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<document id="A1A81F3379A2CDAE6624680D7585ACB0" ID-DOI="10.37828/em.2023.69.2" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246916" ID-ZooBank="urn:lsid:zoobank.org:pub:91B1246B-9D09-454C-8CE8-7EB3CC1C221C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972238801" checkinUser="felipe" docAuthor="Volynkin, Anton V. &amp; Černý, Karel" docDate="2023" docId="6B3517136564FFC8AAE5FC812CCCFAAA" docLanguage="en" docName="EcolMontenegrina.69.10-16.pdf" docOrigin="Ecologica Montenegrina 69" docSource="http://dx.doi.org/10.37828/em.2023.69.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Siccia seima Volynkin &amp; Černý, 2023, sp. n." docType="treatment" docUuid="9880527B-0D26-4EA2-A844-FFD86678ECB8" docUuidSource="ZooBank" docVersion="2" lastPageNumber="15" masterDocId="970C6F6B6565FFCDAA72FFA52D49FFBC" masterDocTitle="Siccia seima, a new species from Cambodia and Thailand (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="16" masterPageNumber="10" pageNumber="11" updateTime="1722979741991" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EA6DF40886423A5B3740DFDAC7EC6E79">Siccia seima, a new species from Cambodia and Thailand (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="11B446CC40D4D056FFFDCF4B88A5654A">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="AA92F3615B63181C504D1933F6367508">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia.</mods:affiliation>
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<mods:namePart id="7F9C549B5347A5B055143D986210433A">Černý, Karel</mods:namePart>
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<mods:affiliation id="14E3B42D43C9D28D1D30D8383CB769CE">Tiergartenstrasse 27, A- 6020 Innsbruck, Austria. E-mail: natura. cerny @ aon. at; https: // orcid. org / 0000 - 0001 - 7207 - 2255 * Corresponding author. E-mail: monstruncusarctia @ gmail. com; https: // orcid. org / 0000 - 0001 - 9447 - 4925</mods:affiliation>
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<taxonomicName id="249CDD866564FFCCAAE5FC812C67FC82" ID-CoL="BXNBG" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[151,302,804,830]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. nov.">
<emphasis id="D1E87A176564FFCCAAE5FC812C67FC82" bold="true" box="[151,302,804,830]" italics="true" pageId="1" pageNumber="11">Siccia seima</emphasis>
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<paragraph id="E323A6056564FFCCAAE5FCC22C21FC3D" blockId="1.[151,1148,804,897]" box="[151,360,871,897]" pageId="1" pageNumber="11">
(
<figureCitation id="7BA7BA806564FFCCAAEDFCC22C40FC3D" box="[159,265,871,897]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 13</figureCitation>
,
<figureCitation id="7BA7BA806564FFCCAB64FCC22C7CFC3D" box="[278,309,871,897]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">11</figureCitation>
,
<figureCitation id="7BA7BA806564FFCCAB30FCC22C16FC3D" box="[322,351,871,897]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="1" pageNumber="11">15</figureCitation>
)
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<emphasis id="D1E87A176564FFCCAAE5FC0F2C1DFC78" bold="true" box="[151,340,938,964]" pageId="1" pageNumber="11">Type material</emphasis>
.
