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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.917.33020" ID-GBIF-Dataset="ad127b3e-6c04-48e9-a653-fcf1dfd30758" ID-PMC="PMC7076068" ID-Pensoft-Pub="1313-2970-917-1" ID-Pensoft-UUID="24D1A40689435E1F874F3C88D19CC77B" ID-PubMed="32206015" ID-ZooBank="4D2DBC43950B40E4B333993C5A76D1EC" ModsDocID="1313-2970-917-1" checkinTime="1583773819196" checkinUser="pensoft" docAuthor="Ospina-Sanchez, Claudia M., Palacios-Vargas, Jose G. &amp; Gonzalez, Grizelle" docDate="2020" docId="2E559586AD58504A98597B515D086689" docLanguage="en" docName="ZooKeys 917: 1-13" docOrigin="ZooKeys 917" docSource="http://dx.doi.org/10.3897/zookeys.917.33020" docTitle="Furculanurida bistribus Ospina-Sánchez &amp; Palacios-Vargas &amp; González 2020, sp. nov." docType="treatment" docVersion="5" id="24D1A40689435E1F874F3C88D19CC77B" lastPageNumber="1" masterDocId="24D1A40689435E1F874F3C88D19CC77B" masterDocTitle="New species of Furculanurida (Collembola, Neanuridae, Pseudachorutinae) from the Luquillo Mountains, Puerto Rico" masterLastPageNumber="13" masterPageNumber="1" pageNumber="1" updateTime="1668168536111" updateUser="ExternalLinkService">
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<mods:title>New species of Furculanurida (Collembola, Neanuridae, Pseudachorutinae) from the Luquillo Mountains, Puerto Rico</mods:title>
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<mods:title id="939724960E908B3B64E0E0D07940D682">New species of Furculanurida (Collembola, Neanuridae, Pseudachorutinae) from the Luquillo Mountains, Puerto Rico</mods:title>
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<mods:namePart>Ospina-Sanchez, Claudia M.</mods:namePart>
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<mods:namePart>Palacios-Vargas, Jose G.</mods:namePart>
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<mods:namePart>Gonzalez, Grizelle</mods:namePart>
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<mods:date>2020</mods:date>
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<treatment ID-GBIF-Taxon="162467296" LSID="urn:lsid:zoobank.org:act:" httpUri="http://treatment.plazi.org/id/2E559586AD58504A98597B515D086689" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="2E559586-AD58-504A-9859-7B515D086689" authority="Ospina-Sánchez &amp; Palacios-Vargas &amp; González, 2020" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus" status="sp. nov.">Furculanurida bistribus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Furculanurida bistribus sp. nov. Specimen in ethanol 96 %." figureDoi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">, 3-8</figureCitation>
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">, 9-13</figureCitation>
<treatment id="2E559586AD58504A98597B515D086689" ID-GBIF-Taxon="162467296" LSID="urn:lsid:zoobank.org:act:" httpUri="http://treatment.plazi.org/id/2E559586AD58504A98597B515D086689" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Collembola" scope_family="Neanuridae" scope_subFamily="Pseudachorutinae">
<subSubSection id="B3908B5F3C542B8FB5EBF3E14DCAA615" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="E4C51B4AEDD81D57FE84426F83AED91C" pageId="0" pageNumber="1">
<taxonomicName id="E00A03E23C8B1DEFE244A08E3C0D7804" LSID="2E559586-AD58-504A-9859-7B515D086689" authority="Ospina-Sánchez &amp; Palacios-Vargas &amp; González, 2020" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Collembola" pageId="0" pageNumber="1" rank="species" species="bistribus" status="sp. nov.">Furculanurida bistribus</taxonomicName>
<taxonomicNameLabel id="7949767E75FB62B2AE90F9BC2B84097C" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="34022BAF1DE903EB0C491E02B361EC12" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Furculanurida bistribus sp. nov. Specimen in ethanol 96 %." figureDoi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation id="D0892FF3DA1917ABA2121198AB5F9CB3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation id="D6DEC0A3AA5EC818064BCB54F8D7CDD2" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">, 3-8</figureCitation>
<figureCitation id="23B97BF37E912E363596AF16D9369ECD" captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">, 9-13</figureCitation>
,
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Furculanurida species with disputed generic placement and their taxonomic history." pageId="0" pageNumber="1">Tables 1</figureCitation>
<figureCitation id="7D824F134105E42BFD5B90BBC3B38A3A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Furculanurida species with disputed generic placement and their taxonomic history." pageId="0" pageNumber="1">Tables 1</figureCitation>
,
<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Main characters of the all known species of Furculanurida, including species moved to other genera." pageId="0" pageNumber="1">2</figureCitation>
<figureCitation id="E2DF998605A828F4055DD48307C8A646" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Main characters of the all known species of Furculanurida, including species moved to other genera." pageId="0" pageNumber="1">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="D95C665CB8C8B901FEB4A1392CD00E64" pageId="0" pageNumber="1" type="type material">
<paragraph id="358C02D91A51CF26C2232D7588DD5737" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="50AA9D1ED1FAF8A2688321C6F28A3701" pageId="0" pageNumber="1">
<emphasis id="5BC6F2158586D27938A6F81C7C34BFB8" bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
(female on slide) and 8
<emphasis bold="true" pageId="0" pageNumber="1">paratypes</emphasis>
