inner striae, external ones almost vanishing; oblique prehumeral margins, almost straight or only slightly concave, gently connected with the arched posthumeral margin, without humeral angles; elytral chaetotaxy with all the pores of the umbilicate marginal series close to the marginal gutter; first four pores in the humeral region, more or less equidistant from each other; the fifth pore at the level of the second discal or slightly anterior (the
<typeStatus id="54D53888FF800936FE4AFEF73E24FEAF" box="[417,516,320,342]" pageId="9" pageNumber="1226" type="holotype">holotype</typeStatus>
is asymmetrical), in a slightly inner position; the distance between the fifth and the sixth pore the same as the distance between the third and fourth; the seventh pore distally three times that distance; the basal setigerous pore present; two discal setigerous pores on the third stria, the second about at the level of the maximum elytral width; the front pore of the apical triangle distant backwards; a shallow apical carina; elytral suture posteriorly divergent, with a small pointed apical protrusion on each elytron. Legs long, especially the femora and tarsi; the anterior tibiae not sulcate; long pubescence on the external distal third of the femora and on the tarsi, shorter and less dense on the tibiae; male protarsi with the first two tarsomeres dilated; tarsal claws developed, long and slender, slightly arcuate. Aedeagus well sclerotized, elongated, similar to that of
<taxonomicName id="4C6EFDA9FF800936FEF1FCFE3DEBFCA7" authorityName="Muller" authorityYear="1913" box="[282,459,841,862]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FEF1FCFE3DEBFCA7" box="[282,459,841,862]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
</taxonomicName>
, but thinner, tubular, with the median lobe almost straight, its axis forming an almost right angle with that of the basal bulb, apex short and folded upwards; styles with four stout setae in the apex; inner sac with thorny scales, armed with a small isotopic copulatory piece, sclerotized, bilobate, comprising two branches basally fused and frontally divergent, giving the copulatory piece the shape of a narrow V. The branches are kept parallel and adherent to each other in retracted position inside the inner sac.
</paragraph>
<caption id="DF11D6A2FF8E0938FF48FBA93874FB75" pageId="7" pageNumber="1224" startId="7.[163,241,1054,1076]" targetBox="[165,1441,198,1013]" targetPageId="7" targetType="figure">
<paragraph id="8BD1862AFF8E0938FF48FBA93874FB75" blockId="7.[163,1444,1054,1164]" pageId="7" pageNumber="1224">
<emphasis id="B91A5A38FF8E0938FF48FBA93D2BFBCA" bold="true" box="[163,267,1054,1076]" pageId="7" pageNumber="1224">Figure 5.</emphasis>
Morphological comparison of the four closely related and syntopically occurring species from Devettisoča jama: A,
<taxonomicName id="4C6EFDA9FF8E0938FF2BFB8C3DCEFBA8" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" box="[192,494,1083,1105]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B91A5A38FF8E0938FF2BFB8C3DCEFBA8" box="[192,494,1083,1105]" italics="true" pageId="7" pageNumber="1224">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C6EFDA9FF8E0938FDFDFB8C3EFCFBA8" box="[534,732,1083,1105]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A5A38FF8E0938FDFDFB8C3E95FBA8" box="[534,693,1083,1105]" italics="true" pageId="7" pageNumber="1224">Aphaenopidius</emphasis>
sp.
</taxonomicName>
; C,
<taxonomicName id="4C6EFDA9FF8E0938FCE8FB8C3FCAFBA8" baseAuthorityName="Krauss" baseAuthorityYear="1906" box="[771,1002,1083,1105]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B91A5A38FF8E0938FCE8FB8C3FCAFBA8" box="[771,1002,1083,1105]" italics="true" pageId="7" pageNumber="1224">Anophthalmus erebus</emphasis>
</taxonomicName>
; and D,
<emphasis id="B91A5A38FF8E0938FBA8FB8C3962FBA8" box="[1091,1346,1083,1105]" italics="true" pageId="7" pageNumber="1224">
<taxonomicName id="4C6EFDA9FF8E0938FBA8FB8C391DFBA8" baseAuthorityName="G. Muller" baseAuthorityYear="1913" box="[1091,1341,1083,1105]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="pretneri">Anophthalmus pretneri</taxonomicName>
.
</emphasis>
Note the differences in the general body plan, size of the head, shape of the mandibles, relative length of the legs and size and shape of claws. Differences in chaetotaxy are presented on the left part of the body as white dots.
</paragraph>
</caption>
<caption id="DF11D6A2FF8E0938FF48F9AC3F4EF995" pageId="7" pageNumber="1224" startId="7.[163,241,1563,1585]" targetBox="[168,1441,1208,1520]" targetPageId="7" targetType="figure">
<paragraph id="8BD1862AFF8E0938FF48F9AC3F4EF995" blockId="7.[163,1443,1563,1644]" pageId="7" pageNumber="1224">
<emphasis id="B91A5A38FF8E0938FF48F9AC3D29F9C9" bold="true" box="[163,265,1563,1585]" pageId="7" pageNumber="1224">Figure 6.</emphasis>
Comparison of the aedeagi and endophallus copulatory piece of the four closely related and syntopically occurring species from Devettisoča jama: A,
<taxonomicName id="4C6EFDA9FF8E0938FDF7F98E3F6DF9B6" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" box="[540,845,1593,1615]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B91A5A38FF8E0938FDF7F98E3F6DF9B6" box="[540,845,1593,1615]" italics="true" pageId="7" pageNumber="1224">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C6EFDA9FF8E0938FC9DF98E381EF9B6" box="[886,1086,1593,1615]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A5A38FF8E0938FC9DF98E3836F9B6" box="[886,1046,1593,1615]" italics="true" pageId="7" pageNumber="1224">Aphaenopidius</emphasis>
sp.
</taxonomicName>
; C,
<taxonomicName id="4C6EFDA9FF8E0938FB8DF98E396FF9B6" baseAuthorityName="Krauss" baseAuthorityYear="1906" box="[1126,1359,1593,1615]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B91A5A38FF8E0938FB8DF98E396FF9B6" box="[1126,1359,1593,1615]" italics="true" pageId="7" pageNumber="1224">Anophthalmus erebus</emphasis>
</taxonomicName>
; and D,
<emphasis id="B91A5A38FF8E0938FF48F9E13DBDF995" box="[163,413,1622,1644]" italics="true" pageId="7" pageNumber="1224">
<taxonomicName id="4C6EFDA9FF8E0938FF48F9E13DB9F995" baseAuthorityName="G. Muller" baseAuthorityYear="1913" box="[163,409,1622,1644]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1224" phylum="Arthropoda" rank="species" species="pretneri">Anophthalmus pretneri</taxonomicName>
.
</emphasis>
All aedeagi are presented in the same scale.
</paragraph>
</caption>
<caption id="DF11D6A2FF810937FCC2FC0E380FFBBD" pageId="8" pageNumber="1225" startId="8.[809,889,953,975]" targetBox="[816,1411,198,911]" targetPageId="8" targetType="figure">
<paragraph id="8BD1862AFF810937FCC2FC0E380FFBBD" blockId="8.[809,1425,953,1092]" pageId="8" pageNumber="1225">
<emphasis id="B91A5A38FF810937FCC2FC0E3FB3FC37" bold="true" box="[809,915,953,975]" pageId="8" pageNumber="1225">Figure 9.</emphasis>
Comparison of shape and length of tarsomeres and claws in the three closely related and syntopically occurring trechines: A,
<taxonomicName id="4C6EFDA9FF810937FBC4FC43394CFBF3" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" box="[1071,1388,1012,1034]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1225" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B91A5A38FF810937FBC4FC43394CFBF3" box="[1071,1388,1012,1034]" italics="true" pageId="8" pageNumber="1225">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C6EFDA9FF810937FCC2FBA6382AFBDE" box="[809,1034,1041,1063]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1225" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A5A38FF810937FCC2FBA63FF1FBDE" box="[809,977,1041,1063]" italics="true" pageId="8" pageNumber="1225">Aphaenopidius</emphasis>
spp.
</taxonomicName>
; and C,
<taxonomicName id="4C6EFDA9FF810937FB80FBA63941FBDE" baseAuthorityName="Krauss" baseAuthorityYear="1906" box="[1131,1377,1041,1063]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1225" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B91A5A38FF810937FB80FBA63941FBDE" box="[1131,1377,1041,1063]" italics="true" pageId="8" pageNumber="1225">Anophthalmus erebus</emphasis>
</taxonomicName>
. All photos in the same scale.
</paragraph>
</caption>
<caption id="DF11D6A2FF810937FF79FAE63CC7FA7C" pageId="8" pageNumber="1225" startId="8.[146,228,1361,1383]" targetBox="[150,754,199,1320]" targetPageId="8" targetType="figure">
<paragraph id="8BD1862AFF810937FF79FAE63CC7FA7C" blockId="8.[145,760,1361,1413]" pageId="8" pageNumber="1225">
<emphasis id="B91A5A38FF810937FF79FAE63D21FA9E" bold="true" box="[146,257,1361,1383]" pageId="8" pageNumber="1225">Figure 7.</emphasis>
<taxonomicName id="4C6EFDA9FF810937FEE7FAE63E68FA9E" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" box="[268,584,1361,1383]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1225" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B91A5A38FF810937FEE7FAE63E68FA9E" box="[268,584,1361,1383]" italics="true" pageId="8" pageNumber="1225">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
, holotype male, habitus.
</paragraph>
</caption>
<caption id="DF11D6A2FF810937FF7AF8833D00F891" pageId="8" pageNumber="1225" startId="8.[145,225,1844,1866]" targetBox="[151,755,1453,1804]" targetPageId="8" targetType="figure">
<paragraph id="8BD1862AFF810937FF7AF8833D00F891" blockId="8.[145,760,1844,1896]" pageId="8" pageNumber="1225">
<emphasis id="B91A5A38FF810937FF7AF8833CDBF8B0" bold="true" box="[145,251,1844,1866]" pageId="8" pageNumber="1225">Figure 8.</emphasis>
Aedeagus of a male holotype of
<taxonomicName id="4C6EFDA9FF810937FDB7F8833D3BF891" authorityName="Delić &amp; Kapla &amp; Colla" authorityYear="2020" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1225" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B91A5A38FF810937FDB7F8833D3BF891" italics="true" pageId="8" pageNumber="1225">Petraphaenops unguiculatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD1862AFF800936FF48FB0D3829FF00" blockId="9.[163,779,1210,1907]" lastBlockId="9.[827,1443,197,311]" pageId="9" pageNumber="1226">
<emphasis id="B91A5A38FF800936FF48FB0D3D38FB36" box="[163,280,1210,1231]" italics="true" pageId="9" pageNumber="1226">Remarks:</emphasis>
From the morphological point of view, characters like the three pairs of supraorbital setae, the absence of humeral angles, the genitalia, etc., suggest that
<taxonomicName id="4C6EFDA9FF800936FED2FAA13DC1FAD2" box="[313,481,1302,1323]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FED2FAA13DC1FAD2" box="[313,481,1302,1323]" italics="true" pageId="9" pageNumber="1226">Petraphaenops</emphasis>
</taxonomicName>
is a new genus related to the syntopic
<taxonomicName id="4C6EFDA9FF800936FED3FA823EA0FAB3" authority="Muller, 1913" authorityName="Muller" authorityYear="1913" box="[312,640,1333,1355]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FED3FA823DC6FAB3" box="[312,486,1333,1354]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
Müller, 1913
</taxonomicName>
, a narrowly distributed, large aphaenopsian genus of isotopic trechine. However, the presently known
<taxonomicName id="4C6EFDA9FF800936FD8BFAC53F2BFA7E" authorityName="Muller" authorityYear="1913" box="[608,779,1394,1415]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FD8BFAC53F2BFA7E" box="[608,779,1394,1415]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
</taxonomicName>
taxa are incomparably larger in size, macrocephalic, with a transverse pronotum clearly wider than it is long and heart-shaped because of the distinct and sinuate lateral margins, with four pronotal setae normally positioned. According to these differential characters and the original description by Müller (1913), the new taxon cannot be included in that genus.
<taxonomicName id="4C6EFDA9FF800936FD8FF9FE3F2BF9A7" box="[612,779,1609,1630]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FD8FF9FE3F2BF9A7" box="[612,779,1609,1630]" italics="true" pageId="9" pageNumber="1226">Petraphaenops</emphasis>
</taxonomicName>
is easily distinguishable from
<taxonomicName id="4C6EFDA9FF800936FDE4F9DF3EE1F984" authorityName="Muller" authorityYear="1913" box="[527,705,1640,1661]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FDE4F9DF3EE1F984" box="[527,705,1640,1661]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
</taxonomicName>
being about half its size, with a thinner head and pronotum, the last totally different in shape and lacking any pronotal setae. This complete lack of setae is not only a differential character from
<taxonomicName id="4C6EFDA9FF800936FE1EF9553E82F90E" authorityName="Muller" authorityYear="1913" box="[501,674,1762,1783]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus">
<emphasis id="B91A5A38FF800936FE1EF9553E82F90E" box="[501,674,1762,1783]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
</taxonomicName>
, but it is generally unusual among southern calcareous Alpine and Dinaric
<taxonomicName id="4C6EFDA9FF800936FEDFF8973DB9F8CF" box="[308,409,1824,1846]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
. Moreover, the general shape of the aedeagus resembles also that of some
<emphasis id="B91A5A38FF800936FD74F8893CE2F88B" italics="true" pageId="9" pageNumber="1226">
<taxonomicName id="4C6EFDA9FF800936FD74F8893F2AF8AA" authorityName="Delarouzee" authorityYear="1859" box="[671,778,1854,1875]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus" sensu="lato">Duvalius</taxonomicName>
<taxonomicNameLabel id="A229E743FF800936FF48F8EA3CE2F88B" box="[163,194,1885,1906]" pageId="9" pageNumber="1226" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C6EFDA9FF800936FF11F8EA3DDFF88A" box="[250,511,1885,1907]" class="Insecta" family="Carabidae" genus="Anophthalmus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A5A38FF800936FF11F8EA3D82F88B" box="[250,418,1885,1906]" italics="true" pageId="9" pageNumber="1226">Anophthalmus</emphasis>
species.
</taxonomicName>
The isotopic copulatory piece also resembles that of
<taxonomicName id="4C6EFDA9FF800936FB93FF723977FF23" box="[1144,1367,197,218]" class="Insecta" family="Carabidae" genus="Aphaenopidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91A5A38FF800936FB93FF723903FF23" box="[1144,1315,197,218]" italics="true" pageId="9" pageNumber="1226">Aphaenopidius</emphasis>
spp.
</taxonomicName>
and of some
<emphasis id="B91A5A38FF800936FC97FF533829FF00" box="[892,1033,228,249]" italics="true" pageId="9" pageNumber="1226">
<taxonomicName id="4C6EFDA9FF800936FC97FF533FC4FF00" authorityName="Delarouzee" authorityYear="1859" box="[892,996,228,249]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="genus" sensu="lato">Duvalius</taxonomicName>
<taxonomicNameLabel id="A229E743FF800936FC01FF533829FF00" box="[1002,1033,228,249]" pageId="9" pageNumber="1226" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD1862AFF800936FCB8FEB53FEDFECF" blockId="9.[827,1443,197,311]" pageId="9" pageNumber="1226">These morphological considerations agree with the genetic ones.</paragraph>
<paragraph id="8BD1862AFF800936FCA8FEC339BBFE72" blockId="9.[835,1435,371,427]" box="[835,1435,371,396]" pageId="9" pageNumber="1226">
<taxonomicName id="4C6EFDA9FF800936FCA8FEC339BBFE72" authority="DELIC ET AL." authorityName="DELIC ET AL." authorityYear="2020" box="[835,1435,371,396]" class="Insecta" family="Carabidae" genus="Petraphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1226" phylum="Arthropoda" rank="species" species="unguiculatus" status="SP.">
<emphasis id="B91A5A38FF800936FCA8FEC338EFFE73" bold="true" box="[835,1231,372,395]" italics="true" pageId="9" pageNumber="1226">
<smallCapsWord id="8D3710F6FF800936FCA8FEC33831FE73" baselines="389,389" box="[835,1041,372,395]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Petraphaenops" pageId="9" pageNumber="1226">PETRAPHAENOPS</smallCapsWord>
<smallCapsWord id="8D3710F6FF800936FBFCFEC038EFFE73" baselines="389" box="[1047,1231,375,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="unguiculatus" pageId="9" pageNumber="1226">UNGUICULATUS</smallCapsWord>
</emphasis>
<smallCapsWord id="8D3710F6FF800936FB3DFEC4393CFE73" baselines="390,390,389" box="[1238,1308,371,396]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Delic" pageId="9" pageNumber="1226">DELIĆ</smallCapsWord>
<emphasis id="B91A5A38FF800936FAC8FECF3943FE72" box="[1315,1379,375,395]" italics="true" pageId="9" pageNumber="1226">
<smallCapsWord id="8D3710F6FF800936FAC8FECF3960FE73" baselines="389" box="[1315,1344,376,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="et" pageId="9" pageNumber="1226">ET</smallCapsWord>
<smallCapsWord id="8D3710F6FF800936FAACFEC03943FE72" baselines="390" box="[1351,1379,375,395]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="al" pageId="9" pageNumber="1226">AL</smallCapsWord>
</emphasis>
.,
<emphasis id="B91A5A38FF800936FA9CFEC039BBFE72" bold="true" box="[1399,1435,371,395]" pageId="9" pageNumber="1226">
<smallCapsWord id="8D3710F6FF800936FA9CFEC039B4FE73" baselines="389" box="[1399,1428,375,394]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="9" pageNumber="1226">SP</smallCapsWord>
.
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,282 @@
<document id="5D2E94A33D11494B135895DDBE51AF2D" ID-ISSN="0024-4082" ID-ZooBank="4D7E449-E29E-4E8F-A215-5E8FA38BC763" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353372604" checkinUser="plazi" docAuthor="Gustafson, Grey T., Michat, Mariano C. &amp; Balke, Michael" docDate="2020" docId="03F787C01E550B28FF0B7F3EFB59FA16" docLanguage="en" docName="zlz161.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Chimerogyrus gigagalea Gustafson, Michat &amp; Balke, 2020, SP. NOV." docType="treatment" docVersion="1" lastPageNumber="1243" masterDocId="FFCEFFB81E5F0B23FFF77D2BFFD5FFB7" masterDocTitle="Burmese amber reveals a new stem lineage of whirligig beetle (Coleoptera: Gyrinidae) based on the larval stage" masterLastPageNumber="1248" masterPageNumber="1232" pageNumber="1242" updateTime="1739358064101" updateUser="GgImagineBatch">
<mods:mods id="331D1CD6399837E2FCFD7FF0CF2A5728" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="22E0E6243BED2FEFA775F04167D05E17">
<mods:title id="EAFAD0CA23362503ADB0750DEDFA3AA1">Burmese amber reveals a new stem lineage of whirligig beetle (Coleoptera: Gyrinidae) based on the larval stage</mods:title>
</mods:titleInfo>
<mods:name id="9741D5D23FDA94F4D8689B495BF34E80" type="personal">
<mods:role id="00BF643CB00D9C2F18CE46B562B0E9EA">
<mods:roleTerm id="5FD47E6158FE0211CFF7CCFC54F7EBFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="529151CE16B4364F9BD8B049BD2E3852">Gustafson, Grey T.</mods:namePart>
</mods:name>
<mods:name id="309D4A1625FE88C38DA23FEC3CEE4E54" type="personal">
<mods:role id="A5376D641D08FD3480AFD7F87A78AAC7">
<mods:roleTerm id="4F0D27CD007CFCE0446DF2F56D9E40B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9504889F2437CFEA6EF3C30D52C042DE">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:name id="F62C5B318FF738ECF91A925AE7F5AAE6" type="personal">
<mods:role id="DA77730091A2C551385DBC245D7E6866">
<mods:roleTerm id="50F2F907FFED653665D6AB509012E55F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="379186AD7192A5F8A507EC41B8AB9A81">Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="FFD560AD565E7AB368600AE4BAE0C1A5">text</mods:typeOfResource>
<mods:relatedItem id="A2DA32DD70BED64324A5A7C79FFCD137" type="host">
<mods:titleInfo id="5E8D43B3A79E7AA77F9EE9489A474F6F">
<mods:title id="852234E689AE49C889BD305D2146B89C">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="FF080C9B247C14031DA8C28254ECA7ED">
<mods:date id="1EEE70EC02E358FC33C8ECAEDAE52E69">2020</mods:date>
<mods:detail id="0E97B5BF3FBDAF96D34BFD86D68D480A" type="volume">
<mods:number id="6A9C9B3B9AE4921F4D4B5710713EAB28">189</mods:number>
</mods:detail>
<mods:extent id="9A4A615667EA1BFA99F6C2ABA5EDC2B2" unit="page">
<mods:start id="103E233EFF12DE6C02E8240FA85B7FD9">1232</mods:start>
<mods:end id="E78143A412745246C126A28B4D4FD424">1248</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="40251120EA569490D85B61DD4E8D1124">journal article</mods:classification>
<mods:identifier id="55B9B04515290C834442EE41228F1F37" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="891FC70A88A4515D807291A3652F7906" type="ZooBank">4D7E449-E29E-4E8F-A215-5E8FA38BC763</mods:identifier>
</mods:mods>
<treatment id="03F787C01E550B28FF0B7F3EFB59FA16" LSID="urn:lsid:plazi:treatment:03F787C01E550B28FF0B7F3EFB59FA16" httpUri="http://treatment.plazi.org/id/03F787C01E550B28FF0B7F3EFB59FA16" lastPageId="11" lastPageNumber="1243" pageId="10" pageNumber="1242">
<subSubSection id="C344655D1E550B29FF0B7F3EFE59FDC5" pageId="10" pageNumber="1242" type="nomenclature">
<paragraph id="8BE136D61E550B29FF0B7F3EFD64FD9A" blockId="10.[252,689,533,557]" box="[252,689,533,557]" pageId="10" pageNumber="1242">
<heading id="D0A981BA1E550B29FF0B7F3EFD64FD9A" box="[252,689,533,557]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="1242" reason="2">
<taxonomicName id="4C5E4D551E550B29FF0B7F3EFD92FD9C" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[252,583,533,557]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1242" phylum="Arthropoda" rank="species" species="gigagalea" status="sp. nov.">
<smallCapsWord id="8D07A00A1E550B29FF0B7F3EFE6FFD9C" baselines="551,550" box="[252,442,533,557]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Chimerogyrus" pageId="10" pageNumber="1242">CHIMEROGYRUS</smallCapsWord>
<smallCapsWord id="8D07A00A1E550B29FE357F33FD92FD9C" baselines="550" box="[450,583,536,555]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="gigagalea" pageId="10" pageNumber="1242">GIGAGALEA</smallCapsWord>
</taxonomicName>
<emphasis id="B92AEAC41E550B29FDB97F33FD64FD9A" bold="true" box="[590,689,533,557]" pageId="10" pageNumber="1242">
<taxonomicNameLabel id="A21957BF1E550B29FDB97F33FD64FD9A" box="[590,689,533,557]" pageId="10" pageNumber="1242" rank="species">
<smallCapsWord id="8D07A00A1E550B29FDB97F33FDBEFD9C" baselines="550" box="[590,619,536,555]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="10" pageNumber="1242">SP</smallCapsWord>
.
<smallCapsWord id="8D07A00A1E550B29FD8E7F33FD7FFD9C" baselines="550" box="[633,682,536,555]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="10" pageNumber="1242">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE136D61E550B29FF547F16FE59FDC5" blockId="10.[163,776,573,626]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF547F16FF01FDE5" box="[163,212,573,594]" italics="true" pageId="10" pageNumber="1242">lsid:</emphasis>
<uri id="FFCF3AD41E550B29FF2C7F16FE59FDC5" pageId="10" pageNumber="1242">
urn:lsid:zoobank.org:act:
<uuid id="FFF80C031E550B29FE197F16FE59FDC5" pageId="10" pageNumber="1242">0B604A53-EED0-4CD1- 9478-40C609EFE8A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C344655D1E550B28FF547FBAFB59FA16" lastPageId="11" lastPageNumber="1243" pageId="10" pageNumber="1242" type="materials_examined">
<paragraph id="8BE136D61E550B29FF547FBAFEA5FD73" blockId="10.[163,778,656,708]" pageId="10" pageNumber="1242">
<materialsCitation id="3B363C8B1E550B29FF547FBAFEB8FD73" country="Myanmar" location="Tanai" municipality="Hukawng valley" pageId="10" pageNumber="1242" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B92AEAC41E550B29FF547FBAFE9FFD12" box="[163,330,656,678]" italics="true" pageId="10" pageNumber="1242">
<typeStatus id="54E588741E550B29FF547FBAFF09FD11" box="[163,220,657,678]" pageId="10" pageNumber="1242">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F34976461E550B29FEAF7FBBFE1AFD12" box="[344,463,656,677]" name="Myanmar" pageId="10" pageNumber="1242">Myanmar</collectingCountry>
,
<collectingRegion id="499AF8341E550B29FE287FBBFD51FD12" box="[479,644,656,678]" country="Myanmar" name="Kachin" pageId="10" pageNumber="1242">Kachin State</collectingRegion>
,
<collectingMunicipality id="6B85ACAC1E550B29FD627FBBFF32FD73" pageId="10" pageNumber="1242">Hukawng valley</collectingMunicipality>
near
<location id="8E81600D1E550B29FEDF7F85FEB8FD73" LSID="urn:lsid:plazi:treatment:03F787C01E550B28FF0B7F3EFB59FA16:8E81600D1E550B29FEDF7F85FEB8FD73" box="[296,365,686,708]" country="Myanmar" municipality="Hukawng valley" name="Tanai" pageId="10" pageNumber="1242" stateProvince="Kachin">Tanai</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE136D61E550B29FF547FC5FEC5FC96" blockId="10.[163,778,749,802]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF547FC5FE90FCB5" box="[163,325,749,771]" italics="true" pageId="10" pageNumber="1242">
<typeStatus id="54E588741E550B29FF547FC5FF0DFCB4" box="[163,216,750,771]" pageId="10" pageNumber="1242">Type</typeStatus>
material:
</emphasis>
Single Burmese amber piece, deposited in ZSMG.
</paragraph>
<paragraph id="8BE136D61E550B29FF547E49FEF3FCCD" blockId="10.[163,294,866,890]" box="[163,294,866,890]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF547E49FEF3FCCD" box="[163,294,866,890]" italics="true" pageId="10" pageNumber="1242">Etymology</emphasis>
</paragraph>
<paragraph id="8BE136D61E550B29FF547EA1FEAAFC4B" blockId="10.[163,780,906,1020]" pageId="10" pageNumber="1242">
The specific epithet is a combination of the Greek γίγας,
<emphasis id="B92AEAC41E550B29FF547E81FF0AFC08" box="[163,223,938,959]" italics="true" pageId="10" pageNumber="1242">gigas</emphasis>
, giant, and Latin
<emphasis id="B92AEAC41E550B29FE5D7E82FE32FC09" box="[426,487,937,958]" italics="true" pageId="10" pageNumber="1242">galea</emphasis>
, an anatomical structure, in reference to the large galea of the new taxon. It is a noun in apposition.
</paragraph>
<paragraph id="8BE136D61E550B29FF54791EFEFAFBFB" blockId="10.[163,303,1077,1100]" box="[163,303,1077,1100]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF54791EFEFAFBFB" box="[163,303,1077,1100]" italics="true" pageId="10" pageNumber="1242">Description</emphasis>
</paragraph>
<paragraph id="8BE136D61E550B29FF547976FEE6FB78" blockId="10.[163,779,1117,1231]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF547976FF08FBC5" box="[163,221,1117,1138]" italics="true" pageId="10" pageNumber="1242">Body</emphasis>
(
<figureCitation id="13652A531E550B29FF057976FEB3FBC4" box="[242,358,1117,1139]" captionStart="Figure 7" captionStartId="11.[145,224,1075,1097]" captionTargetBox="[146,1428,196,1035]" captionTargetId="figure-324@11.[145,1428,195,1036]" captionTargetPageId="11" captionText="Figure 7. Chimerogyrus gigagalea† in situ. A, B, habitus images. Scale bars: 2 mm. A, dorsal view. B, ventral view. C, D, cephalic capsule. C, dorsal view. D, ventral view." pageId="10" pageNumber="1242">Fig. 7A, B</figureCitation>
): Elongate, parallel sided; head and pronotum strongly sclerotized, remainder of thorax and abdomen weakly sclerotized. Body size:
<emphasis id="B92AEAC41E550B29FD8579B0FD57FB07" box="[626,642,1179,1200]" italics="true" pageId="10" pageNumber="1242">L</emphasis>
= 11.5 mm,
<emphasis id="B92AEAC41E550B29FF547991FF6FFB78" box="[163,186,1210,1231]" italics="true" pageId="10" pageNumber="1242">W</emphasis>
= 1.0 mm.
</paragraph>
<paragraph id="8BE136D61E550B29FF5479D3FA40FD71" blockId="10.[163,780,1272,1876]" lastBlockId="10.[827,1443,197,710]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FF5479D3FE67FABA" box="[163,434,1272,1293]" italics="true" pageId="10" pageNumber="1242">Head: Cephalic capsule</emphasis>
(
<figureCitation id="13652A531E550B29FE4879D3FDD7FABA" box="[447,514,1272,1294]" captionStart="Figure 3" captionStartId="5.[145,229,626,648]" captionTargetBox="[145,764,196,583]" captionTargetPageId="5" captionText="Figure 3. Chimerogyrus gigagalea† line drawings of cephalic capsule. Scale bar: 0.16 mm. A, dorsal view. B, ventral view. Abbreviations: FR, frons; PA, parietal." pageId="10" pageNumber="1242">Figs 3</figureCitation>
,
<figureCitation id="13652A531E550B29FDFB79D3FD92FABA" box="[524,583,1272,1293]" captionStart="Figure 7" captionStartId="11.[145,224,1075,1097]" captionTargetBox="[146,1428,196,1035]" captionTargetId="figure-324@11.[145,1428,195,1036]" captionTargetPageId="11" captionText="Figure 7. Chimerogyrus gigagalea† in situ. A, B, habitus images. Scale bars: 2 mm. A, dorsal view. B, ventral view. C, D, cephalic capsule. C, dorsal view. D, ventral view." pageId="10" pageNumber="1242">7C, D</figureCitation>
): subrectangular, lateral margins slightly longer than anterior and posterior margins, without constriction of occipital region; occipital foramen dorsally with narrow margin; occipital suture absent; coronal suture short; frontal sutures U-shaped, extending to antennal bases; frontoclypeus roughly subtriangular, anterior margin divided into three lobes, medial lobe symmetrical, with four equally sized, well-developed teeth; tormae absent; lateral lobes well developed, not projecting anteriorly beyond medial lobe;
<collectionCode id="ED4FAE131E550B29FE727B00FE73F9F7" box="[389,422,1579,1600]" pageId="10" pageNumber="1242">PA</collectionCode>
with six stemmata at each side, four situated dorsally, two ventrally.
<emphasis id="B92AEAC41E550B29FDC97B62FD4BF9E9" box="[574,670,1609,1630]" italics="true" pageId="10" pageNumber="1242">Antenna</emphasis>
(
<figureCitation id="13652A531E550B29FD5A7B62FD28F9E9" box="[685,765,1609,1631]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="10" pageNumber="1242">Fig. 4A</figureCitation>
): elongate and slender; composed of four antennomeres, with I shortest and round; II and III longest, subequal in length, with II cylindrical and straight, III weakly curved; IV shorter, ~3× length of I, weakly curved.
<emphasis id="B92AEAC41E550B29FF547BC9FEC5F940" box="[163,272,1762,1783]" italics="true" pageId="10" pageNumber="1242">Mandible</emphasis>
(
<figureCitation id="13652A531E550B29FEE97BC9FEBAF94F" box="[286,367,1762,1784]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="10" pageNumber="1242">Fig. 4B</figureCitation>
): relatively elongate, curved inward; broadest basally and gradually attenuated apically, ending in sharp apex; inner margin smooth, without denticles or teeth; retinaculum absent.
<emphasis id="B92AEAC41E550B29FD987A15FD1CF8E4" box="[623,713,1854,1875]" italics="true" pageId="10" pageNumber="1242">Maxilla</emphasis>
(
<figureCitation id="13652A531E550B29FD2E7A15FCA2FF6C" box="[729,887,197,1876]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="10" pageNumber="1242">Figs 4C, D</figureCitation>
): well developed, not covering ventral mandibular joint; cardo elongate, subrectangular; stipes long, subtrapezoidal, bearing a lacinia and galea on distal inner margin, distal outer margin with palpifer; dorsally without series of small hook-like setae on internal margin; lacinia relatively small and strongly curved, with sharp apex; posterior margin straight in apical half, dentate basally; galea longer than lacinia, two-segmented, basal segment short and broader, apical segment elongate, tapered towards apex, ~2.5× length of basal segment; palpifer short, palpomerelike in form; maxillary palp elongate and narrow, composed of three palpomeres; I shortest, III longest.
<emphasis id="B92AEAC41E550B29FCCC7F7FFC4EFDDE" box="[827,923,596,617]" italics="true" pageId="10" pageNumber="1242">Labium</emphasis>
(
<figureCitation id="13652A531E550B29FC587F7FFBF9FDDE" box="[943,1068,596,618]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="10" pageNumber="1242">Fig. 4E, F</figureCitation>
): well developed; prementum divided longitudinally into two subcylindrical halves fused basally; labial palps elongate and narrow, two-segmented; segments I and II nearly equal in length.
</paragraph>
<paragraph id="8BE136D61E550B29FCCC7FC5FBE3FBC3" blockId="10.[827,1444,750,1140]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FCCC7FC5FC46FCB4" box="[827,915,750,771]" italics="true" pageId="10" pageNumber="1242">Thorax:</emphasis>
Pronotum slightly larger than subequal meso- and metanotum; protergite well developed, dorsally covering entire segment, with straight anterior margin, rounded lateral and posterior margins, sagittal line evident; membrane separating pro- and mesonotum, with narrow sclerite possessing rounded posterior margin and sagittal line.
<emphasis id="B92AEAC41E550B29FB2E7E8CFADBFC0B" box="[1241,1294,935,956]" italics="true" pageId="10" pageNumber="1242">Legs</emphasis>
(
<figureCitation id="13652A531E550B29FAE87E8DFA41FC0B" box="[1311,1428,934,956]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="10" pageNumber="1242">Fig. 5AE</figureCitation>
): long, slender, composed of six segments; L3 longest, L1 shortest;
<collectionCode id="ED4FAE131E550B29FC3B7ECFFC24FC4E" box="[972,1009,996,1017]" pageId="10" pageNumber="1242">CO</collectionCode>
ovoid, robust,
<collectionCode id="ED4FAE131E550B29FB6F7ECFFB6FFC4E" box="[1176,1210,996,1017]" pageId="10" pageNumber="1242">TR</collectionCode>
short,
<collectionCode id="ED4FAE131E550B29FAFC7ECFFAF8FC4E" box="[1291,1325,996,1017]" pageId="10" pageNumber="1242">FE</collectionCode>
relatively broad,
<collectionCode id="ED4FAE131E550B29FC7D7928FC70FBAF" box="[906,933,1027,1048]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="10" pageNumber="1242">TI</collectionCode>
narrower,
<collectionCode id="ED4FAE131E550B29FBD47929FB90FBA0" box="[1059,1093,1026,1047]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="10" pageNumber="1242">TA</collectionCode>
narrowest and subcylindrical in form,
<collectionCode id="ED4FAE131E550B29FC51790AFC1DFB81" box="[934,968,1057,1078]" pageId="10" pageNumber="1242">PT</collectionCode>
with two long, slender, weakly curved pretarsal claws, posterior claw shorter than anterior claw; spinulae absent.
</paragraph>
<paragraph id="8BE136D61E550B29FCCC79B6FAD7F9C8" blockId="10.[827,1443,1181,1663]" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FCCC79B6FC68FB04" box="[827,957,1181,1203]" italics="true" pageId="10" pageNumber="1242">Abdomen:</emphasis>
Ten-segmented, with segments I VIII similar in form, bearing a tracheal gill at posterolateral angle; segment IX with two tracheal gills at posterolateral angle; segment I similar in size to metanotum; segments IIVII larger and similar in size; segments VIIIIX gradually becoming smaller; tracheal gills slender, plumose with spinulae on anterior and posterior surfaces; gill of segment I smallest, gills of segments IIV larger and similar in size, gills of segments VIIX becoming gradually larger posteriorly, with gills of segment IX largest; segment X small and narrow, arising from posteroventral face of segment IX, without tracheal gills and ventral spinulae, bearing four strongly sclerotized terminal hooks; lateral hooks narrower and more elongate, medial hooks broader and less elongate.
</paragraph>
<paragraph id="8BE136D61E550B28FCCC7B8CFBB4FB5F" blockId="10.[827,1444,1703,1879]" lastBlockId="11.[809,1424,1203,1256]" lastPageId="11" lastPageNumber="1243" pageId="10" pageNumber="1242">
<emphasis id="B92AEAC41E550B29FCCC7B8CFBA6F90B" box="[827,1139,1703,1724]" italics="true" pageId="10" pageNumber="1242">Chaetotaxy: Frontoclypeus:</emphasis>
numerous short, hair-like setae posteriorly.
<emphasis id="B92AEAC41E550B29FC0D7BEDFB8DF96C" box="[1018,1112,1734,1755]" italics="true" pageId="10" pageNumber="1242">Parietal:</emphasis>
basal half with one long hair-like seta (
<collectionCode id="ED4FAE131E550B29FC547BCEFC13F94C" box="[931,966,1765,1787]" pageId="10" pageNumber="1242">PA</collectionCode>
6) evident dorsally.
<emphasis id="B92AEAC41E550B29FB627BCEFB3EF94D" box="[1173,1259,1765,1786]" italics="true" pageId="10" pageNumber="1242">Maxilla</emphasis>
(
<figureCitation id="13652A531E550B29FB0F7BCEFA91F94D" box="[1272,1348,1765,1787]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="10" pageNumber="1242">Fig. 4C</figureCitation>
): base of lacinia bearing two straight hair-like setae of subequal length, anterior (
<collectingCountry id="F34976461E550B29FBF27A09FBE6F88F" box="[1029,1075,1826,1848]" name="Mexico" pageId="10" pageNumber="1242">MX</collectingCountry>
5) longer than posterior (
<collectingCountry id="F34976461E550B29FAAB7A09FA5EF88F" box="[1372,1419,1826,1848]" name="Mexico" pageId="10" pageNumber="1242">MX</collectingCountry>
6).
<emphasis id="B92AEAC41E550B29FCCC7A6AFCBBF8E1" box="[827,878,1857,1878]" italics="true" pageId="10" pageNumber="1242">Legs</emphasis>
(
<figureCitation id="13652A531E550B29FC897A6AFC3AF8E0" box="[894,1007,1857,1879]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="10" pageNumber="1242">Fig. 5AE</figureCitation>
): anterior face of
<collectionCode id="ED4FAE131E550B29FB4E7A6AFB0BF8E1" box="[1209,1246,1857,1878]" pageId="10" pageNumber="1242">CO</collectionCode>
with three large, stout, spine-like setae (
<collectionCode id="ED4FAE131E540B28FE557998FE1EFB7E" box="[418,459,1203,1225]" pageId="11" pageNumber="1243">CO</collectionCode>
6,
<collectionCode id="ED4FAE131E540B28FE127998FDD8FB7F" box="[485,525,1203,1224]" pageId="11" pageNumber="1243">CO</collectionCode>
8 and
<collectionCode id="ED4FAE131E540B28FDA27998FDA9FB7F" box="[597,636,1203,1224]" pageId="11" pageNumber="1243">CO</collectionCode>
9) situated distomedially and one small hair-like seta (
<collectionCode id="ED4FAE131E540B28FD5B79F9FD00FB50" box="[684,725,1234,1255]" pageId="11" pageNumber="1243">CO</collectionCode>
10) distoventrally; posterior face proximally with one hair-like seta (
<collectionCode id="ED4FAE131E540B28FF0A7824FEF1FA93" box="[253,292,1295,1316]" pageId="11" pageNumber="1243">CO</collectionCode>
12) and two hair-like setae (
<collectionCode id="ED4FAE131E540B28FDAD7824FD57FA93" box="[602,642,1295,1316]" pageId="11" pageNumber="1243">CO</collectionCode>
11 and one probably additional or secondary seta) distally; anterior face of
<collectionCode id="ED4FAE131E540B28FF167866FED6FAD5" box="[225,259,1357,1378]" pageId="11" pageNumber="1243">TR</collectionCode>
with one short hair-like seta (
<collectionCode id="ED4FAE131E540B28FDA87867FD51FAD6" box="[607,644,1356,1377]" pageId="11" pageNumber="1243">TR</collectionCode>
3) and one long hair-like seta (
<collectionCode id="ED4FAE131E540B28FE877840FE41FA37" box="[368,404,1387,1408]" pageId="11" pageNumber="1243">TR</collectionCode>
4) ventrodistally; posterior face of
<collectionCode id="ED4FAE131E540B28FF5C78A1FF18FA28" box="[171,205,1418,1439]" pageId="11" pageNumber="1243">TR</collectionCode>
with one short hair-like seta (
<collectionCode id="ED4FAE131E540B28FDEC78A2FDEAFA28" box="[539,575,1417,1439]" pageId="11" pageNumber="1243">TR</collectionCode>
5) ventrodistally; anterior face of
<collectionCode id="ED4FAE131E540B28FEBC7882FEB8FA09" box="[331,365,1449,1470]" pageId="11" pageNumber="1243">FE</collectionCode>
distally with two short spine-like setae, one (
<collectionCode id="ED4FAE131E540B28FEF878ECFEE7FA6B" box="[271,306,1479,1500]" pageId="11" pageNumber="1243">FE</collectionCode>
2) dorsally and another (
<collectionCode id="ED4FAE131E540B28FDBB78ECFDA5FA6B" box="[588,624,1479,1500]" pageId="11" pageNumber="1243">FE</collectionCode>
3) ventrally; posterior face of
<collectionCode id="ED4FAE131E540B28FEBA78CDFEBAFA4C" box="[333,367,1510,1531]" pageId="11" pageNumber="1243">FE</collectionCode>
distally with one longer spine-like seta (
<collectionCode id="ED4FAE131E540B28FF257B2FFF23F9AE" box="[210,246,1540,1561]" pageId="11" pageNumber="1243">FE</collectionCode>
4) and two small spine-like setae (
<collectionCode id="ED4FAE131E540B28FD657B2FFD63F9AD" box="[658,694,1540,1562]" pageId="11" pageNumber="1243">FE</collectionCode>
5 and
<collectionCode id="ED4FAE131E540B28FF667B08FF63F98E" box="[145,182,1571,1593]" pageId="11" pageNumber="1243">FE</collectionCode>
6); anterior face of
<collectionCode id="ED4FAE131E540B28FE657B08FE78F98F" box="[402,429,1571,1592]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
with one long hair-like seta (
<collectionCode id="ED4FAE131E540B28FF6E7B69FF63F9E0" box="[153,182,1602,1623]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
1) proximally and three short spine-like setae distally (
<collectionCode id="ED4FAE131E540B28FF6E7B4BFF60F9C2" box="[153,181,1632,1653]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
2,
<collectionCode id="ED4FAE131E540B28FF3B7B4BFF3DF9C2" box="[204,232,1632,1653]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
3 and
<collectionCode id="ED4FAE131E540B28FEDD7B4BFE90F9C2" box="[298,325,1632,1653]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
4); posterior face of
<collectionCode id="ED4FAE131E540B28FDEA7B4AFDE2F9C1" box="[541,567,1633,1654]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
distally with one long spine-like seta (
<collectionCode id="ED4FAE131E540B28FE817B54FE44F922" box="[374,401,1663,1685]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
5) and two short spine-like setae (
<collectionCode id="ED4FAE131E540B28FF6F7BB5FF66F903" box="[152,179,1694,1716]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
6 and
<collectionCode id="ED4FAE131E540B28FF057BB5FED8F904" box="[242,269,1694,1715]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
7); anterior face of
<collectionCode id="ED4FAE131E540B28FE257BB5FE26F904" box="[466,499,1694,1715]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="11" pageNumber="1243">TA</collectionCode>
with one long spine-like seta at mid-length dorsally (
<collectionCode id="ED4FAE131E540B28FE257B97FE2CF966" box="[466,505,1724,1745]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="11" pageNumber="1243">TA</collectionCode>
1) and two short spine-like setae distally (
<collectionCode id="ED4FAE131E540B28FEAB7BF0FEA3F947" box="[348,374,1755,1776]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
2 and
<collectionCode id="ED4FAE131E540B28FE407BF0FE07F947" box="[439,466,1755,1776]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="1243">TI</collectionCode>
4); posterior face of
<collectionCode id="ED4FAE131E540B28FD577BF0FD14F947" box="[672,705,1755,1776]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="11" pageNumber="1243">TA</collectionCode>
with two short setae distally (
<collectionCode id="ED4FAE131E540B28FE527BD2FE1CF8B8" box="[421,457,1785,1807]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="11" pageNumber="1243">TA</collectionCode>
5 and
<collectionCode id="ED4FAE131E540B28FDFB7BD1FDE4F8A7" box="[524,561,1786,1808]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="11" pageNumber="1243">TA</collectionCode>
6); anterior face of
<collectionCode id="ED4FAE131E540B28FF667A32FF64F899" box="[145,177,1817,1838]" pageId="11" pageNumber="1243">PT</collectionCode>
with one short spine-like seta on basoventral portion (
<collectionCode id="ED4FAE131E540B28FCC67998FC8DFB7F" box="[817,856,1203,1224]" pageId="11" pageNumber="1243">PT</collectionCode>
1).
<emphasis id="B92AEAC41E540B28FC827998FC0AFB7F" box="[885,991,1203,1224]" italics="true" pageId="11" pageNumber="1243">Abdomen</emphasis>
(
<figureCitation id="13652A531E540B28FC077998FB8BFB7F" box="[1008,1118,1203,1225]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="11" pageNumber="1243">Fig. 5F, G</figureCitation>
): segment X with two long hair-like setae paramedially.
</paragraph>
<caption id="DF21665E1E540B28FF667918FD50FBD0" pageId="11" pageNumber="1243" startId="11.[145,224,1075,1097]" targetBox="[146,1428,196,1035]" targetPageId="11" targetType="figure">
<paragraph id="8BE136D61E540B28FF667918FD50FBD0" blockId="11.[145,1425,1075,1127]" pageId="11" pageNumber="1243">
<emphasis id="B92AEAC41E540B28FF667918FF2EFBFE" bold="true" box="[145,251,1075,1097]" pageId="11" pageNumber="1243">Figure 7.</emphasis>
<taxonomicName id="4C5E4D551E540B28FEF17918FDDDFBFE" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[262,520,1075,1097]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1243" phylum="Arthropoda" rank="species" species="gigagalea">
<emphasis id="B92AEAC41E540B28FEF17918FDDDFBFE" box="[262,520,1075,1097]" italics="true" pageId="11" pageNumber="1243">Chimerogyrus gigagalea</emphasis>
</taxonomicName>
<emphasis id="B92AEAC41E540B28FDFF7918FDC6FBFE" bold="true" box="[520,531,1075,1097]" pageId="11" pageNumber="1243"></emphasis>
<emphasis id="B92AEAC41E540B28FDEC7918FD8AFBFE" box="[539,607,1075,1097]" italics="true" pageId="11" pageNumber="1243">in situ</emphasis>
. A, B, habitus images. Scale bars: 2 mm. A, dorsal view. B, ventral view. C, D, cephalic capsule. C, dorsal view. D, ventral view.
</paragraph>
</caption>
<paragraph id="8BE136D61E540B28FCDE780EFBE6FA8A" blockId="11.[809,1075,1317,1341]" box="[809,1075,1317,1341]" pageId="11" pageNumber="1243">
<emphasis id="B92AEAC41E540B28FCDE780EFBE6FA8A" box="[809,1075,1317,1341]" italics="true" pageId="11" pageNumber="1243">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8BE136D61E540B28FCDE7865FB59FA16" blockId="11.[809,1424,1358,1441]" pageId="11" pageNumber="1243">The genus is monotypic. The generic diagnosis provided above separates the species from all other known species based on larvae.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,205 @@
<document id="12AFFD0538CF3A537049048EF229957B" ID-ISSN="0024-4082" ID-ZooBank="4D7E449-E29E-4E8F-A215-5E8FA38BC763" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353372604" checkinUser="plazi" docAuthor="Gustafson, Grey T., Michat, Mariano C. &amp; Balke, Michael" docDate="2020" docId="03F787C01E570B29FC2778F1FE2AFE67" docLanguage="en" docName="zlz161.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Chimerogyrus Gustafson, Michat &amp; Balke, 2020, GEN. NOV." docType="treatment" docVersion="1" lastPageNumber="1242" masterDocId="FFCEFFB81E5F0B23FFF77D2BFFD5FFB7" masterDocTitle="Burmese amber reveals a new stem lineage of whirligig beetle (Coleoptera: Gyrinidae) based on the larval stage" masterLastPageNumber="1248" masterPageNumber="1232" pageNumber="1240" updateTime="1739358064101" updateUser="GgImagineBatch">
<mods:mods id="D03AAF43C01AB81EF478D3574868D6AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="900FE97604D9AEBC36049D0F9B5A8125">
<mods:title id="290FD1B2E246362548790D1B0778B0BC">Burmese amber reveals a new stem lineage of whirligig beetle (Coleoptera: Gyrinidae) based on the larval stage</mods:title>
</mods:titleInfo>
<mods:name id="5EDB97E1574A958441FDA547FD642ECA" type="personal">
<mods:role id="6BFEB09DBA5FC7583167597DCEC23A8B">
<mods:roleTerm id="19FC67700268337D9904DA5368BF1C25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9C26B65A10EDF673DA5C196A8BB6903">Gustafson, Grey T.</mods:namePart>
</mods:name>
<mods:name id="A970AB7FDCB471BC0D660AB9B1E90141" type="personal">
<mods:role id="63FB9C58ABB0BD485292902A9AC80744">
<mods:roleTerm id="233BCB3F1208E1926872980FF056C18A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBF2ED208AEC65FC1AD904C764B3D596">Michat, Mariano C.</mods:namePart>
</mods:name>
<mods:name id="1FEE2A50FF0E67D622E83CA801BA9448" type="personal">
<mods:role id="1E8B18C1850C9107786A19923301D4AD">
<mods:roleTerm id="0C30130E3470F8E5631AA2640D8733B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27DFB88BCAD1EDD03EE907800314D152">Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="A15B343C059DD43CF0151539E3EB1A6C">text</mods:typeOfResource>
<mods:relatedItem id="6285DD9918CA79BC0A2A76DD2AA21B1F" type="host">
<mods:titleInfo id="9051E8E5A30DADBA24DCBDD9D8995E7C">
<mods:title id="B00D9D1DE184EAA1BA065D187DA17732">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="7ACC385A0D649FA7DF39F26DFCD926FC">
<mods:date id="7738CB808D71AADA28E517E372810427">2020</mods:date>
<mods:detail id="DE46A97B2C803D081F2210A0F0119E64" type="volume">
<mods:number id="143963DE9580EBECDB0C1CFC31A3616A">189</mods:number>
</mods:detail>
<mods:extent id="625D84F8518C838D6D2664F15C1F5480" unit="page">
<mods:start id="A54C3E4DD6FB696863024F7C900D7155">1232</mods:start>
<mods:end id="DDE868640ACF4F82CB2832FFBB54703A">1248</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="54769A9F809E6F60924525B7DFC6ABFE">journal article</mods:classification>
<mods:identifier id="E3FEB54C796F3B9FB6E2CABFEA32EDC7" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="70D87A67FDA668FD0292B40AA0722857" type="ZooBank">4D7E449-E29E-4E8F-A215-5E8FA38BC763</mods:identifier>
</mods:mods>
<treatment id="03F787C01E570B29FC2778F1FE2AFE67" LSID="urn:lsid:plazi:treatment:03F787C01E570B29FC2778F1FE2AFE67" httpUri="http://treatment.plazi.org/id/03F787C01E570B29FC2778F1FE2AFE67" lastPageId="10" lastPageNumber="1242" pageId="8" pageNumber="1240">
<subSubSection id="C344655D1E570B2BFC2778F1FBEAF980" pageId="8" pageNumber="1240" type="nomenclature">
<paragraph id="8BE136D61E570B2BFC2778F1FAD8FA45" blockId="8.[976,1293,1498,1522]" box="[976,1293,1498,1522]" pageId="8" pageNumber="1240">
<heading id="D0A981BA1E570B2BFC2778F1FAD8FA45" box="[976,1293,1498,1522]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="1240" reason="2">
<taxonomicName id="4C5E4D551E570B2BFC2778F1FB5BFA47" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[976,1166,1498,1522]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1240" phylum="Arthropoda" rank="genus" status="gen. nov.">
<smallCapsWord id="8D07A00A1E570B2BFC2778F1FB5BFA47" baselines="1516,1515" box="[976,1166,1498,1522]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Chimerogyrus" pageId="8" pageNumber="1240">CHIMEROGYRUS</smallCapsWord>
</taxonomicName>
<emphasis id="B92AEAC41E570B2BFB6178F6FAD8FA45" bold="true" box="[1174,1293,1498,1522]" pageId="8" pageNumber="1240">
<taxonomicNameLabel id="A21957BF1E570B2BFB6178F6FAD8FA45" box="[1174,1293,1498,1522]" pageId="8" pageNumber="1240" rank="genus">
<smallCapsWord id="8D07A00A1E570B2BFB6178F6FB12FA47" baselines="1515" box="[1174,1223,1501,1520]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="gen" pageId="8" pageNumber="1240">GEN</smallCapsWord>
.
<smallCapsWord id="8D07A00A1E570B2BFB2278F6FAD3FA47" baselines="1515" box="[1237,1286,1501,1520]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="8" pageNumber="1240">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE136D61E570B2BFCCC7B28FBEAF980" blockId="8.[827,1443,1539,1591]" pageId="8" pageNumber="1240">
<emphasis id="B92AEAC41E570B2BFCCC7B28FBEAF980" italics="true" pageId="8" pageNumber="1240">
lsid:
<uri id="FFCF3AD41E570B2BFC827B28FBEAF980" pageId="8" pageNumber="1240">
urn:lsid:zoobank.org:act:
<uuid id="FFF80C031E570B2BFB787B28FBEAF980" pageId="8" pageNumber="1240">D7259533-80FC-44A2- B58B-FEDE654E53A5</uuid>
</uri>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C344655D1E570B29FCCC7B7CFE2AFE67" lastPageId="10" lastPageNumber="1242" pageId="8" pageNumber="1240" type="materials_examined">
<paragraph id="8BE136D61E570B2BFCCC7B7CFB9CF93C" blockId="8.[827,1442,1622,1675]" pageId="8" pageNumber="1240">
<emphasis id="B92AEAC41E570B2BFCCC7B7CFACDF9DC" box="[827,1304,1622,1644]" italics="true" pageId="8" pageNumber="1240">
<typeStatus id="54E588741E570B2BFCCC7B7CFCA6F9DB" box="[827,883,1623,1644]" pageId="8" pageNumber="1240">Type</typeStatus>
species:
<taxonomicName id="4C5E4D551E570B2BFC117B7DFAC7F9DC" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[998,1298,1622,1643]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1240" phylum="Arthropoda" rank="species" species="gigagalea">Chimerogyrus gigagalea</taxonomicName>
,
</emphasis>
by current designation, monotypic.
</paragraph>
<paragraph id="8BE136D61E570B2BFCCC7B90FC69F964" blockId="8.[827,956,1723,1747]" box="[827,956,1723,1747]" pageId="8" pageNumber="1240">
<emphasis id="B92AEAC41E570B2BFCCC7B90FC69F964" box="[827,956,1723,1747]" italics="true" pageId="8" pageNumber="1240">Etymology</emphasis>
</paragraph>
<paragraph id="8BE136D61E570B2AFCCC7BCFFD0CFB45" blockId="8.[827,1443,1764,1911]" lastBlockId="9.[145,729,1244,1266]" lastPageId="9" lastPageNumber="1241" pageId="8" pageNumber="1240">
The genus name is Latinized from the Greek χιμαιρα,
<emphasis id="B92AEAC41E570B2BFCCC7A28FC7FF8AF" box="[827,938,1795,1816]" italics="true" pageId="8" pageNumber="1240">chimaira</emphasis>
, a chimera, the amalgamate monster of legend, here in reference to the strange combination of ancestral and derived features of the larval taxon being described, and γύρος,
<emphasis id="B92AEAC41E570B2BFB8D7A4AFB63F8C1" box="[1146,1206,1889,1910]" italics="true" pageId="8" pageNumber="1240">gyros</emphasis>
, meaning turning or spinning, a common ending used in gyrinid names..
</paragraph>
<caption id="DF21665E1E560B2AFF667966FC98FB2A" pageId="9" pageNumber="1241" startId="9.[145,224,1101,1123]" targetBox="[146,1423,197,1060]" targetPageId="9" targetType="figure">
<paragraph id="8BE136D61E560B2AFF667966FC98FB2A" blockId="9.[145,1425,1101,1181]" pageId="9" pageNumber="1241">
<emphasis id="B92AEAC41E560B2AFF667966FF2EFBD5" bold="true" box="[145,251,1101,1123]" pageId="9" pageNumber="1241">Figure 6.</emphasis>
Summary of the results of the phylogenetic analyses. Tree topology derived from Bayesian analysis including
<taxonomicName id="4C5E4D551E560B2AFF667941FE98FB37" box="[145,333,1130,1152]" class="Insecta" family="Gyrinidae" genus="Cretogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="species" species="beuteli">
<emphasis id="B92AEAC41E560B2AFF667941FE98FB37" box="[145,333,1130,1152]" italics="true" pageId="9" pageNumber="1241">Cretogyrus beuteli</emphasis>
</taxonomicName>
†. Boxes at nodes indicate support values across the different analyses either with or without the inclusion of
<taxonomicName id="4C5E4D551E560B2AFF5D79ACFEBDFB2A" box="[170,360,1159,1181]" class="Insecta" family="Gyrinidae" genus="Cretogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="species" species="beuteli">
<emphasis id="B92AEAC41E560B2AFF5D79ACFEBDFB2A" box="[170,360,1159,1181]" italics="true" pageId="9" pageNumber="1241">Cretogyrus beuteli</emphasis>
</taxonomicName>
† based on the key provided at the bottom left.