<materialsCitation id="53F4AC586564FFCCAB1CFC0F29A5FBB4" collectingDate="2006-01-27" collectingDateMax="2006-01-29" collectingDateMin="2006-01-27" collectionCode="ZSM" collectorName="G. Csorba &amp; G. Ronkay &amp; A. Volynkin" country="Cambodia" county="Seima Biodiversity Conservation" elevation="360" latitude="107.063614" location="Area" municipality="Seima" pageId="1" pageNumber="11" specimenCount="133" specimenCount-male="133" stateProvince="Mondolkiri" typeStatus="holotype">
<emphasis id="D1E87A176564FFCCAB1CFC0F2CA9FC78" bold="true" box="[366,480,938,964]" pageId="1" pageNumber="11">
<typeStatus id="3C2718A76564FFCCAB1CFC0F2CA9FC78" box="[366,480,938,964]" pageId="1" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="7BA7BA806564FFCCAB8EFC0F2F1AFC78" box="[508,595,938,964]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 1</figureCitation>
,
<figureCitation id="7BA7BA806564FFCCA81FFC0F2FC4FC78" box="[621,653,938,964]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">11</figureCitation>
): male, “
<collectingCountry id="9B8BE6956564FFCCA97FFC0F2EC2FC78" box="[781,907,938,964]" name="Cambodia" pageId="1" pageNumber="11">Cambodia</collectingCountry>
,
<collectingRegion id="215868E76564FFCCA9D1FC0F2937FC78" box="[931,1150,938,964]" country="Cambodia" name="Mondolkiri" pageId="1" pageNumber="11">Mondolkiri Prov.</collectingRegion>
, |
<collectingCounty id="0A42DE896564FFCCAEDCFC0F2C7DFC5A" pageId="1" pageNumber="11">
<collectingMunicipality id="03473C7F6564FFCCAEDCFC0F29B1FC78" box="[1198,1272,938,964]" pageId="1" pageNumber="11">Seima</collectingMunicipality>
Biodiversity Conservation
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|
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,
<locationDeviation id="F291BF276564FFCCABEBFC692FA4FC5A" box="[409,749,972,998]" firstLocation="Seima" pageId="1" pageNumber="11" secondLocation="O'Rang">between Seima and ORang</locationDeviation>
, | 12°15'44['']
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['']E,
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, |
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</date>
| leg.
<collectorName id="4E69C3D36564FFCCAAB8FC4B2C08FBB4" box="[202,321,1006,1032]" pageId="1" pageNumber="11">G. Csorba</collectorName>
&amp;
<collectorName id="4E69C3D36564FFCCAB14FC4B2CA1FBB4" box="[358,488,1006,1032]" pageId="1" pageNumber="11">G. Ronkay</collectorName>
” / “Slide |
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Arct. | 2019-
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|
<collectorName id="4E69C3D36564FFCCA9E7FC4B2962FBB4" box="[917,1067,1006,1032]" pageId="1" pageNumber="11">A. Volynkin</collectorName>
” (MWM/
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).
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<materialsCitation id="53F4AC586564FFCCAA87FBAA2847FB95" box="[245,1294,1039,1065]" collectionCode="ZSM" collectorName="No." country="Cambodia" pageId="1" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="3C2718A76564FFCCAA87FBAA2C3AFB95" box="[245,371,1039,1065]" pageId="1" pageNumber="11" type="paratype">Paratypes</typeStatus>
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.
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<collectingCountry id="9B8BE6956564FFCCABF0FBAA2F61FB95" box="[386,552,1039,1065]" name="Cambodia" pageId="1" pageNumber="11">CAMBODIA</collectingCountry>
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:
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, the same data as in the
<typeStatus id="3C2718A76564FFCCA9D4FBAA2946FB95" box="[934,1039,1039,1065]" pageId="1" pageNumber="11" type="holotype">holotype</typeStatus>
, gen. prep.
<collectorName id="4E69C3D36564FFCCAEE5FBAA298FFB95" box="[1175,1222,1039,1065]" pageId="1" pageNumber="11">No.</collectorName>
:
<collectionCode id="858D3EC06564FFCCAEA0FBAA2847FB95" box="[1234,1294,1039,1065]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="11" type="Museum">ZSM</collectionCode>
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Arct. 2019-
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<specimenCount id="F59A6D8C6564FFCCAAE5FB942D93FBF7" box="[151,218,1073,1099]" count="132" pageId="1" pageNumber="11" type="male">132♂</specimenCount>
(prepared by
<collectorName id="4E69C3D36564FFCCABF1FB942CBDFBF7" box="[387,500,1073,1099]" pageId="1" pageNumber="11">Volynkin</collectorName>
) (MWM/
<collectionCode id="858D3EC06564FFCCA818FB942FE1FBF7" box="[618,680,1073,1099]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="11" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
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:
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,
<collectingRegion id="215868E76564FFCCA997FB9429EDFBF7" box="[997,1188,1073,1099]" country="Thailand" name="Sakon Nakhon" pageId="1" pageNumber="11">Sakhon Nakhon</collectingRegion>
[
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<collectorName id="4E69C3D36564FFCCAF48FB942DF6FBD1" pageId="1" pageNumber="11">Phy Pan NP</collectorName>
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102°59.91''E,
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<collectorName id="4E69C3D36564FFCCA97FFBF62E35FBD1" box="[781,892,1107,1133]" pageId="1" pageNumber="11">K. Černý</collectorName>
leg., gen. prep.
