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<taxonomicName id="4C2D4D02FF88FFE79B8DE21EAE3015FF" ID-CoL="9X5LT" authorityName="Bai, X. Wang, Zhou, Lin, Meng &amp; Fontoura" authorityYear="2020" box="[151,618,367,394]" class="Heterotardigrada" family="Halechiniscidae" genus="Moebjergarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="clarionclippertonensis">
<emphasis id="B959EA93FF88FFE79B8DE21EAE3015FF" bold="true" box="[151,618,367,394]" italics="true" pageId="16" pageNumber="17">Moebjergarctus clarionclippertonensis</emphasis>
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<bibRefCitation id="EFBC4B70FF88FFE7996BE21EA8E615FC" author="Bai, L. &amp; Wang, X. &amp; Zhou, Y. &amp; Lin, S. &amp; Meng, F. &amp; Fontoura, P." box="[625,1212,367,393]" pageId="16" pageNumber="17" pagination="561 - 575" refId="ref17496" refString="Bai, L., Wang, X., Zhou, Y., Lin, S., Meng, F. &amp; Fontoura, P. (2020) Moebjergarctus clarionclippertonensis, a new abyssal tardigrade (Arthrotardigrada, Halechiniscidae, Euclavarctinae) from the Clarion-Clipperton Fracture Zone, North-East Pacific. Zootaxa, 4755 (3), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4755.3.8" type="journal article" year="2020">
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N=
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:
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, 2 with undetermined sex occurrence: ANDEEP 1,
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;
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bsl
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<figureCitation id="13162A04FF88FFE79BDDE28EAD51166F" box="[199,267,511,538]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
; Supplementary
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<emphasis id="B959EA93FF88FFE79B8DE136AD3D1614" bold="true" box="[151,359,583,609]" pageId="16" pageNumber="17">Short description</emphasis>
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Large and robust tardigrade 148283 µm long and 52.399.7 µm width (n=2). Buccal apparatus with stylets (
<figureCitation id="13162A04FF88FFE79B85E1FEACAD16DC" box="[159,247,655,681]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7B</figureCitation>
). Cephalic cirri with annulated base, smooth portion of the scapus and short flagellum (
<figureCitation id="13162A04FF88FFE79FA2E1FEA97016DC" box="[1208,1322,655,682]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7E, F</figureCitation>
). Each leg with a sensory organ (
<figureCitation id="13162A04FF88FFE79A8BE1C2ADB216B8" box="[401,488,691,717]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7E</figureCitation>
). Internal and external digits are of the same size and wrinkled (
<figureCitation id="13162A04FF88FFE79FA7E1C2A96316B8" box="[1213,1337,691,717]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7C, G</figureCitation>
). Caudal region is covered with a thick cuticular layer shown as a bulge (
<figureCitation id="13162A04FF88FFE79847E1A6AF8C1684" box="[861,982,727,753]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7A, E</figureCitation>
). All the females have a patched seminal receptacle opening (
<figureCitation id="13162A04FF88FFE79A63E18AADAC1760" box="[377,502,763,790]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1703]" captionTargetBox="[193,1394,184,1653]" captionTargetId="figure-17@15.[190,1397,181,1656]" captionTargetPageId="15" captionText="FIGURE 7. Moebjergarctus clarionclippertonensis. A Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note caudal bulge (arrowhead). B Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), dorsal view, note buccal apparatus stylet (arrowhead). C Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), ventral view, note leg II digits (arrowhead). D Moebjergarctus clarionclippertonensis, female, light microscopy (DIC), caudal region, note patched seminal receptacle opening (arrowhead). E Moebjergarctus clarionclippertonensis, female, SEM, sagittal view. F Moebjergarctus clarionclippertonensis, female, SEM, head, dorsal view, note smooth portion of internal cirrus (arrowhead). G Moebjergarctus clarionclippertonensis, female, SEM, leg III digits. H Moebjergarctus clarionclippertonensis, female, SEM, posterior ventro-sagittal view, note seminal receptacle opening patch borders (arrowheads). Abbreviations: an—anus, bu—caudal bulge, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pb— pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385105" httpUri="https://zenodo.org/record/14385105/files/figure.png" pageId="16" pageNumber="17">Fig. 7D, H</figureCitation>
).
</paragraph>
<subSubSection id="C337650AFF88FFE79B8DE032AF84102C" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B923681FF88FFE79B8DE032AD5E1728" blockId="16.[151,1436,835,1114]" box="[151,260,835,861]" pageId="16" pageNumber="17">
<emphasis id="B959EA93FF88FFE79B8DE032AD5E1728" bold="true" box="[151,260,835,861]" pageId="16" pageNumber="17">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF88FFE79BDDE016AF84102C" blockId="16.[151,1436,835,1114]" pageId="16" pageNumber="17">
The identification of this species is certain because of the morphology of cephalic cirri (annulated base, smooth scapus and short flagellum). The internal and external digits of all limbs are the same size that distinguishes this species from
<taxonomicName id="4C2D4D02FF88FFE79A29E0C1AD8817BC" authorityName="Saulenko, Maiorova, Arbizu &amp; Mordukhovich" authorityYear="2022" box="[307,466,943,969]" class="Heterotardigrada" family="Halechiniscidae" genus="Moebjergarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="okhotensis">
<emphasis id="B959EA93FF88FFE79A29E0C1AD8817BC" box="[307,466,943,969]" italics="true" pageId="16" pageNumber="17">M. okhotensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBC4B70FF88FFE79AFEE0DEAEC217BC" author="Bai, L. &amp; Wang, X. &amp; Zhou, Y. &amp; Lin, S. &amp; Meng, F. &amp; Fontoura, P." box="[484,664,943,969]" pageId="16" pageNumber="17" pagination="561 - 575" refId="ref17496" refString="Bai, L., Wang, X., Zhou, Y., Lin, S., Meng, F. &amp; Fontoura, P. (2020) Moebjergarctus clarionclippertonensis, a new abyssal tardigrade (Arthrotardigrada, Halechiniscidae, Euclavarctinae) from the Clarion-Clipperton Fracture Zone, North-East Pacific. Zootaxa, 4755 (3), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4755.3.8" type="journal article" year="2020">
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<emphasis id="B959EA93FF88FFE7990DE0C1AE0917BC" box="[535,595,943,969]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2020
</bibRefCitation>
,
<bibRefCitation id="EFBC4B70FF88FFE799BCE0DEAFC117BC" author="Saulenko, A. A. &amp; Maiorova, A. S. &amp; Martinez Arbizu, P. &amp; Mordukhovich, V. V." box="[678,923,943,969]" pageId="16" pageNumber="17" refId="ref22197" refString="Saulenko, A. A., Maiorova, A. S., Martinez Arbizu, P. &amp; Mordukhovich, V. V. (2022) Deep-sea tardigrades from the NorthWestern Pacific, with descriptions of two new species. Diversity, 14 (12), 1086. https: // doi. org / 10.3390 / d 14121086" type="journal volume" year="2022">
Saulenko
<emphasis id="B959EA93FF88FFE79803E0C1AF0E17BC" box="[793,852,943,969]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2022
</bibRefCitation>
). This species was previously reported from Pacific deep-sea plains with a high abundance of manganese or polymetallic nodules (
<bibRefCitation id="EFBC4B70FF88FFE79F45E0A2A9511798" author="Bai, L. &amp; Wang, X. &amp; Zhou, Y. &amp; Lin, S. &amp; Meng, F. &amp; Fontoura, P." box="[1119,1291,979,1005]" pageId="16" pageNumber="17" pagination="561 - 575" refId="ref17496" refString="Bai, L., Wang, X., Zhou, Y., Lin, S., Meng, F. &amp; Fontoura, P. (2020) Moebjergarctus clarionclippertonensis, a new abyssal tardigrade (Arthrotardigrada, Halechiniscidae, Euclavarctinae) from the Clarion-Clipperton Fracture Zone, North-East Pacific. Zootaxa, 4755 (3), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4755.3.8" type="journal article" year="2020">
<collectingRegion id="49E9F863FF88FFE79F45E0A2A8DD1798" box="[1119,1159,979,1005]" country="Somalia" name="Bay" pageId="16" pageNumber="17">Bai</collectingRegion>
<emphasis id="B959EA93FF88FFE79F94E0A5A89D1798" box="[1166,1223,979,1005]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2020
</bibRefCitation>
). At least the sampling stations of expedition ANDEEP 1 that contained
<taxonomicName id="4C2D4D02FF88FFE79820E089A8041064" authorityName="Bai, X. Wang, Zhou, Lin, Meng &amp; Fontoura" authorityYear="2020" box="[826,1118,1015,1041]" class="Heterotardigrada" family="Halechiniscidae" genus="Moebjergarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="clarionclippertonensis">
<emphasis id="B959EA93FF88FFE79820E089A8041064" box="[826,1118,1015,1041]" italics="true" pageId="16" pageNumber="17">M. clarionclippertonensis</emphasis>
</taxonomicName>
(Shackleton Fracture Zone,
<geoCoordinate id="EE195046FF88FFE79B8DE76CAD4C104D" box="[151,278,1053,1080]" degrees="59" direction="south" minutes="52.30" orientation="latitude" pageId="16" pageNumber="17" precision="9" value="-59.871666">59°52.30S</geoCoordinate>
,
<geoCoordinate id="EE195046FF88FFE79A3EE76CADF6104D" box="[292,428,1053,1080]" degrees="59" direction="west" minutes="57.63" orientation="longitude" pageId="16" pageNumber="17" precision="9" value="-59.9605">59°57.63W</geoCoordinate>
) at the deep sea bottom of the Drake Passage are also close to areas with polymetallic sulfide deposits and hydrothermal vent activity (see discussion for details).
</paragraph>
</subSubSection>
</subSection>
</subSubSection>
</treatment>
</document>

View file

@ -1,119 +0,0 @@
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<paragraph id="8B923681FF88FFE79B8DE6BAAFB4127C" blockId="16.[151,1006,1483,1582]" pageId="16" pageNumber="17">
N=
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occurrence: ANDEEP I,
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<figureCitation id="13162A04FF88FFE79BDDE562AD181258" box="[199,322,1555,1581]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8AD</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF88FFE79AE4E562AE3C125B" box="[510,614,1555,1582]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="16" pageNumber="17">Table 1.5</tableCitation>
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View file

@ -0,0 +1,166 @@
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<heading id="D0DA81EDFF88FFE79B8DE6F2AEC511E8" bold="true" box="[151,671,1411,1438]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C2D4D02FF88FFE79B8DE6F2AEC511E8" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[151,671,1411,1438]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="abyssalis">
<emphasis id="B959EA93FF88FFE79B8DE6F2AEC511E8" bold="true" box="[151,671,1411,1438]" pageId="16" pageNumber="17">
<emphasis id="B959EA93FF88FFE79B8DE6F2AD2911EB" bold="true" box="[151,371,1411,1438]" italics="true" pageId="16" pageNumber="17">Angursa abyssalis</emphasis>
<bibRefCitation id="EFBC4B70FF88FFE79A60E6F2AEC511E8" author="Renaud-Mornant, J." box="[378,671,1411,1437]" pageId="16" pageNumber="17" pagination="161 - 164" refId="ref21762" refString="Renaud-Mornant, J. (1981) Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys, 10, 161 - 164." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
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N=
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occurrence: ANDEEP I,
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,
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.
