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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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Perameloidea
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<bibRefCitation author="Gray, J. E." box="[968,1085,771,795]" pageId="210" pageNumber="210" pagination="337 - 344" refId="ref207550" refString="Gray, J. E. 1825. An outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera apparently appertaining to each tribe. Annals of Philosophy 10 (2): 337 - 344." type="journal article" year="1825">Gray, 1825</bibRefCitation>
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CONTENTS:
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<taxonomicName authorityName="Gill" authorityYear="1872" box="[846,1016,821,845]" class="Mammalia" family="Chaeropodidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Chaeropodidae</taxonomicName>
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<taxonomicName id="4C4617C8F6E968FBD945FC9B199AFC8A" authorityName="Gill" authorityYear="1872" box="[846,1016,821,845]" class="Mammalia" family="Chaeropodidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Chaeropodidae</taxonomicName>
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||||
,
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<taxonomicName authorityName="Gray" authorityYear="1825" box="[1026,1162,821,845]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
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<taxonomicName id="4C4617C8F6E968FBDE09FC9B1EE8FC8A" authorityName="Gray" authorityYear="1825" box="[1026,1162,821,845]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
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, and
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||||
<taxonomicName authorityName="Bensley" authorityYear="1903" box="[684,852,854,878]" class="Mammalia" family="Thylacomyidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Thylacomyidae</taxonomicName>
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<taxonomicName id="4C4617C8F6E968FBD8A7FCF81936FCA9" authorityName="Bensley" authorityYear="1903" box="[684,852,854,878]" class="Mammalia" family="Thylacomyidae" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Thylacomyidae</taxonomicName>
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.
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</paragraph>
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<subSubSection pageId="210" pageNumber="210" type="description">
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<collectionCode box="[716,775,887,911]" pageId="210" pageNumber="210">STEM</collectionCode>
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AGE: 26.1 Mya (95%
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<collectionCode box="[1032,1091,887,910]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="210" pageNumber="210" type="Herbarium">HPD</collectionCode>
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: 20.7–31.8 Mya).
|
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</paragraph>
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<collectionCode box="[716,800,953,977]" pageId="210" pageNumber="210">CROWN</collectionCode>
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AGE: 20.0 Mya (95%
|
||||
<collectionCode box="[1038,1097,954,977]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="210" pageNumber="210" type="Herbarium">HPD</collectionCode>
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: 15.3–24.5 Mya).
|
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</paragraph>
|
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UNAMBIGUOUS CRANIODENTAL
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<collectionCode box="[684,748,1052,1076]" pageId="210" pageNumber="210">SYNAPOMORPHIES</collectionCode>
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: Nasals truncated anterior to lacrimals (char. 3: 0→1; ci = 0.333); lacrimal exposure on orbital rim forming a distinct crest (char. 8: 0→2; ci = 0.118); frontal and squamosal in contact on lateral aspect of braincase (char. 26: 0→1; ci = 0.071); palatine fenestrae present (char. 38: 0→1; ci = 0.071); posterior epitympanic sinus of squamosal present, enclosed laterally (char. 84: 0→1; ci = 0.071); and entocristid indistinct or absent (char. 176: 0→1; ci = 0.077).
|
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</paragraph>
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<paragraph id="8BF96C4BF6E968FBD8C7FCD9188EFC77" blockId="210.[684,1214,821,1738]" pageId="210" pageNumber="210">STEM AGE: 26.1 Mya (95% HPD: 20.7–31.8 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E968FBD8C7FC17188FFC35" blockId="210.[684,1214,821,1738]" pageId="210" pageNumber="210">CROWN AGE: 20.0 Mya (95% HPD: 15.3–24.5 Mya).</paragraph>
|
||||
<paragraph id="8BF96C4BF6E968FBD8C7FC5519B9FA99" blockId="210.[684,1214,821,1738]" pageId="210" pageNumber="210">UNAMBIGUOUS CRANIODENTAL SYNAPOMORPHIES: Nasals truncated anterior to lacrimals (char. 3: 0→1; ci = 0.333); lacrimal exposure on orbital rim forming a distinct crest (char. 8: 0→2; ci = 0.118); frontal and squamosal in contact on lateral aspect of braincase (char. 26: 0→1; ci = 0.071); palatine fenestrae present (char. 38: 0→1; ci = 0.071); posterior epitympanic sinus of squamosal present, enclosed laterally (char. 84: 0→1; ci = 0.071); and entocristid indistinct or absent (char. 176: 0→1; ci = 0.077).</paragraph>
|
||||
</subSubSection>
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<subSubSection id="C35C3FC0F6E968FAD8C7FAC91E05FC16" lastPageId="211" lastPageNumber="211" pageId="210" pageNumber="210" type="discussion">
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<paragraph id="8BF96C4BF6E968FBD8C7FAC91E4EF90D" blockId="210.[684,1214,821,1738]" pageId="210" pageNumber="210">
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COMMENTS: We follow
|
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|
||||
<collectingRegion id="4982A2A9F6E968FBD9DAFAC9199EFAB9" box="[977,1020,1383,1406]" country="Turkey" name="Van" pageId="210" pageNumber="210">Van</collectingRegion>
|
||||
Dyck and Strahan’s (2008)
|
||||
</bibRefCitation>
|
||||
use of Perameloidea to refer only to the peramelemorphian crown clade, as did
|
||||
<bibRefCitation author="Kear, B. P. & K. P. Aplin & M. Westerman" pageId="210" pageNumber="210" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al. (2016)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E968FBDE82FA04195EFA25" author="Kear, B. P. & K. P. Aplin & M. Westerman" pageId="210" pageNumber="210" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al. (2016)</bibRefCitation>
|
||||
. By contrast, other recent studies (e.g.,
|
||||
<bibRefCitation author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[747,1026,1515,1539]" pageId="210" pageNumber="210" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E968FBD8E0FA451E60F9C4" author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[747,1026,1515,1539]" pageId="210" pageNumber="210" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Warburton and Travouillon" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E968FBDE1BFA451917F9E3" author="Warburton and Travouillon" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
) have used Perameloidea to refer to the crown clade plus all taxa more closely related to it than to the fossil genus †
|
||||
<taxonomicName baseAuthorityName="Muirhead and Filan" baseAuthorityYear="1995" box="[1135,1205,1614,1638]" class="Mammalia" family="Yaralidae" genus="Yarala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[1135,1205,1614,1638]" italics="true" pageId="210" pageNumber="210">Yarala</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E968FBDE64F9E01ED7F9A1" baseAuthorityName="Muirhead and Filan" baseAuthorityYear="1995" box="[1135,1205,1614,1638]" class="Mammalia" family="Yaralidae" genus="Yarala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E968FBDE64F9E01ED7F9A1" box="[1135,1205,1614,1638]" italics="true" pageId="210" pageNumber="210">Yarala</emphasis>
|
||||
</taxonomicName>
|
||||
, which has been placed in the family †
|
||||
<taxonomicName box="[1109,1212,1647,1671]" class="Mammalia" family="Yaralidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Yaralidae</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6E968FBDE5EF9C11EDEF940" box="[1109,1212,1647,1671]" class="Mammalia" family="Yaralidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="210" pageNumber="210" phylum="Chordata" rank="family">Yaralidae</taxonomicName>
|
||||
and superfamily †Yaraloidea (
|
||||
<bibRefCitation author="Muirhead, J." box="[1025,1208,1680,1705]" pageId="210" pageNumber="210" pagination="512 - 523" refId="ref217438" refString="Muirhead, J. 2000. Yaraloidea (Marsupialia, Peramelemorphia), a new superfamily of marsupial and a description and analysis of the cranium of the Miocene of Yarala burchfieldi. Journal of Paleontology 74 (3): 512 - 523." type="journal article" year="2000">Muirhead, 2000</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E968FBDE0AF93E1EDAF96E" author="Muirhead, J." box="[1025,1208,1680,1705]" pageId="210" pageNumber="210" pagination="512 - 523" refId="ref217438" refString="Muirhead, J. 2000. Yaraloidea (Marsupialia, Peramelemorphia), a new superfamily of marsupial and a description and analysis of the cranium of the Miocene of Yarala burchfieldi. Journal of Paleontology 74 (3): 512 - 523." type="journal article" year="2000">Muirhead, 2000</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Warburton and Travouillon" box="[684,1057,1714,1738]" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E968FBD8A7F91C1E43F90D" author="Warburton and Travouillon" box="[684,1057,1714,1738]" firstAuthor="Warburton" pageId="210" pageNumber="210" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="211.[108,638,226,1638]" pageId="211" pageNumber="211">
|
||||
<paragraph id="8BF96C4BF6E868FADA87FF4C1B6BFC16" blockId="211.[108,638,226,1638]" pageId="211" pageNumber="211">
|
||||
Within Perameloidea, all our molecular (figs. 27–29) and total-evidence (figs. 32,
|
||||
<figureCitation box="[600,628,259,283]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="211" pageNumber="211">33</figureCitation>
|
||||
<figureCitation id="137D70CEF6E868FAD853FEAD1816FEDC" box="[600,628,259,283]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="211" pageNumber="211">33</figureCitation>
|
||||
) analyses support
|
||||
<taxonomicName authorityName="Gray" authorityYear="1825" box="[323,465,292,316]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6E868FADB48FE8A1BB3FEFB" authorityName="Gray" authorityYear="1825" box="[323,465,292,316]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Reid" authorityYear="1837" box="[536,636,292,315]" class="Mammalia" family="Thylacomyidae" genus="Macrotis" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[536,636,292,315]" italics="true" pageId="211" pageNumber="211">Macrotis</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FAD813FE8A1B50FE9A" authorityName="Reid" authorityYear="1837" class="Mammalia" family="Thylacomyidae" genus="Macrotis" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FAD813FE8A181EFEFC" box="[536,636,292,315]" italics="true" pageId="211" pageNumber="211">Macrotis</emphasis>
|
||||
(Thylacomyidae)
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName authorityName="Bensley" authorityYear="1903" box="[116,299,325,349]" class="Mammalia" family="Thylacomyidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Thylacomyidae</taxonomicName>
|
||||
) as sister taxa, with
|
||||
<taxonomicName authorityName="Ogilby" authorityYear="1838" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
as sister taxa, with
|
||||
<taxonomicName id="4C4617C8F6E868FAD833FEEB1B1CFEB9" authorityName="Ogilby" authorityYear="1838" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FAD833FEEB1AC8FEB9" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
(Chaeropodidae)
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName authorityName="Gill" authorityYear="1872" box="[191,374,358,382]" class="Mammalia" family="Chaeropodidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Chaeropodidae</taxonomicName>
|
||||
) the first perameloid family to diverge. This topology is congruent with most other recent molecular and totalevidence phylogenetic analyses (
|
||||
<bibRefCitation author="Meredith, R. W. & M. Westerman & M. S. Springer" pageId="211" pageNumber="211" pagination="1 - 20" refId="ref216532" refString="Meredith, R. W., M. Westerman, and M. S. Springer. 2008 a. A timescale and phylogeny for " bandicoots " (Peramelemorphia: Marsupialia) based on sequences for five nuclear genes. Molecular Phylogenetics and Evolution 47 (1): 1 - 20." type="journal article" year="2008">Meredith et al., 2008a</bibRefCitation>
|
||||
the first perameloid family to diverge. This topology is congruent with most other recent molecular and totalevidence phylogenetic analyses (
|
||||
<bibRefCitation id="EFD711BAF6E868FAD807FE671B67FDC5" author="Meredith, R. W. & M. Westerman & M. S. Springer" pageId="211" pageNumber="211" pagination="1 - 20" refId="ref216532" refString="Meredith, R. W., M. Westerman, and M. S. Springer. 2008 a. A timescale and phylogeny for " bandicoots " (Peramelemorphia: Marsupialia) based on sequences for five nuclear genes. Molecular Phylogenetics and Evolution 47 (1): 1 - 20." type="journal article" year="2008">Meredith et al., 2008a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Westerman" box="[279,564,490,514]" etAl="et al." firstAuthor="Westerman" pageId="211" pageNumber="211" pagination="97 - 108" refId="ref233182" refString="Westerman, M., et al. 2012. Phylogenetic relationships of living and recently extinct bandicoots based on nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 62 (1): 97 - 108." type="journal article" year="2012">Westerman et al., 2012</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB1CFE451856FDC5" author="Westerman" box="[279,564,490,514]" etAl="et al." firstAuthor="Westerman" pageId="211" pageNumber="211" pagination="97 - 108" refId="ref233182" refString="Westerman, M., et al. 2012. Phylogenetic relationships of living and recently extinct bandicoots based on nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 62 (1): 97 - 108." type="journal article" year="2012">Westerman et al., 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Kear, B. P. & K. P. Aplin & M. Westerman" pageId="211" pageNumber="211" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD84DFE451A91FDE4" author="Kear, B. P. & K. P. Aplin & M. Westerman" pageId="211" pageNumber="211" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al., 2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Travouillon and Phillips" box="[259,623,523,547]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="224 - 256" refId="ref230320" refString="Travouillon, K. J., and M. J. Phillips. 2018. Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species. Zootaxa 4378 (2): 224 - 256." type="journal article" year="2018">Travouillon and Phillips, 2018</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB08FDA5180DFDE4" author="Travouillon and Phillips" box="[259,623,523,547]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="224 - 256" refId="ref230320" refString="Travouillon, K. J., and M. J. Phillips. 2018. Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species. Zootaxa 4378 (2): 224 - 256." type="journal article" year="2018">Travouillon and Phillips, 2018</bibRefCitation>
|
||||
). However, the dated total-evidence analysis of
|
||||
<bibRefCitation author="Travouillon and Phillips" box="[140,497,589,613]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="224 - 256" refId="ref230320" refString="Travouillon, K. J., and M. J. Phillips. 2018. Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species. Zootaxa 4378 (2): 224 - 256." type="journal article" year="2018">Travouillon and Phillips (2018</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADA87FDE31B93FDA2" author="Travouillon and Phillips" box="[140,497,589,613]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="224 - 256" refId="ref230320" refString="Travouillon, K. J., and M. J. Phillips. 2018. Total evidence analysis of the phylogenetic relationships of bandicoots and bilbies (Marsupialia: Peramelemorphia): reassessment of two species and description of a new species. Zootaxa 4378 (2): 224 - 256." type="journal article" year="2018">Travouillon and Phillips (2018</bibRefCitation>
|
||||
: figs. 1E, 2) placed
|
||||
<taxonomicName authorityName="Ogilby" authorityYear="1838" box="[190,308,622,646]" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[190,308,622,646]" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADAB5FDC01B56FD41" authorityName="Ogilby" authorityYear="1838" box="[190,308,622,646]" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADAB5FDC01B56FD41" box="[190,308,622,646]" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
</taxonomicName>
|
||||
sister to
|
||||
<taxonomicName authorityName="Reid" authorityYear="1837" box="[419,514,623,646]" class="Mammalia" family="Thylacomyidae" genus="Macrotis" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[419,514,623,646]" italics="true" pageId="211" pageNumber="211">Macrotis</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADBA8FDC11860FD41" authorityName="Reid" authorityYear="1837" box="[419,514,623,646]" class="Mammalia" family="Thylacomyidae" genus="Macrotis" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADBA8FDC11860FD41" box="[419,514,623,646]" italics="true" pageId="211" pageNumber="211">Macrotis</emphasis>
|
||||
</taxonomicName>
|
||||
, the dated total-evidence analyses of
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al. (2021)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADBA4FD211AD4FD0E" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al. (2021)</bibRefCitation>
|
||||
placed
|
||||
<taxonomicName authorityName="Bensley" authorityYear="1903" box="[273,446,688,712]" class="Mammalia" family="Thylacomyidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Thylacomyidae</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6E868FADB1AFD1E1BDCFD0F" authorityName="Bensley" authorityYear="1903" box="[273,446,688,712]" class="Mammalia" family="Thylacomyidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Thylacomyidae</taxonomicName>
|
||||
outside
|
||||
<taxonomicName authorityName="Ogilby" authorityYear="1838" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FAD82EFD1E1AF0FD2E" authorityName="Ogilby" authorityYear="1838" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FAD82EFD1E1AF0FD2E" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName authorityName="Gray" authorityYear="1825" box="[183,326,722,746]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6E868FADABCFD7C1B24FD2D" authorityName="Gray" authorityYear="1825" box="[183,326,722,746]" class="Mammalia" family="Peramelidae" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="family">Peramelidae</taxonomicName>
|
||||
,
|
||||
<superScript attach="left" box="[331,349,719,736]" fontSize="7" pageId="211" pageNumber="211">31</superScript>
|
||||
<superScript id="7C33C103F6E868FADB40FD611B3FFD27" attach="left" box="[331,349,719,736]" fontSize="7" pageId="211" pageNumber="211">31</superScript>
|
||||
and the molecular phylogenetic analyses of
|
||||
<bibRefCitation author="Travouillon" box="[355,636,755,779]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al. (2019)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB68FD5D181EFCCC" author="Travouillon" box="[355,636,755,779]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al. (2019)</bibRefCitation>
|
||||
placed
|
||||
<taxonomicName authorityName="Ogilby" authorityYear="1838" box="[196,316,788,812]" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[196,316,788,812]" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADACFFCBA1B5EFCEB" authorityName="Ogilby" authorityYear="1838" box="[196,316,788,812]" class="Mammalia" family="Chaeropodidae" genus="Chaeropus" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADACFFCBA1B5EFCEB" box="[196,316,788,812]" italics="true" pageId="211" pageNumber="211">Chaeropus</emphasis>
|
||||
</taxonomicName>
|
||||
either sister to
|
||||
<taxonomicName authorityName="Desmarest" authorityYear="1817" box="[519,608,788,812]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[519,608,788,812]" italics="true" pageId="211" pageNumber="211">Isoodon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FAD80CFCBA1802FCEB" authorityName="Desmarest" authorityYear="1817" box="[519,608,788,812]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FAD80CFCBA1802FCEB" box="[519,608,788,812]" italics="true" pageId="211" pageNumber="211">Isoodon</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" box="[108,219,821,845]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[108,219,821,845]" italics="true" pageId="211" pageNumber="211">Perameles</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADA67FC9B1AB9FC8A" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" box="[108,219,821,845]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADA67FC9B1AB9FC8A" box="[108,219,821,845]" italics="true" pageId="211" pageNumber="211">Perameles</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName box="[257,371,822,845]" class="Mammalia" family="Peramelidae" genus="Peroryctes" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[257,371,822,845]" italics="true" pageId="211" pageNumber="211">Peroryctes</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADB0AFC981B11FC8A" box="[257,371,822,845]" class="Mammalia" family="Peramelidae" genus="Peroryctes" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADB0AFC981B11FC8A" box="[257,371,822,845]" italics="true" pageId="211" pageNumber="211">Peroryctes</emphasis>
|
||||
</taxonomicName>
|
||||
(in an undated maximum likelihood analysis;
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al., 2019</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADBADFCF81AC7FC48" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al., 2019</bibRefCitation>
|
||||
: fig. 15) or sister to
|
||||
<taxonomicName authorityName="Desmarest" authorityYear="1817" box="[403,490,887,911]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[403,490,887,911]" italics="true" pageId="211" pageNumber="211">Isoodon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADB98FCD91B88FC48" authorityName="Desmarest" authorityYear="1817" box="[403,490,887,911]" class="Mammalia" family="Peramelidae" genus="Isoodon" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADB98FCD91B88FC48" box="[403,490,887,911]" italics="true" pageId="211" pageNumber="211">Isoodon</emphasis>
|
||||
</taxonomicName>
|
||||
(in a Bayesian node-dating analysis;
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al., 2019</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADBADFC361AC7FC16" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1 - 56" refId="ref232735" refString="Warburton, N. M., K. J. Travouillon, and A. B. Camens. 2019. Skeletal atlas of the thylacine (Thylacinus cynocephalus). Palaeontologia Electronica 22.2.29 A: 1 - 56." type="book chapter" year="2019">Travouillon et al., 2019</bibRefCitation>
|
||||
: fig. 16).
