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<mods:title>Drymaria veliziae (Caryophyllaceae), a new species from the Andes of Cajamarca (North Peru)</mods:title>
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<mods:title id="0A5DC0D8B41C97C2E5BE249584D822EC">Drymaria veliziae (Caryophyllaceae), a new species from the Andes of Cajamarca (North Peru)</mods:title>
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<mods:namePart>Montesinos-Tubee, Daniel B.</mods:namePart>
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<mods:namePart id="107787AC622B3402349E778854918438">Montesinos-Tubee, Daniel B.</mods:namePart>
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<mods:namePart>Tovar, Carolina</mods:namePart>
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<mods:namePart id="8EEE4F0718F819BB4EB4C20A58C6A79B">Tovar, Carolina</mods:namePart>
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<mods:namePart>Iberico-Vela, Gustavo</mods:namePart>
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<mods:namePart>Montoya-Quino, Juan</mods:namePart>
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<mods:namePart id="6B2FA83C4FE5AD2C01AADBCF3991DDB5">Montoya-Quino, Juan</mods:namePart>
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<mods:namePart>Sanchez-Vega, Isidoro</mods:namePart>
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<mods:namePart id="419FD1B2055907072A606F68892994FC">Sanchez-Vega, Isidoro</mods:namePart>
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<mods:title id="0A4C49EAFAE5E3423BCB0E7F57381E5C">PhytoKeys</mods:title>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:number>140</mods:number>
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<mods:date id="A3DD269C1ED37684F9A9A7BDCE198148">2020</mods:date>
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<mods:number id="21207D626251157854F5ADA30C700564">140</mods:number>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.140.47738</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-140-47</mods:identifier>
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<treatment ID-GBIF-Taxon="162267176" LSID="urn:lsid:plazi:treatment:CB4C39ADD7485373AAC1513FDA856B0D" httpUri="http://treatment.plazi.org/id/CB4C39ADD7485373AAC1513FDA856B0D" lastPageNumber="47" pageId="0" pageNumber="47">
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<taxonomicName LSID="CB4C39AD-D748-5373-AAC1-513FDA856B0D" authority="Montesinos" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae" status="sp. nov.">Drymaria veliziae Montesinos</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
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<treatment id="CB4C39ADD7485373AAC1513FDA856B0D" ID-GBIF-Taxon="162267176" LSID="urn:lsid:plazi:treatment:CB4C39ADD7485373AAC1513FDA856B0D" httpUri="http://treatment.plazi.org/id/CB4C39ADD7485373AAC1513FDA856B0D" lastPageNumber="47" pageId="0" pageNumber="47" scope_family="Caryophyllaceae">
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<subSubSection id="85DE24E2EFDD5037A2080800E6109CA6" pageId="0" pageNumber="47" type="nomenclature">
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<paragraph id="1398D550E5F50F729F335BA83AF0D000" pageId="0" pageNumber="47">
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<taxonomicName id="81E33D801C86C30A7742474DC616D6A9" ID-CoL="6DNC2" LSID="CB4C39AD-D748-5373-AAC1-513FDA856B0D" authority="Montesinos" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae" status="sp. nov.">Drymaria veliziae Montesinos</taxonomicName>
|
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<taxonomicNameLabel id="B6C458621C80C418F7F0CEE4EADD0185" pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
|
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</paragraph>
|
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</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="47" type="materials_examined">
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||||
<paragraph pageId="0" pageNumber="47">Type.</paragraph>
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||||
<paragraph pageId="0" pageNumber="47">
|
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<subSubSection id="CD33AC6DBC7422EDD5B2F7BE08B90FE8" pageId="0" pageNumber="47" type="materials_examined">
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||||
<paragraph id="C83FC298D44B7E82A8452901CF20EA47" pageId="0" pageNumber="47">Type.</paragraph>
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<paragraph id="88D205FFB30418FF45C6078E962F6799" pageId="0" pageNumber="47">
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Peru. Cajamarca: Cajamarca:
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<normalizedToken originalValue="Encañada">Encanada</normalizedToken>
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<normalizedToken id="1001C3C04EE29E6394659614B43AECD0" originalValue="Encañada">Encanada</normalizedToken>
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: Chanta baja, on sandy clay loam soils amongst shrub species and tussock grasslands, close to agricultural lands, 3295 m elev., slope of 60% and rock cover of 35%,
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6°49
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<normalizedToken id="A2BF228140C293583255AE063869B5C8" originalValue="’56’’">'56''</normalizedToken>
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S
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,
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78°30
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<normalizedToken id="3171CA78187D27D637B7D17EFCA14DFA" originalValue="’20’’">'20''</normalizedToken>
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W
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(DMS). 06 June 2009,
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<emphasis italics="true" pageId="0" pageNumber="47">C. Tovar 1058</emphasis>
|
||||
<emphasis id="7D0E96B4FEADBE07B78B70F6F773581D" italics="true" pageId="0" pageNumber="47">C. Tovar 1058</emphasis>
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(holotype CPUN-22705!).
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="47" type="diagnosis">
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<paragraph pageId="0" pageNumber="47">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="47">
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||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<subSubSection id="1A8EFBE48E6574A4F3D9C08CC31D4A9F" pageId="0" pageNumber="47" type="diagnosis">
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<paragraph id="C10E415807679C5AA8D09A372131AAC1" pageId="0" pageNumber="47">Diagnosis.</paragraph>
|
||||
<paragraph id="A05E12033A95D23DDBFAEE6E3362F64D" pageId="0" pageNumber="47">
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||||
<taxonomicName id="65DB77D0A9C86D5C069DEE133321674E" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="0A2EFF84D21FC68FCA24437859D69910" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
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||||
</taxonomicName>
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||||
is similar to
|
||||
<taxonomicName lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
<taxonomicName id="07D797DDC1C4819574219F66276BBACC" lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis id="8DFC46DD333D3D39EEBBE39CE6454FA1" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
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||||
</taxonomicName>
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Mattf. from which it differs in having glands covering the stems and pedicels, leaves with elliptic-ovate form, shorter in size (4-5.5 mm vs. 5-15 mm in
|
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<taxonomicName lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
<taxonomicName id="0C823E0B9BB6E4DEE9C78EAACD203EC4" lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis id="1FF31B9E8DE2C72D3AB73325C5C09F2D" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
</taxonomicName>
|
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), by the leaves arranged in fascicules (vs. simple opposite leaves), stipules in numbers of 14-20 per axis (vs. 2-4 in
|
||||
<taxonomicName lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
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||||
<taxonomicName id="30470DE907AB49CCF8794287779B7491" lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis id="567D345B1ABAB0256C1EB5371DEE13C9" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
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||||
</taxonomicName>
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||||
), pedicel size (1-2 mm long vs. 5-40 mm) and by the capsule size being smaller (1.4-1.6 mm vs. 3-4 mm).
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</paragraph>
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||||
<caption doi="10.3897/phytokeys.140.47738.figure1" httpUri="https://binary.pensoft.net/fig/382681" pageId="0" pageNumber="47" start="Figure 1" startId="F1">
|
||||
<paragraph pageId="0" pageNumber="47">
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||||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 1.</emphasis>
|
||||
<caption id="F7169EA65E0E1F5743620F590EB0D6C7" doi="10.3897/phytokeys.140.47738.figure1" httpUri="https://binary.pensoft.net/fig/382681" pageId="0" pageNumber="47" start="Figure 1" startId="F1">
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||||
<paragraph id="3CC6D92C39AAA5932315A3496828AD88" pageId="0" pageNumber="47">
|
||||
<emphasis id="EC7509CEED80ECBB9FEBB4DA35528C9E" bold="true" pageId="0" pageNumber="47">Figure 1.</emphasis>
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Location map of the type and paratype collection localities. Background: LANDSAT 5TM satellite image (June 2007) where light brown-pink areas represent Jalca grasslands, light green areas are agricultural fields and purple represents mining areas.
