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<mods:affiliation id="0C87CFBBB50DDA1A16AF982AC36F14C8">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:affiliation id="323C46D008A096EA932E4C024315DAB5">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:namePart id="7FBC1AAC38DFE356F07FC804AD8D96AD">Rohal, Melissa</mods:namePart>
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<mods:namePart id="C2BC64856F088A3A564795D331393155">Thistle, David</mods:namePart>
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<treatment id="F64287A2504DFFBA145BFA2DFC6F0E34" ID-DOI="http://doi.org/10.5281/zenodo.3818842" ID-GBIF-Taxon="163993851" ID-Zenodo-Dep="3818842" LSID="urn:lsid:zoobank.org:act:58E8CD4B-12EA-49F0-8B97-167C2F5961B9" httpUri="http://treatment.plazi.org/id/F64287A2504DFFBA145BFA2DFC6F0E34" lastPageId="36" lastPageNumber="36" pageId="27" pageNumber="28">
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<taxonomicName id="B9EB4D37504DFF85145BFA2DFCA60A6A" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[603,888,1523,1550]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hviidarum" status="sp. nov.">
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@ -64,17 +64,17 @@
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<taxonomicNameLabel id="57AC57DD504DFF85157EFA2DFC070A69" box="[894,985,1523,1549]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="E6D02A31504DFF851491F9E5FD370A31" box="[657,745,1595,1621]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="27" pageNumber="28">Figs 10</figureCitation>
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–13,
|
||||
<figureCitation id="E6D02A31504DFF851491F9E5FCC90A31" box="[657,791,1595,1621]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="29.[189,232,1718,1744]" captionStartId-1="30.[189,232,1930,1956]" captionStartId-2="31.[189,232,1752,1778]" captionStartId-3="33.[189,232,1578,1604]" captionTargetBox-0="[287,1297,269,1696]" captionTargetBox-1="[201,1388,269,1911]" captionTargetBox-2="[217,1371,265,1740]" captionTargetId-0="figure@29.[286,1301,265,1696]" captionTargetId-1="figure@30.[200,1389,265,1912]" captionTargetId-2="figure@31.[216,1371,265,1740]" captionTargetId-3="figure@34.[252,1336,265,2020]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="34" captionText-0="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." captionText-1="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." captionText-2="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." captionText-3="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi-0="http://doi.org/10.5281/zenodo.3817843" figureDoi-1="http://doi.org/10.5281/zenodo.3817845" figureDoi-2="http://doi.org/10.5281/zenodo.3817847" figureDoi-3="http://doi.org/10.5281/zenodo.14897716" httpUri-0="https://zenodo.org/record/3817843/files/figure.png" httpUri-1="https://zenodo.org/record/3817845/files/figure.png" httpUri-2="https://zenodo.org/record/3817847/files/figure.png" httpUri-3="https://zenodo.org/record/14897716/files/figure.png" pageId="27" pageNumber="28">Figs 10–13</figureCitation>
|
||||
,
|
||||
<tableCitation id="3369030F504DFF851523F9E5FC7C0A31" box="[803,930,1595,1621]" captionStart-0="Table 8" captionStart-1="Table 9" captionStartId-0="32.[189,258,268,294]" captionStartId-1="33.[189,258,268,294]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="Table 8. Measurements from light microscopy of Echinoderes hviidarum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-5 = maximum sternal width, measured on segment 5 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 9. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes hviidarum sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1 = glandular cell outlet type 1; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; pr = protuberance; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters." httpUri-1="http://table.plazi.org/id/2A94663C5077FFBF16BDFED2FDFB0D9C" pageId="27" pageNumber="28" tableUuid-1="2A94663C5077FFBF16BDFED2FDFB0D9C">Tables 8–9</tableCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -107,13 +107,11 @@ with middorsal spines on segments 6 and 8, and spines in lateroventral positions
|
|||
<emphasis id="4C9FEAA6504AFF8216BDFED5FE740D41" bold="true" box="[189,426,267,293]" pageId="28" pageNumber="29">Material examined</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216EAFEE4FAD90DDA" blockId="28.[189,1399,314,447]" pageId="28" pageNumber="29">
|
||||
<materialsCitation id="CE833CE9504AFF8216EAFEE4FAD90DDA" ID-GBIF-Occurrence="2611490302" collectingDate="2008-09-21" collectionCode="US, NHMD" collectorName="St. & Fluoromount G" country="United States of America" latitude="39.998055" location="West Coast" longLatPrecision="19" longitude="-125.44334" pageId="28" pageNumber="29" specimenCount="1" stateProvince="California" typeStatus="holotype">
|
||||
<paragraph id="7E5436B4504AFF8216EAFEE4FE820D30" blockId="28.[189,1399,314,447]" box="[234,348,314,340]" pageId="28" pageNumber="29">
|
||||
<emphasis id="4C9FEAA6504AFF8216EAFEE4FE820D30" bold="true" box="[234,348,314,340]" pageId="28" pageNumber="29">
|
||||
<typeStatus id="A1508816504AFF8216EAFEE4FE820D30" box="[234,348,314,340]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216BDFE80FAD90DDA" blockId="28.[189,1399,314,447]" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="06FC7624504AFF8216BDFE80FDBA0D13" box="[189,612,349,376]" name="United States of America" pageId="28" pageNumber="29">UNITED STATES OF AMERICA</collectingCountry>
|
||||
: adult ♂,
|
||||
<collectionCode id="18FAAE71504AFF8214E3FE83FCD60D13" box="[739,776,349,375]" country="South Africa" name="University of Stellenbosch" pageId="28" pageNumber="29">US</collectionCode>
|
||||
|
|
@ -139,35 +137,30 @@ on a glass slide,
|
|||
for localities and
|
||||
<tableCitation id="3369030F504AFF82158DFE7AFC3A0DDB" box="[909,996,420,447]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="28" pageNumber="29" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
|
||||
for detailed station data.
|
||||
</paragraph>
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216EAFE39FE550E0F" blockId="28.[189,1399,487,620]" pageId="28" pageNumber="29">
|
||||
<materialsCitation id="CE833CE9504AFF8216EAFE39FE550E0F" ID-GBIF-Occurrence="2611490314" pageId="28" pageNumber="29" typeStatus="paratype">
|
||||
<paragraph id="7E5436B4504AFF8216EAFE39FEB90E65" blockId="28.[189,1399,487,620]" box="[234,359,487,513]" pageId="28" pageNumber="29">
|
||||
<emphasis id="4C9FEAA6504AFF8216EAFE39FEB90E65" bold="true" box="[234,359,487,513]" pageId="28" pageNumber="29">
|
||||
<typeStatus id="A1508816504AFF8216EAFE39FEB90E65" box="[234,359,487,513]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216BDFDD4FE550E0F" blockId="28.[189,1399,487,620]" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="06FC7624504AFF8216BDFDD4FD890E40" box="[189,599,522,548]" name="United States of America" pageId="28" pageNumber="29">UNITED STATES OF AMERICA</collectingCountry>
|
||||
:
|
||||
<specimenCount id="68EDFD3D504AFF821463FDD4FD6B0E40" box="[611,693,522,548]" pageId="28" pageNumber="29" type="female">12 ♀♀</specimenCount>
|
||||
, 9 ♂♂, same collecting data as for
|
||||
<typeStatus id="A1508816504AFF821257FDD4FB1F0E40" box="[1111,1217,522,548]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
, 9 ♂♂, same collecting data as for holotype;
|
||||
<specimenCount id="68EDFD3D504AFF8212CDFDD4FAC10E40" box="[1229,1311,522,548]" pageId="28" pageNumber="29" type="female">11 ♀♀</specimenCount>
|
||||
, 9 ♂♂, St. 5;
|
||||
<specimenCount id="68EDFD3D504AFF821701FDF0FE9D0E2C" box="[257,323,558,584]" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
|
||||
, 1 ♂, St. 6;
|
||||
<specimenCount id="68EDFD3D504AFF8217CEFDF0FE270E2C" box="[462,505,558,584]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
|
||||
, St. 7 (NHMD-213611–213661). All
|
||||
<typeStatus id="A1508816504AFF8215B0FDF0FBFE0E2C" box="[944,1056,558,584]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
|
||||
are mounted in Fluoromount G on glass slides.