<collectorName id="4E69C3D36564FFCCAE31FBF6293BFBD1" box="[1091,1138,1107,1133]" pageId="1" pageNumber="11">No.</collectorName>
: AV3970
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[Province],
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, road 1150, km 17 from
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27.
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.2011, K. Černý leg., gen. prep. No.: AV3969
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.
</paragraph>
</subSubSection>
<subSubSection id="AB86F58E6564FFCFAA87FB7F2FFAFE9E" lastPageId="2" lastPageNumber="12" pageId="1" pageNumber="11" type="diagnosis">
<paragraph id="E323A6056564FFCFAA87FB7F2FFAFE9E" blockId="1.[151,1437,938,2044]" lastBlockId="2.[151,1436,196,290]" lastPageId="2" lastPageNumber="12" pageId="1" pageNumber="11">
<emphasis id="D1E87A176564FFCCAA87FB7F2C27FB48" bold="true" box="[245,366,1242,1268]" pageId="1" pageNumber="11">Diagnosis</emphasis>
. The forewing length is 8.59.0 mm in males and 9.510.0 mm in females.
<taxonomicName id="249CDD866564FFCCAF74FB7F28D5FB48" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[1286,1436,1242,1268]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. nov.">
<emphasis id="D1E87A176564FFCCAF74FB7F28D5FB48" box="[1286,1436,1242,1268]" italics="true" pageId="1" pageNumber="11">Siccia seima</emphasis>
</taxonomicName>
<emphasis id="D1E87A176564FFCCAAE5FB592D92FAAA" bold="true" box="[151,219,1276,1302]" pageId="1" pageNumber="11">
<taxonomicNameLabel id="CADBC76C6564FFCCAAE5FB592D92FAAA" box="[151,219,1276,1302]" pageId="1" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7BA7BA806564FFCCAA99FB592C1CFAAA" box="[235,341,1276,1302]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 13</figureCitation>
) is externally reminiscent of
<taxonomicName id="249CDD866564FFCCA8C6FB592E83FAAA" authority="Butler, 1877" authorityName="Butler" authorityYear="1877" box="[692,970,1276,1302]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="D1E87A176564FFCCA8C6FB592E64FAAA" box="[692,813,1276,1302]" italics="true" pageId="1" pageNumber="11">S. sordida</emphasis>
Butler, 1877
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCA9A9FB59290CFAAA" box="[987,1093,1276,1302]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 46</figureCitation>
) and
<taxonomicName id="249CDD866564FFCCAEFBFB592D95FA8B" authority="(Daniel, 1951)" baseAuthorityName="Daniel" baseAuthorityYear="1951" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="likiangensis">
<emphasis id="D1E87A176564FFCCAEFBFB59287CFAAA" box="[1161,1333,1276,1302]" italics="true" pageId="1" pageNumber="11">S. likiangensis</emphasis>
(Daniel, 1951)
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCAA9EFAB82C2DFA8B" box="[236,356,1309,1335]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 910</figureCitation>
) and differs from the former species in the somewhat larger size, the broader forewing, the more pronounced transverse lines, the more elongate discal spot of the forewing, and the darker hindwing with a more diffuse discal spot. Compared to
<taxonomicName id="249CDD866564FFCCA8C2FAC42E10FAC7" baseAuthorityName="Daniel" baseAuthorityYear="1951" box="[688,857,1377,1403]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="likiangensis">
<emphasis id="D1E87A176564FFCCA8C2FAC42E10FAC7" box="[688,857,1377,1403]" italics="true" pageId="1" pageNumber="11">S. likiangensis</emphasis>
</taxonomicName>
, the new species is markedly smaller in size, and has shorter rami of the male antenna, intense blackish suffusion on the forewing in the antemedial and subterminal areas, and more intense brownish-grey suffusion on the hindwing postmedially. In the male genitalia,
<taxonomicName id="249CDD866564FFCCAB7CFA632C3AFA5C" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[270,371,1478,1504]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. n.">
<emphasis id="D1E87A176564FFCCAB7CFA632C3AFA5C" box="[270,371,1478,1504]" italics="true" pageId="1" pageNumber="11">S. seima</emphasis>
</taxonomicName>