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</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="47" type="description">
|
||||
<paragraph pageId="0" pageNumber="47">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
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<subSubSection id="FBA798F6403B002A4B7F2890A54D76E9" pageId="0" pageNumber="47" type="description">
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<paragraph id="3D9FBE80493DAAC2605563131584B4BB" pageId="0" pageNumber="47">Description.</paragraph>
|
||||
<paragraph id="587F1E261A51274693A9996669FC33AE" pageId="0" pageNumber="47">
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||||
Perennial herb, the taproot woody, stems originating from the root brow spreading or ascending, rarely decumbent, of 20-35(-50) cm long. Stems rigid, greenish-lilac, densely glandular, of about 0.05-0.3 mm long, persistent on mature stems and having scattered plicate trichomes of 1-3 mm long on young stems. Internodes 0.2-5.5 cm long. Leaves opposite, usually forming short fascicules; petioles 0.3-0.9 mm long, partially glaucous, scarcely covered with glands in the margins; blades elliptic to ovate, 4-5.5 mm long
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|
||||
<normalizedToken id="0EBEAE78BFC0802389E8EB2A1CA85207" originalValue="×">x</normalizedToken>
|
||||
1.2-2 mm width, coriaceous, the bases cuneate, decurrent to the petiole, the apex aristate, 1-1.5 mm long, narrowly bearing short glands along the margin, midrib nearly inconspicuous; leaf margins lustrous, revolute and glabrous except at the base; stipules aciculate to linear-lanceolate, aristate, 1.5-3 mm long
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="62DC0806A251CA77E943BBFBBB32AAC0" originalValue="×">x</normalizedToken>
|
||||
0.1-0.4 wide, shorter or equalling the length of the leaves, with glabrous margins and usually verticillate, in numbers of 14-20 per axis, persistent, white translucid to brownish with age, rarely bifid or trifid; bracts opposite, 2.5-3 mm long
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="C3A7E97C88912B0091EFCAD499B11E79" originalValue="×">x</normalizedToken>
|
||||
2 mm width, involute, cupuliform, irregularly ovate, margins covered with scattered glands, surface white coloured with lilac blotches. Flowers except the first formed, axilar and solitary at the end of the branches, base protected by the bracts (in pairs 1 or 2). Pedicels 1-2 mm long, densely glandular and covered with carinate plicate trichomes, rarely aereal, of about 0.2-0.4 mm. Calyx cylindrical-campanulate; sepals 5, equal, 5-6 mm long
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="5939F764F6880DB971B59A0BE5322235" originalValue="×">x</normalizedToken>
|
||||
2-2.2 mm width, glabrous, elliptic-ovate, apex apiculate and aristate, basally truncate, 3-5 nerved; petals 5, 5-7 mm long
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="835E2F60CC5789BC92CFDC3A99D76E76" originalValue="×">x</normalizedToken>
|
||||
1.8-2.2 mm width, bifid about half their length, elliptic, apex rounded, 1-1.2 mm width, 8-10 nerved, constricted at the junction of the lobes; stamens 5, 2-2.2 mm long, anthers oblong, 0.2-0.3 mm long; ovary roundish, 1-1.4 mm long, slightly exceeded by the anthers; style 1-1.2 mm long, trifid about half its length, stigmatic branches twisted or coiled. Capsule ovoid, 1.4-1.6 mm long, 5-8 seeded. Seeds roundish, reniform, 0.6-0.9 mm long
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="656C5B35D84D93F5A0F25E15C5B9D1B7" originalValue="×">x</normalizedToken>
|
||||
0.6-0.8 mm wide, granulate, ventral surface with roundish-acute tubercles, black to dark brown in colour.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/phytokeys.140.47738.figure2" httpUri="https://binary.pensoft.net/fig/382682" pageId="0" pageNumber="47" start="Figure 2" startId="F2">
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 2.</emphasis>
|
||||
<caption id="D76F5C6B747A07B833E44B05782651A6" doi="10.3897/phytokeys.140.47738.figure2" httpUri="https://binary.pensoft.net/fig/382682" pageId="0" pageNumber="47" start="Figure 2" startId="F2">
|
||||
<paragraph id="6568818CA376418279B4DC787C40D6A0" pageId="0" pageNumber="47">
|
||||
<emphasis id="8351B0EE7B0CC5F7C8A7DD81FFDDD736" bold="true" pageId="0" pageNumber="47">Figure 2.</emphasis>
|
||||
Holotype of
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="1F9AA2183EA9ACC17EEEBA985D44237C" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="80CAE5131299E586512E580C192C70CB" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
(CPUN-22705!).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/phytokeys.140.47738.figure3" httpUri="https://binary.pensoft.net/fig/382683" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
|
||||
<caption id="C734FAA23C012D8376EE4F2E9AC50AD4" doi="10.3897/phytokeys.140.47738.figure3" httpUri="https://binary.pensoft.net/fig/382683" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
|
||||
<paragraph id="27147BECCC656894E03E99F71945375B" pageId="0" pageNumber="47">
|
||||
<emphasis id="72837A5CA2B7A8C62B8409CDB88787AC" bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
|
||||
A. Internode and leaf arrangements
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||||
<emphasis id="B5EA2635108B8439D9D166BB400FCF5D" bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||||
glandular stems
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||||
<emphasis id="F864911D6E37EADC64538A4FF6F409DC" bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||||
crown of stipules at the leaf axis
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||||
<emphasis id="DF5D1A6E995A28063BAB771D636ABA4D" bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||||
leaves (adaxial side)
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||||
<emphasis id="11E91971236773B5D19F4BFC5937058E" bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||||
leaves (abaxial side)
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||||
<emphasis id="7973336C447176E725D533EE84D70184" bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||||
axis of leaves
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||||
<emphasis id="97CB600ACE44CD1A67CAB6EBDDD34287" bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||||
bracts at the leaf axis
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||||
<emphasis id="EB91082A4F758A58DEB746B240A28255" bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||||
mature and immature flowers
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">I</emphasis>
|
||||
<emphasis id="A5755B6F2151976CE56957136F936A5F" bold="true" pageId="0" pageNumber="47">I</emphasis>
|
||||
flower detail
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">J</emphasis>
|
||||
<emphasis id="21CF7121CED48822CD370D7A618D416E" bold="true" pageId="0" pageNumber="47">J</emphasis>
|
||||
sepals (adaxial side)
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">K</emphasis>
|
||||
<emphasis id="B37909271E1869BDE89E978714B37ADA" bold="true" pageId="0" pageNumber="47">K</emphasis>
|
||||
sepals (abaxial side)
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">L</emphasis>
|
||||
<emphasis id="4DFBCA3D08DCE9FAD338850B122EEBE2" bold="true" pageId="0" pageNumber="47">L</emphasis>
|
||||
single petal with bifid apex portion
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">M</emphasis>
|
||||
<emphasis id="8663AAA81C1C21484A0115C16FB3B9DE" bold="true" pageId="0" pageNumber="47">M</emphasis>
|
||||
flower with ovary, stamens (indicated with dark lines) and style
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">N</emphasis>
|
||||
<emphasis id="3372C90709CC3D128C3447846281EA50" bold="true" pageId="0" pageNumber="47">N</emphasis>
|
||||
ovary detail with a trifurcated style
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">O</emphasis>
|
||||
<emphasis id="AEDB5B08B342414B4A2395EE66E9DC15" bold="true" pageId="0" pageNumber="47">O</emphasis>
|
||||
seeds.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="47" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="47">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<subSubSection id="5ACE801B95D21A849CE2C89196AED738" pageId="0" pageNumber="47" type="etymology">
|
||||
<paragraph id="9817B6744EAA444D5E9EE13F07BB4F2C" pageId="0" pageNumber="47">Etymology.</paragraph>
|
||||
<paragraph id="CA6768E5DDFF59A780F73815F404696B" pageId="0" pageNumber="47">
|
||||
The epithet "
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">veliziae</emphasis>
|
||||
<emphasis id="61B08C6F6CFFFFBFABC01736FA97D8BD" italics="true" pageId="0" pageNumber="47">veliziae</emphasis>
|
||||
" honours Claudia
|
||||
<normalizedToken originalValue="Véliz">Veliz</normalizedToken>
|
||||
<normalizedToken id="E98F5856CD2810390C8C3EC795BA8BAE" originalValue="Véliz">Veliz</normalizedToken>
|
||||
Rosas (1978-2019), a passionate biologist who devoted her research efforts to the study Peruvian biodiversity. Her deep love of nature, people and travelling inspired her to work throughout Peru, studying freshwater, marine and mountain ecosystems. Her research contributed to the establishment of protected areas and the development of management plans. Claudia dedicated many years to study taricaya turtles in the Amazon, helping local human communities to improve
|
||||
<normalizedToken originalValue="taricayas’">taricayas'</normalizedToken>
|
||||
<normalizedToken id="D0D4D6BFCA0C87C9C21B9C8B268B74F6" originalValue="taricayas’">taricayas'</normalizedToken>
|
||||
management and conservation. She was an excellent and supportive friend, a talented amateur painter and dancer and a keen cyclist.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">Paratype</emphasis>
|
||||
<paragraph id="20A58C969D35BD4583F4B467012723A8" pageId="0" pageNumber="47">
|
||||
<emphasis id="01B6D5C8D33D7B0877C53B061C9CFAD3" bold="true" pageId="0" pageNumber="47">Paratype</emphasis>
|
||||
