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<document id="809AD3DB97A943FEE1B009BA2FD57638" ID-DOI="10.11646/zootaxa.5538.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14611794" ID-ZooBank="AA6CF84D-0E8A-41AA-AF37-E900092B6067" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1736288160019" checkinUser="plazi" docAuthor="Köhler, Gunther, Sameit, Joachim, Seipp, Robert & Geiss, Katharina" docDate="2024" docId="03F387F20008FF8CAEEAFA44FD7D2C99" docLanguage="en" docName="zootaxa.5538.4.1.pdf" docOrigin="Zootaxa 5538 (4)" docSource="https://doi.org/10.11646/zootaxa.5538.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhacodactylus willihenkeli Köhler, Sameit, Seipp & Geiss, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="309" masterDocId="FFCAFF8A000BFF85AE7DFFDFFFCC2C4C" masterDocTitle="A new species of giant gecko of the genus Rhacodactylus from New Caledonia (Squamata, Gekkota, Diplodactylidae)" masterLastPageNumber="321" masterPageNumber="301" pageNumber="304" updateTime="1736427175776" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B4DE90BAF7CC394B97A49E45298C99BE">
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<mods:title id="45D2F79DDF8848849B8C7486519852EB">A new species of giant gecko of the genus Rhacodactylus from New Caledonia (Squamata, Gekkota, Diplodactylidae)</mods:title>
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</mods:titleInfo>
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</mods:role>
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<mods:namePart id="879F256C4D600CFAFA1B1A6181D70EC4">Köhler, Gunther</mods:namePart>
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<mods:nameIdentifier id="A161135E695FC4F2B58FE69F9BA9E2E2" type="ORCID">0000-0002-2563-5331</mods:nameIdentifier>
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<mods:affiliation id="7B8EE015062F5612EBEB84F8BCC2077A">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany</mods:affiliation>
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<mods:nameIdentifier id="B3F4046D3100E993F9FF9685051DFD78" type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
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<mods:roleTerm id="4AF9612B1231B0E3CF2336B33373D007">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1B8023233009CAC78B67E1B08C0A3AA7">Sameit, Joachim</mods:namePart>
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<mods:nameIdentifier id="FB6382E2926353DC34A68AC7CE6E6428" type="ORCID">0009-0004-1922-3906</mods:nameIdentifier>
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<mods:affiliation id="B335D0CFBFFEB112826B9E603B020A0F">Alfred-Döblin-Str. 68, 59192 Bergkamen, Germany</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="8EC578008DBEF089887134EE7C9399FC">Seipp, Robert</mods:namePart>
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<mods:nameIdentifier id="7EA78B6E7CCA9F493CB6C257F93745D7" type="ORCID">0009-0009-2755-2714</mods:nameIdentifier>
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<mods:affiliation id="37B7586F5C65013C8BD754980DE80809">Alt Praunheim 45, 60488 Frankfurt a. M., Germany</mods:affiliation>
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</mods:role>
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<mods:namePart id="821355A48EED9B01E7E0727C51E26FA4">Geiss, Katharina</mods:namePart>
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<mods:nameIdentifier id="DC7DBB68056BC118B8D688115C8C8F94" type="ORCID">0000-0003-3180-352X</mods:nameIdentifier>
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<mods:affiliation id="1F67C421327FFAD94F17D38436374D2C">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany</mods:affiliation>
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<mods:nameIdentifier id="A1C6AC1FC00DDD96B335DC93D2C9157A" type="email">kgeiss@senckenberg.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="EE591FDD6585A614EFBF19051543A35A">text</mods:typeOfResource>
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<mods:title id="37CB78842654E6D3C522BFEEEE800FF0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DDFB9094E8750984583C3F62AFEDDE0E">
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<mods:date id="7210CFF7DD7DFACBB43CD95606BE5186">2024</mods:date>
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<mods:detail id="A8C47840711A02CC86CCCF6BFE4A75C8" type="pubDate">
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<mods:number id="11EBF6B2DE6EE9F23F399F9F5159AB89">2024-11-18</mods:number>
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<mods:detail id="AA1137499D092F95ED4835E9E4E0D483" type="volume">
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<mods:number id="C8905BCBAA43C9FBCC5E43D505CE7B55">5538</mods:number>
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<mods:number id="39DB70FD3F68D2A38D25CF5CC6ED66DB">4</mods:number>
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<mods:start id="388E607A9E90668BCD693CF703642833">301</mods:start>
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<mods:location id="E3A5297DC25443E2AFDF2FCFCF2DAA49">
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<mods:url id="57D0251E6D0BEF4AB4E82C7939A62983">https://doi.org/10.11646/zootaxa.5538.4.1</mods:url>
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</mods:location>
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<mods:classification id="EDA13887FA7679EB30C9AD6836E5E399">journal article</mods:classification>
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<mods:identifier id="285E42BA9C004C5E34290402E9CC0491" type="DOI">10.11646/zootaxa.5538.4.1</mods:identifier>
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|
<mods:identifier id="96DBD055DBC6004D87FB7F854E001C00" type="ISSN">1175-5326</mods:identifier>
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|
<mods:identifier id="DDDED248950E42E28D32B75C83195506" type="Zenodo-Dep">14611794</mods:identifier>
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<mods:identifier id="23DED88935AE0D3F4313230B78E48AA5" type="ZooBank">AA6CF84D-0E8A-41AA-AF37-E900092B6067</mods:identifier>
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</mods:mods>
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<treatment id="03F387F20008FF8CAEEAFA44FD7D2C99" LSID="urn:lsid:plazi:treatment:03F387F20008FF8CAEEAFA44FD7D2C99" httpUri="http://treatment.plazi.org/id/03F387F20008FF8CAEEAFA44FD7D2C99" lastPageId="9" lastPageNumber="309" pageId="3" pageNumber="304">
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<subSubSection id="C340656F0008FF86AEEAFA44FD8F29FA" box="[151,579,1435,1462]" pageId="3" pageNumber="304" type="nomenclature">
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||||||
|
<paragraph id="8BE536E40008FF86AEEAFA44FD8F29FA" blockId="3.[151,579,1435,1462]" box="[151,579,1435,1462]" pageId="3" pageNumber="304">
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||||||
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<heading id="D0AD81880008FF86AEEAFA44FD8F29FA" bold="true" box="[151,579,1435,1462]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
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||||||
|
<emphasis id="B92EEAF60008FF86AEEAFA44FD8F29FA" bold="true" box="[151,579,1435,1462]" pageId="3" pageNumber="304">
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||||||
|
<taxonomicName id="4C5A4D670008FF86AEEAFA44FE1229FA" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[151,478,1435,1462]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="willihenkeli" status="sp. nov.">
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||||||
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<emphasis id="B92EEAF60008FF86AEEAFA44FE1229FA" bold="true" box="[151,478,1435,1462]" italics="true" pageId="3" pageNumber="304">Rhacodactylus willihenkeli</emphasis>
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||||||
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</taxonomicName>
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||||||
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<taxonomicNameLabel id="A21D578D0008FF86AF98FA43FD8F29FA" box="[485,579,1436,1462]" pageId="3" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||||||
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</heading>
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||||||
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</paragraph>
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||||||
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</subSubSection>
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||||||
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<subSubSection id="C340656F0008FF86AEEAFA3FFCE529B5" box="[151,809,1504,1529]" pageId="3" pageNumber="304" type="reference_group">
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||||||
|
<paragraph id="8BE536E40008FF86AEEAFA3FFCE529B5" blockId="3.[151,809,1504,1565]" box="[151,809,1504,1529]" pageId="3" pageNumber="304">
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||||||
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<taxonomicName id="4C5A4D670008FF86AEEAFA3FFD9529B5" authority="(Houailou, N)" baseAuthorityName="Houailou, N" box="[151,601,1504,1529]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="leachianus">
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<emphasis id="B92EEAF60008FF86AEEAFA3FFEE029B4" box="[151,300,1504,1528]" italics="true" pageId="3" pageNumber="304">Rhacodactylus</emphasis>
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cf.
