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<mods:title>Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
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<mods:title id="F7C81C0501D2D1268623D42D646064F7">Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
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<mods:namePart>Will, Kipling</mods:namePart>
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<mods:affiliation>University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kipwill@berkeley.edu</mods:nameIdentifier>
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<mods:namePart id="BD43CC2AA731E04A3B276615364ED86E">Will, Kipling</mods:namePart>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<treatment ID-GBIF-Taxon="182754028" LSID="urn:lsid:zoobank.org:act:6FDFFBC6-C84B-4070-AEC5-D4D39F730356" httpUri="http://treatment.plazi.org/id/8FD37B09F28452A9B8457C6BBA2ADD1B" lastPageNumber="375" pageId="0" pageNumber="375">
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<taxonomicName LSID="http://zoobank.org/6FDFFBC6-C84B-4070-AEC5-D4D39F730356" authority="Will & Guéorguiev, 2021" authorityName="Will & Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis" status="sp. nov.">Coptocarpus amieuensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">Figs 20</figureCitation>
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<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">, 25</figureCitation>
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<figureCitation captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 35</figureCitation>
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<figureCitation captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 53</figureCitation>
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<treatment id="8FD37B09F28452A9B8457C6BBA2ADD1B" ID-GBIF-Taxon="182754028" LSID="urn:lsid:zoobank.org:act:6FDFFBC6-C84B-4070-AEC5-D4D39F730356" httpUri="http://treatment.plazi.org/id/8FD37B09F28452A9B8457C6BBA2ADD1B" lastPageNumber="375" pageId="0" pageNumber="375" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Thryptocerina" scope_tribe="Oodini">
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<taxonomicName id="9BDBB880C34487079A241E504B09701C" ID-CoL="8M8N2" LSID="http://zoobank.org/6FDFFBC6-C84B-4070-AEC5-D4D39F730356" authority="Will & Guéorguiev, 2021" authorityName="Will & Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis" status="sp. nov.">Coptocarpus amieuensis</taxonomicName>
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<taxonomicNameLabel id="373D1638DE49E4ECB05706DD1E2269DB" pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
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<figureCitation id="837DA14DC92D5EA2D047DEA17CD1A696" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">Figs 20</figureCitation>
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<figureCitation id="57340CE1FABA230792C80056C0E0109D" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">, 25</figureCitation>
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<figureCitation id="A962D36192FE5082C5A8202768B91D29" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 35</figureCitation>
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<figureCitation id="859CBBADA7B27A4E79F0187B7A43BC60" captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 53</figureCitation>
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<paragraph pageId="0" pageNumber="375">Material examined.</paragraph>
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<materialsCitation collectingDate="2007-03-04" collectingDateMax="2007-03-16" collectingDateMin="2007-03-04" country="NEW CALEDONIA" elevation="440" latitude="-21.56" location="New Caledonia" longLatPrecision="21" longitude="165.75555" specimenCount="♂" typeStatus="Holotype">
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<paragraph id="330332C143A0D971A5714E56DCBF6400" pageId="0" pageNumber="375">Material examined.</paragraph>
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:
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•
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; "
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/
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<collectingCountry id="40D3AF7D7F97D2CCE3A18FEE3DAFEC9F" name="New Caledonia">NEW CALEDONIA</collectingCountry>
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: Prov Sud. Col