<emphasis id="D1E87A176564FFCCAB0CFA632C8AFA5C" bold="true" box="[382,451,1478,1504]" pageId="1" pageNumber="11">
<taxonomicNameLabel id="CADBC76C6564FFCCAB0CFA632C8AFA5C" box="[382,451,1478,1504]" pageId="1" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7BA7BA806564FFCCABA4FA632F67FA5C" box="[470,558,1478,1504]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">Fig. 11</figureCitation>
) is distinguished from
<taxonomicName id="249CDD866564FFCCA939FA632E8CFA5C" authorityName="Butler" authorityYear="1877" box="[843,965,1478,1504]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="D1E87A176564FFCCA939FA632E8CFA5C" box="[843,965,1478,1504]" italics="true" pageId="1" pageNumber="11">S. sordida</emphasis>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCA9ABFA632978FA5C" box="[985,1073,1478,1504]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">Fig. 12</figureCitation>
) by the somewhat longer and broader phallus, and the longer and broader vesica bearing a longer area of scobination and a single long and robust cornutus whereas
<taxonomicName id="249CDD866564FFCCABA4F9AC2F1FF99F" authorityName="Butler" authorityYear="1877" box="[470,598,1545,1571]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="D1E87A176564FFCCABA4F9AC2F1FF99F" box="[470,598,1545,1571]" italics="true" pageId="1" pageNumber="11">S. sordida</emphasis>
</taxonomicName>
has a row of 24 considerably smaller cornuti. Compared to
<taxonomicName id="249CDD866564FFCCAFF4F9AC2C6AF9F9" baseAuthorityName="Daniel" baseAuthorityYear="1951" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="likiangensis">
<emphasis id="D1E87A176564FFCCAFF4F9AC2C6AF9F9" italics="true" pageId="1" pageNumber="11">S. likiangensis</emphasis>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCAB4AF98E2CDAF9F9" box="[312,403,1579,1605]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">Fig. 14</figureCitation>
), the male genitalia of
<taxonomicName id="249CDD866564FFCCA8CAF98E2E56F9F9" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[696,799,1579,1605]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. n.">
<emphasis id="D1E87A176564FFCCA8CAF98E2E56F9F9" box="[696,799,1579,1605]" italics="true" pageId="1" pageNumber="11">S. seima</emphasis>
</taxonomicName>
<emphasis id="D1E87A176564FFCCA958F98E2E38F9F9" bold="true" box="[810,881,1579,1605]" pageId="1" pageNumber="11">
<taxonomicNameLabel id="CADBC76C6564FFCCA958F98E2E38F9F9" box="[810,881,1579,1605]" pageId="1" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a somewhat longer and medially thicker uncus, a markedly shorter and less sclerotised ventral costal lobe, a shorter dorsal (apical) spur of the cucullus, a shorter and narrower distal saccular process, a considerably longer, broader and more medially curved phallus, and a substantially longer and broader vesica bearing a longer cluster of more robust scobination and a long and robust horn-shaped cornutus which is short thorn-shaped in the congener. The male genitalia structure of the new species is also similar to the externally dissimilar
<taxonomicName id="249CDD866564FFCCAEF2F97128D5F952" authority="Bucsek, 2020" authorityName="Bucsek" authorityYear="2020" box="[1152,1436,1748,1774]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="D1E87A176564FFCCAEF2F97129BCF952" box="[1152,1269,1748,1774]" italics="true" pageId="1" pageNumber="11">S. cinerea</emphasis>
<bibRefCitation id="870DDBF46564FFCCAE8FF97128D5F952" author="Bucsek, K." box="[1277,1436,1748,1774]" pageId="1" pageNumber="11" pagination="19 - 46" refId="ref2329" refString="Bucsek, K. (2020) Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 3. Entomofauna carpathica, 32 (2), 19 - 46. [http: // www. ses. entomology. sk / entomofaunacarpathica / pdf / volume 32 / 32 _ 2 _ 04 _ bucsek _ 2020 _ 1 _ ec. pdf]" type="journal article" year="2020">Bucsek, 2020</bibRefCitation>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCAAEDF9502C5AF8B3" box="[159,275,1781,1807]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="1" pageNumber="11">Figs 7, 8</figureCitation>
,