<emphasis id="2FDE0512CB516640A7265F319823934A" bold="true" pageId="0" pageNumber="1">paratypes</emphasis>
(2 males, 4 females, and 2 juveniles, each one on slides). Puerto Rico, Luquillo, Luquillo Mountains, Pico del Este,
<geoCoordinate degrees="18" direction="north" minutes="16" orientation="latitude" precision="15" seconds="17" value="18.271389">
<geoCoordinate id="5F5E77E448FB3AE37EBC65941F5B7441" degrees="18" direction="north" minutes="16" orientation="latitude" precision="15" seconds="17" value="18.271389">
18°16
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N
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;
<geoCoordinate degrees="65" direction="west" minutes="45" orientation="longitude" precision="15" seconds="40" value="-65.76111">
<geoCoordinate id="21B382EB9436DDD0E36890039A6BFB1A" degrees="65" direction="west" minutes="45" orientation="longitude" precision="15" seconds="40" value="-65.76111">
65°45
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<normalizedToken id="5AD115728A19C74B00F95E037484D3C5" originalValue="’40”">'40&quot;</normalizedToken>
W
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; 987.6 m a.s.l.; ex mosses,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="3C94501AE9C419CF4FE2317EA415BCDA" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="487765464548BDD27185911DFE06E420" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
</taxonomicName>
forest type, from leaf litter and epiphytes, 04 Nov 2014, leg. CM Ospina.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="887A8858CB88E8B9E5F9A170D1AF98F4" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="2DD7390E14FF8C9C718F893C49D4F8CE" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="C53B1178DAA6205445478F7B4731D307" pageId="0" pageNumber="1">
2 females on slides, Puerto Rico, Luquillo, Luquillo Mountains, Pico del Este,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="AFA3C81D4D0CCCEA07D1F109BA7DED89" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="F039D5EB3D2935998EF8E500D236CB2A" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
</taxonomicName>
forest type, epiphyte, 987.6 m a.s.l., 19 May 2015, leg. CM Ospina. 1 female on slide, Puerto Rico, Luquillo, Luquillo Mountains, Pico del Este,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="EE6A75B13E2A5F3711840EDEB3C75137" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="4ED0A901A589F2BB60294FF3F58AFC5B" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
</taxonomicName>
forest type, epiphyte, 987.6 m a.s.l., 11 Feb 2015, leg. CM Ospina. 1 juvenile on slide Puerto Rico, Luquillo, Luquillo Mountains, Pico El Yunque,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="04EEFE99DBE3C60CBBCE86BF8491FF1E" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="9E3866CD97DFC5E96ED2F6A905969D85" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
</taxonomicName>
forest type, leaf litter, 1044.8 m a.s.l., 4 Nov 2014, leg. CM Ospina. 1 female on slide Puerto Rico, Luquillo, Luquillo Mountains, Pico El Yunque,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="56056AD4058CC2D72860EE8B49D15BAE" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="2E1B336057DE4743C65207E3B9D73A70" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
</taxonomicName>
forest type, leaf litter, 1044.8 m a.s.l., 19 May 2015, leg. CM Ospina.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Eyes 3+3 eyes. Post antennal organ in rosette with 5 or 6 vesicles. Ant IV with six sensilla. Seta a0 on head absent. Mandible with four teeth. Dens with three setae. Unguis without internal tooth.</paragraph>
<subSubSection id="B5FB6EFAB6D414542654D3FD47FEB83F" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="8F624BEC337B7465B1A47460EDBD3B26" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="238CC5D569B7318FC3BD7B5652C1F97B" pageId="0" pageNumber="1">Eyes 3+3 eyes. Post antennal organ in rosette with 5 or 6 vesicles. Ant IV with six sensilla. Seta a0 on head absent. Mandible with four teeth. Dens with three setae. Unguis without internal tooth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="5EB4A0050A4BA0D88962BB81E7E6B898" pageId="0" pageNumber="1" type="description">
<paragraph id="310D2DDAC418293F824BAFDE70F8DEA9" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="66A59D070AACF2461F6F6DCE0C139114" pageId="0" pageNumber="1">
Average body length: adults 1009
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="05B8A61E23A199CA9544EA837C10E59A" originalValue="µm">µm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
<emphasis id="35DBE34495241B47A9DEE4AC07676D47" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 5); juveniles 847
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="CD42E08BA21FC07EE5375EEAB296BD22" originalValue="µm">µm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
<emphasis id="A3C284C6F5558E3AED39636CB9F23696" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2). Specimens in ethanol with antenna and abdomen evenly grey, ocular patch dark; head, legs III, and furcula light grey; thorax, legs I and II white to light purple (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Furculanurida bistribus sp. nov. Specimen in ethanol 96 %." figureDoi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1">1</figureCitation>
<figureCitation id="AF120115FA6F72D8C5EC38F0DAFEAA43" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Furculanurida bistribus sp. nov. Specimen in ethanol 96 %." figureDoi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1">1</figureCitation>
). Granulation coarse. Body setae comprising short, smooth and thin setae, and long and smooth sensorial setae.
