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<mods:title id="030906E2436BA39CD7E62461F879622F">The genus Euodynerus Dalla Torre in Europe and the Maghreb (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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<mods:namePart id="85B4B9E8EE14A8E267E892F6C23A87BA">Selis, Marco</mods:namePart>
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<mods:namePart id="5C0E860ADDE13553401641F6A46CD682">Fateryga, Alexander V.</mods:namePart>
<mods:affiliation id="532BF1203C80AC84BDF898C4DFD63C07">T. I. Vyazemsky Karadag Scientific Station-Nature Reserve of RAS-Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nauki Str. 24, Kurortnoye, 298188 Feodosiya, Russia</mods:affiliation>
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<mods:namePart id="2CF164470F428CE9182A5285ABE512CD">Cilia, Giovanni</mods:namePart>
<mods:affiliation id="A077A967B2AD8F08E0FFE5858FFEDF58">CREA Research Centre for Agriculture and Environment (CREA-AA), Via di Corticella 133, 40128, Bologna, Italy</mods:affiliation>
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<mods:date id="E382346CF1446A50B0AA5407028664A2">2024</mods:date>
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<heading id="1B38761762049A3289FDF5B2D493FF60" bold="true" box="[151,916,151,178]" fontSize="11" level="1" pageId="14" pageNumber="165" reason="1">
<taxonomicName id="87CFBAF862049A3289FDF5B2D493FF60" ID-CoL="5H8SR" authority="Bluthgen, 1956" authorityName="Bluthgen" authorityYear="1956" box="[151,916,151,178]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="subSpecies" species="bidentatus" subGenus="Euodynerus" subSpecies="puniceus">
<emphasis id="72BB1D6962049A3289FDF5B2D493FF60" bold="true" box="[151,916,151,178]" pageId="14" pageNumber="165">
<emphasis id="72BB1D6962049A3289FDF5B2D62DFF63" bold="true" box="[151,298,151,178]" italics="true" pageId="14" pageNumber="165">Euodynerus</emphasis>
(
<emphasis id="72BB1D6962049A328851F5B2D6C9FF63" bold="true" box="[315,462,151,178]" italics="true" pageId="14" pageNumber="165">Euodynerus</emphasis>
)
<emphasis id="72BB1D6962049A3288B5F5B2D5D6FF60" bold="true" box="[479,721,151,178]" italics="true" pageId="14" pageNumber="165">bidentatus puniceus</emphasis>
<bibRefCitation id="245EBC8A62049A328BB2F5B2D493FF60" author="Bluthgen, P." box="[728,916,151,177]" pageId="14" pageNumber="165" pagination="313 - 318" refId="ref22022" refString="Bluthgen, P. (1956) Neue oder erwahnenswerte &quot; Eumenidae &quot; aus Italien und Nord-Afrika aus dem Istituto di Entomologia della Universita di Bologna (I. B.) und aus dem Musee Zoologique de Lausanne (M. L.). Bollettino dell'Istituto di Entomologia dell'Universita di Bologna, 21, 313 - 318." type="journal article" year="1956">Blüthgen, 1956</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="08D592F062049A3289FDF59ED626FF04" box="[151,289,187,213]" pageId="14" pageNumber="165" type="description">
<paragraph id="4070C17B62049A3289FDF59ED626FF04" blockId="14.[151,916,151,213]" box="[151,289,187,213]" pageId="14" pageNumber="165">
(
<figureCitation id="D8F4DDFE62049A3289F5F59ED61FFF04" box="[159,280,187,213]" captionStart="FIGURE 4" captionStartId="13.[151,250,1804,1828]" captionTargetBox="[151,1435,181,1780]" captionTargetId="figure-17@13.[151,1435,181,1780]" captionTargetPageId="13" captionText="FIGURE 4. A, C–D, Euodynerus bidentatus bidentatus (Lepeletier). A, D, female from Morocco. C, male from Morocco. B, E, Euodynerus bidentatus puniceus Blüthgen, female from Libya. F–I, Euodynerus caspicus (Morawitz). F, H, male from Astrakhan Province, Russia. G, I, female from Kazakhstan." figureDoi="http://doi.org/10.5281/zenodo.14239447" httpUri="https://zenodo.org/record/14239447/files/figure.png" pageId="14" pageNumber="165">Figs 4B, E</figureCitation>
)
</paragraph>
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<subSubSection id="08D592F062049A3289FDF425D201FEC9" box="[151,1286,255,280]" pageId="14" pageNumber="165" type="reference_group">
<paragraph id="4070C17B62049A3289FDF425D201FEC9" blockId="14.[151,1286,255,280]" box="[151,1286,255,280]" pageId="14" pageNumber="165">
<treatmentCitationGroup id="60DFE65562049A3289FDF425D201FEC9" box="[151,1286,255,280]" pageId="14" pageNumber="165">
<treatmentCitation id="C16EE76A62049A3289FDF425D5BFFEC9" author="Bluthgen, P." box="[151,696,255,280]" page="313" pageId="14" pageNumber="165" year="1956">
<taxonomicName id="87CFBAF862049A3289FDF425D5BFFEC9" ID-CoL="5H8SR" authority="Bluthgen, 1956: 313" authorityName="Bluthgen" authorityPageNumber="313" authorityYear="1956" box="[151,696,255,280]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="subSpecies" species="bidentatus" subSpecies="puniceus">
<emphasis id="72BB1D6962049A3289FDF425D6E7FEC6" box="[151,480,256,280]" italics="true" pageId="14" pageNumber="165">Euodynerus bidentatus puniceus</emphasis>
<materialsCitation id="F0A7CB2662049A32888DF425D5BFFEC9" box="[487,696,255,280]" collectionCode="MZUB" pageId="14" pageNumber="165" specimenCount="1" specimenCount-female="1">
<bibRefCitation id="245EBC8A62049A32888DF425D5BFFEC9" author="Bluthgen, P." box="[487,696,255,280]" pageId="14" pageNumber="165" pagination="313 - 318" refId="ref22022" refString="Bluthgen, P. (1956) Neue oder erwahnenswerte &quot; Eumenidae &quot; aus Italien und Nord-Afrika aus dem Istituto di Entomologia della Universita di Bologna (I. B.) und aus dem Musee Zoologique de Lausanne (M. L.). Bollettino dell'Istituto di Entomologia dell'Universita di Bologna, 21, 313 - 318." type="journal article" year="1956">Blüthgen, 1956: 313</bibRefCitation>
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,
<specimenCount id="56C90AF262049A328BA9F425D5D1FEC6" box="[707,726,256,279]" count="1" pageId="14" pageNumber="165" type="female">♀</specimenCount>
<emphasis id="72BB1D6962049A328BBCF425D5EAFEC6" box="[726,749,256,279]" italics="true" pageId="14" pageNumber="165">—</emphasis>
“Bu el Gheràb (Tripolitanien)” (MZUB [examined]).