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<paragraph id="8B923681FF88FFE79BDDE562AE3C125B" blockId="16.[151,1006,1483,1582]" box="[199,614,1555,1582]" pageId="16" pageNumber="17">
<figureCitation id="13162A04FF88FFE79BDDE562AD181258" box="[199,322,1555,1581]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8AD</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF88FFE79AE4E562AE3C125B" box="[510,614,1555,1582]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="16" pageNumber="17">Table 1.5</tableCitation>
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<emphasis id="B959EA93FF88FFE79B8DE52AAD3D1200" bold="true" box="[151,359,1627,1653]" pageId="16" pageNumber="17">Short description</emphasis>
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<taxonomicName id="4C2D4D02FF88FFE79BDDE50EAD7C12EC" authorityName="Pollock" authorityYear="1979" box="[199,294,1663,1689]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF88FFE79BDDE50EAD7C12EC" box="[199,294,1663,1689]" italics="true" pageId="16" pageNumber="17">Angursa</emphasis>
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with slender body (120180 µm long, 2035 µm width, n=5) and primary clavae longer than cirri
<emphasis id="B959EA93FF88FFE79B8DE5D2ACF212C8" box="[151,168,1699,1725]" italics="true" pageId="16" pageNumber="17">A</emphasis>
. Secondary and tertiary clavae are usually clearly visible (
<figureCitation id="13162A04FF88FFE79848E5D2AFAF12C8" box="[850,1013,1699,1726]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8A, B, B</figureCitation>
<figureCitation id="13162A04FF88FFE798EFE5D2A85F12C8" box="[1013,1029,1699,1725]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="16" pageNumber="17">2</figureCitation>
). Distance between both secondary clavae on the dorsal side of the head is 1.54.0 μm (
<figureCitation id="13162A04FF88FFE799E7E5B6AF201294" box="[765,890,1735,1761]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8B, B</figureCitation>
<figureCitation id="13162A04FF88FFE79860E5B6AFD01294" box="[890,906,1735,1761]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="16" pageNumber="17">2</figureCitation>
). Cirrus
<emphasis id="B959EA93FF88FFE798EAE5B9A85B1294" box="[1008,1025,1736,1761]" italics="true" pageId="16" pageNumber="17">E</emphasis>
short (8.012.0 µm, n=5) (Fig. D). The hemispherical sensory organ with a small apical spine on leg IV is sometimes a bit elongated (as in
<taxonomicName id="4C2D4D02FF88FFE79E27E59AACB4135C" authorityName="Bussau" authorityYear="1992" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="capsula">
<emphasis id="B959EA93FF88FFE79E27E59AACB4135C" italics="true" pageId="16" pageNumber="17">Angursa capsula</emphasis>
</taxonomicName>
) (
<figureCitation id="13162A04FF88FFE79A1DE47EAD05135C" box="[263,351,1807,1833]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF88FFE79B8DE426AF0C13A8" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B923681FF88FFE79B8DE426AD5E1304" blockId="16.[151,1436,1879,2013]" box="[151,260,1879,1905]" pageId="16" pageNumber="17">
<emphasis id="B959EA93FF88FFE79B8DE426AD5E1304" bold="true" box="[151,260,1879,1905]" pageId="16" pageNumber="17">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF88FFE79BDDE40AAF0C13A8" blockId="16.[151,1436,1879,2013]" pageId="16" pageNumber="17">
The two species
<taxonomicName id="4C2D4D02FF88FFE79A64E40AAE5913E0" authorityName="Renaud-Mornant" authorityYear="1981" box="[382,515,1915,1941]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="abyssalis">
<emphasis id="B959EA93FF88FFE79A64E40AAE5913E0" box="[382,515,1915,1941]" italics="true" pageId="16" pageNumber="17">A. abyssalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D02FF88FFE7992DE40AAEF113E0" authorityName="Bussau" authorityYear="1992" box="[567,683,1915,1941]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="17" phylum="Tardigrada" rank="species" species="capsula">
<emphasis id="B959EA93FF88FFE7992DE40AAEF113E0" box="[567,683,1915,1941]" italics="true" pageId="16" pageNumber="17">A. capsula</emphasis>
</taxonomicName>
are quite similar in the shape of the sensory organ of leg IV (Hansen and
<bibRefCitation id="EFBC4B70FF88FFE79BD0E4EEAD2213CC" author="Fujimoto, S. &amp; Hansen, J. G." box="[202,376,1951,1977]" pageId="16" pageNumber="17" pagination="1 - 19" refId="ref19180" refString="Fujimoto, S. &amp; Hansen, J. G. (2019) Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, 510, 1 - 19. https: // doi. org / 10.5852 / ejt. 2019.510" type="journal article" year="2019">Fujimoto 2019</bibRefCitation>
). However, the main identifying trait for this species is the short (less than 4 µm) distance between the secondary clavae and the shorter cirri
<emphasis id="B959EA93FF88FFE799D5E4B5AEBA13A8" box="[719,736,1988,2013]" italics="true" pageId="16" pageNumber="17">E</emphasis>
(
<figureCitation id="13162A04FF88FFE799F5E4B2AF1313A8" box="[751,841,1987,2013]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="16" pageNumber="17">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,154 @@
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<paragraph id="8B923681FF8AFFE59B8DE081AECC107F" blockId="18.[151,662,1007,1034]" box="[151,662,1007,1034]" pageId="18" pageNumber="19">
<heading id="D0DA81EDFF8AFFE59B8DE081AECC107F" bold="true" box="[151,662,1007,1034]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C2D4D02FF8AFFE59B8DE081AECC107F" ID-CoL="66QM4" authority="Villora-Moreno, 1998" authorityName="Villora-Moreno" authorityYear="1998" box="[151,662,1007,1034]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="species" species="antarctica">
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<emphasis id="B959EA93FF8AFFE59B8DE081ADDB107F" bold="true" box="[151,385,1008,1034]" italics="true" pageId="18" pageNumber="19">Angursa antarctica</emphasis>
<bibRefCitation id="EFBC4B70FF8AFFE59A92E09EAECC107F" author="Villora-Moreno, S." box="[392,662,1007,1034]" pageId="18" pageNumber="19" pagination="336 - 341" refId="ref22898" refString="Villora-Moreno, S. (1998) Deep-sea Tardigrada from South Shetland Islands (Antarctica) with description of Angursa antarctica sp. nov. (Arthrotardigrada, Halechiniscidae). Polar Biology, 19, 336 - 341. https: // doi. org / 10.1007 / s 003000050255" type="journal article" year="1998">Víllora-Moreno, 1998</bibRefCitation>
</emphasis>
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<subSubSection id="C337650AFF8AFFE59B8DE742A9601131" pageId="18" pageNumber="19" type="description">
<paragraph id="8B923681FF8AFFE59B8DE742AE3D1004" blockId="18.[151,615,1075,1173]" pageId="18" pageNumber="19">
N=
<specimenCount id="9D2BFD08FF8AFFE59BA1E742AD091038" box="[187,339,1075,1101]" count="62" pageId="18" pageNumber="19" type="generic">62 specimens</specimenCount>
occurrence: ANDEEP I;
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bsl.
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE70AAE3E10E0" blockId="18.[151,615,1075,1173]" box="[199,612,1147,1173]" pageId="18" pageNumber="19">
<figureCitation id="13162A04FF8AFFE59BDDE70AAD1A10E0" box="[199,320,1147,1173]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9AB</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8AFFE59AE6E70AAE3E10E0" box="[508,612,1147,1173]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="18" pageNumber="19">Table 1.7</tableCitation>
</paragraph>
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<heading id="D0DA81EDFF8AFFE59B8DE7CCAD3D10A2" bold="true" box="[151,359,1213,1239]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B959EA93FF8AFFE59B8DE7CCAD3D10A2" bold="true" box="[151,359,1213,1239]" pageId="18" pageNumber="19">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE793A9601131" blockId="18.[151,1437,1213,1348]" pageId="18" pageNumber="19">
Small
<taxonomicName id="4C2D4D02FF8AFFE59A17E793AD361089" authorityName="Pollock" authorityYear="1979" box="[269,364,1250,1276]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF8AFFE59A17E793AD361089" box="[269,364,1250,1276]" italics="true" pageId="18" pageNumber="19">Angursa</emphasis>
</taxonomicName>
with slender body (106150 µm long, 1922 µm width, n=5) and primary clavae shorter than cirri
<emphasis id="B959EA93FF8AFFE59B8DE677ACF21155" box="[151,168,1286,1312]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. Individuals with long cephalic cirri and cirri
<emphasis id="B959EA93FF8AFFE599B1E676AEE61155" box="[683,700,1287,1312]" italics="true" pageId="18" pageNumber="19">E</emphasis>
. Secondary clavae are present (
<figureCitation id="13162A04FF8AFFE59F05E677A8C31155" box="[1055,1177,1286,1312]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9A, A</figureCitation>
<figureCitation id="13162A04FF8AFFE59F83E677A8F21155" box="[1177,1192,1286,1312]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="18" pageNumber="19">2</figureCitation>
). Short hemispherical sensory organs on legs IV present (
<figureCitation id="13162A04FF8AFFE5993AE65BAE201131" box="[544,634,1322,1348]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9A</figureCitation>
). Males have gonopore as in the original description (
<figureCitation id="13162A04FF8AFFE59FCEE65BA9771131" box="[1236,1325,1322,1348]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8AFFE59B8DE61CAE2E1186" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8B923681FF8AFFE59B8DE61CAD5E11F2" blockId="18.[151,1436,1389,1523]" box="[151,260,1389,1415]" pageId="18" pageNumber="19">
<emphasis id="B959EA93FF8AFFE59B8DE61CAD5E11F2" bold="true" box="[151,260,1389,1415]" pageId="18" pageNumber="19">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE6E0AE221186" blockId="18.[151,1436,1389,1523]" pageId="18" pageNumber="19">
<taxonomicName id="4C2D4D02FF8AFFE59BDDE6E0AD0311DF" authorityName="Villora-Moreno" authorityYear="1998" box="[199,345,1425,1451]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="species" species="antarctica">
<emphasis id="B959EA93FF8AFFE59BDDE6E0AD0311DF" box="[199,345,1425,1451]" italics="true" pageId="18" pageNumber="19">A. antarctica</emphasis>
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belongs to the
<taxonomicName id="4C2D4D02FF8AFFE5991DE6E0AEC711DF" authorityName="Renaud-Mornant" authorityYear="1981" box="[519,669,1424,1451]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B959EA93FF8AFFE5991DE6E0AEC711DF" box="[519,669,1424,1451]" italics="true" pageId="18" pageNumber="19">A. lanceolata</emphasis>
</taxonomicName>
group with primary clavae shorter than cirri
<emphasis id="B959EA93FF8AFFE59F80E6E0A8F111DE" box="[1178,1195,1425,1451]" italics="true" pageId="18" pageNumber="19">A</emphasis>
(
<bibRefCitation id="EFBC4B70FF8AFFE59FA6E6E1AC8B11BA" author="Fujimoto, S. &amp; Hansen, J. G." pageId="18" pageNumber="19" pagination="1 - 19" refId="ref19180" refString="Fujimoto, S. &amp; Hansen, J. G. (2019) Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, 510, 1 - 19. https: // doi. org / 10.5852 / ejt. 2019.510" type="journal article" year="2019">Fujimoto &amp; Hansen 2019</bibRefCitation>
). This species is recognisable by hemispherical leg IV sensory organs. Annulated scapi of cirri
<emphasis id="B959EA93FF8AFFE59E11E6C4A94611BB" box="[1291,1308,1461,1486]" italics="true" pageId="18" pageNumber="19">E</emphasis>
sometimes not clearly visible (
<bibRefCitation id="EFBC4B70FF8AFFE59A74E6A9AE311186" author="Villora-Moreno, S." box="[366,619,1496,1523]" pageId="18" pageNumber="19" pagination="336 - 341" refId="ref22898" refString="Villora-Moreno, S. (1998) Deep-sea Tardigrada from South Shetland Islands (Antarctica) with description of Angursa antarctica sp. nov. (Arthrotardigrada, Halechiniscidae). Polar Biology, 19, 336 - 341. https: // doi. org / 10.1007 / s 003000050255" type="journal article" year="1998">Víllora-Moreno, 1998</bibRefCitation>
).