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<emphasis id="B92EEAF60008FF86AF2EFA3FFE0D29B4" box="[339,449,1504,1528]" italics="true" pageId="3" pageNumber="304">leachianus</emphasis>
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(Houailou, N)
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</taxonomicName>
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:
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<bibRefCitation id="EFCB4B150008FF86AC1FFA3EFCE529B5" author="Vences, M. & Henkel, F. - W. & Seipp, R." box="[610,809,1504,1529]" pageId="3" pageNumber="304" pagination="73 - 82" refId="ref9378" refString="Vences, M., Henkel, F. - W. & Seipp, R. (2001) Molekulare Untersuchungen zur Phylogenie und Taxonomie der Neukaledonischen Geckos der Gattung Rhacodactylus (Reptilia: Gekkonidae). Salamandra, 37, 73 - 82." type="journal article" year="2001">
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||||||
|
VENCES
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||||||
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<emphasis id="B92EEAF60008FF86ACCAFA3EFD2829B4" box="[695,740,1504,1528]" italics="true" pageId="3" pageNumber="304">et al</emphasis>
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||||||
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., 2001
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</bibRefCitation>
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||||||
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</paragraph>
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||||||
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</subSubSection>
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||||||
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<subSubSection id="C340656F0008FF86AEEAF9DCFED72A51" box="[151,283,1538,1565]" pageId="3" pageNumber="304" type="description">
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||||||
|
<paragraph id="8BE536E40008FF86AEEAF9DCFED72A51" blockId="3.[151,809,1504,1565]" box="[151,283,1538,1565]" pageId="3" pageNumber="304">
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||||||
|
<figureCitation id="13612A610008FF86AEEAF9DCFF292A51" box="[151,229,1539,1565]" captionStart="FIGURE 2" captionStartId="5.[152,255,1668,1692]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure-20@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 2. Female holotype (SMF 105137, upper individual) and adult male (not preserved,lower individual) of Rhacodactylus willihenkeli sp. nov. in life. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611798" httpUri="https://zenodo.org/record/14611798/files/figure.png" pageId="3" pageNumber="304">Figs. 2</figureCitation>
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,
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<figureCitation id="13612A610008FF86AE8CF9DDFED72A51" box="[241,283,1538,1565]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="7.[152,255,1940,1964]" captionStartId-1="8.[152,255,1066,1090]" captionTargetBox-0="[151,1436,181,1915]" captionTargetBox-1="[151,1436,181,1042]" captionTargetId-0="figure-20@7.[151,1436,181,1916]" captionTargetId-1="figure-21@8.[151,1436,181,1042]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 4. Holotype of Rhacodactylus willihenkeli sp. nov. (SMF 105137). Scale bars equal 40.0 mm in A to D, 10.0 mm in E to H. Photographs by G.K." captionText-1="FIGURE 5. Holotype of Rhacodactylus willihenkeli sp. nov. (SMF 105137). Scale bars equal 10.0 mm. Photographs by G.K." figureDoi-0="http://doi.org/10.5281/zenodo.14611802" figureDoi-1="http://doi.org/10.5281/zenodo.14611804" httpUri-0="https://zenodo.org/record/14611802/files/figure.png" httpUri-1="https://zenodo.org/record/14611804/files/figure.png" pageId="3" pageNumber="304">4–5</figureCitation>
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</paragraph>
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||||||
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</subSubSection>
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||||||
|
<subSubSection id="C340656F0008FF86AEEAF994FAD62B55" pageId="3" pageNumber="304" type="materials_examined">
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||||||
|
<paragraph id="8BE536E40008FF86AEEAF994FBB72AE1" blockId="3.[151,1437,1610,2033]" pageId="3" pageNumber="304">
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||||||
|
<materialsCitation id="3B323CB90008FF86AEEAF994FBB72AE1" collectingDate="1987-09" collectionCode="SMF" collectorName="Friedrich Wilhelm Henkel & Joachim Sameit" country="New Caledonia" elevation="750" location="Bourail" pageId="3" pageNumber="304" specimenCode="SMF 105137" specimenCount="1" typeStatus="holotype">
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||||||
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<emphasis id="B92EEAF60008FF86AEEAF994FEC82A29" bold="true" box="[151,260,1611,1637]" pageId="3" pageNumber="304">
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||||||
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<typeStatus id="54E188460008FF86AEEAF994FEC82A29" box="[151,260,1611,1637]" pageId="3" pageNumber="304" type="holotype">Holotype</typeStatus>
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||||||
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</emphasis>
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||||||
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.
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<specimenCode id="DBFC9E9F0008FF86AF69F994FE602A29" box="[276,428,1610,1637]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="304">SMF 105137</specimenCode>
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, an adult
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<specimenCount id="9D5CFD6D0008FF86AC60F994FDA52A29" box="[541,617,1611,1637]" pageId="3" pageNumber="304" type="female">female</specimenCount>
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from about
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<locationDeviation id="9A572FC60008FF86AC84F995FCE62A29" box="[761,810,1610,1637]" pageId="3" pageNumber="304">12.5</locationDeviation>
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airline
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<locationDeviation id="9A572FC60008FF86ADFEF994FC192A29" box="[899,981,1611,1637]" pageId="3" pageNumber="304">km NE</locationDeviation>
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|
<location id="8E85603F0008FF86ADA3F994FBFE2A29" LSID="urn:lsid:plazi:treatment:03F387F20008FF8CAEEAFA44FD7D2C99:8E85603F0008FF86ADA3F994FBFE2A29" box="[990,1074,1611,1637]" country="New Caledonia" name="Bourail" pageId="3" pageNumber="304">Bourail</location>
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||||||
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(GPS coordinates ca. -21.517°, 165.606°; about
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|
<quantity id="4CA29B010008FF86AF28F9B1FE522AC5" box="[341,414,1646,1673]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="304" unit="m" value="750.0">
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||||||
|
<elevation id="0077D1D70008FF86AF28F9B1FE522AC5" box="[341,414,1646,1673]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="304" unit="m" value="750.0">750 m</elevation>
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</quantity>
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||||||
|
above sea level; see remarks in Geographic Distribution and Conservation), central
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<collectingCountry id="F34D76740008FF86AB1AF9B0FEC72AE1" name="New Caledonia" pageId="3" pageNumber="304">New Caledonia</collectingCountry>
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||||||
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, collected
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||||||
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<date id="FFE410240008FF86AFFEF94CFDF62AE1" box="[387,570,1683,1709]" pageId="3" pageNumber="304" value="1987-09">
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||||||
|
<collectingDate id="EFA0E9CC0008FF86AFFEF94CFDF62AE1" box="[387,570,1683,1709]" pageId="3" pageNumber="304" value="1987-09">September 1987</collectingDate>
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||||||
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</date>
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||||||
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by
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||||||
|
<collectorName id="26AF53320008FF86AC19F94CFC452AE1" box="[612,905,1683,1709]" pageId="3" pageNumber="304">Friedrich Wilhelm Henkel</collectorName>
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||||||
|
und
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||||||
|
<collectorName id="26AF53320008FF86ADBCF94CFBBB2AE1" box="[961,1143,1683,1709]" pageId="3" pageNumber="304">Joachim Sameit</collectorName>
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||||||
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.