|
||||
</paragraph>
|
||||
, St. 7 (NHMD-213611–213661). All paratypes are mounted in Fluoromount G on glass slides.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216EAFD4DFD890EC9" blockId="28.[189,1398,659,756]" box="[234,599,659,685]" pageId="28" pageNumber="29">
|
||||
<emphasis id="4C9FEAA6504AFF8216EAFD4DFD890EC9" bold="true" box="[234,599,659,685]" pageId="28" pageNumber="29">Additional non-type material</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4504AFF8216BDFD69FAD90E90" blockId="28.[189,1398,659,756]" pageId="28" pageNumber="29">
|
||||
<materialsCitation id="CE833CE9504AFF8216BDFD69FAD90E90" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="06FC7624504AFF8216BDFD69FD880EB5" box="[189,598,695,721]" name="United States of America" pageId="28" pageNumber="29">UNITED STATES OF AMERICA</collectingCountry>
|
||||
: 1 ♂, St. 4;
|
||||
<specimenCount id="68EDFD3D504AFF8214E1FD69FCFA0EB5" box="[737,804,695,721]" pageId="28" pageNumber="29" type="female">3 ♀♀</specimenCount>
|
||||
|
|
@ -176,6 +169,7 @@ are mounted in Fluoromount G on glass slides.
|
|||
, 2 ♂♂, St. 6;
|
||||
<specimenCount id="68EDFD3D504AFF8212AEFD69FB2F0EB5" box="[1198,1265,695,721]" pageId="28" pageNumber="29" type="female">3 ♀♀</specimenCount>
|
||||
, 1 ♂, St. 8. All specimens mounted for SEM and stored in the first author’s personal reference collection.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F504AFFBD16BDFCC2FCD70A1B" lastPageId="35" lastPageNumber="36" pageId="28" pageNumber="29" type="description">
|
||||
|
|
@ -198,9 +192,13 @@ A–B, 12A–B, 13A–B). The trunk appears stout in the anterior end, and turns
|
|||
<paragraph id="7E5436B4504AFF8216BDFBC5FC5E0AF6" blockId="28.[189,1399,1051,1682]" pageId="28" pageNumber="29">
|
||||
The head consists of a retractable mouth cone and an introvert (
|
||||
<figureCitation id="E6D02A31504AFF8215CBFBC5FBF60851" box="[971,1064,1051,1077]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="28" pageNumber="29">Figs 11</figureCitation>
|
||||
, 13C). Inner oral styles are present, but their exact number and arrangement could not be determined. The external mouth cone armature consists of nine outer oral styles, each with five basal fringe tips. The introvert sectors are defined by the ten primary spinoscalids in Ring 01. Each primary spinoscalid consists of a basal sheath and a distal end piece with a blunt tip. The basal sheaths have marginal extensions forming two layers of transverse fringes. End pieces are smooth. Rings 02 and 04 have 10 spinoscalids and Ring 03 has 20 (
|
||||
,
|
||||
<figureCitation id="E6D02A31504AFF821238FBC5FBB20851" box="[1080,1132,1051,1077]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="28" pageNumber="29">13C</figureCitation>
|
||||
). Inner oral styles are present, but their exact number and arrangement could not be determined. The external mouth cone armature consists of nine outer oral styles, each with five basal fringe tips. The introvert sectors are defined by the ten primary spinoscalids in Ring 01. Each primary spinoscalid consists of a basal sheath and a distal end piece with a blunt tip. The basal sheaths have marginal extensions forming two layers of transverse fringes. End pieces are smooth. Rings 02 and 04 have 10 spinoscalids and Ring 03 has 20 (
|
||||
<figureCitation id="E6D02A31504AFF8216C5FB2EFEFF096E" box="[197,289,1264,1290]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="28" pageNumber="29">Figs 11</figureCitation>
|
||||
, 13C). All spinoscalids in these rings consist of a basal sheath and a pointed end piece. The basal sheaths terminate in fringed margins in spinoscalids of Rings 02 and 04, whereas the sheaths of Ring 03 have a median spike only. Ring 05 has 20 spinoscalids composed as those in the preceding ring, but with shorter end pieces. A ring of short fringes extends around the introvert in between spinoscalid Rings 04 and 05. Rings 06 and 07 have only 6 spinoscalids each. These scalids are considerably shorter than preceding ones. Well-developed trichoscalids, attached to trichoscalid plates, are present in sectors 2, 4, 5, 7, 8 and 10. Described sector-wise, sectors 1 and 6 are similar, having spinoscalids arranged as two double diamonds. Sectors 2, 4, 8 and 10 all have spinoscalids arranged as a quincunx, located in between a medial anterior spinoscalid (Ring 02) and a trichoscalid plate. The remaining sectors, 3, 5, 7 and 9, are almost identical, having spinoscalids forming a set of double diamonds located anterior to a pair of Ring 07 spinoscalids. However, in sectors 5 and 7, one Ring 07 spinoscalid is missing, due to the presence of a trichoscalid plate that takes up space (
|
||||
,
|
||||
<figureCitation id="E6D02A31504AFF821730FB2EFEBA096E" box="[304,356,1264,1290]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="28" pageNumber="29">13C</figureCitation>
|
||||
). All spinoscalids in these rings consist of a basal sheath and a pointed end piece. The basal sheaths terminate in fringed margins in spinoscalids of Rings 02 and 04, whereas the sheaths of Ring 03 have a median spike only. Ring 05 has 20 spinoscalids composed as those in the preceding ring, but with shorter end pieces. A ring of short fringes extends around the introvert in between spinoscalid Rings 04 and 05. Rings 06 and 07 have only 6 spinoscalids each. These scalids are considerably shorter than preceding ones. Well-developed trichoscalids, attached to trichoscalid plates, are present in sectors 2, 4, 5, 7, 8 and 10. Described sector-wise, sectors 1 and 6 are similar, having spinoscalids arranged as two double diamonds. Sectors 2, 4, 8 and 10 all have spinoscalids arranged as a quincunx, located in between a medial anterior spinoscalid (Ring 02) and a trichoscalid plate. The remaining sectors, 3, 5, 7 and 9, are almost identical, having spinoscalids forming a set of double diamonds located anterior to a pair of Ring 07 spinoscalids. However, in sectors 5 and 7, one Ring 07 spinoscalid is missing, due to the presence of a trichoscalid plate that takes up space (
|
||||
<figureCitation id="E6D02A31504AFF82151CF9A6FCAC0AF6" box="[796,882,1656,1682]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
|
|
@ -216,9 +214,13 @@ in width at their bases. The trichoscalid plates are well-developed and nearly t
|
|||
<paragraph id="7E5436B4504AFF8216BDF89CFAA90B8F" blockId="28.[189,1399,1858,2027]" pageId="28" pageNumber="29">
|
||||
Segment 1 consists of a complete cuticular ring. Sensory spots are located on the anterior half of the segment, but not at the anterior margin, in subdorsal positions (
|
||||
<figureCitation id="E6D02A31504AFF8215A0F8B8FBCF0BE4" box="[928,1041,1894,1920]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="28" pageNumber="29">Figs 10A</figureCitation>
|
||||
, 13D); sensory spots are small and rounded. Glandular cell outlets
|
||||
,
|
||||
<figureCitation id="E6D02A31504AFF82121CF8B8FB8C0BE4" box="[1052,1106,1894,1920]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="28" pageNumber="29">13D</figureCitation>
|
||||
); sensory spots are small and rounded. Glandular cell outlets
|
||||
<typeStatus id="A1508816504AFF821466F854FD470BC0" box="[614,665,1930,1956]" pageId="28" pageNumber="29">type</typeStatus>
|
||||
1 present in middorsal (Fig. 13D) and sublateral positions. In most specimens the segment is completely devoid of cuticular hairs; however, a few specimens had a single transverse row with well-spaced hairs. The posterior segment margin is straight on the dorsal
|
||||
1 present in middorsal (
|
||||
<figureCitation id="E6D02A31504AFF8215BBF857FBF80BC0" box="[955,1062,1929,1956]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="28" pageNumber="29">Fig. 13D</figureCitation>
|
||||
) and sublateral positions. In most specimens the segment is completely devoid of cuticular hairs; however, a few specimens had a single transverse row with well-spaced hairs. The posterior segment margin is straight on the dorsal
|
||||
</paragraph>
|
||||
<caption id="2A94663C504BFF8316BDF968FE270B8F" ID-DOI="http://doi.org/10.5281/zenodo.3817843" ID-Zenodo-Dep="3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1718,1744]" targetBox="[287,1297,269,1696]" targetPageId="29">
|
||||
<paragraph id="7E5436B4504BFF8316BDF968FE270B8F" blockId="29.[189,1399,1717,2027]" pageId="29" pageNumber="30">
|
||||
|
|
@ -259,7 +261,7 @@ Light micrographs showing overviews and details of
|
|||
<emphasis id="4C9FEAA65049FF81158AF907FB450A97" box="[906,1179,1753,1779]" italics="true" pageId="31" pageNumber="32">Echinoderes hviidarum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="57AC57DD5049FF8112A1F907FB230A97" box="[1185,1277,1753,1779]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635).
