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<mods:title id="CE8E9F46B648AFFCFCF51F7217795AD8">Systematics of the colour-polymorphic spider genus Cybaeolus, with comments on the phylogeny of the family Hahniidae (Araneae)</mods:title>
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<mods:namePart id="AA7CB8BDE2287E3BC9C57166D05E6218">Burgo, Agustina</mods:namePart>
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<mods:affiliation id="1F21BBA3171ECF3AC6D9A9188881B8D0">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="C90DEE6E7F6E84B7E91AC94F864C44DA">Catley, Kefyn</mods:namePart>
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<mods:affiliation id="A0F60779C8A1023EB24FBE79F84A508B">Western Carolina University, Cullowhee, NC 28723, United States</mods:affiliation>
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<mods:namePart id="BF2B373D5380569A6EF0ABE7394A89D2">Grismado, Cristian J.</mods:namePart>
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<mods:affiliation id="83C3893AF0AF84F9B81C0C9FB5145CA0">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="BAD023EF14CADDA5958D95C77150A012">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="1EE0EE6E99BE10A7D208BC74F51FA7DB">Museum of Nature Hamburg, Zoology, Leibniz-Institute for the Analysis of Biodiversity Change (LIB), Center for Taxonomy and Morphology, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="97568A3A22D5CE6293F411D7EA4F9890">Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation id="1D65C409CB270ABABCF2FEB44A588E68">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart id="DA1AE45A1E36808D48545B215495AF50">Hormiga, Gustavo</mods:namePart>
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<mods:affiliation id="DFEF39A2C0146508EACCD2C3698FCAA2">Department of Biology, The George Washington University, Washington, DC 20052, United States</mods:affiliation>
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<mods:namePart id="3FA87354FCDB359574B16D7A8597D690">Griswold, Charles</mods:namePart>
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<mods:affiliation id="57E1405B48A3B9523290AC34C7025E4A">California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, United States</mods:affiliation>
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<mods:namePart id="7A14E23E778D46ABDEE7BA0CCBAFB3FE">Martínez, Leonel</mods:namePart>
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<mods:affiliation id="87E169DC7407A596CB4EE1B0E8689989">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="F8850A6E299105AC34DC6CAB1BC2FB1A">Ramírez, Martín J.</mods:namePart>
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<mods:affiliation id="DD813DA383F870494AFAF6955DA5C19E">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:nameIdentifier id="3DFC4F62844883946D7D89E97D5F0506" type="email">ramirez@macn.gov.ar</mods:nameIdentifier>
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<mods:date id="57BF5966CB38B2782FB8831DD985885D">2025</mods:date>
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<mods:number id="8E84592935F0D2AC75E59EFFD7D35A3F">2025-01-10</mods:number>
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<taxonomicName id="7CC74D002763FFA9963EDDF6FDF36DAC" ID-CoL="625HC" authority="Bertkau, 1878" authorityName="Bertkau" authorityYear="1878" box="[304,571,1688,1715]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
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<emphasis id="89B3EA912763FFA9963EDDF6FDF36DAC" bold="true" box="[304,571,1688,1715]" pageId="11" pageNumber="12">
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Hahniidae
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<bibRefCitation id="DF564B722763FFA996AADDF7FDF36DAC" author="Bertkau P" box="[420,571,1688,1715]" pageId="11" pageNumber="12" pagination="351 - 410" refId="ref32954" refString="Bertkau P. Versuch einer naturlichen Anordnung der Spinnen, nebst Bemerkungen zu einzelnen Gattungen. Archiv fur Naturgeschichte 1878; 44: 351 - 410." type="journal article" year="1878">Bertkau, 1878</bibRefCitation>
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</emphasis>
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Hahniidae
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<bibRefCitation id="DF564B722763FFA997E8DDD1FE646DC9" author="Bertkau P" box="[230,428,1727,1751]" pageId="11" pageNumber="12" pagination="351 - 410" refId="ref32954" refString="Bertkau P. Versuch einer naturlichen Anordnung der Spinnen, nebst Bemerkungen zu einzelnen Gattungen. Archiv fur Naturgeschichte 1878; 44: 351 - 410." type="journal article" year="1878">Bertkau 1878: 393</bibRefCitation>
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</taxonomicName>
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(
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genus
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<taxonomicName id="7CC74D002763FFA99532DDD1FF6D6DE9" authority="C. L. Koch, 1841" authorityName="C. L. Koch" authorityYear="1841" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="89B3EA912763FFA99532DDD1FD4F6DC9" box="[572,647,1727,1751]" italics="true" pageId="11" pageNumber="12">Hahnia</emphasis>
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C.L.
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<bibRefCitation id="DF564B722763FFA995B7DDD1FF6D6DE9" author="Koch CL" pageId="11" pageNumber="12" pagination="41 - 131" refId="ref33849" refString="Koch CL. Die Arachniden. Nurnberg Achter Band: C. H. Zeh'sche Buchhandlung, 1841, 41 - 131." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
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</taxonomicName>
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).
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</paragraph>
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<subSubSection id="F3DD65082763FFA99689DC79FC666CAC" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="BB7836832763FFA99689DC79FE2C6C2E" blockId="11.[113,763,1815,1970]" box="[391,484,1815,1840]" pageId="11" pageNumber="12">
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<emphasis id="89B3EA912763FFA99689DC79FE2C6C2E" box="[391,484,1815,1840]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="BB7836832763FFA9977FDC52FC666CAC" blockId="11.[113,763,1815,1970]" lastBlockId="11.[809,1461,1476,1970]" pageId="11" pageNumber="12">
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Members of
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<taxonomicName id="7CC74D002763FFA997F2DC52FEAF6C4A" authorityName="Bertkau" authorityYear="1878" box="[252,359,1852,1876]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Hahniidae</taxonomicName>
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resemble other entelegyne spiders of the RTA clade by having entelegyne genitalia and a retrolateral tibial apophysis in the male palp, but (except for
|
||||
<taxonomicName id="7CC74D002763FFA99582DC15FD276C8D" authorityName="Simon" authorityYear="1884" box="[652,751,1915,1939]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="89B3EA912763FFA99582DC15FD276C8D" box="[652,751,1915,1939]" italics="true" pageId="11" pageNumber="12">Cybaeolus</emphasis>
|
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</taxonomicName>
|
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) differ by having the anterior lateral spinnerets well separated, with the posterior medians advanced in-between them, and the posterior laterals well separated, behind the anterior laterals or, most usually, at their sides, so that all the spinnerets are arranged in a row (
|
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<figureCitation id="23FC2A062763FFA99486DD4CFC236D25" box="[904,1003,1570,1595]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="12.[129,194,1173,1197]" captionStartId-1="13.[114,179,1167,1191]" captionStartId-2="14.[129,194,1128,1152]" captionStartId-3="15.[113,178,1162,1186]" captionTargetBox-0="[161,1441,144,1145]" captionTargetBox-1="[146,1426,144,1140]" captionTargetBox-2="[161,1441,144,1101]" captionTargetBox-3="[146,1426,144,1135]" captionTargetId-0="figure-343@12.[161,1441,144,1145]" captionTargetId-1="figure-448@13.[146,1426,144,1140]" captionTargetId-2="figure-422@14.[161,1441,144,1101]" captionTargetId-3="figure-415@15.[146,1426,144,1135]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="Figure 7. Scanning electron micrographs of spinnerets of Hahnia pusilla (C.L.Koch, 1841), female. A, overview.B, ALS left.C, PMS left.D, PMS right. E, PLS left.F, PLS right.Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-1="Figure 8. Scanning electron micrographs of spinnerets of Hahnia pusilla (C.L.Koch, 1841), male. A, overview. B, ALS left. C, PMS left.D. PLS left. Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Tp, tartipore." captionText-2="Figure 9. Scanning electron micrographs of spinnerets of Neoantistea magna (Keyserling, 1887), female.A, overview.B, ALS right.C, PMS right. D, PLS right. E, same, detail.Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, Piriform gland spigot; Tp, tartipore." captionText-3="Figure 10. Scanning electron micrographs of spinnerets of Neoantistea magna (Keyserling, 1887), male.A, overview. B, ALS left. C, PMS right. D, PLS left.Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Tp, tartipore." pageId="11" pageNumber="12">Figs 7–10</figureCitation>
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||||
). Additionally, they, including
|
||||
<taxonomicName id="7CC74D002763FFA9922CDD4CFA4D6D24" authorityName="Simon" authorityYear="1884" box="[1314,1413,1570,1594]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="89B3EA912763FFA9922CDD4CFA4D6D24" box="[1314,1413,1570,1594]" italics="true" pageId="11" pageNumber="12">Cybaeolus</emphasis>
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</taxonomicName>
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||||
, can be recognized by the male palp with retrolateral tibial apophysis that has a canal, usually curved upwards and with an acute tip bearing regular teeth, a retrolateral patellar apophysis (absent in
|
||||
<taxonomicName id="7CC74D002763FFA99446DDCEFAE26DA6" authority="Schiapelli and Gerschman, 1958" authorityName="Schiapelli and Gerschman" authorityYear="1958" box="[840,1322,1696,1720]" class="Arachnida" family="Hahniidae" genus="Amaloxenops" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912763FFA99446DDCEFC036DA6" box="[840,971,1696,1720]" italics="true" pageId="11" pageNumber="12">Amaloxenops</emphasis>
|
||||
<bibRefCitation id="DF564B722763FFA994DADDCEFAE26DA6" author="Schiapelli RD & Gerschman BS" box="[980,1322,1696,1720]" pageId="11" pageNumber="12" pagination="187 - 231" refId="ref34874" refString="Schiapelli RD, Gerschman BS. Aranas argentinas III. Aranas de Misiones. Revista del Museo Argentino de Ciencias Naturales Bernardino Rivadavia 1958; 3: 187 - 231." type="journal article" year="1958">Schiapelli and Gerschman, 1958</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="7CC74D002763FFA9926DDDCEFC796DC9" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" class="Arachnida" family="Hahniidae" genus="Antistea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912763FFA9926DDDCEFA7B6DA9" box="[1379,1459,1696,1719]" italics="true" pageId="11" pageNumber="12">Antistea</emphasis>
|
||||
<bibRefCitation id="DF564B722763FFA99424DDD1FC796DC9" author="Simon E" box="[810,945,1727,1751]" pageId="11" pageNumber="12" pagination="193 - 380" refId="ref35030" refString="Simon E. Histoire Naturelle des Araignees. Deuxieme Edition, Tome Second. Paris: Roret, 1898, 193 - 380." type="book chapter" year="1898">Simon, 1898</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), and the simple copulatory bulb with the thin embolus circling clockwise (left palp, ventral view), without conductor, and sometimes with a lightly sclerotized median apophysis as a simple lobe (
|
||||
<figureCitation id="23FC2A062763FFA99349DC73FB596C2B" box="[1095,1169,1821,1845]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="11" pageNumber="12">Figs 15</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062763FFA99392DC73FB7E6C2B" box="[1180,1206,1821,1845]" captionStart="Figure 16" captionStartId="21.[115,180,1623,1647]" captionTargetBox="[146,1426,144,1595]" captionTargetId="figure-119@21.[146,1426,144,1595]" captionTargetPageId="21" captionText="Figure 16.Scanning electron micrographs of left male palps of Hahnia pusilla (C.L.Koch, 1841) (A–D) and Neoantistea magna (Keyserling, 1887) (E–H). A, retrolateral view. B, dorsal view.C, ventral view.D, tibia and patella in retrolateral view. E, ventral view.F, dorsal view.G, detail of tibial retroleral apophysis in ventral view. H, same, detail in dorsal view. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, patellar retrolateral apophysis of male palp; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="11" pageNumber="12">16</figureCitation>
|
||||
).