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</materialsCitation>
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||||||
|
</paragraph>
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||||||
|
<paragraph id="8BE536E40008FF86AEBAF968FAD62B55" blockId="3.[151,1437,1610,2033]" pageId="3" pageNumber="304">
|
||||||
|
<materialsCitation id="3B323CB90008FF86AEBAF968FD3D2A9D" box="[199,753,1718,1745]" collectingDate="1987-09" collectionCode="SMF" collectorName="Friedrich Wilhelm Henkel & Joachim Sameit" country="New Caledonia" elevation="750" location="Bourail" pageId="3" pageNumber="304" specimenCode="SMF 106563" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
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||||||
|
<emphasis id="B92EEAF60008FF86AEBAF968FE8A2A9D" bold="true" box="[199,326,1719,1745]" pageId="3" pageNumber="304">
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||||||
|
<typeStatus id="54E188460008FF86AEBAF968FE8D2A9D" box="[199,321,1719,1745]" pageId="3" pageNumber="304" type="paratype">Paratypes</typeStatus>
|
||||||
|
.
|
||||||
|
</emphasis>
|
||||||
|
One
|
||||||
|
<specimenCount id="9D5CFD6D0008FF86AFF1F968FDCA2A9D" box="[396,518,1719,1745]" pageId="3" pageNumber="304" type="adult">adult male</specimenCount>
|
||||||
|
(
|
||||||
|
<specimenCode id="DBFC9E9F0008FF86AC67F968FD7A2A9D" box="[538,694,1718,1745]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="304">SMF 106563</specimenCode>
|
||||||
|
) and
|
||||||
|
</materialsCitation>
|
||||||
|
<materialsCitation id="3B323CB90008FF86AC80F968FDA42B55" collectingDate="1987-09" collectionCode="SMF" collectorName="NE Bourail" country="New Caledonia" elevation="750" location="Friedrich Wilhelm Henkel" pageId="3" pageNumber="304" specimenCode="SMF 107305" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
|
||||||
|
one
|
||||||
|
<specimenCount id="9D5CFD6D0008FF86AD4CF968FC0D2A9D" box="[817,961,1719,1745]" pageId="3" pageNumber="304" type="adult">adult female</specimenCount>
|
||||||
|
(
|
||||||
|
<specimenCode id="DBFC9E9F0008FF86ADA8F968FBBF2A9D" box="[981,1139,1718,1745]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="304">SMF 107305</specimenCode>
|
||||||
|
), same collecting data as holotype, still alive at the time of writing (
|
||||||
|
<figureCitation id="13612A610008FF86AC19F904FD6A2AB9" box="[612,678,1755,1781]" captionStart="FIGURE 6" captionStartId="9.[152,255,1810,1834]" captionTargetBox="[227,1361,237,1786]" captionTargetId="figure-54@9.[227,1361,237,1786]" captionTargetPageId="9" captionText="FIGURE 6. Paratypes of Rhacodactylus willihenkeli sp. nov. in life (still alive at the time of writing). A. Adult male (SMF 106563); B. Adult female (SMF 107305). Photographs by J.S." figureDoi="http://doi.org/10.5281/zenodo.14611806" httpUri="https://zenodo.org/record/14611806/files/figure.png" pageId="3" pageNumber="304">Fig. 6</figureCitation>
|
||||||
|
), to be deposited in the collection of Senckenberg Forschungsinstitut Frankfurt, Germany, after they have died.
|
||||||
|
</materialsCitation>
|
||||||
|
<materialsCitation id="3B323CB90008FF86AC12F920FADB2B55" box="[623,1303,1790,1817]" collectingDate="1987-09" collectionCode="ZFMK" collectorName="NE Bourail" country="Germany" elevation="750" location="Senckenberg Forschungsinstitut Frankfurt" pageId="3" pageNumber="304" specimenCode="ZFMK 73575" specimenCount="1" typeStatus="paratype">
|
||||||
|
<specimenCode id="DBFC9E9F0008FF86AC12F920FCC62B55" box="[623,778,1790,1817]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="304" type="Museum">ZFMK 73575</specimenCode>
|
||||||
|
(adult
|
||||||
|
<specimenCount id="9D5CFD6D0008FF86AD24F920FC582B55" box="[857,916,1791,1817]" pageId="3" pageNumber="304" type="male">male</specimenCount>
|
||||||
|
), same collecting data as holotype
|
||||||
|
</materialsCitation>
|
||||||
|
.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<subSubSection id="C340656F0008FF81AEBAF8FDFD402C99" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304" type="diagnosis">
|
||||||
|
<paragraph id="8BE536E40008FF81AEBAF8FDFD402C99" blockId="3.[151,1437,1610,2033]" lastBlockId="4.[151,1436,150,213]" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304">
|
||||||
|
<emphasis id="B92EEAF60008FF86AEBAF8FDFEF62B70" bold="true" box="[199,314,1826,1852]" pageId="3" pageNumber="304">Diagnosis</emphasis>
|
||||||
|
. A species of the genus
|
||||||
|
<taxonomicName id="4C5A4D670008FF86AC25F8FCFD312B71" authorityName="Fitzinger" authorityYear="1843" box="[600,765,1827,1853]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Chordata" rank="genus">
|
||||||
|
<emphasis id="B92EEAF60008FF86AC25F8FCFD312B71" box="[600,765,1827,1853]" italics="true" pageId="3" pageNumber="304">Rhacodactylus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
(sensu
|
||||||
|
<bibRefCitation id="EFCB4B150008FF86AD2AF8FCFBE22B71" author="Bauer, A. M. & Jackmann, T. R. & Sadlier, R. A. & Whitaker, A. H." box="[855,1070,1827,1853]" pageId="3" pageNumber="304" pagination="1 - 52" refId="ref6356" refString="Bauer, A. M., Jackmann, T. R., Sadlier, R. A. & Whitaker, A. H. (2012 a) Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus). Zootaxa, 3404 (1), 1 - 52. https: // doi. org / 10.11646 / zootaxa. 3404.1.1" type="journal article" year="2012">
|
||||||
|
Bauer
|
||||||
|
<emphasis id="B92EEAF60008FF86ADD9F8FCFC132B71" box="[932,991,1827,1853]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
|
||||||
|
2012a
|
||||||
|
</bibRefCitation>
|
||||||
|
) that differs from all congeners except
|
||||||
|
<taxonomicName id="4C5A4D670008FF86AE9BF898FEB02B2D" box="[230,380,1863,1889]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF60008FF86AE9BF898FEB02B2D" box="[230,380,1863,1889]" italics="true" pageId="3" pageNumber="304">R. leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
by the following combination of characters (1) adult SVL>
|
||||||
|
<quantity id="4CA29B010008FF86AA72F898FBA32B2D" box="[1039,1135,1863,1889]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="304" unit="mm" value="200.0">200 mm</quantity>
|
||||||
|
; (2) skin of head coossified with skull; (3) webbing between digits IV and V present; (4) folds of loose skin on posterior surface of hind limb; (5) folds of skin at mandibular margins.