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,
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el,
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||||
headlamp search"//
|
||||
<normalizedToken originalValue="“EMEC1139863”">"EMEC1139863"</normalizedToken>
|
||||
<normalizedToken id="EA58C56E1ED12DA8BD5539F9365DF4AF" originalValue="“EMEC1139863”">"EMEC1139863"</normalizedToken>
|
||||
</materialsCitation>
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
<typeStatus id="D907607ECFFBC2EB8C921E2C0D7B3CBA">Holotype</typeStatus>
|
||||
pinned, with genitalia in a separate microvial. Specimen relatively well-preserved, but with missing palpomeres, left antennomeres 2-11, right tarsomeres 1-5, left and right mesotibiae and mesotarsomeres 1-5, and left and right metatarsomeres 1-5. Source collection EMEC, deposited MNHN. Type locality as given on label.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="375" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="375">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="375">
|
||||
<subSubSection id="68E3DDCC799FF6187F63BA698727FDD2" pageId="0" pageNumber="375" type="diagnosis">
|
||||
<paragraph id="9C955EC6026A270DF03CEBA67087FB39" pageId="0" pageNumber="375">Diagnosis.</paragraph>
|
||||
<paragraph id="189C24C984D2CC74B59002DF3BA187F0" pageId="0" pageNumber="375">
|
||||
Most similar to
|
||||
<taxonomicName lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
|
||||
<taxonomicName id="3B0A06092CE4D2B80FE722F342E4BB28" lsidName="C. magnus" pageId="0" pageNumber="375" rank="species" species="magnus">
|
||||
<emphasis id="61827FC988C651CCA29E8C31731624C1" italics="true" pageId="0" pageNumber="375">C. magnus</emphasis>
|
||||
</taxonomicName>
|
||||
. However, the latter species has striae 2-5 shallowly impressed from near the elytra base to the apex (vs. 2-4 impressed only in the apical 1/3, and 5 in apical 1/2) and apical lamella of the median lobe more widely rounded at tip (Figs
|
||||
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
<figureCitation id="EAC58CD944A662F088A6A68DD1CB29F5" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">26</figureCitation>
|
||||
<figureCitation id="C2DBC6746FEE325C68DD1B092A280D85" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">26</figureCitation>
|
||||
). Also resembles
|
||||
<taxonomicName lsidName="C. cyllodinus" pageId="0" pageNumber="375" rank="species" species="cyllodinus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">C. cyllodinus</emphasis>
|
||||
<taxonomicName id="3DDC9C7DB66E52FD9755696942F0D238" lsidName="C. cyllodinus" pageId="0" pageNumber="375" rank="species" species="cyllodinus">
|
||||
<emphasis id="76BF8C1283FBC00CD89D5DFEB819851B" italics="true" pageId="0" pageNumber="375">C. cyllodinus</emphasis>
|
||||
</taxonomicName>
|
||||
but in
|
||||
<taxonomicName lsidName="C. amieuensis" pageId="0" pageNumber="375" rank="species" species="amieuensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">C. amieuensis</emphasis>
|
||||
<taxonomicName id="29B71C211EECE92452923D7625E18BCC" lsidName="C. amieuensis" pageId="0" pageNumber="375" rank="species" species="amieuensis">
|
||||
<emphasis id="D284DDE040C4B77F94D8D7E338BDD18F" italics="true" pageId="0" pageNumber="375">C. amieuensis</emphasis>
|
||||
</taxonomicName>
|
||||
the eyes are conical, head lacking a notch deep enough to receive the antenna ventral to the eye and the male metafemur does not have a prominent, ventral tooth. The male aedeagus is distinctly different (Fig.
|
||||
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
<figureCitation id="6CEE844C0F34EDA96AD1CB5558CE09A0" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="375" type="description">
|
||||
<paragraph pageId="0" pageNumber="375">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="375">
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
|
||||
<subSubSection id="D3AB967317CBCCF3243072531810DDDB" pageId="0" pageNumber="375" type="description">
|
||||
<paragraph id="2F6FFC46DA218A263687B07FA8506AC3" pageId="0" pageNumber="375">Description.</paragraph>
|
||||
<paragraph id="5626BC2362D9403B5D14EF1E7722B44B" pageId="0" pageNumber="375">
|
||||
<emphasis id="3AA940799F4427DAB457E041DF4EE7EE" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="1CD155E5ED661A4EA774C5AFBD6EEB9A" italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Small sized for oodines BL: 7.20 mm, BW: 3.50 mm, with ovate, moderately convex body (Fig.