<figureCitation id="7BA7BA806564FFCCAB54F9502C0DF8B3" box="[294,324,1781,1807]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="1" pageNumber="11">13</figureCitation>
) but in
<taxonomicName id="249CDD866564FFCCABC2F9502F5EF8B3" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[432,535,1781,1807]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. n.">
<emphasis id="D1E87A176564FFCCABC2F9502F5EF8B3" box="[432,535,1781,1807]" italics="true" pageId="1" pageNumber="11">S. seima</emphasis>
</taxonomicName>
<emphasis id="D1E87A176564FFCCA856F9502F25F8B3" bold="true" box="[548,620,1781,1807]" pageId="1" pageNumber="11">
<taxonomicNameLabel id="CADBC76C6564FFCCA856F9502F25F8B3" box="[548,620,1781,1807]" pageId="1" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the uncus is shorter and medially thicker, the cucullus lacks the triangular dorsal process, the distal saccular process is shorter, the phallus is longer, broader and more curved, the vesica is longer, broader and bearing a longer cluster of more robust scobination and a markedly longer and thicker cornutus. The female genitalia of
<taxonomicName id="249CDD866564FFCCA95BF8FE2ED9F8C9" authorityName="Volynkin &amp; Černý" authorityYear="2023" box="[809,912,1883,1909]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="seima" status="sp. n.">
<emphasis id="D1E87A176564FFCCA95BF8FE2ED9F8C9" box="[809,912,1883,1909]" italics="true" pageId="1" pageNumber="11">S. seima</emphasis>
</taxonomicName>
<emphasis id="D1E87A176564FFCCA9EEF8FE2EADF8C9" bold="true" box="[924,996,1883,1909]" pageId="1" pageNumber="11">
<taxonomicNameLabel id="CADBC76C6564FFCCA9EEF8FE2EADF8C9" box="[924,996,1883,1909]" pageId="1" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7BA7BA806564FFCCA98AF8FE291BF8C9" box="[1016,1106,1883,1909]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="1" pageNumber="11">Fig. 15</figureCitation>
) are distinguished from
<taxonomicName id="249CDD866564FFCCAFF4F8FE2DB8F82B" authorityName="Butler" authorityYear="1877" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="D1E87A176564FFCCAFF4F8FE2DB8F82B" italics="true" pageId="1" pageNumber="11">S. sordida</emphasis>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCAB71F8D82C13F82B" box="[259,346,1917,1943]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="1" pageNumber="11">Fig. 16</figureCitation>
) and
<taxonomicName id="249CDD866564FFCCABD0F8D82F50F82B" authorityName="Bucsek" authorityYear="2020" box="[418,537,1917,1943]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="D1E87A176564FFCCABD0F8D82F50F82B" box="[418,537,1917,1943]" italics="true" pageId="1" pageNumber="11">S. cinerea</emphasis>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCA85EF8D82FCAF82B" box="[556,643,1917,1943]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="1" pageNumber="11">Fig. 17</figureCitation>
) by the broader ductus bursae, the posterior section of the corpus bursae having a heavily sclerotised concave lateral plate on the left side, the globular anterior section of the corpus bursae (it is elliptical in both the congeners), and the distally broader and granulose appendix bursae. Compared to
<taxonomicName id="249CDD866564FFCCAB4DF8472CA5F840" baseAuthorityName="Daniel" baseAuthorityYear="1951" box="[319,492,2018,2044]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="11" phylum="Arthropoda" rank="species" species="likiangensis">
<emphasis id="D1E87A176564FFCCAB4DF8472CA5F840" box="[319,492,2018,2044]" italics="true" pageId="1" pageNumber="11">S. likiangensis</emphasis>
</taxonomicName>
(
<figureCitation id="7BA7BA806564FFCCA872F8472F13F840" box="[512,602,2018,2044]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="1" pageNumber="11">Fig. 18</figureCitation>
), the female genitalia of the new species have a longer and broader ductus bursae, a well-developed and heavily sclerotised posterior section of the corpus bursae, a larger anterior section of the corpus bursae bearing a second ribbon-like signum ventrally, and a markedly longer, more helicoid and granulose appendix bursae.