</paragraph>
</caption>
<paragraph id="8BE136D61E560B2AFF66780DFEDFFA8A" blockId="9.[145,266,1318,1341]" box="[145,266,1318,1341]" pageId="9" pageNumber="1241">
<emphasis id="B92AEAC41E560B2AFF66780DFEDFFA8A" box="[145,266,1318,1341]" italics="true" pageId="9" pageNumber="1241">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE136D61E560B2AFF667865FD89F8F9" blockId="9.[145,762,1358,1870]" pageId="9" pageNumber="1241">
<taxonomicName id="4C5E4D551E560B2AFF667865FEE8FAD4" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[145,317,1358,1379]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E560B2AFF667865FEE8FAD4" box="[145,317,1358,1379]" italics="true" pageId="9" pageNumber="1241">Chimerogyrus</emphasis>
</taxonomicName>
can be diagnosed by the following combination of characters: cephalic capsule (
<figureCitation id="13652A531E560B2AFD747847FD12FA36" box="[643,711,1388,1410]" captionStart="Figure 3" captionStartId="5.[145,229,626,648]" captionTargetBox="[145,764,196,583]" captionTargetPageId="5" captionText="Figure 3. Chimerogyrus gigagalea† line drawings of cephalic capsule. Scale bar: 0.16 mm. A, dorsal view. B, ventral view. Abbreviations: FR, frons; PA, parietal." pageId="9" pageNumber="1241">Figs 3</figureCitation>
,
<figureCitation id="13652A531E560B2AFD227847FF71FA17" captionStart="Figure 7" captionStartId="11.[145,224,1075,1097]" captionTargetBox="[146,1428,196,1035]" captionTargetId="figure-324@11.[145,1428,195,1036]" captionTargetPageId="11" captionText="Figure 7. Chimerogyrus gigagalea† in situ. A, B, habitus images. Scale bars: 2 mm. A, dorsal view. B, ventral view. C, D, cephalic capsule. C, dorsal view. D, ventral view." pageId="9" pageNumber="1241">7C, D</figureCitation>
) not constricted at level of occipital region;
<collectionCode id="ED4FAE131E560B2AFD6B78A0FD6BFA17" box="[668,702,1419,1440]" pageId="9" pageNumber="1241">FR</collectionCode>
(
<figureCitation id="13652A531E560B2AFD3978A0FF65FA08" captionStart="Figure 3" captionStartId="5.[145,229,626,648]" captionTargetBox="[145,764,196,583]" captionTargetPageId="5" captionText="Figure 3. Chimerogyrus gigagalea† line drawings of cephalic capsule. Scale bar: 0.16 mm. A, dorsal view. B, ventral view. Abbreviations: FR, frons; PA, parietal." pageId="9" pageNumber="1241">Fig. 3A</figureCitation>
) symmetrica,l with four equally sized, well-developed teeth; tormae absent; maxilla (
<figureCitation id="13652A531E560B2AFE1278E3FD86FA69" box="[485,595,1480,1502]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="9" pageNumber="1241">Fig. 4C, D</figureCitation>
) with elongate cardo and stipes; galea longer than lacinia; lacinia small and strongly curved, with inner margin straight apically and weakly dentate basally, apex without indentation; two long setae (
<collectingCountry id="F34976461E560B2AFEFA7B68FEEDF9EE" box="[269,312,1603,1625]" name="Mexico" pageId="9" pageNumber="1241">MX</collectingCountry>
5 and
<collectingCountry id="F34976461E560B2AFE8E7B68FE76F9EE" box="[377,419,1603,1625]" name="Mexico" pageId="9" pageNumber="1241">MX</collectingCountry>
6) subequal in length at base of lacinia; anterior face of
<collectionCode id="ED4FAE131E560B2AFE637B49FE6CF9C0" box="[404,441,1634,1655]" pageId="9" pageNumber="1241">CO</collectionCode>
(
<figureCitation id="13652A531E560B2AFE307B49FDE6F9C0" box="[455,563,1633,1656]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="9" pageNumber="1241">Fig. 5A, B</figureCitation>
) with three large, stout, spine-like setae (
<collectionCode id="ED4FAE131E560B2AFE6C7BABFE16F921" box="[411,451,1664,1686]" pageId="9" pageNumber="1241">CO</collectionCode>
6,
<collectionCode id="ED4FAE131E560B2AFE2B7BABFDD6F922" box="[476,515,1664,1685]" pageId="9" pageNumber="1241">CO</collectionCode>
8 and
<collectionCode id="ED4FAE131E560B2AFDBE7BABFDA4F922" box="[585,625,1664,1685]" pageId="9" pageNumber="1241">CO</collectionCode>
9);
<collectionCode id="ED4FAE131E560B2AFD667BABFD6DF922" box="[657,696,1664,1685]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="9" pageNumber="1241">TA</collectionCode>
1 long and spine like, situated at mid-length dorsally (
<figureCitation id="13652A531E560B2AFD6A7BB4FD3EF902" box="[669,747,1695,1717]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="9" pageNumber="1241">Fig. 5A</figureCitation>
); pretarsal claws (
<figureCitation id="13652A531E560B2AFEC97B95FE73F964" box="[318,422,1725,1748]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="9" pageNumber="1241">Fig. 5AE</figureCitation>
) without spinulae; tracheal gills with spinulae; segment X (
<figureCitation id="13652A531E560B2AFE467BF7FDCDF946" box="[433,536,1756,1778]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="9" pageNumber="1241">Fig. 5F, G</figureCitation>
) with two long, hair-like setae, spinulae absent; terminal hooks (
<figureCitation id="13652A531E560B2AFD777BD0FD25F8A6" box="[640,752,1787,1809]" captionStart="Figure 5" captionStartId="7.[145,225,1683,1705]" captionTargetBox="[241,1328,202,1642]" captionTargetId="figure-28@7.[239,1330,195,1643]" captionTargetPageId="7" captionText="Figure 5. Chimerogyrus gigagalea† line drawings and in situ images. AE, thoracic legs with setae labelled. Scale bar: 0.16 mm. A, left metathoracic leg, anterior view. B, in situ image of left metacoxa in anterior view, showing form of CO8 and CO9. C, right prothoracic leg, posterior view. D, right mesothoracic leg, posterior view. E, right metathoracic leg, posterior view. FH, abdominal segment X and terminal hooks. F, in situ image of abdominal segment X and terminal hooks. G, line drawing of abdominal segment X and terminal hooks. Scale bar: 0.16 mm. H, terminal hooks. Abbreviations: CO, coxa; FE, femur; PT, pretarsus; TA, tarsus; TI, tibia; TR, trochanter." pageId="9" pageNumber="1241">Fig. 5FH</figureCitation>
) asymmetrical, with lateral hooks narrower and elongate, medial hooks broader, less strongly curved.
</paragraph>
<paragraph id="8BE136D61E560B29FCBE79F7FD36FE80" blockId="9.[809,1426,1244,1879]" lastBlockId="10.[163,779,197,464]" lastPageId="10" lastPageNumber="1242" pageId="9" pageNumber="1241">
The form of the maxilla (
<figureCitation id="13652A531E560B2AFB8779F7FB31FB45" box="[1136,1252,1244,1266]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="9" pageNumber="1241">Fig. 4C, D</figureCitation>
) distinguishes
<taxonomicName id="4C5E4D551E560B2AFCDE79D0FC10FAA7" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[809,965,1275,1296]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E560B2AFCDE79D0FC10FAA7" box="[809,965,1275,1296]" italics="true" pageId="9" pageNumber="1241">Chimerogyrus</emphasis>
</taxonomicName>
from all known gyrinid genera.The maxilla of
<taxonomicName id="4C5E4D551E560B2AFCB37831FC35FA98" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[836,992,1306,1327]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E560B2AFCB37831FC35FA98" box="[836,992,1306,1327]" italics="true" pageId="9" pageNumber="1241">Chimerogyrus</emphasis>
</taxonomicName>
is most similar to that of genera of the tribes
<taxonomicName id="4C5E4D551E560B2AFC857813FC0AFAF9" box="[882,991,1336,1358]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1247" phylum="Arthropoda" rank="tribe" tribe="Dineutini">Dineutini</taxonomicName>
and Orectochilini, having an elongate cardo that does not cover the ventral joint of the mandible, with the internal margin of the stipes lacking small hook-like setae dorsally. However, the stipes is longer than that of any currently known dineutine or orectochiline larva, which have relatively short stipes (
<figureCitation id="13652A531E560B2AFB0278F9FABCFA50" box="[1269,1385,1490,1512]" captionStart="Figure 1" captionStartId="3.[145,225,1095,1117]" captionTargetBox="[145,1424,196,1055]" captionTargetPageId="3" captionText="Figure 1. Cephalic capsule of larvae of extant genera of the Dineutini and Gyrinini in dorsal view (to scale), including left maxilla in ventral view (not to scale), except for D, dorsal view of right maxilla. A, Macrogyrus ellipticus (Brullé, 1836). Scale bar: 0.25 mm. B, Dineutus sp. from Africa. Scale bar: 0.5 mm. C, Enhydrus sulcatus (Wiedemann, 1821). Scale bar: 0.50 mm. D, Cretogyrus beuteli† Zhao et al., 2019. Scale bar: 0.20 mm. Drawn from digital images provided by X. Zhao and after Zhao et al., (2019). E, Gyrinus vicinus Aubé, 1838. Scale bar: 0.25 mm. F, Aulonogyrus strigosus (Fabricius, 1801). Scale bar: 0.20 mm.After Beutel &amp; Roughley (1994). Abbreviations:AN, antenna; CA, cardo; FR, frons; GA, galea; LA, lacinia; LAP, labial palp; MN, mandible; MX, maxilla; MXP, maxillary palp; PA, parietal; ST, stipes." pageId="9" pageNumber="1241">Figs 1AD</figureCitation>
,
<figureCitation id="13652A531E560B2AFA8178F9FA56FA50" box="[1398,1411,1490,1511]" captionStart="Figure 2" captionStartId="4.[163,242,918,940]" captionTargetBox="[163,1443,195,875]" captionTargetId="graphics-424@4.[175,1443,195,859]" captionTargetPageId="4" captionText="Figure 2. Cephalic capsule of larvae of extant genera of the Orectochilini in dorsal view (to scale), including left maxilla in ventral view (not to scale). A, Gyretes sp. from Texas. Scale bar: 0.25 mm. B, Gyretes sp. from Texas. Cephalic capsule in ventral view. C, Orectogyrus Régimbart, 1884 sp. 2. Scale bar: 0.25 mm. D, Orectochilus villosus (Müller, 1776). Scale bar: 0.50 mm. Cephalic capsule after Noars (1956) and maxilla after Říha (1952). E, Patrus sp. Scale bar: 0.25 mm. Abbreviations: AN, antenna; CA, cardo; FR, frons; GA, galea; LA, lacinia; LAP, labial palp; MN, mandible; MX, maxilla; MXP, maxillary palp; PA, parietal; ST, stipes." pageId="9" pageNumber="1241">2</figureCitation>
). Furthermore, its maxilla is unlike any known gyrinid larva in that the galea is considerably longer than the small lacinia, rather than being roughly similar in size (as in
<taxonomicName id="4C5E4D551E560B2AFC857B66FB5DF9D5" authority="Macleay, 1825" authorityName="Macleay" authorityYear="1825" box="[882,1160,1612,1634]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1241" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E560B2AFC857B66FC0EF9D5" box="[882,987,1613,1634]" italics="true" pageId="9" pageNumber="1241">Dineutus</emphasis>
Macleay, 1825
</taxonomicName>
;
<figureCitation id="13652A531E560B2AFB617B67FB3CF9D6" box="[1174,1257,1612,1634]" captionStart="Figure 1" captionStartId="3.[145,225,1095,1117]" captionTargetBox="[145,1424,196,1055]" captionTargetPageId="3" captionText="Figure 1. Cephalic capsule of larvae of extant genera of the Dineutini and Gyrinini in dorsal view (to scale), including left maxilla in ventral view (not to scale), except for D, dorsal view of right maxilla. A, Macrogyrus ellipticus (Brullé, 1836). Scale bar: 0.25 mm. B, Dineutus sp. from Africa. Scale bar: 0.5 mm. C, Enhydrus sulcatus (Wiedemann, 1821). Scale bar: 0.50 mm. D, Cretogyrus beuteli† Zhao et al., 2019. Scale bar: 0.20 mm. Drawn from digital images provided by X. Zhao and after Zhao et al., (2019). E, Gyrinus vicinus Aubé, 1838. Scale bar: 0.25 mm. F, Aulonogyrus strigosus (Fabricius, 1801). Scale bar: 0.20 mm.After Beutel &amp; Roughley (1994). Abbreviations:AN, antenna; CA, cardo; FR, frons; GA, galea; LA, lacinia; LAP, labial palp; MN, mandible; MX, maxilla; MXP, maxillary palp; PA, parietal; ST, stipes." pageId="9" pageNumber="1241">Fig. 1B</figureCitation>
) or smaller in size, as in most genera. The form of the lacinia (
<figureCitation id="13652A531E560B2AFACD7B40FA5CF937" box="[1338,1417,1643,1665]" captionStart="Figure 4" captionStartId="6.[163,240,1037,1059]" captionTargetBox="[163,783,196,992]" captionTargetId="graphics-726@6.[191,775,196,613]" captionTargetPageId="6" captionText="Figure 4. Chimerogyrus gigagalea† line drawings. All scale bars: 0.10 mm. A, left antenna in dorsal view. B, mandibles in dorsal view. C, D, left maxilla. C, dorsal view. D, ventral view. E, F, labium. E, dorsal view. F, ventral view. Abbreviations: AN, antenna; LA, labium; MN, mandible; MX, maxilla." pageId="9" pageNumber="1241">Fig. 4C</figureCitation>
) is also unique, being strongly curved, with a straight inner margin apically, becoming dentate proximally and possessing two long setae basally. The presence of two long setae of subequal length basal to the lacinia is similar to larvae of the Orectochilini (
<figureCitation id="13652A531E560B2AFB3C7A2FFADCF8AE" box="[1227,1289,1796,1818]" captionStart="Figure 2" captionStartId="4.[163,242,918,940]" captionTargetBox="[163,1443,195,875]" captionTargetId="graphics-424@4.[175,1443,195,859]" captionTargetPageId="4" captionText="Figure 2. Cephalic capsule of larvae of extant genera of the Orectochilini in dorsal view (to scale), including left maxilla in ventral view (not to scale). A, Gyretes sp. from Texas. Scale bar: 0.25 mm. B, Gyretes sp. from Texas. Cephalic capsule in ventral view. C, Orectogyrus Régimbart, 1884 sp. 2. Scale bar: 0.25 mm. D, Orectochilus villosus (Müller, 1776). Scale bar: 0.50 mm. Cephalic capsule after Noars (1956) and maxilla after Říha (1952). E, Patrus sp. Scale bar: 0.25 mm. Abbreviations: AN, antenna; CA, cardo; FR, frons; GA, galea; LA, lacinia; LAP, labial palp; MN, mandible; MX, maxilla; MXP, maxillary palp; PA, parietal; ST, stipes." pageId="9" pageNumber="1241">Fig. 2</figureCitation>
), but unlike these genera the lacinia is short, somewhat similar to the
<taxonomicName id="4C5E4D551E560B2AFCDE7A6AFC5FF8E0" box="[809,906,1857,1879]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1232" phylum="Arthropoda" rank="tribe" tribe="Gyrinini">Gyrinini</taxonomicName>
, but much narrower and more hook shaped. It can be distinguished from the extinct genus
<taxonomicName id="4C5E4D551E550B29FD657DEDFCDEFF6C" authorityName="Zhao, Zhao, Jarzembowski &amp; Wang" authorityYear="2019" box="[658,779,198,219]" class="Insecta" family="Gyrinidae" genus="Cretogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1242" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E550B29FD657DEDFCDEFF6C" box="[658,779,198,219]" italics="true" pageId="10" pageNumber="1242">Cretogyrus</emphasis>
</taxonomicName>
by having a cephalic capsule without a constriction of the occipital region, four equally sized frontal teeth, the short coronal suture, and by the form of the maxilla.
</paragraph>
<paragraph id="8BE136D61E550B29FF4C7C6BFE2AFE67" blockId="10.[163,779,197,464]" pageId="10" pageNumber="1242">
The setation of the coxae also distinguishes
<taxonomicName id="4C5E4D551E550B29FF547C75FE86FEC4" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[163,339,350,371]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1242" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E550B29FF547C75FE86FEC4" box="[163,339,350,371]" italics="true" pageId="10" pageNumber="1242">Chimerogyrus</emphasis>
</taxonomicName>
from all currently known gyrinid larvae.
<taxonomicName id="4C5E4D551E550B29FEF67C56FE79FE25" authorityName="Gustafson &amp; Michat &amp; Balke" authorityYear="2020" box="[257,428,381,402]" class="Insecta" family="Gyrinidae" genus="Chimerogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1242" phylum="Arthropoda" rank="genus">
<emphasis id="B92AEAC41E550B29FEF67C56FE79FE25" box="[257,428,381,402]" italics="true" pageId="10" pageNumber="1242">Chimerogyrus</emphasis>
</taxonomicName>
has a series of large, stout, spine-like setae (
<collectionCode id="ED4FAE131E550B29FE957CB7FE5FFE05" box="[354,394,412,434]" pageId="10" pageNumber="1242">CO</collectionCode>
6,
<collectionCode id="ED4FAE131E550B29FE557CB7FE1CFE06" box="[418,457,412,433]" pageId="10" pageNumber="1242">CO</collectionCode>
8 and
<collectionCode id="ED4FAE131E550B29FDFA7CB7FDE1FE06" box="[525,564,412,433]" pageId="10" pageNumber="1242">CO</collectionCode>
9) distomedially on the anterior surface of the
<collectionCode id="ED4FAE131E550B29FE247C90FE2EFE67" box="[467,507,443,464]" pageId="10" pageNumber="1242">CO</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,243 @@
<document id="DF8941C5B61357D4EF9E0C8CDB9D326A" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFC6720379F3F943FE65C071" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha disjuncta" docType="treatment" docVersion="1" lastPageNumber="1351" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1351" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="4E8641DEE034E1189AB59F64DFD5DF44" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EEFC6D8E9758CCF31658DA373F2C72FD">
<mods:title id="AAFEDAB08FB3F65758D9278D6BC8DEB6">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="79C7F6E22D6FCBF2E7644B97E15A4F15" type="personal">
<mods:role id="72DCD8729842B7D872058D796717EA03">
<mods:roleTerm id="29414CB45960A0062E8AFA76F1884CD8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB1D0B49620707F8A91DE0A44E254668">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="014124230A5153F7F4CEF11647F5849E" type="personal">
<mods:role id="B163F43731276EDA816FD392E1B1719A">
<mods:roleTerm id="952AAB2D27609B6D5F6A5835971D487F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F2EB28779798AAE6585DDB0549E7E6E">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C6A11509FC68E8847FB7F4F8F930F855">text</mods:typeOfResource>
<mods:relatedItem id="594D5B5ABB8670C76970C58DC4572A86" type="host">
<mods:titleInfo id="E2F9312886844699391537BD069E2B4F">
<mods:title id="308A8C53E7E3B8716D7E747EC8C2F5FE">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="3454FDD33D75D369A1BADCCAE2B09802">
<mods:date id="B8804A6E85D5288EA66D349D28C8C622">2020</mods:date>
<mods:detail id="5B76677336D4EF6B83815FE8EB3E63FA" type="volume">
<mods:number id="99AC7DFA1A5740F0D00DBA39006EDC25">189</mods:number>
</mods:detail>
<mods:extent id="074FA57BC01E7E73E1A2C3DBF875C3D9" unit="page">
<mods:start id="E3AB9CAE8C6A187E8F5999A1E58728DA">1313</mods:start>
<mods:end id="DF37E5F9CC8DD191A8666EC159AE14F3">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="44B652F383BC491BC8FE0A52D7FA4678">journal article</mods:classification>
<mods:identifier id="B4DB38C0D3CD1341EBCD1C59FC5BD96D" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="4B483D3DFEA2E24FF15F188E0A95A159" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFC6720379F3F943FE65C071" LSID="urn:lsid:plazi:treatment:095A87E6FFC6720379F3F943FE65C071" httpUri="http://treatment.plazi.org/id/095A87E6FFC6720379F3F943FE65C071" lastPageId="39" lastPageNumber="1351" pageId="38" pageNumber="1351">
<subSubSection id="C9E9657BFFC6720279F3F943FD7FC1F9" box="[222,683,1710,1735]" pageId="38" pageNumber="1351" type="nomenclature">
<paragraph id="814C36F0FFC6720279F3F943FD7FC1F9" blockId="38.[222,683,1710,1735]" box="[222,683,1710,1735]" pageId="38" pageNumber="1351">
<heading id="DA04819CFFC6720279F3F943FD7FC1F9" box="[222,683,1710,1735]" centered="true" fontSize="9" level="2" pageId="38" pageNumber="1351" reason="2">
<taxonomicName id="46F34D73FFC6720279F3F943FD7FC1F9" ID-CoL="3V6TP" authority="(LINDROTH, 1963)" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[222,683,1710,1735]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B387EAE2FFC6720279F3F943FE0CC1F8" box="[222,472,1711,1734]" italics="true" pageId="38" pageNumber="1351">
<smallCapsWord id="87AAA02CFFC6720279F3F943FE87C1F8" baselines="1728,1729" box="[222,339,1711,1734]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lionepha" pageId="38" pageNumber="1351">LIONEPHA</smallCapsWord>
<smallCapsWord id="87AAA02CFFC672027877F95EFE0CC1F8" baselines="1729" box="[346,472,1714,1734]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="disjuncta" pageId="38" pageNumber="1351">DISJUNCTA</smallCapsWord>
</emphasis>
(
<bibRefCitation id="E5624B01FFC6720278CAF942FD77C1F9" author="Lindroth CH" box="[487,675,1710,1735]" pageId="38" pageNumber="1351" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">
<smallCapsWord id="87AAA02CFFC6720278CAF942FD8AC1F8" baselines="1729,1729" box="[487,606,1710,1735]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lindroth" pageId="38" pageNumber="1351">LINDROTH</smallCapsWord>
, 1963
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC672027836F931FDBBC1C8" box="[283,623,1757,1782]" pageId="38" pageNumber="1351" type="description">
<paragraph id="814C36F0FFC672027836F931FDBBC1C8" blockId="38.[283,623,1757,1782]" box="[283,623,1757,1782]" pageId="38" pageNumber="1351">
(
<figureCitation id="19C82A75FFC67202780EF931FE55C1C8" box="[291,385,1757,1782]" captionStart="Figure 2" captionStartId="2.[144,223,1422,1444]" captionTargetBox="[377,1176,231,1382]" captionTargetId="figure-203@2.[360,1209,195,1382]" captionTargetPageId="2" captionText="Figure 2. Adults of four species of the Lionepha erasa species group. A, L. casta, USA: Oregon: Benton Co., Marys Peak, Alder Creek Falls; B, L. kavanaughi, holotype, USA: Oregon: Wallowa Co., Lostine River Valley; B, L. lindrothi, holotype, USA: California: Tulare Co., outlet of Emerald Lake; D, L. disjuncta, Canada: British Columbia: Summit Creek 28 km E Kootenay Pass. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="38" pageNumber="1351">
<smallCapsWord id="87AAA02CFFC67202780EF931FE8DC1CA" baselines="1776,1775" box="[291,345,1757,1782]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="38" pageNumber="1351">FIGS</smallCapsWord>
2D
</figureCitation>
,
<figureCitation id="19C82A75FFC6720278A3F931FE0BC1C8" box="[398,479,1757,1782]" captionStart="Figure 16" captionStartId="24.[145,223,1304,1326]" captionTargetBox="[148,1421,198,1261]" captionTargetId="figure-261@24.[145,1425,195,1265]" captionTargetPageId="24" captionText="Figure 16. Microsculpture of the elytral disc of four species of the Lionepha erasa species group. Left elytron, region around anterior discal seta on third interval. A, L. casta male; B, L. casta female; C, L. kavanaughi, male; D, L. kavanaughi, female; E, L. lindrothi, male; F, L. lindrothi, female; G, L. disjuncta, male; H, L. disjuncta, female. Scale bar 50 µm." pageId="38" pageNumber="1351">16G, H</figureCitation>
,
<figureCitation id="19C82A75FFC6720278C1F931FDE9C1C8" box="[492,573,1757,1782]" captionStart="Figure 12" captionStartId="20.[145,226,1548,1570]" captionTargetBox="[152,1419,202,1505]" captionTargetId="figure-155@20.[145,1425,195,1508]" captionTargetPageId="20" captionText="Figure 12. Aedeagus of four species of the Lionepha erasa species group, left lateral view. A, L. casta; B, L. casta; C, L. kavanaughi, holotype; D, L. kavanaughi; E, L. lindrothi, holotype; F, L. lindrothi; G, L. disjuncta; H, L. disjuncta. Scale bar 0.1 mm." pageId="38" pageNumber="1351">12G, H</figureCitation>
,
<figureCitation id="19C82A75FFC672027B67F931FDB2C1C8" box="[586,614,1757,1782]" captionStart="Figure 23" captionStartId="39.[164,242,1675,1697]" captionTargetBox="[357,1252,195,1635]" captionTargetId="figure-116@39.[353,1252,195,1635]" captionTargetPageId="39" captionText="Figure 23. Geographic distribution of L. disjuncta. Specimens whose DNA was sequenced are shown in black." pageId="38" pageNumber="1351">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC6720279BCF8E9FDADC066" pageId="38" pageNumber="1351" type="reference_group">
<paragraph id="814C36F0FFC6720279BCF8E9FDADC066" blockId="38.[145,761,1797,1880]" pageId="38" pageNumber="1351">
<treatmentCitationGroup id="A1E311DEFFC6720279BCF8E9FDADC066" pageId="38" pageNumber="1351">
<treatmentCitation id="005210E1FFC6720279BCF8E9FD52C025" author="Lindroth CH" box="[145,646,1797,1819]" page="264" pageId="38" pageNumber="1351" year="1963">
<taxonomicName id="46F34D73FFC6720279BCF8E9FD52C025" ID-CoL="8LXBX" authority="Lindroth, 1963: 264" authorityName="Lindroth" authorityPageNumber="264" authorityYear="1963" box="[145,646,1797,1819]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="disjunctum">
<emphasis id="B387EAE2FFC6720279BCF8E9FE4CC024" box="[145,408,1797,1818]" italics="true" pageId="38" pageNumber="1351">Bembidion disjunctum</emphasis>
<materialsCitation id="319B3CADFFC67202788DF8E9FD52C025" box="[416,646,1797,1819]" collectionCode="MCZ" county="Tuolumne County" location="Sonora Pass" pageId="38" pageNumber="1351" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<bibRefCitation id="E5624B01FFC67202788DF8E9FD52C025" author="Lindroth CH" box="[416,646,1797,1819]" pageId="38" pageNumber="1351" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth, 1963: 264</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="5E488852FFC672027BBCF8E9FD2DC025" box="[657,761,1797,1819]" pageId="38" pageNumber="1351" type="holotype">Holotype</typeStatus>
<specimenCount id="97F5FD79FFC6720279BCF8CBFF75C007" box="[145,161,1831,1849]" count="1" pageId="38" pageNumber="1351" type="male"></specimenCount>
in MCZ (type # 32533), examined. Type locality:
<location id="842C602BFFC6720279BCF8AFFEC9C066" LSID="urn:lsid:plazi:treatment:095A87E6FFC6720379F3F943FE65C071:842C602BFFC6720279BCF8AFFEC9C066" box="[145,285,1859,1880]" county="Tuolumne County" name="Sonora Pass" pageId="38" pageNumber="1351" stateProvince="California">Sonora Pass</location>
,
<collectingCounty id="682D4E7CFFC67202780AF8AEFE20C066" box="[295,500,1858,1880]" pageId="38" pageNumber="1351">Tuolumne County</collectingCounty>
,
<collectingRegion id="4337F812FFC6720278D2F8AEFDA2C066" box="[511,630,1858,1880]" country="United States of America" name="California" pageId="38" pageNumber="1351">California</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC672037A04FF2AFE65C071" lastPageId="39" lastPageNumber="1352" pageId="38" pageNumber="1351" type="diagnosis">
<paragraph id="814C36F0FFC672027A04FF2AFB45C64A" blockId="38.[809,1425,197,832]" pageId="38" pageNumber="1351">
<emphasis id="B387EAE2FFC672027A04FF2AFC72C7E5" box="[809,934,198,219]" italics="true" pageId="38" pageNumber="1351">Diagnosis:</emphasis>
A flat, parallel-sided species with more complete striae than other
<taxonomicName id="46F34D73FFC672027D5EFF08FB36C7C7" authorityName="Casey" authorityYear="1918" box="[1139,1250,228,249]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC672027D5EFF08FB36C7C7" box="[1139,1250,228,249]" italics="true" pageId="38" pageNumber="1351">Lionepha</emphasis>
</taxonomicName>
(
<figureCitation id="19C82A75FFC672027DD9FF08FA98C7C7" box="[1268,1356,228,250]" captionStart="Figure 2" captionStartId="2.[144,223,1422,1444]" captionTargetBox="[377,1176,231,1382]" captionTargetId="figure-203@2.[360,1209,195,1382]" captionTargetPageId="2" captionText="Figure 2. Adults of four species of the Lionepha erasa species group. A, L. casta, USA: Oregon: Benton Co., Marys Peak, Alder Creek Falls; B, L. kavanaughi, holotype, USA: Oregon: Wallowa Co., Lostine River Valley; B, L. lindrothi, holotype, USA: California: Tulare Co., outlet of Emerald Lake; D, L. disjuncta, Canada: British Columbia: Summit Creek 28 km E Kootenay Pass. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="38" pageNumber="1351">Fig. 2D</figureCitation>
), and with relatively strong, slightly transverse elytral microsculpture (
<figureCitation id="19C82A75FFC672027AC4FECDFBBAC609" box="[1001,1134,289,311]" captionStart="Figure 16" captionStartId="24.[145,223,1304,1326]" captionTargetBox="[148,1421,198,1261]" captionTargetId="figure-261@24.[145,1425,195,1265]" captionTargetPageId="24" captionText="Figure 16. Microsculpture of the elytral disc of four species of the Lionepha erasa species group. Left elytron, region around anterior discal seta on third interval. A, L. casta male; B, L. casta female; C, L. kavanaughi, male; D, L. kavanaughi, female; E, L. lindrothi, male; F, L. lindrothi, female; G, L. disjuncta, male; H, L. disjuncta, female. Scale bar 50 µm." pageId="38" pageNumber="1351">Fig. 16G, H</figureCitation>
). Aedeagus (
<figureCitation id="19C82A75FFC672027C29FECDFA5DC609" box="[1284,1417,289,311]" captionStart="Figure 12" captionStartId="20.[145,226,1548,1570]" captionTargetBox="[152,1419,202,1505]" captionTargetId="figure-155@20.[145,1425,195,1508]" captionTargetPageId="20" captionText="Figure 12. Aedeagus of four species of the Lionepha erasa species group, left lateral view. A, L. casta; B, L. casta; C, L. kavanaughi, holotype; D, L. kavanaughi; E, L. lindrothi, holotype; F, L. lindrothi; G, L. disjuncta; H, L. disjuncta. Scale bar 0.1 mm." pageId="38" pageNumber="1351">Fig. 12G, H</figureCitation>
) similar to other members of the
<taxonomicName id="46F34D73FFC672027D86FEACFB3EC66B" baseAuthorityName="LeConte" baseAuthorityYear="1859" box="[1195,1258,320,341]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="erasa">
<emphasis id="B387EAE2FFC672027D86FEACFB3EC66B" box="[1195,1258,320,341]" italics="true" pageId="38" pageNumber="1351">erasa</emphasis>
</taxonomicName>
group, with a more or less ovoid
<collectingCountry id="F9E47660FFC672027AD1FEB3FBF3C64A" box="[1020,1063,351,372]" name="Switzerland" pageId="38" pageNumber="1351">CH</collectingCountry>
1 sclerite.
</paragraph>
<paragraph id="814C36F0FFC672027A6CFE92FB67C47E" blockId="38.[809,1425,197,832]" pageId="38" pageNumber="1351">
In appearance
<taxonomicName id="46F34D73FFC672027D29FE92FB4AC6AC" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[1028,1182,381,403]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B387EAE2FFC672027D29FE92FB4AC6AC" box="[1028,1182,381,403]" italics="true" pageId="38" pageNumber="1351">L. disjuncta</emphasis>
</taxonomicName>
resembles a small member of
<taxonomicName id="46F34D73FFC672027AECFE70FB98C68F" authorityName="Latreille" authorityYear="1802" box="[961,1100,412,433]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC672027AECFE70FB98C68F" box="[961,1100,412,433]" italics="true" pageId="38" pageNumber="1351">Bembidion</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="46F34D73FFC672027DF2FE70FAB7C68F" box="[1247,1379,412,433]" pageId="38" pageNumber="1351" rank="subGenus" subGenus="Plataphus">
<emphasis id="B387EAE2FFC672027DF2FE70FAB7C68F" box="[1247,1379,412,433]" italics="true" pageId="38" pageNumber="1351">Plataphus</emphasis>
</taxonomicName>
, or a
<taxonomicName id="46F34D73FFC672027A6EFE56FAEAC6EE" authority="LeConte, 1863" authorityName="LeConte" authorityYear="1863" box="[835,1342,442,464]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="nebraskense">
<emphasis id="B387EAE2FFC672027A6EFE56FBA7C6F1" box="[835,1139,442,463]" italics="true" pageId="38" pageNumber="1351">Bembidion nebraskense</emphasis>
LeConte, 1863
</taxonomicName>
. From sympatric
<taxonomicName id="46F34D73FFC672027A98FE35FB35C6D0" authorityName="Motschulsky" authorityYear="1864" box="[949,1249,473,494]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="subGenus" subGenus="Plataphus">
<emphasis id="B387EAE2FFC672027A98FE35FBEAC6D0" box="[949,1086,473,494]" italics="true" pageId="38" pageNumber="1351">Bembidion</emphasis>
(
<emphasis id="B387EAE2FFC672027D78FE35FB0CC6D0" box="[1109,1240,473,494]" italics="true" pageId="38" pageNumber="1351">Plataphus</emphasis>
)
</taxonomicName>
specimens it can be distinguished externally by the completely bordered metasternal process (bordered at the sides only in
<taxonomicName id="46F34D73FFC672027AE5FDD9FB96C574" box="[968,1090,565,586]" pageId="38" pageNumber="1351" rank="subGenus" subGenus="Plataphus">
<emphasis id="B387EAE2FFC672027AE5FDD9FB96C574" box="[968,1090,565,586]" italics="true" pageId="38" pageNumber="1351">Plataphus</emphasis>
</taxonomicName>
). The elytral striae are less distinct in
<taxonomicName id="46F34D73FFC672027A96FDB8FB82C557" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[955,1110,596,617]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B387EAE2FFC672027A96FDB8FB82C557" box="[955,1110,596,617]" italics="true" pageId="38" pageNumber="1351">L. disjuncta</emphasis>
</taxonomicName>
. In
<taxonomicName id="46F34D73FFC672027DBDFDB8FAC6C557" box="[1168,1298,596,617]" pageId="38" pageNumber="1351" rank="subGenus" subGenus="Plataphus">
<emphasis id="B387EAE2FFC672027DBDFDB8FAC6C557" box="[1168,1298,596,617]" italics="true" pageId="38" pageNumber="1351">Plataphus</emphasis>
</taxonomicName>
, multiple striae will be easily evident near the elytral apex. The aedeagus is different from
<taxonomicName id="46F34D73FFC672027D85FD7DFAF7C598" box="[1192,1315,657,678]" pageId="38" pageNumber="1351" rank="subGenus" subGenus="Plataphus">
<emphasis id="B387EAE2FFC672027D85FD7DFAF7C598" box="[1192,1315,657,678]" italics="true" pageId="38" pageNumber="1351">Plataphus</emphasis>
</taxonomicName>
, without the evident flagellum present in members of that subgenus. From
<taxonomicName id="46F34D73FFC672027D28FD22FAECC5DD" authorityName="LeConte" authorityYear="1863" box="[1029,1336,718,739]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="nebraskense">
<emphasis id="B387EAE2FFC672027D28FD22FAECC5DD" box="[1029,1336,718,739]" italics="true" pageId="38" pageNumber="1351">Bembidion nebraskense</emphasis>
</taxonomicName>
it can be immediately distinguished by the presence of elytral microsculpture:
<taxonomicName id="46F34D73FFC672027D69FCE0FB27C41F" authorityName="LeConte" authorityYear="1863" box="[1092,1267,780,801]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="nebraskense">
<emphasis id="B387EAE2FFC672027D69FCE0FB27C41F" box="[1092,1267,780,801]" italics="true" pageId="38" pageNumber="1351">B. nebraskense</emphasis>
</taxonomicName>
lacks elytral microsculpture and is thus glossy.
</paragraph>
<paragraph id="814C36F0FFC672027A04FC85FB31C4E4" blockId="38.[809,1425,873,987]" pageId="38" pageNumber="1351">
<emphasis id="B387EAE2FFC672027A04FC85FB87C440" box="[809,1107,873,894]" italics="true" pageId="38" pageNumber="1351">Additional characteristics:</emphasis>
Body length
<quantity id="460B9B15FFC672027DC1FC85FA59C441" box="[1260,1421,873,895]" metricMagnitude="-3" metricUnit="m" metricValue="3.97" metricValueMax="4.32" metricValueMin="3.62" pageId="38" pageNumber="1351" unit="mm" value="3.97" valueMax="4.32" valueMin="3.62">3.624.32 mm</quantity>
. Antenna piceous. Femora rufopiceous or piceous, tibiae rufopiceous or rufous. Hind wings full-sized. Chromosomes of male 24 + X (
<tableCitation id="CC71034BFFC672027DAFFC29FB01C4E5" box="[1154,1237,965,987]" captionStart="Table 5" captionStartId="29.[162,226,196,217]" captionText="Table 5. Chromosome numbers and sex chromosomes of Lionepha males. The Sample column indicates the number of males examined" pageId="38" pageNumber="1351">Table 5</tableCitation>
).
</paragraph>
<paragraph id="814C36F0FFC672027A04FBE8FC6FC255" blockId="38.[809,1424,1028,1387]" pageId="38" pageNumber="1351">
<emphasis id="B387EAE2FFC672027A04FBE8FBE2C327" box="[809,1078,1028,1049]" italics="true" pageId="38" pageNumber="1351">Geographic variation:</emphasis>
This species shows notable variation in 28S, with numerous insertion and deletion events evident (Supporting Information,
<figureCitation id="19C82A75FFC672027A04FB8CFC56C34B" box="[809,898,1120,1142]" captionStart="Figure 1" captionStartId="1.[162,240,1424,1446]" captionTargetBox="[408,1210,236,1367]" captionTargetId="figure-189@1.[395,1210,195,1384]" captionTargetPageId="1" captionText="Figure 1. Adults of three species of the Lionepha erasa species group.A, L. erasa, USA: Oregon: Benton Co., Marys Peak; B, L. australerasa, holotype, USA: California: Amador Co., Oyster Lake, Silver Lake Campground; C, L. probata, USA: Oregon: Klamath Co., Odell Creek near Davis Lake. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="38" pageNumber="1351">Fig. S1</figureCitation>
). Of the ten specimens sequenced, seven have unique indels, and the remaining three share an eighth pattern of insertions and deletions. In addition, the specimen from the Wallowas that was sequenced has a unique amino acid, threonine, within
<emphasis id="B387EAE2FFC672027AA6FB15FC6AC230" box="[907,958,1273,1294]" italics="true" pageId="38" pageNumber="1351">COI</emphasis>
, although other
<taxonomicName id="46F34D73FFC672027D8AFB16FA93C230" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[1191,1351,1273,1295]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B387EAE2FFC672027D8AFB16FA93C230" box="[1191,1351,1273,1295]" italics="true" pageId="38" pageNumber="1351">L. disjuncta</emphasis>
</taxonomicName>
have alanine at that position (base
<quantity id="460B9B15FFC672027DB0FAF4FB3EC210" box="[1181,1258,1304,1326]" metricMagnitude="0" metricUnit="m" metricValue="9.0424" pageId="38" pageNumber="1351" unit="in" value="356.0">356 in</quantity>
the sequence of specimen DNA3848, GenBank accession number
<accessionNumber id="9EA0AB13FFC672027A04FAB9FC7BC255" box="[809,943,1365,1387]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN402196" pageId="38" pageNumber="1351">MN402196</accessionNumber>
).
</paragraph>
<paragraph id="814C36F0FFC672027A04FA78FAAEC15C" blockId="38.[809,1425,1428,1634]" pageId="38" pageNumber="1351">
<emphasis id="B387EAE2FFC672027A04FA78FC6EC297" box="[809,954,1428,1449]" italics="true" pageId="38" pageNumber="1351">Distribution:</emphasis>
A widespread species, from the northern Sierra
<collectingRegion id="4337F812FFC672027A5BFA5EFC19C2F6" box="[886,973,1458,1480]" country="United States of America" name="Nevada" pageId="38" pageNumber="1351">Nevada</collectingRegion>
and
<taxonomicName id="46F34D73FFC672027D29FA5EFB58C2F6" authority="Alps" authorityName="Alps" box="[1028,1164,1458,1480]" class="Arachnida" family="Hubbardiidae" genus="Trinity" higherTaxonomySource="CoL" kingdom="Animalia" order="Schizomida" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="genus">Trinity Alps</taxonomicName>
of northern
<collectingRegion id="4337F812FFC672027C34FA5EFA59C2F6" box="[1305,1421,1458,1480]" country="United States of America" name="California" pageId="38" pageNumber="1351">California</collectingRegion>
, north to southern
<collectingRegion id="4337F812FFC672027D35FA3DFB21C2D9" box="[1048,1269,1489,1511]" country="Canada" name="British Columbia" pageId="38" pageNumber="1351">British Columbia</collectingRegion>
and east to
<collectingRegion id="4337F812FFC672027A04FA1CFC47C13B" box="[809,915,1520,1541]" country="United States of America" name="Montana" pageId="38" pageNumber="1351">Montana</collectingRegion>
(
<figureCitation id="19C82A75FFC672027A88FA1CFC23C13B" box="[933,1015,1520,1542]" captionStart="Figure 23" captionStartId="39.[164,242,1675,1697]" captionTargetBox="[357,1252,195,1635]" captionTargetId="figure-116@39.[353,1252,195,1635]" captionTargetPageId="39" captionText="Figure 23. Geographic distribution of L. disjuncta. Specimens whose DNA was sequenced are shown in black." pageId="38" pageNumber="1351">Fig. 23</figureCitation>
). Found between 650 and
<quantity id="460B9B15FFC672027C15FA1CFA5BC13B" box="[1336,1423,1520,1541]" metricMagnitude="3" metricUnit="m" metricValue="2.93" pageId="38" pageNumber="1351" unit="m" value="2930.0">2930 m</quantity>
in elevation. Most records are from late summer and early fall, with specimens having been collected in June through September, as well as one record from April.
</paragraph>
<paragraph id="814C36F0FFC672037A04F966FE65C071" blockId="38.[809,1425,1674,1880]" lastBlockId="39.[163,778,1757,1871]" lastPageId="39" lastPageNumber="1352" pageId="38" pageNumber="1351">
<emphasis id="B387EAE2FFC672027A04F966FC5BC1A1" box="[809,911,1674,1695]" italics="true" pageId="38" pageNumber="1351">Habitat:</emphasis>
This species is usually encountered as a specimen or two amongst more common
<taxonomicName id="46F34D73FFC672027C3CF945FA45C180" authorityName="Latreille" authorityYear="1802" box="[1297,1425,1705,1726]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1351" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC672027C3CF945FA45C180" box="[1297,1425,1705,1726]" italics="true" pageId="38" pageNumber="1351">Bembidion</emphasis>
</taxonomicName>
on gravel and cobble shores of cold, clear creeks and rivers. There are two localities where numerous specimens have been found: Summit Creek west of Creston,
<collectingRegion id="4337F812FFC672027AB9F8C8FBB6C004" box="[916,1122,1828,1850]" country="Canada" name="British Columbia" pageId="38" pageNumber="1351">British Columbia</collectingRegion>
, and in the
<taxonomicName id="46F34D73FFC672027DDCF8C8FCB9C066" authority="River valley" authorityName="River Valley" authorityYear="1526" class="Insecta" family="Leiodidae" genus="Lostine" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1349" phylum="Arthropoda" rank="genus">Lostine River valley</taxonomicName>
in the Wallowas of
<collectingRegion id="4337F812FFC672027D65F8AEFB4AC069" box="[1096,1182,1858,1879]" country="United States of America" name="Oregon" pageId="38" pageNumber="1351">Oregon</collectingRegion>
. At the latter locality,
<specimenCount id="97F5FD79FFC77203798EF931FEE9C1CD" box="[163,317,1757,1779]" count="36" pageId="39" pageNumber="1352" type="generic">36 specimens</specimenCount>
were found in the drying bed of a small creek in a forest (
<figureCitation id="19C82A75FFC7720378A5F910FE37C02F" box="[392,483,1788,1810]" captionStart="Figure 4" captionStartId="4.[145,225,1687,1709]" captionTargetBox="[146,1428,198,1647]" captionTargetId="figure-27@4.[145,1428,195,1647]" captionTargetPageId="4" captionText="Figure 4. Habitats of Lionepha. Arrows indicate a typical microhabitat. A, habitat of L. tuulukwa and L. casta. USA: Oregon: Benton Co., Marys Peak Road, Alder Creek Falls, 700m, 44.4746°N 123.5286°W, 24 September 2010. B, habitat of L.tuulukwa and L. casta. Lionepha osculans has been found a few kilometres upstream along this same creek. USA: Oregon: Lane Co., Knowles Creek, 6.8 km SE Mapleton, 44.0136°N 123.7880°W, 9 August 2013. C, Environment of L. lindrothi. The beetles were found on dark organic shorelines of a partly shaded outlet stream of this lake, approximately at the point indicated by the arrow. USA: California: Tulare Co., outlet of Emerald Lake, 2814 m, 36.599°N 118.677°W, 21 June 2014." pageId="39" pageNumber="1352">Fig. 4D</figureCitation>
), along with numerous
<taxonomicName id="46F34D73FFC77203798EF8F6FE60C011" box="[163,436,1818,1839]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="1352" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="B387EAE2FFC77203798EF8F6FE60C011" box="[163,436,1818,1839]" italics="true" pageId="39" pageNumber="1352">Lionepha kavanaughi</emphasis>
</taxonomicName>
and
<taxonomicName id="46F34D73FFC7720378D5F8F7FDAEC011" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[504,634,1818,1840]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="1352" phylum="Arthropoda" rank="species" species="probata">
<emphasis id="B387EAE2FFC7720378D5F8F7FDAEC011" box="[504,634,1818,1840]" italics="true" pageId="39" pageNumber="1352">L. probata</emphasis>
</taxonomicName>
, as well as
<taxonomicName id="46F34D73FFC77203798EF8D5FE7EC070" box="[163,426,1849,1870]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="1352" phylum="Arthropoda" rank="species" species="kuprianovii">
<emphasis id="B387EAE2FFC77203798EF8D5FE7EC070" box="[163,426,1849,1870]" italics="true" pageId="39" pageNumber="1352">Bembidion kuprianovii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,221 @@
<document id="D9A869A5A6082D1BE7800A9DD52D33AA" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFC7720C7A8FF931FDE7C019" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha osculans DNA 2638" docType="treatment" docVersion="1" lastPageNumber="1353" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1352" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="2E57AB57187F7C39E44EC49C636D18DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B0DC79B628D7027B8AE076A66D68DF70">
<mods:title id="E07E92CB964727A4163D168867DF1119">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="7D0618E45C130A46D7F35FFE3BAA8582" type="personal">
<mods:role id="B3C95B467550D882A4862A42125D19A3">
<mods:roleTerm id="E9C78D4A07FD5A89A3B918806FC0D071">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F2FF37475B3573EDB8FE2D76210F872">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="67D6BC3481A6738B0042C7C3184A2DE4" type="personal">
<mods:role id="9B5AF06754BBED2B6E7E10221AF2955C">
<mods:roleTerm id="2F0CA5CEA9D01A17B8A9F21CB9197725">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E2E6E3FBD599E5CBDB0989ABF89677BC">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0A147A83AEA69E0CFDFE0F8F7DA90627">text</mods:typeOfResource>
<mods:relatedItem id="7ED75D65936640ECA8BCF1E06B8B3D1B" type="host">
<mods:titleInfo id="C0DDFAFBED4E26336A27F39834A36014">
<mods:title id="B43C64E499764E7448BB02B6AE4E51E3">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="36B813BB9044DA58BDB906E38185DE5D">
<mods:date id="28098391FF072D069AEA84B03ED27512">2020</mods:date>
<mods:detail id="912FE56CC79204A4FDBE93E0F14CFF55" type="volume">
<mods:number id="4E28A85A033E64575D945148099CCFEF">189</mods:number>
</mods:detail>
<mods:extent id="FC88AAC6B7D9833FD673A741DF3C4743" unit="page">
<mods:start id="F8BB7B20667914570C8101BAD373AAC7">1313</mods:start>
<mods:end id="E3072D1A7D2C233567FB455656BDDB06">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7C8DEEF1FDFA7214F4842B36AEA507AC">journal article</mods:classification>
<mods:identifier id="ABD9B88BDEC2BD9FE203F5505F433D33" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="1FC52471BEDA8B688075497588DBAC3D" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFC7720C7A8FF931FDE7C019" LSID="urn:lsid:plazi:treatment:095A87E6FFC7720C7A8FF931FDE7C019" httpUri="http://treatment.plazi.org/id/095A87E6FFC7720C7A8FF931FDE7C019" lastPageId="40" lastPageNumber="1353" pageId="39" pageNumber="1352">
<subSubSection id="C9E9657BFFC772037A8FF931FAEFC1CB" box="[930,1339,1756,1781]" pageId="39" pageNumber="1352" type="nomenclature">
<paragraph id="814C36F0FFC772037A8FF931FAEFC1CB" blockId="39.[930,1339,1756,1781]" box="[930,1339,1756,1781]" pageId="39" pageNumber="1352">
<heading id="DA04819CFFC772037A8FF931FAEFC1CB" box="[930,1339,1756,1781]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="1352" reason="2">
<taxonomicName id="46F34D73FFC772037A8FF931FAEFC1CB" ID-CoL="3V6TW" authority="(CASEY, 1918)" authorityName="DNA" authorityYear="2638" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[930,1339,1756,1781]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="1352" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC772037A8FF931FB47C1CD" box="[930,1171,1757,1780]" italics="true" pageId="39" pageNumber="1352">
<smallCapsWord id="87AAA02CFFC772037A8FF931FBC3C1CD" baselines="1774,1774" box="[930,1047,1757,1780]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lionepha" pageId="39" pageNumber="1352">LIONEPHA</smallCapsWord>
<smallCapsWord id="87AAA02CFFC772037D30F933FB47C1CD" baselines="1774" box="[1053,1171,1759,1779]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="osculans" pageId="39" pageNumber="1352">OSCULANS</smallCapsWord>
</emphasis>
(
<bibRefCitation id="E5624B01FFC772037D8EF930FAE7C1CB" author="Casey TL" box="[1187,1331,1756,1781]" pageId="39" pageNumber="1352" pagination="1 - 223" refId="ref26056" refString="Casey TL. 1918. A review of the North American Bembidiinae. Memoirs on the Coleoptera 8: 1-223." type="journal article" year="1918">
<smallCapsWord id="87AAA02CFFC772037D8EF930FB39C1CD" baselines="1775,1774" box="[1187,1261,1756,1781]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Casey" pageId="39" pageNumber="1352">CASEY</smallCapsWord>
, 1918
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC772037A8FF8E6FAEFC01D" box="[930,1339,1802,1827]" pageId="39" pageNumber="1352" type="description">
<paragraph id="814C36F0FFC772037A8FF8E6FAEFC01D" blockId="39.[930,1339,1802,1827]" box="[930,1339,1802,1827]" pageId="39" pageNumber="1352">
(
<figureCitation id="19C82A75FFC772037A87F8E6FBD3C01D" box="[938,1031,1802,1827]" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="39" pageNumber="1352">
<smallCapsWord id="87AAA02CFFC772037A87F8E6FC34C01C" baselines="1821,1821" box="[938,992,1802,1827]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="39" pageNumber="1352">FIGS</smallCapsWord>
3A
</figureCitation>
,
<figureCitation id="19C82A75FFC772037D39F8E6FBB5C01D" box="[1044,1121,1802,1827]" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="39" pageNumber="1352">17A, B</figureCitation>
,
<figureCitation id="19C82A75FFC772037D43F8E6FB68C01D" box="[1134,1212,1802,1827]" captionStart="Figure 13" captionStartId="21.[163,241,1563,1585]" captionTargetBox="[173,1430,204,1518]" captionTargetId="figure-171@21.[163,1443,195,1523]" captionTargetPageId="21" captionText="Figure 13. Aedeagus of the Lionepha osculans species group, left lateral view. A, L. osculans; B, L. osculans; C, L. sequoiae; D, L. sequoiae; E, L. pseudoerasa; F, L. pseudoerasa; G, L. tuulukwa; H, L. tuulukwa. Scale bar 0.1 mm." pageId="39" pageNumber="1352">13A, B</figureCitation>
;
<figureCitation id="19C82A75FFC772037DE4F8E6FB2CC01D" box="[1225,1272,1802,1827]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="39" pageNumber="1352">19A</figureCitation>
,
<figureCitation id="19C82A75FFC772037C28F8E6FAE7C01D" box="[1285,1331,1802,1827]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="39" pageNumber="1352">24A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC7720C7A16F8DFFD7CC64A" lastPageId="40" lastPageNumber="1353" pageId="39" pageNumber="1352" type="reference_group">
<paragraph id="814C36F0FFC7720C7A16F8DFFD56C7C7" blockId="39.[827,1444,1843,1896]" lastBlockId="40.[145,762,197,372]" lastPageId="40" lastPageNumber="1353" pageId="39" pageNumber="1352">
<treatmentCitationGroup id="A1E311DEFFC7720C7A16F8DFFD56C7C7" lastPageId="40" lastPageNumber="1353" pageId="39" pageNumber="1352">
<treatmentCitation id="005210E1FFC772037A16F8DFFB2AC076" author="Casey TL" box="[827,1278,1843,1864]" page="20" pageId="39" pageNumber="1352" year="1918">
<taxonomicName id="46F34D73FFC772037A16F8DFFB2AC076" ID-CoL="8LXBZ" authority="Casey, 1918: 20" authorityName="Casey" authorityPageNumber="20" authorityYear="1918" box="[827,1278,1843,1864]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="1352" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC772037A16F8DFFBE5C076" box="[827,1073,1843,1864]" italics="true" pageId="39" pageNumber="1352">Bembidion osculans</emphasis>
<bibRefCitation id="E5624B01FFC772037D11F8DFFB2AC076" author="Casey TL" box="[1084,1278,1843,1864]" pageId="39" pageNumber="1352" pagination="1 - 223" refId="ref26056" refString="Casey TL. 1918. A review of the North American Bembidiinae. Memoirs on the Coleoptera 8: 1-223." type="journal article" year="1918">Casey, 1918: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="5E488852FFC772037C26F8DFFA56C076" box="[1291,1410,1843,1864]" pageId="39" pageNumber="1352" type="lectotype">Lectotype</typeStatus>
<specimenCount id="97F5FD79FFC772037CA0F8DAFA48C076" box="[1421,1436,1846,1864]" count="1" pageId="39" pageNumber="1352" type="female"></specimenCount>
, designated by
<treatmentCitation id="005210E1FFC772037AC4F8BEFB34C056" author="Lindroth CH" box="[1001,1248,1873,1896]" page="116" pageId="39" pageNumber="1352" year="1975">
<bibRefCitation id="E5624B01FFC772037AC4F8BEFB34C056" author="Lindroth CH" box="[1001,1248,1873,1896]" pageId="39" pageNumber="1352" pagination="109 - 147" refId="ref26559" refString="Lindroth CH. 1975. Designation of holotypes and lectotypes among ground beetles (Coleoptera, Carabidae) described by Thomas L. Casey. The Coleopterists Bulletin 29: 109-147." type="journal article" year="1975">Lindroth (1975: 116)</bibRefCitation>
</treatmentCitation>
, in USNM (type # 36816), examined. Type locality: Marin County,
<collectingRegion id="4337F812FFC8720C79BCFF08FED2C7C4" box="[145,262,228,250]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
, as restricted by
<bibRefCitation id="E5624B01FFC8720C78EAFF08FDAFC7C4" author="Lindroth CH" box="[455,635,228,250]" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth (1963)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="814C36F0FFC8720C7984FEEEFD7CC64A" blockId="40.[145,762,197,372]" pageId="40" pageNumber="1353">
<treatmentCitationGroup id="A1E311DEFFC8720C7984FEEEFD7CC64A" pageId="40" pageNumber="1353">
<treatmentCitation id="005210E1FFC8720C7984FEEEFD8AC626" author="Casey TL" box="[169,606,258,280]" page="20" pageId="40" pageNumber="1353" year="1918">
<taxonomicName id="46F34D73FFC8720C7984FEEEFD8AC626" ID-CoL="5WJ9X" authority="Casey, 1918: 20" authorityName="Casey" authorityPageNumber="20" authorityYear="1918" box="[169,606,258,280]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="speculum">
<emphasis id="B387EAE2FFC8720C7984FEEEFE4AC629" box="[169,414,258,279]" italics="true" pageId="40" pageNumber="1353">Bembidion speculum</emphasis>
<bibRefCitation id="E5624B01FFC8720C788BFEEFFD8AC626" author="Casey TL" box="[422,606,259,280]" pageId="40" pageNumber="1353" pagination="1 - 223" refId="ref26056" refString="Casey TL. 1918. A review of the North American Bembidiinae. Memoirs on the Coleoptera 8: 1-223." type="journal article" year="1918">Casey, 1918: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="5E488852FFC8720C7B44FEEFFD0FC626" box="[617,731,259,280]" pageId="40" pageNumber="1353" type="lectotype">Lectotype</typeStatus>
<specimenCount id="97F5FD79FFC8720C7BC9FEE9FD27C629" box="[740,755,261,279]" count="1" pageId="40" pageNumber="1353" type="female"></specimenCount>
, designated by
<treatmentCitation id="005210E1FFC8720C7815FECDFDF3C609" author="Lindroth CH" box="[312,551,289,311]" page="116" pageId="40" pageNumber="1353" year="1975">
<bibRefCitation id="E5624B01FFC8720C7815FECDFDF3C609" author="Lindroth CH" box="[312,551,289,311]" pageId="40" pageNumber="1353" pagination="109 - 147" refId="ref26559" refString="Lindroth CH. 1975. Designation of holotypes and lectotypes among ground beetles (Coleoptera, Carabidae) described by Thomas L. Casey. The Coleopterists Bulletin 29: 109-147." type="journal article" year="1975">Lindroth (1975: 116)</bibRefCitation>
</treatmentCitation>
, in USNM (type # 36815), examined. Synonymy established by
<bibRefCitation id="E5624B01FFC8720C7BBEFEACFF0CC64A" author="Lindroth CH" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth (1963)</bibRefCitation>
. Type locality: Marin County,
<collectingRegion id="4337F812FFC8720C7B00FEB2FD70C64A" box="[557,676,350,372]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C79BCFE7FFDE7C019" pageId="40" pageNumber="1353" type="diagnosis">
<paragraph id="814C36F0FFC8720C79BCFE7FFE05C426" blockId="40.[145,762,402,792]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCFE7FFED2C696" box="[145,262,403,424]" italics="true" pageId="40" pageNumber="1353">Diagnosis:</emphasis>
This is the largest species of
<taxonomicName id="46F34D73FFC8720C7B7CFE7EFD6DC699" authorityName="Casey" authorityYear="1918" box="[593,697,402,423]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC8720C7B7CFE7EFD6DC699" box="[593,697,402,423]" italics="true" pageId="40" pageNumber="1353">Lionepha</emphasis>
</taxonomicName>
, with some females reaching nearly
<quantity id="460B9B15FFC8720C78D2FE5DFD96C6F9" box="[511,578,433,455]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="40" pageNumber="1353" unit="mm" value="6.0">6 mm</quantity>
in length. It is also the broadest, with a wide, rounded prothorax (
<figureCitation id="19C82A75FFC8720C7BE2FE3CFF64C53A" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="40" pageNumber="1353">Fig. 3A</figureCitation>
). The elytral microsculpture is more transversely stretched than other
<taxonomicName id="46F34D73FFC8720C78A5FDE1FE20C51C" authorityName="Casey" authorityYear="1918" box="[392,500,525,546]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC8720C78A5FDE1FE20C51C" box="[392,500,525,546]" italics="true" pageId="40" pageNumber="1353">Lionepha</emphasis>
</taxonomicName>
(
<figureCitation id="19C82A75FFC8720C7B29FDE1FD51C51D" box="[516,645,525,547]" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="40" pageNumber="1353">Fig. 17A, B</figureCitation>
), yielding a notable iridescence, especially in the males. Elytra with at least the third stria visible beyond the posterior dorsal puncture and the second stria almost reaching the elytral apex. It tends to be slightly darker, with darker legs, than other members of the
<taxonomicName id="46F34D73FFC8720C7B5EFD4BFD2DC585" authorityName="Casey" authorityYear="1918" box="[627,761,678,700]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC8720C7B5EFD4BFD2DC585" box="[627,761,678,700]" italics="true" pageId="40" pageNumber="1353">L. osculans</emphasis>
</taxonomicName>
group, but some specimens are paler. Aedeagus deep, with a bulbous appearance because of the sinuate ventral surface (
<figureCitation id="19C82A75FFC8720C7865FCEEFE17C426" box="[328,451,770,792]" captionStart="Figure 13" captionStartId="21.[163,241,1563,1585]" captionTargetBox="[173,1430,204,1518]" captionTargetId="figure-171@21.[163,1443,195,1523]" captionTargetPageId="21" captionText="Figure 13. Aedeagus of the Lionepha osculans species group, left lateral view. A, L. osculans; B, L. osculans; C, L. sequoiae; D, L. sequoiae; E, L. pseudoerasa; F, L. pseudoerasa; G, L. tuulukwa; H, L. tuulukwa. Scale bar 0.1 mm." pageId="40" pageNumber="1353">Fig. 13A, B</figureCitation>
).
</paragraph>
<paragraph id="814C36F0FFC8720C79BCFCADFD99C4CE" blockId="40.[145,761,833,1008]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCFCADFE20C468" box="[145,500,833,854]" italics="true" pageId="40" pageNumber="1353">Additional characteristics:</emphasis>
Body length
<quantity id="460B9B15FFC8720C7B9EFCADFF2DC44B" metricMagnitude="-3" metricUnit="m" metricValue="5.29" metricValueMax="5.9" metricValueMin="4.68" pageId="40" pageNumber="1353" unit="mm" value="5.29" valueMax="5.9" valueMin="4.68">4.68 5.90 mm</quantity>
. Antennae dark, piceous, although the first antennomere can be dark rufous on the underside. Legs in most specimens rufopiceous, occasionally rufous, darker at the joints. Hind wings full-sized. Chromosomes of male 24 + X (
<tableCitation id="CC71034BFFC8720C78C7FC36FDEAC4CE" box="[490,574,986,1008]" captionStart="Table 5" captionStartId="29.[162,226,196,217]" captionText="Table 5. Chromosome numbers and sex chromosomes of Lionepha males. The Sample column indicates the number of males examined" pageId="40" pageNumber="1353">Table 5</tableCitation>
).
