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<document id="E785CB043389B39BA4E0A6DE2AAD20D1" ID-CLB-Dataset="305703" ID-DOI="10.11646/zootaxa.5538.6.3" ID-GBIF-Dataset="2f227900-76db-4160-86e5-8b77beb14cd0" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240174" ID-ZooBank="87FF5517-A0EF-4A3A-8FF3-B9F6EEBE715B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732830837656" checkinUser="plazi" docAuthor="Röll, Beate, Pinto, Pedro Vaz &amp; Lobón-Rovira, Javier" docDate="2024" docId="BA0B5D5F52764B761ED7945367FCA14A" docLanguage="en" docName="zootaxa.5538.6.3.pdf" docOrigin="Zootaxa 5538 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5538.6.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lygodactylus" docType="treatment" docVersion="4" lastPageNumber="571" masterDocId="46322527527F4B7C1E4092206439A377" masterDocTitle="A new species of African diurnal dwarf geckos (Gekkonidae: Lygodactylus) from the Lower Guinea rainforest" masterLastPageNumber="574" masterPageNumber="561" pageNumber="570" updateTime="1732924349047" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="76C95AF2941F7B45BB849D4691B5FCFC">A new species of African diurnal dwarf geckos (Gekkonidae: Lygodactylus) from the Lower Guinea rainforest</mods:title>
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<mods:namePart id="B44E81A1F7E497DFDFF6326227E1A9C3">Röll, Beate</mods:namePart>
<mods:nameIdentifier id="A59890E6C2E5AE18FFA569A94320E59A" type="ORCID">0000-0003-4445-9019</mods:nameIdentifier>
<mods:affiliation id="137DD2C589B0EDE4F36DD168B65171A8">Department of General Zoology and Neurobiology, Ruhr-University Bochum, D- 44780 Bochum, Germany. &amp; Institute of Zoology, University of Veterinary Medicine Hannover, Bünteweg 17, D- 30559 Hannover, Germany.</mods:affiliation>
<mods:nameIdentifier id="7D23E15CE4B1F027D10F0CC85CE77011" type="email">beate.roell@rub.de</mods:nameIdentifier>
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<mods:namePart id="E9E3BC3C4C9244B267973A421A23FC54">Pinto, Pedro Vaz</mods:namePart>
<mods:nameIdentifier id="7E21F73FB3562438A4BFB712AA7B0DA8" type="ORCID">0000-0003-4487-1310</mods:nameIdentifier>
<mods:affiliation id="CE96FCCF9E97F0C79A85B6EC2161E2E6">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal. &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal. &amp; TwinLab CIBIO / ISCED, Instituto Superior de Ciências da Educação da Huíla, Rua Sarmento Rodrigues s / n, Lubango, Angola. &amp; Fundação Kissama, Rua 60 Casa 560, Lar do Patriota, Luanda, Angola.</mods:affiliation>
<mods:nameIdentifier id="2D7105EDBCABEA440CDEF31D9CAF5B36" type="email">pedrovazpinto@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CC2730201E5CE6BBEA2C478917801F49">Lobón-Rovira, Javier</mods:namePart>
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<mods:affiliation id="65E443B1CF188618497001EBC49FD773">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal. &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal.</mods:affiliation>
<mods:nameIdentifier id="ED92D69246DD3D65716995289D3EB7DE" type="email">j.lobon.rovira@cibio.up.pt</mods:nameIdentifier>
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<treatment id="BA0B5D5F52764B761ED7945367FCA14A" ID-DOI="http://doi.org/10.5281/zenodo.14248769" ID-GBIF-Taxon="248070712" ID-Zenodo-Dep="14248769" LSID="urn:lsid:plazi:treatment:BA0B5D5F52764B761ED7945367FCA14A" httpUri="http://treatment.plazi.org/id/BA0B5D5F52764B761ED7945367FCA14A" lastPageId="10" lastPageNumber="571" pageId="9" pageNumber="570" scope_family="Gekkonidae" scope_infraClass="Lower">
<subSubSection id="7AB8BFC252764B751ED7945365D8A5F9" box="[151,481,1651,1678]" pageId="9" pageNumber="570" type="nomenclature">
<paragraph id="321DEC4952764B751ED7945365D8A5F9" blockId="9.[151,1437,1651,2038]" box="[151,481,1651,1678]" pageId="9" pageNumber="570">
<heading id="69555B2552764B751ED7945365D8A5F9" bold="true" box="[151,481,1651,1678]" fontSize="11" level="1" pageId="9" pageNumber="570" reason="1">
<emphasis id="00D6305B52764B751ED7945365D8A5F9" bold="true" box="[151,481,1651,1678]" pageId="9" pageNumber="570">
<taxonomicName id="F5A297CA52764B751ED794536517A5F9" ID-CoL="5HP8" authority="Gray" box="[151,302,1651,1678]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="genus">
<emphasis id="00D6305B52764B751ED794536517A5F9" bold="true" box="[151,302,1651,1678]" italics="true" pageId="9" pageNumber="570">Lygodactylus</emphasis>
</taxonomicName>
species groups
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7AB8BFC252764B761E8794B767FCA14A" lastPageId="10" lastPageNumber="571" pageId="9" pageNumber="570" type="discussion">
<paragraph id="321DEC4952764B751E8794B7667EA46A" blockId="9.[151,1437,1651,2038]" pageId="9" pageNumber="570">
The genus
<taxonomicName id="F5A297CA52764B751F0B94B765E6A5C6" box="[331,479,1687,1713]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="genus">
<emphasis id="00D6305B52764B751F0B94B765E6A5C6" box="[331,479,1687,1713]" italics="true" pageId="9" pageNumber="570">Lygodactylus</emphasis>
</taxonomicName>
is generally arranged into different groups, based on a first subdivision provided by
<bibRefCitation id="563391B852764B751EFE949C655FA5A1" author="Pasteur, G." box="[190,358,1723,1750]" pageId="9" pageNumber="570" pagination="1 - 132" refId="ref9476" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards afromalgaches actuels. Travaux de l'Institut Scientifique Cherifien, Serie Zoologie, 29, 1 - 132." type="book chapter" year="1965">Pasteur (1965)</bibRefCitation>
and modified by subsequent authors. The
<taxonomicName id="F5A297CA52764B751D1D949C67DEA5A2" authorityName="Sternfeld" authorityYear="1912" box="[861,999,1723,1749]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52764B751D1D949C67DEA5A2" box="[861,999,1723,1749]" italics="true" pageId="9" pageNumber="570">L. scheffleri</emphasis>
</taxonomicName>
-group is actually composed of seven species, six of them occurring eastward of the Albertine Rift Valley in East Africa, whereas
<taxonomicName id="F5A297CA52764B751AC994C060D6A58E" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1161,1263,1759,1785]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751AC994C060D6A58E" box="[1161,1263,1759,1785]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52764B751AB594C06174A58D" bold="true" box="[1269,1357,1760,1786]" pageId="9" pageNumber="570">
<taxonomicNameLabel id="1BE58D2052764B751AB594C06174A58D" box="[1269,1357,1760,1786]" pageId="9" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs westward in the Lower
<collectingCountry id="4AB5ACD952764B751FE0952365C8A46A" box="[416,497,1795,1821]" name="Guinea-Bissau" pageId="9" pageNumber="570">Guinea</collectingCountry>
region.
</paragraph>
<paragraph id="321DEC4952764B751E8795076018A4A5" blockId="9.[151,1437,1651,2038]" pageId="9" pageNumber="570">
Morphologically,
<taxonomicName id="F5A297CA52764B751FD3950865C4A436" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[403,509,1831,1857]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751FD3950865C4A436" box="[403,509,1831,1857]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52764B751C479508665DA435" bold="true" box="[519,612,1832,1858]" pageId="9" pageNumber="570">
<taxonomicNameLabel id="1BE58D2052764B751C479508665DA435" box="[519,612,1832,1858]" pageId="9" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares two typical scale characters of the group, namely a double row of transversely enlarged subcaudal scales and a semi-divided mental scale, with all other members. However, it differs both in other morphological characters and in coloration. Furthermore,
<taxonomicName id="F5A297CA52764B751A4E9550604FA4FE" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1038,1142,1903,1929]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751A4E9550604FA4FE" box="[1038,1142,1903,1929]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52764B751A3E955060E1A4FD" bold="true" box="[1150,1240,1904,1930]" pageId="9" pageNumber="570">
<taxonomicNameLabel id="1BE58D2052764B751A3E955060E1A4FD" box="[1150,1240,1904,1930]" pageId="9" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs greatly in
<emphasis id="00D6305B52764B751ED795B364F8A4DA" box="[151,193,1939,1965]" italics="true" pageId="9" pageNumber="570">16S</emphasis>
uncorrected pairwise distance (&gt; 10%) from
<taxonomicName id="F5A297CA52764B751CFD95B467B3A4DA" authorityName="Pasteur" authorityYear="1964" box="[701,906,1939,1965]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="laterimaculatus">
<emphasis id="00D6305B52764B751CFD95B467B3A4DA" box="[701,906,1939,1965]" italics="true" pageId="9" pageNumber="570">L. laterimaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F5A297CA52764B751D8195B4601DA4DA" authorityName="Pasteur" authorityYear="1965" box="[961,1060,1940,1965]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="gravis">
<emphasis id="00D6305B52764B751D8195B4601DA4DA" box="[961,1060,1940,1965]" italics="true" pageId="9" pageNumber="570">L. gravis</emphasis>
</taxonomicName>
, well above the species threshold proposed by
<bibRefCitation id="563391B852764B751F689597662DA4A5" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." box="[296,532,1975,2002]" pageId="9" pageNumber="570" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B52764B751FCB959865FAA4A6" box="[395,451,1975,2001]" italics="true" pageId="9" pageNumber="570">et al.</emphasis>
(2021)
</bibRefCitation>
and
<bibRefCitation id="563391B852764B751C0B95976743A4A5" author="Lobon-Rovira, J. &amp; Bauer, A. M. &amp; Vaz Pinto, P. &amp; Trape, J. F. &amp; Conradie, W. &amp; Kusamba, C. &amp; Julio, T. &amp; Cael, G. &amp; Stanley, E. L. &amp; Huges, D. F. &amp; Behangana, M. &amp; Masudi, F. M. &amp; Pauwels, O. S. G. &amp; Greenbaum, E." box="[587,890,1975,2002]" pageId="9" pageNumber="570" pagination="447 - 492" refId="ref8783" refString="Lobon-Rovira, J., Bauer, A. M., Vaz Pinto, P., Trape, J. F., Conradie, W., Kusamba, C., Julio, T., Cael, G., Stanley, E. L., Huges, D. F., Behangana, M., Masudi, F. M., Pauwels, O. S. G. &amp; Greenbaum, E. (2023) Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history. Zoological Journal of the Linnean Society, 201 (2), 447 - 492. [published with pagination in 2024] https: // doi. org / 10.1093 / zoolinnean / zlad 123" type="journal article" year="2023">
Lobón-Rovira
<emphasis id="00D6305B52764B751CB095986710A4A6" box="[752,809,1975,2001]" italics="true" pageId="9" pageNumber="570">et al.</emphasis>
(2023)
</bibRefCitation>
for this genus.
</paragraph>
<paragraph id="321DEC4952764B761E8795FB614EA2DA" blockId="9.[151,1437,1651,2038]" lastBlockId="10.[151,1437,151,573]" lastPageId="10" lastPageNumber="571" pageId="9" pageNumber="570">
Interestingly, the
<taxonomicName id="F5A297CA52764B751FCC95FC662AA482" authorityName="Sternfeld" authorityYear="1912" box="[396,531,2011,2037]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52764B751FCC95FC662AA482" box="[396,531,2011,2037]" italics="true" pageId="9" pageNumber="570">L. scheffleri</emphasis>
</taxonomicName>
-group shares the double row of enlarged subcaudals with
<taxonomicName id="F5A297CA52764B751AEB95FC6127A482" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1195,1310,2011,2037]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52764B751AEB95FC6127A482" box="[1195,1310,2011,2037]" italics="true" pageId="9" pageNumber="570">L. fischeri</emphasis>
</taxonomicName>
of western and central Africa, a species sometimes confused with
<taxonomicName id="F5A297CA52754B761CBA92B76746A3C6" authorityName="Sternfeld" authorityYear="1912" box="[762,895,151,177]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52754B761CBA92B76746A3C6" box="[762,895,151,177]" italics="true" pageId="10" pageNumber="571">L. scheffleri</emphasis>
</taxonomicName>
and
<taxonomicName id="F5A297CA52754B761DF592B7600BA3C6" authorityName="Matschie" authorityYear="1892" box="[949,1074,151,177]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="conradti">
<emphasis id="00D6305B52754B761DF592B7600BA3C6" box="[949,1074,151,177]" italics="true" pageId="10" pageNumber="571">L. conradti</emphasis>
</taxonomicName>
. However,
<taxonomicName id="F5A297CA52754B761AF292B7611DA3C6" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1202,1316,151,177]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761AF292B7611DA3C6" box="[1202,1316,151,177]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
is actually a member of a different group (the
<taxonomicName id="F5A297CA52754B761C69929B66A5A3A2" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[553,668,187,213]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761C69929B66A5A3A2" box="[553,668,187,213]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
-group), which originally contained two other West African species, namely
<taxonomicName id="F5A297CA52754B761EB292FF65F8A38E" authority="Tornier" authorityName="Tornier" authorityYear="1902" box="[242,449,223,249]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="conraui">
<emphasis id="00D6305B52754B761EB292FF655EA38F" box="[242,359,223,248]" italics="true" pageId="10" pageNumber="571">L. conraui</emphasis>
Tornier
</taxonomicName>
and
<taxonomicName id="F5A297CA52754B761FB992FF66D3A38E" authority="(Peters)" baseAuthorityName="Peters" baseAuthorityYear="1881" box="[505,746,223,249]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="thomensis">
<emphasis id="00D6305B52754B761FB992FF66B1A38E" box="[505,648,223,249]" italics="true" pageId="10" pageNumber="571">L. thomensis</emphasis>
(Peters)
</taxonomicName>
.
<bibRefCitation id="563391B852754B761CB692FF67A2A38E" author="Pasteur, G." box="[758,923,222,249]" pageId="10" pageNumber="571" pagination="1 - 132" refId="ref9476" refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards afromalgaches actuels. Travaux de l'Institut Scientifique Cherifien, Serie Zoologie, 29, 1 - 132." type="book chapter" year="1965">Pasteur (1965)</bibRefCitation>
erected this group as the only member of his ‘phylum guinéen’ due to the distribution of these species in western Africa. However, in doing so he deliberately ignored that the members of this group are inconsistent in typical scale characters: while
<taxonomicName id="F5A297CA52754B761A2B930760E5A236" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1131,1244,295,321]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761A2B930760E5A236" box="[1131,1244,295,321]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
has a double row of enlarged subcaudals and a semi-divided mental, both
<taxonomicName id="F5A297CA52754B761D45936B6740A213" authorityName="Tornier" authorityYear="1902" box="[773,889,331,356]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="conraui">
<emphasis id="00D6305B52754B761D45936B6740A213" box="[773,889,331,356]" italics="true" pageId="10" pageNumber="571">L. conraui</emphasis>
</taxonomicName>
and
<taxonomicName id="F5A297CA52754B761DED936B6003A212" baseAuthorityName="Peters" baseAuthorityYear="1881" box="[941,1082,331,357]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="thomensis">
<emphasis id="00D6305B52754B761DED936B6003A212" box="[941,1082,331,357]" italics="true" pageId="10" pageNumber="571">L. thomensis</emphasis>
</taxonomicName>
have only one median row with strongly enlarged subcaudals and a large, undivided mental. Judging from these scale characters,
<taxonomicName id="F5A297CA52754B761A97934F6171A2FE" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1239,1352,367,393]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761A97934F6171A2FE" box="[1239,1352,367,393]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
fits the species of the
<emphasis id="00D6305B52754B761F7793B365FFA2DA" box="[311,454,403,429]" italics="true" pageId="10" pageNumber="571">L. scheffleri-</emphasis>
group much better. Indeed, it would seem justified to split up the ‘
<taxonomicName id="F5A297CA52754B761AEA93B36125A2DA" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1194,1308,403,429]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761AEA93B36125A2DA" box="[1194,1308,403,429]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
’-group.
</paragraph>
<paragraph id="321DEC4952754B761E87939767FCA14A" blockId="10.[151,1437,151,573]" pageId="10" pageNumber="571">
To resolve the crucial points discussed above, a revision of the
<taxonomicName id="F5A297CA52754B761DD893976026A2A6" authorityName="Sternfeld" authorityYear="1912" box="[920,1055,439,465]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52754B761DD893976026A2A6" box="[920,1055,439,465]" italics="true" pageId="10" pageNumber="571">L. scheffleri</emphasis>
</taxonomicName>
-group is necessary. To date, only three members of the
<taxonomicName id="F5A297CA52754B761FCD93FB662AA282" authorityName="Sternfeld" authorityYear="1912" box="[397,531,475,501]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52754B761FCD93FB662AA282" box="[397,531,475,501]" italics="true" pageId="10" pageNumber="571">L. scheffleri</emphasis>
</taxonomicName>
-group (
<emphasis id="00D6305B52754B761C2F93FB6796A282" box="[623,943,475,501]" italics="true" pageId="10" pageNumber="571">
<taxonomicName id="F5A297CA52754B761C2F93FB66EFA283" authorityName="Pasteur" authorityYear="1965" box="[623,726,475,500]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="gravis">L. gravis</taxonomicName>
,
<taxonomicName id="F5A297CA52754B761CA193FB6796A282" authorityName="Pasteur" authorityYear="1964" box="[737,943,475,501]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="laterimaculatus">L. laterimaculatus</taxonomicName>
</emphasis>
and
<taxonomicName id="F5A297CA52754B761DA793FB6077A282" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[999,1102,475,501]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52754B761DA793FB6077A282" box="[999,1102,475,501]" italics="true" pageId="10" pageNumber="571">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52754B761A1593FB6096A282" bold="true" box="[1109,1199,475,501]" pageId="10" pageNumber="571">
<taxonomicNameLabel id="1BE58D2052754B761A1593FB6096A282" box="[1109,1199,475,501]" pageId="10" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) have been analyzed genetically. For all other members of the
<taxonomicName id="F5A297CA52754B761CCC93DF672EA16E" authorityName="Sternfeld" authorityYear="1912" box="[652,791,511,537]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52754B761CCC93DF672EA16E" box="[652,791,511,537]" italics="true" pageId="10" pageNumber="571">L. scheffleri</emphasis>
</taxonomicName>
-group, including its candidate member—
<taxonomicName id="F5A297CA52754B761B4293DF6140A16E" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[1282,1401,511,537]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="10" pageNumber="571" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B52754B761B4293DF6140A16E" box="[1282,1401,511,537]" italics="true" pageId="10" pageNumber="571">L. fischeri</emphasis>
</taxonomicName>
—, sequence data are not yet available, but are most desirable for a revision.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="00D6305B527C4B7F1ED7976B65B2A611" bold="true" box="[151,395,1355,1382]" italics="true" pageId="3" pageNumber="564">Lygodactylus lobeke</emphasis>
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(
<figureCitation id="AA99F0CC527C4B7F1EDF974F6535A6FE" box="[159,268,1391,1417]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1933,1957]" captionStartId-1="5.[151,250,1755,1779]" captionStartId-2="6.[151,250,1844,1868]" captionTargetBox-0="[151,1435,1281,1910]" captionTargetBox-1="[151,1435,181,1732]" captionTargetBox-2="[151,1436,885,1821]" captionTargetId-0="figure-565@4.[151,1435,1281,1910]" captionTargetId-1="figure-20@5.[151,1435,181,1732]" captionTargetId-2="figure-350@6.[151,1436,885,1821]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2: Lygodactylus lobeke sp. nov. in life, shortly after capture. Female holotype: lateral (a) and dorsal views (b). Male paratype: lateral (c) and dorsal views (d). Photographs of holotype: ambient light, analog (Kodachrome 200) (2002), photographs of paratype: flash, digital (Canon EOS 6D Mark II) (2023). Photographs by B.R. (a, b) and P.VP. (c, d)." captionText-1="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." captionText-2="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi-0="http://doi.org/10.5281/zenodo.14240178" figureDoi-1="http://doi.org/10.5281/zenodo.14240180" figureDoi-2="http://doi.org/10.5281/zenodo.14240182" httpUri-0="https://zenodo.org/record/14240178/files/figure.png" httpUri-1="https://zenodo.org/record/14240180/files/figure.png" httpUri-2="https://zenodo.org/record/14240182/files/figure.png" pageId="3" pageNumber="564">Figs. 2–4</figureCitation>
,
<figureCitation id="AA99F0CC527C4B7F1F57974F651FA6FE" box="[279,294,1391,1417]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="3" pageNumber="564">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC2527C4B7F1ED7978E6609A692" pageId="3" pageNumber="564" type="reference_group">
<paragraph id="321DEC49527C4B7F1ED7978E6732A6B1" blockId="3.[151,779,1454,1509]" box="[151,779,1454,1478]" pageId="3" pageNumber="564">
<taxonomicName id="F5A297CA527C4B7F1ED7978E6561A6B1" box="[151,344,1454,1478]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="undefined-B">
<emphasis id="00D6305B527C4B7F1ED7978E6524A6B1" box="[151,285,1454,1478]" italics="true" pageId="3" pageNumber="564">Lygodactylus</emphasis>
sp. B
</taxonomicName>
(
<bibRefCitation id="563391B8527C4B7F1F25978E6631A6B1" author="Roll, B. &amp; Prohl, H. &amp; Hoffmann, K. - P." box="[357,520,1454,1478]" pageId="3" pageNumber="564" pagination="327 - 335" refId="ref9837" refString="Roll, B., Prohl, H. &amp; Hoffmann, K. - P. (2010) Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata, Gekkonidae). Molecular Phylogenetics and Evolution, 56 (1), 327 - 335. https: // doi. org / 10.1016 / j. ympev. 2010.02.002" type="journal article" year="2010">
Röll
<emphasis id="00D6305B527C4B7F1FD9978F65FCA6B1" box="[409,453,1454,1478]" italics="true" pageId="3" pageNumber="564">et al</emphasis>
. 2010
</bibRefCitation>
;
<bibRefCitation id="563391B8527C4B7F1C52978E673DA6B1" author="Mezzasalma, M. &amp; Andreone, F. &amp; Aprea, G. &amp; Glaw, F. &amp; Odierna, G. &amp; Guarino, F. M." box="[530,772,1454,1478]" pageId="3" pageNumber="564" pagination="42 - 54" refId="ref9202" refString="Mezzasalma, M., Andreone, F., Aprea, G., Glaw, F., Odierna, G. &amp; Guarino, F. M. (2017) Molecular phylogeny, biogeography and chromosome evolution of Malagasy geckos of the genus Lygodactylus (Squamata, Gekkonidae). Zoological Scripta, 46, 42 - 54. https: // doi. org / 10.1111 / zsc. 12188" type="journal article" year="2017">
Mezzasalma
<emphasis id="00D6305B527C4B7F1CD7978F66F3A6B1" box="[663,714,1454,1478]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2017
</bibRefCitation>
)
</paragraph>
<paragraph id="321DEC49527C4B7F1ED797ED6609A692" blockId="3.[151,779,1454,1509]" box="[151,560,1485,1509]" pageId="3" pageNumber="564">
<taxonomicName id="F5A297CA527C4B7F1ED797ED656DA692" box="[151,340,1485,1509]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="undefined-6">
<emphasis id="00D6305B527C4B7F1ED797ED6524A692" box="[151,285,1485,1509]" italics="true" pageId="3" pageNumber="564">Lygodactylus</emphasis>
sp. 6
</taxonomicName>
(
<bibRefCitation id="563391B8527C4B7F1F2197ED6610A692" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." box="[353,553,1485,1509]" pageId="3" pageNumber="564" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B527C4B7F1FFC97EE65D6A692" box="[444,495,1485,1509]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2021
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC2527C4B7F1ED7942E664FA5AC" pageId="3" pageNumber="564" type="materials_examined">
<paragraph id="321DEC49527C4B7F1ED7942E67A3A518" blockId="3.[151,1437,1550,2043]" pageId="3" pageNumber="564">
<materialsCitation id="82CAE614527C4B7F1ED7942E67A3A518" ID-GBIF-Occurrence="4986907301" collectingDate="2002-03-26" collectionCode="ZSM" collectorName="Beate Roll &amp; Alexander Pelzer &amp; Mathias Heinze." country="Cameroon" county="Region East" elevation="500" latitude="2.4557" location="Lobeke region" longLatPrecision="7" longitude="15.4203" municipality="Goumela" pageId="3" pageNumber="564" specimenCode="ZSM 59/2024" specimenCount="1" typeStatus="holotype">
<emphasis id="00D6305B527C4B7F1ED7942E6532A55F" bold="true" box="[151,267,1550,1576]" pageId="3" pageNumber="564">
<typeStatus id="ED1952EB527C4B7F1ED7942E653FA55F" box="[151,262,1550,1576]" pageId="3" pageNumber="564" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="62044432527C4B7F1F52942E6588A55E" box="[274,433,1550,1577]" pageId="3" pageNumber="564">ZSM 59/2024</specimenCode>
,
<specimenCount id="24A427C0527C4B7F1FFC942E6673A55F" box="[444,586,1550,1576]" pageId="3" pageNumber="564" type="adult">adult female</specimenCount>
, collected in the
<location id="377DBA92527C4B7F1D47942E6792A55F" LSID="urn:lsid:plazi:treatment:BA0B5D5F527C4B751ED7976B60F5A1BB:377DBA92527C4B7F1D47942E6792A55F" box="[775,939,1550,1576]" country="Cameroon" county="Region East" latitude="2.4557" longLatPrecision="7" longitude="15.4203" municipality="Goumela" name="Lobeke region" pageId="3" pageNumber="564">Lobeke region</location>
, south of
<collectingMunicipality id="D2797633527C4B7F1A59942E60B9A55F" box="[1049,1152,1550,1576]" pageId="3" pageNumber="564">Goumela</collectingMunicipality>
(Koumela),
<collectingCounty id="DB7C94C5527C4B7F1B4D942E61A1A55F" box="[1293,1432,1550,1576]" pageId="3" pageNumber="564">Region East</collectingCounty>
, south-eastern
<collectingCountry id="4AB5ACD9527C4B7F1F7494126595A53B" box="[308,428,1586,1612]" name="Cameroon" pageId="3" pageNumber="564">Cameroon</collectingCountry>
, at geographical coordinates
<geoCoordinate id="57968A8E527C4B7F1CB594116760A53B" box="[757,857,1585,1612]" degrees="2.4557" direction="north" orientation="latitude" pageId="3" pageNumber="564" precision="5" value="2.4557">N2.4557</geoCoordinate>
,
<geoCoordinate id="57968A8E527C4B7F1D25941167EDA53B" box="[869,980,1585,1612]" degrees="15.4203" direction="east" orientation="longitude" pageId="3" pageNumber="564" precision="5" value="15.4203">E15.4203</geoCoordinate>
, about
<elevation id="B98F0B7A527C4B7F1A669411611BA53B" box="[1062,1314,1585,1612]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="564" unit="m" value="500.0">
<quantity id="F55A41AC527C4B7F1A6694116057A53B" box="[1062,1134,1585,1612]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="564" unit="m" value="500.0">500 m</quantity>
above sea level
</elevation>
(a.s.l.), on
<date id="461CCA89527C4B7F1ED794756576A507" box="[151,335,1621,1648]" pageId="3" pageNumber="564" value="2002-03-26">
<collectingDate id="56583361527C4B7F1ED794756576A507" box="[151,335,1621,1648]" pageId="3" pageNumber="564" value="2002-03-26">
26
<superScript id="C5D74101527C4B7F1EF3947564F9A514" attach="left" box="[179,192,1621,1635]" fontSize="6" pageId="3" pageNumber="564">th</superScript>
March 2002
</collectingDate>
</date>
by
<collectorName id="9F57899F527C4B7F1F39947665CAA518" box="[377,499,1621,1648]" pageId="3" pageNumber="564">Beate Röll</collectorName>
,
<collectorName id="9F57899F527C4B7F1FBC94756687A518" box="[508,702,1621,1647]" pageId="3" pageNumber="564">Alexander Pelzer</collectorName>
&amp;
<collectorName id="9F57899F527C4B7F1CA2947567A3A518" box="[738,922,1621,1647]" pageId="3" pageNumber="564">Mathias Heinze.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="321DEC49527C4B7F1E879459664FA5AC" blockId="3.[151,1437,1550,2043]" pageId="3" pageNumber="564">
<materialsCitation id="82CAE614527C4B7F1E879459664AA5AC" ID-GBIF-Occurrence="4986907302" collectingDate="2023-06-06" collectionCode="FKH" collectorName="Pedro Vaz Pinto &amp; Michael Mills" country="Angola" elevation="221" latitude="-4.6253" location="Sanga Wanda" longLatPrecision="7" longitude="12.4583" municipality="Mayombe National Park" pageId="3" pageNumber="564" specimenCode="FKH 1259" specimenCount="1" stateProvince="Cabinda Province" typeStatus="paratype">
<emphasis id="00D6305B527C4B7F1E8794596502A5E4" bold="true" box="[199,315,1657,1683]" pageId="3" pageNumber="564">
<typeStatus id="ED1952EB527C4B7F1E879459650FA5E4" box="[199,310,1657,1683]" pageId="3" pageNumber="564" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="62044432527C4B7F1F06945A65FDA5E3" box="[326,452,1657,1684]" pageId="3" pageNumber="564">FKH 1259</specimenCode>
, probably subadult, but sexually mature male, collected in
<location id="377DBA92527C4B7F1A3D94596125A5E4" LSID="urn:lsid:plazi:treatment:BA0B5D5F527C4B751ED7976B60F5A1BB:377DBA92527C4B7F1A3D94596125A5E4" box="[1149,1308,1657,1683]" country="Angola" latitude="-4.6253" longLatPrecision="7" longitude="12.4583" municipality="Mayombe National Park" name="Sanga Wanda" pageId="3" pageNumber="564" stateProvince="Cabinda Province">Sanga Wanda</location>
,
<collectingMunicipality id="D2797633527C4B7F1B6B9459650CA5C0" pageId="3" pageNumber="564">Mayombe National Park</collectingMunicipality>
,
<collectingRegion id="F06622AB527C4B7F1F0094BD6633A5C0" box="[320,522,1693,1719]" country="Angola" name="Cabinda" pageId="3" pageNumber="564">Cabinda Province</collectingRegion>
,
<collectingCountry id="4AB5ACD9527C4B7F1C5494BD6650A5C0" box="[532,617,1693,1719]" name="Angola" pageId="3" pageNumber="564">Angola</collectingCountry>
, at geographical coordinates
<geoCoordinate id="57968A8E527C4B7F1DF094BD6036A5CF" box="[944,1039,1693,1720]" degrees="4.6253" direction="south" orientation="latitude" pageId="3" pageNumber="564" precision="5" value="-4.6253">S4.6253</geoCoordinate>
,
<geoCoordinate id="57968A8E527C4B7F1A5A94BD60B0A5CF" box="[1050,1161,1693,1720]" degrees="12.4583" direction="east" orientation="longitude" pageId="3" pageNumber="564" precision="5" value="12.4583">E12.4583</geoCoordinate>
,
<elevation id="B98F0B7A527C4B7F1AD494BD6121A5C0" box="[1172,1304,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="3" pageNumber="564" unit="m" value="221.0">
<quantity id="F55A41AC527C4B7F1AD494BD60E2A5CF" box="[1172,1243,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="3" pageNumber="564" unit="m" value="221.0">221 m</quantity>
a.s.l.
