Add updates up until 2025-01-16 15:23:39

This commit is contained in:
ggserver 2025-01-16 15:29:43 +00:00
parent 1fe81f8f8a
commit d639d84c0e
5 changed files with 955 additions and 325 deletions

View file

@ -0,0 +1,351 @@
<document id="7770773C4881C16335C454E9421FCEDA" ID-DOI="10.3897/zookeys.1223.135292" ID-ZooBank="49B9232E-01DD-426E-ABB8-9C02DA071776" ID-publisher-id="135292" URI-arpha="ED31D2FD-08B7-5E1C-A694-DD8451177BB7" URI-zoobank="http://zoobank.org/49B9232E-01DD-426E-ABB8-9C02DA071776" article-type="research-article" checkinTime="1737040972054" checkinUser="pensoft" docAuthor="Shepherd, Bart, Pinheiro, Hudson T., Najeeb, Ahmed, Rocha, Claudia R. &amp; Rocha, Luiz A." docDate="2025" docId="0EADD278842A56CE835756F979292AAE" docLanguage="en" docName="ZooKeys 1223: 333-344" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/135292/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Plectranthias raki Shepherd, Pinheiro, Najeeb, Rocha &amp; Rocha, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="344" masterDocId="ED31D2FD08B75E1CA694DD8451177BB7" masterDocTitle="Plectranthias raki (Teleostei, Serranidae), a new species of perchlet from mesophotic coral ecosystems of the Maldives" masterLastPageNumber="344" masterPageNumber="333" pageNumber="333" updateTime="1737041322759" updateUser="ExternalLinkService">
<mods:mods id="7771782C18AB5AC39C766AC79894DAFB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1777343C359132B59F59B84F7E959187">
<mods:title id="533D2D0C5C494AC6D76DA8AAEFCD69D1">Plectranthias raki (Teleostei, Serranidae), a new species of perchlet from mesophotic coral ecosystems of the Maldives</mods:title>
</mods:titleInfo>
<mods:name id="AD73E1F94F86C55EA21021819B7D9DCA" type="personal">
<mods:role id="A81DCE913831D8E1296C07FAA3B43529">
<mods:roleTerm id="9B3B9BE123E5102B5AACEFD6778BA42B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A377F19E1D9D9319FFBF9D17B003A3DF">Shepherd, Bart</mods:namePart>
<mods:nameIdentifier id="47EF9B7020124D781CF9C7316536E5FC" type="ORCID">0000-0002-8918-1551</mods:nameIdentifier>
<mods:affiliation id="6AB035E9CF947BF4AB21BFDBC3B7CB83">Steinhart Aquarium, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:name id="5370BDF41F51BFE986C0FAFE0436329F" type="personal">
<mods:role id="28932CC2C45E5C1EF5C7F85749E2C57B">
<mods:roleTerm id="F6A4C1879EF2A5DA74D4BAC85619F116">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AB2B98810A206DF95B3CC7A724DD039">Pinheiro, Hudson T.</mods:namePart>
<mods:nameIdentifier id="4045FE468C9C262298E5CA386B45E421" type="ORCID">0000-0002-3143-1474</mods:nameIdentifier>
<mods:affiliation id="47DCC0D98D20EE6B144693B6FB0F6983">Steinhart Aquarium, California Academy of Sciences, San Francisco, CA 94118, USA &amp; Department of Ichthyology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:name id="572A10B47FBE3325691A899B895775F0" type="personal">
<mods:role id="48803B6F2816A501041CB596BBF8CED1">
<mods:roleTerm id="2B182A546F22A145626553C8EBE9986D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA2C01B8934527E19E25F84634EF90C1">Najeeb, Ahmed</mods:namePart>
<mods:affiliation id="3D1BFD7A33A7ED9FD6A7768A80B78717">Steinhart Aquarium, California Academy of Sciences, San Francisco, CA 94118, USA &amp; Center for Marine Biology, University of São Paulo, São Sebastião, SP 11600, Brazil</mods:affiliation>
</mods:name>
<mods:name id="76FCE51BBF9B11141F08C40FA029269C" type="personal">
<mods:role id="685E794D6918721BBF934A724032F772">
<mods:roleTerm id="2A1CEBF6D77F0D087DC580CB4949AEEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D487CAF97E3403D74F4F2F27036D8509">Rocha, Claudia R.</mods:namePart>
<mods:nameIdentifier id="1A43C54CA8C9310354A272492A634A35" type="ORCID">0000-0001-8069-6535</mods:nameIdentifier>
<mods:affiliation id="7A9B591F04C938DC37E37123F873660B">Steinhart Aquarium, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:name id="E337C92DAF510F671551B8A60FE6BCA4" type="personal">