: Peru: Cajamarca: Hualgayoc: Hualgayoc, less than 1 km from the Goldfield mine, surrounded by agricultural fields downslope, found on sandy clay loam soils, 3715 m elev., 6°46
|
||||
<normalizedToken originalValue="’43’’">'43''</normalizedToken>
|
||||
<normalizedToken id="B108FAD6D0D1B50B411A60E4D383C6E2" originalValue="’43’’">'43''</normalizedToken>
|
||||
S (DMS) and 78°37
|
||||
<normalizedToken originalValue="’5’’">'5''</normalizedToken>
|
||||
<normalizedToken id="51D7C884718A211A510C680517F87821" originalValue="’5’’">'5''</normalizedToken>
|
||||
W (DMS), 100% slope and 5% rock cover. 01 June 2009,
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">C. Tovar 909</emphasis>
|
||||
<emphasis id="2D2B256B82039CB8D210B5076E735A52" italics="true" pageId="0" pageNumber="47">C. Tovar 909</emphasis>
|
||||
(CPUN-22858!).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">Ecology and distribution</emphasis>
|
||||
<paragraph id="6C2A676536E4DAEF24A705E913F4CF1F" pageId="0" pageNumber="47">
|
||||
<emphasis id="65D4512E0A7249E6CE2ED6DC1B906054" bold="true" pageId="0" pageNumber="47">Ecology and distribution</emphasis>
|
||||
:
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="D484AE0136CF352719EF72F9EF41EC24" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="BCC11AC424A6426F7D834781F3E649C5" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
grows on steep mountain cliffs (slope 60-100%) on sandy clay loam soils at an elevation of 3295-3715 m on the eastern slopes of the Jalca, on the headwaters of the Llaucano River, tributary of the
|
||||
<normalizedToken originalValue="Marañon">Maranon</normalizedToken>
|
||||
<normalizedToken id="330824DA1E0DFA9F919553E3896C8E0C" originalValue="Marañon">Maranon</normalizedToken>
|
||||
River. Climatic characteristics for the localities of the type and paratype, extracted from the CHELSA climatology (
|
||||
<bibRefCitation DOI="https://doi.org/10.1038/sdata.2017.122" author="Karger, DN" journalOrPublisher="Scientific Data" pageId="0" pageNumber="47" pagination=": 170122" refId="B8" refString="Karger, DN, Conrad, O, Bohner, J, Kawohl, T, Kreft, H, Soria-Auza, RW, Zimmermann, N, Linder, HP, Kessler, M, 2017. Climatologies at high resolution for the earth's land surface areas. Scientific Data 4 (1): 170122, DOI: https://doi.org/10.1038/sdata.2017.122" title="Climatologies at high resolution for the earth's land surface areas." url="https://doi.org/10.1038/sdata.2017.122" volume="4" year="2017">Karger et al. 2017</bibRefCitation>
|
||||
<bibRefCitation id="78B097457C6AEDC52EAC89BB0A8D755C" DOI="https://doi.org/10.1038/sdata.2017.122" author="Karger, DN" journalOrPublisher="Scientific Data" pageId="0" pageNumber="47" pagination=": 170122" refId="B8" refString="Karger, DN, Conrad, O, Bohner, J, Kawohl, T, Kreft, H, Soria-Auza, RW, Zimmermann, N, Linder, HP, Kessler, M, 2017. Climatologies at high resolution for the earth's land surface areas. Scientific Data 4 (1): 170122, DOI: https://doi.org/10.1038/sdata.2017.122" title="Climatologies at high resolution for the earth's land surface areas." url="https://doi.org/10.1038/sdata.2017.122" volume="4" year="2017">Karger et al. 2017</bibRefCitation>
|
||||
), show mean annual temperatures in these areas are 8.5-11.5°C, with minimum temperatures estimated between 1.8 and 5°C. Total annual precipitation ranges from 900 to 1200 mm with driest months receiving 16-28 mm. Other species found in the two localities were
|
||||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Hieracium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hieracium peruanum" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="peruanum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Hieracium peruanum</emphasis>
|
||||
<taxonomicName id="FE933BDAC22415603977353F8DC7EB38" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hieracium peruanum" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="peruanum">
|
||||
<emphasis id="685C119B81257003175DC5437F010FD8" italics="true" pageId="0" pageNumber="47">Hieracium peruanum</emphasis>
|
||||
</taxonomicName>
|
||||
Fr. (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="319E14885FF51FAA22F83EE40470EF76" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Hypochaeris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hypochaeris taraxacoides" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="taraxacoides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Hypochaeris taraxacoides</emphasis>
|
||||
<taxonomicName id="E7816F7C1EA31311D6AEC1FCC278E50B" class="Magnoliopsida" family="Asteraceae" genus="Hypochaeris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hypochaeris taraxacoides" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="taraxacoides">
|
||||
<emphasis id="494FE4C39BA79827CC82B7FC8782181D" italics="true" pageId="0" pageNumber="47">Hypochaeris taraxacoides</emphasis>
|
||||
</taxonomicName>
|
||||
(Meyen & Walp.) Ball (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="E15A92D88C2FDE040D4B450EA1028BC4" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
) and
|
||||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Calamagrostis</emphasis>
|
||||
<taxonomicName id="16ED65F3C96E559B0BD47F127E36BBFB" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="8B1A4E5C6B493B23EC2B1F2C2F5D728D" italics="true" pageId="0" pageNumber="47">Calamagrostis</emphasis>
|
||||
</taxonomicName>
|
||||
spp. (
|
||||
<taxonomicName family="Poaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Poaceae</taxonomicName>
|
||||
<taxonomicName id="B4FA24F1C98EDD7DC3393C51F4E7C45A" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||||
). In the type locality of
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="53E8C8DE780B4F80D27C87EB172B95BB" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="3C4B7286D6237F68FDDB6E82519534A4" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Quadrat where the new species was found. Chanta baja, Encanada District, 3295 m elev. Photo by Carolina Tovar." figureDoi="10.3897/phytokeys.140.47738.figure4" httpUri="https://binary.pensoft.net/fig/382684" pageId="0" pageNumber="47">4</figureCitation>
|
||||
<figureCitation id="D94D9E28675E4CAC36F0F8AD50E57998" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Quadrat where the new species was found. Chanta baja, Encanada District, 3295 m elev. Photo by Carolina Tovar." figureDoi="10.3897/phytokeys.140.47738.figure4" httpUri="https://binary.pensoft.net/fig/382684" pageId="0" pageNumber="47">4</figureCitation>
|
||||
), it has been observed that it grows associated with shrubs (e.g.
|
||||
<taxonomicName authorityName="S. F. Blake & Sherff" authorityYear="1925" class="Magnoliopsida" family="Asteraceae" genus="Coreopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coreopsis senaria" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="senaria">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Coreopsis senaria</emphasis>
|
||||
<taxonomicName id="55CDC6900AB304E1CA6DCFEF033B3CE8" authorityName="S. F. Blake & Sherff" authorityYear="1925" class="Magnoliopsida" family="Asteraceae" genus="Coreopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coreopsis senaria" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="senaria">
|
||||
<emphasis id="89F229752AFC200065E5EB656A69F8FE" italics="true" pageId="0" pageNumber="47">Coreopsis senaria</emphasis>
|
||||
</taxonomicName>
|
||||
S.F. Blake & Sherff (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="5CDDD7666A26C2315812D9A0E9E0F3B0" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Achyrocline" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achyrocline celosioides" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="celosioides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Achyrocline celosioides</emphasis>
|
||||
<taxonomicName id="25D571316D8DDB7E9F8A60B30558BAFD" class="Magnoliopsida" family="Asteraceae" genus="Achyrocline" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achyrocline celosioides" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="celosioides">
|
||||
<emphasis id="B3D11602D73FD6D9D51D4C87523C04BF" italics="true" pageId="0" pageNumber="47">Achyrocline celosioides</emphasis>
|
||||
</taxonomicName>
|
||||
(Kunth) DC. (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="4ADF03C8F987EFF852C3E5AB0618EDD0" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Ageratina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ageratina cutervensis" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="cutervensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Ageratina cutervensis</emphasis>
|
||||
<taxonomicName id="FD1EC56ED9D40415CB305B3E50B6ADB9" class="Magnoliopsida" family="Asteraceae" genus="Ageratina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ageratina cutervensis" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="cutervensis">
|
||||
<emphasis id="90E072A89563B076D42E5095D795A7F1" italics="true" pageId="0" pageNumber="47">Ageratina cutervensis</emphasis>
|
||||
</taxonomicName>
|
||||
(Hieron.) R.M. King & H. Rob. (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="12BB2B2F7D31BFC985849E7040BE4181" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
)), tussock grasses (
|
||||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Calamagrostis</emphasis>
|
||||
<taxonomicName id="4E3D8999C9BA7048F605FF9A0331981E" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="494404E3C7124C7696CA433B464FAB98" italics="true" pageId="0" pageNumber="47">Calamagrostis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Festuca</emphasis>
|
||||
<taxonomicName id="A37BEDC840659BADA1C726BF5A296FB6" class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="14C95CFE9DA08636EE35B947C98B6790" italics="true" pageId="0" pageNumber="47">Festuca</emphasis>
|
||||
</taxonomicName>
|
||||
spp. (
|
||||
<taxonomicName family="Poaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Poaceae</taxonomicName>
|
||||
<taxonomicName id="47B55E0DBFEB328BE902CDF09FBE5D2D" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||||
)) and an orchid (e.g.