|
||||
</paragraph>
|
||||
<paragraph blockId="211.[108,638,226,1638]" pageId="211" pageNumber="211">
|
||||
<paragraph id="8BF96C4BF6E868FADA87FC751BEDFA99" blockId="211.[108,638,226,1638]" pageId="211" pageNumber="211">
|
||||
We identified a number of unambiguous cranidodental synapomorphies as characterizing Perameloidea, although most of these are homoplastic. Perhaps the most notable of these is truncation of the nasals anterior to the lacrimals, a trait that is present in all known perameloids (and arose homoplastically in
|
||||
<taxonomicName baseAuthorityName="Gadow" baseAuthorityYear="1892" box="[108,224,1218,1241]" class="Mammalia" family="Notoryctidae" genus="Notoryctes" kingdom="Animalia" order="Notoryctemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[108,224,1218,1241]" italics="true" pageId="211" pageNumber="211">Notoryctes</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADA67FB6C1A82FB1E" baseAuthorityName="Gadow" baseAuthorityYear="1892" box="[108,224,1218,1241]" class="Mammalia" family="Notoryctidae" genus="Notoryctes" kingdom="Animalia" order="Notoryctemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADA67FB6C1A82FB1E" box="[108,224,1218,1241]" italics="true" pageId="211" pageNumber="211">Notoryctes</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Beck & Voss & Jansa" authorityYear="2022" box="[286,382,1218,1241]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[286,382,1218,1241]" italics="true" pageId="211" pageNumber="211">Potorous</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADB15FB6C1B1CFB1E" authorityName="Beck & Voss & Jansa" authorityYear="2022" box="[286,382,1218,1241]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADB15FB6C1B1CFB1E" box="[286,382,1218,1241]" italics="true" pageId="211" pageNumber="211">Potorous</emphasis>
|
||||
</taxonomicName>
|
||||
) but that is absent in nonperameloid peramelemorphians, all of which are “short-snouted” (
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1528 - 1546" refId="ref230485" refString="Travouillon, K. J., Y. Gurovich, R. M. D. Beck, and J. Muirhead. 2010. An exceptionally well-preserved short-snouted bandicoot (Marsupialia; Peramelemorphia) from Riversleigh's Oligo-Miocene deposits, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 30 (5): 1528 - 1546." type="journal article" year="2010">Travouillon et al., 2010</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADBA7FAAA1AC7FAFA" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="1528 - 1546" refId="ref230485" refString="Travouillon, K. J., Y. Gurovich, R. M. D. Beck, and J. Muirhead. 2010. An exceptionally well-preserved short-snouted bandicoot (Marsupialia; Peramelemorphia) from Riversleigh's Oligo-Miocene deposits, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 30 (5): 1528 - 1546." type="journal article" year="2010">Travouillon et al., 2010</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Travouillon" box="[185,256,1317,1341]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="153 - 168" refId="ref230601" refString="Travouillon, K. J., Y. Gurovich, M. Archer, S. J. Hand, and J. Muirhead. 2013 b. The genus Galadi: three new bandicoots (Marsupialia, Peramelemorphia) from Riversleigh's Miocene deposits, Northwestern Queensland, Australia. Journal of Vertebrate Paleontology 33 (1): 153 - 168." type="journal article" year="2013" yearSuffix="b">2013b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADAB2FA8B1B62FAFA" author="Travouillon" box="[185,256,1317,1341]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="153 - 168" refId="ref230601" refString="Travouillon, K. J., Y. Gurovich, M. Archer, S. J. Hand, and J. Muirhead. 2013 b. The genus Galadi: three new bandicoots (Marsupialia, Peramelemorphia) from Riversleigh's Miocene deposits, Northwestern Queensland, Australia. Journal of Vertebrate Paleontology 33 (1): 153 - 168." type="journal article" year="2013" yearSuffix="b">2013b</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Travouillon" box="[275,347,1317,1341]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="141 - 167" refId="ref230854" refString="Travouillon, K. J., M. Archer, S. J. Hand, and J. Muirhead. 2015 b. Sexually dimorphic bandicoots (Marsupialia: Peramelemorphia) from the Oligo-Miocene of Australia, first cranial ontogeny for fossil bandicoots and new species descriptions. Journal of Mammalian Evolution 22: 141 - 167." type="journal article" year="2015" yearSuffix="b">2015b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB18FA8B1B39FAFA" author="Travouillon" box="[275,347,1317,1341]" firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="141 - 167" refId="ref230854" refString="Travouillon, K. J., M. Archer, S. J. Hand, and J. Muirhead. 2015 b. Sexually dimorphic bandicoots (Marsupialia: Peramelemorphia) from the Oligo-Miocene of Australia, first cranial ontogeny for fossil bandicoots and new species descriptions. Journal of Mammalian Evolution 22: 141 - 167." type="journal article" year="2015" yearSuffix="b">2015b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Gurovich, Y. & K. J. Travouillon & R. M. D. Beck & J. Muirhead & M. Archer" box="[367,631,1317,1341]" pageId="211" pageNumber="211" pagination="265 - 290" refId="ref208146" refString="Gurovich, Y., K. J. Travouillon, R. M. D. Beck, J. Muirhead, and M. Archer. 2014. Biogeographical implications of a new mouse-sized fossil bandicoot (Marsupialia: Peramelemorphia) occupying a dasyurid-like ecological niche across Australia. Journal of Systematic Palaeontology 12 (3): 265 - 290." type="journal article" year="2014">Gurovich et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB64FA8B1815FAFA" author="Gurovich, Y. & K. J. Travouillon & R. M. D. Beck & J. Muirhead & M. Archer" box="[367,631,1317,1341]" pageId="211" pageNumber="211" pagination="265 - 290" refId="ref208146" refString="Gurovich, Y., K. J. Travouillon, R. M. D. Beck, J. Muirhead, and M. Archer. 2014. Biogeographical implications of a new mouse-sized fossil bandicoot (Marsupialia: Peramelemorphia) occupying a dasyurid-like ecological niche across Australia. Journal of Systematic Palaeontology 12 (3): 265 - 290." type="journal article" year="2014">Gurovich et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[108,387,1350,1374]" pageId="211" pageNumber="211" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADA67FAE81BE1FA99" author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[108,387,1350,1374]" pageId="211" pageNumber="211" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="211.[108,638,226,1638]" lastBlockId="211.[684,1212,226,977]" pageId="211" pageNumber="211">
|
||||
<paragraph id="8BF96C4BF6E868FADA86FAC91E05FC16" blockId="211.[108,638,226,1638]" lastBlockId="211.[684,1212,226,977]" pageId="211" pageNumber="211">
|
||||
The oldest putative perameloid is †
|
||||
<emphasis italics="true" pageId="211" pageNumber="211">Bulbadon warburtonae</emphasis>
|
||||
<emphasis id="B932B059F6E868FAD818FAC91A9EFA67" italics="true" pageId="211" pageNumber="211">Bulbadon warburtonae</emphasis>
|
||||
from the late Oligocene Ditjimanka Local Fauna (Faunal Zone B) of the Etadunna Formation, which is known from a single partial mandible, and has been tentatively identified as a thylacomyid (
|
||||
<bibRefCitation author="Travouillon" box="[367,625,1548,1572]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al., 2021</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FADB64F9A21813F9E3" author="Travouillon" box="[367,625,1548,1572]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al., 2021</bibRefCitation>
|
||||
). A best-fit age-model of paleomagnetic data indicates that the Etadunna Formation spans 23.6–26.1 Mya (
|
||||
<bibRefCitation author="Metzger, C. A. & G. J. Retallack" box="[868,1201,226,250]" pageId="211" pageNumber="211" pagination="871 - 885" refId="ref216925" refString="Metzger, C. A., and G. J. Retallack. 2010. Paleosol record of Neogene climate change in the Australian outback. Australian Journal of Earth Sciences 57: 871 - 885." type="journal article" year="2010">Metzger and Retallack, 2010</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD96FFF4C1ED3FF3D" author="Metzger, C. A. & G. J. Retallack" box="[868,1201,226,250]" pageId="211" pageNumber="211" pagination="871 - 885" refId="ref216925" refString="Metzger, C. A., and G. J. Retallack. 2010. Paleosol record of Neogene climate change in the Australian outback. Australian Journal of Earth Sciences 57: 871 - 885." type="journal article" year="2010">Metzger and Retallack, 2010</bibRefCitation>
|
||||
), so, if †
|
||||
<emphasis box="[763,868,259,283]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
<emphasis id="B932B059F6E868FAD8F0FEAD1906FEDC" box="[763,868,259,283]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
is a thylacomyid, then it predates our estimate for the crown age of Perameloidea. However, †
|
||||
<emphasis box="[929,1034,325,349]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
<emphasis id="B932B059F6E868FAD9AAFEEB1E68FE9A" box="[929,1034,325,349]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
did not form a clade with other thylacomyids in the dated total-evidence analyses of
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al. (2021)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD9E6FE291894FE07" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="109 - 125" refId="ref231075" refString="Travouillon, K. J., R. M. D. Beck, and J. A. Case. 2021. Upper Oligocene - lower-Middle Miocene peramelemorphians from the Etadunna, Namba and Wipajiri formations of South Australia. Alcheringa: an Australasian Journal of Palaeontology 45 (1): 109 - 125." type="journal article" year="2021">Travouillon et al. (2021)</bibRefCitation>
|
||||
and a lack of resolution meant that a position for †
|
||||
<emphasis box="[847,954,457,481]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
<emphasis id="B932B059F6E868FAD944FE6719D8FE26" box="[847,954,457,481]" italics="true" pageId="211" pageNumber="211">Bulbadon</emphasis>
|
||||
outside Perameloidea could not be ruled out in their analyses. Two taxa from the middle Miocene of Riversleigh World Heritage Area, both known only from isolated teeth, may also be early perameloids (
|
||||
<bibRefCitation author="Travouillon" box="[691,988,622,647]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">Travouillon et al., 2014a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD8B8FDC019BEFD40" author="Travouillon" box="[691,988,622,647]" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">Travouillon et al., 2014a</bibRefCitation>
|
||||
): †
|
||||
<taxonomicName authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[1026,1212,622,646]" class="Mammalia" family="Peramelidae" genus="Crash" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="species">
|
||||
<emphasis box="[1026,1212,622,646]" italics="true" pageId="211" pageNumber="211">Crash bandicoot</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADE09FDC01EDEFD41" authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[1026,1212,622,646]" class="Mammalia" family="Peramelidae" genus="Crash" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="species" species="bandicoot">
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<emphasis id="B932B059F6E868FADE09FDC01EDEFD41" box="[1026,1212,622,646]" italics="true" pageId="211" pageNumber="211">Crash bandicoot</emphasis>
|
||||
</taxonomicName>
|
||||
(which appears to be a peramelid), and †
|
||||
<taxonomicName authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" class="Mammalia" family="Thylacomyidae" genus="Liyamayi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="species">
|
||||
<emphasis italics="true" pageId="211" pageNumber="211">Liyamayi dayi</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADE8FFD3E197FFD0F" authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" class="Mammalia" family="Thylacomyidae" genus="Liyamayi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="species" species="dayi">
|
||||
<emphasis id="B932B059F6E868FADE8FFD3E197FFD0F" italics="true" pageId="211" pageNumber="211">Liyamayi dayi</emphasis>
|
||||
</taxonomicName>
|
||||
(which was described as a thylacomyid, although its position in published phylogenies is highly labile;
|
||||
<bibRefCitation author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">
|
||||
<bibRefCitation id="EFD711BAF6E868FAD9F9FD5D1893FCEB" author="Travouillon" etAl="et al." firstAuthor="Travouillon" pageId="211" pageNumber="211" pagination="375 - 382" refId="ref230665" refString="Travouillon, K. J., S. J. Hand, M. Archer, and K. H. Black. 2014 a. Earliest modern bandicoot and bilby (Marsupialia, Peramelidae and Thylacomyidae) from the Miocene of the Riversleigh World Heritage Area, northwestern Queensland, Australia. Journal of Vertebrate Paleontology 34 (2): 375 - 382." type="journal article" year="2014" yearSuffix="a">
|
||||
Travouillon et al., 2014
|
||||
<bibRefCitation author="a" box="[741,837,788,812]" firstAuthor="a" pageId="211" pageNumber="211" pagination="1408" refId="ref237143" refString="Yates, A. M. 2015 b. New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia. PeerJ 3: e 1408." type="journal article" year="2015" yearSuffix="b">
|
||||
a, 2015
|
||||
<bibRefCitation author="b" box="[823,907,788,812]" firstAuthor="b" pageId="211" pageNumber="211" pagination="1408" refId="ref237143" refString="Yates, A. M. 2015 b. New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia. PeerJ 3: e 1408." type="journal article" year="2021" yearSuffix="b">b, 2021</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD8EEFCBA1893FCEB" author="a" box="[741,753,788,812]" firstAuthor="a" pageId="211" pageNumber="211" pagination="1408" refId="ref237143" refString="Yates, A. M. 2015 b. New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia. PeerJ 3: e 1408." type="journal article" year="2015" yearSuffix="b">a</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
, 2015
|
||||
<bibRefCitation id="EFD711BAF6E868FAD93CFCBA19E9FCEB" author="b" box="[823,907,788,812]" firstAuthor="b" pageId="211" pageNumber="211" pagination="1408" refId="ref237143" refString="Yates, A. M. 2015 b. New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia. PeerJ 3: e 1408." type="journal article" year="2021" yearSuffix="b">b, 2021</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[921,1208,788,812]" pageId="211" pageNumber="211" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD992FCBA1EDAFCEB" author="Chamberlain, P. M. & K. J. Travouillon & M. Archer & S. J. Hand" box="[921,1208,788,812]" pageId="211" pageNumber="211" pagination="197 - 206" refId="ref197024" refString="Chamberlain, P. M., K. J. Travouillon, M. Archer, and S. J. Hand. 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa: an Australasian Journal of Palaeontology 40 (2): 197 - 206." type="journal article" year="2015">Chamberlain et al., 2015</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Kear, B. P. & K. P. Aplin & M. Westerman" box="[684,873,821,845]" pageId="211" pageNumber="211" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6E868FAD8A7FC9B190BFC8A" author="Kear, B. P. & K. P. Aplin & M. Westerman" box="[684,873,821,845]" pageId="211" pageNumber="211" pagination="37537" refId="ref210965" refString="Kear, B. P., K. P. Aplin, and M. Westerman. 2016. Bandicoot fossils and DNA elucidate lineage antiquity amongst xeric-adapted Australasian marsupials. Scientific Reports 6: 37537." type="journal article" year="2016">Kear et al., 2016</bibRefCitation>
|
||||
). If †
|
||||
<taxonomicName authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[932,995,821,845]" class="Mammalia" family="Peramelidae" genus="Crash" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[932,995,821,845]" italics="true" pageId="211" pageNumber="211">Crash</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FAD9AFFC9B1981FC8A" authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[932,995,821,845]" class="Mammalia" family="Peramelidae" genus="Crash" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FAD9AFFC9B1981FC8A" box="[932,995,821,845]" italics="true" pageId="211" pageNumber="211">Crash</emphasis>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[1068,1170,822,845]" class="Mammalia" family="Thylacomyidae" genus="Liyamayi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[1068,1170,822,845]" italics="true" pageId="211" pageNumber="211">Liyamayi</emphasis>
|
||||
<taxonomicName id="4C4617C8F6E868FADE27FC981EF0FC8A" authorityName="Travouillon, Hand, Archer & Black" authorityYear="2014" box="[1068,1170,822,845]" class="Mammalia" family="Thylacomyidae" genus="Liyamayi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Peramelemorphia" pageId="211" pageNumber="211" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6E868FADE27FC981EF0FC8A" box="[1068,1170,822,845]" italics="true" pageId="211" pageNumber="211">Liyamayi</emphasis>
|
||||
</taxonomicName>
|
||||
are indeed early perameloids, then their middle Miocene age is broadly congruent with our middle to early Miocene age estimate for the onset of diversfication of Perameloidea.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,218 +1,206 @@
|
|||
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<treatment id="03EFDD5DF6F268E3DABAFC29182BFD41" ID-DOI="http://doi.org/10.5281/zenodo.6974209" ID-Zenodo-Dep="6974209" LSID="urn:lsid:plazi:treatment:03EFDD5DF6F268E3DABAFC29182BFD41" httpUri="http://treatment.plazi.org/id/03EFDD5DF6F268E3DABAFC29182BFD41" lastPageId="202" lastPageNumber="201" pageId="201" pageNumber="201">
|
||||
<subSubSection id="C35C3FC0F6F268E0DABAFC291858FC67" box="[177,570,903,928]" pageId="201" pageNumber="201" type="nomenclature">
|
||||
<paragraph id="8BF96C4BF6F268E0DABAFC291858FC67" blockId="201.[177,570,903,928]" box="[177,570,903,928]" pageId="201" pageNumber="201">
|
||||
<heading id="D0B1DB27F6F268E0DABAFC291858FC67" box="[177,570,903,928]" centered="true" fontSize="10" level="2" pageId="201" pageNumber="201" reason="4">
|
||||
<taxonomicName id="4C4617C8F6F268E0DABAFC291858FC67" authority="Archer et al., 1988" authorityName="Archer" authorityYear="1988" box="[177,570,903,928]" genus="Yalkaparidontia" kingdom="Animalia" pageId="201" pageNumber="201" rank="genus">
|
||||
†Yalkaparidontia
|
||||
<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[369,570,903,928]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DB7AFC291858FC67" author="Archer, M. & S. Hand & H. Godthelp" box="[369,570,903,928]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection box="[140,435,953,977]" pageId="201" pageNumber="201" type="synonymic_list">
|
||||
<paragraph blockId="201.[108,638,953,1738]" box="[140,435,953,977]" pageId="201" pageNumber="201">
|
||||
<subSubSection id="C35C3FC0F6F268E0DA87FC171BD1FC16" box="[140,435,953,977]" pageId="201" pageNumber="201" type="synonymic_list">
|
||||
<paragraph id="8BF96C4BF6F268E0DA87FC171BD1FC16" blockId="201.[108,638,953,1738]" box="[140,435,953,977]" pageId="201" pageNumber="201">
|
||||
CONTENTS: †
|
||||
<emphasis box="[285,435,953,977]" italics="true" pageId="201" pageNumber="201">
|
||||
<taxonomicName box="[285,432,953,977]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">Yalkaparidon</taxonomicName>
|
||||
<emphasis id="B932B059F6F268E0DB16FC171BD1FC16" box="[285,435,953,977]" italics="true" pageId="201" pageNumber="201">
|
||||
<taxonomicName id="4C4617C8F6F268E0DB16FC171BD2FC16" box="[285,432,953,977]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">Yalkaparidon</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="201" pageNumber="201" type="description">
|
||||
<paragraph blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">
|
||||
<collectionCode box="[140,199,986,1010]" pageId="201" pageNumber="201">STEM</collectionCode>
|
||||
AGE: 39.9 Mya (95%
|
||||
<collectionCode box="[456,515,987,1010]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="201" pageNumber="201" type="Herbarium">HPD</collectionCode>
|
||||
: 30.7–49.7 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="201.[108,638,953,1738]" box="[140,457,1052,1076]" pageId="201" pageNumber="201">
|
||||
<collectionCode box="[140,224,1052,1076]" pageId="201" pageNumber="201">CROWN</collectionCode>
|
||||
AGE: Not applicable.
|
||||
</paragraph>
|
||||
<paragraph blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">
|
||||
UNAMBIGUOUS CRANIODENTAL
|
||||
<collectionCode box="[108,171,1119,1143]" pageId="201" pageNumber="201">AUTAPOMORPHIES</collectionCode>
|
||||
: Postglenoid process greatly reduced or absent (char. 75: 0→1; ci = 0.200); mandible usually with one mental foramen (char. 98: 1→0; ci = 0.063); second and third upper incisors entirely lacking enamel (char. 105: 0→1; ci = 1.000); first upper premolar absent (char. 114: 0→1; ci = 0.200); protocone absent (char. 141: 0→1; ci = 1.000); talonid greatly reduced or absent (char. 166: 0→1; ci = 0.500).