<mods:role id="E563AE709306D236E2685DE41603B571">
<mods:roleTerm id="375A1ACD312DA6C8E167599AC46043F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16538880C6DD074C0EF70B9A2FA7DFC1">Rocha, Luiz A.</mods:namePart>
<mods:nameIdentifier id="422DC14E1D9088DEA00E77BC0535CE5E" type="ORCID">0000-0003-4011-569X</mods:nameIdentifier>
<mods:affiliation id="B7FC7FCD3C9BAF9F9662993B8B3AFF5E">Steinhart Aquarium, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="04E6CC449F49D64129FB1CB3412EFF7B">text</mods:typeOfResource>
<mods:relatedItem id="06FAFDF8899887D346B1EC1718CC3579" type="host">
<mods:titleInfo id="3601D8E593E9EC01A4B13B6261EFB071">
<mods:title id="1A6B77348C0F0F0913592FD8ADEA4C7F">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="6CD98CF65892E00D65F9CF072154D80E">
<mods:date id="74E81BEAA11DB99812499DFCE54D3138">2025</mods:date>
<mods:detail id="A240DC649D5F2971F8AF3C48ABB30802" type="pubDate">
<mods:number id="0704778C91DD7613FEA551C183E70F5C">2025-01-16</mods:number>
</mods:detail>
<mods:detail id="4F252E95307911671BC4593B8FF7EC2D" type="volume">
<mods:number id="0C4AB1A148AFC19E3A5A2D81845D1ED8">1223</mods:number>
</mods:detail>
<mods:extent id="B230B86DF1CDCA38D48EC15F3EAE7904" unit="page">
<mods:start id="6BC866049FA0C8A4876A168729DD6E91">333</mods:start>
<mods:end id="0571A006606648FDED5A5A8B98F3B601">344</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="85144955D8F650D6F1FA0C56BB5E39C5">journal article</mods:classification>
<mods:identifier id="AD04BF1E726C3F21793A2FC9503B5131" type="DOI">10.3897/zookeys.1223.135292</mods:identifier>
<mods:identifier id="92DB0EA192DC6984A42FE066C83B9505" type="ZooBank">49B9232E-01DD-426E-ABB8-9C02DA071776</mods:identifier>
</mods:mods>
<treatment id="0EADD278842A56CE835756F979292AAE" ID-DOI="http://doi.org/10.5281/zenodo.14674064" ID-Zenodo-Dep="14674064" ID-arpha="0EADD278-842A-56CE-8357-56F979292AAE" ID-zoobank="https://zoobank.org/9B0CEAD6-829B-4BE7-B82C-2AB8E3EBDE77" LSID="urn:lsid:plazi:treatment:0EADD278842A56CE835756F979292AAE" LSID-ZBK="urn:lsid:zoobank.org:act:9B0CEAD6-829B-4BE7-B82C-2AB8E3EBDE77" httpUri="http://treatment.plazi.org/id/0EADD278842A56CE835756F979292AAE">
<subSubSection id="25DB1F1F17B9F4A621302638EC96B010" type="nomenclature">
<paragraph id="41DDDFDCAB034BAE1B50114C618110B5">
<taxonomicName id="4E81A595C68DE21C60599DCD9A20A29A" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki" status="sp. nov.">
<emphasis id="D1FD423F06A5AF5A4023A54F86928A97" italics="true">Plectranthias raki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BD742A5C64AED1E658E3FB04D66B9AB1" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6811FEE903DE067C294917C3A5BDF10B" type="description">
<paragraph id="72C54AB27140D6B11FE6B8987601BC71">
<figureCitation id="953CB736366CA4FA0621C19FD7E9019B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (left) and paratype (right) of Plectranthias raki sp. nov. Holotype CAS - ICH 248439 66.2 mm SL, shortly after collection (A), preserved specimen (B) and x-radiograph (C). Paratype LACM 61827, 70.4 mm SL, shortly after collection (D), preserved specimen (E) and x-radiograph (F). Photos: A, D by Luiz Rocha. B, C, E, F by Jon Fong." figureDoi="10.3897/zookeys.1223.135292.figure1" httpUri="https://binary.pensoft.net/fig/1221256">Figs 1</figureCitation>
,
<figureCitation id="A9008362C7914E40DC34D51BE663036F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living specimen (not retained) of Plectranthias raki sp. nov. photographed at 110 m depth at Dhaalu Atoll, Maldives. Photo by Luiz Rocha." figureDoi="10.3897/zookeys.1223.135292.figure2" httpUri="https://binary.pensoft.net/fig/1221257">2</figureCitation>
,
<tableCitation id="F8824E03407A5EE41636DD1150ACC97B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for Plectranthias raki sp. nov., expressed as a percentage of standard length." httpUri="http://table.plazi.org/id/79DB6D23893AF1CB3450DDCAB2145434" tableUuid="79DB6D23893AF1CB3450DDCAB2145434">Table 1 Dhivehi common name: Raki bureki English common name: Maldivian Perchlet</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGQAE" type="type locality">