</paragraph>
</subSubSection>
<caption id="B7E3F68D6567FFCFAAE5F80D2D94F840" ID-DOI="http://doi.org/10.5281/zenodo.13246920" ID-Zenodo-Dep="13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="2" pageNumber="12" startId="2.[151,237,1960,1983]" targetBox="[159,1424,302,1928]" targetPageId="2" targetType="figure">
<paragraph id="E323A6056567FFCFAAE5F80D2D94F840" blockId="2.[151,1437,1960,2044]" pageId="2" pageNumber="12">
<emphasis id="D1E87A176567FFCFAAE5F80D2C66F803" bold="true" box="[151,303,1960,1983]" pageId="2" pageNumber="12">Figures 110</emphasis>
.
<taxonomicName id="249CDD866567FFCFAB31F80D2C88F803" box="[323,449,1960,1983]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D1E87A176567FFCFAB31F80D2CCDF803" box="[323,388,1960,1983]" italics="true" pageId="2" pageNumber="12">Siccia</emphasis>
spp.
</taxonomicName>
: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after
<bibRefCitation id="870DDBF46567FFCFA99BF8622935F862" author="Bucsek, K." box="[1001,1148,1991,2014]" pageId="2" pageNumber="12" pagination="19 - 46" refId="ref2329" refString="Bucsek, K. (2020) Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 3. Entomofauna carpathica, 32 (2), 19 - 46. [http: // www. ses. entomology. sk / entomofaunacarpathica / pdf / volume 32 / 32 _ 2 _ 04 _ bucsek _ 2020 _ 1 _ ec. pdf]" type="journal article" year="2020">Bucsek 2020</bibRefCitation>
); 8 in CKB (after
<bibRefCitation id="870DDBF46567FFCFAF3CF8622D98F840" author="Bucsek, K." pageId="2" pageNumber="12" pagination="19 - 46" refId="ref2329" refString="Bucsek, K. (2020) Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 3. Entomofauna carpathica, 32 (2), 19 - 46. [http: // www. ses. entomology. sk / entomofaunacarpathica / pdf / volume 32 / 32 _ 2 _ 04 _ bucsek _ 2020 _ 1 _ ec. pdf]" type="journal article" year="2020">Bucsek 2020</bibRefCitation>
).
</paragraph>
</caption>
<caption id="B7E3F68D6566FFCEAAE5F83E2C87F86C" ID-DOI="http://doi.org/10.5281/zenodo.13246922" ID-Zenodo-Dep="13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="3" pageNumber="13" startId="3.[151,237,1947,1970]" targetBox="[162,1436,189,1910]" targetPageId="3" targetType="figure">
<paragraph id="E323A6056566FFCEAAE5F83E2C87F86C" blockId="3.[151,1436,1947,2000]" pageId="3" pageNumber="13">
<emphasis id="D1E87A176566FFCEAAE5F83E2C71F80E" bold="true" box="[151,312,1947,1970]" pageId="3" pageNumber="13">Figures 1114</emphasis>
.
<taxonomicName id="249CDD866566FFCEAB35F83E2C89F80E" box="[327,448,1947,1970]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D1E87A176566FFCEAB35F83E2CC1F80E" box="[327,392,1947,1970]" italics="true" pageId="3" pageNumber="13">Siccia</emphasis>
spp.
</taxonomicName>
: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after
<bibRefCitation id="870DDBF46566FFCEAB41F81C2C8BF86C" author="Bucsek, K." box="[307,450,1977,2000]" pageId="3" pageNumber="13" pagination="19 - 46" refId="ref2329" refString="Bucsek, K. (2020) Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 3. Entomofauna carpathica, 32 (2), 19 - 46. [http: // www. ses. entomology. sk / entomofaunacarpathica / pdf / volume 32 / 32 _ 2 _ 04 _ bucsek _ 2020 _ 1 _ ec. pdf]" type="journal article" year="2020">Bucsek 2020</bibRefCitation>
).
</paragraph>
</caption>
<caption id="B7E3F68D6561FFC9AAE5F83E2C6CF86C" ID-DOI="http://doi.org/10.5281/zenodo.13246926" ID-Zenodo-Dep="13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="4" pageNumber="14" startId="4.[151,237,1947,1970]" targetBox="[234,1402,190,1910]" targetPageId="4" targetType="figure">
<paragraph id="E323A6056561FFC9AAE5F83E2C6CF86C" blockId="4.[151,1436,1947,2000]" pageId="4" pageNumber="14">
<emphasis id="D1E87A176561FFC9AAE5F83E2C76F80E" bold="true" box="[151,319,1947,1970]" pageId="4" pageNumber="14">Figures 1518</emphasis>
.