|
||||
</paragraph>
|
||||
<subSubSection id="C35C3FC0F6F268E0DA87FC741B34FAB8" pageId="201" pageNumber="201" type="description">
|
||||
<paragraph id="8BF96C4BF6F268E0DA87FC741ACEFBD4" blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">STEM AGE: 39.9 Mya (95% HPD: 30.7–49.7 Mya).</paragraph>
|
||||
<paragraph id="8BF96C4BF6F268E0DA87FBB21BABFBF3" blockId="201.[108,638,953,1738]" box="[140,457,1052,1076]" pageId="201" pageNumber="201">CROWN AGE: Not applicable.</paragraph>
|
||||
<paragraph id="8BF96C4BF6F268E0DA87FB931B34FAB8" blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">UNAMBIGUOUS CRANIODENTAL AUTAPOMORPHIES: Postglenoid process greatly reduced or absent (char. 75: 0→1; ci = 0.200); mandible usually with one mental foramen (char. 98: 1→0; ci = 0.063); second and third upper incisors entirely lacking enamel (char. 105: 0→1; ci = 1.000); first upper premolar absent (char. 114: 0→1; ci = 0.200); protocone absent (char. 141: 0→1; ci = 1.000); talonid greatly reduced or absent (char. 166: 0→1; ci = 0.500).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201" type="discussion">
|
||||
<paragraph blockId="201.[108,638,953,1738]" lastBlockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
|
||||
<subSubSection id="C35C3FC0F6F268E3DA87FA26182BFD41" lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201" type="discussion">
|
||||
<paragraph id="8BF96C4BF6F268E0DA87FA261958FD2D" blockId="201.[108,638,953,1738]" lastBlockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
|
||||
COMMENTS: The peculiar †
|
||||
<taxonomicName box="[458,607,1416,1440]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[458,607,1416,1440]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0DBC1FA26183DFA67" box="[458,607,1416,1440]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0DBC1FA26183DFA67" box="[458,607,1416,1440]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
is known only from two described species (†
|
||||
<taxonomicName class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
|
||||
<emphasis italics="true" pageId="201" pageNumber="201">Y. coheni</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D86CFA041AD1FA25" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species" species="coheni">
|
||||
<emphasis id="B932B059F6F268E0D86CFA041AD1FA25" italics="true" pageId="201" pageNumber="201">Y. coheni</emphasis>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<taxonomicName box="[256,351,1483,1506]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
|
||||
<emphasis box="[256,351,1483,1506]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0DB0BFA651B3DFA25" box="[256,351,1483,1506]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species" species="jonesi">
|
||||
<emphasis id="B932B059F6F268E0DB0BFA651B3DFA25" box="[256,351,1483,1506]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
|
||||
</taxonomicName>
|
||||
) from late Oligocene to middle Miocene sites at Riversleigh World Heritage Area in northern
|
||||
<collectingCountry box="[370,471,1548,1572]" name="Australia" pageId="201" pageNumber="201">Australia</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F268E0DB79F9A21BB5F9E3" box="[370,471,1548,1572]" name="Australia" pageId="201" pageNumber="201">Australia</collectingCountry>
|
||||
(
|
||||
<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DBECF9A21AC7F982" author="Archer, M. & S. Hand & H. Godthelp" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[176,360,1581,1605]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DABBF9831B0AF982" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[176,360,1581,1605]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
). †
|
||||
<taxonomicName box="[389,612,1581,1605]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
|
||||
<emphasis box="[389,612,1581,1605]" italics="true" pageId="201" pageNumber="201">Yalkaparidon coheni</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0DB8EF9831806F982" box="[389,612,1581,1605]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species" species="coheni">
|
||||
<emphasis id="B932B059F6F268E0DB8EF9831806F982" box="[389,612,1581,1605]" italics="true" pageId="201" pageNumber="201">Yalkaparidon coheni</emphasis>
|
||||
</taxonomicName>
|
||||
is by far the better known of the two species, being represented by partial cranial material, whereas †
|
||||
<taxonomicName box="[221,314,1681,1704]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
|
||||
<emphasis box="[221,314,1681,1704]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0DAD6F93F1B58F96F" box="[221,314,1681,1704]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species" species="jonesi">
|
||||
<emphasis id="B932B059F6F268E0DAD6F93F1B58F96F" box="[221,314,1681,1704]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
|
||||
</taxonomicName>
|
||||
is known only from a fragmentary right mandible (
|
||||
<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[406,631,1714,1738]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DB9DF91C1815F90D" author="Archer, M. & S. Hand & H. Godthelp" box="[406,631,1714,1738]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,897,226,250]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D8A7FF4C19E3FF3D" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,897,226,250]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
). †
|
||||
<taxonomicName box="[937,1088,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[937,1088,226,250]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D9A2FF4C1E22FF3D" box="[937,1088,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D9A2FF4C1E22FF3D" box="[937,1088,226,250]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
combines some relatively plesiomorphic features of the skull (particularly of the basicranium) with a highly derived dentition that includes an enlarged “gliriform” anteriormost lower incisor and zalambdodont molars (
|
||||
<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[996,1209,391,415]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D9EFFE291EDBFE58" author="Archer, M. & S. Hand & H. Godthelp" box="[996,1209,391,415]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D." box="[686,818,424,448]" pageId="201" pageNumber="201" pagination="1 - 17" refId="ref192600" refString="Beck, R. M. D. 2009. Was the Oligo-Miocene Australian metatherian Yalkaparidon a ' mammalian woodpecker'? Biological Journal of the Linnean Society 97: 1 - 17." type="journal article" year="2009">Beck, 2009</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D8A5FE061950FE07" author="Beck, R. M. D." box="[686,818,424,448]" pageId="201" pageNumber="201" pagination="1 - 17" refId="ref192600" refString="Beck, R. M. D. 2009. Was the Oligo-Miocene Australian metatherian Yalkaparidon a ' mammalian woodpecker'? Biological Journal of the Linnean Society 97: 1 - 17." type="journal article" year="2009">Beck, 2009</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[834,1039,424,448]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D949FE061E6DFE07" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[834,1039,424,448]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
). Of the other unambiguous craniodental apomorphies identified here, extreme reduction of the postglenoid process likely reflects extensive anteroposterior movement of the lower jaw (a trait also seen, for example, in members of the placental clade Glires;
|
||||
<bibRefCitation author="Cope, E. D." box="[738,877,623,647]" pageId="201" pageNumber="201" pagination="3 - 11" refId="ref198605" refString="Cope, E. D. 1888. The mechanical causes of the origin of the dentition of the Rodentia. American Naturalist 22: 3 - 11." type="journal article" year="1888">Cope, 1888</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D8E9FDC1190FFD40" author="Cope, E. D." box="[738,877,623,647]" pageId="201" pageNumber="201" pagination="3 - 11" refId="ref198605" refString="Cope, E. D. 1888. The mechanical causes of the origin of the dentition of the Rodentia. American Naturalist 22: 3 - 11." type="journal article" year="1888">Cope, 1888</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Druzinsky, R. E." box="[895,1093,622,646]" pageId="201" pageNumber="201" pagination="323 - 349" refId="ref200607" refString="Druzinsky, R. E. 2015. The oral apparatus of rodents: variations on the theme of a gnawing machine. In P. G. Cox and L. Hautier (editors), Evolution of the rodents: 323 - 349. Cambridge: Cambridge University Press." type="book chapter" year="2015">Druzinsky, 2015</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D974FDC01E27FD41" author="Druzinsky, R. E." box="[895,1093,622,646]" pageId="201" pageNumber="201" pagination="323 - 349" refId="ref200607" refString="Druzinsky, R. E. 2015. The oral apparatus of rodents: variations on the theme of a gnawing machine. In P. G. Cox and L. Hautier (editors), Evolution of the rodents: 323 - 349. Cambridge: Cambridge University Press." type="book chapter" year="2015">Druzinsky, 2015</bibRefCitation>
|
||||
), whereas absence of enamel from I2 and I3 is an autapomorphy that we have not seen in any other metatherian.
|
||||
</paragraph>
|
||||
<paragraph blockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
|
||||
<paragraph id="8BF96C4BF6F268E0D8C7FD5D190FFB92" blockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
|
||||
Based on its unusual combination of features, we agree with
|
||||
<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[929,1159,788,812]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al. (1988)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D9AAFCBA1EE5FCEB" author="Archer, M. & S. Hand & H. Godthelp" box="[929,1159,788,812]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al. (1988)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,884,821,845]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D8A7FC9B1916FC8A" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,884,821,845]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
that †
|
||||
<taxonomicName box="[960,1106,821,845]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[960,1106,821,845]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D9CBFC9B1E30FC8A" box="[960,1106,821,845]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D9CBFC9B1E30FC8A" box="[960,1106,821,845]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
warrants classification within its own family and order. †
|
||||
<taxonomicName box="[698,844,887,911]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[698,844,887,911]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D8B1FCD9192EFC48" box="[698,844,887,911]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D8B1FCD9192EFC48" box="[698,844,887,911]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
is the only definitive member of this order currently known, although the enigmatic †
|
||||
<taxonomicName box="[767,1056,953,977]" class="Mammalia" family="Yingabalanaridae" genus="Yingabalanara" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
|
||||
<emphasis box="[767,1056,953,977]" italics="true" pageId="201" pageNumber="201">Yingabalanara richardsoni</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D8F4FC171E42FC16" box="[767,1056,953,977]" class="Mammalia" family="Yingabalanaridae" genus="Yingabalanara" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species" species="richardsoni">
|
||||
<emphasis id="B932B059F6F268E0D8F4FC171E42FC16" box="[767,1056,953,977]" italics="true" pageId="201" pageNumber="201">Yingabalanara richardsoni</emphasis>
|
||||
</taxonomicName>
|
||||
(known from two lower molars from the early Miocene of Riversleigh;
|
||||
<bibRefCitation author="Archer, M. & R. G. Every & H. Godthelp & S. Hand & K. Scally" box="[823,1033,1019,1043]" pageId="201" pageNumber="201" pagination="193 - 202" refId="ref190672" refString="Archer, M., R. G. Every, H. Godthelp, S. Hand, and K. Scally. 1990. Yingabalanaridae, a new family of enigmatic mammals from Tertiary deposits of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 28: 193 - 202." type="journal article" year="1990">Archer et al., 1990</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D93CFC551E6BFBD4" author="Archer, M. & R. G. Every & H. Godthelp & S. Hand & K. Scally" box="[823,1033,1019,1043]" pageId="201" pageNumber="201" pagination="193 - 202" refId="ref190672" refString="Archer, M., R. G. Every, H. Godthelp, S. Hand, and K. Scally. 1990. Yingabalanaridae, a new family of enigmatic mammals from Tertiary deposits of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 28: 193 - 202." type="journal article" year="1990">Archer et al., 1990</bibRefCitation>
|
||||
) may be a dentally plesiomorphic yalkaparidontian (see
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 155</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DE8DFBB21903FB92" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 155</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="201.[684,1214,226,1738]" lastBlockId="202.[108,638,226,646]" lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201">
|
||||
<paragraph id="8BF96C4BF6F268E3D8C6FBF1182BFD41" blockId="201.[684,1214,226,1738]" lastBlockId="202.[108,638,226,646]" lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201">
|
||||
Our undated (fig. 32) and dated (
|
||||
<figureCitation box="[1127,1205,1118,1143]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="201" pageNumber="201">fig. 33</figureCitation>
|
||||
<figureCitation id="137D70CEF6F268E0DE6CFBF01ED7FBB0" box="[1127,1205,1118,1143]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="201" pageNumber="201">fig. 33</figureCitation>
|
||||
) total-evidence analyses both recover a clade that unites †
|
||||
<taxonomicName box="[776,923,1184,1208]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[776,923,1184,1208]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D903FB0E19F9FB7F" box="[776,923,1184,1208]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D903FB0E19F9FB7F" box="[776,923,1184,1208]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
with paucituberculatans. However, isolated tarsals tentatively referred to †
|
||||
<taxonomicName box="[697,841,1251,1275]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[697,841,1251,1275]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D8B2FB4D192BFB3C" box="[697,841,1251,1275]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D8B2FB4D192BFB3C" box="[697,841,1251,1275]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[879,1067,1251,1275]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D964FB4D1E49FB3C" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[879,1067,1251,1275]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
show greater derived similarities to australidelphians than to paucituberculatans and Beck et al.’s (2014) accompanying phylogenetic analyses also supported australidelphian affinities for this taxon (see also
|
||||
<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[795,995,1416,1440]" pageId="201" pageNumber="201" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">Beck et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D910FA261981FA67" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[795,995,1416,1440]" pageId="201" pageNumber="201" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">Beck et al., 2016</bibRefCitation>
|
||||
). Nevertheless, the subsequent phylogenetic analyses of
|
||||
<bibRefCitation author="Beck, R. M. D." pageId="201" pageNumber="201" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck (2017a)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DE8FFA071960FA25" author="Beck, R. M. D." pageId="201" pageNumber="201" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck (2017a)</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation author="Zimicz and Goin" box="[837,1127,1482,1506]" firstAuthor="Zimicz" pageId="201" pageNumber="201" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin (2020)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0D94EFA641E05FA25" author="Zimicz and Goin" box="[837,1127,1482,1506]" firstAuthor="Zimicz" pageId="201" pageNumber="201" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin (2020)</bibRefCitation>
|
||||
, which included †
|
||||
<taxonomicName box="[813,963,1515,1539]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[813,963,1515,1539]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F268E0D926FA4519A1F9C4" box="[813,963,1515,1539]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F268E0D926FA4519A1F9C4" box="[813,963,1515,1539]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
and used character scores from these referred tarsals also placed this taxon in a clade with paucituberculatans (including argyrolagids, a fossil group not included in our study; but see
|
||||
<bibRefCitation author="Abello, M. A. & A. M. Candela" pageId="201" pageNumber="201" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F268E0DE08F9C11945F96E" author="Abello, M. A. & A. M. Candela" pageId="201" pageNumber="201" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
|
||||
). Although this clade is worthy of further investigation, we suspect that placement of †
|
||||
<taxonomicName box="[149,292,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[149,292,226,250]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F168E3DA9EFF4C1B46FF3D" box="[149,292,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F168E3DA9EFF4C1B46FF3D" box="[149,292,226,250]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
with paucituberculatans largely reflects convergent acquisitions of a gliriform lower incisor. Current evidence (including the analyses presented here) indicates that diprotodontians evolved a similar lower incisor independently of paucituberculatans, so it is plausible that †
|
||||
<taxonomicName box="[174,319,424,448]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[174,319,424,448]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F168E3DAA5FE061B5DFE07" box="[174,319,424,448]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F168E3DAA5FE061B5DFE07" box="[174,319,424,448]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
||||
</taxonomicName>
|
||||
represents a third origin of this tooth
|
||||
<typeStatus box="[226,276,458,482]" pageId="202" pageNumber="202">type</typeStatus>
|
||||
<typeStatus id="54FDD2E9F6F168E3DAE9FE641B76FE25" box="[226,276,458,482]" pageId="202" pageNumber="202">type</typeStatus>
|
||||
. Based on our results and those of other recent studies (
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[375,564,490,514]" pageId="202" pageNumber="202" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F168E3DB7CFE441856FDC5" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[375,564,490,514]" pageId="202" pageNumber="202" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
||||
,
|
||||
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<bibRefCitation id="EFD711BAF6F168E3D84AFE44181AFDC5" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[577,632,490,514]" pageId="202" pageNumber="202" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D." box="[108,244,523,547]" pageId="202" pageNumber="202" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck, 2017a</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F168E3DA67FDA51A96FDE4" author="Beck, R. M. D." box="[108,244,523,547]" pageId="202" pageNumber="202" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck, 2017a</bibRefCitation>
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;
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<bibRefCitation author="Abello, M. A. & A. M. Candela" box="[256,546,523,547]" pageId="202" pageNumber="202" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F168E3DB0BFDA51840FDE4" author="Abello, M. A. & A. M. Candela" box="[256,546,523,547]" pageId="202" pageNumber="202" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
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;
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<bibRefCitation author="Zimicz and Goin" firstAuthor="Zimicz" pageId="202" pageNumber="202" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin, 2020</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F168E3D824FDA51B47FD83" author="Zimicz and Goin" firstAuthor="Zimicz" pageId="202" pageNumber="202" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin, 2020</bibRefCitation>
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), we consider the higher-level affinities of †
|
||||
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<emphasis box="[273,423,589,613]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
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<emphasis id="B932B059F6F168E3DB1AFDE31BC5FDA2" box="[273,423,589,613]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
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to be uncertain, beyond its being a member of
|
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<taxonomicName authorityName=", Illiger" authorityYear="1811" box="[448,581,622,646]" class="Mammalia" kingdom="Animalia" order="Marsupialia" pageId="202" pageNumber="202" phylum="Chordata" rank="order">Marsupialia</taxonomicName>
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.
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Agreodontia
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CONTENTS:
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,
|
||||
<taxonomicName authorityName="Kirsch" authorityYear="1977" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7DE13F9A318BCF982" authorityName="Kirsch" authorityYear="1977" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
, and
|
||||
<taxonomicName baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[794,991,1581,1605]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7D911F98319BDF982" baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[794,991,1581,1605]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="207" lastPageNumber="207" pageId="206" pageNumber="206" type="materials_examined">
|
||||
<paragraph blockId="206.[684,1212,1548,1738]" pageId="206" pageNumber="206">
|
||||
<collectionCode box="[716,775,1615,1639]" pageId="206" pageNumber="206">STEM</collectionCode>
|
||||
<subSubSection id="C35C3FC0F6F568E6D8C7F9E1183FFEFB" lastPageId="207" lastPageNumber="207" pageId="206" pageNumber="206" type="materials_examined">
|
||||
<paragraph id="8BF96C4BF6F568E7D8C7F9E1188EF94F" blockId="206.[684,1212,1548,1738]" pageId="206" pageNumber="206">
|
||||
<collectionCode id="ED57F48EF6F568E7D8C7F9E11965F9A0" box="[716,775,1615,1639]" pageId="206" pageNumber="206">STEM</collectionCode>
|
||||
AGE: 48.0 Mya (95%
|
||||
<collectionCode box="[1032,1091,1615,1638]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="206" pageNumber="206" type="Herbarium">HPD</collectionCode>
|
||||
<collectionCode id="ED57F48EF6F568E7DE03F9E11E21F9A1" box="[1032,1091,1615,1638]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="206" pageNumber="206" type="Herbarium">HPD</collectionCode>
|
||||
: 44.3–50.9 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[684,1212,1548,1738]" pageId="206" pageNumber="206">
|
||||
<collectionCode box="[716,800,1681,1705]" pageId="206" pageNumber="206">CROWN</collectionCode>
|
||||
<paragraph id="8BF96C4BF6F568E7D8C7F93F188FF90D" blockId="206.[684,1212,1548,1738]" pageId="206" pageNumber="206">
|
||||
<collectionCode id="ED57F48EF6F568E7D8C7F93F1942F96E" box="[716,800,1681,1705]" pageId="206" pageNumber="206">CROWN</collectionCode>
|
||||
AGE: 45.7 Mya (95%
|
||||
<collectionCode box="[1038,1097,1681,1704]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="206" pageNumber="206" type="Herbarium">HPD</collectionCode>
|
||||
<collectionCode id="ED57F48EF6F568E7DE05F93F1E2BF96F" box="[1038,1097,1681,1704]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="206" pageNumber="206" type="Herbarium">HPD</collectionCode>
|
||||
: 42.3–49.2 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
<paragraph id="8BF96C4BF6F468E6DA87FF4C183FFEFB" blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
UNAMBIGUOUS CRANIODENTAL
|
||||
<collectionCode box="[108,171,259,283]" pageId="207" pageNumber="207">SYNAPOMORPHIES</collectionCode>
|
||||
<collectionCode id="ED57F48EF6F468E6DBE0FF4C1AC9FEDC" pageId="207" pageNumber="207">SYNAPOMORPHIES</collectionCode>
|
||||
: Median parietal suture at least partially fused in subadults (char. 25: 0→1; ci = 0.143).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="207" pageNumber="207" type="discussion">
|
||||
<paragraph blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
<subSubSection id="C35C3FC0F6F468E6DA87FEEB1992F90D" pageId="207" pageNumber="207" type="discussion">
|
||||
<paragraph id="8BF96C4BF6F468E6DA87FEEB1847FAFA" blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
COMMENTS:
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[281,472,325,349]" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB12FEEB1BBAFE9A" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[281,472,325,349]" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
|
||||
gave the name Agreodontia to a clade comprising the Australian orders
|
||||
<taxonomicName baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[232,423,391,415]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F468E6DAE3FE291BC5FE58" baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[232,423,391,415]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
,
|
||||
<taxonomicName baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[433,631,391,415]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F468E6DBBAFE291815FE58" baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[433,631,391,415]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
, and
|
||||
<taxonomicName authorityName="Kirsch" authorityYear="1977" box="[160,371,424,448]" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F468E6DAABFE061B11FE07" authorityName="Kirsch" authorityYear="1977" box="[160,371,424,448]" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
, which they defined as the most inclusive clade including
|
||||
<taxonomicName authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species">
|
||||
<emphasis italics="true" pageId="207" pageNumber="207">Perameles nasuta</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6D805FE671AD7FDC5" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species" species="nasuta">
|
||||
<emphasis id="B932B059F6F468E6D805FE671AD7FDC5" italics="true" pageId="207" pageNumber="207">Perameles nasuta</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName baseAuthorityName="Stirling" baseAuthorityYear="1889" box="[195,408,490,514]" class="Mammalia" family="Notoryctidae" genus="Notoryctes" kingdom="Animalia" order="Notoryctemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species">
|
||||