|
||||
<figureCitation captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">20</figureCitation>
|
||||
<figureCitation id="568DDD93407736F5BC5B9DE3C7E72AE0" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.63775.figures18-21" httpUri="https://binary.pensoft.net/fig/554053" pageId="0" pageNumber="375">20</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
|
||||
<emphasis id="0BF9CED4A4BBB299A2CB6597103D7672" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="02CAE3F5B5B59B4B8C9831A31FE58148" italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Dorsal and ventral surface primarily black, with pronotum sides brownish black, fore part of head and femora dark reddish brown, antennomeres 1-3 and tibiae reddish brown, palpi, antennomeres 4-12 and tarsomeres yellowish brown. Integument moderately glossy, without spectral iridescence.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
|
||||
<emphasis id="6B76F7B52A1746C17FD4E2107711E5D7" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="74F5A9F8F0F7344F05BD94BCD01EA54B" italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head, pronotum and elytra with isodiametric meshes well impressed, prosternum in front and abdominal ventrites in middle with slightly transverse meshes, prosternal process with isodiametric meshes less impressed than on dorsal surface, rest of ventral surface with scarcely-visible sculpticells or sculpticells not apparent. Micropunctation evident on head, pronotum, prosternum medially (including prosternal process) and medial parts of abdominal ventrites, more or less uniform, lacking on remaining part of ventral surface; abdominal ventrite 1 with some large punctures at sides, ventrites 2-5 at sides moderately rugose.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Chaetotaxy</emphasis>
|
||||
<emphasis id="1A095B8B7C1666F15396CAE96EA4C1A6" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="B559F212BB5E9F06CCC26A81ED4CEB36" italics="true" pageId="0" pageNumber="375">Chaetotaxy</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Labrum with six setae distributed more or less evenly across width each in its own socket. Clypeal setae present. Supraorbital setae absent. Elytron without discal setiferous punctures. Abdominal ventrites 1-5 without ambulatory setae.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">Head.</emphasis>
|
||||
<emphasis id="8FDE1E82FDD166AE87D6610C71E738C4" bold="true" pageId="0" pageNumber="375">Head.</emphasis>
|
||||
Less than 2 1/2
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="393F002E1E661FE58EFA5770DFCE6A8D" originalValue="×">x</normalizedToken>
|
||||
as wide as pronotum (Suppl. material 2: Table S1). Eye relatively large, moderately protruded, EyW/HW: 1.47. Labrum anterior margin straight. Frontoclypeal suture present medially, almost obliterated at sides. Antenna moderately long, with last segment not exceeding pronotal base and pubescence starting from segment 4. Mentum tooth with paramedial border distinct.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
|
||||
<emphasis id="CB6611BD649496EE526AB386EB73896C" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="C12EA425D4CDD70F89EA339802057E05" italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Pronotum ~ 1 2/3 wider than long (PW/PL: 1.70); width at apex less than 2
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="8501C752CE461B788D92810C76E90FA1" originalValue="×">x</normalizedToken>
|
||||
at widest point (PW/PA: 2.16). Disc with middle line very fine, without apical transverse impression; anterior angles slightly, widely rounded at tips, anterior margin concave, anterior submarginal sulcus present, interrupted in middle. Prosternal process rhomboidal, finely bordered at apex, not bordered between coxae. Mesosternum anteriorly with two prominent, symmetrical tubercles separated by a space smaller or equal to diameter of one of them. Metepisternum wider than long, with lateral margin barely convex, coadunation with epipleuron long, extended along most of length.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">Elytra.</emphasis>
|
||||
<emphasis id="2ACDF6445D809AD201AAE7E8ACC4EF82" bold="true" pageId="0" pageNumber="375">Elytra.</emphasis>