</paragraph>
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</treatment>
</document>

View file

@ -1,108 +0,0 @@
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N=
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occurrence: ANDEEP I;
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bsl.
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; Supplementary
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<paragraph id="8B923681FF8AFFE59B8DE3E6AE5414C4" blockId="18.[151,526,151,178]" box="[151,526,151,178]" pageId="18" pageNumber="19">
<heading id="D0DA81EDFF8AFFE59B8DE3E6AE5414C4" bold="true" box="[151,526,151,178]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="B959EA93FF8AFFE59B8DE3E6AD3E14C7" bold="true" box="[151,356,151,178]" italics="true" pageId="18" pageNumber="19">Angursa capsula</emphasis>
<bibRefCitation id="EFBC4B70FF8AFFE59A71E3E6AE5414C4" author="Bussau, C." box="[363,526,151,177]" pageId="18" pageNumber="19" pagination="79 - 91" refId="ref18111" refString="Bussau, C. (1992) New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta, 21 (1), 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau, 1992</bibRefCitation>
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N=
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occurrence: ANDEEP I,
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,
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,
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;
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bsl
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.
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE253AE3F1549" blockId="18.[151,1006,218,316]" box="[199,613,290,316]" pageId="18" pageNumber="19">
<figureCitation id="13162A04FF8AFFE59BDDE253AD181549" box="[199,322,290,316]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="18" pageNumber="19">Fig. 8 EF</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8AFFE59AE7E253AE3F1549" box="[509,613,290,316]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="18" pageNumber="19">Table 1.6</tableCitation>
</paragraph>
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<emphasis id="B959EA93FF8AFFE59B8DE215AD3D150B" bold="true" box="[151,359,356,382]" pageId="18" pageNumber="19">Short description</emphasis>
</heading>
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<taxonomicName id="4C2D4D02FF8AFFE59BDDE2F8AD7C15D6" authorityName="Pollock" authorityYear="1979" box="[199,294,393,419]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF8AFFE59BDDE2F8AD7C15D6" box="[199,294,393,419]" italics="true" pageId="18" pageNumber="19">Angursa</emphasis>
</taxonomicName>
with a slender body (120176 µm long, 2030 µm width, n=5) and primary clavae longer than cirri
<emphasis id="B959EA93FF8AFFE59E9EE2F8A9CF15D6" box="[1412,1429,393,419]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. Distance between both secondary clavae on the dorsal side of the head is 6.311.0 μm (
<figureCitation id="13162A04FF8AFFE59F71E2DDA8BB15B2" box="[1131,1249,428,455]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="18" pageNumber="19">Fig. 8E, E</figureCitation>
<figureCitation id="13162A04FF8AFFE59FFBE2DCA8AA15B2" box="[1249,1264,429,455]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="18" pageNumber="19">2</figureCitation>
). Capsules-like sensory organs on legs IV (
<figureCitation id="13162A04FF8AFFE59AD2E2A1AE44159E" box="[456,542,464,491]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="18" pageNumber="19">Fig. 8F</figureCitation>
). Cirri
<emphasis id="B959EA93FF8AFFE59977E2A0AE24159F" box="[621,638,465,490]" italics="true" pageId="18" pageNumber="19">E</emphasis>
long (13.019.0 µm, n=5) (
<figureCitation id="13162A04FF8AFFE598ADE2A1A854159E" box="[951,1038,464,491]" captionStart="FIGURE 8" captionStartId="17.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@17.[151,1435,181,1750]" captionTargetPageId="17" captionText="FIGURE 8. Angursa bicuspis group. A Angursa abyssalis, male, light microscopy (DIC), ventral view. B female, light microscopy (DIC), head, dorsal view. B2 female, light microscopy (DIC), head, dorsal view, secondary clavae indicated. C male, light microscopy (DIC), dorsal view. D female, light microscopy (DIC), caudal region. E Angursa capsula, female, light microscopy (DIC), dorsal view. E2 Angursa capsula, female, light microscopy (DIC), dorsal view, secondary clavae indicated. F Angursa capsula, female, light microscopy (DIC), ventral view. Abbreviations: cE—cirrus E, ec—external cirrus, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385109" httpUri="https://zenodo.org/record/14385109/files/figure.png" pageId="18" pageNumber="19">Fig. 8E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8AFFE59B8DE162ADA616EC" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8B923681FF8AFFE59B8DE162AD5E1658" blockId="18.[151,1436,531,666]" box="[151,260,531,557]" pageId="18" pageNumber="19">
<emphasis id="B959EA93FF8AFFE59B8DE162AD5E1658" bold="true" box="[151,260,531,557]" pageId="18" pageNumber="19">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE146ADA616EC" blockId="18.[151,1436,531,666]" pageId="18" pageNumber="19">
One of the most difficult species to identify because of unclear borders of the secondary clavae. On the sagittal position of the specimen we mostly used the length of cirri
<emphasis id="B959EA93FF8AFFE59824E12DAF151600" box="[830,847,604,629]" italics="true" pageId="18" pageNumber="19">E</emphasis>
as a species-specific trait, which are longer than 11 µm (Hansen &amp;
<bibRefCitation id="EFBC4B70FF8AFFE59A5FE10EADAA16EC" author="Fujimoto, S. &amp; Hansen, J. G." box="[325,496,639,665]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref19180" refString="Fujimoto, S. &amp; Hansen, J. G. (2019) Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, 510, 1 - 19. https: // doi. org / 10.5852 / ejt. 2019.510" type="journal article" year="2019">Fujimoto 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,119 +0,0 @@
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N=
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; Supplementary
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<subSubSection id="C337650AFF8AFFE59B8DE50DA906131C" pageId="18" pageNumber="19" type="description">
<paragraph id="8B923681FF8AFFE59B8DE50DAF9712CF" blockId="18.[151,973,1660,1759]" pageId="18" pageNumber="19">
N=
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occurrence: ANDEEP I,
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,
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;
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bsl
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.
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE5B5AE1312AA" blockId="18.[151,973,1660,1759]" box="[199,585,1732,1759]" pageId="18" pageNumber="19">
<figureCitation id="13162A04FF8AFFE59BDDE5B5AD7C12AA" box="[199,294,1732,1759]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9 C</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8AFFE59AFBE5B5AE1312AA" box="[481,585,1732,1759]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="18" pageNumber="19">Table 1.8</tableCitation>
</paragraph>
<paragraph id="8B923681FF8AFFE59B8DE476AD3D1354" blockId="18.[151,1437,1799,1898]" box="[151,359,1799,1825]" pageId="18" pageNumber="19">
<heading id="D0DA81EDFF8AFFE59B8DE476AD3D1354" bold="true" box="[151,359,1799,1825]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B959EA93FF8AFFE59B8DE476AD3D1354" bold="true" box="[151,359,1799,1825]" pageId="18" pageNumber="19">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE45AA906131C" blockId="18.[151,1437,1799,1898]" pageId="18" pageNumber="19">
<taxonomicName id="4C2D4D02FF8AFFE59BDDE45AAD7C1330" authorityName="Pollock" authorityYear="1979" box="[199,294,1835,1861]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF8AFFE59BDDE45AAD7C1330" box="[199,294,1835,1861]" italics="true" pageId="18" pageNumber="19">Angursa</emphasis>
</taxonomicName>
with slender body (101150 µm long, 15.724.0 µm width, n=4) and primary clavae shorter than cirri
<emphasis id="B959EA93FF8AFFE59B8DE43EACF2131C" box="[151,168,1871,1897]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. Secondary and tertiary clavae present. Elongated leg IV sensory organs. Cirri
<emphasis id="B959EA93FF8AFFE59F04E421A875131C" box="[1054,1071,1872,1897]" italics="true" pageId="18" pageNumber="19">E</emphasis>
with lance-shaped ending.
</paragraph>
</subSubSection>
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<paragraph id="8B923681FF8AFFE59B8DE4E6AD5E13C4" blockId="18.[151,1436,1943,2042]" box="[151,260,1943,1969]" pageId="18" pageNumber="19">
<emphasis id="B959EA93FF8AFFE59B8DE4E6AD5E13C4" bold="true" box="[151,260,1943,1969]" pageId="18" pageNumber="19">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8AFFE59BDDE4CAA805138C" blockId="18.[151,1436,1943,2042]" pageId="18" pageNumber="19">
<taxonomicName id="4C2D4D02FF8AFFE59BDDE4CAAD0713A0" authorityName="Renaud-Mornant" authorityYear="1981" box="[199,349,1979,2005]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B959EA93FF8AFFE59BDDE4CAAD0713A0" box="[199,349,1979,2005]" italics="true" pageId="18" pageNumber="19">A. lanceolata</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C2D4D02FF8AFFE59910E4CAAEC513A0" authorityName="Renaud-Mornant" authorityYear="1981" box="[522,671,1979,2005]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="18" pageNumber="19" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B959EA93FF8AFFE59910E4CAAEC513A0" box="[522,671,1979,2005]" italics="true" pageId="18" pageNumber="19">A. lanceolata</emphasis>
</taxonomicName>
group with primary clavae shorter than cirri
<emphasis id="B959EA93FF8AFFE59F81E4CAA8F613A0" box="[1179,1196,1979,2005]" italics="true" pageId="18" pageNumber="19">A</emphasis>
(
<bibRefCitation id="EFBC4B70FF8AFFE59FA7E4CAAC8B138C" author="Fujimoto, S. &amp; Hansen, J. G." pageId="18" pageNumber="19" pagination="1 - 19" refId="ref19180" refString="Fujimoto, S. &amp; Hansen, J. G. (2019) Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, 510, 1 - 19. https: // doi. org / 10.5852 / ejt. 2019.510" type="journal article" year="2019">Fujimoto &amp; Hansen 2019</bibRefCitation>
). Species with easily recognisable lance-shaped cirri
<emphasis id="B959EA93FF8AFFE5983DE491AF62138C" box="[807,824,2016,2041]" italics="true" pageId="18" pageNumber="19">E</emphasis>
(
<bibRefCitation id="EFBC4B70FF8AFFE5985DE4AEA808138C" author="Renaud-Mornant, J." box="[839,1106,2015,2041]" pageId="18" pageNumber="19" pagination="161 - 164" refId="ref21762" refString="Renaud-Mornant, J. (1981) Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys, 10, 161 - 164." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,117 +0,0 @@
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N=
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occurrence: ANDEEP I,
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.