|
||||||
|
<taxonomicName id="4C5A4D670008FF86AC1CF850FC5D2BE5" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[609,913,1935,1961]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60008FF86AC1CF850FC5D2BE5" box="[609,913,1935,1961]" italics="true" pageId="3" pageNumber="304">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
differs from
|
||||||
|
<taxonomicName id="4C5A4D670008FF86AA5AF850FB0C2BE5" box="[1063,1216,1935,1961]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF60008FF86AA5AF850FB0C2BE5" box="[1063,1216,1935,1961]" italics="true" pageId="3" pageNumber="304">R. leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
by having a darker overall coloration with almost no discernable pattern (versus a wide range of color variation, usually with a bold reticulated body pattern and also usually with distinct pale ocellated blotches along flanks in
|
||||||
|
<taxonomicName id="4C5A4D670008FF86AADEF808FAF72BBD" box="[1187,1339,2007,2033]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="3" pageNumber="304" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF60008FF86AADEF808FAF72BBD" box="[1187,1339,2007,2033]" italics="true" pageId="3" pageNumber="304">R. leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
;
|
||||||
|
<figureCitation id="13612A610008FF86AB34F808FA5B2BBD" box="[1353,1431,2007,2033]" captionStart="FIGURE 2" captionStartId="5.[152,255,1668,1692]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure-20@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 2. Female holotype (SMF 105137, upper individual) and adult male (not preserved,lower individual) of Rhacodactylus willihenkeli sp. nov. in life. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611798" httpUri="https://zenodo.org/record/14611798/files/figure.png" pageId="3" pageNumber="304">Figs. 2</figureCitation>
|
||||||
|
,
|
||||||
|
<figureCitation id="13612A61000FFF81AEEAFF48FF6A2CFD" box="[151,166,151,177]" captionStart="FIGURE 3" captionStartId="6.[152,255,1692,1716]" captionTargetBox="[151,1436,741,1669]" captionTargetId="figure-314@6.[151,1436,741,1669]" captionTargetPageId="6" captionText="FIGURE 3. Specimens of Rhacodactylus leachianus in life (not preserved). (A) from southern Grande Terre; (B) from Ile de Pins; (C) from Ilot Brosse; (D) from Ile de Nuami. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611800" httpUri="https://zenodo.org/record/14611800/files/figure.png" pageId="4" pageNumber="305">3</figureCitation>
|
||||||
|
,
|
||||||
|
<figureCitation id="13612A61000FFF81AEC9FF48FF082CFD" box="[180,196,151,177]" captionStart="FIGURE 6" captionStartId="9.[152,255,1810,1834]" captionTargetBox="[227,1361,237,1786]" captionTargetId="figure-54@9.[227,1361,237,1786]" captionTargetPageId="9" captionText="FIGURE 6. Paratypes of Rhacodactylus willihenkeli sp. nov. in life (still alive at the time of writing). A. Adult male (SMF 106563); B. Adult female (SMF 107305). Photographs by J.S." figureDoi="http://doi.org/10.5281/zenodo.14611806" httpUri="https://zenodo.org/record/14611806/files/figure.png" pageId="4" pageNumber="305">6</figureCitation>
|
||||||
|
). In addition, the two taxa are genetically divergent with a genetic distance of 5.0% in the 16S gene fragment and 8.5% in the ND2 fragment, respectively.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<caption id="DF25666C000FFF81AEE5F8B7FB422BA0" ID-DOI="http://doi.org/10.5281/zenodo.14611796" ID-Zenodo-Dep="14611796" httpUri="https://zenodo.org/record/14611796/files/figure.png" pageId="4" pageNumber="305" startId="4.[152,255,1896,1920]" targetBox="[403,1184,239,1864]" targetPageId="4" targetType="figure">
|
||||||
|
<paragraph id="8BE536E4000FFF81AEE5F8B7FB422BA0" blockId="4.[151,1437,1896,2028]" pageId="4" pageNumber="305">
|
||||||
|
<emphasis id="B92EEAF6000FFF81AEE5F8B7FED02BCC" bold="true" box="[152,284,1896,1920]" pageId="4" pageNumber="305">FIGURE 1.</emphasis>
|
||||||
|
Phylogenetic tree of diplodactylid geckoes related to
|
||||||
|
<taxonomicName id="4C5A4D67000FFF81AD23F8B7FBA72BCC" box="[862,1131,1896,1920]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="4" pageNumber="305" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF6000FFF81AD23F8B7FBA72BCC" box="[862,1131,1896,1920]" italics="true" pageId="4" pageNumber="305">Rhacodactylus leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
from a maximum-likelihood analysis of DNA sequences of eight nuclear (ACM4, C-MOS, KIAA1549, KIF24, MXRA5, PDC, RAG1 and RAG2) and two mitochondrial markers (16S, ND2 and assigned tRNAs). A scale bar is indicated. The numbers at nodes are bootstrap values (left) and Bayesian posterior probabilities (right) but only bootstrap values higher than 70 are noted.