|
||||
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Masdevallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Masdevallia" order="Asparagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Masdevallia</emphasis>
|
||||
<taxonomicName id="3550B387A5D46D593206100D9F184AC5" class="Liliopsida" family="Orchidaceae" genus="Masdevallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Masdevallia" order="Asparagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="05544109531CE0F63D9E03A02ECEA246" italics="true" pageId="0" pageNumber="47">Masdevallia</emphasis>
|
||||
</taxonomicName>
|
||||
spp.), amongst others. The locality of the paratype, being at higher altitude, had less species richness and associated species with
|
||||
<taxonomicName lsidName="D. velizzii" pageId="0" pageNumber="47" rank="species" species="velizzii">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. velizzii</emphasis>
|
||||
<taxonomicName id="D012155188D626BB6007DE881DFEBC69" lsidName="D. velizzii" pageId="0" pageNumber="47" rank="species" species="velizzii">
|
||||
<emphasis id="35F955285AACA67F29DB3118B0C365BF" italics="true" pageId="0" pageNumber="47">D. velizzii</emphasis>
|
||||
</taxonomicName>
|
||||
were
|
||||
<taxonomicName authorityName="Hieron" authorityYear="1895" class="Magnoliopsida" family="Geraniaceae" genus="Geranium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Geranium peruvianum" order="Geraniales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="peruvianum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Geranium peruvianum</emphasis>
|
||||
<taxonomicName id="E50B5D4D17C67C6D16BFA3161625F1AD" authorityName="Hieron" authorityYear="1895" class="Magnoliopsida" family="Geraniaceae" genus="Geranium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Geranium peruvianum" order="Geraniales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="peruvianum">
|
||||
<emphasis id="D45D157DD83E46D9A1599AE3A27EDC5B" italics="true" pageId="0" pageNumber="47">Geranium peruvianum</emphasis>
|
||||
</taxonomicName>
|
||||
Hieron. (
|
||||
<taxonomicName family="Geraniaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Geraniaceae</taxonomicName>
|
||||
<taxonomicName id="9C0ED284E3F6C409742BC9478A01F72C" class="Magnoliopsida" family="Geraniaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Geraniales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Geraniaceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Calceolariaceae" genus="Calceolaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calceolaria concava" order="Lamiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="concava">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Calceolaria concava</emphasis>
|
||||
<taxonomicName id="78092F62243915732B9623F18D03EA7B" class="Magnoliopsida" family="Calceolariaceae" genus="Calceolaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calceolaria concava" order="Lamiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="concava">
|
||||
<emphasis id="E375CF0C8C6BBC4D647666D8509A1E29" italics="true" pageId="0" pageNumber="47">Calceolaria concava</emphasis>
|
||||
</taxonomicName>
|
||||
Molau (
|
||||
<taxonomicName family="Calceolariaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Calceolariaceae</taxonomicName>
|
||||
<taxonomicName id="901BBFB637B6AC1FE6B64459087C642D" class="Magnoliopsida" family="Calceolariaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Calceolariaceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Chrysactinium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chrysactinium acaule" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="acaule">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Chrysactinium acaule</emphasis>
|
||||
<taxonomicName id="9787920086493DD26A8067F7B38741C3" class="Magnoliopsida" family="Asteraceae" genus="Chrysactinium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chrysactinium acaule" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="acaule">
|
||||
<emphasis id="6A2A4676A7418E8EDDC6F172949CFB3F" italics="true" pageId="0" pageNumber="47">Chrysactinium acaule</emphasis>
|
||||
</taxonomicName>
|
||||
(Kunth) Wedd. (
|
||||
<taxonomicName family="Asteraceae" lsidName="" pageId="0" pageNumber="47" rank="family">Asteraceae</taxonomicName>
|
||||
<taxonomicName id="000F8DF1BAF0EDD8B54EEBDB5524F14D" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euphorbia huanchahana" order="Malpighiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="huanchahana">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Euphorbia huanchahana</emphasis>
|
||||
<taxonomicName id="72667131C99A068855132D1D6481FF2B" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euphorbia huanchahana" order="Malpighiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="huanchahana">
|
||||
<emphasis id="F6CBD563AD59F37D8000867BC4BACA28" italics="true" pageId="0" pageNumber="47">Euphorbia huanchahana</emphasis>
|
||||
</taxonomicName>
|
||||
(Klotzsch & Garcke) Boiss. (
|
||||
<taxonomicName family="Euphorbiaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Euphorbiaceae</taxonomicName>
|
||||
<taxonomicName id="86C85548F2A8F84EA3941F845097629A" class="Magnoliopsida" family="Euphorbiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malpighiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
|
||||
), amongst others.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/phytokeys.140.47738.figure4" httpUri="https://binary.pensoft.net/fig/382684" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
|
||||
<caption id="411C21A451AE77B583E967C48787C3E0" doi="10.3897/phytokeys.140.47738.figure4" httpUri="https://binary.pensoft.net/fig/382684" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
|
||||
<paragraph id="2449615A8F9EC19A58DFAB382A773526" pageId="0" pageNumber="47">
|
||||
<emphasis id="E7E2B56FCB170693CA493A9C1E2D6140" bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
|
||||
Quadrat where the new species was found. Chanta baja,
|
||||
<normalizedToken originalValue="Encañada">Encanada</normalizedToken>
|
||||
<normalizedToken id="BB95249FA4B45539935AB917480CC29B" originalValue="Encañada">Encanada</normalizedToken>
|
||||
District, 3295 m elev. Photo by Carolina Tovar.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="47" type="taxonomical notes">
|
||||
<paragraph pageId="0" pageNumber="47">Taxonomical notes.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<subSubSection id="736BAD0777BCAD846C7F5052547E489F" pageId="0" pageNumber="47" type="taxonomical notes">
|
||||
<paragraph id="253E650A5832061A61BE22F91D644D24" pageId="0" pageNumber="47">Taxonomical notes.</paragraph>
|
||||
<paragraph id="4547AF571A7DE613617C9B9410EB933D" pageId="0" pageNumber="47">
|
||||
On the basis of the classification proposed by
|
||||
<bibRefCitation DOI="https://doi.org/10.2307/2394953" author="Duke, JA" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="47" pagination="173 - 268" refId="B2" refString="Duke, JA, 1961. Preliminary revision of the genus Drymaria. Annals of the Missouri Botanical Garden 48 (3): 173 - 268, DOI: https://doi.org/10.2307/2394953" title="Preliminary revision of the genus Drymaria." url="https://doi.org/10.2307/2394953" volume="48" year="1961">Duke (1961)</bibRefCitation>
|
||||
<bibRefCitation id="144CA65BA957144C1F35B9F8FD7EE379" DOI="https://doi.org/10.2307/2394953" author="Duke, JA" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="47" pagination="173 - 268" refId="B2" refString="Duke, JA, 1961. Preliminary revision of the genus Drymaria. Annals of the Missouri Botanical Garden 48 (3): 173 - 268, DOI: https://doi.org/10.2307/2394953" title="Preliminary revision of the genus Drymaria." url="https://doi.org/10.2307/2394953" volume="48" year="1961">Duke (1961)</bibRefCitation>
|
||||
,
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="BB1D54FE90BCAC518FA97BBB9AFB1E9F" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="C14C81EEE096BB0B901CB951B1648552" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
would belong to the ser.
|
||||
<taxonomicName authorityName="Mattfeld" authorityYear="1936" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="series" series="Frutescens">Frutescens</taxonomicName>
|
||||
<taxonomicName id="F8CE902A3932E69E2752CC068F4D5548" authorityName="Mattfeld" authorityYear="1936" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="series" series="Frutescens">Frutescens</taxonomicName>
|
||||
Duke sharing the leaf shape (linear to lanceolate) glandular pedicels, the number of sepal nervadures (3-5) and petals bifid which are not tapered to the claw.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<paragraph id="7714016E3C71DE74E9B602B37C96A06D" pageId="0" pageNumber="47">
|
||||
<taxonomicName id="860C2488A17E9152B00A81CDD7E82E3B" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="7B50DD53831C00A8BAF3504F66D1CF75" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
is morphologically similar to
|
||||
<taxonomicName lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
<taxonomicName id="F3015FD7CAB8BA08CA1615C0E0DC8066" lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis id="2614F26B4911CCFE42067843EC22EED3" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
</taxonomicName>
|
||||
Mattf. (1936: 438-439) but differs in having glands covering the stems and pedicels, leaves with elliptic-ovate form, shorter in size, by the leaves arranged in fascicules (vs. simple opposite leaves), stipules larger numbers per axis, shorter pedicel size and smaller capsule size.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<paragraph id="7DAE7180300A1DF6D8486FD59165FEFE" pageId="0" pageNumber="47">
|
||||
Furthermore,
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="FE6DBB3B45C14009F5209F6AFFBAE8F9" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="7BE83D3D5453C9117726DE306D4C5A4B" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
<taxonomicName id="CEF3579AA62CC01F7F2A39C7F8E6D72A" lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis id="D88E2988E5E6D4CFDFC5C70D3C661575" italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
</taxonomicName>
|
||||
Mattf. (1936: 436-437) by the plant habit, the glabrous surface of the leaves (vs. presence of glands and puberulent trichomes in
|
||||
<taxonomicName lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
<taxonomicName id="6109CC7B539C316486305960979E897D" lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis id="BAB1AB7F8613DF5A1D55ED61A4EC8C37" italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
</taxonomicName>
|
||||
), bifid or trifid stipules (vs. entire), shorter stamen size (2-2.2 mm vs. 4-6 mm), shorter style size (1-1.2 mm vs. 1-2.5 mm), capsule size shorter (1.4-1.6 vs. 2.5-3.5 mm) and seed size (0.6-0.9 vs. 0.9-1.3 mm in
|
||||
<taxonomicName lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
<taxonomicName id="3AB50767969B2DC383B178283B94FBBF" lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis id="81579D38AE718D1C8906DBBDBDB95872" italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<paragraph id="A12D76639C77D47A0FCFE7AC93549B52" pageId="0" pageNumber="47">
|
||||
The new species is further differentiated from
|
||||
<taxonomicName lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
<taxonomicName id="E8FC84F4C71AC2A0E46E59F7066ED3FE" lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis id="A8FEFFC4EC5062EB5DBD7E90E488EF67" italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
</taxonomicName>
|
||||
Willd. ex Schult. (1819: 406) by the stipule form (bifid to trifid vs. entire), shorter bract size (2.5-3 mm vs. 3-5 mm in
|
||||
<taxonomicName lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
<taxonomicName id="CF547ACDA876A4568C87ED5556E316DB" lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis id="EE89EA6027B36D37A52E5E3BFC195292" italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
</taxonomicName>
|
||||
), sepals glabrous (vs. glabrous to densely glandular-puberulent) and shorter capsule size and form (1.4-1.6 mm, ovoid vs. 3-5 mm long, ellipsoid).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<paragraph id="C72F9E0D6FAFCB8E78DFEDAECC875009" pageId="0" pageNumber="47">
|
||||
An updating of the diagnostic key for the ser.