</elevation>
, on
<date id="461CCA89527C4B7F1B0794BD64F6A5AC" pageId="3" pageNumber="564" value="2023-06-06">
<collectingDate id="56583361527C4B7F1B0794BD64F6A5AC" pageId="3" pageNumber="564" value="2023-06-06">
6
<superScript id="C5D74101527C4B7F1B1594BD615BA5DC" attach="left" box="[1365,1378,1693,1707]" fontSize="6" pageId="3" pageNumber="564">th</superScript>
June 2023
</collectingDate>
</date>
by
<collectorName id="9F57899F527C4B7F1EB994E16594A5AC" box="[249,429,1729,1756]" pageId="3" pageNumber="564">Pedro Vaz Pinto</collectorName>
&amp;
<collectorName id="9F57899F527C4B7F1F9194E1664AA5AC" box="[465,627,1729,1755]" pageId="3" pageNumber="564">Michael Mills</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC2527C4B781E8794C565D7A0BE" lastPageId="4" lastPageNumber="565" pageId="3" pageNumber="564" type="diagnosis">
<paragraph id="321DEC49527C4B7F1E8794C56637A4F8" blockId="3.[151,1437,1550,2043]" pageId="3" pageNumber="564">
<emphasis id="00D6305B527C4B7F1E8794C5657BA588" bold="true" box="[199,322,1765,1791]" pageId="3" pageNumber="564">Diagnosis.</emphasis>
A moderately sized dwarf gecko, clearly belonging to the
<taxonomicName id="F5A297CA527C4B7F1DAA94C6604BA588" authorityName="Sternfeld" authorityYear="1912" box="[1002,1138,1765,1791]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527C4B7F1DAA94C6604BA588" box="[1002,1138,1765,1791]" italics="true" pageId="3" pageNumber="564">L. scheffleri</emphasis>
</taxonomicName>
-group on the basis of the double series of transversely enlarged subcaudals (
<figureCitation id="AA99F0CC527C4B7F1C9B95296712A454" box="[731,811,1801,1827]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1779]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure-20@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." figureDoi="http://doi.org/10.5281/zenodo.14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="3" pageNumber="564">Fig. 3c</figureCitation>
) and phylogenetic results (
<figureCitation id="AA99F0CC527C4B7F1A2395296090A453" box="[1123,1193,1801,1828]" captionStart="FIGURE 5" captionStartId="7.[151,250,1870,1894]" captionTargetBox="[155,1436,1394,1846]" captionTargetId="figure-561@7.[151,1436,1389,1846]" captionTargetPageId="7" captionText="FIGURE 5: Phylogenetic relationships of Lygodactylus lobeke sp. nov. (Left) Maximum likelihood tree based on partitioned multigene analysis constructed with iQtree, including information from Gippner et al. (2021), plus all available samples of the L. scheffleri group, with collapsed groups defined by Gippner et al. (2021). (Right) Detailed phylogenetic placement of Lygodactylus lobeke sp. nov. within the L. scheffleri group. Support values are shown above (PP = Bayesian Inference posterior probabilities) and below (BS = Maximum Likelihood bootstrap values) the nodes." figureDoi="http://doi.org/10.5281/zenodo.14240184" httpUri="https://zenodo.org/record/14240184/files/figure.png" pageId="3" pageNumber="564">Fig. 5</figureCitation>
). In its coloration,
<taxonomicName id="F5A297CA527C4B7F1BC6952A64D9A430" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527C4B7F1BC6952A64D9A430" italics="true" pageId="3" pageNumber="564">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527C4B7F1EA6950D6506A430" bold="true" box="[230,319,1837,1863]" pageId="3" pageNumber="564">
<taxonomicNameLabel id="1BE58D20527C4B7F1EA6950D6506A430" box="[230,319,1837,1863]" pageId="3" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from most other species of the
<taxonomicName id="F5A297CA527C4B7F1CAA950D6757A430" authorityName="Sternfeld" authorityYear="1912" box="[746,878,1837,1863]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527C4B7F1CAA950D6757A430" box="[746,878,1837,1863]" italics="true" pageId="3" pageNumber="564">L. scheffleri</emphasis>
</taxonomicName>
-group either by lacking conspicuous dark patches on shoulders and/or on the sides (as in
<emphasis id="00D6305B527C4B7F1C1D957167FCA41D" box="[605,965,1872,1898]" italics="true" pageId="3" pageNumber="564">
<taxonomicName id="F5A297CA527C4B7F1C1D957166D1A41D" authorityName="Sternfeld" authorityYear="1912" box="[605,744,1872,1898]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="scheffleri">L. scheffleri</taxonomicName>
,
<taxonomicName id="F5A297CA527C4B7F1CB5957167FCA41D" authorityName="Pasteur" authorityYear="1964" box="[757,965,1872,1898]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="laterimaculatus">L. laterimaculatus</taxonomicName>
</emphasis>
) or lacking a colored underside (as in
<emphasis id="00D6305B527C4B7F1BC6957165C4A4F9" italics="true" pageId="3" pageNumber="564">
<taxonomicName id="F5A297CA527C4B7F1BC69571653DA4F9" authorityName="Pasteur" authorityYear="1995" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="broadleyi">L. broadleyi</taxonomicName>
,
<taxonomicName id="F5A297CA527C4B7F1F4F955565B6A4F9" authorityName="Matschie" authorityYear="1892" box="[271,399,1908,1934]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="conradti">L. conradti</taxonomicName>
,
<taxonomicName id="F5A297CA527C4B7F1FDA955565C4A4F9" authorityName="Pasteur" authorityYear="1965" box="[410,509,1909,1934]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="gravis">L. gravis</taxonomicName>
</emphasis>
).
</paragraph>
<paragraph id="321DEC49527C4B7F1E8795B86662A48D" blockId="3.[151,1437,1550,2043]" pageId="3" pageNumber="564">
<emphasis id="00D6305B527C4B7F1E8795B865DAA4C5" bold="true" box="[199,483,1944,1970]" pageId="3" pageNumber="564">Comparative diagnosis.</emphasis>
It differs from
<taxonomicName id="F5A297CA527C4B7F1CD895B9671BA4C5" authorityName="Pasteur" authorityYear="1995" box="[664,802,1944,1970]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="broadleyi">
<emphasis id="00D6305B527C4B7F1CD895B9671BA4C5" box="[664,802,1944,1970]" italics="true" pageId="3" pageNumber="564">L. broadleyi</emphasis>
</taxonomicName>
by the color of the underside (whitish in belly and tail
<emphasis id="00D6305B527C4B7F1ED7959D64E6A4A1" box="[151,223,1981,2006]" italics="true" pageId="3" pageNumber="564">versus</emphasis>
yellow and red), by a higher longitudinally count of dorsal scales (240–250
<emphasis id="00D6305B527C4B7F1A71959D6040A4A1" box="[1073,1145,1981,2006]" italics="true" pageId="3" pageNumber="564">versus</emphasis>
212–235) and by a higher number of precloacal pores (9
<emphasis id="00D6305B527C4B7F1FAE95C1660FA48D" box="[494,566,2017,2042]" italics="true" pageId="3" pageNumber="564">versus</emphasis>
6).
</paragraph>
<paragraph id="321DEC49527B4B781E8792B76048A38E" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
It differs from
<taxonomicName id="F5A297CA527B4B781F3092B765D7A3C6" authorityName="Matschie" authorityYear="1892" box="[368,494,151,177]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="conradti">
<emphasis id="00D6305B527B4B781F3092B765D7A3C6" box="[368,494,151,177]" italics="true" pageId="4" pageNumber="565">L. conradti</emphasis>
</taxonomicName>
by the color of the underside (whitish
<emphasis id="00D6305B527B4B781DEA92B767CBA3C7" box="[938,1010,151,176]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
bright yellow in males), by lacking a prominent light dorsolateral stripe, by a different number of supralabials (7
<emphasis id="00D6305B527B4B781DA5929B6014A3A3" box="[997,1069,187,212]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
8), by a higher number of dorsal scales (240–250
<emphasis id="00D6305B527B4B781F1392FF65A2A38F" box="[339,411,223,248]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
218) and by a higher number of precloacal pores (9
<emphasis id="00D6305B527B4B781DA792FF6016A38F" box="[999,1071,223,248]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
5–7).
</paragraph>
<paragraph id="321DEC49527B4B781E87932360A4A2FE" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
It differs from
<taxonomicName id="F5A297CA527B4B781F33932365E1A26B" authorityName="Pasteur" authorityYear="1965" box="[371,472,259,284]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="gravis">
<emphasis id="00D6305B527B4B781F33932365E1A26B" box="[371,472,259,284]" italics="true" pageId="4" pageNumber="565">L. gravis</emphasis>
</taxonomicName>
by smaller size (SVL of
<quantity id="F55A41AC527B4B781CBD9322675AA26A" box="[765,867,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="4" pageNumber="565" unit="mm" value="32.5">32.5 mm</quantity>
<emphasis id="00D6305B527B4B781D2C9323678DA26B" box="[876,948,259,284]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
<quantity id="F55A41AC527B4B781DFD93236005A26A" box="[957,1084,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="4" pageNumber="565" unit="mm" value="40.0" valueMax="42.0" valueMin="38.0">38–42 mm</quantity>
), by the color of the underside (whitish
<emphasis id="00D6305B527B4B781EB993076578A237" box="[249,321,295,320]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
yellow to red in females and males), by the color of the scales of the pseudo-escutcheon
<emphasis id="00D6305B527B4B781B5E93076165A237" box="[1310,1372,295,320]" italics="true" pageId="4" pageNumber="565">sensu</emphasis>
<bibRefCitation id="563391B8527B4B781B2393076560A212" author="Bons, J. &amp; Pasteur, G." pageId="4" pageNumber="565" pagination="2547 - 2550" refId="ref7617" refString="Bons, J. &amp; Pasteur, G. (1977) Solution histologique a un probleme de taxonomie herpetologique interessant les rapports paleobiologiques de l'Amerique du Sud de de l' Afrique. Comptes Rendus de l' Academie des Sciences, Serie D, 284 (24), 2547 - 2550." type="journal article" year="1977">Bons &amp; Pasteur (1977)</bibRefCitation>
(see description of
<typeStatus id="ED1952EB527B4B781C77936B66A0A212" box="[567,665,331,357]" pageId="4" pageNumber="565" type="paratype">paratype</typeStatus>
) (shiny yellowish
<emphasis id="00D6305B527B4B781D29936B6788A213" box="[873,945,331,356]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
beige), by a higher number of dorsal scales (240–250
<emphasis id="00D6305B527B4B781F4A934F656BA2FF" box="[266,338,367,392]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
158–195) and by a slightly higher number of precloacal pores (9
<emphasis id="00D6305B527B4B781A71934F6040A2FF" box="[1073,1145,367,392]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
8).
</paragraph>
<paragraph id="321DEC49527B4B781E8793B367A1A282" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
It differs from
<taxonomicName id="F5A297CA527B4B781F3093B365CFA2DA" authorityName="Sternfeld" authorityYear="1912" box="[368,502,403,429]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527B4B781F3093B365CFA2DA" box="[368,502,403,429]" italics="true" pageId="4" pageNumber="565">L. scheffleri</emphasis>
</taxonomicName>
by lacking dark spots on the neck and on the flanks in males, by a higher number of dorsal scales (240–250
<emphasis id="00D6305B527B4B781FFE9397663FA2A7" box="[446,518,439,464]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
163–166), by a higher number of labials (7 infralabials
<emphasis id="00D6305B527B4B781A3A939760FBA2A7" box="[1146,1218,439,464]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
5–6, 7 supralabials
<emphasis id="00D6305B527B4B781ED793FB64E6A283" box="[151,223,475,500]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
5–6) and by a higher number of precloacal pores (9
<emphasis id="00D6305B527B4B781D6B93FB674AA283" box="[811,883,475,500]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
6).
</paragraph>
<paragraph id="321DEC49527B4B781E8793DF67EAA116" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
It differs from
<taxonomicName id="F5A297CA527B4B781F2E93DF6604A16E" authorityName="Pasteur" authorityYear="1964" box="[366,573,511,537]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="laterimaculatus">
<emphasis id="00D6305B527B4B781F2E93DF6604A16E" box="[366,573,511,537]" italics="true" pageId="4" pageNumber="565">L. laterimaculatus</emphasis>
</taxonomicName>
by the lack of large dark blotches on the flanks in males, by the color of the scales of the pseudo-escutcheon (shiny yellowish
<emphasis id="00D6305B527B4B781C8890036729A14B" box="[712,784,547,572]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
brown or dark brown), by a higher number of infralabials (7
<emphasis id="00D6305B527B4B781EF6906764C7A117" box="[182,254,583,608]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
5–6) and by a higher number of precloacal pores (9
<emphasis id="00D6305B527B4B781D0A906767ABA117" box="[842,914,583,608]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
6–7).
</paragraph>
<paragraph id="321DEC49527B4B781E87904B6622A04E" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
Other members of the
<taxonomicName id="F5A297CA527B4B781F9E904B6653A1F2" authorityName="Sternfeld" authorityYear="1912" box="[478,618,619,645]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527B4B781F9E904B6653A1F2" box="[478,618,619,645]" italics="true" pageId="4" pageNumber="565">L. scheffleri</emphasis>
</taxonomicName>
-group include two poorly known taxa,
<taxonomicName id="F5A297CA527B4B781A0E904B615FA1F3" authorityName="Pasteur" authorityYear="1964" box="[1102,1382,619,645]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="subSpecies" species="scheffleri" subSpecies="compositus">
<emphasis id="00D6305B527B4B781A0E904B615FA1F3" box="[1102,1382,619,645]" italics="true" pageId="4" pageNumber="565">L. scheffleri compositus</emphasis>
</taxonomicName>
and
<taxonomicName id="F5A297CA527B4B781ED790AF6503A1DE" authorityName="Pasteur" authorityYear="1964" box="[151,314,655,681]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="ulugurensis">
<emphasis id="00D6305B527B4B781ED790AF6503A1DE" box="[151,314,655,681]" italics="true" pageId="4" pageNumber="565">L. ulugurensis</emphasis>
</taxonomicName>
, originally described as a subspecies of
<taxonomicName id="F5A297CA527B4B781D5090AF67A0A1DE" authorityName="Sternfeld" authorityYear="1912" box="[784,921,655,681]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527B4B781D5090AF67A0A1DE" box="[784,921,655,681]" italics="true" pageId="4" pageNumber="565">L. scheffleri</emphasis>
</taxonomicName>
. The taxon
<emphasis id="00D6305B527B4B781A6790AF609CA1DF" box="[1063,1189,655,680]" italics="true" pageId="4" pageNumber="565">compositus</emphasis>
is characterized by a large dark blotch on the neck and seven precloacal pores in males.
<taxonomicName id="F5A297CA527B4B781DD790936044A1BA" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[919,1149,691,717]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781DD790936044A1BA" box="[919,1149,691,717]" italics="true" pageId="4" pageNumber="565">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781AC6909360D8A1BA" bold="true" box="[1158,1249,691,717]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781AC6909360D8A1BA" box="[1158,1249,691,717]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the taxon
<taxonomicName id="F5A297CA527B4B781E9D90F76566A186" authorityName="Pasteur" authorityYear="1964" box="[221,351,727,753]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="ulugurensis">
<emphasis id="00D6305B527B4B781E9D90F76566A186" box="[221,351,727,753]" italics="true" pageId="4" pageNumber="565">ulugurensis</emphasis>
</taxonomicName>
by the color pattern of the throat (stippled with dark spots
<emphasis id="00D6305B527B4B781DBB90F7607AA187" box="[1019,1091,727,752]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
a dark median line flanked by convergent lines), by the numbers of post- and postpostmental scales (3 and 5
<emphasis id="00D6305B527B4B781A4490DB6075A063" box="[1028,1100,763,788]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
2 and 4) and a higher number of precloacal pores (9
<emphasis id="00D6305B527B4B781FD1913F65E0A04F" box="[401,473,799,824]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
5–6).
</paragraph>
<paragraph id="321DEC49527B4B781E87916365D7A0BE" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
<taxonomicName id="F5A297CA527B4B781E8791636595A02A" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[199,428,835,861]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781E8791636595A02A" box="[199,428,835,861]" italics="true" pageId="4" pageNumber="565">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781FF391636634A02A" bold="true" box="[435,525,835,861]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781FF391636634A02A" box="[435,525,835,861]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="F5A297CA527B4B781CE19163672BA02A" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[673,786,835,861]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B527B4B781CE19163672BA02A" box="[673,786,835,861]" italics="true" pageId="4" pageNumber="565">L. fischeri</emphasis>
</taxonomicName>
(
<typeStatus id="ED1952EB527B4B781D61916367BEA02A" box="[801,903,835,861]" pageId="4" pageNumber="565" type="holotype">holotype</typeStatus>
, BMNH 1946.8.22.45), currently placed within the
<taxonomicName id="F5A297CA527B4B781E839147650EA0F6" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[195,311,871,897]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="fischeri">
<emphasis id="00D6305B527B4B781E839147650EA0F6" box="[195,311,871,897]" italics="true" pageId="4" pageNumber="565">L. fischeri</emphasis>
</taxonomicName>
-group, by the lack of a large dark blotch above the shoulders and of several dark blotches on the flanks, by numbers of labial scales (7 supralabials
<emphasis id="00D6305B527B4B781C8F91AB672EA0D3" box="[719,791,907,932]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
9, 7 infralabials
<emphasis id="00D6305B527B4B781D9591AB6024A0D3" box="[981,1053,907,932]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
6) and a slightly lower number of precloacal pores (9
<emphasis id="00D6305B527B4B781F33918F6582A0BF" box="[371,443,943,968]" italics="true" pageId="4" pageNumber="565">versus</emphasis>
10).
