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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitat and large-sized plant of Begonia giganticaulis D. K. Tian &amp; W. G. Wang, sp. nov. A habitat showing plants (arrows indicate) growing along stream bank B flowering plant growing along slope of valley C one of the tallest individuals with Dr. Dai-Ke Tian. (Photos A by Dai-Ke Tian B by Shi-Wei Guo C by Qing-Gong Mao)." figureDoi="10.3897/phytokeys.187.75854.figure1" httpUri="https://binary.pensoft.net/fig/629006" pageId="0" pageNumber="189">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Begonia giganticaulis D. K. Tian &amp; W. G. Wang, sp. nov. A one of the single tallest plants cut into four sections B main stem base C stems showing colour of nodal cross-sections D main stem with much expanded node and whitish-green lines or spots E expanded node on terminal branch F, G male plant branches showing inflorescences and different colours H female branches I adaxially (left) nearly glabrous and abaxially sparse hairs on veins (right, arrows indicate) on blade surfaces J stipules showing shape and colour. (Photo F by Wen-Guang Wang; others by Dai-Ke Tian)." figureDoi="10.3897/phytokeys.187.75854.figure2" httpUri="https://binary.pensoft.net/fig/629007" pageId="0" pageNumber="189">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Flower and fruit morphology of B. giganticaulis compared with its close species B. longifolia and B. acetosella A-H Begonia giganticaulis A staminate flowers with pinkish outer tepals B staminate flowers with white tepals C, D pistillate flower E ovary sections showing different colour F fruits on branch G, H dorsal and front views of fruits I-K B. longifolia I flowering and fruiting branch J fruits showing short horns K ovary dissection L-O B. acetosella L staminate flower M Pistillate flower N, O fruits with short horns or wings. (Photos C by Shi-Wei Guo E (left) L, M &amp; O by Wen-Guang Wang; others by Dai-Ke Tian)." figureDoi="10.3897/phytokeys.187.75854.figure3" httpUri="https://binary.pensoft.net/fig/629008" pageId="0" pageNumber="189">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Illustration of Begonia giganticaulis D. K. Tian &amp; W. G. Wang, sp. nov. (Drawn by Mr. Zhi-Min Li) A male flowering branches B female flowering branches C main stem line spots, much expanded node and internode base D expanded node and internode base on small upper branches E leaf blade F leaf (abaxial), showing sparse hairs on veins G stipule H staminate flowers I, J pistillate flower K side view of androecium L stamens M ovary and stigmas N fruit O dissection of ovary showing placentae." figureDoi="10.3897/phytokeys.187.75854.figure4" httpUri="https://binary.pensoft.net/fig/629009" pageId="0" pageNumber="189">
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<materialsCitation collectingDate="2020-09-10" country="China" county="Autonomous Region" elevation="1320" latitude="29.3525" location="Beibeng town" longLatPrecision="20" longitude="95.18917" municipality="Medog county" specimenCount="♀" stateProvince="Xizang" typeStatus="holotype">
<emphasis bold="true" pageId="0" pageNumber="189">
<collectingCountry name="China">China</collectingCountry>
<paragraph id="5CA119389FC6297853DFD42B808C4AB2" pageId="0" pageNumber="189">
<materialsCitation id="7F2212FBCABC490D9F8924C66FA452A1" collectingDate="2020-09-10" country="China" county="Autonomous Region" elevation="1320" latitude="29.3525" location="Beibeng town" longLatPrecision="20" longitude="95.18917" municipality="Medog county" specimenCount="♀" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="55B1CEF716DBB84F225812BCFD10DE1D" bold="true" pageId="0" pageNumber="189">
<collectingCountry id="44B68EF438DE4ACAF0AA033ECC181EBE" name="China">China</collectingCountry>
</emphasis>
.
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
<collectingRegion id="9B67AEDF2152565EA91AE23E25E0A8EE" country="China" name="Xizang">Xizang</collectingRegion>
(
<collectingRegion country="China" name="Xizang">Tibet</collectingRegion>
<collectingRegion id="98E43775BA98D08AE0C8C1149FE6A5E1" country="China" name="Xizang">Tibet</collectingRegion>
)
<collectingCounty>Autonomous Region</collectingCounty>
<collectingCounty id="5A8B897A28E120923D675BD9E830D678">Autonomous Region</collectingCounty>
:
<collectingMunicipality>
<normalizedToken originalValue="Mêdog">Medog</normalizedToken>
<collectingMunicipality id="5D18EDEF070C7D83716DDDBA2FE01761">
<normalizedToken id="B7F60AE654B882EC1D8BDBC1B8490C0F" originalValue="Mêdog">Medog</normalizedToken>
county
</collectingMunicipality>
(墨脱县),
<location LSID="urn:lsid:plazi:treatment:2D720B89EF535D5488FA0D47E8F10C53:DDB7FED97F165491874AD18C4D307C2A" country="China" county="Autonomous Region" latitude="29.3525" longLatPrecision="20" longitude="95.18917" municipality="Medog county" name="Beibeng town" stateProvince="Xizang">Beibeng town</location>
<location id="DDB7FED97F165491874AD18C4D307C2A" LSID="urn:lsid:plazi:treatment:2D720B89EF535D5488FA0D47E8F10C53:DDB7FED97F165491874AD18C4D307C2A" country="China" county="Autonomous Region" latitude="29.3525" longLatPrecision="20" longitude="95.18917" municipality="Medog county" name="Beibeng town" stateProvince="Xizang">Beibeng town</location>
(背崩乡),
<location LSID="urn:lsid:plazi:treatment:2D720B89EF535D5488FA0D47E8F10C53:DC8C202644C0BAE73EAF5906A5896E06" country="China" county="Autonomous Region" latitude="29.3525" longLatPrecision="20" longitude="95.18917" municipality="Medog county" name="Baimu Xiri river" stateProvince="Xizang">Baimu Xiri river</location>
<location id="DC8C202644C0BAE73EAF5906A5896E06" LSID="urn:lsid:plazi:treatment:2D720B89EF535D5488FA0D47E8F10C53:DC8C202644C0BAE73EAF5906A5896E06" country="China" county="Autonomous Region" latitude="29.3525" longLatPrecision="20" longitude="95.18917" municipality="Medog county" name="Baimu Xiri river" stateProvince="Xizang">Baimu Xiri river</location>
(白母西日河), forest slope of river valley or
<normalizedToken originalValue="waters">water's</normalizedToken>
<normalizedToken id="CA781B1A82909205ABC05C6CA7147744" originalValue="waters">water's</normalizedToken>
edge along stream,
<geoCoordinate degrees="29" direction="north" minutes="21" orientation="latitude" precision="15" seconds="9" value="29.3525">29°21'9&quot;N</geoCoordinate>