|
||||
, holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD-213635).
|
||||
<emphasis id="4C9FEAA65049FF81172EF8C1FE9D0B5D" bold="true" box="[302,323,1823,1849]" pageId="31" pageNumber="32">A</emphasis>
|
||||
. Dorsal overview.
|
||||
<emphasis id="4C9FEAA65049FF811425F8C1FDE70B5D" bold="true" box="[549,569,1823,1849]" pageId="31" pageNumber="32">B</emphasis>
|
||||
|
|
@ -485,7 +487,9 @@ Measurements from light microscopy of
|
|||
</paragraph>
|
||||
<paragraph id="7E5436B45076FFBE16BDFA7EFE3D09B9" blockId="32.[189,1398,1440,1501]" pageId="32" pageNumber="33">and ventral sides, but with sublateral concave indentations, terminating in a pectinate fringe with short, triangular saw-tooth tips.</paragraph>
|
||||
<paragraph id="7E5436B45076FFBE16BDF9DBFC420B38" blockId="32.[189,1399,1541,1884]" pageId="32" pageNumber="33">
|
||||
Segment 2 consists of a complete cuticular ring. Pachycyclus of the anterior segment margin is relatively thin, middorsally interrupted. Sensory spots are located in middorsal (Fig. 13D), midlateral and ventromedial positions (
|
||||
Segment 2 consists of a complete cuticular ring. Pachycyclus of the anterior segment margin is relatively thin, middorsally interrupted. Sensory spots are located in middorsal (
|
||||
<figureCitation id="E6D02A315076FFBE127AF9F6FB360A26" box="[1146,1256,1576,1602]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="32" pageNumber="33">Fig. 13D</figureCitation>
|
||||
), midlateral and ventromedial positions (
|
||||
<figureCitation id="E6D02A315076FFBE1407F995FD840A01" box="[519,602,1611,1637]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="32" pageNumber="33">Fig. 10</figureCitation>
|
||||
A–B). Sensory spots very minute, but with rather long, thin marginal cuticular hairs; all sensory spots from this segment to segment 10 have the same appearance. Glandular cell outlets
|
||||
<typeStatus id="A15088165076FFBE174AF94CFEA30AC8" box="[330,381,1682,1708]" pageId="32" pageNumber="33">type</typeStatus>
|
||||
|
|
@ -494,41 +498,7 @@ A–B). Sensory spots very minute, but with rather long, thin marginal cuticular
|
|||
<paragraph id="7E5436B45076FFBF16BDF85DFE2C0B8F" blockId="32.[189,1399,1923,2020]" lastBlockId="33.[189,1399,1578,2027]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||||
Segment 3, and remaining segments, consist of one tergal and two sternal plates (
|
||||
<figureCitation id="E6D02A315076FFBE1291F85DFB350BF9" box="[1169,1259,1923,1949]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="32" pageNumber="33">Figs 10</figureCitation>
|
||||
A–B, 12B). Pachycyclus of the anterior segment margin of medium thickness, interrupted at the tergosternal junctions and middorsally. No conspicuous cuticular structures present. Fairly long hairs are arranged in two to
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF9F4FECC0A20" bold="true" box="[189,274,1578,1604]" pageId="33" pageNumber="34">Fig. 13</emphasis>
|
||||
(opposite page). Scanning electron micrographs showing overviews and details of
|
||||
<taxonomicName id="B9EB4D375077FFBF12E5F9F4FEE80A0C" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="33" pageNumber="34" phylum="Cephalorhyncha" rank="species" species="hviidarum" status="sp. nov.">
|
||||
<emphasis id="4C9FEAA65077FFBF12E5F9F4FEE80A0C" italics="true" pageId="33" pageNumber="34">Echinoderes hviidarum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="57AC57DD5077FFBF1745F990FE790A0C" box="[325,423,1614,1640]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="4C9FEAA65077FFBF17B6F993FE150A03" bold="true" box="[438,459,1613,1639]" pageId="33" pageNumber="34">A</emphasis>
|
||||
. Dorsal overview.
|
||||
<emphasis id="4C9FEAA65077FFBF14C2F990FD080A0C" bold="true" box="[706,726,1614,1640]" pageId="33" pageNumber="34">B</emphasis>
|
||||
. Ventral overview.
|
||||
<emphasis id="4C9FEAA65077FFBF15D1F993FC380A03" bold="true" box="[977,998,1613,1639]" pageId="33" pageNumber="34">C</emphasis>
|
||||
. Introvert sector 6 (middorsal).
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF9AFFF0C0AEF" bold="true" box="[189,210,1649,1675]" pageId="33" pageNumber="34">D</emphasis>
|
||||
. Segments 1 to 3, dorsal view.
|
||||
<emphasis id="4C9FEAA65077FFBF1469F9AFFDA30AEF" bold="true" box="[617,637,1649,1675]" pageId="33" pageNumber="34">E</emphasis>
|
||||
. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8.
|
||||
<emphasis id="4C9FEAA65077FFBF1435F94AFD990ACA" bold="true" box="[565,583,1684,1710]" pageId="33" pageNumber="34">F</emphasis>
|
||||
. Detail showing the typical stout laterodorsal tube on segment 9.
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF969FF0A0AB5" bold="true" box="[189,212,1719,1745]" pageId="33" pageNumber="34">G</emphasis>
|
||||
. Detail showing laterodorsal tube on segment 10, with associated papillary structure.
|
||||
<emphasis id="4C9FEAA65077FFBF12BFF969FB080AB5" bold="true" box="[1215,1238,1719,1745]" pageId="33" pageNumber="34">H</emphasis>
|
||||
.
|
||||
<specimenCount id="68EDFD3D5077FFBF12E5F966FB250AB6" box="[1253,1275,1720,1746]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
|
||||
, segments 10 to 11, ventral view.
|
||||
<emphasis id="4C9FEAA65077FFBF17DEF905FE370A91" bold="true" box="[478,489,1755,1781]" pageId="33" pageNumber="34">I</emphasis>
|
||||
. Detail showing the less common slender laterodorsal tube on segment 9.