</paragraph>
<caption doi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
<caption id="D87520CD5931751D6E61D4589ADFCA8A" doi="10.3897/zookeys.917.33020.figure1" httpUri="https://binary.pensoft.net/fig/387222" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph id="EB7C13FE0D724132C0BF6809C36408DD" pageId="0" pageNumber="1">
<emphasis id="C7E40E0B5ABFEE24EAEC0171B30461EE" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName id="EEB76367E91B2064EA77391BF35B0739" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis id="9A3390616A7C19CD5EA77A0B3898E7BE" italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
</taxonomicName>
sp. nov. Specimen in ethanol 96%.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<paragraph id="F1D784EF622306C10A24716DBCAA982A" pageId="0" pageNumber="1">
Head: antenna shorter (0.6) than head diagonal. Ant III and IV fused dorsally, ventral separation clearly marked. Ant IV dorsally with trilobed apical vesicle, six subcylindrical thin sensilla and 14 long setae; subapical organite present; dorsoexternal microsensillum absent (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
<figureCitation id="FE67EDAFE09C30035B119215680C94B8" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
); no sensorial field ventrally on Ant IV (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">4</figureCitation>
<figureCitation id="B98CFD94792036DC458F6C61171E7E75" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">4</figureCitation>
); Ant III sense organ with two small internal slightly bent sensilla, two subcylindrical guard sensilla, Sgv larger than Sgd, ventral microsensillum present (Figs
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
<figureCitation id="5F07EA1CB281C45D44E00D27DFC33034" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
,
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">5</figureCitation>
<figureCitation id="FA82453710FB1F7469BA77A85B809E38" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">5</figureCitation>
); Sgd apically displaced, towards Ant IV, aligned to S2 and S3; Ant II with 10 setae; Ant I with seven setae (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
<figureCitation id="76F66928D43DF08003639242723CB143" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">3</figureCitation>
). Eyes 3+3 on a pigmented eyepatch; PAO with five or six vesicles disposed as a rosette (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
<figureCitation id="F044BAF7F285C5CD3706A5DE21FE5610" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
). Head dorsal chaetotaxy as in Figure
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
<figureCitation id="3A613CCFA4914FA71435EC7AAC494015" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
; seta a0 absent; d row with 4 setae, sd row with three setae; setae Oc 1-3 present; c setae absent; p1-3 setae present. Buccal cone elongate, labium with complete chaetotaxy, A to G setae, C and D apically displaced (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">6</figureCitation>
<figureCitation id="E2AC01CDA9AFBCE2DAE7079458D4347C" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">6</figureCitation>
). Pre-labral/Labral chaetotaxy 4/2322 (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">7</figureCitation>
<figureCitation id="2942F51A66401AFBB451DB596CA7909F" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">7</figureCitation>
). Mandible with four teeth, two apical short and subequal, one medial and one basal large and subequal; maxilla styliform with one lamella (Fig.