<geoCoordinate id="CB67B4B65C753F8827F2B337AD0480A6" degrees="29" direction="north" minutes="21" orientation="latitude" precision="15" seconds="9" value="29.3525">29°21'9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="11" orientation="longitude" precision="15" seconds="21" value="95.18917">95°11'21&quot;E</geoCoordinate>
<geoCoordinate id="1F0E0DBAEA424BCF01F6FFF1ADF0DFAB" degrees="95" direction="east" minutes="11" orientation="longitude" precision="15" seconds="21" value="95.18917">95°11'21&quot;E</geoCoordinate>
, elev.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">1320 m</elevation>
<quantity id="C2F4E160C04CCB8AD5871F092E4B4D21" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">
<elevation id="A4CB67274DFB771D7EEDD7FEC0A846BB" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="1320.0">1320 m</elevation>
</quantity>
,
<collectingDate value="2020-09-10">10 September 2020</collectingDate>
<collectingDate id="13D60F72628E3D1273CF0BE6E466C2D9" value="2020-09-10">10 September 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu, TDK4773-A</emphasis>
<emphasis id="2CD19AE42C5AFA70FE7A43F51B087FF5" italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu, TDK4773-A</emphasis>
(
<typeStatus>holotype</typeStatus>
<typeStatus id="A2BF20EA85C189218721F9A7754F0DBF">holotype</typeStatus>
CSH! Barcode number: 0180561,
<specimenCount></specimenCount>
<specimenCount id="5E2DF6916F8131E14AADCCE8001F67EB"></specimenCount>
)
</materialsCitation>
</paragraph>
<caption doi="10.3897/phytokeys.187.75854.figure1" httpUri="https://binary.pensoft.net/fig/629006" pageId="0" pageNumber="189" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figure 1.</emphasis>
<caption id="2FA0B5A123103CBF2EA052F771D04E3C" doi="10.3897/phytokeys.187.75854.figure1" httpUri="https://binary.pensoft.net/fig/629006" pageId="0" pageNumber="189" start="Figure 1" startId="F1">
<paragraph id="692BE212E2CE57C4F4EF256E4A91EDDA" pageId="0" pageNumber="189">
<emphasis id="D82EFF3AF245120506CB733D66EA9703" bold="true" pageId="0" pageNumber="189">Figure 1.</emphasis>
Habitat and large-sized plant of
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<taxonomicName id="BF1D517205A76CB87F9B430D0E365EFB" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="9ABE5111ED7633A745CF94CC9FE37992" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
D.K.Tian &amp; W.G.Wang, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="189">A</emphasis>
<emphasis id="4454694343277BE0724B948FB31259A1" bold="true" pageId="0" pageNumber="189">A</emphasis>
habitat showing plants (arrows indicate) growing along stream bank
<emphasis bold="true" pageId="0" pageNumber="189">B</emphasis>
<emphasis id="7091548543118F6C739D78612FE1698B" bold="true" pageId="0" pageNumber="189">B</emphasis>
flowering plant growing along slope of valley
<emphasis bold="true" pageId="0" pageNumber="189">C</emphasis>
<emphasis id="AC0C150A08695E979F8F5B51E9835DED" bold="true" pageId="0" pageNumber="189">C</emphasis>
one of the tallest individuals with Dr. Dai-Ke Tian. (Photos
<emphasis bold="true" pageId="0" pageNumber="189">A</emphasis>
<emphasis id="EA14ECAFC19EC298E5A2F0523252B184" bold="true" pageId="0" pageNumber="189">A</emphasis>
by Dai-Ke Tian
<emphasis bold="true" pageId="0" pageNumber="189">B</emphasis>
<emphasis id="E652192D6E5F27D03117A7F34C97F019" bold="true" pageId="0" pageNumber="189">B</emphasis>
by Shi-Wei Guo
<emphasis bold="true" pageId="0" pageNumber="189">C</emphasis>
<emphasis id="4AF90BCDB908D7E95A80D063CE9F1141" bold="true" pageId="0" pageNumber="189">C</emphasis>
by Qing-Gong Mao).
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.187.75854.figure2" httpUri="https://binary.pensoft.net/fig/629007" pageId="0" pageNumber="189" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figure 2.</emphasis>
<caption id="E94C36C2430F041FE39C7774E1CEC186" doi="10.3897/phytokeys.187.75854.figure2" httpUri="https://binary.pensoft.net/fig/629007" pageId="0" pageNumber="189" start="Figure 2" startId="F2">
<paragraph id="F10FE1A9A1C3B3DAFDB9145B2132D621" pageId="0" pageNumber="189">
<emphasis id="67A6989C74B2D9808DFB7CA2033719EA" bold="true" pageId="0" pageNumber="189">Figure 2.</emphasis>
Morphology of
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<taxonomicName id="526E6DEA1FBC92A9E9AABA7A55C83608" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="3E900F48F73E9518FBAD39E9803C0529" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
D.K.Tian &amp; W.G.Wang, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="189">A</emphasis>
<emphasis id="C2BD6B8A529473BD5D70F4ED225D03B1" bold="true" pageId="0" pageNumber="189">A</emphasis>
one of the single tallest plants cut into four sections
<emphasis bold="true" pageId="0" pageNumber="189">B</emphasis>
<emphasis id="FFA4C4F3ED14255FE38A6CBEB971127E" bold="true" pageId="0" pageNumber="189">B</emphasis>
main stem base
<emphasis bold="true" pageId="0" pageNumber="189">C</emphasis>
<emphasis id="2F6CE59F4428C6A728C6199562492A79" bold="true" pageId="0" pageNumber="189">C</emphasis>
stems showing colour of nodal cross-sections
<emphasis bold="true" pageId="0" pageNumber="189">D</emphasis>
<emphasis id="4B0899FFCA6A29E9C219106FD0B4140B" bold="true" pageId="0" pageNumber="189">D</emphasis>
main stem with much expanded node and whitish-green lines or spots
<emphasis bold="true" pageId="0" pageNumber="189">E</emphasis>
<emphasis id="B780D7E77D95065E1308C8C94099AFC3" bold="true" pageId="0" pageNumber="189">E</emphasis>
expanded node on terminal branch
<emphasis bold="true" pageId="0" pageNumber="189">F, G</emphasis>
<emphasis id="7FA5D21CA619CC42E2F91D92C81DC2C0" bold="true" pageId="0" pageNumber="189">F, G</emphasis>
male plant branches showing inflorescences and different colours
<emphasis bold="true" pageId="0" pageNumber="189">H</emphasis>
<emphasis id="E3F072D8664574F37334811A6F3EC8BE" bold="true" pageId="0" pageNumber="189">H</emphasis>
female branches
<emphasis bold="true" pageId="0" pageNumber="189">I</emphasis>
<emphasis id="916BECF66D1CAC7B4B282BC366854E6D" bold="true" pageId="0" pageNumber="189">I</emphasis>
adaxially (left) nearly glabrous and abaxially sparse hairs on veins (right, arrows indicate) on blade surfaces
<emphasis bold="true" pageId="0" pageNumber="189">J</emphasis>
<emphasis id="43DDA592E5F12E3E57AC837B8F9656D0" bold="true" pageId="0" pageNumber="189">J</emphasis>
stipules showing shape and colour. (Photo
<emphasis bold="true" pageId="0" pageNumber="189">F</emphasis>
<emphasis id="328D5247EB64F2B6095DF8414BD516C2" bold="true" pageId="0" pageNumber="189">F</emphasis>
by Wen-Guang Wang; others by Dai-Ke Tian).