<taxonomicName id="249CDD866561FFC9AB24F83E2C9FF80E" box="[342,470,1947,1970]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D1E87A176561FFC9AB24F83E2CDEF80E" box="[342,407,1947,1970]" italics="true" pageId="4" pageNumber="14">Siccia</emphasis>
spp.
</taxonomicName>
: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM.
</paragraph>
</caption>
<subSubSection id="AB86F58E6560FFC8AA87FF6129D6FB33" pageId="5" pageNumber="15" type="description">
<paragraph id="E323A6056560FFC8AA87FF6129D6FB33" blockId="5.[151,1437,196,1302]" pageId="5" pageNumber="15">
<emphasis id="D1E87A176560FFC8AA87FF612CCFFF62" bold="true" box="[245,390,196,222]" pageId="5" pageNumber="15">Description</emphasis>
.
<emphasis id="D1E87A176560FFC8ABEFFF612E7AFF62" bold="true" box="[413,819,196,222]" pageId="5" pageNumber="15">External morphology of adults</emphasis>
(
<figureCitation id="7BA7BA806560FFC8A938FF612EF5FF62" box="[842,956,196,222]" captionStart="Figures 110" captionStartId="2.[151,237,1960,1983]" captionTargetBox="[159,1424,302,1928]" captionTargetId="figure-47@2.[159,1427,302,1941]" captionTargetPageId="2" captionText="Figures 110. Siccia spp.: adults. Depositories of the specimens: 1, 9 and 10 in MWM/ZSM; 2, 3, 5 and 6 in MWM/ZSM; 4 in NHMUK (©The Trustees of NHMUK); 7 in SNM (after Bucsek 2020); 8 in CKB (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246920" httpUri="https://zenodo.org/record/13246920/files/figure.png" pageId="5" pageNumber="15">Figs 13</figureCitation>
). Head off-white, suffused with black scales. Antenna pale brown, shortly bipectinate in male and filiform in female. Thorax off-white, suffused with black scales. Forewing ground colour of-white with black irroration. Pattern elements slate black, diffuse. Subbasal line interrupted into two spots: larger one on costa and smaller one medially. Antemedial area intensely suffused with slate black between vein R and anal margin. Antemedial line sinuous, dentate medially, dilated into trapezoidal spots on wing margins. Cell with medial round spot between antemedial and medial lines, and reniform discal spot. Medial line irregularly sinuous and dentate, strongly bent outward medially and strongly dilated into trapezoidal spot on costa. Postmedial line irregularly dentate, strongly bent outward medially, strongly dilated into trapezoidal spot on costa, and almost touching medial line at anal margin with medial area intensely suffused these lines. Subterminal line parallel postmedial one, diffuse, strongly dilated into large trapezoidal subapical spot on costa. Terminal line interrupted into spots between veins. Forewing cilia off-white with diffuse blackish spot medially. Hindwing and its cilia pale brownish-grey; discal spot semilunar, indistinct. Abdomen brownish-grey, male valvae covered with hair-like pale ochreous scales; female corethrogyne also pale ochreous.
<emphasis id="D1E87A176560FFC8A932FDDE2EBAFD29" bold="true" box="[832,1011,635,661]" pageId="5" pageNumber="15">Male genitalia</emphasis>
(
<figureCitation id="7BA7BA806560FFC8AE71FDDE2910FD29" box="[1027,1113,635,661]" captionStart="Figures 1114" captionStartId="3.[151,237,1947,1970]" captionTargetBox="[162,1436,189,1910]" captionTargetId="figure-10@3.[151,1436,189,1928]" captionTargetPageId="3" captionText="Figures 1114. Siccia spp.: male genitalia. Depositories of the specimens dissected: 11, 12 and 14 in MWM/ZSM; 13 in SNM (after Bucsek 2020)." figureDoi="http://doi.org/10.5281/zenodo.13246922" httpUri="https://zenodo.org/record/13246922/files/figure.png" pageId="5" pageNumber="15">Fig. 11</figureCitation>
). Uncus cylindrical, almost straight, medially slightly dilated, distally tapered and with downcurved claw-shaped tip. Arms of tegumen strongly dilated posteriorly and fused in posterior third. Vinculum moreo к less equal in length to tegumen, with heavily sclerotised U-shaped saccus. Valva lobular, medially dilated and distally tapered. Costa with rounded lobe-like proximal ventral lobe. Cucullus with two thorn-like processes apically and ventrally, and short dorsal protrusion subapically. Sacculus narrow with short triangular distal process directed dorsally. Transtilla ribbon-like. Juxta weakly sclerotised, trapezoidal. Phallus gradually upcurved, with very short coecum. Vesica broad sack-like, with outer half covered with robust scobination and short lateral diverticulum basally bearing long and robust thorn-like cornutus. Basal plate of vesica ejaculatorius long, ribbon-like.