<paragraph id="58E6400B140E09C88ED2261DE914618C">
<heading id="ED59BE969768162892F8830A51279D43" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="192296F257390A642F4D98F24EBFCFC3">
<collectingCountry id="E9E971B7CDD6B545E9209CC381908F4B" name="Maldives">Maldives</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELQAE" type="material">
<paragraph id="74347479B646C8BB0FEE6DA4941D2584">
<heading id="304FE3EE06CA2CFEEB4BCFADCA956673" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="27F8DF4B529DC571C2DF3FDEAEA21B0A">
<materialsCitation id="2D94A12E615808BCD725825D578C6437" collectingDate="2022-12-09" collectionCode="CAS, MV" collectorName="GenBank PQ &amp; B Shepherd &amp; HT Pinheiro &amp; Bell &amp; LA Rocha" country="Maldives" county="Rasdhoo Atoll" elevation="118" latitude="4.256111" location="Location" longLatPrecision="21" longitude="72.98333" municipality="Kuramathi Outer Reef" specimenCode="ICH 248439" specimenCount="1" typeStatus="holotype">
<emphasis id="868342853AF8EC1505F73E64A45FA6FE" bold="true">
<emphasis id="9CEE9CDE4B8943178B63297072BF75DF" bold="true" italics="true">
<typeStatus id="606949356C9E57843050D5E373CAEAFB" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<named-content id="517A6C1F8D32A2D29479EF688C0526F3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences ichthyological collection">
<collectionCode id="DCF371E6365DA5FC76B5B0FB285A5CE1" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</named-content>
-
<specimenCode id="78D9FABDBE71941BA54802F3E91F56AF" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">ICH 248439</specimenCode>
(Field number LAR 2951)
<quantity id="831F92ACAB28E7E096C336D164AA047B" metricMagnitude="-2" metricUnit="m" metricValue="6.62" unit="mm" value="66.2">66.2 mm</quantity>
<abbrev id="ABBRID0E1QAE" xlink_title="standard length">SL</abbrev>
,
<collectorName id="61AFCF661AEDAD4EFC095F87AEC8D262">
GenBank
<ext-link id="10971349BEF9F9A12F5B03A217BE5E35" ext-link-type="gen" xlink_href="PQ416576" xlink_type="simple">PQ 416576</ext-link>
</collectorName>
.
<location id="67EE23DB464F17110515B454D0339C49" LSID="urn:lsid:plazi:treatment:0EADD278842A56CE835756F979292AAE:67EE23DB464F17110515B454D0339C49" country="Maldives" county="Rasdhoo Atoll" latitude="4.256111" longLatPrecision="21" longitude="72.98333" municipality="Kuramathi Outer Reef" name="Location">Location</location>
:
<collectingMunicipality id="1A99E38397078F56111CD00DA70F8264">Kuramathi Outer Reef</collectingMunicipality>
,
<collectingCounty id="4518DADABD2AAD557E31DB0B711162F0">Rasdhoo Atoll</collectingCounty>
,
<collectingCountry id="1F3BA58AFDAA2A10B64AC2BFFE9E4F0B" name="Maldives">Maldives</collectingCountry>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[72.983333,4.256111]}">
<geoCoordinate id="6ABF79A69665611490A9F3EDE42886AB" degrees="4" direction="north" minutes="15" orientation="latitude" precision="15" seconds="22" value="4.256111">4 ° 15 ' 22 &quot; N</geoCoordinate>
,
<geoCoordinate id="2B0BD47CB13C2E905B8127452F7252AC" degrees="72" direction="east" minutes="59" orientation="longitude" precision="15" seconds="00" value="72.98333">72 ° 59 ' 00 &quot; E</geoCoordinate>
</named-content>
, depth of collection
<quantity id="8D617BB19E47B884CFD716D20F40E1C5" metricMagnitude="2" metricUnit="m" metricValue="1.18" unit="m" value="118.0">
<elevation id="49C011A18E47EF8E4FD78C1F9C7B426C" metricMagnitude="2" metricUnit="m" metricValue="1.18" unit="m" value="118.0">118 m</elevation>
</quantity>
, collected with hand nets by
<collectorName id="1C859B2663D900DEF3BEF7CE952EE77A">B Shepherd</collectorName>
,
<collectorName id="3B8044FDDC66D92E79E5BFB97F5AB9A2">HT Pinheiro</collectorName>
,