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.187.75854.figure3" httpUri="https://binary.pensoft.net/fig/629008" pageId="0" pageNumber="189" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figure 3.</emphasis>
<caption id="2322B43ED6F4C6E0DA8DABD8C00C7CEF" doi="10.3897/phytokeys.187.75854.figure3" httpUri="https://binary.pensoft.net/fig/629008" pageId="0" pageNumber="189" start="Figure 3" startId="F3">
<paragraph id="6A5FCA7B5956D876D549182F2E61E1F5" pageId="0" pageNumber="189">
<emphasis id="3142E7BB56F5E549769841EBFD1F27E4" bold="true" pageId="0" pageNumber="189">Figure 3.</emphasis>
Flower and fruit morphology of
<taxonomicName lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
<taxonomicName id="65B7449770F48C7C4056B01645ACF8F1" lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis id="F0431E4C1992656D412AAF72961A9E83" italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
</taxonomicName>
compared with its close species
<taxonomicName lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
<taxonomicName id="66E13A504B6DE1C9DA1FA63CDBC53099" lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis id="442C4E536600A99E4A2D34B06E3B0258" italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
<taxonomicName id="F09DEC7D90C0CB08156652F90C9C4D7E" lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis id="C5CCB63AE081D531B2D558E734B1EC45" italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="189">A-H</emphasis>
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<emphasis id="0E773AF7DA1D67F43E7ED5ECBB780249" bold="true" pageId="0" pageNumber="189">A-H</emphasis>
<taxonomicName id="8589C4E5589BC2EAA84A30D27646D49B" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="B8696350B692EA41D7730154B303E86A" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="189">A</emphasis>
<emphasis id="A46FC6F2492C3F432A9EEFC4074751F5" bold="true" pageId="0" pageNumber="189">A</emphasis>
staminate flowers with pinkish outer tepals
<emphasis bold="true" pageId="0" pageNumber="189">B</emphasis>
<emphasis id="FBA2637CCA7E0557022EFBED43D860AB" bold="true" pageId="0" pageNumber="189">B</emphasis>
staminate flowers with white tepals
<emphasis bold="true" pageId="0" pageNumber="189">C, D</emphasis>
<emphasis id="FFF234C70492BD58E48C2B6035699FE6" bold="true" pageId="0" pageNumber="189">C, D</emphasis>
pistillate flower
<emphasis bold="true" pageId="0" pageNumber="189">E</emphasis>
<emphasis id="703973840013F1F13597662D8E9C8B7D" bold="true" pageId="0" pageNumber="189">E</emphasis>
ovary sections showing different colour
<emphasis bold="true" pageId="0" pageNumber="189">F</emphasis>
<emphasis id="066AB1283974F722FDDBD9EF92CDA715" bold="true" pageId="0" pageNumber="189">F</emphasis>
fruits on branch
<emphasis bold="true" pageId="0" pageNumber="189">G, H</emphasis>
<emphasis id="1A5243FCF24B3DA59155AAAB07DFF1EB" bold="true" pageId="0" pageNumber="189">G, H</emphasis>
dorsal and front views of fruits
<emphasis bold="true" pageId="0" pageNumber="189">I-K</emphasis>
<taxonomicName lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
<emphasis id="4BAC11ECB5659666CED10E2AF553FA1A" bold="true" pageId="0" pageNumber="189">I-K</emphasis>
<taxonomicName id="9395F7F6AFE6DC9C71458B73F8A796B2" lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis id="FDF225263701AB06BEF78D1C37AEF187" italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="189">I</emphasis>
<emphasis id="724E890192A208696EC92640FE2FD4FF" bold="true" pageId="0" pageNumber="189">I</emphasis>
flowering and fruiting branch
<emphasis bold="true" pageId="0" pageNumber="189">J</emphasis>
<emphasis id="99917140E491E9952FA2FBA213345833" bold="true" pageId="0" pageNumber="189">J</emphasis>
fruits showing short horns
<emphasis bold="true" pageId="0" pageNumber="189">K</emphasis>
<emphasis id="634E85CF7A74C8BC98BDED31A73EAF8C" bold="true" pageId="0" pageNumber="189">K</emphasis>
ovary dissection
<emphasis bold="true" pageId="0" pageNumber="189">L-O</emphasis>
<taxonomicName lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
<emphasis id="C225AC813D98AC280BFDDE036D9E4DD9" bold="true" pageId="0" pageNumber="189">L-O</emphasis>
<taxonomicName id="F0532F687C0C798454F37245E6B55316" lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis id="4B98853D7DDE7852F65E618657F5C5AB" italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="189">L</emphasis>
<emphasis id="ACE5DE63562CFF0E92AD2DCB17729239" bold="true" pageId="0" pageNumber="189">L</emphasis>
staminate flower
<emphasis bold="true" pageId="0" pageNumber="189">M</emphasis>
<emphasis id="2908B6F9E9739CE861FD3560E8EBF780" bold="true" pageId="0" pageNumber="189">M</emphasis>
Pistillate flower
<emphasis bold="true" pageId="0" pageNumber="189">N, O</emphasis>
<emphasis id="35D495BA4E84CB919DDD0E487E13849A" bold="true" pageId="0" pageNumber="189">N, O</emphasis>
fruits with short horns or wings. (Photos
<emphasis bold="true" pageId="0" pageNumber="189">C</emphasis>
<emphasis id="CDC21A976A1F2F57FA9F7E3ACC7ACE9A" bold="true" pageId="0" pageNumber="189">C</emphasis>
by Shi-Wei Guo
<emphasis bold="true" pageId="0" pageNumber="189">E</emphasis>
<emphasis id="9EBD7BC4954B172A6DA2215E3716B995" bold="true" pageId="0" pageNumber="189">E</emphasis>
(left)
<emphasis bold="true" pageId="0" pageNumber="189">L, M</emphasis>
<emphasis id="7A06AD3E55650DB5BCD03FF21DAE47CE" bold="true" pageId="0" pageNumber="189">L, M</emphasis>
&amp;
<emphasis bold="true" pageId="0" pageNumber="189">O</emphasis>
<emphasis id="C509234CD4ADCF36DC35090016375292" bold="true" pageId="0" pageNumber="189">O</emphasis>
by Wen-Guang Wang; others by Dai-Ke Tian).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="diagnosis">
<paragraph pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="0B459DD9F08A89E1D27E5C477EF97B7A" pageId="0" pageNumber="189" type="diagnosis">
<paragraph id="ECC384D45FFB92AC6C62CB8659A1ED36" pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph id="E7B27FEF89F444CC3E5381BA88889325" pageId="0" pageNumber="189">
The new species is mostly similar to
<taxonomicName lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
<taxonomicName id="13E595817BC707F66BDB1E123D7516F8" lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis id="E71D1E5D20A1963552E84CFA98DB4761" italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
<taxonomicName id="9A4581DEFC84143BB8B3D0B068F3AF6B" lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis id="5A718B666F31F131A8344870F0D84CDF" italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
</taxonomicName>
, but clearly differs from the former mainly by its dioecious (vs. monoecious), taller (to 4 m vs. less than 2 m) plants, longer (vs. shorter) inflorescence, and unique shape of fruits, and differs from the latter mainly by its taller (to 4 m vs. less than 2 m) plants, late and longer (Jun. to Oct. vs. Mar. to Apr.) flowering time, longer (6-20 mm vs. 5-12 mm) pedicel, 6 (vs. 4) tepals of pistillate flower and 3 (vs. 4)-loculed ovary (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological comparison of Begonia giganticaulis, B. longifolia and B. acetosella." httpUri="http://table.plazi.org/id/22D325F41DBAD6A0E6FC7DC3CA1598C8" pageId="0" pageNumber="189" tableUuid="22D325F41DBAD6A0E6FC7DC3CA1598C8">3</tableCitation>
<tableCitation id="018815DBCEE387A0062941B7F21A779E" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological comparison of Begonia giganticaulis, B. longifolia and B. acetosella." httpUri="http://table.plazi.org/id/22D325F41DBAD6A0E6FC7DC3CA1598C8" pageId="0" pageNumber="189" tableUuid="22D325F41DBAD6A0E6FC7DC3CA1598C8">3</tableCitation>
, Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Flower and fruit morphology of B. giganticaulis compared with its close species B. longifolia and B. acetosella A-H Begonia giganticaulis A staminate flowers with pinkish outer tepals B staminate flowers with white tepals C, D pistillate flower E ovary sections showing different colour F fruits on branch G, H dorsal and front views of fruits I-K B. longifolia I flowering and fruiting branch J fruits showing short horns K ovary dissection L-O B. acetosella L staminate flower M Pistillate flower N, O fruits with short horns or wings. (Photos C by Shi-Wei Guo E (left) L, M &amp; O by Wen-Guang Wang; others by Dai-Ke Tian)." figureDoi="10.3897/phytokeys.187.75854.figure3" httpUri="https://binary.pensoft.net/fig/629008" pageId="0" pageNumber="189">3</figureCitation>