<figureCitation captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">8</figureCitation>
<figureCitation id="91750E6322671F2F156937D60BF3258A" captionStart="Figures 3–8" captionStartId="F3" captionText="Figures 3 - 8. Furculanurida bistribus sp. nov. 3 Ant I-IV, dorsal view 4 Ant IV and III, ventral view 5 Sensorial Organ in Ant. III 6 Labium 7 Labrum 8 Maxilla and mandible." figureDoi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1">8</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
<caption id="DC9A93308070DA6614D16744A2DAACAE" doi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="B1B87DF8F9C548345F07ED917C1D703B" pageId="0" pageNumber="1">
<emphasis id="15B31FD8BC8BD2A03187A6AFAE523FEF" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName id="50F41C06D2878F8C7FFCB8D22051CC57" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis id="868DE3CDD72E8E08B44C8EA389A47BB8" italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
</taxonomicName>
sp. nov. Dorsal chaetotaxy of body (female).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1" start="Figures 3–8" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 3-8.</emphasis>
<taxonomicName authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
<caption id="6B95F96A73C44A3C760007EBD607CE9C" doi="10.3897/zookeys.917.33020.figures3-8" httpUri="https://binary.pensoft.net/fig/387224" pageId="0" pageNumber="1" start="Figures 3–8" startId="F3">
<paragraph id="D6F2FBB440C55950767E5D22DB1F7E54" pageId="0" pageNumber="1">
<emphasis id="B003C7ED81E4D8C0937AC307E76E7805" bold="true" pageId="0" pageNumber="1">Figures 3-8.</emphasis>
<taxonomicName id="D657836E77827195C802282997668FC4" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis id="820D719CB4CF9FB890D825A9F98041D1" italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
<emphasis id="AA75451999CBA845E6CC0ECD18F97D0C" bold="true" pageId="0" pageNumber="1">3</emphasis>
Ant I-IV, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
<emphasis id="881B3E7FC01397961D0B72F404F5FDDB" bold="true" pageId="0" pageNumber="1">4</emphasis>
Ant IV and III, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
<emphasis id="597D548293F8932A3281DDFB64AA27F3" bold="true" pageId="0" pageNumber="1">5</emphasis>
Sensorial Organ in Ant. III
<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
<emphasis id="F46E22231BC92AC77ABD0EE8E2E5B7AF" bold="true" pageId="0" pageNumber="1">6</emphasis>
Labium
<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
<emphasis id="6736B372976875F2D93216958B308392" bold="true" pageId="0" pageNumber="1">7</emphasis>
Labrum
<emphasis bold="true" pageId="0" pageNumber="1">8</emphasis>
<emphasis id="B79FC0E90C627BEEF5F30C997F533998" bold="true" pageId="0" pageNumber="1">8</emphasis>
Maxilla and mandible.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<paragraph id="3F0FE5542FDA357E5E2378FFBE5BA4FE" pageId="0" pageNumber="1">
Dorsal chaetotaxy: ordinary setae smooth, distributed as in Figure
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
<figureCitation id="662C05E38377A744A2365184BAE5C773" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Furculanurida bistribus sp. nov. Dorsal chaetotaxy of body (female)." figureDoi="10.3897/zookeys.917.33020.figure2" httpUri="https://binary.pensoft.net/fig/387223" pageId="0" pageNumber="1">2</figureCitation>
. Th I with 2+2 setae; Th II and III with one dorsolateral seta posteriorly displaced. Sensory setae (s) clearly differentiated, in position p3 and p6 in Th II and III and in position p3 in Abd I-V; S-chaetotaxy formula = 022/11111.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="7C00C43835BFCDF7D190CD57ADD4AEF4" pageId="0" pageNumber="1">
Legs chaetotaxy: subcoxae 1, two; subcoxae 2, one; Cx, three; Tr, four; Fe, 10, and Ti 19 setae (Fig.
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">9</figureCitation>
<figureCitation id="45A33A7E398657B06838FE7D699617E9" captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">9</figureCitation>
). Seta M present between B4 and B5 without displacement. Tenent hair acuminate. Claws without teeth, unguis larger than Ti (ratio tibiotarsus: unguis = 1:1.2:); unguiculus absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="ACCF71DB13617017E0921456F1A99925" pageId="0" pageNumber="1">
Ventral chaetotaxy as in Figure
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">10</figureCitation>
<figureCitation id="A4F6FBD40F3B372CA06366F541054DEF" captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">10</figureCitation>
. VT with 3+3 setae; tenaculum with 3+3 teeth and without setae; furcula well developed, manubrium with 14 setae, dens with three setae, mucro straight with a broad hook-like apex (Fig.
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">11</figureCitation>
<figureCitation id="F7AED770EB87D4E6E596EF347994B030" captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Furculanurida bistribus sp. nov. 9 Tibiotarsus II 10 Ventral quetotaxy 11 Manubrium and furcula 12 Female genital plate 13 Male genital plate." figureDoi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1">11</figureCitation>
). Ratio mucro: dens = 1: 1.3. Female genital plate with 2+2 pregenital setae, four circumgenital setae, and 1+1 eugenital setae (Fig.
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). Male genital plate with 2+2 pregenital setae, ten circumgenital setae, and 4+4 eugenital setae (Fig.
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).