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<figureCitation id="13162A04FF8AFFE59BDDE5B5AD7C12AA" box="[199,294,1732,1759]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="18" pageNumber="19">Fig. 9 C</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8AFFE59AFBE5B5AE1312AA" box="[481,585,1732,1759]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="18" pageNumber="19">Table 1.8</tableCitation>
</paragraph>
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@ -0,0 +1,143 @@
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<taxonomicName id="4C2D4D02FF8CFFE39B8DE06EAE8D174C" ID-CoL="53C2G" authority="Kristensen &amp; Higgins, 1984" authorityName="Kristensen &amp; Higgins" authorityYear="1984" box="[151,727,799,825]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="qivitoq">
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<emphasis id="B959EA93FF8CFFE39B8DE06EAD2D174C" bold="true" box="[151,375,799,825]" italics="true" pageId="20" pageNumber="21">Styraconyx qivitoq</emphasis>
<bibRefCitation id="EFBC4B70FF8CFFE39A65E06EAE8D174C" author="Kristensen, R. M. &amp; Higgins, R. P." box="[383,727,799,825]" pageId="20" pageNumber="21" pagination="1 - 40" refId="ref20640" refString="Kristensen, R. M. &amp; Higgins, R. P. (1984) Revision of Styraconyx (Tardigrada: Halechiniscidae) with descriptions of two new species from Disko Bay, West Greenland. Smithsonian Contributions to Zoology, 391, 1 - 40. https: // doi. org / 10.5479 / si. 00810282.391" type="journal article" year="1984">Kristensen &amp; Higgins, 1984</bibRefCitation>
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<paragraph id="8B923681FF8CFFE39B8DE016AF4317D0" blockId="20.[151,793,871,969]" pageId="20" pageNumber="21">
N=
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:
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occurrence:
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;
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bsl
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.
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<paragraph id="8B923681FF8CFFE39BDDE0DEAED817BC" blockId="20.[151,793,871,969]" box="[199,642,943,969]" pageId="20" pageNumber="21">
<figureCitation id="13162A04FF8CFFE39BDDE0DEAD0B17BC" box="[199,337,943,969]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Figs 10AE</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8CFFE39917E0DEAED817BC" box="[525,642,943,969]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="20" pageNumber="21">Table 1.10</tableCitation>
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<emphasis id="B959EA93FF8CFFE39B8DE086AD3D1064" bold="true" box="[151,359,1015,1041]" pageId="20" pageNumber="21">Short description</emphasis>
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We observed only dwarf males (72108 µm long) unique for this species. Ovoid primary clavae and long cephalic cirri present (
<figureCitation id="13162A04FF8CFFE39A8DE74EAE58102C" box="[407,514,1087,1113]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10A</figureCitation>
). Secondary clavae are present (
<figureCitation id="13162A04FF8CFFE3986EE74EAF87102C" box="[884,989,1087,1113]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10B</figureCitation>
). Neck region with folded cuticle (
<figureCitation id="13162A04FF8CFFE39E75E74EAC901008" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10C</figureCitation>
). All digits are covered with a cuticular hood. Peduncles are large (
<figureCitation id="13162A04FF8CFFE398B2E712A8551008" box="[936,1039,1123,1149]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10D</figureCitation>
). Inconspicuous tubular-shape male gonopore is present (
<figureCitation id="13162A04FF8CFFE39A98E7F6ADBD10D4" box="[386,487,1159,1185]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8CFFE39B8DE7BEAECD1144" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B923681FF8CFFE39B8DE7BEAD5E109C" blockId="20.[151,1436,1231,1330]" box="[151,260,1231,1257]" pageId="20" pageNumber="21">
<emphasis id="B959EA93FF8CFFE39B8DE7BEAD5E109C" bold="true" box="[151,260,1231,1257]" pageId="20" pageNumber="21">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8CFFE39BDDE782AECD1144" blockId="20.[151,1436,1231,1330]" pageId="20" pageNumber="21">
All the described characters are species-specific and correspond well to the original description (
<bibRefCitation id="EFBC4B70FF8CFFE39E1FE782AD601144" author="Kristensen, R. M. &amp; Higgins, R. P." pageId="20" pageNumber="21" pagination="1 - 40" refId="ref20640" refString="Kristensen, R. M. &amp; Higgins, R. P. (1984) Revision of Styraconyx (Tardigrada: Halechiniscidae) with descriptions of two new species from Disko Bay, West Greenland. Smithsonian Contributions to Zoology, 391, 1 - 40. https: // doi. org / 10.5479 / si. 00810282.391" type="journal article" year="1984">Kristensen &amp; Higgins, 1984</bibRefCitation>
), so the identification is sound.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,117 +0,0 @@
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N=
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occurrence:
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; Supplementary
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<heading id="D0DA81EDFF8CFFE39B8DE3E6AE5A14C4" bold="true" box="[151,512,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C2D4D02FF8CFFE39B8DE3E6AE5A14C4" ID-CoL="5V2M5" authority="Bussau, 1992" authorityName="Bussau" authorityYear="1992" box="[151,512,151,178]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="lingua">
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<emphasis id="B959EA93FF8CFFE39B8DE3E6AD0F14C7" bold="true" box="[151,341,151,178]" italics="true" pageId="20" pageNumber="21">Angursa lingua</emphasis>
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N=
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occurrence: ANDEEP I,
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,
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;
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bsl
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.
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<figureCitation id="13162A04FF8CFFE39BDDE256AD191537" box="[199,323,295,322]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="20" pageNumber="21">Figs 9D, E</figureCitation>
; Supplementary
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<emphasis id="B959EA93FF8CFFE39B8DE21EAD3D15FC" bold="true" box="[151,359,367,393]" pageId="20" pageNumber="21">Short description</emphasis>
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<taxonomicName id="4C2D4D02FF8CFFE39BDDE2E2AD7C15D8" authorityName="Pollock" authorityYear="1979" box="[199,294,403,429]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF8CFFE39BDDE2E2AD7C15D8" box="[199,294,403,429]" italics="true" pageId="20" pageNumber="21">Angursa</emphasis>
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with slender body (137172 µm long, n=5) and primary clavae shorter than cirri
<emphasis id="B959EA93FF8CFFE39F87E2E2A8F415D8" box="[1181,1198,403,429]" italics="true" pageId="20" pageNumber="21">A</emphasis>
(
<figureCitation id="13162A04FF8CFFE39FA6E2E2A94E15D8" box="[1212,1300,403,429]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="20" pageNumber="21">Fig. 9D</figureCitation>
). Secondary and tertiary clavae present. Elongated leg IV sensory organs. Cirri
<emphasis id="B959EA93FF8CFFE39898E2C9AFC915A4" box="[898,915,440,465]" italics="true" pageId="20" pageNumber="21">E</emphasis>
with tongue-shaped ending (
<figureCitation id="13162A04FF8CFFE39FC3E2C6A96A15A4" box="[1241,1328,439,465]" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="20" pageNumber="21">Fig. 9E</figureCitation>
).
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<emphasis id="B959EA93FF8CFFE39B8DE28EAD5E166C" bold="true" box="[151,260,511,537]" pageId="20" pageNumber="21">Remarks</emphasis>
</paragraph>
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<taxonomicName id="4C2D4D02FF8CFFE39BDDE152AD7D1648" authorityName="Bussau" authorityYear="1992" box="[199,295,547,573]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="lingua">
<emphasis id="B959EA93FF8CFFE39BDDE152AD7D1648" box="[199,295,547,573]" italics="true" pageId="20" pageNumber="21">A.lingua</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C2D4D02FF8CFFE39AD4E152AE381648" authorityName="Renaud-Mornant" authorityYear="1981" box="[462,610,547,573]" class="Heterotardigrada" family="Halechiniscidae" genus="Angursa" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B959EA93FF8CFFE39AD4E152AE381648" box="[462,610,547,573]" italics="true" pageId="20" pageNumber="21">A. lanceolata</emphasis>
</taxonomicName>
group with primary clavae shorter than cirri
<emphasis id="B959EA93FF8CFFE39F4BE152A8381648" box="[1105,1122,547,573]" italics="true" pageId="20" pageNumber="21">A</emphasis>
(
<bibRefCitation id="EFBC4B70FF8CFFE39F68E152A9D41648" author="Fujimoto, S. &amp; Hansen, J. G." box="[1138,1422,547,574]" pageId="20" pageNumber="21" pagination="1 - 19" refId="ref19180" refString="Fujimoto, S. &amp; Hansen, J. G. (2019) Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, 510, 1 - 19. https: // doi. org / 10.5852 / ejt. 2019.510" type="journal article" year="2019">Fujimoto &amp; Hansen 2019</bibRefCitation>
). Species with easily recognisable traits, such as tongue-shaped cirri
<emphasis id="B959EA93FF8CFFE3989FE139AFCC1614" box="[901,918,584,609]" italics="true" pageId="20" pageNumber="21">E</emphasis>
and elongated sensory organs on legs IV (
<figureCitation id="13162A04FF8CFFE39E75E136ACE216F0" captionStart="FIGURE 9" captionStartId="19.[151,250,1773,1797]" captionTargetBox="[154,1432,184,1747]" captionTargetId="figure-17@19.[151,1435,181,1750]" captionTargetPageId="19" captionText="FIGURE 9. Angursa lanceolata group. A Angursa antarctica, male, light microscopy (DIC), dorsal view. A2 Angursa antarctica, male, light microscopy (DIC), dorsal view, secondary clavae indicated. B Angursa antarctica, male, light microscopy (DIC), ventral view, note male gonopore (arrowhead). C Angursa lanceolata, female, light microscopy (DIC), ventral view. D Angursa lingua, female, light microscopy (DIC), dorsal view. E Angursa lingua, light microscopy (DIC), sagittal view, note cE (arrowhead). Abbreviations: cA—cirrus A, cE—cirrus E, pb—pharyngeal bulb, pc—primary clava, sc—secondary clava, se—leg sensory organ, tc—tertiary clava." figureDoi="http://doi.org/10.5281/zenodo.14385111" httpUri="https://zenodo.org/record/14385111/files/figure.png" pageId="20" pageNumber="21">Fig. 9E</figureCitation>
) (
<bibRefCitation id="EFBC4B70FF8CFFE39BD5E11AAD3116F0" author="Bussau, C." box="[207,363,619,645]" pageId="20" pageNumber="21" pagination="79 - 91" refId="ref18111" refString="Bussau, C. (1992) New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta, 21 (1), 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau, 1992</bibRefCitation>
).