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF920FF120B7C" bold="true" box="[189,204,1790,1816]" pageId="33" pageNumber="34">J</emphasis>
|
||||
. ♂, segments 10 to 11, dorsal view.
|
||||
<emphasis id="4C9FEAA65077FFBF1478F920FD510B7C" bold="true" box="[632,655,1790,1816]" pageId="33" pageNumber="34">K</emphasis>
|
||||
.
|
||||
<specimenCount id="68EDFD3D5077FFBF149EF920FD6A0B7C" box="[670,692,1790,1816]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
|
||||
, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet
|
||||
<typeStatus id="A15088165077FFBF16BDF8B6FF2E0BE6" box="[189,240,1896,1922]" pageId="33" pageNumber="34">type</typeStatus>
|
||||
1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot.
|
||||
A–B, 12B). Pachycyclus of the anterior segment margin of medium thickness, interrupted at the tergosternal junctions and middorsally. No conspicuous cuticular structures present. Fairly long hairs are arranged in two to.
|
||||
</paragraph>
|
||||
<caption id="2A94663C5077FFBF16BDFED2FDFB0D9C" ID-Table-UUID="2A94663C5077FFBF16BDFED2FDFB0D9C" httpUri="http://table.plazi.org/id/2A94663C5077FFBF16BDFED2FDFB0D9C" pageId="33" pageNumber="34" targetBox="[189,1399,557,1482]" targetIsTable="true" targetPageId="33">
|
||||
<paragraph id="7E5436B45077FFBF16BDFED2FDFB0D9C" blockId="33.[189,1399,267,504]" pageId="33" pageNumber="34">
|
||||
|
|
@ -612,6 +582,44 @@ Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD
|
|||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<caption id="2A94663C5077FFBF16BDF9F4FE310B8F" ID-DOI="http://doi.org/10.5281/zenodo.14897716" ID-Zenodo-Dep="14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,232,1578,1604]" subCaptionStartIDs="33.[1082,1253,1790,1816]" subCaptionStarts="Abbr" targetBox="[252,1336,267,2020]" targetPageId="34">
|
||||
<paragraph id="7E5436B45077FFBF16BDF9F4FE310B8F" blockId="33.[189,1399,1578,2027]" pageId="33" pageNumber="34">
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF9F4FECC0A20" bold="true" box="[189,274,1578,1604]" pageId="33" pageNumber="34">Fig. 13</emphasis>
|
||||
(opposite page). Scanning electron micrographs showing overviews and details of
|
||||
<taxonomicName id="B9EB4D375077FFBF12E5F9F4FEE80A0C" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="33" pageNumber="34" phylum="Cephalorhyncha" rank="species" species="hviidarum" status="sp. nov.">
|
||||
<emphasis id="4C9FEAA65077FFBF12E5F9F4FEE80A0C" italics="true" pageId="33" pageNumber="34">Echinoderes hviidarum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="57AC57DD5077FFBF1745F990FE790A0C" box="[325,423,1614,1640]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="4C9FEAA65077FFBF17B6F993FE150A03" bold="true" box="[438,459,1613,1639]" pageId="33" pageNumber="34">A</emphasis>
|
||||
. Dorsal overview.
|
||||
<emphasis id="4C9FEAA65077FFBF14C2F990FD080A0C" bold="true" box="[706,726,1614,1640]" pageId="33" pageNumber="34">B</emphasis>
|
||||
. Ventral overview.
|
||||
<emphasis id="4C9FEAA65077FFBF15D1F993FC380A03" bold="true" box="[977,998,1613,1639]" pageId="33" pageNumber="34">C</emphasis>
|
||||
. Introvert sector 6 (middorsal).
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF9AFFF0C0AEF" bold="true" box="[189,210,1649,1675]" pageId="33" pageNumber="34">D</emphasis>
|
||||
. Segments 1 to 3, dorsal view.
|
||||
<emphasis id="4C9FEAA65077FFBF1469F9AFFDA30AEF" bold="true" box="[617,637,1649,1675]" pageId="33" pageNumber="34">E</emphasis>
|
||||
. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8.
|
||||
<emphasis id="4C9FEAA65077FFBF1435F94AFD990ACA" bold="true" box="[565,583,1684,1710]" pageId="33" pageNumber="34">F</emphasis>
|
||||
. Detail showing the typical stout laterodorsal tube on segment 9.
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF969FF0A0AB5" bold="true" box="[189,212,1719,1745]" pageId="33" pageNumber="34">G</emphasis>
|
||||
. Detail showing laterodorsal tube on segment 10, with associated papillary structure.
|
||||
<emphasis id="4C9FEAA65077FFBF12BFF969FB080AB5" bold="true" box="[1215,1238,1719,1745]" pageId="33" pageNumber="34">H</emphasis>
|
||||
.
|
||||
<specimenCount id="68EDFD3D5077FFBF12E5F966FB250AB6" box="[1253,1275,1720,1746]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
|
||||
, segments 10 to 11, ventral view.
|
||||
<emphasis id="4C9FEAA65077FFBF17DEF905FE370A91" bold="true" box="[478,489,1755,1781]" pageId="33" pageNumber="34">I</emphasis>
|
||||
. Detail showing the less common slender laterodorsal tube on segment 9.
|
||||
<emphasis id="4C9FEAA65077FFBF16BDF920FF120B7C" bold="true" box="[189,204,1790,1816]" pageId="33" pageNumber="34">J</emphasis>
|
||||
. ♂, segments 10 to 11, dorsal view.
|
||||
<emphasis id="4C9FEAA65077FFBF1478F920FD510B7C" bold="true" box="[632,655,1790,1816]" pageId="33" pageNumber="34">K</emphasis>
|
||||
.
|
||||
<specimenCount id="68EDFD3D5077FFBF149EF920FD6A0B7C" box="[670,692,1790,1816]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
|
||||
, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet
|
||||
<typeStatus id="A15088165077FFBF16BDF8B6FF2E0BE6" box="[189,240,1896,1922]" pageId="33" pageNumber="34">type</typeStatus>
|
||||
1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="7E5436B45075FFBD16BDFED5FC4E0D2E" blockId="35.[189,1398,267,330]" pageId="35" pageNumber="36">
|
||||
Glandular cell outlets
|
||||
<typeStatus id="A15088165075FFBD17CEFED2FDDF0D42" box="[462,513,268,294]" pageId="35" pageNumber="36">type</typeStatus>
|
||||
|
|
@ -637,7 +645,9 @@ A–B, 12A–D, 13A). Minute tubes present in lateral accessory positions (
|
|||
<figureCitation id="E6D02A315075FFBD152DFDC1FC430E5D" box="[813,925,543,569]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10B</figureCitation>
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD15ADFDC1FC3F0E5D" box="[941,993,543,569]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">12D</figureCitation>
|
||||
, 13E). Middorsal spine is usually long, reaching the terminal segment (
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD15F0FDC1FBFD0E5D" box="[1008,1059,543,569]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13E</figureCitation>
|
||||
). Middorsal spine is usually long, reaching the terminal segment (
|
||||
<figureCitation id="E6D02A315075FFBD147BFD9CFD300E39" box="[635,750,578,605]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10A</figureCitation>
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD14FBFD9DFCEC0E39" box="[763,818,579,605]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">12A</figureCitation>
|
||||
|
|
@ -658,11 +668,19 @@ Segment 9 with acicular spines in lateroventral positions (
|
|||
<figureCitation id="E6D02A315075FFBD1736FD2EFE780F6E" box="[310,422,752,778]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10A</figureCitation>
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD17B2FD2EFE3B0F6E" box="[434,485,752,778]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">12C</figureCitation>
|
||||
, 13A, F, I); in most specimens (including the
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD17F1FD2EFD840F6F" box="[497,602,752,779]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13A, F, I</figureCitation>
|
||||
); in most specimens (including the
|
||||
<typeStatus id="A15088165075FFBD15F9FD2EFBBC0F6F" box="[1017,1122,752,779]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
|
||||
) the tubes are short and stout (Fig. 13F), without differentiated bases or lateral wings, but a few specimens showed longer and more slender tubes, but still without bases or wings (Fig. 13I). The dimorphism is not related to gender or locality. Sensory spots present in paradorsal (
|
||||
) the tubes are short and stout (
|
||||
<figureCitation id="E6D02A315075FFBD1707FCCAFEB10F4A" box="[263,367,788,814]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13F</figureCitation>
|
||||
), without differentiated bases or lateral wings, but a few specimens showed longer and more slender tubes, but still without bases or wings (
|
||||
<figureCitation id="E6D02A315075FFBD1528FCE6FC540F36" box="[808,906,824,850]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13I</figureCitation>
|
||||
). The dimorphism is not related to gender or locality. Sensory spots present in paradorsal (
|
||||