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC2527B4B7B1E8791F2654EA04E" lastPageId="7" lastPageNumber="568" pageId="4" pageNumber="565" type="description">
<paragraph id="321DEC49527B4B781E8791F267B9A79E" blockId="4.[151,1437,151,1257]" pageId="4" pageNumber="565">
<emphasis id="00D6305B527B4B781E8791F2654BA09B" bold="true" box="[199,370,978,1004]" pageId="4" pageNumber="565">Phylogenetics.</emphasis>
<taxonomicName id="F5A297CA527B4B781F3791F36663A09A" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[375,602,979,1005]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781F3791F36663A09A" box="[375,602,979,1005]" italics="true" pageId="4" pageNumber="565">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781C1F91F36681A09A" bold="true" box="[607,696,979,1005]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781C1F91F36681A09A" box="[607,696,979,1005]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="F5A297CA527B4B781D1C91F367D8A09A" authorityName="Sternfeld" authorityYear="1912" box="[860,993,979,1005]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B527B4B781D1C91F367D8A09A" box="[860,993,979,1005]" italics="true" pageId="4" pageNumber="565">L. scheffleri</emphasis>
</taxonomicName>
-group as identified by
<bibRefCitation id="563391B8527B4B781AA091F361A5A09A" author="Roll, B. &amp; Prohl, H. &amp; Hoffmann, K. - P." box="[1248,1436,979,1005]" pageId="4" pageNumber="565" pagination="327 - 335" refId="ref9837" refString="Roll, B., Prohl, H. &amp; Hoffmann, K. - P. (2010) Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata, Gekkonidae). Molecular Phylogenetics and Evolution, 56 (1), 327 - 335. https: // doi. org / 10.1016 / j. ympev. 2010.02.002" type="journal article" year="2010">
Röll
<emphasis id="00D6305B527B4B781B5691F36175A09A" box="[1302,1356,979,1005]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
(2010)
</bibRefCitation>
and confirmed by
<bibRefCitation id="563391B8527B4B781F2491D7666AA766" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." box="[356,595,1015,1041]" pageId="4" pageNumber="565" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B527B4B781F8791D765C6A766" box="[455,511,1015,1041]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
(2021)
</bibRefCitation>
. The result of the phylogenetic analysis presented here recovered
<taxonomicName id="F5A297CA527B4B781B7691D761A5A766" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1334,1436,1015,1041]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781B7691D761A5A766" box="[1334,1436,1015,1041]" italics="true" pageId="4" pageNumber="565">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781ED7963B64C9A742" bold="true" box="[151,240,1051,1077]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781ED7963B64C9A742" box="[151,240,1051,1077]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as an independent clade (PP: 1, BS: 100;
<figureCitation id="AA99F0CC527B4B781CFA963B66C2A742" box="[698,763,1050,1077]" captionStart="FIGURE 5" captionStartId="7.[151,250,1870,1894]" captionTargetBox="[155,1436,1394,1846]" captionTargetId="figure-561@7.[151,1436,1389,1846]" captionTargetPageId="7" captionText="FIGURE 5: Phylogenetic relationships of Lygodactylus lobeke sp. nov. (Left) Maximum likelihood tree based on partitioned multigene analysis constructed with iQtree, including information from Gippner et al. (2021), plus all available samples of the L. scheffleri group, with collapsed groups defined by Gippner et al. (2021). (Right) Detailed phylogenetic placement of Lygodactylus lobeke sp. nov. within the L. scheffleri group. Support values are shown above (PP = Bayesian Inference posterior probabilities) and below (BS = Maximum Likelihood bootstrap values) the nodes." figureDoi="http://doi.org/10.5281/zenodo.14240184" httpUri="https://zenodo.org/record/14240184/files/figure.png" pageId="4" pageNumber="565">Fig. 5</figureCitation>
), sister of the other members found East to the Albertine Rift (namely,
<taxonomicName id="F5A297CA527B4B781F41961F65F7A72E" authorityName="Pasteur" authorityYear="1964" box="[257,462,1087,1113]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="laterimaculatus">
<emphasis id="00D6305B527B4B781F41961F65F7A72E" box="[257,462,1087,1113]" italics="true" pageId="4" pageNumber="565">L. laterimaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="F5A297CA527B4B781C43961F665FA72F" authorityName="Pasteur" authorityYear="1965" box="[515,614,1087,1112]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="gravis">
<emphasis id="00D6305B527B4B781C43961F665FA72F" box="[515,614,1087,1112]" italics="true" pageId="4" pageNumber="565">L. gravis</emphasis>
</taxonomicName>
), in agreement with the results found by
<bibRefCitation id="563391B8527B4B781A70961F60C9A72E" author="Roll, B. &amp; Prohl, H. &amp; Hoffmann, K. - P." box="[1072,1264,1087,1113]" pageId="4" pageNumber="565" pagination="327 - 335" refId="ref9837" refString="Roll, B., Prohl, H. &amp; Hoffmann, K. - P. (2010) Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata, Gekkonidae). Molecular Phylogenetics and Evolution, 56 (1), 327 - 335. https: // doi. org / 10.1016 / j. ympev. 2010.02.002" type="journal article" year="2010">
Röll
<emphasis id="00D6305B527B4B781A27961F60A6A72E" box="[1127,1183,1087,1113]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
(2010)
</bibRefCitation>
and
<bibRefCitation id="563391B8527B4B781B66961F6530A70A" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." pageId="4" pageNumber="565" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B527B4B781BC8961F648DA70A" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
(2021)
</bibRefCitation>
. However, as several members within this group still lack genetic material, the phylogenetic hypothesis of this group remains unclear and needs further investigation.
<taxonomicName id="F5A297CA527B4B781D1096A7678EA7D6" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[848,951,1159,1185]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781D1096A7678EA7D6" box="[848,951,1159,1185]" italics="true" pageId="4" pageNumber="565">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781DFE96A76021A7D6" bold="true" box="[958,1048,1159,1185]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781DFE96A76021A7D6" box="[958,1048,1159,1185]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other members of the group by c. 9.01% and 12.99%, respectively in
<emphasis id="00D6305B527B4B781CEA968A66EDA7B3" box="[682,724,1194,1220]" italics="true" pageId="4" pageNumber="565">16S</emphasis>
uncorrected p-distance. The two known specimens of
<taxonomicName id="F5A297CA527B4B781B76968B61A5A7B2" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1334,1436,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781B76968B61A5A7B2" box="[1334,1436,1195,1221]" italics="true" pageId="4" pageNumber="565">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781ED796EF64C8A79E" bold="true" box="[151,241,1231,1257]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781ED796EF64C8A79E" box="[151,241,1231,1257]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differ by 3.5% (
<emphasis id="00D6305B527B4B781FEA96EE65EDA79F" box="[426,468,1230,1256]" italics="true" pageId="4" pageNumber="565">16S</emphasis>
uncorrected p-distance) to each other.
</paragraph>
<caption id="66DDBCC1527B4B781ED795AD6108A49A" ID-DOI="http://doi.org/10.5281/zenodo.14240178" ID-Zenodo-Dep="14240178" httpUri="https://zenodo.org/record/14240178/files/figure.png" pageId="4" pageNumber="565" startId="4.[151,250,1933,1957]" targetBox="[151,1435,1281,1910]" targetPageId="4" targetType="figure">
<paragraph id="321DEC49527B4B781ED795AD6108A49A" blockId="4.[151,1437,1933,2030]" pageId="4" pageNumber="565">
<emphasis id="00D6305B527B4B781ED795AD6520A4D2" bold="true" box="[151,281,1933,1957]" pageId="4" pageNumber="565">FIGURE 2:</emphasis>
<taxonomicName id="F5A297CA527B4B781F6395AD65CCA4D2" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[291,501,1933,1957]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527B4B781F6395AD65CCA4D2" box="[291,501,1933,1957]" italics="true" pageId="4" pageNumber="565">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527B4B781FBF95AE666AA4D1" bold="true" box="[511,595,1934,1958]" pageId="4" pageNumber="565">
<taxonomicNameLabel id="1BE58D20527B4B781FBF95AE666AA4D1" box="[511,595,1934,1958]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life, shortly after capture. Female holotype: lateral (a) and dorsal views (b). Male paratype: lateral (c) and dorsal views (d). Photographs of holotype: ambient light, analog (Kodachrome 200) (2002), photographs of paratype: flash, digital (Canon EOS 6D Mark II) (2023). Photographs by B.R. (a, b) and P.VP. (c, d).
</paragraph>
</caption>
<caption id="66DDBCC1527A4B791ED794FB65F3A4DF" ID-DOI="http://doi.org/10.5281/zenodo.14240180" ID-Zenodo-Dep="14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="5" pageNumber="566" startId="5.[151,250,1755,1779]" targetBox="[151,1435,181,1732]" targetPageId="5" targetType="figure">
<paragraph id="321DEC49527A4B791ED794FB65F3A4DF" blockId="5.[151,1437,1755,1960]" pageId="5" pageNumber="566">
<emphasis id="00D6305B527A4B791ED794FB652FA584" bold="true" box="[151,278,1755,1779]" pageId="5" pageNumber="566">FIGURE 3:</emphasis>
<taxonomicName id="F5A297CA527A4B791F5C94FC65D2A583" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[284,491,1756,1780]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B527A4B791F5C94FC65D2A583" box="[284,491,1756,1780]" italics="true" pageId="5" pageNumber="566">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B527A4B791FB294FC667AA583" bold="true" box="[498,579,1756,1780]" pageId="5" pageNumber="566">
<taxonomicNameLabel id="1BE58D20527A4B791FB294FC667AA583" box="[498,579,1756,1780]" pageId="5" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d).
</paragraph>
</caption>
<paragraph id="321DEC4952794B7A1E8792B664F3A38E" blockId="6.[151,1437,150,861]" pageId="6" pageNumber="567">
<emphasis id="00D6305B52794B7A1E8792B6661FA3C6" bold="true" box="[199,550,150,177]" pageId="6" pageNumber="567">
Description of the
<typeStatus id="ED1952EB52794B7A1FF892B76618A3C6" box="[440,545,151,177]" pageId="6" pageNumber="567" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female, in good state of preservation, tongue removed for genetic investigations (
<figureCitation id="AA99F0CC52794B7A1F01929B6590A3A2" box="[321,425,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1844,1868]" captionTargetBox="[151,1436,885,1821]" captionTargetId="figure-350@6.[151,1436,885,1821]" captionTargetPageId="6" captionText="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi="http://doi.org/10.5281/zenodo.14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="6" pageNumber="567">Fig 4a–d</figureCitation>
). SVL
<quantity id="F55A41AC52794B7A1FB6929A6662A3A2" box="[502,603,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="6" pageNumber="567" unit="mm" value="32.5">32.5 mm</quantity>
, TL
<quantity id="F55A41AC52794B7A1CCE929B66E7A3A2" box="[654,734,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="567" unit="mm" value="32.0">32 mm</quantity>
, tail doubly regenerated (last 18 +
<quantity id="F55A41AC52794B7A1A20929A609DA3A2" box="[1120,1188,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="567" unit="mm" value="5.0">5 mm</quantity>
), HW
<quantity id="F55A41AC52794B7A1AAB929A617DA3A2" box="[1259,1348,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="567" unit="mm" value="5.2">5.2 mm</quantity>
; HL
<quantity id="F55A41AC52794B7A1B39929B64FFA38E" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="567" unit="mm" value="7.7">7.7 mm</quantity>
.
</paragraph>
<paragraph id="321DEC4952794B7A1E879323661DA1F2" blockId="6.[151,1437,150,861]" pageId="6" pageNumber="567">
Rostral scale slightly smaller in width than the mental scale. Posterior margin of the nostril exactly above the suture of rostral and first supralabial scale; nostril surrounded by following scales: rostral and first supralabial, one postnasal, one supranasal and nasal. Two internasal scales. Seven infralabial scales, seven supralabial scales (right and left sides, respectively). Mental scale bell-shaped and semi-divided with sutures (
<figureCitation id="AA99F0CC52794B7A1A2D934F60F8A2FE" box="[1133,1217,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1779]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure-20@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." figureDoi="http://doi.org/10.5281/zenodo.14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="6" pageNumber="567">Fig. 3a</figureCitation>
). Three postmental scales, outer scales of same shape and size, separated by a smaller one. Five postpostmental scales. Gulars decrease gradually in size until the posterior margin of the eyes, after that the gulars are distinctly smaller and similar in size. Number of dorsal scales 246, number of ventral scales 98. Nine precloacal spots corresponding to precloacal pores in males. Fourth toe with four pairs of subdigital lamellae. Original part of tail with two median rows of subcaudal scales distinctly enlarged transversely (
<figureCitation id="AA99F0CC52794B7A1C929003671FA14A" box="[722,806,547,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1779]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure-20@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." figureDoi="http://doi.org/10.5281/zenodo.14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="6" pageNumber="567">Fig. 3c</figureCitation>
). Scales of the regenerated parts (regenerates 1 and 2) below irregularly arranged. Tip of tail (regenerate 2) tapered, caudal adhesive pad consisting of small irregularly arranged scales with setae (
<figureCitation id="AA99F0CC52794B7A1F89904B662EA1F2" box="[457,535,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1844,1868]" captionTargetBox="[151,1436,885,1821]" captionTargetId="figure-350@6.[151,1436,885,1821]" captionTargetPageId="6" captionText="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi="http://doi.org/10.5281/zenodo.14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="6" pageNumber="567">Fig 4d</figureCitation>
).
</paragraph>
<paragraph id="321DEC4952794B7A1E8790AF60F2A02A" blockId="6.[151,1437,150,861]" pageId="6" pageNumber="567">
In life (after capture), the
<typeStatus id="ED1952EB52794B7A1FAF90AF6668A1DE" box="[495,593,655,681]" pageId="6" pageNumber="567" type="holotype">holotype</typeStatus>
had a fawn to beige-brownish ground coloration on dorsum and tail, with a slightly lighter streak dorsolaterally (
<figureCitation id="AA99F0CC52794B7A1C15909366FEA1BA" box="[597,711,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1933,1957]" captionTargetBox="[151,1435,1281,1910]" captionTargetId="figure-565@4.[151,1435,1281,1910]" captionTargetPageId="4" captionText="FIGURE 2: Lygodactylus lobeke sp. nov. in life, shortly after capture. Female holotype: lateral (a) and dorsal views (b). Male paratype: lateral (c) and dorsal views (d). Photographs of holotype: ambient light, analog (Kodachrome 200) (2002), photographs of paratype: flash, digital (Canon EOS 6D Mark II) (2023). Photographs by B.R. (a, b) and P.VP. (c, d)." figureDoi="http://doi.org/10.5281/zenodo.14240178" httpUri="https://zenodo.org/record/14240178/files/figure.png" pageId="6" pageNumber="567">Fig. 2a–b</figureCitation>
). Additionally, head, dorsum, flanks und upper side of the legs exhibited dark and light mottling.A dark line extended from the tip of the snout through the eye and ended above the ear opening. Several rows of light, partly dark-edged spots were arranged longitudinally on the dorsum. The short original part of the tail had two adjoining light spots followed by a light cross-band. Throat, belly and underside of the tail were whitish, the throat was stippled with dark spots. (see also
<emphasis id="00D6305B52794B7A1DEE91636082A02A" box="[942,1211,835,861]" italics="true" pageId="6" pageNumber="567">Results: Natural history</emphasis>
.)
</paragraph>
<caption id="66DDBCC152794B7A1ED7951464EDA4CF" ID-DOI="http://doi.org/10.5281/zenodo.14240182" ID-Zenodo-Dep="14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="6" pageNumber="567" startId="6.[151,250,1844,1868]" targetBox="[151,1436,885,1821]" targetPageId="6" targetType="figure">
<paragraph id="321DEC4952794B7A1ED7951464EDA4CF" blockId="6.[151,1437,1844,1976]" pageId="6" pageNumber="567">
<emphasis id="00D6305B52794B7A1ED79514652CA43B" bold="true" box="[151,277,1844,1868]" pageId="6" pageNumber="567">FIGURE 4:</emphasis>
<taxonomicName id="F5A297CA52794B7A1F5B951465D3A43B" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[283,490,1844,1868]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="6" pageNumber="567" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52794B7A1F5B951465D3A43B" box="[283,490,1844,1868]" italics="true" pageId="6" pageNumber="567">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52794B7A1FB095146678A43B" bold="true" box="[496,577,1844,1868]" pageId="6" pageNumber="567">
<taxonomicNameLabel id="1BE58D2052794B7A1FB095146678A43B" box="[496,577,1844,1868]" pageId="6" pageNumber="567" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h).
</paragraph>
</caption>
<paragraph id="321DEC4952784B7B1E8792B7657DA38E" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
After 17 years in ethanol, the
<typeStatus id="ED1952EB52784B7B1C5792B76640A3C6" box="[535,633,151,177]" pageId="7" pageNumber="568" type="holotype">holotype</typeStatus>
is faded and appears almost homogenous fawn. However, the dark line from the tip of the snout in direction of the ear opening and the light spots as well as the light crossband on the tail are still visible.
</paragraph>
<paragraph id="321DEC4952784B7B1E8793226018A212" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
<emphasis id="00D6305B52784B7B1E8793226619A26A" bold="true" box="[199,544,258,285]" pageId="7" pageNumber="568">
Description of the
<typeStatus id="ED1952EB52784B7B1FEE93236622A26A" box="[430,539,259,285]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
.
</emphasis>
Subadult male in moderate state of preservation (
<figureCitation id="AA99F0CC52784B7B1A29932360E5A26A" box="[1129,1244,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1844,1868]" captionTargetBox="[151,1436,885,1821]" captionTargetId="figure-350@6.[151,1436,885,1821]" captionTargetPageId="6" captionText="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi="http://doi.org/10.5281/zenodo.14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="7" pageNumber="568">Fig. 4e–h</figureCitation>
). Visceral cavity opened for tissue removal for genetic analyses. Tail broken close to the body and with necrotic tissue of older date shortly before the tail tip. SVL
<quantity id="F55A41AC52784B7B1FB5936A6662A212" box="[501,603,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="7" pageNumber="568" unit="mm" value="26.5">26.5 mm</quantity>
, TL
<quantity id="F55A41AC52784B7B1CCE936A66CDA212" box="[654,756,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="7" pageNumber="568" unit="mm" value="25.7">25.7 mm</quantity>
, HW
<quantity id="F55A41AC52784B7B1D75936A67B4A212" box="[821,909,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="568" unit="mm" value="5.1">5.1 mm</quantity>
, HL
<quantity id="F55A41AC52784B7B1D84936A6024A212" box="[964,1053,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="568" unit="mm" value="9.5">9.5 mm</quantity>
.
</paragraph>
<paragraph id="321DEC4952784B7B1E87934F600AA16E" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
The scalation of the
<typeStatus id="ED1952EB52784B7B1FEB934F6632A2FE" box="[427,523,367,393]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
is the same as in the
<typeStatus id="ED1952EB52784B7B1CBA934F6765A2FE" box="[762,860,367,393]" pageId="7" pageNumber="568" type="holotype">holotype</typeStatus>
(
<figureCitation id="AA99F0CC52784B7B1D2B934F67E3A2FE" box="[875,986,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1779]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure-20@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." figureDoi="http://doi.org/10.5281/zenodo.14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="7" pageNumber="568">Fig. 3b, d</figureCitation>
). Due to epidermal injuries the numbers of dorsal and ventral scales cannot be counted. Tail with two rows of distinctly enlarged subcaudals. Typically arranged caudal adhesive lamellae are missing. The male
<typeStatus id="ED1952EB52784B7B1D5B93976742A2A6" box="[795,891,439,465]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
has nine precloacal pores (
<figureCitation id="AA99F0CC52784B7B1AEB939760C7A2A6" box="[1195,1278,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1779]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure-20@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURE 3: Lygodactylus lobeke sp. nov. in life. (a) Chin and (c) precloacal region and proximal part of the tail of the female holotype. Precloacal region with 9 precloacal spots, corresponding to the precloacal pores in the male specimen. Arrowheads in (c) mark row of precloacal scales with spots. Original part of tail with double row of enlarged subcaudal scales, two of which are marked with circles. Inset: same, contrast enhanced. (b) Chin and (d) precloacal region and pseudo-escutcheon (yellow scales) of the male paratype. Precloacal region with 9 precloacal pores. Arrowheads in (d) mark row of precloacal pores. Photographs by B.R. (a, c) and P.VP. (b, d)." figureDoi="http://doi.org/10.5281/zenodo.14240180" httpUri="https://zenodo.org/record/14240180/files/figure.png" pageId="7" pageNumber="568">Fig. 3d</figureCitation>
). The pseudo-escutcheon, also called ss-glands or ss-scales (
<bibRefCitation id="563391B852784B7B1CD893FB6777A282" author="Maderson, P. F. A." box="[664,846,475,501]" pageId="7" pageNumber="568" pagination="1 - 35" refId="ref8961" refString="Maderson, P. F. A. (1968) The epidermal glands of Lygodactylus (Gekkonidae, Lacertilia). Breviora, 288, 1 - 35." type="journal article" year="1968">Maderson 1968</bibRefCitation>
;
<bibRefCitation id="563391B852784B7B1D1B93FB606AA282" author="Bons, J. &amp; Pasteur, G." box="[859,1107,475,501]" pageId="7" pageNumber="568" pagination="2547 - 2550" refId="ref7617" refString="Bons, J. &amp; Pasteur, G. (1977) Solution histologique a un probleme de taxonomie herpetologique interessant les rapports paleobiologiques de l'Amerique du Sud de de l' Afrique. Comptes Rendus de l' Academie des Sciences, Serie D, 284 (24), 2547 - 2550." type="journal article" year="1977">Bons &amp; Pasteur 1977</bibRefCitation>
), consists of shiny yellowish scales in the precloacal and postcloacal region and on the inner sides of the thighs.
</paragraph>
<paragraph id="321DEC4952784B7B1E8790036523A1DE" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
The coloration of the live male
<typeStatus id="ED1952EB52784B7B1C6F900366B6A14A" box="[559,655,547,573]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
(after capture) corresponds largely to that of the
<typeStatus id="ED1952EB52784B7B1A809003611BA14A" box="[1216,1314,547,573]" pageId="7" pageNumber="568" type="holotype">holotype</typeStatus>
except for the numbers of longitudinally rows of spots on the dorsum (
<figureCitation id="AA99F0CC52784B7B1D6F906767A4A116" box="[815,925,583,609]" captionStart="FIGURE 2" captionStartId="4.[151,250,1933,1957]" captionTargetBox="[151,1435,1281,1910]" captionTargetId="figure-565@4.[151,1435,1281,1910]" captionTargetPageId="4" captionText="FIGURE 2: Lygodactylus lobeke sp. nov. in life, shortly after capture. Female holotype: lateral (a) and dorsal views (b). Male paratype: lateral (c) and dorsal views (d). Photographs of holotype: ambient light, analog (Kodachrome 200) (2002), photographs of paratype: flash, digital (Canon EOS 6D Mark II) (2023). Photographs by B.R. (a, b) and P.VP. (c, d)." figureDoi="http://doi.org/10.5281/zenodo.14240178" httpUri="https://zenodo.org/record/14240178/files/figure.png" pageId="7" pageNumber="568">Fig. 2c–d</figureCitation>
). In the
<typeStatus id="ED1952EB52784B7B1DB690676060A116" box="[1014,1113,583,609]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
, there were two rows of light spots, partially edged dark brown towards the spine. The freshly broken tail had indistinct, nearly equidistant light crossbands.