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</paragraph>
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<subSubSection id="08D592F062049A3289FDF460D543FC71" pageId="14" pageNumber="165" type="distribution">
<paragraph id="4070C17B62049A3289FDF460D5B5FE79" blockId="14.[151,1437,325,928]" pageId="14" pageNumber="165">
<emphasis id="72BB1D6962049A3289FDF460D636FE8E" bold="true" box="[151,305,325,351]" pageId="14" pageNumber="165">Distribution.</emphasis>
<collectingCountry id="38D881EB62049A328851F463D678FEB1" box="[315,383,326,352]" name="Libya" pageId="14" pageNumber="165">Libya</collectingCountry>
, currently known only from the typical locality of Bi’r Bu al Ghurab (
<bibRefCitation id="245EBC8A62049A328DD3F463D26FFEB0" author="Bluthgen, P." box="[1209,1384,326,353]" pageId="14" pageNumber="165" pagination="313 - 318" refId="ref22022" refString="Bluthgen, P. (1956) Neue oder erwahnenswerte &quot; Eumenidae &quot; aus Italien und Nord-Afrika aus dem Istituto di Entomologia della Universita di Bologna (I. B.) und aus dem Musee Zoologique de Lausanne (M. L.). Bollettino dell'Istituto di Entomologia dell'Universita di Bologna, 21, 313 - 318." type="journal article" year="1956">Blüthgen 1956</bibRefCitation>
). As reported above,
<bibRefCitation id="245EBC8A62049A32883AF44FD557FE54" author="Giordani Soika, A." box="[336,592,362,389]" pageId="14" pageNumber="165" pagination="235 - 267" refId="ref23302" refString="Giordani Soika, A. (1953 [1952]) Hymenopteres recoltes par une Mission suisse au Maroc (1947). Eumeninae. Bulletin de la Societe des sciences naturelles du Maroc, 32, 235 - 267." type="book chapter" year="1953">Giordani Soika (1953)</bibRefCitation>
recorded
<taxonomicName id="87CFBAF862049A328BAEF44ED45DFE55" authorityName="MA Tafraoute" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[708,858,362,388]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="72BB1D6962049A328BAEF44ED45DFE55" box="[708,858,362,388]" italics="true" pageId="14" pageNumber="165">E. bidentatus</emphasis>
</taxonomicName>
from
<collectingCountry id="38D881EB62049A328AC8F44FD4E4FE55" box="[930,995,362,388]" name="Libya" pageId="14" pageNumber="165">Libya</collectingCountry>
(Gharyan and Cyrenaica), but the real identity of those records remains to be assessed.
</paragraph>
<paragraph id="4070C17B62049A3289ADF497D543FC71" blockId="14.[151,1437,325,928]" pageId="14" pageNumber="165">
<emphasis id="72BB1D6962049A3289ADF497D616FE1D" bold="true" box="[199,273,434,460]" pageId="14" pageNumber="165">Notes.</emphasis>
<bibRefCitation id="245EBC8A62049A328876F497D6DFFE1C" author="Bluthgen, P." box="[284,472,434,461]" pageId="14" pageNumber="165" pagination="313 - 318" refId="ref22022" refString="Bluthgen, P. (1956) Neue oder erwahnenswerte &quot; Eumenidae &quot; aus Italien und Nord-Afrika aus dem Istituto di Entomologia della Universita di Bologna (I. B.) und aus dem Musee Zoologique de Lausanne (M. L.). Bollettino dell'Istituto di Entomologia dell'Universita di Bologna, 21, 313 - 318." type="journal article" year="1956">Blüthgen (1956)</bibRefCitation>
described this taxon as a Libyan subspecies of the western Maghrebi
<taxonomicName id="87CFBAF862049A328C7CF497D69DFE21" authority="(Lepeletier)" authorityName="MA Tafraoute" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="72BB1D6962049A328C7CF497D60BFE21" italics="true" pageId="14" pageNumber="165">Euodynerus bidentatus</emphasis>
(Lepeletier)
</taxonomicName>
, differentiating it by the reduced and ferruginous markings only (
<figureCitation id="D8F4DDFE62049A328D05F4F3D3EDFE21" box="[1135,1258,470,496]" captionStart="FIGURE 4" captionStartId="13.[151,250,1804,1828]" captionTargetBox="[151,1435,181,1780]" captionTargetId="figure-17@13.[151,1435,181,1780]" captionTargetPageId="13" captionText="FIGURE 4. A, C–D, Euodynerus bidentatus bidentatus (Lepeletier). A, D, female from Morocco. C, male from Morocco. B, E, Euodynerus bidentatus puniceus Blüthgen, female from Libya. F–I, Euodynerus caspicus (Morawitz). F, H, male from Astrakhan Province, Russia. G, I, female from Kazakhstan." figureDoi="http://doi.org/10.5281/zenodo.14239447" httpUri="https://zenodo.org/record/14239447/files/figure.png" pageId="14" pageNumber="165">Figs 4B, E</figureCitation>
). Other than the pattern, the comparison of
<specimenCount id="56C90AF262049A3288A1F4DFD54DFDC5" box="[459,586,506,532]" count="2" pageId="14" pageNumber="165" type="female">two female</specimenCount>
specimens of
<taxonomicName id="87CFBAF862049A328B9BF4DED4F4FDC5" authorityName="Bluthgen" authorityYear="1956" box="[753,1011,506,532]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="subSpecies" species="bidentatus" subSpecies="puniceus">
<emphasis id="72BB1D6962049A328B9BF4DED4F4FDC5" box="[753,1011,506,532]" italics="true" pageId="14" pageNumber="165">E. bidentatus puniceus</emphasis>
</taxonomicName>
, including the