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<subSubSection id="C337650AFF8CFFE39B8DE6F2AFAD11E8" box="[151,1015,1411,1438]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B923681FF8CFFE39B8DE6F2AFAD11E8" blockId="20.[151,1015,1411,1438]" box="[151,1015,1411,1438]" pageId="20" pageNumber="21">
<heading id="D0DA81EDFF8CFFE39B8DE6F2AFAD11E8" bold="true" box="[151,1015,1411,1438]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C2D4D02FF8CFFE39B8DE6F2AFAD11E8" authority="Fujimoto, Suzuki, Ito, Tamura &amp; Tsujimoto, 2020" authorityName="Fujimoto, Suzuki, Ito, Tamura &amp; Tsujimoto" authorityYear="2020" box="[151,1015,1411,1438]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="takeshii">
<emphasis id="B959EA93FF8CFFE39B8DE6F2AFAD11E8" bold="true" box="[151,1015,1411,1438]" pageId="20" pageNumber="21">
<emphasis id="B959EA93FF8CFFE39B8DE6F2ADDF11EB" bold="true" box="[151,389,1411,1438]" italics="true" pageId="20" pageNumber="21">Styraconyx takeshii</emphasis>
<bibRefCitation id="EFBC4B70FF8CFFE39A96E6F2AFAD11E8" author="Fujimoto, S. &amp; Suzuki, A. C. &amp; Ito, M. &amp; Tamura, T. &amp; Tsujimoto, M." box="[396,1015,1411,1437]" pageId="20" pageNumber="21" pagination="679 - 693" refId="ref19235" refString="Fujimoto, S., Suzuki, A. C., Ito, M., Tamura, T. &amp; Tsujimoto, M. (2020) Marine tardigrades from Lutzow-Holm Bay, East Antarctica with the description of a new species. Polar Biology, 43, 679 - 693. https: // doi. org / 10.1007 / s 00300 - 020 - 02671 - w" type="journal article" year="2020">Fujimoto, Suzuki, Ito, Tamura &amp; Tsujimoto, 2020</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8CFFE39B8DE6BAADB2135C" pageId="20" pageNumber="21" type="description">
<paragraph id="8B923681FF8CFFE39B8DE6BAAE2A127C" blockId="20.[151,629,1483,1581]" pageId="20" pageNumber="21">
N=
<specimenCount id="9D2BFD08FF8CFFE39BDAE6BAAD641190" box="[192,318,1483,1509]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
: female occurrence: ANDEEP
<collectionCode id="ED3CAE44FF8CFFE39ADEE681AD80127F" box="[452,474,1520,1546]" pageId="20" pageNumber="21">II</collectionCode>
;
<elevation id="0000D1B2FF8CFFE39AFFE69EAE36127C" box="[485,620,1519,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="20" pageNumber="21" unit="m" value="1109.0">
<quantity id="4CD59B64FF8CFFE39AFFE69EAE1B127F" box="[485,577,1519,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="20" pageNumber="21" unit="m" value="1109.0">1,109 m</quantity>
bsl
</elevation>
.
</paragraph>
<paragraph id="8B923681FF8CFFE39BDDE562AE2F1258" blockId="20.[151,629,1483,1581]" box="[199,629,1555,1581]" pageId="20" pageNumber="21">
<figureCitation id="13162A04FF8CFFE39BDDE562AD1E1258" box="[199,324,1555,1581]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Figs 10FI</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8CFFE3991AE562AE2F1258" box="[512,629,1555,1581]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="20" pageNumber="21">Table 1.11</tableCitation>
</paragraph>
<paragraph id="8B923681FF8CFFE39B8DE52AAD3D1200" blockId="20.[151,1437,1627,1833]" box="[151,359,1627,1653]" pageId="20" pageNumber="21">
<heading id="D0DA81EDFF8CFFE39B8DE52AAD3D1200" bold="true" box="[151,359,1627,1653]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B959EA93FF8CFFE39B8DE52AAD3D1200" bold="true" box="[151,359,1627,1653]" pageId="20" pageNumber="21">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF8CFFE39BDDE50EADB2135C" blockId="20.[151,1437,1627,1833]" pageId="20" pageNumber="21">
Adult female (145 µm long, n=1) with the typical character combination of this species. Dorsal cuticle undulated (
<figureCitation id="13162A04FF8CFFE39A0DE5D2AD2712C8" box="[279,381,1699,1725]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10F</figureCitation>
). Primary clavae elongated (
<figureCitation id="13162A04FF8CFFE399DEE5D2AF7012C8" box="[708,810,1699,1725]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10F</figureCitation>
). Large secondary clavae are present (
<figureCitation id="13162A04FF8CFFE39FFBE5D2A91612C8" box="[1249,1356,1699,1725]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10G</figureCitation>
). Head cirri are divided into three regions (
<figureCitation id="13162A04FF8CFFE39939E5B6AED61294" box="[547,652,1735,1761]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10G</figureCitation>
). Thick seminal receptacle duct openings with no terminal swelling as lateral projections (
<figureCitation id="13162A04FF8CFFE39A6BE59AAD801370" box="[369,474,1771,1797]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10H</figureCitation>
). Digits with proximal pads and small peduncles (
<figureCitation id="13162A04FF8CFFE39F17E59AA8301370" box="[1037,1130,1771,1797]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10I</figureCitation>
). Leg IV sensory organs as elongated papillae (
<figureCitation id="13162A04FF8CFFE39A69E47EAD86135C" box="[371,476,1807,1833]" captionStart="FIGURE 10" captionStartId="21.[151,250,1711,1735]" captionTargetBox="[180,1407,184,1684]" captionTargetId="figure-17@21.[177,1410,181,1687]" captionTargetPageId="21" captionText="FIGURE 10. Genus Styraconyx. A Styraconyx qivitoq, male, light microscopy (DIC), dorsal view. B Styraconyx qivitoq, male, light microscopy (DIC), head, dorsal view, note secondary clava (arrowhead). C Styraconyx qivitoq, male, light microscopy (DIC), head, ventral view, note neck folds (arrowhead). D Styraconyx qivitoq, male, light microscopy (DIC), leg III, note digit covered with a cuticle hood (arrowhead). E Styraconyx qivitoq, male, light microscopy (DIC), caudal region, note male gonopore (arrowhead). F Styraconyx takeshii, female, light microscopy (DIC), sagittal view, note undulated dorsal cuticle (arrowhead). G Styraconyx takeshii, female, light microscopy (DIC), sagittal view. H Styraconyx takeshii, female, light microscopy (DIC), sagittal view. I Styraconyx takeshii, female, light microscopy (DIC), sagittal view. Abbreviations: cA—cirrus A, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mc—median cirrus, pc—primary clava, pe—peduncle, pp—proximal pad, se—leg sensory organ, sro—seminal receptacle opening." figureDoi="http://doi.org/10.5281/zenodo.14385113" httpUri="https://zenodo.org/record/14385113/files/figure.png" pageId="20" pageNumber="21">Fig. 10H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8CFFE39B8DE426A88213CC" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B923681FF8CFFE39B8DE426AD5E1304" blockId="20.[151,1436,1879,1977]" box="[151,260,1879,1905]" pageId="20" pageNumber="21">
<emphasis id="B959EA93FF8CFFE39B8DE426AD5E1304" bold="true" box="[151,260,1879,1905]" pageId="20" pageNumber="21">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8CFFE39BDDE40AA88213CC" blockId="20.[151,1436,1879,1977]" pageId="20" pageNumber="21">
<taxonomicName id="4C2D4D02FF8CFFE39BDDE40AADF813E0" authorityName="Fujimoto, Suzuki, Ito, Tamura &amp; Tsujimoto" authorityYear="2020" box="[199,418,1915,1941]" class="Heterotardigrada" family="Halechiniscidae" genus="Styraconyx" kingdom="Animalia" order="Arthrotardigrada" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="takeshii">
<emphasis id="B959EA93FF8CFFE39BDDE40AADF813E0" box="[199,418,1915,1941]" italics="true" pageId="20" pageNumber="21">Styraconyx takeshii</emphasis>
</taxonomicName>
is well recognisable by the shape of dorsal cuticle with transversal ridges, elongated form of primary clavae, small peduncles and robust seminal receptacle opening (
<bibRefCitation id="EFBC4B70FF8CFFE398FBE4EEA89613CC" author="Fujimoto, S. &amp; Suzuki, A. C. &amp; Ito, M. &amp; Tamura, T. &amp; Tsujimoto, M." box="[993,1228,1951,1977]" pageId="20" pageNumber="21" pagination="679 - 693" refId="ref19235" refString="Fujimoto, S., Suzuki, A. C., Ito, M., Tamura, T. &amp; Tsujimoto, M. (2020) Marine tardigrades from Lutzow-Holm Bay, East Antarctica with the description of a new species. Polar Biology, 43, 679 - 693. https: // doi. org / 10.1007 / s 00300 - 020 - 02671 - w" type="journal article" year="2020">
Fujimoto
<emphasis id="B959EA93FF8CFFE39F4AE4D1A8D313CC" box="[1104,1161,1951,1977]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,115 +0,0 @@
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<mods:title id="D1B2E29A05807E63308851495C0A99C9">Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean</mods:title>
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<mods:affiliation id="946B578ABD6040D6F03B98E8C44E2E98">VN Karazin Kharkiv National University, Svobody Square, 4, 61022, Kharkiv, Ukraine. &amp; German Centre for Marine Biodiversity Research (DZMB), Senckenberg am Meer, Südstrand 44, 26382, Wilhelmshaven, Germany.</mods:affiliation>
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<mods:namePart id="64B287E6FC21CB8F914DCC20A095EFDD">Schmidt-Rhaesa, Andreas</mods:namePart>
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<mods:affiliation id="8BED505B4C385AB77B7D6C92735AE7A9">ZTM, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<subSubSection id="C337650AFF8EFFE19B8DE256AE481614" pageId="22" pageNumber="23" type="description">
<paragraph id="8B923681FF8EFFE19B8DE256AFF91510" blockId="22.[151,931,295,393]" pageId="22" pageNumber="23">
N=
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:
<specimenCount id="9D2BFD08FF8EFFE19A4EE256ADE21534" box="[340,440,295,321]" count="1" pageId="22" pageNumber="23" type="female">1 female</specimenCount>
,
<specimenCount id="9D2BFD08FF8EFFE19AD9E256AE461534" box="[451,540,295,321]" count="4" pageId="22" pageNumber="23" type="male">4 males</specimenCount>
, 3 with undetermined sex occurrence: ANDEEP I,
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;
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bsl
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.