</paragraph>
<paragraph id="321DEC4952784B7B1E879093654EA04E" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
After almost one year in formalin and ethanol, the head of the
<typeStatus id="ED1952EB52784B7B1DE79093603EA1BA" box="[935,1031,691,717]" pageId="7" pageNumber="568" type="paratype">paratype</typeStatus>
has become dark (
<figureCitation id="AA99F0CC52784B7B1AA690936163A1BA" box="[1254,1370,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1844,1868]" captionTargetBox="[151,1436,885,1821]" captionTargetId="figure-350@6.[151,1436,885,1821]" captionTargetPageId="6" captionText="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi="http://doi.org/10.5281/zenodo.14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="7" pageNumber="568">Fig. 4e, g</figureCitation>
). The upper sides of the body and the tail have a grey-brownish ground coloration with some dark mottling. The pattern consisting of light, dark-edged blotches is hardly visible. The modified scales of the pseudo-escutcheon have lost their color (
<figureCitation id="AA99F0CC52784B7B1F5A913F6553A04E" box="[282,362,799,825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1844,1868]" captionTargetBox="[151,1436,885,1821]" captionTargetId="figure-350@6.[151,1436,885,1821]" captionTargetPageId="6" captionText="FIGURE 4: Lygodactylus lobeke sp. nov. type material in preservative. a) to d) female holotype (ZSM 56/2024): a) dorsal and b) ventral views; c) dorsal view of head, d) ventral view of (regenerated) tail tip. e) to h) male paratype (FKH 1259): e) dorsal and f) ventral views; g) dorsal view of head, h) ventral view of (regenerated) tail tip. Photographs by B.R. (a–d) and P. VP. (e–h)." figureDoi="http://doi.org/10.5281/zenodo.14240182" httpUri="https://zenodo.org/record/14240182/files/figure.png" pageId="7" pageNumber="568">Fig. 4f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC252784B7B1E87916367B5A742" pageId="7" pageNumber="568" type="etymology">
<paragraph id="321DEC4952784B7B1E87916367B5A742" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
<emphasis id="00D6305B52784B7B1E8791636577A02A" bold="true" box="[199,334,835,861]" pageId="7" pageNumber="568">Etymology.</emphasis>
The name ‘lobeke’ is a noun in apposition and refers to the Lobeke region in south-eastern
<collectingCountry id="4AB5ACD952784B7B1ED791476536A0F6" box="[151,271,871,897]" name="Cameroon" pageId="7" pageNumber="568">Cameroon</collectingCountry>
, where the
<typeStatus id="ED1952EB52784B7B1FD1914765CAA0F6" box="[401,499,871,897]" pageId="7" pageNumber="568" type="holotype">holotype</typeStatus>
of this species was found. The species is named in honor of this tropical rainforest region, which is also home of the Lobeke National Park (Parc National de Lobéké). The name indirectly honors Mathias Heinze (German Agency for Technical Cooperation [GTZ], at that time Yokadouma), who was one of the ‘architects’ of Lobeke National Park and suggested the name in
<date id="461CCA8952784B7B1DE191F36067A09A" box="[929,1118,979,1005]" pageId="7" pageNumber="568" value="2022-09">September 2022</date>
. Mathias died suddenly and unexpectedly on
<date id="461CCA8952784B7B1F1991D7660DA766" box="[345,564,1014,1041]" pageId="7" pageNumber="568" value="2022-12-06">
6
<superScript id="C5D7410152784B7B1F2791D6654DA773" attach="left" box="[359,372,1014,1028]" fontSize="6" pageId="7" pageNumber="568">th</superScript>
December 2022
</date>
. As common names we suggest ‘Lobeke Dwarf Gecko’ in English, ‘Lobeke-Zwerggecko’ in German, and ‘Osga-Anã de Lobeke’ in Portuguese.
</paragraph>
</subSubSection>
<subSubSection id="7AB8BFC252784B7B1E87961E661EA622" pageId="7" pageNumber="568" type="distribution">
<paragraph id="321DEC4952784B7B1E87961E661EA622" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="568">
<emphasis id="00D6305B52784B7B1E87961E6558A72F" bold="true" box="[199,353,1086,1112]" pageId="7" pageNumber="568">Distribution.</emphasis>
<taxonomicName id="F5A297CA52784B7B1F27961F6672A72E" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[359,587,1087,1113]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52784B7B1F27961F6672A72E" box="[359,587,1087,1113]" italics="true" pageId="7" pageNumber="568">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52784B7B1C11961F6693A72E" bold="true" box="[593,682,1087,1113]" pageId="7" pageNumber="568">
<taxonomicNameLabel id="1BE58D2052784B7B1C11961F6693A72E" box="[593,682,1087,1113]" pageId="7" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from two localities of the western rim of the
<collectingCountry id="4AB5ACD952784B7B1B4B961F616FA72E" box="[1291,1366,1087,1113]" name="Democratic Republic of the Congo" pageId="7" pageNumber="568">Congo</collectingCountry>
Basin in Central Africa with a linear distance of about
<quantity id="F55A41AC52784B7B1C8396426720A70A" box="[707,793,1122,1149]" metricMagnitude="5" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="568" unit="km" value="850.0">850 km</quantity>
from each other (
<figureCitation id="AA99F0CC52784B7B1DA796436013A70A" box="[999,1066,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,956,980]" captionTargetBox="[158,1419,381,933]" captionTargetId="figure-134@2.[158,1430,381,933]" captionTargetPageId="2" captionText="FIGURE 1: Habitats and geographical distribution of Lygodactylus lobeke sp. nov. on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Photos of habitats from (a) type locality (holotype) near Goumela, southeastern Cameroon, and (b) second locality (paratype) near Sanga Wanda, Mayombe National Park, Cabinda Province, Angola. Photographs by B.R. (a) and J.L-R. (b)." figureDoi="http://doi.org/10.5281/zenodo.14240176" httpUri="https://zenodo.org/record/14240176/files/figure.png" pageId="7" pageNumber="568">Fig. 1</figureCitation>
). The
<collectingCountry id="4AB5ACD952784B7B1A38964360FAA70A" box="[1144,1219,1123,1149]" name="Democratic Republic of the Congo" pageId="7" pageNumber="568">Congo</collectingCountry>
Basin includes the forests between the western edge of the Albertine Rift highlands and the Gulf of
<collectingCountry id="4AB5ACD952784B7B1A7E96A760ABA7D6" box="[1086,1170,1159,1185]" name="Guinea-Bissau" pageId="7" pageNumber="568">Guinea</collectingCountry>
, comprising the largest contiguous African rainforest block. Specifically, both localities of
<taxonomicName id="F5A297CA52784B7B1DE6968B6029A7B2" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[934,1040,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52784B7B1DE6968B6029A7B2" box="[934,1040,1195,1221]" italics="true" pageId="7" pageNumber="568">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52784B7B1A5B968B6041A7B2" bold="true" box="[1051,1144,1195,1221]" pageId="7" pageNumber="568">
<taxonomicNameLabel id="1BE58D2052784B7B1A5B968B6041A7B2" box="[1051,1144,1195,1221]" pageId="7" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lie in the Lower
<collectingCountry id="4AB5ACD952784B7B1B0B968B61A5A7B2" box="[1355,1436,1195,1221]" name="Guinea-Bissau" pageId="7" pageNumber="568">Guinea</collectingCountry>
rainforest that constitutes the western component of the
<collectingCountry id="4AB5ACD952784B7B1D5D96EF6751A79E" box="[797,872,1231,1257]" name="Democratic Republic of the Congo" pageId="7" pageNumber="568">Congo</collectingCountry>
Basin (
<bibRefCitation id="563391B852784B7B1D8296EF60AAA79E" author="Hardy, O. J. &amp; Born, C. &amp; Budde, K. &amp; Dainou, K. &amp; Dauby, G. &amp; Duminil, J. &amp; Ewedje, E. - E. B. K. &amp; Gomez, C. &amp; Heuertzu, M. &amp; Koffi, G. K. &amp; Lowe, A. J. &amp; Micheneau, C. &amp; Ndiade-Bourobou, D. &amp; Pineiro, R. &amp; Poncet, V." box="[962,1171,1231,1257]" pageId="7" pageNumber="568" pagination="284 - 296" refId="ref8251" refString="Hardy, O. J., Born, C., Budde, K., Dainou, K., Dauby, G., Duminil, J., Ewedje, E. - E. B. K., Gomez, C., Heuertzu, M., Koffi, G. K., Lowe, A. J., Micheneau, C., Ndiade-Bourobou, D., Pineiro, R. &amp; Poncet, V. (2013) Comparative phylogeography of African rain forest trees: A review of genetic signatures of vegetation history in the Guineo-Congolian region. Comptes Rendus Geoscience, 345, 284 - 296. https: // doi. org / 10.1016 / j. crte. 2013.05.001" type="journal article" year="2013">
Hardy
<emphasis id="00D6305B52784B7B1A5396EF6074A79E" box="[1043,1101,1231,1257]" italics="true" pageId="7" pageNumber="568">et al.</emphasis>
2013
</bibRefCitation>
). As
<taxonomicName id="F5A297CA52784B7B1A8E96EF610EA79E" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1230,1335,1231,1257]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52784B7B1A8E96EF610EA79E" box="[1230,1335,1231,1257]" italics="true" pageId="7" pageNumber="568">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52784B7B1B0196EF61A5A79E" bold="true" box="[1345,1436,1231,1257]" pageId="7" pageNumber="568">
<taxonomicNameLabel id="1BE58D2052784B7B1B0196EF61A5A79E" box="[1345,1436,1231,1257]" pageId="7" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
apparently is an arboreal and forest-dependent species, its area of distribution presumably includes further parts of the Lower
<collectingCountry id="4AB5ACD952784B7B1F78973765B0A646" box="[312,393,1303,1329]" name="Guinea-Bissau" pageId="7" pageNumber="568">Guinea</collectingCountry>
region, possibly including
<collectingCountry id="4AB5ACD952784B7B1C879737672DA646" box="[711,788,1303,1329]" name="Gabon" pageId="7" pageNumber="568">Gabon</collectingCountry>
,
<collectingCountry id="4AB5ACD952784B7B1D63973767CDA646" box="[803,1012,1303,1329]" name="Equatorial Guinea" pageId="7" pageNumber="568">Equatorial Guinea</collectingCountry>
,
<collectingCountry id="4AB5ACD952784B7B1DB89737613DA646" box="[1016,1284,1303,1329]" name="Republic of the Congo" pageId="7" pageNumber="568">Republic of the Congo</collectingCountry>
, and western
<collectingCountry id="4AB5ACD952784B7B1ED7971B661BA622" box="[151,546,1339,1365]" name="Democratic Republic of the Congo" pageId="7" pageNumber="568">Democratic Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="66DDBCC152784B7B1ED7956E67EEA481" ID-DOI="http://doi.org/10.5281/zenodo.14240184" ID-Zenodo-Dep="14240184" httpUri="https://zenodo.org/record/14240184/files/figure.png" pageId="7" pageNumber="568" startId="7.[151,250,1870,1894]" targetBox="[155,1436,1394,1846]" targetPageId="7" targetType="figure">
<paragraph id="321DEC4952784B7B1ED7956E67EEA481" blockId="7.[151,1436,1870,2038]" pageId="7" pageNumber="568">
<emphasis id="00D6305B52784B7B1ED7956E652FA411" bold="true" box="[151,278,1870,1894]" pageId="7" pageNumber="568">FIGURE 5:</emphasis>
Phylogenetic relationships of
<taxonomicName id="F5A297CA52784B7B1C0E956E6727A411" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[590,798,1870,1894]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52784B7B1C0E956E6727A411" box="[590,798,1870,1894]" italics="true" pageId="7" pageNumber="568">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52784B7B1D65956E674EA411" bold="true" box="[805,887,1870,1894]" pageId="7" pageNumber="568">
<taxonomicNameLabel id="1BE58D2052784B7B1D65956E674EA411" box="[805,887,1870,1894]" pageId="7" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Left) Maximum likelihood tree based on partitioned multigene analysis constructed with iQtree, including information from
<bibRefCitation id="563391B852784B7B1DC69552605FA4FD" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." box="[902,1126,1906,1930]" pageId="7" pageNumber="568" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B52784B7B1DA295536021A4FD" box="[994,1048,1906,1930]" italics="true" pageId="7" pageNumber="568">et al.</emphasis>
(2021)
</bibRefCitation>
, plus all available samples of the
<taxonomicName id="F5A297CA52784B7B1EFF95B76503A4D9" authorityName="Sternfeld" authorityYear="1912" box="[191,314,1942,1966]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="scheffleri">
<emphasis id="00D6305B52784B7B1EFF95B76503A4D9" box="[191,314,1942,1966]" italics="true" pageId="7" pageNumber="568">L. scheffleri</emphasis>
</taxonomicName>
group, with collapsed groups defined by
<bibRefCitation id="563391B852784B7B1CAF95B667F6A4D9" author="Gippner, S. &amp; Travers, S. L. &amp; Scherz, M. D. &amp; Colston, T. J. &amp; Lyra, M. L. &amp; Ashwini, V. M. &amp; Multzsch, M. &amp; Nielson, S. V. &amp; Rancilhac, L. &amp; Glaw, F. &amp; Bauer, A. M. &amp; Vences, M." box="[751,975,1942,1966]" pageId="7" pageNumber="568" refId="ref8048" refString="Gippner, S., Travers, S. L., Scherz, M. D., Colston, T. J., Lyra, M. L., Ashwini, V. M., Multzsch, M., Nielson, S. V., Rancilhac, L., Glaw, F., Bauer, A. M. &amp; Vences, M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal volume" year="2021">
Gippner
<emphasis id="00D6305B52784B7B1D0B95B767B9A4D9" box="[843,896,1942,1966]" italics="true" pageId="7" pageNumber="568">et al.</emphasis>
(2021)
</bibRefCitation>
. (Right) Detailed phylogenetic placement of
<taxonomicName id="F5A297CA52784B7B1ED7959A655CA4A5" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[151,357,1978,2002]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="7" pageNumber="568" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52784B7B1ED7959A655CA4A5" box="[151,357,1978,2002]" italics="true" pageId="7" pageNumber="568">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52784B7B1F2B959A6582A4A5" bold="true" box="[363,443,1978,2002]" pageId="7" pageNumber="568">
<taxonomicNameLabel id="1BE58D2052784B7B1F2B959A6582A4A5" box="[363,443,1978,2002]" pageId="7" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
within the
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<emphasis id="00D6305B52784B7B1C6C959B669DA4A5" box="[556,676,1978,2002]" italics="true" pageId="7" pageNumber="568">L. scheffleri</emphasis>
</taxonomicName>
group. Support values are shown above (PP = Bayesian Inference posterior probabilities) and below (BS = Maximum Likelihood bootstrap values) the nodes.
</paragraph>
</caption>
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<paragraph id="321DEC4952774B741ED795086647A4FF" blockId="8.[151,1436,1832,1928]" pageId="8" pageNumber="569">
<emphasis id="00D6305B52774B741ED79508652DA437" bold="true" box="[151,276,1832,1856]" pageId="8" pageNumber="569">FIGURE 6:</emphasis>
<taxonomicName id="F5A297CA52774B741F59950865DFA437" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[281,486,1832,1856]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="8" pageNumber="569" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52774B741F59950865DFA437" box="[281,486,1832,1856]" italics="true" pageId="8" pageNumber="569">Lygodactylus lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52774B741FAB95096602A436" bold="true" box="[491,571,1833,1857]" pageId="8" pageNumber="569">
<taxonomicNameLabel id="1BE58D2052774B741FAB95096602A436" box="[491,571,1833,1857]" pageId="8" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R.
</paragraph>
</caption>
<subSubSection id="7AB8BFC252764B751E8792B660F5A1BB" pageId="9" pageNumber="570" type="biology_ecology">
<paragraph id="321DEC4952764B751E8792B66711A213" blockId="9.[151,1437,150,717]" pageId="9" pageNumber="570">
<emphasis id="00D6305B52764B751E8792B665BEA3C7" bold="true" box="[199,391,150,176]" pageId="9" pageNumber="570">Natural history.</emphasis>
The species apparently inhabits humid rainforests. The
<typeStatus id="ED1952EB52764B751A4892B66053A3C7" box="[1032,1130,150,176]" pageId="9" pageNumber="570" type="holotype">holotype</typeStatus>
was found during the day, resting under a sliver of bark at a height of about
<quantity id="F55A41AC52764B751CF5929A66CEA3A2" box="[693,759,186,213]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="570" unit="m" value="1.5">1.5 m</quantity>
. Careful search with binoculars for several days did not yield a second specimen. The species also seemed to be unknown to local people. The
<typeStatus id="ED1952EB52764B751A7392FF60AAA38E" box="[1075,1171,223,249]" pageId="9" pageNumber="570" type="paratype">paratype</typeStatus>
has been found among leaf litter while preparing a camp site in the forest. It may have fallen from the tree canopy above. Subsequent attempts to find more specimens in the area also did not produce results. Presumably,
<taxonomicName id="F5A297CA52764B751A0F9307608CA237" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[1103,1205,294,320]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751A0F9307608CA237" box="[1103,1205,294,320]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<emphasis id="00D6305B52764B751AFC9306612CA237" bold="true" box="[1212,1301,294,320]" pageId="9" pageNumber="570">
<taxonomicNameLabel id="1BE58D2052764B751AFC9306612CA237" box="[1212,1301,294,320]" pageId="9" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
inhabits the canopy region and therefore is rarely met in lower heights.
</paragraph>
<paragraph id="321DEC4952764B751E87934E60F5A1BB" blockId="9.[151,1437,150,717]" pageId="9" pageNumber="570">
The
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of
<taxonomicName id="F5A297CA52764B751F38934F65E5A2FF" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[376,476,366,392]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751F38934F65E5A2FF" box="[376,476,366,392]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BE58D2052764B751FA0934E6601A2FE" box="[480,568,366,393]" pageId="9" pageNumber="570" rank="species">
<emphasis id="00D6305B52764B751FA0934E6608A2FF" bold="true" box="[480,561,366,392]" pageId="9" pageNumber="570">sp. nov</emphasis>
.
</taxonomicNameLabel>
lived for about 5 years in a laboratory colony of several species of
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<emphasis id="00D6305B52764B751B41934E61ACA2FF" box="[1281,1429,366,392]" italics="true" pageId="9" pageNumber="570">Lygodactylus</emphasis>
</taxonomicName>
. Thus, the animal could be observed much more thoroughly than had been possible in the natural habitat. Like several other species of the genus
<taxonomicName id="F5A297CA52764B751C509396669DA2A7" box="[528,676,438,464]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="genus">
<emphasis id="00D6305B52764B751C509396669DA2A7" box="[528,676,438,464]" italics="true" pageId="9" pageNumber="570">Lygodactylus</emphasis>
</taxonomicName>
(
<bibRefCitation id="563391B852764B751CF29396671CA2A7" author="Roll, B." box="[690,805,438,464]" pageId="9" pageNumber="570" refId="ref9814" refString="Roll, B. (2013) Tagaktive Zwerggeckos der Gattung Lygodactylus. Natur und Tier Verlag, Munster, 120 pp." type="book" year="2013">Röll 2013</bibRefCitation>
,
<bibRefCitation id="563391B852764B751D6F939667E7A2A7" author="Roll, B. &amp; Sanchez, M. &amp; Gippner, S. &amp; Bauer, A. M. &amp; Travers, S. L. &amp; Glaw, F. &amp; Hawlitschek, O. &amp; Vences, M." box="[815,990,438,464]" pageId="9" pageNumber="570" pagination="232 - 250" refId="ref9898" refString="Roll, B., Sanchez, M., Gippner, S., Bauer, A. M., Travers, S. L., Glaw, F., Hawlitschek, O. &amp; Vences, M. (2023). Phylogeny of dwarf geckos of the genus Lygodactylus (Gekkonidae) in the Western Indian Ocean. Zootaxa, 5311 (2), 232 - 250. https: // doi. org / 10.11646 / zootaxa. 5311.2.4" type="journal article" year="2023">
Röll
<emphasis id="00D6305B52764B751D25939767A5A2A7" box="[869,924,438,464]" italics="true" pageId="9" pageNumber="570">et al.</emphasis>
2023
</bibRefCitation>
), it possessed distinct ‘mood dependent’ colorations, comprising a ‘neutral’, a ‘distress’, a ‘display’, and a ‘pyjamas’ coloration (
<figureCitation id="AA99F0CC52764B751ADD93FA60DFA283" box="[1181,1254,474,500]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="9" pageNumber="570">Fig. 6</figureCitation>
). In the neutral coloration, the ground color was beige to light brown and the dorsolateral streak was only indistinct (
<figureCitation id="AA99F0CC52764B751AB893DE6170A16F" box="[1272,1353,510,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="9" pageNumber="570">Fig. 6c</figureCitation>
). In the display coloration,
<taxonomicName id="F5A297CA52764B751F2E900365ECA14B" authorityName="Röll &amp; Pinto &amp; Lobón-Rovira" authorityYear="2024" box="[366,469,546,572]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="9" pageNumber="570" phylum="Chordata" rank="species" species="lobeke" status="sp. nov.">
<emphasis id="00D6305B52764B751F2E900365ECA14B" box="[366,469,546,572]" italics="true" pageId="9" pageNumber="570">L. lobeke</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BE58D2052764B751F9B9002660FA14A" box="[475,566,546,573]" pageId="9" pageNumber="570" rank="species">
<emphasis id="00D6305B52764B751F9B90026609A14B" bold="true" box="[475,560,546,572]" pageId="9" pageNumber="570">sp. nov</emphasis>
.
</taxonomicNameLabel>
has a fawn ground coloration. Dorsal pattern elements, e.g., light patches, are hard to recognize (
<figureCitation id="AA99F0CC52764B751F2790666583A117" box="[359,442,582,608]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="9" pageNumber="570">Fig. 6d</figureCitation>
). In response to stress, the animal became darker, the dorsal pattern was richer in contrast and the dorsolateral streak was clearly visible (‘distress’ coloration,
<figureCitation id="AA99F0CC52764B751DC9904A67CEA1F3" box="[905,1015,618,644]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="9" pageNumber="570">Fig. 6a–b</figureCitation>
). At night, the animal generally rested on leaves of plants in the terrarium. Asleep, it showed a characteristic ‘pyjamas’ coloration with conspicuously whitish lateral bands extending from the neck over the flanks and the tail to the tail tip (
<figureCitation id="AA99F0CC52764B751A2C90926087A1BB" box="[1132,1214,690,716]" captionStart="FIGURE 6" captionStartId="8.[151,250,1832,1856]" captionTargetBox="[151,1435,181,1807]" captionTargetId="figure-23@8.[151,1435,181,1808]" captionTargetPageId="8" captionText="FIGURE 6: Lygodactylus lobeke sp. nov. female holotype in life: ‘Mood dependent’ colorations. (a) and (b) ‘distress’ coloration (compare coloration shortly after capture, see Fig. 2a); (c) ‘neutral’ coloration; (d) ‘display’ coloration; (e) ‘pyjamas’ coloration (resting at night on a leaf). Photographs by B.R." figureDoi="http://doi.org/10.5281/zenodo.14240186" httpUri="https://zenodo.org/record/14240186/files/figure.png" pageId="9" pageNumber="570">Fig. 6e</figureCitation>
).