<emphasis box="[195,408,490,514]" italics="true" pageId="207" pageNumber="207">Notoryctes typhlops</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6DAC8FE451BFAFDC5" baseAuthorityName="Stirling" baseAuthorityYear="1889" box="[195,408,490,514]" class="Mammalia" family="Notoryctidae" genus="Notoryctes" kingdom="Animalia" order="Notoryctemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species" species="typhlops">
|
||||
<emphasis id="B932B059F6F468E6DAC8FE451BFAFDC5" box="[195,408,490,514]" italics="true" pageId="207" pageNumber="207">Notoryctes typhlops</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName baseAuthorityName="Kerr" baseAuthorityYear="1792" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species">
|
||||
<emphasis italics="true" pageId="207" pageNumber="207">Dasyurus maculatus</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6DBDCFE451ACDFDE4" baseAuthorityName="Kerr" baseAuthorityYear="1792" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="species" species="maculatus">
|
||||
<emphasis id="B932B059F6F468E6DBDCFE451ACDFDE4" italics="true" pageId="207" pageNumber="207">Dasyurus maculatus</emphasis>
|
||||
</taxonomicName>
|
||||
, but excluding
|
||||
<taxonomicName authority="(Beck et al., 2014: 132)" baseAuthorityName="Beck" baseAuthorityPageNumber="132" baseAuthorityYear="2014" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="207" pageNumber="207" phylum="Chordata" rank="species">
|
||||
<emphasis box="[348,566,523,547]" italics="true" pageId="207" pageNumber="207">Phalanger orientalis</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6DB57FDA51B51FD83" authority="(Beck et al., 2014: 132)" baseAuthorityName="Beck" baseAuthorityPageNumber="132" baseAuthorityYear="2014" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="207" pageNumber="207" phylum="Chordata" rank="species" species="orientalis">
|
||||
<emphasis id="B932B059F6F468E6DB57FDA51854FDE4" box="[348,566,523,547]" italics="true" pageId="207" pageNumber="207">Phalanger orientalis</emphasis>
|
||||
(
|
||||
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 132</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D84FFDA51B4EFD83" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 132</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Monophyly of Agreodontia has been consistently supported by analyses of nuclear and combined nuclear and mitochondrial sequence data (e.g.,
|
||||
<bibRefCitation author="Amrine-Madsen, H." pageId="207" pageNumber="207" pagination="186 - 196" refId="ref188949" refString="Amrine-Madsen, H., et al. 2003 b. Nuclear gene sequences provide evidence for the monophyly of australidelphian marsupials. Molecular Phylogenetics and Evolution 28 (2): 186 - 196." type="journal article" year="2003">Amrine-Madsen et al., 2003b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB88FD211AD0FD0F" author="Amrine-Madsen, H." pageId="207" pageNumber="207" pagination="186 - 196" refId="ref188949" refString="Amrine-Madsen, H., et al. 2003 b. Nuclear gene sequences provide evidence for the monophyly of australidelphian marsupials. Molecular Phylogenetics and Evolution 28 (2): 186 - 196." type="journal article" year="2003">Amrine-Madsen et al., 2003b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Phillips" box="[190,408,688,712]" etAl="et al." firstAuthor="Phillips" pageId="207" pageNumber="207" pagination="122 - 137" refId="ref221314" refString="Phillips, M. J., P. A. McLenachan, C. Down, G. C. Gibb, and D. Penny. 2006. Combined mitochondrial and nuclear DNA sequences resolve the interrelations of the major Australasian marsupial radiations. Systematic Biology 55 (1): 122 - 137." type="journal article" year="2006">Phillips et al., 2006</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DAB5FD1E1BFAFD0F" author="Phillips" box="[190,408,688,712]" etAl="et al." firstAuthor="Phillips" pageId="207" pageNumber="207" pagination="122 - 137" refId="ref221314" refString="Phillips, M. J., P. A. McLenachan, C. Down, G. C. Gibb, and D. Penny. 2006. Combined mitochondrial and nuclear DNA sequences resolve the interrelations of the major Australasian marsupial radiations. Systematic Biology 55 (1): 122 - 137." type="journal article" year="2006">Phillips et al., 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D." box="[420,557,688,713]" pageId="207" pageNumber="207" pagination="175 - 189" refId="ref192516" refString="Beck, R. M. D. 2008 a. A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89 (1): 175 - 189." type="journal article" year="2008">Beck, 2008a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DBAFFD1E184FFD0E" author="Beck, R. M. D." box="[420,557,688,713]" pageId="207" pageNumber="207" pagination="175 - 189" refId="ref192516" refString="Beck, R. M. D. 2008 a. A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89 (1): 175 - 189." type="journal article" year="2008">Beck, 2008a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Meredith, R. W. & M. Westerman & J. A. Case & M. S. Springer" pageId="207" pageNumber="207" pagination="1 - 36" refId="ref216587" refString="Meredith, R. W., M. Westerman, J. A. Case, and M. S. Springer. 2008 b. A phylogeny and timescale for marsupial evolution based on sequences for five nuclear genes. Journal of Mammalian Evolution 15 (1): 1 - 36." type="journal article" year="2008">Meredith et al., 2008b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D832FD1F1B53FD2D" author="Meredith, R. W. & M. Westerman & J. A. Case & M. S. Springer" pageId="207" pageNumber="207" pagination="1 - 36" refId="ref216587" refString="Meredith, R. W., M. Westerman, J. A. Case, and M. S. Springer. 2008 b. A phylogeny and timescale for marsupial evolution based on sequences for five nuclear genes. Journal of Mammalian Evolution 15 (1): 1 - 36." type="journal article" year="2008">Meredith et al., 2008b</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Meredith, R. W. & C. Krajewski & M. Westerman & M. S. Springer" box="[319,386,722,746]" pageId="207" pageNumber="207" pagination="383 - 406" refId="ref216745" refString="Meredith, R. W., C. Krajewski, M. Westerman, and M. S. Springer. 2009 c. Relationships and divergence times among the orders and families of Marsupialia. Museum of Northern Arizona Bulletin 65: 383 - 406." type="journal article" year="2009">
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB34FD7C1BE0FD2D" author="Meredith, R. W. & C. Krajewski & M. Westerman & M. S. Springer" box="[319,386,722,746]" pageId="207" pageNumber="207" pagination="383 - 406" refId="ref216745" refString="Meredith, R. W., C. Krajewski, M. Westerman, and M. S. Springer. 2009 c. Relationships and divergence times among the orders and families of Marsupialia. Museum of Northern Arizona Bulletin 65: 383 - 406." type="journal article" year="2009">
|
||||
2009
|
||||
<bibRefCitation author="Rodgers, J. C." box="[375,457,722,746]" pageId="207" pageNumber="207" refId="ref224441" refString="Rodgers, J. C. 2011. Comparative morphology of the vestibular semicircular canals in therian mammals. Ph. D. dissertation, Faculty of the Graduate School, University of Texas at Austin, Austin." type="book" year="2011">c, 2011</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB7CFD7C1BE0FD2D" author="Rodgers, J. C." box="[375,386,722,746]" pageId="207" pageNumber="207" refId="ref224441" refString="Rodgers, J. C. 2011. Comparative morphology of the vestibular semicircular canals in therian mammals. Ph. D. dissertation, Faculty of the Graduate School, University of Texas at Austin, Austin." type="book" year="2011">c</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
, 2011;
|
||||
<bibRefCitation id="EFD711BAF6F468E6DBDCFD7C1AC1FCCC" author="Mitchell, K. J." pageId="207" pageNumber="207" pagination="2322 - 2330" refId="ref217043" refString="Mitchell, K. J., et al. 2014. Molecular phylogeny, biogeography, and habitat preference evolution of marsupials. Molecular Biology and Evolution 31 (9): 2322 - 2330." type="journal article" year="2014">Mitchell et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Mitchell, K. J." pageId="207" pageNumber="207" pagination="2322 - 2330" refId="ref217043" refString="Mitchell, K. J., et al. 2014. Molecular phylogeny, biogeography, and habitat preference evolution of marsupials. Molecular Biology and Evolution 31 (9): 2322 - 2330." type="journal article" year="2014">Mitchell et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DAA5FD5D1BEDFCCC" author="Duchene, D. A." box="[174,399,755,779]" pageId="207" pageNumber="207" pagination="400 - 412" refId="ref200659" refString="Duchene, D. A., et al. 2018. Analysis of phylogenomic tree space resolves relationships among marsupial families. Systematic Biology 67 (3): 400 - 412." type="journal article" year="2018">Duchêne et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Duchene, D. A." box="[174,399,755,779]" pageId="207" pageNumber="207" pagination="400 - 412" refId="ref200659" refString="Duchene, D. A., et al. 2018. Analysis of phylogenomic tree space resolves relationships among marsupial families. Systematic Biology 67 (3): 400 - 412." type="journal article" year="2018">Duchêne et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Alvarez-Carretero, S." pageId="207" pageNumber="207" pagination="263 - 267" refId="ref188636" refString="Alvarez-Carretero, S., et al. 2021. A species-level timeline of mammal evolution integrating phylogenomic data. Nature 602: 263 - 267." type="journal article" year="2021">Álvarez-Carretero et al., 2021</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB91FD5D1AAEFCEB" author="Alvarez-Carretero, S." pageId="207" pageNumber="207" pagination="263 - 267" refId="ref188636" refString="Alvarez-Carretero, S., et al. 2021. A species-level timeline of mammal evolution integrating phylogenomic data. Nature 602: 263 - 267." type="journal article" year="2021">Álvarez-Carretero et al., 2021</bibRefCitation>
|
||||
), but it is supported only by mitochondrial sequence data when base composition is corrected for (
|
||||
<bibRefCitation author="Nilsson" box="[265,477,854,878]" etAl="et al." firstAuthor="Nilsson" pageId="207" pageNumber="207" pagination="189 - 196" refId="ref219362" refString="Nilsson, M. A., U. Arnason, P. B. S. Spencer, and A. Janke. 2004. Marsupial relationships and a timeline for marsupial radiation in South Gondwana. Gene 340: 189 - 196." type="journal article" year="2004">Nilsson et al., 2004</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DB02FCF81BBFFCA9" author="Nilsson" box="[265,477,854,878]" etAl="et al." firstAuthor="Nilsson" pageId="207" pageNumber="207" pagination="189 - 196" refId="ref219362" refString="Nilsson, M. A., U. Arnason, P. B. S. Spencer, and A. Janke. 2004. Marsupial relationships and a timeline for marsupial radiation in South Gondwana. Gene 340: 189 - 196." type="journal article" year="2004">Nilsson et al., 2004</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Phillips" etAl="et al." firstAuthor="Phillips" pageId="207" pageNumber="207" pagination="122 - 137" refId="ref221314" refString="Phillips, M. J., P. A. McLenachan, C. Down, G. C. Gibb, and D. Penny. 2006. Combined mitochondrial and nuclear DNA sequences resolve the interrelations of the major Australasian marsupial radiations. Systematic Biology 55 (1): 122 - 137." type="journal article" year="2006">Phillips et al., 2006</bibRefCitation>
|
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). A single, uncontradicted retrotransposon insertion also supports agreodontian monophyly, but this does not represent statistically significant support (
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). Recent morphological analyses vary as to whether they recover Agreodontia or not (e.g.,
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<bibRefCitation id="EFD711BAF6F468E6DB6FFB0E1872FB7E" author="Carneiro, L. M." box="[356,528,1184,1209]" pageId="207" pageNumber="207" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">Carneiro, 2019</bibRefCitation>
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), but this clade has been recovered by most total-evidence analyses (
|
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<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[217,423,1251,1275]" pageId="207" pageNumber="207" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
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<bibRefCitation id="EFD711BAF6F468E6DAD2FB4D1BC5FB3C" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[217,423,1251,1275]" pageId="207" pageNumber="207" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
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Beck et al., 2008
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<bibRefCitation author="Rambaut, A. & M. A. Suchard & D. Xie & A. J. Drummond" box="[411,493,1251,1275]" pageId="207" pageNumber="207" refId="ref223023" refString="Rambaut, A., M. A. Suchard, D. Xie, and A. J. Drummond. 2014. Tracer v 1.6." type="book" year="2014">a, 2014</bibRefCitation>
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</bibRefCitation>
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,
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<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[506,562,1251,1275]" pageId="207" pageNumber="207" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
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, 2014,
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<bibRefCitation id="EFD711BAF6F468E6DBF1FB4D1850FB3C" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[506,562,1251,1275]" pageId="207" pageNumber="207" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D." pageId="207" pageNumber="207" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6D84BFB4D1AC6FADB" author="Beck, R. M. D." pageId="207" pageNumber="207" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
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,
|
||||
<bibRefCitation author="Beck, R. M. D." box="[180,249,1284,1308]" pageId="207" pageNumber="207" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">2017a</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6DABFFAAA1A9BFADB" author="Beck, R. M. D." box="[180,249,1284,1308]" pageId="207" pageNumber="207" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">2017a</bibRefCitation>
|
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;
|
||||
<bibRefCitation author="Maga, A. M. & R. M. D. Beck" box="[264,520,1284,1308]" pageId="207" pageNumber="207" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6DB03FAAA186AFADB" author="Maga, A. M. & R. M. D. Beck" box="[264,520,1284,1308]" pageId="207" pageNumber="207" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
) with the exception of that of
|
||||
<bibRefCitation author="Asher, R. J. & I. Horovitz & M. R. Sanchez-Villagra" box="[334,545,1317,1341]" pageId="207" pageNumber="207" pagination="240 - 250" refId="ref191802" refString="Asher, R. J., I. Horovitz, and M. R. Sanchez-Villagra. 2004. First combined cladistic analysis of marsupial mammal interrelationships. Molecular Phylogenetics and Evolution 33: 240 - 250." type="journal article" year="2004">Asher et al. (2004)</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6DB45FA8B1843FAFA" author="Asher, R. J. & I. Horovitz & M. R. Sanchez-Villagra" box="[334,545,1317,1341]" pageId="207" pageNumber="207" pagination="240 - 250" refId="ref191802" refString="Asher, R. J., I. Horovitz, and M. R. Sanchez-Villagra. 2004. First combined cladistic analysis of marsupial mammal interrelationships. Molecular Phylogenetics and Evolution 33: 240 - 250." type="journal article" year="2004">Asher et al. (2004)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
<paragraph id="8BF96C4BF6F468E6DA87FAE81B19F90D" blockId="207.[108,637,226,1738]" pageId="207" pageNumber="207">
|
||||
Our results conform to this general pattern of inconsistent support: whereas our nuclear (fig. 27), combined nuclear and mitochondrial (fig. 29), and total evidence (figs. 32,
|
||||
<figureCitation box="[453,480,1449,1473]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="207" pageNumber="207">33</figureCitation>
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<figureCitation id="137D70CEF6F468E6DBCEFA071B82FA06" box="[453,480,1449,1473]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="207" pageNumber="207">33</figureCitation>
|
||||
) analyses support monophyly of Agreodontia, our mitochondrial (fig. 28) and morphological (figs. 30, 31) analyses do not. Only partial fusion of the median parietal suture before adulthood optimizes as an unambiguous craniodental synapomorphy of this clade in our dated total-evidence analysis, but this transformation is reversed within
|
||||
<taxonomicName baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[186,375,1714,1738]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
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<taxonomicName id="4C4617C8F6F468E6DAB1F91C1B15F90D" baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[186,375,1714,1738]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph blockId="207.[684,1213,226,1738]" pageId="207" pageNumber="207">
|
||||
<paragraph id="8BF96C4BF6F468E6D8C7FF4C19F4FC48" blockId="207.[684,1213,226,1738]" pageId="207" pageNumber="207">
|
||||
Putative peramelemorphians have been reported from the earliest Eocene Tingamarra Local Fauna (
|
||||
<bibRefCitation author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" box="[843,1091,292,316]" pageId="207" pageNumber="207" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6D940FE8A1E21FEFB" author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" box="[843,1091,292,316]" pageId="207" pageNumber="207" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
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;
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<bibRefCitation author="Archer, M. & H. Godthelp & S. J. Hand" pageId="207" pageNumber="207" pagination="193 - 200" refId="ref190726" refString="Archer, M., H. Godthelp, and S. J. Hand. 1993 a. Early Eocene marsupial from Australia. Kaupia 3: 193 - 200." type="journal article" year="1993">Archer et al., 1993a</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6DE5AFE8A197BFE9A" author="Archer, M. & H. Godthelp & S. J. Hand" pageId="207" pageNumber="207" pagination="193 - 200" refId="ref190726" refString="Archer, M., H. Godthelp, and S. J. Hand. 1993 a. Early Eocene marsupial from Australia. Kaupia 3: 193 - 200." type="journal article" year="1993">Archer et al., 1993a</bibRefCitation>
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<bibRefCitation author="Muirhead, J." box="[805,986,325,349]" pageId="207" pageNumber="207" refId="ref217366" refString="Muirhead, J. 1994. Systematics, evolution and palaeobiology of recent and fossil bandicoots (Marsupialia: Peramelemorphia). Ph. D. dissertation, School of Biological Science, University of New South Wales, Sydney." type="book" year="1994">Muirhead, 1994</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6D92EFEEB19B8FE9A" author="Muirhead, J." box="[805,986,325,349]" pageId="207" pageNumber="207" refId="ref217366" refString="Muirhead, J. 1994. Systematics, evolution and palaeobiology of recent and fossil bandicoots (Marsupialia: Peramelemorphia). Ph. D. dissertation, School of Biological Science, University of New South Wales, Sydney." type="book" year="1994">Muirhead, 1994</bibRefCitation>
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;
|
||||
<bibRefCitation author="Bassarova, M. & M. Archer" box="[998,1207,325,349]" pageId="207" pageNumber="207" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D9EDFEEB1ED5FE9A" author="Bassarova, M. & M. Archer" box="[998,1207,325,349]" pageId="207" pageNumber="207" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[685,888,358,382]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8A6FEC8191AFEB9" author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[685,888,358,382]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
). Given the published early Eocene (54.6 Mya) radiometric date for Tingamarra (
|
||||
<bibRefCitation author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" box="[774,1036,424,448]" pageId="207" pageNumber="207" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
|
||||
), this material would represent the oldest known record of the Agreodontian crown clade and would markedly predate our estimate for the most recent common ancestor of Agreodontia (45.7 Mya; 95%
|
||||
<collectionCode box="[819,880,590,613]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="207" pageNumber="207" type="Herbarium">HPD</collectionCode>
|
||||
: 42.3–49.2 Mya). However, examination of these Tingamarran specimens by
|
||||
<collectionCode box="[727,749,656,679]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="207" pageNumber="207">R</collectionCode>
|
||||
.M.D.B. revealed some similarities to bunodont, nonperamelemorphian metatherians from the Palaeogene of South America and
|
||||
<collectingCountry box="[684,790,755,779]" name="Australia" pageId="207" pageNumber="207">Australia</collectingCountry>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D90DFE061E6EFE07" author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" box="[774,1036,424,448]" pageId="207" pageNumber="207" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
|
||||
), this material would represent the oldest known record of the Agreodontian crown clade and would markedly predate our estimate for the most recent common ancestor of Agreodontia (45.7 Mya; 95% HPD: 42.3–49.2 Mya). However, examination of these Tingamarran specimens by R.M.D.B. revealed some similarities to bunodont, nonperamelemorphian metatherians from the Palaeogene of South America and
|
||||
<collectingCountry id="F3512CDBF6F468E6D8A7FD5D1974FCCC" box="[684,790,755,779]" name="Australia" pageId="207" pageNumber="207">Australia</collectingCountry>
|
||||
(e.g.,
|
||||
<taxonomicName authorityName="Crochet & Sigé" authorityYear="1993" box="[877,992,755,779]" class="Mammalia" family="Didelphidae" genus="Chulpasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[877,992,755,779]" italics="true" pageId="207" pageNumber="207">Chulpasia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6D966FD5D1982FCCC" authorityName="Crochet & Sigé" authorityYear="1993" box="[877,992,755,779]" class="Mammalia" family="Didelphidae" genus="Chulpasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F468E6D966FD5D1982FCCC" box="[877,992,755,779]" italics="true" pageId="207" pageNumber="207">Chulpasia</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="Goin & Candela" authorityYear="1996" box="[1014,1154,755,779]" class="Mammalia" family="Bonapartheriidae" genus="Rosendolops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="207" pageNumber="207" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[1014,1154,755,779]" italics="true" pageId="207" pageNumber="207">Rosendolops</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F468E6D9FDFD5D1EE0FCCC" authorityName="Goin & Candela" authorityYear="1996" box="[1014,1154,755,779]" class="Mammalia" family="Bonapartheriidae" genus="Rosendolops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="207" pageNumber="207" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F468E6D9FDFD5D1EE0FCCC" box="[1014,1154,755,779]" italics="true" pageId="207" pageNumber="207">Rosendolops</emphasis>
|
||||
</taxonomicName>
|
||||
; see
|
||||
<bibRefCitation author="Archer, M. & H. Godthelp & S. J. Hand" box="[684,918,788,812]" pageId="207" pageNumber="207" pagination="193 - 200" refId="ref190726" refString="Archer, M., H. Godthelp, and S. J. Hand. 1993 a. Early Eocene marsupial from Australia. Kaupia 3: 193 - 200." type="journal article" year="1993">Archer et al., 1993a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8A7FCBA19F4FCEB" author="Archer, M. & H. Godthelp & S. J. Hand" box="[684,918,788,812]" pageId="207" pageNumber="207" pagination="193 - 200" refId="ref190726" refString="Archer, M., H. Godthelp, and S. J. Hand. 1993 a. Early Eocene marsupial from Australia. Kaupia 3: 193 - 200." type="journal article" year="1993">Archer et al., 1993a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Crochet, J. - Y. & B. Sige" box="[933,1207,788,812]" pageId="207" pageNumber="207" pagination="97 - 107" refId="ref198980" refString="Crochet, J. - Y., and B. Sige. 1993. Les mammiferes de Chulpas (Formation Umayo, transition Cretace-Tertiare, Perou). Donnees preliminaires. Documents du Laboratoire de Geologie de Lyon 125: 97 - 107." type="journal article" year="1993">Crochet and Sigé, 1993</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D9AEFCBA1ED5FCEB" author="Crochet, J. - Y. & B. Sige" box="[933,1207,788,812]" pageId="207" pageNumber="207" pagination="97 - 107" refId="ref198980" refString="Crochet, J. - Y., and B. Sige. 1993. Les mammiferes de Chulpas (Formation Umayo, transition Cretace-Tertiare, Perou). Donnees preliminaires. Documents du Laboratoire de Geologie de Lyon 125: 97 - 107." type="journal article" year="1993">Crochet and Sigé, 1993</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Goin, F. J. & A. M. Candela" box="[684,967,821,845]" pageId="207" pageNumber="207" pagination="292 - 296" refId="ref205936" refString="Goin, F. J., and A. M. Candela. 1996. A new early Eocene polydolopimorphian (Mammalia, Marsupialia) from Patagonia. Journal of Vertebrate Paleontology 16 (2): 292 - 296." type="journal article" year="1996">Goin and Candela, 1996</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8A7FC9B19A5FC8A" author="Goin, F. J. & A. M. Candela" box="[684,967,821,845]" pageId="207" pageNumber="207" pagination="292 - 296" refId="ref205936" refString="Goin, F. J., and A. M. Candela. 1996. A new early Eocene polydolopimorphian (Mammalia, Marsupialia) from Patagonia. Journal of Vertebrate Paleontology 16 (2): 292 - 296." type="journal article" year="1996">Goin and Candela, 1996</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Sige" box="[980,1166,821,845]" etAl="et al." firstAuthor="Sige" pageId="207" pageNumber="207" pagination="813 - 823" refId="ref227404" refString="Sige, B., et al. 2009. Chulpasia and Thylacotinga, late Paleocene-earliest Eocene trans-Antarctic Gondwanan bunodont marsupials: New data from Australia. Geobios 42 (6): 813 - 823." type="journal article" year="2009">Sigé et al., 2009</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D9DFFC9B1EECFC8A" author="Sige" box="[980,1166,821,845]" etAl="et al." firstAuthor="Sige" pageId="207" pageNumber="207" pagination="813 - 823" refId="ref227404" refString="Sige, B., et al. 2009. Chulpasia and Thylacotinga, late Paleocene-earliest Eocene trans-Antarctic Gondwanan bunodont marsupials: New data from Australia. Geobios 42 (6): 813 - 823." type="journal article" year="2009">Sigé et al., 2009</bibRefCitation>
|
||||
), so we are not convinced that they represent peramelemorphians.