|
||||||
|
</paragraph>
|
||||||
|
</caption>
|
||||||
|
<caption id="DF25666C000EFF80AEE5F95BFD532A8C" ID-DOI="http://doi.org/10.5281/zenodo.14611798" ID-Zenodo-Dep="14611798" httpUri="https://zenodo.org/record/14611798/files/figure.png" pageId="5" pageNumber="306" startId="5.[152,255,1668,1692]" targetBox="[151,1436,181,1644]" targetPageId="5" targetType="figure">
|
||||||
|
<paragraph id="8BE536E4000EFF80AEE5F95BFD532A8C" blockId="5.[151,1436,1668,1728]" pageId="5" pageNumber="306">
|
||||||
|
<emphasis id="B92EEAF6000EFF80AEE5F95BFEDA2AD0" bold="true" box="[152,278,1668,1692]" pageId="5" pageNumber="306">FIGURE 2.</emphasis>
|
||||||
|
Female holotype (SMF 105137, upper individual) and adult male (not preserved, lower individual) of
|
||||||
|
<taxonomicName id="4C5A4D67000EFF80AB7AF95BFEC12A8C" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="willihenkeli" status="sp. nov.">
|
||||||
|
<emphasis id="B92EEAF6000EFF80AB7AF95BFEC12A8C" italics="true" pageId="5" pageNumber="306">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
<emphasis id="B92EEAF6000EFF80AF69F977FEA82A8C" bold="true" box="[276,356,1704,1728]" pageId="5" pageNumber="306">
|
||||||
|
<taxonomicNameLabel id="A21D578D000EFF80AF69F977FEA82A8C" box="[276,356,1704,1728]" pageId="5" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
|
||||||
|
</emphasis>
|
||||||
|
in life. Photographs by F.W.H.
|
||||||
|
</paragraph>
|
||||||
|
</caption>
|
||||||
|
<subSubSection id="C340656F000EFF8DAEBAF8D7FB8F2957" lastPageId="8" lastPageNumber="309" pageId="5" pageNumber="306" type="description">
|
||||||
|
<paragraph id="8BE536E4000EFF83AEBAF8D7FE292E81" blockId="5.[151,1437,1800,2042]" lastBlockId="6.[151,1437,151,717]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
|
||||||
|
<emphasis id="B92EEAF6000EFF80AEBAF8D7FDD52B6E" bold="true" box="[199,537,1800,1826]" pageId="5" pageNumber="306">
|
||||||
|
Description of the
|
||||||
|
<typeStatus id="54E18846000EFF80AFCFF8D7FDD52B6E" box="[434,537,1800,1826]" pageId="5" pageNumber="306" type="holotype">holotype</typeStatus>
|
||||||
|
</emphasis>
|
||||||
|
(
|
||||||
|
<figureCitation id="13612A61000EFF80AC50F8D7FD4C2B6E" box="[557,640,1800,1826]" captionStart="FIGURE 4" captionStartId="7.[152,255,1940,1964]" captionTargetBox="[151,1436,181,1915]" captionTargetId="figure-20@7.[151,1436,181,1916]" captionTargetPageId="7" captionText="FIGURE 4. Holotype of Rhacodactylus willihenkeli sp. nov. (SMF 105137). Scale bars equal 40.0 mm in A to D, 10.0 mm in E to H. Photographs by G.K." figureDoi="http://doi.org/10.5281/zenodo.14611802" httpUri="https://zenodo.org/record/14611802/files/figure.png" pageId="5" pageNumber="306">Figs. 4</figureCitation>
|
||||||
|
and
|
||||||
|
<figureCitation id="13612A61000EFF80ACBDF8D7FD1C2B6F" box="[704,720,1800,1827]" captionStart="FIGURE 5" captionStartId="8.[152,255,1066,1090]" captionTargetBox="[151,1436,181,1042]" captionTargetId="figure-21@8.[151,1436,181,1042]" captionTargetPageId="8" captionText="FIGURE 5. Holotype of Rhacodactylus willihenkeli sp. nov. (SMF 105137). Scale bars equal 10.0 mm. Photographs by G.K." figureDoi="http://doi.org/10.5281/zenodo.14611804" httpUri="https://zenodo.org/record/14611804/files/figure.png" pageId="5" pageNumber="306">5</figureCitation>
|
||||||
|
). Adult female (SVL 220.0 mm) as indicated by absence of hemipenial bulge and femoral and precloacal pores. Body robust, slightly depressed. Head triangular with a blunt snout, large (ratio HL/SVL 0.23), wide (ratio HW/HL 0.73) and depressed (ratio HH/HL 0.47), weakly offset from neck; interorbital/frontal region somewhat depressed, nasal region with a distinct, shallow indentation; canthus weakly inflated; snout long (ratio SL/HL 0.87). Scales on dorsum of snout slightly larger than of those on occipital region. Eye small (ratio EYD/HL 0.22); pupil oval, margins crenellated. Ear opening small, horizontally elongate (ratio EarL/HL 0.07); eye to ear distance much longer than diameter of eyes (ratio EED/EYD 1.92). Rostral and mental region damaged with scar tissue. Scales in 2–4 rows medial to infralabials somewhat enlarged and elongate. Dorsal scales heterogeneous with flat tiny scales interspersed among even smaller scales; ventral scales about half the size of dorsals, smooth, flattened, juxtaposed, slightly enlarged on posterior abdomen, in precloacal region and on base of thighs. Approximately 282 scale rows around midbody. Ventrolateral skin folds present. Scales on palms and soles smooth, flattened. Forelimbs and hind limbs short and thick (ratio FAL/SVL 0.13; ratio SHL/SVL 0.15), axillary pocket weakly developed. Digits short, all bearing robust and strongly recurved claws. Relative length of digits of manus: IV> III> V> II> I, and of pes: IV> III> V> II> I; digits moderately webbed. Subdigital lamellae of digits unpaired, except for distal portion of 4
|
||||||
|
<superScript id="7C2F9BAC000DFF83ACCCFE4DFD722DEC" attach="left" box="[689,702,402,416]" fontSize="6" pageId="6" pageNumber="307">th</superScript>
|
||||||
|
finger on right side. Claw of digit I, manus and pes, with medial apical scansor. Lamellar counts (including all scales> twice size of scales at base of digits) from right (and left) sides 8-11-14-18-10 (7-9-15-16-10) manus and 9-15-17-20-16 (9-13-16-18-17) pes. No precloacal pores or femoral pores. Regenerated tail 38% of SVL, slender, round in cross section. Coloration in life, based on
|
||||||
|
<figureCitation id="13612A61000DFF83AB78FE20FA852E55" box="[1285,1353,511,537]" captionStart="FIGURE 2" captionStartId="5.[152,255,1668,1692]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure-20@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 2. Female holotype (SMF 105137, upper individual) and adult male (not preserved,lower individual) of Rhacodactylus willihenkeli sp. nov. in life. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611798" httpUri="https://zenodo.org/record/14611798/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
|
||||||
|
(upper portion), was recorded as follows: Dorsal and lateral ground color of head body, and limbs Light Drab (268) with suffusions of Hair Brown (277) and Citrine (119) and with scattered Sulphur Yellow (91) and Medium Greenish Yellow (88) splotches; dorsal surface of tail base Glaucous (291), regenerated portion of tail Brownish Olive (292) with Smoky White (261) splotches; ring around eye Sulphur Yellow (91); iris Light Neutral Gray (297) with Pale Neutral Gray (296) splotches.