<figureCitation id="E6D02A315075FFBD14FDFC82FCB70F12" box="[765,873,860,886]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">Fig. 12C</figureCitation>
|
||||
), laterodorsal (Fig. 13F, I) and ventrolateral (
|
||||
), laterodorsal (
|
||||
<figureCitation id="E6D02A315075FFBD121DFC82FB420F12" box="[1053,1180,860,886]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13F, I</figureCitation>
|
||||
) and ventrolateral (
|
||||
<figureCitation id="E6D02A315075FFBD16C6FCA1FEEB0FFE" box="[198,309,895,922]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">Fig. 12D</figureCitation>
|
||||
) positions, and glandular cell outlets type
|
||||
<quantity id="B9139B515075FFBD1541FC5EFCAC0FFD" box="[833,882,895,922]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1 in</quantity>
|
||||
|
|
@ -673,20 +691,30 @@ paradorsal positions. Small rounded sieve plates are present in sublateral posit
|
|||
<paragraph id="7E5436B45075FFBD16BDFBD7FB010926" blockId="35.[189,1399,1033,1346]" pageId="35" pageNumber="36">
|
||||
Segment 10 with laterodorsal tubes near posterior segment margin (
|
||||
<figureCitation id="E6D02A315075FFBD15F4FBD7FB510840" box="[1012,1167,1033,1060]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10A, D</figureCitation>
|
||||
, 13G, J); tubes are very minute and sometimes appear to have an even shorter, protruding papillary structure located next to them (Fig. 13G), but due to their minute size, their presence was often hard to confirm. Sensory spots present in ventrolateral positions (
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD129EFBD7FB2E0840" box="[1182,1264,1033,1060]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13G, J</figureCitation>
|
||||
); tubes are very minute and sometimes appear to have an even shorter, protruding papillary structure located next to them (
|
||||
<figureCitation id="E6D02A315075FFBD172DFB8FFE42080F" box="[301,412,1105,1131]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13G</figureCitation>
|
||||
), but due to their minute size, their presence was often hard to confirm. Sensory spots present in ventrolateral positions (
|
||||
<figureCitation id="E6D02A315075FFBD14A2FBABFCE708EB" box="[674,825,1141,1167]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10B, E</figureCitation>
|
||||
, 13H). Glandular cell outlets
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD1549FBABFC5E08EB" box="[841,896,1141,1167]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13H</figureCitation>
|
||||
). Glandular cell outlets
|
||||
<typeStatus id="A15088165075FFBD12A7FBABFB0408EB" box="[1191,1242,1141,1167]" pageId="35" pageNumber="36">type</typeStatus>
|
||||
1 present as two middorsal ones; other glandular cell outlets
|
||||
<typeStatus id="A15088165075FFBD1500FB47FCED08D7" box="[768,819,1177,1203]" pageId="35" pageNumber="36">type</typeStatus>
|
||||
1 not observed. Segment almost devoid of hairs (Fig. 13H, K), except for a few short ones on the lateral sides. The posterior segment margin of the tergal plate is straight, with rather shorter fringe tips than those on the tergal plate. The margins of the sternal plates have considerably longer fringe tips and extend midventrally to a point that almost reaches the posterior margin of the terminal segment (Fig. 13H). Pachycycli as on preceding segment.
|
||||
1 not observed. Segment almost devoid of hairs (
|
||||
<figureCitation id="E6D02A315075FFBD16C6FB62FE8D08B3" box="[198,339,1212,1239]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13H, K</figureCitation>
|
||||
), except for a few short ones on the lateral sides. The posterior segment margin of the tergal plate is straight, with rather shorter fringe tips than those on the tergal plate. The margins of the sternal plates have considerably longer fringe tips and extend midventrally to a point that almost reaches the posterior margin of the terminal segment (
|
||||
<figureCitation id="E6D02A315075FFBD14ACFAF6FCC40926" box="[684,794,1320,1346]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">Fig. 13H</figureCitation>
|
||||
). Pachycycli as on preceding segment.
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B45075FFBD16BDFAB4FCD70A1B" blockId="35.[189,1399,1386,1663]" pageId="35" pageNumber="36">
|
||||
Segment 11 with lateral terminal spines (
|
||||
<figureCitation id="E6D02A315075FFBD1493FAB4FD3B09E0" box="[659,741,1386,1412]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||||
A–C). Males with three pairs of penile spines (
|
||||
<figureCitation id="E6D02A315075FFBD12FCFAB4FA8E09E0" box="[1276,1360,1386,1412]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10</figureCitation>
|
||||
A– B, 12F, 13J); dorsal and ventral penile spines are thin, flexible tubes, whereas the median ones are thicker, conical, and stout; females with short lateral terminal accessory spines (
|
||||
AB, 12F, 13J); dorsal and ventral penile spines are thin, flexible tubes, whereas the median ones are thicker, conical, and stout; females with short lateral terminal accessory spines (
|
||||
<figureCitation id="E6D02A315075FFBD1258FA6FFB7009A8" box="[1112,1198,1457,1484]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10</figureCitation>
|
||||
D–E, 12G, 13H). Sensory spots without long marginal hairs present in paradorsal positions. Glandular cell outlets
|
||||
<typeStatus id="A15088165075FFBD1344FA08FAA90994" box="[1348,1399,1494,1520]" pageId="35" pageNumber="36">type</typeStatus>
|
||||
|
|
@ -694,11 +722,15 @@ D–E, 12G, 13H). Sensory spots without long marginal hairs present in paradorsa
|
|||
<figureCitation id="E6D02A315075FFBD12DAFA27FAAC0A70" box="[1242,1394,1529,1556]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10A, D</figureCitation>
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD16BDF9C3FF310A53" box="[189,239,1565,1591]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">12E</figureCitation>
|
||||
, 13J–K). The segment is completely devoid of cuticular hairs, but densely covered with short hairlike extensions, in middorsal to laterodorsal areas. Tergal extensions are short and pointed, with an extra tooth at the inferior margin (
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD16FBF9C3FEC90A53" box="[251,279,1565,1591]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13</figureCitation>
|
||||
J–K). The segment is completely devoid of cuticular hairs, but densely covered with short hairlike extensions, in middorsal to laterodorsal areas. Tergal extensions are short and pointed, with an extra tooth at the inferior margin (
|
||||
<figureCitation id="E6D02A315075FFBD140AF9BBFDBB0A1B" box="[522,613,1637,1663]" captionStart="Fig" captionStartId="29.[189,232,1718,1744]" captionTargetBox="[287,1297,269,1696]" captionTargetId="figure@29.[286,1301,265,1696]" captionTargetPageId="29" captionText="Fig. 10. Line art illustrations of Echinoderes hviidarum sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; si = sieve plate; slgco1 = sublateral glandular cell outlet type 1; slss = sublateral sensory spot; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817843" httpUri="https://zenodo.org/record/3817843/files/figure.png" pageId="35" pageNumber="36">Figs 10</figureCitation>
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD1471F9BBFD7E0A1B" box="[625,672,1637,1663]" captionStart="Fig" captionStartId="31.[189,232,1752,1778]" captionTargetBox="[217,1371,265,1740]" captionTargetId="figure@31.[216,1371,265,1740]" captionTargetPageId="31" captionText="Fig. 12. Light micrographs showing overviews and details of Echinoderes hviidarum sp. nov., holotype, ♂ (A–B, E–F) (NHMD-213610), paratype, ♂ (C–D) (NHMD-213614) and paratype, ♀ (G) (NHMD- 213635). A. Dorsal overview. B. Ventral overview. C. Segments 6 to 9, dorsal view. D. Segments 8 to 11, ventral view. E. ♂, segments 10 to 11, dorsal view. F. ♂, segments 10 to 11, ventral view. G. ♀, segments 10 to 11, ventral view. Abbreviations: lat = lateral accessory tube; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; pdss = paradorsal sensory spot; pe = penile spines; pr = protuberance; sdss = subdorsal sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817847" httpUri="https://zenodo.org/record/3817847/files/figure.png" pageId="35" pageNumber="36">12F</figureCitation>
|
||||
, 13J–K).