<typeStatus id="9F747FD962049A328DCAF4DFD202FDC5" box="[1184,1285,506,532]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
, with several specimens of the nominotypical subspecies revealed two additional subtle differences in sculpture: clypeus with more defined punctures less tending to form longitudinal striae and mesoscutum and metasoma with slightly sparser punctures. In addition, DNA barcoding of one of the
<specimenCount id="56C90AF262049A328B95F743D4A0FD51" box="[767,935,614,640]" count="2" pageId="14" pageNumber="165" type="generic">two specimens</specimenCount>
resulted in a sequence presenting a genetic distance of 6.64% from the nominotypical subspecies (
<figureCitation id="D8F4DDFE62049A328A67F7AFD452FD75" box="[781,853,650,676]" captionStart="FIGURE 2" captionStartId="11.[151,250,1908,1932]" captionTargetBox="[272,1316,192,1873]" captionTargetId="figure-17@11.[261,1327,181,1884]" captionTargetPageId="11" captionText="FIGURE 2. Phylogenetic tree (Neighbor-Joining) of Euodynerus (Euodynerus) based on the mitochondrial COI gene. Ancistrocerus gazella (Panzer) was used as an outgroup. Numbers adjacent to branches represent bootstrap support (values of &lt;75 are omitted); only the species occurring in the considered area are highlighted. Countries are abbreviated using ISO 3166-1 alpha-2 codes. Continues in Fig. 3." figureDoi="http://doi.org/10.5281/zenodo.14239443" httpUri="https://zenodo.org/record/14239443/files/figure.png" pageId="14" pageNumber="165">Fig. 2</figureCitation>
). The weak morphological differences and the low genetic distance would suggest synonymizing the subspecies
<emphasis id="72BB1D6962049A328A56F78AD4A7FD19" box="[828,928,687,712]" italics="true" pageId="14" pageNumber="165">puniceus</emphasis>
with the nominotypical one, however, the two subspecies do not seem to present intermediate forms as regards the chromatic pattern and the subspecies
<emphasis id="72BB1D6962049A328C4AF7F6D283FD3D" box="[1312,1412,723,748]" italics="true" pageId="14" pageNumber="165">puniceus</emphasis>
is currently known only from a very isolated locality in the Libyan desert, thus indicating a certain degree of isolation between the two populations. Since this could be a case analogous to what was observed in the
<taxonomicName id="87CFBAF862049A328C6FF63FD6E2FC89" authority="(de Saussure)" baseAuthorityName="de Saussure" baseAuthorityYear="1856" class="Insecta" family="Eumenidae" genus="Stenodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="fastidiosissimus">
<emphasis id="72BB1D6962049A328C6FF63FD64FFC89" italics="true" pageId="14" pageNumber="165">Stenodynerus fastidiosissimus</emphasis>
(de Saussure)
</taxonomicName>
/
<taxonomicName id="87CFBAF862049A328887F61BD426FC89" authority="Giordani Soika" authorityName="Giordani Soika" box="[493,801,830,856]" class="Insecta" family="Eumenidae" genus="Stenodynerus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="72BB1D6962049A328887F61BD575FC89" box="[493,626,830,856]" italics="true" pageId="14" pageNumber="165">S. rufescens</emphasis>
Giordani Soika
</taxonomicName>
pair (
<bibRefCitation id="245EBC8A62049A328A0BF61BD310FC89" author="Selis, M. &amp; Cilia, G. &amp; Wood, T. J. &amp; Soon, V." box="[865,1047,830,856]" pageId="14" pageNumber="165" pagination="34 - 56" refId="ref25682" refString="Selis, M., Cilia, G., Wood, T. J. &amp; Soon, V. (2024) Taxonomic revision of the Stenodynerus fastidiosissimus species-group in Western Europe and North Africa (Hymenoptera: Vespidae: Eumeninae). Zootaxa, 5418 (1), 34 - 56. https: // doi. org / 10.11646 / zootaxa. 5418.1.2" type="journal article" year="2024">
Selis
<emphasis id="72BB1D6962049A328AF4F61AD4D1FC89" box="[926,982,830,856]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
2024
</bibRefCitation>
), in which the females do not show clear morphological differences, we prefer to maintain the subspecies
<emphasis id="72BB1D6962049A328AD4F646D325FCAD" box="[958,1058,867,892]" italics="true" pageId="14" pageNumber="165">puniceus</emphasis>
pending the availability of more material and the discovery of its male.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="B0DCE8CC6CA431EF83D30776F2C059C8">The genus Euodynerus Dalla Torre in Europe and the Maghreb (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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<mods:namePart id="3C4D726893C95906F81DD1D016D7094A">Selis, Marco</mods:namePart>
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<mods:namePart id="F9B23FF8D0AA0EF5DDF940B4353771C1">Fateryga, Alexander V.</mods:namePart>
<mods:affiliation id="CAA5DA94C009002C465F0F3B3A0D4306">T. I. Vyazemsky Karadag Scientific Station-Nature Reserve of RAS-Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nauki Str. 24, Kurortnoye, 298188 Feodosiya, Russia</mods:affiliation>
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<heading id="1B38761762189A2E89FDF38FD419F915" bold="true" box="[151,798,1706,1733]" fontSize="11" level="1" pageId="18" pageNumber="169" reason="1">