</paragraph>
<paragraph id="814C36F0FFC8720C79BCFBF5FD57C3B5" blockId="40.[145,761,1049,1163]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCFBF5FE46C311" box="[145,402,1049,1071]" italics="true" pageId="40" pageNumber="1353">Geographic variation:</emphasis>
<materialsCitation id="319B3CADFFC8720C78B2FBF5FDABC3B5" location="Oregon" pageId="40" pageNumber="1353" specimenCount="6" stateProvince="Oregon">
The
<specimenCount id="97F5FD79FFC8720C78F9FBF5FDA3C311" box="[468,631,1049,1071]" count="6" pageId="40" pageNumber="1353" type="generic">six specimens</specimenCount>
sequenced from
<collectingRegion id="4337F812FFC8720C79E2FBD4FEF7C373" box="[207,291,1080,1101]" country="United States of America" name="Oregon" pageId="40" pageNumber="1353">Oregon</collectingRegion>
have three unique bases within the 28S gene; this is evident in the 28S tree in
<figureCitation id="19C82A75FFC8720C7B6BFBBAFD73C352" box="[582,679,1110,1132]" captionStart="Figure 5" captionStartId="13.[163,243,1682,1704]" captionTargetBox="[284,1321,197,1642]" captionTargetId="graphics-1213@13.[783,1129,198,1636]" captionTargetPageId="13" captionText="Figure 5. Maximum likelihood trees for 28S and COI. Branch length is shown proportional to relative divergence, as estimated by IQ-TREE; scale bars indicate 0.01 units. Outgroups not depicted." pageId="40" pageNumber="1353">Figure 5</figureCitation>
, where the
<collectingRegion id="4337F812FFC8720C7990FB99FEC4C3B4" box="[189,272,1141,1162]" country="United States of America" name="Oregon" pageId="40" pageNumber="1353">Oregon</collectingRegion>
specimens form a distinct clade
</materialsCitation>
.
</paragraph>
<paragraph id="814C36F0FFC8720C79BCFB43FDCAC200" blockId="40.[145,761,1198,1343]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCFB43FF18C3FA" box="[145,204,1199,1220]" italics="true" pageId="40" pageNumber="1353">Note:</emphasis>
<bibRefCitation id="E5624B01FFC8720C79F5FB42FEA2C3FD" author="Hering D" box="[216,374,1198,1220]" pageId="40" pageNumber="1353" pagination="161 - 170" refId="ref26330" refString="Hering D. 1998. Riparian beetles (Coleoptera) along a small stream in the Oregon Coast Range and their interactions with the aquatic environment. Coleopterists Bulletin 52: 161-170." type="journal article" year="1998">Hering (1998)</bibRefCitation>
reported on the food consumed by
<taxonomicName id="46F34D73FFC8720C79BBFB21FEAEC3DC" authorityName="Casey" authorityYear="1918" box="[150,378,1229,1250]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC8720C79BBFB21FEAEC3DC" box="[150,378,1229,1250]" italics="true" pageId="40" pageNumber="1353">Bembidion osculans</emphasis>
</taxonomicName>
on Knowles Creek in
<collectingRegion id="4337F812FFC8720C7B54FB21FD1AC3DC" box="[633,718,1229,1250]" country="United States of America" name="Oregon" pageId="40" pageNumber="1353">Oregon</collectingRegion>
; we examined a selection of specimens from his study, and the majority belong to
<taxonomicName id="46F34D73FFC8720C78B9FAE6FDA7C221" box="[404,627,1290,1311]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="tuulukwa">
<emphasis id="B387EAE2FFC8720C78B9FAE6FDA7C221" box="[404,627,1290,1311]" italics="true" pageId="40" pageNumber="1353">Lionepha tuulukwa</emphasis>
</taxonomicName>
, with a few specimens belonging to
<taxonomicName id="46F34D73FFC8720C78B2FAC6FDCAC200" authorityName="Casey" authorityYear="1918" box="[415,542,1321,1343]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC8720C78B2FAC6FDCAC200" box="[415,542,1321,1343]" italics="true" pageId="40" pageNumber="1353">L. osculans</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="814C36F0FFC8720C79BCFA84FE44C14C" blockId="40.[145,761,1384,1651]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCFA84FEFDC243" box="[145,297,1384,1405]" italics="true" pageId="40" pageNumber="1353">Distribution:</emphasis>
A widespread species, common in the Sierra
<collectingRegion id="4337F812FFC8720C79CFFA6AFEEFC2A2" box="[226,315,1414,1436]" country="United States of America" name="Nevada" pageId="40" pageNumber="1353">Nevada</collectingRegion>
and coastal areas of
<collectingRegion id="4337F812FFC8720C7B19FA6AFD78C2A2" box="[564,684,1414,1436]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
, north to
<collectingRegion id="4337F812FFC8720C7981FA49FEE0C285" box="[172,308,1445,1467]" country="United States of America" name="Washington" pageId="40" pageNumber="1353">Washington</collectingRegion>
and
<collectingRegion id="4337F812FFC8720C7844FA49FE7EC285" box="[361,426,1445,1467]" country="United States of America" name="Idaho" pageId="40" pageNumber="1353">Idaho</collectingRegion>
(
<figureCitation id="19C82A75FFC8720C7894FA49FDC2C284" box="[441,534,1445,1467]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="40" pageNumber="1353">Fig. 24A</figureCitation>
). As noted by
<bibRefCitation id="E5624B01FFC8720C7B9FFA49FE59C2E7" author="Erwin TL &amp; Kavanaugh DH" pageId="40" pageNumber="1353" pagination="33 - 72" refId="ref26201" refString="Erwin TL, Kavanaugh DH. 1981. Systematics and zoogeography of Bembidion Latreille: 1. The carlhi and erasum groups of western North America (Coleoptera: Carabidae, Bembidiini). Entomologica Scandinavica Supplement 15: 33-72." type="journal article" year="1981">Erwin &amp; Kavanaugh (1981)</bibRefCitation>
, the record from the Olympic Peninsula in
<collectingRegion id="4337F812FFC8720C7810FA0EFE05C2C6" box="[317,465,1506,1528]" country="United States of America" name="Washington" pageId="40" pageNumber="1353">Washington</collectingRegion>
is somewhat doubtful. Commonly encountered between 0 and
<quantity id="460B9B15FFC8720C7B4DF9EDFD6CC128" box="[608,696,1537,1558]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="1353" unit="m" value="2000.0">2000 m</quantity>
, with a few specimens found up to
<quantity id="460B9B15FFC8720C78D4F9CCFD82C10B" box="[505,598,1568,1589]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="40" pageNumber="1353" unit="m" value="2400.0">2400 m</quantity>
. Found in all months of the year except February; most common in the middle of summer.
</paragraph>
<paragraph id="814C36F0FFC8720C79BCF97AFDE7C019" blockId="40.[145,762,1686,1831]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C79BCF97AFF2EC195" box="[145,250,1686,1707]" italics="true" pageId="40" pageNumber="1353">Habitat:</emphasis>
By far the commonest large
<taxonomicName id="46F34D73FFC8720C7B50F97AFD26C195" authorityName="Casey" authorityYear="1918" box="[637,754,1686,1707]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC8720C7B50F97AFD26C195" box="[637,754,1686,1707]" italics="true" pageId="40" pageNumber="1353">Lionepha</emphasis>
</taxonomicName>
, found in a variety of habitats associated with water, including along the shores of creeks, especially in forests and on the edge of melting snowfields in open conifer forests in the Sierra
<collectingRegion id="4337F812FFC8720C78F9F8FDFDFBC019" box="[468,559,1809,1831]" country="United States of America" name="Nevada" pageId="40" pageNumber="1353">Nevada</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,163 @@
<document id="7D0CE0C3FABAF142B7C12A0CAC5FC56B" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFC8720C7A51FF29FC09C2B0" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha sequoiae" docType="treatment" docVersion="1" lastPageNumber="1353" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1353" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="F233F750F9D576150C0D6B5356D260F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7399359EA2F3ECCDDEE60F3E4AFF951C">
<mods:title id="A2C9A4775F504E50DA7EFFFA1E160232">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="22AB2E5AD7CCE3E71857C6C1A7744616" type="personal">
<mods:role id="482AEE9C17CEDBB2A63D8B4B304E8925">
<mods:roleTerm id="DB1ACCFB511D5843B9D05B5FDA7DB217">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8CCFDD808A46ADFDC6C7AC67479E4A6D">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="314272A65A7581024751A316B700E185" type="personal">
<mods:role id="D6C5CE0996CB3EC4C9504F9D661DC85F">
<mods:roleTerm id="8F92E21D7E99FE8FBE3DB6FDFC9F54A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FB9A5B585132C919EFD34603215759E">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C94C1057505583472FF37797A8EF49E0">text</mods:typeOfResource>
<mods:relatedItem id="1FB0F4A2E6D6BF48151528292F619138" type="host">
<mods:titleInfo id="227F94424DA429DA70D8FE81343949EF">
<mods:title id="DF76F7420AFE8ECDFC29F73842D8854B">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="8230C6A71A2C7279E0D91E85C6CF8332">
<mods:date id="07CAA8B3DEAB64F4A328543F2CD07122">2020</mods:date>
<mods:detail id="C3B5E3328E0A1FC5925A5318B3FBBCED" type="volume">
<mods:number id="822F56B5590DB3B28B7437E9E7B4C7EA">189</mods:number>
</mods:detail>
<mods:extent id="5C4CBB9C55F66C7C54A2C034C781EB98" unit="page">
<mods:start id="407B684CCF0F388D7ADA794817AF2CD5">1313</mods:start>
<mods:end id="C9E1CDFF21BD52F966B3239832ADAF2B">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="40164800CBDAA65A5091A030F865F5D3">journal article</mods:classification>
<mods:identifier id="8003CF479E05DF64D79CD87025FE4475" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="1DF2A2C34B296E34421C326375CBFDC3" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFC8720C7A51FF29FC09C2B0" LSID="urn:lsid:plazi:treatment:095A87E6FFC8720C7A51FF29FC09C2B0" httpUri="http://treatment.plazi.org/id/095A87E6FFC8720C7A51FF29FC09C2B0" lastPageNumber="1353" pageId="40" pageNumber="1353">
<subSubSection id="C9E9657BFFC8720C7A51FF29FAE9C7E3" box="[892,1341,196,221]" pageId="40" pageNumber="1353" type="nomenclature">
<paragraph id="814C36F0FFC8720C7A51FF29FAE9C7E3" blockId="40.[892,1341,196,221]" box="[892,1341,196,221]" pageId="40" pageNumber="1353">
<heading id="DA04819CFFC8720C7A51FF29FAE9C7E3" box="[892,1341,196,221]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="1353" reason="2">
<taxonomicName id="46F34D73FFC8720C7A51FF29FAE9C7E3" ID-CoL="3V6TZ" authority="(LINDROTH, 1963)" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[892,1341,196,221]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="sequoiae">
<emphasis id="B387EAE2FFC8720C7A51FF29FBBDC7E5" box="[892,1129,197,220]" italics="true" pageId="40" pageNumber="1353">
<smallCapsWord id="87AAA02CFFC8720C7A51FF29FC25C7E5" baselines="214,214" box="[892,1009,197,220]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lionepha" pageId="40" pageNumber="1353">LIONEPHA</smallCapsWord>
<smallCapsWord id="87AAA02CFFC8720C7AD5FF2BFBBDC7E5" baselines="214" box="[1016,1129,199,219]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sequoiae" pageId="40" pageNumber="1353">SEQUOIAE</smallCapsWord>
</emphasis>
(
<bibRefCitation id="E5624B01FFC8720C7D54FF28FAE0C7E3" author="Lindroth CH" box="[1145,1332,196,221]" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">
<smallCapsWord id="87AAA02CFFC8720C7D54FF28FB24C7E5" baselines="215,214" box="[1145,1264,196,221]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lindroth" pageId="40" pageNumber="1353">LINDROTH</smallCapsWord>
, 1963
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C7ABDFF1CFAFDC637" box="[912,1321,240,265]" pageId="40" pageNumber="1353" type="description">
<paragraph id="814C36F0FFC8720C7ABDFF1CFAFDC637" blockId="40.[912,1321,240,265]" box="[912,1321,240,265]" pageId="40" pageNumber="1353">
(
<figureCitation id="19C82A75FFC8720C7AB4FF1CFC21C637" box="[921,1013,240,265]" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="40" pageNumber="1353">
<smallCapsWord id="87AAA02CFFC8720C7AB4FF1CFC1BC639" baselines="259,258" box="[921,975,240,265]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="40" pageNumber="1353">FIGS</smallCapsWord>
3B
</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D2FFF1CFB84C637" box="[1026,1104,240,265]" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="40" pageNumber="1353">17C, D</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D70FF1CFB7EC637" box="[1117,1194,240,265]" captionStart="Figure 13" captionStartId="21.[163,241,1563,1585]" captionTargetBox="[173,1430,204,1518]" captionTargetId="figure-171@21.[163,1443,195,1523]" captionTargetPageId="21" captionText="Figure 13. Aedeagus of the Lionepha osculans species group, left lateral view. A, L. osculans; B, L. osculans; C, L. sequoiae; D, L. sequoiae; E, L. pseudoerasa; F, L. pseudoerasa; G, L. tuulukwa; H, L. tuulukwa. Scale bar 0.1 mm." pageId="40" pageNumber="1353">13C, D</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D9AFF1CFB31C637" box="[1207,1253,240,265]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="40" pageNumber="1353">19B</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7DDFFF1CFAF5C637" box="[1266,1313,240,265]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="40" pageNumber="1353">24B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C7A04FECAFB86C647" pageId="40" pageNumber="1353" type="reference_group">
<paragraph id="814C36F0FFC8720C7A04FECAFB86C647" blockId="40.[809,1425,294,377]" pageId="40" pageNumber="1353">
<treatmentCitationGroup id="A1E311DEFFC8720C7A04FECAFB86C647" pageId="40" pageNumber="1353">
<treatmentCitation id="005210E1FFC8720C7A04FECAFB3BC602" author="Lindroth CH" box="[809,1263,294,316]" page="260" pageId="40" pageNumber="1353" year="1963">
<taxonomicName id="46F34D73FFC8720C7A04FECAFB3BC602" ID-CoL="8ML9B" authority="Lindroth, 1963: 260" authorityName="Lindroth" authorityPageNumber="260" authorityYear="1963" box="[809,1263,294,316]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="sequoiae">
<emphasis id="B387EAE2FFC8720C7A04FECAFBDEC605" box="[809,1034,294,315]" italics="true" pageId="40" pageNumber="1353">Bembidion sequoiae</emphasis>
<materialsCitation id="319B3CADFFC8720C7D3DFECAFB3BC602" box="[1040,1263,294,316]" collectionCode="MCZ" location="Sequoia National Park" pageId="40" pageNumber="1353" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<bibRefCitation id="E5624B01FFC8720C7D3DFECAFB3BC602" author="Lindroth CH" box="[1040,1263,294,316]" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth, 1963: 260</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="5E488852FFC8720C7DD5FECAFA8AC602" box="[1272,1374,294,316]" pageId="40" pageNumber="1353" type="holotype">Holotype</typeStatus>
<specimenCount id="97F5FD79FFC8720C7C49FEC5FAA0C605" box="[1380,1396,297,315]" count="1" pageId="40" pageNumber="1353" type="male"></specimenCount>
in MCZ (type # 32532), examined. Type locality:
<location id="842C602BFFC8720C7C1BFEA8FC19C647" LSID="urn:lsid:plazi:treatment:095A87E6FFC8720C7A51FF29FC09C2B0:842C602BFFC8720C7C1BFEA8FC19C647" name="Sequoia National Park" pageId="40" pageNumber="1353" stateProvince="California">Sequoia National Park</location>
,
<collectingRegion id="4337F812FFC8720C7AFAFE8FFB9AC647" box="[983,1102,355,377]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C7A04FE74FC09C2B0" pageId="40" pageNumber="1353" type="diagnosis">
<paragraph id="814C36F0FFC8720C7A04FE74FB9CC5FE" blockId="40.[809,1425,407,705]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C7A04FE74FC71C693" box="[809,933,408,429]" italics="true" pageId="40" pageNumber="1353">Diagnosis:</emphasis>
The most distinctive external feature of this moderately large
<taxonomicName id="46F34D73FFC8720C7D1FFE5AFB74C6F5" authorityName="Casey" authorityYear="1918" box="[1074,1184,438,459]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC8720C7D1FFE5AFB74C6F5" box="[1074,1184,438,459]" italics="true" pageId="40" pageNumber="1353">Lionepha</emphasis>
</taxonomicName>
is the exceptionally small basal protarsomeres in males (
<figureCitation id="19C82A75FFC8720C7DF8FE38FAE1C6D7" box="[1237,1333,468,490]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="40" pageNumber="1353">Fig. 19B</figureCitation>
), which are only slightly wider than the second protarsomeres, much smaller than in other
<taxonomicName id="46F34D73FFC8720C7DB8FDFEFADCC519" authorityName="Casey" authorityYear="1918" box="[1173,1288,530,551]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="genus">
<emphasis id="B387EAE2FFC8720C7DB8FDFEFADCC519" box="[1173,1288,530,551]" italics="true" pageId="40" pageNumber="1353">Lionepha</emphasis>
</taxonomicName>
(see other images in
<figureCitation id="19C82A75FFC8720C7AB1FDDCFC3CC57B" box="[924,1000,560,582]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="40" pageNumber="1353">Fig. 19</figureCitation>
). Elytra not iridescent, as sculpticells are not sufficiently transverse (
<figureCitation id="19C82A75FFC8720C7DBDFDA3FADAC55A" box="[1168,1294,591,613]" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="40" pageNumber="1353">Fig. 17C, D</figureCitation>
). Aedeagus with broad apex, and with clearly visible dark scales on the internal sac membranes, yielding a speckled appearance (
<figureCitation id="19C82A75FFC8720C7A96FD47FBECC5FE" box="[955,1080,683,705]" captionStart="Figure 13" captionStartId="21.[163,241,1563,1585]" captionTargetBox="[173,1430,204,1518]" captionTargetId="figure-171@21.[163,1443,195,1523]" captionTargetPageId="21" captionText="Figure 13. Aedeagus of the Lionepha osculans species group, left lateral view. A, L. osculans; B, L. osculans; C, L. sequoiae; D, L. sequoiae; E, L. pseudoerasa; F, L. pseudoerasa; G, L. tuulukwa; H, L. tuulukwa. Scale bar 0.1 mm." pageId="40" pageNumber="1353">Fig. 13C, D</figureCitation>
).
</paragraph>
<paragraph id="814C36F0FFC8720C7A04FD06FA45C465" blockId="40.[809,1426,745,860]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C7A04FD06FB85C5C1" box="[809,1105,746,767]" italics="true" pageId="40" pageNumber="1353">Additional characteristics:</emphasis>
Body length
<quantity id="460B9B15FFC8720C7DC1FD05FA59C5C1" box="[1260,1421,745,767]" metricMagnitude="-3" metricUnit="m" metricValue="4.63" metricValueMax="4.95" metricValueMin="4.31" pageId="40" pageNumber="1353" unit="mm" value="4.63" valueMax="4.95" valueMin="4.31">4.314.95 mm</quantity>
. Antenna piceous, with first antennomere rufous, at least on the underside. Legs rufous, with darker joints. Hind wings full-sized. Chromosomes of male 24 + X (
<tableCitation id="CC71034BFFC8720C7C1FFCAAFA56C465" box="[1330,1410,837,860]" captionStart="Table 5" captionStartId="29.[162,226,196,217]" captionText="Table 5. Chromosome numbers and sex chromosomes of Lionepha males. The Sample column indicates the number of males examined" pageId="40" pageNumber="1353">Table 5</tableCitation>
).
</paragraph>
<paragraph id="814C36F0FFC8720C7A04FC60FA9DC388" blockId="40.[809,1426,908,1206]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C7A04FC60FC1EC49F" box="[809,970,908,929]" italics="true" pageId="40" pageNumber="1353">Distribution:</emphasis>
This species has been found in the Sierra
<collectingRegion id="4337F812FFC8720C7A57FC47FC07C4FF" box="[890,979,939,961]" country="United States of America" name="Nevada" pageId="40" pageNumber="1353">Nevada</collectingRegion>
of
<collectingRegion id="4337F812FFC8720C7AD5FC47FBA5C4FF" box="[1016,1137,939,961]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
, the Cascades of
<collectingRegion id="4337F812FFC8720C7C11FC47FA45C4FE" box="[1340,1425,939,960]" country="United States of America" name="Oregon" pageId="40" pageNumber="1353">Oregon</collectingRegion>
and north to Vancouver,
<collectingRegion id="4337F812FFC8720C7D6BFC26FAC4C4DE" box="[1094,1296,970,992]" country="Canada" name="British Columbia" pageId="40" pageNumber="1353">British Columbia</collectingRegion>
(
<figureCitation id="19C82A75FFC8720C7C0CFC26FA56C4E1" box="[1313,1410,970,992]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="40" pageNumber="1353">Fig. 24B</figureCitation>
). There are, in addition, two low-elevation records from coastal
<collectingRegion id="4337F812FFC8720C7AAEFBEBFC2EC323" box="[899,1018,1031,1053]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
(Sonoma County); although one of these records (from
<quantity id="460B9B15FFC8720C7D39FBC9FBAAC302" box="[1044,1150,1061,1084]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="40" pageNumber="1353" unit="mi" value="2.5">2.5 miles</quantity>
N of Cazadero on King Ridge, Big Austin Creek; CAS) is relatively recent, both records are unexpected and should be confirmed. Found from
<quantity id="460B9B15FFC8720C7AECFB6EFBB6C3A9" box="[961,1122,1154,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.13" metricValueMax="2.23" metricValueMin="0.03" pageId="40" pageNumber="1353" unit="m" value="1130.0" valueMax="2230.0" valueMin="30.0">30 to 2230 m</quantity>
in elevation. Specimens were collected from March through September.
</paragraph>
<paragraph id="814C36F0FFC8720C7A04FB33FC09C2B0" blockId="40.[809,1425,1247,1422]" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C7A04FB33FC58C3CA" box="[809,908,1247,1268]" italics="true" pageId="40" pageNumber="1353">Habitat:</emphasis>
<materialsCitation id="319B3CADFFC8720C7AB7FB33FC0CC2B0" location="Cascades" municipality="School Creek" pageId="40" pageNumber="1353" specimenCount="80" stateProvince="Oregon">
Found along the gravel and cobble shores of creeks in forests or in forest clearings. Over
<specimenCount id="97F5FD79FFC8720C7C58FB12FC77C20C" count="80" pageId="40" pageNumber="1353" type="generic">80 specimens</specimenCount>
were found along partly shaded areas of
<collectingMunicipality id="6128AC8AFFC8720C7A04FAD7FC12C26F" box="[809,966,1339,1361]" pageId="40" pageNumber="1353">School Creek</collectingMunicipality>
in the
<location id="842C602BFFC8720C7D33FAD7FB58C26F" LSID="urn:lsid:plazi:treatment:095A87E6FFC8720C7A51FF29FC09C2B0:842C602BFFC8720C7D33FAD7FB58C26F" box="[1054,1164,1339,1361]" municipality="School Creek" name="Cascades" pageId="40" pageNumber="1353" stateProvince="Oregon">Cascades</location>
of
<collectingRegion id="4337F812FFC8720C7D9EFAD7FAD8C26E" box="[1203,1292,1339,1360]" country="United States of America" name="Oregon" pageId="40" pageNumber="1353">Oregon</collectingRegion>
, with most found at the gravel and cobble edge of a splash pool below a culvert
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,184 @@
<document id="AE238471EC46AFB60FF2E04CAFD2E739" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFC8720E7A49FA32FD03C7C4" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha pseudoerasa" docType="treatment" docVersion="1" lastPageNumber="1355" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1353" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="34F2F875A4D92439075C2BFD84D50638" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5C663C151A614E3FB3FF9D8282BEBF15">
<mods:title id="12C3657010808B74B9116A228172656C">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="6FCBA4DBDD6532FE930D17801E6B93D4" type="personal">
<mods:role id="DC67CBC65671EA90D424F5EAD770EE3E">
<mods:roleTerm id="5AF9BFCE7BD3D5E3B6B0CD84694BAD81">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33D697C9FA960916B7CA561029D213AF">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="70B856C4361766D247796B9501A56D26" type="personal">
<mods:role id="6133F73A0AE39B441F1E542AFF700EA4">
<mods:roleTerm id="D970BF0AE67335367A06D59F3B9DD4B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBC3A2E12048F6E13F4F4BD4E8B3052B">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="24DCD62DBA49584F278B6AECF9DC17DD">text</mods:typeOfResource>
<mods:relatedItem id="019D82C9CF85DD4D0BC1DABABE57D74B" type="host">
<mods:titleInfo id="ECF009EE01AA514FD6CA63072DB97F2D">
<mods:title id="73061E2626C733D9051FF2956650FA54">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="3EF37C4F5246A02D3A8A65030C0A25D4">
<mods:date id="C68607D8151DC367386B8DF6CBB60BB7">2020</mods:date>
<mods:detail id="BFCE94D57AA984496A70BA1F437A5BAF" type="volume">
<mods:number id="CAB3B896D8C0584B52F4890BF6D1045C">189</mods:number>
</mods:detail>
<mods:extent id="C08D8FD902D061CC5B5FBCDAC17CB444" unit="page">
<mods:start id="CAA987BDD133DB69F04D5F3A722C3582">1313</mods:start>
<mods:end id="B818DD12370984D15C3F655CD2CE0BC0">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0A41B3BE52303DA8872ECEA369C8781F">journal article</mods:classification>
<mods:identifier id="D7195E32FA393B4D9C3C1454AC8E0AFB" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="EEECAB3C54714E939A52A289315A37C9" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFC8720E7A49FA32FD03C7C4" LSID="urn:lsid:plazi:treatment:095A87E6FFC8720E7A49FA32FD03C7C4" httpUri="http://treatment.plazi.org/id/095A87E6FFC8720E7A49FA32FD03C7C4" lastPageId="42" lastPageNumber="1355" pageId="40" pageNumber="1353">
<subSubSection id="C9E9657BFFC8720C7A49FA32FA81C2C8" box="[868,1365,1501,1526]" pageId="40" pageNumber="1353" type="nomenclature">
<paragraph id="814C36F0FFC8720C7A49FA32FA81C2C8" blockId="40.[868,1365,1501,1526]" box="[868,1365,1501,1526]" pageId="40" pageNumber="1353">
<heading id="DA04819CFFC8720C7A49FA32FA81C2C8" box="[868,1365,1501,1526]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="1353" reason="2">
<taxonomicName id="46F34D73FFC8720C7A49FA32FA81C2C8" ID-CoL="3V6TY" authority="(LINDROTH, 1963)" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[868,1365,1501,1526]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="pseudoerasa">
<emphasis id="B387EAE2FFC8720C7A49FA32FB55C2CB" box="[868,1153,1502,1525]" italics="true" pageId="40" pageNumber="1353">
<smallCapsWord id="87AAA02CFFC8720C7A49FA32FC0DC2CB" baselines="1519,1520" box="[868,985,1502,1525]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lionepha" pageId="40" pageNumber="1353">LIONEPHA</smallCapsWord>
<smallCapsWord id="87AAA02CFFC8720C7ACDFA0DFB55C2CB" baselines="1520" box="[992,1153,1505,1525]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="pseudoerasa" pageId="40" pageNumber="1353">PSEUDOERASA</smallCapsWord>
</emphasis>
(
<bibRefCitation id="E5624B01FFC8720C7DBCFA31FA98C2C8" author="Lindroth CH" box="[1169,1356,1501,1526]" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">
<smallCapsWord id="87AAA02CFFC8720C7DBCFA31FADCC2CB" baselines="1520,1520" box="[1169,1288,1501,1526]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lindroth" pageId="40" pageNumber="1353">LINDROTH</smallCapsWord>
, 1963
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C7AB8F9E0FAF0C11B" box="[917,1316,1548,1573]" pageId="40" pageNumber="1353" type="description">
<paragraph id="814C36F0FFC8720C7AB8F9E0FAF0C11B" blockId="40.[917,1316,1548,1573]" box="[917,1316,1548,1573]" pageId="40" pageNumber="1353">
(
<figureCitation id="19C82A75FFC8720C7AB0F9E0FC2DC11B" box="[925,1017,1548,1573]" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="40" pageNumber="1353">
<smallCapsWord id="87AAA02CFFC8720C7AB0F9E0FC07C11D" baselines="1567,1566" box="[925,979,1548,1573]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="40" pageNumber="1353">FIGS</smallCapsWord>
3C
</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D2BF9E0FB84C11B" box="[1030,1104,1548,1573]" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="40" pageNumber="1353">17E, F</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D70F9E0FB72C11B" box="[1117,1190,1548,1573]" captionStart="Figure 13" captionStartId="21.[163,241,1563,1585]" captionTargetBox="[173,1430,204,1518]" captionTargetId="figure-171@21.[163,1443,195,1523]" captionTargetPageId="21" captionText="Figure 13. Aedeagus of the Lionepha osculans species group, left lateral view. A, L. osculans; B, L. osculans; C, L. sequoiae; D, L. sequoiae; E, L. pseudoerasa; F, L. pseudoerasa; G, L. tuulukwa; H, L. tuulukwa. Scale bar 0.1 mm." pageId="40" pageNumber="1353">13E, F</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7D9EF9E0FB35C11B" box="[1203,1249,1548,1573]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="40" pageNumber="1353">19C</figureCitation>
,
<figureCitation id="19C82A75FFC8720C7DC3F9E0FAC8C11B" box="[1262,1308,1548,1573]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="40" pageNumber="1353">24C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720C7A04F9D0FAF2C1AE" pageId="40" pageNumber="1353" type="reference_group">
<paragraph id="814C36F0FFC8720C7A04F9D0FAF2C1AE" blockId="40.[809,1425,1596,1833]" pageId="40" pageNumber="1353">
<treatmentCitationGroup id="A1E311DEFFC8720C7A04F9D0FAF2C1AE" pageId="40" pageNumber="1353">
<treatmentCitation id="005210E1FFC8720C7A04F9D0FA5FC16C" author="Lindroth CH" box="[809,1419,1596,1618]" page="260" pageId="40" pageNumber="1353" year="1963">
<taxonomicName id="46F34D73FFC8720C7A04F9D0FA5FC16C" ID-CoL="8M6HQ" authority="Lindroth, 1963: 260" authorityName="Lindroth" authorityPageNumber="260" authorityYear="1963" box="[809,1419,1596,1618]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1353" phylum="Arthropoda" rank="species" species="pseudoerasum">
<emphasis id="B387EAE2FFC8720C7A04F9D0FBA2C16F" box="[809,1142,1596,1617]" italics="true" pageId="40" pageNumber="1353">Bembidion pseudoerasum</emphasis>
<materialsCitation id="319B3CADFFC8720C7DA9F9D0FA5FC16C" box="[1156,1419,1596,1618]" collectionCode="USNM" location="County" municipality="Truckee" pageId="40" pageNumber="1353" specimenCount="1" specimenCount-male="1" stateProvince="Nevada" typeStatus="holotype">
<bibRefCitation id="E5624B01FFC8720C7DA9F9D0FA5FC16C" author="Lindroth CH" box="[1156,1419,1596,1618]" pageId="40" pageNumber="1353" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth, 1963: 260</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="5E488852FFC8720C7A04F9B7FC45C14F" box="[809,913,1627,1649]" pageId="40" pageNumber="1353" type="holotype">Holotype</typeStatus>
<specimenCount id="97F5FD79FFC8720C7AB4F9B2FC7DC14E" box="[921,937,1630,1648]" count="1" pageId="40" pageNumber="1353" type="male"></specimenCount>
in USNM (type # 76638), examined. Type locality:
<collectingMunicipality id="6128AC8AFFC8720C7AA5F996FC32C1AE" box="[904,998,1658,1680]" pageId="40" pageNumber="1353">Truckee</collectingMunicipality>
,
<collectingRegion id="4337F812FFC8720C7ADCF996FB9CC1AE" box="[1009,1096,1658,1680]" country="United States of America" name="Nevada" pageId="40" pageNumber="1353">Nevada</collectingRegion>
<location id="842C602BFFC8720C7D62F996FB75C1B1" LSID="urn:lsid:plazi:treatment:095A87E6FFC8720E7A49FA32FD03C7C4:842C602BFFC8720C7D62F996FB75C1B1" box="[1103,1185,1658,1679]" municipality="Truckee" name="County" pageId="40" pageNumber="1353" stateProvince="Nevada">County</location>
,
<collectingRegion id="4337F812FFC8720C7D81F996FAF7C1AE" box="[1196,1315,1658,1680]" country="United States of America" name="California" pageId="40" pageNumber="1353">California</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFC8720E7A04F975FD03C7C4" lastPageId="42" lastPageNumber="1355" pageId="40" pageNumber="1353" type="diagnosis">
<paragraph id="814C36F0FFC8720D7A04F975FEAEC157" blockId="40.[809,1425,1596,1833]" lastBlockId="41.[163,780,1466,1795]" lastPageId="41" lastPageNumber="1354" pageId="40" pageNumber="1353">
<emphasis id="B387EAE2FFC8720C7A04F975FC77C190" box="[809,931,1689,1710]" italics="true" pageId="40" pageNumber="1353">Diagnosis:</emphasis>
This species is most easily distinguished externally by the reduced elytral striation, with third stria not visible beyond the posterior dorsal puncture. Elytra only slightly iridescent. Prothoracic sides moderately rounded, in most specimens with the posterior portion being almost parallel sided (
<figureCitation id="19C82A75FFC9720D7BCDFA56FF16C2D0" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="41" pageNumber="1354">Fig. 3C</figureCitation>
). Basal protarsomeres of male large, significantly wider than the second tarsomere (
<figureCitation id="19C82A75FFC9720D7B04FA1BFD5CC133" box="[553,648,1527,1549]" captionStart="Figure 19" captionStartId="28.[145,227,1848,1870]" captionTargetBox="[163,1425,1163,1806]" captionTargetId="figure-362@28.[145,1425,1160,1809]" captionTargetPageId="28" captionText="Figure 19. Dorsal view of male protarsi of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa; D, L. tuulukwa." pageId="41" pageNumber="1354">Fig. 19C</figureCitation>
). Aedeagus with ventral surface gently curved (
<figureCitation id="19C82A75FFC9720D7B6FF9FAFD14C112" box="[578,704,1558,1580]" captionStart="Figure 11" captionStartId="19.[165,245,1848,1870]" captionTargetBox="[232,1370,923,1806]" captionTargetId="figure-191@19.[227,1379,918,1809]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of three species of the Lionepha erasa species group, left lateral view. A, L. erasa; B, L. erasa; C, L. australerasa; D, L. australerasa, holotype; E, L. probata; F, L. probata. Scale bar 0.1 mm." pageId="41" pageNumber="1354">Fig. 11E, F</figureCitation>
), with internal sac sclerites reduced (although not as much so as in
<taxonomicName id="46F34D73FFC9720D79CDF9B8FEBFC156" box="[224,363,1619,1641]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="tuulukwa">
<emphasis id="B387EAE2FFC9720D79CDF9B8FEBFC156" box="[224,363,1619,1641]" italics="true" pageId="41" pageNumber="1354">L. tuulukwa</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="D58C6678FFC9720D798EFAB6FAD7C2B0" pageId="41" pageNumber="1354" startId="41.[163,242,1370,1392]" targetBox="[166,1443,195,1330]" targetPageId="41" targetType="figure">
<paragraph id="814C36F0FFC9720D798EFAB6FAD7C2B0" blockId="41.[163,1443,1370,1422]" pageId="41" pageNumber="1354">
<emphasis id="B387EAE2FFC9720D798EFAB6FECDC251" bold="true" box="[163,281,1370,1392]" pageId="41" pageNumber="1354">Figure 24.</emphasis>
Geographic distribution of members of the
<taxonomicName id="46F34D73FFC9720D7BC1FAB7FC65C24F" authorityName="DNA" authorityYear="2638" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[748,945,1371,1393]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC9720D7BC1FAB7FC65C24F" box="[748,945,1371,1393]" italics="true" pageId="41" pageNumber="1354">Lionepha osculans</emphasis>
</taxonomicName>
species group. A,
<taxonomicName id="46F34D73FFC9720D7D40FAB7FB30C24F" authorityName="DNA" authorityYear="2638" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[1133,1252,1371,1393]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC9720D7D40FAB7FB30C24F" box="[1133,1252,1371,1393]" italics="true" pageId="41" pageNumber="1354">L. osculans</emphasis>
</taxonomicName>
; B,
<taxonomicName id="46F34D73FFC9720D7C21FAB7FA55C24F" authorityName="Lindroth" authorityYear="1963" box="[1292,1409,1371,1393]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="sequoiae">
<emphasis id="B387EAE2FFC9720D7C21FAB7FA55C24F" box="[1292,1409,1371,1393]" italics="true" pageId="41" pageNumber="1354">L. sequoiae</emphasis>
</taxonomicName>
; C,
<taxonomicName id="46F34D73FFC9720D798EFA95FEEAC2B0" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[163,318,1400,1422]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="pseudoerasa">
<emphasis id="B387EAE2FFC9720D798EFA95FEEAC2B0" box="[163,318,1400,1422]" italics="true" pageId="41" pageNumber="1354">L. pseudoerasa</emphasis>
</taxonomicName>
(circles) and
<taxonomicName id="46F34D73FFC9720D78E7FA95FD9CC2B0" box="[458,584,1400,1422]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="tuulukwa">
<emphasis id="B387EAE2FFC9720D78E7FA95FD9CC2B0" box="[458,584,1400,1422]" italics="true" pageId="41" pageNumber="1354">L. tuulukwa</emphasis>
</taxonomicName>
(stars). Specimens whose DNA was sequenced are shown in black.
</paragraph>
</caption>
<paragraph id="814C36F0FFC9720D7996F99FFC7EC156" blockId="41.[163,780,1466,1795]" lastBlockId="41.[827,1443,1466,1641]" pageId="41" pageNumber="1354">
In comparison to
<taxonomicName id="46F34D73FFC9720D78B4F99FFDFBC1B9" box="[409,559,1650,1672]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="tuulukwa">
<emphasis id="B387EAE2FFC9720D78B4F99FFDFBC1B9" box="[409,559,1650,1672]" italics="true" pageId="41" pageNumber="1354">L. tuulukwa</emphasis>
</taxonomicName>
, the prothorax is relatively wider compared to the head (
<figureCitation id="19C82A75FFC9720D7B59F97DFD13C198" box="[628,711,1681,1703]" captionStart="Figure 3" captionStartId="3.[163,243,1395,1417]" captionTargetBox="[403,1212,195,1329]" captionTargetId="figure-160@3.[390,1216,195,1355]" captionTargetPageId="3" captionText="Figure 3. Adults of the Lionepha osculans species group. A, L. osculans, USA: California: Tehama Co., Nanny Creek; B, L. sequoiae, USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir; C, L. pseudoerasa, USA: California: El Dorado Co., Strawberry Creek at Sciots Camp; D, L. tuulukwa, USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="41" pageNumber="1354">Fig. 3C</figureCitation>
), and the elytral microsculpture is more transverse (
<figureCitation id="19C82A75FFC9720D7B85F943FF67C1DA" captionStart="Figure 17" captionStartId="25.[162,241,1305,1327]" captionTargetBox="[166,1440,199,1263]" captionTargetId="figure-189@25.[163,1443,195,1266]" captionTargetPageId="25" captionText="Figure 17. Microsculpture of the elytral disc of the Lionepha osculans species group. Left elytron, region around anterior discal seta on third interval. A, L. osculans, male; B, L. osculans, female; C, L. sequoiae, male; D, L. sequoiae, female; E, L. pseudoerasa, male; F, L. pseudoerasa, female; G, L. tuulukwa, male; H, L. tuulukwa, female. Scale bar 50 µm." pageId="41" pageNumber="1354">Fig. 17E, F</figureCitation>
). In comparison to
<taxonomicName id="46F34D73FFC9720D78BCF923FDC5C1DD" authorityName="DNA" authorityYear="2638" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[401,529,1742,1764]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="species" species="osculans">
<emphasis id="B387EAE2FFC9720D78BCF923FDC5C1DD" box="[401,529,1742,1764]" italics="true" pageId="41" pageNumber="1354">L. osculans</emphasis>
</taxonomicName>
, narrower and with a less rounded prothorax, and with less iridescent elytra.
<emphasis id="B387EAE2FFC9720D7A16FA56FB4AC2F1" box="[827,1182,1466,1487]" italics="true" pageId="41" pageNumber="1354">Additional characteristics:</emphasis>
Body length
<quantity id="460B9B15FFC9720D7C70FA56FC72C2D0" metricMagnitude="-3" metricUnit="m" metricValue="4.83" metricValueMax="5.25" metricValueMin="4.41" pageId="41" pageNumber="1354" unit="mm" value="4.83" valueMax="5.25" valueMin="4.41">4.41 5.25 mm</quantity>
, with most specimens&gt;
<quantity id="460B9B15FFC9720D7DE2FA35FAFAC2D0" box="[1231,1326,1497,1519]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="41" pageNumber="1354" unit="mm" value="4.6">4.6 mm</quantity>
. Antenna piceous, with underside of first antennomere rufous. Legs rufous or dark rufous, with darker joints. Hind wings full-sized. Male with 25 chromosomes (
<tableCitation id="CC71034BFFC9720D7A6EF9BFFC4EC157" box="[835,922,1619,1641]" captionStart="Table 5" captionStartId="29.[162,226,196,217]" captionText="Table 5. Chromosome numbers and sex chromosomes of Lionepha males. The Sample column indicates the number of males examined" pageId="41" pageNumber="1354">Table 5</tableCitation>
).
</paragraph>
<paragraph id="814C36F0FFC9720D7A16F97EFB1FC03D" blockId="41.[827,1443,1682,1796]" pageId="41" pageNumber="1354">
<emphasis id="B387EAE2FFC9720D7A16F97EFC36C199" box="[827,994,1682,1703]" italics="true" pageId="41" pageNumber="1354">Distribution:</emphasis>
This species is only known from the Sierra
<collectingRegion id="4337F812FFC9720D7AE9F95DFBF5C1F9" box="[964,1057,1713,1735]" country="United States of America" name="Nevada" pageId="41" pageNumber="1354">Nevada</collectingRegion>
and
<taxonomicName id="46F34D73FFC9720D7D4EF95DFB2EC1F9" authority="Alps" authorityName="Alps" box="[1123,1274,1713,1735]" class="Arachnida" family="Hubbardiidae" genus="Trinity" higherTaxonomySource="CoL" kingdom="Animalia" order="Schizomida" pageId="41" pageNumber="1354" phylum="Arthropoda" rank="genus">Trinity Alps</taxonomicName>
of
<collectingRegion id="4337F812FFC9720D7C0BF95DFA76C1F9" box="[1318,1442,1713,1735]" country="United States of America" name="California" pageId="41" pageNumber="1354">California</collectingRegion>
(
<figureCitation id="19C82A75FFC9720D7A6EF923FC75C1DA" box="[835,929,1743,1765]" captionStart="Figure 24" captionStartId="41.[163,242,1370,1392]" captionTargetBox="[166,1443,195,1330]" captionTargetId="figure-202@41.[163,1443,195,1331]" captionTargetPageId="41" captionText="Figure 24. Geographic distribution of members of the Lionepha osculans species group. A, L. osculans; B, L. sequoiae; C, L. pseudoerasa (circles) and L. tuulukwa (stars). Specimens whose DNA was sequenced are shown in black." pageId="41" pageNumber="1354">Fig. 24C</figureCitation>
). A mid-elevation species, found from
<quantity id="460B9B15FFC9720D7C62F923FC44C03D" metricMagnitude="3" metricUnit="m" metricValue="1.96" metricValueMax="2.7199999999999998" metricValueMin="1.2" pageId="41" pageNumber="1354" unit="m" value="1960.0" valueMax="2720.0" valueMin="1200.0">1200 to 2720 m</quantity>
, from May through August.
</paragraph>
<paragraph id="814C36F0FFCA720E79BCFF29FD03C7C4" blockId="42.[145,760,197,250]" pageId="42" pageNumber="1355">
<emphasis id="B387EAE2FFCA720E79BCFF29FF25C7E4" box="[145,241,197,218]" italics="true" pageId="42" pageNumber="1355">Habitat:</emphasis>
This rarely collected species has been found on sand and silt near streams and ponds in forests.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,68 @@
<document id="E450A95365DE50777E36B0AD117794CA" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFF9723D79E8F8EBFE35C064" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha probata" docType="treatment" docVersion="1" lastPageNumber="1338" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1338" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="8D23D1249AF44E2E6D8B68EE500537F6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5540F3112D201C9ECA9988F3D30C0CBC">
<mods:title id="68A0FB07DFE379725B94D7A12DB9C12A">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="2BF9DCE87760549EC7CD2ED30B184392" type="personal">
<mods:role id="8FAA830E382E42B1E8122FDECB0BA6BD">
<mods:roleTerm id="FD0026AF7441AD3E4A608109780A43E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4EB36DF6A5A86657F8B827D9D660F2FA">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="C547B69A23B358D0B255D1128916DA3F" type="personal">
<mods:role id="AF0249F3B85EEEFDC18E5195D5FFF996">
<mods:roleTerm id="EA9315D57D177215BC282D7187AE8353">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="829FEBA06F39147DC6F37CD0C9464797">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5FDC35671DDF919368E318A69D5CF41E">text</mods:typeOfResource>
<mods:relatedItem id="7665870FABDF4DCE950A933CBE72AE4D" type="host">
<mods:titleInfo id="E2636FBA2205AA37562623014CB499D5">
<mods:title id="1F567604B6F9CEA3719D5034A9FD3244">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E47937C04351862E6B26E8614A5BFA3C">
<mods:date id="381D3C5191C2E9CD8D7474BE0C8AE748">2020</mods:date>
<mods:detail id="FAFB26C79C777D11FF9E32D0E3211D7F" type="volume">
<mods:number id="83C39B5611F011617A215904346C025D">189</mods:number>
</mods:detail>
<mods:extent id="E28259E4AD073C48E1B343095C2C6A96" unit="page">
<mods:start id="0A98C5A404EC6DA1E941FC2727BBAAC5">1313</mods:start>
<mods:end id="714B50137B35BAB9DFD17FDCFA3F9F2C">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A65303C7D76343B0FA2BF37584035307">journal article</mods:classification>
<mods:identifier id="0ACAB9E5F06CB0BABB720D799D0BCC07" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="12665B2E7E2AA4FF9B1ADA0AE080A300" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFF9723D79E8F8EBFE35C064" LSID="urn:lsid:plazi:treatment:095A87E6FFF9723D79E8F8EBFE35C064" httpUri="http://treatment.plazi.org/id/095A87E6FFF9723D79E8F8EBFE35C064" lastPageNumber="1338" pageId="25" pageNumber="1338">
<subSubSection id="C9E9657BFFF9723D79E8F8EBFE03C025" box="[197,471,1798,1820]" pageId="25" pageNumber="1338" type="nomenclature">
<paragraph id="814C36F0FFF9723D79E8F8EBFE03C025" blockId="25.[197,489,1737,1882]" box="[197,471,1798,1820]" pageId="25" pageNumber="1338">
<heading id="DA04819CFFF9723D79E8F8EBFE03C025" box="[197,471,1798,1820]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="1338" reason="2">
<taxonomicName id="46F34D73FFF9723D79E8F8EBFE03C025" ID-CoL="3V6TX" authority="(Casey, 1918)" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[197,471,1798,1820]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1338" phylum="Arthropoda" rank="species" species="probata">
<emphasis id="B387EAE2FFF9723D79E8F8EBFEEFC025" box="[197,315,1798,1820]" italics="true" pageId="25" pageNumber="1338">L. probata</emphasis>
(
<bibRefCitation id="E5624B01FFF9723D7867F8EAFE1BC025" author="Casey TL" box="[330,463,1798,1819]" pageId="25" pageNumber="1338" pagination="1 - 223" refId="ref26056" refString="Casey TL. 1918. A review of the North American Bembidiinae. Memoirs on the Coleoptera 8: 1-223." type="journal article" year="1918">Casey, 1918</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFF9723D79E8F8CAFE35C064" pageId="25" pageNumber="1338" type="reference_group">
<paragraph id="814C36F0FFF9723D79E8F8CAFE6EC004" blockId="25.[197,489,1737,1882]" box="[197,442,1829,1851]" pageId="25" pageNumber="1338">
<taxonomicName id="46F34D73FFF9723D79E8F8CAFE6EC004" ID-CoL="3V6TM" authority="(Casey, 1918)" baseAuthorityName="Casey" baseAuthorityYear="1918" box="[197,442,1829,1851]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1338" phylum="Arthropoda" rank="species" species="casta">
<emphasis id="B387EAE2FFF9723D79E8F8CAFECAC005" box="[197,286,1830,1851]" italics="true" pageId="25" pageNumber="1338">L. casta</emphasis>
(
<bibRefCitation id="E5624B01FFF9723D7800F8C9FE66C004" author="Casey TL" box="[301,434,1829,1850]" pageId="25" pageNumber="1338" pagination="1 - 223" refId="ref26056" refString="Casey TL. 1918. A review of the North American Bembidiinae. Memoirs on the Coleoptera 8: 1-223." type="journal article" year="1918">Casey, 1918</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="814C36F0FFF9723D79E8F8A8FE35C064" blockId="25.[197,489,1737,1882]" box="[197,481,1860,1882]" pageId="25" pageNumber="1338">
<taxonomicName id="46F34D73FFF9723D79E8F8A8FE35C064" authority="Maddison" authorityName="Maddison" authorityYear="2020" box="[197,481,1860,1882]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1338" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="B387EAE2FFF9723D79E8F8A8FEBFC067" box="[197,363,1860,1881]" italics="true" pageId="25" pageNumber="1338">L. kavanaughi</emphasis>
Maddison
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,60 @@
<document id="87FE1CD0C0DF451A0060F77DD5F2ABB7" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFF9723D7A70FA58FB71C2D9" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha lindrothi Maddison &amp; Sproul 2020" docType="treatment" docVersion="1" lastPageNumber="1338" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1338" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="D338E1AB2CD95BD96FDAEF5E94E91D87" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D271D9BB55A876D576BBF293DFD2CFBA">
<mods:title id="3AC7542E47A11BC04B765FC07834426D">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="AA19768549A2202F615B44D37DE6E983" type="personal">
<mods:role id="70AC80DF73B67C9918BA218648BDBECC">
<mods:roleTerm id="F837AA4D4FBA2B9C71A9BB5C1F9E2644">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="386D04BF20C7DDFBEF9B95C1B988DBD6">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="389D323704FDD7E7DC0E723F1BB9B64A" type="personal">
<mods:role id="451D6B643E898C75F1A816E4D57D0F57">
<mods:roleTerm id="CEA395C7F205943C7C6D7B316F4EED0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0BBEDFC91806C7E2705664620539CF8">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="BBFF18DDFCD50464002E7B1C82614DD4">text</mods:typeOfResource>
<mods:relatedItem id="67C5447E19227E26BECDA9FB0A10588D" type="host">
<mods:titleInfo id="FE5560554282B230EE4200897D8B55F0">
<mods:title id="0DFB59F88452F28C77F0DC13821DD264">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="A86EBA48F5B74536F21893F79DBA6D60">
<mods:date id="A15FF500F125CC183781CC46B6EF7698">2020</mods:date>
<mods:detail id="0C393AEE1F0B59CF0E85C3113304A70C" type="volume">
<mods:number id="405EA5527506427E117B14F44F40686B">189</mods:number>
</mods:detail>
<mods:extent id="8A671F2F5C2450539DA1A7C737883980" unit="page">
<mods:start id="383487882A25637A48F5A499A9E0229C">1313</mods:start>
<mods:end id="DCD225CDFABAADA2429BAC51AB6ED925">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8085AFFBBF40A95C0CD0BC9CF11B25CD">journal article</mods:classification>
<mods:identifier id="F084D0340C65E5EA327B8D0D22EF5D0A" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="89234DC7C6BD804FF5D786840EAE3DE2" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFF9723D7A70FA58FB71C2D9" LSID="urn:lsid:plazi:treatment:095A87E6FFF9723D7A70FA58FB71C2D9" httpUri="http://treatment.plazi.org/id/095A87E6FFF9723D7A70FA58FB71C2D9" lastPageNumber="1338" pageId="25" pageNumber="1338">
<subSubSection id="C9E9657BFFF9723D7A70FA58FB11C2F4" box="[861,1221,1460,1482]" pageId="25" pageNumber="1338" type="nomenclature">
<paragraph id="814C36F0FFF9723D7A70FA58FB11C2F4" blockId="25.[861,1221,1460,1512]" box="[861,1221,1460,1482]" pageId="25" pageNumber="1338">
<heading id="DA04819CFFF9723D7A70FA58FB11C2F4" box="[861,1221,1460,1482]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="1338" reason="2">
<taxonomicName id="46F34D73FFF9723D7A70FA58FB11C2F4" authority="Maddison &amp; Sproul" authorityName="Maddison &amp; Sproul" authorityYear="2020" box="[861,1221,1460,1482]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1338" phylum="Arthropoda" rank="species" species="lindrothi">
<emphasis id="B387EAE2FFF9723D7A70FA58FC35C2F7" box="[861,993,1460,1481]" italics="true" pageId="25" pageNumber="1338">L. lindrothi</emphasis>
Maddison &amp; Sproul
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFF9723D7A70FA3FFB71C2D9" box="[861,1189,1490,1512]" pageId="25" pageNumber="1338" type="reference_group">
<paragraph id="814C36F0FFF9723D7A70FA3FFB71C2D9" blockId="25.[861,1221,1460,1512]" box="[861,1189,1490,1512]" pageId="25" pageNumber="1338">
<taxonomicName id="46F34D73FFF9723D7A70FA3FFB71C2D9" ID-CoL="3V6TP" authority="(Lindroth, 1963)" baseAuthorityName="Lindroth" baseAuthorityYear="1963" box="[861,1189,1490,1512]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1338" phylum="Arthropoda" rank="species" species="disjuncta">
<emphasis id="B387EAE2FFF9723D7A70FA3FFC31C2D9" box="[861,997,1490,1512]" italics="true" pageId="25" pageNumber="1338">L. disjuncta</emphasis>
(
<bibRefCitation id="E5624B01FFF9723D7AD9FA3EFB49C2D6" author="Lindroth CH" box="[1012,1181,1490,1512]" pageId="25" pageNumber="1338" pagination="201 - 408" refId="ref26528" refString="Lindroth CH. 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica Supplementum XXIV: 201-408." type="book chapter" year="1963">Lindroth, 1963</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,78 @@
<document id="DAEFDE57BEA5E59211CD06FE958FAD46" ID-ISSN="0024-4082" ID-ZooBank="BF69699-4A1E-47DD-848A-D2FC000FFE0A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353624339" checkinUser="plazi" docAuthor="Maddison, David R. &amp; Sproul, John S." docDate="2020" docId="095A87E6FFFB723F783CF930FD64C00E" docLanguage="en" docName="zlz167.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Lionepha erasa" docType="treatment" docVersion="1" lastPageNumber="1340" masterDocId="F563FF9EFFE07224792DFFECFFD4C73E" masterDocTitle="Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)" masterLastPageNumber="1358" masterPageNumber="1313" pageNumber="1340" updateTime="1739358424447" updateUser="GgImagineBatch">
<mods:mods id="875639FE20714507FCFD284758626B64" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4E3C39B18756A85ED283CAAA65CADC7F">
<mods:title id="A5B33EB929F90B7D5C2934382A5989F6">Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="1462E2C702CE6F89A8C5606431E7DC23" type="personal">
<mods:role id="76C7DECB92F938CAF9413E2678485E06">
<mods:roleTerm id="7938C9540ED13469B5FEED89110A7AAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6ABE00BB20316907875012EFBA06B6B4">Maddison, David R.</mods:namePart>
</mods:name>
<mods:name id="44019B9213DD658F96E600C5E9AABC33" type="personal">
<mods:role id="1CDD4829D8AA19D0135D2A6E4F512BCD">
<mods:roleTerm id="526C9A17B03664A94A590DA388FF09E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E7B631CD24C6BE910D881960767CB23">Sproul, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="38D05AD28D528225BCA3257FB6057795">text</mods:typeOfResource>
<mods:relatedItem id="D9B186F39F54D723A5057A7B02B31874" type="host">
<mods:titleInfo id="9C8E8C359C916F9FFC3F0ED74D13A821">
<mods:title id="F0BF4DFE3995E98BD65003329B9308CC">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="3DB70978384B620F5D0C1C08B10B1A16">
<mods:date id="3F2666B788936ADD849F84A04327E272">2020</mods:date>
<mods:detail id="E4DAD108E6385CBA0EF162671F16E52E" type="volume">
<mods:number id="C171C59CDF6F7D334F8E5A2A7F412A53">189</mods:number>
</mods:detail>
<mods:extent id="AA0CAE14B81F5DEBECFB32C8B1F931E0" unit="page">
<mods:start id="68B74AB70FA89401CFAAD345765F07F3">1313</mods:start>
<mods:end id="4A21A64687E8666B89EBF45C21FCF3F7">1358</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="DED30BC1EDB15A51C795FFEFB6BFC8E5">journal article</mods:classification>
<mods:identifier id="7F1B7C938BF694C1BF3BC3018E67B129" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="0044B3EC6457E145322AC90B25846CC3" type="ZooBank">BF69699-4A1E-47DD-848A-D2FC000FFE0A</mods:identifier>
</mods:mods>
<treatment id="095A87E6FFFB723F783CF930FD64C00E" LSID="urn:lsid:plazi:treatment:095A87E6FFFB723F783CF930FD64C00E" httpUri="http://treatment.plazi.org/id/095A87E6FFFB723F783CF930FD64C00E" lastPageNumber="1340" pageId="27" pageNumber="1340">
<subSubSection id="C9E9657BFFFB723F783CF930FD49C1CA" box="[273,669,1755,1780]" pageId="27" pageNumber="1340" type="nomenclature">
<paragraph id="814C36F0FFFB723F783CF930FD49C1CA" blockId="27.[273,669,1755,1780]" box="[273,669,1755,1780]" pageId="27" pageNumber="1340">
<heading id="DA04819CFFFB723F783CF930FD49C1CA" box="[273,669,1755,1780]" centered="true" fontSize="9" level="2" pageId="27" pageNumber="1340" reason="2">
<taxonomicName id="46F34D73FFFB723F783CF930FD49C1CA" ID-CoL="3V6TQ" authority="(LECONTE, 1859)" baseAuthorityName="LeConte" baseAuthorityYear="1859" box="[273,669,1755,1780]" class="Insecta" family="Carabidae" genus="Lionepha" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1340" phylum="Arthropoda" rank="species" species="erasa">
<emphasis id="B387EAE2FFFB723F783CF930FE01C1CC" box="[273,469,1756,1779]" italics="true" pageId="27" pageNumber="1340">
<smallCapsWord id="87AAA02CFFFB723F783CF930FE52C1CC" baselines="1773,1773" box="[273,390,1756,1779]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Lionepha" pageId="27" pageNumber="1340">LIONEPHA</smallCapsWord>
<smallCapsWord id="87AAA02CFFFB723F78A0F932FE01C1CC" baselines="1773" box="[397,469,1758,1778]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="erasa" pageId="27" pageNumber="1340">ERASA</smallCapsWord>
</emphasis>
(
<bibRefCitation id="E5624B01FFFB723F78C9F937FD41C1CA" author="LeConte JL" box="[484,661,1755,1780]" pageId="27" pageNumber="1340" pagination="69 - 90" refId="ref26498" refString="LeConte JL. 1859. Catalogue of the Coleoptera of Fort Tejon, California. Proceedings of the Academy of Natural Sciences of Philadelphia 11: 69-90." type="journal article" year="1859">
<smallCapsWord id="87AAA02CFFFB723F78C9F937FD9BC1CC" baselines="1774,1773" box="[484,591,1755,1780]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Leconte" pageId="27" pageNumber="1340">LECONTE</smallCapsWord>
, 1859
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9E9657BFFFB723F79D0F8FBFD64C00E" box="[253,688,1815,1840]" pageId="27" pageNumber="1340" type="description">
<paragraph id="814C36F0FFFB723F79D0F8FBFD64C00E" blockId="27.[253,688,1815,1840]" box="[253,688,1815,1840]" pageId="27" pageNumber="1340">
(
<figureCitation id="19C82A75FFFB723F7828F8FBFEB6C00E" box="[261,354,1815,1840]" captionStart="Figure 1" captionStartId="1.[162,240,1424,1446]" captionTargetBox="[408,1210,236,1367]" captionTargetId="figure-189@1.[395,1210,195,1384]" captionTargetPageId="1" captionText="Figure 1. Adults of three species of the Lionepha erasa species group.A, L. erasa, USA: Oregon: Benton Co., Marys Peak; B, L. australerasa, holotype, USA: California: Amador Co., Oyster Lake, Silver Lake Campground; C, L. probata, USA: Oregon: Klamath Co., Odell Creek near Davis Lake. Images copyright David R. Maddison 2019, released under a Creative Commons CC-BY 4.0 license. Scale bar 1 mm." pageId="27" pageNumber="1340">
<smallCapsWord id="87AAA02CFFFB723F7828F8FBFEEFC010" baselines="1834,1833" box="[261,315,1815,1840]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="27" pageNumber="1340">FIGS</smallCapsWord>
1A
</figureCitation>
,
<figureCitation id="19C82A75FFFB723F7842F8FBFE68C00E" box="[367,444,1815,1840]" captionStart="Figure 15" captionStartId="23.[162,242,1038,1060]" captionTargetBox="[166,1440,198,995]" captionTargetId="figure-385@23.[163,1443,195,999]" captionTargetPageId="23" captionText="Figure 15. Microsculpture of the elytral disc of three species of the Lionepha erasa species group. Left elytron, region around anterior discal seta on third interval. A, L. erasa male; B, L. erasa female; C, L.australerasa, male; D, L.australerasa, female; E, L. probata, male; F, L. probata, female. Scale bar 50 µm." pageId="27" pageNumber="1340">15A, B</figureCitation>
,
<figureCitation id="19C82A75FFFB723F78E7F8FBFDC2C00E" box="[458,534,1815,1840]" captionStart="Figure 11" captionStartId="19.[165,245,1848,1870]" captionTargetBox="[232,1370,923,1806]" captionTargetId="figure-191@19.[227,1379,918,1809]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of three species of the Lionepha erasa species group, left lateral view. A, L. erasa; B, L. erasa; C, L. australerasa; D, L. australerasa, holotype; E, L. probata; F, L. probata. Scale bar 0.1 mm." pageId="27" pageNumber="1340">11A, B</figureCitation>
,
<figureCitation id="19C82A75FFFB723F7B09F8FBFD86C00E" box="[548,594,1815,1840]" captionStart="Figure 14" captionStartId="22.[145,223,1675,1697]" captionTargetBox="[230,1339,203,1631]" captionTargetId="figure-78@22.[226,1344,195,1635]" captionTargetPageId="22" captionText="Figure 14. Aedeagus of six species of the Lionepha erasa species group, left lateral closeup view, with sclerite CH1 outlined, and the nub, when present, marked with an arrow. A, L. erasa; B, L. australerasa, holotype; C, L. probata; D, L. casta; E, L. kavanaughi, holotype; F, L. lindrothi." pageId="27" pageNumber="1340">14A</figureCitation>
,
<figureCitation id="19C82A75FFFB723F7B72F8FBFDABC00E" box="[607,639,1815,1840]" captionStart="Figure 9" captionStartId="18.[145,223,1675,1697]" captionTargetBox="[340,1230,195,1634]" captionTargetId="figure-75@18.[340,1230,195,1635]" captionTargetPageId="18" captionText="Figure 9. Female reproductive tract of the Lionepha erasa species group, dorsal view.Arrows in A and B indicate the dorsal microtrichial patch of the bursa. A, L. probata; B, L. erasa; C, L. kavanaughi; D, L. lindrothi; E, L. kavanaughi, closeup; F, L. lindrothi, closeup. Scale bars 100 µm." pageId="27" pageNumber="1340">9B</figureCitation>
,
<figureCitation id="19C82A75FFFB723F7BA1F8FBFD7CC00E" box="[652,680,1815,1840]" captionStart="Figure 20" captionStartId="31.[163,244,1506,1528]" captionTargetBox="[163,787,195,1467]" captionTargetId="figure-541@31.[163,787,195,1467]" captionTargetPageId="31" captionText="Figure 20. Geographic distribution of L. erasa (circles) and L. australerasa (stars). Specimens whose DNA was sequenced are shown in black." pageId="27" pageNumber="1340">20</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,42 +1,45 @@
<document id="E75F7D02600F9E6291F0A6006A4BEEDB" ID-DOI="10.11646/zootaxa.5584.3.11" ID-ISSN="1175-5326" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739311124461" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2025" docId="4C478527C95D083BFF12FF36FE06BB7E" docLanguage="en" docName="zootaxa.5584.3.11.pdf" docOrigin="Zootaxa 5584 (3)" docSource="https://doi.org/10.11646/zootaxa.5584.3.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hapalogenius fuscipennis" docType="treatment" docVersion="2" lastPageNumber="449" masterDocId="B07EFD5FC95F0839FF85FF8DFF89B85D" masterDocTitle="Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar" masterLastPageNumber="450" masterPageNumber="447" pageNumber="449" updateTime="1739311747773" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7F942C045EF691BE15C89FD6FA213EDF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="285191564E902AE346FE9FF508ED169D">
<mods:title id="9F020B23557B6C063A04BD2C08206956">Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar</mods:title>
<document id="561B792243313F3318CCFDCF991D8926" ID-CLB-Dataset="308096" ID-DOI="10.11646/zootaxa.5584.3.11" ID-GBIF-Dataset="dc8d6b61-cb06-4121-8eae-3209883c1b75" ID-ISSN="1175-5326" ID-Zenodo-Dep="14858040" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739311124461" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2025" docId="4C478527C95D083BFF12FF36FE06BB7E" docLanguage="en" docName="zootaxa.5584.3.11.pdf" docOrigin="Zootaxa 5584 (3)" docSource="https://doi.org/10.11646/zootaxa.5584.3.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hapalogenius fuscipennis" docType="treatment" docVersion="4" lastPageNumber="449" masterDocId="B07EFD5FC95F0839FF85FF8DFF89B85D" masterDocTitle="Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar" masterLastPageNumber="450" masterPageNumber="447" pageNumber="449" updateTime="1739358349669" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F21C493CB2EB6CE84DE1952BD5D39C1D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F41157DA664EE0133E5B78274B193699">
<mods:title id="6D4A8B5233300C470D02C2F46440D5AB">Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="72070F423AA07FD7D2F25B4DBA9889F2" type="personal">
<mods:role id="E72956FA98909AB1D85A4C6E09B57526">
<mods:roleTerm id="97269746BE6E43B8CEFE81BA4F9FC2C4">Author</mods:roleTerm>
<mods:name id="001CFB0D40172FB1FE4CC812D72008F6" type="personal">
<mods:role id="E556A67762047430D7B298CA09335334">
<mods:roleTerm id="DC9D9583B8D8D28B4C63C46C7723BB03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26C2444C83049766C59BB4BDCAFF16E3">Jordal, Bjarte H.</mods:namePart>
<mods:namePart id="80CB3B11732381E2EFB75269C96C0D44">Jordal, Bjarte H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4D44352187D8F56B3332EC3E7D9A8FD2">text</mods:typeOfResource>
<mods:relatedItem id="C751DAFE8B6433DD2D734220AC969A01" type="host">
<mods:titleInfo id="EC87258472C1E21E883BD9CD9655C763">
<mods:title id="5D451BBCA4615DF9E2478932B729C3E4">Zootaxa</mods:title>
<mods:typeOfResource id="64B0E3EF5A00B0FC909D6C58664095AB">text</mods:typeOfResource>
<mods:relatedItem id="A417D3757ED5A7395C6126E15E48367D" type="host">
<mods:titleInfo id="841E0D07060B9346CA74DB1106D334AA">
<mods:title id="550DAFF493273DDB1E2A8B9A7191C1AE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A587A7E426BC7DDC3E91504D235B44FA">
<mods:date id="C0CB123EB6FD5B238CF75DBF1234DE76">2025</mods:date>
<mods:detail id="22562D71B50D57EF701A0905B313FB0C" type="pubDate">
<mods:number id="44584531539CD3ECA1A1DD35CABDD342">2025-02-10</mods:number>
<mods:part id="076039814BC68F123E90E3FE36F948FF">
<mods:date id="CCA3F711C076553BBFC179B3B5E7CADF">2025</mods:date>
<mods:detail id="AC5070BF4B6204263C7219A8001F1ADF" type="pubDate">
<mods:number id="2BDE9ADB9E4E3451898E16D228BAAC6E">2025-02-10</mods:number>
</mods:detail>
<mods:detail id="C7C57636F3FA240391438964646A21BC" type="volume">
<mods:number id="DDE334C7F6DC48EE3E4E4E0F17398010">5584</mods:number>
<mods:detail id="08942B174A5F816BE115C0978EABB433" type="volume">
<mods:number id="88F68465059E0821DCAB624B0ADFC926">5584</mods:number>
</mods:detail>
<mods:detail id="0BDC6D55946F910AD6874939FE1FCB9B" type="issue">
<mods:number id="57DAD8E87F45C9FA5899D50F8571AE05">3</mods:number>
<mods:detail id="1AB94ABF53472B2CFB7A6F0E9EB2DC90" type="issue">
<mods:number id="C9ACD669FD3C78007A50C7B65CAD5465">3</mods:number>
</mods:detail>
<mods:extent id="9680D56C2EFFE6E8537307419E94696A" unit="page">
<mods:start id="DEC05008B0D10880DF494169BF86544E">447</mods:start>
<mods:end id="21571620C52DA3E41818AD26513C05E5">450</mods:end>
<mods:extent id="9B016C617E3E7BA6782F5AB0CAC94FBD" unit="page">
<mods:start id="FFD3580B15BF189DBCACB7E1EAB8A13A">447</mods:start>
<mods:end id="60ECC052BF6976CCF8CB950869E3D7B6">450</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="80892FA2E3431E43B4DE70CED50B0F44">
<mods:url id="4BD3BBC5741592D3D1F063D8169D428A">https://doi.org/10.11646/zootaxa.5584.3.11</mods:url>
<mods:location id="3D1158C2B1B5F6D30643A929381EC18C">
<mods:url id="726DDD675828AA5F7C58B7F3E94C3339">https://doi.org/10.11646/zootaxa.5584.3.11</mods:url>
</mods:location>
<mods:classification id="72A827553855A149A8110C5B3528F51A">journal article</mods:classification>
<mods:identifier id="D8817A30555A1319B53EB94F2BE05E6C" type="DOI">10.11646/zootaxa.5584.3.11</mods:identifier>
<mods:identifier id="33CBE7A4F6B8DE1BCF4C9F69C6DF3114" type="ISSN">1175-5326</mods:identifier>
<mods:classification id="12CF9E62F71CB338ECAB203AD3677E4C">journal article</mods:classification>
<mods:identifier id="91D83E7D481D933D3DB5C2EDCFAFF1CE" type="CLB-Dataset">308096</mods:identifier>
<mods:identifier id="335E2E4D6741E776E3CDA22AE2F4F19D" type="DOI">10.11646/zootaxa.5584.3.11</mods:identifier>
<mods:identifier id="5F825B8E5B1240FD2CED2A48D634AE5F" type="GBIF-Dataset">dc8d6b61-cb06-4121-8eae-3209883c1b75</mods:identifier>
<mods:identifier id="E95602CA59E5CE646E93B107F60D4C0D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="64128F84B0AF50CBA87D5940C4B55146" type="Zenodo-Dep">14858040</mods:identifier>
</mods:mods>
<treatment id="4C478527C95D083BFF12FF36FE06BB7E" ID-DOI="http://doi.org/10.5281/zenodo.14853372" ID-Zenodo-Dep="14853372" LSID="urn:lsid:plazi:treatment:4C478527C95D083BFF12FF36FE06BB7E" httpUri="http://treatment.plazi.org/id/4C478527C95D083BFF12FF36FE06BB7E" lastPageNumber="449" pageId="2" pageNumber="449">
<subSubSection id="8CF467BAC95D083BFF12FF36FDDFB888" box="[151,598,187,214]" pageId="2" pageNumber="449" type="nomenclature">
@ -54,7 +57,7 @@
<subSubSection id="8CF467BAC95D083BFF12FF6CFE8FB8A4" box="[151,262,225,249]" pageId="2" pageNumber="449" type="description">
<paragraph id="C4513431C95D083BFF12FF6CFE8FB8A4" blockId="2.[151,1437,151,804]" box="[151,262,225,249]" pageId="2" pageNumber="449">
(
<figureCitation id="5CD528B4C95D083BFF1AFF6CFF77B8A4" box="[159,254,225,249]" captionStart="FIGURES 16" captionStartId="1.[151,264,1970,1994]" captionTargetBox="[193,1435,318,1941]" captionTargetId="figure-88@1.[151,1436,308,1944]" captionTargetPageId="1" captionText="FIGURES 16. Dorsal, lateral and front view of Glochiphorus globosus, syntype (13), and Glochiphorus alienus, paratype (46)." pageId="2" pageNumber="449">Figs 46</figureCitation>
<figureCitation id="5CD528B4C95D083BFF1AFF6CFF77B8A4" box="[159,254,225,249]" captionStart="FIGURES 16" captionStartId="1.[151,264,1970,1994]" captionTargetBox="[193,1435,318,1941]" captionTargetId="figure-88@1.[151,1436,308,1944]" captionTargetPageId="1" captionText="FIGURES 16. Dorsal, lateral and front view of Glochiphorus globosus, syntype (13), and Glochiphorus alienus, paratype (46)." figureDoi="http://doi.org/10.5281/zenodo.14858042" httpUri="https://zenodo.org/record/14858042/files/figure.png" pageId="2" pageNumber="449">Figs 46</figureCitation>
)
</paragraph>
</subSubSection>
@ -152,7 +155,7 @@
]
</materialsCitation>
.