|
||||
<taxonomicName id="7CC74D002763FFA993C3DC73FAF86C2B" authorityName="Simon" authorityYear="1884" box="[1229,1328,1821,1845]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912763FFA993C3DC73FAF86C2B" box="[1229,1328,1821,1845]" italics="true" pageId="11" pageNumber="12">Cybaeolus</emphasis>
|
||||
</taxonomicName>
|
||||
has grouped spinnerets (
|
||||
<figureCitation id="23FC2A062763FFA994ADDC52FC306C4A" box="[931,1016,1852,1877]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="8.[129,194,1268,1292]" captionStartId-1="9.[113,178,1508,1532]" captionStartId-2="10.[129,194,1249,1273]" captionStartId-3="11.[113,178,1348,1372]" captionTargetBox-0="[161,1441,144,1241]" captionTargetBox-1="[146,1426,144,1480]" captionTargetBox-2="[161,1441,144,1221]" captionTargetBox-3="[146,1426,144,1321]" captionTargetId-0="figure-387@8.[161,1441,144,1241]" captionTargetId-1="figure-265@9.[146,1426,144,1480]" captionTargetId-2="figure-436@10.[161,1441,144,1221]" captionTargetId-3="figure-255@11.[146,1426,144,1321]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 3. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, female. A, overview.B, ALS left. C, overview of PMS. D, PLS left (several Ac spigot shafts broken).Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-1="Figure 4. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, male.A, overview. B, PMS left.C, ALS left. D, PLS left. E, epiandrum.Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-2="Figure 5. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), female. A, overview.B, ALS right.C, PMS left. D, PLS right. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Pi, piriform gland spigot; TS, tracheal spiracle." captionText-3="Figure 6. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), male. A, overview.B, ALS left. C, overview of PMS. D, PLS right.E, epiandrum. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot." pageId="11" pageNumber="12">Figs 3–6</figureCitation>
|
||||
), but the male genitalia have the mentioned diagnostic characters;
|
||||
<taxonomicName id="7CC74D002763FFA99302DC32FB476C6A" authorityName="Schiapelli and Gerschman" authorityYear="1958" box="[1036,1167,1884,1908]" class="Arachnida" family="Hahniidae" genus="Amaloxenops" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912763FFA99302DC32FB476C6A" box="[1036,1167,1884,1908]" italics="true" pageId="11" pageNumber="12">Amaloxenops</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="7CC74D002763FFA993B1DC32FAC76C6D" authorityName="Simon" authorityYear="1898" box="[1215,1295,1884,1907]" class="Arachnida" family="Hahniidae" genus="Antistea" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912763FFA993B1DC32FAC76C6D" box="[1215,1295,1884,1907]" italics="true" pageId="11" pageNumber="12">Antistea</emphasis>
|
||||
</taxonomicName>
|
||||
lack the patellar apophysis, but the tibial apophysis and the spinnerets are typical of the family.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="EFB8660B2764FFAE978FDFFBFA8E6FFB" pageId="12" pageNumber="13" startId="12.[129,194,1173,1197]" targetBox="[161,1441,144,1145]" targetPageId="12" targetType="figure">
|
||||
<paragraph id="BB7836832764FFAE978FDFFBFA8E6FFB" blockId="12.[129,1435,1173,1254]" pageId="12" pageNumber="13">
|
||||
<emphasis id="89B3EA912764FFAE978FDFFBFF106FB3" bold="true" box="[129,216,1173,1197]" pageId="12" pageNumber="13">Figure 7.</emphasis>
|
||||
Scanning electron micrographs of spinnerets of
|
||||
<taxonomicName id="7CC74D002764FFAE9593DFFBFC1B6FB3" authority="(C. L. Koch, 1841)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1841" box="[669,979,1173,1197]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pusilla">
|
||||
<emphasis id="89B3EA912764FFAE9593DFFBFCEB6FB3" box="[669,803,1173,1197]" italics="true" pageId="12" pageNumber="13">Hahnia pusilla</emphasis>
|
||||
(C.L.
|
||||
<bibRefCitation id="DF564B722764FFAE9458DFFBFC046FB3" author="Koch CL" box="[854,972,1173,1197]" pageId="12" pageNumber="13" pagination="41 - 131" refId="ref33849" refString="Koch CL. Die Arachniden. Nurnberg Achter Band: C. H. Zeh'sche Buchhandlung, 1841, 41 - 131." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female. A, overview. B, ALS left. C, PMS left. D, PMS right. E, PLS left. F, PLS right. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="F3DD65082764FFAE96A0DE78FD876DB2" pageId="12" pageNumber="13" type="discussion">
|
||||
<paragraph id="BB7836832764FFAE96A0DE78FE146E31" blockId="12.[128,778,1302,1708]" box="[430,476,1302,1327]" pageId="12" pageNumber="13">
|
||||
<emphasis id="89B3EA912764FFAE96A0DE78FE146E31" box="[430,476,1302,1327]" italics="true" pageId="12" pageNumber="13">Note</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832764FFAE978FDE55FD876DB2" blockId="12.[128,778,1302,1708]" pageId="12" pageNumber="13">
|
||||
The family
|
||||
<taxonomicName id="7CC74D002764FFAE97F2DE55FEAF6E4D" authorityName="Bertkau" authorityYear="1878" box="[252,359,1339,1363]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Hahniidae</taxonomicName>
|
||||
traditionally listed only representatives with spinnerets in the characteristic arrangement of a transverse line, until
|
||||
<bibRefCitation id="DF564B722764FFAE97EADE14FE586E8C" author="Lehtinen PT" box="[228,400,1402,1426]" pageId="12" pageNumber="13" pagination="199 - 468" refId="ref33957" refString="Lehtinen PT. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 1967; 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
|
||||
expanded the family limits to include
|
||||
<taxonomicName id="7CC74D002764FFAE978EDEF7FF2B6EAF" authorityName="Simon" authorityYear="1884" box="[128,227,1433,1457]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE978EDEF7FF2B6EAF" box="[128,227,1433,1457]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
||||
</taxonomicName>
|
||||
, with grouped spinnerets, plus several genera now placed in
|
||||
<taxonomicName id="7CC74D002764FFAE97EADED7FE986ECF" box="[228,336,1465,1489]" class="Arachnida" family="Cybaeidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cybaeidae</taxonomicName>
|
||||
and Cicurinidae (see:
|
||||
<bibRefCitation id="DF564B722764FFAE9533DED7FCCC6ECF" author="Wheeler WC & Coddington JA & Crowley LM" box="[573,772,1465,1489]" pageId="12" pageNumber="13" pagination="574 - 616" refId="ref35325" refString="Wheeler WC, Coddington JA, Crowley LM et al. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 2017; 33: 574 - 616." type="journal article" year="2017">
|
||||
Wheeler
|
||||
<emphasis id="89B3EA912764FFAE9594DED4FD026ECF" box="[666,714,1465,1489]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="DF564B722764FFAE978FDEB6FE936EEE" author="Castellucci F & Scharff N & Luchetti A" box="[129,347,1496,1520]" pageId="12" pageNumber="13" pagination="107833 - 9" refId="ref32981" refString="Castellucci F, Scharff N, Luchetti A. Molecular systematics and phylogenetics of the spider genus Mastigusa Menge, 1854 (Araneae, Cybaeidae). Molecular Phylogenetics and Evolution 2023; 186: 107833 - 9. https: // doi. org / 10.1016 / j. ympev. 2023.107833" type="journal article" year="2023">
|
||||
Castellucci
|
||||
<emphasis id="89B3EA912764FFAE97FBDEB7FEEB6EEE" box="[245,291,1496,1520]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="DF564B722764FFAE966BDEB6FDE26EEE" author="Gorneau JA & Crews SC & Cala-Riquelme F" box="[357,554,1496,1520]" pageId="12" pageNumber="13" pagination="1 - 18" refId="ref33475" refString="Gorneau JA, Crews SC, Cala-Riquelme F et al. Webs of intrigue: museum genomics elucidate relationships of the marronoid spider clade (Araneae). Insect Systematics and Diversity 2023; 7: 1 - 18." type="journal article" year="2023">
|
||||
Gorneau
|
||||
<emphasis id="89B3EA912764FFAE96CADEB7FE3A6EEE" box="[452,498,1496,1520]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
). Some hahniids have the posterior lateral spinnerets behind the anterior laterals (e.g.
|
||||
<taxonomicName id="7CC74D002764FFAE978CDD79FE8F6D31" authority="Roth, 1967" authorityName="Roth" authorityYear="1967" box="[130,327,1559,1583]" class="Arachnida" family="Hahniidae" genus="Lizarba" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE978CDD79FF186D31" box="[130,208,1559,1583]" italics="true" pageId="12" pageNumber="13">Lizarba</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE97DADD79FE8F6D31" author="Roth VD" box="[212,327,1559,1583]" pageId="12" pageNumber="13" pagination="297 - 345" refId="ref34792" refString="Roth VD. A review of the South American spiders of the family Agelenidae (Arachnida, Araneae). Bulletin of the American Museum of Natural History 1967; 134: 297 - 345." type="journal article" year="1967">Roth, 1967</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="89B3EA912764FFAE965CDD76FE646D31" box="[338,428,1560,1583]" italics="true" pageId="12" pageNumber="13">Paramito</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE96BCDD79FD7A6D31" author="Duperre N & Tapia E" box="[434,690,1559,1583]" pageId="12" pageNumber="13" pagination="53 - 111" refId="ref33170" refString="Duperre N, Tapia E. First record of the family Hahniidae in Ecuador with description of thirteen new species and three new genera (Araneae: Hahniidae). Taxonomy 2024; 4: 53 - 111. https: // doi. org / 10.3390 / taxonomy 4010005" type="journal article" year="2024">Dupérré and Tapia, 2024</bibRefCitation>
|
||||
), others slightly behind and at the sides (e.g.