<taxonomicName id="87CFBAF862189A2E89FDF38FD419F915" ID-CoL="6GY3W" authority="Bluthgen, 1942" authorityName="Bluthgen. A" authorityYear="1942" box="[151,798,1706,1733]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="hellenicus" subGenus="Euodynerus">
<emphasis id="72BB1D6962189A2E89FDF38FD419F915" bold="true" box="[151,798,1706,1733]" pageId="18" pageNumber="169">
<emphasis id="72BB1D6962189A2E89FDF38FD62DF914" bold="true" box="[151,298,1706,1733]" italics="true" pageId="18" pageNumber="169">Euodynerus</emphasis>
(
<emphasis id="72BB1D6962189A2E8851F38FD6C9F914" bold="true" box="[315,462,1706,1733]" italics="true" pageId="18" pageNumber="169">Euodynerus</emphasis>
)
<emphasis id="72BB1D6962189A2E88B5F38FD55CF914" bold="true" box="[479,603,1706,1733]" italics="true" pageId="18" pageNumber="169">hellenicus</emphasis>
<bibRefCitation id="245EBC8A62189A2E8B08F38ED419F915" author="Bluthgen, P." box="[610,798,1706,1733]" pageId="18" pageNumber="169" pagination="300 - 322" refId="ref21813" refString="Bluthgen, P. (1942) Neue palaarktische Faltenwespen (Hymenoptera, Vespidae, Eumenidinae). Mitteilungen aus dem Zoologischen Museum in Berlin, 25, 300 - 322." type="journal article" year="1942">Blüthgen, 1942</bibRefCitation>
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(
<figureCitation id="D8F4DDFE62189A2E89F5F3EAD61CF938" box="[159,283,1743,1769]" captionStart="FIGURE 8" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[200,1386,192,1918]" captionTargetId="figure-20@20.[200,1386,181,1918]" captionTargetPageId="20" captionText="FIGURE 8. A–F, Euodynerus hellenicus Blüthgen. A, male from Bulgaria. B, E, female from Spain. C, F, female from Iran. D, male from Spain. G–M, Euodynerus semisaecularis (Dalla Torre). G, J, male from Greece. H, L, female from Crete, Greece. I, M, female from Afghanistan. K, male from Armenia." figureDoi="http://doi.org/10.5281/zenodo.14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="18" pageNumber="169">Figs 8A–F</figureCitation>
;
<figureCitation id="D8F4DDFE62189A2E884CF3EAD65DF938" box="[294,346,1743,1769]" captionStart="FIGURE 14" captionStartId="30.[151,250,1557,1581]" captionTargetBox="[153,1423,182,1517]" captionTargetId="figure-20@30.[152,1436,181,1533]" captionTargetPageId="30" captionText="FIGURE 14. Male genitalia, from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Euodynerus bidentatus (Lepeletier). B, E. caspicus (Morawitz). C, E. curictensis Blüthgen. D, E. dantici (Rossi). E, E. disconotatus (Lichtenstein). F, E. fastidiosus (de Saussure). G, E. hellenicus Blüthgen. H, E. semisaecularis (Dalla Torre). I, E. variegatus (Fabricius). J, E. velutinus Blüthgen. K, E. bidentoides (Giordani Soika). L, E. notatus (Jurine). M, E. posticus (Herrich-Schaeffer). N, E. quadrifasciatus (Fabricius)." figureDoi="http://doi.org/10.5281/zenodo.14239467" httpUri="https://zenodo.org/record/14239467/files/figure.png" pageId="18" pageNumber="169">14G</figureCitation>
;
<figureCitation id="D8F4DDFE62189A2E880FF3EAD6BEF938" box="[357,441,1743,1769]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="18" pageNumber="169">16D, H</figureCitation>
)
</paragraph>
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<paragraph id="4070C17B62189A2E89FDF236D352F8FA" blockId="18.[151,1333,1811,1866]" box="[151,1109,1811,1835]" pageId="18" pageNumber="169">
<treatmentCitationGroup id="60DFE65562189A2E89FDF236D352F8FA" box="[151,1109,1811,1835]" pageId="18" pageNumber="169">
<treatmentCitation id="C16EE76A62189A2E89FDF236D5A5F8FA" author="Bluthgen, P." box="[151,674,1811,1835]" page="301" pageId="18" pageNumber="169" year="1942">
<taxonomicName id="87CFBAF862189A2E89FDF236D5A5F8FA" ID-CoL="6GY3W" authority="Bluthgen, 1942: 301" authorityName="Bluthgen" authorityPageNumber="301" authorityYear="1942" box="[151,674,1811,1835]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="subSpecies" species="dantici" subSpecies="hellenicus">
<emphasis id="72BB1D6962189A2E89FDF236D6CDF8FA" box="[151,458,1811,1835]" italics="true" pageId="18" pageNumber="169">Euodynerus dantici hellenicus</emphasis>
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<bibRefCitation id="245EBC8A62189A2E88BAF236D5A5F8FA" author="Bluthgen, P." box="[464,674,1811,1835]" pageId="18" pageNumber="169" pagination="300 - 322" refId="ref21813" refString="Bluthgen, P. (1942) Neue palaarktische Faltenwespen (Hymenoptera, Vespidae, Eumenidinae). Mitteilungen aus dem Zoologischen Museum in Berlin, 25, 300 - 322." type="journal article" year="1942">Blüthgen, 1942: 301</bibRefCitation>
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,
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,
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<emphasis id="72BB1D6962189A2E8BB5F231D5F1F8FA" box="[735,758,1812,1835]" italics="true" pageId="18" pageNumber="169">—</emphasis>
“Amorgos (Kykladen)” (NHMW).