|
||||
<taxonomicName lsidName="" pageId="0" pageNumber="47" rank="series" series="Frutescens">Frutescens</taxonomicName>
|
||||
<taxonomicName id="6ABFED89FB05A928886A1AACCC52DBE5" lsidName="" pageId="0" pageNumber="47" rank="series" series="Frutescens">Frutescens</taxonomicName>
|
||||
, as proposed by
|
||||
<bibRefCitation DOI="https://doi.org/10.2307/2394953" author="Duke, JA" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="47" pagination="173 - 268" refId="B2" refString="Duke, JA, 1961. Preliminary revision of the genus Drymaria. Annals of the Missouri Botanical Garden 48 (3): 173 - 268, DOI: https://doi.org/10.2307/2394953" title="Preliminary revision of the genus Drymaria." url="https://doi.org/10.2307/2394953" volume="48" year="1961">Duke (1961</bibRefCitation>
|
||||
<bibRefCitation id="466B4C83387155E8A0B63F5C3334C418" DOI="https://doi.org/10.2307/2394953" author="Duke, JA" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="47" pagination="173 - 268" refId="B2" refString="Duke, JA, 1961. Preliminary revision of the genus Drymaria. Annals of the Missouri Botanical Garden 48 (3): 173 - 268, DOI: https://doi.org/10.2307/2394953" title="Preliminary revision of the genus Drymaria." url="https://doi.org/10.2307/2394953" volume="48" year="1961">Duke (1961</bibRefCitation>
|
||||
: 214) follows:
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<table pageId="0" pageNumber="47">
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">1</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals present and glabrous, sepals (3-)5-nerved</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">2</emphasis>
|
||||
<paragraph id="C09AA92A9A516C468DEA850324878840" pageId="0" pageNumber="47">
|
||||
<table id="D598AE391EB0636996D1BD433756C7DF" pageId="0" pageNumber="47">
|
||||
<tr id="E6BDA1FB59388616F473F329D41506F4" pageId="0" pageNumber="47">
|
||||
<td id="E7DF983864F05304DCDFDB9AA802F35A" colspan="1" pageId="0" pageNumber="47" rowspan="1">1</td>
|
||||
<td id="5D54D45CDE3B1F1B69FF404057FE9129" colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals present and glabrous, sepals (3-)5-nerved</td>
|
||||
<td id="D916EADCA464B2E4579CF3831A4F4E0F" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis id="2E7940D7AB16067A86152181B4FF1970" bold="true" pageId="0" pageNumber="47">2</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">edicels and sepals present or absent, glabrous or glandular, sepals 3-nerved</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">3</emphasis>
|
||||
<tr id="4C16ED6F0955D7A07403A6CD4DE9EC2B" pageId="0" pageNumber="47">
|
||||
<td id="FE5B55B4BFB55B15C744A1928AD8A8F8" colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td id="6F91FFE607496F67EDED75273FCDBB9B" colspan="1" pageId="0" pageNumber="47" rowspan="1">edicels and sepals present or absent, glabrous or glandular, sepals 3-nerved</td>
|
||||
<td id="3B802337FDF09990B4CBC427E0B2A378" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis id="D5F5494DAFE9FE05EE8179A08AF7E438" bold="true" pageId="0" pageNumber="47">3</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">2</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Leaves imbricate, closely appressed to the stems, 2-6 mm long, 1-1.5 mm broad, basally clasping and pungently acute; the sepals mostly 5-nerved; petals tapered to the claw</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName lsidName="D. frutescens" pageId="0" pageNumber="47" rank="species" species="frutescens">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">D. frutescens</emphasis>
|
||||
<tr id="D4DFEF13D1D87B20EB6FD5ECC2F9AD9B" pageId="0" pageNumber="47">
|
||||
<td id="67C972C2F9A903FE8FE8983AF02CE9A0" colspan="1" pageId="0" pageNumber="47" rowspan="1">2</td>
|
||||
<td id="E7CEC519C7F55D384B63C9F53A5B4692" colspan="1" pageId="0" pageNumber="47" rowspan="1">Leaves imbricate, closely appressed to the stems, 2-6 mm long, 1-1.5 mm broad, basally clasping and pungently acute; the sepals mostly 5-nerved; petals tapered to the claw</td>
|
||||
<td id="F46A44273EDF671410AC6574A98040BB" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName id="D9D2F6A55E82425AFC457C521B1EADA3" lsidName="D. frutescens" pageId="0" pageNumber="47" rank="species" species="frutescens">
|
||||
<emphasis id="02BDDE34767E331CB44A0DAF510825C2" bold="true" italics="true" pageId="0" pageNumber="47">D. frutescens</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Leaves not imbricate, 4-12 mm long, 2-6 mm broad, apically acute to aristately acuminate; the sepals 3-4-nerved; petals not tapered to the claw</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
<tr id="1F3D14ABE3144494DFE6E482839CF240" pageId="0" pageNumber="47">
|
||||
<td id="C9C93F86EC0CFD8B6269859E7C9FA5B0" colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td id="28853FCCC99769D6D76F509F2A05CBA1" colspan="1" pageId="0" pageNumber="47" rowspan="1">Leaves not imbricate, 4-12 mm long, 2-6 mm broad, apically acute to aristately acuminate; the sepals 3-4-nerved; petals not tapered to the claw</td>
|
||||
<td id="9E474AFF47B25CA92C093819CAA577AA" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName id="728A9CA0A51E2C0C56711ECAD8B78A10" lsidName="D. stereophylla" pageId="0" pageNumber="47" rank="species" species="stereophylla">
|
||||
<emphasis id="D9BA7E569029C074754E9D025622902D" bold="true" italics="true" pageId="0" pageNumber="47">D. stereophylla</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals usually glandular, leaves glabrous, apically aristate-attenuate, aristate and basally cuneate; stipules present; seeds with domical or conical tubercles</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="47">4</emphasis>
|
||||
<tr id="F0A6165BADF18D79EE1DB925A534FAFC" pageId="0" pageNumber="47">
|
||||
<td id="E617DAA551E592BA6D6A2C01ECE6A611" colspan="1" pageId="0" pageNumber="47" rowspan="1">3</td>
|
||||
<td id="3A2E9FED0EDF269353A4E8A6CACCBE2C" colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals usually glandular, leaves glabrous, apically aristate-attenuate, aristate and basally cuneate; stipules present; seeds with domical or conical tubercles</td>
|
||||
<td id="D5ED7329EE66C870A4B4F4BDA0B22C9C" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<emphasis id="4734B5055BF3B15E23482512B6EFF9B5" bold="true" pageId="0" pageNumber="47">4</emphasis>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals glabrous to densely glandular-puberulent; leaves apically acute and marginally entire, densely glandular-puberulent; stipules entire, apparently fused or occasionally absent; seeds without domical tubercles</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
<tr id="CD395BF9457B57CDE4CDCEE328B2D584" pageId="0" pageNumber="47">
|
||||
<td id="4DF526628757ACD5A2A8C89BDE0142CC" colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td id="BC647072DFF80820B42698E1EC6D499A" colspan="1" pageId="0" pageNumber="47" rowspan="1">Pedicels and sepals glabrous to densely glandular-puberulent; leaves apically acute and marginally entire, densely glandular-puberulent; stipules entire, apparently fused or occasionally absent; seeds without domical tubercles</td>
|
||||
<td id="D8BA4A140B1679E2D1E1DF380ECE3CE4" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName id="1ADFE1BDB10E5135BC13D91D788C9F26" lsidName="D. stellarioides" pageId="0" pageNumber="47" rank="species" species="stellarioides">
|
||||
<emphasis id="7CD218FF2F35047760907598E38D4D07" bold="true" italics="true" pageId="0" pageNumber="47">D. stellarioides</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Bracts absent, simple opposite leaves, pedicels 5-40 mm long, stipules copiously shorter than the leaves; glabrous pedicels and stems, short number of pedicels per axis (2-4)</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
<tr id="AB377AABBB46A514F12053C6F2314B66" pageId="0" pageNumber="47">
|
||||
<td id="874E763494C03A5A5ECFB8A2E4CA3052" colspan="1" pageId="0" pageNumber="47" rowspan="1">4</td>
|
||||
<td id="39B04FAFD509FABF00C7C6CA46B69C57" colspan="1" pageId="0" pageNumber="47" rowspan="1">Bracts absent, simple opposite leaves, pedicels 5-40 mm long, stipules copiously shorter than the leaves; glabrous pedicels and stems, short number of pedicels per axis (2-4)</td>
|
||||
<td id="91A9EB703D389DE81EE82BF0BC967184" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName id="A3456FC284DE46DC72557D9A80FEB765" lsidName="D. auriculipetala" pageId="0" pageNumber="47" rank="species" species="auriculipetala">
|
||||
<emphasis id="3FA951DD4F7D64A5756B67A22E8776AC" bold="true" italics="true" pageId="0" pageNumber="47">D. auriculipetala</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="0" pageNumber="47">
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Bracts present; leaves arranged in fascicules; pedicels 1-2 mm long, glands covering the stems and pedicels; leaves with elliptic-ovate form, large number of pedicels per axis (14-20)</td>
|
||||
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName lsidName="D. veliziae" pageId="0" pageNumber="47" rank="species" species="veliziae">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">D. veliziae</emphasis>
|
||||
<tr id="EBEA123E6AC97317FC739BC453367496" pageId="0" pageNumber="47">
|
||||
<td id="21CF2AEA5EB257650CCB11514DFFCD00" colspan="1" pageId="0" pageNumber="47" rowspan="1">-</td>
|
||||
<td id="620F5115F11F86482DD2D4610D1808D1" colspan="1" pageId="0" pageNumber="47" rowspan="1">Bracts present; leaves arranged in fascicules; pedicels 1-2 mm long, glands covering the stems and pedicels; leaves with elliptic-ovate form, large number of pedicels per axis (14-20)</td>
|
||||
<td id="6532F59ED9E7010C06105CA06F730027" colspan="1" pageId="0" pageNumber="47" rowspan="1">
|
||||
<taxonomicName id="EC3D51ACD4CD4893FF55CCE4C89832C7" lsidName="D. veliziae" pageId="0" pageNumber="47" rank="species" species="veliziae">
|
||||
<emphasis id="FB67C74842BE1FC322A70EC9125FF116" bold="true" italics="true" pageId="0" pageNumber="47">D. veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="47" type="conservation status">