</paragraph>
<caption doi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1" start="Figures 9–13" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 9-13.</emphasis>
<taxonomicName authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
<caption id="825F42A82AC52F10ACD1804A6F69142F" doi="10.3897/zookeys.917.33020.figures9-13" httpUri="https://binary.pensoft.net/fig/387225" pageId="0" pageNumber="1" start="Figures 9–13" startId="F4">
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<emphasis id="AE15986BE0EF66E9315E31B4CA32C742" bold="true" pageId="0" pageNumber="1">Figures 9-13.</emphasis>
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<emphasis id="7EAE80D845C17EBDE6D4B89C025BCF9D" italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
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sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">9</emphasis>
<emphasis id="AFC8C35E2A36E03F40E569ACCEAED238" bold="true" pageId="0" pageNumber="1">9</emphasis>
Tibiotarsus II
<emphasis bold="true" pageId="0" pageNumber="1">10</emphasis>
<emphasis id="773BAB953A22F540DC919B40D06F6181" bold="true" pageId="0" pageNumber="1">10</emphasis>
Ventral quetotaxy
<emphasis bold="true" pageId="0" pageNumber="1">11</emphasis>
<emphasis id="4B26085A0D241E4E81DF98AA916FFDB4" bold="true" pageId="0" pageNumber="1">11</emphasis>
Manubrium and furcula
<emphasis bold="true" pageId="0" pageNumber="1">12</emphasis>
<emphasis id="F917E040B1F8A665DB3FE9958A4A55EB" bold="true" pageId="0" pageNumber="1">12</emphasis>
Female genital plate
<emphasis bold="true" pageId="0" pageNumber="1">13</emphasis>
<emphasis id="933C9FE2AEDCB9B77DB9C7B88111998A" bold="true" pageId="0" pageNumber="1">13</emphasis>
Male genital plate.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Bistribus" lsidName="Bistribus" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bistribus</emphasis>
<subSubSection id="E355DBCA5C1A83653D23B9B2B677AEB9" pageId="0" pageNumber="1" type="etymology">
<paragraph id="BCB18D3784BF7C2651BA9BDBB6A04F37" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="B3333ED05237A241DCD88B1932DCA802" pageId="0" pageNumber="1">
<taxonomicName id="64060308BD4DC3E81C8716D4E311F50E" genus="Bistribus" lsidName="Bistribus" pageId="0" pageNumber="1" rank="genus">
<emphasis id="A6CE6EA7B2A93D7F97B7BA49E8D966A2" italics="true" pageId="0" pageNumber="1">Bistribus</emphasis>
</taxonomicName>
, Latin for two times three, in reference to the presence of 3+3 eyes and 3+3 setae on dens, diagnostic characters of the species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="D2A75376DD680F63A14EEC60B0EB6567" pageId="0" pageNumber="1" type="distribution">
<paragraph id="5CDCBEACD7BCC4C5A6CCF423A653B624" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="15B01AE7C45936EEFCBD6522FB342E0E" pageId="0" pageNumber="1">
This species is only the
<taxonomicName authorityName="Massoud" authorityYear="1967" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida</emphasis>
<taxonomicName id="A79861AB685E47E9F16A975A679C27D1" authorityName="Massoud" authorityYear="1967" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="A81658E5C79B29215878CB71CECF918D" italics="true" pageId="0" pageNumber="1">Furculanurida</emphasis>
</taxonomicName>
known from the Luquillo Mountains in the
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
<taxonomicName id="FAB5192A5A076ECCC842E9EA3C5C8496" class="Magnoliopsida" family="Bignoniaceae" genus="Tabebuia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tabebuia rigida" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="7F95D397F3BFA6B83F96319E4CD693C9" italics="true" pageId="0" pageNumber="1">Tabebuia rigida</emphasis>
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forest type, on Pico del Este
<geoCoordinate degrees="18" direction="north" minutes="16" orientation="latitude" precision="15" seconds="17" value="18.271389">
<geoCoordinate id="CE1C5470A31226132C7069B49AFF43CF" degrees="18" direction="north" minutes="16" orientation="latitude" precision="15" seconds="17" value="18.271389">
18°16
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N
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;
<geoCoordinate degrees="65" direction="west" minutes="45" orientation="longitude" precision="15" seconds="40" value="-65.76111">
<geoCoordinate id="09ADEE875F9EB9312A795CA3BAF6E068" degrees="65" direction="west" minutes="45" orientation="longitude" precision="15" seconds="40" value="-65.76111">
65°45
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W
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; 987.6 m a.s.l. and Pico El Yunque
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<geoCoordinate id="E7DA312BD83A30AAF5CF869FD0ADB5B5" degrees="18" direction="north" minutes="18" orientation="latitude" precision="15" seconds="37" value="18.310278">
18°18
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<normalizedToken id="B1C49573AC1F3D423A44B6F351D65D8B" originalValue="’37”">'37&quot;</normalizedToken>
N
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;
<geoCoordinate degrees="65" direction="west" minutes="47" orientation="longitude" precision="15" seconds="26" value="-65.79056">
<geoCoordinate id="560DE6A6C761B7367460FE02D86046CC" degrees="65" direction="west" minutes="47" orientation="longitude" precision="15" seconds="26" value="-65.79056">
65°47
<normalizedToken originalValue="’26”">'26&quot;</normalizedToken>
<normalizedToken id="6DD748CD92AC7C4CF32547A2BAEA3059" originalValue="’26”">'26&quot;</normalizedToken>
W
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; 1044.8 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
<subSubSection id="21CBA0DC5C2474EFE274A2F0A2659A50" pageId="0" pageNumber="1" type="ecology">
<paragraph id="FF1A65EAED88E08289762C4171F8DC68" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="A3929107988F1FD76D91E862D48A48AB" pageId="0" pageNumber="1">
<taxonomicName id="0AA53EF35FA194E3469BAF82475333D9" authorityName="Ospina-Sánchez &amp; Palacios-Vargas &amp; González" authorityYear="2020" class="Collembola" family="Neanuridae" genus="Furculanurida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Furculanurida bistribus" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bistribus">
<emphasis id="B6DCFAC66BD826E2CB9353BDEF3167B3" italics="true" pageId="0" pageNumber="1">Furculanurida bistribus</emphasis>
</taxonomicName>
sp. nov. was extracted from leaf litter and mosses in both dry and rainy seasons during November 2014, and May and August 2015.