|
||||
Slightly longer than wide (EL/EW: 1.11). Basal margin forming distinct denticle at shoulder, ended medially just before level of parascutellar puncture. Humeral submarginal carina present. Apical sinuation shallow. Parascutellar striole absent, striae 1-5 present at apex, ended well before basal margin; striae 6 and 7 well impressed throughout, 7 reaching basal margin, 6 ended near but not reaching margin. Elytral intervals 1-6 where evident at apex, convex, interval 7 slightly convex in anterior 1/2, clearly convex in posterior 1/2, interval 8 convex throughout, from 2-3
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="D77C989F266E7C92B14202311BFBAF29" originalValue="×">x</normalizedToken>
|
||||
narrower than interval 7. Granulation in marginal furrow evident in first 1/5 and last 3/5, lacking in second 1/5.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Legs</emphasis>
|
||||
<emphasis id="320879EDD6AA4F7518F67286F66B66C0" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="D23324571733BA9E15F4659D55066194" italics="true" pageId="0" pageNumber="375">Legs</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Male protarsomeres 1-3 asymmetrically expanded, beneath with adhesive setae (tarsomeres 2 and 3 fully covered, tarsomere 1 covered only on apical 1/4), each of them transverse, wider than long.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
|
||||
<emphasis id="CC3C43287F9109B0D48383F21DD32021" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="CC930F78BF1BB31BDA685FE30A864CC6" italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Female specimens unknown.
|
||||
<emphasis bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
|
||||
<emphasis id="486EA32CAA71169DEC85A8E679541022" bold="true" pageId="0" pageNumber="375">
|
||||
<emphasis id="06406640B17FBCA652E3BCB90D860A04" italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Median lobe of aedeagus in lateral view long, curved ventrally, with rather thin apex (Fig.
|
||||
<figureCitation captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">35</figureCitation>
|
||||
<figureCitation id="3B4662E303F4303213C0BEC89A753747" captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">35</figureCitation>
|
||||
); lobe in dorsal view, with apical lamella well bent to left, with left side clearly concave and right side slightly sinuate (Fig.
|
||||
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
<figureCitation id="A68B306EBB39B1DB8FFC8D76CF3403EE" captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">25</figureCitation>
|
||||
); basal bulb rounded dorsally; sclerotized portion of the endophallus with three sclerites, most clearly visible from ventral side.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="375" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="375">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="375">
|
||||
<subSubSection id="17CA4EA971416E01158E6A52ED47A801" pageId="0" pageNumber="375" type="etymology">
|
||||
<paragraph id="5F0ADBD583B973302ED90F68F12574FA" pageId="0" pageNumber="375">Etymology.</paragraph>
|
||||
<paragraph id="B7DB8980E37C6136B00D3926F77BE8CC" pageId="0" pageNumber="375">
|
||||
The specific epithet
|
||||
<taxonomicName authorityName="Will & Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="375">Coptocarpus amieuensis</emphasis>
|
||||
<taxonomicName id="3BD4529851105CF793B386ECD58A5601" authorityName="Will & Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Coptocarpus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocarpus amieuensis" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="amieuensis">
|
||||
<emphasis id="618D9514B847AC89C5F16197564F32CB" italics="true" pageId="0" pageNumber="375">Coptocarpus amieuensis</emphasis>
|
||||
</taxonomicName>
|
||||
is based on the type locality Col
|
||||
<normalizedToken originalValue="d’Amieu">d'Amieu</normalizedToken>
|
||||
<normalizedToken id="34A6F6201A851DB9986410548F85FDD6" originalValue="d’Amieu">d'Amieu</normalizedToken>
|
||||
and is treated as an adjective.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,178 +1,178 @@
|
|||
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7.