|
||||
<taxonomicName id="7CC74D002764FFAE96FBDD58FF7D6D70" authority="Mello-Leitao, 1942" authorityName="Mello-Leitao" authorityYear="1942" class="Arachnida" family="Hahniidae" genus="Austrohahnia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE96FBDD58FDB26D50" box="[501,634,1590,1614]" italics="true" pageId="12" pageNumber="13">Austrohahnia</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE9571DD58FF7D6D70" author="Mello-Leitao CF" pageId="12" pageNumber="13" pagination="381 - 426" refId="ref34125" refString="Mello-Leitao CF. Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (nueva serie, Zoologia) 1942; 2: 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), and in many the posterior laterals are more at the side of the anterior laterals; this variation forms a continuum. The above diagnosis is provided to reflect those changes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
383
data/33/6E/87/336E87952764FFAC930BDD08FA946E7B.xml
Normal file
383
data/33/6E/87/336E87952764FFAC930BDD08FA946E7B.xml
Normal file
|
|
@ -0,0 +1,383 @@
|
|||
<document id="5AA34EC784FBA9E22F81D676C0286A36" ID-DOI="10.1093/zoolinnean/zlae165" ID-ISSN="0024-4082" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1738158451996" checkinUser="plazi" docAuthor="Burgo, Agustina, Catley, Kefyn, Grismado, Cristian J., Dupérré, Nadine, Benjamin, Suresh P., Hormiga, Gustavo, Griswold, Charles, Martínez, Leonel & Ramírez, Martín J." docDate="2025" docId="336E87952764FFAC930BDD08FA946E7B" docLanguage="en" docName="zlae165.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae165" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Cybaeolus Simon 1884" docType="treatment" docVersion="1" lastPageNumber="15" masterDocId="CF57FFED2768FFA2970EDB6EFFC86B1E" masterDocTitle="Systematics of the colour-polymorphic spider genus Cybaeolus, with comments on the phylogeny of the family Hahniidae (Araneae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="13" updateTime="1738922663959" updateUser="carolina">
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<mods:mods id="3C723A0666BA176E10F5EC27F75D0AB4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A876E29FAAF1AC83BBF50731BE88C76D">
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<mods:title id="F39653665E06BD6AE426A8545A4053B2">Systematics of the colour-polymorphic spider genus Cybaeolus, with comments on the phylogeny of the family Hahniidae (Araneae)</mods:title>
|
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|
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<mods:name id="C6F870A495EADDF8036CE1EAD5250FEF" type="personal">
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<mods:role id="81C09B7B700455BEBBFE07CC57A4DFC2">
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<mods:roleTerm id="15376B6E22FA45B1F5BFD4E12392E76E">Author</mods:roleTerm>
|
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</mods:role>
|
||||
<mods:namePart id="E66514253606C0AA9999AB9075CDE62C">Burgo, Agustina</mods:namePart>
|
||||
<mods:affiliation id="C98CFDC0BAAD2260AAD64A77CF83512B">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="9D0DCE44592002C4C3B14B230FD74930" type="personal">
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<mods:role id="629FA3001A963DDAA79104C35601031C">
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<mods:roleTerm id="F6E680F02054AA70B2B6F8F4DB1E41E4">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="B0CB1375E15544616C44C52BD24F0EB1">Catley, Kefyn</mods:namePart>
|
||||
<mods:affiliation id="432EA8FB1BAAC13764F0AF62E4E1F22E">Western Carolina University, Cullowhee, NC 28723, United States</mods:affiliation>
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</mods:name>
|
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<mods:name id="34CCFD4B620EC054D9C549000F8D9048" type="personal">
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<mods:roleTerm id="D0CB435984519B3B3A13E41A7A01563D">Author</mods:roleTerm>
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</mods:role>
|
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<mods:namePart id="A893A800916B8D6522CE7894C3920C0E">Grismado, Cristian J.</mods:namePart>
|
||||
<mods:affiliation id="08B5BAA9B57D4542F98B427E4520B09E">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
|
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</mods:name>
|
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<mods:namePart id="1A433CED217C29F3C6A4D50BEC1CAE23">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="F25D36B6A6A74655BA605EDD19D9BE48">Museum of Nature Hamburg, Zoology, Leibniz-Institute for the Analysis of Biodiversity Change (LIB), Center for Taxonomy and Morphology, 20146 Hamburg, Germany</mods:affiliation>
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<mods:affiliation id="29F34E73655C139A05F6D68DA9FB0C28">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart id="B65F689531E48ACFDBE2557104EDCDE4">Hormiga, Gustavo</mods:namePart>
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<mods:affiliation id="1D651B2B0DE95159F54873D111ADE7BD">Department of Biology, The George Washington University, Washington, DC 20052, United States</mods:affiliation>
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<mods:affiliation id="CC25DE9A4CE5A32F29C7CB471ED7456E">California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, United States</mods:affiliation>
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<mods:namePart id="F0FB68E7BDDEAC5396645E3906AC293F">Martínez, Leonel</mods:namePart>
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<mods:affiliation id="127D64017B99766B3DF3BA52ACD80D72">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:affiliation id="371044AA28430282CFFD59467121A6BF">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:nameIdentifier id="198CC39F11DAD991C87A0118DF2F17E0" type="email">ramirez@macn.gov.ar</mods:nameIdentifier>
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<mods:title id="2831E320AF0DBFEB9B1EFD2809C789EB">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="E5C49D04167E4BF8E626E34D232E5904">2025</mods:date>
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<mods:number id="5309C46495770B97B143FD628420F2D5">2025-01-10</mods:number>
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<mods:number id="6A59C961A5980981BBB2F0E72392B924">203</mods:number>
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<mods:number id="565CED954A39034EAD80655FBC82805C">1</mods:number>
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<mods:url id="6ECD6FDFDB0C97EC62D6B3D44CC829BC">https://doi.org/10.1093/zoolinnean/zlae165</mods:url>
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<mods:identifier id="2B8CF153C86C5DE34494EBDDDE5052A2" type="ISSN">0024-4082</mods:identifier>
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<treatment id="336E87952764FFAC930BDD08FA946E7B" LSID="urn:lsid:plazi:treatment:336E87952764FFAC930BDD08FA946E7B" httpUri="http://treatment.plazi.org/id/336E87952764FFAC930BDD08FA946E7B" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="F3DD65082764FFAE930BDD08FB3E6D9F" box="[1029,1270,1638,1665]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="BB7836832764FFAE930BDD08FB3E6D9F" blockId="12.[825,1475,1638,1890]" box="[1029,1270,1638,1665]" pageId="12" pageNumber="13">
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<taxonomicName id="7CC74D002764FFAE930BDD08FB3E6D9F" ID-CoL="3XSB" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[1029,1270,1638,1665]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="89B3EA912764FFAE930BDD08FB3E6D9F" bold="true" box="[1029,1270,1638,1665]" pageId="12" pageNumber="13">
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<emphasis id="89B3EA912764FFAE930BDD08FBA46D9F" bold="true" box="[1029,1132,1638,1665]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
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<bibRefCitation id="DF564B722764FFAE937EDD08FB3E6D9F" author="Simon E" box="[1136,1270,1638,1665]" pageId="12" pageNumber="13" pagination="117 - 44" refId="ref34941" refString="Simon E. Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France 1884; 9: 117 - 44." type="journal article" year="1884">Simon, 1884</bibRefCitation>
|
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</emphasis>
|
||||
</taxonomicName>
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection id="F3DD65082764FFAE9437DDE3FB686C7F" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="BB7836832764FFAE9437DDE3FB3F6DDB" blockId="12.[825,1475,1638,1890]" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="7CC74D002764FFAE9437DDE3FBBE6DBB" authority="Simon 1884: 125" authorityName="Simon" authorityPageNumber="125" authorityYear="1884" box="[825,1142,1677,1701]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9437DDE3FC546DBB" box="[825,924,1677,1701]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
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<bibRefCitation id="DF564B722764FFAE94A0DDE3FBBE6DBB" author="Simon E" box="[942,1142,1677,1701]" pageId="12" pageNumber="13" pagination="117 - 44" refId="ref34941" refString="Simon E. Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France 1884; 9: 117 - 44." type="journal article" year="1884">Simon 1884: 125</bibRefCitation>
|
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</taxonomicName>
|
||||
(
|
||||
<typeStatus id="647C88212764FFAE939EDDE3FB096DBB" box="[1168,1217,1677,1701]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species by monotypy
|
||||
<taxonomicName id="7CC74D002764FFAE9437DDC2FBB96DDB" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[825,1137,1708,1733]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pusillus">
|
||||
<emphasis id="89B3EA912764FFAE9437DDC2FC226DDA" box="[825,1002,1708,1732]" italics="true" pageId="12" pageNumber="13">Cybaeolus pusillus</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE94FEDDC2FBB96DDB" author="Simon E" box="[1008,1137,1708,1733]" pageId="12" pageNumber="13" pagination="117 - 44" refId="ref34941" refString="Simon E. Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France 1884; 9: 117 - 44." type="journal article" year="1884">Simon, 1884</bibRefCitation>
|
||||
</taxonomicName>
|
||||
); 1898: 244.