|
||||||
|
</paragraph>
|
||||||
|
<caption id="DF25666C000DFF83AEE5F943FC452A94" ID-DOI="http://doi.org/10.5281/zenodo.14611800" ID-Zenodo-Dep="14611800" httpUri="https://zenodo.org/record/14611800/files/figure.png" pageId="6" pageNumber="307" startId="6.[152,255,1692,1716]" targetBox="[151,1436,741,1669]" targetPageId="6" targetType="figure">
|
||||||
|
<paragraph id="8BE536E4000DFF83AEE5F943FC452A94" blockId="6.[151,1436,1692,1752]" pageId="6" pageNumber="307">
|
||||||
|
<emphasis id="B92EEAF6000DFF83AEE5F943FED52AF8" bold="true" box="[152,281,1692,1716]" pageId="6" pageNumber="307">FIGURE 3.</emphasis>
|
||||||
|
Specimens of
|
||||||
|
<taxonomicName id="4C5A4D67000DFF83AFCCF943FD772AF8" box="[433,699,1692,1716]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF6000DFF83AFCCF943FD772AF8" box="[433,699,1692,1716]" italics="true" pageId="6" pageNumber="307">Rhacodactylus leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
in life (not preserved). (A) from southern Grande Terre; (B) from Ile de Pins; (C) from Ilot Brosse; (D) from Ile de Nuami. Photographs by F.W.H.
|
||||||
|
</paragraph>
|
||||||
|
</caption>
|
||||||
|
<paragraph id="8BE536E4000DFF83AEBAF8F5FC262BB5" blockId="6.[151,1436,1834,2041]" pageId="6" pageNumber="307">
|
||||||
|
<emphasis id="B92EEAF6000DFF83AEBAF8F5FDC62B08" bold="true" box="[199,522,1834,1860]" pageId="6" pageNumber="307">Genomic characterization.</emphasis>
|
||||||
|
Whole genome sequencing and assembly yielded 298,315,598 Illumina short reads with a total data size of 102.4 Gb. The k-mer analysis resulted in an estimated genome size of 1.56 Gb and a heterozygosity of 0.6%. The mitochondrial genome was assembled into one linear contig with a total length of 15,957 bp (accession number OR288172). A total of 37 genes could be annotated, including all 13 protein coding genes, 2 rRNAs and 22 tRNAs. Pairwise alignment to the complete mitochondrial genome of
|
||||||
|
<taxonomicName id="4C5A4D67000DFF83AACBF865FA502B98" box="[1206,1436,1978,2004]" class="Squamata" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" pageId="6" pageNumber="307" phylum="Chordata" rank="species" species="ciliatus">
|
||||||
|
<emphasis id="B92EEAF6000DFF83AACBF865FA502B98" box="[1206,1436,1978,2004]" italics="true" pageId="6" pageNumber="307">Correlophus ciliatus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
(GenBank accession No. OQ506627) revealed a pairwise identity of 83.9%.
|
||||||
|
</paragraph>
|
||||||
|
<caption id="DF25666C000CFF82AEE5F84BFE702B9C" ID-DOI="http://doi.org/10.5281/zenodo.14611802" ID-Zenodo-Dep="14611802" httpUri="https://zenodo.org/record/14611802/files/figure.png" pageId="7" pageNumber="308" startId="7.[152,255,1940,1964]" targetBox="[151,1436,181,1915]" targetPageId="7" targetType="figure">
|
||||||
|
<paragraph id="8BE536E4000CFF82AEE5F84BFE702B9C" blockId="7.[151,1437,1940,2000]" pageId="7" pageNumber="308">
|
||||||
|
<emphasis id="B92EEAF6000CFF82AEE5F84BFED42BE0" bold="true" box="[152,280,1940,1964]" pageId="7" pageNumber="308">FIGURE 4.</emphasis>
|
||||||
|
Holotype of
|
||||||
|
<taxonomicName id="4C5A4D67000CFF82AFE2F84BFD7D2BE0" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[415,689,1940,1964]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="willihenkeli" status="sp. nov.">
|
||||||
|
<emphasis id="B92EEAF6000CFF82AFE2F84BFD7D2BE0" box="[415,689,1940,1964]" italics="true" pageId="7" pageNumber="308">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
<emphasis id="B92EEAF6000CFF82ACC5F84BFCC42BE0" bold="true" box="[696,776,1940,1964]" pageId="7" pageNumber="308">
|
||||||
|
<taxonomicNameLabel id="A21D578D000CFF82ACC5F84BFCC42BE0" box="[696,776,1940,1964]" pageId="7" pageNumber="308" rank="species">sp. nov.</taxonomicNameLabel>
|
||||||
|
</emphasis>
|
||||||
|
(SMF 105137). Scale bars equal 40.0 mm in A to D, 10.0 mm in E to H. Photographs by G.K.
|
||||||
|
</paragraph>
|
||||||
|
</caption>
|
||||||
|
<caption id="DF25666C0003FF8DAEE5FBF5FA55280E" ID-DOI="http://doi.org/10.5281/zenodo.14611804" ID-Zenodo-Dep="14611804" box="[152,1433,1066,1090]" httpUri="https://zenodo.org/record/14611804/files/figure.png" pageId="8" pageNumber="309" startId="8.[152,255,1066,1090]" targetBox="[151,1436,181,1042]" targetPageId="8" targetType="figure">
|
||||||
|
<paragraph id="8BE536E40003FF8DAEE5FBF5FA55280E" blockId="8.[152,1433,1066,1090]" box="[152,1433,1066,1090]" pageId="8" pageNumber="309">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEE5FBF5FED4280E" bold="true" box="[152,280,1066,1090]" pageId="8" pageNumber="309">FIGURE 5.</emphasis>
|
||||||
|
Holotype of
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DAFE2FBF5FD7C280E" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[415,688,1066,1090]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli" status="sp. nov.">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAFE2FBF5FD7C280E" box="[415,688,1066,1090]" italics="true" pageId="8" pageNumber="309">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
<emphasis id="B92EEAF60003FF8DACCAFBF5FCCB280E" bold="true" box="[695,775,1066,1090]" pageId="8" pageNumber="309">
|
||||||
|
<taxonomicNameLabel id="A21D578D0003FF8DACCAFBF5FCCB280E" box="[695,775,1066,1090]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
|
||||||
|
</emphasis>
|
||||||
|
(SMF 105137). Scale bars equal 10.0 mm. Photographs by G.K.