<figureCitation id="3841381EF0BB063E009434BC9A0E90C5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Flower and fruit morphology of B. giganticaulis compared with its close species B. longifolia and B. acetosella A-H Begonia giganticaulis A staminate flowers with pinkish outer tepals B staminate flowers with white tepals C, D pistillate flower E ovary sections showing different colour F fruits on branch G, H dorsal and front views of fruits I-K B. longifolia I flowering and fruiting branch J fruits showing short horns K ovary dissection L-O B. acetosella L staminate flower M Pistillate flower N, O fruits with short horns or wings. (Photos C by Shi-Wei Guo E (left) L, M &amp; O by Wen-Guang Wang; others by Dai-Ke Tian)." figureDoi="10.3897/phytokeys.187.75854.figure3" httpUri="https://binary.pensoft.net/fig/629008" pageId="0" pageNumber="189">3</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="22D325F41DBAD6A0E6FC7DC3CA1598C8" httpUri="http://table.plazi.org/id/22D325F41DBAD6A0E6FC7DC3CA1598C8" pageId="0" pageNumber="189" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Table 3.</emphasis>
<caption id="22D325F41DBAD6A0E6FC7DC3CA1598C8" ID-Table-UUID="22D325F41DBAD6A0E6FC7DC3CA1598C8" httpUri="http://table.plazi.org/id/22D325F41DBAD6A0E6FC7DC3CA1598C8" pageId="0" pageNumber="189" start="Table 3" startId="T3">
<paragraph id="BE08981D3B6542C8038F6C425E804381" pageId="0" pageNumber="189">
<emphasis id="16399510C0282C7E1679083E837AB4BF" bold="true" pageId="0" pageNumber="189">Table 3.</emphasis>
Morphological comparison of
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<taxonomicName id="55C05A029B83571C31DE196A33091340" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="1D56616E11279B63CBA4CD936C0B1A9F" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
<taxonomicName id="EED3699A658918AAF865BAAB1F72D42A" lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis id="3A0F331843C2D0DD1AADB99ADCF7074E" italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
<taxonomicName id="6E86504D8D88175463C85CB6D7D6CFC4" lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis id="686FDFAE533F9E986C2AD37EC2C2A3F4" italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="189">
<table pageId="0" pageNumber="189">
<tr pageId="0" pageNumber="189">
<th colspan="1" pageId="0" pageNumber="189" rowspan="1">Character</th>
<th colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
<paragraph id="11584122EB8213330E499C78D5537995" pageId="0" pageNumber="189">
<table id="B8E3615DA5AA7ED0CBEDBBBD0715EE63" pageId="0" pageNumber="189">
<tr id="DADA38AC020599EFDC2546FE95911CBA" pageId="0" pageNumber="189">
<th id="CEF63BEF5B1A50EC3DFF6F8DCDABE723" colspan="1" pageId="0" pageNumber="189" rowspan="1">Character</th>
<th id="76609F4194EAFFD70A48DCFB7736C176" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName id="2A85C0A8EB26FD77CBAC8ACE1C939B4D" lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis id="84CD884FD58B2F9877F9C547F750FA68" italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
<th id="D1D6FFB09B948A1603D361C131CF18D2" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName id="A0FB25EB13BF278838FD13ACA6AA66FC" lsidName="B. longifolia" pageId="0" pageNumber="189" rank="species" species="longifolia">
<emphasis id="639041FF03AE782219FB9E46E8166280" italics="true" pageId="0" pageNumber="189">B. longifolia</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
<th id="3F9A83CF93E4A5043C38E0D55B51E246" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<taxonomicName id="DB8E4FD674C90BF5405C5D2C1F94C154" lsidName="B. acetosella" pageId="0" pageNumber="189" rank="species" species="acetosella">
<emphasis id="6BEAACE3BA929991754EA7143CA3FD88" italics="true" pageId="0" pageNumber="189">B. acetosella</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Plant</emphasis>
<tr id="E877D208DA36A7E8FA5B2ED5F436FFA7" pageId="0" pageNumber="189">
<td id="B0F1DE6A7A3358C4B91D64AD028B6C33" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="D201EB7C81265B3252AC3973FA51CAF5" bold="true" pageId="0" pageNumber="189">Plant</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="D675FDDCF066E8D65B8A3CD5B965A9B8" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="C27AC87433BE6C21877742D53D52F414" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="BA689CC86B6825F218A84298EDE59AB4" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">sexuality</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">dioecious</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">monoecious</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">dioecious</td>
<tr id="DDBFAF053A3310877EB052081ACC46A3" pageId="0" pageNumber="189">
<td id="594C27836DDA371F2CBEF00B27D11523" colspan="1" pageId="0" pageNumber="189" rowspan="1">sexuality</td>
<td id="66924D94DCC1761B55F28550FC9598E5" colspan="1" pageId="0" pageNumber="189" rowspan="1">dioecious</td>
<td id="C9EAF1BE41919B778DCF41F04DDF65B6" colspan="1" pageId="0" pageNumber="189" rowspan="1">monoecious</td>
<td id="63FB1FC7B26E2BDC421C44723FD63BF0" colspan="1" pageId="0" pageNumber="189" rowspan="1">dioecious</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">height (m)</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">up to 4</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">less than 2</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">less than 2</td>
<tr id="96850B3D8EC3A4B3A2B644E5B1CD8142" pageId="0" pageNumber="189">
<td id="8F901B4963DBEFF7D324D51D3D6A14A1" colspan="1" pageId="0" pageNumber="189" rowspan="1">height (m)</td>
<td id="0982F4FB55C4E2AF30F2A54C944FB374" colspan="1" pageId="0" pageNumber="189" rowspan="1">up to 4</td>
<td id="B1B6657B0F320B3C111E227F70991650" colspan="1" pageId="0" pageNumber="189" rowspan="1">less than 2</td>
<td id="328317A3185FC901962732C4BDCFA3CC" colspan="1" pageId="0" pageNumber="189" rowspan="1">less than 2</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Petiole</emphasis>
<tr id="D0DD1309BA22A3062777C8F2F920184E" pageId="0" pageNumber="189">
<td id="A8F81C7A3FB7178AA25088580901C9D7" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="D8A032607EF0AABB9360B6DBF5903F22" bold="true" pageId="0" pageNumber="189">Petiole</emphasis>
length (cm)
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">0.7-7</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">1-12</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">1-10</td>
<td id="511E89A60BCACF0AA7A94A40FC4DA9B1" colspan="1" pageId="0" pageNumber="189" rowspan="1">0.7-7</td>
<td id="9DEAA20661A15DBC0A72F3A27304728A" colspan="1" pageId="0" pageNumber="189" rowspan="1">1-12</td>
<td id="A87C42DB997CBAC0954DAA57CF0C71C8" colspan="1" pageId="0" pageNumber="189" rowspan="1">1-10</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Leaf blade surface</emphasis>
<tr id="650273DFFA2E6AEA8E875DD2EF239943" pageId="0" pageNumber="189">
<td id="5EC8CF70CC3CD80B318224297797D2E8" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="4E94DE4A60D53AA263C17160A97F9394" bold="true" pageId="0" pageNumber="189">Leaf blade surface</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">muriculate</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">glabrous to less muriculate</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">muriculate to hirsutulous</td>
<td id="404D86CC283D2506C8C0A1A89F2F2B83" colspan="1" pageId="0" pageNumber="189" rowspan="1">muriculate</td>
<td id="50A9FC2E097CF847DDC96711413F2D9D" colspan="1" pageId="0" pageNumber="189" rowspan="1">glabrous to less muriculate</td>
<td id="B6B90449EA3F9F38E669798898B35F44" colspan="1" pageId="0" pageNumber="189" rowspan="1">muriculate to hirsutulous</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Inflorescence</emphasis>
<tr id="A093DCDD7B24484939403E055C2A8266" pageId="0" pageNumber="189">
<td id="65688367E9F27B39EC37F840B72DF3E5" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="B2D0A4EE1074D3B3F66B806A53F388BA" bold="true" pageId="0" pageNumber="189">Inflorescence</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="A61B057613E9DE679D5864A6A1344C45" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="35DFC0E62E57123C6A05D44D26EDC2C0" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="EB2C61948820A395F247B1387E211AC6" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">peduncle length (mm)</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">7-15</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">4-10</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">2-10</td>
<tr id="AD0A6AD39DAEFBDE7955A99429054958" pageId="0" pageNumber="189">
<td id="4C30A01CAA1AF3C6CA504B061A1763EA" colspan="1" pageId="0" pageNumber="189" rowspan="1">peduncle length (mm)</td>
<td id="902003CAF5168AEEB66B961A3216AC41" colspan="1" pageId="0" pageNumber="189" rowspan="1">7-15</td>
<td id="C20B426FAF8EBBE74FD2B7435DDFAA40" colspan="1" pageId="0" pageNumber="189" rowspan="1">4-10</td>