<materialsCitation id="74863E6CC95D083BFED8FE45FE48BA5E" collectingDate="1973-02-08" collectionCode="NHMW" collectorName="S. Afr. Transvaal &amp; Soutpan &amp; Pretoria &amp; Endrody-Younga" country="Democratic Republic of the Congo" latitude="-25.25" longLatPrecision="748" longitude="28.05" pageId="2" pageNumber="449" specimenCount="1" stateProvince="Sud-Kivu" typeStatus="paratype">
<materialsCitation id="74863E6CC95D083BFED8FE45FE48BA5E" collectingDate="1973-02-08" collectionCode="NHMW" collectorName="Endrody-Younga" country="Democratic Republic of the Congo" latitude="-25.25" longLatPrecision="748" longitude="28.05" pageId="2" pageNumber="449" specimenCount="1" stateProvince="Sud-Kivu" typeStatus="paratype">
<typeStatus id="1B558A93C95D083BFED8FE45FE33B9BD" box="[349,442,456,480]" pageId="2" pageNumber="449" type="paratype">Paratype</typeStatus>
<taxonomicName id="03EE4FB2C95D083BFE45FE45FDA4B9BD" authorityName="Schedl" authorityYear="1982" box="[448,557,456,480]" class="Insecta" family="Curculionidae" genus="Glochiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="alienus">
<emphasis id="F69AE823C95D083BFE45FE45FDA4B9BD" box="[448,557,456,480]" italics="true" pageId="2" pageNumber="449">G. alienus</emphasis>

View file

@ -1,42 +1,45 @@
<document id="08FDFEEF395CC6620D2126F13CD3B246" ID-DOI="10.11646/zootaxa.5584.3.11" ID-ISSN="1175-5326" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739311124461" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2025" docId="4C478527C95F0838FF12FB6CFAE9B946" docLanguage="en" docName="zootaxa.5584.3.11.pdf" docOrigin="Zootaxa 5584 (3)" docSource="https://doi.org/10.11646/zootaxa.5584.3.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glochiphorus Strohmeyer" docType="treatment" docVersion="2" lastPageNumber="447" masterDocId="B07EFD5FC95F0839FF85FF8DFF89B85D" masterDocTitle="Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar" masterLastPageNumber="450" masterPageNumber="447" pageNumber="447" updateTime="1739311747773" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B554FACAD12AD60CBFD29E32D2627CA4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="335B84217DAA0CD5CD34D729816F4482">
<mods:title id="911E550537381FE4E77F79EF941BD671">Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar</mods:title>
<document id="68838A6AAD00642FB4E0D5527F5F8ACD" ID-CLB-Dataset="308096" ID-DOI="10.11646/zootaxa.5584.3.11" ID-GBIF-Dataset="dc8d6b61-cb06-4121-8eae-3209883c1b75" ID-ISSN="1175-5326" ID-Zenodo-Dep="14858040" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739311124461" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2025" docId="4C478527C95F0838FF12FB6CFAE9B946" docLanguage="en" docName="zootaxa.5584.3.11.pdf" docOrigin="Zootaxa 5584 (3)" docSource="https://doi.org/10.11646/zootaxa.5584.3.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glochiphorus Strohmeyer" docType="treatment" docVersion="4" lastPageNumber="447" masterDocId="B07EFD5FC95F0839FF85FF8DFF89B85D" masterDocTitle="Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar" masterLastPageNumber="450" masterPageNumber="447" pageNumber="447" updateTime="1739358349669" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E307E3DBD67433B652FCA7E3B51D52DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C89C49C806D3A6D2C4904AE15404DC15">
<mods:title id="89D486700C3DDACB4BE470D34BD88ADB">Glochiphorus Strohmeyer (Coleoptera, Scolytinae) - a monotypic genus endemic to Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="8A15A2CE9C0D4D3344AA032D5A2BAE45" type="personal">
<mods:role id="A50BFAE31D62D7172ED23238632A9F38">
<mods:roleTerm id="8BFE7FDEEA42B824BDC0B4F0AB07F7FE">Author</mods:roleTerm>
<mods:name id="94F5C58DF9654897BD3D690344A00999" type="personal">
<mods:role id="6F539BF317CF247CF3769ECA42268FAE">
<mods:roleTerm id="7DFF9909F34F0338B8BC2259B37828DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31131E534B112D9933329AAA43FE4FF7">Jordal, Bjarte H.</mods:namePart>
<mods:namePart id="5C7661C2AE5F9AFAE02DEDBAD4BECFDC">Jordal, Bjarte H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EAD336B4AA2BA6913ED221E5E5BCDEE5">text</mods:typeOfResource>
<mods:relatedItem id="F147A823F73C5E95620E86950EF53CF1" type="host">
<mods:titleInfo id="8CBD1E206C176A61C729912713EDACD3">
<mods:title id="D63E0166FB3F45E82246C8B889AC9E66">Zootaxa</mods:title>
<mods:typeOfResource id="89FD4908FC3DCFEC1170C716379D751B">text</mods:typeOfResource>
<mods:relatedItem id="6D8DC432C231F297996A207010CDD865" type="host">
<mods:titleInfo id="AFF4E9710A39D1B26BB61228C20DE499">
<mods:title id="5D2F126818AB6BC290F4AD928BC6B745">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="20FB7274D96402F39E3F54FDFAD8EC16">
<mods:date id="0342427DA4830FE16A87CEDCB86771C9">2025</mods:date>
<mods:detail id="7CC4F5C8A4702E653A7578CAE5B00051" type="pubDate">
<mods:number id="E72C78C581F99A4BE99F9A50948A5C7F">2025-02-10</mods:number>
<mods:part id="7F2FE166889B21B7B3895F10B110F378">
<mods:date id="467B3EDE90A341595291CD0DDA63E944">2025</mods:date>
<mods:detail id="50F032062FD5477333014C71AE765D6F" type="pubDate">
<mods:number id="1392B5E1B7FE11CC1A9B9E0B0DDB58A2">2025-02-10</mods:number>
</mods:detail>
<mods:detail id="356157509AE1909B8441045FE379ED59" type="volume">
<mods:number id="84F1F3EBCA6E5FAE25C0164E9525F920">5584</mods:number>
<mods:detail id="9B49EEA2D6C7443C6B9484A7C10E80B1" type="volume">
<mods:number id="7A648A4644FBE39659E3747BCC0AD510">5584</mods:number>
</mods:detail>
<mods:detail id="2C7472EA173FE9037ACBF5AD8586D517" type="issue">
<mods:number id="40ECCBDF38AC05E237B70D43B699383D">3</mods:number>
<mods:detail id="B92944EFF41070C90020EEF514E5845B" type="issue">
<mods:number id="72446795E18818F90EBB19849F7620AD">3</mods:number>
</mods:detail>
<mods:extent id="0376B9BD4EE3902B7F0151E36B7EF572" unit="page">
<mods:start id="C7CF10727A78E8DB07213DF7EE4BBDAB">447</mods:start>
<mods:end id="AB6E550B3938B611CA1EEEE42A032F3B">450</mods:end>
<mods:extent id="AF2CEC8B1DB2B51B6DEFA9EFC8CB312C" unit="page">
<mods:start id="A623AF2C4FD28E3F9EA2873F9C3517DF">447</mods:start>
<mods:end id="EFBCD68A4554C492D7DD2D7F715CF30B">450</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E45653908F583ABF2C8451A03772634D">
<mods:url id="85D0D4210D581A38D4A7D62F58E25F19">https://doi.org/10.11646/zootaxa.5584.3.11</mods:url>
<mods:location id="61DA6620994910BB839F54F7663E236E">
<mods:url id="65CE6A98795BA647BC527935A906A3C5">https://doi.org/10.11646/zootaxa.5584.3.11</mods:url>
</mods:location>
<mods:classification id="58A47F24B43B9242DE68B3D2A5ED4A6A">journal article</mods:classification>
<mods:identifier id="CB9D766792ED3682B2D7621FD5D91183" type="DOI">10.11646/zootaxa.5584.3.11</mods:identifier>
<mods:identifier id="1058279EB8E8003C8E638D562C3E247F" type="ISSN">1175-5326</mods:identifier>
<mods:classification id="7DEE609059D297EAA9D86825905C57AD">journal article</mods:classification>
<mods:identifier id="9CF7D19495BC4EEF688916EF4695EFCA" type="CLB-Dataset">308096</mods:identifier>
<mods:identifier id="C852276AB71845A1928E3B50E125A05B" type="DOI">10.11646/zootaxa.5584.3.11</mods:identifier>
<mods:identifier id="8F0EDCAECECD18A08818FE9DAFCBE91E" type="GBIF-Dataset">dc8d6b61-cb06-4121-8eae-3209883c1b75</mods:identifier>
<mods:identifier id="8E8B4FF3A591EB0EF5FCE1D5C428D9C9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="95302504D81BCA09A5B8B4584FCAB27F" type="Zenodo-Dep">14858040</mods:identifier>
</mods:mods>
<treatment id="4C478527C95F0838FF12FB6CFAE9B946" ID-DOI="http://doi.org/10.5281/zenodo.14853368" ID-Zenodo-Dep="14853368" LSID="urn:lsid:plazi:treatment:4C478527C95F0838FF12FB6CFAE9B946" httpUri="http://treatment.plazi.org/id/4C478527C95F0838FF12FB6CFAE9B946" lastPageId="1" lastPageNumber="447" pageId="0" pageNumber="447">
<subSubSection id="8CF467BAC95F0839FF12FB6CFE50BCA1" box="[151,473,1249,1276]" pageId="0" pageNumber="447" type="nomenclature">
@ -62,20 +65,22 @@ species:
<bibRefCitation id="A07F49C0C95F0839FDB6FA85FCA0BD7D" author="Strohmeyer, H." box="[563,809,1288,1312]" pageId="0" pageNumber="447" pagination="126 - 132" refId="ref1907" refString="Strohmeyer, H. (1910) Neue Borkenkafer aus Abyssenyen, Madagaskar, Indien und Tasmania. Entomologische Blatter, 6, 126-132." type="journal article" year="1910">Strohmeyer, 1910: 127</bibRefCitation>
</taxonomicName>
, monotypic (
<figureCitation id="5CD528B4C95F0839FC31FA85FB9BBD7D" box="[948,1042,1288,1312]" captionStart="FIGURES 16" captionStartId="1.[151,264,1970,1994]" captionTargetBox="[193,1435,318,1941]" captionTargetId="figure-88@1.[151,1436,308,1944]" captionTargetPageId="1" captionText="FIGURES 16. Dorsal, lateral and front view of Glochiphorus globosus, syntype (13), and Glochiphorus alienus, paratype (46)." pageId="0" pageNumber="447">Figs 13</figureCitation>
<figureCitation id="5CD528B4C95F0839FC31FA85FB9BBD7D" box="[948,1042,1288,1312]" captionStart="FIGURES 16" captionStartId="1.[151,264,1970,1994]" captionTargetBox="[193,1435,318,1941]" captionTargetId="figure-88@1.[151,1436,308,1944]" captionTargetPageId="1" captionText="FIGURES 16. Dorsal, lateral and front view of Glochiphorus globosus, syntype (13), and Glochiphorus alienus, paratype (46)." figureDoi="http://doi.org/10.5281/zenodo.14858042" httpUri="https://zenodo.org/record/14858042/files/figure.png" pageId="0" pageNumber="447">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8CF467BAC95F0839FF42FAA1FECDBD3A" pageId="0" pageNumber="447" type="materials_examined">
<paragraph id="C4513431C95F0839FF42FAA1FECDBD3A" blockId="0.[151,1437,1214,2032]" pageId="0" pageNumber="447">
<emphasis id="F69AE823C95F0839FF42FAA1FEE5BD1E" bold="true" box="[199,364,1323,1348]" inLineHeading="true" pageId="0" pageNumber="447" reason="1">Type material.</emphasis>
<materialsCitation id="74863E6CC95F0839FEF7FAA1FEB6BD3A" collectionCode="NHMW" collectorName="de l'Androy &amp; Dr J. Decorse" country="Madagascar" location="Madagascar" pageId="0" pageNumber="447" specimenCount="1" typeStatus="syntype">
<materialsCitation id="74863E6CC95F0839FEF7FAA1FEB6BD3A" collectionCode="NHMW" collectorName="Dr J. Decorse" country="Madagascar" location="region de l'Androy" municipality="Ambovombe" pageId="0" pageNumber="447" specimenCount="1" typeStatus="syntype">
<typeStatus id="1B558A93C95F0839FEF7FAA1FE51BD19" box="[370,472,1324,1348]" pageId="0" pageNumber="447" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="BCF974A1C95F0839FE64FAA1FDEDBD19" box="[481,612,1324,1348]" name="Madagascar" pageId="0" pageNumber="447">Madagascar</collectingCountry>
, region
<collectorName id="691B51E7C95F0839FD32FAA1FCB0BD19" box="[695,825,1324,1348]" pageId="0" pageNumber="447">de lAndroy</collectorName>
, Ambovombe [-25.16, 46.08], fev. &amp; mars,
,
<location id="C13162EAC95F0839FDE8FAA1FCB0BD19" LSID="urn:lsid:plazi:treatment:4C478527C95F0838FF12FB6CFAE9B946:C13162EAC95F0839FDE8FAA1FCB0BD19" box="[621,825,1324,1348]" country="Madagascar" municipality="Ambovombe" name="region de l'Androy" pageId="0" pageNumber="447">region de lAndroy</location>
,
<collectingMunicipality id="2435AE4BC95F0839FCC4FAA1FC45BD19" box="[833,972,1324,1348]" pageId="0" pageNumber="447">Ambovombe</collectingMunicipality>
[-25.16, 46.08], fev. &amp; mars,
<collectorName id="691B51E7C95F0839FA80FAA1FA11BD1E" box="[1285,1432,1324,1347]" pageId="0" pageNumber="447">Dr J. Decorse</collectorName>
, 1901 [
<collectionCode id="A2FFACF4C95F0839FF5CFADDFEB0BD3A" box="[217,313,1360,1383]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="447" type="Museum">NHMW</collectionCode>

View file

@ -0,0 +1,590 @@
<document id="F0CF66759023DE8234A1BD259E7515F5" ID-ISSN="0024-4082" ID-ZooBank="A81CEE6-9D1C-49A9-A9D1-FB0501198097" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353570418" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, García-París, Mario, Ruiz, José L. &amp; Recuero, Ernesto" docDate="2020" docId="5568A03458681100FF449BEDFA25FC3A" docLanguage="en" docName="zlz164.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Berberomeloe comunero SANCHEZ-VIALAS ET AL. 2020, SP. NOV." docType="treatment" docVersion="1" lastPageNumber="1274" masterDocId="A951D84C587E1119FFFC9D51FFAAFF8E" masterDocTitle="Patterns of morphological diversification in giant Berberomeloe blister beetles (Coleoptera: Meloidae) reveal an unexpected taxonomic diversity concordant with mtDNA phylogenetic structure" masterLastPageNumber="1312" masterPageNumber="1249" pageNumber="1271" updateTime="1739358454141" updateUser="GgImagineBatch">
<mods:mods id="2F81B0CDDFB1E1D51852EA78339BB9F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C933CC10589FC70FB80314E746E8BCBD">
<mods:title id="89B3353030783793E00CCD93F1C2C117">Patterns of morphological diversification in giant Berberomeloe blister beetles (Coleoptera: Meloidae) reveal an unexpected taxonomic diversity concordant with mtDNA phylogenetic structure</mods:title>
</mods:titleInfo>
<mods:name id="AB07884B4E8D0BB504097B4D383082ED" type="personal">
<mods:role id="FA591584CE8536CEFFA55E6CE8A301DC">
<mods:roleTerm id="08F70D61DF839E2A6748DF388FFD05D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1E1235B246DCC726FDDEFCA15F736A3">Sánchez-Vialas, Alberto</mods:namePart>
</mods:name>
<mods:name id="03B8DBB7F341535F7010C4842380A32C" type="personal">
<mods:role id="0E1C97D6F27E55C620B1A88369AE81BC">
<mods:roleTerm id="28AFDA8F9D0924560D2B3DA93D5B26B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09642C4A26240F8A182E0A2CC08BAB6F">García-París, Mario</mods:namePart>
</mods:name>
<mods:name id="66CB7CDE104A91BC9408CFBE002C8307" type="personal">
<mods:role id="F26FDF828B1BBC4C313CCAB801D7479C">
<mods:roleTerm id="2E93033D559D6349F161A1E8BAA45E29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="01D2D8AC737BDB135F5981E94D27ABCA">Ruiz, José L.</mods:namePart>
</mods:name>
<mods:name id="F26A6C0D7D183BE38953991FBA3D7B64" type="personal">
<mods:role id="A24618E6B39007379C4CEBB96535DECD">
<mods:roleTerm id="7C05E2B24AFABB808BE5416769784AFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8092EC708E925A97E4CBC095C4CDEAE1">Recuero, Ernesto</mods:namePart>
</mods:name>
<mods:typeOfResource id="CC11BEFB9B9D27D9C30DA55FC6BB28EB">text</mods:typeOfResource>
<mods:relatedItem id="B8304C837122EAF9E91364890AD13B54" type="host">
<mods:titleInfo id="791965A3FBCDD4CE1F1E753C80B613B9">
<mods:title id="B1A8A2C996A8BBC35675EE48AA4FEEAF">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="C2919B073266C5A386B31E294568E438">
<mods:date id="879A9F7EF67DBCEDDED9471E0BA64660">2020</mods:date>
<mods:detail id="CD2118EEB84B3FEEAF0BB30310B04F5C" type="volume">
<mods:number id="0781132E15EFB039633164C18BCC9CF1">189</mods:number>
</mods:detail>
<mods:extent id="BD1E78815BB3495701D2C78287763B0B" unit="page">
<mods:start id="A322FD1F9C130DB0E5E5DE0706978BB9">1249</mods:start>
<mods:end id="A436C1608BC84914D56E9EC0AC672EF9">1312</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E7BB3F59C46E40622A2D5B86D92A2918">journal article</mods:classification>
<mods:identifier id="77582EE30997462E7CAD4E277B45EA7E" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="4FDAA9570D73D46C2568A50D89698833" type="ZooBank">A81CEE6-9D1C-49A9-A9D1-FB0501198097</mods:identifier>
</mods:mods>
<treatment id="5568A03458681100FF449BEDFA25FC3A" LSID="urn:lsid:plazi:treatment:5568A03458681100FF449BEDFA25FC3A" httpUri="http://treatment.plazi.org/id/5568A03458681100FF449BEDFA25FC3A" lastPageId="25" lastPageNumber="1274" pageId="22" pageNumber="1271">
<subSubSection id="95DB42A95868110FFF449BEDFE7BF8B9" pageId="22" pageNumber="1271" type="nomenclature">
<paragraph id="DD7E11225868110FFF449BEDFD8AF97C" blockId="22.[184,722,1723,1779]" pageId="22" pageNumber="1271">
<heading id="8636A64E5868110FFF449BEDFD8AF97C" centered="true" fontSize="9" level="2" pageId="22" pageNumber="1271" reason="2">
<taxonomicName id="1AC16AA15868110FFF449BEDFE1AF97C" authority="SANCHEZ-VIALAS ET AL." authorityName="SANCHEZ-VIALAS ET AL." authorityYear="2020" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="comunero" status="sp. nov.">
<emphasis id="EFB5CD305868110FFF449BEDFDADF95C" bold="true" box="[184,519,1724,1747]" italics="true" pageId="22" pageNumber="1271">
<smallCapsWord id="DB9887FE5868110FFF449BEDFED0F95C" baselines="1741,1741" box="[184,378,1724,1747]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Berberomeloe" pageId="22" pageNumber="1271">BERBEROMELOE</smallCapsWord>
<smallCapsWord id="DB9887FE5868110FFE7D9BEEFDADF95C" baselines="1741" box="[385,519,1727,1746]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="comunero" pageId="22" pageNumber="1271">COMUNERO</smallCapsWord>
</emphasis>
<smallCapsWord id="DB9887FE5868110FFDF29BEAFD78F95D" baselines="1742,1742,1742,1742,1742" box="[526,722,1723,1748]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Sanchez-Vialas" pageId="22" pageNumber="1271">SÁNCHEZ-VIALAS</smallCapsWord>
<emphasis id="EFB5CD305868110FFE959B8FFE1AF97C" box="[361,432,1755,1778]" italics="true" pageId="22" pageNumber="1271">
<smallCapsWord id="DB9887FE5868110FFE959B8FFE2CF97E" baselines="1771" box="[361,390,1758,1776]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="et" pageId="22" pageNumber="1271">ET</smallCapsWord>
<smallCapsWord id="DB9887FE5868110FFE719B8FFE03F97C" baselines="1773" box="[397,425,1758,1778]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="al" pageId="22" pageNumber="1271">AL</smallCapsWord>
.
</emphasis>
</taxonomicName>
,
<emphasis id="EFB5CD305868110FFE419B8CFD8AF97C" bold="true" box="[445,544,1754,1778]" pageId="22" pageNumber="1271">
<taxonomicNameLabel id="F486704B5868110FFE419B8CFD8AF97C" box="[445,544,1754,1778]" pageId="22" pageNumber="1271" rank="species">
<smallCapsWord id="DB9887FE5868110FFE419B8CFE70F97E" baselines="1771" box="[445,474,1757,1776]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="22" pageNumber="1271">SP</smallCapsWord>
.
<smallCapsWord id="DB9887FE5868110FFE149B8CFDB3F97E" baselines="1771" box="[488,537,1757,1776]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="22" pageNumber="1271">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DD7E11225868110FFF6D9A52FE7BF8B9" blockId="22.[145,761,1794,1908]" pageId="22" pageNumber="1271">
LSID:
<uri id="A9501D205868110FFF199A53FE7BF8B9" pageId="22" pageNumber="1271">
urn:lsid:zoobank.org:act:
<uuid id="A9672BF75868110FFDEA9A53FE7BF8B9" pageId="22" pageNumber="1271">CF638D73-6A93- 4FAE-88D1-D548C0214D29</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="95DB42A958681100FF6D9A11FA25FC3A" lastPageId="25" lastPageNumber="1274" pageId="22" pageNumber="1271" type="materials_examined">
<paragraph id="DD7E11225868110FFF6D9A11FAE0FEFA" blockId="22.[145,761,1794,1908]" lastBlockId="22.[809,1425,197,372]" pageId="22" pageNumber="1271">
<materialsCitation id="6DA91B7F5868110FFF6D9A11FC69FF74" collectingDate="1999-04-24" collectionCode="MNCN" collectorName="Montejo de la Sierra &amp; M. Garcia-Paris" country="Spain" location="Montejo de la Sierra" municipality="Male" pageId="22" pageNumber="1271" specimenCount="1" stateProvince="Madrid" typeStatus="holotype">
<emphasis id="EFB5CD305868110FFF6D9A11FF57F8DB" box="[145,253,1856,1877]" italics="true" pageId="22" pageNumber="1271">
<typeStatus id="027AAF805868110FFF6D9A11FF5DF8DB" box="[145,247,1856,1877]" pageId="22" pageNumber="1271" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingMunicipality id="3D1A8B585868110FFEF69A11FEEDF8D8" box="[266,327,1856,1878]" pageId="22" pageNumber="1271">Male</collectingMunicipality>
:
<collectingCountry id="A5D651B25868110FFEAD9A11FE3CF8D8" box="[337,406,1856,1878]" name="Spain" pageId="22" pageNumber="1271">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC05868110FFE5E9A11FE53F8D8" box="[418,505,1856,1878]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
,
<collectorName id="703474F45868110FFDF99A11FD5EF8D8" box="[517,756,1856,1878]" pageId="22" pageNumber="1271">
<location id="D81E47F95868110FFDF99A11FD5EF8D8" LSID="urn:lsid:plazi:treatment:5568A03458681100FF449BEDFA25FC3A:D81E47F95868110FFDF99A11FD5EF8D8" box="[517,756,1856,1878]" country="Spain" municipality="Male" name="Montejo de la Sierra" pageId="22" pageNumber="1271" stateProvince="Madrid">Montejo de la Sierra</location>
</collectorName>
, pastizal junto al pueblo,
<date id="A97F37E25868110FFE5F9A0EFD96F8FA" box="[419,572,1886,1908]" pageId="22" pageNumber="1271" value="1999-04-24">
<collectingDate id="B93BCE0A5868110FFE5F9A0EFD96F8FA" box="[419,572,1886,1908]" pageId="22" pageNumber="1271" value="1999-04-24">24 April 1999</collectingDate>
</date>
,
<collectorName id="703474F45868110FFDBA9A0EFD53F8FA" box="[582,761,1887,1908]" pageId="22" pageNumber="1271">M. García-París</collectorName>
leg. [printed white label];
<collectionCode id="BBD089E75868110FFBB09D94FB0AFF54" box="[1100,1184,197,218]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 231431 [printed white label]
</materialsCitation>
;
<materialsCitation id="6DA91B7F5868110FFC289DB5FAECFEFA" country="Spain" county="Preserved" location="Museo Nacional de Ciencias Naturales" municipality="Entomological Collection" pageId="22" pageNumber="1271" specimenCount="1" stateProvince="Madrid" typeStatus="holotype">
<typeStatus id="027AAF805868110FFC289DB5FBFCFF74" box="[980,1110,228,250]" pageId="22" pageNumber="1271" type="holotype">Holotypus</typeStatus>
,
<taxonomicName id="1AC16AA15868110FFB9A9DB5FAF0FE96" authority="Sanchez-Vialas, Garcia-Paris, Ruiz &amp; Recuero" authorityName="Sanchez-Vialas, Garcia-Paris, Ruiz &amp; Recuero" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="comunero">
<emphasis id="EFB5CD305868110FFB9A9DB5FA3AFF74" box="[1126,1424,228,250]" italics="true" pageId="22" pageNumber="1271">Berberomeloe comunero</emphasis>
Sánchez-Vialas, García-París, Ruiz &amp; Recuero
</taxonomicName>
des. 2019 [printed white label].
<collectingCounty id="341F69AE5868110FFB7D9C70FB56FEB9" box="[1153,1276,289,311]" pageId="22" pageNumber="1271">Preserved</collectingCounty>
in absolute ethanol, held at the
<collectingMunicipality id="3D1A8B585868110FFBE09C11FAEEFED8" box="[1052,1348,320,342]" pageId="22" pageNumber="1271">Entomological Collection</collectingMunicipality>
of the
<location id="D81E47F95868110FFCD59C0EFB49FEFA" LSID="urn:lsid:plazi:treatment:5568A03458681100FF449BEDFA25FC3A:D81E47F95868110FFCD59C0EFB49FEFA" box="[809,1251,350,372]" country="Spain" county="Preserved" municipality="Entomological Collection" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" stateProvince="Madrid">Museo Nacional de Ciencias Naturales</location>
,
<collectingRegion id="1F05DFC05868110FFB129C0FFAECFEFA" box="[1262,1350,350,372]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="DD7E112258681101FCD59CCFFD72FB3F" blockId="22.[809,1425,413,1907]" lastBlockId="24.[145,762,198,1201]" lastPageId="24" lastPageNumber="1273" pageId="22" pageNumber="1271">
<emphasis id="EFB5CD305868110FFCD59CCFFC01FE3D" box="[809,939,414,435]" italics="true" pageId="22" pageNumber="1271">
<typeStatus id="027AAF805868110FFCD59CCFFC0EFE3D" box="[809,932,414,435]" pageId="22" pageNumber="1271" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="A5D651B25868110FFC469CCCFBAEFE3D" box="[954,1028,413,435]" name="Spain" pageId="22" pageNumber="1271">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC05868110FFBE89CCCFBDBFE3D" box="[1044,1137,413,435]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Montejo de la Sierra, pastizal junto al pueblo,
<date id="A97F37E25868110FFB989CEDFABBFE5F" box="[1124,1297,444,466]" pageId="22" pageNumber="1271" value="1999-04-24">24 April 1999</date>
, MAB166, M. García-París leg. [white label, printed];
<collectionCode id="BBD089E75868110FFACC9C8AFA28FE7E" box="[1328,1410,475,496]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_ Ent 231431 [white label, printed] (male, preserved in ethanol).―
<collectingCountry id="A5D651B25868110FFC569F49FC47FDA0" box="[938,1005,536,558]" name="Spain" pageId="22" pageNumber="1271">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC05868110FFC099F49FBE0FDA0" box="[1013,1098,536,558]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Montejo de la Sierra, pastizal junto al pueblo,
<date id="A97F37E25868110FFC179F66FB24FDC2" box="[1003,1166,566,588]" pageId="22" pageNumber="1271" value="1999-04-24">24 April 1999</date>
,
<collectionCode id="BBD089E75868110FFB609F66FB77FDC2" box="[1180,1245,567,588]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="22" pageNumber="1271">NTM</collectionCode>
120, M. García-París leg. [white label, printed];
<collectionCode id="BBD089E75868110FFB559F04FB54FDE4" box="[1193,1278,597,618]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 231433 (female, extracted genitalia, preserved dry).―
<collectingCountry id="A5D651B25868110FFABB9F25FA26FD04" box="[1351,1420,628,650]" name="Spain" pageId="22" pageNumber="1271">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC05868110FFCD59FC3FCD5FD26" box="[809,895,658,680]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Montejo de la Sierra, pastizal junto al pueblo,
<date id="A97F37E25868110FFCD59FE0FC72FD48" box="[809,984,689,711]" pageId="22" pageNumber="1271" value="1999-04-24">24 April 1999</date>
, M. García-París leg. [white label, printed];
<collectionCode id="BBD089E75868110FFC739F81FC48FD6B" box="[911,994,720,741]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 231432 [white label, printed]; 118 [white label, handwritten] (female, extracted genitalia, three legs separated, preserved dry).―
<collectingRegion id="1F05DFC05868110FFB129E5CFAEFFCAD" box="[1262,1349,781,803]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Rivas de Jarama (Urbanizaciones),
<date id="A97F37E25868110FFB839E7DFAA5FCCF" box="[1151,1295,812,834]" pageId="22" pageNumber="1271" value="2009-06-05">5 June 2009</date>
, mel09044, M. Gª-París &amp; G. Gª-Martín leg. [white label, printed];
<collectionCode id="BBD089E75868110FFCD59E38FC2AFCF0" box="[809,896,873,894]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 231434 [white label, printed] (female, preserved in ethanol).―Área de Montarco, Rivas- Vaciamadrid (
<collectingRegion id="1F05DFC05868110FFC3B9EF7FBB7FC32" box="[967,1053,934,956]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
),
<date id="A97F37E25868110FFBCC9EF6FB60FC32" box="[1072,1226,934,956]" pageId="22" pageNumber="1271" value="2006-04-13">13 April 2006</date>
, J. I. López-Colón leg. [white label, handwritten];
<collectionCode id="BBD089E75868110FFB5A9E94FB56FC54" box="[1190,1276,965,986]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 231435 [white label, printed] (preserved in ethanol).―
<collectingRegion id="1F05DFC05868110FFACB9EB5FA26FC74" box="[1335,1420,996,1018]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Alcalá de Henares,
<date id="A97F37E25868110FFBF89953FB3DFB99" box="[1028,1175,1026,1048]" pageId="22" pageNumber="1271" value="2001-05-16">16 May 2001</date>
[white label, printed];
<collectionCode id="BBD089E75868110FFCD59970FCD7FBB8" box="[809,893,1057,1078]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 233312 (preserved dry).―
<collectingRegion id="1F05DFC05868110FFB1B9970FA97FBB9" box="[1255,1341,1057,1079]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Alcalá de Henares,
<date id="A97F37E25868110FFC329911FBDDFBD8" box="[974,1143,1088,1110]" pageId="22" pageNumber="1271" value="2001-05-16">16 May 2001</date>
[white label, printed];
<collectionCode id="BBD089E75868110FFCD5990FFCD7FBFD" box="[809,893,1118,1139]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 233313 (preserved dry).―
<collectingRegion id="1F05DFC05868110FFB1B990FFA97FBFA" box="[1255,1341,1118,1140]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Alcalá de Henares,
<date id="A97F37E25868110FFC37992CFBC5FB1C" box="[971,1135,1149,1171]" pageId="22" pageNumber="1271" value="2001-05-16">16 May 2001</date>
[white label, printed];
<collectionCode id="BBD089E75868110FFCD599CDFC2FFB3F" box="[809,901,1180,1201]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 233314 (preserved dry).―
<collectingRegion id="1F05DFC05868110FFAD799CDFA20FB3C" box="[1323,1418,1180,1202]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Leganés,
<date id="A97F37E25868110FFC6399EBFB9DFB41" box="[927,1079,1210,1232]" pageId="22" pageNumber="1271" value="1990-06-08">8 June 1990</date>
[white label, handwritten];
<taxonomicName id="1AC16AA15868110FFCD59988FB8DFB61" authority="Linneo" authorityName="Linneo" box="[809,1063,1241,1263]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD305868110FFCD59988FC61FB60" box="[809,971,1241,1262]" italics="true" pageId="22" pageNumber="1271">Meloe majalis</emphasis>
Linneo
</taxonomicName>
; ex. Colección M. de los Mozos;
<collectionCode id="BBD089E75868110FFCD599A9FCD4FA83" box="[809,894,1272,1293]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 173819 [white label, printed] (preserved dry).―
<collectingRegion id="1F05DFC05868110FFC819847FC7EFAA2" box="[893,980,1302,1324]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Barrio del Pilar,
<date id="A97F37E25868110FFB589847FAEFFAA5" box="[1188,1349,1302,1324]" pageId="22" pageNumber="1271" value="1980-04-21">21 April 1980</date>
, M. de los Mozos leg., entre hierbas [white label, handwritten];
<taxonomicName id="1AC16AA15868110FFCD59805FB8FFAE4" authority="Linneo" authorityName="Linneo" box="[809,1061,1364,1386]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD305868110FFCD59805FC66FAE7" box="[809,972,1364,1385]" italics="true" pageId="22" pageNumber="1271">Meloe majalis</emphasis>
Linneo
</taxonomicName>
[white label, handwritten]; ex. Colección M. de los Mozos;
<collectionCode id="BBD089E75868110FFBA29823FB18FA09" box="[1118,1202,1394,1415]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 173820 [white label, printed] (preserved dry).―5 exx.
<collectionCode id="BBD089E75868110FFAF998C0FAF1FA28" box="[1285,1371,1425,1446]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 232449 to
<collectionCode id="BBD089E75868110FFC5898E1FC53FA4B" box="[932,1017,1456,1477]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 232453 [white label, printed];
<collectingRegion id="1F05DFC05868110FFCD5989FFC2FFA6A" box="[809,901,1486,1508]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Braojos,
<date id="A97F37E25868110FFBFF989FFB1AFA6D" box="[1027,1200,1486,1508]" pageId="22" pageNumber="1271" value="2012-04-24">24 April 2012</date>
, F. A. Montes leg. [white label, printed] (preserved dry).―
<collectingRegion id="1F05DFC05868110FFAFC98BCFAF2F98D" box="[1280,1368,1517,1539]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, San Fernando de Henares,
<date id="A97F37E25868110FFBD39B5DFB6FF9AF" box="[1071,1221,1548,1570]" pageId="22" pageNumber="1271" value="2011-05-16">16 May 2011</date>
, J. I. López-Colón leg. [white label, printed];
<collectionCode id="BBD089E75868110FFBA79B7BFB1AF9B1" box="[1115,1200,1578,1599]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 232454 [white label, printed] (preserved dry).―
<collectingRegion id="1F05DFC05868110FFB319B18FA83F9D1" box="[1229,1321,1609,1631]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Ciudad Universitaria, 14 April [white label, handwritten];
<taxonomicName id="1AC16AA15868110FFCD59BD7FB5AF912" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[809,1264,1670,1692]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD305868110FFCD59BD7FB87F915" box="[809,1069,1670,1691]" italics="true" pageId="22" pageNumber="1271">Berberomeloe majalis</emphasis>
<bibRefCitation id="B9506CD35868110FFBCB9BD7FB5AF912" author="Linnaeus C" box="[1079,1264,1670,1692]" pageId="22" pageNumber="1271" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
[white label, printed];
<collectionCode id="BBD089E75868110FFC669BF4FC5BF934" box="[922,1009,1701,1722]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 233310 [white label, printed] (preserved dry).―
<collectingCountry id="A5D651B25868110FFBFF9B95FBE3F954" box="[1027,1097,1732,1754]" name="Spain" pageId="22" pageNumber="1271">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC05868110FFBAA9B95FB04F954" box="[1110,1198,1732,1754]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, Rivas del Jarama,
<date id="A97F37E25868110FFCD59BB3FC79F979" box="[809,979,1762,1784]" pageId="22" pageNumber="1271" value="1988-03-13">13 March 1988</date>
, F. A. Montes leg. [white label, printed];
<taxonomicName id="1AC16AA15868110FFCD59A50FB7BF899" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[809,1233,1793,1815]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="1271" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD305868110FFCD59A50FBB6F898" box="[809,1052,1793,1814]" italics="true" pageId="22" pageNumber="1271">Berberomeloe majalis</emphasis>
<bibRefCitation id="B9506CD35868110FFBDF9A50FB7BF899" author="Linnaeus C" box="[1059,1233,1793,1815]" pageId="22" pageNumber="1271" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
, M. Gª-París det. 98 [white label, printed]; 342 [white label, handwritten];
<collectionCode id="BBD089E75868110FFCD59A6FFCD4F8DD" box="[809,894,1854,1875]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="22" pageNumber="1271" type="Museum">MNCN</collectionCode>
_Ent 233311 [white label, printed] (preserved dry).―
<collectingRegion id="1F05DFC05868110FFC829A0CFC7CF8FD" box="[894,982,1885,1907]" country="Spain" name="Madrid" pageId="22" pageNumber="1271">Madrid</collectingRegion>
, El Molar [white label, handwritten];
<taxonomicName id="1AC16AA15869110EFF5F9A73FDE1F8B6" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[163,587,1826,1848]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1272" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD305869110EFF5F9A73FE3CF8B9" box="[163,406,1826,1847]" italics="true" pageId="23" pageNumber="1272">Berberomeloe majalis</emphasis>
<bibRefCitation id="B9506CD35869110EFE619A73FDE1F8B6" author="Linnaeus C" box="[413,587,1826,1848]" pageId="23" pageNumber="1272" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
, M. Gª-París det. 98; 571 [white label, handwritten];
<collectionCode id="BBD089E75869110EFDD09A10FDD5F8D8" box="[556,639,1857,1878]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="23" pageNumber="1272" type="Museum">MNCN</collectionCode>
_Ent 233309 [white label, printed] (preserved dry).―2 exx. labelled:
<collectingRegion id="1F05DFC05869110EFCC79A73FC38F8B6" box="[827,914,1826,1848]" country="Spain" name="Madrid" pageId="23" pageNumber="1272">Madrid</collectingRegion>
, Lozoyuela,
<date id="A97F37E25869110EFBDD9A73FB14F8B9" box="[1057,1214,1826,1848]" pageId="23" pageNumber="1272" value="2002-06-20">20 June 2002</date>
, I. Martínez-Solano &amp; I. Sánchez [white label, handwritten];
<collectionCode id="BBD089E75869110EFAE59A10FAC4F8D8" box="[1305,1390,1857,1878]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="23" pageNumber="1272" type="Museum">MNCN</collectionCode>
_Ent 250993 and
<collectionCode id="BBD089E75869110EFC309A31FB8BF8FB" box="[972,1057,1888,1909]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="23" pageNumber="1272" type="Museum">MNCN</collectionCode>
_Ent 250994, respectively [white labels, printed] (preserved in ethanol).―
<collectingRegion id="1F05DFC058661101FD649D97FD59FF52" box="[664,755,198,220]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, Montejo de la Sierra,
<date id="A97F37E258661101FE719DB4FD87FF74" box="[397,557,229,251]" pageId="24" pageNumber="1273" value="1999-04-24">24 April 1999</date>
,
<collectionCode id="BBD089E758661101FDC59DB4FDDFFF74" box="[569,629,229,250]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="24" pageNumber="1273">NTM</collectionCode>
121 [white label, handwritten];
<collectionCode id="BBD089E758661101FE829C55FE78FE97" box="[382,466,260,281]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 250995 [white label, printed] (preserved in ethanol).―
<collectingCountry id="A5D651B258661101FDDC9C73FDCFFEB6" box="[544,613,290,312]" name="Spain" pageId="24" pageNumber="1273">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC058661101FD8E9C73FD60FEB6" box="[626,714,290,312]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, 4.5 Km E/SE de Rivas Vaciamadrid,
<quantity id="1A39BCC758661101FDFF9C10FDE3FED8" box="[515,585,321,343]" metricMagnitude="2" metricUnit="m" metricValue="5.76" pageId="24" pageNumber="1273" unit="m" value="576.0">576 m</quantity>
,
<geoCoordinate id="B8F577E558661101FDA99C10FD5EFED9" box="[597,756,321,343]" degrees="40" direction="north" minutes="18" orientation="latitude" pageId="24" pageNumber="1273" precision="1" seconds="54.46" value="40.31513">40°1854.46N</geoCoordinate>
,
<geoCoordinate id="B8F577E558661101FF6D9C0EFE8CFEFB" box="[145,294,351,373]" degrees="3" direction="west" minutes="34" orientation="longitude" pageId="24" pageNumber="1273" precision="1" seconds="22.57" value="-3.572936">3°3422.57W</geoCoordinate>
,
<date id="A97F37E258661101FED29C31FE0BFEFB" box="[302,417,352,373]" pageId="24" pageNumber="1273" value="2009-05-11">
11/
<collectionCode id="BBD089E758661101FEB19C31FECBFEFB" box="[333,353,352,373]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="1273" type="Museum">V</collectionCode>
/2009
</date>
, I. Martínez-Solano, C. Settani, E. Recuero leg. [white label, handwritten];
<collectionCode id="BBD089E758661101FD8D9C2FFD6EFE1D" box="[625,708,382,403]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 250996 [white label, printed] (female, preserved in ethanol).―
<collectingCountry id="A5D651B258661101FEE19CEDFECFFE5C" box="[285,357,444,466]" name="Spain" pageId="24" pageNumber="1273">Spain</collectingCountry>
,
<collectingRegion id="1F05DFC058661101FE889CEDFE65FE5C" box="[372,463,444,466]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, Colmenar de Oreja, La Aldehuela,
<quantity id="1A39BCC758661101FEEC9C8BFEFFFE7E" box="[272,341,474,496]" metricMagnitude="2" metricUnit="m" metricValue="7.26" pageId="24" pageNumber="1273" unit="m" value="726.0">726 m</quantity>
,
<geoCoordinate id="B8F577E558661101FE9C9C8BFDAAFE61" box="[352,512,474,495]" degrees="40" direction="north" minutes="04" orientation="latitude" pageId="24" pageNumber="1273" precision="1" seconds="43.74" value="40.078815">40°0443.74N</geoCoordinate>
,
<geoCoordinate id="B8F577E558661101FDF69C8BFD31FE7E" box="[522,667,474,496]" degrees="3" direction="east" minutes="23" orientation="longitude" pageId="24" pageNumber="1273" precision="1" seconds="57.14" value="3.3992054">3°2357.14O</geoCoordinate>
,
<date id="A97F37E258661101FD5A9C8BFF63FD80" pageId="24" pageNumber="1273" value="2012-05-15">15 May 2012</date>
, mel12012, M. Gª-París leg. [white label, printed];
<collectionCode id="BBD089E758661101FF6D9F49FF4EFDA3" box="[145,228,536,557]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 250997 [white label, printed] (preserved in ethanol).―
<collectingRegion id="1F05DFC058661101FEEE9F67FECDFDC2" box="[274,359,566,588]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, Rivas de Jarama (Urbanizaciones),
<date id="A97F37E258661101FF6D9F04FE89FDE4" box="[145,291,597,619]" pageId="24" pageNumber="1273" value="2009-06-05">5 June 2009</date>
, mel09041, M. Gª-París &amp; G. Gª-Martín leg. [white label, printed];
<collectionCode id="BBD089E758661101FE3F9F25FDB2FD07" box="[451,536,628,649]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 250998 [white label, printed] (preserved in ethanol).―
<collectingRegion id="1F05DFC058661101FDA49FC3FD04FD26" box="[600,686,658,680]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, Rivas de Jarama (Urbanizaciones),
<date id="A97F37E258661101FE1B9FE0FDDDFD48" box="[487,631,689,711]" pageId="24" pageNumber="1273" value="2009-06-05">5 June 2009</date>
, mel09042, M. Gª-París &amp; G. Gª-Martín leg. [white label, printed];
<collectionCode id="BBD089E758661101FF6D9FBFFF4EFC8D" box="[145,228,750,771]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 250999 [white label, printed] (preserved in ethanol).―
<collectingRegion id="1F05DFC058661101FEEE9E5CFECDFCAD" box="[274,359,781,803]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
, Rivas de Jarama (Urbanizaciones),
<date id="A97F37E258661101FF6D9E7AFE89FCCF" box="[145,291,811,833]" pageId="24" pageNumber="1273" value="2009-06-05">5 June 2009</date>
, mel09043, M. Gª-París &amp; G. Gª-Martín leg. [white label, printed];
<collectionCode id="BBD089E758661101FE3F9E1BFDB2FCD1" box="[451,536,842,863]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 251000 [white label, printed] (preserved in ethanol).―3 exx. labelled: Segovia,Prádena,
<date id="A97F37E258661101FEAB9ED9FE79FC13" box="[343,467,903,925]" pageId="24" pageNumber="1273" value="2015-05-07">7May2015</date>
,M.García-Parísleg.[white label, handwritten];
<collectionCode id="BBD089E758661101FE819EF7FE7BFC35" box="[381,465,934,955]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 251001,
<collectionCode id="BBD089E758661101FD939EF7FD69FC35" box="[623,707,934,955]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 251002 and
<collectionCode id="BBD089E758661101FED89E94FED0FC54" box="[292,378,965,986]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="24" pageNumber="1273" type="Museum">MNCN</collectionCode>
_Ent 251003 respectively [white labels, printed] (preserved in ethanol). All
<typeStatus id="027AAF8058661101FD799EB5FD53FC77" box="[645,761,996,1017]" pageId="24" pageNumber="1273" type="paratype">paratypes</typeStatus>
labelled:
<typeStatus id="027AAF8058661101FF079953FED8FB99" box="[251,370,1026,1047]" pageId="24" pageNumber="1273" type="paratype">Paratypus</typeStatus>
,
<taxonomicName id="1AC16AA158661101FE829953FDEDFBB8" authority="Sanchez-Vialas, Garcia-Paris, Ruiz &amp; Recuero" authorityName="Sanchez-Vialas, Garcia-Paris, Ruiz &amp; Recuero" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="1273" phylum="Arthropoda" rank="species" species="comunero">
<emphasis id="EFB5CD3058661101FE829953FD21FB96" box="[382,651,1026,1048]" italics="true" pageId="24" pageNumber="1273">Berberomeloe comunero</emphasis>
Sánchez-Vialas, García-París, Ruiz &amp; Recuero
</taxonomicName>
, des. 2019 [red labels for dry-preserved specimens, and white labels for ethanol-preserved specimens, all printed]. All specimens are held at the Entomological Collection of the Museo Nacional de Ciencias Naturales,
<collectingRegion id="1F05DFC058661101FD8199CAFD7EFB3F" box="[637,724,1179,1201]" country="Spain" name="Madrid" pageId="24" pageNumber="1273">Madrid</collectingRegion>
.