|
||||
</paragraph>
|
||||
<paragraph blockId="207.[684,1213,226,1738]" pageId="207" pageNumber="207">
|
||||
<paragraph id="8BF96C4BF6F468E6D8C7FC361992F90D" blockId="207.[684,1213,226,1738]" pageId="207" pageNumber="207">
|
||||
A major gap in the Australian fossil record after the early Eocene (
|
||||
<bibRefCitation author="Bassarova, M. & M. Archer" box="[975,1209,953,977]" pageId="207" pageNumber="207" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D9C4FC171EDBFC16" author="Bassarova, M. & M. Archer" box="[975,1209,953,977]" pageId="207" pageNumber="207" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[684,901,986,1011]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8A7FC7519E7FC35" author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[684,901,986,1011]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Woodhead" box="[921,1204,986,1010]" etAl="et al." firstAuthor="Woodhead" pageId="207" pageNumber="207" pagination="153 - 167" refId="ref235654" refString="Woodhead, J., et al. 2014. Developing a radiometricallydated chronologic sequence for Neogene biotic change in Australia, from the Riversleigh World Heritage Area of Queensland. Gondwana Research 29 (1): 153 - 167." type="journal article" year="2014">Woodhead et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D992FC741ED6FC35" author="Woodhead" box="[921,1204,986,1010]" etAl="et al." firstAuthor="Woodhead" pageId="207" pageNumber="207" pagination="153 - 167" refId="ref235654" refString="Woodhead, J., et al. 2014. Developing a radiometricallydated chronologic sequence for Neogene biotic change in Australia, from the Riversleigh World Heritage Area of Queensland. Gondwana Research 29 (1): 153 - 167." type="journal article" year="2014">Woodhead et al., 2014</bibRefCitation>
|
||||
) means that the oldest definitive agreodontians are from the late Oligocene, which is much younger than our estimate for the age of the most recent common ancestor of Agreodontia. Specifically, representatives of
|
||||
<taxonomicName baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F468E6DE11FBD118BCFB7F" baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
and
|
||||
<taxonomicName baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[804,1001,1184,1208]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F468E6D92FFB0E198BFB7F" baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[804,1001,1184,1208]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="207" pageNumber="207" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
are known from multiple late Oligocene sites in central
|
||||
<collectingCountry name="Australia" pageId="207" pageNumber="207">Australia</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F468E6DE6CFB6C18AAFB3C" name="Australia" pageId="207" pageNumber="207">Australia</collectingCountry>
|
||||
and at Riversleigh World Heritage Area (
|
||||
<bibRefCitation author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[692,894,1284,1308]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8BFFAAA191CFADB" author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[692,894,1284,1308]" pageId="207" pageNumber="207" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Wroe, S." box="[908,1044,1284,1308]" pageId="207" pageNumber="207" pagination="102 - 123" refId="ref236472" refString="Wroe, S. 2003. Australian marsupial carnivores: recent advances in palaeontology. In M. Jones, C. Dickman, and M. Archer (editors), Predators with pouches: the biology of marsupial carnivores: 102 - 123. Collingwood, Australia: CSIRO Publishing." type="book chapter" year="2003">Wroe, 2003</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D987FAAA1E76FADB" author="Wroe, S." box="[908,1044,1284,1308]" pageId="207" pageNumber="207" pagination="102 - 123" refId="ref236472" refString="Wroe, S. 2003. Australian marsupial carnivores: recent advances in palaeontology. In M. Jones, C. Dickman, and M. Archer (editors), Predators with pouches: the biology of marsupial carnivores: 102 - 123. Collingwood, Australia: CSIRO Publishing." type="book chapter" year="2003">Wroe, 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Archer, M. & S. J. Hand" pageId="207" pageNumber="207" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer et al., 2006</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DE29FAAA1887FAFA" author="Archer, M. & S. J. Hand" pageId="207" pageNumber="207" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer et al., 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Archer, M. & S. J. Hand" box="[753,1025,1317,1341]" pageId="207" pageNumber="207" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer and Hand, 2006</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8FAFA8B1E63FAFA" author="Archer, M. & S. J. Hand" box="[753,1025,1317,1341]" pageId="207" pageNumber="207" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer and Hand, 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Warburton and Travouillon" firstAuthor="Warburton" pageId="207" pageNumber="207" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6DE06FA8B19EBFA99" author="Warburton and Travouillon" firstAuthor="Warburton" pageId="207" pageNumber="207" pagination="151 - 181" refId="ref232688" refString="Warburton, N. M., and K. J. Travouillon. 2016. The biology and palaeontology of the Peramelemorphia: a review of current knowledge and future research directions. Australian Journal of Zoology 64 (3): 151 - 181." type="journal article" year="2016">Warburton and Travouillon, 2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Kealy, S. & R. M. D. Beck" box="[927,1208,1350,1374]" pageId="207" pageNumber="207" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck, 2017</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D994FAE81EDAFA99" author="Kealy, S. & R. M. D. Beck" box="[927,1208,1350,1374]" pageId="207" pageNumber="207" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck, 2017</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Eldridge, M. D. B. & R. M. D. Beck & D. A. Croft & K. J. Travouillon & B. J. Fox" box="[684,929,1383,1407]" pageId="207" pageNumber="207" pagination="802 - 837" refId="ref200965" refString="Eldridge, M. D. B., R. M. D. Beck, D. A. Croft, K. J. Travouillon, and B. J. Fox. 2019. An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria). Journal of Mammalogy 100 (3): 802 - 837." type="journal article" year="2019">Eldridge et al., 2019</bibRefCitation>
|
||||
). A single partial upper molar of an alleged thylacinid (
|
||||
<collectionCode box="[1056,1121,1416,1440]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="207" pageNumber="207">NTM</collectionCode>
|
||||
P2815- 10;
|
||||
<bibRefCitation author="Murray, P. F. & D. Megirian" box="[726,1059,1449,1474]" pageId="207" pageNumber="207" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
|
||||
) and a single upper molar of a probable notoryctemorphian (
|
||||
<collectionCode box="[693,758,1515,1539]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="207" pageNumber="207">NTM</collectionCode>
|
||||
P2815-6;
|
||||
<bibRefCitation author="Murray, P. F. & D. Megirian" box="[875,1209,1515,1540]" pageId="207" pageNumber="207" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8A7FAC919C3FAB8" author="Eldridge, M. D. B. & R. M. D. Beck & D. A. Croft & K. J. Travouillon & B. J. Fox" box="[684,929,1383,1407]" pageId="207" pageNumber="207" pagination="802 - 837" refId="ref200965" refString="Eldridge, M. D. B., R. M. D. Beck, D. A. Croft, K. J. Travouillon, and B. J. Fox. 2019. An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria). Journal of Mammalogy 100 (3): 802 - 837." type="journal article" year="2019">Eldridge et al., 2019</bibRefCitation>
|
||||
). A single partial upper molar of an alleged thylacinid (NTM P2815- 10;
|
||||
<bibRefCitation id="EFD711BAF6F468E6D8DDFA041E41FA06" author="Murray, P. F. & D. Megirian" box="[726,1059,1449,1474]" pageId="207" pageNumber="207" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
|
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) and a single upper molar of a probable notoryctemorphian (NTM P2815-6;
|
||||
<bibRefCitation id="EFD711BAF6F468E6D960FA421EDBF9C4" author="Murray, P. F. & D. Megirian" box="[875,1209,1515,1540]" pageId="207" pageNumber="207" pagination="211 - 228" refId="ref218942" refString="Murray, P. F., and D. Megirian. 2006 b. The Pwerte Marnte Marnte Local Fauna: a new vertebrate assemblage of presumed Oligocene age from the Northern Territory of Australia. Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 211 - 228." type="book chapter" year="2006">Murray and Megirian, 2006b</bibRefCitation>
|
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;
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[686,942,1548,1572]" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 151</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F468E6D8A5F9A219CCF9E3" author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[686,942,1548,1572]" pageId="207" pageNumber="207" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 151</bibRefCitation>
|
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,
|
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<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[957,1072,1548,1572]" pageId="207" pageNumber="207" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016: 166</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F468E6D9B6F9A21E52F9E3" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[957,1072,1548,1572]" pageId="207" pageNumber="207" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016: 166</bibRefCitation>
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) are known from the Pwerte Marnte Marnte Local Fauna in the
|
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<collectingRegion box="[764,988,1614,1638]" country="Australia" name="Northern Territory" pageId="207" pageNumber="207">Northern Territory</collectingRegion>
|
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<collectingRegion id="4982A2A9F6F468E6D8F7F9E019BEF9A1" box="[764,988,1614,1638]" country="Australia" name="Northern Territory" pageId="207" pageNumber="207">Northern Territory</collectingRegion>
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, which is probably slightly older than the central Australian and Riversleigh sites (
|
||||
<bibRefCitation author="Megirian, D. & G. J. Prideaux & P. F. Murray & N. Smit" box="[894,1151,1680,1705]" pageId="207" pageNumber="207" pagination="658 - 671" refId="ref216431" refString="Megirian, D., G. J. Prideaux, P. F. Murray, and N. Smit. 2010. An Australian land mammal age biochronological scheme. Paleobiology 36 (4): 658 - 671." type="journal article" year="2010">Megirian et al., 2010</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F468E6D975F93E1E1DF96E" author="Megirian, D. & G. J. Prideaux & P. F. Murray & N. Smit" box="[894,1151,1680,1705]" pageId="207" pageNumber="207" pagination="658 - 671" refId="ref216431" refString="Megirian, D., G. J. Prideaux, P. F. Murray, and N. Smit. 2010. An Australian land mammal age biochronological scheme. Paleobiology 36 (4): 658 - 671." type="journal article" year="2010">Megirian et al., 2010</bibRefCitation>
|
||||
), but which is still late Oligocene.
|
||||
</paragraph>
|
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|
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|
|
|
|||
|
|
@ -1,350 +1,334 @@
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Australidelphia
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CONTENTS:
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,
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|
||||
<taxonomicName id="4C4617C8F6F668E4DBD2FC55181AFBD4" authorityName="Owen" authorityYear="1866" box="[473,632,1019,1043]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="Ameghino" authorityYear="1889" box="[108,294,1052,1076]" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F668E4DA67FBB21B44FBF3" authorityName="Ameghino" authorityYear="1889" box="[108,294,1052,1076]" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
,
|
||||
<taxonomicName authorityName="Kirsch" authorityYear="1977" box="[329,556,1052,1076]" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F668E4DB42FBB2184EFBF3" authorityName="Kirsch" authorityYear="1977" box="[329,556,1052,1076]" class="Mammalia" kingdom="Animalia" order="Notoryctemorphia" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Notoryctemorphia</taxonomicName>
|
||||
, and
|
||||
<taxonomicName baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[108,305,1085,1109]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F668E4DA67FB931B53FB92" baseAuthorityName="Travouillon and Phillips" baseAuthorityYear="2018" box="[108,305,1085,1109]" class="Mammalia" kingdom="Animalia" order="Peramelemorphia" pageId="205" pageNumber="205" phylum="Chordata" rank="order">Peramelemorphia</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="205" pageNumber="205" type="description">
|
||||
<paragraph blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">
|
||||
<collectionCode box="[140,199,1119,1143]" pageId="205" pageNumber="205">STEM</collectionCode>
|
||||
AGE: 55.1 Mya (95%
|
||||
<collectionCode box="[456,515,1119,1142]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="205" pageNumber="205" type="Herbarium">HPD</collectionCode>
|
||||
: 54.6–56.6 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">
|
||||
<collectionCode box="[140,224,1185,1209]" pageId="205" pageNumber="205">CROWN</collectionCode>
|
||||
AGE: 48.0 Mya (95%
|
||||
<collectionCode box="[462,521,1185,1208]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="205" pageNumber="205" type="Herbarium">HPD</collectionCode>
|
||||
: 44.3–50.9 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">
|
||||
UNAMBIGUOUS CRANIODENTAL
|
||||
<collectionCode box="[108,170,1284,1308]" pageId="205" pageNumber="205">SYNAPOMORPHIES</collectionCode>
|
||||
: None.
|
||||
</paragraph>
|
||||
<subSubSection id="C35C3FC0F6F668E4DA87FBF11A98FADC" pageId="205" pageNumber="205" type="description">
|
||||
<paragraph id="8BF96C4BF6F668E4DA87FBF11ACEFB5F" blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">STEM AGE: 55.1 Mya (95% HPD: 54.6–56.6 Mya).</paragraph>
|
||||
<paragraph id="8BF96C4BF6F668E4DA87FB0F1ACFFB1D" blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">CROWN AGE: 48.0 Mya (95% HPD: 44.3–50.9 Mya).</paragraph>
|
||||
<paragraph id="8BF96C4BF6F668E4DA87FB4D1A98FADC" blockId="205.[108,636,1019,1738]" pageId="205" pageNumber="205">UNAMBIGUOUS CRANIODENTAL SYNAPOMORPHIES: None.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="206" lastPageNumber="206" pageId="205" pageNumber="205" type="discussion">
|
||||
<paragraph blockId="205.[108,636,1019,1738]" lastBlockId="205.[684,1213,226,1738]" pageId="205" pageNumber="205">
|
||||
<subSubSection id="C35C3FC0F6F668E7DA87FA8B1E66FA67" lastPageId="206" lastPageNumber="206" pageId="205" pageNumber="205" type="discussion">
|
||||
<paragraph id="8BF96C4BF6F668E4DA87FA8B1936FC48" blockId="205.[108,636,1019,1738]" lastBlockId="205.[684,1213,226,1738]" pageId="205" pageNumber="205">
|
||||
COMMENTS: Recent phylogenetic analyses based on molecular data (e.g.,
|
||||
<bibRefCitation author="Phillips" etAl="et al." firstAuthor="Phillips" pageId="205" pageNumber="205" pagination="1533 - 1538" refId="ref221252" refString="Phillips, M. J., Y. H. Lin, G. L. Harrison, and D. Penny. 2001. Mitochondrial genomes of a bandicoot and a brushtail possum confirm the monophyly of australidelphian marsupials. Proceedings of the Royal Society of London B, Biological Sciences 268 (1475): 1533 - 1538." type="journal article" year="2001">Phillips et al., 2001</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DBD0FAE81AC1FAB8" author="Phillips" etAl="et al." firstAuthor="Phillips" pageId="205" pageNumber="205" pagination="1533 - 1538" refId="ref221252" refString="Phillips, M. J., Y. H. Lin, G. L. Harrison, and D. Penny. 2001. Mitochondrial genomes of a bandicoot and a brushtail possum confirm the monophyly of australidelphian marsupials. Proceedings of the Royal Society of London B, Biological Sciences 268 (1475): 1533 - 1538." type="journal article" year="2001">Phillips et al., 2001</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Amrine-Madsen, H." box="[177,510,1383,1407]" pageId="205" pageNumber="205" pagination="186 - 196" refId="ref188949" refString="Amrine-Madsen, H., et al. 2003 b. Nuclear gene sequences provide evidence for the monophyly of australidelphian marsupials. Molecular Phylogenetics and Evolution 28 (2): 186 - 196." type="journal article" year="2003">Amrine-Madsen et al., 2003b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DABAFAC91B9CFAB8" author="Amrine-Madsen, H." box="[177,510,1383,1407]" pageId="205" pageNumber="205" pagination="186 - 196" refId="ref188949" refString="Amrine-Madsen, H., et al. 2003 b. Nuclear gene sequences provide evidence for the monophyly of australidelphian marsupials. Molecular Phylogenetics and Evolution 28 (2): 186 - 196." type="journal article" year="2003">Amrine-Madsen et al., 2003b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson" etAl="et al." firstAuthor="Nilsson" pageId="205" pageNumber="205" pagination="189 - 196" refId="ref219362" refString="Nilsson, M. A., U. Arnason, P. B. S. Spencer, and A. Janke. 2004. Marsupial relationships and a timeline for marsupial radiation in South Gondwana. Gene 340: 189 - 196." type="journal article" year="2004">Nilsson et al., 2004</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F668E4D800FAC91AA8FA67" author="Nilsson" etAl="et al." firstAuthor="Nilsson" pageId="205" pageNumber="205" pagination="189 - 196" refId="ref219362" refString="Nilsson, M. A., U. Arnason, P. B. S. Spencer, and A. Janke. 2004. Marsupial relationships and a timeline for marsupial radiation in South Gondwana. Gene 340: 189 - 196." type="journal article" year="2004">Nilsson et al., 2004</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Phillips" box="[213,425,1416,1440]" etAl="et al." firstAuthor="Phillips" pageId="205" pageNumber="205" pagination="122 - 137" refId="ref221314" refString="Phillips, M. J., P. A. McLenachan, C. Down, G. C. Gibb, and D. Penny. 2006. Combined mitochondrial and nuclear DNA sequences resolve the interrelations of the major Australasian marsupial radiations. Systematic Biology 55 (1): 122 - 137." type="journal article" year="2006">Phillips et al., 2006</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DADEFA261BCBFA67" author="Phillips" box="[213,425,1416,1440]" etAl="et al." firstAuthor="Phillips" pageId="205" pageNumber="205" pagination="122 - 137" refId="ref221314" refString="Phillips, M. J., P. A. McLenachan, C. Down, G. C. Gibb, and D. Penny. 2006. Combined mitochondrial and nuclear DNA sequences resolve the interrelations of the major Australasian marsupial radiations. Systematic Biology 55 (1): 122 - 137." type="journal article" year="2006">Phillips et al., 2006</bibRefCitation>
|
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;
|
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<bibRefCitation author="Beck, R. M. D." box="[436,570,1416,1440]" pageId="205" pageNumber="205" pagination="175 - 189" refId="ref192516" refString="Beck, R. M. D. 2008 a. A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89 (1): 175 - 189." type="journal article" year="2008">Beck, 2008a</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F668E4DBBFFA261858FA67" author="Beck, R. M. D." box="[436,570,1416,1440]" pageId="205" pageNumber="205" pagination="175 - 189" refId="ref192516" refString="Beck, R. M. D. 2008 a. A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89 (1): 175 - 189." type="journal article" year="2008">Beck, 2008a</bibRefCitation>
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;
|
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<bibRefCitation author="Meredith, R. W. & M. Westerman & J. A. Case & M. S. Springer" pageId="205" pageNumber="205" pagination="1 - 36" refId="ref216587" refString="Meredith, R. W., M. Westerman, J. A. Case, and M. S. Springer. 2008 b. A phylogeny and timescale for marsupial evolution based on sequences for five nuclear genes. Journal of Mammalian Evolution 15 (1): 1 - 36." type="journal article" year="2008">Meredith et al., 2008b</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D84DFA271B59FA06" author="Meredith, R. W. & M. Westerman & J. A. Case & M. S. Springer" pageId="205" pageNumber="205" pagination="1 - 36" refId="ref216587" refString="Meredith, R. W., M. Westerman, J. A. Case, and M. S. Springer. 2008 b. A phylogeny and timescale for marsupial evolution based on sequences for five nuclear genes. Journal of Mammalian Evolution 15 (1): 1 - 36." type="journal article" year="2008">Meredith et al., 2008b</bibRefCitation>