<td id="152B02403FBF37BD519730EE46915FFA" colspan="1" pageId="0" pageNumber="189" rowspan="1">2-10</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">flower number</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">1-11</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">1-11(15)</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">1-3(5)</td>
<tr id="D78058247AB0488C2428F06FB79A1C83" pageId="0" pageNumber="189">
<td id="091276534E5FD6B495187DB9F776B273" colspan="1" pageId="0" pageNumber="189" rowspan="1">flower number</td>
<td id="D32D82A3CD0390BBA853B4385147DF0D" colspan="1" pageId="0" pageNumber="189" rowspan="1">1-11</td>
<td id="5C3464ED5F3081FEDFF43942F6A27F77" colspan="1" pageId="0" pageNumber="189" rowspan="1">1-11(15)</td>
<td id="A00048191FC9F8AAF75132E9321AC983" colspan="1" pageId="0" pageNumber="189" rowspan="1">1-3(5)</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Tepal number of pistillate flower</emphasis>
<tr id="47F37AD00A0577AE550D98209C761DE1" pageId="0" pageNumber="189">
<td id="F91F56D5AEFE95936728D190A7A731CF" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="3239D8869D781573B36A48855BCA3E91" bold="true" pageId="0" pageNumber="189">Tepal number of pistillate flower</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">4</td>
<td id="9E3F9A0D83F33B7EA5A8EFBF93E79D52" colspan="1" pageId="0" pageNumber="189" rowspan="1">6</td>
<td id="B3EBF8355A245AC10CADC8336137B627" colspan="1" pageId="0" pageNumber="189" rowspan="1">6</td>
<td id="957940FE8C6FDE17C5DBE1EC7D0C7166" colspan="1" pageId="0" pageNumber="189" rowspan="1">4</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Tepal colour</emphasis>
<tr id="6751EE5238F99A8C4EC25F98DABB6A83" pageId="0" pageNumber="189">
<td id="BF8B197A0B6E0D85D33691847C25FC1E" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="6377863F299F4581D5C45305A4A8DD05" bold="true" pageId="0" pageNumber="189">Tepal colour</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">pinkish to white</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">pinkish to white</td>
<td id="438EDED0F0739846BF6D739267A36E45" colspan="1" pageId="0" pageNumber="189" rowspan="1">pinkish to white</td>
<td id="897BFBCDF5F5F5C37B121B1058E80A95" colspan="1" pageId="0" pageNumber="189" rowspan="1">white</td>
<td id="97225E23E2659B0DC005C714D6A29FAF" colspan="1" pageId="0" pageNumber="189" rowspan="1">pinkish to white</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Ovary</emphasis>
<tr id="08D352983798252BD697B51AFD9F9A6F" pageId="0" pageNumber="189">
<td id="280F911A0B0F87A8709D4BF2DA11B6BF" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="BF898C530C8D3711A74C5CF1D01594AE" bold="true" pageId="0" pageNumber="189">Ovary</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">3-loculed</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">3-loculed</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">4-loculed</td>
<td id="EC560FFBF024EBE54EE1E39588AD0C39" colspan="1" pageId="0" pageNumber="189" rowspan="1">3-loculed</td>
<td id="5B643EF277EDBE9297E940A6E8F919A2" colspan="1" pageId="0" pageNumber="189" rowspan="1">3-loculed</td>
<td id="5DB97C27E6E601816538CC27BACBE45A" colspan="1" pageId="0" pageNumber="189" rowspan="1">4-loculed</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Pedicel length</emphasis>
<tr id="610C4BBA9BB72A60D4C9FF74B6F9D2E5" pageId="0" pageNumber="189">
<td id="A5B8C3FEEF5450133925677692F9D62C" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="7E20B83238AC3F11280804D78CAD1734" bold="true" pageId="0" pageNumber="189">Pedicel length</emphasis>
(mm)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="FEFE612103F6EA04525DBE7AB074FA25" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="CB3DE9FB213D08FA90CF10C6BED5FA12" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
<td id="9147F21C2B360D84EF89327B286E3907" colspan="1" isEmpty="true" pageId="0" pageNumber="189" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">male flower</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">10-20</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">5-12</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">8-12</td>
<tr id="E2FCC68C8C952E818AD77E0E0FEBC027" pageId="0" pageNumber="189">
<td id="4CB1D073D7523284BDCA0CE3A3315404" colspan="1" pageId="0" pageNumber="189" rowspan="1">male flower</td>
<td id="F2E577CF76B0CC007B1D17CF660D7166" colspan="1" pageId="0" pageNumber="189" rowspan="1">10-20</td>
<td id="8992841561DECEBBA7E4DEB13A087F28" colspan="1" pageId="0" pageNumber="189" rowspan="1">5-12</td>
<td id="2B52D6A67A8D4226F6D38F26083381F9" colspan="1" pageId="0" pageNumber="189" rowspan="1">8-12</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">female flower</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">6-12</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">5-12</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">5-10</td>
<tr id="71AC352DAFDC58A31226E08CF309A706" pageId="0" pageNumber="189">
<td id="334ABC51D3699FD8D976E1EFDFC3CAF3" colspan="1" pageId="0" pageNumber="189" rowspan="1">female flower</td>
<td id="77E97D5E4B69528717FEC47EF7AEBAA5" colspan="1" pageId="0" pageNumber="189" rowspan="1">6-12</td>
<td id="854DBBBB721B90A35F2543B57B6F4BBA" colspan="1" pageId="0" pageNumber="189" rowspan="1">5-12</td>
<td id="637D39617FFC668AA04AAA53E33D92B8" colspan="1" pageId="0" pageNumber="189" rowspan="1">5-10</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Fruit horn or wing</emphasis>
<tr id="DA1703A5537DF8A0B82859C1D0726D47" pageId="0" pageNumber="189">
<td id="B3843C02463ADCB65D552798C60F4F14" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="06536ABE84E5A6EECAA7DB4EF711C01A" bold="true" pageId="0" pageNumber="189">Fruit horn or wing</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">none to rarely short crest</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">none to short crest or horns</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">short to long horns or wings</td>
<td id="600F038B0C5EF4150D8C9448BAA04371" colspan="1" pageId="0" pageNumber="189" rowspan="1">none to rarely short crest</td>
<td id="F047E69483BCC7A1179BE6E05F866DFA" colspan="1" pageId="0" pageNumber="189" rowspan="1">none to short crest or horns</td>
<td id="3DACD285A834E7C391E80D0D297BCC65" colspan="1" pageId="0" pageNumber="189" rowspan="1">short to long horns or wings</td>
</tr>
<tr pageId="0" pageNumber="189">
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="189">Flowering time</emphasis>
<tr id="8270EE78E522FE40F7EE9DC23C28D87B" pageId="0" pageNumber="189">
<td id="8855CCEEFFA9A90AAF08DC3618F4A5AC" colspan="1" pageId="0" pageNumber="189" rowspan="1">
<emphasis id="C0108B3372A851C0A6D5D6A61DF91D40" bold="true" pageId="0" pageNumber="189">Flowering time</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">June-October</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">June-December</td>
<td colspan="1" pageId="0" pageNumber="189" rowspan="1">March-April</td>
<td id="5AE6FAC434F297888E033A4425BB9AB0" colspan="1" pageId="0" pageNumber="189" rowspan="1">June-October</td>
<td id="60BB14AF291C5306CE55F865007D7908" colspan="1" pageId="0" pageNumber="189" rowspan="1">June-December</td>
<td id="17F350FA633DC04851EA97210141AA55" colspan="1" pageId="0" pageNumber="189" rowspan="1">March-April</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="description">
<paragraph pageId="0" pageNumber="189">Description.</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Herb</emphasis>
<subSubSection id="97CADE7FC66F26EF8B084DEE41E9C27C" pageId="0" pageNumber="189" type="description">
<paragraph id="091F21D739046CF45705DCFC92EFF708" pageId="0" pageNumber="189">Description.</paragraph>
<paragraph id="ECC95445D3A75CD51C08644C44AA75C1" pageId="0" pageNumber="189">
<emphasis id="B9E695ACFCE80BF9F0B95DC619321C19" bold="true" pageId="0" pageNumber="189">Herb</emphasis>
perennial, evergreen, to 4 m tall, dioecious.
<emphasis bold="true" pageId="0" pageNumber="189">Rhizome</emphasis>
<emphasis id="49DA470657E8BAA2B41C43D9B73509D0" bold="true" pageId="0" pageNumber="189">Rhizome</emphasis>
short, stout, nearly unbranched, reddish brown, to 12 cm thick.
<emphasis bold="true" pageId="0" pageNumber="189">Stem</emphasis>
<emphasis id="D2D30CF7CA26BABBBB2E5C53E761C14D" bold="true" pageId="0" pageNumber="189">Stem</emphasis>
erect, reddish brown or green, glabrous, internodes to 5 cm thick, with many longitudinally fusiform whitish spots, cross section of stem often reddish brown, nodes notably enlarged, to 7 cm thick, with unequally oval to round whitish spots, many shrubby branches on the upper part of main stem.