|
||||
,
|
||||
<figureCitation id="E6D02A315075FFBD14ABF9BBFD190A1B" box="[683,711,1637,1663]" captionStart="Fig" captionStartId="33.[189,232,1578,1604]" captionTargetId="figure@34.[252,1336,265,2020]" captionTargetPageId="34" captionText="Fig. 13 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hviidarum sp. nov. A. Dorsal overview. B. Ventral overview. C. Introvert sector 6 (middorsal). D. Segments 1 to 3, dorsal view. E. Detail showing attachment of lateroventral spine and lateral accessory tube on segment 8. F. Detail showing the typical stout laterodorsal tube on segment 9. G. Detail showing laterodorsal tube on segment 10, with associated papillary structure. H. ♀, segments 10 to 11, ventral view. I. Detail showing the less common slender laterodorsal tube on segment 9. J. ♂, segments 10 to 11, dorsal view. K. ♀, segments 10 to 11, dorsal view. Abbreviations: lat = lateral accessory tube; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mdp = middorsal placid; mds = middorsal spine; mdss = middorsal sensory spot; pap = papillary structure; pe = penile spines; pr = protuberance; psp = primary spinoscalid; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlss = ventrolateral sensory spot" figureDoi="http://doi.org/10.5281/zenodo.14897716" httpUri="https://zenodo.org/record/14897716/files/figure.png" pageId="35" pageNumber="36">13</figureCitation>
|
||||
J–K).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F5075FFBA16BDF979FC6F0E34" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="discussion">
|
||||
|
|
|
|||
|
|
@ -1,61 +1,61 @@
|
|||
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<figureCitation id="E6D02A31505DFF9514A0FE8AFCD70D0A" box="[672,777,340,366]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="12.[189,232,1791,1817]" captionStartId-1="13.[189,232,1683,1709]" captionStartId-2="15.[189,232,1576,1602]" captionTargetBox-0="[261,1323,265,1769]" captionTargetBox-1="[236,1351,265,1655]" captionTargetId-0="figure@12.[260,1327,265,1769]" captionTargetId-1="figure@13.[236,1351,265,1656]" captionTargetId-2="figure@16.[195,1389,265,2019]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="16" captionText-0="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." captionText-1="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." captionText-2="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi-0="http://doi.org/10.5281/zenodo.3817831" figureDoi-1="http://doi.org/10.5281/zenodo.3817833" figureDoi-2="http://doi.org/10.5281/zenodo.3817835" httpUri-0="https://zenodo.org/record/3817831/files/figure.png" httpUri-1="https://zenodo.org/record/3817833/files/figure.png" httpUri-2="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">
|
||||
<figureCitation id="E6D02A31505DFF9514A0FE8AFD370D0A" box="[672,745,340,366]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
|
||||
–6
|
||||
</figureCitation>
|
||||
<figureCitation id="E6D02A31505DFF9514A0FE8AFCD70D0A" box="[672,777,340,366]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="12.[189,232,1791,1817]" captionStartId-1="13.[189,232,1683,1709]" captionStartId-2="15.[189,232,1576,1602]" captionTargetBox-0="[261,1323,265,1769]" captionTargetBox-1="[236,1351,265,1655]" captionTargetId-0="figure@12.[260,1327,265,1769]" captionTargetId-1="figure@13.[236,1351,265,1656]" captionTargetId-2="figure@16.[195,1389,265,2019]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="16" captionText-0="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." captionText-1="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." captionText-2="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi-0="http://doi.org/10.5281/zenodo.3817831" figureDoi-1="http://doi.org/10.5281/zenodo.3817833" figureDoi-2="http://doi.org/10.5281/zenodo.3817835" httpUri-0="https://zenodo.org/record/3817831/files/figure.png" httpUri-1="https://zenodo.org/record/3817833/files/figure.png" httpUri-2="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">Figs 4–6</figureCitation>
|
||||
,
|
||||
<tableCitation id="3369030F505DFF951514FE8AFC4A0D0A" box="[788,916,340,366]" captionStart-0="Table 4" captionStart-1="Table 5" captionStart-2="Table 6" captionStartId-0="15.[189,258,268,294]" captionStartId-1="17.[189,258,268,294]" captionStartId-2="23.[189,258,268,294]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionTargetPageId-2="23" captionText-0="Table 4. Measurements from light microscopy of Echinoderes dubiosus sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 5. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes dubiosus sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco 1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; pf = papillary flap; po = pore; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters." captionText-2="Table 6. Measurements from light microscopy of Echinoderes hamiltonorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." httpUri-0="http://table.plazi.org/id/2A94663C5059FF9116BDFED2FB6F0DD6" pageId="11" pageNumber="12" tableUuid-0="2A94663C5059FF9116BDFED2FB6F0DD6">Tables 4–6</tableCitation>
|
||||
</paragraph>
|
||||
|
|
@ -120,13 +117,11 @@ is derived from the Latin ‘dubium’, because of its very close resemblance wi
|
|||
<emphasis id="4C9FEAA6505DFF9516BDFCA9FE740FF5" bold="true" box="[189,426,887,913]" pageId="11" pageNumber="12">Material examined</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<materialsCitation id="CE833CE9505DFF9516EAFC79FAC50849" ID-GBIF-Occurrence="2611490312" collectingDate="2008-09-20" collectionCode="US" collectorName="St. & Fluoromount G & See" country="United States of America" latitude="39.999443" location="West Coast" longLatPrecision="19" longitude="-125.87417" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
|
||||
<paragraph id="7E5436B4505DFF9516EAFC79FE820FA5" blockId="11.[189,1399,935,1070]" box="[234,348,935,961]" pageId="11" pageNumber="12">
|
||||
<paragraph id="7E5436B4505DFF9516EAFC79FAC50849" blockId="11.[189,1399,935,1070]" pageId="11" pageNumber="12">
|
||||
<materialsCitation id="CE833CE9505DFF9516EAFC79FAC50849" ID-GBIF-Occurrence="2611490312" collectingDate="2008-09-20" collectionCode="US" country="United States of America" latitude="39.999443" location="West Coast" longLatPrecision="19" longitude="-125.87417" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
|
||||
<emphasis id="4C9FEAA6505DFF9516EAFC79FE820FA5" bold="true" box="[234,348,935,961]" pageId="11" pageNumber="12">
|
||||
<typeStatus id="A1508816505DFF9516EAFC79FE820FA5" box="[234,348,935,961]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4505DFF9516BDFC15FAC50849" blockId="11.[189,1399,935,1070]" pageId="11" pageNumber="12">
|
||||
<collectingCountry id="06FC7624505DFF9516BDFC15FDBA0F81" box="[189,612,971,997]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||||
: adult
|
||||
<specimenCount id="68EDFD3D505DFF9514BAFC15FD0E0F81" box="[698,720,971,997]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
|
||||
|
|
@ -153,22 +148,19 @@ is derived from the Latin ‘dubium’, because of its very close resemblance wi
|
|||
for localities and
|
||||
<tableCitation id="3369030F505DFF9515A1FBCDFC260849" box="[929,1016,1043,1069]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="11" pageNumber="12" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
|
||||
for detailed station data.