<emphasis id="D1E87A176560FFC8AB5CFC0F2F4BFC78" bold="true" box="[302,514,938,964]" pageId="5" pageNumber="15">Female genitalia</emphasis>
(
<figureCitation id="7BA7BA806560FFC8A86AFC0F2F3CFC78" box="[536,629,938,964]" captionStart="Figures 1518" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[234,1402,190,1910]" captionTargetId="figure-10@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1518. Siccia spp.: female genitalia. Depositories of the specimens dissected: 15 in CKC; 1618 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13246926" httpUri="https://zenodo.org/record/13246926/files/figure.png" pageId="5" pageNumber="15">Fig. 15</figureCitation>
). Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses long and thin, apophysis posterioris ca. twice longer than apophysis anterioris. Ductus bursae sclerotised, dorso-ventrally flattened, funnel-like dilated posteriorly. Posterior section of corpus bursae weakly sclerotised, with heavily sclerotised concave lateral plate on left side, and elongate cluster of minute spinules on right side at base of appendix bursae. Anterior section of corpus bursae globular, membranous, with rugose ribbon-like signum ventrally and elliptical serrulate signum dorsally. Appendix bursae tubular, helicoid, covered with granuli, situated postero-laterally on right side, directed laterally.
</paragraph>
</subSubSection>
<subSubSection id="AB86F58E6560FFC8AA87FB332C55FB6E" pageId="5" pageNumber="15" type="distribution">
<paragraph id="E323A6056560FFC8AA87FB332C55FB6E" blockId="5.[151,1437,196,1302]" pageId="5" pageNumber="15">
<emphasis id="D1E87A176560FFC8AA87FB332CC6FB0C" bold="true" box="[245,399,1174,1200]" pageId="5" pageNumber="15">Distribution</emphasis>
.
<collectingCountry id="9B8BE6956560FFC8ABD7FB332F54FB0C" box="[421,541,1174,1200]" name="Cambodia" pageId="5" pageNumber="15">Cambodia</collectingCountry>
(
<collectingRegion id="215868E76560FFC8A846FB332E7CFB0C" box="[564,821,1174,1200]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="15">Mondolkiri Province</collectingRegion>
) and
<collectingCountry id="9B8BE6956560FFC8A9F6FB332EA4FB0C" box="[900,1005,1174,1200]" name="Thailand" pageId="5" pageNumber="15">Thailand</collectingCountry>
(
<collectingRegion id="215868E76560FFC8AE71FB332980FB0C" box="[1027,1225,1174,1200]" country="Thailand" name="Sakon Nakhon" pageId="5" pageNumber="15">Sakhon Nakhon</collectingRegion>
and
<collectingRegion id="215868E76560FFC8AF62FB3328D4FB0C" box="[1296,1437,1174,1200]" country="Thailand" name="Chiang Rai" pageId="5" pageNumber="15">Chiang Rai</collectingRegion>
Provinces).
</paragraph>
</subSubSection>
<subSubSection id="AB86F58E6560FFC8AA87FB7F2CCCFAAA" pageId="5" pageNumber="15" type="etymology">
<paragraph id="E323A6056560FFC8AA87FB7F2CCCFAAA" blockId="5.[151,1437,196,1302]" pageId="5" pageNumber="15">
<emphasis id="D1E87A176560FFC8AA87FB7F2C34FB48" bold="true" box="[245,381,1242,1268]" pageId="5" pageNumber="15">Etymology</emphasis>
. The specific epithet is homonymic of the
<typeStatus id="3C2718A76560FFC8A907FB7F2EE1FB48" box="[885,936,1242,1268]" pageId="5" pageNumber="15">type</typeStatus>
locality. The name is a noun in apposition to the generic name.
</paragraph>
</subSubSection>
</treatment>
</document>