</paragraph>
<caption id="89BE41AA5869110EFF5F9BD4FDC4F97D" pageId="23" pageNumber="1272" startId="23.[163,241,1669,1691]" targetBox="[271,1337,198,1628]" targetPageId="23" targetType="figure">
<paragraph id="DD7E11225869110EFF5F9BD4FDC4F97D" blockId="23.[163,1443,1669,1779]" pageId="23" pageNumber="1272">
<emphasis id="EFB5CD305869110EFF5F9BD4FEBDF914" bold="true" box="[163,279,1669,1691]" pageId="23" pageNumber="1272">Figure 10.</emphasis>
A, B, habitat of
<taxonomicName id="1AC16AA15869110EFE439BD4FD38F915" box="[447,658,1669,1691]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1272" phylum="Arthropoda" rank="species" species="castuo">
<emphasis id="EFB5CD305869110EFE439BD4FD38F915" box="[447,658,1669,1691]" italics="true" pageId="23" pageNumber="1272">Berberomeloe castuo</emphasis>
</taxonomicName>
, Spain. A, dehesa ecosystem with cattle, Plasencia, Cáceres; B, Jarandilla de la Vera, Cáceres. D, E, specimens of
<taxonomicName id="1AC16AA15869110EFDE79BF5FDD2F937" box="[539,632,1700,1721]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1272" phylum="Arthropoda" rank="species" species="castuo">
<emphasis id="EFB5CD305869110EFDE79BF5FDD2F937" box="[539,632,1700,1721]" italics="true" pageId="23" pageNumber="1272">B. castuo</emphasis>
</taxonomicName>
from Ramacastañas, Ávila and Agallas, Salamanca, respectively. C, specimen of
<taxonomicName id="1AC16AA15869110EFF409B90FEB3F958" box="[188,281,1729,1750]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1272" phylum="Arthropoda" rank="species" species="castuo">
<emphasis id="EFB5CD305869110EFF409B90FEB3F958" box="[188,281,1729,1750]" italics="true" pageId="23" pageNumber="1272">B. castuo</emphasis>
</taxonomicName>
showing defensive behaviour from Cabezas Rubias, Huelva; note the lack of the coloured abdominal transverse bars, typical from the southern populations.
</paragraph>
</caption>
<paragraph id="DD7E112258661101FF6D998BFDBBFA26" blockId="24.[145,761,1242,1448]" pageId="24" pageNumber="1273">
<emphasis id="EFB5CD3058661101FF6D998BFEBFFB61" box="[145,277,1242,1263]" italics="true" pageId="24" pageNumber="1273">Etymology:</emphasis>
This species is named in remembrance of the ballad Los Comuneros writen by Luis López Álvarez, published in 1972, and set to music by the folk music group Nuevo Mester de Juglaría in 1976. This ballad narrates the tragic historical events that took place between 1520 and
<quantity id="1A39BCC758661101FE109822FDE9FA07" box="[492,579,1395,1417]" metricMagnitude="1" metricUnit="m" metricValue="3.8658799999999998" pageId="24" pageNumber="1273" unit="in" value="1522.0">1522 in</quantity>
Castille, in the area where the new species occur.
</paragraph>
<paragraph id="DD7E112258661100FF6D9880FD95FF74" blockId="24.[145,761,1489,1909]" lastBlockId="25.[163,778,197,250]" lastPageId="25" lastPageNumber="1274" pageId="24" pageNumber="1273">
<emphasis id="EFB5CD3058661101FF6D9880FE6FFA68" box="[145,453,1489,1510]" italics="true" pageId="24" pageNumber="1273">
Description of the
<typeStatus id="027AAF8058661101FE9C9880FE15FA68" box="[352,447,1489,1510]" pageId="24" pageNumber="1273" type="holotype">holotype</typeStatus>
:
</emphasis>
Length (frons to posterior margin of elytra),
<quantity id="1A39BCC758661101FE8F98A1FE78F98B" box="[371,466,1520,1541]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="1273" unit="mm" value="7.3">7.3 mm</quantity>
. Total length (including abdomen) of preserved
<typeStatus id="027AAF8058661101FE569B5FFDBBF9AA" box="[426,529,1550,1572]" pageId="24" pageNumber="1273" type="holotype">holotype</typeStatus>
,
<quantity id="1A39BCC758661101FDE29B5FFDDEF9AA" box="[542,628,1550,1572]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="24" pageNumber="1273" unit="mm" value="31.0">31 mm</quantity>
. Maximum width,
<quantity id="1A39BCC758661101FF149B7CFE9BF9CC" box="[232,305,1581,1603]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="1273" unit="mm" value="5.0">5 mm</quantity>
. Moderately wide orange transverse bar in the posterior edge of each tergum except VII and VIII, which are entirely black. Tegument finely microreticulated, semi-glossy. Head maximum width,
<quantity id="1A39BCC758661101FF6D9BF6FF5AF933" box="[145,240,1703,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="1273" unit="mm" value="2.4">2.4 mm</quantity>
. Surface covered by numerous, uniformly distributed punctures. Head punctures from medium-sized to large, rounded, deep marked, close to each other, confluent in temples. A longitudinal midline is impressed on the upper half of the frons, but faded at the vertex. Minimum interorbital distance,
<quantity id="1A39BCC758661101FD659A11FD5EF8D8" box="[665,756,1856,1878]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="1273" unit="mm" value="1.5">1.5 mm</quantity>
. Clypeus
<quantity id="1A39BCC758661101FF0E9A0EFEECF8FB" box="[242,326,1887,1909]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="1273" unit="mm" value="1.1">1.1 mm</quantity>
wide by
<quantity id="1A39BCC758661101FE5A9A0EFE50F8FB" box="[422,506,1887,1909]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="1273" unit="mm" value="0.4">0.4 mm</quantity>
long. Labrum 1.1 wide by
<quantity id="1A39BCC758661101FCB59D94FC37FF54" box="[841,925,197,218]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="1273" unit="mm" value="0.4">0.4 mm</quantity>
long. Antennae with antennomeres widened apically but not strongly serrated, with short black vestiture, mostly decumbent and with a few sparse erect setae, longer and semi-erect on antennomeres III; antennomere I slightly widened apically, subcylindrical (length:
<quantity id="1A39BCC758661101FC7A9C0FFC77FEFD" box="[902,989,350,372]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="1273" unit="mm" value="0.5">0.5 mm</quantity>
); II short, cylindrical (length:
<quantity id="1A39BCC758661101FAD29C0FFA2FFEFD" box="[1326,1413,350,371]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="1273" unit="mm" value="0.2">0.2 mm</quantity>
); III (length:
<quantity id="1A39BCC758661101FC4E9C2CFBA1FE1C" box="[946,1035,381,402]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="1273" unit="mm" value="0.4">0.4 mm</quantity>
) subcylindrical, slightly widened apically, rectangular; IV (length:
<quantity id="1A39BCC758661101FB359CCDFA8FFE3F" box="[1225,1317,412,433]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) shorter than III but equally wide, subrectangular;
<collectionCode id="BBD089E758661101FAE29CEAFA9AFE5E" box="[1310,1328,443,464]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="1273" type="Museum">V</collectionCode>
(length:
<quantity id="1A39BCC758661101FCD59C88FC2BFE60" box="[809,897,473,494]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, wider than
<collectionCode id="BBD089E758661101FB569C88FB62FE60" box="[1194,1224,473,494]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="24" pageNumber="1273">VI</collectionCode>
, with a wide and rounded apical tooth on the inner edge;
<collectionCode id="BBD089E758661101FAF39CA9FA86FD83" box="[1295,1324,504,525]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="24" pageNumber="1273">VI</collectionCode>
(length:
<quantity id="1A39BCC758661101FCD59F47FC2CFDA5" box="[809,902,534,555]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, with a rounded apical tooth on the inner edge; VII (length:
<quantity id="1A39BCC758661101FB679F64FB5FFDC4" box="[1179,1269,565,586]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, with an apical tooth on the inner edge; VIII (length:
<quantity id="1A39BCC758661101FCD59F23FCD5FD09" box="[809,895,626,647]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, narrow than VII, weakly dentate on the apex of the inner edge; IX (length:
<quantity id="1A39BCC758661101FAD19FC0FA22FD28" box="[1325,1416,657,678]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, similar to VIII, dentate on inner edge; X (length:
<quantity id="1A39BCC758661101FC5A9F9FFC55FD6D" box="[934,1023,718,739]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="1273" unit="mm" value="0.3">0.3 mm</quantity>
) trapezoidal, apical tooth slightly acute; XI (length:
<quantity id="1A39BCC758661101FBF59FBCFBCCFC8C" box="[1033,1126,749,770]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="1273" unit="mm" value="0.4">0.4 mm</quantity>
) subconical, moderately wide, notched on apex. Pronotum subquadrate with subparallel sides, slightly narrower towards the base (anterior side of pronotum:
<quantity id="1A39BCC758661101FB969E18FB69FCD0" box="[1130,1219,841,862]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="1273" unit="mm" value="2.1">2.1 mm</quantity>
; posterior edge of pronotum:
<quantity id="1A39BCC758661101FC559E39FC46FCF3" box="[937,1004,872,893]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="1273" unit="mm" value="2.0">2 mm</quantity>
; pronotum length on sagittal plane:
<quantity id="1A39BCC758661101FCD59ED7FC2AFC15" box="[809,896,902,923]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="1273" unit="mm" value="1.7">1.7 mm</quantity>
); fore angles rounded; anterior margin curved, posterior margin slightly arcuate; with an impressed longitudinal midline through the pronotum length and diffuse lateral depressions. Pronotum surface densely and homogenously punctate; punctures of various sizes, mostly large, rounded, and deep, close to each other, confluent and forming a corrugated or reticulated pattern. Dorsal surface of pronotum almost glabrous, with a short isolated seta in each puncture; anterior margin, adjacent to the neck, with numerous, moderately long setae. Elytra imbricated basally (the right over the left), length:
<quantity id="1A39BCC758661101FBAB9986FB07FB63" box="[1111,1197,1239,1261]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="24" pageNumber="1273" unit="mm" value="4.1">4.1 mm</quantity>
; tegument glabrous, slightly corrugated longitudinally with impressed irregular vermicular lines, with scarce, weakly marked, dispersed punctures. Distal margin of tergites I
<collectionCode id="BBD089E758661101FA8E9865FA25FAC7" box="[1394,1423,1332,1353]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="24" pageNumber="1273">VI</collectionCode>
with an orange transverse bar, moderately narrow in comparison with the total width of the terga (width of the second tergum:
<quantity id="1A39BCC758661101FBCC98DEFB21FA2B" box="[1072,1163,1423,1445]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="24" pageNumber="1273" unit="mm" value="4.8">4.8 mm</quantity>
; width transverse bar of the second tergum:
<quantity id="1A39BCC758661101FBDB98FFFB2AFA4D" box="[1063,1152,1454,1475]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="24" pageNumber="1273" unit="mm" value="2.7">2.7 mm</quantity>
). Last ventrite notched. Metafemur longer than metatibia (metafemur length:
<quantity id="1A39BCC758661101FC7D98BAFC70F98F" box="[897,986,1515,1537]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="24" pageNumber="1273" unit="mm" value="2.5">2.5 mm</quantity>
; metatibia length:
<quantity id="1A39BCC758661101FB4498BAFB57F98E" box="[1208,1277,1515,1536]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="1273" unit="mm" value="2.0">2 mm</quantity>
). Metatarsal tarsomeres length, from inner to apical: 2.2, 0.4, 0.4,
<quantity id="1A39BCC758661101FCD59B78FC28F9B0" box="[809,898,1577,1598]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="1273" unit="mm" value="0.7">0.7 mm</quantity>
. Genitalia with tegmen brownish; moderately elongated, slender both on dorsal and lateral views. Phallobase longer than wide, length similar to the parameres; maximum width at the middle. Parameres longer than wide, basally cylindrical; distal third formed by parameral lobes; setae present on middle region of parameres. Parameral lobes separated by a longitudinal notch that extends to the middle of the dorsal surface of the parameres; apexes rounded. Median lobe long, robust, flattened, truncated at the apex in lateral view, with two acute ventral hooks; ventral hooks similar and close to each other, separated from apex. Endophallic hook visible.
</paragraph>
<paragraph id="DD7E112258671100FF5F9C73FE51FE7E" blockId="25.[163,779,290,496]" pageId="25" pageNumber="1274">
<emphasis id="EFB5CD3058671100FF5F9C73FF56FEB9" box="[163,252,290,311]" italics="true" pageId="25" pageNumber="1274">Female:</emphasis>
Similar to male, but with the last abdominal ventrite rounded, not emarginated in its posterior margin and antennomeres less widened apically. Studied specimens present the inner surface of the bursa copulatrix with few and small sclerotized spicules, sometimes absent (similar to those of
<taxonomicName id="1AC16AA158671100FF5F9C8AFEA3FE61" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[163,265,474,496]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="indalo">
<emphasis id="EFB5CD3058671100FF5F9C8AFEA3FE61" box="[163,265,474,496]" italics="true" pageId="25" pageNumber="1274">B. indalo</emphasis>
</taxonomicName>
, as figured in
<figureCitation id="45FA0DA758671100FE509C8BFE46FE7E" box="[428,492,474,496]" captionStart="Figure 8" captionStartId="20.[145,223,1260,1282]" captionTargetBox="[344,1228,202,1213]" captionTargetId="figure-302@20.[337,1233,195,1220]" captionTargetPageId="20" captionText="Figure 8. Female genitalia of (A) Berberomeloe castuo and (B) B. indalo, showing the valvifera and style over the inner side of the vagina. An arrow shows the place where sclerotized spicules are present (A) or absent (B)." pageId="25" pageNumber="1274">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="DD7E112258671100FF5F9F48FEF7FB5C" blockId="25.[163,780,537,1234]" pageId="25" pageNumber="1274">
<emphasis id="EFB5CD3058671100FF5F9F48FE8DFDA0" box="[163,295,537,558]" italics="true" pageId="25" pageNumber="1274">Variability:</emphasis>
Body length (frons to posterior border of elytra) highly variable, from
<quantity id="1A39BCC758671100FE0D9F69FD21FDC3" box="[497,651,568,589]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="2.0" metricValueMin="0.73" pageId="25" pageNumber="1274" unit="mm" value="13.65" valueMax="20.0" valueMin="7.3">7.3 to 20 mm</quantity>
; maximum total length among preserved, studied specimens,
<quantity id="1A39BCC758671100FF5F9F24FF5FFD05" box="[163,245,629,651]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="1274" unit="mm" value="54.0">54 mm</quantity>
. Morphometric variability is shown in
<tableCitation id="9043249958671100FD4D9F24FCAFFD04" box="[689,773,629,651]" captionStart="Table 2" captionStartId="16.[145,209,196,217]" captionText="Table 2. Mean and standard deviation (SD) of the morphometric measurements (in mm) from studied Berberomeloe specimens (excluding B. laevigatus and B. insignis)" pageId="25" pageNumber="1274">Table 2</tableCitation>
. Some specimens have a single, small, red spot on the frons, diffuse and faintly marked. Colour of transverse bars in live specimens vary from orange to blood-red, but constant within each population. Populations composed exclusively of entirely black-coloured specimens are found in
<collectingRegion id="1F05DFC058671100FEEF9E7CFEC1FCCD" box="[275,363,813,835]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
(Rivas de Jarama, Valdepiélagos, Talamanca, Daganzo de Arriba, Vicálvaro) and western Guadalajara (Uceda) (
<bibRefCitation id="B9506CD358671100FE5C9E3AFDC2FC0E" author="Bravo C &amp; Mas-Peinado P &amp; Bautista LM &amp; Blanco G &amp; Alonso JC &amp; Garcia-Paris M" box="[416,616,874,896]" pageId="25" pageNumber="1274" pagination="790 - 804" refId="ref39536" refString="Bravo C, Mas-Peinado P, Bautista LM, Blanco G, Alonso JC, Garcia-Paris M. 2017. Cantharidin is conserved across phylogeographic lineages and present in both morphs of Iberian Berberomeloe blister beetles (Coleoptera, Meloidae). Zoological Journal of the Linnean Society 180: 790-804." type="journal article" year="2017">
Bravo
<emphasis id="EFB5CD3058671100FE179E3AFD8EFCF1" box="[491,548,874,896]" italics="true" pageId="25" pageNumber="1274">et al.</emphasis>
, 2017
</bibRefCitation>
; authors pers. obs.).
<taxonomicName id="1AC16AA158671100FF239ED8FE4CFC11" box="[223,486,905,927]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="comunero">
<emphasis id="EFB5CD3058671100FF239ED8FE4CFC11" box="[223,486,905,927]" italics="true" pageId="25" pageNumber="1274">Berberomeloe comunero</emphasis>
</taxonomicName>
presents a high variability in the width of coloured transverse tergal bars, but on average they are relatively wide (ratio total width of the second tergum/width of the transverse bar of the second tergum between 1.32 and 2.34, average = 1.8, SD = 0.29,
<emphasis id="EFB5CD3058671100FED99972FE93FBB6" box="[293,313,1059,1080]" italics="true" pageId="25" pageNumber="1274">N</emphasis>
= 31). Transverse bars are narrower in mountain populations from the provinces of
<collectingRegion id="1F05DFC058671100FD489910FCA0FBD9" box="[692,778,1089,1111]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
and Segovia. Specimens with deeper punctures on the pronotum, forming a strongly corrugated pattern, are found over most part of the province of
<collectingRegion id="1F05DFC058671100FD8399CCFD7CFB3D" box="[639,726,1181,1203]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
and eastern Segovia.
</paragraph>
<paragraph id="DD7E112258671100FF5F99A9FCA2F935" blockId="25.[163,780,1272,1723]" pageId="25" pageNumber="1274">
<emphasis id="EFB5CD3058671100FF5F99A9FD30FA83" box="[163,666,1272,1293]" italics="true" pageId="25" pageNumber="1274">
Diagnosis:
<taxonomicName id="1AC16AA158671100FEB599A9FD30FA83" box="[329,666,1272,1293]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="comunero">Berberomeloe comunero</taxonomicName>
</emphasis>
<materialsCitation id="6DA91B7F58671100FD5199A9FCAFF935" collectionCode="VI, V" pageId="25" pageNumber="1274" specimenCount="2" typeStatus="holotype">
can be distinguished from all other
<taxonomicName id="1AC16AA158671100FDF59847FCA0FAA2" box="[521,778,1302,1324]" pageId="25" pageNumber="1274">
<emphasis id="EFB5CD3058671100FDF59847FD01FAA5" box="[521,683,1302,1323]" italics="true" pageId="25" pageNumber="1274">Berberomeloe</emphasis>
species
</taxonomicName>
by the following combination of characters (
<figureCitation id="45FA0DA758671100FD509864FD56FAC4" box="[684,764,1333,1355]" captionStart="Figure 11" captionStartId="26.[145,223,1660,1682]" captionTargetBox="[250,1319,201,1621]" captionTargetId="figure-64@26.[250,1319,195,1621]" captionTargetPageId="26" captionText="Figure 11. Habitus and morphological details of Berberomeloe comunero from Pueblade la Sierra, Madrid (A) and Canencia, Madrid (BI). A, dorsal view of a female; B, male antenna; C, D, E, dorsal, ventral, lateral view of male genitalia, showing parameres and phallobase; F, lateral view of aedeagus; G, SEM micrographs of distal part of aedeagus; H, SEM micrographs of pronotum, dorsal view; I, SEM micrographs of head, frontal view." pageId="25" pageNumber="1274">Fig. 11</figureCitation>
): (1) medium-sized to moderately narrow, coloured, transverse tergal bars on the distal margin of tergites I
<collectionCode id="BBD089E758671100FF4798C0FF70FA28" box="[187,218,1425,1446]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="25" pageNumber="1274">VI</collectionCode>
; (2) punctures on head large to medium-sized, rounded, deep and close to each other; (3) pronotum surface densely and homogeneously punctate, with
<specimenCount id="CBC7DAAB58671100FF5F98BCFEB9F98C" box="[163,275,1517,1538]" count="2" pageId="25" pageNumber="1274" type="generic" typeStatus="types">two types</specimenCount>
of puncturation simultaneously, large and medium-sized, confluent or subconfluent, often forming a corrugated pattern; (4) fore angles of pronotum rounded and not especially expanded; (5) apex of the median lobe of male genitalia obliquely truncated in lateral view; and (6) antennomere XI in males wide,
<collectionCode id="BBD089E758671100FD179BD6FD57F912" box="[747,765,1671,1692]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="25" pageNumber="1274" type="Museum">V</collectionCode>
VII in males weakly expanded at its inner apical side
</materialsCitation>
.
</paragraph>
<paragraph id="DD7E112258671100FF5F9BB3FAFBFCD1" blockId="25.[163,779,1761,1906]" lastBlockId="25.[827,1444,197,863]" pageId="25" pageNumber="1274">
<emphasis id="EFB5CD3058671100FF5F9BB3FE7EF89B" italics="true" pageId="25" pageNumber="1274">
Distribution and notes on natural history:
<taxonomicName id="1AC16AA158671100FF5F9A51FE7EF89B" box="[163,468,1792,1813]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="comunero">Berberomeloe comunero</taxonomicName>
</emphasis>
is a euryecious species distributed over most of the northern Central Iberian Plateau, in the provinces of
<collectingRegion id="1F05DFC058671100FDF99A6CFDF4F8DD" box="[517,606,1853,1875]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
(except at its south-eastern edge), Burgos, Segovia, Guadalajara and most of Valladolid. According to the known localities, it occupies a wide elevational range from
<quantity id="1A39BCC758671100FCC79C53FC2AFE96" box="[827,896,258,280]" metricMagnitude="2" metricUnit="m" metricValue="5.22" pageId="25" pageNumber="1274" unit="m" value="522.0">522 m</quantity>
at Balcón del Tajo (
<collectingRegion id="1F05DFC058671100FBA39C53FB1EFE96" box="[1119,1204,258,280]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
) to
<quantity id="1A39BCC758671100FB229C53FA98FE96" box="[1246,1330,258,280]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="25" pageNumber="1274" unit="m" value="1440.0">1440 m</quantity>
at Puerto de Somosierra (
<collectingRegion id="1F05DFC058671100FC099C70FBFAFEB9" box="[1013,1104,289,311]" country="Spain" name="Madrid" pageId="25" pageNumber="1274">Madrid</collectingRegion>
). It inhabits a wide range of geological formations, from limestone and gypsum of the highlands of the Northern Plateau to granite and gneisses at Sierra de Guadarrama, while micaceous schists, slates and quartzites from Somosierra to Sierra de Ayllón (
<bibRefCitation id="B9506CD358671100FBEC9CEBFAFDFE41" author="Rivas-Martinez S &amp; Fernandez-Gonzalez F &amp; Mata D &amp; Pizarro JM" box="[1040,1367,442,464]" pageId="25" pageNumber="1274" pagination="3 - 132" refId="ref43670" refString="Rivas-Martinez S, Fernandez-Gonzalez F, Sanchez- Mata D, Pizarro JM. 1990. Vegetacion de la Sierra de Guadarrama. Itinera Geobotanica 4: 3-132." type="journal article" year="1990">
Rivas-Martínez
<emphasis id="EFB5CD3058671100FB2E9CEAFAA4FE41" box="[1234,1294,442,464]" italics="true" pageId="25" pageNumber="1274">et al.</emphasis>
, 1990
</bibRefCitation>
;
<bibRefCitation id="B9506CD358671100FA999CEAFCD8FE60" author="Vera JA" pageId="25" pageNumber="1274" refId="ref44945" refString="Vera JA. 2004. Geologia de Espana. Madrid: Sociedad Geologica de Espana-Instituto Geologico y Minero de Espana, 890." type="journal volume" year="2004">Vera, 2004</bibRefCitation>
). It occupies the meso- and supra-Mediterranean bioclimatic levels (see:
<bibRefCitation id="B9506CD358671100FBB09CA6FAE3FD83" author="Rivas-Martinez S" box="[1100,1353,503,525]" pageId="25" pageNumber="1274" refId="ref43617" refString="Rivas-Martinez S. 1987. Memoria del mapa de las series de vegetacion de Espana 1: 400.000. Madrid: ICONA." type="book" year="1987">Rivas-Martínez, 1987</bibRefCitation>
;
<bibRefCitation id="B9506CD358671100FAA49CA6FBE8FDA5" author="Rivas-Martinez S &amp; Diaz TE &amp; Fernandez-Gonzalez F &amp; Izco J &amp; Loidi J &amp; Lousa M &amp; Penas A" pageId="25" pageNumber="1274" pagination="5 - 432" refId="ref43700" refString="Rivas-Martinez S, Diaz TE, Fernandez-Gonzalez F, Izco J, Loidi J, Lousa M, Penas A. 2002. Vascular plant communities of Spain and Portugal. Addenda to the syntaxonomical checklist of 2001. Part I. Itinera Geobotanica 15: 5-432." type="journal article" year="2002">
Rivas-Martínez
<emphasis id="EFB5CD3058671100FC4A9F46FC5CFDA5" box="[950,1014,534,556]" italics="true" pageId="25" pageNumber="1274">et al.</emphasis>
, 2002
</bibRefCitation>
).
<taxonomicName id="1AC16AA158671100FBA29F47FA2EFDA2" box="[1118,1412,534,556]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="comunero">
<emphasis id="EFB5CD3058671100FBA29F47FA2EFDA2" box="[1118,1412,534,556]" italics="true" pageId="25" pageNumber="1274">Berberomeloe comunero</emphasis>
</taxonomicName>
is found over an extensive variety of open habitats (
<figureCitation id="45FA0DA758671100FCB89F02FC32FDE7" box="[836,920,595,617]" captionStart="Figure 12" captionStartId="27.[163,241,1658,1680]" captionTargetBox="[260,1347,198,1618]" captionTargetId="figure-83@27.[259,1347,195,1618]" captionTargetPageId="27" captionText="Figure 12. A, specimen of Berberomeloe comunero from Pinilla de los Moros, Burgos. B, C, habitat of B. comunero, Spain; B, Cantalejo, Segovia; C, Tielmes, Madrid." pageId="25" pageNumber="1274">Fig. 12</figureCitation>
), such as cereal fields (mostly wheat and barley), gypsum hills with dispersed
<taxonomicName id="1AC16AA158671100FB219F23FA09FD09" box="[1245,1443,626,647]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="1274" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis id="EFB5CD3058671100FB219F23FA09FD09" box="[1245,1443,626,647]" italics="true" pageId="25" pageNumber="1274">Quercus coccifera</emphasis>
</taxonomicName>
L. trees, steppes, juniper groves and open oak (
<taxonomicName id="1AC16AA158671100FABA9FC0FB8BFD4B" authority="Lam." authorityName="Lam." class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="1274" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="EFB5CD3058671100FABA9FC0FC79FD4A" italics="true" pageId="25" pageNumber="1274">Quercus rotundifolia</emphasis>
Lam.
</taxonomicName>
,
<taxonomicName id="1AC16AA158671100FBCD9FE1FA80FD4B" authority="Willd." authorityName="Willd." box="[1073,1322,687,709]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="1274" phylum="Tracheophyta" rank="species" species="pyrenaica">
<emphasis id="EFB5CD3058671100FBCD9FE1FB7BFD4B" box="[1073,1233,688,709]" italics="true" pageId="25" pageNumber="1274">Q. pyrenaica</emphasis>
Willd.
</taxonomicName>
) and ash (
<taxonomicName id="1AC16AA158671100FCBF9F9EFBD5FD6A" authority="Vahl" authorityName="Vahl" box="[835,1151,718,740]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="25" pageNumber="1274" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="EFB5CD3058671100FCBF9F9EFBEAFD6D" box="[835,1088,718,740]" italics="true" pageId="25" pageNumber="1274">Fraxinus angustifolia</emphasis>
Vahl
</taxonomicName>
) forests (
<bibRefCitation id="B9506CD358671100FB119F9FFC71FC8C" author="Rivas-Martinez S &amp; Costa M" pageId="25" pageNumber="1274" pagination="193 - 224" refId="ref43640" refString="Rivas-Martinez S, Costa M. 1970. Comunidades gipsicolas del centro de Espana. Anales del Instituto Botanico A. J. Cavanilles 27: 193-224." type="journal article" year="1970">Rivas-Martínez &amp; Costa, 1970</bibRefCitation>
;
<bibRefCitation id="B9506CD358671100FC159FBCFAB7FC8C" author="Rivas-Martinez S &amp; Fernandez-Gonzalez F &amp; Mata D &amp; Pizarro JM" box="[1001,1309,749,771]" pageId="25" pageNumber="1274" pagination="3 - 132" refId="ref43670" refString="Rivas-Martinez S, Fernandez-Gonzalez F, Sanchez- Mata D, Pizarro JM. 1990. Vegetacion de la Sierra de Guadarrama. Itinera Geobotanica 4: 3-132." type="journal article" year="1990">
Rivas-Martínez
<emphasis id="EFB5CD3058671100FB5D9FBCFB73FC8C" box="[1185,1241,749,770]" italics="true" pageId="25" pageNumber="1274">et al.</emphasis>
, 1990
</bibRefCitation>
). Biological aspects of this species are expected to be similar to the ones described for the genus by
<bibRefCitation id="B9506CD358671100FB579E7BFAFAFCB1" author="Bologna MA" box="[1195,1360,810,832]" pageId="25" pageNumber="1274" pagination="359 - 366" refId="ref39320" refString="Bologna MA. 1989. Berberomeloe, a new west Mediterranean genus of Lyttini for Meloe majalis Linee (Coleoptera, Meloidae). Systematics and bionomics. Bolletino di Zoologia 55: 359-366." type="journal article" year="1989">Bologna (1989</bibRefCitation>
,
<bibRefCitation id="B9506CD358671100FAA19E7BFA3EFCB1" author="Bologna MA" box="[1373,1428,810,831]" pageId="25" pageNumber="1274" refId="ref39357" refString="Bologna MA. 1991. Fauna d´Italia. XXVIII. Coleoptera Meloidae. Bologna: Edizioni Calderini." type="book" year="1991">1991</bibRefCitation>
). Adult specimens are found from March to July.
</paragraph>
<paragraph id="DD7E112258671100FCB29ECDFA25FC3A" blockId="25.[846,1423,923,978]" box="[846,1423,923,948]" pageId="25" pageNumber="1274">
<taxonomicName id="1AC16AA158671100FCB29ECDFA23FC3D" authority="SANCHEZ-VIALAS ET AL." authorityName="SANCHEZ-VIALAS ET AL." authorityYear="2020" box="[846,1417,923,948]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1274" phylum="Arthropoda" rank="species" species="indalo" status="SP. NOV.">
<emphasis id="EFB5CD3058671100FCB29ECDFBDAFC3C" bold="true" box="[846,1136,924,947]" italics="true" pageId="25" pageNumber="1274">
<smallCapsWord id="DB9887FE58671100FCB29ECDFBBAFC3C" baselines="941,941" box="[846,1040,924,947]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Berberomeloe" pageId="25" pageNumber="1274">BERBEROMELOE</smallCapsWord>
<smallCapsWord id="DB9887FE58671100FBEB9ECEFBDAFC3C" baselines="941" box="[1047,1136,927,946]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="indalo" pageId="25" pageNumber="1274">INDALO</smallCapsWord>
</emphasis>
<smallCapsWord id="DB9887FE58671100FB849ECAFA96FC3D" baselines="942,942,942,942,942" box="[1144,1340,923,948]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Sanchez-Vialas" pageId="25" pageNumber="1274">SÁNCHEZ-VIALAS</smallCapsWord>
<emphasis id="EFB5CD3058671100FABF9EF1FA23FC3D" box="[1347,1417,924,947]" italics="true" pageId="25" pageNumber="1274">
<smallCapsWord id="DB9887FE58671100FABF9EF1FACAFC3C" baselines="941" box="[1347,1376,928,946]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="et" pageId="25" pageNumber="1274">ET</smallCapsWord>
<smallCapsWord id="DB9887FE58671100FA9A9ECEFA28FC3D" baselines="942" box="[1382,1410,927,947]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="al" pageId="25" pageNumber="1274">AL</smallCapsWord>
.
</emphasis>
</taxonomicName>
,
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,820 @@
<document id="041A4A7AE13BD98E76E2FCC3DE46F81C" ID-ISSN="0024-4082" ID-ZooBank="A81CEE6-9D1C-49A9-A9D1-FB0501198097" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353570418" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, García-París, Mario, Ruiz, José L. &amp; Recuero, Ernesto" docDate="2020" docId="5568A03458731109FC8A9A42FAD6FA54" docLanguage="en" docName="zlz164.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Berberomeloe majalis" docType="treatment" docVersion="1" lastPageNumber="1265" masterDocId="A951D84C587E1119FFFC9D51FFAAFF8E" masterDocTitle="Patterns of morphological diversification in giant Berberomeloe blister beetles (Coleoptera: Meloidae) reveal an unexpected taxonomic diversity concordant with mtDNA phylogenetic structure" masterLastPageNumber="1312" masterPageNumber="1249" pageNumber="1262" updateTime="1739358454141" updateUser="GgImagineBatch">
<mods:mods id="ABB6E7E54DBDD703EA012DE2E69100CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ED4360B4D241F9A5FE4B780C57C0D3A9">
<mods:title id="077891401EE5B714FB177DA97A7A1589">Patterns of morphological diversification in giant Berberomeloe blister beetles (Coleoptera: Meloidae) reveal an unexpected taxonomic diversity concordant with mtDNA phylogenetic structure</mods:title>
</mods:titleInfo>
<mods:name id="8CA36BBD7604AE560EE2FE041902ABA0" type="personal">
<mods:role id="098FEA3C8899AB816EA814B60769FD6D">
<mods:roleTerm id="69191FA1E03C9C65E7271C57EC2196B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E361DBC50E33B9ED179794133B4536B8">Sánchez-Vialas, Alberto</mods:namePart>
</mods:name>
<mods:name id="BACAF70B38BE7E3073880E72B8DF90C3" type="personal">
<mods:role id="F81A8B7B854FB018B3FB8EDC6F3B416D">
<mods:roleTerm id="7189A8F8AC420006BB4820A941C61453">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4CDC5AB899ADDD244ECE134AFDE0B7FF">García-París, Mario</mods:namePart>
</mods:name>
<mods:name id="F3F83BE822BE992C38C536811CC971EF" type="personal">
<mods:role id="12333AF546E3F28C0631B7D7BCAFCA43">
<mods:roleTerm id="ED1198A083468F576E4F993B939BC5AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C3E374158FEF58B6A00A3B3D2F5042C4">Ruiz, José L.</mods:namePart>
</mods:name>
<mods:name id="936711E9461BD3C960FFC5D170103E46" type="personal">
<mods:role id="951D67ABFC82E3E4D8AE5A1B673E121D">
<mods:roleTerm id="51C1554FEED1B2AD7FF5C39EF4421C8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69A89193FB642D004DF548E7AE1CA1FE">Recuero, Ernesto</mods:namePart>
</mods:name>
<mods:typeOfResource id="7CBBA33A29B5AF2A78E410468BB65FC9">text</mods:typeOfResource>
<mods:relatedItem id="E6302A9DF628A1498E07693A1771F98D" type="host">
<mods:titleInfo id="9395834252DD9AC00E32DCE117EC2030">
<mods:title id="DFB8AA95175C0A60105EE91B2FB306E1">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="FD6BBEF56B9F471D491A529420ECDF03">
<mods:date id="BF3B75D3F6136C2327843B4C28323301">2020</mods:date>
<mods:detail id="6B6AD3790D15C5B4A1BDD1A98DEAF667" type="volume">
<mods:number id="B0FBE1E51E7BD55567B57435079DB1AC">189</mods:number>
</mods:detail>
<mods:extent id="54C9DE67175BBA04A73B1BC89510B62D" unit="page">
<mods:start id="44101C34D11B80E932CD0C54348639BE">1249</mods:start>
<mods:end id="E1A6E5CC191BEFA78D0700A4A240EAC0">1312</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F5DC530930B427BD231C6B3E75DFC106">journal article</mods:classification>
<mods:identifier id="23CB9F925E44399BBBB0339FCF683BE0" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="DD17775E90EA318B0C98C8551D6768B7" type="ZooBank">A81CEE6-9D1C-49A9-A9D1-FB0501198097</mods:identifier>
</mods:mods>
<treatment id="5568A03458731109FC8A9A42FAD6FA54" LSID="urn:lsid:plazi:treatment:5568A03458731109FC8A9A42FAD6FA54" httpUri="http://treatment.plazi.org/id/5568A03458731109FC8A9A42FAD6FA54" lastPageId="16" lastPageNumber="1265" pageId="13" pageNumber="1262">
<subSubSection id="95DB42A958731114FC8A9A42FACDF8A5" box="[886,1383,1810,1835]" pageId="13" pageNumber="1262" type="nomenclature">
<paragraph id="DD7E112258731114FC8A9A42FACDF8A5" blockId="13.[886,1383,1810,1835]" box="[886,1383,1810,1835]" pageId="13" pageNumber="1262">
<heading id="8636A64E58731114FC8A9A42FACDF8A5" box="[886,1383,1810,1835]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="1262" reason="2">
<taxonomicName id="1AC16AA158731114FC8A9A42FACDF8A5" authority="(LINNAEUS, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[886,1383,1810,1835]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1262" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058731114FC8A9A42FB3FF8A4" box="[886,1173,1811,1834]" italics="true" pageId="13" pageNumber="1262">
<smallCapsWord id="DB9887FE58731114FC8A9A42FB85F8A4" baselines="1828,1829" box="[886,1071,1811,1834]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Berberomeloe" pageId="13" pageNumber="1262">BERBEROMELOE</smallCapsWord>
<smallCapsWord id="DB9887FE58731114FBCA9A47FB3FF8A4" baselines="1829" box="[1078,1173,1814,1834]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="majalis" pageId="13" pageNumber="1262">MAJALIS</smallCapsWord>
</emphasis>
(
<bibRefCitation id="B9506CD358731114FB599A43FAF5F8A5" author="Linnaeus C" box="[1189,1375,1810,1835]" pageId="13" pageNumber="1262" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">
<smallCapsWord id="DB9887FE58731114FB599A43FAB0F8A4" baselines="1829,1829" box="[1189,1306,1810,1835]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Linnaeus" pageId="13" pageNumber="1262">LINNAEUS</smallCapsWord>
, 1758
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95DB42A958731117FCC79A6AFEAFFEFA" lastPageId="14" lastPageNumber="1263" pageId="13" pageNumber="1262" type="reference_group">
<paragraph id="DD7E112258731114FCC79A6AFAE3F8E1" blockId="13.[827,1442,1851,1903]" pageId="13" pageNumber="1262">
<treatmentCitationGroup id="FDD1360C58731114FCC79A6AFAE3F8E1" pageId="13" pageNumber="1262">
<treatmentCitation id="5C60373358731114FCC79A6AFB70F8DE" author="Linnaeus C" box="[827,1242,1851,1873]" page="419" pageId="13" pageNumber="1262" year="1758">
<taxonomicName id="1AC16AA158731114FCC79A6AFB70F8DE" authority="Linnaeus, 1758: 419" authorityName="Linnaeus" authorityPageNumber="419" authorityYear="1758" box="[827,1242,1851,1873]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1262" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058731114FCC79A6AFC4AF8DE" box="[827,992,1851,1872]" italics="true" pageId="13" pageNumber="1262">Meloe majalis</emphasis>
<bibRefCitation id="B9506CD358731114FC149A6AFB70F8DE" author="Linnaeus C" box="[1000,1242,1851,1873]" pageId="13" pageNumber="1262" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758: 419</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Terra typica: in Europa australiore; in America…; in Africa….
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD7E112258701117FF6D9D94FE24FF77" blockId="14.[145,761,197,372]" pageId="14" pageNumber="1263">
<treatmentCitationGroup id="FDD1360C58701117FF6D9D94FE24FF77" pageId="14" pageNumber="1263">
<treatmentCitation id="5C60373358701117FF6D9D94FD5FFF54" author="Escherich K" box="[145,757,197,219]" page="334" pageId="14" pageNumber="1263" year="1889">
<taxonomicName id="1AC16AA158701117FF6D9D94FD5FFF54" authority="Escherich, 1889: 334" authorityName="Escherich" authorityPageNumber="334" authorityYear="1889" box="[145,757,197,219]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="variety" species="majalis" variety="sanguineus">
<emphasis id="EFB5CD3058701117FF6D9D94FE9FFF54" box="[145,309,197,218]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
var.
<emphasis id="EFB5CD3058701117FE8D9D97FE5CFF55" box="[369,502,198,219]" italics="true" pageId="14" pageNumber="1263">sanguineus</emphasis>
<bibRefCitation id="B9506CD358701117FE029D94FD5FFF54" author="Escherich K" box="[510,757,197,219]" pageId="14" pageNumber="1263" pagination="333 - 335" refId="ref40560" refString="Escherich K. 1889. Nachtrage und Berichtigungen zum Catalogus Coleopterorum von Gemminger und Harold, betreffend die Gattung Meloe. Deutsche Entomologische Zeitschrift 1889: 333-335." type="journal article" year="1889">Escherich, 1889: 334</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Terra typica: Cuenca.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD7E112258701117FF6D9C53FF5DFEB8" blockId="14.[145,761,197,372]" pageId="14" pageNumber="1263">
<treatmentCitationGroup id="FDD1360C58701117FF6D9C53FF5DFEB8" pageId="14" pageNumber="1263">
<taxonomicName id="1AC16AA158701117FF6D9C53FDCCFE96" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[145,614,258,280]" class="Insecta" family="Meloidae" genus="Trichomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FF6D9C53FE23FE99" box="[145,393,258,279]" italics="true" pageId="14" pageNumber="1263">Trichomeloe majalis</emphasis>
(
<bibRefCitation id="B9506CD358701117FE619C53FDF0FE96" author="Linnaeus C" box="[413,602,258,280]" pageId="14" pageNumber="1263" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="5C60373358701117FD889C53FF58FEB8" author="MacSwain JW" page="71" pageId="14" pageNumber="1263" year="1956">
<bibRefCitation id="B9506CD358701117FD889C53FF58FEB8" author="MacSwain JW" pageId="14" pageNumber="1263" pagination="1 - 182" refId="ref42239" refString="MacSwain JW. 1956. A classification of the first instar larvae of the Meloidae (Coleoptera). University of California Publications in Entomology 12: 1-182." type="journal article" year="1956">MacSwain, 1956: 71</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD7E112258701117FF6D9C11FEAFFEFA" blockId="14.[145,761,197,372]" pageId="14" pageNumber="1263">
<treatmentCitationGroup id="FDD1360C58701117FF6D9C11FEAFFEFA" pageId="14" pageNumber="1263">
<taxonomicName id="1AC16AA158701117FF6D9C11FDD4FEDB" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[145,638,320,342]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FF6D9C11FE34FEDB" box="[145,414,320,341]" italics="true" pageId="14" pageNumber="1263">Berberomeloe majalis</emphasis>
(
<bibRefCitation id="B9506CD358701117FE4F9C11FDD8FED8" author="Linnaeus C" box="[435,626,320,342]" pageId="14" pageNumber="1263" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="5C60373358701117FD729C11FEAAFEFA" author="Bologna MA" page="361" pageId="14" pageNumber="1263" year="1989">
<bibRefCitation id="B9506CD358701117FD729C11FEAAFEFA" author="Bologna MA" pageId="14" pageNumber="1263" pagination="359 - 366" refId="ref39320" refString="Bologna MA. 1989. Berberomeloe, a new west Mediterranean genus of Lyttini for Meloe majalis Linee (Coleoptera, Meloidae). Systematics and bionomics. Bolletino di Zoologia 55: 359-366." type="journal article" year="1989">Bologna, 1989: 361</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95DB42A958701109FF6D9CFCFAD6FA54" lastPageId="16" lastPageNumber="1265" pageId="14" pageNumber="1263" type="materials_examined">
<paragraph id="DD7E112258701117FF6D9CFCFE1DFE4B" blockId="14.[145,439,429,453]" box="[145,439,429,453]" pageId="14" pageNumber="1263">
<emphasis id="EFB5CD3058701117FF6D9CFCFE1DFE4B" box="[145,439,429,453]" italics="true" pageId="14" pageNumber="1263">Type material examined</emphasis>
</paragraph>
<paragraph id="DD7E112258701117FF6D9C84FEF4FDE8" blockId="14.[145,761,469,798]" pageId="14" pageNumber="1263">
<typeStatus id="027AAF8058701117FF6D9C84FEA8FE64" box="[145,258,469,490]" pageId="14" pageNumber="1263" type="syntype">Syntypes</typeStatus>
of
<taxonomicName id="1AC16AA158701117FED39C84FD00FE65" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[303,682,469,491]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FED39C84FE74FE64" box="[303,478,469,490]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
<bibRefCitation id="B9506CD358701117FE159C84FD00FE65" author="Linnaeus C" box="[489,682,469,491]" pageId="14" pageNumber="1263" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
(
<collectionCode id="BBD089E758701117FD419C84FD59FE64" box="[701,755,469,490]" pageId="14" pageNumber="1263">LSL</collectionCode>
, photographs) (not sexed). LINN 6662, labelled: 2
<taxonomicName id="1AC16AA158701117FF6D9F43FF40FDA9" authorityName="Linnaeus" authorityYear="1758" box="[145,234,530,551]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FF6D9F43FF40FDA9" box="[145,234,530,551]" italics="true" pageId="14" pageNumber="1263">majalis</emphasis>
</taxonomicName>
[handwritten white label]; 2 [printed white label]. LINN 6663, not labelled.
<typeStatus id="027AAF8058701117FE009F63FDC4FDC9" box="[508,622,562,583]" pageId="14" pageNumber="1263" type="lectotype">Lectotype</typeStatus>
: LINN 6662 (designated here).
</paragraph>
<paragraph id="DD7E112258701117FF559F3FFD84FC93" blockId="14.[145,761,469,798]" pageId="14" pageNumber="1263">
<typeStatus id="027AAF8058701117FF559F3FFEA5FD0A" box="[169,271,622,644]" pageId="14" pageNumber="1263" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="1AC16AA158701117FED29F3FFF62FD2C" authority="Escherich, 1889" authorityName="Escherich" authorityYear="1889" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="variety" species="majalis" variety="sanguineus">
<emphasis id="EFB5CD3058701117FED29F3FFE60FD0D" box="[302,458,622,643]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
var.
<emphasis id="EFB5CD3058701117FE019F3EFDD4FD0A" box="[509,638,623,644]" italics="true" pageId="14" pageNumber="1263">sanguineus</emphasis>
<bibRefCitation id="B9506CD358701117FD7F9F3FFF62FD2C" author="Escherich K" pageId="14" pageNumber="1263" pagination="333 - 335" refId="ref40560" refString="Escherich K. 1889. Nachtrage und Berichtigungen zum Catalogus Coleopterorum von Gemminger und Harold, betreffend die Gattung Meloe. Deutsche Entomologische Zeitschrift 1889: 333-335." type="journal article" year="1889">Escherich, 1889</bibRefCitation>
</taxonomicName>
(
<collectionCode id="BBD089E758701117FF289FDCFE92FD2C" box="[212,312,653,674]" pageId="14" pageNumber="1263">MNHAT</collectionCode>
) (not sexed). Specimen labelled: Cuenca Castilien 87. K. [printed white label];
<taxonomicName id="1AC16AA158701117FD949FFDFF7FFD6E" authority="Lucas" authorityName="Lucas" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="maculifrons">
<emphasis id="EFB5CD3058701117FD949FFDFD53FD4F" box="[616,761,684,705]" italics="true" pageId="14" pageNumber="1263">maculifrons</emphasis>
Lucas
</taxonomicName>
[handwritten white label];
<taxonomicName id="1AC16AA158701117FDFB9F9BFEA6FD71" authority="Escherich" authorityName="Escherich" authorityYear="1889" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="variety" species="majalis" variety="sanguineus">
<emphasis id="EFB5CD3058701117FDFB9F9BFDF6FD51" box="[519,604,714,735]" italics="true" pageId="14" pageNumber="1263">majalis</emphasis>
v.
<emphasis id="EFB5CD3058701117FD849F9AFD53FD6E" box="[632,761,715,736]" italics="true" pageId="14" pageNumber="1263">sanguineus</emphasis>
Escherich
</taxonomicName>
[handwritten white label]. Figured in
<bibRefCitation id="B9506CD358701117FF6D9E59FE77FC93" author="Garcia-Paris M &amp; Ruiz JL &amp; Alonso-Zarazaga MA" box="[145,477,776,797]" pageId="14" pageNumber="1263" pagination="165 - 212" refId="ref41109" refString="Garcia-Paris M, Ruiz JL, Alonso-Zarazaga MA. 2010. Catalogo sinonimico de los taxones ibero-baleares de la familia Meloidae (Coleoptera). Graellsia 66: 165-212." type="journal article" year="2010">
García-París
<emphasis id="EFB5CD3058701117FEDB9E59FECAFC93" box="[295,352,776,797]" italics="true" pageId="14" pageNumber="1263">et al.</emphasis>
(2010: 170
</bibRefCitation>
, fig. 2).
</paragraph>
<paragraph id="DD7E112258701117FF6D9E16FD25FA71" blockId="14.[145,761,838,1657]" pageId="14" pageNumber="1263">
<emphasis id="EFB5CD3058701117FF6D9E16FEA2FCF5" italics="true" pageId="14" pageNumber="1263">
<typeStatus id="027AAF8058701117FF6D9E16FEBCFCD2" box="[145,278,839,860]" pageId="14" pageNumber="1263" type="lectotype">Lectotype</typeStatus>
designation and nomenclatural comments:
</emphasis>
The type series of
<taxonomicName id="1AC16AA158701117FE189E34FF64FC14" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FE189E34FD2BFCF4" box="[484,641,869,890]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
<bibRefCitation id="B9506CD358701117FD759E34FF64FC14" author="Linnaeus C" pageId="14" pageNumber="1263" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
consists of two dry-preserved specimens (
<typeStatus id="027AAF8058701117FF659EF2FEA3FC36" box="[153,265,931,952]" pageId="14" pageNumber="1263" type="syntype">syntypes</typeStatus>
) held in the collection of the Linnean Society of London at Burlington House. We examined several photographs of both specimens (available at http://linnean-online.org/), which are catalogued as LINN 6662 and LINN 6663. Revision of qualitative features indicates that LINN 6662 and LINN 6663 do not represent the same taxonomic unit, differing mainly in the width of the tergal red bars and in the punctures of the pronotum. While the identity of LINN 6663 is difficult to ascertain without manipulation and close examination of the specimen, the specimen LINN 6662 can be morphologically assigned to the central-southern Iberian plateau clade (lineage A). So, under the provisions of article 74 of the International Code of Zoological Nomenclature and Declaration 44, Amendment of article 74.7.3 (
<bibRefCitation id="B9506CD358701117FDE0983FFD08FA0D" author="ICZN" box="[540,674,1390,1411]" pageId="14" pageNumber="1263" refId="ref41760" refString="ICZN. 1999. International code of Zoological nomenclature, 4 th edn. London: International Trust for Zoological Nomenclature, c / o The Natural History Museum, xxix + 306." type="journal volume" year="1999">ICZN, 1999</bibRefCitation>
,
<bibRefCitation id="B9506CD358701117FD4D983FFD43FA0D" author="ICZN" box="[689,745,1390,1411]" pageId="14" pageNumber="1263" refId="ref41794" refString="ICZN. 2003. Declaration 44. Amendment of article 74.7. 3. Bulletin of Zoological Nomenclature 60: 263." type="journal volume" year="2003">2003</bibRefCitation>
), we designate the specimen LINN 6662 as
<typeStatus id="027AAF8058701117FD7298DCFD52FA2D" box="[654,760,1421,1443]" pageId="14" pageNumber="1263" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="1AC16AA158701117FF5398FDFDB2FA4C" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[175,536,1452,1474]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FF5398FDFEFDFA4F" box="[175,343,1452,1473]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
<bibRefCitation id="B9506CD358701117FE9C98FDFDB2FA4C" author="Linnaeus C" box="[352,536,1452,1474]" pageId="14" pageNumber="1263" refId="ref42088" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 10 th edn. Vol. 1. Stockholm: L. Salvius, 824." type="journal volume" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
, fixing the unique bearer of the name to the taxon corresponding to lineage A (central-southern Iberian plateau).
</paragraph>
<paragraph id="DD7E112258701117FF559B59FEB6F9F7" blockId="14.[145,761,838,1657]" pageId="14" pageNumber="1263">
Likewise, the study of the
<typeStatus id="027AAF8058701117FE289B59FD9FF990" box="[468,565,1544,1566]" pageId="14" pageNumber="1263" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1AC16AA158701117FDAA9B59FE70F9B5" authority="Escherich, 1889" authorityName="Escherich" authorityYear="1889" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="variety" species="majalis" variety="sanguineus">
<emphasis id="EFB5CD3058701117FDAA9B59FD62F993" box="[598,712,1544,1565]" italics="true" pageId="14" pageNumber="1263">B. majalis</emphasis>
var.
<emphasis id="EFB5CD3058701117FF6D9B76FEBCF9B2" box="[145,278,1575,1596]" italics="true" pageId="14" pageNumber="1263">sanguineus</emphasis>
<bibRefCitation id="B9506CD358701117FEE39B77FE70F9B5" author="Escherich K" box="[287,474,1574,1596]" pageId="14" pageNumber="1263" pagination="333 - 335" refId="ref40560" refString="Escherich K. 1889. Nachtrage und Berichtigungen zum Catalogus Coleopterorum von Gemminger und Harold, betreffend die Gattung Meloe. Deutsche Entomologische Zeitschrift 1889: 333-335." type="journal article" year="1889">Escherich, 1889</bibRefCitation>
</taxonomicName>
confirms the synonymy of this
<taxonomicName id="1AC16AA158701117FF189B14FEDFF9D5" box="[228,373,1605,1627]" pageId="14" pageNumber="1263" rank="variety" variety="with">variety with</taxonomicName>
<emphasis id="EFB5CD3058701117FE819B17FDB5F9D5" box="[381,543,1605,1627]" italics="true" pageId="14" pageNumber="1263">
<taxonomicName id="1AC16AA158701117FE819B17FE5EF9D4" authorityName="Linnaeus" authorityYear="1758" box="[381,500,1605,1627]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">B. majalis</taxonomicName>
<taxonomicNameLabel id="F486704B58701117FE019B17FDB5F9D5" box="[509,543,1606,1627]" pageId="14" pageNumber="1263" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
(see:
<bibRefCitation id="B9506CD358701117FD989B14FEA6F9F7" author="Garcia-Paris M &amp; Ruiz JL &amp; Alonso-Zarazaga MA" pageId="14" pageNumber="1263" pagination="165 - 212" refId="ref41109" refString="Garcia-Paris M, Ruiz JL, Alonso-Zarazaga MA. 2010. Catalogo sinonimico de los taxones ibero-baleares de la familia Meloidae (Coleoptera). Graellsia 66: 165-212." type="journal article" year="2010">
García-París
<emphasis id="EFB5CD3058701117FF6D9B35FF63F9F7" box="[145,201,1636,1657]" italics="true" pageId="14" pageNumber="1263">et al.</emphasis>
, 2010
</bibRefCitation>
).
</paragraph>
<paragraph id="DD7E112258701117FF6D9BF2FDA7F978" blockId="14.[145,761,1698,1904]" pageId="14" pageNumber="1263">
<emphasis id="EFB5CD3058701117FF6D9BF2FDF3F936" box="[145,601,1698,1720]" italics="true" pageId="14" pageNumber="1263">
Description:
<taxonomicName id="1AC16AA158701117FED69BF3FD87F939" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[298,557,1698,1719]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" sensu="stricto" species="majalis">Berberomeloe majalis</taxonomicName>
<taxonomicNameLabel id="F486704B58701117FDCA9BF2FDF3F936" box="[566,601,1699,1720]" pageId="14" pageNumber="1263" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
presents the general traits of the genus (see:
<bibRefCitation id="B9506CD358701117FDF49B90FD01F958" author="Bologna MA" box="[520,683,1729,1751]" pageId="14" pageNumber="1263" pagination="359 - 366" refId="ref39320" refString="Bologna MA. 1989. Berberomeloe, a new west Mediterranean genus of Lyttini for Meloe majalis Linee (Coleoptera, Meloidae). Systematics and bionomics. Bolletino di Zoologia 55: 359-366." type="journal article" year="1989">Bologna, 1989</bibRefCitation>
,
<bibRefCitation id="B9506CD358701117FD459B90FD5BF958" author="Bologna MA" box="[697,753,1729,1750]" pageId="14" pageNumber="1263" refId="ref39357" refString="Bologna MA. 1991. Fauna d´Italia. XXVIII. Coleoptera Meloidae. Bologna: Edizioni Calderini." type="book" year="1991">1991</bibRefCitation>
) and it is characterized as follows:
</paragraph>
<paragraph id="DD7E112258701117FF559BAFFBCEFBFD" blockId="14.[145,761,1698,1904]" lastBlockId="14.[809,1426,197,1139]" pageId="14" pageNumber="1263">
<materialsCitation id="6DA91B7F58701117FF559BAFFBCAFBFD" collectionCode="V, VI" pageId="14" pageNumber="1263" specimenCount="2" typeStatus="holotype">
Length (frons to posterior border of elytra),
<quantity id="1A39BCC758701117FF6D9A4CFEA7F8BC" box="[145,269,1821,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="14" pageNumber="1263" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">1020 mm</quantity>
; maximum total length among preserved, studied specimens,
<quantity id="1A39BCC758701117FE899A6DFE62F8DF" box="[373,456,1852,1874]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="1263" unit="mm" value="60.0">60 mm</quantity>
. General coloration black, usually with red or orange transverse bars on the posterior margin of each abdominal terga, except VII and VIII. Tegument of the head and pronotum finely microreticulated, semi-glossy. Head with punctures dispersed, similar to those of pronotum but sparser; medium-sized punctures over the temples; middle line subtle but perceptible at the vertex. Antennae (male) with antennomeres IIIX subtrapezoidal, not especially expanded at inner apexes; antennomeres
<collectionCode id="BBD089E758701117FAD89CCDFA9CFE3F" box="[1316,1334,412,433]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="1263" type="Museum">V</collectionCode>
and
<collectionCode id="BBD089E758701117FA899CCDFA3BFE3F" box="[1397,1425,412,433]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="1263">VI</collectionCode>
subdentiform, with a yellowish, distinct but diffused area in the inner apex; antennomere XI of normal width, widely cleft at the apex, with the inner end pointed. Pronotum subquadrate, with lateral sides weakly or barely convergent in its posterior two-thirds; presence of
<specimenCount id="CBC7DAAB58701117FCBB9F05FC13FDE7" box="[839,953,596,617]" count="2" pageId="14" pageNumber="1263" type="generic" typeStatus="types">two types</specimenCount>
of punctures, small and medium-sized, all over its surface; puncture density low to medium, mostly covering the sides and fore angles; middle longitudinal line finely impressed. Elytral surface with shallow and dispersed punctures; tegument with fine but marked longitudinal corrugations. Abdomen black, usually with a wide red-orange coloured transverse bar at the caudal margin of each terga, except VII and VIII. Male genitalia (
<figureCitation id="45FA0DA758701117FC1C9E18FBFAFCD1" box="[992,1104,841,863]" captionStart="Figure 6" captionStartId="15.[164,242,1646,1668]" captionTargetBox="[273,1332,224,1606]" captionTargetId="figure-72@15.[273,1332,195,1606]" captionTargetPageId="15" captionText="Figure 6. Habitus and morphological details of Berberomeloe majalis from Villafranca de los Caballeros, Ciudad Real (A) and La Guardia, Toledo (BJ). A, dorsal view of a female; B, male antenna; C, D, E, dorsal, ventral and lateral view of male genitalia, showing parameres and phallobase; F, lateral view of aedeagus; G, SEM micrographs of distal part of aedeagus; H, SEM micrographs of two last antennomeres; I, SEM micrographs of pronotum, dorsal view; J, SEM micrographs of head, frontal view." pageId="14" pageNumber="1263">Fig. 6CF</figureCitation>
) with tegmen (parameres + phallobasis) evenly sclerotized; parameres moderately elongated, length similar or a little longer than phallobase, with distal lobes relatively elongated in lateral view; median lobe (aedeagus) with two similar, acute ventral hooks far from the apex, and well separated between each other; apex obliquely truncated in lateral view; endophallic hook slender. Female genitalia with sclerotized spicules in the inner side of the bursa copulatrix
</materialsCitation>
.