<collectionCode id="83649A8B7F7CBD574DDDA3E9D118670B" country="0" name="University of Montana Museum">MV</collectionCode>
<collectorName id="A1FCDADAC82B6CFA822B68DA89CC579C">Bell</collectorName>
, and
<collectorName id="6E9C07C0FA81A2676520150280C946BE">LA Rocha</collectorName>
,
<date id="86CFFBD923BD071F1D241A660410CBD1" value="2022-12-09">
<collectingDate id="565E8809B879AED6BF68D10C2E0D505D" value="2022-12-09">9 December 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="8865A36500307A156D50716DB06B36BB" collectingDate="2022-12-09" collectionCode="LACM" collectorName="GenBank PQ &amp; Same" country="Maldives" county="Rasdhoo Atoll" elevation="118" latitude="4.256111" location="Location" longLatPrecision="21" longitude="72.98333" municipality="Kuramathi Outer Reef" specimenCode="LACM 61827" specimenCount="1" typeStatus="paratype">
<emphasis id="50FA8234E6221376621D4AEC7D7E2386" bold="true">
<emphasis id="9A6DB3DF1CE857F5D717A2B09E798B76" bold="true" italics="true">
<typeStatus id="B0350B12B3078000F9A91977EB149174" type="paratype">Paratype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="0D3F21146271FBD64AB1490FF7351057" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">
<named-content id="CA52718BBCFEA850A98BBE70B3444F25" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Natural History Museum of Los Angeles County">LACM</named-content>
61827
</specimenCode>
(Field number LAR 2952)
<quantity id="4E5EAC791ACBA3ADEC27782839B47D7D" metricMagnitude="-2" metricUnit="m" metricValue="7.04" unit="mm" value="70.4">70.4 mm</quantity>
<abbrev id="ABBRID0ETRAE" xlink_title="standard length">SL</abbrev>
,
<collectorName id="6724988E6EE34B03C40198F45BF73CE2">
GenBank
<ext-link id="A9707D76AADE9BA383CE9B56405DD3E2" ext-link-type="gen" xlink_href="PQ416577" xlink_type="simple">PQ 416577</ext-link>
</collectorName>
.
<collectorName id="FF3397E4160CF7ADB6D613FD7742787E">Same</collectorName>
collection data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3RAE" type="diagnosis">
<paragraph id="1E67ADBA73599DA4438583A9CDAB681B">
<heading id="33B0A58B9EC3F4E8398D117DF7E5F3E4" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="74A7579C53533645F568911D0DDCBFCC">
<taxonomicName id="BD26EC7E79B5F729113FA0D94B27A99E" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki" status="sp. nov.">
<emphasis id="AB888B7E06A854BFB5489EF6E9703679" italics="true">Plectranthias raki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="872E2727B87BFBF14B4CA9EB11C86816" rank="species">sp. nov.</taxonomicNameLabel>
is unique in both morphology and coloration. The following combination of characters distinguishes it from all known congeners: dorsal fin X, 15; anal-fin rays III, 7; pectoral-fin rays 13 | 13 (13 | 12), all unbranched; principal caudal-fin rays 9 + 8; lateral line complete with 3032 tubed scales; 3 supraneural bones, predorsal formula 0 / 0 + 0 / 2 / 1 + 1 / 1 / 1 /; gill rakers 5 + 12; circumpeduncular scales 1112; oblique rows of scales on cheek 8 (7); longest dorsal spine the 4
<superScript id="ADD62ABD35F72D0CA89D80877C8F2339">th</superScript>
or 5
<superScript id="D3D41947BB3DEEE707CC3B208C31CBFB">th</superScript>
; no fleshy tips on the dorsal-fin spines; no antrorse serrations on preopercle. Live coloration consisting of a series of irregularly shaped patches of red-orange along dorsal two-thirds of body; patches divided into two indistinct rows by the lateral line; patches red-orange dorsally and posteriorly, becoming more yellow-orange anteriorly and ventrally, golden yellow on opercle and maxilla; anal fin pointed, mostly white proximally, yellow distally, with two yellow spots approximately one-third orbit diameter at base of third spine and fifth and sixth soft rays; two irregularly shaped yellow-orange spots, approximately one-third orbit diameter, located on either side of centermost caudal-fin rays; small orange spot, approximately one-half orbit diameter, at base of pelvic fin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERSAE" type="description">