<emphasis bold="true" pageId="0" pageNumber="189">Stipule</emphasis>
<emphasis id="A50C6C5AA198675B210C9591BF39888F" bold="true" pageId="0" pageNumber="189">Stipule</emphasis>
long-triangular, light green or pinkish green, 9-25
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9E8D491C320434A73C13C921B1046E68" originalValue="×">x</normalizedToken>
2-8 mm, glabrous, margin entire, dorsal ridge pinkish, apex acuminate with arista 4-6 mm long.
<emphasis bold="true" pageId="0" pageNumber="189">Petioles</emphasis>
<emphasis id="7A348CED11A2FAC6D5A162229D6D85BF" bold="true" pageId="0" pageNumber="189">Petioles</emphasis>
green, pink to red, glabrous, 7-22 cm long, 1-3 mm thick.
<emphasis bold="true" pageId="0" pageNumber="189">Leaf blade</emphasis>
<emphasis id="E9A05085D28A1D3E3E36058B8517A2B2" bold="true" pageId="0" pageNumber="189">Leaf blade</emphasis>
ovate-lanceolate to lanceolate, 4-19
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="EEF6DF2FAA4C1D6FA79D2A179AEF2CE1" originalValue="×">x</normalizedToken>
0.8-8 cm, adaxial green, muriculate to nearly glabrous, adaxial veins slightly concave; abaxial greyish green, veins usually red, convex, main veins sparsely and obliquely strigose; base obliquely cordate, margin shallowly and remotely denticulate, apex long caudate;
<emphasis bold="true" pageId="0" pageNumber="189">Inflorescence</emphasis>
<emphasis id="7F39975CAC7C5B079A69D2C5F66A9AFE" bold="true" pageId="0" pageNumber="189">Inflorescence</emphasis>
dichasial cyme, axillary, short, 3-5 cm long, unbranched to branched once, rachis glabrous, green, pinkish green to red, base usually red-brown, 7-15 mm long, 1-1.5 mm thick, 3-11 male flowers or 1-5 female per inflorescence.
<emphasis bold="true" pageId="0" pageNumber="189">Bract</emphasis>
<emphasis id="06473C701C464C53674B7395DCB9B85A" bold="true" pageId="0" pageNumber="189">Bract</emphasis>
often caducous, pinkish green, long triangular, glabrous, ca. 6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="287689A0674EFD2629221E5A9012A07E" originalValue="×">x</normalizedToken>
3 mm, apex acuminate; bracteoles smaller.
<emphasis bold="true" pageId="0" pageNumber="189">Staminate flower</emphasis>
<emphasis id="D48F0EFC0D481E07A9BB6DF08AEDA974" bold="true" pageId="0" pageNumber="189">Staminate flower</emphasis>
: pedicel glabrous, white, whitish or pinkish green, 10-14 mm long, ca.1 mm thick; corolla 18-24 mm in diameter; tepals 4, subequal, glabrous, outer 2, obovate, 9-14
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="22692C8197566DEE94461FCD1F1CCDD9" originalValue="×">x</normalizedToken>
6-9 mm, apex obtuse, adaxially white and middle-upper part abaxially pink, or pure white for some individuals, longitudinal veins unapparent; inner 2, pure white, obovate to obovate-lanceolate, 8-13
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="D67C7B9D8FCC44AACA443E44BDCD9B4E" originalValue="×">x</normalizedToken>
5-7 mm, apex obtuse; androecium nearly actinomorphic, ca. 5 mm long, 6-7 mm in diam; stamens 48-60, filaments free, 1-2 mm long; anther yellow, 2-3 mm long, apex obtuse or nearly so.
<emphasis bold="true" pageId="0" pageNumber="189">Pistillate flower</emphasis>
<emphasis id="4559D089D965568E6046C2F308BA8F7E" bold="true" pageId="0" pageNumber="189">Pistillate flower</emphasis>
: pedicel white or green-white, 6-12 mm long, 0.8-1 mm thick; corolla 20-25 mm, tepals 6, rarely 4, glabrous, outer 3 (rarely 2), obovate or long obovate, thick and rigid, 12-18
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="0BEEA1DF9C663F6DFD080EE9B443369F" originalValue="×">x</normalizedToken>
7-10 mm, adaxial surface nearly white, distinctly concave, abaxially pink on middle-upper part, inner 3 (rarely 2), obovate-lanceolate to oblanceolate or long elliptical, slightly narrower than outer tepals, 10-19
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="4E0C077BC1863442A91D33759A51F998" originalValue="×">x</normalizedToken>
6-8 mm, white, glabrous, apex obtuse; styles + stigmas 5 mm long, 7-8 mm wide; styles 3, free; stigmas yellow, nearly U-shaped, each side spirally twisted 1.5 circles; ovary pink or green, with white convex spots; placentation axile, 3-loculed, each placenta 2-branched.
<emphasis bold="true" pageId="0" pageNumber="189">Peduncle</emphasis>
<emphasis id="1F6A091F5CCC262A5C12B12D243FD70C" bold="true" pageId="0" pageNumber="189">Peduncle</emphasis>
green to pinkish green, glabrous, 8-12 mm long, ca. 1 mm thick.
<emphasis bold="true" pageId="0" pageNumber="189">Fruit</emphasis>
<emphasis id="53F35589F0FC4E72679B5A43D4DCB114" bold="true" pageId="0" pageNumber="189">Fruit</emphasis>
red, pink or green, glabrous, triangular-gyroscopic, 8-11
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="10C61088A7CC57B02DE30B01C2722A3B" originalValue="×">x</normalizedToken>
1-12 mm wide, concave between two placentas, wingless to occasionally short ridged, apex with beak 3-4 mm long. Flowering Jun.-Oct., fruiting Jul.-Dec.
</paragraph>
<caption doi="10.3897/phytokeys.187.75854.figure4" httpUri="https://binary.pensoft.net/fig/629009" pageId="0" pageNumber="189" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figure 4.</emphasis>
<caption id="9D130415311FE132678EA48884DBA602" doi="10.3897/phytokeys.187.75854.figure4" httpUri="https://binary.pensoft.net/fig/629009" pageId="0" pageNumber="189" start="Figure 4" startId="F4">
<paragraph id="6004ADC78F58B0F5F1594A2D7D5E0920" pageId="0" pageNumber="189">
<emphasis id="F890871EBFB5AA5311C20AC98ED7BAA9" bold="true" pageId="0" pageNumber="189">Figure 4.</emphasis>
Illustration of
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<taxonomicName id="97A2E1545CA3A0DF75510F99C274FFB5" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="DDA7C44F8D78555F7BCB2450548D8CCF" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
D.K.Tian &amp; W.G.Wang, sp. nov. (Drawn by Mr. Zhi-Min Li)
<emphasis bold="true" pageId="0" pageNumber="189">A</emphasis>
<emphasis id="4187943081660AA19DAE7E7E3D0E8DD1" bold="true" pageId="0" pageNumber="189">A</emphasis>
male flowering branches
<emphasis bold="true" pageId="0" pageNumber="189">B</emphasis>
<emphasis id="40270D1EB61196581944E0392249EB86" bold="true" pageId="0" pageNumber="189">B</emphasis>
female flowering branches
<emphasis bold="true" pageId="0" pageNumber="189">C</emphasis>
<emphasis id="6E28FFD8AC90DED20AA692529DE42337" bold="true" pageId="0" pageNumber="189">C</emphasis>
main stem line spots, much expanded node and internode base
<emphasis bold="true" pageId="0" pageNumber="189">D</emphasis>
<emphasis id="F69FD96BD57BB8DA7E940F8EDAEC0499" bold="true" pageId="0" pageNumber="189">D</emphasis>
expanded node and internode base on small upper branches
<emphasis bold="true" pageId="0" pageNumber="189">E</emphasis>
<emphasis id="2BAD8B8424D45793732003DDF80E49A0" bold="true" pageId="0" pageNumber="189">E</emphasis>
leaf blade
<emphasis bold="true" pageId="0" pageNumber="189">F</emphasis>
<emphasis id="EAA6DE7B94547F6B9EE889DADCB0E993" bold="true" pageId="0" pageNumber="189">F</emphasis>
leaf (abaxial), showing sparse hairs on veins
<emphasis bold="true" pageId="0" pageNumber="189">G</emphasis>
<emphasis id="64D0F008C145AA3F3E740BFAEE7289FB" bold="true" pageId="0" pageNumber="189">G</emphasis>