</paragraph>

119
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Vitsika manifesta
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<paragraph pageId="83" pageNumber="84" type="mainText">(Figs 102-104, Map 156)</paragraph>
<paragraph pageId="83" pageNumber="84" type="mainText">WORKER (holotype in parentheses). TL 3.3-4.1 (4.0), HL 0.72-0.90 (0.84), HW 0.62-0.76 (0.72), CI 83-88 (86), SL 0.56-0.68 (0.62), SI 80-90 (86), PW 0.48-0.62 (0.58), WL 0.92-1.18 (1.04) (15 measured).</paragraph>
<paragraph pageId="83" pageNumber="84" type="mainText">Eye with 5-8 rows of ommatidia, and with 6-10 ommatidia in the longest row; EL 0.15-0.20 (EL/HW 0.23-0.28). MfL 0.68-0.88 (MfL/HW 1.21-1.32). Oblique costulae on mesopleuron conspicuous. Diameter of annulus of propodeal spiracle is usually slightly greater than the thickness of the propodeal spine at its midlength. Propodeal declivity transversely costulate. In posterior view the petiole node tall and slender, its dorsal margin rising to a narrowly rounded median peak. Height of petiole node in posterior view (from midpoint of the dorsal margin of the foramen to the highest point of the node) 1.10-1.23 x its maximum width. In dorsal view postpetiole 1.07-1.15 x broader than long; maximum width of postpetiole 1.00-1.10 x the length of a propodeal spine; maximum width of postpetiole 0.90-1.04 x the distance between the apices of the propodeal spines. Side of petiole node usually, but not always, with longitudinal costulae; when present the costulae tend to overlap onto the posterior surface of the node, at least posterolaterally. Length of postpetiole node less than the height of the segment. Disc of postpetiole mostly smooth, but longitudinal costulae are usually visible laterodorsally. Full adult colour rich reddish brown to dark brown.</paragraph>
<paragraph pageId="83" pageNumber="84" type="mainText">QUEEN (gyne). Ergatoid (e.g. CASENT0914197), alate, and intermediate (e.g. CASENT0914198) forms are known. Ergatoid: HL 0.86, HW 0.75, CI 87, SL 0.64, SI 85, PW 0.55, WL 1.12, MfL 0.84. The ergatoid has three ocelli. The mesosoma is almost worker-like except that the mesonotum is slightly enlarged compared to the worker. The mesopleural sulcus is absent. Intermediate: HL 0.84, HW 0.74, CI 88, SL 0.60, SI 81, PW 0.56, maximum length of mesoscutum 0.40, maximum width of mesoscutum 0.40, WL 1.08, MfL 0.78. Three ocelli present. Mesosoma with a full complement of flight sclerites, but the individual sclerites small, considerably reduced from size seen in alate queen, and without trace of wing articulations. Mesopleuron with a transverse sulcus. Alate: HL 0.87, HW 0.79, CI 91, SL 0.64, SI 81, PW 0.68, maximum length of mesoscutum 0.70, maximum width of mesoscutum 0.68, WL 1.30, MfL 0.86. Three ocelli present. Mesosoma with a full complement of full-sized flight sclerites, with wing articulation present. Mesopleuron with a broad transverse sulcus. Note that in all forms most measurements are within the worker range, but in the fully alate form PW and WL are greatly increased. Also, the mesoscutum is much larger in the alate than in the intermediate.</paragraph>
<paragraph pageId="83" pageNumber="84" type="mainText">MALE: unknown.</paragraph>
<subSubSection id="0A69213BCA920227CCCD2AC7D25AC783" type="description">
<paragraph id="28A4B22E12ED25E8C44B6808D0DF80E2" pageId="83" pageNumber="84" type="mainText">(Figs 102-104, Map 156)</paragraph>
<paragraph id="6FB221EE8AFE29DA4E6C80B4913598B2" pageId="83" pageNumber="84" type="mainText">WORKER (holotype in parentheses). TL 3.3-4.1 (4.0), HL 0.72-0.90 (0.84), HW 0.62-0.76 (0.72), CI 83-88 (86), SL 0.56-0.68 (0.62), SI 80-90 (86), PW 0.48-0.62 (0.58), WL 0.92-1.18 (1.04) (15 measured).</paragraph>