|
||||
</paragraph>
|
||||
</materialsCitation>
|
||||
<paragraph id="7E5436B4505DFF9516EAFB8BFEB9080B" blockId="11.[189,1399,1109,1244]" box="[234,359,1109,1135]" pageId="11" pageNumber="12">
|
||||
<materialsCitation id="CE833CE9505DFF9516EAFB8BFEB9080B" ID-GBIF-Occurrence="2611490308" box="[234,359,1109,1135]" pageId="11" pageNumber="12" typeStatus="paratype">
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4505DFF9516EAFB8BFBCC08B8" blockId="11.[189,1399,1109,1244]" pageId="11" pageNumber="12">
|
||||
<materialsCitation id="CE833CE9505DFF9516EAFB8BFB4508F7" ID-GBIF-Occurrence="2611490308" pageId="11" pageNumber="12" typeStatus="paratype">
|
||||
<emphasis id="4C9FEAA6505DFF9516EAFB8BFEB9080B" bold="true" box="[234,359,1109,1135]" pageId="11" pageNumber="12">
|
||||
<typeStatus id="A1508816505DFF9516EAFB8BFEB9080B" box="[234,359,1109,1135]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4505DFF9516BDFBA4FBCC08B8" blockId="11.[189,1399,1109,1244]" pageId="11" pageNumber="12">
|
||||
<collectingCountry id="06FC7624505DFF9516BDFBA4FD8508F7" box="[189,603,1145,1172]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||||
:
|
||||
<specimenCount id="68EDFD3D505DFF951468FBA7FD4B08F0" box="[616,661,1145,1172]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||||
, 1 ♂, same collecting data as for
|
||||
<typeStatus id="A1508816505DFF95122DFBA7FB4908F7" box="[1069,1175,1145,1171]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||||
;
|
||||
, 1 ♂, same collecting data as for holotype;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="CE833CE9505DFF9512A4FBA7FBCC08B8" ID-GBIF-Occurrence="2611490308" pageId="11" pageNumber="12" typeStatus="paratype">
|
||||
<specimenCount id="68EDFD3D505DFF9512A4FBA7FB3708F0" box="[1188,1257,1145,1172]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||||
, St. 1;
|
||||
<specimenCount id="68EDFD3D505DFF951341FBA7FAB108F0" box="[1345,1391,1145,1172]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||||
|
|
@ -178,20 +170,21 @@ for detailed station data.
|
|||
<specimenCount id="68EDFD3D505DFF951785FB40FE1608DC" box="[389,456,1182,1208]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||||
, St. 5; 1 ♂, St. 6;
|
||||
<specimenCount id="68EDFD3D505DFF95149FFB40FD3C08DC" box="[671,738,1182,1208]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||||
, 1 ♂, St. 8 (NHMD-213584–213595). All
|
||||
<typeStatus id="A1508816505DFF9512DAFB40FA9408DC" box="[1242,1354,1182,1208]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
|
||||
are mounted in Fluoromount G, nine on glass slides and three on H-S slides.
|
||||
, 1 ♂, St. 8 (NHMD-213584–213595). All paratypes are mounted in Fluoromount G, nine on glass slides and three on H-S slides.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4505DFF9516EAFADAFD89097A" blockId="11.[189,1399,1284,1382]" box="[234,599,1284,1310]" pageId="11" pageNumber="12">
|
||||
<emphasis id="4C9FEAA6505DFF9516EAFADAFD89097A" bold="true" box="[234,599,1284,1310]" pageId="11" pageNumber="12">Additional non-type material</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4505DFF9516BDFAF7FD6A0902" blockId="11.[189,1399,1284,1382]" pageId="11" pageNumber="12">
|
||||
<materialsCitation id="CE833CE9505DFF9516BDFAF7FD6A0902" pageId="11" pageNumber="12">
|
||||
<collectingCountry id="06FC7624505DFF9516BDFAF7FD870926" box="[189,601,1320,1347]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||||
:
|
||||
<specimenCount id="68EDFD3D505DFF951464FAF6FD4F0927" box="[612,657,1320,1347]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||||
, 1 ♂, St. 6;
|
||||
<specimenCount id="68EDFD3D505DFF951524FAF6FC8F0927" box="[804,849,1320,1347]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||||
, St. 7. All mounted for SEM and stored in the first author’s personal reference collection.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F505DFF8F16BDFA51FCBB0AC2" lastPageId="17" lastPageNumber="18" pageId="11" pageNumber="12" type="description">
|
||||
|
|
|
|||
149
data/F6/42/87/F64287A2506AFFA217C0FB36FBA80B8F.xml
Normal file
149
data/F6/42/87/F64287A2506AFFA217C0FB36FBA80B8F.xml
Normal file
|
|
@ -0,0 +1,149 @@
|
|||
<document id="40666D2E88B5B01218ABC21240E71DA7" ID-CLB-Dataset="22296" ID-DOI="10.5852/ejt.2018.456" ID-GBIF-Dataset="0566a11e-d6a3-4283-89c7-c7feea96fdbf" ID-Zenodo-Dep="3817823" ID-ZooBank="DE1B1DEE-9871-4803-9F67-025F2B439560" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1588965964057" checkinUser="valdenar" docAuthor="Sørensen, Martin V., Rohal, Melissa & Thistle, David" docDate="2018" docId="F64287A2506AFFA217C0FB36FBA80B8F" docLanguage="en" docName="ejt-456_sorensen_rohal_thistle.pdf.imf" docOrigin="European Journal of Taxonomy 456" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes hakaiensis Herranz, Yangel & Leander 2017" docType="treatment" docVersion="1" lastPageNumber="61" masterDocId="0A7BFFDA5056FF9E1600FFDEFFDE0C64" masterDocTitle="Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific" masterLastPageNumber="75" masterPageNumber="1" pageNumber="61" updateTime="1740014107622" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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||||
<mods:mods id="4116E281B553DF384F4A1BFF00E2B623" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="377490B364AA66FE16C4983A43684A73">
|
||||
<mods:title id="4894CA8AC5B2A39931078E0A574D6366">Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="B61566032E34225FDD99CED8CAC75862" type="personal">
|
||||
<mods:role id="774B1464B330820913CF02B57BC3C4D3">
|
||||
<mods:roleTerm id="F989844EB8355210EBCD818E8721DDFF">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="C4A794B5AFCCD7CD5C113C137A980915">Sørensen, Martin V.</mods:namePart>
|
||||
<mods:nameIdentifier id="7C3619DC9483045BA09CE5860E707981" type="LSID">4143D650-12FC-4914-93F5-2C39339A7156</mods:nameIdentifier>
|
||||
<mods:affiliation id="990DF7C35749925B77F7B603755DCFEA">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
|
||||
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<mods:namePart id="C0A757A83AFEE453081F59D4549E8871">Rohal, Melissa</mods:namePart>
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<mods:namePart id="FD0F17FBD3159ACEB66F529BED2F7371">Thistle, David</mods:namePart>
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<mods:nameIdentifier id="A5BBBC41868FC376B70C3CE22D166480" type="email">dthistle@fsu.edu</mods:nameIdentifier>
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<mods:title id="402DCF5C7DC6C78BD7C2B2C3B11A2814">European Journal of Taxonomy</mods:title>
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<mods:part id="014522098A8002EA776B1CEA3CB269D0">
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<mods:date id="A48E6F45C4283DED3F99653BC644909E">2018</mods:date>
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<treatment id="F64287A2506AFFA217C0FB36FBA80B8F" ID-DOI="http://doi.org/10.5281/zenodo.3818834" ID-GBIF-Taxon="163993847" ID-Zenodo-Dep="3818834" LSID="urn:lsid:plazi:treatment:F64287A2506AFFA217C0FB36FBA80B8F" httpUri="http://treatment.plazi.org/id/F64287A2506AFFA217C0FB36FBA80B8F" lastPageNumber="61" pageId="60" pageNumber="61" scope_class="Cyclorhagida" scope_family="Echinoderidae" scope_order="Echinorhagata" scope_phylum="Kinorhyncha">
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<heading id="251C81D8506AFFA217C0FB36FBAD0966" box="[448,1139,1256,1283]" centered="true" fontSize="11" level="2" pageId="60" pageNumber="61" reason="2">
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<taxonomicName id="B9EB4D37506AFFA217C0FB36FBAD0966" authority="Herranz, Yangel & Leander, 2017" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[448,1139,1256,1283]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
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<emphasis id="4C9FEAA6506AFFA217C0FB36FD3E0967" bold="true" box="[448,736,1256,1283]" italics="true" pageId="60" pageNumber="61">Echinoderes hakaiensis</emphasis>
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Herranz, Yangel & Leander, 2017
|
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</taxonomicName>
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</paragraph>
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<emphasis id="4C9FEAA6506AFFA216BDFAF5FE740921" bold="true" box="[189,426,1323,1349]" pageId="60" pageNumber="61">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4506AFFA216BDFA83FBDD0981" blockId="60.[189,1399,1370,1509]" pageId="60" pageNumber="61">
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<collectingCountry id="06FC7624506AFFA216BDFA83FD840912" box="[189,602,1372,1399]" name="United States of America" pageId="60" pageNumber="61">UNITED STATES OF AMERICA</collectingCountry>
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:
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<specimenCount id="68EDFD3D506AFFA21466FA82FD660912" box="[614,696,1372,1398]" pageId="60" pageNumber="61" type="adult">1 adult</specimenCount>
|
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♂,
|
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<collectionCode id="18FAAE71506AFFA214E7FA82FCD20912" box="[743,780,1372,1398]" country="South Africa" name="University of Stellenbosch" pageId="60" pageNumber="61">US</collectionCode>
|
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West Coast,
|
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<collectingRegion id="BC2FF856506AFFA215AAFA82FBFA0912" box="[938,1060,1372,1398]" country="United States of America" name="California" pageId="60" pageNumber="61">California</collectingRegion>
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, off Monterey,
|
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<geoCoordinate id="1BDF5073506AFFA212DBFA84FAAC0913" box="[1243,1394,1370,1399]" degrees="36" direction="north" minutes="40" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="52" value="36.681114">36°40′52″ N</geoCoordinate>
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,
|
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<geoCoordinate id="1BDF5073506AFFA216BDFAA1FEB809F8" box="[189,358,1407,1436]" degrees="122" direction="west" minutes="49" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="37" value="-122.82694">122°49′37″ W</geoCoordinate>
|
||||
, St. 6, 2719 m deep,
|
||||
<date id="0A551074506AFFA21465FA5FFCD809FF" box="[613,774,1409,1435]" pageId="60" pageNumber="61" value="2008-09-24">24 Sep. 2008</date>
|
||||
, mounted in Fluoromount G (NHMD-223924); 1 ♂, same collecting data as for preceding, mounted for SEM and stored in the first author’s personal reference collection. See
|
||||
<figureCitation id="E6D02A31506AFFA21771FA15FE680981" box="[369,438,1483,1509]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="60" pageNumber="61">Fig. 1</figureCitation>
|
||||
for localities and
|
||||
<tableCitation id="3369030F506AFFA21489FA15FD3E0981" box="[649,736,1483,1509]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="60" pageNumber="61" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
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for detailed station data.