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832764FFAE9437DDA3FBE86C3C" blockId="12.[825,1475,1638,1890]" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="7CC74D002764FFAE9437DDA3FB816DFA" authority="Simon 1904: 110" authorityName="Simon" authorityPageNumber="110" authorityYear="1904" box="[825,1097,1740,1764]" class="Arachnida" family="Hahniidae" genus="Mevianes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9437DDA3FC5C6DFA" box="[825,916,1741,1764]" italics="true" pageId="12" pageNumber="13">Mevianes</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE9497DDA2FB816DFA" author="Simon E" box="[921,1097,1740,1764]" pageId="12" pageNumber="13" pagination="83 - 114" refId="ref35103" refString="Simon E. Etude sur les arachnides du Chili recueillis en 1900, 1901 et 1902, par MM. C. Porter, Dr Delfin, Barcey Wilson et Edwards. Annales de la Societe Entomologique de Belgique 1904; 48: 83 - 114." type="journal article" year="1904">Simon 1904: 110</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<typeStatus id="647C88212764FFAE9358DDA2FB4F6DFA" box="[1110,1159,1740,1764]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="7CC74D002764FFAE93D7DDA3FA746DFA" authority="Simon, 1904" authorityName="Simon" authorityYear="1904" box="[1241,1468,1740,1764]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="delfini">
|
||||
<emphasis id="89B3EA912764FFAE93D7DDA3FAFE6DFA" box="[1241,1334,1740,1764]" italics="true" pageId="12" pageNumber="13">M. delfini</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE9235DDA2FA746DFA" author="Simon E" box="[1339,1468,1740,1764]" pageId="12" pageNumber="13" pagination="83 - 114" refId="ref35103" refString="Simon E. Etude sur les arachnides du Chili recueillis en 1900, 1901 et 1902, par MM. C. Porter, Dr Delfin, Barcey Wilson et Edwards. Annales de la Societe Entomologique de Belgique 1904; 48: 83 - 114." type="journal article" year="1904">Simon, 1904</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, designated by
|
||||
<bibRefCitation id="DF564B722764FFAE94D1DD85FB336C1D" author="Petrunkevitch A" box="[991,1275,1771,1795]" pageId="12" pageNumber="13" pagination="1 - 270" refId="ref34672" refString="Petrunkevitch A. Systema Aranearum. Transactions of the Connecticut Academy of Arts and Sciences 1928; 29: 1 - 270." type="journal article" year="1928">Petrunkevitch (1928: 93)</bibRefCitation>
|
||||
. Synonymized by
|
||||
<bibRefCitation id="DF564B722764FFAE9437DC64FBD46C3C" author="Lehtinen PT" box="[825,1052,1802,1826]" pageId="12" pageNumber="13" pagination="199 - 468" refId="ref33957" refString="Lehtinen PT. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 1967; 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967: 226)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832764FFAE9437DC44FB686C7F" blockId="12.[825,1475,1638,1890]" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="7CC74D002764FFAE9437DC44FBF56C5C" authority="Simon 1902: 20" authorityName="Simon" authorityPageNumber="20" authorityYear="1902" box="[825,1085,1834,1858]" class="Arachnida" family="Dictynidae" genus="Clitistes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9437DC44FC4D6C5C" box="[825,901,1834,1858]" italics="true" pageId="12" pageNumber="13">Clitistes</emphasis>
|
||||
Simon 1902: 20
|
||||
</taxonomicName>
|
||||
(
|
||||
<typeStatus id="647C88212764FFAE9341DC44FB486C5C" box="[1103,1152,1834,1858]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species by monotypy
|
||||
<taxonomicName id="7CC74D002764FFAE9278DC44FBD86C7F" authority="Simon 1902" authorityName="Simon" authorityYear="1902" class="Arachnida" family="Dictynidae" genus="Clitistes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="velutinus">
|
||||
<emphasis id="89B3EA912764FFAE9278DC44FC476C7F" italics="true" pageId="12" pageNumber="13">Clitistes velutinus</emphasis>
|
||||
Simon 1902
|
||||
</taxonomicName>
|
||||
), synon. nov..
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="F3DD65082764FFAF9345DCECFD6D6EBB" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
|
||||
<paragraph id="BB7836832764FFAE9345DCECFB796C85" blockId="12.[825,1474,1922,2015]" box="[1099,1201,1922,1947]" pageId="12" pageNumber="13">
|
||||
<emphasis id="89B3EA912764FFAE9345DCECFB796C85" box="[1099,1201,1922,1947]" italics="true" pageId="12" pageNumber="13">Diagnosis</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832764FFAF9437DCC9FD6D6EBB" blockId="12.[825,1474,1922,2015]" lastBlockId="13.[113,762,1295,1445]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||||
Species of
|
||||
<taxonomicName id="7CC74D002764FFAE94B8DCC9FBD16CA1" authorityName="Simon" authorityYear="1884" box="[950,1049,1959,1983]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE94B8DCC9FBD16CA1" box="[950,1049,1959,1983]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
||||
</taxonomicName>
|
||||
resemble other hahniids by having a retrolateral hook-shaped tibial process with a groove through which (presumably) the embolus slides (
|
||||
<figureCitation id="23FC2A062765FFAF9528DE61FD5D6E39" box="[550,661,1295,1319]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="13" pageNumber="14">Fig. 15I–K</figureCitation>
|
||||
), but are distinguished by having a retrolateral patellar process with two or more tips (
|
||||
<figureCitation id="23FC2A062765FFAF97F3DE20FE9C6E78" box="[253,340,1358,1382]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="13" pageNumber="14">Fig. 15D</figureCitation>
|
||||
), and spinnerets arranged in a group with the
|
||||
<collectionCode id="DDD6AE462765FFAF9796DE00FF036E98" box="[152,203,1390,1414]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="13" pageNumber="14">PMS</collectionCode>
|
||||
posterior to the
|
||||
<collectionCode id="DDD6AE462765FFAF967ADE00FE696E98" box="[372,417,1390,1414]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
(
|
||||
<figureCitation id="23FC2A062765FFAF96BCDE03FDC16E9B" box="[434,521,1389,1414]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="8.[129,194,1268,1292]" captionStartId-1="9.[113,178,1508,1532]" captionStartId-2="10.[129,194,1249,1273]" captionStartId-3="11.[113,178,1348,1372]" captionTargetBox-0="[161,1441,144,1241]" captionTargetBox-1="[146,1426,144,1480]" captionTargetBox-2="[161,1441,144,1221]" captionTargetBox-3="[146,1426,144,1321]" captionTargetId-0="figure-387@8.[161,1441,144,1241]" captionTargetId-1="figure-265@9.[146,1426,144,1480]" captionTargetId-2="figure-436@10.[161,1441,144,1221]" captionTargetId-3="figure-255@11.[146,1426,144,1321]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 3. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, female. A, overview.B, ALS left. C, overview of PMS. D, PLS left (several Ac spigot shafts broken).Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-1="Figure 4. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, male.A, overview. B, PMS left.C, ALS left. D, PLS left. E, epiandrum.Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-2="Figure 5. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), female. A, overview.B, ALS right.C, PMS left. D, PLS right. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Pi, piriform gland spigot; TS, tracheal spiracle." captionText-3="Figure 6. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), male. A, overview.B, ALS left. C, overview of PMS. D, PLS right.E, epiandrum. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot." pageId="13" pageNumber="14">Figs 3–6</figureCitation>
|
||||
), instead of arranged in a line or having the
|
||||
<collectionCode id="DDD6AE462765FFAF9636DEE3FEA36EBB" box="[312,363,1421,1445]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="13" pageNumber="14">PMS</collectionCode>
|
||||
advanced in between the
|
||||
<collectionCode id="DDD6AE462765FFAF957CDEE3FD696EBB" box="[626,673,1421,1445]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="EFB8660B2765FFAF977CDFE1FF036FC1" pageId="13" pageNumber="14" startId="13.[114,179,1167,1191]" targetBox="[146,1426,144,1140]" targetPageId="13" targetType="figure">
|
||||
<paragraph id="BB7836832765FFAF977CDFE1FF036FC1" blockId="13.[113,1448,1167,1247]" pageId="13" pageNumber="14">
|
||||
<emphasis id="89B3EA912765FFAF977CDFE1FF026FB9" bold="true" box="[114,202,1167,1191]" pageId="13" pageNumber="14">Figure 8.</emphasis>
|
||||
Scanning electron micrographs of spinnerets of
|
||||
<taxonomicName id="7CC74D002765FFAF9580DFE1FC0C6FB6" authority="(C. L. Koch, 1841)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1841" box="[654,964,1167,1192]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusilla">
|
||||
<emphasis id="89B3EA912765FFAF9580DFE1FCDC6FB9" box="[654,788,1167,1191]" italics="true" pageId="13" pageNumber="14">Hahnia pusilla</emphasis>
|
||||
(C.L.