<paragraph id="461C1828607E67FEF0D56693344942C5" pageId="83" pageNumber="84" type="mainText">Eye with 5-8 rows of ommatidia, and with 6-10 ommatidia in the longest row; EL 0.15-0.20 (EL/HW 0.23-0.28). MfL 0.68-0.88 (MfL/HW 1.21-1.32). Oblique costulae on mesopleuron conspicuous. Diameter of annulus of propodeal spiracle is usually slightly greater than the thickness of the propodeal spine at its midlength. Propodeal declivity transversely costulate. In posterior view the petiole node tall and slender, its dorsal margin rising to a narrowly rounded median peak. Height of petiole node in posterior view (from midpoint of the dorsal margin of the foramen to the highest point of the node) 1.10-1.23 x its maximum width. In dorsal view postpetiole 1.07-1.15 x broader than long; maximum width of postpetiole 1.00-1.10 x the length of a propodeal spine; maximum width of postpetiole 0.90-1.04 x the distance between the apices of the propodeal spines. Side of petiole node usually, but not always, with longitudinal costulae; when present the costulae tend to overlap onto the posterior surface of the node, at least posterolaterally. Length of postpetiole node less than the height of the segment. Disc of postpetiole mostly smooth, but longitudinal costulae are usually visible laterodorsally. Full adult colour rich reddish brown to dark brown.</paragraph>
<paragraph id="CE465CE3C106F5481EA2C8C600128431" pageId="83" pageNumber="84" type="mainText">QUEEN (gyne). Ergatoid (e.g. CASENT0914197), alate, and intermediate (e.g. CASENT0914198) forms are known. Ergatoid: HL 0.86, HW 0.75, CI 87, SL 0.64, SI 85, PW 0.55, WL 1.12, MfL 0.84. The ergatoid has three ocelli. The mesosoma is almost worker-like except that the mesonotum is slightly enlarged compared to the worker. The mesopleural sulcus is absent. Intermediate: HL 0.84, HW 0.74, CI 88, SL 0.60, SI 81, PW 0.56, maximum length of mesoscutum 0.40, maximum width of mesoscutum 0.40, WL 1.08, MfL 0.78. Three ocelli present. Mesosoma with a full complement of flight sclerites, but the individual sclerites small, considerably reduced from size seen in alate queen, and without trace of wing articulations. Mesopleuron with a transverse sulcus. Alate: HL 0.87, HW 0.79, CI 91, SL 0.64, SI 81, PW 0.68, maximum length of mesoscutum 0.70, maximum width of mesoscutum 0.68, WL 1.30, MfL 0.86. Three ocelli present. Mesosoma with a full complement of full-sized flight sclerites, with wing articulation present. Mesopleuron with a broad transverse sulcus. Note that in all forms most measurements are within the worker range, but in the fully alate form PW and WL are greatly increased. Also, the mesoscutum is much larger in the alate than in the intermediate.</paragraph>
<paragraph id="7E6A1837252977D44EA69C46CE1CDB55" pageId="83" pageNumber="84" type="mainText">MALE: unknown.</paragraph>
</subSubSection>
<subSubSection type="materials_examined">
<paragraph pageId="83" pageNumber="84" type="mainText">Holotype worker, Madagascar: Prov. Antsiranana, P.N. Marojejy, 25.7 km. 32° NNE Andapa, 21-23.xi.2003, 1575 m., 14°26.7'S, 49°44.5'E, sifted litter, montane rainforest, BLF 9242(9), CASENT0040789 (Fisher et al.) (CASC).</paragraph>
<paragraph pageId="83" pageNumber="84" type="mainText">Paratypes. 7 workers with same data as holotype, and all BLF 9242; (L0) CASENT0040797, (4) CASENT0040784, (6) CASENT0040790, (10) CASENT0040791, (13) CASENT0040792, (16) CASENT0040793, (22) CASENT0040795 (CASC, BMNH).</paragraph>
<subSubSection id="58FF78978010958387F0AEC2FD3E7440" type="materials_examined">
<paragraph id="311AA70B3D7AC9E164909F20F955C65D" pageId="83" pageNumber="84" type="mainText">Holotype worker, Madagascar: Prov. Antsiranana, P.N. Marojejy, 25.7 km. 32° NNE Andapa, 21-23.xi.2003, 1575 m., 14°26.7'S, 49°44.5'E, sifted litter, montane rainforest, BLF 9242(9), CASENT0040789 (Fisher et al.) (CASC).</paragraph>
<paragraph id="9947690377638421EA11554C982039B0" pageId="83" pageNumber="84" type="mainText">Paratypes. 7 workers with same data as holotype, and all BLF 9242; (L0) CASENT0040797, (4) CASENT0040784, (6) CASENT0040790, (10) CASENT0040791, (13) CASENT0040792, (16) CASENT0040793, (22) CASENT0040795 (CASC, BMNH).</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="mainText">
<subSubSection id="B39401283B7BF3D383A88D8723951FB0" type="discussion">
<paragraph id="02C7188992B0107F2107302D9BA4C73F" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="mainText">
As seen in posterior view, the high, narrow petiole node, whose dorsal margin rises to a narrowly rounded median peak, is immediately diagnostic of