|
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,
|
||||
<bibRefCitation author="Meredith, R. W. & C. Krajewski & M. Westerman & M. S. Springer" box="[330,396,1449,1473]" pageId="205" pageNumber="205" pagination="383 - 406" refId="ref216745" refString="Meredith, R. W., C. Krajewski, M. Westerman, and M. S. Springer. 2009 c. Relationships and divergence times among the orders and families of Marsupialia. Museum of Northern Arizona Bulletin 65: 383 - 406." type="journal article" year="2009">
|
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<bibRefCitation id="EFD711BAF6F668E4DB41FA071BEEFA06" author="Meredith, R. W. & C. Krajewski & M. Westerman & M. S. Springer" box="[330,396,1449,1473]" pageId="205" pageNumber="205" pagination="383 - 406" refId="ref216745" refString="Meredith, R. W., C. Krajewski, M. Westerman, and M. S. Springer. 2009 c. Relationships and divergence times among the orders and families of Marsupialia. Museum of Northern Arizona Bulletin 65: 383 - 406." type="journal article" year="2009">
|
||||
2009
|
||||
<bibRefCitation author="Rodgers, J. C." box="[385,466,1449,1473]" pageId="205" pageNumber="205" refId="ref224441" refString="Rodgers, J. C. 2011. Comparative morphology of the vestibular semicircular canals in therian mammals. Ph. D. dissertation, Faculty of the Graduate School, University of Texas at Austin, Austin." type="book" year="2011">c, 2011</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DB8AFA071BEEFA06" author="Rodgers, J. C." box="[385,396,1449,1473]" pageId="205" pageNumber="205" refId="ref224441" refString="Rodgers, J. C. 2011. Comparative morphology of the vestibular semicircular canals in therian mammals. Ph. D. dissertation, Faculty of the Graduate School, University of Texas at Austin, Austin." type="book" year="2011">c</bibRefCitation>
|
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</bibRefCitation>
|
||||
, 2011;
|
||||
<bibRefCitation id="EFD711BAF6F668E4DBEBFA071AC1FA25" author="Nilsson" etAl="et al." firstAuthor="Nilsson" pageId="205" pageNumber="205" pagination="1000436" refId="ref219407" refString="Nilsson, M. A., et al. 2010. Tracking marsupial evolution using archaic genomic retroposon insertions. PLoS Biology 8 (7): e 1000436." type="journal article" year="2010">Nilsson et al., 2010</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Nilsson" etAl="et al." firstAuthor="Nilsson" pageId="205" pageNumber="205" pagination="1000436" refId="ref219407" refString="Nilsson, M. A., et al. 2010. Tracking marsupial evolution using archaic genomic retroposon insertions. PLoS Biology 8 (7): e 1000436." type="journal article" year="2010">Nilsson et al., 2010</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DABBFA641BF0FA25" author="Mitchell, K. J." box="[176,402,1482,1506]" pageId="205" pageNumber="205" pagination="2322 - 2330" refId="ref217043" refString="Mitchell, K. J., et al. 2014. Molecular phylogeny, biogeography, and habitat preference evolution of marsupials. Molecular Biology and Evolution 31 (9): 2322 - 2330." type="journal article" year="2014">Mitchell et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Mitchell, K. J." box="[176,402,1482,1506]" pageId="205" pageNumber="205" pagination="2322 - 2330" refId="ref217043" refString="Mitchell, K. J., et al. 2014. Molecular phylogeny, biogeography, and habitat preference evolution of marsupials. Molecular Biology and Evolution 31 (9): 2322 - 2330." type="journal article" year="2014">Mitchell et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DB94FA64181AFA25" author="Gallus, S. & A. Janke & V. Kumar & M. A. Nilsson" box="[415,632,1482,1506]" pageId="205" pageNumber="205" pagination="985 - 992" refId="ref204175" refString="Gallus, S., A. Janke, V. Kumar, and M. A. Nilsson. 2015 a. Disentangling the relationship of the Australian marsupial orders using retrotransposon and evolutionary network analyses. Genome Biology and Evolution 7 (4): 985 - 992." type="journal article" year="2015">Gallus et al., 2015a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Gallus, S. & A. Janke & V. Kumar & M. A. Nilsson" box="[415,632,1482,1506]" pageId="205" pageNumber="205" pagination="985 - 992" refId="ref204175" refString="Gallus, S., A. Janke, V. Kumar, and M. A. Nilsson. 2015 a. Disentangling the relationship of the Australian marsupial orders using retrotransposon and evolutionary network analyses. Genome Biology and Evolution 7 (4): 985 - 992." type="journal article" year="2015">Gallus et al., 2015a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DA67FA451B39F9C4" author="Duchene, D. A." box="[108,347,1515,1539]" pageId="205" pageNumber="205" pagination="400 - 412" refId="ref200659" refString="Duchene, D. A., et al. 2018. Analysis of phylogenomic tree space resolves relationships among marsupial families. Systematic Biology 67 (3): 400 - 412." type="journal article" year="2018">Duchêne et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Duchene, D. A." box="[108,347,1515,1539]" pageId="205" pageNumber="205" pagination="400 - 412" refId="ref200659" refString="Duchene, D. A., et al. 2018. Analysis of phylogenomic tree space resolves relationships among marsupial families. Systematic Biology 67 (3): 400 - 412." type="journal article" year="2018">Duchêne et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Alvarez-Carretero, S." pageId="205" pageNumber="205" pagination="263 - 267" refId="ref188636" refString="Alvarez-Carretero, S., et al. 2021. A species-level timeline of mammal evolution integrating phylogenomic data. Nature 602: 263 - 267." type="journal article" year="2021">Álvarez-Carretero et al., 2021</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DB61FA421AC0F9E3" author="Alvarez-Carretero, S." pageId="205" pageNumber="205" pagination="263 - 267" refId="ref188636" refString="Alvarez-Carretero, S., et al. 2021. A species-level timeline of mammal evolution integrating phylogenomic data. Nature 602: 263 - 267." type="journal article" year="2021">Álvarez-Carretero et al., 2021</bibRefCitation>
|
||||
) and others based on total-evidence datasets (e.g.,
|
||||
<bibRefCitation author="Asher, R. J. & I. Horovitz & M. R. Sanchez-Villagra" box="[212,414,1581,1605]" pageId="205" pageNumber="205" pagination="240 - 250" refId="ref191802" refString="Asher, R. J., I. Horovitz, and M. R. Sanchez-Villagra. 2004. First combined cladistic analysis of marsupial mammal interrelationships. Molecular Phylogenetics and Evolution 33: 240 - 250." type="journal article" year="2004">Asher et al., 2004</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DADFF9831BFCF982" author="Asher, R. J. & I. Horovitz & M. R. Sanchez-Villagra" box="[212,414,1581,1605]" pageId="205" pageNumber="205" pagination="240 - 250" refId="ref191802" refString="Asher, R. J., I. Horovitz, and M. R. Sanchez-Villagra. 2004. First combined cladistic analysis of marsupial mammal interrelationships. Molecular Phylogenetics and Evolution 33: 240 - 250." type="journal article" year="2004">Asher et al., 2004</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[429,632,1581,1605]" pageId="205" pageNumber="205" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
|
||||
<bibRefCitation id="EFD711BAF6F668E4DBA6F983181AF982" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[429,632,1581,1605]" pageId="205" pageNumber="205" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
|
||||
Beck et al., 2008
|
||||
<bibRefCitation author="Rambaut, A. & M. A. Suchard & D. Xie & A. J. Drummond" pageId="205" pageNumber="205" refId="ref223023" refString="Rambaut, A., M. A. Suchard, D. Xie, and A. J. Drummond. 2014. Tracer v 1.6." type="book" year="2014">a, 2014</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D867F983181AF982" author="Rambaut, A. & M. A. Suchard & D. Xie & A. J. Drummond" box="[620,632,1581,1605]" pageId="205" pageNumber="205" refId="ref223023" refString="Rambaut, A., M. A. Suchard, D. Xie, and A. J. Drummond. 2014. Tracer v 1.6." type="book" year="2014">a</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[179,234,1614,1638]" pageId="205" pageNumber="205" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
|
||||
, 2014,
|
||||
<bibRefCitation id="EFD711BAF6F668E4DAB8F9E01A88F9A1" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[179,234,1614,1638]" pageId="205" pageNumber="205" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Horovitz, I." box="[250,489,1614,1639]" pageId="205" pageNumber="205" pagination="8278" refId="ref209439" refString="Horovitz, I., et al. 2009. Cranial anatomy of the earliest marsupials and the origin of opossums. PLoS One 4 (12): e 8278." type="journal article" year="2009">Horovitz et al., 2009</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DAF1F9E01B8BF9A0" author="Horovitz, I." box="[250,489,1614,1639]" pageId="205" pageNumber="205" pagination="8278" refId="ref209439" refString="Horovitz, I., et al. 2009. Cranial anatomy of the earliest marsupials and the origin of opossums. PLoS One 4 (12): e 8278." type="journal article" year="2009">Horovitz et al., 2009</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D." box="[505,632,1614,1639]" pageId="205" pageNumber="205" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DBF2F9E0181AF9A0" author="Beck, R. M. D." box="[505,632,1614,1639]" pageId="205" pageNumber="205" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Maga, A. M. & R. M. D. Beck" box="[108,352,1647,1672]" pageId="205" pageNumber="205" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4DA67F9DE1B02F94F" author="Maga, A. M. & R. M. D. Beck" box="[108,352,1647,1672]" pageId="205" pageNumber="205" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
) have consistently recovered monophyly of Australidelphia, and australidelphian monophyly has also been found in most (e.g.,
|
||||
<bibRefCitation author="Horovitz, I. & M. R. Sanchez-Villagra" box="[805,1208,226,250]" pageId="205" pageNumber="205" pagination="181 - 212" refId="ref209364" refString="Horovitz, I., and M. R. Sanchez-Villagra. 2003. A morphological analysis of marsupial mammal higherlevel phylogenetic relationships. Cladistics 19: 181 - 212." type="journal article" year="2003">Horovitz and Sánchez-Villagra, 2003</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D92EFF4C1EDAFF3D" author="Horovitz, I. & M. R. Sanchez-Villagra" box="[805,1208,226,250]" pageId="205" pageNumber="205" pagination="181 - 212" refId="ref209364" refString="Horovitz, I., and M. R. Sanchez-Villagra. 2003. A morphological analysis of marsupial mammal higherlevel phylogenetic relationships. Cladistics 19: 181 - 212." type="journal article" year="2003">Horovitz and Sánchez-Villagra, 2003</bibRefCitation>
|
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;
|
||||
<bibRefCitation author="Sanchez-Villagra" box="[685,1020,259,283]" etAl="et al." firstAuthor="Sanchez-Villagra" pageId="205" pageNumber="205" pagination="83 - 97" refId="ref226509" refString="Schmelzle, T., M. R. Sanchez-Villagra, and W. Maier. 2007. Vestibular labyrinth diversity in diprotodontian marsupial mammals. Mammal Study 32: 83 - 97." type="journal article" year="2007">Sánchez-Villagra et al., 2007</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F668E4D8A6FEAD199EFEDC" author="Sanchez-Villagra" box="[685,1020,259,283]" etAl="et al." firstAuthor="Sanchez-Villagra" pageId="205" pageNumber="205" pagination="83 - 97" refId="ref226509" refString="Schmelzle, T., M. R. Sanchez-Villagra, and W. Maier. 2007. Vestibular labyrinth diversity in diprotodontian marsupial mammals. Mammal Study 32: 83 - 97." type="journal article" year="2007">Sánchez-Villagra et al., 2007</bibRefCitation>
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;
|
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<bibRefCitation author="Horovitz, I." pageId="205" pageNumber="205" pagination="109 - 141" refId="ref209395" refString="Horovitz, I., et al. 2008. The anatomy of Herpetotherium cf. fugax Cope, 1873, a metatherian from the Oligocene of North America. Palaeontographica Abteilung A 284 (4 - 6): 109 - 141." type="journal article" year="2008">Horovitz et al., 2008</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DE06FEAD1881FEFB" author="Horovitz, I." pageId="205" pageNumber="205" pagination="109 - 141" refId="ref209395" refString="Horovitz, I., et al. 2008. The anatomy of Herpetotherium cf. fugax Cope, 1873, a metatherian from the Oligocene of North America. Palaeontographica Abteilung A 284 (4 - 6): 109 - 141." type="journal article" year="2008">Horovitz et al., 2008</bibRefCitation>
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,
|
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<bibRefCitation author="Horovitz, I." box="[754,809,292,316]" pageId="205" pageNumber="205" pagination="8278" refId="ref209439" refString="Horovitz, I., et al. 2009. Cranial anatomy of the earliest marsupials and the origin of opossums. PLoS One 4 (12): e 8278." type="journal article" year="2009">2009</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D8F9FE8A194BFEFB" author="Horovitz, I." box="[754,809,292,316]" pageId="205" pageNumber="205" pagination="8278" refId="ref209439" refString="Horovitz, I., et al. 2009. Cranial anatomy of the earliest marsupials and the origin of opossums. PLoS One 4 (12): e 8278." type="journal article" year="2009">2009</bibRefCitation>
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;
|
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<bibRefCitation author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[823,1047,292,316]" pageId="205" pageNumber="205" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D93CFE8A1E75FEFB" author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[823,1047,292,316]" pageId="205" pageNumber="205" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
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;
|
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<bibRefCitation author="Carneiro, L. M. & E. V. Oliveira" pageId="205" pageNumber="205" pagination="355 - 372" refId="ref196481" refString="Carneiro, L. M., and E. V. Oliveira. 2017 a. Systematic affinities of the extinct metatherian Eobrasilia coutoi Simpson, 1947, a South American early Eocene Stagodontidae: implications for " Eobrasiliinae. " Revista Brasileira de Paleontologia 20 (3): 355 - 372." type="journal article" year="2017">Carneiro and Oliveira, 2017a</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DE2DFE8A1937FE9A" author="Carneiro, L. M. & E. V. Oliveira" pageId="205" pageNumber="205" pagination="355 - 372" refId="ref196481" refString="Carneiro, L. M., and E. V. Oliveira. 2017 a. Systematic affinities of the extinct metatherian Eobrasilia coutoi Simpson, 1947, a South American early Eocene Stagodontidae: implications for " Eobrasiliinae. " Revista Brasileira de Paleontologia 20 (3): 355 - 372." type="journal article" year="2017">Carneiro and Oliveira, 2017a</bibRefCitation>
|
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;
|
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<bibRefCitation author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[866,1096,325,349]" pageId="205" pageNumber="205" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al., 2018</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D969FEEB1E2AFE9A" author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[866,1096,325,349]" pageId="205" pageNumber="205" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al., 2018</bibRefCitation>
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;
|
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<bibRefCitation author="Carneiro, L. M." pageId="205" pageNumber="205" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">Carneiro, 2019</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4DE5EFEEB1881FEB9" author="Carneiro, L. M." pageId="205" pageNumber="205" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">Carneiro, 2019</bibRefCitation>
|
||||
) but not all (e.g.,
|
||||
<bibRefCitation author="Ladeveze, S. & C. de Muizon" pageId="205" pageNumber="205" pagination="746 - 784" refId="ref212638" refString="Ladeveze, S., and C. de Muizon. 2010. Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the late Palaeocene of Itaborai (Brazil) based on teeth and petrosal bones. Zoological Journal of the Linnean Society 159 (3): 746 - 784." type="journal article" year="2010">Ladevèze and Muizon, 2010</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F668E4D9B0FEC81881FE58" author="Ladeveze, S. & C. de Muizon" pageId="205" pageNumber="205" pagination="746 - 784" refId="ref212638" refString="Ladeveze, S., and C. de Muizon. 2010. Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the late Palaeocene of Itaborai (Brazil) based on teeth and petrosal bones. Zoological Journal of the Linnean Society 159 (3): 746 - 784." type="journal article" year="2010">Ladevèze and Muizon, 2010</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Wilson" box="[751,960,391,415]" etAl="et al." firstAuthor="Wilson" pageId="205" pageNumber="205" pagination="13734" refId="ref234583" refString="Wilson, G. P., E. G. Ekdale, J. W. Hoganson, J. J. Calede, and A. Vander Linden. 2016. A large carnivorous mammal from the Late Cretaceous and the North American origin of marsupials. Nature Communications 7: 13734." type="journal article" year="2016">Wilson et al., 2016</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D8E4FE2919A2FE58" author="Wilson" box="[751,960,391,415]" etAl="et al." firstAuthor="Wilson" pageId="205" pageNumber="205" pagination="13734" refId="ref234583" refString="Wilson, G. P., E. G. Ekdale, J. W. Hoganson, J. J. Calede, and A. Vander Linden. 2016. A large carnivorous mammal from the Late Cretaceous and the North American origin of marsupials. Nature Communications 7: 13734." type="journal article" year="2016">Wilson et al., 2016</bibRefCitation>
|
||||
) recent morphological analyses. This overall pattern is confirmed here: our morphological analyses did not recover australidelphian monophyly (figs. 30, 31), whereas it was strongly supported in all our molecular (figs. 27–29) and total evidence (figs. 32,
|
||||
<figureCitation box="[1074,1102,556,580]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="205" pageNumber="205">33</figureCitation>
|
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<figureCitation id="137D70CEF6F668E4DE39FD821E2CFD83" box="[1074,1102,556,580]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="205" pageNumber="205">33</figureCitation>
|
||||
) analyses. Given that
|
||||
<bibRefCitation author="Szalay" box="[810,969,589,613]" firstAuthor="Szalay" pageId="205" pageNumber="205" pagination="621 - 640" refId="ref228988" refString="Szalay, F. S. 1982 a. A new appraisal of marsupial phylogeny and classification. In M. Archer (editor), Carnivorous marsupials: 621 - 640. Mosman, New South Wales: Royal Zoological Society of New South Wales." type="book chapter" year="1982" yearSuffix="a">Szalay (1982a)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4D921FDE319ABFDA2" author="Szalay" box="[810,969,589,613]" firstAuthor="Szalay" pageId="205" pageNumber="205" pagination="621 - 640" refId="ref228988" refString="Szalay, F. S. 1982 a. A new appraisal of marsupial phylogeny and classification. In M. Archer (editor), Carnivorous marsupials: 621 - 640. Mosman, New South Wales: Royal Zoological Society of New South Wales." type="book chapter" year="1982" yearSuffix="a">Szalay (1982a)</bibRefCitation>
|
||||
first proposed monophyly of Australidelphia based primarily on shared derived features of the tarsus (specifically the presence of a continuous lower ankle joint, and a tripartite calcaneocuboid facet; see also
|
||||
<bibRefCitation author="Szalay" box="[684,820,755,779]" firstAuthor="Szalay" pageId="205" pageNumber="205" refId="ref229131" refString="Szalay, F. S. 1994. Evolutionary history of the marsupials and an analysis of osteological characters, Cambridge: Cambridge University Press." type="book" year="1994">Szalay, 1994</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F668E4D8A7FD5D1956FCCC" author="Szalay" box="[684,820,755,779]" firstAuthor="Szalay" pageId="205" pageNumber="205" refId="ref229131" refString="Szalay, F. S. 1994. Evolutionary history of the marsupials and an analysis of osteological characters, Cambridge: Cambridge University Press." type="book" year="1994">Szalay, 1994</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D." box="[833,959,755,779]" pageId="205" pageNumber="205" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F668E4D94AFD5D19DDFCCC" author="Beck, R. M. D." box="[833,959,755,779]" pageId="205" pageNumber="205" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
||||
), the addition of postcranial characters to our craniodental matrix may ultimately result in morphological support for Australidelphia (as in Horovitz and SánchezVillagra, 2003).