|
||||
<paragraph pageId="0" pageNumber="47">Conservation status.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="47">
|
||||
<subSubSection id="5137C88269976AC1F619F1081ACE8D89" pageId="0" pageNumber="47" type="conservation status">
|
||||
<paragraph id="473BB9D6AEE1F975A140C14FB9DE9CBC" pageId="0" pageNumber="47">Conservation status.</paragraph>
|
||||
<paragraph id="197FA37EB6BA76CD7202815F54169DB4" pageId="0" pageNumber="47">
|
||||
Only the two localities referring to holotype and paratype are currently known for
|
||||
<taxonomicName authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
<taxonomicName id="71F75388384ACA37E71ACC6005B51CD0" authorityName="Montesinos" authorityYear="2020" class="Magnoliopsida" family="Caryophyllaceae" genus="Drymaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Drymaria veliziae" order="Caryophyllales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="veliziae">
|
||||
<emphasis id="FCBAE0CCA8A33CB9F2EAA7BBCDAA5BDB" italics="true" pageId="0" pageNumber="47">Drymaria veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
(these localities are separated by about 12 km). The surrounding areas are characterised by various types of human activities, for example, agriculture, land conversion, forestry with exotic species, slash burning, natural resource extraction, amongst others (Figure
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Location map of the type and paratype collection localities. Background: LANDSAT 5 TM satellite image (June 2007) where light brown-pink areas represent Jalca grasslands, light green areas are agricultural fields and purple represents mining areas." figureDoi="10.3897/phytokeys.140.47738.figure1" httpUri="https://binary.pensoft.net/fig/382681" pageId="0" pageNumber="47">1</figureCitation>
|
||||
<figureCitation id="84420D9FB673F02C99346C0E98E43712" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Location map of the type and paratype collection localities. Background: LANDSAT 5 TM satellite image (June 2007) where light brown-pink areas represent Jalca grasslands, light green areas are agricultural fields and purple represents mining areas." figureDoi="10.3897/phytokeys.140.47738.figure1" httpUri="https://binary.pensoft.net/fig/382681" pageId="0" pageNumber="47">1</figureCitation>
|
||||
). Land use change occurred between 1987 and 2007 with a reduction of the 25% of grasslands and an increasing of landscape fragmentation (see
|
||||
<bibRefCitation DOI="https://doi.org/10.1016/j.landurbplan.2012.12.003" author="Tovar, C" journalOrPublisher="Landscape and Urban Planning" pageId="0" pageNumber="47" pagination="40 - 49" refId="B17" refString="Tovar, C, Seijmonsbergen, AC, Duivenvoorden, JF, 2013. Monitoring land use and land cover change in mountain regions: An example in the Jalca grasslands of the Peruvian Andes. Landscape and Urban Planning 112: 40 - 49, DOI: https://doi.org/10.1016/j.landurbplan.2012.12.003" title="Monitoring land use and land cover change in mountain regions: An example in the Jalca grasslands of the Peruvian Andes." url="https://doi.org/10.1016/j.landurbplan.2012.12.003" volume="112" year="2013">Tovar et al. 2013</bibRefCitation>
|
||||
<bibRefCitation id="D52198755316679D3A48272F59986FDB" DOI="https://doi.org/10.1016/j.landurbplan.2012.12.003" author="Tovar, C" journalOrPublisher="Landscape and Urban Planning" pageId="0" pageNumber="47" pagination="40 - 49" refId="B17" refString="Tovar, C, Seijmonsbergen, AC, Duivenvoorden, JF, 2013. Monitoring land use and land cover change in mountain regions: An example in the Jalca grasslands of the Peruvian Andes. Landscape and Urban Planning 112: 40 - 49, DOI: https://doi.org/10.1016/j.landurbplan.2012.12.003" title="Monitoring land use and land cover change in mountain regions: An example in the Jalca grasslands of the Peruvian Andes." url="https://doi.org/10.1016/j.landurbplan.2012.12.003" volume="112" year="2013">Tovar et al. 2013</bibRefCitation>
|
||||
). The type specimen was collected on a Jalca patch surrounded by agricultural fields (
|
||||
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vicia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vicia faba" order="Fabales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="faba">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Vicia faba</emphasis>
|
||||
<taxonomicName id="F2D25BE7CADE58D126164A1A0E2918D4" class="Magnoliopsida" family="Fabaceae" genus="Vicia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vicia faba" order="Fabales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="faba">
|
||||
<emphasis id="7963A8A05E76542FD2FA9869A0AFDBDA" italics="true" pageId="0" pageNumber="47">Vicia faba</emphasis>
|
||||
</taxonomicName>
|
||||
L. (
|
||||
<taxonomicName family="Fabaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Fabaceae</taxonomicName>
|
||||
<taxonomicName id="CE1D04C50BF2D64F1BA402CE7D92EDA8" class="Magnoliopsida" family="Fabaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum tuberosum" order="Solanales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="tuberosum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Solanum tuberosum</emphasis>
|
||||
<taxonomicName id="E31F684181A9DCDB239F42496E5DE15B" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum tuberosum" order="Solanales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="tuberosum">
|
||||
<emphasis id="04BFCA9AA8FBC059D4E03CC14E5E20FB" italics="true" pageId="0" pageNumber="47">Solanum tuberosum</emphasis>
|
||||
</taxonomicName>
|
||||
L. (
|
||||
<taxonomicName family="Solanaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Solanaceae</taxonomicName>
|
||||
<taxonomicName id="809CE64464C7B2333F817F093207464D" class="Magnoliopsida" family="Solanaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
|
||||
),
|
||||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Zea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Zea mays" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="mays">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">Zea mays</emphasis>
|
||||
<taxonomicName id="D2379CF9CC19AEA02CE85528BEDA40E5" class="Liliopsida" family="Poaceae" genus="Zea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Zea mays" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="mays">
|
||||
<emphasis id="85B6EA48D38C9799F92263AE72A0AB97" italics="true" pageId="0" pageNumber="47">Zea mays</emphasis>
|
||||
</taxonomicName>
|
||||
L. (
|
||||
<taxonomicName family="Poaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Poaceae</taxonomicName>
|
||||
<taxonomicName id="540510B30693E137411F21E127CFEF5D" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
|
||||
)), while the paratype was collected in a smaller patch less than 1 km distant from a mining area developed after 1987. A total of 110 vegetation plots were sampled across the Jalca in 2007 (
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/j.1744-7429.2011.00820.x" author="Tovar, C" journalOrPublisher="Biotropica" pageId="0" pageNumber="47" pagination="321 - 330" refId="B16" refString="Tovar, C, Duivenvoorden, JF, Sanchez-Vega, I, Seijmonsbergen, AH, 2012. Recent changes in patch characteristics and plant communities in the Jalca grasslands of the Peruvian Andes. Biotropica 44 (3): 321 - 330, DOI: https://doi.org/10.1111/j.1744-7429.2011.00820.x" title="Recent changes in patch characteristics and plant communities in the Jalca grasslands of the Peruvian Andes." url="https://doi.org/10.1111/j.1744-7429.2011.00820.x" volume="44" year="2012">Tovar et al. 2012</bibRefCitation>
|
||||
<bibRefCitation id="AD902D160597AA54F48380F35FD6431D" DOI="https://doi.org/10.1111/j.1744-7429.2011.00820.x" author="Tovar, C" journalOrPublisher="Biotropica" pageId="0" pageNumber="47" pagination="321 - 330" refId="B16" refString="Tovar, C, Duivenvoorden, JF, Sanchez-Vega, I, Seijmonsbergen, AH, 2012. Recent changes in patch characteristics and plant communities in the Jalca grasslands of the Peruvian Andes. Biotropica 44 (3): 321 - 330, DOI: https://doi.org/10.1111/j.1744-7429.2011.00820.x" title="Recent changes in patch characteristics and plant communities in the Jalca grasslands of the Peruvian Andes." url="https://doi.org/10.1111/j.1744-7429.2011.00820.x" volume="44" year="2012">Tovar et al. 2012</bibRefCitation>
|
||||
) and the new species was found in only two of them. Using the criteria B1a and B1b of the
|
||||
<bibRefCitation author="IUCN" journalOrPublisher="Flora of North America North of Mexico 12 + vols. New York and Oxford, vol. 5" pageId="0" pageNumber="47" publicationUrl="http://www.iucnredlist.org" refId="B6" refString="IUCN, 2019. The IUCN Red List of Threatened Species. Version 2019-2. http://www.iucnredlist.org" title="The IUCN Red List of Threatened Species. Version 2019 - 2." url="http://www.iucnredlist.org" year="2019">IUCN (2019)</bibRefCitation>
|
||||
<bibRefCitation id="2A71FEC8EC513359A76CF34ECE933312" author="IUCN" journalOrPublisher="Flora of North America North of Mexico 12 + vols. New York and Oxford, vol. 5" pageId="0" pageNumber="47" publicationUrl="http://www.iucnredlist.org" refId="B6" refString="IUCN, 2019. The IUCN Red List of Threatened Species. Version 2019-2. http://www.iucnredlist.org" title="The IUCN Red List of Threatened Species. Version 2019 - 2." url="http://www.iucnredlist.org" year="2019">IUCN (2019)</bibRefCitation>
|
||||
, we assessed
|
||||
<taxonomicName lsidName="D. veliziae" pageId="0" pageNumber="47" rank="species" species="veliziae">
|
||||
<emphasis italics="true" pageId="0" pageNumber="47">D. veliziae</emphasis>
|
||||
<taxonomicName id="2CC37C81AD1EF022AEF548F7470F6612" lsidName="D. veliziae" pageId="0" pageNumber="47" rank="species" species="veliziae">
|
||||
<emphasis id="174D92B1416BAE14B0FA3BE4378F3A33" italics="true" pageId="0" pageNumber="47">D. veliziae</emphasis>
|
||||
</taxonomicName>
|
||||
as Endangered species (EN).