<paragraph id="A686E74CFE397925A7D6B8E18F4EA941">
<heading id="23E6DA034E193859CC7E49412CA9543E" reason="title">Description.</heading>
</paragraph>
<paragraph id="5F14242563D61B3DB1A82B1FBFC71927">Dorsal rays X, 15, all segmented rays branched; anal rays III, 7, all segmented rays branched; pectoral-fin rays 13 | 13 (13 | 12), all unbranched; pectoral fin moderately long, longest ray reaching to vertical above midpoint of anal fin; pelvic-fin I, 5; upper procurrent caudal-fin rays 6; lower procurrent caudal-fin rays 4; principal caudal-fin rays 9 + 8; branched caudal-fin rays 9 + 7 (8 +?); lateral line complete with 32 (30) tubed scales on the left side; scales above lateral line to origin of dorsal fin 3; scales above lateral line to base of middle dorsal spine 2; scales below lateral line to origin of anal fin 10; oblique rows of scales on cheek 8 (7); circumpeduncular scales 11 (12); gill rakers 5 + 12, the upper 4 and lower 3 rudiments; pseudobranchial filaments 18 (15); branchiostegal rays 7. Vertebrae 10 + 16; supraneural (predorsal) bones 3; predorsal formula 0 / 0 + 0 / 2 / 1 + 1 / 1 / 1 /; dorsal pterygiophores in interneural spaces 913 1 / 1 / 1 + 1 / 1 + 1 / 1; no trisegmental pterygiophores associated with dorsal fin; terminal dorsal pterygiophore in interneural space 18; no trisegmental pterygiophores associated with anal fin; terminal anal pterygiophore in interhaemal space 5; ribs present on vertebrae 3 through 10; epineurals present on vertebrae 1 through 12 (possibly 13); parhypural and hypurals autogenous; well-developed hypurapophysis on parhypural; epurals 3; single uroneural (posterior uroneural absent); ventral tip of cleithrum with well-developed posteroventral process; proximal tip of first anal-fin pterygiophore near distal tips of parapophyses on vertebra 10.</paragraph>
<paragraph id="30B1708FED6D1F6ADE281A2C99623AA2">
Body moderately deep, the depth
<quantity id="0C39DD6D02E49764366A62B16B72B16E" metricMagnitude="-2" metricUnit="m" metricValue="6.858" unit="in" value="2.7">2.7 in</quantity>
<abbrev id="ABBRID0EYSAE" xlink_title="standard length">SL</abbrev>
, and compressed, the width
<quantity id="4C7D672E0EA2AD1025A4CBF5AFCE8E43" metricMagnitude="-2" metricUnit="m" metricValue="5.588" unit="in" value="2.2">2.2 in</quantity>
depth; head fairly short, 2.2 (2.1) in
<abbrev id="ABBRID0E3SAE" xlink_title="standard length">SL</abbrev>
; dorsal fin originates at a vertical line just above the third lateral line scale; dorsal fin continuous and notched between the spinous and soft portions to about half of the length of the first soft ray; dorsal-fin spines without fleshy tabs on the tips; dorsal-fin base length 3.2 (2.1) in
<abbrev id="ABBRID0EATAE" xlink_title="standard length">SL</abbrev>
; the fourth and fifth dorsal spines the longest and the same length in the
<typeStatus id="87C59679382287E9762C1B14E99DBD75" type="holotype">holotype</typeStatus>
(the fourth the longest on the
<typeStatus id="439A2D1A4E0207E72EBEDB1AFE588BD2" type="paratype">paratype</typeStatus>
); third dorsal-fin soft ray the longest, with extended filament; anal-fin base length 6.0 (5.7) in
<abbrev id="ABBRID0EETAE" xlink_title="standard length">SL</abbrev>
; second anal-fin spine the longest; anal fin pointed with the third segmented ray the longest; pectoral fin moderately long and pointed, 2.9 (2.7) in
<abbrev id="ABBRID0EITAE" xlink_title="standard length">SL</abbrev>
; pelvic fin relatively short, 4.0 (4.2) in
<abbrev id="ABBRID0EMTAE" xlink_title="standard length">SL</abbrev>
, and not reaching anus; caudal fin slightly emarginate, with several filaments on elongated ray branches (caudal fin in
<typeStatus id="C0BE4D3D06760CA99F0B417C46A9CE2E" type="paratype">paratype</typeStatus>
damaged); caudal peduncle length 3.9 (5.1) in HL; caudal peduncle depth 3.7 (4.2) in HL.