|
||||||
|
</paragraph>
|
||||||
|
</caption>
|
||||||
|
<paragraph id="8BE536E40003FF8DAEBAFB4BFB8F2957" blockId="8.[151,1437,1172,2027]" pageId="8" pageNumber="309">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEBAFB4BFEF328E2" bold="true" box="[199,319,1172,1198]" pageId="8" pageNumber="309">Variation.</emphasis>
|
||||||
|
Coloration in life of an adult male, based on
|
||||||
|
<figureCitation id="13612A610003FF8DAD48FB4BFCB428E3" box="[821,888,1172,1199]" captionStart="FIGURE 2" captionStartId="5.[152,255,1668,1692]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure-20@5.[151,1436,181,1644]" captionTargetPageId="5" captionText="FIGURE 2. Female holotype (SMF 105137, upper individual) and adult male (not preserved,lower individual) of Rhacodactylus willihenkeli sp. nov. in life. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611798" httpUri="https://zenodo.org/record/14611798/files/figure.png" pageId="8" pageNumber="309">Fig. 2</figureCitation>
|
||||||
|
, was recorded as follows: Dorsal ground color of head body, and limbs Pale Cinnamon (55) with Verona Brown (37) streaks on head and suffusions on body, grading into Olive Horn Color (16) in flank region; labial region and ring around eye Sulphur Yellow (80); iris Smoky White (261) with Smoke Gray (266) stipples. See
|
||||||
|
<figureCitation id="13612A610003FF8DAC03FADFFD732956" box="[638,703,1280,1306]" captionStart="FIGURE 6" captionStartId="9.[152,255,1810,1834]" captionTargetBox="[227,1361,237,1786]" captionTargetId="figure-54@9.[227,1361,237,1786]" captionTargetPageId="9" captionText="FIGURE 6. Paratypes of Rhacodactylus willihenkeli sp. nov. in life (still alive at the time of writing). A. Adult male (SMF 106563); B. Adult female (SMF 107305). Photographs by J.S." figureDoi="http://doi.org/10.5281/zenodo.14611806" httpUri="https://zenodo.org/record/14611806/files/figure.png" pageId="8" pageNumber="309">Fig. 6</figureCitation>
|
||||||
|
for photos in life of the
|
||||||
|
<typeStatus id="54E188460003FF8DADADFADEFBF32957" box="[976,1087,1281,1307]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||||||
|
.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<subSubSection id="C340656F0003FF8DAEBAFAFAFBF729E6" pageId="8" pageNumber="309" type="etymology">
|
||||||
|
<paragraph id="8BE536E40003FF8DAEBAFAFAFBF729E6" blockId="8.[151,1437,1172,2027]" pageId="8" pageNumber="309">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEBAFAFAFE842973" bold="true" box="[199,328,1317,1343]" pageId="8" pageNumber="309">Etymology</emphasis>
|
||||||
|
. The species name
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DAC4BFAFBFD752972" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[566,697,1316,1342]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAC4BFAFBFD752972" box="[566,697,1316,1342]" italics="true" pageId="8" pageNumber="309">willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
honors Friedrich Wilhelm Henkel, nickname “Willi”, who has studied geckos of the genus
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DAFB6FA97FDBC292E" authorityName="Fitzinger" authorityYear="1843" box="[459,624,1352,1378]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAFB6FA97FDBC292E" box="[459,624,1352,1378]" italics="true" pageId="8" pageNumber="309">Rhacodactylus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
in their natural habitat during about a dozen excursions to
|
||||||
|
<collectingCountry id="F34D76740003FF8DAA8CFA96FA51292E" box="[1265,1437,1352,1379]" name="New Caledonia" pageId="8" pageNumber="309">New Caledonia</collectingCountry>
|
||||||
|
and while keeping and breeding these fascinating animals in terrariums for decades. Friedrich Wilhelm Henkel coauthored research articles as well as a popular science book about these geckoes.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<subSubSection id="C340656F0003FF8DAEBAFA6AFCE62A5A" pageId="8" pageNumber="309" type="biology_ecology">
|
||||||
|
<paragraph id="8BE536E40003FF8DAEBAFA6AFCE62A5A" blockId="8.[151,1437,1172,2027]" pageId="8" pageNumber="309">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEBAFA6AFE0F2983" bold="true" box="[199,451,1460,1487]" pageId="8" pageNumber="309">Natural history notes</emphasis>
|
||||||
|
. At the time of collection of the
|
||||||
|
<typeStatus id="54E188460003FF8DAD57FA6BFC402982" box="[810,908,1460,1486]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
|
||||||
|
of
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DADD2FA6BFB112982" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[943,1245,1460,1486]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60003FF8DADD2FA6BFB112982" box="[943,1245,1460,1486]" italics="true" pageId="8" pageNumber="309">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
, the type locality was located in a large, forested area in the mountains (
|
||||||
|
<figureCitation id="13612A610003FF8DAD72FA07FC9229BF" box="[783,862,1496,1523]" captionStart="FIGURE 7" captionStartId="10.[152,255,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-21@10.[151,1436,181,1121]" captionTargetPageId="10" captionText="FIGURE 7. Habitat of Rhacodactylus willihenkeli sp. nov. near the type locality. Photographs by F.W.H." figureDoi="http://doi.org/10.5281/zenodo.14611808" httpUri="https://zenodo.org/record/14611808/files/figure.png" pageId="8" pageNumber="309">Figs. 7</figureCitation>
|
||||||
|
and
|
||||||
|
<figureCitation id="13612A610003FF8DADE4FA06FC6529BF" box="[921,937,1497,1523]" captionStart="FIGURE 8" captionStartId="11.[152,255,1938,1962]" captionTargetBox="[151,1436,237,1915]" captionTargetId="figure-51@11.[151,1436,237,1915]" captionTargetPageId="11" captionText="FIGURE 8. Landscape views from the type locality of Rhacodactylus willihenkeli sp. nov. A. View in eastern direction.Arrow points to Canala Bay on the east coast of central New Caledonia; B. View in unknown direction. Photographs by J.S." figureDoi="http://doi.org/10.5281/zenodo.14611810" httpUri="https://zenodo.org/record/14611810/files/figure.png" pageId="8" pageNumber="309">8</figureCitation>
|
||||||
|
). Henkel and Sameit were accompanying a group of lumberjacks, checking the fallen trees for geckos.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<subSubSection id="C340656F0003FF8DAEBAF9FFFDEA2B17" pageId="8" pageNumber="309" type="distribution">
|
||||||
|
<paragraph id="8BE536E40003FF8DAEBAF9FFFDEA2B17" blockId="8.[151,1437,1172,2027]" pageId="8" pageNumber="309">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEBAF9FFFD062A76" bold="true" box="[199,714,1568,1595]" pageId="8" pageNumber="309">Geographic Distribution and Conservation</emphasis>
|
||||||
|
. As currently known,
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DADB8F9FFFB392A76" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[965,1269,1568,1594]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60003FF8DADB8F9FFFB392A76" box="[965,1269,1568,1594]" italics="true" pageId="8" pageNumber="309">Rhacodactylus willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