stipule
<emphasis bold="true" pageId="0" pageNumber="189">H</emphasis>
<emphasis id="FB9C943B87EBF01D5B37F7326EAA6604" bold="true" pageId="0" pageNumber="189">H</emphasis>
staminate flowers
<emphasis bold="true" pageId="0" pageNumber="189">I, J</emphasis>
<emphasis id="128EA0AD03B3C4552714A310E0C6C51B" bold="true" pageId="0" pageNumber="189">I, J</emphasis>
pistillate flower
<emphasis bold="true" pageId="0" pageNumber="189">K</emphasis>
<emphasis id="143583FB48A51ACDD65891935B99A1E1" bold="true" pageId="0" pageNumber="189">K</emphasis>
side view of androecium
<emphasis bold="true" pageId="0" pageNumber="189">L</emphasis>
<emphasis id="6AE1EDF27FCB55C7D9876F3810FF72C5" bold="true" pageId="0" pageNumber="189">L</emphasis>
stamens
<emphasis bold="true" pageId="0" pageNumber="189">M</emphasis>
<emphasis id="D906B710CD90708E777A0A02A8641DD1" bold="true" pageId="0" pageNumber="189">M</emphasis>
ovary and stigmas
<emphasis bold="true" pageId="0" pageNumber="189">N</emphasis>
<emphasis id="706BA83A8E4D37BD8F9DCC8B42130185" bold="true" pageId="0" pageNumber="189">N</emphasis>
fruit
<emphasis bold="true" pageId="0" pageNumber="189">O</emphasis>
<emphasis id="62943DCB027D040FBB5428461A63A4C6" bold="true" pageId="0" pageNumber="189">O</emphasis>
dissection of ovary showing placentae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="additional specimen examined">
<paragraph pageId="0" pageNumber="189">Additional specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">China. Xizang</emphasis>
<subSubSection id="08180359795658E71CD5C091CD121043" pageId="0" pageNumber="189" type="additional specimen examined">
<paragraph id="C7C5AFDC65B9FC0FE90016482A820BD1" pageId="0" pageNumber="189">Additional specimen examined.</paragraph>
<paragraph id="F9B5BC098EBF984C20F8E9D7698F7A9E" pageId="0" pageNumber="189">
<emphasis id="7DD8C718BA50C0CFFF9488204237D941" bold="true" pageId="0" pageNumber="189">China. Xizang</emphasis>
:
<normalizedToken originalValue="Mêdog">Medog</normalizedToken>
<normalizedToken id="99967FE4EC06EEAA92D4D467073A3353" originalValue="Mêdog">Medog</normalizedToken>
County (墨脱县), Beibeng Town (背崩乡), Baimu Xiri River (白母西日河), forest slope of river valley or
<normalizedToken originalValue="waters">water's</normalizedToken>
<normalizedToken id="1E1404867C09C3E057F7C8807161FA75" originalValue="waters">water's</normalizedToken>
edge along stream,
<geoCoordinate degrees="29" direction="north" minutes="21" orientation="latitude" precision="15" seconds="9" value="29.3525">29°21'9&quot;N</geoCoordinate>
<geoCoordinate id="2407C63197D09F4859CA15E12C84A24A" degrees="29" direction="north" minutes="21" orientation="latitude" precision="15" seconds="9" value="29.3525">29°21'9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="11" orientation="longitude" precision="15" seconds="21" value="95.18917">95°11'21&quot;E</geoCoordinate>
<geoCoordinate id="D97BDF179D9683188B1C59AFACD2DE54" degrees="95" direction="east" minutes="11" orientation="longitude" precision="15" seconds="21" value="95.18917">95°11'21&quot;E</geoCoordinate>
, elev. 1320 m, 10 September 2020,
<emphasis italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4773-B</emphasis>
<emphasis id="D2C6759F1BEDAD89716631B95F472C03" italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4773-B</emphasis>
(paratype CSH!, ♂);
<geoCoordinate degrees="29" direction="north" minutes="20" orientation="latitude" precision="15" seconds="0" value="29.333334">29°20'0&quot;N</geoCoordinate>
<geoCoordinate id="5BB52466C07759DBB9D2BCDCF92064C5" degrees="29" direction="north" minutes="20" orientation="latitude" precision="15" seconds="0" value="29.333334">29°20'0&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="49" value="95.180275">95°10'49&quot;E</geoCoordinate>
<geoCoordinate id="5BF62DBE9AF9BA0BA3EC3B35E3548C6B" degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="49" value="95.180275">95°10'49&quot;E</geoCoordinate>
, elev. 1110 m, 10 September 2020,
<emphasis italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4765-A, TDK4765-B</emphasis>
<emphasis id="0717186F01454EEC820D3B217B667C47" italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4765-A, TDK4765-B</emphasis>
, (paratype CSH!);
<geoCoordinate degrees="29" direction="north" minutes="18" orientation="latitude" precision="15" seconds="32" value="29.308887">29°18'32&quot;N</geoCoordinate>
<geoCoordinate id="DE96F716C68DF0A9BEBFC4C57AB7EB0B" degrees="29" direction="north" minutes="18" orientation="latitude" precision="15" seconds="32" value="29.308887">29°18'32&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="38" value="95.17722">95°10'38&quot;E</geoCoordinate>
<geoCoordinate id="6D59DA06DDD78AA0820F051504B4D08E" degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="38" value="95.17722">95°10'38&quot;E</geoCoordinate>
, elev. 980 m, 10 September 2020,
<emphasis italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4777</emphasis>
<emphasis id="CC30A74DFA22779C5C0DA47C54A70E67" italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Fang Wen, Qing-Gong Mao, &amp; Zhu Lu TDK4777</emphasis>
(paratype CSH!); near Ani Bridge (阿尼桥),
<geoCoordinate degrees="29" direction="north" minutes="17" orientation="latitude" precision="1" seconds="8.41" value="29.28567">29°17'8.41&quot;N</geoCoordinate>
<geoCoordinate id="FF28629CB63B1F87CAE30EBF806977FC" degrees="29" direction="north" minutes="17" orientation="latitude" precision="1" seconds="8.41" value="29.28567">29°17'8.41&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="10" orientation="longitude" precision="1" seconds="3.23" value="95.167564">95°10'3.23&quot;E</geoCoordinate>
<geoCoordinate id="119D34DBFC1E7CCAEF09108FDE27CA73" degrees="95" direction="east" minutes="10" orientation="longitude" precision="1" seconds="3.23" value="95.167564">95°10'3.23&quot;E</geoCoordinate>
, elev. 810 m, 3 July 2020,
<emphasis italics="true" pageId="0" pageNumber="189">Wen-Guang Wang, You-Yun Li, Xing-Da Ma, &amp; Jian-Yong Shen, WWG 2014</emphasis>
<emphasis id="2C0F032135F3B3B6333ECA9973C03116" italics="true" pageId="0" pageNumber="189">Wen-Guang Wang, You-Yun Li, Xing-Da Ma, &amp; Jian-Yong Shen, WWG 2014</emphasis>
(paratype, HITBC!),
<emphasis italics="true" pageId="0" pageNumber="189">WWG 2015</emphasis>
<emphasis id="48A69498279A46123266C55F90CF4085" italics="true" pageId="0" pageNumber="189">WWG 2015</emphasis>
(paratype HITBC!); elev. 1100 m, 16 September 1974,
<emphasis italics="true" pageId="0" pageNumber="189">anonymous 2608</emphasis>
<emphasis id="1F84C3AC1FFDE45408E27D2C8D249FD0" italics="true" pageId="0" pageNumber="189">anonymous 2608</emphasis>
(paratype PE!); elev. 800-1400 m, 30 June 1980,
<emphasis italics="true" pageId="0" pageNumber="189">Wei-Lie Chen 10809</emphasis>
<emphasis id="324C050EAAB614AEF49E74A893480804" italics="true" pageId="0" pageNumber="189">Wei-Lie Chen 10809</emphasis>
(PE!); near No. 2 Bridge,
<geoCoordinate degrees="29" direction="north" minutes="16" orientation="latitude" precision="15" seconds="42" value="29.278334">29°16'42&quot;N</geoCoordinate>
<geoCoordinate id="D3F2FA4092947C5271A0DFB35D39404B" degrees="29" direction="north" minutes="16" orientation="latitude" precision="15" seconds="42" value="29.278334">29°16'42&quot;N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="49" value="95.180275">95°10'49&quot;E</geoCoordinate>