</paragraph>
<paragraph id="20C900CF4102CE95EEB5595FD55E62AD">
Morphometric values are summarized in Table
<tableCitation id="8CD80941FD11693367FA9976A7858301" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for Plectranthias raki sp. nov., expressed as a percentage of standard length." httpUri="http://table.plazi.org/id/79DB6D23893AF1CB3450DDCAB2145434" tableUuid="79DB6D23893AF1CB3450DDCAB2145434">1</tableCitation>
.
</paragraph>
<paragraph id="4615D38BBA69FCDB3D0D97AE1FE526A6">
Head of moderate size, 2.2 (2.1) in
<abbrev id="ABBRID0EYTAE" xlink_title="standard length">SL</abbrev>
; snout pointed, 3.4 (3.9) in HL; mouth relatively large, terminal and oblique, the posterior margin of maxilla reaching to vertical line almost directly beneath the center of pupil; upper jaw length 2.3 (2.5) in HL; maxilla expanded posteriorly; upper jaw with one large canine on each corner; 5 irregular rows of villiform teeth, largest in back, the teeth on the last row as big as the front canines; lower jaw with one small canine on either side of symphysis and 45 irregular rows of villiform teeth, the largest in the back; a pair of enlarged canines on each side of the dentary; teeth on the last row as big as the canines; palatine with 4 rows of small villiform teeth; vomer with 5 rows of villiform teeth.
</paragraph>
<paragraph id="04CBD8F486FFC7DFB82F261A19DA0D9B">Opercle with 3 spines, the middle one the largest, sharp and pointed, and terminating most posteriorly, the upper one obscured by scales; posterior margin of preopercle with 26 serrae; ventral margin of preopercle smooth and without conspicuous antrorse or retrorse spines; posterior margins of interopercle and subopercle smooth, obscured by scales; posttemporal with 3 small serrations; lower margin of infraorbital 1 smooth; anterior nostril located close to the anterior margin of orbit, with a small flap; posterior nostril located adjacent to anterior border of orbit, without flap.</paragraph>
<paragraph id="A769D77D984A3C21F9B595B6D618057F">
Scales ctenoid with peripheral cteni; lateral line broadly arched over pectoral fin, following body contour beneath dorsal fin to caudal-fin base; scales between eyes; no scales on maxilla, chin, mandible, lower part of snout, or branchiostegal rays; scales on head starting above the center of the eyes; triangular shaped patch of 6 rows of scales on pectoral fin, extending approximately ¼ ⅓ length of fin, extending furthest on 7
<superScript id="5C1C3735AA197B5E95E7772410C16908">th</superScript>
and 8
<superScript id="77AD302E9B4191072C2A568A9CE19E5E">th</superScript>
rays; scales on basal fourth of caudal fin; anal fin with one row of scales along anterior half of base; all other fins without scales.