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<paragraph id="8B923681FF8EFFE19BDDE21EAE1015FC" blockId="22.[151,931,295,393]" box="[199,586,367,393]" pageId="22" pageNumber="23">
<figureCitation id="13162A04FF8EFFE19BDDE21EAD4315FC" box="[199,281,367,393]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF8EFFE19ACEE21EAE1015FC" box="[468,586,367,393]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="22" pageNumber="23">Table 1.12</tableCitation>
</paragraph>
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<heading id="D0DA81EDFF8EFFE19B8DE2C6AD3D15A4" bold="true" box="[151,359,439,465]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B959EA93FF8EFFE19B8DE2C6AD3D15A4" bold="true" box="[151,359,439,465]" pageId="22" pageNumber="23">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF8EFFE19BDDE2AAAE481614" blockId="22.[151,1436,439,610]" pageId="22" pageNumber="23">
Tardigrades with slender body and bell-shaped outer epicuticle (
<figureCitation id="13162A04FF8EFFE19885E2AAA8731580" box="[927,1065,475,501]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11A, C</figureCitation>
). Short primary clavae and large secondary clavae present (
<figureCitation id="13162A04FF8EFFE19ADAE28EAE7D166C" box="[448,551,511,537]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11A</figureCitation>
). Sexual dimorphism concerning body size (female is 336 µm long and 79 µm wide, n=1; males are 165173 µm long and 40 µm wide, n=3) (
<figureCitation id="13162A04FF8EFFE1987AE152AF981648" box="[864,962,547,573]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11A</figureCitation>
compared with
<figureCitation id="13162A04FF8EFFE19F62E152A8861648" box="[1144,1244,547,573]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11C</figureCitation>
). Sensory organs on each leg present (
<figureCitation id="13162A04FF8EFFE19A64E136AE5C1614" box="[382,518,583,609]" captionStart="FIGURE 11" captionStartId="23.[151,250,1773,1797]" captionTargetBox="[154,1432,241,1747]" captionTargetId="figure-17@23.[151,1435,181,1750]" captionTargetPageId="23" captionText="FIGURE 11. Tholoarctus oleseni. A Tholoarctus oleseni, female, light microscopy (DIC), dorsal view. B Tholoarctus oleseni, female, light microscopy (DIC), leg II (right) and leg III (left) region, dorsal view. C Tholoarctus oleseni, male, light microscopy (DIC), ventral view. D Tholoarctus oleseni, male, SEM, ventro-sagittal view. D2 Tholoarctus oleseni, male, SEM, leg III, note sensory organ (arrowhead).Abbreviations: an—anus, cA—cirrus A, cE—cirrus E, ec—external cirrus, ed—external digit, go— gonopore, ic—internal cirrus, id—internal digit, mc—median cirrus, oe—outer epicuticle, pc—primary clava, pi—cuticular pilars, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385117" httpUri="https://zenodo.org/record/14385117/files/figure.png" pageId="22" pageNumber="23">Fig. 11B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF8EFFE19B8DE1FEACB3174C" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8B923681FF8EFFE19B8DE1FEAD5E16DC" blockId="22.[151,1437,655,825]" box="[151,260,655,681]" pageId="22" pageNumber="23">
<emphasis id="B959EA93FF8EFFE19B8DE1FEAD5E16DC" bold="true" box="[151,260,655,681]" pageId="22" pageNumber="23">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF8EFFE19BDDE1C2ACB3174C" blockId="22.[151,1437,655,825]" pageId="22" pageNumber="23">
Tardigrades with clear sexual dimorphism in size (mature females are twice as big as males). Short spine-like sensory organ present also on the legs II and
<collectionCode id="ED3CAE44FF8EFFE19980E1A9AEE31687" box="[666,697,728,754]" pageId="22" pageNumber="23">III</collectionCode>
. Outer epicuticle with various levels of inflation so we measured length and width by inner cuticle borders, which corresponded to the original description by
<bibRefCitation id="EFBC4B70FF8EFFE19FFFE18DACBF174C" author="Jorgensen, A. &amp; Boesgaard, T. M. &amp; Mobjerg, N. &amp; Kristensen, R. M." pageId="22" pageNumber="23" pagination="401 - 433" refId="ref20236" refString="Jorgensen, A., Boesgaard, T. M., Mobjerg, N. &amp; Kristensen, R. M. (2014) The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada. Zootaxa, 3802 (4), 401 - 433. https: // doi. org / 10.1078 / 0044 - 5231 - 00033" type="journal article" year="2014">
Jørgensen
<emphasis id="B959EA93FF8EFFE19E7AE18DA9C71760" box="[1376,1437,763,789]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,119 +0,0 @@
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<emphasis id="B959EA93FF8EFFE19B8DE3AEADDC148F" bold="true" box="[151,390,223,250]" italics="true" pageId="22" pageNumber="23">Tholoarctus oleseni</emphasis>
Jørgensen, Boesgaard, Møbjerg, &amp; Kristensen, 2014
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N=
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@ -1,84 +1,84 @@
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<paragraph id="8B923681FF91FFFE9B8DE58CA8481362" blockId="9.[151,1042,1789,1816]" box="[151,1042,1789,1816]" pageId="9" pageNumber="10">
<heading id="D0DA81EDFF91FFFE9B8DE58CA8481362" bold="true" box="[151,1042,1789,1816]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="4C2D4D02FF91FFFE9B8DE58CA8481362" authority="Bartels, Fontoura, Nelson &amp; Kaczmarek, 2024" authorityName="Bartels, Fontoura, Nelson &amp; Kaczmarek" authorityYear="2024" box="[151,1042,1789,1816]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="species" species="wyedeleinorum">
@ -89,6 +89,8 @@
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF91FFFE9B8DE434ADCC13D2" pageId="9" pageNumber="10" type="multiple">
<paragraph id="8B923681FF91FFFE9B8DE434ADCC13D2" blockId="9.[151,1436,1861,2032]" pageId="9" pageNumber="10">
N=
<specimenCount id="9D2BFD08FF91FFFE9BA6E434AD13132A" box="[188,329,1861,1887]" count="2" pageId="9" pageNumber="10" type="generic">2 specimens</specimenCount>
@ -178,7 +180,8 @@ and external cirri all originate from a common lateral bulge of the head (
<figureCitation id="13162A04FF92FFFD9802E049AF281726" box="[792,882,824,851]" captionStart="FIGURE 5" captionStartId="12.[151,250,1680,1704]" captionTargetBox="[161,1427,184,1654]" captionTargetId="figure-19@12.[158,1430,181,1657]" captionTargetPageId="12" captionText="FIGURE 5. Batillipes wyedeleinorum. A female, light microscopy (DIC), ventro-sagittal view, note leg III digit arrangement (arrowhead). B female, light microscopy (DIC), ventro-sagittal view. C female, light microscopy (DIC), ventro-sagittal view, note body projections (arrowheads). D female, light microscopy (DIC), caudal region, note leg IV digits arrangement. D2 female, light microscopy (DIC), leg IV sensory organ, note tuft of additional filaments (arrowhead). E female, light microscopy (DIC), caudal region, note smooth area (arrowhead). F Batillipes wyedeleinorum, juvenile, light microscopy (DIC), sagittal view, note body projections (arrowheads). G female, SEM, ventro-sagittal view, note cirrus A tip (arrowhead). H female, SEM, ventral view, cephalic region, note cephalic cirri tips (arrowheads). I female, SEM, ventral view, leg II region, note folded digits (arrowhead). J female, SEM, caudal region, ventral view, note cuticular pillars (arrowhead). Abbreviations: an—anus, ca—caudal appendage, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mo—mouth opening, lp—leg projection, pb—pharyngeal bulb, pc—primary clava, pr—body projection, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385097" httpUri="https://zenodo.org/record/14385097/files/figure.png" pageId="10" pageNumber="11">Fig. 5B</figureCitation>
). Sensory organs of legs I and II are about 7 µm long, of legs III to 10 µm. Sensory organs of legs IV with a cirrophore are 18 µm long and have frayed tips (
<figureCitation id="13162A04FF92FFFD9E70E02DAC8417EF" captionStart="FIGURE 5" captionStartId="12.[151,250,1680,1704]" captionTargetBox="[161,1427,184,1654]" captionTargetId="figure-19@12.[158,1430,181,1657]" captionTargetPageId="12" captionText="FIGURE 5. Batillipes wyedeleinorum. A female, light microscopy (DIC), ventro-sagittal view, note leg III digit arrangement (arrowhead). B female, light microscopy (DIC), ventro-sagittal view. C female, light microscopy (DIC), ventro-sagittal view, note body projections (arrowheads). D female, light microscopy (DIC), caudal region, note leg IV digits arrangement. D2 female, light microscopy (DIC), leg IV sensory organ, note tuft of additional filaments (arrowhead). E female, light microscopy (DIC), caudal region, note smooth area (arrowhead). F Batillipes wyedeleinorum, juvenile, light microscopy (DIC), sagittal view, note body projections (arrowheads). G female, SEM, ventro-sagittal view, note cirrus A tip (arrowhead). H female, SEM, ventral view, cephalic region, note cephalic cirri tips (arrowheads). I female, SEM, ventral view, leg II region, note folded digits (arrowhead). J female, SEM, caudal region, ventral view, note cuticular pillars (arrowhead). Abbreviations: an—anus, ca—caudal appendage, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mo—mouth opening, lp—leg projection, pb—pharyngeal bulb, pc—primary clava, pr—body projection, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385097" httpUri="https://zenodo.org/record/14385097/files/figure.png" pageId="10" pageNumber="11">Figs 5D, D</figureCitation>
2). Bases of legs I and legs II have a small (1.52.0 µm) dome-shaped projection. On legs I, digits 2 (4.0 µm) and 4 (5.5 µm) are the shortest (considering digit 1 the most cephalically), digits 3 (10.5 µm) and 5 (11.0 µm) are the longest and digits 1 (6.0 µm) and 6 (7.0 µm) are medium sized. The same pattern of digits on legs II and III (
<figureCitation id="13162A04FF92FFFD9BC4E0F1ACB717EF" box="[222,237,896,922]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="10" pageNumber="11">2</figureCitation>
). Bases of legs I and legs II have a small (1.52.0 µm) dome-shaped projection. On legs I, digits 2 (4.0 µm) and 4 (5.5 µm) are the shortest (considering digit 1 the most cephalically), digits 3 (10.5 µm) and 5 (11.0 µm) are the longest and digits 1 (6.0 µm) and 6 (7.0 µm) are medium sized. The same pattern of digits on legs II and III (
<figureCitation id="13162A04FF92FFFD9E75E0B9ACE71072" captionStart="FIGURE 5" captionStartId="12.[151,250,1680,1704]" captionTargetBox="[161,1427,184,1654]" captionTargetId="figure-19@12.[158,1430,181,1657]" captionTargetPageId="12" captionText="FIGURE 5. Batillipes wyedeleinorum. A female, light microscopy (DIC), ventro-sagittal view, note leg III digit arrangement (arrowhead). B female, light microscopy (DIC), ventro-sagittal view. C female, light microscopy (DIC), ventro-sagittal view, note body projections (arrowheads). D female, light microscopy (DIC), caudal region, note leg IV digits arrangement. D2 female, light microscopy (DIC), leg IV sensory organ, note tuft of additional filaments (arrowhead). E female, light microscopy (DIC), caudal region, note smooth area (arrowhead). F Batillipes wyedeleinorum, juvenile, light microscopy (DIC), sagittal view, note body projections (arrowheads). G female, SEM, ventro-sagittal view, note cirrus A tip (arrowhead). H female, SEM, ventral view, cephalic region, note cephalic cirri tips (arrowheads). I female, SEM, ventral view, leg II region, note folded digits (arrowhead). J female, SEM, caudal region, ventral view, note cuticular pillars (arrowhead). Abbreviations: an—anus, ca—caudal appendage, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mo—mouth opening, lp—leg projection, pb—pharyngeal bulb, pc—primary clava, pr—body projection, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385097" httpUri="https://zenodo.org/record/14385097/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). On legs IV, digits 3 (6.0 µm) and 4 (8.0 µm) are the shortest, and digits 1 (11.0 µm), 2 (14.5 µm), 5 (14.0 µm), 6 (11.0 µm) are the longest (