</paragraph>
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<emphasis id="00CCA0270023B925FF4D828300C0FF04" bold="true" box="[151,507,151,178]" pageId="5" pageNumber="353">
<taxonomicName id="F5B807B60023B925FF4D828300ADFF07" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[151,406,151,178]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="353" phylum="Arthropoda" rank="species" species="ketambe" status="sp. nov.">
<emphasis id="00CCA0270023B925FF4D828300ADFF07" bold="true" box="[151,406,151,178]" italics="true" pageId="5" pageNumber="353">Thelcticopis ketambe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BFF1D5C0023B925FE47828300C0FF04" box="[413,507,151,177]" pageId="5" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0023B925FF4D82AF0053FF63" box="[151,360,187,214]" pageId="5" pageNumber="353" type="description">
<paragraph id="32077C350023B925FF4D82AF0053FF63" blockId="5.[151,507,151,214]" box="[151,360,187,214]" pageId="5" pageNumber="353">
<figureCitation id="AA8360B00023B925FF4D82AF002CFF60" box="[151,279,187,213]" captionStart-0="FIGURES 13–18" captionStart-1="FIGURES 19–24" captionStartId-0="4.[151,264,1740,1764]" captionStartId-1="5.[151,264,1394,1418]" captionTargetBox-0="[156,1426,248,1717]" captionTargetBox-1="[151,1429,381,1364]" captionTargetId-0="figure-30@4.[151,1436,237,1717]" captionTargetId-1="figure-56@5.[151,1436,381,1371]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 13–18. Thelcticopis ketambe sp. nov., male. 13–14 Habitus (13 dorsal, 14 ventral); 15–18 Palp (15 dorsal, 16, prolateral, 17 ventral, 18 retrolateral). Scale bars: 13–14=10 mm; 15–18=1 mm. Abbreviations: CS—cymbial scopula, RB— retrolateral bulge of cymbium." captionText-1="FIGURES 19–24. Thelcticopis ketambe sp. nov., male. 19–21 Palpal tibia (19 dorsal, 20 retrolateral, 21 ventral); 22–24 Bulb (22 prolateral, 23 ventral, 24 retrolateral). Scale bars: 0.5 mm. Abbreviations: C—conductor, DE—distal embolic apophysis, dRTA—dorsalretrolateraltibial apophysis,E—embolus,PE—prolateralembolicapophysis,RE—retrolateralembolic apophysis, RK—retrolateral keel, RTS—retrolateral tibial spine, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.14239947" figureDoi-1="http://doi.org/10.5281/zenodo.14239949" httpUri-0="https://zenodo.org/record/14239947/files/figure.png" httpUri-1="https://zenodo.org/record/14239949/files/figure.png" pageId="5" pageNumber="353">Figs 13–21</figureCitation>
,
<figureCitation id="AA8360B00023B925FEF882AF0053FF63" box="[290,360,187,214]" captionStart="FIGURES 25–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[151,1436,885,1419]" captionTargetId="figure-340@6.[151,1436,885,1419]" captionTargetPageId="6" captionText="FIGURES 25–26. Distribution records of Thelcticopis spp. from Sumatra, Mentawai Islands and adjacent Malay Peninsula: T. karnyi (1), T. lizae sp. nov. (2), T. ketambe sp. nov. (3), T. klossi (4), T. orichalcea (5), T. modesta (6)." figureDoi="http://doi.org/10.5281/zenodo.14239953" httpUri="https://zenodo.org/record/14239953/files/figure.png" pageId="5" pageNumber="353">25–26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0023B925FF4D8317003AFED0" pageId="5" pageNumber="353" type="materials_examined">
<paragraph id="32077C350023B925FF4D8317003AFED0" blockId="5.[151,1436,259,357]" pageId="5" pageNumber="353">
<emphasis id="00CCA0270023B925FF4D8317007EFEA8" bold="true" box="[151,325,259,285]" pageId="5" pageNumber="353">Type material.</emphasis>
<materialsCitation id="82D076680023B925FE91831702C6FEF4" collectionCode="ISEA" country="Indonesia" elevation="450" latitude="3.6833334" location="Ketambe village" longLatPrecision="1307" longitude="97.65" pageId="5" pageNumber="353" specimenCount="1" specimenCount-male="1" stateProvince="Aceh" typeStatus="holotype">
<emphasis id="00CCA0270023B925FE9183170213FEA8" bold="true" box="[331,808,259,286]" pageId="5" pageNumber="353">
<collectingCountry id="4AAF3CA50023B925FE91831700D5FEA8" box="[331,494,259,285]" name="Indonesia" pageId="5" pageNumber="353">INDONESIA</collectingCountry>
:
<emphasis id="00CCA0270023B925FE2083170213FEA8" bold="true" box="[506,808,259,286]" italics="true" pageId="5" pageNumber="353">
<collectingRegion id="F07CB2D70023B925FE2083170399FEA8" box="[506,674,259,286]" country="Indonesia" name="Aceh" pageId="5" pageNumber="353">Aceh Province</collectingRegion>
(
<collectingRegion id="F07CB2D70023B925FD6883170223FEA8" box="[690,792,259,285]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="353">Sumatra</collectingRegion>
):
</emphasis>
</emphasis>
<typeStatus id="ED03C2970023B925FCF5831702ACFEA8" box="[815,919,259,285]" pageId="5" pageNumber="353" type="holotype">Holotype</typeStatus>
<specimenCount id="24BEB7BC0023B925FC478310028FFEAB" box="[925,948,260,286]" count="1" pageId="5" pageNumber="353" type="male">♂</specimenCount>
,
<location id="37672AEE0023B925FC1A83170543FEA8" LSID="urn:lsid:plazi:treatment:BA11CD230023B926FF4D828302B1FCE8:37672AEE0023B925FC1A83170543FEA8" box="[960,1144,259,285]" country="Indonesia" latitude="3.6833334" longLatPrecision="1307" longitude="97.65" name="Ketambe village" pageId="5" pageNumber="353" stateProvince="Aceh">Ketambe village</location>
[
<geoCoordinate id="578C1AF20023B925FB5F831105D3FE95" box="[1157,1256,261,288]" degrees="03" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="353" precision="925" value="3.6833334">03°41′N</geoCoordinate>
,
<geoCoordinate id="578C1AF20023B925FB2983110469FE95" box="[1267,1362,261,288]" degrees="97" direction="east" minutes="39" orientation="longitude" pageId="5" pageNumber="353" precision="925" value="97.65">97°39′E</geoCoordinate>
],
<quantity id="F540D1D00023B925FABE831701D8FEF7" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="353" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="B9959B060023B925FABE831701D8FEF7" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="353" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400– 500 m</elevation>
</quantity>
, 1988 (precise date unknown), unknown collector (
<collectionCode id="54A9E4F00023B925FCEC83330243FEF4" box="[822,888,295,321]" pageId="5" pageNumber="353">ISEA</collectionCode>
, 001.9100)
</materialsCitation>
.
<materialsCitation id="82D076680023B925FBD0833C01C6FED0" collectionCode="IBSS" country="Indonesia" elevation="450" latitude="3.6833334" location="Ketambe village" longLatPrecision="1307" longitude="97.65" pageId="5" pageNumber="353" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<typeStatus id="ED03C2970023B925FBD0833C054BFEF7" box="[1034,1136,296,322]" pageId="5" pageNumber="353" type="paratype">Paratype</typeStatus>
:
<specimenCount id="24BEB7BC0023B925FBA48333059AFEF4" box="[1150,1185,295,321]" count="1" pageId="5" pageNumber="353" type="male">1♂</specimenCount>
(
<collectionCode id="54A9E4F00023B925FB69833305C8FEF4" box="[1203,1267,295,321]" pageId="5" pageNumber="353">IBSS</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="66C72CBD0023B925FF4D8766003CF9AE" ID-DOI="http://doi.org/10.5281/zenodo.14239949" ID-Zenodo-Dep="14239949" httpUri="https://zenodo.org/record/14239949/files/figure.png" pageId="5" pageNumber="353" startId="5.[151,264,1394,1418]" targetBox="[151,1429,381,1364]" targetPageId="5" targetType="figure">
<paragraph id="32077C350023B925FF4D8766003CF9AE" blockId="5.[151,1437,1394,1563]" pageId="5" pageNumber="353">
<emphasis id="00CCA0270023B925FF4D8766006EFA3E" bold="true" box="[151,341,1394,1419]" pageId="5" pageNumber="353">FIGURES 19–24.</emphasis>
<taxonomicName id="F5B807B60023B925FE8187670309FA3E" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[347,562,1395,1419]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="353" phylum="Arthropoda" rank="species" species="ketambe" status="sp. nov.">
<emphasis id="00CCA0270023B925FE8187670309FA3E" box="[347,562,1395,1419]" italics="true" pageId="5" pageNumber="353">Thelcticopis ketambe</emphasis>
</taxonomicName>
<emphasis id="00CCA0270023B925FDE3876703B0FA3E" bold="true" box="[569,651,1395,1419]" pageId="5" pageNumber="353">
<taxonomicNameLabel id="1BFF1D5C0023B925FDE3876703B0FA3E" box="[569,651,1395,1419]" pageId="5" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 19–21 Palpal tibia (19 dorsal, 20 retrolateral, 21 ventral); 22–24 Bulb (22 prolateral, 23 ventral, 24 retrolateral). Scale bars: 0.5 mm. Abbreviations: C—conductor, DE—distal embolic apophysis, dRTA—dorsal retrolateral tibial apophysis, E—embolus, PE—prolateral embolic apophysis, RE—retrolateral embolic apophysis, RK—retrolateral keel, RTS—retrolateral tibial spine, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe.
</paragraph>
</caption>
<subSubSection id="7AA22FBE0023B925FF1D84790370F91E" pageId="5" pageNumber="353" type="etymology">
<paragraph id="32077C350023B925FF1D84790370F91E" blockId="5.[151,1437,1645,2031]" pageId="5" pageNumber="353">
<emphasis id="00CCA0270023B925FF1D84790075F932" bold="true" box="[199,334,1645,1671]" pageId="5" pageNumber="353">Etymology.</emphasis>
The specific epithet is derived from Ketambe Village, near which the
<typeStatus id="ED03C2970023B925FB5D8479058CF932" box="[1159,1207,1645,1671]" pageId="5" pageNumber="353">type</typeStatus>
locality of the new species is situated; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0023B925FF1D84A1002DF812" pageId="5" pageNumber="353" type="diagnosis">
<paragraph id="32077C350023B925FF1D84A1002DF812" blockId="5.[151,1437,1645,2031]" pageId="5" pageNumber="353">
<emphasis id="00CCA0270023B925FF1D84A10079F97A" bold="true" box="[199,322,1717,1743]" pageId="5" pageNumber="353">Diagnosis.</emphasis>
The male of new species differs from those of all congeners except the Chinese species
<taxonomicName id="F5B807B60023B925FAF984A20071F946" authority="Lin &amp; Li, 2024" authorityName="Lin &amp; Li" authorityYear="2024" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="353" phylum="Arthropoda" rank="species" species="chongzu">
<emphasis id="00CCA0270023B925FAF984A204A7F97A" box="[1315,1436,1717,1743]" italics="true" pageId="5" pageNumber="353">T. chongzu</emphasis>
<bibRefCitation id="562901C40023B925FF4D84CD0071F946" author="Lin, Y. J. &amp; Li, S. Q. &amp; Mo, H. L. &amp; Wang, X. H." box="[151,330,1753,1779]" pageId="5" pageNumber="353" pagination="4 - 98" refId="ref4905" refString="Lin, Y. J., Li, S. Q., Mo, H. L. &amp; Wang, X. H. (2024) Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam. Zoological Systematics, 49, 4 - 98. https: // doi. org / 10.11865 / zs. 2024101" type="journal article" year="2024">
Lin &amp; Li,
<quantity id="F540D1D00023B925FEC884CD0071F946" box="[274,330,1753,1779]" metricMagnitude="1" metricUnit="m" metricValue="5.14096" pageId="5" pageNumber="353" unit="in" value="2024.0">2024</quantity>
</bibRefCitation>
</taxonomicName>
in
<bibRefCitation id="562901C40023B925FEA984CD030FF946" author="Lin, Y. J. &amp; Li, S. Q. &amp; Mo, H. L. &amp; Wang, X. H." box="[371,564,1753,1779]" pageId="5" pageNumber="353" pagination="4 - 98" refId="ref4905" refString="Lin, Y. J., Li, S. Q., Mo, H. L. &amp; Wang, X. H. (2024) Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam. Zoological Systematics, 49, 4 - 98. https: // doi. org / 10.11865 / zs. 2024101" type="journal article" year="2024">
Lin
<emphasis id="00CCA0270023B925FE7E84CE00DBF946" box="[420,480,1753,1779]" italics="true" pageId="5" pageNumber="353">et al.</emphasis>
(2024)
</bibRefCitation>
and
<taxonomicName id="F5B807B60023B925FDAA84CE0280F946" authority="Cai &amp; Zhong, 2021" authorityName="Cai &amp; Zhong" authorityYear="2021" box="[624,955,1753,1779]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="353" phylum="Arthropoda" rank="species" species="pinmi">
<emphasis id="00CCA0270023B925FDAA84CE03F4F946" box="[624,719,1754,1779]" italics="true" pageId="5" pageNumber="353">T. pinmi</emphasis>
<bibRefCitation id="562901C40023B925FD0384CD0280F946" author="Cai, C. H. &amp; Lin, Y. J. &amp; Zhong, Y." box="[729,955,1753,1779]" pageId="5" pageNumber="353" pagination="1 - 8" refId="ref4335" refString="Cai, C. H., Lin, Y. J. &amp; Zhong, Y. (2021) Description of a new species of Thelcticopis Karsch (Araneae, Sparassidae) from Guangxi Province, China. Biodiversity Data Journal, 9 (e 67437), 1 - 8. https: // doi. org / 10.3897 / BDJ. 9. e 67437" type="journal article" year="2021">
Cai &amp; Zhong,
<quantity id="F540D1D00023B925FC5984CD0280F946" box="[899,955,1753,1779]" metricMagnitude="1" metricUnit="m" metricValue="5.13334" pageId="5" pageNumber="353" unit="in" value="2021.0">2021</quantity>
</bibRefCitation>
</taxonomicName>
in
<bibRefCitation id="562901C40023B925FC3F84CD059EF946" author="Cai, C. H. &amp; Lin, Y. J. &amp; Zhong, Y." box="[997,1189,1753,1779]" pageId="5" pageNumber="353" pagination="1 - 8" refId="ref4335" refString="Cai, C. H., Lin, Y. J. &amp; Zhong, Y. (2021) Description of a new species of Thelcticopis Karsch (Araneae, Sparassidae) from Guangxi Province, China. Biodiversity Data Journal, 9 (e 67437), 1 - 8. https: // doi. org / 10.3897 / BDJ. 9. e 67437" type="journal article" year="2021">
Cai
<emphasis id="00CCA0270023B925FBCC84CE056AF946" box="[1046,1105,1753,1779]" italics="true" pageId="5" pageNumber="353">et al.</emphasis>
(2021)
</bibRefCitation>
in having a complex embolus (E) possessing several apophyses. The male of
<taxonomicName id="F5B807B60023B925FCD184EA02BFF8A2" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[779,900,1789,1815]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="353" phylum="Arthropoda" rank="species" species="ketambe" status="sp. nov.">
<emphasis id="00CCA0270023B925FCD184EA02BFF8A2" box="[779,900,1789,1815]" italics="true" pageId="5" pageNumber="353">T. ketambe</emphasis>
</taxonomicName>
<emphasis id="00CCA0270023B925FC5184E902DFF8A2" bold="true" box="[907,996,1789,1815]" pageId="5" pageNumber="353">
<taxonomicNameLabel id="1BFF1D5C0023B925FC5184E902DFF8A2" box="[907,996,1789,1815]" pageId="5" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from both species by an absence of median retrolateral tibial apophysis, instead of which is retrolateral tibial spine (RTS) present, blunt ventral (vRTA) and wide dorsal (dRTA) retrolateral tibial apophyses [vs. median retrolateral tibial apophysis present, vRTA pointed, dRTA narrow, whip-like; cf.
<figureCitation id="AA8360B00023B925FD3D857D025EF836" box="[743,869,1897,1923]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1394,1418]" captionTargetBox="[151,1429,381,1364]" captionTargetId="figure-56@5.[151,1436,381,1371]" captionTargetPageId="5" captionText="FIGURES 19–24. Thelcticopis ketambe sp. nov., male. 19–21 Palpal tibia (19 dorsal, 20 retrolateral, 21 ventral); 22–24 Bulb (22 prolateral, 23 ventral, 24 retrolateral). Scale bars: 0.5 mm. Abbreviations: C—conductor, DE—distal embolic apophysis, dRTA—dorsalretrolateraltibial apophysis,E—embolus,PE—prolateralembolicapophysis,RE—retrolateralembolic apophysis, RK—retrolateral keel, RTS—retrolateral tibial spine, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi="http://doi.org/10.5281/zenodo.14239949" httpUri="https://zenodo.org/record/14239949/files/figure.png" pageId="5" pageNumber="353">Figs 19–21</figureCitation>
and 51B in
<bibRefCitation id="562901C40023B925FC28857D0596F836" author="Lin, Y. J. &amp; Li, S. Q. &amp; Mo, H. L. &amp; Wang, X. H." box="[1010,1197,1897,1923]" pageId="5" pageNumber="353" pagination="4 - 98" refId="ref4905" refString="Lin, Y. J., Li, S. Q., Mo, H. L. &amp; Wang, X. H. (2024) Thirty-eight spider species (Arachnida: Araneae) from China, Indonesia, Japan and Vietnam. Zoological Systematics, 49, 4 - 98. https: // doi. org / 10.11865 / zs. 2024101" type="journal article" year="2024">
Lin
<emphasis id="00CCA0270023B925FBFB857E0561F836" box="[1057,1114,1897,1923]" italics="true" pageId="5" pageNumber="353">et al.</emphasis>
(2024)
</bibRefCitation>
and fig. 3B in
<bibRefCitation id="562901C40023B925FA83857D0030F812" author="Cai, C. H. &amp; Lin, Y. J. &amp; Zhong, Y." pageId="5" pageNumber="353" pagination="1 - 8" refId="ref4335" refString="Cai, C. H., Lin, Y. J. &amp; Zhong, Y. (2021) Description of a new species of Thelcticopis Karsch (Araneae, Sparassidae) from Guangxi Province, China. Biodiversity Data Journal, 9 (e 67437), 1 - 8. https: // doi. org / 10.3897 / BDJ. 9. e 67437" type="journal article" year="2021">
Cai
<emphasis id="00CCA0270023B925FA52857E0196F812" italics="true" pageId="5" pageNumber="353">et al</emphasis>
. (2021)
</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0023B926FF1D85A500A5FC8C" lastPageId="6" lastPageNumber="354" pageId="5" pageNumber="353" type="description">
<paragraph id="32077C350023B926FF1D85A50373FE64" blockId="5.[151,1437,1645,2031]" lastBlockId="6.[151,1437,151,861]" lastPageId="6" lastPageNumber="354" pageId="5" pageNumber="353">
<emphasis id="00CCA0270023B925FF1D85A50097F87E" bold="true" box="[199,428,1969,1995]" pageId="5" pageNumber="353">Description. Male:</emphasis>
Total length 17.0. Carapace: 8.5 long, 6.8 wide. Opisthosoma: 9.1 long, 5.7 wide. Coloration: Carapace and chelicerae brown. Labium and endites dark brown. Sternum brown. Coxae brown. Palps: Fe–Ti yellowish brown, Ta brown. Legs yellowish brown. Opisthosoma and spinnerets gray. Carapace, chelicerae, palps, legs and abdomen with light setae. Eye sizes and interdistances: AME 0.54, ALE 0.36, PME 0.27, PLE 0.31, AME–AME 0.47, AME–ALE 0.41, PME–PME 0.94, PME–PLE 0.86, AME–PME 0.24, ALE–PLE 0.36. Clypeus height at AME 0.43. Clypeus height at ALE 0.29. Leg and palp measurements: Palp: 3.0, 1.5, 1.6, -, 3.3 (9.4). Leg I: 8.3, 3.9, 8.1, 6.8, 1.8 (28.9). Leg II: 8.1, 3.8, 7.2, 6.0, 1.7 (26.8). Leg III: 6.3, 3.0, 5.0, 4.5, 1.5 (20.3). Leg IV: 8.1, 2.9, 6.9, 7.3, 2.0 (27.2). Leg formula: 1423. Leg and palp spination: Palp: Fe d3 p1 r1; Pa r1; Ti p2. Leg I: Fe d2 p3 r3; Ti d1 p3 r2 v2-2-2-2-2-2; Mt p2 r2 v2. Leg II: Fe d2 p3 r3; Ti d1 p2 r2 v2-2-2-2-2-1; Mt p2 r2 v2. Leg III: Fe d2 p3 r2; Ti d1 p2 r2 v2-2-2; Mt p3 r3 v2-1. Leg IV: Fe d2 p3 r1; Ti d1 p2 r2 v2-2-2; Mt p3 r3 v2-1. Chelicerae with 3 promarginal and 6 retromarginal teeth.
</paragraph>
<paragraph id="32077C350020B926FF1D83CF0010FCA0" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="354">
Male palp as shown in
<figureCitation id="AA8360B00020B926FE1C83CF037EFE40" box="[454,581,474,501]" captionStart-0="FIGURES 13–18" captionStart-1="FIGURES 19–24" captionStartId-0="4.[151,264,1740,1764]" captionStartId-1="5.[151,264,1394,1418]" captionTargetBox-0="[156,1426,248,1717]" captionTargetBox-1="[151,1429,381,1364]" captionTargetId-0="figure-30@4.[151,1436,237,1717]" captionTargetId-1="figure-56@5.[151,1436,381,1371]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 13–18. Thelcticopis ketambe sp. nov., male. 13–14 Habitus (13 dorsal, 14 ventral); 15–18 Palp (15 dorsal, 16, prolateral, 17 ventral, 18 retrolateral). Scale bars: 13–14=10 mm; 15–18=1 mm. Abbreviations: CS—cymbial scopula, RB— retrolateral bulge of cymbium." captionText-1="FIGURES 19–24. Thelcticopis ketambe sp. nov., male. 19–21 Palpal tibia (19 dorsal, 20 retrolateral, 21 ventral); 22–24 Bulb (22 prolateral, 23 ventral, 24 retrolateral). Scale bars: 0.5 mm. Abbreviations: C—conductor, DE—distal embolic apophysis, dRTA—dorsalretrolateraltibial apophysis,E—embolus,PE—prolateralembolicapophysis,RE—retrolateralembolic apophysis, RK—retrolateral keel, RTS—retrolateral tibial spine, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.14239947" figureDoi-1="http://doi.org/10.5281/zenodo.14239949" httpUri-0="https://zenodo.org/record/14239947/files/figure.png" httpUri-1="https://zenodo.org/record/14239949/files/figure.png" pageId="6" pageNumber="354">Figs 15–24</figureCitation>
. Tibia twice as long as wide. Dorsal retrolateral tibial apophysis (dRTA) with triangular tip. Ventral retrolateral tibial apophysis (
<emphasis id="00CCA0270020B926FD1983EB03C4FDAC" box="[707,767,511,537]" italics="true" pageId="6" pageNumber="354">vRTA</emphasis>
) cone-shaped. Retrolateral tibial spine (RTS) short and stout. Ventro-distal lobe (VDL) short, semicircular. Cymbium 1.2 times longer than wide, with rectangular retrolateral bulge (RB) and well developed cymbial scopula (CS) disto-dorsally. Bulb 1.2 times longer than wide. Tegulum (T) heart-shaped. Spermophor (Sp) S-shaped in retrolateral view. Tegular apophysis (TA) basally membranous, arising from T in 3-o’clock position, with mediad apex. Conductor (C) arising distally, with triangular retrolateral margin. Embolus (E) arising in 9-o’clock position, complex, with retrolateral keel (RK) and three apophyses: beak-shaped prolateral apophysis (PE), rounded retrolateral one (RE) and isosceles triangle-shaped distal one (DE). Tip of E beak-shaped.