|
||||
<bibRefCitation id="DF564B722765FFAF9448DFE1FC756FB6" author="Koch CL" box="[838,957,1167,1192]" pageId="13" pageNumber="14" pagination="41 - 131" refId="ref33849" refString="Koch CL. Die Arachniden. Nurnberg Achter Band: C. H. Zeh'sche Buchhandlung, 1841, 41 - 131." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, male. A, overview. B, ALS left. C, PMS left. D. PLS left. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Tp, tartipore.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="F3DD65082765FFAC968EDEABFE2A6C4A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||||
<paragraph id="BB7836832765FFAF968EDEABFE246EC0" blockId="13.[384,492,1477,1502]" box="[384,492,1477,1502]" pageId="13" pageNumber="14">
|
||||
<emphasis id="89B3EA912765FFAF968EDEABFE246EC0" box="[384,492,1477,1502]" italics="true" pageId="13" pageNumber="14">Description</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832765FFAF977FDE85FCA36D81" blockId="13.[112,764,1515,1978]" lastBlockId="13.[809,1461,1295,1978]" pageId="13" pageNumber="14">
|
||||
Small spiders (1.50–3.50) with eight eyes (
|
||||
<figureCitation id="23FC2A062765FFAF9536DE85FD586D1D" box="[568,656,1515,1539]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="13" pageNumber="14">Fig. 15A</figureCitation>
|
||||
), anterior eye row slightly procurved in anterior view, posterior row slightly procurved to straight in dorsal view,
|
||||
<collectionCode id="DDD6AE462765FFAF96E7DD44FDE96D5C" box="[489,545,1578,1602]" pageId="13" pageNumber="14">AME</collectionCode>
|
||||
normally developed, PME separated by one diameter of PME, clypeus no greater than 2.5× diameter of
|
||||
<collectionCode id="DDD6AE462765FFAF962BDD06FE976D9E" box="[293,351,1640,1664]" pageId="13" pageNumber="14">AME</collectionCode>
|
||||
. Colour patterns of prosoma, and especially of opisthosoma, extremely polymorphic (
|
||||
<figureCitation id="23FC2A062765FFAF9566DDE9FD036DBE" box="[616,715,1671,1696]" captionStart="Figure 21" captionStartId="26.[129,194,1772,1796]" captionTargetBox="[327,1276,144,1744]" captionTargetId="figure-6@26.[327,1276,144,1744]" captionTargetPageId="26" captionText="Figure 21. Cybaeolus pusillus Simon, 1884, female.A, dorsal habitus. B, epigyne, ventral view.C, ventral habitus (A–C, CASENT 9039051). D, epigyne cleared, ventral view (MACN-Ar 43242). E, epigyne cleared, dorsal view, variant with copulatory openings separated (MACN-Ar 29766). F, epigyne cleared, dorsal view, extreme variant with copulatory openings well separated (MACN-Ar 41386). G, female holotype, dorsal habitus. H, same, ventral habitus.I, same, dissected epigyne, dorsal view. J, same, epigyne cleared, dorsal view." pageId="13" pageNumber="14">Figs. 21A</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF95D5DDE6FD3D6DBE" box="[731,757,1672,1696]" captionStart="Figure 22" captionStartId="27.[113,178,1353,1377]" captionTargetBox="[146,1426,144,1326]" captionTargetId="figure-270@27.[146,1426,144,1326]" captionTargetPageId="27" captionText="Figure 22. Colour polymorphism in Cybaeolus pusillus Simon, 1884, grouped by locality and collection event.A, Chile, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43965). B, Chile, Los Lagos, Parque Nacional Alerce Andino (MACN-Ar 43952). C, Argentina, Río Negro, Parque Nacional Nahuel Huapi (MACN-Ar 43386).D, Chile, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43971). E, Chile, Los Lagos, Parque Nacional (MACN-Ar 43970).F, Chile, Magallanes, Punta Arenas (MACN-Ar 43980). G, Chile, Los Lagos, Alerce Andino (MACN-Ar 43967). H, Chile, Los Lagos, Alerce Andino (MACN-Ar 43957).I, Chile, Los Ríos, Alerce Costero (MACN-Ar 43961). J, Chile, Magallanes, Cueva del Milodón (MACN-Ar 43976, MACN-Ar 43978). K, Chile, Los Ríos, Reserva Costera Valdiviana (CASENT 9039051, CASENT 9039054, CASENT 9034002)." pageId="13" pageNumber="14">22</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF977FDDC9FF556DA1" box="[113,157,1703,1727]" captionStart="Figure 23" captionStartId="29.[113,178,1404,1428]" captionTargetBox="[146,1426,144,1376]" captionTargetId="figure-284@29.[146,1426,144,1376]" captionTargetPageId="29" captionText="Figure 23. Cybaeolus rastellus (Roth, 1967), female.A, dorsal habitus. B, epigyne, ventral view.C, ventral habitus.D, epigyne cleared, ventral view." pageId="13" pageNumber="14">23A</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF97A7DDC9FF0B6DA1" box="[169,195,1703,1727]" captionStart="Figure 24" captionStartId="30.[130,195,1562,1586]" captionTargetBox="[161,1441,144,1534]" captionTargetId="figure-180@30.[161,1441,144,1534]" captionTargetPageId="30" captionText="Figure 24.Colour polymorphism in Cybaeolus rastellus (Roth, 1967) in Chile, grouped by locality and collection event. A, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43969). B, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43966). C, Los Lagos, Parque Nacional Alerce Andino (MACN-Ar 43968).D, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43954). E, Los Lagos, Parque Nacional Alerce (MACN-Ar 43958). F, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43973). G, Los Lagos, Parque Nacional Alerce Andino (MACN-Ar 43953). H, Los Lagos, Parque Nacional Alerce Andino (MACN-Ar 43959).I, CLos Ríos, Parque Nacional Alerce Costero (MACN-Ar 43962)." pageId="13" pageNumber="14">24</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF97DEDDC9FF346DA1" box="[208,252,1703,1727]" captionStart="Figure 25" captionStartId="34.[129,194,1738,1762]" captionTargetBox="[161,1441,144,1711]" captionTargetId="figure-95@34.[161,1441,144,1711]" captionTargetPageId="34" captionText="Figure 25. Cybaeolus delfini (Simon 1904), female (A–E, MACN-Ar 42819. G, MACN-Ar 42750). A, eyes, dorsal view.B, prosoma, anterior view. C, habitus lateral. D, habitus dorsal. E, epigyne, ventral view. F, habitus ventral. G, epigyne cleared, ventral view." pageId="13" pageNumber="14">25D</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF9606DDC9FEF96DA1" box="[264,305,1703,1727]" captionStart="Figure 26" captionStartId="35.[113,178,1523,1547]" captionTargetBox="[146,1426,144,1495]" captionTargetId="figure-195@35.[146,1426,144,1496]" captionTargetPageId="35" captionText="Figure 26. Cybaeolus delfini (Simon, 1904), female.A, epigyne from variant with transversal proximal copulatory ducts, ventral view (MACN-Ar 43232). B, same, cleared, ventral view, arrow to proximal copulatory duct. C, genitalia of variant with transversal proximal copulatory duct and longer posterior loop (arrows; AMNH LAM-0559). D, genitalia of intermediate variant (AMNH, LAM-0558). E, holotype, dorsal habitus. F, same, ventral habitus. G, same, epigyne cleared, dorsal view.E, same, schematic drawing." pageId="13" pageNumber="14">26E</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF9633DDC9FE9F6DA1" box="[317,343,1703,1727]" captionStart="Figure 27" captionStartId="36.[129,194,1772,1796]" captionTargetBox="[181,1421,144,1744]" captionTargetId="figure-6@36.[181,1421,144,1744]" captionTargetPageId="36" captionText="Figure 27. Colour polymorphism in Cybaeolus delfini (Simon, 1904), grouped by locality and collection event. A, Argentina, Neuquén: Parque Nacional Lanín (MACN-Ar 43581). B, Chile, La Araucanía, Parque Nacional Nahuelbuta (MACN-Ar 43232). C, Chile, Los Ríos, Reserva Costera Valdiviana, (CASENT 9039052, CASENT 9034004). D, Argentina, Santa Cruz, Chaltén (MACN-Ar 43327). E, Argentina, Neuquén, Parque Nacional Lanín (MACN-Ar 43343). F, Chile, Los Lagos, Parque Nacional Puyehue (MACN-Ar 43972)." pageId="13" pageNumber="14">27</figureCitation>
|
||||
). Carapace light brown, with or without thoracic markings, without foveal furrow. Opisthosoma oval or elongate, it can be dark, pale, orange, or brown, mottled, with or without chevrons. Chelicerae with three or four promarginal and three retromarginal teeth, male with or without stridulatory file made of minute bumps, rastellum (a row of rake setae with thickened bases;
|
||||
<figureCitation id="23FC2A062765FFAF962ADC0DFEB56C65" box="[292,381,1891,1915]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">Fig. 18C</figureCitation>
|
||||
) present in females of
|
||||
<taxonomicName id="7CC74D002765FFAF957CDC0DFD046C65" authorityName="Simon" authorityYear="1904" box="[626,716,1891,1915]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfini">
|
||||
<emphasis id="89B3EA912765FFAF957CDC0DFD046C65" box="[626,716,1891,1915]" italics="true" pageId="13" pageNumber="14">C. delfini</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="7CC74D002765FFAF977FDCECFF156C84" baseAuthorityName="Roth" baseAuthorityYear="1967" box="[113,221,1922,1946]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rastellus">
|
||||
<emphasis id="89B3EA912765FFAF977FDCECFF156C84" box="[113,221,1922,1946]" italics="true" pageId="13" pageNumber="14">C. rastellus</emphasis>
|
||||
</taxonomicName>
|
||||
, absent in
|
||||
<taxonomicName id="7CC74D002765FFAF9645DCECFE676C84" authorityName="Simon" authorityYear="1884" box="[331,431,1922,1946]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusillus">
|
||||
<emphasis id="89B3EA912765FFAF9645DCECFE676C84" box="[331,431,1922,1946]" italics="true" pageId="13" pageNumber="14">C. pusillus</emphasis>
|
||||
</taxonomicName>
|
||||
. Endites not convergent, serrula in a transverse line (
|
||||
<figureCitation id="23FC2A062765FFAF9632DCCFFE466CA4" box="[316,398,1953,1978]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">Fig. 18B</figureCitation>
|
||||
), posterior external margin variably inflated or projecting in males. Labium slightly wider than long, anterior edge straight, posterior slightly convex. Sternum shieldshaped, weakly concave anteriorly at base of labium. Male palp with simple retrolateral patellar apophysis with two or more distal thorns (
|
||||
<figureCitation id="23FC2A062765FFAF94B2DEE3FBDD6EBB" box="[956,1045,1421,1445]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="13" pageNumber="14">Fig. 15D</figureCitation>
|
||||
), RTA acute, curved, furrowed, tip with regular teeth pointing backwards (
|
||||
<figureCitation id="23FC2A062765FFAF939CDEC2FACD6EDA" box="[1170,1285,1452,1476]" captionStart="Figure 15" captionStartId="20.[130,195,1169,1193]" captionTargetBox="[161,1441,144,1141]" captionTargetId="figure-197@20.[161,1441,144,1141]" captionTargetPageId="20" captionText="Figure 15.Scanning electron micrographs of male Cybaeolus rastellus (Roth, 1967). A, prosoma in anterior view. B, detail of left chelicera in anterior view, showing rake setae with slightly expanded bases. C, left endite, showing posterior lateral edge protruding.D, right palpal patella in dorsal view showing patellar apophysis.E, left palp in ventral view. F, same in dorsal view. G, same in retrolateral view.H, detail of copulatory bulb in ventral view.I, detail of tibial retrolateral apophysis in retrolateral view.J, detail of male palp in retrolateral view. K, same, closeup showing the cymbial groove. L, same, dorsal view, closeup showing the groove on the RTA. Abbreviations: CyGv, cymbial groove of male palp; E, embolus of male palp; MA, median apophysis of male palp; PA, male palpal patellar retrolateral apophysis; RTA retrolateral tibial apophysis of male palp; RTAGv, groove on the retrolateral tibial apophysis of male palp." pageId="13" pageNumber="14">Figs 15I–K</figureCitation>
|
||||
); a swollen area near the RTA base bears several thick setae. Cymbium oval, with retrolateral furrow leading to RTA (
|
||||
<figureCitation id="23FC2A062765FFAF9391DE85FB266D1D" box="[1183,1262,1515,1539]" captionStart="Figure 17" captionStartId="22.[128,193,1502,1526]" captionTargetBox="[161,1441,144,1474]" captionTargetId="figure-216@22.[161,1441,144,1474]" captionTargetPageId="22" captionText="Figure 17. Scanning electron micrographs of female Cybaeolus pusillus Simon, 1884. A, left chelicera, posterior view.B, same, detail of promargin and fang, ventral view. C, claws of left leg I. D, tarsus I, dorsal view.E, metatarsal sensory organ of left leg I. F, trichobothrium of tarsus I. G, tarsal organ I. H, epigyne, note copulatory openings obstructed by mating plugs. I, vulva, dorsal view. Abbreviations: COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">Fig. 17J</figureCitation>
|
||||
). Copulatory bulb with disc-shaped tegulum, with a membranous patch where the fleshy median apophysis arises in
|
||||
<taxonomicName id="7CC74D002765FFAF938ADD44FB146D5F" authorityName="Simon" authorityYear="1904" box="[1156,1244,1577,1601]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="delfini">
|
||||
<emphasis id="89B3EA912765FFAF938ADD44FB146D5F" box="[1156,1244,1577,1601]" italics="true" pageId="13" pageNumber="14">C. delfini</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<emphasis id="89B3EA912765FFAF9200DD44FA496D5F" box="[1294,1409,1577,1601]" italics="true" pageId="13" pageNumber="14">
|
||||
<taxonomicName id="7CC74D002765FFAF9200DD44FAB26D5F" baseAuthorityName="Roth" baseAuthorityYear="1967" box="[1294,1402,1577,1601]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rastellus">C. rastellus</taxonomicName>
|
||||
;
|
||||
</emphasis>
|
||||
only the membranous patch present in
|
||||
<taxonomicName id="7CC74D002765FFAF939BDD27FB356D7F" authorityName="Simon" authorityYear="1884" box="[1173,1277,1609,1633]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pusillus">
|
||||
<emphasis id="89B3EA912765FFAF939BDD27FB356D7F" box="[1173,1277,1609,1633]" italics="true" pageId="13" pageNumber="14">C. pusillus</emphasis>
|
||||
</taxonomicName>
|
||||
. Embolus arising proximally from tegulum, rotating clockwise (left palp, ventral view).