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|
||||
<paragraph id="325B2FA2E14E7644219A929A9D167195" pageId="0" pageNumber="193">
|
||||
"
|
||||
<taxonomicName authorityName="Stichel" authorityYear="1906" class="Insecta" family="Papilionidae" genus="Parnassius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parnassius apollo subsp. pumilus" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="apollo" subSpecies="pumilus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="193">Parnassius apollo pumilus</emphasis>
|
||||
<taxonomicName id="CEB13D34D864FB59E767B0581B5F49E8" authorityName="Stichel" authorityYear="1906" class="Insecta" family="Papilionidae" genus="Parnassius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parnassius apollo subsp. pumilus" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="apollo" subSpecies="pumilus">
|
||||
<emphasis id="49A5458A8DED62C9D2078CBC963EFF48" italics="true" pageId="0" pageNumber="193">Parnassius apollo pumilus</emphasis>
|
||||
</taxonomicName>
|
||||
Stich. forma bispupillata" Turati, 1918 Atti. Soc. Ital. Sci. Nat. 57: 40.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="193" type="current combination">
|
||||
<paragraph pageId="0" pageNumber="193">Current combination.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="193">
|
||||
<emphasis bold="true" pageId="0" pageNumber="193">
|
||||
<taxonomicName authorityName="Stichel" authorityYear="1906" class="Insecta" family="Papilionidae" genus="Parnassius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parnassius apollo subsp. pumilus" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="apollo" subSpecies="pumilus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="193">Parnassius apollo pumilus</emphasis>
|
||||
<subSubSection id="11BBD1585A318F882705006939A17F01" pageId="0" pageNumber="193" type="current combination">
|
||||
<paragraph id="5BF288802AFCBF107C7F1845F55275CA" pageId="0" pageNumber="193">Current combination.</paragraph>
|
||||
<paragraph id="63851256BC3F1A2EDB859F4AF61EE0B9" pageId="0" pageNumber="193">
|
||||
<emphasis id="9111E64FF837EE8FFAAA80D5E1291BA6" bold="true" pageId="0" pageNumber="193">
|
||||
<taxonomicName id="381376871E10612204EEB051CC03691B" authorityName="Stichel" authorityYear="1906" class="Insecta" family="Papilionidae" genus="Parnassius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parnassius apollo subsp. pumilus" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="apollo" subSpecies="pumilus">
|
||||
<emphasis id="64C7B35550247680466F0350BCBF10B8" italics="true" pageId="0" pageNumber="193">Parnassius apollo pumilus</emphasis>
|
||||
</taxonomicName>
|
||||
f. bispupillata Turati, 1918
|
||||
</emphasis>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="193" type="current status">
|
||||
<paragraph pageId="0" pageNumber="193">Current status.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="193">Infrasubspecific and hence unavailable name.</paragraph>
|
||||
<subSubSection id="354D8820DA37A60F7151C0664F36A455" pageId="0" pageNumber="193" type="current status">
|
||||
<paragraph id="23A6414F7D79AE7210B679155CA6D4E1" pageId="0" pageNumber="193">Current status.</paragraph>
|
||||
<paragraph id="8F073573022699F646FBC71B96FAA19B" pageId="0" pageNumber="193">Infrasubspecific and hence unavailable name.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="193" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="193">Original material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="193">
|
||||
<materialsCitation accessionNumber="ZMH 61588" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. Warnecke" elevation="1800" specimenCount="1">
|
||||
<subSubSection id="F270E7460DD62289044F0361B9F9CE77" pageId="0" pageNumber="193" type="materials_examined">
|
||||
<paragraph id="5CB9DC7B3B99CC96883D0BAD402F26AC" pageId="0" pageNumber="193">Original material.</paragraph>
|
||||
<paragraph id="BC908DBCF2D0FB7CA4D98F5CFE86DB2A" pageId="0" pageNumber="193">
|
||||
<materialsCitation id="73858A7E038FE5B14BC26EFB063BE5E3" accessionNumber="ZMH 61588" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. Warnecke" elevation="1800" specimenCount="1">
|
||||
Labelled as
|
||||
<normalizedToken originalValue="“Cotype”">"Cotype"</normalizedToken>
|
||||
<normalizedToken id="09C5FCE3367FAC615C2B9173B81C4BA7" originalValue="“Cotype”">"Cotype"</normalizedToken>
|
||||
1? (
|
||||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61588">ZMH 61588</accessionNumber>
|
||||
<accessionNumber id="76D8B4A776A2C8EE4A653D7F4557F64B" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61588">ZMH 61588</accessionNumber>
|
||||
) (Fig.