</paragraph>
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<emphasis id="00CCA0270020B926FF1D810B0013FC8C" bold="true" box="[199,296,799,825]" pageId="6" pageNumber="354">Female:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0020B926FF1D815602B1FCE8" box="[199,906,834,861]" pageId="6" pageNumber="354" type="distribution">
<paragraph id="32077C350020B926FF1D815602B1FCE8" blockId="6.[151,1437,151,861]" box="[199,906,834,861]" pageId="6" pageNumber="354">
<emphasis id="00CCA0270020B926FF1D8156005AFCE9" bold="true" box="[199,353,834,860]" pageId="6" pageNumber="354">Distribution.</emphasis>
Only known from the
<typeStatus id="ED03C2970020B926FDBA815703ABFCE8" box="[608,656,835,861]" pageId="6" pageNumber="354">type</typeStatus>
locality (
<figureCitation id="AA8360B00020B926FD2081570245FCE8" box="[762,894,834,861]" captionStart="FIGURES 25–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[151,1436,885,1419]" captionTargetId="figure-340@6.[151,1436,885,1419]" captionTargetPageId="6" captionText="FIGURES 25–26. Distribution records of Thelcticopis spp. from Sumatra, Mentawai Islands and adjacent Malay Peninsula: T. karnyi (1), T. lizae sp. nov. (2), T. ketambe sp. nov. (3), T. klossi (4), T. orichalcea (5), T. modesta (6)." figureDoi="http://doi.org/10.5281/zenodo.14239953" httpUri="https://zenodo.org/record/14239953/files/figure.png" pageId="6" pageNumber="354">Figs 25–26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

229
data/BA/11/CD/BA11CD230027B924FF4D84FE02B1FF60.xml Normal file
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@ -0,0 +1,229 @@
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<mods:title id="2E6DCB26CADE411BA3621A13D68103E8">A survey of the Sumatran Sparassidae (Araneae). 1. Two new Thelcticopis species</mods:title>
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<mods:affiliation id="AE79479FFD1776D52F4EAD302509584D">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
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<mods:affiliation id="DDE4CD68F45E921DC24A5B3B317D140F">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia</mods:affiliation>
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<paragraph id="32077C350027B921FF4D84FE00F6F8B0" blockId="1.[151,461,1770,1833]" box="[151,461,1770,1797]" pageId="1" pageNumber="349">
<heading id="694FCB590027B921FF4D84FE00F6F8B0" bold="true" box="[151,461,1770,1797]" fontSize="11" level="1" pageId="1" pageNumber="349" reason="1">
<emphasis id="00CCA0270027B921FF4D84FE00F6F8B0" bold="true" box="[151,461,1770,1797]" pageId="1" pageNumber="349">
<taxonomicName id="F5B807B60027B921FF4D84FE0053F8B0" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[151,360,1770,1797]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="349" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="00CCA0270027B921FF4D84FE0053F8B0" bold="true" box="[151,360,1770,1797]" italics="true" pageId="1" pageNumber="349">Thelcticopis lizae</emphasis>
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<taxonomicNameLabel id="1BFF1D5C0027B921FEAA84FF00F6F8B0" box="[368,461,1771,1797]" pageId="1" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="7AA22FBE0027B921FF4D851B0061F89C" box="[151,346,1806,1833]" pageId="1" pageNumber="349" type="description">
<paragraph id="32077C350027B921FF4D851B0061F89C" blockId="1.[151,461,1770,1833]" box="[151,346,1806,1833]" pageId="1" pageNumber="349">
<figureCitation id="AA8360B00027B921FF4D851B0033F89C" box="[151,264,1807,1833]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="2.[151,264,1567,1591]" captionStartId-1="3.[151,264,1139,1163]" captionTargetBox-0="[160,1425,191,1543]" captionTargetBox-1="[151,1433,183,1110]" captionTargetId-0="figure-19@2.[151,1436,181,1543]" captionTargetId-1="figure-19@3.[151,1436,181,1116]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Thelcticopis lizae sp. nov., male. 1–2 Habitus (1 dorsal, 2 ventral); 3–6 Palp (3 dorsal, 4 prolateral, 5 ventral, 6 retrolateral). Scale bars: 1–2=5 mm; 3–6=1 mm. Abbreviations: CS—cymbial scopula, RB—retrolateral bulge of cymbium." captionText-1="FIGURES 7–12. Thelcticopis lizae sp. nov., male. 7–9 Palpal tibia (7 dorsal, 8 retrolateral, 9 ventral); 10–12 Bulb (10 prolateral, 11 ventral, 12 retrolateral). Scale bars: 0.5 mm. Abbreviations: bpC—basal process of conductor, C—conductor, E—embolus, RTA—retrolateral tibial apophysis, Se—bunch of setae, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.14239939" figureDoi-1="http://doi.org/10.5281/zenodo.14239943" httpUri-0="https://zenodo.org/record/14239939/files/figure.png" httpUri-1="https://zenodo.org/record/14239943/files/figure.png" pageId="1" pageNumber="349">Figs 1–12</figureCitation>
,
<figureCitation id="AA8360B00027B921FECE851A0061F89C" box="[276,346,1806,1833]" captionStart="FIGURES 25–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[151,1436,885,1419]" captionTargetId="figure-340@6.[151,1436,885,1419]" captionTargetPageId="6" captionText="FIGURES 25–26. Distribution records of Thelcticopis spp. from Sumatra, Mentawai Islands and adjacent Malay Peninsula: T. karnyi (1), T. lizae sp. nov. (2), T. ketambe sp. nov. (3), T. klossi (4), T. orichalcea (5), T. modesta (6)." figureDoi="http://doi.org/10.5281/zenodo.14239953" httpUri="https://zenodo.org/record/14239953/files/figure.png" pageId="1" pageNumber="349">25–26</figureCitation>
</paragraph>
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<paragraph id="32077C350027B921FF4D854302CDF820" blockId="1.[151,1436,1878,2013]" pageId="1" pageNumber="349">
<emphasis id="00CCA0270027B921FF4D8543007EF8C5" bold="true" box="[151,325,1878,1905]" pageId="1" pageNumber="349">Type material.</emphasis>
<materialsCitation id="82D076680027B921FE96854202CDF820" collectionCode="ISEA" country="Indonesia" elevation="1400" latitude="3.9833333" location="Kedah village" longLatPrecision="1307" longitude="97.25" pageId="1" pageNumber="349" specimenCount="1" specimenCount-male="1" stateProvince="Aceh" typeStatus="holotype">
<emphasis id="00CCA0270027B921FE9685420210F8C4" bold="true" box="[332,811,1878,1905]" pageId="1" pageNumber="349">
<collectingCountry id="4AAF3CA50027B921FE96854200D4F8C5" box="[332,495,1878,1904]" name="Indonesia" pageId="1" pageNumber="349">INDONESIA</collectingCountry>
:
<emphasis id="00CCA0270027B921FE2185420210F8C4" bold="true" box="[507,811,1878,1905]" italics="true" pageId="1" pageNumber="349">
<collectingRegion id="F07CB2D70027B921FE218542039EF8C4" box="[507,677,1878,1905]" country="Indonesia" name="Aceh" pageId="1" pageNumber="349">Aceh Province</collectingRegion>
(
<collectingRegion id="F07CB2D70027B921FD6F85430220F8C4" box="[693,795,1879,1905]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="349">Sumatra</collectingRegion>
):
</emphasis>
</emphasis>
<typeStatus id="ED03C2970027B921FCE8854302A1F8C4" box="[818,922,1879,1905]" pageId="1" pageNumber="349" type="holotype">Holotype</typeStatus>
<specimenCount id="24BEB7BC0027B921FC7885430282F8C4" box="[930,953,1879,1905]" count="1" pageId="1" pageNumber="349" type="male">♂</specimenCount>
,
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, [
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<elevation id="B9959B060027B921FA8C854301D5F820" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="1" pageNumber="349" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300– 1500 m</elevation>
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, 1988 (precise date unknown), unknown collector (
<collectionCode id="54A9E4F00027B921FCF4856F024BF820" box="[814,880,1915,1941]" pageId="1" pageNumber="349">ISEA</collectionCode>
, 001.9099).
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</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0027B921FF1D858B03A3F868" pageId="1" pageNumber="349" type="etymology">
<paragraph id="32077C350027B921FF1D858B03A3F868" blockId="1.[151,1436,1878,2013]" pageId="1" pageNumber="349">
<emphasis id="00CCA0270027B921FF1D858B0075F80C" bold="true" box="[199,334,1951,1977]" pageId="1" pageNumber="349">Etymology.</emphasis>
The specific name is a matronym in honor of the senior author’s wife, Elizaveta A. Nepaeva (Novoaltaysk,
<collectingCountry id="4AAF3CA50027B921FEE685D700B7F868" box="[316,396,1987,2013]" name="Russia" pageId="1" pageNumber="349">Russia</collectingCountry>
); name in genitive case.
</paragraph>
</subSubSection>
<caption id="66C72CBD0024B922FF4D840B0444F9EE" ID-DOI="http://doi.org/10.5281/zenodo.14239939" ID-Zenodo-Dep="14239939" httpUri="https://zenodo.org/record/14239939/files/figure.png" pageId="2" pageNumber="350" startId="2.[151,264,1567,1591]" targetBox="[160,1425,191,1543]" targetPageId="2" targetType="figure">
<paragraph id="32077C350024B922FF4D840B0444F9EE" blockId="2.[151,1437,1567,1627]" pageId="2" pageNumber="350">
<emphasis id="00CCA0270024B922FF4D840B0001F982" bold="true" box="[151,314,1567,1591]" pageId="2" pageNumber="350">FIGURES 1–6.</emphasis>
<taxonomicName id="F5B807B60024B922FE9A840B00C9F982" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[320,498,1567,1591]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="00CCA0270024B922FE9A840B00C9F982" box="[320,498,1567,1591]" italics="true" pageId="2" pageNumber="350">Thelcticopis lizae</emphasis>
</taxonomicName>
<emphasis id="00CCA0270024B922FE22840B0373F982" bold="true" box="[504,584,1567,1591]" pageId="2" pageNumber="350">
<taxonomicNameLabel id="1BFF1D5C0024B922FE22840B0373F982" box="[504,584,1567,1591]" pageId="2" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 1–2 Habitus (1 dorsal, 2 ventral); 3–6 Palp (3 dorsal, 4 prolateral, 5 ventral, 6 retrolateral). Scale bars: 1–2=5 mm; 3–6=1 mm. Abbreviations: CS—cymbial scopula, RB—retrolateral bulge of cymbium.
</paragraph>
</caption>
<subSubSection id="7AA22FBE0024B922FF1D84B40317F84D" pageId="2" pageNumber="350" type="diagnosis">
<paragraph id="32077C350024B922FF1D84B40317F84D" blockId="2.[151,1437,1696,2040]" pageId="2" pageNumber="350">
<emphasis id="00CCA0270024B922FF1D84B40079F90F" bold="true" box="[199,322,1696,1722]" pageId="2" pageNumber="350">Diagnosis.</emphasis>
This new species is similar to Indian
<taxonomicName id="F5B807B60024B922FD2E84B50519F90F" authority="Pocock, 1901" authorityName="Pocock" authorityYear="1901" box="[756,1058,1696,1722]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="00CCA0270024B922FD2E84B50245F90F" box="[756,894,1696,1722]" italics="true" pageId="2" pageNumber="350">T. bicornuta</emphasis>
<bibRefCitation id="562901C40024B922FC5C84B40519F90F" author="Pocock, R. I." box="[902,1058,1696,1722]" pageId="2" pageNumber="350" pagination="478 - 498" refId="ref5108" refString="Pocock, R. I. (1901) Descriptions of some new species of spiders from British India. Journal of the Bombay Natural History Society, 13, 478 - 498." type="journal article" year="1901">Pocock, 1901</bibRefCitation>
</taxonomicName>
, Taiwanese
<taxonomicName id="F5B807B60024B922FB7784B5001DF96B" authority="Zhu &amp; Zhong, 2020" authorityName="Zhu &amp; Zhong" authorityYear="2020" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="dahanensis">
<emphasis id="00CCA0270024B922FB7784B50473F90F" box="[1197,1352,1696,1722]" italics="true" pageId="2" pageNumber="350">T. dahanensis</emphasis>
<bibRefCitation id="562901C40024B922FA8B84B4001DF96B" author="Zhu, Y. &amp; Lin, Y. J. &amp; Zhong, Y." pageId="2" pageNumber="350" pagination="105 - 115" refId="ref5634" refString="Zhu, Y., Lin, Y. J. &amp; Zhong, Y. (2020) Two new and one newly recorded species of Thelcticopis Karsch, 1884 (Araneae, Sparassidae) from China. ZooKeys, 940, 105 - 115. https: // doi. org / 10.3897 / zookeys. 940.50764" type="journal article" year="2020">Zhu &amp; Zhong, 2020</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F5B807B60024B922FE8684D00346F96B" authority="(L. Koch, 1875)" baseAuthorityName="L. Koch" baseAuthorityYear="1875" box="[348,637,1731,1758]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="severa">
<emphasis id="00CCA0270024B922FE8684D000FBF968" box="[348,448,1732,1757]" italics="true" pageId="2" pageNumber="350">T. severa</emphasis>
(L.
<bibRefCitation id="562901C40024B922FE3584D7034FF96B" author="Koch, L." box="[495,628,1731,1758]" pageId="2" pageNumber="350" pagination="577 - 740" refId="ref4819" refString="Koch, L. (1875) Die Arachniden Australiens, nach der Natur beschrieben und abgebildet [Erster Theil. Lieferung 12 - 16]. Bauer &amp; Raspe, Nurnberg, pp. 577 - 740, pls. 44 - 65. https: // doi. org / 10.5962 / bhl. title. 121660" type="book chapter" year="1875">Koch, 1875</bibRefCitation>
)
</taxonomicName>
, which is widespread from
<collectingCountry id="4AAF3CA50024B922FC6A84D002CFF96B" box="[944,1012,1732,1758]" name="South Korea" pageId="2" pageNumber="350">Korea</collectingCountry>
to
<collectingCountry id="4AAF3CA50024B922FBC284D00569F96B" box="[1048,1106,1732,1758]" name="Laos" pageId="2" pageNumber="350">Laos</collectingCountry>
. The males of all four species have a simple whip-like embolus (E) in combination with a bunch of long parallel setae (Se) on the palpal tibia, which is twice as long as the retrolateral tibial apophysis (RTA). The male of
<taxonomicName id="F5B807B60024B922FC34851E0589F891" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[1006,1202,1802,1828]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="00CCA0270024B922FC34851E0589F891" box="[1006,1202,1802,1828]" italics="true" pageId="2" pageNumber="350">Thelcticopis lizae</emphasis>
</taxonomicName>
<emphasis id="00CCA0270024B922FB62851E042AF891" bold="true" box="[1208,1297,1802,1828]" pageId="2" pageNumber="350">
<taxonomicNameLabel id="1BFF1D5C0024B922FB62851E042AF891" box="[1208,1297,1802,1828]" pageId="2" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from those of
<taxonomicName id="F5B807B60024B922FF23853A00B9F8F2" authorityName="Pocock" authorityYear="1901" box="[249,386,1837,1863]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="00CCA0270024B922FF23853A00B9F8F2" box="[249,386,1837,1863]" italics="true" pageId="2" pageNumber="350">T. bicornuta</emphasis>
</taxonomicName>
and
<taxonomicName id="F5B807B60024B922FE60853A0324F8F2" baseAuthorityName="L. Koch" baseAuthorityYear="1875" box="[442,543,1838,1863]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="severa">
<emphasis id="00CCA0270024B922FE60853A0324F8F2" box="[442,543,1838,1863]" italics="true" pageId="2" pageNumber="350">T. severa</emphasis>
</taxonomicName>
by a distally positioned RTA which runs parallel to the longitudinal axis of the tibia [vs. medially or proximally located RTA with its basal part perpendicular to the tibia’s length axis; cf.
<figureCitation id="AA8360B00024B922FAE6854404A7F8DF" box="[1340,1436,1872,1898]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1139,1163]" captionTargetBox="[151,1433,183,1110]" captionTargetId="figure-19@3.[151,1436,181,1116]" captionTargetPageId="3" captionText="FIGURES 7–12. Thelcticopis lizae sp. nov., male. 7–9 Palpal tibia (7 dorsal, 8 retrolateral, 9 ventral); 10–12 Bulb (10 prolateral, 11 ventral, 12 retrolateral). Scale bars: 0.5 mm. Abbreviations: bpC—basal process of conductor, C—conductor, E—embolus, RTA—retrolateral tibial apophysis, Se—bunch of setae, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi="http://doi.org/10.5281/zenodo.14239943" httpUri="https://zenodo.org/record/14239943/files/figure.png" pageId="2" pageNumber="350">Figs 7–9</figureCitation>
and figs 5A–B in
<bibRefCitation id="562901C40024B922FEB885670365F83B" author="Sankaran, P. M. &amp; Sherwood, D. &amp; Jager, P." box="[354,606,1907,1934]" pageId="2" pageNumber="350" pagination="301 - 338" refId="ref5218" refString="Sankaran, P. M., Sherwood, D. &amp; Jager, P. (2024) On the identity of species of the huntsman spider genus Thelcticopis Karsch, 1884 (Araneae: Sparassidae: Sparianthinae) from India, Myanmar, Pakistan, and Sri Lanka. Zootaxa, 5463 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 5463.3.1" type="journal article" year="2024">
Sankaran
<emphasis id="00CCA0270024B922FE088560033FF838" box="[466,516,1907,1933]" italics="true" pageId="2" pageNumber="350">et al</emphasis>
. (2024)
</bibRefCitation>
and figs 2B–D in
<bibRefCitation id="562901C40024B922FCEB856702CCF83B" author="Zhu, Y. &amp; Lin, Y. J. &amp; Zhong, Y." box="[817,1015,1907,1934]" pageId="2" pageNumber="350" pagination="105 - 115" refId="ref5634" refString="Zhu, Y., Lin, Y. J. &amp; Zhong, Y. (2020) Two new and one newly recorded species of Thelcticopis Karsch, 1884 (Araneae, Sparassidae) from China. ZooKeys, 940, 105 - 115. https: // doi. org / 10.3897 / zookeys. 940.50764" type="journal article" year="2020">
Zhu
<emphasis id="00CCA0270024B922FCBC856002A3F838" box="[870,920,1907,1933]" italics="true" pageId="2" pageNumber="350">et al</emphasis>
. (2020)
</bibRefCitation>
]. The male of the new species differs from that of
<taxonomicName id="F5B807B60024B922FEFF85830084F804" authorityName="Zhu &amp; Zhong" authorityYear="2020" box="[293,447,1943,1969]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="dahanensis">
<emphasis id="00CCA0270024B922FEFF85830084F804" box="[293,447,1943,1969]" italics="true" pageId="2" pageNumber="350">T. dahanensis</emphasis>
</taxonomicName>
by the non-bifurcated RTA, relatively short E ending at 1-o’clock, and medially located conductor (C) [vs. bifurcated RTA, relatively long E ending at 3-o’clock and C located retrolaterally; cf.
<figureCitation id="AA8360B00024B922FAF485AE04A7F861" box="[1326,1436,1978,2004]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1139,1163]" captionTargetBox="[151,1433,183,1110]" captionTargetId="figure-19@3.[151,1436,181,1116]" captionTargetPageId="3" captionText="FIGURES 7–12. Thelcticopis lizae sp. nov., male. 7–9 Palpal tibia (7 dorsal, 8 retrolateral, 9 ventral); 10–12 Bulb (10 prolateral, 11 ventral, 12 retrolateral). Scale bars: 0.5 mm. Abbreviations: bpC—basal process of conductor, C—conductor, E—embolus, RTA—retrolateral tibial apophysis, Se—bunch of setae, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi="http://doi.org/10.5281/zenodo.14239943" httpUri="https://zenodo.org/record/14239943/files/figure.png" pageId="2" pageNumber="350">Figs 8, 11</figureCitation>
and figs 1B, D in
<bibRefCitation id="562901C40024B922FE8485CA0325F84D" author="Zhu, Y. &amp; Lin, Y. J. &amp; Zhong, Y." box="[350,542,2014,2040]" pageId="2" pageNumber="350" pagination="105 - 115" refId="ref5634" refString="Zhu, Y., Lin, Y. J. &amp; Zhong, Y. (2020) Two new and one newly recorded species of Thelcticopis Karsch, 1884 (Araneae, Sparassidae) from China. ZooKeys, 940, 105 - 115. https: // doi. org / 10.3897 / zookeys. 940.50764" type="journal article" year="2020">
Zhu
<emphasis id="00CCA0270024B922FE4885CB00FFF84D" box="[402,452,2014,2040]" italics="true" pageId="2" pageNumber="350">et al</emphasis>
. (2020)
</bibRefCitation>
].
</paragraph>
</subSubSection>
<caption id="66C72CBD0025B923FF4D8667003CFB4D" ID-DOI="http://doi.org/10.5281/zenodo.14239943" ID-Zenodo-Dep="14239943" httpUri="https://zenodo.org/record/14239943/files/figure.png" pageId="3" pageNumber="351" startId="3.[151,264,1139,1163]" targetBox="[151,1433,183,1110]" targetPageId="3" targetType="figure">
<paragraph id="32077C350025B923FF4D8667003CFB4D" blockId="3.[151,1437,1139,1272]" pageId="3" pageNumber="351">
<emphasis id="00CCA0270025B923FF4D8667007DFB3E" bold="true" box="[151,326,1139,1163]" pageId="3" pageNumber="351">FIGURES 7–12.</emphasis>
<taxonomicName id="F5B807B60025B923FE91866000C0FB39" authorityName="Fomichev &amp; Omelko" authorityYear="2024" box="[331,507,1140,1164]" class="Arachnida" family="Sparassidae" genus="Thelcticopis" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="351" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="00CCA0270025B923FE91866000C0FB39" box="[331,507,1140,1164]" italics="true" pageId="3" pageNumber="351">Thelcticopis lizae</emphasis>
</taxonomicName>
<emphasis id="00CCA0270025B923FDDA86600374FB39" bold="true" box="[512,591,1140,1164]" pageId="3" pageNumber="351">
<taxonomicNameLabel id="1BFF1D5C0025B923FDDA86600374FB39" box="[512,591,1140,1164]" pageId="3" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 7–9 Palpal tibia (7 dorsal, 8 retrolateral, 9 ventral); 10–12 Bulb (10 prolateral, 11 ventral, 12 retrolateral). Scale bars: 0.5 mm. Abbreviations: bpC—basal process of conductor, C—conductor, E—embolus, RTA—retrolateral tibial apophysis, Se—bunch of setae, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe.
</paragraph>
</caption>
<subSubSection id="7AA22FBE0025B924FF1D872F00A5FF04" lastPageId="4" lastPageNumber="352" pageId="3" pageNumber="351" type="description">
<paragraph id="32077C350025B923FF1D872F045FF903" blockId="3.[151,1437,1339,2035]" pageId="3" pageNumber="351">
<emphasis id="00CCA0270025B923FF1D872F0098FAE3" bold="true" box="[199,419,1339,1366]" pageId="3" pageNumber="351">Description. Male:</emphasis>
Total length 13.0. Carapace: 5.8 long, 4.8 wide. Opisthosoma:6.1 long, 3.5 wide. Coloration: Carapace and chelicerae brown. Labium and endites dark brown. Sternum brown. Coxae yellow brown. Palps: Fe–Pa yellowish brown, Ti–Ta brown. Legs: Fe yellowish brown, Pa–Mt brown, Tr yellowish brown. Abdomen brown dorsally, gray ventrally. Spinnerets brown gray. Carapace, chelicerae, palps, legs and opisthosoma with light setae. Eye sizes and interdistances: AME 0.41, ALE 0.26, PME 0.21, PLE 0.36, AME–AME 0.26, AME–ALE 0.24, PME–PME 0.57, PME–PLE 0.51, AME–PME 0.21, ALE–PLE 0.17. Clypeus height at AME 0.36. Clypeus height at ALE 0.14. Leg and palp measurements: Palp: 2.2, 1.1, 1.1, -, 2.4 (6.8). Leg I: 5.7, 2.8, 5.7, 4.7, 1.3 (20.2). Leg II: 5.8, 2.6, 5.2, 4.4, 1.2 (19.2). Leg III: 4.8, 2.2, 3.8, 3.4, 1.2 (15.4). Leg IV: 5.7, 2.1, 4.3, 5.3, 1.5 (18.9). Leg formula: 1243. Leg and palp spination: Palp: Fe d3 p1r1; Pa p1 r1; Ti p3. Leg I: Fe d2 p3 r3; Ti p2 r2 v2-2-2-2-2-2; Mt p2 r2 v2. Leg II: Fe d2 p3 r3; Ti p2 r2 v2-2-2-2-2-2; Mt p3 r3 v2. Leg III: Fe d2 p3 r2; Ti p2 r2 v2-2-2; Mt p3 r3 v2-1-1. Leg IV: Fe d2 p3 r1; Ti p2 r2 v2-2-2; Mt p3 r3 v2-2. Chelicerae with 3 promarginal and 5 retromarginal teeth.
</paragraph>
<paragraph id="32077C350025B923FF1D84AB0038F846" blockId="3.[151,1437,1339,2035]" pageId="3" pageNumber="351">
Male palp as shown in
<figureCitation id="AA8360B00025B923FE1E84AB030EF96C" box="[452,565,1727,1753]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStartId-0="2.[151,264,1567,1591]" captionStartId-1="3.[151,264,1139,1163]" captionTargetBox-0="[160,1425,191,1543]" captionTargetBox-1="[151,1433,183,1110]" captionTargetId-0="figure-19@2.[151,1436,181,1543]" captionTargetId-1="figure-19@3.[151,1436,181,1116]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–6. Thelcticopis lizae sp. nov., male. 1–2 Habitus (1 dorsal, 2 ventral); 3–6 Palp (3 dorsal, 4 prolateral, 5 ventral, 6 retrolateral). Scale bars: 1–2=5 mm; 3–6=1 mm. Abbreviations: CS—cymbial scopula, RB—retrolateral bulge of cymbium." captionText-1="FIGURES 7–12. Thelcticopis lizae sp. nov., male. 7–9 Palpal tibia (7 dorsal, 8 retrolateral, 9 ventral); 10–12 Bulb (10 prolateral, 11 ventral, 12 retrolateral). Scale bars: 0.5 mm. Abbreviations: bpC—basal process of conductor, C—conductor, E—embolus, RTA—retrolateral tibial apophysis, Se—bunch of setae, Sp—spermophor, T—tegulum, TA—tegular apophysis, VDL—ventrodistal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.14239939" figureDoi-1="http://doi.org/10.5281/zenodo.14239943" httpUri-0="https://zenodo.org/record/14239939/files/figure.png" httpUri-1="https://zenodo.org/record/14239943/files/figure.png" pageId="3" pageNumber="351">Figs 3–12</figureCitation>
. Tibia slightly longer than wide. Retrolateral tibial apophysis (RTA) consisting of single prong, twice shorter than tibia. Ventro-distal lobe (VDL) short, triangular. Base of RTA with three stiff parallel fused setae forming a single bunch (Se). Cymbium 1.7 times longer than wide, with a pronounced retrolateral bulge (RB) and well developed cymbial scopula (CS) disto-dorsally. Bulb 1.4 times longer than wide. Tegulum (T) large and complex, with spermophor (Sp) forming S-shape in ventral view. Tegular apophysis (TA) arising from T at 3-o’clock position, with mediad apex. Conductor (C) arising medially, rectangular in ventral view, not covering the apex of embolus in ventral view. Basal process of conductor (bpC) almost triangular. Embolus (E) relatively short, originating prolaterally at 8-o’clock, with slightly widened tip, this latter resting into a recess on a prolateral side of C.