<figureCitation id="13162A04FF92FFFD9AC6E761AE6D105E" box="[476,567,1040,1067]" captionStart="FIGURE 5" captionStartId="12.[151,250,1680,1704]" captionTargetBox="[161,1427,184,1654]" captionTargetId="figure-19@12.[158,1430,181,1657]" captionTargetPageId="12" captionText="FIGURE 5. Batillipes wyedeleinorum. A female, light microscopy (DIC), ventro-sagittal view, note leg III digit arrangement (arrowhead). B female, light microscopy (DIC), ventro-sagittal view. C female, light microscopy (DIC), ventro-sagittal view, note body projections (arrowheads). D female, light microscopy (DIC), caudal region, note leg IV digits arrangement. D2 female, light microscopy (DIC), leg IV sensory organ, note tuft of additional filaments (arrowhead). E female, light microscopy (DIC), caudal region, note smooth area (arrowhead). F Batillipes wyedeleinorum, juvenile, light microscopy (DIC), sagittal view, note body projections (arrowheads). G female, SEM, ventro-sagittal view, note cirrus A tip (arrowhead). H female, SEM, ventral view, cephalic region, note cephalic cirri tips (arrowheads). I female, SEM, ventral view, leg II region, note folded digits (arrowhead). J female, SEM, caudal region, ventral view, note cuticular pillars (arrowhead). Abbreviations: an—anus, ca—caudal appendage, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mo—mouth opening, lp—leg projection, pb—pharyngeal bulb, pc—primary clava, pr—body projection, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385097" httpUri="https://zenodo.org/record/14385097/files/figure.png" pageId="10" pageNumber="11">Fig. 5D</figureCitation>
@ -224,5 +227,51 @@ Bartels
</paragraph>
<paragraph id="8B923681FF92FFFD9BDDE6EDA9DB11C3" blockId="10.[151,1437,1256,1463]" box="[199,1409,1436,1463]" pageId="10" pageNumber="11">A possible ecological difference is evidenced by our new geographic and bathymetric record of this species.</paragraph>
</subSubSection>
<subSection id="E2A22F6AFF92FFFD9B8DE579A9151383" pageId="10" pageNumber="11" type="multiple">
<subSubSection id="C337650AFF92FFFD9B8DE579AE6B128F" pageId="10" pageNumber="11" type="description">
<paragraph id="8B923681FF92FFFD9B8DE579ADA11257" blockId="10.[151,527,1544,1607]" box="[151,507,1544,1570]" pageId="10" pageNumber="11">
<heading id="D0DA81EDFF92FFFD9B8DE579ADA11257" bold="true" box="[151,507,1544,1570]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B959EA93FF92FFFD9B8DE579ADA11257" bold="true" box="[151,507,1544,1570]" pageId="10" pageNumber="11">Short description of juvenile</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF92FFFD9B8DE55DAE551233" blockId="10.[151,527,1544,1607]" box="[151,527,1580,1607]" pageId="10" pageNumber="11">
<figureCitation id="13162A04FF92FFFD9B8DE55DACB11232" box="[151,235,1580,1607]" captionStart="FIGURE 5" captionStartId="12.[151,250,1680,1704]" captionTargetBox="[161,1427,184,1654]" captionTargetId="figure-19@12.[158,1430,181,1657]" captionTargetPageId="12" captionText="FIGURE 5. Batillipes wyedeleinorum. A female, light microscopy (DIC), ventro-sagittal view, note leg III digit arrangement (arrowhead). B female, light microscopy (DIC), ventro-sagittal view. C female, light microscopy (DIC), ventro-sagittal view, note body projections (arrowheads). D female, light microscopy (DIC), caudal region, note leg IV digits arrangement. D2 female, light microscopy (DIC), leg IV sensory organ, note tuft of additional filaments (arrowhead). E female, light microscopy (DIC), caudal region, note smooth area (arrowhead). F Batillipes wyedeleinorum, juvenile, light microscopy (DIC), sagittal view, note body projections (arrowheads). G female, SEM, ventro-sagittal view, note cirrus A tip (arrowhead). H female, SEM, ventral view, cephalic region, note cephalic cirri tips (arrowheads). I female, SEM, ventral view, leg II region, note folded digits (arrowhead). J female, SEM, caudal region, ventral view, note cuticular pillars (arrowhead). Abbreviations: an—anus, ca—caudal appendage, cE—cirrus E, ec—external cirrus, go—gonopore, ic—internal cirrus, mo—mouth opening, lp—leg projection, pb—pharyngeal bulb, pc—primary clava, pr—body projection, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385097" httpUri="https://zenodo.org/record/14385097/files/figure.png" pageId="10" pageNumber="11">Fig. 5F</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF92FFFD9ABDE55DAE551233" box="[423,527,1580,1606]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="10" pageNumber="11">Table 1.1</tableCitation>
</paragraph>
<paragraph id="8B923681FF92FFFD9B8DE505AE6B128F" blockId="10.[151,1436,1652,1786]" pageId="10" pageNumber="11">
Four-toed juvenile 123.5 µm long with punctuated cuticle. Cephalic cirri with lance-shaped tips. Each leg with a sensory organ. Lateral projections on the neck region, between leg II and leg
<collectionCode id="ED3CAE44FF92FFFD98F1E5E8A85D12C6" box="[1003,1031,1689,1715]" pageId="10" pageNumber="11">III</collectionCode>
are conical; projections between leg I and leg II, as well as between leg
<collectionCode id="ED3CAE44FF92FFFD9907E5CCAE6312A2" box="[541,569,1725,1751]" pageId="10" pageNumber="11">III</collectionCode>
and leg IV are blunt. On each leg digit 2 is shorter than the other. Short single conical caudal appendage presented.
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF92FFFD9B8DE459A9161383" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B923681FF92FFFD9B8DE459AD5E1337" blockId="10.[151,1437,1832,2039]" box="[151,260,1832,1858]" pageId="10" pageNumber="11">
<emphasis id="B959EA93FF92FFFD9B8DE459AD5E1337" bold="true" box="[151,260,1832,1858]" pageId="10" pageNumber="11">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF92FFFD9BDDE43CA9151383" blockId="10.[151,1437,1832,2039]" pageId="10" pageNumber="11">
We identified this specimen as
<taxonomicName id="4C2D4D02FF92FFFD992DE43DAF021313" box="[567,856,1868,1894]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="wyedeleinorum">
<emphasis id="B959EA93FF92FFFD992DE43DAF021313" box="[567,856,1868,1894]" italics="true" pageId="10" pageNumber="11">Batillipes wyedeleinorum</emphasis>
</taxonomicName>
because of overall morphological similarity. This individual has comparably shorter cirri
<emphasis id="B959EA93FF92FFFD9955E400AE3A13FF" box="[591,608,1905,1930]" italics="true" pageId="10" pageNumber="11">E</emphasis>
, a sensory organ on the leg IV and more elongated suction disks on digits. This kind of minor dissimilarities between mature and juvenile stages are known for other species of
<taxonomicName id="4C2D4D02FF92FFFD9E2BE4E5AE5813A7" authority="(Kristensen &amp; Mackness 2000)" baseAuthorityName="Kristensen &amp; Mackness" baseAuthorityYear="2000" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF92FFFD9E2BE4E5A9C613DB" box="[1329,1436,1940,1966]" italics="true" pageId="10" pageNumber="11">Batillipes</emphasis>
(
<bibRefCitation id="EFBC4B70FF92FFFD9B85E4C9ADA013A7" author="Kristensen, R. M. &amp; Mackness, B. S." box="[159,506,1976,2002]" pageId="10" pageNumber="11" pagination="73 - 87" refId="ref20700" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen &amp; Mackness 2000</bibRefCitation>
)
</taxonomicName>
. Furthermore, both specimens occur in a similar habitat, the Antarctic deep sea bottom, which is a rather unusual and unique habitat for species of the genus
<taxonomicName id="4C2D4D02FF92FFFD98E3E4ADA83E1383" baseAuthorityName="Kristensen &amp; Mackness" baseAuthorityYear="2000" box="[1017,1124,2012,2038]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="genus">
<emphasis id="B959EA93FF92FFFD98E3E4ADA83E1383" box="[1017,1124,2012,2038]" italics="true" pageId="10" pageNumber="11">Batillipes</emphasis>
</taxonomicName>
(but see discussion).
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>

View file

@ -1,121 +0,0 @@
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<mods:title id="8067A87B9720AA8F767216066BEF2EA1">Towards a better understanding of deep-sea tardigrade biogeography: numerous new records from the Southern Ocean</mods:title>
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; Supplementary
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View file

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<figureCitation id="13162A04FF95FFFA9BDDE74EAD51102C" box="[199,267,1087,1113]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF95FFFA9ADDE74EAE75102C" box="[455,559,1087,1113]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="13" pageNumber="14">Table 1.3</tableCitation>
</paragraph>
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<emphasis id="B959EA93FF95FFFA9B8DE7F6AD3D10D4" bold="true" box="[151,359,1159,1185]" pageId="13" pageNumber="14">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF95FFFA9BDDE7DDA8D01120" blockId="13.[151,1436,1159,1510]" pageId="13" pageNumber="14">
Large tardigrades (205294 µm, n=2) of elongated body, with coelomocytes and other spherical cells. Secondary clavae have clearly visible ventral side shape as in original description (
<figureCitation id="13162A04FF95FFFA98AFE7BEA875109C" box="[949,1071,1231,1257]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6G, G</figureCitation>
<figureCitation id="13162A04FF95FFFA9F35E7BEA864109C" box="[1071,1086,1231,1257]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="13" pageNumber="14">2</figureCitation>
). On the dorsal side the median cirrus is placed between the secondary clavae (
<figureCitation id="13162A04FF95FFFA99A9E782AF511178" box="[691,779,1267,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6F</figureCitation>
). Claws without heteromorphy. Accessory spines only on the internal claws of leg IV (
<figureCitation id="13162A04FF95FFFA9AFBE666AE671144" box="[481,573,1303,1329]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6H</figureCitation>
). Juvenile (205 µm) with internal claws and sometimes with external claws developing inside the femur (
<figureCitation id="13162A04FF95FFFA9AFAE64AAE751120" box="[480,559,1339,1365]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6I</figureCitation>
). The female has a gonopore and seminal receptacles.
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF95FFFA9B8DE6F2AFEC1190" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B923681FF95FFFA9B8DE6F2AD5E11E8" blockId="13.[151,1436,1159,1510]" box="[151,260,1411,1437]" pageId="13" pageNumber="14">
<emphasis id="B959EA93FF95FFFA9B8DE6F2AD5E11E8" bold="true" box="[151,260,1411,1437]" pageId="13" pageNumber="14">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF95FFFA9BDDE6D9AFE01190" blockId="13.[151,1436,1159,1510]" pageId="13" pageNumber="14">
<taxonomicName id="4C2D4D02FF95FFFA9BDDE6D9AD6511B4" authorityName="Renaud-Mornant" authorityYear="1974" box="[199,319,1447,1474]" class="Heterotardigrada" family="Coronarctidae" genus="Coronarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="tenellus">
<emphasis id="B959EA93FF95FFFA9BDDE6D9AD6511B4" box="[199,319,1447,1474]" italics="true" pageId="13" pageNumber="14">C. tenellus</emphasis>
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is recognisable by the secondary clavae shape and median cirrus position. All representatives have accessory spines on each leg IV internal claw (
<bibRefCitation id="EFBC4B70FF95FFFA99B8E6BAAFF41190" author="Renaud-Mornant, J." box="[674,942,1483,1509]" pageId="13" pageNumber="14" pagination="3087 - 3090" refId="ref21670" refString="Renaud-Mornant, J. (1974) Une nouvelle famille de Tardigrades marins abyssaux: les Coronarctidae fam. nov. (Heterotardigrada). Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences, Paris, 278, 3087 - 3090." type="journal article" year="1974">Renaud-Mornant, 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B959EA93FF95FFFA9B8DE256ADFC1537" bold="true" box="[151,422,295,322]" italics="true" pageId="13" pageNumber="14">Coronarctus dissimilis</emphasis>
<bibRefCitation id="EFBC4B70FF95FFFA9AB4E257A8D71534" author="Gomes-Junior, E. &amp; Santos, E. &amp; da Rocha, C. M. &amp; Santos, P. J. &amp; Fontoura, P." box="[430,1165,294,321]" pageId="13" pageNumber="14" refId="ref19620" refString="Gomes-Junior, E., Santos, E., da Rocha, C. M., Santos, P. J. &amp; Fontoura, P. (2020) The deep-sea genus Coronarctus (Tardigrada, Arthrotardigrada) in Brazil, south-western Atlantic Ocean, with the description of three new species. Diversity, 12 (2), 63. https: // doi. org / 10.3390 / d 12020063" type="journal volume" year="2020">Gomes-Júnior, Santos, da Rocha, Santos &amp; Fontoura, 2020</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B923681FF95FFFA9B8DE21EAE9615D8" blockId="13.[151,716,367,465]" pageId="13" pageNumber="14">
N=
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: female occurrence:
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;
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bsl
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.