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<treatmentCitationGroup id="60DFE65562189A2E89FDF217D232F89B" box="[151,1333,1842,1866]" pageId="18" pageNumber="169">
<treatmentCitation id="C16EE76A62189A2E89FDF217D5B7F898" author="Gusenleitner, J." box="[151,688,1842,1866]" page="77" pageId="18" pageNumber="169" year="1972">
<taxonomicName id="87CFBAF862189A2E89FDF217D5B7F898" ID-CoL="3CGMS" authority="Gusenleitner, 1972: 77" authorityName="Gusenleitner" authorityPageNumber="77" authorityYear="1972" box="[151,688,1842,1866]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="subSpecies" species="hellenicus" subSpecies="vechti">
<emphasis id="72BB1D6962189A2E89FDF217D6C7F89B" box="[151,448,1842,1866]" italics="true" pageId="18" pageNumber="169">Euodynerus hellenicus vechti</emphasis>
<materialsCitation id="F0A7CB2662189A2E88ACF217D5B7F898" box="[454,688,1842,1866]" collectionCode="RMNH" country="Spain" location="Badajoz" municipality="Merida" pageId="18" pageNumber="169" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="245EBC8A62189A2E88ACF217D5B7F898" author="Gusenleitner, J." box="[454,688,1842,1866]" pageId="18" pageNumber="169" pagination="73 - 78" refId="ref23689" refString="Gusenleitner, J. (1972) Bemerkenswertes uber Faltenwespen IV. (Diploptera, Hymenoptera). Nachrichtenblatt der Bayerischen Entomologen, 21 (4), 73 - 78." type="journal article" year="1972">Gusenleitner, 1972: 77</bibRefCitation>
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,
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,
<specimenCount id="56C90AF262189A2E8BB3F217D5EBF898" box="[729,748,1842,1865]" count="1" pageId="18" pageNumber="169" type="male">♂</specimenCount>
<emphasis id="72BB1D6962189A2E8B86F216D404F89B" box="[748,771,1843,1866]" italics="true" pageId="18" pageNumber="169">—</emphasis>
“
<collectingCountry id="38D881EB62189A2E8A66F217D465F89B" box="[780,866,1842,1866]" name="Spain" pageId="18" pageNumber="169">Spanien</collectingCountry>
,
<location id="451097A062189A2E8A06F217D4C5F89B" LSID="urn:lsid:plazi:treatment:C866706D62189A2989FDF38FD5A2FECC:451097A062189A2E8A06F217D4C5F89B" box="[876,962,1842,1866]" country="Spain" municipality="Merida" name="Badajoz" pageId="18" pageNumber="169">Badajoz</location>
,
<collectingMunicipality id="A0145B0162189A2E8AA6F217D31FF89B" box="[972,1048,1842,1866]" pageId="18" pageNumber="169">Merida</collectingMunicipality>
” (
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female RMNH).
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<emphasis id="72BB1D6962189A2E89FDF25DD636F843" bold="true" box="[151,305,1912,1938]" pageId="18" pageNumber="169">Distribution.</emphasis>
Occurring in the Iberian Peninsula, in the area ranging from the Balkan Peninsula to
<collectingCountry id="38D881EB62189A2E8D8EF25DD26AF843" box="[1252,1389,1912,1938]" name="Afghanistan" pageId="18" pageNumber="169">Afghanistan</collectingCountry>
and descending into the Levant (
<bibRefCitation id="245EBC8A62189A2E88BEF2B9D5A0F867" author="Gusenleitner, J." box="[468,679,1948,1974]" pageId="18" pageNumber="169" pagination="5 - 107" refId="ref23878" refString="Gusenleitner, J. (2013) Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae). Linzer biologische Beitrage, 45 (1), 5 - 107." type="journal article" year="2013">Gusenleitner 2013</bibRefCitation>
; present data).
</paragraph>
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<paragraph id="4070C17B62199A2F89FDF251D59FF829" blockId="19.[151,1436,1908,2041]" pageId="19" pageNumber="170">
<emphasis id="72BB1D6962199A2F89FDF251D656F85D" bold="true" box="[151,337,1908,1933]" pageId="19" pageNumber="170">FIGURE 7. A–H</emphasis>
,
<taxonomicName id="87CFBAF862199A2F8808F251D5F8F85D" authority="(Lichtenstein)" authorityName="UZ Uchkulach" baseAuthorityName="Lichtenstein" baseAuthorityYear="1884" box="[354,767,1908,1932]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="disconotatus">
<emphasis id="72BB1D6962199A2F8808F251D563F85D" box="[354,612,1908,1932]" italics="true" pageId="19" pageNumber="170">Euodynerus disconotatus</emphasis>
(Lichtenstein)
</taxonomicName>
.
<emphasis id="72BB1D6962199A2F8A61F251D41AF85D" bold="true" box="[779,797,1908,1932]" pageId="19" pageNumber="170">A</emphasis>
, male from Azerbaijan.
<emphasis id="72BB1D6962199A2F8D75F250D337F85C" bold="true" box="[1055,1072,1909,1933]" pageId="19" pageNumber="170">B</emphasis>
,
<emphasis id="72BB1D6962199A2F8D2AF250D356F85C" bold="true" box="[1088,1105,1909,1933]" pageId="19" pageNumber="170">E</emphasis>
, male from Italy.
<emphasis id="72BB1D6962199A2F8C78F251D223F85D" bold="true" box="[1298,1316,1908,1932]" pageId="19" pageNumber="170">C</emphasis>
,
<emphasis id="72BB1D6962199A2F8C5EF251D24FF85D" bold="true" box="[1332,1352,1908,1932]" pageId="19" pageNumber="170">G</emphasis>
, female from Italy.
<emphasis id="72BB1D6962199A2F8861F2BCD61AF860" bold="true" box="[267,285,1945,1969]" pageId="19" pageNumber="170">D</emphasis>
, female from Greece.
<emphasis id="72BB1D6962199A2F8B6CF2BCD512F860" bold="true" box="[518,533,1945,1969]" pageId="19" pageNumber="170">F</emphasis>
, female from Sicily, Italy.
<emphasis id="72BB1D6962199A2F8A46F2BCD447F860" bold="true" box="[812,832,1945,1969]" pageId="19" pageNumber="170">H</emphasis>
, female from Azerbaijan.