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832765FFAC944BDDC9FE2A6C4A" blockId="13.[809,1461,1295,1978]" lastBlockId="14.[128,779,1257,1876]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||||
Epigyne with paired copulatory openings, usually blocked with mating plugs (
|
||||
<figureCitation id="23FC2A062765FFAF930FDDA8FBAC6DC0" box="[1025,1124,1734,1758]" captionStart="Figure 17" captionStartId="22.[128,193,1502,1526]" captionTargetBox="[161,1441,144,1474]" captionTargetId="figure-216@22.[161,1441,144,1474]" captionTargetPageId="22" captionText="Figure 17. Scanning electron micrographs of female Cybaeolus pusillus Simon, 1884. A, left chelicera, posterior view.B, same, detail of promargin and fang, ventral view. C, claws of left leg I. D, tarsus I, dorsal view.E, metatarsal sensory organ of left leg I. F, trichobothrium of tarsus I. G, tarsal organ I. H, epigyne, note copulatory openings obstructed by mating plugs. I, vulva, dorsal view. Abbreviations: COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">Figs 17H</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062765FFAF937ADDA8FB696DC0" box="[1140,1185,1734,1758]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">18G</figureCitation>
|
||||
). Copulatory ducts long, leading to spherical spermathecae, fertilization ducts short, on posterior margin; accessory bulb unremarkable, restricted to small bump with gland ducts near the copulatory opening (observed only in
|
||||
<taxonomicName id="7CC74D002765FFAF94B6DC2AFBEA6C45" baseAuthorityName="Roth" baseAuthorityYear="1967" box="[952,1058,1859,1883]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rastellus">
|
||||
<emphasis id="89B3EA912765FFAF94B6DC2AFBEA6C45" box="[952,1058,1859,1883]" italics="true" pageId="13" pageNumber="14">C. rastellus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<figureCitation id="23FC2A062765FFAF9320DC2DFBB26C42" box="[1070,1146,1859,1884]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="13" pageNumber="14">Fig.18I</figureCitation>
|
||||
). Spinnerets (
|
||||
<figureCitation id="23FC2A062765FFAF9204DC2DFA976C45" box="[1290,1375,1859,1884]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStartId-0="8.[129,194,1268,1292]" captionStartId-1="9.[113,178,1508,1532]" captionStartId-2="10.[129,194,1249,1273]" captionStartId-3="11.[113,178,1348,1372]" captionTargetBox-0="[161,1441,144,1241]" captionTargetBox-1="[146,1426,144,1480]" captionTargetBox-2="[161,1441,144,1221]" captionTargetBox-3="[146,1426,144,1321]" captionTargetId-0="figure-387@8.[161,1441,144,1241]" captionTargetId-1="figure-265@9.[146,1426,144,1480]" captionTargetId-2="figure-436@10.[161,1441,144,1221]" captionTargetId-3="figure-255@11.[146,1426,144,1321]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 3. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, female. A, overview.B, ALS left. C, overview of PMS. D, PLS left (several Ac spigot shafts broken).Abbreviations:Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-1="Figure 4. Scanning electron micrographs of spinnerets of Cybaeolus pusillus Simon, 1884, male.A, overview. B, PMS left.C, ALS left. D, PLS left. E, epiandrum.Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot; Tp, tartipore." captionText-2="Figure 5. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), female. A, overview.B, ALS right.C, PMS left. D, PLS right. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; Pi, piriform gland spigot; TS, tracheal spiracle." captionText-3="Figure 6. Scanning electron micrographs of spinnerets of Cybaeolus rastellus (Roth, 1967), male. A, overview.B, ALS left. C, overview of PMS. D, PLS right.E, epiandrum. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, piriform gland spigot." pageId="13" pageNumber="14">Figs 3–6</figureCitation>
|
||||
) in relatively compact group,
|
||||
<collectionCode id="DDD6AE462765FFAF9305DC0DFBFE6C65" box="[1035,1078,1891,1915]" pageId="13" pageNumber="14">PLS</collectionCode>
|
||||
70–80° from
|
||||
<collectionCode id="DDD6AE462765FFAF93CCDC0DFB396C65" box="[1218,1265,1891,1915]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
,
|
||||
<collectionCode id="DDD6AE462765FFAF93F7DC0DFAE46C65" box="[1273,1324,1891,1915]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="13" pageNumber="14">PMS</collectionCode>
|
||||
40–50° from
|
||||
<collectionCode id="DDD6AE462765FFAF9424DCECFC916C84" box="[810,857,1922,1946]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
.
|
||||
<collectionCode id="DDD6AE462765FFAF946FDCECFC466C84" box="[865,910,1922,1946]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
conical with wide base,
|
||||
<collectionCode id="DDD6AE462765FFAF9371DCECFB626C84" box="[1151,1194,1922,1946]" pageId="13" pageNumber="14">PLS</collectionCode>
|
||||
and
|
||||
<collectionCode id="DDD6AE462765FFAF93D4DCECFAC56C84" box="[1242,1293,1922,1946]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="13" pageNumber="14">PMS</collectionCode>
|
||||
cylindrical,
|
||||
<collectionCode id="DDD6AE462765FFAF9286DCECFA7B6C84" box="[1416,1459,1922,1946]" pageId="13" pageNumber="14">PLS</collectionCode>
|
||||
equal in length to
|
||||
<collectionCode id="DDD6AE462765FFAF94EADCCCFBDB6CA4" box="[996,1043,1954,1978]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
, distal segment of
|
||||
<collectionCode id="DDD6AE462765FFAF93DFDCCCFB366CA4" box="[1233,1278,1954,1978]" pageId="13" pageNumber="14">ALS</collectionCode>
|
||||
very short; distal segment of
|
||||
<collectionCode id="DDD6AE462766FFAC97F4DF87FEED6E1F" box="[250,293,1257,1281]" pageId="14" pageNumber="15">PLS</collectionCode>
|
||||
very short,
|
||||
<collectionCode id="DDD6AE462766FFAC96AADF87FE076E1F" box="[420,463,1257,1281]" pageId="14" pageNumber="15">PLS</collectionCode>
|
||||
not extended, length equal to or smaller than
|
||||
<collectionCode id="DDD6AE462766FFAC9622DE66FE936E3E" box="[300,347,1288,1312]" pageId="14" pageNumber="15">ALS</collectionCode>
|
||||
. Colulus as two patches of setae. Spigot morphology (examined with SEM in adults of
|
||||
<taxonomicName id="7CC74D002766FFAC9553DE46FD0A6E21" authorityName="Simon" authorityYear="1884" box="[605,706,1319,1343]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pusillus">
|
||||
<emphasis id="89B3EA912766FFAC9553DE46FD0A6E21" box="[605,706,1319,1343]" italics="true" pageId="14" pageNumber="15">C. pusillus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="7CC74D002766FFAC95FDDE46FF186E41" baseAuthorityName="Roth" baseAuthorityYear="1967" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rastellus">
|
||||
<emphasis id="89B3EA912766FFAC95FDDE46FF186E41" italics="true" pageId="14" pageNumber="15">C. rastellus</emphasis>
|
||||
</taxonomicName>
|
||||
):
|
||||
<collectionCode id="DDD6AE462766FFAC97E4DE29FEDF6E41" box="[234,279,1351,1375]" pageId="14" pageNumber="15">ALS</collectionCode>
|
||||
with one major ampullate gland spigot flanked by a nubbin, and 10–18 piriform gland spigots.
|
||||
<collectionCode id="DDD6AE462766FFAC957CDE08FD6D6E60" box="[626,677,1382,1406]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="14" pageNumber="15">PMS</collectionCode>
|
||||
with one minor ampullate gland spigot, two cylindrical gland spigots in female, from five to eight aciniform gland spigots.