<geoCoordinate id="CE6A18F3B4884636483050C6B38AE512" degrees="95" direction="east" minutes="10" orientation="longitude" precision="15" seconds="49" value="95.180275">95°10'49&quot;E</geoCoordinate>
, elev. 810 m, 1 October 2017,
<emphasis italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Yan Xiao, Xin Zhong, Li-Zhi Tian &amp; Zhu Lu TDK3429</emphasis>
<emphasis id="F01A19FE5ABC57587B888B80F69620F0" italics="true" pageId="0" pageNumber="189">Dai-Ke Tian, Yan Xiao, Xin Zhong, Li-Zhi Tian &amp; Zhu Lu TDK3429</emphasis>
(paratype CSH!); Beibeng to Hanmi (汗密), elev. 840 m, 7 August 2010, South Tibet Expedition Team (藏南队),
<emphasis italics="true" pageId="0" pageNumber="189">Xiao-Hua Jin, Shu-Dong Zhang, Zhong-Yang Li, Bao-Cheng Wu, Xian-Yun Mu, Jing Li &amp; Wei-Tao Jin, STET2304</emphasis>
<emphasis id="00CDFA3ABF0695DAA42C4DD445A6C3B7" italics="true" pageId="0" pageNumber="189">Xiao-Hua Jin, Shu-Dong Zhang, Zhong-Yang Li, Bao-Cheng Wu, Xian-Yun Mu, Jing Li &amp; Wei-Tao Jin, STET2304</emphasis>
(paratype PE!); Hanmi to Maniweng (马尼翁), elev. 800-1000 m, 6 August 1974,
<emphasis italics="true" pageId="0" pageNumber="189">Qingzang Team 74-4114</emphasis>
<emphasis id="CA3A485793EA5D2D21493C4246CDCF39" italics="true" pageId="0" pageNumber="189">Qingzang Team 74-4114</emphasis>
(paratype PE!); elev. 1200 m, 24 June 1983,
<emphasis italics="true" pageId="0" pageNumber="189">Bo-Sheng Li &amp; Shu-Zhi Chen 05229</emphasis>
<emphasis id="806CB8F7E5569DF490239CCF8DCE3571" italics="true" pageId="0" pageNumber="189">Bo-Sheng Li &amp; Shu-Zhi Chen 05229</emphasis>
(paratype PE!); Maniweng to Ani Bridge, elev. 700-1000 m, 3 August 1972,
<emphasis italics="true" pageId="0" pageNumber="189">Tibet Expedition Team, Institute of Biology 1631</emphasis>
<emphasis id="16DBD90ECE0D7134F3DFC5D111084C29" italics="true" pageId="0" pageNumber="189">Tibet Expedition Team, Institute of Biology 1631</emphasis>
(paratype HNWP!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="distribution">
<paragraph pageId="0" pageNumber="189">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="78C7150CBBE408258B5091FAB0553F0B" pageId="0" pageNumber="189" type="distribution">
<paragraph id="1B4AD87716A0955648C9E39CC8208D68" pageId="0" pageNumber="189">Distribution and habitat.</paragraph>
<paragraph id="9194B6FE80E86A047117D856C5CA4BCF" pageId="0" pageNumber="189">
Currently known from at least two localities in
<normalizedToken originalValue="Mêdog">Medog</normalizedToken>
<normalizedToken id="8CEA59AA7E1C04B026B178BE15B3C8CF" originalValue="Mêdog">Medog</normalizedToken>
, southern Xizang (Tibet), China (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of B. giganticaulis (triangles) from southern Xizang, China." figureDoi="10.3897/phytokeys.187.75854.figure5" httpUri="https://binary.pensoft.net/fig/629010" pageId="0" pageNumber="189">5</figureCitation>
<figureCitation id="AF08D285A00C2EDEBDD374B87BAF11BE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of B. giganticaulis (triangles) from southern Xizang, China." figureDoi="10.3897/phytokeys.187.75854.figure5" httpUri="https://binary.pensoft.net/fig/629010" pageId="0" pageNumber="189">5</figureCitation>
). It grows on the slopes under forest along streams, elevation 450-1400 m.
</paragraph>
<caption doi="10.3897/phytokeys.187.75854.figure5" httpUri="https://binary.pensoft.net/fig/629010" pageId="0" pageNumber="189" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figure 5.</emphasis>
<caption id="05CE5CEDD2F00D7B5C2A6B289E6CACA3" doi="10.3897/phytokeys.187.75854.figure5" httpUri="https://binary.pensoft.net/fig/629010" pageId="0" pageNumber="189" start="Figure 5" startId="F5">
<paragraph id="CE0CFBB2749913ACDEA03D9AA4747405" pageId="0" pageNumber="189">
<emphasis id="7451D9EA7864BF5AAED71FE9E9339942" bold="true" pageId="0" pageNumber="189">Figure 5.</emphasis>
Distribution of
<taxonomicName lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
<taxonomicName id="CB3C4F172BB9B11AC026F52A433B6EF7" lsidName="B. giganticaulis" pageId="0" pageNumber="189" rank="species" species="giganticaulis">
<emphasis id="4585B02D8E8775C0D294544058DB8989" italics="true" pageId="0" pageNumber="189">B. giganticaulis</emphasis>
</taxonomicName>
(triangles) from southern Xizang, China.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="conservation status">
<paragraph pageId="0" pageNumber="189">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="189">
<taxonomicName authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
<subSubSection id="51E4077DE18F1DE026A966D7B23C1146" pageId="0" pageNumber="189" type="conservation status">
<paragraph id="62249E6698A57DFFBF676E9F11B1C422" pageId="0" pageNumber="189">Conservation status.</paragraph>
<paragraph id="366EB02CA302B357D2F3F34747EE296A" pageId="0" pageNumber="189">
<taxonomicName id="C34E953B5B79BB58DB7709112E7CFB23" authorityName="D. K. Tian &amp; W. G. Wang" authorityYear="2021" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia giganticaulis" order="Cucurbitales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="giganticaulis">
<emphasis id="BE3823FEEC6E4354DAF732A2953BB099" italics="true" pageId="0" pageNumber="189">Begonia giganticaulis</emphasis>
</taxonomicName>
is currently found in at least two localities in
<normalizedToken originalValue="Mêdog">Medog</normalizedToken>
<normalizedToken id="A1124C637DFCBE0BD7950CDD5B90D443" originalValue="Mêdog">Medog</normalizedToken>
of Tibet. Additional populations might be discovered when more surveys are conducted in China-India border region. However, based on current data, it should be categorised as Endangered: B2a (
<bibRefCitation author="IUCN" journalOrPublisher="Mitteilungen aus dem Institut fuer Allgemeine Botanik in Hamburg" pageId="0" pageNumber="189" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B30" refString="IUCN, 2019. Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
<bibRefCitation id="C5005972E85ED1F0DC6A0B6D742C5E71" author="IUCN" journalOrPublisher="Mitteilungen aus dem Institut fuer Allgemeine Botanik in Hamburg" pageId="0" pageNumber="189" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B30" refString="IUCN, 2019. Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
) due to &lt;500 km2 area of occupancy with severely fragmented habitat consisting of &lt;5 known populations totally under 1000 individuals by estimation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="etymology">
<paragraph pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="189">The specific epithet refers to the huge (very tall and thick stem) plant size of the new species, which is the tallest begonia in Asia.</paragraph>
<subSubSection id="4A6DF221ADD308C6FDCD0E699BDBED52" pageId="0" pageNumber="189" type="etymology">
<paragraph id="464D80E91DCAB8113A733B7B12E45FB3" pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph id="2B235E30ADAB06E7C46CA785D0744DB3" pageId="0" pageNumber="189">The specific epithet refers to the huge (very tall and thick stem) plant size of the new species, which is the tallest begonia in Asia.</paragraph>
</subSubSection>
</treatment>
</document>