<emphasis id="72BB1D6962199A2F8D25F2BDD37EF861" bold="true" box="[1103,1145,1944,1968]" pageId="19" pageNumber="170">I–O</emphasis>
,
<taxonomicName id="87CFBAF862199A2F8DEDF2BDD7FAF804" authority="(de Saussure)" authorityName="UZ Uchkulach" baseAuthorityName="de Saussure" baseAuthorityYear="1853" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="fastidiosus">
<emphasis id="72BB1D6962199A2F8DEDF2BDD273F861" box="[1159,1396,1944,1968]" italics="true" pageId="19" pageNumber="170">Euodynerus fastidiosus</emphasis>
(de Saussure)
</taxonomicName>
.
<emphasis id="72BB1D6962199A2F8862F298D615F804" bold="true" box="[264,274,1981,2005]" pageId="19" pageNumber="170">I</emphasis>
,
<emphasis id="72BB1D6962199A2F884AF298D636F804" bold="true" box="[288,305,1981,2005]" pageId="19" pageNumber="170">L</emphasis>
, male from Italy.
<emphasis id="72BB1D6962199A2F8884F298D6FCF804" bold="true" box="[494,507,1981,2005]" pageId="19" pageNumber="170">J</emphasis>
, male from Greece.
<emphasis id="72BB1D6962199A2F8BBBF298D5E2F804" bold="true" box="[721,741,1981,2005]" pageId="19" pageNumber="170">K</emphasis>
, female from Crete, Greece.
<emphasis id="72BB1D6962199A2F8D7FF298D32AF804" bold="true" box="[1045,1069,1981,2005]" pageId="19" pageNumber="170">M</emphasis>
, female from Italy.
<emphasis id="72BB1D6962199A2F8D97F298D208F804" bold="true" box="[1277,1295,1981,2005]" pageId="19" pageNumber="170">N</emphasis>
, female from Crimea, Russia.
<emphasis id="72BB1D6962199A2F8855F2C5D654F829" bold="true" box="[319,339,2016,2040]" pageId="19" pageNumber="170">O</emphasis>
, female from Dagestan, Russia.
</paragraph>
</caption>
<caption id="14B091F3621E9A2889FDF2B2D5B3F826" ID-DOI="http://doi.org/10.5281/zenodo.14239455" ID-Zenodo-Dep="14239455" httpUri="https://zenodo.org/record/14239455/files/figure.png" pageId="20" pageNumber="171" startId="20.[151,250,1943,1967]" targetBox="[200,1386,192,1918]" targetPageId="20" targetType="figure">
<paragraph id="4070C17B621E9A2889FDF2B2D5B3F826" blockId="20.[151,1436,1943,2039]" pageId="20" pageNumber="171">
<emphasis id="72BB1D69621E9A2889FDF2B2D641F87E" bold="true" box="[151,326,1943,1967]" pageId="20" pageNumber="171">FIGURE 8. A–F</emphasis>
,
<taxonomicName id="87CFBAF8621E9A288838F2B2D5B0F87E" authority="Bluthgen. A" authorityName="Bluthgen. A" authorityYear="1942" box="[338,695,1943,1967]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="hellenicus">
<emphasis id="72BB1D69621E9A288838F2B2D53FF87E" box="[338,568,1943,1967]" italics="true" pageId="20" pageNumber="171">Euodynerus hellenicus</emphasis>
Blüthgen.
<emphasis id="72BB1D69621E9A288BCFF2B2D5B0F87E" bold="true" box="[677,695,1943,1967]" pageId="20" pageNumber="171">A</emphasis>
</taxonomicName>
, male from Bulgaria.
<emphasis id="72BB1D69621E9A288AFEF2B2D4A2F87E" bold="true" box="[916,933,1943,1967]" pageId="20" pageNumber="171">B</emphasis>
,
<emphasis id="72BB1D69621E9A288ADBF2B2D4C5F87E" bold="true" box="[945,962,1943,1967]" pageId="20" pageNumber="171">E</emphasis>
, female from Spain.
<emphasis id="72BB1D69621E9A288DFEF2B2D3A1F87E" bold="true" box="[1172,1190,1943,1967]" pageId="20" pageNumber="171">C</emphasis>
,
<emphasis id="72BB1D69621E9A288DD8F2B2D3C6F87E" bold="true" box="[1202,1217,1943,1967]" pageId="20" pageNumber="171">F</emphasis>
, female from Iran.
<emphasis id="72BB1D69621E9A288CEEF2B2D291F87E" bold="true" box="[1412,1430,1943,1967]" pageId="20" pageNumber="171">D</emphasis>
, male from Spain.
<emphasis id="72BB1D69621E9A288821F29ED683F802" bold="true" box="[331,388,1979,2003]" pageId="20" pageNumber="171">G–M</emphasis>
,
<taxonomicName id="87CFBAF8621E9A2888FAF29ED435F802" authority="(Dalla Torre)" authorityName="UZ Kokand Sands" baseAuthorityName="Dalla Torre" baseAuthorityYear="1889" box="[400,818,1979,2003]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="semisaecularis">
<emphasis id="72BB1D69621E9A2888FAF29ED5A1F802" box="[400,678,1979,2003]" italics="true" pageId="20" pageNumber="171">Euodynerus semisaecularis</emphasis>
(Dalla Torre)
</taxonomicName>
.
<emphasis id="72BB1D69621E9A288A51F29ED448F802" bold="true" box="[827,847,1979,2003]" pageId="20" pageNumber="171">G</emphasis>
,
<emphasis id="72BB1D69621E9A288A31F29ED46FF802" bold="true" box="[859,872,1979,2003]" pageId="20" pageNumber="171">J</emphasis>
, male from Greece.
<emphasis id="72BB1D69621E9A288D5CF29ED34DF802" bold="true" box="[1078,1098,1979,2003]" pageId="20" pageNumber="171">H</emphasis>
,
<emphasis id="72BB1D69621E9A288D3CF29ED360F802" bold="true" box="[1110,1127,1979,2003]" pageId="20" pageNumber="171">L</emphasis>
, female from Crete, Greece.