</paragraph>
<paragraph id="DD7E112258701116FCD599C8FE4AF8FE" blockId="14.[809,1426,1177,1904]" lastBlockId="15.[163,778,1822,1905]" lastPageId="15" lastPageNumber="1264" pageId="14" pageNumber="1263">
<emphasis id="EFB5CD3058701117FCD599C8FC05FB20" box="[809,943,1177,1198]" italics="true" pageId="14" pageNumber="1263">Variability:</emphasis>
Body length highly variable, from
<quantity id="1A39BCC758701117FAA999C8FCD1FB43" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="14" pageNumber="1263" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10 to 20 mm</quantity>
. Morphometric variability is shown in
<tableCitation id="9043249958701117FACB99E6FA21FB43" box="[1335,1419,1207,1229]" captionStart="Table 2" captionStartId="16.[145,209,196,217]" captionText="Table 2. Mean and standard deviation (SD) of the morphometric measurements (in mm) from studied Berberomeloe specimens (excluding B. laevigatus and B. insignis)" pageId="14" pageNumber="1263">Table 2</tableCitation>
. Colour intensity of the transverse bar variable, from light-red to blood-red. Populations composed only by entirely black specimens have been recorded in some geographically localized areas in the province of Ciudad Real (Piedrabuena and Carrión de Calatrava) (
<bibRefCitation id="B9506CD358701117FCCE9821FBAFFA0B" author="Bravo C &amp; Mas-Peinado P &amp; Bautista LM &amp; Blanco G &amp; Alonso JC &amp; Garcia-Paris M" box="[818,1029,1391,1413]" pageId="14" pageNumber="1263" pagination="790 - 804" refId="ref39536" refString="Bravo C, Mas-Peinado P, Bautista LM, Blanco G, Alonso JC, Garcia-Paris M. 2017. Cantharidin is conserved across phylogeographic lineages and present in both morphs of Iberian Berberomeloe blister beetles (Coleoptera, Meloidae). Zoological Journal of the Linnean Society 180: 790-804." type="journal article" year="2017">
Bravo
<emphasis id="EFB5CD3058701117FC7D9821FC16FA0A" box="[897,956,1391,1413]" italics="true" pageId="14" pageNumber="1263">et al.</emphasis>
, 2017
</bibRefCitation>
). Some specimens show a single, small red spot on the frons, diffuse and little marked (e.g.
<typeStatus id="027AAF8058701117FC9B98FCFC7BFA4D" box="[871,977,1453,1475]" pageId="14" pageNumber="1263" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1AC16AA158701117FC0398FCFA2AFA4C" authorityName="Escherich" authorityYear="1889" box="[1023,1408,1453,1474]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="variety" species="majalis" variety="sanguineus">
<emphasis id="EFB5CD3058701117FC0398FCFB04FA4C" box="[1023,1198,1453,1474]" italics="true" pageId="14" pageNumber="1263">Meloe majalis</emphasis>
var.
<emphasis id="EFB5CD3058701117FB0F98FCFA2AFA4C" box="[1267,1408,1453,1474]" italics="true" pageId="14" pageNumber="1263">sanguineus</emphasis>
</taxonomicName>
). A slight variation in the density of the puncturation of head and pronotum was observed across populations.
<taxonomicName id="1AC16AA158701117FCD59B58FB9DF990" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[809,1079,1545,1566]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058701117FCD59B58FB9DF990" box="[809,1079,1545,1566]" italics="true" pageId="14" pageNumber="1263">Berberomeloe majalis</emphasis>
</taxonomicName>
presents variability in the width of coloured transverse tergal bars, but on average they are relatively wide (ratio total width of the second tergum/width of the transverse bar of the second tergum between 1.04 and 2.03, average = 1.6, SD = 0.27,
<emphasis id="EFB5CD3058701117FC4B9BF2FC61F936" box="[951,971,1699,1720]" italics="true" pageId="14" pageNumber="1263">N</emphasis>
= 23). Specimens from Penyagolosa (Castellón) show wider coloured transverse tergal bars and deeper punctures. Individuals recently emerged from pupal stage present a non-expanded abdomen, almost completely covered by the elytra; abdomen expands and increases in volume as they start feeding (as reported by
<bibRefCitation id="B9506CD358701117FC1A9A0BFB4DF8E1" author="Cortes-Fossati F" box="[998,1255,1882,1904]" pageId="14" pageNumber="1263" pagination="15 - 20" refId="ref39990" refString="Cortes-Fossati F. 2018 a. Meloidos, ' curitas' y cantaridina: apuntes sobre Berberomeloe majalis (Linnaeus, 1758), un coleoptero peculiar. Sociedad Gaditana de Historia Natural 6: 15-20." type="journal article" year="2018">Cortés-Fossati, 2018a</bibRefCitation>
for
<taxonomicName id="1AC16AA158701117FAE69A0AFA20F8FE" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[1306,1418,1883,1904]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1263" phylum="Arthropoda" rank="species" species="payoyo">
<emphasis id="EFB5CD3058701117FAE69A0AFA20F8FE" box="[1306,1418,1883,1904]" italics="true" pageId="14" pageNumber="1263">B. payoyo</emphasis>
</taxonomicName>
; pers. obs., under laboratory conditions). This pattern of individual abdominal size change is present in all the species of
<taxonomicName id="1AC16AA158711116FEBD9A0AFE73F8FE" authorityName="Bologna" authorityYear="1989" box="[321,473,1883,1904]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD3058711116FEBD9A0AFE73F8FE" box="[321,473,1883,1904]" italics="true" pageId="15" pageNumber="1264">Berberomeloe</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="89BE41AA58711116FF589B3FFA09F952" pageId="15" pageNumber="1264" startId="15.[164,242,1646,1668]" targetBox="[273,1332,224,1606]" targetPageId="15" targetType="figure">
<paragraph id="DD7E112258711116FF589B3FFA09F952" blockId="15.[163,1444,1646,1756]" pageId="15" pageNumber="1264">
<emphasis id="EFB5CD3058711116FF589B3FFEA7F90D" bold="true" box="[164,269,1646,1668]" pageId="15" pageNumber="1264">Figure 6.</emphasis>
Habitus and morphological details of
<taxonomicName id="1AC16AA158711116FD5D9B3FFC28F90A" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[673,898,1646,1668]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1264" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD3058711116FD5D9B3FFC28F90A" box="[673,898,1646,1668]" italics="true" pageId="15" pageNumber="1264">Berberomeloe majalis</emphasis>
</taxonomicName>
from Villafranca de los Caballeros, Ciudad Real (A) and La Guardia, Toledo (BJ). A, dorsal view of a female; B, male antenna; C, D, E, dorsal, ventral and lateral view of male genitalia, showing parameres and phallobase; F, lateral view of aedeagus; G, SEM micrographs of distal part of aedeagus; H, SEM micrographs of two last antennomeres; I, SEM micrographs of pronotum, dorsal view; J, SEM micrographs of head, frontal view.
</paragraph>
</caption>
<paragraph id="DD7E112258711109FCC79A4FFD75FA4C" blockId="15.[827,1443,1822,1905]" lastBlockId="16.[145,761,1084,1474]" lastPageId="16" lastPageNumber="1265" pageId="15" pageNumber="1264">
<emphasis id="EFB5CD3058711116FCC79A4FFB1AF8BD" box="[827,1200,1822,1843]" italics="true" pageId="15" pageNumber="1264">
Diagnosis:
<taxonomicName id="1AC16AA158711116FC469A4FFB1AF8BD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[954,1200,1822,1843]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1264" phylum="Arthropoda" rank="species" species="majalis">Berberomeloe majalis</taxonomicName>
</emphasis>
<materialsCitation id="6DA91B7F58711109FB4B9A4FFD71FA4C" collectionCode="VI" lastPageId="16" lastPageNumber="1265" pageId="15" pageNumber="1264" specimenCount="2" typeStatus="holotype">
can be distinguished from all other species of
<taxonomicName id="1AC16AA158711116FBAC9A6DFB4DF8DF" authorityName="Bologna" authorityYear="1989" box="[1104,1255,1852,1873]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD3058711116FBAC9A6DFB4DF8DF" box="[1104,1255,1852,1873]" italics="true" pageId="15" pageNumber="1264">Berberomeloe</emphasis>
</taxonomicName>
by the following combination of characters (
<figureCitation id="45FA0DA758711116FB809A0AFB14F8FF" box="[1148,1214,1883,1905]" captionStart="Figure 6" captionStartId="15.[164,242,1646,1668]" captionTargetBox="[273,1332,224,1606]" captionTargetId="figure-72@15.[273,1332,195,1606]" captionTargetPageId="15" captionText="Figure 6. Habitus and morphological details of Berberomeloe majalis from Villafranca de los Caballeros, Ciudad Real (A) and La Guardia, Toledo (BJ). A, dorsal view of a female; B, male antenna; C, D, E, dorsal, ventral and lateral view of male genitalia, showing parameres and phallobase; F, lateral view of aedeagus; G, SEM micrographs of distal part of aedeagus; H, SEM micrographs of two last antennomeres; I, SEM micrographs of pronotum, dorsal view; J, SEM micrographs of head, frontal view." pageId="15" pageNumber="1264">Fig. 6</figureCitation>
): (1) abdomen with red-coloured transverse bars of medium to large width in the posterior margin of tergites I
<collectionCode id="BBD089E7586E1109FDCB990AFDFFFBFE" box="[567,597,1115,1136]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="16" pageNumber="1265">VI</collectionCode>
; (2) punctures on the head small to medium-sized, rounded, shallow, isolated from each other, but relatively dense; (3) pronotum subquadrate with rounded, not particularly expanded, fore angles and with lateral margins weakly converging backwards; (4) pronotum surface usually homogeneously punctured, with
<specimenCount id="CBC7DAAB586E1109FD7F9842FD52FAA6" box="[643,760,1299,1320]" count="2" pageId="16" pageNumber="1265" type="generic" typeStatus="types">two types</specimenCount>
of puncturation, small and medium-sized, generally non-confluent; (5) male genitalia with the apex of the median lobe obliquely truncated in lateral view; (6) male antennomere XI wide; and (7) male antennomeres VII and IX moderately expanded on the inner apical side
</materialsCitation>
.
</paragraph>
<caption id="89BE41AA586E1109FF6D9D95FD0FFF79" pageId="16" pageNumber="1265" startId="16.[145,209,196,217]" targetType="table">
<paragraph id="DD7E1122586E1109FF6D9D95FD0FFF79" blockId="16.[145,1363,196,247]" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FF6D9D95FF41FF57" bold="true" box="[145,235,196,217]" pageId="16" pageNumber="1265">Table 2.</emphasis>
Mean and standard deviation (SD) of the morphometric measurements (in mm) from studied
<taxonomicName id="1AC16AA1586E1109FB3B9D95FAF9FF54" authorityName="Bologna" authorityYear="1989" box="[1223,1363,196,218]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FB3B9D95FAF9FF54" box="[1223,1363,196,218]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
specimens (excluding
<taxonomicName id="1AC16AA1586E1109FE899DB3FE51FF79" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" baseAuthorityName="Olivier" baseAuthorityYear="1797" box="[373,507,225,247]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="EFB5CD30586E1109FE899DB3FE51FF79" box="[373,507,225,247]" italics="true" pageId="16" pageNumber="1265">B. laevigatus</emphasis>
</taxonomicName>
and
<taxonomicName id="1AC16AA1586E1109FDD39DB3FD37FF79" baseAuthorityName="Charpentier" baseAuthorityYear="1818" box="[559,669,225,247]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="EFB5CD30586E1109FDD39DB3FD37FF79" box="[559,669,225,247]" italics="true" pageId="16" pageNumber="1265">B. insignis</emphasis>
</taxonomicName>
)
</paragraph>
</caption>
<paragraph id="DD7E1122586E1109FDE99C4AFA24FC77" pageId="16" pageNumber="1265">
<table id="AFC1E382586EEEE6FF6D9C4AFA24FC77" box="[145,1422,283,1017]" gridcols="13" gridrows="22" pageId="16" pageNumber="1265">
<tr id="63F11360586EEEE6FF6D9C4AFA24FEBF" box="[145,1422,283,305]" gridrow="0" pageId="16" pageNumber="1265" rowspan-0="1" rowspan-12="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
<th id="20207A1C586EEEE6FDE99C4AFD19FEBF" box="[533,691,283,305]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FDE99C4AFD19FEBF" authorityName="Bologna" authorityYear="1989" box="[533,691,283,305]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FDE99C4AFD19FEBF" bold="true" box="[533,691,283,305]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</th>
<th id="20207A1C586EEEE6FD079C4AFC33FEBF" box="[763,921,283,305]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FD079C4AFC33FEBF" authorityName="Bologna" authorityYear="1989" box="[763,921,283,305]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FD079C4AFC33FEBF" bold="true" box="[763,921,283,305]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</th>
<th id="20207A1C586EEEE6FC1D9C4AFBD5FEBF" box="[993,1151,283,305]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FC1D9C4AFBD5FEBF" authorityName="Bologna" authorityYear="1989" box="[993,1151,283,305]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FC1D9C4AFBD5FEBF" bold="true" box="[993,1151,283,305]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</th>
<th id="20207A1C586EEEE6FB349C4AFACCFEBF" box="[1224,1382,283,305]" colspan="2" colspanRight="1" gridcol="10" gridrow="0" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FB349C4AFACCFEBF" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[1224,1382,283,305]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="castuo">
<emphasis id="EFB5CD30586E1109FB349C4AFACCFEBF" bold="true" box="[1224,1382,283,305]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="63F11360586EEEE6FF6D9C69FA24FEC0" box="[145,1422,312,334]" gridrow="1" pageId="16" pageNumber="1265" rowspan-0="1" rowspan-11="1" rowspan-12="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="20207A1C586EEEE6FDE99C69FDCBFEC0" box="[533,609,312,334]" gridcol="1" gridrow="1" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FDE99C68FDCBFEC0" bold="true" box="[533,609,313,334]" italics="true" pageId="16" pageNumber="1265">castuo</emphasis>
</td>
<td id="20207A1C586EEEE6FD079C69FCFEFEC0" box="[763,852,312,334]" gridcol="4" gridrow="1" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FD079C68FCE0FEC0" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[763,842,313,334]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="payoyo">
<emphasis id="EFB5CD30586E1109FD079C68FCE0FEC0" bold="true" box="[763,842,313,334]" italics="true" pageId="16" pageNumber="1265">payoyo</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FC1D9C69FB87FEC0" box="[993,1069,312,334]" gridcol="7" gridrow="1" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FC1D9C69FB87FEC0" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[993,1069,312,334]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="indalo">
<emphasis id="EFB5CD30586E1109FC1D9C69FB87FEC0" bold="true" box="[993,1069,312,334]" italics="true" pageId="16" pageNumber="1265">indalo</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FB349C69FAA3FEC0" box="[1224,1289,312,334]" gridcol="10" gridrow="1" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FB349C69FB55FEC0" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[1224,1279,312,334]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="yebli">
<emphasis id="EFB5CD30586E1109FB349C69FB55FEC0" bold="true" box="[1224,1279,312,334]" italics="true" pageId="16" pageNumber="1265">yebli</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="63F11360586EEEE6FF6D9C3EFA24FE0B" box="[145,1422,367,389]" gridrow="2" pageId="16" pageNumber="1265" rowspan-0="1">
<td id="20207A1C586EEEE6FDE99C3EFDCBFE0B" box="[533,609,367,389]" gridcol="1" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FDE99C21FDFCFE0B" bold="true" box="[533,598,368,389]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FD899C3EFD19FE0B" box="[629,691,367,389]" gridcol="2" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD899C3EFD3DFE0B" bold="true" box="[629,663,367,389]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FD3E9C3EFD71FE0B" box="[706,731,367,389]" gridcol="3" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD3E9C21FD7FFE0B" bold="true" box="[706,725,368,389]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FD079C3EFCFEFE0B" box="[763,852,367,389]" gridcol="4" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD079C21FC96FE0B" bold="true" box="[763,828,368,389]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FCA09C3EFC33FE0B" box="[860,921,367,389]" gridcol="5" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FCA09C3EFCD4FE0B" bold="true" box="[860,894,367,389]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FC549C3EFC6BFE0B" box="[936,961,367,389]" gridcol="6" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FC549C21FC11FE0B" bold="true" box="[936,955,368,389]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FC1D9C3EFB87FE0B" box="[993,1069,367,389]" gridcol="7" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FC1D9C21FB88FE0B" bold="true" box="[993,1058,368,389]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FBBE9C3EFBD5FE0B" box="[1090,1151,367,389]" gridcol="8" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FBBE9C3EFBCEFE0B" bold="true" box="[1090,1124,367,389]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FB729C3EFB02FE0B" box="[1166,1192,367,389]" gridcol="9" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FB729C21FB0BFE0B" bold="true" box="[1166,1185,368,389]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FB349C3EFAA3FE0B" box="[1224,1289,367,389]" gridcol="10" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FB349C21FAA3FE0B" bold="true" box="[1224,1289,368,389]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FAD59C3EFACCFE0B" box="[1321,1382,367,389]" gridcol="11" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FAD59C3EFAE1FE0B" bold="true" box="[1321,1355,367,389]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FA899C3EFA24FE0B" box="[1397,1422,367,389]" gridcol="12" gridrow="2" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FA899C21FA22FE0B" bold="true" box="[1397,1416,368,389]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
</tr>
<tr id="63F11360586EEEE6FF6D9CF6FA24FE33" box="[145,1422,423,445]" gridrow="3" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9CF6FE67FE33" box="[145,461,423,445]" gridcol="0" gridrow="3" pageId="16" pageNumber="1265">Pronotum length</th>
<td id="20207A1C586EEEE6FDE99CF6FDCBFE33" box="[533,609,423,445]" gridcol="1" gridrow="3" pageId="16" pageNumber="1265">2.74</td>
<td id="20207A1C586EEEE6FD899CF6FD19FE33" box="[629,691,423,445]" gridcol="2" gridrow="3" pageId="16" pageNumber="1265">0.66</td>
<td id="20207A1C586EEEE6FD3E9CF6FD71FE33" box="[706,731,423,445]" gridcol="3" gridrow="3" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FD079CF6FCFEFE33" box="[763,852,423,445]" gridcol="4" gridrow="3" pageId="16" pageNumber="1265">3.11</td>
<td id="20207A1C586EEEE6FCA09CF6FC33FE33" box="[860,921,423,445]" gridcol="5" gridrow="3" pageId="16" pageNumber="1265">0.7</td>
<td id="20207A1C586EEEE6FC549CF6FC6BFE33" box="[936,961,423,445]" gridcol="6" gridrow="3" pageId="16" pageNumber="1265">21</td>
<td id="20207A1C586EEEE6FC1D9CF6FB87FE33" box="[993,1069,423,445]" gridcol="7" gridrow="3" pageId="16" pageNumber="1265">3.21</td>
<td id="20207A1C586EEEE6FBBE9CF6FBD5FE33" box="[1090,1151,423,445]" gridcol="8" gridrow="3" pageId="16" pageNumber="1265">0.47</td>
<td id="20207A1C586EEEE6FB729CF6FB02FE33" box="[1166,1192,423,445]" gridcol="9" gridrow="3" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349CF6FAA3FE33" box="[1224,1289,423,445]" gridcol="10" gridrow="3" pageId="16" pageNumber="1265">2.64</td>
<td id="20207A1C586EEEE6FAD59CF6FACCFE33" box="[1321,1382,423,445]" gridcol="11" gridrow="3" pageId="16" pageNumber="1265">0.33</td>
<td id="20207A1C586EEEE6FA899CF6FA24FE33" box="[1397,1422,423,445]" gridcol="12" gridrow="3" pageId="16" pageNumber="1265">9</td>
</tr>
<tr id="63F11360586EEEE6FF6D9C94FA24FE55" box="[145,1422,453,475]" gridrow="4" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9C94FE67FE55" box="[145,461,453,475]" gridcol="0" gridrow="4" pageId="16" pageNumber="1265">Pronotum proximal width</th>
<td id="20207A1C586EEEE6FDE99C94FDCBFE55" box="[533,609,453,475]" gridcol="1" gridrow="4" pageId="16" pageNumber="1265">3.6</td>
<td id="20207A1C586EEEE6FD899C94FD19FE55" box="[629,691,453,475]" gridcol="2" gridrow="4" pageId="16" pageNumber="1265">0.89</td>
<td id="20207A1C586EEEE6FD3E9C94FD71FE55" box="[706,731,453,475]" gridcol="3" gridrow="4" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FD079C94FCFEFE55" box="[763,852,453,475]" gridcol="4" gridrow="4" pageId="16" pageNumber="1265">4.38</td>
<td id="20207A1C586EEEE6FCA09C94FC33FE55" box="[860,921,453,475]" gridcol="5" gridrow="4" pageId="16" pageNumber="1265">1.08</td>
<td id="20207A1C586EEEE6FC549C94FC6BFE55" box="[936,961,453,475]" gridcol="6" gridrow="4" pageId="16" pageNumber="1265">21</td>
<td id="20207A1C586EEEE6FC1D9C94FB87FE55" box="[993,1069,453,475]" gridcol="7" gridrow="4" pageId="16" pageNumber="1265">4.53</td>
<td id="20207A1C586EEEE6FBBE9C94FBD5FE55" box="[1090,1151,453,475]" gridcol="8" gridrow="4" pageId="16" pageNumber="1265">0.58</td>
<td id="20207A1C586EEEE6FB729C94FB02FE55" box="[1166,1192,453,475]" gridcol="9" gridrow="4" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349C94FAA3FE55" box="[1224,1289,453,475]" gridcol="10" gridrow="4" pageId="16" pageNumber="1265">3.47</td>
<td id="20207A1C586EEEE6FAD59C94FACCFE55" box="[1321,1382,453,475]" gridcol="11" gridrow="4" pageId="16" pageNumber="1265">0.41</td>
<td id="20207A1C586EEEE6FA899C94FA24FE55" box="[1397,1422,453,475]" gridcol="12" gridrow="4" pageId="16" pageNumber="1265">9</td>
</tr>
<tr id="63F11360586EEEE6FF6D9CB2FA24FE77" box="[145,1422,483,505]" gridrow="5" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9CB2FE67FE77" box="[145,461,483,505]" gridcol="0" gridrow="5" pageId="16" pageNumber="1265">Pronotum distal width</th>
<td id="20207A1C586EEEE6FDE99CB2FDCBFE77" box="[533,609,483,505]" gridcol="1" gridrow="5" pageId="16" pageNumber="1265">3.27</td>
<td id="20207A1C586EEEE6FD899CB2FD19FE77" box="[629,691,483,505]" gridcol="2" gridrow="5" pageId="16" pageNumber="1265">0.79</td>
<td id="20207A1C586EEEE6FD3E9CB2FD71FE77" box="[706,731,483,505]" gridcol="3" gridrow="5" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FD079CB2FCFEFE77" box="[763,852,483,505]" gridcol="4" gridrow="5" pageId="16" pageNumber="1265">4</td>
<td id="20207A1C586EEEE6FCA09CB2FC33FE77" box="[860,921,483,505]" gridcol="5" gridrow="5" pageId="16" pageNumber="1265">0.98</td>
<td id="20207A1C586EEEE6FC549CB2FC6BFE77" box="[936,961,483,505]" gridcol="6" gridrow="5" pageId="16" pageNumber="1265">21</td>
<td id="20207A1C586EEEE6FC1D9CB2FB87FE77" box="[993,1069,483,505]" gridcol="7" gridrow="5" pageId="16" pageNumber="1265">3.92</td>
<td id="20207A1C586EEEE6FBBE9CB2FBD5FE77" box="[1090,1151,483,505]" gridcol="8" gridrow="5" pageId="16" pageNumber="1265">0.55</td>
<td id="20207A1C586EEEE6FB729CB2FB02FE77" box="[1166,1192,483,505]" gridcol="9" gridrow="5" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349CB2FAA3FE77" box="[1224,1289,483,505]" gridcol="10" gridrow="5" pageId="16" pageNumber="1265">3.25</td>
<td id="20207A1C586EEEE6FAD59CB2FACCFE77" box="[1321,1382,483,505]" gridcol="11" gridrow="5" pageId="16" pageNumber="1265">0.44</td>
<td id="20207A1C586EEEE6FA899CB2FA24FE77" box="[1397,1422,483,505]" gridcol="12" gridrow="5" pageId="16" pageNumber="1265">9</td>
</tr>
<tr id="63F11360586EEEE6FF6D9F50FA24FD99" box="[145,1422,513,535]" gridrow="6" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9F50FE67FD99" box="[145,461,513,535]" gridcol="0" gridrow="6" pageId="16" pageNumber="1265">Head width</th>
<td id="20207A1C586EEEE6FDE99F50FDCBFD99" box="[533,609,513,535]" gridcol="1" gridrow="6" pageId="16" pageNumber="1265">4.37</td>
<td id="20207A1C586EEEE6FD899F50FD19FD99" box="[629,691,513,535]" gridcol="2" gridrow="6" pageId="16" pageNumber="1265">1.09</td>
<td id="20207A1C586EEEE6FD3E9F50FD71FD99" box="[706,731,513,535]" gridcol="3" gridrow="6" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FD079F50FCFEFD99" box="[763,852,513,535]" gridcol="4" gridrow="6" pageId="16" pageNumber="1265">5.1</td>
<td id="20207A1C586EEEE6FCA09F50FC33FD99" box="[860,921,513,535]" gridcol="5" gridrow="6" pageId="16" pageNumber="1265">1.24</td>
<td id="20207A1C586EEEE6FC549F50FC6BFD99" box="[936,961,513,535]" gridcol="6" gridrow="6" pageId="16" pageNumber="1265">21</td>
<td id="20207A1C586EEEE6FC1D9F50FB87FD99" box="[993,1069,513,535]" gridcol="7" gridrow="6" pageId="16" pageNumber="1265">5.43</td>
<td id="20207A1C586EEEE6FBBE9F50FBD5FD99" box="[1090,1151,513,535]" gridcol="8" gridrow="6" pageId="16" pageNumber="1265">0.77</td>
<td id="20207A1C586EEEE6FB729F50FB02FD99" box="[1166,1192,513,535]" gridcol="9" gridrow="6" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349F50FAA3FD99" box="[1224,1289,513,535]" gridcol="10" gridrow="6" pageId="16" pageNumber="1265">4.13</td>
<td id="20207A1C586EEEE6FAD59F50FACCFD99" box="[1321,1382,513,535]" gridcol="11" gridrow="6" pageId="16" pageNumber="1265">0.6</td>
<td id="20207A1C586EEEE6FA899F50FA24FD99" box="[1397,1422,513,535]" gridcol="12" gridrow="6" pageId="16" pageNumber="1265">9</td>
</tr>
<tr id="63F11360586EEEE6FF6D9F4EFA24FDBB" box="[145,1422,543,565]" gridrow="7" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9F4EFE67FDBB" box="[145,461,543,565]" gridcol="0" gridrow="7" pageId="16" pageNumber="1265">Width of the second tergal bar</th>
<td id="20207A1C586EEEE6FDE99F4EFDCBFDBB" box="[533,609,543,565]" gridcol="1" gridrow="7" pageId="16" pageNumber="1265">4.01</td>
<td id="20207A1C586EEEE6FD899F4EFD19FDBB" box="[629,691,543,565]" gridcol="2" gridrow="7" pageId="16" pageNumber="1265">1.16</td>
<td id="20207A1C586EEEE6FD3E9F4EFD71FDBB" box="[706,731,543,565]" gridcol="3" gridrow="7" pageId="16" pageNumber="1265">28</td>
<td id="20207A1C586EEEE6FD079F4EFCFEFDBB" box="[763,852,543,565]" gridcol="4" gridrow="7" pageId="16" pageNumber="1265">4.68</td>
<td id="20207A1C586EEEE6FCA09F4EFC33FDBB" box="[860,921,543,565]" gridcol="5" gridrow="7" pageId="16" pageNumber="1265">1.09</td>
<td id="20207A1C586EEEE6FC549F4EFC6BFDBB" box="[936,961,543,565]" gridcol="6" gridrow="7" pageId="16" pageNumber="1265">18</td>
<td id="20207A1C586EEEE6FC1D9F4EFB87FDBB" box="[993,1069,543,565]" gridcol="7" gridrow="7" pageId="16" pageNumber="1265">7.99</td>
<td id="20207A1C586EEEE6FBBE9F4EFBD5FDBB" box="[1090,1151,543,565]" gridcol="8" gridrow="7" pageId="16" pageNumber="1265">1.47</td>
<td id="20207A1C586EEEE6FB729F4EFB02FDBB" box="[1166,1192,543,565]" gridcol="9" gridrow="7" pageId="16" pageNumber="1265">15</td>
<td id="20207A1C586EEEE6FB349F4EFAA3FDBB" box="[1224,1289,543,565]" gridcol="10" gridrow="7" pageId="16" pageNumber="1265">4.28</td>
<td id="20207A1C586EEEE6FAD59F4EFACCFDBB" box="[1321,1382,543,565]" gridcol="11" gridrow="7" pageId="16" pageNumber="1265">0.78</td>
<td id="20207A1C586EEEE6FA899F4EFA24FDBB" box="[1397,1422,543,565]" gridcol="12" gridrow="7" pageId="16" pageNumber="1265">7</td>
</tr>
<tr id="63F11360586EEEE6FF6D9F6CFA24FDDD" box="[145,1422,573,595]" gridrow="8" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9F6CFE67FDDD" box="[145,461,573,595]" gridcol="0" gridrow="8" pageId="16" pageNumber="1265">Width of the third tergal bar</th>
<td id="20207A1C586EEEE6FDE99F6CFDCBFDDD" box="[533,609,573,595]" gridcol="1" gridrow="8" pageId="16" pageNumber="1265">4.31</td>
<td id="20207A1C586EEEE6FD899F6CFD19FDDD" box="[629,691,573,595]" gridcol="2" gridrow="8" pageId="16" pageNumber="1265">1.21</td>
<td id="20207A1C586EEEE6FD3E9F6CFD71FDDD" box="[706,731,573,595]" gridcol="3" gridrow="8" pageId="16" pageNumber="1265">28</td>
<td id="20207A1C586EEEE6FD079F6CFCFEFDDD" box="[763,852,573,595]" gridcol="4" gridrow="8" pageId="16" pageNumber="1265">4.97</td>
<td id="20207A1C586EEEE6FCA09F6CFC33FDDD" box="[860,921,573,595]" gridcol="5" gridrow="8" pageId="16" pageNumber="1265">1.27</td>
<td id="20207A1C586EEEE6FC549F6CFC6BFDDD" box="[936,961,573,595]" gridcol="6" gridrow="8" pageId="16" pageNumber="1265">19</td>
<td id="20207A1C586EEEE6FC1D9F6CFB87FDDD" box="[993,1069,573,595]" gridcol="7" gridrow="8" pageId="16" pageNumber="1265">8.39</td>
<td id="20207A1C586EEEE6FBBE9F6CFBD5FDDD" box="[1090,1151,573,595]" gridcol="8" gridrow="8" pageId="16" pageNumber="1265">1.34</td>
<td id="20207A1C586EEEE6FB729F6CFB02FDDD" box="[1166,1192,573,595]" gridcol="9" gridrow="8" pageId="16" pageNumber="1265">15</td>
<td id="20207A1C586EEEE6FB349F6CFAA3FDDD" box="[1224,1289,573,595]" gridcol="10" gridrow="8" pageId="16" pageNumber="1265">4.75</td>
<td id="20207A1C586EEEE6FAD59F6CFACCFDDD" box="[1321,1382,573,595]" gridcol="11" gridrow="8" pageId="16" pageNumber="1265">0.76</td>
<td id="20207A1C586EEEE6FA899F6CFA24FDDD" box="[1397,1422,573,595]" gridcol="12" gridrow="8" pageId="16" pageNumber="1265">8</td>
</tr>
<tr id="63F11360586EEEE6FF6D9F0AFA24FDFF" box="[145,1422,603,625]" gridrow="9" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9F0AFE67FDFF" box="[145,461,603,625]" gridcol="0" gridrow="9" pageId="16" pageNumber="1265">Longitude of metafemur</th>
<td id="20207A1C586EEEE6FDE99F0AFDCBFDFF" box="[533,609,603,625]" gridcol="1" gridrow="9" pageId="16" pageNumber="1265">4.07</td>
<td id="20207A1C586EEEE6FD899F0AFD19FDFF" box="[629,691,603,625]" gridcol="2" gridrow="9" pageId="16" pageNumber="1265">1</td>
<td id="20207A1C586EEEE6FD3E9F0AFD71FDFF" box="[706,731,603,625]" gridcol="3" gridrow="9" pageId="16" pageNumber="1265">25</td>
<td id="20207A1C586EEEE6FD079F0AFCFEFDFF" box="[763,852,603,625]" gridcol="4" gridrow="9" pageId="16" pageNumber="1265">4.71</td>
<td id="20207A1C586EEEE6FCA09F0AFC33FDFF" box="[860,921,603,625]" gridcol="5" gridrow="9" pageId="16" pageNumber="1265">1.05</td>
<td id="20207A1C586EEEE6FC549F0AFC6BFDFF" box="[936,961,603,625]" gridcol="6" gridrow="9" pageId="16" pageNumber="1265">20</td>
<td id="20207A1C586EEEE6FC1D9F0AFB87FDFF" box="[993,1069,603,625]" gridcol="7" gridrow="9" pageId="16" pageNumber="1265">5.24</td>
<td id="20207A1C586EEEE6FBBE9F0AFBD5FDFF" box="[1090,1151,603,625]" gridcol="8" gridrow="9" pageId="16" pageNumber="1265">0.7</td>
<td id="20207A1C586EEEE6FB729F0AFB02FDFF" box="[1166,1192,603,625]" gridcol="9" gridrow="9" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349F0AFAA3FDFF" box="[1224,1289,603,625]" gridcol="10" gridrow="9" pageId="16" pageNumber="1265">3.75</td>
<td id="20207A1C586EEEE6FAD59F0AFACCFDFF" box="[1321,1382,603,625]" gridcol="11" gridrow="9" pageId="16" pageNumber="1265">0.41</td>
<td id="20207A1C586EEEE6FA899F0AFA24FDFF" box="[1397,1422,603,625]" gridcol="12" gridrow="9" pageId="16" pageNumber="1265">6</td>
</tr>
<tr id="63F11360586EEEE6FF6D9F28FA24FD01" box="[145,1422,633,655]" gridrow="10" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9F28FE67FD01" box="[145,461,633,655]" gridcol="0" gridrow="10" pageId="16" pageNumber="1265">Longitude of elytra</th>
<td id="20207A1C586EEEE6FDE99F28FDCBFD01" box="[533,609,633,655]" gridcol="1" gridrow="10" pageId="16" pageNumber="1265">7.59</td>
<td id="20207A1C586EEEE6FD899F28FD19FD01" box="[629,691,633,655]" gridcol="2" gridrow="10" pageId="16" pageNumber="1265">1.68</td>
<td id="20207A1C586EEEE6FD3E9F28FD71FD01" box="[706,731,633,655]" gridcol="3" gridrow="10" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FD079F28FCFEFD01" box="[763,852,633,655]" gridcol="4" gridrow="10" pageId="16" pageNumber="1265">9.22</td>
<td id="20207A1C586EEEE6FCA09F28FC33FD01" box="[860,921,633,655]" gridcol="5" gridrow="10" pageId="16" pageNumber="1265">2.07</td>
<td id="20207A1C586EEEE6FC549F28FC6BFD01" box="[936,961,633,655]" gridcol="6" gridrow="10" pageId="16" pageNumber="1265">20</td>
<td id="20207A1C586EEEE6FC1D9F28FB87FD01" box="[993,1069,633,655]" gridcol="7" gridrow="10" pageId="16" pageNumber="1265">9.44</td>
<td id="20207A1C586EEEE6FBBE9F28FBD5FD01" box="[1090,1151,633,655]" gridcol="8" gridrow="10" pageId="16" pageNumber="1265">1.36</td>
<td id="20207A1C586EEEE6FB729F28FB02FD01" box="[1166,1192,633,655]" gridcol="9" gridrow="10" pageId="16" pageNumber="1265">17</td>
<td id="20207A1C586EEEE6FB349F28FAA3FD01" box="[1224,1289,633,655]" gridcol="10" gridrow="10" pageId="16" pageNumber="1265">7.48</td>
<td id="20207A1C586EEEE6FAD59F28FACCFD01" box="[1321,1382,633,655]" gridcol="11" gridrow="10" pageId="16" pageNumber="1265">1.04</td>
<td id="20207A1C586EEEE6FA899F28FA24FD01" box="[1397,1422,633,655]" gridcol="12" gridrow="10" pageId="16" pageNumber="1265">9</td>
</tr>
<tr id="63F11360586EEEE6FF6D9FCEFA24FD3B" box="[145,1422,671,693]" gridrow="11" pageId="16" pageNumber="1265" rowspan-0="1" rowspan-12="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
<td id="20207A1C586EEEE6FDE99FCEFD19FD3B" box="[533,691,671,693]" colspan="2" colspanRight="1" gridcol="1" gridrow="11" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FDE99FCEFD19FD3B" authorityName="Bologna" authorityYear="1989" box="[533,691,671,693]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FDE99FCEFD19FD3B" bold="true" box="[533,691,671,693]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FD079FCEFC33FD3B" box="[763,921,671,693]" colspan="2" colspanRight="1" gridcol="4" gridrow="11" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FD079FCEFC33FD3B" authorityName="Bologna" authorityYear="1989" box="[763,921,671,693]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FD079FCEFC33FD3B" bold="true" box="[763,921,671,693]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FC1D9FCEFBD5FD3B" box="[993,1151,671,693]" colspan="2" colspanRight="1" gridcol="7" gridrow="11" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FC1D9FCEFBD5FD3B" authorityName="Bologna" authorityYear="1989" box="[993,1151,671,693]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="EFB5CD30586E1109FC1D9FCEFBD5FD3B" bold="true" box="[993,1151,671,693]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FB349FCEFACCFD3B" box="[1224,1382,671,693]" colspan="2" colspanRight="1" gridcol="10" gridrow="11" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FB349FCEFACCFD3B" box="[1224,1382,671,693]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1306" phylum="Arthropoda" rank="species" species="maculifrons">
<emphasis id="EFB5CD30586E1109FB349FCEFACCFD3B" bold="true" box="[1224,1382,671,693]" italics="true" pageId="16" pageNumber="1265">Berberomeloe</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="63F11360586EEEE6FF6D9FEDFA24FD5C" box="[145,1422,700,722]" gridrow="12" pageId="16" pageNumber="1265" rowspan-0="1" rowspan-12="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
<td id="20207A1C586EEEE6FDE99FEDFD19FD5C" box="[533,691,700,722]" colspan="2" colspanRight="1" gridcol="1" gridrow="12" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FDE99FEDFD09FD5C" bold="true" box="[533,675,700,722]" italics="true" pageId="16" pageNumber="1265">maculifrons</emphasis>
</td>
<td id="20207A1C586EEEE6FD079FEDFCFEFD5C" box="[763,852,700,722]" gridcol="4" gridrow="12" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FD079FEDFCFEFD5C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[763,852,700,722]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD30586E1109FD079FEDFCFEFD5C" bold="true" box="[763,852,700,722]" italics="true" pageId="16" pageNumber="1265">majalis</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FC1D9FEDFBD5FD5C" box="[993,1151,700,722]" colspan="2" colspanRight="1" gridcol="7" gridrow="12" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FC1D9FECFBF9FD5C" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[993,1107,701,722]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="comunero">
<emphasis id="EFB5CD30586E1109FC1D9FECFBF9FD5C" bold="true" box="[993,1107,701,722]" italics="true" pageId="16" pageNumber="1265">comunero</emphasis>
</taxonomicName>
</td>
<td id="20207A1C586EEEE6FB349FEDFACCFD5C" box="[1224,1382,700,722]" colspan="2" colspanRight="1" gridcol="10" gridrow="12" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FB349FEDFAEEFD5C" authorityName="Sánchez-Vialas &amp; García-París &amp; Ruiz &amp; Recuero" authorityYear="2020" box="[1224,1348,700,722]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="EFB5CD30586E1109FB349FEDFAEEFD5C" bold="true" box="[1224,1348,700,722]" italics="true" pageId="16" pageNumber="1265">tenebrosus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="63F11360586EEEE6FF6D9FB9FA24FD70" box="[145,1422,744,766]" gridrow="13" pageId="16" pageNumber="1265" rowspan-0="1">
<td id="20207A1C586EEEE6FDE99FB9FDCBFD70" box="[533,609,744,766]" gridcol="1" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FDE99FB9FDFCFD73" bold="true" box="[533,598,744,765]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FD899FB9FD19FD70" box="[629,691,744,766]" gridcol="2" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD899FB9FD3DFD70" bold="true" box="[629,663,744,766]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FD3E9FB9FD71FD70" box="[706,731,744,766]" gridcol="3" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD3E9FB9FD7FFD73" bold="true" box="[706,725,744,765]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FD079FB9FCFEFD70" box="[763,852,744,766]" gridcol="4" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FD079FB9FC96FD73" bold="true" box="[763,828,744,765]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FCA09FB9FC33FD70" box="[860,921,744,766]" gridcol="5" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FCA09FB9FCD4FD70" bold="true" box="[860,894,744,766]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FC549FB9FC6BFD70" box="[936,961,744,766]" gridcol="6" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FC549FB9FC11FD73" bold="true" box="[936,955,744,765]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FC1D9FB9FB87FD70" box="[993,1069,744,766]" gridcol="7" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FC1D9FB9FB88FD73" bold="true" box="[993,1058,744,765]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FBBE9FB9FBD5FD70" box="[1090,1151,744,766]" gridcol="8" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FBBE9FB9FBCEFD70" bold="true" box="[1090,1124,744,766]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FB729FB9FB02FD70" box="[1166,1192,744,766]" gridcol="9" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FB729FB9FB0BFD73" bold="true" box="[1166,1185,744,765]" pageId="16" pageNumber="1265">N</emphasis>
</td>
<td id="20207A1C586EEEE6FB349FB9FAA3FD70" box="[1224,1289,744,766]" gridcol="10" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FB349FB9FAA3FD73" bold="true" box="[1224,1289,744,765]" pageId="16" pageNumber="1265">Mean</emphasis>
</td>
<td id="20207A1C586EEEE6FAD59FB9FACCFD70" box="[1321,1382,744,766]" gridcol="11" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FAD59FB9FAE1FD70" bold="true" box="[1321,1355,744,766]" pageId="16" pageNumber="1265">SD</emphasis>
</td>
<td id="20207A1C586EEEE6FA899FB9FA24FD70" box="[1397,1422,744,766]" gridcol="12" gridrow="13" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FA899FB9FA22FD73" bold="true" box="[1397,1416,744,765]" italics="true" pageId="16" pageNumber="1265">N</emphasis>
</td>
</tr>
<tr id="63F11360586EEEE6FF6D9E40FA24FCA9" box="[145,1422,785,807]" gridrow="14" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9E40FE67FCA9" box="[145,461,785,807]" gridcol="0" gridrow="14" pageId="16" pageNumber="1265">Pronotum length</th>
<td id="20207A1C586EEEE6FDE99E40FDCBFCA9" box="[533,609,785,807]" gridcol="1" gridrow="14" pageId="16" pageNumber="1265">2.79</td>
<td id="20207A1C586EEEE6FD899E40FD19FCA9" box="[629,691,785,807]" gridcol="2" gridrow="14" pageId="16" pageNumber="1265">0.53</td>
<td id="20207A1C586EEEE6FD3E9E40FD71FCA9" box="[706,731,785,807]" gridcol="3" gridrow="14" pageId="16" pageNumber="1265">39</td>
<td id="20207A1C586EEEE6FD079E40FCFEFCA9" box="[763,852,785,807]" gridcol="4" gridrow="14" pageId="16" pageNumber="1265">2.83</td>
<td id="20207A1C586EEEE6FCA09E40FC33FCA9" box="[860,921,785,807]" gridcol="5" gridrow="14" pageId="16" pageNumber="1265">0.51</td>
<td id="20207A1C586EEEE6FC549E40FC6BFCA9" box="[936,961,785,807]" gridcol="6" gridrow="14" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FC1D9E40FB87FCA9" box="[993,1069,785,807]" gridcol="7" gridrow="14" pageId="16" pageNumber="1265">2.81</td>
<td id="20207A1C586EEEE6FBBE9E40FBD5FCA9" box="[1090,1151,785,807]" gridcol="8" gridrow="14" pageId="16" pageNumber="1265">0.6</td>
<td id="20207A1C586EEEE6FB729E40FB02FCA9" box="[1166,1192,785,807]" gridcol="9" gridrow="14" pageId="16" pageNumber="1265">44</td>
<td id="20207A1C586EEEE6FB349E40FAA3FCA9" box="[1224,1289,785,807]" gridcol="10" gridrow="14" pageId="16" pageNumber="1265">2.98</td>
<td id="20207A1C586EEEE6FAD59E40FACCFCA9" box="[1321,1382,785,807]" gridcol="11" gridrow="14" pageId="16" pageNumber="1265">0.5</td>
<td id="20207A1C586EEEE6FA899E40FA24FCA9" box="[1397,1422,785,807]" gridcol="12" gridrow="14" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9E7EFA24FCCB" box="[145,1422,815,837]" gridrow="15" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9E7EFE67FCCB" box="[145,461,815,837]" gridcol="0" gridrow="15" pageId="16" pageNumber="1265">Pronotum proximal width</th>
<td id="20207A1C586EEEE6FDE99E7EFDCBFCCB" box="[533,609,815,837]" gridcol="1" gridrow="15" pageId="16" pageNumber="1265">3.72</td>
<td id="20207A1C586EEEE6FD899E7EFD19FCCB" box="[629,691,815,837]" gridcol="2" gridrow="15" pageId="16" pageNumber="1265">0.74</td>
<td id="20207A1C586EEEE6FD3E9E7EFD71FCCB" box="[706,731,815,837]" gridcol="3" gridrow="15" pageId="16" pageNumber="1265">39</td>
<td id="20207A1C586EEEE6FD079E7EFCFEFCCB" box="[763,852,815,837]" gridcol="4" gridrow="15" pageId="16" pageNumber="1265">3.69</td>
<td id="20207A1C586EEEE6FCA09E7EFC33FCCB" box="[860,921,815,837]" gridcol="5" gridrow="15" pageId="16" pageNumber="1265">0.66</td>
<td id="20207A1C586EEEE6FC549E7EFC6BFCCB" box="[936,961,815,837]" gridcol="6" gridrow="15" pageId="16" pageNumber="1265">28</td>
<td id="20207A1C586EEEE6FC1D9E7EFB87FCCB" box="[993,1069,815,837]" gridcol="7" gridrow="15" pageId="16" pageNumber="1265">3.63</td>
<td id="20207A1C586EEEE6FBBE9E7EFBD5FCCB" box="[1090,1151,815,837]" gridcol="8" gridrow="15" pageId="16" pageNumber="1265">0.86</td>
<td id="20207A1C586EEEE6FB729E7EFB02FCCB" box="[1166,1192,815,837]" gridcol="9" gridrow="15" pageId="16" pageNumber="1265">44</td>
<td id="20207A1C586EEEE6FB349E7EFAA3FCCB" box="[1224,1289,815,837]" gridcol="10" gridrow="15" pageId="16" pageNumber="1265">4.07</td>
<td id="20207A1C586EEEE6FAD59E7EFACCFCCB" box="[1321,1382,815,837]" gridcol="11" gridrow="15" pageId="16" pageNumber="1265">0.64</td>
<td id="20207A1C586EEEE6FA899E7EFA24FCCB" box="[1397,1422,815,837]" gridcol="12" gridrow="15" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9E1CFA24FCED" box="[145,1422,845,867]" gridrow="16" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9E1CFE67FCED" box="[145,461,845,867]" gridcol="0" gridrow="16" pageId="16" pageNumber="1265">Pronotum distal width</th>
<td id="20207A1C586EEEE6FDE99E1CFDCBFCED" box="[533,609,845,867]" gridcol="1" gridrow="16" pageId="16" pageNumber="1265">3.42</td>
<td id="20207A1C586EEEE6FD899E1CFD19FCED" box="[629,691,845,867]" gridcol="2" gridrow="16" pageId="16" pageNumber="1265">0.66</td>
<td id="20207A1C586EEEE6FD3E9E1CFD71FCED" box="[706,731,845,867]" gridcol="3" gridrow="16" pageId="16" pageNumber="1265">39</td>
<td id="20207A1C586EEEE6FD079E1CFCFEFCED" box="[763,852,845,867]" gridcol="4" gridrow="16" pageId="16" pageNumber="1265">3.35</td>
<td id="20207A1C586EEEE6FCA09E1CFC33FCED" box="[860,921,845,867]" gridcol="5" gridrow="16" pageId="16" pageNumber="1265">0.59</td>
<td id="20207A1C586EEEE6FC549E1CFC6BFCED" box="[936,961,845,867]" gridcol="6" gridrow="16" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FC1D9E1CFB87FCED" box="[993,1069,845,867]" gridcol="7" gridrow="16" pageId="16" pageNumber="1265">3.35</td>
<td id="20207A1C586EEEE6FBBE9E1CFBD5FCED" box="[1090,1151,845,867]" gridcol="8" gridrow="16" pageId="16" pageNumber="1265">0.8</td>
<td id="20207A1C586EEEE6FB729E1CFB02FCED" box="[1166,1192,845,867]" gridcol="9" gridrow="16" pageId="16" pageNumber="1265">44</td>
<td id="20207A1C586EEEE6FB349E1CFAA3FCED" box="[1224,1289,845,867]" gridcol="10" gridrow="16" pageId="16" pageNumber="1265">3.38</td>
<td id="20207A1C586EEEE6FAD59E1CFACCFCED" box="[1321,1382,845,867]" gridcol="11" gridrow="16" pageId="16" pageNumber="1265">0.6</td>
<td id="20207A1C586EEEE6FA899E1CFA24FCED" box="[1397,1422,845,867]" gridcol="12" gridrow="16" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9E3AFA24FC0F" box="[145,1422,875,897]" gridrow="17" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9E3AFE67FC0F" box="[145,461,875,897]" gridcol="0" gridrow="17" pageId="16" pageNumber="1265">Head width</th>
<td id="20207A1C586EEEE6FDE99E3AFDCBFC0F" box="[533,609,875,897]" gridcol="1" gridrow="17" pageId="16" pageNumber="1265">4.46</td>
<td id="20207A1C586EEEE6FD899E3AFD19FC0F" box="[629,691,875,897]" gridcol="2" gridrow="17" pageId="16" pageNumber="1265">0.98</td>
<td id="20207A1C586EEEE6FD3E9E3AFD71FC0F" box="[706,731,875,897]" gridcol="3" gridrow="17" pageId="16" pageNumber="1265">39</td>
<td id="20207A1C586EEEE6FD079E3AFCFEFC0F" box="[763,852,875,897]" gridcol="4" gridrow="17" pageId="16" pageNumber="1265">4.48</td>
<td id="20207A1C586EEEE6FCA09E3AFC33FC0F" box="[860,921,875,897]" gridcol="5" gridrow="17" pageId="16" pageNumber="1265">0.82</td>
<td id="20207A1C586EEEE6FC549E3AFC6BFC0F" box="[936,961,875,897]" gridcol="6" gridrow="17" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FC1D9E3AFB87FC0F" box="[993,1069,875,897]" gridcol="7" gridrow="17" pageId="16" pageNumber="1265">4.41</td>
<td id="20207A1C586EEEE6FBBE9E3AFBD5FC0F" box="[1090,1151,875,897]" gridcol="8" gridrow="17" pageId="16" pageNumber="1265">1.04</td>
<td id="20207A1C586EEEE6FB729E3AFB02FC0F" box="[1166,1192,875,897]" gridcol="9" gridrow="17" pageId="16" pageNumber="1265">44</td>
<td id="20207A1C586EEEE6FB349E3AFAA3FC0F" box="[1224,1289,875,897]" gridcol="10" gridrow="17" pageId="16" pageNumber="1265">4.57</td>
<td id="20207A1C586EEEE6FAD59E3AFACCFC0F" box="[1321,1382,875,897]" gridcol="11" gridrow="17" pageId="16" pageNumber="1265">0.8</td>
<td id="20207A1C586EEEE6FA899E3AFA24FC0F" box="[1397,1422,875,897]" gridcol="12" gridrow="17" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9ED8FA24FC11" box="[145,1422,905,927]" gridrow="18" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9ED8FE67FC11" box="[145,461,905,927]" gridcol="0" gridrow="18" pageId="16" pageNumber="1265">Width of the second tergal bar</th>
<td id="20207A1C586EEEE6FDE99ED8FDCBFC11" box="[533,609,905,927]" gridcol="1" gridrow="18" pageId="16" pageNumber="1265">6.05</td>
<td id="20207A1C586EEEE6FD899ED8FD19FC11" box="[629,691,905,927]" gridcol="2" gridrow="18" pageId="16" pageNumber="1265">1.2</td>
<td id="20207A1C586EEEE6FD3E9ED8FD71FC11" box="[706,731,905,927]" gridcol="3" gridrow="18" pageId="16" pageNumber="1265">33</td>
<td id="20207A1C586EEEE6FD079ED8FCFEFC11" box="[763,852,905,927]" gridcol="4" gridrow="18" pageId="16" pageNumber="1265">4.72</td>
<td id="20207A1C586EEEE6FCA09ED8FC33FC11" box="[860,921,905,927]" gridcol="5" gridrow="18" pageId="16" pageNumber="1265">1.34</td>
<td id="20207A1C586EEEE6FC549ED8FC6BFC11" box="[936,961,905,927]" gridcol="6" gridrow="18" pageId="16" pageNumber="1265">24</td>
<td id="20207A1C586EEEE6FC1D9ED8FB87FC11" box="[993,1069,905,927]" gridcol="7" gridrow="18" pageId="16" pageNumber="1265">4.97</td>
<td id="20207A1C586EEEE6FBBE9ED8FBD5FC11" box="[1090,1151,905,927]" gridcol="8" gridrow="18" pageId="16" pageNumber="1265">1.43</td>
<td id="20207A1C586EEEE6FB729ED8FB02FC11" box="[1166,1192,905,927]" gridcol="9" gridrow="18" pageId="16" pageNumber="1265">35</td>
<td id="20207A1C586EEEE6FB349ED8FAA3FC11" box="[1224,1289,905,927]" gridcol="10" gridrow="18" pageId="16" pageNumber="1265">0</td>
<td id="20207A1C586EEEE6FAD59ED8FACCFC11" box="[1321,1382,905,927]" gridcol="11" gridrow="18" pageId="16" pageNumber="1265">0</td>
<td id="20207A1C586EEEE6FA899ED8FA24FC11" box="[1397,1422,905,927]" gridcol="12" gridrow="18" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9EF6FA24FC33" box="[145,1422,935,957]" gridrow="19" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9EF6FE67FC33" box="[145,461,935,957]" gridcol="0" gridrow="19" pageId="16" pageNumber="1265">Width of the third tergal bar</th>
<td id="20207A1C586EEEE6FDE99EF6FDCBFC33" box="[533,609,935,957]" gridcol="1" gridrow="19" pageId="16" pageNumber="1265">6.47</td>
<td id="20207A1C586EEEE6FD899EF6FD19FC33" box="[629,691,935,957]" gridcol="2" gridrow="19" pageId="16" pageNumber="1265">1.35</td>
<td id="20207A1C586EEEE6FD3E9EF6FD71FC33" box="[706,731,935,957]" gridcol="3" gridrow="19" pageId="16" pageNumber="1265">32</td>
<td id="20207A1C586EEEE6FD079EF6FCFEFC33" box="[763,852,935,957]" gridcol="4" gridrow="19" pageId="16" pageNumber="1265">5.21</td>
<td id="20207A1C586EEEE6FCA09EF6FC33FC33" box="[860,921,935,957]" gridcol="5" gridrow="19" pageId="16" pageNumber="1265">1.37</td>
<td id="20207A1C586EEEE6FC549EF6FC6BFC33" box="[936,961,935,957]" gridcol="6" gridrow="19" pageId="16" pageNumber="1265">24</td>
<td id="20207A1C586EEEE6FC1D9EF6FB87FC33" box="[993,1069,935,957]" gridcol="7" gridrow="19" pageId="16" pageNumber="1265">5.4</td>
<td id="20207A1C586EEEE6FBBE9EF6FBD5FC33" box="[1090,1151,935,957]" gridcol="8" gridrow="19" pageId="16" pageNumber="1265">1.51</td>
<td id="20207A1C586EEEE6FB729EF6FB02FC33" box="[1166,1192,935,957]" gridcol="9" gridrow="19" pageId="16" pageNumber="1265">35</td>
<td id="20207A1C586EEEE6FB349EF6FAA3FC33" box="[1224,1289,935,957]" gridcol="10" gridrow="19" pageId="16" pageNumber="1265">0</td>
<td id="20207A1C586EEEE6FAD59EF6FACCFC33" box="[1321,1382,935,957]" gridcol="11" gridrow="19" pageId="16" pageNumber="1265">0</td>
<td id="20207A1C586EEEE6FA899EF6FA24FC33" box="[1397,1422,935,957]" gridcol="12" gridrow="19" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9E94FA24FC55" box="[145,1422,965,987]" gridrow="20" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9E94FE67FC55" box="[145,461,965,987]" gridcol="0" gridrow="20" pageId="16" pageNumber="1265">Longitude of metafemur</th>
<td id="20207A1C586EEEE6FDE99E94FDCBFC55" box="[533,609,965,987]" gridcol="1" gridrow="20" pageId="16" pageNumber="1265">4.52</td>
<td id="20207A1C586EEEE6FD899E94FD19FC55" box="[629,691,965,987]" gridcol="2" gridrow="20" pageId="16" pageNumber="1265">0.75</td>
<td id="20207A1C586EEEE6FD3E9E94FD71FC55" box="[706,731,965,987]" gridcol="3" gridrow="20" pageId="16" pageNumber="1265">24</td>
<td id="20207A1C586EEEE6FD079E94FCFEFC55" box="[763,852,965,987]" gridcol="4" gridrow="20" pageId="16" pageNumber="1265">4.36</td>
<td id="20207A1C586EEEE6FCA09E94FC33FC55" box="[860,921,965,987]" gridcol="5" gridrow="20" pageId="16" pageNumber="1265">0.75</td>
<td id="20207A1C586EEEE6FC549E94FC6BFC55" box="[936,961,965,987]" gridcol="6" gridrow="20" pageId="16" pageNumber="1265">18</td>
<td id="20207A1C586EEEE6FC1D9E94FB87FC55" box="[993,1069,965,987]" gridcol="7" gridrow="20" pageId="16" pageNumber="1265">4.1</td>
<td id="20207A1C586EEEE6FBBE9E94FBD5FC55" box="[1090,1151,965,987]" gridcol="8" gridrow="20" pageId="16" pageNumber="1265">0.86</td>
<td id="20207A1C586EEEE6FB729E94FB02FC55" box="[1166,1192,965,987]" gridcol="9" gridrow="20" pageId="16" pageNumber="1265">33</td>
<td id="20207A1C586EEEE6FB349E94FAA3FC55" box="[1224,1289,965,987]" gridcol="10" gridrow="20" pageId="16" pageNumber="1265">4.36</td>
<td id="20207A1C586EEEE6FAD59E94FACCFC55" box="[1321,1382,965,987]" gridcol="11" gridrow="20" pageId="16" pageNumber="1265">0.71</td>
<td id="20207A1C586EEEE6FA899E94FA24FC55" box="[1397,1422,965,987]" gridcol="12" gridrow="20" pageId="16" pageNumber="1265">11</td>
</tr>
<tr id="63F11360586EEEE6FF6D9EB2FA24FC77" box="[145,1422,995,1017]" gridrow="21" pageId="16" pageNumber="1265">
<th id="20207A1C586EEEE6FF6D9EB2FE67FC77" box="[145,461,995,1017]" gridcol="0" gridrow="21" pageId="16" pageNumber="1265">Longitude of elytra</th>
<td id="20207A1C586EEEE6FDE99EB2FDCBFC77" box="[533,609,995,1017]" gridcol="1" gridrow="21" pageId="16" pageNumber="1265">8</td>
<td id="20207A1C586EEEE6FD899EB2FD19FC77" box="[629,691,995,1017]" gridcol="2" gridrow="21" pageId="16" pageNumber="1265">1.71</td>
<td id="20207A1C586EEEE6FD3E9EB2FD71FC77" box="[706,731,995,1017]" gridcol="3" gridrow="21" pageId="16" pageNumber="1265">35</td>
<td id="20207A1C586EEEE6FD079EB2FCFEFC77" box="[763,852,995,1017]" gridcol="4" gridrow="21" pageId="16" pageNumber="1265">7.88</td>
<td id="20207A1C586EEEE6FCA09EB2FC33FC77" box="[860,921,995,1017]" gridcol="5" gridrow="21" pageId="16" pageNumber="1265">1.48</td>
<td id="20207A1C586EEEE6FC549EB2FC6BFC77" box="[936,961,995,1017]" gridcol="6" gridrow="21" pageId="16" pageNumber="1265">29</td>
<td id="20207A1C586EEEE6FC1D9EB2FB87FC77" box="[993,1069,995,1017]" gridcol="7" gridrow="21" pageId="16" pageNumber="1265">7.79</td>
<td id="20207A1C586EEEE6FBBE9EB2FBD5FC77" box="[1090,1151,995,1017]" gridcol="8" gridrow="21" pageId="16" pageNumber="1265">1.67</td>
<td id="20207A1C586EEEE6FB729EB2FB02FC77" box="[1166,1192,995,1017]" gridcol="9" gridrow="21" pageId="16" pageNumber="1265">44</td>
<td id="20207A1C586EEEE6FB349EB2FAA3FC77" box="[1224,1289,995,1017]" gridcol="10" gridrow="21" pageId="16" pageNumber="1265">7.78</td>
<td id="20207A1C586EEEE6FAD59EB2FACCFC77" box="[1321,1382,995,1017]" gridcol="11" gridrow="21" pageId="16" pageNumber="1265">1.08</td>
<td id="20207A1C586EEEE6FA899EB2FA24FC77" box="[1397,1422,995,1017]" gridcol="12" gridrow="21" pageId="16" pageNumber="1265">11</td>
</tr>
</table>
</paragraph>
<paragraph id="DD7E1122586E1109FF6D98B9FAB9FB64" blockId="16.[145,762,1512,1902]" lastBlockId="16.[809,1426,1084,1412]" pageId="16" pageNumber="1265">
<emphasis id="EFB5CD30586E1109FF6D98B9FF4EF992" italics="true" pageId="16" pageNumber="1265">
Distribution and notes on natural history:
<taxonomicName id="1AC16AA1586E1109FD9998B9FF4EF992" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="majalis">Berberomeloe majalis</taxonomicName>
</emphasis>
occurs in most of the central and eastern regions of the Iberian Peninsula, covering the Meseta Sur and most of the Sistema Ibérico Meridional Mountain Range, over the following Spanish provinces (
<figureCitation id="45FA0DA7586E1109FDB19B32FD24F9F6" box="[589,654,1635,1657]" captionStart="Figure 2" captionStartId="8.[146,224,1637,1659]" captionTargetBox="[305,1265,554,1598]" captionTargetId="figure-112@8.[305,1265,554,1598]" captionTargetPageId="8" captionText="Figure 2. Map of the Iberian Peninsula and north-western Africa showing the sampling localities of sequenced specimens of Berberomeloe." pageId="16" pageNumber="1265">Fig. 2</figureCitation>
):
<collectingRegion id="1F05DFC0586E1109FD589B32FD53F9F7" box="[676,761,1635,1657]" country="Spain" name="Madrid" pageId="16" pageNumber="1265">Madrid</collectingRegion>
(limited to the south-eastern border), central and eastern Toledo and Ciudad Real, Albacete, Cuenca, Guadalajara (mostly in the south-east), Teruel, Castellón,
<collectingRegion id="1F05DFC0586E1109FD6B9BEFFD52F95A" box="[663,760,1726,1748]" country="Spain" name="Comunitat Valenciana" pageId="16" pageNumber="1265">Valencia</collectingRegion>
and Jaén (north of the province). Distributional limits are yet to be defined in detail. It occurs across a wide spectrum of substrates, from limestone and gypsum soils to granitic formations (see:
<bibRefCitation id="B9506CD3586E1109FE139A6BFDC8F8C0" author="Vera JA" box="[495,610,1849,1871]" pageId="16" pageNumber="1265" refId="ref44945" refString="Vera JA. 2004. Geologia de Espana. Madrid: Sociedad Geologica de Espana-Instituto Geologico y Minero de Espana, 890." type="journal volume" year="2004">Vera, 2004</bibRefCitation>
). It is present in different habitat
<typeStatus id="027AAF80586E1109FE939A09FE06F8E3" box="[367,428,1880,1901]" pageId="16" pageNumber="1265">types</typeStatus>
, mostly open areas, including cereal fields, steppes, Mediterranean shrublands and Mediterranean open forests (
<figureCitation id="45FA0DA7586E1109FB84990BFB13FBFE" box="[1144,1209,1114,1136]" captionStart="Figure 7" captionStartId="17.[164,243,1666,1688]" captionTargetBox="[266,1340,198,1626]" captionTargetId="figure-69@17.[265,1340,195,1626]" captionTargetPageId="17" captionText="Figure 7. AC, habitat of Berberomeloe majalis, Spain. A, Pozo Airón, La Almarcha, Cuenca; B, Pajares, Ciudad Real; C, Bascuñana, Cuenca. D, newly emerged specimen from Tembleque, Toledo; note the unexpanded abdomen. E, specimen from Penyagolosa, Castelló; unlike most populations, specimens from this region present characteristic wide transverse abdominal bars. F, Berberomeloe majalis courtship, Alcazar de San Juan, Ciudad Real." pageId="16" pageNumber="1265">Fig. 7</figureCitation>
). Most records are located within the meso-Mediterranean bioclimatic level and, to a lesser extent, at the supra-Mediterranean, in regions with ombrotype from dry to humid (see:
<bibRefCitation id="B9506CD3586E1109FAB599E7FC65FB64" author="Rivas-Martinez S" pageId="16" pageNumber="1265" refId="ref43617" refString="Rivas-Martinez S. 1987. Memoria del mapa de las series de vegetacion de Espana 1: 400.000. Madrid: ICONA." type="book" year="1987">Rivas-Martínez, 1987</bibRefCitation>
;
<bibRefCitation id="B9506CD3586E1109FC209984FAAFFB64" author="Rivas-Martinez S &amp; Diaz TE &amp; Fernandez-Gonzalez F &amp; Izco J &amp; Loidi J &amp; Lousa M &amp; Penas A" box="[988,1285,1237,1259]" pageId="16" pageNumber="1265" pagination="5 - 432" refId="ref43700" refString="Rivas-Martinez S, Diaz TE, Fernandez-Gonzalez F, Izco J, Loidi J, Lousa M, Penas A. 2002. Vascular plant communities of Spain and Portugal. Addenda to the syntaxonomical checklist of 2001. Part I. Itinera Geobotanica 15: 5-432." type="journal article" year="2002">
Rivas-Martínez
<emphasis id="EFB5CD30586E1109FB729987FB6EFB64" box="[1166,1220,1237,1259]" italics="true" pageId="16" pageNumber="1265">et al.</emphasis>
, 2002
</bibRefCitation>
).