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,165 +1,164 @@
|
|||
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||||
<mods:titleInfo>
|
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<mods:title>The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n ..</mods:title>
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<mods:title id="F6642C38A8E5F5727426892527E27DA4">The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n ..</mods:title>
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<mods:namePart id="04687AE39D19F6AA5CF6AAF0781DA762">Fisher, Brian</mods:namePart>
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<mods:date>2014</mods:date>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6137090" ID-GBIF-Taxon="100060543" ID-Zenodo-Dep="6137090" LSID="urn:lsid:plazi:treatment:CB7C011E94CE7F180D112E97B282DAA3" httpUri="http://treatment.plazi.org/id/CB7C011E94CE7F180D112E97B282DAA3" lastPageNumber="52" pageNumber="50">
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<subSubSection type="nomenclature">
|
||||
<paragraph pageId="49" pageNumber="50" type="mainText">
|
||||
<taxonomicName authority="Bolton & Fisher" authorityName="Bolton & Fisher" class="Hexapoda" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="diminuta">
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<treatment id="CB7C011E94CE7F180D112E97B282DAA3" ID-DOI="http://doi.org/10.5281/zenodo.6137090" ID-GBIF-Taxon="100060543" ID-Zenodo-Dep="6137090" LSID="urn:lsid:plazi:treatment:CB7C011E94CE7F180D112E97B282DAA3" httpUri="http://treatment.plazi.org/id/CB7C011E94CE7F180D112E97B282DAA3" lastPageNumber="52" pageNumber="50" scope_family="Formicidae" scope_order="Hymenoptera">
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||||
<subSubSection id="D592B00FB825AA57BB945886566E3140" type="nomenclature">
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||||
<paragraph id="83F63BC5E1A67EECA95474322C423AA2" pageId="49" pageNumber="50" type="mainText">
|
||||
<taxonomicName id="23564E47357DBDC44151AF8FBB6BC3EF" authority="Bolton & Fisher" authorityName="Bolton & Fisher" genus="Royidris" pageId="49" pageNumber="50" rank="species" species="diminuta">
|
||||
Royidris diminuta
|
||||
<authority>Bolton & Fisher</authority>
|
||||
<authority id="529F40A50A7B275BDFBCF01E8D811CE0">Bolton & Fisher</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="698AB3E5BA7CFA81D66675498B34FBA6" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph pageId="49" pageNumber="50" type="mainText">(Figs 48-50, Map 138)</paragraph>
|
||||
<paragraph pageId="49" pageNumber="50" type="mainText">WORKER (holotype in parentheses). TL 2.8-3.0 (3.0), HL 0.66-0.73 (0.72), HW 0.50-0.58 (0.58), CI 76-81 (81), SL 0.66-0.73 (0.70), SI 121-130 (121), PW 0.34-0.38 (0.37), WL 0.88-0.96 (0.96) (7 measured). EL 0.16-0.18 (EL/HW 0.30-0.32). MfL 0.66-0.76 (MfL/HW 1.25-1.33, MfL/MfH 3.36-3.80).</paragraph>
|
||||
<subSubSection id="D16582D5F1AD918DF26B9445CEABC473" type="description">
|
||||
<paragraph id="79B148D323919BF29086896D96F217D3" pageId="49" pageNumber="50" type="mainText">(Figs 48-50, Map 138)</paragraph>
|
||||
<paragraph id="7AACC4B50508D3AF40245C775C289D4E" pageId="49" pageNumber="50" type="mainText">WORKER (holotype in parentheses). TL 2.8-3.0 (3.0), HL 0.66-0.73 (0.72), HW 0.50-0.58 (0.58), CI 76-81 (81), SL 0.66-0.73 (0.70), SI 121-130 (121), PW 0.34-0.38 (0.37), WL 0.88-0.96 (0.96) (7 measured). EL 0.16-0.18 (EL/HW 0.30-0.32). MfL 0.66-0.76 (MfL/HW 1.25-1.33, MfL/MfH 3.36-3.80).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption pageId="50" pageNumber="51">
|
||||
<paragraph pageId="50" pageNumber="51" type="caption">
|
||||
<pageBreakToken pageId="50" pageNumber="51" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="FCC857A88944C6FA924B36C65A6AF522" type="multiple">
|
||||
<caption id="AAC2D2392B9F532A5FA7C0AD88CEF37F" pageId="50" pageNumber="51">
|
||||
<paragraph id="EDC37C6F7B8932C047C964C71D332C52" pageId="50" pageNumber="51" type="caption">
|
||||
<pageBreakToken id="BC4E8386B6777AAF630139280FAFC651" pageId="50" pageNumber="51" start="start">FIGURES</pageBreakToken>
|
||||
48-50. Lateral, full face and dorsal view of body.
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diminuta">Royidris diminuta</taxonomicName>
|
||||
<taxonomicName id="AFD92860007703FAD4930839F9010DA0" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="diminuta">Royidris diminuta</taxonomicName>
|
||||
holotype worker CASENT0002420.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph pageId="50" pageNumber="51" type="mainText">
|
||||
<subSubSection id="E11DD95F4E5B316FAC32DD09D248206B" type="discussion">
|
||||
<paragraph id="222F58CFD842A50AE40445D1819275A9" pageId="50" pageNumber="51" type="mainText">
|
||||
Matching the description of
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="F6F68B558BE06BD7BB9F27AC6E904035" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
, and with critical indices falling within the same ranges, but
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
<taxonomicName id="99F677201DEC639EE7C3ADAFEEFC5D5E" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
much smaller in absolute measurements. For comparison of measurements see Table 2.