|
||||
<figureCitation captionStart="Plate 2" captionStartId="F2" captionText="Plate 2. 7. Parnassius apollo pumilus f. bispupillata Turati, 1918; 8. Parnssius apollo pumilus f. appendiculata Turati, 1918; 9. Parnassius apollo ab. lacrimans Marschner, 1911; 10. Parnassius apollo oenipontanus Stauder & Felkel, 1921; 11. Parnassius apollo serpentinicus Mayer, 1925 12. Parnassius apollo ab. marschneri Bryk, 1911. a. Dorsal view, b. ventral view." httpUri="https://binary.pensoft.net/fig/579711" pageId="0" pageNumber="193">7</figureCitation>
|
||||
<figureCitation id="C0262CF8F907BA354F1E14D06B13DFA1" captionStart="Plate 2" captionStartId="F2" captionText="Plate 2. 7. Parnassius apollo pumilus f. bispupillata Turati, 1918; 8. Parnssius apollo pumilus f. appendiculata Turati, 1918; 9. Parnassius apollo ab. lacrimans Marschner, 1911; 10. Parnassius apollo oenipontanus Stauder & Felkel, 1921; 11. Parnassius apollo serpentinicus Mayer, 1925 12. Parnassius apollo ab. marschneri Bryk, 1911. a. Dorsal view, b. ventral view." httpUri="https://binary.pensoft.net/fig/579711" pageId="0" pageNumber="193">7</figureCitation>
|
||||
). "Calabrien / [illegible],
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
|
||||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
|
||||
<quantity id="8C42C9471CFFFA51ED10A64064B277FF" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
|
||||
<elevation id="44949B8858E35880E1DF5F7A26DC6C32" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
|
||||
</quantity>
|
||||
/ Stauder, 5.7.20" //
|
||||
<normalizedToken originalValue="“bispupillata”">"bispupillata"</normalizedToken>
|
||||
<normalizedToken id="E5067306EBBEA9F87C18828E322144E7" originalValue="“bispupillata”">"bispupillata"</normalizedToken>
|
||||
//
|
||||
<normalizedToken originalValue="“bispupillata”">"bispupillata"</normalizedToken>
|
||||
<normalizedToken id="FCC438945BDC92CCDC06C56EC8376525" originalValue="“bispupillata”">"bispupillata"</normalizedToken>
|
||||
//
|
||||
<normalizedToken originalValue="“Cotype”">"Cotype"</normalizedToken>
|
||||
<normalizedToken id="13CA10FE18A0F911D317FB19BABB7351" originalValue="“Cotype”">"Cotype"</normalizedToken>
|
||||
// "Sammlung /
|
||||
<collectorName>G. Warnecke</collectorName>
|
||||
/ Eing. Nr. 5, 1949" // "
|
||||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61588">ZMH 61588</accessionNumber>
|
||||
<collectorName id="BBB4031BAA1980B8A6D7D93F7F56FC1C">G. Warnecke</collectorName>
|
||||
/ Eing. Nr. 5, 1949" // "
|
||||
<accessionNumber id="7011F47DCC64A2CA5E574009C9B95C6E" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61588">ZMH 61588</accessionNumber>
|
||||
"
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="193" type="original locality">
|
||||
<paragraph pageId="0" pageNumber="193">Original locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="193">Italy: Calabria.</paragraph>
|
||||
<subSubSection id="9FE51CDB63F7796F91970FB43AAD196D" pageId="0" pageNumber="193" type="original locality">
|
||||
<paragraph id="E102A0C367DD2A95BFAF904F0C507E0D" pageId="0" pageNumber="193">Original locality.</paragraph>
|
||||
<paragraph id="39D54083FDC3CD9079450C8F0C659109" pageId="0" pageNumber="193">Italy: Calabria.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="193" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="193">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="193">
|
||||