</paragraph>
<paragraph id="DD7E1122586E1109FCBD99A5FAF4FA0A" blockId="16.[809,1426,1084,1412]" pageId="16" pageNumber="1265">
Biological aspects of
<taxonomicName id="1AC16AA1586E1109FBD499A5FB32FA87" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1064,1176,1268,1289]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="majalis">
<emphasis id="EFB5CD30586E1109FBD499A5FB32FA87" box="[1064,1176,1268,1289]" italics="true" pageId="16" pageNumber="1265">B. majalis</emphasis>
</taxonomicName>
are poorly known, but they are expected to be similar to the ones described generically for
<emphasis id="EFB5CD30586E1109FC2C9863FBCDFAC8" box="[976,1127,1329,1351]" italics="true" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FC2C9863FBE8FAC8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[976,1090,1329,1351]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="majalis">B. majalis</taxonomicName>
<taxonomicNameLabel id="F486704B586E1109FBBB9860FBCDFAC8" box="[1095,1127,1329,1350]" pageId="16" pageNumber="1265" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
(previous to this study) by
<bibRefCitation id="B9506CD3586E1109FCD59801FBABFAEB" author="Beauregard H" box="[809,1025,1360,1382]" pageId="16" pageNumber="1265" refId="ref39114" refString="Beauregard H. 1890. Les insectes vesicants. Paris: Felix Alcan." type="book" year="1890">Beauregard (1890)</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FBF29801FB19FAEB" author="Bologna MA" box="[1038,1203,1360,1382]" pageId="16" pageNumber="1265" pagination="359 - 366" refId="ref39320" refString="Bologna MA. 1989. Berberomeloe, a new west Mediterranean genus of Lyttini for Meloe majalis Linee (Coleoptera, Meloidae). Systematics and bionomics. Bolletino di Zoologia 55: 359-366." type="journal article" year="1989">Bologna (1989</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FB3D9801FB52FAEB" author="Bologna MA" box="[1217,1272,1360,1381]" pageId="16" pageNumber="1265" refId="ref39357" refString="Bologna MA. 1991. Fauna d´Italia. XXVIII. Coleoptera Meloidae. Bologna: Edizioni Calderini." type="book" year="1991">1991</bibRefCitation>
),
<bibRefCitation id="B9506CD3586E1109FAF29801FA20FAEB" author="Cros A" box="[1294,1418,1360,1381]" pageId="16" pageNumber="1265" pagination="45 - 48" refId="ref40093" refString="Cros A. 1912. Moeurs et evolution du Meloe majalis L. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord 3: 45-48, 70 - 72, 93 - 96, 154 - 161, 181 - 190, 209 - 215." type="journal article" year="1912">Cros (1912</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FCD5983EFCCAFA0A" author="Cros A" box="[809,864,1391,1412]" pageId="16" pageNumber="1265" pagination="16 - 26" refId="ref40142" refString="Cros A. 1913. Moeurs et evolution du Meloe majalis L. (suite et fin). Moeurs et evolution du Meloe majalis L. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord 4: 16-26." type="journal article" year="1913">1913</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FC90983EFC09FA0A" author="Cros A" box="[876,931,1391,1412]" pageId="16" pageNumber="1265" pagination="133 - 155" refId="ref40185" refString="Cros A. 1921. Notes sur les larves primaires des Meloidae avec indication des larves nouvelles (2 e serie). Annales de la Societe Entomologique de France 90: 133-155." type="journal article" year="1921">1921</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FC4C983EFC4DFA0A" author="Cros A" box="[944,999,1391,1412]" pageId="16" pageNumber="1265" pagination="27 - 58" refId="ref40221" refString="Cros A. 1928. Essai sur la forme contractee (hypnotheque ou pseudonymphe) des larves des Meloidae. Annales de la Societe Entomologique de France 97: 27-58." type="journal article" year="1928">1928</bibRefCitation>
) and
<bibRefCitation id="B9506CD3586E1109FBD4983FFAA1FA0A" author="Gorriz Munoz RJ" box="[1064,1291,1390,1412]" pageId="16" pageNumber="1265" pagination="71 - 73" refId="ref41526" refString="Gorriz Munoz RJ. 1878. Sin titulo [Observaciones sobre Meloe majalis]. Actas de la Sociedad Espanola de Historia Natural 7: 71-73." type="journal article" year="1878">Górriz Muñoz (1878</bibRefCitation>
,
<bibRefCitation id="B9506CD3586E1109FAE4983EFAE5FA0A" author="Gorriz Munoz RJ" box="[1304,1359,1391,1412]" pageId="16" pageNumber="1265" refId="ref41555" refString="Gorriz Munoz RJ. 1882. Ensayo para la monografia de los coleopteros meloidos indigenas con aplicacion a las ciencias medicas. Zaragoza: Imprenta y libreria de J. Sanz." type="book" year="1882">1882</bibRefCitation>
).
</paragraph>
<paragraph id="DD7E1122586E1109FCC29890FAD6FA54" blockId="16.[830,1404,1473,1528]" box="[830,1404,1473,1498]" pageId="16" pageNumber="1265">
<taxonomicName id="1AC16AA1586E1109FCC29890FADCFA57" authority="SANCHEZ-VIALAS ET AL." authorityName="SANCHEZ-VIALAS ET AL." authorityYear="2020" box="[830,1398,1473,1498]" class="Insecta" family="Meloidae" genus="Berberomeloe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1265" phylum="Arthropoda" rank="species" species="castuo" status="SP. NOV.">
<emphasis id="EFB5CD30586E1109FCC29890FBCEFA59" bold="true" box="[830,1124,1473,1496]" italics="true" pageId="16" pageNumber="1265">
<smallCapsWord id="DB9887FE586E1109FCC29890FBAAFA59" baselines="1490,1490" box="[830,1024,1473,1496]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Berberomeloe" pageId="16" pageNumber="1265">BERBEROMELOE</smallCapsWord>
<smallCapsWord id="DB9887FE586E1109FBFB9895FBCEFA59" baselines="1490" box="[1031,1124,1476,1495]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="castuo" pageId="16" pageNumber="1265">CASTUO</smallCapsWord>
</emphasis>
<smallCapsWord id="DB9887FE586E1109FB979895FA82FA56" baselines="1491,1492,1491" box="[1131,1320,1473,1498]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sanchez-vialas" pageId="16" pageNumber="1265">SÁNCHEZ-VIALAS</smallCapsWord>
<emphasis id="EFB5CD30586E1109FAD39894FADCFA57" box="[1327,1398,1474,1497]" italics="true" pageId="16" pageNumber="1265">
<smallCapsWord id="DB9887FE586E1109FAD39894FAE6FA59" baselines="1490" box="[1327,1356,1477,1495]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="et" pageId="16" pageNumber="1265">ET</smallCapsWord>
<smallCapsWord id="DB9887FE586E1109FAAF9895FAC5FA56" baselines="1491" box="[1363,1391,1476,1496]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="al" pageId="16" pageNumber="1265">AL</smallCapsWord>
.
</emphasis>
</taxonomicName>
,
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,296 @@
<document id="3715A73BB4DC73940452DF7F95ADC0E1" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB61090277187F952C35BFC57" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Tanytarsus ramus Gilka &amp; Zakrzewska 2020, SP. NOV." docType="treatment" docVersion="1" lastPageNumber="1407" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1403" updateTime="1739357823542" updateUser="GgImagineBatch">
<mods:mods id="0818073AC80770C48E9B2DB73F5A0E11" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CF4B7A9B7E75324B769FB681665F83D">
<mods:title id="AF217BBE6BEE4D2BB6624F99EA5B9554">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="D9F24CD88805DF605B8FEFCE98CE949F" type="personal">
<mods:role id="F1381CECE5E9CB78CF10D11349D1288A">
<mods:roleTerm id="E1B388B8BFE1FB8382BAFF2DA0103580">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB75AC61D5E90EA60A918FC22E325330">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="6A170958A37C5ABE4F37D880CFB5DE52" type="personal">
<mods:role id="C01251464A42FB0F9C37C1AE3024E756">
<mods:roleTerm id="E6C22AFA9B2F7CEC0DB9D47FF0632931">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9251BDB15C3645B6862ABE88A76963A">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="5C76BEAD5A4C62C0D687B0F01C8A1676" type="personal">
<mods:role id="107C4EA0D6FB3360073857AD46B99635">
<mods:roleTerm id="45B5A59D7322AE89F34C09C1C60AB31C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C90C81E9E04D9E9884BF4605FA207CB1">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="DDE875C11A0D427FF79B3A06D7E7ACFF" type="personal">
<mods:role id="7905F92F65FBBA22C12E7DA74F73FB56">
<mods:roleTerm id="C289AD6C7E810A955C06DA74769C0EF3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF2FD74FD089D381F01E89E9541C178F">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="1476471AFFD591FF552F8E75B64086DE">text</mods:typeOfResource>
<mods:relatedItem id="FAD5E4E6850BAEDF08BF5096BFD5B81E" type="host">
<mods:titleInfo id="0842FF36C1E586C77CE0B3ABB34E0745">
<mods:title id="A0AB5E09F064578757CF5D0E8DC83C23">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="D7F0CF72C64C0C2CD1FD4DB95112BEF8">
<mods:date id="3C4B66D8AA74F4E72DECA17BBDF31BBB">2020</mods:date>
<mods:detail id="5A3D8CF8FD9ACC1AD34FF31D82C47A1E" type="volume">
<mods:number id="2B777D1B539D2FE1BA0EDE9E4FAC9C79">189</mods:number>
</mods:detail>
<mods:extent id="C35B50DF47106DC67628D143D145D8CC" unit="page">
<mods:start id="E5FF8C1CCFC2F2533EDEC688CE1B6E75">1398</mods:start>
<mods:end id="A33EE34C2A48AB0B88E0A40D03032A10">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5051C12F3A563648B95C18D54554D8F0">journal article</mods:classification>
<mods:identifier id="2107F44A5BF105D599A8558C94240372" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="C99072329B836F59A4E2A837FF1DEF9E" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB61090277187F952C35BFC57" LSID="urn:lsid:plazi:treatment:7C3D87FBB61090277187F952C35BFC57" httpUri="http://treatment.plazi.org/id/7C3D87FBB61090277187F952C35BFC57" lastPageId="9" lastPageNumber="1407" pageId="5" pageNumber="1403">
<subSubSection id="BC8E6566B610902B7187F952C3E2FFE5" box="[146,759,1739,1764]" pageId="5" pageNumber="1403" type="nomenclature">
<paragraph id="F42B36EDB610902B7187F952C3E2FFE5" blockId="5.[146,759,1739,1764]" box="[146,759,1739,1764]" pageId="5" pageNumber="1403">
<heading id="AF638181B610902B7187F952C3E2FFE5" box="[146,759,1739,1764]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="1403" reason="2">
<taxonomicName id="33944D6EB610902B7187F952C398FFE4" ID-CoL="9HM8P" authority="Gilka &amp; Zakrzewska" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[146,653,1739,1764]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1416" phylum="Arthropoda" rank="species" species="ramus" status="sp. nov.">
<emphasis id="C6E0EAFFB610902B7187F952C090FFE7" bold="true" box="[146,389,1739,1762]" italics="true" pageId="5" pageNumber="1403">
<smallCapsWord id="F2CDA031B610902B7187F952C038FFE7" baselines="1756,1756" box="[146,301,1739,1762]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsus" pageId="5" pageNumber="1403">TANYTARSUS</smallCapsWord>
<smallCapsWord id="F2CDA031B610902B7021F957C090FFE7" baselines="1756" box="[308,389,1742,1761]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="ramus" pageId="5" pageNumber="1403">RAMUS</smallCapsWord>
</emphasis>
<smallCapsWord id="F2CDA031B610902B7099F952C0C1FFE4" baselines="1758,1757" box="[396,468,1739,1764]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gilka" pageId="5" pageNumber="1403">GIŁKA</smallCapsWord>
&amp;
<smallCapsWord id="F2CDA031B610902B70E3F952C398FFE4" baselines="1758,1757" box="[502,653,1739,1764]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zakrzewska" pageId="5" pageNumber="1403">ZAKRZEWSKA</smallCapsWord>
</taxonomicName>
<emphasis id="C6E0EAFFB610902B7381F957C3E2FFE5" bold="true" box="[660,759,1739,1763]" pageId="5" pageNumber="1403">
<taxonomicNameLabel id="DDD35784B610902B7381F957C3E2FFE5" box="[660,759,1739,1763]" pageId="5" pageNumber="1403" rank="species">
<smallCapsWord id="F2CDA031B610902B7381F957C3A4FFE7" baselines="1756" box="[660,689,1742,1761]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="5" pageNumber="1403">SP</smallCapsWord>
.
<smallCapsWord id="F2CDA031B610902B73AAF957C3E5FFE7" baselines="1756" box="[703,752,1742,1761]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="5" pageNumber="1403">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B61090277028F895C35BFC57" lastPageId="9" lastPageNumber="1407" pageId="5" pageNumber="1403" type="description">
<paragraph id="F42B36EDB610902B7028F895C359FE23" blockId="5.[317,588,1804,1829]" box="[317,588,1804,1829]" pageId="5" pageNumber="1403">
(
<figureCitation id="6CAF2A68B610902B7050F895C0ABFE23" box="[325,446,1804,1829]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="5" pageNumber="1403">
<smallCapsWord id="F2CDA031B610902B7050F895C06EFE22" baselines="1823,1823" box="[325,379,1804,1829]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="5" pageNumber="1403">FIGS</smallCapsWord>
4AF
</figureCitation>
,
<figureCitation id="6CAF2A68B610902B70DEF895C312FE23" box="[459,519,1804,1829]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="5" pageNumber="1403">5AF</figureCitation>
,
<figureCitation id="6CAF2A68B610902B7301F895C350FE23" box="[532,581,1804,1829]" captionStart="Figure 11" captionStartId="16.[163,242,1861,1883]" captionTargetBox="[227,1379,195,1821]" captionTargetId="figure-26@16.[227,1379,195,1821]" captionTargetPageId="16" captionText="Figure 11. Fourier transform infrared spectroscopy (attenuated total reflectance) spectra obtained from Cambay amber chunks with the examined inclusions (early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India)." pageId="5" pageNumber="1403">11D</figureCitation>
)
</paragraph>
<paragraph id="F42B36EDB610902B7184F8ACC0D9FE6F" blockId="5.[145,760,1845,1897]" pageId="5" pageNumber="1403">
<emphasis id="C6E0EAFFB610902B7184F8ACC1C1FE4C" box="[145,212,1845,1866]" italics="true" pageId="5" pageNumber="1403">LSID</emphasis>
:
<uri id="80053AEFB610902B71CFF8ACC0D9FE6F" pageId="5" pageNumber="1403">
urn:lsid:zoobank.org:act:
<uuid id="80320C38B610902B7306F8ACC0D9FE6F" pageId="5" pageNumber="1403">D8802738-369F- 4D19-823F-379C31FFA80B</uuid>
</uri>
</paragraph>
<paragraph id="F42B36EDB610902B723CFAE3C2C6FC94" blockId="5.[809,979,1402,1426]" box="[809,979,1402,1426]" pageId="5" pageNumber="1403">
<emphasis id="C6E0EAFFB610902B723CFAE3C2C6FC94" box="[809,979,1402,1426]" italics="true" pageId="5" pageNumber="1403">Type material</emphasis>
</paragraph>
<paragraph id="F42B36EDB610902B723CFA3BC436FF57" blockId="5.[809,1425,1442,1617]" pageId="5" pageNumber="1403">
<materialsCitation id="44FC3CB0B610902B723CFA3BC436FF57" country="India" county="Tadkeshwar" location="Eocene" municipality="Mya" pageId="5" pageNumber="1403" specimenCount="1" stateProvince="Gujarat" typeStatus="holotype">
<typeStatus id="2B2F884FB610902B723CFA3BC282FCBE" box="[809,919,1442,1464]" pageId="5" pageNumber="1403" type="holotype">Holotype</typeStatus>
, inventory no.
<emphasis id="C6E0EAFFB610902B7544FA3AC5EEFCB1" box="[1105,1275,1442,1464]" italics="true" pageId="5" pageNumber="1403">BSIP Tad-521</emphasis>
: adult male (tarsi of left foreleg and of both mid- and hindlegs missing;
<figureCitation id="6CAF2A68B610902B729BFA46C2EBFCF3" box="[910,1022,1503,1525]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="5" pageNumber="1403">Fig. 4A, B</figureCitation>
) in a
<quantity id="336C9B08B610902B7555FA79C594FCF3" box="[1088,1153,1504,1525]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="1403" unit="mm" value="7.0">7 mm</quantity>
×
<quantity id="336C9B08B610902B7588FA46C5CBFCF3" box="[1181,1246,1503,1525]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="1403" unit="mm" value="5.0">5 mm</quantity>
×
<quantity id="336C9B08B610902B75EEFA46C445FCF3" box="[1275,1360,1503,1525]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="1403" unit="mm" value="2.5">2.5 mm</quantity>
piece of amber preserved in a cubicoid plastic mass (early
<location id="F14B6036B610902B723CF984C26AFF34" LSID="urn:lsid:plazi:treatment:7C3D87FBB61090277187F952C35BFC57:F14B6036B610902B723CF984C26AFF34" box="[809,895,1565,1586]" country="India" county="Tadkeshwar" municipality="Mya" name="Eocene" pageId="5" pageNumber="1403" stateProvince="Gujarat">Eocene</location>
, ~54
<collectingMunicipality id="144FAC97B610902B72D4F984C2E2FF34" box="[961,1015,1565,1586]" pageId="5" pageNumber="1403">Mya</collectingMunicipality>
,
<collectingCounty id="1D4A4E61B610902B7516F984C580FF35" box="[1027,1173,1565,1587]" pageId="5" pageNumber="1403">Tadkeshwar</collectingCounty>
mine,
<collectingRegion id="3650F80FB610902B75FCF984C49EFF34" box="[1257,1419,1565,1587]" country="India" name="Gujarat" pageId="5" pageNumber="1403">Gujarat state</collectingRegion>
,
<collectingCountry id="8C83767DB610902B723CF9A2C272FF57" box="[809,871,1595,1617]" name="India" pageId="5" pageNumber="1403">India</collectingCountry>
; IR spectrum no. 14257 IAA,
<figureCitation id="6CAF2A68B610902B75A1F9A2C406FF57" box="[1204,1299,1595,1617]" captionStart="Figure 11" captionStartId="16.[163,242,1861,1883]" captionTargetBox="[227,1379,195,1821]" captionTargetId="figure-26@16.[227,1379,195,1821]" captionTargetPageId="16" captionText="Figure 11. Fourier transform infrared spectroscopy (attenuated total reflectance) spectra obtained from Cambay amber chunks with the examined inclusions (early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India)." pageId="5" pageNumber="1403">Fig. 11D</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="F42B36EDB610902B723CF9E3C2BFFF94" blockId="5.[809,938,1658,1682]" box="[809,938,1658,1682]" pageId="5" pageNumber="1403">
<emphasis id="C6E0EAFFB610902B723CF9E3C2BFFF94" box="[809,938,1658,1682]" italics="true" pageId="5" pageNumber="1403">Etymology</emphasis>
</paragraph>
<paragraph id="F42B36EDB610902B723CF93AC487FFD1" blockId="5.[809,1426,1698,1751]" pageId="5" pageNumber="1403">
From Latin
<taxonomicName id="33944D6EB610902B72A5F93AC2ECFFBE" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[944,1017,1699,1720]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="1403" phylum="Arthropoda" rank="species" species="ramus">
<emphasis id="C6E0EAFFB610902B72A5F93AC2ECFFBE" box="[944,1017,1699,1720]" italics="true" pageId="5" pageNumber="1403">ramus</emphasis>
</taxonomicName>
, branch or twig, refering to the frail, simply branched median volsellae. Noun in apposition.
</paragraph>
<paragraph id="F42B36EDB610902B723CF889C2B7FE21" blockId="5.[809,930,1808,1831]" box="[809,930,1808,1831]" pageId="5" pageNumber="1403">
<emphasis id="C6E0EAFFB610902B723CF889C2B7FE21" box="[809,930,1808,1831]" italics="true" pageId="5" pageNumber="1403">Diagnosis</emphasis>
</paragraph>
<paragraph id="F42B36EDB610902B723CF8AEC453FE6A" blockId="5.[809,1425,1847,1900]" pageId="5" pageNumber="1403">Wing vein Sc extending well beyond RM; FCu placed well distal to RM; distance between ends of R</paragraph>
<paragraph id="F42B36EDB610902B7453F8FFC45BFE74" box="[1350,1358,1894,1906]" pageId="5" pageNumber="1403">1</paragraph>
<paragraph id="F42B36EDB610902B745BF8CEC47BFE6A" blockId="5.[809,1425,1847,1900]" box="[1358,1390,1879,1900]" pageId="5" pageNumber="1403">R</paragraph>
<paragraph id="F42B36EDB610902B747BF8FFC484FE74" box="[1390,1425,1894,1906]" pageId="5" pageNumber="1403">2 + 3</paragraph>
<caption id="A0EB6665B613902871B6FB6FC5DEFC2C" pageId="6" pageNumber="1404" startId="6.[163,242,1270,1292]" targetBox="[163,1412,195,1230]" targetPageId="6" targetType="figure">
<paragraph id="F42B36EDB613902871B6FB6FC5DEFC2C" blockId="6.[163,1443,1270,1322]" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB613902871B6FB6FC019FC0D" bold="true" box="[163,268,1270,1292]" pageId="6" pageNumber="1404">Figure 3.</emphasis>
<taxonomicName id="33944D6EB61390287002FB6EC0D8FC0A" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[279,461,1270,1292]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="1404" phylum="Arthropoda" rank="species" species="forfex">
<emphasis id="C6E0EAFFB61390287002FB6EC0D8FC0A" box="[279,461,1270,1292]" italics="true" pageId="6" pageNumber="1404">Tanytarsus forfex</emphasis>
</taxonomicName>
, adult male,
<emphasis id="C6E0EAFFB61390287342FB6FC3EAFC0A" box="[599,767,1270,1292]" italics="true" pageId="6" pageNumber="1404">BSIP Tad-568 a</emphasis>
, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, B, hypopygium in dorsal aspect. C, median volsella. Volsellae: green, superior; red, median; blue, inferior.
</paragraph>
</caption>
<paragraph id="F42B36EDB613902871B6FACFC219FF44" blockId="6.[163,780,1366,1602]" pageId="6" pageNumber="1404">
and R
<subScript id="681034A8B613902871F0FAFCC011FC77" attach="left" box="[229,260,1381,1394]" fontSize="5" pageId="6" pageNumber="1404">2 + 3</subScript>
R
<subScript id="681034A8B61390287036FAFCC056FC77" attach="left" box="[291,323,1381,1394]" fontSize="5" pageId="6" pageNumber="1404">4 + 5</subScript>
unequal, VR
<subScript id="681034A8B613902870C7FAFCC0C9FC77" attach="left" box="[466,476,1381,1393]" fontSize="5" pageId="6" pageNumber="1404">C</subScript>
~5. Tibial combs and spurs stout, combs fan shaped, only one comb with spur on mid-leg, each comb with spur on hindleg. Anal tergite with three spinulae in a row, placed in pit surrounded by crest-like ridges. Anal point slender, long, reaching apex of inferior volsella. Superior volsella pipe shaped. Stem of median volsella twig shaped, branched into two simple, slender lamellae. Inferior volsella parallel sided.
</paragraph>
<paragraph id="F42B36EDB613902871B6F9E2C03AFF94" blockId="6.[163,303,1659,1682]" box="[163,303,1659,1682]" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB613902871B6F9E2C03AFF94" box="[163,303,1659,1682]" italics="true" pageId="6" pageNumber="1404">Description</emphasis>
</paragraph>
<paragraph id="F42B36EDB613902871B6F93AC01BFFD1" blockId="6.[163,779,1699,1751]" pageId="6" pageNumber="1404">
Adult male (
<emphasis id="C6E0EAFFB61390287025F93DC051FFBF" box="[304,324,1700,1721]" italics="true" pageId="6" pageNumber="1404">N</emphasis>
= 1). Total length ~
<quantity id="336C9B08B61390287337F93AC36CFFBF" box="[546,633,1699,1721]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="1404" unit="mm" value="2.1">2.1 mm</quantity>
; wing length ~840 μm.
</paragraph>
<paragraph id="F42B36EDB613902871B6F898C288FC8C" blockId="6.[163,779,1793,1907]" lastBlockId="6.[827,1443,1366,1418]" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB613902871B6F898C1F1FE10" box="[163,228,1793,1814]" italics="true" pageId="6" pageNumber="1404">Head</emphasis>
(
<figureCitation id="6CAF2A68B613902871EFF898C043FE10" box="[250,342,1793,1814]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB613902871EFF898C043FE10" box="[250,342,1793,1814]" italics="true" pageId="6" pageNumber="1404">Fig. 4B</emphasis>
</figureCitation>
): Eyes bare, slightly produced in dorsomedian position. Frontal tubercles not observed. Antenna with 13 flagellomeres, AR unmeasurable, plume fully developed. Length of palpomeres 25 (in micrometres): ~25, 105, 105 and 169. At least 12 strong clypeals.
</paragraph>
<paragraph id="F42B36EDB6139028722EFA29C509FCE3" blockId="6.[827,1442,1456,1509]" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB6139028722EFA29C286FCC3" box="[827,915,1456,1477]" italics="true" pageId="6" pageNumber="1404">Thorax:</emphasis>
Dc at least nine, Pa at least one, other thoracic setae unobservable.
</paragraph>
<paragraph id="F42B36EDB6139028722EF997C46DFE1F" blockId="6.[827,1443,1550,1817]" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB6139028722EF997C26DFF25" box="[827,888,1550,1571]" italics="true" pageId="6" pageNumber="1404">Wing</emphasis>
(
<figureCitation id="6CAF2A68B6139028729CF997C2CAFF25" box="[905,991,1550,1571]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB6139028729CF997C2CAFF25" box="[905,991,1550,1571]" italics="true" pageId="6" pageNumber="1404">Fig. 4C</emphasis>
</figureCitation>
): Anal lobe relatively well developed, broadest at two-thirds of length, width 265 μm, length/ width ratio 3.17. Sc extending well beyond RM. RM slightly oblique relative to M and R. R
<subScript id="681034A8B613902875CEF9E0C5F6FF83" attach="left" box="[1243,1251,1657,1669]" fontSize="5" pageId="6" pageNumber="1404">1</subScript>
and R
<subScript id="681034A8B6139028743FF9E0C45DFF83" attach="left" box="[1322,1352,1657,1670]" fontSize="5" pageId="6" pageNumber="1404">2 + 3</subScript>
arcuate in distal section. FCu placed well distal to RM; VR
<subScript id="681034A8B6139028749AF901C4B7FFA2" attach="left" box="[1423,1442,1688,1700]" fontSize="5" pageId="6" pageNumber="1404">Cu</subScript>
1.35. Veins ending as follows (in order from base to tip): An, Sc, Cu
<subScript id="681034A8B613902872B9F94CC2A1FFE7" attach="left" box="[940,948,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">1</subScript>
, R
<subScript id="681034A8B613902872C7F94CC2CFFFE7" attach="left" box="[978,986,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">1</subScript>
, R +
<subScript id="681034A8B613902872E2F94CC503FFE7" attach="left" box="[1015,1046,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">2 3</subScript>
, M +
<subScript id="681034A8B6139028752CF94CC54DFFE7" attach="left" box="[1081,1112,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">3 4</subScript>
, R +
<subScript id="681034A8B61390287560F94CC580FFE7" attach="left" box="[1141,1173,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">4 5</subScript>
, M +
<subScript id="681034A8B613902875A2F94CC5C2FFE7" attach="left" box="[1207,1239,1749,1761]" fontSize="5" pageId="6" pageNumber="1404">1 2</subScript>
; distance between ends of R
<subScript id="681034A8B613902872B1F96DC2B9FE06" attach="left" box="[932,940,1780,1792]" fontSize="5" pageId="6" pageNumber="1404">1</subScript>
R
<subScript id="681034A8B613902872D9F96DC2F8FE06" attach="left" box="[972,1005,1780,1792]" fontSize="5" pageId="6" pageNumber="1404">2 + 3</subScript>
and R
<subScript id="681034A8B6139028752DF96DC54CFE06" attach="left" box="[1080,1113,1780,1792]" fontSize="5" pageId="6" pageNumber="1404">2 + 3</subScript>
R
<subScript id="681034A8B6139028756DF96DC58CFE06" attach="left" box="[1144,1177,1780,1792]" fontSize="5" pageId="6" pageNumber="1404">4 + 5</subScript>
unequal; VR
<subScript id="681034A8B61390287438F96DC422FE06" attach="left" box="[1325,1335,1780,1792]" fontSize="5" pageId="6" pageNumber="1404">C</subScript>
~5. Wing membrane covered with macrotrichia in distal half.
</paragraph>
<paragraph id="F42B36EDB6139027722EF8A4C3C6F873" blockId="6.[827,1443,1852,1905]" lastBlockId="9.[145,761,198,374]" lastPageId="9" lastPageNumber="1407" pageId="6" pageNumber="1404">
<emphasis id="C6E0EAFFB6139028722EF8A4C262FE54" box="[827,887,1853,1874]" italics="true" pageId="6" pageNumber="1404">Legs:</emphasis>
Tibia of foreleg with stout spur ~25 μm long. Tibial combs of mid leg fan shaped, teeth ≤ 25 μm long, only one comb bearing slender spur ~55 μm long (
<figureCitation id="6CAF2A68B61C9027718CFF7CC01BF9FC" box="[153,270,229,251]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="9" pageNumber="1407">Fig. 4D, E</figureCitation>
); combs of hindleg broadly separated, fan shaped; one comb broad, with teeth ≤ 25 μm long, bearing spur ~50 μm long; second comb slender, teeth ≤ 30 μm long, bearing spur ~60 μm long (
<figureCitation id="6CAF2A68B61C90277379FED8C3A9F850" box="[620,700,321,343]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="9" pageNumber="1407">Fig. 4F</figureCitation>
). For the leg segment lengths and leg ratios, see
<tableCitation id="B9160356B61C9027736CFEF9C3D9F873" box="[633,716,352,374]" captionStart="Table 3" captionStartId="9.[143,207,1645,1666]" captionText="Table 3. Leg segment lengths (in micrometres) and leg ratios of male Tanytarsus ramus" pageId="9" pageNumber="1407">Table 3</tableCitation>
.
</paragraph>
<caption id="A0EB6665B61290297184F958C203FE14" pageId="7" pageNumber="1405" startId="7.[145,225,1729,1751]" targetBox="[145,1425,195,1689]" targetPageId="7" targetType="figure">
<paragraph id="F42B36EDB61290297184F958C203FE14" blockId="7.[145,1425,1729,1810]" pageId="7" pageNumber="1405">
<emphasis id="C6E0EAFFB61290297184F958C1E9FFD0" bold="true" box="[145,252,1729,1751]" pageId="7" pageNumber="1405">Figure 4.</emphasis>
<taxonomicName id="33944D6EB6129029701DF95BC0D8FFD1" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[264,461,1730,1751]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="1405" phylum="Arthropoda" rank="species" species="ramus">
<emphasis id="C6E0EAFFB6129029701DF95BC0D8FFD1" box="[264,461,1730,1751]" italics="true" pageId="7" pageNumber="1405">Tanytarsus ramus</emphasis>
</taxonomicName>
, adult male,
<emphasis id="C6E0EAFFB61290297349F958C3E0FFD1" box="[604,757,1729,1751]" italics="true" pageId="7" pageNumber="1405">BSIP Tad-521</emphasis>
, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg.
</paragraph>
</caption>
<caption id="A0EB6665B61D902671B6F958C03AFE14" pageId="8" pageNumber="1406" startId="8.[163,242,1729,1751]" targetBox="[163,1443,195,1689]" targetPageId="8" targetType="figure">
<paragraph id="F42B36EDB61D902671B6F958C03AFE14" blockId="8.[163,1443,1729,1810]" pageId="8" pageNumber="1406">
<emphasis id="C6E0EAFFB61D902671B6F958C019FFD0" bold="true" box="[163,268,1729,1751]" pageId="8" pageNumber="1406">Figure 5.</emphasis>
<taxonomicName id="33944D6EB61D9026700DF95BC0CCFFD1" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[280,473,1730,1751]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="1406" phylum="Arthropoda" rank="species" species="ramus">
<emphasis id="C6E0EAFFB61D9026700DF95BC0CCFFD1" box="[280,473,1730,1751]" italics="true" pageId="8" pageNumber="1406">Tanytarsus ramus</emphasis>
</taxonomicName>
, adult male,
<emphasis id="C6E0EAFFB61D90267371F958C3ECFFD1" box="[612,761,1729,1751]" italics="true" pageId="8" pageNumber="1406">BSIP Tad-521</emphasis>
, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior.
</paragraph>
</caption>
<paragraph id="F42B36EDB61C90277184FE03C326FA9F" blockId="9.[145,762,409,921]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277184FE03C031F8A9" box="[145,292,410,431]" italics="true" pageId="9" pageNumber="1407">Hypopygium</emphasis>
(
<figureCitation id="6CAF2A68B61C9027702FFE03C0BEF8A8" box="[314,427,409,431]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C9027702FFE03C0BEF8A8" box="[314,427,409,431]" italics="true" pageId="9" pageNumber="1407">Fig. 5AF</emphasis>
</figureCitation>
): Gonostylus slightly longer than gonocoxite, ~85 μm long, tapering towards pointed apex. Anal tergite semicircular, bearing three spinulae arranged in a row, placed in rounded, longitudinal pit, its crest-like ridges tapering towards base of anal point; several setae on each side. Anal point long, reaching apex of inferior volsella, slender, parallel sided with blunt tip; severalstrongposterolateralsetaeoneachside(
<figureCitation id="6CAF2A68B61C9027739DFDF6C3FFFB83" box="[648,746,623,645]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig.5A,B</figureCitation>
). Superior volsella pipe shaped, with oval basal part, distinctly narrowed at mid-length, distal part elongate with apex blunt; several short setae at median margin of basal part (
<figureCitation id="6CAF2A68B61C90277039FD73C08EFA06" box="[300,411,746,768]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig. 5B, D</figureCitation>
). Digitus not observed. Stem of median volsella ~20 μm long, posteromedially directed, twig shaped, branched into two simple, slender lamellae (
<figureCitation id="6CAF2A68B61C9027718CFCDFC034FA5A" box="[153,289,838,860]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig. 5C, E, F</figureCitation>
). Inferior volsella reaching half the length of the gonostylus, nearly parallel sided, with distal half armed, bearing numerous stout setae.
</paragraph>
<paragraph id="F42B36EDB61C90277184FC5BC1EBFADC" blockId="9.[145,254,962,986]" box="[145,254,962,986]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277184FC5BC1EBFADC" box="[145,254,962,986]" italics="true" pageId="9" pageNumber="1407">Remarks</emphasis>
</paragraph>
<paragraph id="F42B36EDB61C90277184FC72C35BFC57" blockId="9.[145,762,1002,1362]" pageId="9" pageNumber="1407">
<taxonomicName id="33944D6EB61C90277184FC72C07CFD06" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[145,361,1003,1024]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="species" species="ramus">
<emphasis id="C6E0EAFFB61C90277184FC72C07CFD06" box="[145,361,1003,1024]" italics="true" pageId="9" pageNumber="1407">Tanytarsus ramus</emphasis>
</taxonomicName>
differs from all known species of the genus by a set of distinct characters as follows: the VR
<subScript id="681034A8B61C902771F4FBAEC1FEFD45" attach="left" box="[225,235,1079,1091]" fontSize="5" pageId="9" pageNumber="1407">C</subScript>
ratio is nearly five (
<figureCitation id="6CAF2A68B61C902770CFFBB1C338FD3B" box="[474,557,1064,1086]" captionStart="Figure 4" captionStartId="7.[145,225,1729,1751]" captionTargetBox="[145,1425,195,1689]" captionTargetId="figure-26@7.[145,1425,195,1689]" captionTargetPageId="7" captionText="Figure 4. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, wing. D, tibial armature of mid-leg. E, spurless comb of mid-leg magnified approximately three times relative to D. F, tibial combs and spurs of hindleg." pageId="9" pageNumber="1407">Fig. 4C</figureCitation>
), the anal tergite has a couple of spinulae placed in a row in a large pit, and the anal point is slender and extraordinarily long (
<figureCitation id="6CAF2A68B61C9027718CFB1DC019FD9F" box="[153,268,1155,1178]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig. 5A, B</figureCitation>
); the pipe shaped superior volsella should be considered a key feature (
<figureCitation id="6CAF2A68B61C902770F4FB3BC341FDBE" box="[481,596,1186,1208]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig. 5B, D</figureCitation>
), whereas the digitus, typical of
<taxonomicName id="33944D6EB61C90277043FB5BC0C0FDD1" authorityName="van der Wulp" authorityYear="1874" box="[342,469,1218,1239]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E0EAFFB61C90277043FB5BC0C0FDD1" box="[342,469,1218,1239]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
</taxonomicName>
, is not observed (probably absent). The most peculiar structure diagnostic of
<taxonomicName id="33944D6EB61C90277184FB66C1E0FC12" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[145,245,1279,1300]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="species" species="ramus">
<emphasis id="C6E0EAFFB61C90277184FB66C1E0FC12" box="[145,245,1279,1300]" italics="true" pageId="9" pageNumber="1407">T. ramus</emphasis>
</taxonomicName>
is the hypopygial median volsella (
<figureCitation id="6CAF2A68B61C90277396FB67C3E5FC12" box="[643,752,1278,1300]" captionStart="Figure 5" captionStartId="8.[163,242,1729,1751]" captionTargetBox="[163,1443,195,1689]" captionTargetId="figure-25@8.[163,1443,195,1689]" captionTargetPageId="8" captionText="Figure 5. Tanytarsus ramus, adult male, BSIP Tad-521, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. AC, hypopygium in dorsal (A, B) and ventral (C) aspect. D, superior volsella. E, F, median volsella. Volsellae: green, superior; blue, inferior." pageId="9" pageNumber="1407">Fig. 5E, F</figureCitation>
) of a shape perceived as a plesiomorphy (see Discussion: Evolution of the diagnostic structures).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,97 @@
<document id="BBB0ECB66838C013013965D97F5904FF" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB611902A7004FD0CC3FAFA34" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Tanytarsus van der Wulp 1874" docType="treatment" docVersion="2" lastPageNumber="1402" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1402" updateTime="1739358162056" updateUser="ExternalLinkService">
<mods:mods id="36CDF63D7E14C3DEF9BFBDE3847E9655" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4514114B8D3198274C743E892C91D261">
<mods:title id="7805D12C19421609FBCC32D02B54D1DE">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="3F877344A163A3086F4886A9474140CA" type="personal">
<mods:role id="D446F32CAC25DEDE6BDF851E688E4C6E">
<mods:roleTerm id="15A5BD5537BD977F6714A5122D2DCB7D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF26C06EB62EB4B5116A3CA29D68CDE8">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="B23DE7F0B62884CACF1CD2B761DC425E" type="personal">
<mods:role id="D8C8692BECD9C88BA248CBFF55C8463D">
<mods:roleTerm id="CA67090EE5AE211F545A6E23E5FC0D35">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D2A26C27B8F9D4302D213097A14822D">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="84622AEF4D5DF469F688E95A7DEB8E41" type="personal">
<mods:role id="779FE45DD9AA0D2730CABBF6052BEC06">
<mods:roleTerm id="1F89AA0E39006EF9C0832BD888C16CA5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F981D1256075DE2C788157A524096A73">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="26DCF15AD46DE2FA19C7264F882708B6" type="personal">
<mods:role id="5B331EC5C28F2A06A0365E575E16BA8E">
<mods:roleTerm id="EB832A8186B27DA824F3F3C48B31BA37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D8109ADC63879A4BF9E02933B93452C">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="21659C9F43D70EFEE72101882AABC2E0">text</mods:typeOfResource>
<mods:relatedItem id="023644F5327AB59A210083EDD792BFD6" type="host">
<mods:titleInfo id="B3A220808BC4F839DD7B15213CBC617D">
<mods:title id="97C9A75E15138703F35315A34C37C1F2">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="049521A24DB2DCB6E8916D6640B41CE6">
<mods:date id="F9401C0A19FD05E277EF76F2D99626CB">2020</mods:date>
<mods:detail id="AA81F90E253BACD7C45F3073FF49A1B1" type="volume">
<mods:number id="A30BB82BE7144B26AA3CD6E63736EC50">189</mods:number>
</mods:detail>
<mods:extent id="4AD55A69ACE5A17FC47F06D3F3337E24" unit="page">
<mods:start id="EFFC77B7CAFDF4E3301553C50AD13D6C">1398</mods:start>
<mods:end id="099DF01A33BA5F00EBC444E1B0BE00A3">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1CC8121EDF23180E2D66F2033BFD8BA1">journal article</mods:classification>
<mods:identifier id="6F2F16BAE8F20C60488C66AD521340BE" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="47D50A277470E75353BE08542657BB28" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB611902A7004FD0CC3FAFA34" ID-DOI="http://doi.org/10.5281/zenodo.14858048" ID-Zenodo-Dep="14858048" LSID="urn:lsid:plazi:treatment:7C3D87FBB611902A7004FD0CC3FAFA34" httpUri="http://treatment.plazi.org/id/7C3D87FBB611902A7004FD0CC3FAFA34" lastPageNumber="1402" pageId="4" pageNumber="1402">
<subSubSection id="BC8E6566B611902A7004FD0CC389FBAB" box="[273,668,660,685]" pageId="4" pageNumber="1402" type="nomenclature">
<paragraph id="F42B36EDB611902A7004FD0CC389FBAB" blockId="4.[273,668,660,685]" box="[273,668,660,685]" pageId="4" pageNumber="1402">
<heading id="AF638181B611902A7004FD0CC389FBAB" box="[273,668,660,685]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="1402" reason="2">
<taxonomicName id="33944D6EB611902A7004FD0CC389FBAB" ID-CoL="7SJQ" authority="VAN DER WULP, 1874" authorityName="van der Wulp" authorityYear="1874" box="[273,668,660,685]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="1416" phylum="Arthropoda" rank="genus">
<smallCapsWord id="F2CDA031B611902A7004FD0CC0B0FBAA" baselines="678,679" box="[273,421,661,684]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsus" pageId="4" pageNumber="1402">
<emphasis id="C6E0EAFFB611902A7004FD0CC0B0FBAA" box="[273,421,661,684]" italics="true" pageId="4" pageNumber="1402">TANYTARSUS</emphasis>
</smallCapsWord>
<smallCapsWord id="F2CDA031B611902A70B9FD01C0CDFBAA" baselines="679" box="[428,472,664,684]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="van" pageId="4" pageNumber="1402">VAN</smallCapsWord>
<smallCapsWord id="F2CDA031B611902A70F5FD01C318FBAA" baselines="679" box="[480,525,664,684]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="der" pageId="4" pageNumber="1402">DER</smallCapsWord>
<smallCapsWord id="F2CDA031B611902A7306FD0DC34DFBAA" baselines="679,679" box="[531,600,660,685]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Wulp" pageId="4" pageNumber="1402">WULP</smallCapsWord>
, 1874
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B611902A71B6FD44C3FAFA34" pageId="4" pageNumber="1402" type="materials_examined">
<paragraph id="F42B36EDB611902A71B6FD44C219FBF5" blockId="4.[163,780,733,755]" box="[163,780,733,755]" pageId="4" pageNumber="1402">
<emphasis id="C6E0EAFFB611902A71B6FD44C03AFBF4" box="[163,303,733,754]" italics="true" pageId="4" pageNumber="1402">
<typeStatus id="2B2F884FB611902A71B6FD44C1C2FBF4" box="[163,215,733,754]" pageId="4" pageNumber="1402">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="33944D6EB611902A702EFD44C21DFBF5" authority="(van der Wulp, 1858)" baseAuthorityName="van der Wulp" baseAuthorityYear="1858" box="[315,776,733,755]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="1402" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="C6E0EAFFB611902A702EFD44C309FBF4" box="[315,540,733,754]" italics="true" pageId="4" pageNumber="1402">Tanytarsus signatus</emphasis>
(van der Wulp, 1858)
</taxonomicName>
.