</paragraph>
<paragraph id="8B923681FF95FFFA9BDDE2C6AE7515A4" blockId="13.[151,716,367,465]" box="[199,559,439,465]" pageId="13" pageNumber="14">
<figureCitation id="13162A04FF95FFFA9BDDE2C6AD5115A4" box="[199,267,439,465]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF95FFFA9ADDE2C6AE7515A4" box="[455,559,439,465]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="13" pageNumber="14">Table 1.2</tableCitation>
</paragraph>
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<heading id="D0DA81EDFF95FFFA9B8DE28EAD3D166C" bold="true" box="[151,359,511,537]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B959EA93FF95FFFA9B8DE28EAD3D166C" bold="true" box="[151,359,511,537]" pageId="13" pageNumber="14">Short description</emphasis>
</heading>
</paragraph>
<paragraph id="8B923681FF95FFFA9BDDE155A96816DC" blockId="13.[151,1437,511,682]" pageId="13" pageNumber="14">
Large female (273 µm, n=1) with claw heteromorphy (
<figureCitation id="13162A04FF95FFFA9822E152AFCE1648" box="[824,916,547,573]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6A</figureCitation>
). On the dorsal side a median cirrus is placed close between the posterior edges of the secondary clavae (
<figureCitation id="13162A04FF95FFFA9859E136AFC41614" box="[835,926,583,609]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6B</figureCitation>
). Edges of secondary clavae are also clearly visible on the ventral side (
<figureCitation id="13162A04FF95FFFA9ADBE11AAE6D16F0" box="[449,567,619,645]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6C, C</figureCitation>
<figureCitation id="13162A04FF95FFFA992DE11AAE1C16F0" box="[567,582,619,645]" captionStart="FIGURE 2" captionStartId="8.[151,250,1863,1887]" captionTargetBox="[183,1395,226,1814]" captionTargetId="graphics-20@8.[186,1402,183,1838]" captionText="FIGURE 2. Absolute abundances of Tardigrada from all 130 analysed core samples of expeditions ANDEEP-1 to 3 and ANDEEP-SYSTCO to the Southern Ocean plotted against depth. Samples were taken at different working areas in the Cape Basin, in the Lazarev Sea, in the Weddell Sea, in the Ona Basin, and at different working stations in the Drake Passage" pageId="13" pageNumber="14">2</figureCitation>
). Both external and internal claws are with accessory spines on each leg (
<figureCitation id="13162A04FF95FFFA9E75E11AAC8416DC" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6B, D</figureCitation>
). The presence of gonopore and seminal receptacles allows us to be secure with sex determination.
</paragraph>
</subSubSection>
<subSubSection id="C337650AFF95FFFA9B8DE1A6ADB21728" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B923681FF95FFFA9B8DE1A6AD5E1684" blockId="13.[151,1436,727,861]" box="[151,260,727,753]" pageId="13" pageNumber="14">
<emphasis id="B959EA93FF95FFFA9B8DE1A6AD5E1684" bold="true" box="[151,260,727,753]" pageId="13" pageNumber="14">Remarks</emphasis>
</paragraph>
<paragraph id="8B923681FF95FFFA9BDDE18DADB61728" blockId="13.[151,1436,727,861]" pageId="13" pageNumber="14">
<taxonomicName id="4C2D4D02FF95FFFA9BDDE18DAD0E1760" authorityName="Gomes-Junior, Santos, da Rocha, Santos &amp; Fontoura" authorityYear="2020" box="[199,340,763,790]" class="Heterotardigrada" family="Coronarctidae" genus="Coronarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dissimilis">
<emphasis id="B959EA93FF95FFFA9BDDE18DAD0E1760" box="[199,340,763,790]" italics="true" pageId="13" pageNumber="14">C. dissimilis</emphasis>
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is recognisable by the secondary clavae shape and median cirrus placed on the posterior end of secondary clavae. This representative exhibits claw heteromorphy and accessory spines as in the original description of
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Gomes-Júnior
<emphasis id="B959EA93FF95FFFA9A41E035ADC91728" box="[347,403,835,861]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2020)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,115 +0,0 @@
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<mods:identifier id="BA7D6992DCBEBF90CCB1051F7C2E2C26" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8132F0C45217078C7B3418B55DDC83E8" type="Zenodo-Dep">14385085</mods:identifier>
<mods:identifier id="6684B7DDBB3AAD264FAA0CE9726B3768" type="ZooBank">F6372401-5C79-487C-A8C1-DBDEE7C71671</mods:identifier>
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<treatment id="03848797FF95FFFA9B8DE256AE7515A4" ID-DOI="http://doi.org/10.5281/zenodo.14385174" ID-Zenodo-Dep="14385174" LSID="urn:lsid:plazi:treatment:03848797FF95FFFA9B8DE256AE7515A4" httpUri="http://treatment.plazi.org/id/03848797FF95FFFA9B8DE256AE7515A4" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="C337650AFF95FFFA9B8DE256AE9615D8" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B923681FF95FFFA9B8DE256A8D71534" blockId="13.[151,1165,294,322]" box="[151,1165,294,322]" pageId="13" pageNumber="14">
<heading id="D0DA81EDFF95FFFA9B8DE256A8D71534" bold="true" box="[151,1165,294,322]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="4C2D4D02FF95FFFA9B8DE256A8D71534" ID-CoL="9X6RK" authority="Gomes-Junior, Santos, da Rocha, Santos &amp; Fontoura, 2020" authorityName="Gomes-Junior, Santos, da Rocha, Santos &amp; Fontoura" authorityYear="2020" box="[151,1165,294,322]" class="Heterotardigrada" family="Coronarctidae" genus="Coronarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dissimilis">
<emphasis id="B959EA93FF95FFFA9B8DE256A8D71534" bold="true" box="[151,1165,294,322]" pageId="13" pageNumber="14">
<emphasis id="B959EA93FF95FFFA9B8DE256ADFC1537" bold="true" box="[151,422,295,322]" italics="true" pageId="13" pageNumber="14">Coronarctus dissimilis</emphasis>
<bibRefCitation id="EFBC4B70FF95FFFA9AB4E257A8D71534" author="Gomes-Junior, E. &amp; Santos, E. &amp; da Rocha, C. M. &amp; Santos, P. J. &amp; Fontoura, P." box="[430,1165,294,321]" pageId="13" pageNumber="14" refId="ref19620" refString="Gomes-Junior, E., Santos, E., da Rocha, C. M., Santos, P. J. &amp; Fontoura, P. (2020) The deep-sea genus Coronarctus (Tardigrada, Arthrotardigrada) in Brazil, south-western Atlantic Ocean, with the description of three new species. Diversity, 12 (2), 63. https: // doi. org / 10.3390 / d 12020063" type="journal volume" year="2020">Gomes-Júnior, Santos, da Rocha, Santos &amp; Fontoura, 2020</bibRefCitation>
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</paragraph>
<paragraph id="8B923681FF95FFFA9B8DE21EAE9615D8" blockId="13.[151,716,367,465]" pageId="13" pageNumber="14">
N=
<materialsCitation id="3B453CDCFF95FFFA9BDAE21EAE9215D8" collectionCode="ANDEEP-SYSTCO" elevation="2981" pageId="13" pageNumber="14" specimenCount="1">
<specimenCount id="9D2BFD08FF95FFFA9BDAE21EAD6415FC" box="[192,318,367,393]" count="1" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
: female occurrence:
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;
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bsl
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.
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<subSubSection id="C337650AFF95FFFA9BDDE2C6AE7515A4" box="[199,559,439,465]" pageId="13" pageNumber="14" type="description">
<paragraph id="8B923681FF95FFFA9BDDE2C6AE7515A4" blockId="13.[151,716,367,465]" box="[199,559,439,465]" pageId="13" pageNumber="14">
<figureCitation id="13162A04FF95FFFA9BDDE2C6AD5115A4" box="[199,267,439,465]" captionStart="FIGURE 6" captionStartId="14.[151,250,1648,1672]" captionTargetBox="[206,1382,185,1622]" captionTargetId="figure-19@14.[203,1385,182,1625]" captionTargetPageId="14" captionText="FIGURE 6. Genus Coronarctus. A Coronarctus dissimilis, female, light microscopy (DIC), ventro-sagittal view. B Coronarctus dissimilis, female, light microscopy (DIC), head, dorsal view, note leg I accessory spines on the external and internal claws (arrowheads). C Coronarctus dissimilis, female, light microscopy (DIC), head ventral view. C2 Coronarctus dissimilis, female, light microscopy (DIC), head ventral view, secondary clava indicated. D Coronarctus dissimilis, female, light microscopy (DIC), leg IV, note leg IV accessory spines on the both external and internal claws (arrowheads). E Coronarctus tenellus, male, light microscopy (DIC), ventral position. F Coronarctus tenellus, male, light microscopy (DIC), head, dorsal view. G Coronarctus tenellus, male, light microscopy (DIC), head, ventral view. G2 Coronarctus tenellus, male, light microscopy (DIC), head, ventral view, secondary clava indicated. H Coronarctus tenellus, light microscopy (DIC), leg IV, note leg IV accessory spines just on the both internal claw (arrowhead). I Coronarctus tenellus, juvenile, light microscopy (DIC), sagittal position, note leg IV with two internal claws and two growing external claws (arrowhead).Abbreviations: bt—buccal tube, cE—cirrus E, mc—median cirrus, pb—pharyngeal bulb, sc—secondary clava, se—leg sensory organ." figureDoi="http://doi.org/10.5281/zenodo.14385101" httpUri="https://zenodo.org/record/14385101/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
; Supplementary
<tableCitation id="C6AF033AFF95FFFA9ADDE2C6AE7515A4" box="[455,559,439,465]" captionStart="TABLE 1" captionStartId="3.[152,244,469,495]" captionText="TABLE 1. All corer deployments of expeditions ANDEEP 1, 2, 3 and ANDEEP-SYSTCO that contained Tardigrada. For two deployments at the Shona Ridge, no information about position and depth was available. GKG: boxcorer with 0.25 m2 sampling area, MUC (A): multicorer with 70.85 cm2 sampling area per core tube, MUC (B): multicorer with 69.36 cm2 sampling area per core tube, MUC (C): multicorer with 25.5 cm2 sampling area per core tube" pageId="13" pageNumber="14">Table 1.2</tableCitation>
</paragraph>
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