</paragraph>
<paragraph id="9A4954A2D21A286663B7D4A3521FAEEF">
<emphasis id="F4E22BECC92579F098640F9FEA8ADFAE" bold="true" italics="true">Coloration when fresh</emphasis>
: (Figs
<figureCitation id="2561936E5ACEABD48866F123FACD876E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (left) and paratype (right) of Plectranthias raki sp. nov. Holotype CAS - ICH 248439 66.2 mm SL, shortly after collection (A), preserved specimen (B) and x-radiograph (C). Paratype LACM 61827, 70.4 mm SL, shortly after collection (D), preserved specimen (E) and x-radiograph (F). Photos: A, D by Luiz Rocha. B, C, E, F by Jon Fong." figureDoi="10.3897/zookeys.1223.135292.figure1" httpUri="https://binary.pensoft.net/fig/1221256">1 A, C</figureCitation>
,
<figureCitation id="C13FC27B202035DE1C08A688B37E5D67" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Living specimen (not retained) of Plectranthias raki sp. nov. photographed at 110 m depth at Dhaalu Atoll, Maldives. Photo by Luiz Rocha." figureDoi="10.3897/zookeys.1223.135292.figure2" httpUri="https://binary.pensoft.net/fig/1221257">2</figureCitation>
) Body pinkish-white with a series of irregularly shaped patches of red-orange along dorsal two-thirds of body; patches red-orange dorsally and posteriorly, becoming more yellow-orange anteriorly and ventrally; patches are golden yellow on opercle, especially on maxilla; one indistinct yellow stripe crossing opercle diagonally from bottom edge of eye; indistinct orange-red stripe extending from anterior margin of eye to upper lip; orbit white with indistinct yellow stripe through pupil, a continuation of the red-orange stripe originating at the snout tip; sharp break in orange-colored body-patches along lateral line, forming two distinct rows; this is especially pronounced in living specimens; patches above lateral line smaller and associated with pairs or trios of dorsal-fin spines and rays, creating an alternating pattern of five orange spots interspersed with white areas along the base of dorsal fin; patches below the lateral line are larger and more irregular in shape and distribution anteriorly, more rectangular and uniform on posterior half of body; throat and belly white; dorsal fin white at base; interspinous membranes of dorsal fin mostly yellow, white proximally, hyaline distally; membranes of soft dorsal mostly white with yellow stripe roughly midway from body and following contour of body; dorsal spines yellow, rays with yellow tips; anal fin mostly white proximally, yellow distally, with two yellow spots approximately one-third orbit diameter at base of third spine and fifth and sixth soft rays; anal-fin spines white; pelvic fins white proximally, hyaline distally, with yellow-orange spot at base of spine; pectoral fin white to hyaline, with orange spot, approximately one-half diameter of orbit, at fin base; caudal fin white at origin, becoming pale yellow distally, with two irregularly shaped yellow-orange spots, approximately one-third orbit diameter, located on either side of centermost fin rays; additional, smaller spots of similar coloration distally; tips of caudal-fin rays yellow; proximal half of dorsalmost and ventralmost caudal-fin rays yellow-orange.
</paragraph>
<caption id="E53958E7E3475D19EA5D720B230D9E89" ID-DOI="10.3897/zookeys.1223.135292.figure1" ID-arpha="CCD122FA-DFC6-556A-846F-177A69EB5476" httpUri="https://binary.pensoft.net/fig/1221256" startId="F1">
<paragraph id="9189B542F912DE927A1672CC05CB444F">
<label id="6DB2C80B2FA8349E705A5CF92F007E76">Figure 1.</label>
</paragraph>
<paragraph id="1B4ACC28DAAB3D9CF20C87F1A466F7CB">
Holotype (left) and paratype (right) of
<taxonomicName id="D1011C1B3422EBF7FE4E755405B5D5F3" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki" status="sp. nov.">
<emphasis id="D920407D7F0BA56F0E620C875171F12E" italics="true">Plectranthias raki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F6C118412D5AC92A5411E29FADB26703" rank="species">sp. nov.</taxonomicNameLabel>
Holotype
<named-content id="27ADC33DADB871C38535941A9CA60936" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences ichthyological collection">CAS</named-content>
- ICH 248439 66.2 mm
<abbrev id="ABBRID0EJVAE" xlink_title="standard length">SL</abbrev>
, shortly after collection (
<emphasis id="7C532555AC7DAA8810D6FF75CE9760A6" bold="true">A</emphasis>
), preserved specimen (
<emphasis id="531F8114EBA4698681723AC661FB4193" bold="true">B</emphasis>
) and x-radiograph (
<emphasis id="FA55D42BB68D9D9EB0F22E6803272526" bold="true">C</emphasis>
). Paratype
<named-content id="498DEA873873BA531037AEFFA1362489" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Natural History Museum of Los Angeles County">LACM</named-content>
61827, 70.4 mm
<abbrev id="ABBRID0EXVAE" xlink_title="standard length">SL</abbrev>
, shortly after collection (
<emphasis id="58A49982FA6688F2E3FADC9939546F0F" bold="true">D</emphasis>
), preserved specimen (
<emphasis id="E3AC6B7135328801904EAB4DBB29E31A" bold="true">E</emphasis>
) and x-radiograph (
<emphasis id="CD910B2AA3D6CA7C228047DFFC19222B" bold="true">F</emphasis>
). Photos:
<emphasis id="1EFD2DED55FD241C444B9D62F235300D" bold="true">A, D</emphasis>
by Luiz Rocha.
<emphasis id="02D426B25F6D0B076D4ACE01AC89F642" bold="true">B, C, E, F</emphasis>
by Jon Fong.