<emphasis id="72BB1D69621E9A288CE6F29ED291F802" bold="true" box="[1420,1430,1979,2003]" pageId="20" pageNumber="171">I</emphasis>
,
<emphasis id="72BB1D69621E9A2889FDF2FAD7A8F826" bold="true" box="[151,175,2015,2039]" pageId="20" pageNumber="171">M</emphasis>
, female from Afghanistan.
<emphasis id="72BB1D69621E9A2888ACF2FAD6DDF826" bold="true" box="[454,474,2015,2039]" pageId="20" pageNumber="171">K</emphasis>
, male from Armenia.
</paragraph>
</caption>
<paragraph id="4070C17B621F9A2989ADF5B2D5A2FECC" blockId="21.[151,1437,151,285]" pageId="21" pageNumber="172">
<emphasis id="72BB1D69621F9A2989ADF5B2D616FF60" bold="true" box="[199,273,151,177]" pageId="21" pageNumber="172">Notes.</emphasis>
This species shows a disjunct distribution, being known from the Iberian Peninsula in the west and from an area ranging from the Balkan Peninsula to
<collectingCountry id="38D881EB621F9A298B87F59ED471FF04" box="[749,886,187,213]" name="Afghanistan" pageId="21" pageNumber="172">Afghanistan</collectingCountry>
in the east, with a gap in the Italian Peninsula. Morphological comparison and genetic data confirm the conspecificity of the two populations, as no morphological and genetic differences were observed (
<figureCitation id="D8F4DDFE621F9A298B39F426D590FECC" box="[595,663,259,285]" captionStart="FIGURE 3" captionStartId="12.[151,250,1320,1344]" captionTargetBox="[159,1430,194,1283]" captionTargetId="figure-20@12.[151,1436,181,1296]" captionTargetPageId="12" captionText="FIGURE 3. Phylogenetic tree (Neighbor-Joining) of Euodynerus (Euodynerus) based on the mitochondrial COI gene. Ancistrocerus gazella (Panzer) was used as an outgroup. Numbers adjacent to branches represent bootstrap support (values of &lt;75 are omitted); only the species occurring in the considered area are highlighted. Countries are abbreviated using ISO 3166-1 alpha-2 codes. Continuation of Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.14239445" httpUri="https://zenodo.org/record/14239445/files/figure.png" pageId="21" pageNumber="172">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="1B387617621D9A2B89FDF7ABD416FD79" bold="true" box="[151,785,654,681]" fontSize="11" level="1" pageId="23" pageNumber="174" reason="1">
<taxonomicName id="87CFBAF8621D9A2B89FDF7ABD416FD79" ID-CoL="3CGMT" authority="Bluthgen, 1951" authorityName="Bluthgen. I" authorityYear="1951" box="[151,785,654,681]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="velutinus" subGenus="Euodynerus">
<emphasis id="72BB1D69621D9A2B89FDF7ABD416FD79" bold="true" box="[151,785,654,681]" pageId="23" pageNumber="174">
<emphasis id="72BB1D69621D9A2B89FDF7ABD62DFD78" bold="true" box="[151,298,654,681]" italics="true" pageId="23" pageNumber="174">Euodynerus</emphasis>
(
<emphasis id="72BB1D69621D9A2B8851F7ABD6C9FD78" bold="true" box="[315,462,654,681]" italics="true" pageId="23" pageNumber="174">Euodynerus</emphasis>
)
<emphasis id="72BB1D69621D9A2B88B5F7ABD549FD78" bold="true" box="[479,590,654,681]" italics="true" pageId="23" pageNumber="174">velutinus</emphasis>
Blüthgen, 1951
</emphasis>
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<subSubSection id="08D592F0621D9A2B89FDF796D68DFD1C" box="[151,394,691,717]" pageId="23" pageNumber="174" type="description">
<paragraph id="4070C17B621D9A2B89FDF796D68DFD1C" blockId="23.[151,785,654,717]" box="[151,394,691,717]" pageId="23" pageNumber="174">
(
<figureCitation id="D8F4DDFE621D9A2B89F5F796D61EFD1C" box="[159,281,691,717]" captionStart="FIGURE 9" captionStartId="22.[151,250,1908,1932]" captionTargetBox="[177,1411,192,1883]" captionTargetId="figure-20@22.[177,1411,181,1883]" captionTargetPageId="22" captionText="FIGURE 9. A–H, Euodynerus variegatus (Fabricius). A, E–F, female from Morocco. B, H, female from Egypt (E. variegatus kruegeri (von Schulthess), syn. nov.). C, G, female from Israel (E. variegatus kruegeri (von Schulthess), syn. nov.). D, male from Sardinia, Italy. I–M, Euodynerus velutinus Blüthgen. I, male from Azerbaijan, J, L, male from Greece. K, female from Greece. M, female from Dagestan, Russia." figureDoi="http://doi.org/10.5281/zenodo.14239457" httpUri="https://zenodo.org/record/14239457/files/figure.png" pageId="23" pageNumber="174">Figs 9I–M</figureCitation>
;
<figureCitation id="D8F4DDFE621D9A2B884FF796D649FD1C" box="[293,334,691,717]" captionStart="FIGURE 14" captionStartId="30.[151,250,1557,1581]" captionTargetBox="[153,1423,182,1517]" captionTargetId="figure-20@30.[152,1436,181,1533]" captionTargetPageId="30" captionText="FIGURE 14. Male genitalia, from left to right: aedeagus in ventral view, aedeagus in lateral view, digitus+volsella. A, Euodynerus bidentatus (Lepeletier). B, E. caspicus (Morawitz). C, E. curictensis Blüthgen. D, E. dantici (Rossi). E, E. disconotatus (Lichtenstein). F, E. fastidiosus (de Saussure). G, E. hellenicus Blüthgen. H, E. semisaecularis (Dalla Torre). I, E. variegatus (Fabricius). J, E. velutinus Blüthgen. K, E. bidentoides (Giordani Soika). L, E. notatus (Jurine). M, E. posticus (Herrich-Schaeffer). N, E. quadrifasciatus (Fabricius)." figureDoi="http://doi.org/10.5281/zenodo.14239467" httpUri="https://zenodo.org/record/14239467/files/figure.png" pageId="23" pageNumber="174">14J</figureCitation>