|
||||
<collectionCode id="DDD6AE462766FFAC95ABDECBFD186EA3" box="[677,720,1445,1469]" pageId="14" pageNumber="15">PLS</collectionCode>
|
||||
with one cylindrical gland spigot in female, from six to 11 aciniform gland spigots. Tracheal spiracle wide, slightly separated from spinnerets, leading to two thick trunks, each ramifying in three bundles of tracheoles, all limited to opisthosoma (observed from dissection of
|
||||
<taxonomicName id="7CC74D002766FFAC960ADD2CFEA66D47" baseAuthorityName="Roth" baseAuthorityYear="1967" box="[260,366,1601,1625]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rastellus">
|
||||
<emphasis id="89B3EA912766FFAC960ADD2CFEA66D47" box="[260,366,1601,1625]" italics="true" pageId="14" pageNumber="15">C. rastellus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<figureCitation id="23FC2A062766FFAC9674DD2FFE056D47" box="[378,461,1601,1625]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="14" pageNumber="15">Fig. 18E</figureCitation>
|
||||
). Leg formula 1243. Spination: femora I with dorsal and prolateral spines, patellae with two dorsal spines, tibiae I and II with dorsal and prolateral spines, tibiae II and IV with dorsal, prolateral, ventral, and retrolateral spines, metatarsi III and IV with prolateral, retrolateral, and ventral spines. Trichobothria with longitudinal fingerprint striae on proximal plate (
|
||||
<figureCitation id="23FC2A062766FFAC9646DD93FE556C0B" box="[328,413,1789,1813]" captionStart="Figure 17" captionStartId="22.[128,193,1502,1526]" captionTargetBox="[161,1441,144,1474]" captionTargetId="figure-216@22.[161,1441,144,1474]" captionTargetPageId="22" captionText="Figure 17. Scanning electron micrographs of female Cybaeolus pusillus Simon, 1884. A, left chelicera, posterior view.B, same, detail of promargin and fang, ventral view. C, claws of left leg I. D, tarsus I, dorsal view.E, metatarsal sensory organ of left leg I. F, trichobothrium of tarsus I. G, tarsal organ I. H, epigyne, note copulatory openings obstructed by mating plugs. I, vulva, dorsal view. Abbreviations: COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="14" pageNumber="15">Fig. 17F</figureCitation>
|
||||
), arranged on tarsi d0-1-0 or d0-1- 1-0 (
|
||||
<figureCitation id="23FC2A062766FFAC97BADC73FED86C2B" box="[180,272,1821,1845]" captionStart="Figure 17" captionStartId="22.[128,193,1502,1526]" captionTargetBox="[161,1441,144,1474]" captionTargetId="figure-216@22.[161,1441,144,1474]" captionTargetPageId="22" captionText="Figure 17. Scanning electron micrographs of female Cybaeolus pusillus Simon, 1884. A, left chelicera, posterior view.B, same, detail of promargin and fang, ventral view. C, claws of left leg I. D, tarsus I, dorsal view.E, metatarsal sensory organ of left leg I. F, trichobothrium of tarsus I. G, tarsal organ I. H, epigyne, note copulatory openings obstructed by mating plugs. I, vulva, dorsal view. Abbreviations: COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="14" pageNumber="15">Figs 17D</figureCitation>
|
||||
,
|
||||
<figureCitation id="23FC2A062766FFAC9612DC73FE816C2B" box="[284,329,1821,1845]" captionStart="Figure 18" captionStartId="23.[113,178,1443,1467]" captionTargetBox="[146,1426,144,1415]" captionTargetId="figure-134@23.[146,1426,144,1415]" captionTargetPageId="23" captionText="Figure 18. Scanning electron micrographs of female Cybaeolus rastellus (Roth, 1967). A, prosoma, anterior view. B, detail of endite showing the serrula, dorsal view.C, left chelicera, ventral view, arrow to expanded bases of rake setae making the rastellum. D, right tarsus I, dorsal view. E, tracheal system. F, claws of right leg I, retrolateral view. G, epigyne, note copulatory opening obstructed by mating plug.H, vulva, anteriordorsal view. I, same, detail of start of copulatory duct showing small accessory bulb and gland ducts. Abbreviations: AB, accessory bulb; COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="14" pageNumber="15">18D</figureCitation>
|
||||
), metatarsi d0-1-0 and tibiae d0-1-0. Tarsal organ a simple keyhole (
|
||||
<figureCitation id="23FC2A062766FFAC9677DC52FE196C4A" box="[377,465,1852,1876]" captionStart="Figure 17" captionStartId="22.[128,193,1502,1526]" captionTargetBox="[161,1441,144,1474]" captionTargetId="figure-216@22.[161,1441,144,1474]" captionTargetPageId="22" captionText="Figure 17. Scanning electron micrographs of female Cybaeolus pusillus Simon, 1884. A, left chelicera, posterior view.B, same, detail of promargin and fang, ventral view. C, claws of left leg I. D, tarsus I, dorsal view.E, metatarsal sensory organ of left leg I. F, trichobothrium of tarsus I. G, tarsal organ I. H, epigyne, note copulatory openings obstructed by mating plugs. I, vulva, dorsal view. Abbreviations: COp, copulatory opening; TO, tarsal organ; Tr, trichobothria." pageId="14" pageNumber="15">Fig. 17G</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="EFB8660B2766FFAC978FDF06FA8F6FA7" pageId="14" pageNumber="15" startId="14.[129,194,1128,1152]" targetBox="[161,1441,144,1101]" targetPageId="14" targetType="figure">
|
||||
<paragraph id="BB7836832766FFAC978FDF06FA8F6FA7" blockId="14.[129,1430,1128,1209]" pageId="14" pageNumber="15">
|
||||
<emphasis id="89B3EA912766FFAC978FDF06FF106F9F" bold="true" box="[129,216,1128,1153]" pageId="14" pageNumber="15">Figure 9.</emphasis>
|
||||
Scanning electron micrographs of spinnerets of
|
||||
<taxonomicName id="7CC74D002766FFAC9593DF07FBCC6F9F" authority="(Keyserling, 1887)" baseAuthorityName="Keyserling" baseAuthorityYear="1887" box="[669,1028,1129,1153]" class="Arachnida" family="Hahniidae" genus="Neoantistea" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="magna">
|
||||
<emphasis id="89B3EA912766FFAC9593DF07FC826F9E" box="[669,842,1129,1152]" italics="true" pageId="14" pageNumber="15">Neoantistea magna</emphasis>
|
||||
(
|
||||
<bibRefCitation id="DF564B722766FFAC945BDF07FC346F9F" author="Keyserling E" box="[853,1020,1129,1153]" pageId="14" pageNumber="15" pagination="421 - 90" refId="ref33794" refString="Keyserling E. Neue Spinnen aus America. VII. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 1887; 37: 421 - 90." type="journal article" year="1887">Keyserling, 1887</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, female. A, overview. B, ALS right. C, PMS right. D, PLS right. E, same, detail. Abbreviations: Ac, aciniform gland spigot; AT, anal tubercle; Cy, cylindrical gland spigot; MAP, major ampullate gland spigot; mAP, minor ampullate gland spigot; N, nubbin (atrophied spigot); Pi, Piriform gland spigot; Tp, tartipore.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="F3DD65082766FFAC9686DC1CFEE26CD1" pageId="14" pageNumber="15" type="taxon_list">
|
||||
<paragraph id="BB7836832766FFAC9686DC1CFDC96C92" blockId="14.[129,777,1906,1999]" box="[392,513,1906,1932]" pageId="14" pageNumber="15">
|
||||
<emphasis id="89B3EA912766FFAC9686DC1CFDC96C92" box="[392,513,1906,1932]" italics="true" pageId="14" pageNumber="15">Composition</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832766FFAC978FDCF6FEE26CD1" blockId="14.[129,777,1906,1999]" pageId="14" pageNumber="15">
|
||||
Three species:
|
||||
<taxonomicName id="7CC74D002766FFAC963FDCF6FE276CAE" authorityName="Simon" authorityYear="1884" box="[305,495,1944,1968]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pusillus">
|
||||
<emphasis id="89B3EA912766FFAC963FDCF6FE276CAE" box="[305,495,1944,1968]" italics="true" pageId="14" pageNumber="15">Cybaeolus pusillus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="7CC74D002766FFAC9509DCF6FD036CAE" baseAuthorityName="Roth" baseAuthorityYear="1967" box="[519,715,1944,1968]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rastellus">
|
||||
<emphasis id="89B3EA912766FFAC9509DCF6FD036CAE" box="[519,715,1944,1968]" italics="true" pageId="14" pageNumber="15">Cybaeolus rastellus</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="7CC74D002766FFAC978FDCD9FEEC6CD1" authorityName="Simon" authorityYear="1904" box="[129,292,1975,1999]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="delfini">
|
||||
<emphasis id="89B3EA912766FFAC978FDCD9FEEC6CD1" box="[129,292,1975,1999]" italics="true" pageId="14" pageNumber="15">Cybaeolus delfini</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="F3DD65082766FFAC934ADF86FA946E7B" pageId="14" pageNumber="15" type="distribution">
|
||||
<paragraph id="BB7836832766FFAC934ADF86FB7F6E1C" blockId="14.[1092,1207,1256,1282]" box="[1092,1207,1256,1282]" pageId="14" pageNumber="15">
|
||||
<emphasis id="89B3EA912766FFAC934ADF86FB7F6E1C" box="[1092,1207,1256,1282]" italics="true" pageId="14" pageNumber="15">Distribution</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="BB7836832766FFAC9437DE61FA946E7B" blockId="14.[825,1474,1295,1382]" pageId="14" pageNumber="15">
|
||||
In temperate forests and shrublands of
|
||||
<collectingCountry id="C3D076132766FFAC93D9DE61FAD96E39" box="[1239,1297,1295,1319]" name="Chile" pageId="14" pageNumber="15">Chile</collectingCountry>
|
||||
, from
|
||||
<collectingRegion id="7903F8612766FFAC9259DE61FC7A6E58" country="Chile" name="Antofagasta" pageId="14" pageNumber="15">Región de Antofagasta</collectingRegion>
|
||||
to
|
||||
<collectingRegion id="7903F8612766FFAC94D9DE40FB776E58" box="[983,1215,1326,1350]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="14" pageNumber="15">Región de Magallanes</collectingRegion>
|
||||
, and adjacent
|
||||
<collectingCountry id="C3D076132766FFAC9259DE40FA776E58" box="[1367,1471,1326,1350]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
|
||||
, from
|
||||
<collectingRegion id="7903F8612766FFAC947EDE20FBE76E7B" box="[880,1071,1357,1382]" country="Argentina" name="Neuquen" pageId="14" pageNumber="15">Neuquén province</collectingRegion>
|
||||
to
|
||||
<collectingRegion id="7903F8612766FFAC9341DE23FA906E7B" box="[1103,1368,1357,1382]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="14" pageNumber="15">Tierra del Fuego province</collectingRegion>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
195
data/33/6E/87/336E87952764FFAE9631DDA5FBF66D59.xml
Normal file
195
data/33/6E/87/336E87952764FFAE9631DDA5FBF66D59.xml
Normal file
|
|
@ -0,0 +1,195 @@
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<mods:title id="2F2B677D4EDED57BD1AA8A1D5D3E62A9">Systematics of the colour-polymorphic spider genus Cybaeolus, with comments on the phylogeny of the family Hahniidae (Araneae)</mods:title>
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<mods:affiliation id="40B6DE810B8BC340CBD649F635D42977">Western Carolina University, Cullowhee, NC 28723, United States</mods:affiliation>
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<mods:namePart id="FE970541CDE74367B6408E29ECCE6BE9">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="2AF35B0067637C6F058AB60C7930FD82">Museum of Nature Hamburg, Zoology, Leibniz-Institute for the Analysis of Biodiversity Change (LIB), Center for Taxonomy and Morphology, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="FCD046BF236CDF50EEC8C19D306BE0DD">Hormiga, Gustavo</mods:namePart>
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<mods:affiliation id="BFA704149F5BF0F1466218CA9D0CD6B8">Department of Biology, The George Washington University, Washington, DC 20052, United States</mods:affiliation>
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<mods:affiliation id="C64E760FD73E545B2C50C538D1F98CA5">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:affiliation id="EA6201B6D1263CDE177869FE20FCAF4E">Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’ - CONICET, Division of Arachnology, Av. Angel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:nameIdentifier id="1CC6095C30DC7268E48D9FC3432EE781" type="email">ramirez@macn.gov.ar</mods:nameIdentifier>
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<mods:date id="84BD63569336370F2B13A64FAE77D68C">2025</mods:date>
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<mods:number id="5DD9AFC06AF379F57A978A55AC466A82">2025-01-10</mods:number>
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<mods:number id="F96BB938A5065F733E3ADC8757F7448A">203</mods:number>
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<mods:url id="C64BDACB27773680BC6046EA7A69BB69">https://doi.org/10.1093/zoolinnean/zlae165</mods:url>
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<emphasis id="89B3EA912764FFAE9631DDA5FD826DFB" bold="true" box="[319,586,1739,1766]" pageId="12" pageNumber="13">
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Hahniinae
|
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<bibRefCitation id="DF564B722764FFAE96BDDDA2FD826DFB" author="Bertkau P" box="[435,586,1739,1766]" pageId="12" pageNumber="13" pagination="351 - 410" refId="ref32954" refString="Bertkau P. Versuch einer naturlichen Anordnung der Spinnen, nebst Bemerkungen zu einzelnen Gattungen. Archiv fur Naturgeschichte 1878; 44: 351 - 410." type="journal article" year="1878">Bertkau, 1878</bibRefCitation>
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Hahniinae
|
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<bibRefCitation id="DF564B722764FFAE97FBDD9CFE736C14" author="Bertkau P" box="[245,443,1778,1802]" pageId="12" pageNumber="13" pagination="351 - 410" refId="ref32954" refString="Bertkau P. Versuch einer naturlichen Anordnung der Spinnen, nebst Bemerkungen zu einzelnen Gattungen. Archiv fur Naturgeschichte 1878; 44: 351 - 410." type="journal article" year="1878">Bertkau 1878: 393</bibRefCitation>
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(
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genus
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<taxonomicName id="7CC74D002764FFAE9545DD9CFF7D6C34" authority="C. L. Koch, 1841" authorityName="C. L. Koch" authorityYear="1841" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="89B3EA912764FFAE9545DD9CFD5E6C14" box="[587,662,1778,1802]" italics="true" pageId="12" pageNumber="13">Hahnia</emphasis>
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C.L.