is known only from its type locality in central
|
||||||
|
<collectingCountry id="F34D76740003FF8DAF85F99AFD6A2A12" box="[504,678,1604,1631]" name="New Caledonia" pageId="8" pageNumber="309">New Caledonia</collectingCountry>
|
||||||
|
(
|
||||||
|
<figureCitation id="13612A610003FF8DACC8F99BFD352A13" box="[693,761,1604,1631]" captionStart="FIGURE 9" captionStartId="12.[152,255,1083,1107]" captionTargetBox="[177,1411,181,1059]" captionTargetId="figure-21@12.[177,1411,181,1059]" captionTargetPageId="12" captionText="FIGURE 9. Map indicating collecting localities of samples used in genetic analyses. Green square = Rhacodactylus willihenkeli sp. nov.; red circles = R. leachianus." figureDoi="http://doi.org/10.5281/zenodo.14611812" httpUri="https://zenodo.org/record/14611812/files/figure.png" pageId="8" pageNumber="309">Fig. 9</figureCitation>
|
||||||
|
). Unfortunately, at the time of collection of the
|
||||||
|
<typeStatus id="54E188460003FF8DAB6DF99BFAB92A12" box="[1296,1397,1604,1630]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
|
||||||
|
, no GPS device was available and, therefore, the precise coordinates are unknown. From the type locality, they could see both coasts, and at night the lights of the city of Houaïlou. FWH and JS started their excursion from Bourail and it took them 1.5 h along a dirt road to reach the place where they camped and collected the
|
||||||
|
<typeStatus id="54E188460003FF8DAA85F96FFA962A86" box="[1272,1370,1712,1738]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
|
||||||
|
of
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DABF9F96EFED62AA2" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60003FF8DABF9F96EFED62AA2" italics="true" pageId="8" pageNumber="309">R. willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
. On a landscape photo taken at the type locality (
|
||||||
|
<figureCitation id="13612A610003FF8DAD3DF90BFC5D2AA3" box="[832,913,1748,1775]" captionStart="FIGURE 8" captionStartId="11.[152,255,1938,1962]" captionTargetBox="[151,1436,237,1915]" captionTargetId="figure-51@11.[151,1436,237,1915]" captionTargetPageId="11" captionText="FIGURE 8. Landscape views from the type locality of Rhacodactylus willihenkeli sp. nov. A. View in eastern direction.Arrow points to Canala Bay on the east coast of central New Caledonia; B. View in unknown direction. Photographs by J.S." figureDoi="http://doi.org/10.5281/zenodo.14611810" httpUri="https://zenodo.org/record/14611810/files/figure.png" pageId="8" pageNumber="309">Fig. 8a</figureCitation>
|
||||||
|
), a narrow bay, enclosed by mountain ridges is visible. This bay clearly corresponds to Canala Bay on the east coast of central
|
||||||
|
<collectingCountry id="F34D76740003FF8DAA69F926FB092B5E" box="[1044,1221,1784,1811]" name="New Caledonia" pageId="8" pageNumber="309">New Caledonia</collectingCountry>
|
||||||
|
. Thus, we consider the locality information (i.e., northern
|
||||||
|
<collectingCountry id="F34D76740003FF8DAC3BF8C2FD3A2B7A" box="[582,758,1820,1847]" name="New Caledonia" pageId="8" pageNumber="309">New Caledonia</collectingCountry>
|
||||||
|
, west of Houaïlou) given by
|
||||||
|
<bibRefCitation id="EFCB4B150003FF8DAA46F8C2FAD72B7B" author="Vences, M. & Henkel, F. - W. & Seipp, R." box="[1083,1307,1820,1847]" pageId="8" pageNumber="309" pagination="73 - 82" refId="ref9378" refString="Vences, M., Henkel, F. - W. & Seipp, R. (2001) Molekulare Untersuchungen zur Phylogenie und Taxonomie der Neukaledonischen Geckos der Gattung Rhacodactylus (Reptilia: Gekkonidae). Salamandra, 37, 73 - 82." type="journal article" year="2001">
|
||||||
|
Vences
|
||||||
|
<emphasis id="B92EEAF60003FF8DAAECF8C2FB062B7A" box="[1169,1226,1820,1846]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
|
||||||
|
(2001)
|
||||||
|
</bibRefCitation>
|
||||||
|
for their
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DABF9F8C2FEF82B16" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" isUncertain="true" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="leachianus">
|
||||||
|
<emphasis id="B92EEAF60003FF8DABF9F8C2FA592B7A" box="[1412,1429,1821,1846]" italics="true" pageId="8" pageNumber="309">R</emphasis>
|
||||||
|
. cf.
|
||||||
|
<emphasis id="B92EEAF60003FF8DAEC6F89FFEF82B16" box="[187,308,1856,1882]" italics="true" pageId="8" pageNumber="309">leachianus</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
sample as erroneous.
|
||||||
|
</paragraph>
|
||||||
|
</subSubSection>
|
||||||
|
<subSubSection id="C340656F0003FF8CAEBAF8BBFD7D2C99" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="conservation">
|
||||||
|
<paragraph id="8BE536E40003FF8CAEBAF8BBFD7D2C99" blockId="8.[151,1437,1172,2027]" lastBlockId="9.[151,1436,150,213]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
|
||||||
|
Given the little we know about this species, we classify
|
||||||
|
<taxonomicName id="4C5A4D670003FF8DAD51F8BAFC012B32" authorityName="Köhler & Sameit & Seipp & Geiss" authorityYear="2024" box="[812,973,1892,1918]" class="Squamata" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="willihenkeli">
|
||||||
|
<emphasis id="B92EEAF60003FF8DAD51F8BAFC012B32" box="[812,973,1892,1918]" italics="true" pageId="8" pageNumber="309">R. willihenkeli</emphasis>
|
||||||
|
</taxonomicName>
|
||||||
|
as Data Deficient based on the
|
||||||
|
<collectionCode id="ED4BAE210003FF8DAB58F8BAFAA52B33" box="[1317,1385,1893,1919]" pageId="8" pageNumber="309">IUCN</collectionCode>
|
||||||
|
Red List Categories and Criteria (
|
||||||
|
<specimenCode id="DBFC9E9F0003FF8DAFA3F856FDA52BEF" box="[478,617,1929,1955]" collectionCode="IUCN" pageId="8" pageNumber="309">IUCN 2012</specimenCode>
|
||||||
|
). This species is oviparous and has been bred in captivity. The following observations were made by JS who has bred about 15 individuals of this species: Females lay three or four clutches of
|
||||||
|
<specimenCount id="9D5CFD6D0003FF8DAECBF80EFED72BA7" box="[182,283,2001,2027]" count="2" pageId="8" pageNumber="309" type="egg">two eggs</specimenCount>
|
||||||
|
each per year. After an incubation period of about 90 days the juveniles hatch with a
|
||||||
|
<collectionCode id="ED4BAE210003FF8DAA9DF80EFAD92BA7" box="[1248,1301,2001,2027]" pageId="8" pageNumber="309">SVL</collectionCode>
|
||||||
|
of about
|
||||||
|
<quantity id="4CA29B010003FF8CABFDF80FFF0A2CFD" lastPageId="9" lastPageNumber="310" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="309" unit="mm" value="60.0">60 mm</quantity>
|
||||||
|
. The adults that are still alive have been kept by JS for almost 40 years now and are probably more than 50 years old since these were collected as adults in 1987.
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</paragraph>
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</subSubSection>
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|
</treatment>
|
||||||
|
</document>
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Reference in a new issue