;
<figureCitation id="D8F4DDFE621D9A2B8830F796D685FD1C" box="[346,386,691,717]" captionStart="FIGURE 16" captionStartId="32.[151,250,995,1019]" captionTargetBox="[151,1435,181,971]" captionTargetId="figure-20@32.[151,1435,181,971]" captionTargetPageId="32" captionText="FIGURE 16. A–B, male mandible in frontal view (arrowhead indicating normal or modified tooth). C–G, tegula in dorsal view. H–I, apex of T2 in dorsal view (arrowhead indicating lamellar margin). J–K, male hind femur in frontal view (arrowhead in J indicating basal expansion). L–N, apex of male metasoma in lateral view. A, Euodynerus notatus (Jurine). B, E. posticus (Herrich-Schaeffer). C, E. caspicus (Morawitz). D, H, E. hellenicus Blüthgen. E, I, E. dantici (Rossi). F, L, E. fastidiosus (de Saussure). G, N, E. disconotatus (Lichtenstein). J, E. velutinus Blüthgen. K, E. curictensis Blüthgen. M, E. semisaecularis (Dalla Torre)." figureDoi="http://doi.org/10.5281/zenodo.14239471" httpUri="https://zenodo.org/record/14239471/files/figure.png" pageId="23" pageNumber="174">16J</figureCitation>
)
</paragraph>
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<taxonomicName id="87CFBAF8621D9A2B89FDF7D2D6FCFCDE" ID-CoL="3CGMT" authority="&quot; (Kostylev)" baseAuthorityName="Kostylev" box="[151,507,759,783]" class="Insecta" family="Eumenidae" genus="Euodynerus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="72BB1D69621D9A2B89FDF7D2D671FCDE" box="[151,374,759,783]" italics="true" pageId="23" pageNumber="174">Euodynerus velutinus</emphasis>
“(Kostylev)
</taxonomicName>
”
<treatmentCitation id="C16EE76A621D9A2B8B7AF7D2D5F0FCDE" author="Bluthgen, P." box="[528,759,759,783]" page="172" pageId="23" pageNumber="174" year="1951">
<bibRefCitation id="245EBC8A621D9A2B8B7AF7D2D5F0FCDE" author="Bluthgen, P." box="[528,759,759,783]" pageId="23" pageNumber="174" pagination="166 - 201" refId="ref21879" refString="Bluthgen, P. (1951 a) Neue oder bemerkenswerte palaarktische Faltenwespen aus der Zoologischen Staatssammlung in Munchen (Hym. Vespidae, Eumenidinae). Mitteilungen der Munchner Entomologischen Gesellschaft, 41, 166 - 201." type="journal article" year="1951">Blüthgen, 1951a: 172</bibRefCitation>
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, 195, footnote,
<specimenCount id="56C90AF2621D9A2B8ACEF7DDD4B0FCDE" box="[932,951,760,783]" count="1" pageId="23" pageNumber="174" type="female">♀</specimenCount>
,
<specimenCount id="56C90AF2621D9A2B8AA2F7DDD4DCFCDE" box="[968,987,760,783]" count="1" pageId="23" pageNumber="174" type="male">♂</specimenCount>
<emphasis id="72BB1D69621D9A2B8AB1F7DDD4F5FCDE" box="[987,1010,760,783]" italics="true" pageId="23" pageNumber="174">—</emphasis>
“Süd-Morea;
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<collectingRegion id="820B0F99621D9A2B8DEBF7D2D3FEFCDE" box="[1153,1273,759,783]" country="Greece" name="Peloponnisos" pageId="23" pageNumber="174">Peloponnes</collectingRegion>
;
<collectingRegion id="820B0F99621D9A2B8C6DF7D2D24EFCDE" box="[1287,1353,759,783]" country="Greece" name="Attiki" pageId="23" pageNumber="174">Attika</collectingRegion>
” (
<collectionCode id="26DE59BE621D9A2B8C0EF7D2D29BFCDE" box="[1380,1436,759,783]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="23" pageNumber="174" type="Museum">ZMB</collectionCode>
[examined])
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.
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</paragraph>
</subSubSection>
<subSubSection id="08D592F0621D9A2B89FDF679D618FC4B" pageId="23" pageNumber="174" type="distribution">
<paragraph id="4070C17B621D9A2B89FDF679D618FC4B" blockId="23.[151,1436,860,922]" pageId="23" pageNumber="174">
<emphasis id="72BB1D69621D9A2B89FDF679D636FCA7" bold="true" box="[151,305,860,886]" pageId="23" pageNumber="174">Distribution.</emphasis>
From the
<collectingRegion id="820B0F99621D9A2B88F5F679D6E9FCA7" box="[415,494,860,886]" country="Turkmenistan" name="Balkan Welayaty" pageId="23" pageNumber="174">Balkan</collectingRegion>
Peninsula in the west to the Crimean Peninsula and the Caucasus in the east (
<bibRefCitation id="245EBC8A621D9A2B8C52F679D615FC4B" author="Fateryga, A. V. &amp; Proshchalykin, M. Yu. &amp; Maharramov, M. M. &amp; Astafurova, Yu. V." pageId="23" pageNumber="174" pagination="36 - 60" refId="ref23016" refString="Fateryga, A. V., Proshchalykin, M. Yu., Maharramov, M. M. &amp; Astafurova, Yu. V. (2021) New records of solitary vespid wasps (Hymenoptera: Vespidae: Masarinae and Eumeninae s. l.) from the Nakhchivan Autonomous Republic of Azerbaijan. Zootaxa, 5027 (1), 36 - 60. https: // doi. org / 10.11646 / zootaxa. 5027.1.2" type="journal article" year="2021">
Fateryga
<emphasis id="72BB1D69621D9A2B89FDF6A4D7C8FC4B" box="[151,207,896,922]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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