|
||||
</paragraph>
|
||||
<paragraph blockId="205.[684,1213,226,1738]" lastBlockId="206.[108,637,226,1738]" lastPageId="206" lastPageNumber="206" pageId="205" pageNumber="205">
|
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<paragraph id="8BF96C4BF6F668E7D8C6FC371BD9FEDC" blockId="205.[684,1213,226,1738]" lastBlockId="206.[108,637,226,1738]" lastPageId="206" lastPageNumber="206" pageId="205" pageNumber="205">
|
||||
No craniodental feature optimizes as an unambiguous synapomorphy of Australidelphia in our dated total-evidence analysis, but three optimize as synapomorphies under Accelerated Transformation—extracranial course of mandibular nerve fully enclosed by medial outgrowths of the auditory bulla (char. 52: 0→1; ci = 0.231); posterior limb of ectotympanic in contact with, but suturally distinct from, pars canalicularis of the petrosal and/or posttympanic process of the squamosal (char. 60: 0→1; ci = 0.333); and prootic canal foramen on tympanic face of petrosal absent (char. 69: 0→1; ci = 0.083)—but all three traits show high levels of homoplasy and undergo subsequent reversals within Australidelphia (see file S
|
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the online supplement). Our failure to identify compelling craniodental synapomorphies for Australidelphia likely reflects the fact that the ancestral australidelphian probably had a relatively generalized cranium and dentition that was little different from the plesiomorphic marsupial condition (see
|
||||
<bibRefCitation author="Szalay" box="[847,1033,1647,1672]" firstAuthor="Szalay" page="346" pageId="205" pageNumber="205" refId="ref229131" refString="Szalay, F. S. 1994. Evolutionary history of the marsupials and an analysis of osteological characters, Cambridge: Cambridge University Press." type="book" year="1994">Szalay, 1994: 346</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F668E4D944F9C11E6BF940" author="Szalay" box="[847,1033,1647,1672]" firstAuthor="Szalay" page="346" pageId="205" pageNumber="205" refId="ref229131" refString="Szalay, F. S. 1994. Evolutionary history of the marsupials and an analysis of osteological characters, Cambridge: Cambridge University Press." type="book" year="1994">Szalay, 1994: 346</bibRefCitation>
|
||||
) and that different lineages within Australidelphia subsequently evolved very disparate apomorphies of the dentition, cranium, or both (see also comments by
|
||||
<bibRefCitation author="Archer, M." box="[217,431,259,283]" pageId="206" pageNumber="206" pagination="633 - 808" refId="ref190081" refString="Archer, M. 1984 c. The Australian marsupial radiation. In M. Archer and G. Clayton (editors), Vertebrate zoogeography and evolution in Australasia: 633 - 808. Perth: Hesperian Press." type="book chapter" year="1984">Archer, 1984c: 782</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7DAD2FEAD1BCDFEDC" author="Archer, M." box="[217,431,259,283]" pageId="206" pageNumber="206" pagination="633 - 808" refId="ref190081" refString="Archer, M. 1984 c. The Australian marsupial radiation. In M. Archer and G. Clayton (editors), Vertebrate zoogeography and evolution in Australasia: 633 - 808. Perth: Hesperian Press." type="book chapter" year="1984">Archer, 1984c: 782</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[108,637,226,1738]" pageId="206" pageNumber="206">
|
||||
<paragraph id="8BF96C4BF6F568E7DA86FE8A1BDCFC8A" blockId="206.[108,637,226,1738]" pageId="206" pageNumber="206">
|
||||
Our estimate for the first split within Australidelphia is in the early to middle Eocene. This is younger than the early or middle Paleocene †
|
||||
<taxonomicName authorityName="Marshall & de Muizon" authorityYear="1988" box="[185,418,391,415]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="species">
|
||||
<emphasis box="[185,418,391,415]" italics="true" pageId="206" pageNumber="206">Khasia cordillerensis</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7DAB2FE291BC0FE58" authorityName="Marshall & de Muizon" authorityYear="1988" box="[185,418,391,415]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="species" species="cordillerensis">
|
||||
<emphasis id="B932B059F6F568E7DAB2FE291BC0FE58" box="[185,418,391,415]" italics="true" pageId="206" pageNumber="206">Khasia cordillerensis</emphasis>
|
||||
</taxonomicName>
|
||||
, a taxon that was originally described as a microbiotherian (
|
||||
<bibRefCitation author="Marshall, L. G. & C. de Muizon" pageId="206" pageNumber="206" pagination="23 - 55" refId="ref215460" refString="Marshall, L. G., and C. de Muizon. 1988. The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall and Muizon, 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D848FE061B11FE26" author="Marshall, L. G. & C. de Muizon" pageId="206" pageNumber="206" pagination="23 - 55" refId="ref215460" refString="Marshall, L. G., and C. de Muizon. 1988. The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall and Muizon, 1988</bibRefCitation>
|
||||
; see also
|
||||
<bibRefCitation author="Muizon, C. de" box="[476,631,457,481]" pageId="206" pageNumber="206" pagination="575 - 624" refId="ref217663" refString="Muizon, C. de. 1991. La fauna de mamiferos de Tiupampa (Paleoceno Inferior, Formacion Santa Lucia), Bolivia. In R. Suarez-Soruco (editor), Fosiles y facies de Bolivia, vol. 1. Vertebrados: 575 - 624. Santa Cruz, Bolivia: Revista Technica de Yacimientos Petroliferos Fiscales Bolivianos." type="book chapter" year="1991">Muizon, 1991</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBD7FE671815FE26" author="Muizon, C. de" box="[476,631,457,481]" pageId="206" pageNumber="206" pagination="575 - 624" refId="ref217663" refString="Muizon, C. de. 1991. La fauna de mamiferos de Tiupampa (Paleoceno Inferior, Formacion Santa Lucia), Bolivia. In R. Suarez-Soruco (editor), Fosiles y facies de Bolivia, vol. 1. Vertebrados: 575 - 624. Santa Cruz, Bolivia: Revista Technica de Yacimientos Petroliferos Fiscales Bolivianos." type="book chapter" year="1991">Muizon, 1991</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Cozzuol, M. A. & F. Goin & M. De Los Reyes & A. Ranzi" box="[109,302,490,514]" pageId="206" pageNumber="206" pagination="663 - 667" refId="ref198725" refString="Cozzuol, M. A., F. Goin, M. De Los Reyes, and A. Ranzi. 2006. The oldest species of Didelphis (Mammalia, Marsupialia, Didelphidae), from the late Miocene of Amazonia. Journal of Mammalogy 87 (4): 663 - 667." type="journal article" year="2006">Goin et al., 2006</bibRefCitation>
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||||
<bibRefCitation id="EFD711BAF6F568E7DA66FE441B4CFDC5" author="Cozzuol, M. A. & F. Goin & M. De Los Reyes & A. Ranzi" box="[109,302,490,514]" pageId="206" pageNumber="206" pagination="663 - 667" refId="ref198725" refString="Cozzuol, M. A., F. Goin, M. De Los Reyes, and A. Ranzi. 2006. The oldest species of Didelphis (Mammalia, Marsupialia, Didelphidae), from the late Miocene of Amazonia. Journal of Mammalogy 87 (4): 663 - 667." type="journal article" year="2006">Goin et al., 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Muizon, C. de & S. Ladeveze & C. Selva & R. Vignaud & F. Goussard" box="[315,537,490,514]" pageId="206" pageNumber="206" pagination="363 - 459" refId="ref218213" refString="Muizon, C. de, S. Ladeveze, C. Selva, R. Vignaud, and F. Goussard. 2018. Allqokirus australis (Sparassodonta, Metatheria) from the early Palaeocene of Tiupampa (Bolivia) and the rise of the metatherian carnivorous radiation in South America. Geodiversitas 40 (16): 363 - 459." type="journal article" year="2018">Muizon et al., 2018</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DB30FE44187BFDC5" author="Muizon, C. de & S. Ladeveze & C. Selva & R. Vignaud & F. Goussard" box="[315,537,490,514]" pageId="206" pageNumber="206" pagination="363 - 459" refId="ref218213" refString="Muizon, C. de, S. Ladeveze, C. Selva, R. Vignaud, and F. Goussard. 2018. Allqokirus australis (Sparassodonta, Metatheria) from the early Palaeocene of Tiupampa (Bolivia) and the rise of the metatherian carnivorous radiation in South America. Geodiversitas 40 (16): 363 - 459." type="journal article" year="2018">Muizon et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Ladeveze, S. & C. Selva & C. de Muizon" pageId="206" pageNumber="206" pagination="1463 - 1479" refId="ref212907" refString="Ladeveze, S., C. Selva, and C. de Muizon. 2020. What are " opossum-like " fossils? The phylogeny of herpetotheriid and peradectid metatherians, based on new features from the petrosal anatomy. Journal of Systematic Palaeontology 18 (17): 1463 - 1479." type="journal article" year="2020">Muizon and Ladevèze, 2020</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D82CFE441B2CFDE4" author="Ladeveze, S. & C. Selva & C. de Muizon" pageId="206" pageNumber="206" pagination="1463 - 1479" refId="ref212907" refString="Ladeveze, S., C. Selva, and C. de Muizon. 2020. What are " opossum-like " fossils? The phylogeny of herpetotheriid and peradectid metatherians, based on new features from the petrosal anatomy. Journal of Systematic Palaeontology 18 (17): 1463 - 1479." type="journal article" year="2020">Muizon and Ladevèze, 2020</bibRefCitation>
|
||||
) and hence a crown-clade australidelphian. However, as noted above, several subsequent authors have argued that †
|
||||
<taxonomicName class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7D84EFDE31AE8FD41" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F568E7D84EFDE31AE8FD41" italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
</taxonomicName>
|
||||
is more likely a “pediomyoid” (
|
||||
<bibRefCitation author="Oliveira and Goin" firstAuthor="Oliveira" pageId="206" pageNumber="206" pagination="299 - 320" refId="ref220101" refString="Oliveira, E. V., and F. J. Goin. 2006. Marsupiais do inicio do Terciario do Brasil: origem, irradiacao e historia biogeografica. In N. C. Caceres and E. L. A. Monteiro Filho (editors), Os Marsupiais do Brasil: biologia, ecologia e evolucao: 299 - 320. Campo Grande: UFMS." type="book chapter" year="2006">Oliveira and Goin, 2006</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBE4FDC01A88FD60" author="Oliveira and Goin" firstAuthor="Oliveira" pageId="206" pageNumber="206" pagination="299 - 320" refId="ref220101" refString="Oliveira, E. V., and F. J. Goin. 2006. Marsupiais do inicio do Terciario do Brasil: origem, irradiacao e historia biogeografica. In N. C. Caceres and E. L. A. Monteiro Filho (editors), Os Marsupiais do Brasil: biologia, ecologia e evolucao: 299 - 320. Campo Grande: UFMS." type="book chapter" year="2006">Oliveira and Goin, 2006</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Goin, F. J. & M. O. Woodburne & A. N. Zimicz & G. M. Martin & L. Chornogubsky" box="[244,428,655,679]" pageId="206" pageNumber="206" refId="ref207099" refString="Goin, F. J., M. O. Woodburne, A. N. Zimicz, G. M. Martin, and L. Chornogubsky. 2016. A brief history of South American metatherians: Evolutionary contexts and intercontinental dispersals. Dordrecht: Springer." type="book" year="2016">Goin et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DAFFFD211BCEFD60" author="Goin, F. J. & M. O. Woodburne & A. N. Zimicz & G. M. Martin & L. Chornogubsky" box="[244,428,655,679]" pageId="206" pageNumber="206" refId="ref207099" refString="Goin, F. J., M. O. Woodburne, A. N. Zimicz, G. M. Martin, and L. Chornogubsky. 2016. A brief history of South American metatherians: Evolutionary contexts and intercontinental dispersals. Dordrecht: Springer." type="book" year="2016">Goin et al., 2016</bibRefCitation>
|
||||
), a hypothesis supported by the morphological phylogenetic analysis of
|
||||
<bibRefCitation author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[190,440,722,746]" pageId="206" pageNumber="206" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al. (2018)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DAB5FD7C1BDAFD2D" author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[190,440,722,746]" pageId="206" pageNumber="206" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al. (2018)</bibRefCitation>
|
||||
. †
|
||||
<taxonomicName box="[467,543,722,746]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[467,543,722,746]" italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7DBD8FD7C187DFD2D" box="[467,543,722,746]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F568E7DBD8FD7C187DFD2D" box="[467,543,722,746]" italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
</taxonomicName>
|
||||
has not been included here because it is represented only by dental specimens (
|
||||
<bibRefCitation author="Marshall, L. G. & C. de Muizon" pageId="206" pageNumber="206" pagination="23 - 55" refId="ref215460" refString="Marshall, L. G., and C. de Muizon. 1988. The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall and Muizon, 1988</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBDFFCBA1B6AFC8A" author="Marshall, L. G. & C. de Muizon" pageId="206" pageNumber="206" pagination="23 - 55" refId="ref215460" refString="Marshall, L. G., and C. de Muizon. 1988. The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall and Muizon, 1988</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Muizon, C. de" box="[276,434,821,845]" pageId="206" pageNumber="206" pagination="575 - 624" refId="ref217663" refString="Muizon, C. de. 1991. La fauna de mamiferos de Tiupampa (Paleoceno Inferior, Formacion Santa Lucia), Bolivia. In R. Suarez-Soruco (editor), Fosiles y facies de Bolivia, vol. 1. Vertebrados: 575 - 624. Santa Cruz, Bolivia: Revista Technica de Yacimientos Petroliferos Fiscales Bolivianos." type="book chapter" year="1991">Muizon, 1991</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DB1FFC9B1BD0FC8A" author="Muizon, C. de" box="[276,434,821,845]" pageId="206" pageNumber="206" pagination="575 - 624" refId="ref217663" refString="Muizon, C. de. 1991. La fauna de mamiferos de Tiupampa (Paleoceno Inferior, Formacion Santa Lucia), Bolivia. In R. Suarez-Soruco (editor), Fosiles y facies de Bolivia, vol. 1. Vertebrados: 575 - 624. Santa Cruz, Bolivia: Revista Technica de Yacimientos Petroliferos Fiscales Bolivianos." type="book chapter" year="1991">Muizon, 1991</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[108,637,226,1738]" pageId="206" pageNumber="206">
|
||||
<paragraph id="8BF96C4BF6F568E7DA86FCF8183CF9C4" blockId="206.[108,637,226,1738]" pageId="206" pageNumber="206">
|
||||
If †
|
||||
<taxonomicName box="[181,258,854,878]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[181,258,854,878]" italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7DABEFCF81B60FCA9" box="[181,258,854,878]" class="Mammalia" family="Microbiotheriidae" genus="Khasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F568E7DABEFCF81B60FCA9" box="[181,258,854,878]" italics="true" pageId="206" pageNumber="206">Khasia</emphasis>
|
||||
</taxonomicName>
|
||||
is discounted, the oldest known australidelphian is probably †
|
||||
<taxonomicName class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="species">
|
||||
<emphasis italics="true" pageId="206" pageNumber="206">Djarthia murgonensis</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7DBC9FCD91AD2FC77" class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="species" species="murgonensis">
|
||||
<emphasis id="B932B059F6F568E7DBC9FCD91AD2FC77" italics="true" pageId="206" pageNumber="206">Djarthia murgonensis</emphasis>
|
||||
</taxonomicName>
|
||||
from the Tingamarra fossil site in eastern
|
||||
<collectingCountry box="[155,261,953,977]" name="Australia" pageId="206" pageNumber="206">Australia</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F568E7DA90FC171B67FC16" box="[155,261,953,977]" name="Australia" pageId="206" pageNumber="206">Australia</collectingCountry>
|
||||
, which has been radiometrically dated as earliest Eocene (~54.6 Mya; (
|
||||
<bibRefCitation author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" pageId="206" pageNumber="206" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D835FC741B53FBD4" author="Godthelp, H. & M. Archer & R. L. Cifelli & S. J. Hand & C. F. Gilkeson" pageId="206" pageNumber="206" pagination="514 - 516" refId="ref205513" refString="Godthelp, H., M. Archer, R. L. Cifelli, S. J. Hand, and C. F. Gilkeson. 1992. Earliest known Australian Tertiary mammal fauna. Nature 356: 514 - 516." type="journal article" year="1992">Godthelp et al., 1992</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Godthelp, H. & S. Wroe & M. Archer" box="[320,567,1019,1043]" pageId="206" pageNumber="206" pagination="289 - 313" refId="ref205557" refString="Godthelp, H., S. Wroe, and M. Archer. 1999. A new marsupial from the early Eocene Tingamarra Local Fauna of Murgon, southeastern Queensland: a prototypical Australian marsupial? Journal of Mammalian Evolution 6 (3): 289 - 313." type="journal article" year="1999">Godthelp et al., 1999</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DB4BFC551855FBD4" author="Godthelp, H. & S. Wroe & M. Archer" box="[320,567,1019,1043]" pageId="206" pageNumber="206" pagination="289 - 313" refId="ref205557" refString="Godthelp, H., S. Wroe, and M. Archer. 1999. A new marsupial from the early Eocene Tingamarra Local Fauna of Murgon, southeastern Queensland: a prototypical Australian marsupial? Journal of Mammalian Evolution 6 (3): 289 - 313." type="journal article" year="1999">Godthelp et al., 1999</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D84EFC551A98FBF3" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
). We have not included †
|
||||
<taxonomicName authorityName="Godthelp, Wroe & Archer" authorityYear="1999" box="[540,636,1052,1076]" class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[540,636,1052,1076]" italics="true" pageId="206" pageNumber="206">Djarthia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7D817FBB2181EFBF3" authorityName="Godthelp, Wroe & Archer" authorityYear="1999" box="[540,636,1052,1076]" class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F568E7D817FBB2181EFBF3" box="[540,636,1052,1076]" italics="true" pageId="206" pageNumber="206">Djarthia</emphasis>
|
||||
</taxonomicName>
|
||||
here due to its incompleteness (the only craniodental remains are incomplete dental specimens and isolated petrosals;
|
||||
<bibRefCitation author="Godthelp, H. & S. Wroe & M. Archer" pageId="206" pageNumber="206" pagination="289 - 313" refId="ref205557" refString="Godthelp, H., S. Wroe, and M. Archer. 1999. A new marsupial from the early Eocene Tingamarra Local Fauna of Murgon, southeastern Queensland: a prototypical Australian marsupial? Journal of Mammalian Evolution 6 (3): 289 - 313." type="journal article" year="1999">Godthelp et al., 1999</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBCFFBD11AC7FB7E" author="Godthelp, H. & S. Wroe & M. Archer" pageId="206" pageNumber="206" pagination="289 - 313" refId="ref205557" refString="Godthelp, H., S. Wroe, and M. Archer. 1999. A new marsupial from the early Eocene Tingamarra Local Fauna of Murgon, southeastern Queensland: a prototypical Australian marsupial? Journal of Mammalian Evolution 6 (3): 289 - 313." type="journal article" year="1999">Godthelp et al., 1999</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[180,392,1184,1209]" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DABFFB0E1BEAFB7E" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[180,392,1184,1209]" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
), but isolated tarsals referred to this taxon exhibit characteristic australidelphian synapomorphies (
|
||||
<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBF1FB4D1AD5FADB" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">Beck et al., 2008a</bibRefCitation>
|
||||
). However, †
|
||||
<taxonomicName authorityName="Godthelp, Wroe & Archer" authorityYear="1999" box="[360,461,1284,1308]" class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis box="[360,461,1284,1308]" italics="true" pageId="206" pageNumber="206">Djarthia</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7DB63FAAA1BAFFADB" authorityName="Godthelp, Wroe & Archer" authorityYear="1999" box="[360,461,1284,1308]" class="Mammalia" family="Marsupialia" genus="Djarthia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="206" pageNumber="206" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B932B059F6F568E7DB63FAAA1BAFFADB" box="[360,461,1284,1308]" italics="true" pageId="206" pageNumber="206">Djarthia</emphasis>
|
||||
</taxonomicName>
|
||||
falls outside crown-clade Australidelphia in most published analyses (with the notable exception of
|
||||
<bibRefCitation author="Maga, A. M. & R. M. D. Beck" pageId="206" pageNumber="206" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D834FAE81B7CFAB8" author="Maga, A. M. & R. M. D. Beck" pageId="206" pageNumber="206" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
: fig. 38, in which it is sister to
|
||||
<taxonomicName baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[110,312,1416,1440]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7DA65FA261B5AFA67" baseAuthorityName="Kealy and Beck" baseAuthorityYear="2017" box="[110,312,1416,1440]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
|
||||
), and its position is unresolved with respect to the crown clade in others (
|
||||
<bibRefCitation author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[156,358,1482,1506]" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
|
||||
<bibRefCitation id="EFD711BAF6F568E7DA97FA641B04FA25" author="Beck, R. M. D. & H. Godthelp & V. Weisbecker & M. Archer & S. J. Hand" box="[156,358,1482,1506]" pageId="206" pageNumber="206" pagination="1858" refId="ref192866" refString="Beck, R. M. D., H. Godthelp, V. Weisbecker, M. Archer, and S. J. Hand. 2008 a. Australia's oldest marsupial fossils and their biogeographical implications. PLoS One 3 (3): e 1858." type="journal article" year="2008">
|
||||
Beck et al., 2008
|
||||
<bibRefCitation author="Rambaut, A. & M. A. Suchard & D. Xie & A. J. Drummond" box="[346,427,1482,1506]" pageId="206" pageNumber="206" refId="ref223023" refString="Rambaut, A., M. A. Suchard, D. Xie, and A. J. Drummond. 2014. Tracer v 1.6." type="book" year="2014">a, 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DB51FA641B04FA25" author="Rambaut, A. & M. A. Suchard & D. Xie & A. J. Drummond" box="[346,358,1482,1506]" pageId="206" pageNumber="206" refId="ref223023" refString="Rambaut, A., M. A. Suchard, D. Xie, and A. J. Drummond. 2014. Tracer v 1.6." type="book" year="2014">a</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[438,495,1482,1506]" pageId="206" pageNumber="206" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
|
||||
, 2014,
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBBDFA641B8DFA25" author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[438,495,1482,1506]" pageId="206" pageNumber="206" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Beck, R. M. D." box="[506,631,1482,1506]" pageId="206" pageNumber="206" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DBF1FA641815FA25" author="Beck, R. M. D." box="[506,631,1482,1506]" pageId="206" pageNumber="206" pagination="715 - 729" refId="ref192633" refString="Beck, R. M. D. 2012. An ' ameridelphian' marsupial from the early Eocene of Australia supports a complex model of Southern Hemisphere marsupial biogeography. Naturwissenschaften 99 (9): 715 - 729." type="journal article" year="2012">Beck, 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[109,332,1515,1539]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DA66FA421B2EF9C4" author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[109,332,1515,1539]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Maga, A. M. & R. M. D. Beck" box="[344,593,1515,1540]" pageId="206" pageNumber="206" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7DB53FA421833F9C4" author="Maga, A. M. & R. M. D. Beck" box="[344,593,1515,1540]" pageId="206" pageNumber="206" pagination="0181712" refId="ref214955" refString="Maga, A. M., and R. M. D. Beck. 2017. Skeleton of an unusual, cat-sized marsupial relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44 - 43 million years ago) of Turkey. PLoS One 12 (8): e 0181712." type="journal article" year="2017">Maga and Beck, 2017</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[108,637,226,1738]" lastBlockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
<paragraph id="8BF96C4BF6F568E7DA87F9A219ECFE58" blockId="206.[108,637,226,1738]" lastBlockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
The next-oldest definitive australidelphian remains are isolated tarsals from the La Barda locality in
|
||||
<collectingCountry box="[225,337,1614,1638]" name="Argentina" pageId="206" pageNumber="206">Argentina</collectingCountry>
|
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<collectingCountry id="F3512CDBF6F568E7DAEAF9E01B33F9A1" box="[225,337,1614,1638]" name="Argentina" pageId="206" pageNumber="206">Argentina</collectingCountry>
|
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, which has been radiometrically dated as middle Eocene (between ~48 and 43 Mya;
|
||||
<bibRefCitation author="Tejedor" box="[200,405,1680,1705]" etAl="et al." firstAuthor="Tejedor" pageId="206" pageNumber="206" pagination="1 - 43" refId="ref229849" refString="Tejedor, M. F., et al. 2009. New early Eocene mammalian fauna from western Patagonia, Argentina. American Museum Novitates 3638: 1 - 43." type="journal article" year="2009">Tejedor et al., 2009</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7DAC3F93E1BF7F96E" author="Tejedor" box="[200,405,1680,1705]" etAl="et al." firstAuthor="Tejedor" pageId="206" pageNumber="206" pagination="1 - 43" refId="ref229849" refString="Tejedor, M. F., et al. 2009. New early Eocene mammalian fauna from western Patagonia, Argentina. American Museum Novitates 3638: 1 - 43." type="journal article" year="2009">Tejedor et al., 2009</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[415,625,1680,1704]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7DB94F93F1813F96F" author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[415,625,1680,1704]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al., 2016</bibRefCitation>
|
||||
). The La Barda australidelphian tarsals fell within
|
||||
<taxonomicName authorityName="Owen" authorityYear="1866" box="[686,842,226,250]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7D8A5FF4C1928FF3D" authorityName="Owen" authorityYear="1866" box="[686,842,226,250]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||||
(members of which are otherwise known only from
|
||||
<collectingCountry box="[896,999,259,283]" name="Australia" pageId="206" pageNumber="206">Australia</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F568E7D98BFEAD1985FEDC" box="[896,999,259,283]" name="Australia" pageId="206" pageNumber="206">Australia</collectingCountry>
|
||||
, New
|
||||
<collectingCountry box="[1074,1156,259,283]" name="Guinea-Bissau" pageId="206" pageNumber="206">Guinea</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F568E7DE39FEAD1EE6FEDC" box="[1074,1156,259,283]" name="Guinea-Bissau" pageId="206" pageNumber="206">Guinea</collectingCountry>
|
||||
, and adjacent islands) in the phylogenetic analysis of
|
||||
<bibRefCitation author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[684,917,325,349]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al. (2016)</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D8A7FEEB19F7FE9A" author="Lorente, M. & L. Chornogubsky & F. J. Goin" box="[684,917,325,349]" pageId="206" pageNumber="206" pagination="533 - 547" refId="ref213633" refString="Lorente, M., L. Chornogubsky, and F. J. Goin. 2016. On the existence of non-microbiotherian australidelphian marsupials (Diprotodontia) in the Eocene of Patagonia. Palaeontology 59 (4): 533 - 547." type="journal article" year="2016">Lorente et al. (2016)</bibRefCitation>
|
||||
, but we consider that this biogeographically anomalous relationship warrants further testing.