<bibRefCitation author="Turati, E" journalOrPublisher="Atti della Societa italiana di scienze naturali" pageId="0" pageNumber="193" pagination="28 - 89" refId="B88" refString="Turati, E, 1918. Variabilita' Del Parnassius apollo pumilus Stich. e ricerche sull'origine dei Parnassius (Note Biologico-Critiche). Atti della Societa italiana di scienze naturali 57: 28 - 89" title="Variabilita' Del Parnassius apollo pumilus Stich. e ricerche sull'origine dei Parnassius (Note Biologico-Critiche)." volume="57" year="1918">Turati (1918)</bibRefCitation>
|
||||
<subSubSection id="473C97C88487BD3AA75D8396D55B0B6C" pageId="0" pageNumber="193" type="remarks">
|
||||
<paragraph id="AF7ECCED4E94C4CD1BFFC4D1563FD924" pageId="0" pageNumber="193">Remarks.</paragraph>
|
||||
<paragraph id="A462B79185B34ECDE8ABF40033420303" pageId="0" pageNumber="193">
|
||||
<bibRefCitation id="7386D350ED6BCC2DCFF67BAEE7498E2B" author="Turati, E" journalOrPublisher="Atti della Societa italiana di scienze naturali" pageId="0" pageNumber="193" pagination="28 - 89" refId="B88" refString="Turati, E, 1918. Variabilita' Del Parnassius apollo pumilus Stich. e ricerche sull'origine dei Parnassius (Note Biologico-Critiche). Atti della Societa italiana di scienze naturali 57: 28 - 89" title="Variabilita' Del Parnassius apollo pumilus Stich. e ricerche sull'origine dei Parnassius (Note Biologico-Critiche)." volume="57" year="1918">Turati (1918)</bibRefCitation>
|
||||
proposed this name as a form of
|
||||
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. a. subsp. pumilus" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="a." subSpecies="pumilus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="193">P. a. pumilus</emphasis>
|
||||
<taxonomicName id="567A6F93EF5AFFCFE6008B8038F958A1" class="Insecta" kingdom="Animalia" lsidName="P. a. subsp. pumilus" order="Coleoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="a." subSpecies="pumilus">
|
||||
<emphasis id="F916774ED3C0C8AF702A7FEEE8FCD418" italics="true" pageId="0" pageNumber="193">P. a. pumilus</emphasis>
|
||||
</taxonomicName>
|
||||
Stichel, 1906. Therefore, as stated by article 45.6.1 (
|
||||
<bibRefCitation author="ICZN" journalOrPublisher="Internationale Entomologische Zeitschrift" pageId="0" pageNumber="193" publicationUrl="http://www.iczn.org/iczn/index.jsp" refId="B47" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. London. [available online at http://www.iczn.org/iczn/index.jsp" title="International Code of Zoological Nomenclature. Fourth edition. London. [available online at" url="http://www.iczn.org/iczn/index.jsp" year="1999">ICZN 1999</bibRefCitation>
|
||||
<bibRefCitation id="A52477C7267D343D02EE7103CCFD737A" author="ICZN" journalOrPublisher="Internationale Entomologische Zeitschrift" pageId="0" pageNumber="193" publicationUrl="http://www.iczn.org/iczn/index.jsp" refId="B47" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. London. [available online at http://www.iczn.org/iczn/index.jsp" title="International Code of Zoological Nomenclature. Fourth edition. London. [available online at" url="http://www.iczn.org/iczn/index.jsp" year="1999">ICZN 1999</bibRefCitation>
|
||||
) it is deemed to be an infrasubspecific name (if the author expressly gave it infrasubspecific rank) and is hence unavailable.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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