</paragraph>
</caption>
<caption id="C57D4E64561C60CDFE6CFF96675BCE6A" ID-DOI="10.3897/zookeys.1223.135292.figure2" ID-arpha="A180073C-14D6-5A46-8BE3-821D3636EB77" httpUri="https://binary.pensoft.net/fig/1221257" startId="F2">
<paragraph id="3C35D9379C600E33AF47B3A040C12F5A">
<label id="F9F24EDE9727CD51DE777A8F745098F5">Figure 2.</label>
</paragraph>
<paragraph id="1924339585619DFC4D07BE79902F5C26">
Living specimen (not retained) of
<taxonomicName id="51AE3DB54C93196DCB56D59DDAF487D4" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki" status="sp. nov.">
<emphasis id="9B0BE9F718C7E0EE5A143B073E1FB364" italics="true">Plectranthias raki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBD402D42A35D9C100AB5564A1E71E65" rank="species">sp. nov.</taxonomicNameLabel>
photographed at 110 m depth at Dhaalu Atoll, Maldives. Photo by Luiz Rocha.
</paragraph>
</caption>
<paragraph id="8770AF4B97952A6122EFF53811E73DDB">
<emphasis id="086B24937A53EA6D7DA387D8270F686B" bold="true" italics="true">Color in alcohol</emphasis>
: Uniform pale golden-brown with no distinct markings. Scattered melanophores on the nape and along base of dorsal fin (denser on the nape).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXAE" type="etymology">
<paragraph id="4724EA9D2545F874D837272143E149ED">
<heading id="EC5D5C196DC15C5D524021BB5195C40B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5796F6D7C948F1230CDB7C37112DEA82">
The species name,
<taxonomicName id="2000BEF058040E721E5DA64EF27A477F" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki">
<emphasis id="DAF5AC85A74EEAA7B69C7531DD40723A" italics="true">raki</emphasis>
</taxonomicName>
, means “ feeling shy to confront people ” in the Dhivehi language. This was chosen because
<taxonomicName id="16B2C54ADEA66AE865D4EDE31DD87BC8" authorityName="Bleeker" authorityYear="1873" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="genus">
<emphasis id="120647DFE5E46723C3F76BEAC23BDABC" italics="true">Plectranthias</emphasis>
</taxonomicName>
are shy by nature and typically hide from us when we are conducting surveys. To be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6XAE" type="distribution">
<paragraph id="1A186D8679E82554AFA344F078B4531C">
<heading id="A38166E4522501C669F371105CA0C453" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="BC90FCD059178B3082F3E7DA5D2B9A04">
<taxonomicName id="D1BB86901D17C5B08B1EE851EC8F7AD1" authorityName="Shepherd &amp; Pinheiro &amp; Najeeb &amp; Rocha &amp; Rocha" authorityYear="2025" family="Serranidae" genus="Plectranthias" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="raki" status="sp. nov.">
<emphasis id="21430DA9BF6D842725D691169AAAB185" italics="true">Plectranthias raki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30615954DA5953A4D073304A00A337BE" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the
<collectingCountry id="9516DF54F83D85C90EA9F1B5598032FF" name="Maldives">Maldives</collectingCountry>
, where it is likely widespread. It was seen inhabiting small holes of reef walls at several locations between Rasdhoo (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETYAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[72.950000,4.250000]}">
<geoCoordinate id="4D54AABE71090F441E2B3F1A8B644790" degrees="4" direction="north" minutes="15" orientation="latitude" precision="925" value="4.25">4 ° 15 ' N</geoCoordinate>
,
<geoCoordinate id="D35705941EF87B7090A0F98FF6C2BBB7" degrees="72" direction="east" minutes="57" orientation="longitude" precision="925" value="72.95">72 ° 57 ' E</geoCoordinate>
</named-content>
) and Dhaalu atolls (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2YAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[72.850000,2.683333]}">
<geoCoordinate id="8F6F5F86E51666B0E167E37BBF19E363" degrees="2" direction="north" minutes="41" orientation="latitude" precision="925" value="2.6833334">2 ° 41 ' N</geoCoordinate>
,
<geoCoordinate id="7933CEB3EECB0C03C6510E2DD033E7FD" degrees="72" direction="east" minutes="51" orientation="longitude" precision="925" value="72.85">72 ° 51 ' E</geoCoordinate>
</named-content>
) at approximately
<quantity id="8F4C7AB195C97006C15180E49BCE56B9" metricMagnitude="2" metricUnit="m" metricValue="1.125" metricValueMax="1.25" metricValueMin="1.0" unit="m" value="112.5" valueMax="125.0" valueMin="100.0">100 to 125 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>