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
<paragraph id="8BF96C4BF6F568E7D8C7FE061E28FB91" blockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
The woodburnodontid microbiotherian †
|
||||
<taxonomicName box="[698,939,457,481]" class="Mammalia" family="Woodburnodontidae" genus="Woodburnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="species">
|
||||
<emphasis box="[698,939,457,481]" italics="true" pageId="206" pageNumber="206">Woodburnodon casei</emphasis>
|
||||
<taxonomicName id="4C4617C8F6F568E7D8B1FE6719C9FE26" box="[698,939,457,481]" class="Mammalia" family="Woodburnodontidae" genus="Woodburnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="species" species="casei">
|
||||
<emphasis id="B932B059F6F568E7D8B1FE6719C9FE26" box="[698,939,457,481]" italics="true" pageId="206" pageNumber="206">Woodburnodon casei</emphasis>
|
||||
</taxonomicName>
|
||||
from the Cucullaea I Allomember of the La Meseta Formation on Seymour Island, off the Antarctic Penninsula, is a more compelling candidate for the oldest definitive crown-clade australidelphian because it preserves distinctive dental features that are characteristic of microbiotherians (
|
||||
<bibRefCitation author="Goin, F. J." pageId="206" pageNumber="206" pagination="597 - 603" refId="ref206658" refString="Goin, F. J., et al. 2007 c. New marsupial (Mammalia) from the Eocene of Antarctica, and the origins and affinities of the Microbiotheria. Revista de la Asociacion Geologica Argentina 62 (4): 597 - 603." type="journal article" year="2007">Goin et al., 2007c</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DE31FD21188DFD0E" author="Goin, F. J." pageId="206" pageNumber="206" pagination="597 - 603" refId="ref206658" refString="Goin, F. J., et al. 2007 c. New marsupial (Mammalia) from the Eocene of Antarctica, and the origins and affinities of the Microbiotheria. Revista de la Asociacion Geologica Argentina 62 (4): 597 - 603." type="journal article" year="2007">Goin et al., 2007c</bibRefCitation>
|
||||
), and it has been recovered within
|
||||
<taxonomicName authorityName="Ameghino" authorityYear="1889" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7DE7BFD1E196CFD2D" authorityName="Ameghino" authorityYear="1889" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
in several phylogenetic analyses (
|
||||
<bibRefCitation author="Carneiro, L. M. & E. V. Oliveira" pageId="206" pageNumber="206" pagination="497 - 507" refId="ref196536" refString="Carneiro, L. M., and E. V. Oliveira. 2017 b. The Eocene South American metatherian Zeusdelphys complicatus is not a protodidelphid but a hatcheriform: Paleobiogeographic implications. Acta Palaeontologica Polonica 62 (3): 497 - 507." type="journal article" year="2017">Carneiro and Oliveira, 2017b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DE82FD7C19AAFCCC" author="Carneiro, L. M. & E. V. Oliveira" pageId="206" pageNumber="206" pagination="497 - 507" refId="ref196536" refString="Carneiro, L. M., and E. V. Oliveira. 2017 b. The Eocene South American metatherian Zeusdelphys complicatus is not a protodidelphid but a hatcheriform: Paleobiogeographic implications. Acta Palaeontologica Polonica 62 (3): 497 - 507." type="journal article" year="2017">Carneiro and Oliveira, 2017b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[981,1208,755,779]" pageId="206" pageNumber="206" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al., 2018</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D9DEFD5D1EDAFCCC" author="Carneiro, L. M. & E. V. Oliveira & F. J. Goin" box="[981,1208,755,779]" pageId="206" pageNumber="206" pagination="120 - 131" refId="ref196583" refString="Carneiro, L. M., E. V. Oliveira, and F. J. Goin. 2018. Austropediomys marshalli gen. et sp. nov., a new Pediomyoidea (Mammalia, Metatheria) from the Paleogene of Brazil: paleobiogeographic implications. Revista Brasileira de Paleontologia 21 (2): 120 - 131." type="journal article" year="2018">Carneiro et al., 2018</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Carneiro, L. M." box="[686,865,788,812]" pageId="206" pageNumber="206" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">Carneiro, 2019</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D8A5FCBA1903FCEB" author="Carneiro, L. M." box="[686,865,788,812]" pageId="206" pageNumber="206" pagination="20180440" refId="ref196433" refString="Carneiro, L. M. 2019. A new protodidelphid (Mammalia, Marsupialia, Didelphimorphia) from the Itaborai Basin and its implications for the evolution of the Protodidelphidae. Anais da Academia Brasileira de Ciencias 91 (Suppl. 2): e 20180440." type="journal article" year="2019">Carneiro, 2019</bibRefCitation>
|
||||
). The age of the Cucullaea I Allomember has proved controversial (
|
||||
<bibRefCitation author="Crame, J. A." pageId="206" pageNumber="206" pagination="114743" refId="ref198824" refString="Crame, J. A., et al. 2014. The early origin of the Antarctic marine fauna and its evolutionary implications. PLoS One 9 (12): e 114743." type="journal article" year="2014">Crame et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DE5DFC9B196FFCA9" author="Crame, J. A." pageId="206" pageNumber="206" pagination="114743" refId="ref198824" refString="Crame, J. A., et al. 2014. The early origin of the Antarctic marine fauna and its evolutionary implications. PLoS One 9 (12): e 114743." type="journal article" year="2014">Crame et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Kemp, D. B." box="[795,998,854,878]" pageId="206" pageNumber="206" pagination="583 - 586" refId="ref211003" refString="Kemp, D. B., et al. 2014. A cool temperate climate on the Antarctic Peninsula through the latest Cretaceous to early Paleogene. Geology 42 (7): 583 - 586." type="journal article" year="2014">Kemp et al., 2014</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7D910FCF81984FCA9" author="Kemp, D. B." box="[795,998,854,878]" pageId="206" pageNumber="206" pagination="583 - 586" refId="ref211003" refString="Kemp, D. B., et al. 2014. A cool temperate climate on the Antarctic Peninsula through the latest Cretaceous to early Paleogene. Geology 42 (7): 583 - 586." type="journal article" year="2014">Kemp et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Gelfo, J. N. & G. M. Lopez & S. N. Santillana" box="[1012,1209,854,878]" pageId="206" pageNumber="206" pagination="445 - 455" refId="ref204712" refString="Gelfo, J. N., G. M. Lopez, and S. N. Santillana. 2017. Eocene ungulate mammals from west Antarctica: implications from their fossil record and a new species. Antarctic Science 29 (05): 445 - 455." type="journal article" year="2017">Gelfo et al., 2017</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F568E7D9FFFCF81EDBFCA9" author="Gelfo, J. N. & G. M. Lopez & S. N. Santillana" box="[1012,1209,854,878]" pageId="206" pageNumber="206" pagination="445 - 455" refId="ref204712" refString="Gelfo, J. N., G. M. Lopez, and S. N. Santillana. 2017. Eocene ungulate mammals from west Antarctica: implications from their fossil record and a new species. Antarctic Science 29 (05): 445 - 455." type="journal article" year="2017">Gelfo et al., 2017</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Gelfo, J. N. & F. J. Goin & N. Bauza & M. Reguero" box="[684,738,887,911]" pageId="206" pageNumber="206" pagination="274 - 292" refId="ref204762" refString="Gelfo, J. N., F. J. Goin, N. Bauza, and M. Reguero. 2019. The fossil record of Antarctic land mammals: commented review and hypotheses for future research. Advances in Polar Science 30 (3): 274 - 292." type="journal article" year="2019">2019</bibRefCitation>
|
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<bibRefCitation id="EFD711BAF6F568E7D8A7FCD91880FC48" author="Gelfo, J. N. & F. J. Goin & N. Bauza & M. Reguero" box="[684,738,887,911]" pageId="206" pageNumber="206" pagination="274 - 292" refId="ref204762" refString="Gelfo, J. N., F. J. Goin, N. Bauza, and M. Reguero. 2019. The fossil record of Antarctic land mammals: commented review and hypotheses for future research. Advances in Polar Science 30 (3): 274 - 292." type="journal article" year="2019">2019</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Zimicz, A. N. & F. J. Goin" box="[749,931,887,911]" pageId="206" pageNumber="206" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Goin et al., 2020</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D8E6FCD919C1FC48" author="Zimicz, A. N. & F. J. Goin" box="[749,931,887,911]" pageId="206" pageNumber="206" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Goin et al., 2020</bibRefCitation>
|
||||
), but it now appears to be about 40 Mya (
|
||||
<bibRefCitation author="Douglas, P. M." box="[846,1062,920,944]" pageId="206" pageNumber="206" pagination="6582 - 6587" refId="ref200562" refString="Douglas, P. M., et al. 2014. Pronounced zonal heterogeneity in Eocene southern high-latitude sea surface temperatures. Proceedings of the National Academy of Sciences of the United States of America 111 (18): 6582 - 6587." type="journal article" year="2014">Douglas et al., 2014</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F568E7D945FC361E44FC77" author="Douglas, P. M." box="[846,1062,920,944]" pageId="206" pageNumber="206" pagination="6582 - 6587" refId="ref200562" refString="Douglas, P. M., et al. 2014. Pronounced zonal heterogeneity in Eocene southern high-latitude sea surface temperatures. Proceedings of the National Academy of Sciences of the United States of America 111 (18): 6582 - 6587." type="journal article" year="2014">Douglas et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Amenabar, C. R. & M. Montes & F. Nozal & S. Santillana" pageId="206" pageNumber="206" pagination="351 - 366" refId="ref188835" refString="Amenabar, C. R., M. Montes, F. Nozal, and S. Santillana. 2019. Dinoflagellate cysts of the La Meseta Formation (middle to late Eocene), Antarctic Peninsula: implications for biostratigraphy, palaeoceanography and palaeoenvironment. Geological Magazine 157 (3): 351 - 366." type="journal article" year="2019">Amenábar et al., 2019</bibRefCitation>
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<bibRefCitation id="EFD711BAF6F568E7DE3BFC36196EFC16" author="Amenabar, C. R. & M. Montes & F. Nozal & S. Santillana" pageId="206" pageNumber="206" pagination="351 - 366" refId="ref188835" refString="Amenabar, C. R., M. Montes, F. Nozal, and S. Santillana. 2019. Dinoflagellate cysts of the La Meseta Formation (middle to late Eocene), Antarctic Peninsula: implications for biostratigraphy, palaeoceanography and palaeoenvironment. Geological Magazine 157 (3): 351 - 366." type="journal article" year="2019">Amenábar et al., 2019</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Mors, T. & M. Reguero & D. Vasilyan" box="[793,986,953,977]" pageId="206" pageNumber="206" pagination="5051" refId="ref217290" refString="Mors, T., M. Reguero, and D. Vasilyan. 2020. First fossil frog from Antarctica: implications for Eocene high latitude climate conditions and Gondwanan cosmopolitanism of Australobatrachia. Scientific Reports 10: 5051." type="journal article" year="2020">Mörs et al., 2020</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D912FC1419B8FC16" author="Mors, T. & M. Reguero & D. Vasilyan" box="[793,986,953,977]" pageId="206" pageNumber="206" pagination="5051" refId="ref217290" refString="Mors, T., M. Reguero, and D. Vasilyan. 2020. First fossil frog from Antarctica: implications for Eocene high latitude climate conditions and Gondwanan cosmopolitanism of Australobatrachia. Scientific Reports 10: 5051." type="journal article" year="2020">Mörs et al., 2020</bibRefCitation>
|
||||
). This postdates our estimate for the first divergence within Australidelphia (see above) and also for the split between
|
||||
<taxonomicName authorityName="Ameghino" authorityYear="1889" box="[684,852,1052,1076]" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
<taxonomicName id="4C4617C8F6F568E7D8A7FBB21936FBF3" authorityName="Ameghino" authorityYear="1889" box="[684,852,1052,1076]" class="Mammalia" kingdom="Animalia" order="Microbiotheria" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Microbiotheria</taxonomicName>
|
||||
and
|
||||
<taxonomicName authorityName="Owen" authorityYear="1866" box="[918,1078,1052,1076]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||||
(median = 45.6 Mya; 95%
|
||||
<collectionCode box="[852,911,1086,1109]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="206" pageNumber="206" type="Herbarium">HPD</collectionCode>
|
||||
: 41.4–48.8 Mya).
|
||||
<taxonomicName id="4C4617C8F6F568E7D99DFBB21E54FBF3" authorityName="Owen" authorityYear="1866" box="[918,1078,1052,1076]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="206" pageNumber="206" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||||
(median = 45.6 Mya; 95% HPD: 41.4–48.8 Mya).
|
||||
</paragraph>
|
||||
<paragraph blockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
<paragraph id="8BF96C4BF6F568E7D8C7FBF01E66FA67" blockId="206.[684,1214,226,1441]" pageId="206" pageNumber="206">
|
||||
Older putative microbiotherians have been reported from the early Eocene (probably 51.4– 56.0 Mya;
|
||||
<bibRefCitation author="Clyde, W. C." box="[802,999,1184,1209]" pageId="206" pageNumber="206" pagination="289 - 306" refId="ref197816" refString="Clyde, W. C., et al. 2014. New age constraints for the Salamanca Formation and lower Rio Chico Group in the western San Jorge Basin, Patagonia, Argentina: implications for Cretaceous-Paleogene extinction recovery and land mammal age correlations. Geological Society of America Bulletin 126 (3 - 4): 289 - 306." type="journal article" year="2014">Clyde et al., 2014</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D929FB0E1985FB7E" author="Clyde, W. C." box="[802,999,1184,1209]" pageId="206" pageNumber="206" pagination="289 - 306" refId="ref197816" refString="Clyde, W. C., et al. 2014. New age constraints for the Salamanca Formation and lower Rio Chico Group in the western San Jorge Basin, Patagonia, Argentina: implications for Cretaceous-Paleogene extinction recovery and land mammal age correlations. Geological Society of America Bulletin 126 (3 - 4): 289 - 306." type="journal article" year="2014">Clyde et al., 2014</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Woodburne" etAl="et al." firstAuthor="Woodburne" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="a">
|
||||
<bibRefCitation id="EFD711BAF6F568E7D9FFFB0E1892FB1D" author="Woodburne" etAl="et al." firstAuthor="Woodburne" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="a">
|
||||
Woodburne et al., 2014
|
||||
<bibRefCitation author="a" box="[740,837,1218,1242]" etAl="et al." firstAuthor="a" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="b">a, 2014b</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D8EFFB6C1892FB1D" author="a" box="[740,752,1218,1242]" etAl="et al." firstAuthor="a" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="b">a</bibRefCitation>
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Krause, J. M." box="[854,1075,1218,1242]" pageId="206" pageNumber="206" pagination="886 - 903" refId="ref212274" refString="Krause, J. M., et al. 2017. New age constraints for early Paleogene strata of central Patagonia, Argentina: Implications for the timing of South American Land Mammal Ages. Geological Society of America Bulletin 129 (7 - 8): 886 - 903." type="journal article" year="2017">Krause et al., 2017</bibRefCitation>
|
||||
, 2014b;
|
||||
<bibRefCitation id="EFD711BAF6F568E7D95DFB6C1E51FB1D" author="Krause, J. M." box="[854,1075,1218,1242]" pageId="206" pageNumber="206" pagination="886 - 903" refId="ref212274" refString="Krause, J. M., et al. 2017. New age constraints for early Paleogene strata of central Patagonia, Argentina: Implications for the timing of South American Land Mammal Ages. Geological Society of America Bulletin 129 (7 - 8): 886 - 903." type="journal article" year="2017">Krause et al., 2017</bibRefCitation>
|
||||
) Las
|
||||
<collectingCountry box="[1144,1211,1218,1242]" name="Indonesia" pageId="206" pageNumber="206">Flores</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F568E7DE73FB6C1ED9FB1D" box="[1144,1211,1218,1242]" name="Indonesia" pageId="206" pageNumber="206">Flores</collectingCountry>
|
||||
Local Fauna of southern
|
||||
<collectingCountry box="[958,1067,1251,1275]" name="Argentina" pageId="206" pageNumber="206">Argentina</collectingCountry>
|
||||
<collectingCountry id="F3512CDBF6F568E7D9B5FB4D1E49FB3C" box="[958,1067,1251,1275]" name="Argentina" pageId="206" pageNumber="206">Argentina</collectingCountry>
|
||||
(
|
||||
<bibRefCitation author="Goin, F. J." box="[1083,1208,1251,1275]" pageId="206" pageNumber="206" pagination="30 - 42" refId="ref205827" refString="Goin, F. J. 2003. Early marsupial radiations in South America. In M. Jones, C. Dickman, and M. Archer (editors), Predators with pouches: the biology of carnivorous marsupials: 30 - 42. Collingwood, Australia: CSIRO (Commonwealth Scientific and Industrial Research Organization)." type="book chapter" year="2003">Goin, 2003</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DE30FB4D1EDAFB3C" author="Goin, F. J." box="[1083,1208,1251,1275]" pageId="206" pageNumber="206" pagination="30 - 42" refId="ref205827" refString="Goin, F. J. 2003. Early marsupial radiations in South America. In M. Jones, C. Dickman, and M. Archer (editors), Predators with pouches: the biology of carnivorous marsupials: 30 - 42. Collingwood, Australia: CSIRO (Commonwealth Scientific and Industrial Research Organization)." type="book chapter" year="2003">Goin, 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Zimicz" box="[685,831,1284,1308]" firstAuthor="Zimicz" pageId="206" pageNumber="206" refId="ref237684" refString="Zimicz, A. N. 2012. Ecomorfologia de los marsupiales paleogenos de America del Sur. Ph. D. dissertation, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata." type="book" year="2012">Zimicz, 2012</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D8A6FAAA195DFADB" author="Zimicz" box="[685,831,1284,1308]" firstAuthor="Zimicz" pageId="206" pageNumber="206" refId="ref237684" refString="Zimicz, A. N. 2012. Ecomorfologia de los marsupiales paleogenos de America del Sur. Ph. D. dissertation, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata." type="book" year="2012">Zimicz, 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Woodburne" box="[843,1117,1284,1308]" etAl="et al." firstAuthor="Woodburne" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="a">Woodburne et al., 2014a</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7D940FAAA1E3FFADB" author="Woodburne" box="[843,1117,1284,1308]" etAl="et al." firstAuthor="Woodburne" pageId="206" pageNumber="206" pagination="109 - 119" refId="ref235572" refString="Woodburne, M. O., et al. 2014 a. Revised timing of the South American early Paleogene land mammal ages. Journal of South American Earth Sciences 54: 109 - 119." type="journal article" year="2014" yearSuffix="a">Woodburne et al., 2014a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Goin, F. J. & M. O. Woodburne & A. N. Zimicz & G. M. Martin & L. Chornogubsky" pageId="206" pageNumber="206" refId="ref207099" refString="Goin, F. J., M. O. Woodburne, A. N. Zimicz, G. M. Martin, and L. Chornogubsky. 2016. A brief history of South American metatherians: Evolutionary contexts and intercontinental dispersals. Dordrecht: Springer." type="book" year="2016">Goin et al., 2016</bibRefCitation>
|
||||
<bibRefCitation id="EFD711BAF6F568E7DE61FAAA196DFAFA" author="Goin, F. J. & M. O. Woodburne & A. N. Zimicz & G. M. Martin & L. Chornogubsky" pageId="206" pageNumber="206" refId="ref207099" refString="Goin, F. J., M. O. Woodburne, A. N. Zimicz, G. M. Martin, and L. Chornogubsky. 2016. A brief history of South American metatherians: Evolutionary contexts and intercontinental dispersals. Dordrecht: Springer." type="book" year="2016">Goin et al., 2016</bibRefCitation>
|
||||
), which is incongruent with our estimate for the time of the Microbiotheria-Diprotodontia split, but these potentially important fossils have yet to be described.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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