|
||||
</paragraph>
|
||||
<caption ID-Table-UUID="4F405ADD2A51BE0C5B4620F63F46740B" httpUri="http://table.plazi.org/id/4F405ADD2A51BE0C5B4620F63F46740B" pageId="50" pageNumber="51">
|
||||
<paragraph pageId="50" pageNumber="51" type="caption">
|
||||
<caption id="4F405ADD2A51BE0C5B4620F63F46740B" ID-Table-UUID="4F405ADD2A51BE0C5B4620F63F46740B" httpUri="http://table.plazi.org/id/4F405ADD2A51BE0C5B4620F63F46740B" pageId="50" pageNumber="51">
|
||||
<paragraph id="63CCA2CE46CF83CFF4837738E4CAD881" pageId="50" pageNumber="51" type="caption">
|
||||
TABLE 2. Comparison of measurements between
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diminuta">Royidris diminuta</taxonomicName>
|
||||
<taxonomicName id="745B5E13C1738380E1A398FC8F2E0ED3" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="diminuta">Royidris diminuta</taxonomicName>
|
||||
and
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="0875C36CBCC16FDE32659FB08D81B211" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="50" pageNumber="51" type="table">
|
||||
<table border="1" pageId="50" pageNumber="51">
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td isEmpty="true" pageId="50" pageNumber="51">-</td>
|
||||
<td pageId="50" pageNumber="51">
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
<paragraph id="6180F3BA8496266F654B56833F5EB812" pageId="50" pageNumber="51" type="table">
|
||||
<table id="69CE9D3F84E88985D58F2C91C584105F" border="1" pageId="50" pageNumber="51">
|
||||
<tr id="B2E96237DBED280F4B39AA103E33BD13" pageId="50" pageNumber="51">
|
||||
<td id="DC32B4920CEEFBB961C0FB6BF061F941" isEmpty="true" pageId="50" pageNumber="51">-</td>
|
||||
<td id="8B38D74E5356088A56C40838F3BF2DA4" pageId="50" pageNumber="51">
|
||||
<taxonomicName id="A9FD362E88F71245AD25F349D40AB28F" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
</td>
|
||||
<td pageId="50" pageNumber="51">
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<td id="7A33B79AF17D6DF5B4AFAA4A7BCA0CC0" pageId="50" pageNumber="51">
|
||||
<taxonomicName id="2D1E8182D51AF847348298D2DF5390DB" genus="Royidris" pageId="50" pageNumber="51" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
</td>
|
||||
</tr>
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td pageId="50" pageNumber="51">HL</td>
|
||||
<td pageId="50" pageNumber="51">0.66-0.73</td>
|
||||
<td pageId="50" pageNumber="51">0.80-0.98</td>
|
||||
<tr id="61028E19FE301E4C24570D6B360F3DE9" pageId="50" pageNumber="51">
|
||||
<td id="61B8994AEA019D9C2EA8A8AF14C31830" pageId="50" pageNumber="51">HL</td>
|
||||
<td id="E19F8214355868EBF74B82791E932E31" pageId="50" pageNumber="51">0.66-0.73</td>
|
||||
<td id="F7CB3B771048E6D3395BB0161CBEABC6" pageId="50" pageNumber="51">0.80-0.98</td>
|
||||
</tr>
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td pageId="50" pageNumber="51">HW</td>
|
||||
<td pageId="50" pageNumber="51">0.50-0.58</td>
|
||||
<td pageId="50" pageNumber="51">0.65-0.81</td>
|
||||
<tr id="DA246E61DD2F028E0F15420B4072C69D" pageId="50" pageNumber="51">
|
||||
<td id="F206C2A0CF10D25B4D900DF9186BC2B5" pageId="50" pageNumber="51">HW</td>
|
||||
<td id="814F96081CB6EA4D7D7A072E6FDA4BBD" pageId="50" pageNumber="51">0.50-0.58</td>
|
||||
<td id="0108F2A9819C3C7BB9D716D4EF8B2454" pageId="50" pageNumber="51">0.65-0.81</td>
|
||||
</tr>
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td pageId="50" pageNumber="51">SL</td>
|
||||
<td pageId="50" pageNumber="51">0.66-0.73</td>
|
||||
<td pageId="50" pageNumber="51">0. 86 - 1.00</td>
|
||||
<tr id="A27DD53B0598A794FDC2A4CC9A546C36" pageId="50" pageNumber="51">
|
||||
<td id="9BE714A0444694DAAA3A42237DC4FE40" pageId="50" pageNumber="51">SL</td>
|
||||
<td id="29987B96AD4146814CE726353016A764" pageId="50" pageNumber="51">0.66-0.73</td>
|
||||
<td id="6CA1E5817612B76AF0F16F6AF46E377A" pageId="50" pageNumber="51">0. 86 - 1.00</td>
|
||||
</tr>
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td pageId="50" pageNumber="51">PW</td>
|
||||
<td pageId="50" pageNumber="51">0.34-0.38</td>
|
||||
<td pageId="50" pageNumber="51">0.43-0.51</td>
|
||||
<tr id="900D6F6336EF2D4D82CB589E9E51ACF8" pageId="50" pageNumber="51">
|
||||
<td id="DD8E0DF36B3BDE401012D42F8D69FC8D" pageId="50" pageNumber="51">PW</td>
|
||||
<td id="AABC19D94EC44AA7EC653DA35A88359F" pageId="50" pageNumber="51">0.34-0.38</td>
|
||||
<td id="47D1A30FD0CC2C80044513030EF004AF" pageId="50" pageNumber="51">0.43-0.51</td>
|
||||
</tr>
|
||||
<tr pageId="50" pageNumber="51">
|
||||
<td pageId="50" pageNumber="51">MfL</td>
|
||||
<td pageId="50" pageNumber="51">0.66-0.76</td>
|
||||
<td pageId="50" pageNumber="51">0. 88 - 1.10</td>
|
||||
<tr id="58EB9C6295D51D1BE6F8E6CAFEE0D4A0" pageId="50" pageNumber="51">
|
||||
<td id="42CE301E412C52365073FB82A0660FE2" pageId="50" pageNumber="51">MfL</td>
|
||||
<td id="9434CED97F4013733C433A03F7724280" pageId="50" pageNumber="51">0.66-0.76</td>
|
||||
<td id="9497C96089B4C93FCA7C7353702E1E16" pageId="50" pageNumber="51">0. 88 - 1.10</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">
|
||||
<pageBreakToken pageId="51" pageNumber="52" start="start">QUEEN</pageBreakToken>
|
||||
<subSubSection id="FBDCA3A6F4D76DE2D70E883EF13447C8" type="description">
|
||||
<paragraph id="617F86CF1E3EC6157FB01F2D409DA0A5" pageId="51" pageNumber="52" type="mainText">
|
||||
<pageBreakToken id="4360A7C821402B26A8E3082BAA78DF4E" pageId="51" pageNumber="52" start="start">QUEEN</pageBreakToken>
|
||||
(gyne). Alate when virgin. Very much larger than the worker. TL 6.8, HL 1.25, HW 1.21, CI 97, SL 1.04, SI 86, PW 0.90, WL 2.08, EL 0.36 (EL/HW 0.30), MfL 1.22 (MfL/HW 1.01, MfL/MfH 4.36) (1 measured).
|
||||
</paragraph>
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">MALE: unknown.</paragraph>
|
||||
<paragraph id="22ECA7CD6530896AEC8B9695B65635F4" pageId="51" pageNumber="52" type="mainText">MALE: unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">Holotype worker (top specimen of three on pin), Madagascar: Prov. Toliara, Mahafaly Plateau, 6.2 km. 74° ENE Itampolo, 24°39.2'S, 43°59.8'E, 80 m., 21-25.ii.2002, BLF 5814, CASENT0002420, under stone, spiny forest/thicket (Fisher et al.) (CASC).</paragraph>
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">Paratypes. 2 workers on same pin as holotype, 2 workers with same data but CASENT0002421, 1 worker with same data but CASENT0009802 (CASC).</paragraph>
|
||||
<subSubSection id="261A247020AEF926680AB2CE3AF5D577" type="materials_examined">
|
||||
<paragraph id="47F4CDC24FBE200B5189AAD24115DCBD" pageId="51" pageNumber="52" type="mainText">Holotype worker (top specimen of three on pin), Madagascar: Prov. Toliara, Mahafaly Plateau, 6.2 km. 74° ENE Itampolo, 24°39.2'S, 43°59.8'E, 80 m., 21-25.ii.2002, BLF 5814, CASENT0002420, under stone, spiny forest/thicket (Fisher et al.) (CASC).</paragraph>
|
||||
<paragraph id="CAC97C9155B9D93F4532387B74802479" pageId="51" pageNumber="52" type="mainText">Paratypes. 2 workers on same pin as holotype, 2 workers with same data but CASENT0002421, 1 worker with same data but CASENT0009802 (CASC).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">
|
||||
<subSubSection id="33D804D2E1957344971EF547CF348B05" type="discussion">
|
||||
<paragraph id="612234F386C5667F6954131E0449DAEC" pageId="51" pageNumber="52" type="mainText">
|
||||
This small form is separated from
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="CC11C90A1A1D1ACB815D10C9FF201E39" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
with some reservations. In terms of general morphology and critical indices, workers of the two are similar, but in terms of absolute dimensions there is a distinct and persistent gap between them in the ranges of all measurements, with no trace of intermediates. No species of
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Royidris</taxonomicName>
|
||||
<taxonomicName id="2CE74635641D8FC63F576E4BD6936D16" genus="Royidris" pageId="51" pageNumber="52" rank="genus">Royidris</taxonomicName>
|
||||
show any signs of worker dimorphism, but the possibility that the
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
<taxonomicName id="E3022658A2F50F15D0EA59EE2B01FBD2" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
workers represent the undersized first brood of new foundress queens of
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="93213C7EA2972F829A8705E7EE679D1D" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
cannot be entirely ruled out. An alternative hypothesis, that
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
<taxonomicName id="596B732CD003FFBCD3AD1E7AFE3C3CD2" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
,
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="etiolata">etiolata</taxonomicName>
|
||||
<taxonomicName id="0199B1811E434BB01F5C898596CB3FCB" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="etiolata">etiolata</taxonomicName>
|
||||
and
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="98AA40C3D872CA975676BB25F3888551" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
constitute a single species, results in a monomorphic species with SI 115-158, an exceptionally broad range. All three species are found in sympatry in the Mahafaly Plateau.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="biology_ecology">
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">
|
||||
<subSubSection id="4414BA43A7A9AD7728E2FC2852086B00" type="biology_ecology">
|
||||
<paragraph id="6F84D959A1C71D2533459D92C8379802" pageId="51" pageNumber="52" type="mainText">
|
||||
Both series of
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
<taxonomicName id="2B4A0B3EB84CCBB8B14D0FCD1EF3471D" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="diminuta">diminuta</taxonomicName>
|
||||
, which nests under stones in spiny forest, were included under
|
||||
<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
<taxonomicName id="D96506379274D1E9CBBC726A550CEB62" genus="Royidris" pageId="51" pageNumber="52" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
|
||||
by Heterick (2006).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph pageId="51" pageNumber="52" type="mainText">Non-paratypic material examined. Madagascar: Prov. Toliara, Mahafaly Plateau, ENE Itampolo (Fisher et al.).</paragraph>
|
||||
<subSubSection id="7AA42136FA1D73D176AACF9A530AAECD" type="materials_examined">
|
||||
<paragraph id="883204FE6B23746BA4EC4CFE8C9FF4DB" pageId="51" pageNumber="52" type="mainText">Non-paratypic material examined. Madagascar: Prov. Toliara, Mahafaly Plateau, ENE Itampolo (Fisher et al.).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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