|
||||
<bibRefCitation id="DF564B722764FFAE95C6DD9CFF7D6C34" author="Koch CL" pageId="12" pageNumber="13" pagination="41 - 131" refId="ref33849" refString="Koch CL. Die Arachniden. Nurnberg Achter Band: C. H. Zeh'sche Buchhandlung, 1841, 41 - 131." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
|
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</taxonomicName>
|
||||
).
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</paragraph>
|
||||
<paragraph id="BB7836832764FFAE978FDC5FFE8C6C76" blockId="12.[129,777,1739,1897]" pageId="12" pageNumber="13">
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<taxonomicName id="7CC74D002764FFAE978FDC5FFE1A6C57" authority="Lehtinen 1967: 226" authorityName="Lehtinen" authorityYear="1967" box="[129,466,1841,1865]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cybaeolinae">
|
||||
Cybaeolinae
|
||||
<bibRefCitation id="DF564B722764FFAE9608DC5FFE1A6C57" author="Lehtinen PT" box="[262,466,1841,1865]" pageId="12" pageNumber="13" pagination="199 - 468" refId="ref33957" refString="Lehtinen PT. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 1967; 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967: 226</bibRefCitation>
|
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</taxonomicName>
|
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(
|
||||
<typeStatus id="647C88212764FFAE96D1DC5FFDD86C57" box="[479,528,1841,1865]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
genus
|
||||
<taxonomicName id="7CC74D002764FFAE9557DC5FFF7D6C76" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9557DC5FFD746C57" box="[601,700,1841,1865]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
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<bibRefCitation id="DF564B722764FFAE95CFDC5FFF7D6C76" author="Simon E" pageId="12" pageNumber="13" pagination="117 - 44" refId="ref34941" refString="Simon E. Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France 1884; 9: 117 - 44." type="journal article" year="1884">Simon, 1884</bibRefCitation>
|
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</taxonomicName>
|
||||
), synon. nov..
|
||||
</paragraph>
|
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|
||||
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<emphasis id="89B3EA912764FFAE96A0DCE8FE146C81" box="[430,476,1926,1951]" italics="true" pageId="12" pageNumber="13">Note</emphasis>
|
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</paragraph>
|
||||
<paragraph id="BB7836832764FFAE978FDCC2FBF66D59" blockId="12.[129,778,1964,2020]" lastBlockId="12.[825,1474,1301,1607]" pageId="12" pageNumber="13">
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<bibRefCitation id="DF564B722764FFAE978FDCC2FEE76CDA" author="Lehtinen PT" box="[129,303,1964,1988]" pageId="12" pageNumber="13" pagination="199 - 468" refId="ref33957" refString="Lehtinen PT. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 1967; 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
|
||||
originally included in
|
||||
<taxonomicName id="7CC74D002764FFAE9516DCC2FD5F6CDA" authorityName="Lehtinen" authorityYear="1967" box="[536,663,1964,1988]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cybaeolinae">Cybaeolinae</taxonomicName>
|
||||
the genera
|
||||
<taxonomicName id="7CC74D002764FFAE978CDCA5FEBF6CFA" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[130,375,1995,2020]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE978CDCA5FF2D6CFD" box="[130,229,1995,2019]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE97E1DCA5FEBF6CFA" author="Simon E" box="[239,375,1995,2020]" pageId="12" pageNumber="13" pagination="117 - 44" refId="ref34941" refString="Simon E. Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France 1884; 9: 117 - 44." type="journal article" year="1884">Simon, 1884</bibRefCitation>
|
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</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="7CC74D002764FFAE9687DCA5FD936CFD" authority="Roth, 1967" authorityName="Roth" authorityYear="1967" box="[393,603,1995,2019]" class="Arachnida" family="Hahniidae" genus="Lizarba" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9687DCA5FE1F6CFD" box="[393,471,1995,2019]" italics="true" pageId="12" pageNumber="13">Lizarba</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE96ECDCA5FD936CFD" author="Roth VD" box="[482,603,1995,2019]" pageId="12" pageNumber="13" pagination="297 - 345" refId="ref34792" refString="Roth VD. A review of the South American spiders of the family Agelenidae (Arachnida, Araneae). Bulletin of the American Museum of Natural History 1967; 134: 297 - 345." type="journal article" year="1967">Roth, 1967</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, and tentatively
|
||||
<taxonomicName id="7CC74D002764FFAE9437DE7BFB4F6E33" authority="Mello-Leitao, 1942" authorityName="Mello-Leitao" authorityYear="1942" box="[825,1159,1301,1325]" class="Arachnida" family="Hahniidae" genus="Austrohahnia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9437DE7BFC766E33" box="[825,958,1301,1325]" italics="true" pageId="12" pageNumber="13">Austrohahnia</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE94CADE7BFB4F6E33" author="Mello-Leitao CF" box="[964,1159,1301,1325]" pageId="12" pageNumber="13" pagination="381 - 426" refId="ref34125" refString="Mello-Leitao CF. Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (nueva serie, Zoologia) 1942; 2: 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. This last genus was tentatively included by
|
||||
<bibRefCitation id="DF564B722764FFAE94BADE5AFB5D6E53" author="Lehtinen PT" box="[948,1173,1332,1357]" pageId="12" pageNumber="13" pagination="199 - 468" refId="ref33957" refString="Lehtinen PT. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 1967; 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967: 217)</bibRefCitation>
|
||||
without having seen the
|
||||
<typeStatus id="647C88212764FFAE9281DE5BFA766E53" box="[1423,1470,1333,1357]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
, but he thought that it might be a synonym of
|
||||
<taxonomicName id="7CC74D002764FFAE921BDE3AFABE6E72" authorityName="Simon" authorityYear="1884" box="[1301,1398,1364,1388]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE921BDE3AFABE6E72" box="[1301,1398,1364,1388]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation id="DF564B722764FFAE928CDE3AFC706E95" author="Rubio GD & Lo-Man-Hung NF & Iuri HA" pageId="12" pageNumber="13" pagination="106 - 16" refId="ref34828" refString="Rubio GD, Lo-Man-Hung NF, Iuri HA. A revision of the Neotropical genus Austrohahnia Mello-Leitao (Araneae, Hahniidae). Zootaxa 2014; 3894: 106 - 16. http: // dx. doi. org / 10.11646 / zootaxa. 3894.1.8" type="journal article" year="2014">
|
||||
Rubio
|
||||
<emphasis id="89B3EA912764FFAE9437DE1AFCA06E95" box="[825,872,1395,1419]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||||
(2014)
|
||||
</bibRefCitation>
|
||||
revised the genus
|
||||
<taxonomicName id="7CC74D002764FFAE937ADE1DFA0A6E95" authority="Mello-Leitao, 1942" authorityName="Mello-Leitao" authorityYear="1942" box="[1140,1474,1395,1419]" class="Arachnida" family="Hahniidae" genus="Austrohahnia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE937ADE1DFB316E95" box="[1140,1273,1395,1419]" italics="true" pageId="12" pageNumber="13">Austrohahnia</emphasis>
|
||||
<bibRefCitation id="DF564B722764FFAE93F0DE1DFA0A6E95" author="Mello-Leitao CF" box="[1278,1474,1395,1419]" pageId="12" pageNumber="13" pagination="381 - 426" refId="ref34125" refString="Mello-Leitao CF. Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (nueva serie, Zoologia) 1942; 2: 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and transferred it to Hahniinae. Our phylogenetic analysis indicates that
|
||||
<taxonomicName id="7CC74D002764FFAE9492DEDCFC376ED4" authorityName="Simon" authorityYear="1884" box="[924,1023,1458,1482]" class="Arachnida" family="Hahniidae" genus="Cybaeolus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="89B3EA912764FFAE9492DEDCFC376ED4" box="[924,1023,1458,1482]" italics="true" pageId="12" pageNumber="13">Cybaeolus</emphasis>
|
||||
</taxonomicName>
|
||||
is nested within
|
||||
<taxonomicName id="7CC74D002764FFAE93A9DEDCFADC6ED4" authorityName="Bertkau" authorityYear="1878" box="[1191,1300,1458,1482]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Hahniidae</taxonomicName>
|
||||
, along with other genera typically considered hahniines, thus we synonymize
|
||||
<taxonomicName id="7CC74D002764FFAE9437DE9EFC706D16" authorityName="Lehtinen" authorityYear="1967" box="[825,952,1520,1544]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cybaeolinae">Cybaeolinae</taxonomicName>
|
||||
with the older name Hahniinae. For a future revision of the classification of the family,
|
||||
<taxonomicName id="7CC74D002764FFAE93C4DD7EFA816D36" authorityName="Lehtinen" authorityYear="1967" box="[1226,1353,1552,1576]" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cybaeolinae">Cybaeolinae</taxonomicName>
|
||||
remains an available subfamily name.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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