</paragraph>
<paragraph id="32077C350022B924FF1D828300A5FF04" blockId="4.[199,906,151,213]" box="[199,414,151,177]" pageId="4" pageNumber="352">
<emphasis id="00CCA0270022B924FF1D82830013FF04" bold="true" box="[199,296,151,177]" pageId="4" pageNumber="352">Female:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="7AA22FBE0022B924FF1D82AE02B1FF60" box="[199,906,186,213]" pageId="4" pageNumber="352" type="distribution">
<paragraph id="32077C350022B924FF1D82AE02B1FF60" blockId="4.[199,906,151,213]" box="[199,906,186,213]" pageId="4" pageNumber="352">
<emphasis id="00CCA0270022B924FF1D82AE005AFF61" bold="true" box="[199,353,186,212]" pageId="4" pageNumber="352">Distribution.</emphasis>
Only known from the
<typeStatus id="ED03C2970022B924FDBA82AF03ABFF60" box="[608,656,187,213]" pageId="4" pageNumber="352">type</typeStatus>
locality (
<figureCitation id="AA8360B00022B924FD2082AF0245FF60" box="[762,894,186,213]" captionStart="FIGURES 25–26" captionStartId="6.[151,264,1442,1466]" captionTargetBox="[151,1436,885,1419]" captionTargetId="figure-340@6.[151,1436,885,1419]" captionTargetPageId="6" captionText="FIGURES 25–26. Distribution records of Thelcticopis spp. from Sumatra, Mentawai Islands and adjacent Malay Peninsula: T. karnyi (1), T. lizae sp. nov. (2), T. ketambe sp. nov. (3), T. klossi (4), T. orichalcea (5), T. modesta (6)." figureDoi="http://doi.org/10.5281/zenodo.14239953" httpUri="https://zenodo.org/record/14239953/files/figure.png" pageId="4" pageNumber="352">Figs 25–26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

254
data/BA/5C/D3/BA5CD33291E47170983F34AE88C510E2.xml
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@ -1,219 +1,219 @@
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<treatment ID-GBIF-Taxon="152047455" LSID="urn:lsid:plazi:treatment:BA5CD33291E47170983F34AE88C510E2" httpUri="http://treatment.plazi.org/id/BA5CD33291E47170983F34AE88C510E2" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection pageId="1" pageNumber="2" type="nomenclature">
<paragraph pageId="1" pageNumber="2">
<taxonomicName LSID="http://species-id.net/wiki/Hoplatessara_luxuriosa" authority="Silvestri, 1895" authorityName="Silvestri" authorityYear="1895" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa (Silvestri, 1895)</taxonomicName>
<treatment id="BA5CD33291E47170983F34AE88C510E2" ID-GBIF-Taxon="152047455" LSID="urn:lsid:plazi:treatment:BA5CD33291E47170983F34AE88C510E2" httpUri="http://treatment.plazi.org/id/BA5CD33291E47170983F34AE88C510E2" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
<subSubSection id="6D60686B63981C3017C488DA5227538D" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="2B6E69C45B6F46459D49E15D92B27A2D" pageId="1" pageNumber="2">
<taxonomicName id="B2BF8E2B79475DECFB3491BB4768EAA5" ID-CoL="3MJW3" LSID="http://species-id.net/wiki/Hoplatessara_luxuriosa" authority="Silvestri, 1895" authorityName="Silvestri" authorityYear="1895" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa (Silvestri, 1895)</taxonomicName>
Figs 1-3, 5-8
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="reference_group">
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation author="Silvestri, F" journalOrPublisher="Annali del Museo Civico di Storia Naturale, Genova" pageId="5" pageNumber="6" pagination="619 - 659" title="Chilopodi e Diplopodi della Papuasia." volume="34" year="1895">Silvestri 1895</bibRefCitation>
<subSubSection id="92DC4EB01B96BB0C88F5674E111B1F76" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="3698BE43DA7AD6771A137E9E1E557460" pageId="1" pageNumber="2">
<taxonomicName id="0838ED67627004CD2B18EE0A0BC06E8E" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation id="9289A730C6C0D33FCE554DD34294340C" author="Silvestri, F" journalOrPublisher="Annali del Museo Civico di Storia Naturale, Genova" pageId="5" pageNumber="6" pagination="619 - 659" title="Chilopodi e Diplopodi della Papuasia." volume="34" year="1895">Silvestri 1895</bibRefCitation>
: 643.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation author="Attems, C" journalOrPublisher="Mathematisch-Naturwissenschaftliche Klasse" pageId="4" pageNumber="5" pagination="221 - 482" title="System der Polydesmiden. I Theil. Denkschriften der kaiserlichen Akademie der Wissenschaften." volume="67" year="1898">Attems 1898</bibRefCitation>
<paragraph id="79053DBA4C7F9A4FB1AF8FA5A4B66015" pageId="1" pageNumber="2">
<taxonomicName id="42314ECB58A8F9AF96BD48880A28CEDC" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation id="BAB232484E28473746118B1C38E92AE9" author="Attems, C" journalOrPublisher="Mathematisch-Naturwissenschaftliche Klasse" pageId="4" pageNumber="5" pagination="221 - 482" title="System der Polydesmiden. I Theil. Denkschriften der kaiserlichen Akademie der Wissenschaften." volume="67" year="1898">Attems 1898</bibRefCitation>
: 318.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation author="Attems, C" journalOrPublisher="Archiv fuer Naturgeschichte (A)" pageId="5" pageNumber="6" pagination="1 - 398" title="Die indo-australischen Myriopoden." volume="80" year="1914">Attems 1914</bibRefCitation>
<paragraph id="86142F7D40D4DDE332BC0F2BDC51C903" pageId="1" pageNumber="2">
<taxonomicName id="B24AD637A26379F54309A39240D89FBB" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation id="62DA18BD4E4C0F1E735F277AD958677F" author="Attems, C" journalOrPublisher="Archiv fuer Naturgeschichte (A)" pageId="5" pageNumber="6" pagination="1 - 398" title="Die indo-australischen Myriopoden." volume="80" year="1914">Attems 1914</bibRefCitation>
: 221.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Strongylosoma</taxonomicName>
<paragraph id="E8C3038B99A4C5C0486887E7D2BF3047" pageId="1" pageNumber="2">
<taxonomicName id="DC162FF24BE90C47455C7E9E3B38F9D7" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Strongylosoma</taxonomicName>
(?) luxuriosum
<bibRefCitation author="Attems, C" journalOrPublisher="Zoologie" pageId="5" pageNumber="6" pagination="1 - 37" title="Myriopoden von Neu-Guinea II, gesammelt waehrend der Expeditionen 1904 - 1909. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la nouvelle-guinee en 1912 et 1913." volume="13" year="1915">Attems 1915</bibRefCitation>
<bibRefCitation id="0C434E82A5C4E98DDB8C0F5581C41B8E" author="Attems, C" journalOrPublisher="Zoologie" pageId="5" pageNumber="6" pagination="1 - 37" title="Myriopoden von Neu-Guinea II, gesammelt waehrend der Expeditionen 1904 - 1909. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la nouvelle-guinee en 1912 et 1913." volume="13" year="1915">Attems 1915</bibRefCitation>
: 22.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="5" pageNumber="6" pagination="1 - 269" title="The Myriopoda of the Australian region." volume="64" year="1920">Chamberlin 1920</bibRefCitation>
<paragraph id="5C15664ADABFFD5E881A0045534A88D4" pageId="1" pageNumber="2">
<taxonomicName id="DE6DA5EEA2EDDC0714E9CB051A1E3F11" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Strongylosoma luxuriosum</taxonomicName>
<bibRefCitation id="C8A5E391C987B1FB31BDC2BDA4F16A3E" author="Chamberlin, RV" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="5" pageNumber="6" pagination="1 - 269" title="The Myriopoda of the Australian region." volume="64" year="1920">Chamberlin 1920</bibRefCitation>
: 119.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Antichiropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antichiropus luxuriosus" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosus">Antichiropus luxuriosus</taxonomicName>
<bibRefCitation author="Attems, C" journalOrPublisher="Das Tierreich" pageId="5" pageNumber="6" pagination="1 - 300" title="Myriapoda. 3. Polydesmoidea. I. Fam. Strongylosomidae." volume="68" year="1937">Attems 1937</bibRefCitation>
<paragraph id="0FAD288EDD815C90DA946CAFBA410D09" pageId="1" pageNumber="2">
<taxonomicName id="C4DEE60E690964C6402E480D9D3B398F" class="Diplopoda" family="Paradoxosomatidae" genus="Antichiropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antichiropus luxuriosus" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosus">Antichiropus luxuriosus</taxonomicName>
<bibRefCitation id="877CAF695506893327CD822FC90827A4" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="5" pageNumber="6" pagination="1 - 300" title="Myriapoda. 3. Polydesmoidea. I. Fam. Strongylosomidae." volume="68" year="1937">Attems 1937</bibRefCitation>
: 271. [comb. n.]
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Hoplatessara luxuriosum</taxonomicName>
<paragraph id="9FEF975F5D84C3F0F726540E6798B5B5" pageId="1" pageNumber="2">
<taxonomicName id="99B08D27BC12F9A6E189062248DC213C" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosum" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosum">Hoplatessara luxuriosum</taxonomicName>
[sic]
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel 1956</bibRefCitation>
<bibRefCitation id="70E6564F6D1BBEADEB39D7875F7803C6" author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel 1956</bibRefCitation>
: 184; figs 1-4, p. 186. [comb. n.]
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="5" pageNumber="6" pagination="343 - 379" title="A revision of the Papuan Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa." volume="110" year="1967">Jeekel 1967</bibRefCitation>
<paragraph id="A8C1026DB52FDF3CB7FC458007E2E192" pageId="1" pageNumber="2">
<taxonomicName id="DF7BF5B0CB3F97B159DE70469EFDD6BE" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation id="B518E734CCCCA9E595249A332D22300B" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="5" pageNumber="6" pagination="343 - 379" title="A revision of the Papuan Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa." volume="110" year="1967">Jeekel 1967</bibRefCitation>
: 377. [Lectotype chosen]
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="C. A. W. Jeekel, Rotterdam" pageId="5" pageNumber="6" title="On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
<paragraph id="19D770396114BCA44E7BFE392AA1FC36" pageId="1" pageNumber="2">
<taxonomicName id="59550925B022B22618517AB4A2B4E2E4" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation id="7764915E92E2E2A6146CCD8A39E3CE20" author="Jeekel, CAW" journalOrPublisher="C. A. W. Jeekel, Rotterdam" pageId="5" pageNumber="6" title="On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
: 23, 25.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Verslagen en Technische Gegevens, Instituut voor Taxonomische Zooelogie (Zooelogisch Museum), Universiteit van Amsterdam" pageId="5" pageNumber="6" pagination="1 - 59" title="Australia Expedition 1980; legit C. A. W. Jeekel and A. M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania." volume="30" year="1981">Jeekel 1981</bibRefCitation>
<paragraph id="B39ABEF4211F68109EAE5EA5DCE53372" pageId="1" pageNumber="2">
<taxonomicName id="48901326027AE7BC7610D5B91CEC0981" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation id="BBD730B52752874058D1F8687E68B757" author="Jeekel, CAW" journalOrPublisher="Verslagen en Technische Gegevens, Instituut voor Taxonomische Zooelogie (Zooelogisch Museum), Universiteit van Amsterdam" pageId="5" pageNumber="6" pagination="1 - 59" title="Australia Expedition 1980; legit C. A. W. Jeekel and A. M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania." volume="30" year="1981">Jeekel 1981</bibRefCitation>
: 51.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Records of the Australian Museum" pageId="5" pageNumber="6" pagination="19 - 44" title="Millipedes from Australia, 6: Australiosomatini from Victoria (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://australianmuseum.net.au/journal/Jeekel-1984-Rec-Aust-Mus-361-1944" volume="36" year="1984">Jeekel 1984</bibRefCitation>
<paragraph id="539EB10CCEA3F95648D48DB33F3F8A29" pageId="1" pageNumber="2">
<taxonomicName id="8712D53EFDD5CFD5ED3D43BF59797EC0" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation id="7E6AAFD5CF827FC0C637D4F0C75F6B6E" author="Jeekel, CAW" journalOrPublisher="Records of the Australian Museum" pageId="5" pageNumber="6" pagination="19 - 44" title="Millipedes from Australia, 6: Australiosomatini from Victoria (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://australianmuseum.net.au/journal/Jeekel-1984-Rec-Aust-Mus-361-1944" volume="36" year="1984">Jeekel 1984</bibRefCitation>
: 38, 43.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="5" pageNumber="6" pagination="28 - 45" title="Millipedes from Australia, 16: Further notes on Australiosomatinae from New South Wales with descriptions of a new genus and four new species (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="6" year="2003">Jeekel 2003</bibRefCitation>
<paragraph id="F7B0A47A3E32C46A27F4E95C4E0DD79C" pageId="1" pageNumber="2">
<taxonomicName id="25365A6A50054EA431DC718A537B0DE8" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<bibRefCitation id="5AC27EC5C854ED1136F8778884F735DA" author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="5" pageNumber="6" pagination="28 - 45" title="Millipedes from Australia, 16: Further notes on Australiosomatinae from New South Wales with descriptions of a new genus and four new species (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="6" year="2003">Jeekel 2003</bibRefCitation>
: 28.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="lectotype">
<paragraph pageId="1" pageNumber="2">Lectotype.</paragraph>
<paragraph pageId="1" pageNumber="2">
<subSubSection id="9E73F3B1C44B9A7C4C8CA35853168229" pageId="1" pageNumber="2" type="lectotype">
<paragraph id="724E4949E76C1F4EF3DFEFB08815C09D" pageId="1" pageNumber="2">Lectotype.</paragraph>
<paragraph id="6B8C9D6CFD12331214464C087DB3D876" pageId="1" pageNumber="2">
Male, lacking segments 8 and 9, Sorong, New Guinea [see Discussion], L.M.
<normalizedToken originalValue="D’Albertis">D'Albertis</normalizedToken>
<normalizedToken id="4315D3107924AA35E2CE76592922D835" originalValue="D’Albertis">D'Albertis</normalizedToken>
, MCG.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="paralectotypes">
<paragraph pageId="1" pageNumber="2">Paralectotypes.</paragraph>
<paragraph pageId="1" pageNumber="2">1 female, 1 female fragment, 1 male fragment, same details, MCG.</paragraph>
<subSubSection id="6C963F5433DE4B0FBA552908B77A047A" pageId="1" pageNumber="2" type="paralectotypes">
<paragraph id="96B1B40D581F82F751492C3BE67ED584" pageId="1" pageNumber="2">Paralectotypes.</paragraph>
<paragraph id="4867F8B4B2284CA64BA988239B954AB4" pageId="1" pageNumber="2">1 female, 1 female fragment, 1 male fragment, same details, MCG.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="material examined">
<paragraph pageId="1" pageNumber="2">Material examined.</paragraph>
<paragraph pageId="1" pageNumber="2">
<subSubSection id="BD9C1F936DA4D3FD7017B59A2CF78C2F" pageId="1" pageNumber="2" type="material examined">
<paragraph id="265EA50FAED043C63F588B8A94601571" pageId="1" pageNumber="2">Material examined.</paragraph>
<paragraph id="49621965BAB358B42889F1F64A9A5C36" pageId="1" pageNumber="2">
1 male, Sunny Corner State Forest near Bathurst, NSW,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-33.4">33°24'S</geoCoordinate>
<geoCoordinate id="7A15705CCFDFB145C596B075F8A1DB51" direction="south" orientation="latitude" precision="925" value="-33.4">33°24'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="149.85">149°51'E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="2BCE294B28B424FF51CB765605D65619" direction="east" orientation="longitude" precision="925" value="149.85">149°51'E</geoCoordinate>
<normalizedToken id="E5A7877CDB5A93DDFCCB2FF04BF1BBB0" originalValue="±">+/-</normalizedToken>
2 km, 4 December 1972, J.S. Disney, under pine log, AM KS.18542; 1 male, near Merrill, SW of Crookwell, NSW [locality text not on label],
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-34.666668">34°40'S</geoCoordinate>
<geoCoordinate id="B3462D19A456AFF9A931053F90D03B8E" direction="south" orientation="latitude" precision="925" value="-34.666668">34°40'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="149.28334">149°17'E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="096C6A98EAD3FFC4EDF2B94A7950CC45" direction="east" orientation="longitude" precision="925" value="149.28334">149°17'E</geoCoordinate>
<normalizedToken id="DC0623B069B2CBC5F228E88F79168010" originalValue="±">+/-</normalizedToken>
2 km, 20 April 1990, [collector uncertain], AM KS.106310; 6 males, 2 females, 1 juvenile, 5 km NE of Colo, NSW [locality text not on label],
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-33.75">33°45'S</geoCoordinate>
<geoCoordinate id="F0FC55D6150409D492ECABC260512EB8" direction="south" orientation="latitude" precision="925" value="-33.75">33°45'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="149.28334">149°17'E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="8E12DF1287110AF0937BA5D8AC731D2B" direction="east" orientation="longitude" precision="925" value="149.28334">149°17'E</geoCoordinate>
<normalizedToken id="471BF8288A08A7088C8FDF2C0FCD5B0F" originalValue="±">+/-</normalizedToken>
2 km, 653 m a.s.l., 20 April 1990, L. Kirwan, AM KS.106320; 2 males, Sunny Corner near Bathurst, NSW,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-33.4">33°24'S</geoCoordinate>
<geoCoordinate id="4B765082BFEF86BD7B7D80BBADCC0213" direction="south" orientation="latitude" precision="925" value="-33.4">33°24'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="149.88333">149°53'E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="E2986FF642D67451B488F001A6385020" direction="east" orientation="longitude" precision="925" value="149.88333">149°53'E</geoCoordinate>
<normalizedToken id="7D233E3F2DB16E745D7E39B23679D2E9" originalValue="±">+/-</normalizedToken>
2 km, 24-27 January 1997, S.J. Fellenberg, AM KS.96088; 1 male, 1 female, 1 stadium 7 female, Daylight Creek Road near Sunny Corner, NSW,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-33.364166">33°21'51&quot;S</geoCoordinate>
<geoCoordinate id="9B3C8140FFA104E5CD57F35ABB1522D2" direction="south" orientation="latitude" precision="15" value="-33.364166">33°21'51&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="149.89417">149°53'39&quot;E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="17E20681DF29B198B0616CA7157874A5" direction="east" orientation="longitude" precision="15" value="149.89417">149°53'39&quot;E</geoCoordinate>
<normalizedToken id="D1B976B3EFAF2ACA41073DD565387F88" originalValue="±">+/-</normalizedToken>
100 m, 1050 m a.s.l., 30 April 2013, R. Mesibov and T. Moule, AM KS.120531; 1 male, 1 female, Sunny Corner Road near Sunny Corner, NSW,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-33.399445">33°23'58&quot;S</geoCoordinate>
<geoCoordinate id="43DF5B3D58947D3FFEB69757B5D277D2" direction="south" orientation="latitude" precision="15" value="-33.399445">33°23'58&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="149.9061">149°54'22&quot;E</geoCoordinate>
<normalizedToken originalValue="±">+/-</normalizedToken>
<geoCoordinate id="3826874A3495975503FA5E23C011DE9F" direction="east" orientation="longitude" precision="15" value="149.9061">149°54'22&quot;E</geoCoordinate>
<normalizedToken id="B08FBC027CE94E1B55C7A43704EFB74C" originalValue="±">+/-</normalizedToken>
25 m, 1210 m a.s.l., same date and collectors, AM KS.120532.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph pageId="1" pageNumber="2">Description.</paragraph>
<paragraph pageId="1" pageNumber="2">
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel (1956</bibRefCitation>
<subSubSection id="6C17AFD68363121EE5B1198112A0B088" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="63A4D8497489EBFF81DDE439754F7473" pageId="1" pageNumber="2">Description.</paragraph>
<paragraph id="BE1AF947746524A21954917F3584A749" pageId="1" pageNumber="2">
<bibRefCitation id="43FFC72E940AC892DF0486DB35283F42" author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel (1956</bibRefCitation>
,
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="5" pageNumber="6" pagination="343 - 379" title="A revision of the Papuan Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa." volume="110" year="1967">1967</bibRefCitation>
<bibRefCitation id="0A6EDA29B887EE50D21E5E0E72473CF3" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="5" pageNumber="6" pagination="343 - 379" title="A revision of the Papuan Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa." volume="110" year="1967">1967</bibRefCitation>
) gave admirably complete redescriptions of the specimens examined by
<bibRefCitation author="Silvestri, F" journalOrPublisher="Annali del Museo Civico di Storia Naturale, Genova" pageId="5" pageNumber="6" pagination="619 - 659" title="Chilopodi e Diplopodi della Papuasia." volume="34" year="1895">Silvestri (1895)</bibRefCitation>
<bibRefCitation id="080527DD6C8F2847C2C176114A0B1F26" author="Silvestri, F" journalOrPublisher="Annali del Museo Civico di Storia Naturale, Genova" pageId="5" pageNumber="6" pagination="619 - 659" title="Chilopodi e Diplopodi della Papuasia." volume="34" year="1895">Silvestri (1895)</bibRefCitation>
, and here we add only a few details:
</paragraph>
<paragraph lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<paragraph id="068B8AEFA60E8DC915DCF859BA343C49" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
Live and freshly preserved males and females (Figs 1, 2) with pale brownish-yellow ground colour, lighter ventrally; darker brown on prozonites and anterior portion of metazonites, darkest at waist and dorsally; head dark brown dorsally, lightening ven
<pageBreakToken pageId="2" pageNumber="3" start="start">trally</pageBreakToken>
<pageBreakToken id="8645756064BB48287D3F0F8625FA8E68" pageId="2" pageNumber="3" start="start">trally</pageBreakToken>
; collum dark brown ringed with pale brownish-yellow; antennae dark brown, lighter basally; legs with coxae, prefemur and basal portion of femur pale yellow, and postfemur, tibia and tarsus brown, darkening distally; preanal ring dark brown with
<pageBreakToken pageId="3" pageNumber="4" start="start">pale</pageBreakToken>
<pageBreakToken id="68092AB54E95D0DD18DD4C52636CBEC5" pageId="3" pageNumber="4" start="start">pale</pageBreakToken>
yellow epiproct; hypoproct light brown, anal valves dark brown ringed with pale brownish-yellow.