</paragraph>
<paragraph id="F42B36EDB611902A71AAFC83C3FAFA34" blockId="4.[191,751,793,848]" box="[191,751,793,818]" pageId="4" pageNumber="1402">
<taxonomicName id="33944D6EB611902A71AAFC83C3D1FA37" authority="Gilka &amp; Zakrzewska" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[191,708,793,818]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="1416" phylum="Arthropoda" rank="species" species="forfex" status="sp. nov.">
<emphasis id="C6E0EAFFB611902A71AAFC83C0A9FA36" bold="true" box="[191,444,794,817]" italics="true" pageId="4" pageNumber="1402">
<smallCapsWord id="F2CDA031B611902A71AAFC83C04CFA36" baselines="811,811" box="[191,345,794,817]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsus" pageId="4" pageNumber="1402">TANYTARSUS</smallCapsWord>
<smallCapsWord id="F2CDA031B611902A7075FC84C0A9FA36" baselines="811" box="[352,444,797,816]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="forfex" pageId="4" pageNumber="1402">FORFEX</smallCapsWord>
</emphasis>
<smallCapsWord id="F2CDA031B611902A70D6FC80C31FFA37" baselines="812,812" box="[451,522,793,818]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gilka" pageId="4" pageNumber="1402">GIŁKA</smallCapsWord>
&amp;
<smallCapsWord id="F2CDA031B611902A7338FC80C3D1FA37" baselines="812,812" box="[557,708,793,818]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zakrzewska" pageId="4" pageNumber="1402">ZAKRZEWSKA</smallCapsWord>
</taxonomicName>
<emphasis id="C6E0EAFFB611902A73DEFC84C3FAFA34" bold="true" box="[715,751,794,818]" pageId="4" pageNumber="1402">
<taxonomicNameLabel id="DDD35784B611902A73DEFC84C3FAFA34" box="[715,751,794,818]" pageId="4" pageNumber="1402">
<smallCapsWord id="F2CDA031B611902A73DEFC84C3FDFA36" baselines="811" box="[715,744,797,816]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="4" pageNumber="1402">SP</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,94 @@
<document id="F15BFB8FBC13B83CF45781EEE4362849" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB617902C71E4FC76C3A1FDA2" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Chironomidae NEWMAN 1834" docType="treatment" docVersion="1" lastPageNumber="1400" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1400" updateTime="1739357823542" updateUser="GgImagineBatch">
<mods:mods id="39CD124E44F7B62EB2F996691A33779A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="83361CA969F3FC05AD48241ECF0E1CAB">
<mods:title id="E9EFB902D9A2ADB29742A2E280D7F2E7">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="521B6C47D9952C643E75C211D6AF4B31" type="personal">
<mods:role id="9E02E304C29C74FBDE488DAC4A668AE3">
<mods:roleTerm id="F74BD34894CD62B981C4EC6FF149A238">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBB8220DC46F0B3389B13264B56E9698">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="29D3044471DA66959C7A90E8630793BB" type="personal">
<mods:role id="008FD53F8CF66E234170835D87391CA7">
<mods:roleTerm id="978821EA2866535DA85569B92D6C7353">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B8D7F5B7F51DE8D26261B9D75ABC894">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="D8ADBEB8BC0692454D00272ADCC96D3D" type="personal">
<mods:role id="9FF7F409CDC0EC4CAB4723DE60A86AF9">
<mods:roleTerm id="C7B9615CB65096E1EEE44722B2AFDDC7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECDD69018F8F8AF45781058B2BD65F56">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="E46A9FA26A8F1959682AFBDCB67DA316" type="personal">
<mods:role id="FBC923893E1E5B235853BCA324DF2580">
<mods:roleTerm id="8CF7DF21ED735592DA385CBB88E88B19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="657ED7EB382AB8F2D0CD3997C9FB3B51">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="26E386E9AF366A5359A7868BBA650D34">text</mods:typeOfResource>
<mods:relatedItem id="7E12422A2AA53FA2261C66A65B51F6D7" type="host">
<mods:titleInfo id="B309D3B338E6223517E1F5A996AD2625">
<mods:title id="218F37D8CB91642F16686043FC61BF9A">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="D92AB60F73B6545D8E7D337CB2315A3C">
<mods:date id="0A7567BABB8DF95BC74356CD78E5BF04">2020</mods:date>
<mods:detail id="2D019177136ABFE3E2F7D6F31B5AA50E" type="volume">
<mods:number id="D5D19AEDE4E31E96AE5F494E68FF14DD">189</mods:number>
</mods:detail>
<mods:extent id="8584249A232008714E95AF2FCCA0AFE8" unit="page">
<mods:start id="5544937FCDE80AB7F49B6E21D96F7733">1398</mods:start>
<mods:end id="3F3299211D9E4A3592A85C4167933AF0">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9BEF9E7763B59F32FA2D3136C7428E79">journal article</mods:classification>
<mods:identifier id="3F82594A83743746D2D5F02D30EBE53C" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="BACA7CAC9712817FB9D66155EB0C2937" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB617902C71E4FC76C3A1FDA2" LSID="urn:lsid:plazi:treatment:7C3D87FBB617902C71E4FC76C3A1FDA2" httpUri="http://treatment.plazi.org/id/7C3D87FBB617902C71E4FC76C3A1FDA2" lastPageNumber="1400" pageId="2" pageNumber="1400">
<subSubSection id="BC8E6566B617902C71E4FC76C3A9FD0E" box="[241,700,1007,1032]" pageId="2" pageNumber="1400" type="nomenclature">
<paragraph id="F42B36EDB617902C71E4FC76C3A9FD0E" blockId="2.[241,700,1007,1032]" box="[241,700,1007,1032]" pageId="2" pageNumber="1400">
<heading id="AF638181B617902C71E4FC76C3A9FD0E" box="[241,700,1007,1032]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="1400" reason="8">
<smallCapsWord id="F2CDA031B617902C71E4FC76C053FD01" baselines="1026,1026" box="[241,326,1007,1032]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Family" pageId="2" pageNumber="1400">FAMILY</smallCapsWord>
:
<taxonomicName id="33944D6EB617902C7041FC76C3A9FD0E" ID-CoL="84X" ID-ENA="7149" authority="NEWMAN, 1834" authorityName="NEWMAN" authorityYear="1834" box="[340,700,1007,1032]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1400" phylum="Arthropoda" rank="family">
<smallCapsWord id="F2CDA031B617902C7041FC76C313FD01" baselines="1026,1026" box="[340,518,1007,1032]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Chironomidae" pageId="2" pageNumber="1400">CHIRONOMIDAE</smallCapsWord>
<smallCapsWord id="F2CDA031B617902C7318FC76C362FD01" baselines="1026,1026" box="[525,631,1007,1032]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Newman" pageId="2" pageNumber="1400">NEWMAN</smallCapsWord>
, 1834
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B617902C71C9FB87C3A1FDA2" pageId="2" pageNumber="1400" type="description">
<paragraph id="F42B36EDB617902C71C9FB87C3C4FD31" blockId="2.[220,721,1054,1079]" box="[220,721,1054,1079]" pageId="2" pageNumber="1400">
<smallCapsWord id="F2CDA031B617902C71C9FB87C04EFD30" baselines="1073,1073" box="[220,347,1054,1079]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Subfamily" pageId="2" pageNumber="1400">SUBFAMILY</smallCapsWord>
:
<taxonomicName id="33944D6EB617902C707CFB87C3C4FD31" authority="NEWMAN, 1834" authorityName="NEWMAN" authorityYear="1834" box="[361,721,1054,1079]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1400" phylum="Arthropoda" rank="subFamily" subFamily="Chironominae">
<smallCapsWord id="F2CDA031B617902C707CFB87C309FD30" baselines="1073,1073" box="[361,540,1054,1079]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Chironominae" pageId="2" pageNumber="1400">CHIRONOMINAE</smallCapsWord>
<smallCapsWord id="F2CDA031B617902C7336FB87C398FD30" baselines="1073,1073" box="[547,653,1054,1079]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Newman" pageId="2" pageNumber="1400">NEWMAN</smallCapsWord>
, 1834
</taxonomicName>
</paragraph>
<paragraph id="F42B36EDB617902C7007FBCCC38EFD68" blockId="2.[274,667,1109,1134]" box="[274,667,1109,1134]" pageId="2" pageNumber="1400">
<smallCapsWord id="F2CDA031B617902C7007FBCCC042FD6A" baselines="1128,1127" box="[274,343,1109,1134]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tribe" pageId="2" pageNumber="1400">TRIBE</smallCapsWord>
:
<taxonomicName id="33944D6EB617902C7076FBCCC38EFD68" authority="ZAVREL, 1917" authorityName="ZAVREL" authorityYear="1917" box="[355,667,1109,1134]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1400" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">
<smallCapsWord id="F2CDA031B617902C7076FBCCC0EFFD6A" baselines="1128,1127" box="[355,506,1109,1134]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsini" pageId="2" pageNumber="1400">TANYTARSINI</smallCapsWord>
<smallCapsWord id="F2CDA031B617902C7314FBCCC343FD6A" baselines="1128,1127,1127,1127" box="[513,598,1109,1134]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zavrel" pageId="2" pageNumber="1400">ZAVŘEL</smallCapsWord>
, 1917
</taxonomicName>
</paragraph>
<paragraph id="F42B36EDB617902C71ECFB12C3A1FDA2" blockId="2.[249,692,1163,1188]" box="[249,692,1163,1188]" pageId="2" pageNumber="1400">
<smallCapsWord id="F2CDA031B617902C71ECFB12C07FFDA5" baselines="1182,1182" box="[249,362,1163,1188]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Subtribe" pageId="2" pageNumber="1400">SUBTRIBE</smallCapsWord>
:
<taxonomicName id="33944D6EB617902C7063FB12C306FDA5" box="[374,531,1163,1188]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1417" phylum="Arthropoda" rank="subTribe" subTribe="Tanytarsina">
<smallCapsWord id="F2CDA031B617902C7063FB12C306FDA5" baselines="1182,1182" box="[374,531,1163,1188]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsina" pageId="2" pageNumber="1400">TANYTARSINA</smallCapsWord>
</taxonomicName>
<smallCapsWord id="F2CDA031B617902C730FFB12C37AFDA5" baselines="1182,1182,1181,1182" box="[538,623,1163,1188]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zavrel" pageId="2" pageNumber="1400">ZAVŘEL</smallCapsWord>
, 1917
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
<document id="00A8CC746E1394BE423CA8197BFD06C0" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB617902C722BFF5DC5DFFB6B" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Gujaratomyia miripes Gilka &amp; Zakrzewska 2018" docType="treatment" docVersion="1" lastPageNumber="1400" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1400" updateTime="1739357823542" updateUser="GgImagineBatch">
<mods:mods id="32185D1F2857548EFE41AC4926F59558" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F10D9CBE39660A4CF4729DE3EEEB2F08">
<mods:title id="DC60178D08ED5C5256951A1F68D0741C">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="BE7997D9FEBD3816AAC0D8381E17F9CD" type="personal">
<mods:role id="3225F916AA20B2C2614E537871BC39A8">
<mods:roleTerm id="F59DECD4772821A45426969BD81C11B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BACEDFE29D8D21FDF17C919CFD5C4496">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="AAF15489A6BCE318A786982D6FCD25C5" type="personal">
<mods:role id="35EB342FBD1360E6FFC9B12CFD1068B5">
<mods:roleTerm id="072B36F8B3DC3B6D4A3CE797AB351538">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BCB514A8F2E1A212DCCD02505C355892">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="8394CDFC517D0CBAE25424D65CCFC14D" type="personal">
<mods:role id="2FB52DA3BC83ADE26C2024966F078729">
<mods:roleTerm id="B5FF744510409D2E5D33C15550D247CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5466A8898587BADE2FF2CA82A68AA8CE">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="959E4F93E963AA515F17A9708440C7C8" type="personal">
<mods:role id="51D1C5A3187008226E2AA7C96C69DEBD">
<mods:roleTerm id="B6D0ED1C2599F58F58F77AA7E4FCEF5F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3EF10E43442C1CD468F961943E467576">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="0DB110BA6B0B19EBAB312BB7085B7A69">text</mods:typeOfResource>
<mods:relatedItem id="7869F17002CB234945702E8FDF60DB6A" type="host">
<mods:titleInfo id="61CB87E5784ECD9B357B73A581E86D51">
<mods:title id="4CABFA7470CA7ADF0F78400A5369E6C2">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="B1D43DAAD2B9AEBCF963A006A4BA383B">
<mods:date id="4BA119F8C3730BE51F53819758A6C6F0">2020</mods:date>
<mods:detail id="C76174F4F39C23BE1C9286ECD165DCDC" type="volume">
<mods:number id="6EEB60F2031BABC0F8EA307D57821AC1">189</mods:number>
</mods:detail>
<mods:extent id="D3BDE1CD31B7AC9350495A3782D95F34" unit="page">
<mods:start id="C913B8D48BC22DC99E1A5D2E3FEF8218">1398</mods:start>
<mods:end id="6EEE1B69609258241D83890B77A23C8E">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7C7ACD9D3A7858E77408A2AC7F9DAFC8">journal article</mods:classification>
<mods:identifier id="1E950508AC9FD136E03C714EFEAA74E9" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="4F770BAD291D97320D9A3F9307EED17D" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB617902C722BFF5DC5DFFB6B" LSID="urn:lsid:plazi:treatment:7C3D87FBB617902C722BFF5DC5DFFB6B" httpUri="http://treatment.plazi.org/id/7C3D87FBB617902C722BFF5DC5DFFB6B" lastPageNumber="1400" pageId="2" pageNumber="1400">
<subSubSection id="BC8E6566B617902C722BFF5DC48AF9DB" box="[830,1439,196,221]" pageId="2" pageNumber="1400" type="nomenclature">
<paragraph id="F42B36EDB617902C722BFF5DC48AF9DB" blockId="2.[830,1439,196,221]" box="[830,1439,196,221]" pageId="2" pageNumber="1400">
<heading id="AF638181B617902C722BFF5DC48AF9DB" box="[830,1439,196,221]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="1400" reason="2">
<taxonomicName id="33944D6EB617902C722BFF5DC48AF9DB" ID-CoL="8C8Q2" authority="GILKA &amp; ZAKRZEWSKA, 2018" authorityName="Gilka &amp; Zakrzewska" authorityYear="2018" box="[830,1439,196,221]" class="Insecta" family="Chironomidae" genus="Gujaratomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1400" phylum="Arthropoda" rank="species" species="miripes">
<emphasis id="C6E0EAFFB617902C722BFF5DC547F9DD" box="[830,1106,196,219]" italics="true" pageId="2" pageNumber="1400">
<smallCapsWord id="F2CDA031B617902C722BFF5DC2FBF9DD" baselines="213,214" box="[830,1006,196,219]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gujaratomyia" pageId="2" pageNumber="1400">GUJARATOMYIA</smallCapsWord>
<smallCapsWord id="F2CDA031B617902C72E3FF5EC547F9DD" baselines="214" box="[1014,1106,199,219]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="miripes" pageId="2" pageNumber="1400">MIRIPES</smallCapsWord>
</emphasis>
<smallCapsWord id="F2CDA031B617902C754CFF5DC5B4F9DD" baselines="215,214" box="[1113,1185,196,221]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gilka" pageId="2" pageNumber="1400">GIŁKA</smallCapsWord>
&amp;
<bibRefCitation id="90054B1CB617902C75D1FF5DC48AF9DB" author="Zakrzewska M &amp; Stebner F &amp; Puchalski M &amp; Singh H &amp; Gilka W" box="[1220,1439,196,221]" pageId="2" pageNumber="1400" pagination="255 - 261" refId="ref16888" refString="Zakrzewska M, Stebner F, Puchalski M, Singh H, Gilka W. 2018. A peculiar leg structure in the first non-biting midge described from Cambay amber, India (Diptera: Chironomidae). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107: 255-261." type="journal article" year="2018">
<smallCapsWord id="F2CDA031B617902C75D1FF5DC44FF9DD" baselines="215,214" box="[1220,1370,196,221]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zakrzewska" pageId="2" pageNumber="1400">ZAKRZEWSKA</smallCapsWord>
, 2018
</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B617902C72E9FF6BC5DFFB6B" pageId="2" pageNumber="1400" type="description">
<paragraph id="F42B36EDB617902C72E9FF6BC5F4F80D" blockId="2.[1020,1249,242,267]" box="[1020,1249,242,267]" pageId="2" pageNumber="1400">
(
<figureCitation id="6CAF2A68B617902C7510FF6BC568F80D" box="[1029,1149,242,267]" captionStart="Figure 1" captionStartId="3.[145,224,1655,1677]" captionTargetBox="[145,1425,195,1616]" captionTargetId="figure-26@3.[145,1425,195,1616]" captionTargetPageId="3" captionText="Figure 1. Gujaratomyia miripes, adult male, BSIP Tad-610, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, apical part of hindleg tibia. D, head. E, scutellum with strong setae. F, hypopygium in dorsal aspect." pageId="2" pageNumber="1400">
<smallCapsWord id="F2CDA031B617902C7510FF6BC52EF80C" baselines="261,261" box="[1029,1083,242,267]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="2" pageNumber="1400">FIGS</smallCapsWord>
1AF
</figureCitation>
,
<figureCitation id="6CAF2A68B617902C759EFF6BC5CDF80D" box="[1163,1240,242,267]" captionStart="Figure 11" captionStartId="16.[163,242,1861,1883]" captionTargetBox="[227,1379,195,1821]" captionTargetId="figure-26@16.[227,1379,195,1821]" captionTargetPageId="16" captionText="Figure 11. Fourier transform infrared spectroscopy (attenuated total reflectance) spectra obtained from Cambay amber chunks with the examined inclusions (early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India)." pageId="2" pageNumber="1400">11A, B</figureCitation>
)
</paragraph>
<paragraph id="F42B36EDB617902C722EFEA3C537F854" blockId="2.[827,1058,314,338]" box="[827,1058,314,338]" pageId="2" pageNumber="1400">
<emphasis id="C6E0EAFFB617902C722EFEA3C537F854" box="[827,1058,314,338]" italics="true" pageId="2" pageNumber="1400">Material examined</emphasis>
</paragraph>
<paragraph id="F42B36EDB617902C722EFEFAC5DFFB6B" blockId="2.[827,1443,355,622]" pageId="2" pageNumber="1400">
<materialsCitation id="44FC3CB0B617902C722EFEFAC5D3FB6B" country="India" county="Tadkeshwar" location="Adult" municipality="Mya" pageId="2" pageNumber="1400" specimenCount="2" stateProvince="Gujarat" typeStatus="holotype">
<location id="F14B6036B617902C722EFEFAC26EF87F" LSID="urn:lsid:plazi:treatment:7C3D87FBB617902C722BFF5DC5DFFB6B:F14B6036B617902C722EFEFAC26EF87F" box="[827,891,355,377]" country="India" county="Tadkeshwar" municipality="Mya" name="Adult" pageId="2" pageNumber="1400" stateProvince="Gujarat">Adult</location>
male, inventory no. BSIP Tad-610 (left: antenna, wing, foreleg, tarsi of mid- and hindlegs and tarsus of right foreleg damaged;
<figureCitation id="6CAF2A68B617902C756EFE39C5E3F8B0" box="[1147,1270,416,438]" captionStart="Figure 1" captionStartId="3.[145,224,1655,1677]" captionTargetBox="[145,1425,195,1616]" captionTargetId="figure-26@3.[145,1425,195,1616]" captionTargetPageId="3" captionText="Figure 1. Gujaratomyia miripes, adult male, BSIP Tad-610, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India. A, inclusion in amber. B, habitus. C, apical part of hindleg tibia. D, head. E, scutellum with strong setae. F, hypopygium in dorsal aspect." pageId="2" pageNumber="1400">Fig. 1A, B</figureCitation>
) preserved in an
<quantity id="336C9B08B617902C7275FE26C2A2F8D2" box="[864,951,447,469]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="1400" unit="mm" value="8.5">8.5 mm</quantity>
×
<quantity id="336C9B08B617902C72C3FE26C53BF8D2" box="[982,1070,447,469]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="1400" unit="mm" value="4.5">4.5 mm</quantity>
×
<quantity id="336C9B08B617902C7558FE26C5B1F8D2" box="[1101,1188,447,469]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="1400" unit="mm" value="3.5">3.5 mm</quantity>
piece of amber (early
<location id="F14B6036B617902C722EFE47C284F8F5" LSID="urn:lsid:plazi:treatment:7C3D87FBB617902C722BFF5DC5DFFB6B:F14B6036B617902C722EFE47C284F8F5" box="[827,913,478,499]" country="India" county="Tadkeshwar" municipality="Mya" name="Eocene" pageId="2" pageNumber="1400" stateProvince="Gujarat">Eocene</location>
, ~54
<collectingMunicipality id="144FAC97B617902C72C6FE47C51CF8F5" box="[979,1033,478,499]" pageId="2" pageNumber="1400">Mya</collectingMunicipality>
,
<collectingCounty id="1D4A4E61B617902C7500FE47C5B2F8F2" box="[1045,1191,478,500]" pageId="2" pageNumber="1400">Tadkeshwar</collectingCounty>
mine,
<collectingRegion id="3650F80FB617902C75EEFE47C488F8F5" box="[1275,1437,478,500]" country="India" name="Gujarat" pageId="2" pageNumber="1400">Gujarat state</collectingRegion>
,
<collectingCountry id="8C83767DB617902C722EFE65C26CFB14" box="[827,889,508,530]" name="India" pageId="2" pageNumber="1400">India</collectingCountry>
; IR spectrum no. 14258 IAA;
<figureCitation id="6CAF2A68B617902C75DDFE65C432FB17" box="[1224,1319,508,530]" captionStart="Figure 11" captionStartId="16.[163,242,1861,1883]" captionTargetBox="[227,1379,195,1821]" captionTargetId="figure-26@16.[227,1379,195,1821]" captionTargetPageId="16" captionText="Figure 11. Fourier transform infrared spectroscopy (attenuated total reflectance) spectra obtained from Cambay amber chunks with the examined inclusions (early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India)." pageId="2" pageNumber="1400">Fig. 11A</figureCitation>
).
<typeStatus id="2B2F884FB617902C7429FE65C4B7FB14" box="[1340,1442,508,530]" pageId="2" pageNumber="1400" type="holotype">Holotype</typeStatus>
and
<typeStatus id="2B2F884FB617902C7266FD82C2F6FB36" box="[883,995,539,560]" pageId="2" pageNumber="1400" type="paratype">paratype</typeStatus>
, inventory no.
<emphasis id="C6E0EAFFB617902C75B7FD82C47FFB37" box="[1186,1386,539,561]" italics="true" pageId="2" pageNumber="1400">BSIP Tad-607 a</emphasis>
and b, syninclusions (see
<bibRefCitation id="90054B1CB617902C7551FDA3C47EFB49" author="Zakrzewska M &amp; Stebner F &amp; Puchalski M &amp; Singh H &amp; Gilka W" box="[1092,1387,570,592]" pageId="2" pageNumber="1400" pagination="255 - 261" refId="ref16888" refString="Zakrzewska M, Stebner F, Puchalski M, Singh H, Gilka W. 2018. A peculiar leg structure in the first non-biting midge described from Cambay amber, India (Diptera: Chironomidae). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107: 255-261." type="journal article" year="2018">
Zakrzewska
<emphasis id="C6E0EAFFB617902C75F4FDA3C435FB49" box="[1249,1312,570,591]" italics="true" pageId="2" pageNumber="1400">et al.</emphasis>
, 2018
</bibRefCitation>
); IR spectrum no. 14259 IAA (
<figureCitation id="6CAF2A68B617902C7549FDC1C5AFFB6B" box="[1116,1210,600,622]" captionStart="Figure 11" captionStartId="16.[163,242,1861,1883]" captionTargetBox="[227,1379,195,1821]" captionTargetId="figure-26@16.[227,1379,195,1821]" captionTargetPageId="16" captionText="Figure 11. Fourier transform infrared spectroscopy (attenuated total reflectance) spectra obtained from Cambay amber chunks with the examined inclusions (early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat, India)." pageId="2" pageNumber="1400">Fig. 11B</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,95 @@
<document id="717422E3AD254B9972AA964A8754F509" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB61890237010FBEAC3EDFC29" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Stempellinella Brundin 1947" docType="treatment" docVersion="2" lastPageNumber="1411" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1411" updateTime="1739358162056" updateUser="ExternalLinkService">
<mods:mods id="788F0CD5B15806909320F90F11E9EFD5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="50A3689D49035748AFFCB280CACCBF30">
<mods:title id="47464CEE1313B316A543094F8DCD8F51">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="BFFA9EB31A8A9B211FD19E377069E7BE" type="personal">
<mods:role id="E4C0BD525569011BF0954DED13A578EB">
<mods:roleTerm id="FC687514EC21BF1E7D6FDEC996B29C09">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="254F8F613BD9BD699BD0B050E987ACBA">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="087915DBDCE0902A85610519CB40FC6F" type="personal">
<mods:role id="FBD726A2BEFEA9F8823E092FD4D5903A">
<mods:roleTerm id="E938596BF44F84F68B545409BDE14178">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CBF51FC4FC658B7DE7032C4E85DA135">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="9AB65547CD97CBFBD2994B2F970AC675" type="personal">
<mods:role id="9D398AA23C270F47990514D5F0E3A00D">
<mods:roleTerm id="3426F1DEF0FFF0A6B4D502E99D4A707A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAA0E8AAEE08A34D7E6372DF0EF8EC94">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="91437A294E54E3D5C178C41A35DF742D" type="personal">
<mods:role id="C28D95C6FAE8A37162EB70DF9EEC791D">
<mods:roleTerm id="D29C1AEE7FDE277C0513C886F81AD270">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E63DCDCCB05DBF13716E2E7E88E2E5E">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="B5FD1AA7A36738747D1EC8CE28D742B4">text</mods:typeOfResource>
<mods:relatedItem id="41FB65C865CB008B710391E8149B068E" type="host">
<mods:titleInfo id="A93BDB9B341C3F3518F6FAED8AAE60A4">
<mods:title id="6C90D0BA2467196F70C90D656736849D">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="0AF4E92A6F044E45365F08C18586F3BD">
<mods:date id="F1C60E45E5990D1FC03FF2A9838972A2">2020</mods:date>
<mods:detail id="D9DB408F8C00CCF741BDFCCF307F9A20" type="volume">
<mods:number id="1C52A6CED4682C64D90F3776EF925F1D">189</mods:number>
</mods:detail>
<mods:extent id="895B46CB493A5164DBC96B1D23413955" unit="page">
<mods:start id="7F474FF1A769D52CB64C46C571E53F50">1398</mods:start>
<mods:end id="40E31D733D88C668ACA90FC8ADDF02D9">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="35135C70AC9AA7E3CDA7DA4B7380575F">journal article</mods:classification>
<mods:identifier id="81930F9C8510E1E6A74932BDDB0E9207" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="DC29DD821D9492E6F4127F25944BDD3E" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB61890237010FBEAC3EDFC29" ID-DOI="http://doi.org/10.5281/zenodo.14858052" ID-Zenodo-Dep="14858052" LSID="urn:lsid:plazi:treatment:7C3D87FBB61890237010FBEAC3EDFC29" httpUri="http://treatment.plazi.org/id/7C3D87FBB61890237010FBEAC3EDFC29" lastPageNumber="1411" pageId="13" pageNumber="1411">
<subSubSection id="BC8E6566B61890237010FBEAC391FD8A" box="[261,644,1139,1164]" pageId="13" pageNumber="1411" type="nomenclature">
<paragraph id="F42B36EDB61890237010FBEAC391FD8A" blockId="13.[261,644,1139,1164]" box="[261,644,1139,1164]" pageId="13" pageNumber="1411">
<heading id="AF638181B61890237010FBEAC391FD8A" box="[261,644,1139,1164]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="1411" reason="2">
<taxonomicName id="33944D6EB61890237010FBEAC391FD8A" ID-CoL="7MCT" authority="BRUNDIN, 1947" authorityName="Brundin" authorityYear="1947" box="[261,644,1139,1164]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1411" phylum="Arthropoda" rank="genus">
<smallCapsWord id="F2CDA031B61890237010FBEAC0DBFD8D" baselines="1156,1158" box="[261,462,1139,1163]" lowerCaseFontSize="10" mainFontSize="10" normCase="title" normString="Stempellinella" pageId="13" pageNumber="1411">
<emphasis id="C6E0EAFFB61890237010FBEAC0DBFD8D" box="[261,462,1139,1163]" italics="true" pageId="13" pageNumber="1411">STEMPELLINELLA</emphasis>
</smallCapsWord>
<smallCapsWord id="F2CDA031B618902370C1FBEAC32AFD8D" baselines="1158,1158" box="[468,575,1139,1164]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Brundin" pageId="13" pageNumber="1411">BRUNDIN</smallCapsWord>
, 1947
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B61890237184FB25C3EDFC29" pageId="13" pageNumber="1411" type="materials_examined">
<paragraph id="F42B36EDB61890237184FB25C1C2FDE9" blockId="13.[145,760,1211,1263]" pageId="13" pageNumber="1411">
<emphasis id="C6E0EAFFB61890237184FB25C03DFDD7" box="[145,296,1212,1233]" italics="true" pageId="13" pageNumber="1411">
<typeStatus id="2B2F884FB61890237184FB25C1DDFDD7" box="[145,200,1212,1233]" pageId="13" pageNumber="1411">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="33944D6EB61890237023FB22C1C6FDE9" authority="(Goetghebuer, 1921)" baseAuthorityName="Goetghebuer" baseAuthorityYear="1921" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1411" phylum="Arthropoda" rank="species" species="saltuum">
<emphasis id="C6E0EAFFB61890237023FB22C35EFDD6" box="[310,587,1211,1232]" italics="true" pageId="13" pageNumber="1411">Stempellinella saltuum</emphasis>
(Goetghebuer, 1921)
</taxonomicName>
.
</paragraph>
<paragraph id="F42B36EDB61890237187FA8EC3EDFC29" blockId="13.[146,760,1303,1358]" box="[146,760,1303,1328]" pageId="13" pageNumber="1411">
<taxonomicName id="33944D6EB61890237187FA8EC3D8FC29" authority="Gilka &amp; Zakrzewska" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[146,717,1303,1328]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1416" phylum="Arthropoda" rank="species" species="pollex" status="sp. nov.">
<emphasis id="C6E0EAFFB61890237187FA8EC0D0FC28" bold="true" box="[146,453,1303,1327]" italics="true" pageId="13" pageNumber="1411">
<smallCapsWord id="F2CDA031B61890237187FA8EC076FC28" baselines="1321,1321" box="[146,355,1303,1327]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Stempellinella" pageId="13" pageNumber="1411">STEMPELLINELLA</smallCapsWord>
<smallCapsWord id="F2CDA031B6189023707EFA82C0D0FC28" baselines="1321" box="[363,453,1307,1326]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="pollex" pageId="13" pageNumber="1411">POLLEX</smallCapsWord>
</emphasis>
<smallCapsWord id="F2CDA031B618902370D9FA8EC301FC29" baselines="1322,1322" box="[460,532,1303,1328]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gilka" pageId="13" pageNumber="1411">GIŁKA</smallCapsWord>
&amp;
<smallCapsWord id="F2CDA031B61890237322FA8EC3D8FC29" baselines="1322,1322" box="[567,717,1303,1328]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zakrzewska" pageId="13" pageNumber="1411">ZAKRZEWSKA</smallCapsWord>
</taxonomicName>
<emphasis id="C6E0EAFFB618902373C1FA83C3EDFC29" bold="true" box="[724,760,1303,1327]" pageId="13" pageNumber="1411">
<taxonomicNameLabel id="DDD35784B618902373C1FA83C3EDFC29" box="[724,760,1303,1327]" pageId="13" pageNumber="1411">
<smallCapsWord id="F2CDA031B618902373C1FA83C3E4FC2B" baselines="1320" box="[724,753,1306,1325]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="13" pageNumber="1411">SP</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,187 @@
<document id="21D8BCC095B87DF091BA7AE171D302F1" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB61C90277075FAE0C421FD74" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Tanytarsus SPP." docType="treatment" docVersion="1" lastPageNumber="1407" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1407" updateTime="1739357823542" updateUser="GgImagineBatch">
<mods:mods id="E25FB90A673ED7A606190956766E54A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F879F22DCAAB89D166A9D09BBA55AFA9">
<mods:title id="FF6036DC7BCA47FDC72F0EDD5ACA4464">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="2CE863127244FA4A56C437B21A822E59" type="personal">
<mods:role id="12E4C40089ADA4D725C6EC03A61FC496">
<mods:roleTerm id="FF4B37BB5E53B890AA319CCF0ABDD99A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32DE7187AAE7A8DEA0BD7DACA8418AD5">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="A2490D664C023D43E40F93F254C82AD6" type="personal">
<mods:role id="2D789A014AFB6F4B2E28289E87914BE7">
<mods:roleTerm id="18BA071748363BDF79CE73F4EF86193B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99D70BA1C41ABE035BD96FDE4999E496">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="4A5F4BE2314BFE759A016B8FA5DD7D22" type="personal">
<mods:role id="47317340A12A9141DA7BC7B017FB11A8">
<mods:roleTerm id="E2D7F7145D06F87A06A2D1CCFA8F00FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0194F0B517A7010814BBC95ABA6E22DF">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="3AFEE72ECE92FADC23157EBC34EE675B" type="personal">
<mods:role id="253373B227E28E6451BF8CF63220A0E8">
<mods:roleTerm id="F51E8C1C1190922EB2AC0A0A79148AEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF38CA1BEBC7EB8CDDC41A2E3ABB737E">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="39C3FABD158CFB52175A7A38C4EBB40C">text</mods:typeOfResource>
<mods:relatedItem id="64611F6C6AF7AF6885E352D8D7B6BAB1" type="host">
<mods:titleInfo id="85132F2387364B261CD7CF2DAE9113D9">
<mods:title id="94B88AFA6C640D3FD7633CA70A107A67">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E58C83F829FD6EA6B4D2ADE16F188BDF">
<mods:date id="6B30C13B96C0A99D2E0E2965F790FB92">2020</mods:date>
<mods:detail id="089E1A8DD8B4201347D272F83E75AFD9" type="volume">
<mods:number id="FF080DE8CC10512E009AD599CCDC6D92">189</mods:number>
</mods:detail>
<mods:extent id="F9E9050D2EC716F8EE9C1162F91EF116" unit="page">
<mods:start id="204017F74C29906B6E9EE97B81B2A272">1398</mods:start>
<mods:end id="0A684689D72D36598DD4B8C2AE4A3411">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4BD622C735F2597016171A67FEEDA132">journal article</mods:classification>
<mods:identifier id="6832CEEB2820CA6033511B9FCE911D5A" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="B5CB32F7E549C700FDE01CE765F2CA22" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB61C90277075FAE0C421FD74" LSID="urn:lsid:plazi:treatment:7C3D87FBB61C90277075FAE0C421FD74" httpUri="http://treatment.plazi.org/id/7C3D87FBB61C90277075FAE0C421FD74" lastPageNumber="1407" pageId="9" pageNumber="1407">
<subSubSection id="BC8E6566B61C90277075FAE0C33FFC97" box="[352,554,1400,1425]" pageId="9" pageNumber="1407" type="nomenclature">
<paragraph id="F42B36EDB61C90277075FAE0C33FFC97" blockId="9.[352,554,1400,1425]" box="[352,554,1400,1425]" pageId="9" pageNumber="1407">
<heading id="AF638181B61C90277075FAE0C33FFC97" box="[352,554,1400,1425]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="1407" reason="2">
<taxonomicName id="33944D6EB61C90277075FAE0C33FFC97" ID-CoL="7SJQ" authority="SPP." authorityName="SPP." box="[352,554,1400,1425]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<smallCapsWord id="F2CDA031B61C90277075FAE0C0E1FC96" baselines="1418,1419" box="[352,500,1401,1424]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tanytarsus" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277075FAE0C0E1FC96" box="[352,500,1401,1424]" italics="true" pageId="9" pageNumber="1407">TANYTARSUS</emphasis>
</smallCapsWord>
<smallCapsWord id="F2CDA031B61C902770E9FAE5C331FC96" baselines="1419" box="[508,548,1404,1424]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="spp" pageId="9" pageNumber="1407">SPP</smallCapsWord>
.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B61C90277097FA2DC421FD74" pageId="9" pageNumber="1407" type="description">
<paragraph id="F42B36EDB61C90277097FA2DC312FCCB" blockId="9.[386,519,1460,1485]" box="[386,519,1460,1485]" pageId="9" pageNumber="1407">
(
<figureCitation id="6CAF2A68B61C9027709FFA2DC0EAFCCB" box="[394,511,1460,1485]" captionStart="Figure 6" captionStartId="10.[163,242,1290,1312]" captionTargetBox="[163,1443,195,1250]" captionTargetId="figure-240@10.[163,1443,195,1250]" captionTargetPageId="10" captionText="Figure 6. Tanytarsus spp., adult males, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India. A, B, Tanytarsus sp. 1, BSIP Tad-533. C, D, Tanytarsus sp. 2, BSIP Tad-883 a. E, F, Tanytarsus sp. 3, BSIP Tad-884. A, C, E, inclusion in amber. B, D, F, habitus." pageId="9" pageNumber="1407">
<smallCapsWord id="F2CDA031B61C9027709FFA2DC0A6FCCA" baselines="1479,1479" box="[394,435,1460,1485]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Fig" pageId="9" pageNumber="1407">FIG</smallCapsWord>
. 6AF
</figureCitation>
)
</paragraph>
<paragraph id="F42B36EDB61C90277184FA45C06DFCF2" blockId="9.[145,376,1500,1524]" box="[145,376,1500,1524]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277184FA45C06DFCF2" box="[145,376,1500,1524]" italics="true" pageId="9" pageNumber="1407">Material examined</emphasis>
</paragraph>
<paragraph id="F42B36EDB61C90277184F99FC500F8D6" blockId="9.[145,760,1541,1594]" lastBlockId="9.[809,1426,197,464]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277184F99FC027FF1D" box="[145,306,1542,1563]" italics="true" pageId="9" pageNumber="1407">Ta n y t a r s u s</emphasis>
s p. 1, o n e a d u l t m a l e i n a
<materialsCitation id="44FC3CB0B61C902773FFF99CC504F8D6" country="India" county="Tadkeshwar" latitude="8.5" location="Coleoptera" municipality="Mya" pageId="9" pageNumber="1407" specimenCount="2" specimenCount-adult="2" stateProvince="Gujarat">
<geoCoordinate id="91A0502AB61C902773FFF99CC1A1FF3F" degrees="8.5" direction="north" orientation="latitude" pageId="9" pageNumber="1407" precision="5555" value="8.5">
n
<quantity id="336C9B08B61C90277184F9BAC1A1FF3F" box="[145,180,1571,1593]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="1407" unit="mm" value="8.5">8.5</quantity>
</geoCoordinate>
mm ×
<quantity id="336C9B08B61C9027701DF9BDC05FFF3F" box="[264,330,1572,1594]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="1407" unit="mm" value="6.0">6 mm</quantity>
×
<quantity id="336C9B08B61C9027707FF9BAC0D7FF3F" box="[362,450,1571,1593]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="1407" unit="mm" value="3.5">3.5 mm</quantity>
piece of amber (inventory no. BSIP Tad-533;
<figureCitation id="6CAF2A68B61C9027750FFF5CC58EF9DD" box="[1050,1179,197,219]" captionStart="Figure 6" captionStartId="10.[163,242,1290,1312]" captionTargetBox="[163,1443,195,1250]" captionTargetId="figure-240@10.[163,1443,195,1250]" captionTargetPageId="10" captionText="Figure 6. Tanytarsus spp., adult males, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India. A, B, Tanytarsus sp. 1, BSIP Tad-533. C, D, Tanytarsus sp. 2, BSIP Tad-883 a. E, F, Tanytarsus sp. 3, BSIP Tad-884. A, C, E, inclusion in amber. B, D, F, habitus." pageId="9" pageNumber="1407">Fig. 6A, B</figureCitation>
);
<taxonomicName id="33944D6EB61C902775A2FF5FC49EF9DC" box="[1207,1419,197,219]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="C6E0EAFFB61C902775A2FF5FC456F9DD" box="[1207,1347,198,219]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
sp. 2
</taxonomicName>
,
<specimenCount id="E292FD64B61C9027723CFF7DC289F9FC" box="[809,924,228,250]" count="1" pageId="9" pageNumber="1407" type="adult">one adult</specimenCount>
male in a
<quantity id="336C9B08B61C90277537FF7DC568F9FF" box="[1058,1149,228,250]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="1407" unit="mm" value="6.5">6.5 mm</quantity>
×
<quantity id="336C9B08B61C9027758AFF7DC5F6F9FF" box="[1183,1251,228,250]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="1407" unit="mm" value="5.0">5 mm</quantity>
×
<quantity id="336C9B08B61C90277413FF7DC45FF9FF" box="[1286,1354,228,249]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="1407" unit="mm" value="4.0">4 mm</quantity>
piece of amber (inventory no. BSIP Tad-883 a;
<figureCitation id="6CAF2A68B61C90277424FE9BC229F831" captionStart="Figure 6" captionStartId="10.[163,242,1290,1312]" captionTargetBox="[163,1443,195,1250]" captionTargetId="figure-240@10.[163,1443,195,1250]" captionTargetPageId="10" captionText="Figure 6. Tanytarsus spp., adult males, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India. A, B, Tanytarsus sp. 1, BSIP Tad-533. C, D, Tanytarsus sp. 2, BSIP Tad-883 a. E, F, Tanytarsus sp. 3, BSIP Tad-884. A, C, E, inclusion in amber. B, D, F, habitus." pageId="9" pageNumber="1407">Fig. 6C, D</figureCitation>
), syninclusion:
<location id="F14B6036B61C90277513FEB8C599F831" LSID="urn:lsid:plazi:treatment:7C3D87FBB61C90277075FAE0C421FD74:F14B6036B61C90277513FEB8C599F831" box="[1030,1164,289,311]" country="India" county="Tadkeshwar" latitude="8.5" municipality="Mya" name="Coleoptera" pageId="9" pageNumber="1407" stateProvince="Gujarat">Coleoptera</location>
(inventory no. BSIP Tad-883 b);
<taxonomicName id="33944D6EB61C902772ABFED9C599F853" box="[958,1164,320,341]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="C6E0EAFFB61C902772ABFED9C553F853" box="[958,1094,320,341]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
sp. 3
</taxonomicName>
,
<specimenCount id="E292FD64B61C90277589FED9C405F850" box="[1180,1296,320,342]" count="1" pageId="9" pageNumber="1407" type="adult">one adult</specimenCount>
male in a
<quantity id="336C9B08B61C9027723CFEC7C278F872" box="[809,877,350,372]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="1407" unit="mm" value="6.0">6 mm</quantity>
×
<quantity id="336C9B08B61C9027729BFEC6C2C7F872" box="[910,978,351,372]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="1407" unit="mm" value="3.0">3 mm</quantity>
×
<quantity id="336C9B08B61C902772E6FEC6C52DF872" box="[1011,1080,351,372]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="1407" unit="mm" value="1.0">1 mm</quantity>
piece of amber preserved in a cubicoid plastic mass (inventory no. BSIP Tad-884;
<figureCitation id="6CAF2A68B61C9027723CFE05C283F8B7" box="[809,918,412,434]" captionStart="Figure 6" captionStartId="10.[163,242,1290,1312]" captionTargetBox="[163,1443,195,1250]" captionTargetId="figure-240@10.[163,1443,195,1250]" captionTargetPageId="10" captionText="Figure 6. Tanytarsus spp., adult males, early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India. A, B, Tanytarsus sp. 1, BSIP Tad-533. C, D, Tanytarsus sp. 2, BSIP Tad-883 a. E, F, Tanytarsus sp. 3, BSIP Tad-884. A, C, E, inclusion in amber. B, D, F, habitus." pageId="9" pageNumber="1407">Fig. 6E, F</figureCitation>
);
<location id="F14B6036B61C902772BEFE05C550F8B7" LSID="urn:lsid:plazi:treatment:7C3D87FBB61C90277075FAE0C421FD74:F14B6036B61C902772BEFE05C550F8B7" box="[939,1093,412,434]" country="India" county="Tadkeshwar" latitude="8.5" municipality="Mya" name="Early Eocene" pageId="9" pageNumber="1407" stateProvince="Gujarat">Early Eocene</location>
, ~54
<collectingMunicipality id="144FAC97B61C90277595FE05C5A1F8B7" box="[1152,1204,412,433]" pageId="9" pageNumber="1407">Mya</collectingMunicipality>
,
<collectingCounty id="1D4A4E61B61C902775ABFE05C45EF8B4" box="[1214,1355,412,434]" pageId="9" pageNumber="1407">Tadkeshwar</collectingCounty>
mine,
<collectingRegion id="3650F80FB61C9027723CFE23C2D1F8D6" box="[809,964,442,464]" country="India" name="Gujarat" pageId="9" pageNumber="1407">Gujarat state</collectingRegion>
,
<collectingCountry id="8C83767DB61C902772C5FE23C504F8D6" box="[976,1041,442,464]" name="India" pageId="9" pageNumber="1407">India</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="F42B36EDB61C9027723CFD90C283FB27" blockId="9.[809,918,521,545]" box="[809,918,521,545]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C9027723CFD90C283FB27" box="[809,918,521,545]" italics="true" pageId="9" pageNumber="1407">Remarks</emphasis>
</paragraph>
<paragraph id="F42B36EDB61C9027723CFDA8C5A1FD34" blockId="9.[809,1426,561,1074]" pageId="9" pageNumber="1407">
The above
<specimenCount id="E292FD64B61C902772ACFDA8C59EFB41" box="[953,1163,561,583]" count="3" pageId="9" pageNumber="1407" type="generic">three specimens</specimenCount>
are ascribed to the genus
<taxonomicName id="33944D6EB61C90277260FDC8C2E2FB60" box="[885,1015,593,614]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E0EAFFB61C90277260FDC8C2E2FB60" box="[885,1015,593,614]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
</taxonomicName>
on the basis of the head, wing and hypopygium characters (cf.
<bibRefCitation id="90054B1CB61C9027759DFDF7C495FB82" author="Stebner F &amp; Baranov V &amp; Zakrzewska M &amp; Singh H &amp; Gilka W" box="[1160,1408,622,644]" pageId="9" pageNumber="1407" pagination="154 - 161" refId="ref16522" refString="Stebner F, Baranov V, Zakrzewska M, Singh H, Gilka W. 2017. The Chironomidae diversity based on records from early Eocene Cambay amber, India, with implications on habitats of fossil Diptera. Palaeogeography, Palaeoclimatology, Palaeoecology 475: 154-161." type="journal article" year="2017">
Stebner
<emphasis id="C6E0EAFFB61C902775E1FDF6C426FB85" box="[1268,1331,622,644]" italics="true" pageId="9" pageNumber="1407">et al.</emphasis>
, 2017
</bibRefCitation>
). However, most of the diagnostic structures are weakly observable, deformed or insufficiently preserved to define the individuals definitively to species level. Nevertheless, these males differ distinctly from each other and from those designated as new species described in this paper, thus they most probably belong to another species. Therefore, with presumably five species,
<taxonomicName id="33944D6EB61C902772D9FCFDC542FA7F" box="[972,1111,868,889]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E0EAFFB61C902772D9FCFDC542FA7F" box="[972,1111,868,889]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
</taxonomicName>
is the most species-rich genus among the
<collectingCountry id="8C83767DB61C902772EAFC1BC575FA9E" box="[1023,1120,898,920]" name="India" pageId="9" pageNumber="1407">Cambay</collectingCountry>
<taxonomicName id="33944D6EB61C9027757DFC1BC5E2FA9E" box="[1128,1271,898,920]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
. The specific diversity proportions within the tribe may thus be similar relative to the extant fauna: five
<taxonomicName id="33944D6EB61C9027741DFC59C49FFAD3" box="[1288,1418,960,981]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E0EAFFB61C9027741DFC59C49FFAD3" box="[1288,1418,960,981]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
</taxonomicName>
/ eight
<taxonomicName id="33944D6EB61C9027726CFC46C501FAF2" box="[889,1044,990,1012]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
in
<collectingCountry id="8C83767DB61C9027755FFC46C5A6FAF2" box="[1098,1203,990,1012]" name="India" pageId="9" pageNumber="1407">Cambay</collectingCountry>
amber vs. ~360
<taxonomicName id="33944D6EB61C9027723CFC67C2BCFD15" box="[809,937,1022,1043]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<emphasis id="C6E0EAFFB61C9027723CFC67C2BCFD15" box="[809,937,1022,1043]" italics="true" pageId="9" pageNumber="1407">Tanytarsus</emphasis>
</taxonomicName>
/~700
<taxonomicName id="33944D6EB61C902772FAFC64C563FD15" box="[1007,1142,1021,1043]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
extant species (Giłka W, Zakrzewska M, unpublished data).
</paragraph>
<paragraph id="F42B36EDB61C90277290FBC0C421FD74" blockId="9.[901,1332,1113,1138]" box="[901,1332,1113,1138]" pageId="9" pageNumber="1407">
<smallCapsWord id="F2CDA031B61C90277290FBC0C2E3FD76" baselines="1132,1131" box="[901,1014,1113,1138]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Subtribe" pageId="9" pageNumber="1407">SUBTRIBE</smallCapsWord>
:
<taxonomicName id="33944D6EB61C90277516FBC0C421FD74" authority="SAETHER, 1977" authorityName="SAETHER" authorityYear="1977" box="[1027,1332,1113,1138]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="tribe" tribe="Zavreliina">
<smallCapsWord id="F2CDA031B61C90277516FBC0C59DFD76" baselines="1132,1131" box="[1027,1160,1113,1138]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zavreliina" pageId="9" pageNumber="1407">ZAVRELIINA</smallCapsWord>
<bibRefCitation id="90054B1CB61C9027759AFBC0C421FD74" author="Saether OA" box="[1167,1332,1113,1138]" pageId="9" pageNumber="1407" pagination="1 - 209" refId="ref16249" refString="Saether OA. 1977. Female genitalia in Chironomidae and other Nematocera: morphology, phylogenies, keys. Bulletin of Fisheries Research Board of Canada 197: 1-209." type="journal article" year="1977">
<smallCapsWord id="F2CDA031B61C9027759AFBC0C5FAFD76" baselines="1132,1131" box="[1167,1263,1113,1138]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Saether" pageId="9" pageNumber="1407">SAETHER</smallCapsWord>
, 1977
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,97 @@
<document id="983A0F0A87E2823C5E9CD50C672C25C9" ID-ISSN="0024-4082" ID-ZooBank="4FC7C16-C9EB-49F5-9E63-208751C04B65" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353359319" checkinUser="plazi" docAuthor="Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa &amp; Giłka, Wojciech" docDate="2020" docId="7C3D87FBB61C90277276FB11C492FC13" docLanguage="en" docName="zlz159.pdf" docOrigin="Zoological Journal of the Linnean Society 189" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Stempellina Thienemann &amp; Bause 1913" docType="treatment" docVersion="2" lastPageNumber="1407" masterDocId="8004FF83B615902E7115FF99C115F906" masterDocTitle="Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber" masterLastPageNumber="1425" masterPageNumber="1398" pageNumber="1407" updateTime="1739358162056" updateUser="ExternalLinkService">
<mods:mods id="4C2069D20FA9E9A1C77C529971B0E599" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B87456EA1D05CBD88BD8EDEF7AF2D86C">
<mods:title id="E91D6A904D1710709DB98E042EF2D6E8">Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber</mods:title>
</mods:titleInfo>
<mods:name id="39755439836C3D1441119E493B9D3912" type="personal">
<mods:role id="828AFDDA944F53E2C73A9EB326BF7738">
<mods:roleTerm id="E3512017EE4EDD12EC06395977AA2AD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="261020F1042D72936E55EE721959EDC3">Zakrzewska, Marta</mods:namePart>
</mods:name>
<mods:name id="7016A8E14D0B7A44FC110697E743A4A6" type="personal">
<mods:role id="6A4777B793789207F2AEC765A65C30FB">
<mods:roleTerm id="876668B23FFFF50E933936B6372F08A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E20D03BEC296D9AE145CF8952D5A2832">Singh, Hukam</mods:namePart>
</mods:name>
<mods:name id="4CC6B48B9EEC3EDF6AB7EAF2B5083BEB" type="personal">
<mods:role id="F9723E4C73BF856A2DB2A74FBE62F1C4">
<mods:roleTerm id="40D9F7FC6F4FCF6A6D7C34CB9C7A35AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9738B33AAAFF906133269E591C8782F">Wagner-Wysiecka, Ewa</mods:namePart>
</mods:name>
<mods:name id="FBB3320AC7377484819A346B1B9EAD5E" type="personal">
<mods:role id="24897A13022F73CF3A15FE3B5401D42F">
<mods:roleTerm id="0B08F33035CCE219E04C43CEB01B48E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9E0E4044D0657462A53D1C9750FFBBD">Giłka, Wojciech</mods:namePart>
</mods:name>
<mods:typeOfResource id="8A74D2FD1056D9A6906E133D3D407D42">text</mods:typeOfResource>
<mods:relatedItem id="9F7DC11AF9A97B543795EAC5FC2D0097" type="host">
<mods:titleInfo id="DE2FCEBBD98702437557B01D2FC5465A">
<mods:title id="C0D88BFA30D92F19DC2CE1D5591E65FE">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="25125491DF9A582BB325EEC69D19E7C6">
<mods:date id="F7A82C055383970D1EF1BA0D72B03E44">2020</mods:date>
<mods:detail id="6F38F779A4A265C6373C23D746CDD2B5" type="volume">
<mods:number id="3A0B3C055B4BB5EC6B8562E62317E2FC">189</mods:number>
</mods:detail>
<mods:extent id="E9AF6BBE8C510F44F1A7B05D3D5770D4" unit="page">
<mods:start id="0A5056EAA3366AABC4DB9307DF54321C">1398</mods:start>
<mods:end id="8A046B53AAEFC6CE49D317018B5D4715">1425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D0C3AE77C921FCCE3A7259D8871427F3">journal article</mods:classification>
<mods:identifier id="E4CEA76EDE7D10C35EB6EA4BAE00C7FD" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="72F9C8722E2CDB9A16DB2E84C28A1082" type="ZooBank">4FC7C16-C9EB-49F5-9E63-208751C04B65</mods:identifier>
</mods:mods>
<treatment id="7C3D87FBB61C90277276FB11C492FC13" ID-DOI="http://doi.org/10.5281/zenodo.14858050" ID-Zenodo-Dep="14858050" LSID="urn:lsid:plazi:treatment:7C3D87FBB61C90277276FB11C492FC13" httpUri="http://treatment.plazi.org/id/7C3D87FBB61C90277276FB11C492FC13" lastPageNumber="1407" pageId="9" pageNumber="1407">
<subSubSection id="BC8E6566B61C90277276FB11C443FDA6" box="[867,1366,1159,1184]" pageId="9" pageNumber="1407" type="nomenclature">
<paragraph id="F42B36EDB61C90277276FB11C443FDA6" blockId="9.[867,1366,1159,1184]" box="[867,1366,1159,1184]" pageId="9" pageNumber="1407">
<heading id="AF638181B61C90277276FB11C443FDA6" box="[867,1366,1159,1184]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="1407" reason="2">
<taxonomicName id="33944D6EB61C90277276FB11C443FDA6" ID-CoL="7MCS" authority="THIENEMANN &amp; BAUSE, 1913" authorityName="Thienemann &amp; Bause" authorityYear="1913" box="[867,1366,1159,1184]" class="Insecta" family="Chironomidae" genus="Stempellina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="genus">
<smallCapsWord id="F2CDA031B61C90277276FB11C517FD99" baselines="1177,1178" box="[867,1026,1160,1183]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Stempellina" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C90277276FB11C517FD99" box="[867,1026,1160,1183]" italics="true" pageId="9" pageNumber="1407">STEMPELLINA</emphasis>
</smallCapsWord>
<smallCapsWord id="F2CDA031B61C9027751DFB1EC5B6FD99" baselines="1178,1178" box="[1032,1187,1159,1184]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Thienemann" pageId="9" pageNumber="1407">THIENEMANN</smallCapsWord>
&amp;
<smallCapsWord id="F2CDA031B61C902775D0FB1EC404FD99" baselines="1178,1178" box="[1221,1297,1159,1184]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Bause" pageId="9" pageNumber="1407">BAUSE</smallCapsWord>
, 1913
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BC8E6566B61C9027723CFB58C492FC13" pageId="9" pageNumber="1407" type="materials_examined">
<paragraph id="F42B36EDB61C9027723CFB58C441FDD3" blockId="9.[809,1364,1216,1238]" box="[809,1364,1216,1238]" pageId="9" pageNumber="1407">
<emphasis id="C6E0EAFFB61C9027723CFB58C2ACFDD3" box="[809,953,1216,1238]" italics="true" pageId="9" pageNumber="1407">
<typeStatus id="2B2F884FB61C9027723CFB58C24BFDD0" box="[809,862,1217,1238]" pageId="9" pageNumber="1407">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="33944D6EB61C902772D9FB59C45AFDD3" authority="(Kieffer, 1911)" baseAuthorityName="Kieffer" baseAuthorityYear="1911" box="[972,1359,1216,1238]" class="Insecta" family="Chironomidae" genus="Stempellina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1407" phylum="Arthropoda" rank="species" species="bausei">
<emphasis id="C6E0EAFFB61C902772D9FB59C5B0FDD3" box="[972,1189,1216,1237]" italics="true" pageId="9" pageNumber="1407">Stempellina bausei</emphasis>
(Kieffer, 1911)
</taxonomicName>
.
</paragraph>
<paragraph id="F42B36EDB61C90277226FB64C492FC13" blockId="9.[819,1415,1277,1331]" box="[819,1415,1277,1302]" pageId="9" pageNumber="1407">
<taxonomicName id="33944D6EB61C90277226FB64C449FC12" authority="Gilka &amp; Zakrzewska" authorityName="Gilka &amp; Zakrzewska" authorityYear="2020" box="[819,1372,1277,1302]" class="Insecta" family="Chironomidae" genus="Stempellina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="1416" phylum="Arthropoda" rank="species" species="stebneri" status="sp. nov.">
<emphasis id="C6E0EAFFB61C90277226FB64C541FC15" bold="true" box="[819,1108,1277,1301]" italics="true" pageId="9" pageNumber="1407">
<smallCapsWord id="F2CDA031B61C90277226FB64C2CCFC15" baselines="1295,1294" box="[819,985,1277,1301]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Stempellina" pageId="9" pageNumber="1407">STEMPELLINA</smallCapsWord>
<smallCapsWord id="F2CDA031B61C902772F5FA99C541FC15" baselines="1294" box="[992,1108,1280,1299]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="stebneri" pageId="9" pageNumber="1407">STEBNERI</smallCapsWord>
</emphasis>
<smallCapsWord id="F2CDA031B61C9027754EFB64C5B6FC12" baselines="1296,1295" box="[1115,1187,1277,1302]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gilka" pageId="9" pageNumber="1407">GIŁKA</smallCapsWord>
&amp;
<smallCapsWord id="F2CDA031B61C902775D3FB64C449FC12" baselines="1296,1295" box="[1222,1372,1277,1302]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zakrzewska" pageId="9" pageNumber="1407">ZAKRZEWSKA</smallCapsWord>
</taxonomicName>
<emphasis id="C6E0EAFFB61C90277476FA99C492FC13" bold="true" box="[1379,1415,1277,1301]" pageId="9" pageNumber="1407">
<taxonomicNameLabel id="DDD35784B61C90277476FA99C492FC13" box="[1379,1415,1277,1301]" pageId="9" pageNumber="1407">
<smallCapsWord id="F2CDA031B61C90277476FA99C495FC15" baselines="1294" box="[1379,1408,1280,1299]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="9" pageNumber="1407">SP</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,122 @@
<document id="B8AA7C3E024AAAC7A2BF0CA9536F65E5" ID-DOI="10.1093/zoolinnean/zlz160" ID-ISSN="0024-4082" ID-Zenodo-Dep="14858057" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1739353363407" checkinUser="plazi" docAuthor="Falniowski, Andrzej, Heller, Joseph, Cameron, Robert A D, Pokryszko, Beata M, Osikowski, Artur, Rysiewska, Aleksandra &amp; Hofman, Sebastian" docDate="2020" docId="921887F80D698F54FC90FB6A2049F237" docLanguage="en" docName="zlz160.pdf" docOrigin="Zoological Journal of the Linnean Society 190 (2)" docSource="https://academic.oup.com/zoolinnean/article/190/2/483/5788722" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Melanopsis astropaliae" docType="treatment" docVersion="2" lastPageNumber="497" masterDocId="6E21FF800D678F5AFFB9FFA9245BF57E" masterDocTitle="Melanopsidae (Caenogastropoda: Cerithioidea) from the eastern Mediterranean: another case of morphostatic speciation" masterLastPageNumber="507" masterPageNumber="483" pageNumber="497" updateTime="1739358232563" updateUser="ExternalLinkService">
<mods:mods id="8B8921E536B02FE82015743403F375CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="37ADB96378562F2351C8D313135BA1CE">
<mods:title id="D1F3DFE441CAC197A2E4042DAFADA918">Melanopsidae (Caenogastropoda: Cerithioidea) from the eastern Mediterranean: another case of morphostatic speciation</mods:title>
</mods:titleInfo>
<mods:name id="6A0D934B3394D23D88407BE2461BC10A" type="personal">
<mods:role id="49F3260D9338BBF81516A5156C372DED">
<mods:roleTerm id="3AE06C4D529E2740B4C9FB506AFEA930">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BD9EE86E40B8267FC68F06997694F6F">Falniowski, Andrzej</mods:namePart>
</mods:name>
<mods:name id="4D197436C7EDFFDD71F75651A18224B1" type="personal">
<mods:role id="92DA476453265DF13A94B7E3683CD98B">
<mods:roleTerm id="994E27E5A5ADDBF2DBE7C5DEAD7F601B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2D403BD2A97A8B9A087BF633B250C17">Heller, Joseph</mods:namePart>
</mods:name>
<mods:name id="40AB300C43E9C6EEBA4687A7A6EB6D50" type="personal">
<mods:role id="75B80C55F5CCDB9FF07DECABE881D070">
<mods:roleTerm id="4A1DB0147DFE9BF983D47D26623955CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5B480F6C593972FF4232E07D218DAA4">Cameron, Robert A D</mods:namePart>
</mods:name>
<mods:name id="3EB25624846EE9EA8059FA1BDE3984F8" type="personal">
<mods:role id="89051B6CBEF800D08B9F98DB94B59F16">
<mods:roleTerm id="DA29BD4E0005EFF38E07636444A2935C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="948A852151EE0D39DA9281118ACA45B1">Pokryszko, Beata M</mods:namePart>
</mods:name>
<mods:name id="44BAF2E083496C184F597F474A21BC07" type="personal">
<mods:role id="9A00905EB4D09129F8128A6A8D340397">
<mods:roleTerm id="8B102FD6874D346FC661980CFB0384D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56F6863381B83164C68CF4B99F098456">Osikowski, Artur</mods:namePart>
</mods:name>
<mods:name id="4977177270C9D417A28DD20F416C62D9" type="personal">
<mods:role id="0979FA0192A78AF3D8A5A3F871804BA2">
<mods:roleTerm id="BF3E0BFB5A90A500C5DB5629B22E6395">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F6DD3A779822CAB880EA54EECE83894">Rysiewska, Aleksandra</mods:namePart>
</mods:name>
<mods:name id="BF4D6A9AA2581F71C68A31816D28247C" type="personal">
<mods:role id="559E815630E03AF3A34531EE2BBA936A">
<mods:roleTerm id="32F2BF5F74D0585072310BC41E3FA148">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B848205E2B2D6CF574E70DE94B64ECC">Hofman, Sebastian</mods:namePart>
</mods:name>
<mods:typeOfResource id="23DB426AD617E732975B14B86BEAA260">text</mods:typeOfResource>
<mods:relatedItem id="50BFB98AE1BCA1373A6578A3011D6599" type="host">
<mods:titleInfo id="824D2F36E374AD2E9A89CA416050D446">
<mods:title id="513C3F5C32E1F3A39C9C51D01BF87E30">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="1C86C7094B7735749BAD77BC9A01D109">
<mods:date id="851D98F34341A6EB8370FB7B67783C43">2020</mods:date>
<mods:detail id="7C5B0F89395DA8A5311D8FC55137A4B2" type="pubDate">
<mods:number id="C94990CCA833815FF552B28F9565EA31">2020-10-31</mods:number>
</mods:detail>
<mods:detail id="A0019E7330D38BF4FCF341647EB695C3" type="volume">
<mods:number id="45B41B17060AA5297A28FD0D1FA862D9">190</mods:number>
</mods:detail>
<mods:detail id="C31803789707C509A1E8E995B13C3AAE" type="issue">
<mods:number id="565DF9037AB297F4F5BA3A254DDF64F9">2</mods:number>
</mods:detail>
<mods:extent id="ACD8A13CE60C9BA5DC99423C5E822554" unit="page">
<mods:start id="EC000BAE2924BCCE411AB8401512B915">483</mods:start>
<mods:end id="3882067BC79BB20EC64D6F79BCD7AD8C">507</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B97587BFC2A8595D27FB91E17E6CD2D5">
<mods:url id="0CFA1851941D2D3F86254C627249A3A9">https://academic.oup.com/zoolinnean/article/190/2/483/5788722</mods:url>
</mods:location>
<mods:classification id="C2CD7A9F2AEC7F49AF01D272821CD6F4">journal article</mods:classification>
<mods:identifier id="B89C9EA7F31085D6F5C4981FA668F405" type="DOI">10.1093/zoolinnean/zlz160</mods:identifier>
<mods:identifier id="231E525493F672F2C800A3F683322EAA" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="A64303BE99BCBAD2FE68E46B43978490" type="Zenodo-Dep">14858057</mods:identifier>
</mods:mods>
<treatment id="921887F80D698F54FC90FB6A2049F237" ID-DOI="http://doi.org/10.5281/zenodo.14858061" ID-Zenodo-Dep="14858061" LSID="urn:lsid:plazi:treatment:921887F80D698F54FC90FB6A2049F237" httpUri="http://treatment.plazi.org/id/921887F80D698F54FC90FB6A2049F237" lastPageNumber="497" pageId="14" pageNumber="497">
<subSubSection id="52AB65650D698F54FC90FB6A27CEF184" pageId="14" pageNumber="497" type="nomenclature">
<paragraph id="1A0E36EE0D698F54FC90FB6A217AF1A4" blockId="14.[809,1313,1219,1275]" box="[809,1313,1219,1243]" pageId="14" pageNumber="497">
<heading id="414681820D698F54FC90FB6A217AF1A4" box="[809,1313,1219,1243]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="497" reason="2">
<taxonomicName id="DDB14D6D0D698F54FC90FB6A217AF1A4" ID-CoL="7S4RM" authority="(Gambetta, 1929)" baseAuthorityName="Gambetta" baseAuthorityYear="1929" box="[809,1313,1219,1243]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="497" phylum="Mollusca" rank="species" species="astropaliae">
Melanopsis astropaliae
<emphasis id="28C5EAFC0D698F54FBF4FB6A217AF1A4" box="[1101,1313,1219,1243]" italics="true" pageId="14" pageNumber="497">
(
<bibRefCitation id="7E204B1F0D698F54FBEFFB6A214CF1A5" author="Gambetta L" box="[1110,1303,1219,1243]" pageId="14" pageNumber="497" pagination="45 - 117" refId="ref12251" refString="Gambetta L. 1929. Molluschi. Archivio Zoologico Italiano 13: 45-117." type="journal article" year="1929">Gambetta, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="1A0E36EE0D698F54FC90FB4A27CEF184" blockId="14.[809,1313,1219,1275]" box="[809,917,1251,1275]" pageId="14" pageNumber="497">
<emphasis id="28C5EAFC0D698F54FC90FB4A27CEF184" box="[809,917,1251,1275]" italics="true" pageId="14" pageNumber="497">(clade B)</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="52AB65650D698F54FC90FAA52049F237" pageId="14" pageNumber="497" type="description">
<paragraph id="1A0E36EE0D698F54FC90FAA52049F237" blockId="14.[809,1425,1291,1865]" pageId="14" pageNumber="497">
<bibRefCitation id="7E204B1F0D698F54FC90FAA52003F05F" author="Gambetta L" box="[809,1112,1291,1313]" pageId="14" pageNumber="497" pagination="45 - 117" refId="ref12251" refString="Gambetta L. 1929. Molluschi. Archivio Zoologico Italiano 13: 45-117." type="journal article" year="1929">Gambetta (1929: 104)</bibRefCitation>
described
<taxonomicName id="DDB14D6D0D698F54FB40FAA52174F041" authority="Gambetta, 1929" authorityName="Gambetta" authorityYear="1929" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="497" phylum="Mollusca" rank="variety" species="praemorsa" variety="astropaliae">
<emphasis id="28C5EAFC0D698F54FB40FAA527FDF03E" italics="true" pageId="14" pageNumber="497">Melanopsis praemorsa</emphasis>
var.
<emphasis id="28C5EAFC0D698F54FC5AFA832033F041" box="[995,1128,1322,1343]" italics="true" pageId="14" pageNumber="497">astropaliae</emphasis>
<bibRefCitation id="7E204B1F0D698F54FBC8FA832174F041" author="Gambetta L" box="[1137,1327,1322,1344]" pageId="14" pageNumber="497" pagination="45 - 117" refId="ref12251" refString="Gambetta L. 1929. Molluschi. Archivio Zoologico Italiano 13: 45-117." type="journal article" year="1929">Gambetta, 1929</bibRefCitation>
</taxonomicName>
(for the original description see the Supporting Information, File S1) from Stampalia [Astypalaia Island], between Naxos and Samos, thus within the range of our clade II (Naxos, Samos, Tinos). He described the completely smooth shell, black and purplish without brick brown. Considering the intraspecies colour variability in
<taxonomicName id="DDB14D6D0D698F54FC90F9A827EAF368" authorityName="Ferussac in de Ferussac &amp; de Ferussac" authorityYear="1807" box="[809,945,1537,1558]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="497" phylum="Mollusca" rank="genus">
<emphasis id="28C5EAFC0D698F54FC90F9A827EAF368" box="[809,945,1537,1558]" italics="true" pageId="14" pageNumber="497">Melanopsis</emphasis>
</taxonomicName>
(Gambertta described it in detail), his description could be applied to our gastropods from Naxos, Samos and Tinos. The soft-parts coloration in the population from Naxos correspond with the one given by Gambetta for
<taxonomicName id="DDB14D6D0D698F54FBC7F9D2215CF3EE" baseAuthorityName="Gambetta" baseAuthorityYear="1929" box="[1150,1287,1659,1680]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="497" phylum="Mollusca" rank="species" species="astropaliae">
<emphasis id="28C5EAFC0D698F54FBC7F9D2215CF3EE" box="[1150,1287,1659,1680]" italics="true" pageId="14" pageNumber="497">astropaliae</emphasis>
</taxonomicName>
. Certainly, there is too much emphasis on the shell coloration in his description, considering the wide variability of this character, clearly visible in our photographs. This clade is molecularly distinct, and Gambettas description is the only one published for a
<taxonomicName id="DDB14D6D0D698F54FAB6F8BC21CBF254" authorityName="Ferussac in de Ferussac &amp; de Ferussac" authorityYear="1807" box="[1295,1424,1813,1834]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="497" phylum="Mollusca" rank="genus">
<emphasis id="28C5EAFC0D698F54FAB6F8BC21CBF254" box="[1295,1424,1813,1834]" italics="true" pageId="14" pageNumber="497">Melanopsis</emphasis>
</taxonomicName>
taxon in this region.
</paragraph>
</subSubSection>
</treatment>
</document>