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="manifesta">manifesta</taxonomicName>
<taxonomicName id="1DC76E994D972247E0F73150E8749B34" genus="Vitsika" pageId="83" pageNumber="84" rank="species" species="manifesta">manifesta</taxonomicName>
. The height of the node in posterior view (from midpoint of the dorsal margin of the foramen to the highest point of the node) is 1.10-1.23 x its maximum width; i.e. the node
<pageBreakToken pageId="84" pageNumber="85" start="start">is</pageBreakToken>
<pageBreakToken id="886A7DFA00E7263A37AED1B1AD6A77B5" pageId="84" pageNumber="85" start="start">is</pageBreakToken>
always higher than wide. In all other species of
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">Vitsika</taxonomicName>
<taxonomicName id="7A8E4B9998D78CB9523037A70E8ECBB3" genus="Vitsika" pageId="83" pageNumber="84" rank="genus">Vitsika</taxonomicName>
, with the node in posterior view the dorsal margin is always flat to broadly, shallowly convex (concave in
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="incisura">incisura</taxonomicName>
<taxonomicName id="86DC07FC06E5297258135FACB6BA219A" genus="Vitsika" pageId="83" pageNumber="84" rank="species" species="incisura">incisura</taxonomicName>
). In the remainder of the genus the total range of height of node is 0.66-1.10 x its maximum width. Only
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="labes">labes</taxonomicName>
<taxonomicName id="683552AACDDB4115DFDA5443C89A2ECF" genus="Vitsika" pageId="83" pageNumber="84" rank="species" species="labes">labes</taxonomicName>
has an upper value of 1.10, but here the postpetiole node in profile is distinctly longer than the height of the segment.
</paragraph>
</subSubSection>
<subSubSection type="multiple">
<caption pageId="84" pageNumber="85">
<paragraph pageId="84" pageNumber="85" type="caption">
<subSubSection id="34C8351A355D7C1CC9776C9F2EB7D0E5" type="multiple">
<caption id="28818F030766D06B27D49C17E4389691" pageId="84" pageNumber="85">
<paragraph id="60FE4B0EDD3E5FF20A0DFF5CA930F220" pageId="84" pageNumber="85" type="caption">
FIGURES 102-104. Lateral, full face and dorsal view of body.
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="manifesta">Vitsika manifesta</taxonomicName>
<taxonomicName id="2537787291ABA2E2A479E98C89CCA592" genus="Vitsika" pageId="84" pageNumber="85" rank="species" species="manifesta">Vitsika manifesta</taxonomicName>
holotype worker CASENT0040789.
</paragraph>
</caption>
</subSubSection>
<subSubSection type="reference_group">
<paragraph pageId="84" pageNumber="85" type="mainText">
<subSubSection id="34CFB3E0B4DC227DF710A82AA718610F" type="reference_group">
<paragraph id="3C05E5E83DD8E9BEE5F435456634505D" pageId="84" pageNumber="85" type="mainText">
All material of
<taxonomicName authority="Bolton &amp; Fisher, 2014" authorityName="Bolton &amp; Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Vitsika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="manifesta">manifesta</taxonomicName>
<taxonomicName id="BE430DB897AA319B7309C0C62CADE936" genus="Vitsika" pageId="84" pageNumber="85" rank="species" species="manifesta">manifesta</taxonomicName>
is from montane shrubland and rainforest, at considerable altitude (1565-2000 m.). Specimens have been retrieved from litter samples and yellow pan traps.
</paragraph>
</subSubSection>
<subSubSection type="materials_examined">
<paragraph pageId="84" pageNumber="85" type="mainText">Non-paratypic material examined. Madagascar: Prov. Antsiranana, P.N. Marojejy, NNE Andapa (Fisher et al.); Prov. Antsiranana, RNI Marojejy, NW Manantenina (E.L. Quinter); Prov. Antsiranana, WSW Befingotra, Res. Anjanaharibe-Sud (B.L. Fisher).</paragraph>
<subSubSection id="D439EF4BB6550344455F8B021945F8C2" type="materials_examined">
<paragraph id="F61211792958328148655D16D69519B3" pageId="84" pageNumber="85" type="mainText">Non-paratypic material examined. Madagascar: Prov. Antsiranana, P.N. Marojejy, NNE Andapa (Fisher et al.); Prov. Antsiranana, RNI Marojejy, NW Manantenina (E.L. Quinter); Prov. Antsiranana, WSW Befingotra, Res. Anjanaharibe-Sud (B.L. Fisher).</paragraph>
</subSubSection>
</treatment>
</document>

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