</paragraph>
<paragraph pageId="3" pageNumber="4">Small pleural keels on female rings 2-4, more prominent on rings 3 and 4; traces of keels on male rings 2-4.</paragraph>
<paragraph pageId="3" pageNumber="4">Male ring 6 sternite with transverse brushes of long setae between legpairs 6 and 7. Male ring 5 sternite with sparse transverse brush of long setae between legpair 5, well-separated from sternal lamella between legpair 4.</paragraph>
<paragraph pageId="3" pageNumber="4">Spiracles on diplosegments of males and females well-separated (Fig. 3); posterior spiracle crater-like, anterior spiracle rim oval (long axis more or less dorsoventral), dorsal portion of rim extended posterolaterally around emergent, finely textured, subspherical spiracle.</paragraph>
<paragraph pageId="3" pageNumber="4">
<paragraph id="72F71EFBAF5B4DCAB64C75DC16417E00" pageId="3" pageNumber="4">Small pleural keels on female rings 2-4, more prominent on rings 3 and 4; traces of keels on male rings 2-4.</paragraph>
<paragraph id="0AF0579B7FF5B8862372424C9D1575D0" pageId="3" pageNumber="4">Male ring 6 sternite with transverse brushes of long setae between legpairs 6 and 7. Male ring 5 sternite with sparse transverse brush of long setae between legpair 5, well-separated from sternal lamella between legpair 4.</paragraph>
<paragraph id="ACC2EB015AD0BDC8C964B4C216266154" pageId="3" pageNumber="4">Spiracles on diplosegments of males and females well-separated (Fig. 3); posterior spiracle crater-like, anterior spiracle rim oval (long axis more or less dorsoventral), dorsal portion of rim extended posterolaterally around emergent, finely textured, subspherical spiracle.</paragraph>
<paragraph id="04FDFC1D823281DF04801D9A407E18FA" pageId="3" pageNumber="4">
Gonopods (Figs 5-8) as described by
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel (1956)</bibRefCitation>
<bibRefCitation id="B68ACEAF5D7BB73EEEB5FA0F3E909EF1" author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel (1956)</bibRefCitation>
and illustrated in posterior view (Fig. 4 in
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel 1956</bibRefCitation>
<bibRefCitation id="25ECD1701263B421910530217F4F7939" author="Jeekel, CAW" journalOrPublisher="Amsterdam" pageId="5" pageNumber="6" pagination="184 - 188" title="On the generic status of Strongylosoma luxuriosum Silvestri 1894 from New Guinea (Diplopoda, Polydesmida, Strongylosomidae)." volumeTitle="Entomologische Berichten" year="1956">Jeekel 1956</bibRefCitation>
); the slight bulge on the medial side of the femoral process is partly obscured in posterior view (Fig. 5) and is shown more clearly in anterior view (Fig. 7).
</paragraph>
<caption pageId="3" pageNumber="4">
<paragraph pageId="3" pageNumber="4">
<caption id="DE39BB081F257441B13263DF52B1E643" pageId="3" pageNumber="4">
<paragraph id="64C616DB1A9EB39CB0A762F9EEAEB920" pageId="3" pageNumber="4">
Figures 5-8.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<taxonomicName id="9088DEACFF688F583193E3ACE9AFEA50" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
(Silvestri, 1895), male ex AM KS.120532. Posterior (5), medial (6), anterior (7) and lateral (8) views of left gonopod, not to same scale. Scale bar in Fig. 6 = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="distribution">
<paragraph pageId="3" pageNumber="4">Distribution and habitat.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="BA27FE10A6DF59B0773714026F6E2A18" pageId="3" pageNumber="4" type="distribution">
<paragraph id="59C9DCB5591F8DFBB667B664B5500174" pageId="3" pageNumber="4">Distribution and habitat.</paragraph>
<paragraph id="E78A4C8596F511C8CB42C88173324E9C" pageId="3" pageNumber="4">
Occurs in the high country west of the Blue Mountains in New South Wales (Fig. 9) in grassy eucalypt forest (Fig. 4) and plantations of
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus radiata" order="Pinales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="radiata">Pinus radiata</taxonomicName>
<taxonomicName id="13B1789082C73009091AF18965A42A19" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus radiata" order="Pinales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="radiata">Pinus radiata</taxonomicName>
, where adults were found in 2013 sheltering under logs and small pieces of fallen wood or bark. We have not yet confirmed by further collecting the two 1990 localities south of Bathurst (Fig. 9); if these are correct, the north-south range of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<taxonomicName id="AB92B1D38D6AFB55199634BA1EA08172" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
is ca 150 km.
</paragraph>
<caption pageId="3" pageNumber="4">
<paragraph pageId="3" pageNumber="4">
<caption id="C02436D34BF004412001A545A9AEF46F" pageId="3" pageNumber="4">
<paragraph id="E35353099B6D90C695D01886C883C1E7" pageId="3" pageNumber="4">
Figure 9. Localities for
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
<taxonomicName id="90FC732077A6C1C09560138BCD0DD12B" class="Diplopoda" family="Paradoxosomatidae" genus="Hoplatessara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplatessara luxuriosa" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="luxuriosa">Hoplatessara luxuriosa</taxonomicName>
(Silvestri, 1895) as of 30 April 2013 (black squares). Inset shows location of main map. Base map from the Spatial Information Exchange, New South Wales Department of Finance and Services (http://maps.six.nsw.gov.au/ [accessed 9 July 2013]), showing named localities mentioned in text.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="remarks">
<paragraph pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="18A20B0D29BE538187E270F555F82504" pageId="3" pageNumber="4" type="remarks">
<paragraph id="9CD66AF5B3D6E6776B7B7C2A72A7453E" pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph id="FD55DFEF9E98FCE2269F7EDEF39B6BCF" pageId="3" pageNumber="4">
The structure of the anterior spiracle on diplosegments (Fig. 3) is very similar to that found in other australiosomatine paradoxosomatids [see figs 2A, 2B, 3C in
<bibRefCitation author="Mesibov, R" journalOrPublisher="Soil Organisms" pageId="5" pageNumber="6" pagination="531 - 542" title="New and little-used morphological characters in Polydesmida (Diplopoda)." volume="81" year="2009">Mesibov (2009)</bibRefCitation>
<bibRefCitation id="C0D9BD8D8DC01AA89D98FDEAEC56828F" author="Mesibov, R" journalOrPublisher="Soil Organisms" pageId="5" pageNumber="6" pagination="531 - 542" title="New and little-used morphological characters in Polydesmida (Diplopoda)." volume="81" year="2009">Mesibov (2009)</bibRefCitation>
].
</paragraph>
</subSubSection>

220
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<mods:role id="923B2E29DD81049140DC964B9BCDCE94">
<mods:roleTerm id="BF067C41F949009C6D023143F8CB9E40">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gates, Genevieve</mods:namePart>
<mods:namePart id="03215A6B79407BA8BE89E55A3D304990">Gates, Genevieve</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="F4A6ABCC43A0E20C8E165D95CF9B6D53" type="personal">
<mods:role id="3C76BCDBEAF9F30ACC6D9A95F04B959C">
<mods:roleTerm id="BD8E7E51E8D28223FFB74F47534E4CC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Du, Ping</mods:namePart>
<mods:namePart id="0081638450593F25EE8FE6D0A61C1128">Du, Ping</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>MycoKeys</mods:title>
<mods:typeOfResource id="931BD4609140A3F05F79A622C05DEFAA">text</mods:typeOfResource>
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<mods:title id="9743842CA84B07AA498CE0AC64C79918">MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>41</mods:number>
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<mods:date id="734E361B7B7AB860686EE5BC18945D00">2018</mods:date>
<mods:detail id="980A4FF1B9C385C8F00B3301E2CC9A05" type="volume">
<mods:number id="1016FC034C66E43A1C68B6B03658F535">41</mods:number>
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<mods:extent unit="page">
<mods:start>29</mods:start>
<mods:end>38</mods:end>
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<mods:start id="C8C8817472A6D68B2FD78E38061AAE13">29</mods:start>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.41.28485</mods:url>
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<mods:url id="951D40CB4E658E20E8E18B2D17F117EE">http://dx.doi.org/10.3897/mycokeys.41.28485</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.41.28485</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049--29</mods:identifier>
<mods:classification id="8EC9AAC5424F414FABC4E6053BC85C8A">journal article</mods:classification>
<mods:identifier id="C78FECA40E5626A784A78762C6677436" type="DOI">http://dx.doi.org/10.3897/mycokeys.41.28485</mods:identifier>
<mods:identifier id="BB0602D0E359B26B4416C673C0805384" type="Pensoft-Pub">1314-4049--29</mods:identifier>
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<treatment ID-GBIF-Taxon="154473944" LSID="urn:lsid:plazi:treatment:BA9D653572819D03B4725CD843B29FF5" httpUri="http://treatment.plazi.org/id/BA9D653572819D03B4725CD843B29FF5" lastPageId="6" lastPageNumber="35" pageId="4" pageNumber="33">
<subSubSection pageId="4" pageNumber="33" type="nomenclature">
<paragraph pageId="4" pageNumber="33">
<taxonomicName LSID="MB827073" authority="Y. C. Dai, G. M. Gates, X. H. Ji &amp; P. Du" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica Y.C. Dai, G.M. Gates, X.H. Ji &amp; P. Du</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="33">sp. nov.</taxonomicNameLabel>
<treatment id="BA9D653572819D03B4725CD843B29FF5" ID-GBIF-Taxon="154473944" LSID="urn:lsid:plazi:treatment:BA9D653572819D03B4725CD843B29FF5" httpUri="http://treatment.plazi.org/id/BA9D653572819D03B4725CD843B29FF5" lastPageId="6" lastPageNumber="35" pageId="4" pageNumber="33" scope_family="Hericiaceae" scope_phylum="Basidiomycota">
<subSubSection id="1C7FDA3F3322061030D315A6E61084CC" pageId="4" pageNumber="33" type="nomenclature">
<paragraph id="51E589491ED059D888E02EBF52FC6BB0" pageId="4" pageNumber="33">
<taxonomicName id="8F76E3350249D9A37DB28571B9BEABB8" ID-CoL="6CKDF" LSID="MB827073" authority="Y. C. Dai, G. M. Gates, X. H. Ji &amp; P. Du" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica Y.C. Dai, G.M. Gates, X.H. Ji &amp; P. Du</taxonomicName>
<taxonomicNameLabel id="58CE1010540204A9540893A7D5E6EEFF" pageId="4" pageNumber="33">sp. nov.</taxonomicNameLabel>
Figs 1, 2
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="diagnosis">
<paragraph pageId="4" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="33">
<subSubSection id="9907DBC914424809F674FC88F1E17A60" pageId="4" pageNumber="33" type="diagnosis">
<paragraph id="29490533D165C647E20F054E45994875" pageId="4" pageNumber="33">Diagnosis.</paragraph>
<paragraph id="A3391A0F51287DBBE46944BF45216075" pageId="4" pageNumber="33">
Differs from other
<taxonomicName class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis" order="Russulales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="genus">Dentipellis</taxonomicName>
<taxonomicName id="C7D1ADBC0318142B77E802F184CE92D3" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis" order="Russulales" pageId="4" pageNumber="33" phylum="Basidiomycota" rank="genus">Dentipellis</taxonomicName>
species by its gloeoplerous hyphae and gloeocystidia that become dark blue in
<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
<normalizedToken id="9E9FE3743C1C7B26F6B17EA9F4C1F4A7" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent and the presence of chlamydospores in subiculum.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="holotype">
<paragraph pageId="4" pageNumber="33">Holotype.</paragraph>
<paragraph pageId="4" pageNumber="33">
<subSubSection id="F25617ADCC1E402AFF0DB9B8A89954EF" pageId="4" pageNumber="33" type="holotype">
<paragraph id="2D6AD2756C319D8A2C404F6FAD48002D" pageId="4" pageNumber="33">Holotype.</paragraph>
<paragraph id="CA3C0A663FEEA11F218A72AE770F12D3" pageId="4" pageNumber="33">
AUSTRALIA. Tasmania: Arve River Streamside Reserve,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-43.166668">43°10'S</geoCoordinate>
<geoCoordinate id="AC9659A2E851AEA2B4C18D65EDF0FDED" direction="south" orientation="latitude" precision="925" value="-43.166668">43°10'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="92" value="146.80833">146°48.5'E</geoCoordinate>
<geoCoordinate id="9FBF06B39E749854F612562DFF244031" direction="east" orientation="longitude" precision="92" value="146.80833">146°48.5'E</geoCoordinate>
, elev. 160 m, on fallen trunk of
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
<taxonomicName id="70B2AB8E6A715C7557952AE7747AFE7E" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
sp., 15 May 2018, Dai 18767 (M, isotype in BJFC; ITS GenBank accession number: MH571699, 28S GenBank accession number: MH571702).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="etymology">
<paragraph pageId="4" pageNumber="33">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="33">
<taxonomicName class="Insecta" family="Curculionidae" genus="Tasmanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanica" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">Tasmanica</taxonomicName>
<subSubSection id="D80CE0FF9CFCBCADCA2788AA1595D5B5" pageId="4" pageNumber="33" type="etymology">
<paragraph id="46C3DA7FBC6C44766427007B3C43C9F5" pageId="4" pageNumber="33">Etymology.</paragraph>
<paragraph id="4A1DEFADC04894ED8A98AD9F96C86CCA" pageId="4" pageNumber="33">
<taxonomicName id="88DFE9387FB363669925156FD463788B" class="Insecta" family="Curculionidae" genus="Tasmanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanica" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">Tasmanica</taxonomicName>
(Lat.): referring to the species collected from Tasmania of Australia.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="basidiomata">
<paragraph pageId="4" pageNumber="33">Basidiomata.</paragraph>
<paragraph pageId="4" pageNumber="33">Annual, resupinate, inseparable from substratum, soft corky, without odour or taste when fresh, fragile upon drying, up to 100 cm long, 40 cm wide and 3.5 mm thick at centre. Hymenophore with spines, cream when fresh and cinnamon when dry, spines up to 3 mm long, 2-3 per mm across base, soft corky to fragile, a few glued at tips when dry; margin distinct, white, fibrillous to cottony, up to 5 mm wide; spines, cream, becoming fragile and clay-buff when dry, up to 3 mm long. Subiculum very thin, soft corky, white to cream, &lt;1 mm thick.</paragraph>
<subSubSection id="AA53672FECCC6D84DF05FD798CF35D9A" pageId="4" pageNumber="33" type="basidiomata">
<paragraph id="4451EC9C7FD8558764E8348A9FC59856" pageId="4" pageNumber="33">Basidiomata.</paragraph>
<paragraph id="264B0DAEB564D359741BEF7114F4135B" pageId="4" pageNumber="33">Annual, resupinate, inseparable from substratum, soft corky, without odour or taste when fresh, fragile upon drying, up to 100 cm long, 40 cm wide and 3.5 mm thick at centre. Hymenophore with spines, cream when fresh and cinnamon when dry, spines up to 3 mm long, 2-3 per mm across base, soft corky to fragile, a few glued at tips when dry; margin distinct, white, fibrillous to cottony, up to 5 mm wide; spines, cream, becoming fragile and clay-buff when dry, up to 3 mm long. Subiculum very thin, soft corky, white to cream, &lt;1 mm thick.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="hyphal structure">
<paragraph pageId="4" pageNumber="33">Hyphal structure.</paragraph>
<paragraph pageId="4" pageNumber="33">
<subSubSection id="B4C22C8B0A50C535C5B8BC1D1DC09D4D" pageId="4" pageNumber="33" type="hyphal structure">
<paragraph id="EF0EF9AFC0D568D3D7168183DFBF9AB5" pageId="4" pageNumber="33">Hyphal structure.</paragraph>
<paragraph id="003ED1E828E5DDA7C47A057299A913B9" pageId="4" pageNumber="33">
Hyphal system monomitic; generative hyphae with clamp connections,
<normalizedToken originalValue="IKI–">IKI-</normalizedToken>
<normalizedToken id="3F5DDF6BE05E33DCCD8E212057C3CCAA" originalValue="IKI–">IKI-</normalizedToken>
, CB+; the colour and size unchanged in KOH.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="subiculum">
<paragraph pageId="4" pageNumber="33">Subiculum.</paragraph>
<paragraph pageId="4" pageNumber="33">
<subSubSection id="12D7CD4304A3F3516D16CDD6D2BAE08A" pageId="4" pageNumber="33" type="subiculum">
<paragraph id="C45DA3E935ECB947DD63227B61A50984" pageId="4" pageNumber="33">Subiculum.</paragraph>
<paragraph id="D5B2CDFD21F684AC3CCA34345819B001" pageId="4" pageNumber="33">
Generative hyphae colourless, thin- to slightly thick-walled, frequently branched, flexuous, interwoven, 3-4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="63008A59EBF8DCCEF59901CA534B79C3" originalValue="μm">μm</normalizedToken>
in diam. Gloeoplerous hyphae occasionally present, dark blue in
<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
<normalizedToken id="D1E2F477AD131FEB42191A9286829ACE" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent. Chlamydospores present, ellipsoid, thick-walled, 5-5.6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="E928962688EFDCACF11B0EED7FDA1449" originalValue="×">x</normalizedToken>
2.8-3.3
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="F18841303A121BDE4757C74AA3C6B222" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="33" type="hymenophoral trama">
<paragraph pageId="4" pageNumber="33">Hymenophoral trama.</paragraph>
<paragraph pageId="4" pageNumber="33">
<subSubSection id="F551835B77FE615FBBA2AE851FE014E3" pageId="4" pageNumber="33" type="hymenophoral trama">
<paragraph id="C050805E481423403E8538B8569091CF" pageId="4" pageNumber="33">Hymenophoral trama.</paragraph>
<paragraph id="5743B48D96EE8CA1C3F8AB6434CB9A1E" pageId="4" pageNumber="33">
Generative hyphae colourless, thin-walled, frequently branched, straight, parallel along the spines, 2.8-4
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="B7D5395CA71D275309928AA51E1F325A" originalValue="μm">μm</normalizedToken>
in diam. Gloeocystidia abundant, colourless, thin- to slightly thick-walled, clavate, contents oily and dark blue in
<normalizedToken originalValue="Melzer’s">Melzer's</normalizedToken>
<normalizedToken id="F9708C9276415B86A99CA40AEE6AAD9F" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent, rooting deep from the trama, up to a few hundred microns long, the cystidia-like apical part 30-45
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="D4A6AE1DB7F144AAC234B06EEA78F312" originalValue="×">x</normalizedToken>
5-8
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="A0AF9C2BD56BBE52448158402CBEEA5D" originalValue="μm">μm</normalizedToken>
. Oily material abundant amongst trama.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="hymenium">
<paragraph pageId="4" pageNumber="33">Hymenium.</paragraph>
<paragraph lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
<subSubSection id="23E8403FDBDB3EF7C15FFF758E171881" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="hymenium">
<paragraph id="ACAD663815C26AB2F4563B97B64DBE5A" pageId="4" pageNumber="33">Hymenium.</paragraph>
<paragraph id="930B5636643B73546549E523D99819C8" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
Cystidioles colorless, thin-walled, ventricose with elongated apical portion, bearing some irregular crystals, 30-45
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="71F94F234D23CD018E587AB076C5E009" originalValue="×">x</normalizedToken>
5-8
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="44B19F260CED6C35C1A7EA65AD3F950A" originalValue="μm">μm</normalizedToken>
; basidia clavate with four
<pageBreakToken pageId="5" pageNumber="34" start="start">sterigmata</pageBreakToken>
<pageBreakToken id="19B0F56E51B2FCB59874B97694078765" pageId="5" pageNumber="34" start="start">sterigmata</pageBreakToken>
and a basal clamp connection, 20-26
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="86181DF4F6F8DCF85F10DAAB42401486" originalValue="×">x</normalizedToken>
3-4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="52DD2DDA6BC701643585299C447436A0" originalValue="μm">μm</normalizedToken>
. Basidiospores ellipsoid, coloruless, thin-walled, densely echinulate, IKI+, CB+, (3.4
<normalizedToken originalValue="–)3.5–4.5(–">-)3.5-4.5(-</normalizedToken>
<normalizedToken id="8370CEAA131261B3AFABFB9C4D327307" originalValue="–)3.5–4.5(–">-)3.5-4.5(-</normalizedToken>
4.8)
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="EA7CD59A592E453226DD074B935F5044" originalValue="×">x</normalizedToken>
2.4
<normalizedToken originalValue="–3.2(–">-3.2(-</normalizedToken>
<normalizedToken id="C11F0A1B144F21411D9D9D47E25045CB" originalValue="–3.2(–">-3.2(-</normalizedToken>
3.5)
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="D793D1574E469341417479ECB8948FEE" originalValue="μm">μm</normalizedToken>
, L = 3.99
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="095B6B14E0634BF8CD69A2872CC62736" originalValue="μm">μm</normalizedToken>
, W = 2.92
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="4D3C607D146FD822AEC13F01E5EF8776" originalValue="μm">μm</normalizedToken>
, Q = 1.36-1.39 (n = 90/3).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="materials_examined">
<paragraph pageId="5" pageNumber="34">Additional specimens examined (paratypes).</paragraph>
<paragraph lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
<subSubSection id="CF820A073BAE167C2A29B4C5D5B554BF" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="materials_examined">
<paragraph id="3C716E7F861C7AD2536EAB5C6086A87D" pageId="5" pageNumber="34">Additional specimens examined (paratypes).</paragraph>
<paragraph id="AD25320A86B8153F13F3FA576C78053B" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
AUSTRALIA. Tasmania: Arve River Streamside Reserve, on fallen trunk of
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
<taxonomicName id="7A74CAE3558346C4FE5914CF3F837C1C" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
sp., 15 May 2018, Dai 18768 (M, duplicate in BJFC; ITS GenBank accession number: MH571700, 28S GenBank accession
<pageBreakToken pageId="6" pageNumber="35" start="start">number</pageBreakToken>
<pageBreakToken id="02F46C295784EB5FDBE2A4AC7918AAC1" pageId="6" pageNumber="35" start="start">number</pageBreakToken>
: MH571703); Mt Field National Park,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-42.683334">42°41'S</geoCoordinate>
<geoCoordinate id="7E5C09A8298BDD0BDA08C9E3873CED98" direction="south" orientation="latitude" precision="925" value="-42.683334">42°41'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="146.7">146°42'E</geoCoordinate>
<geoCoordinate id="5D71170E3A17A59CF0ED61FCBA1C2A28" direction="east" orientation="longitude" precision="925" value="146.7">146°42'E</geoCoordinate>
, elev., 180 m, on fallen trunk of
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="6" pageNumber="35" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
<taxonomicName id="063C53DC2BD6A91EA0E6405031FEFEBA" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="6" pageNumber="35" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
sp., 14 May 2018, Dai 18737 (M, duplicate in BJFC; ITS GenBank accession number: MH571698, 28S GenBank accession number: MH571701).
</paragraph>
<caption pageId="6" pageNumber="35">
<paragraph pageId="6" pageNumber="35">
<caption id="A206986D92400FAAD20D6BC3C8C5E4F1" pageId="6" pageNumber="35">
<paragraph id="785EFC5D3E472450D338969B657413EB" pageId="6" pageNumber="35">
Figure 1. Strict consensus tree illustrating the phylogenetic position of
<taxonomicName class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
<taxonomicName id="02CD972307912464F6600EDF580D1AB1" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
, generated by the maximum parsimony method based on ITS+28S sequence data. Branches are labelled with parsimony bootstrap values ≥50% and Bayesian posterior probabilities ≥0.95.
<taxonomicName class="Agaricomycetes" family="Bondarzewiaceae" genus="Bondarzewia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Bondarzewia podocarpi" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="podocarpi">Bondarzewia podocarpi</taxonomicName>
<taxonomicName id="C7E87CD6556C1791E2AC966BC3B7F6B7" class="Agaricomycetes" family="Bondarzewiaceae" genus="Bondarzewia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Bondarzewia podocarpi" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="podocarpi">Bondarzewia podocarpi</taxonomicName>
and
<taxonomicName lsidName="B. occidentalis" pageId="6" pageNumber="35" rank="species" species="occidentalis">B. occidentalis</taxonomicName>
<taxonomicName id="E5F57F41214928A4F35306A4A2FEE180" lsidName="B. occidentalis" pageId="6" pageNumber="35" rank="species" species="occidentalis">B. occidentalis</taxonomicName>
are used to root the tree. Branch lengths reflect expected changes per site as indicated by the scale.
</paragraph>
</caption>
<caption pageId="6" pageNumber="35">
<paragraph pageId="6" pageNumber="35">
<caption id="32BDABACAA37125B8567C23CB1CBAD7D" pageId="6" pageNumber="35">
<paragraph id="751131A550644F9FAB4A95163758206C" pageId="6" pageNumber="35">
Figure 2. A fresh basidiocarp of
<taxonomicName class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
<taxonomicName id="76E489456E9C3BA145A9B569BF3145E8" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
(holotype). Scale bar: 1 cm.
</paragraph>
</caption>
<caption pageId="6" pageNumber="35">
<paragraph pageId="6" pageNumber="35">
<caption id="67BADCCA3CEC9171F12A111E04E862DF" pageId="6" pageNumber="35">
<paragraph id="973975379529ED47C1F48849056FB886" pageId="6" pageNumber="35">
Figure 3. Microscopic structures of
<taxonomicName class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
<taxonomicName id="14BA0C2F2645EDF9FDDCB80395C4A08C" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dentipellis tasmanica" order="Russulales" pageId="6" pageNumber="35" phylum="Basidiomycota" rank="species" species="tasmanica">Dentipellis tasmanica</taxonomicName>
(holotype). a Basidiospores b Basidia and basidioles c Gloeocystidia and Cystidioles d Hyphae from trama e Hyphae from subiculum.
</paragraph>
</caption>