|
||||
</materialsCitation>
|
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F506AFFA216BDF9D1FBA80B8F" pageId="60" pageNumber="61" type="discussion">
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<emphasis id="4C9FEAA6506AFFA216BDF9D1FBF70A4D" bold="true" box="[189,1065,1551,1577]" pageId="60" pageNumber="61">
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||||
Notes on distribution, morphology and comparison with
|
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<typeStatus id="A1508816506AFFA21582F9D1FC660A4D" box="[898,952,1551,1577]" pageId="60" pageNumber="61">type</typeStatus>
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material
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4506AFFA216BDF9E1FAB90B5C" blockId="60.[189,1398,1599,1848]" pageId="60" pageNumber="61">
|
||||
The two recorded specimens show a very close resemblance with
|
||||
<taxonomicName id="B9EB4D37506AFFA215DFF99EFBA10A3D" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[991,1151,1599,1625]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
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<emphasis id="4C9FEAA6506AFFA215DFF99EFBA10A3D" box="[991,1151,1599,1625]" italics="true" pageId="60" pageNumber="61">E. hakaiensis</emphasis>
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||||
</taxonomicName>
|
||||
, a species described from the coast of
|
||||
<collectingRegion id="BC2FF856506AFFA21788F9BAFD880A1A" box="[392,598,1636,1662]" country="Canada" name="British Columbia" pageId="60" pageNumber="61">British Columbia</collectingRegion>
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,
|
||||
<collectingCountry id="06FC7624506AFFA21460F9BAFD660A1A" box="[608,696,1636,1662]" name="Canada" pageId="60" pageNumber="61">Canada</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="1A7A4B45506AFFA214C6F9BAFC730A1A" author="Herranz M. & Yangel E. & Leander B." box="[710,941,1636,1662]" pageId="60" pageNumber="61" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
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Herranz
|
||||
<emphasis id="4C9FEAA6506AFFA2152DF9BBFCB90A1A" box="[813,871,1636,1662]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
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2018
|
||||
</bibRefCitation>
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||||
). The single specimen mounted for LM has a trunk length of
|
||||
<quantity id="B9139B51506AFFA217BAF954FDC60AC0" box="[442,536,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="60" pageNumber="61" unit="mm" value="320.0">320 µm</quantity>
|
||||
, which is very close to the mean length of the type specimens (
|
||||
<quantity id="B9139B51506AFFA21308F954FAB70AC0" box="[1288,1385,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="3.2399999999999998" pageId="60" pageNumber="61" unit="mm" value="324.0">324 µm</quantity>
|
||||
). Moreover, spine and segment lengths are within the range of the type material. The distribution of spines, tubes, sensory spots and glandular cell outlets correspond in detail to the original description given by
|
||||
<bibRefCitation id="1A7A4B45506AFFA21730F927FDEF0B77" author="Herranz M. & Yangel E. & Leander B." box="[304,561,1785,1811]" pageId="60" pageNumber="61" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
|
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Herranz
|
||||
<emphasis id="4C9FEAA6506AFFA21798F924FE110B77" box="[408,463,1785,1811]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
|
||||
. (2018)
|
||||
</bibRefCitation>
|
||||
, and re-examination of
|
||||
<typeStatus id="A1508816506AFFA21553F927FC1D0B77" box="[851,963,1785,1811]" pageId="60" pageNumber="61" type="paratype">paratypes</typeStatus>
|
||||
deposited at
|
||||
<collectionCode id="18FAAE71506AFFA21267F927FB1F0B77" box="[1127,1217,1785,1811]" pageId="60" pageNumber="61">NHMD</collectionCode>
|
||||
confirmed this compliance. The specimens are not illustrated herein, since no new or different structures were found.
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4506AFFA216BDF8BFFBA80B8F" blockId="60.[189,1399,1889,2027]" pageId="60" pageNumber="61">
|
||||
The only considerable difference regards the habitat.
|
||||
<taxonomicName id="B9EB4D37506AFFA2154BF8BFFBBA0B1F" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[843,1124,1889,1915]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
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<emphasis id="4C9FEAA6506AFFA2154BF8BFFBBA0B1F" box="[843,1124,1889,1915]" italics="true" pageId="60" pageNumber="61">Echinoderes hakaiensis</emphasis>
|
||||
</taxonomicName>
|
||||
was described from a depth of
|
||||
<quantity id="B9139B51506AFFA21724F859FE420BC5" box="[292,412,1927,1953]" metricMagnitude="2" metricUnit="m" metricValue="1.14" metricValueMax="1.4" metricValueMin="0.88" pageId="60" pageNumber="61" unit="m" value="114.0" valueMax="140.0" valueMin="88.0">88–140 m</quantity>
|
||||
, and is thus a part of the shelf or sublittoral fauna. The recorded specimens originate from the bathyal fauna, which suggests that the species is not limited by depth preferences, but can be found along the American west coast, throughout the sublittoral to bathyal zones.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
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</document>
|
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|
|
@ -1,149 +1,63 @@
|
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<mods:affiliation id="52DF2C99F91C6725B03717A75FA92BFE">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:affiliation id="ED3E7CF4D11F4E3B2F7EA99DA214A6CE">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<treatment id="F64287A2506AFFDF17C0FB36FBF20A86" ID-DOI="http://doi.org/10.5281/zenodo.3818834" ID-GBIF-Taxon="163993847" ID-Zenodo-Dep="3818834" LSID="urn:lsid:plazi:treatment:F64287A2506AFFDF17C0FB36FBF20A86" httpUri="http://treatment.plazi.org/id/F64287A2506AFFDF17C0FB36FBF20A86" lastPageId="65" lastPageNumber="66" pageId="60" pageNumber="61">
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<taxonomicName id="B9EB4D37506AFFA217C0FB36FBAD0966" authority="Herranz, Yangel & Leander, 2017" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[448,1139,1256,1283]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
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<emphasis id="4C9FEAA6506AFFA217C0FB36FD3E0967" bold="true" box="[448,736,1256,1283]" italics="true" pageId="60" pageNumber="61">Echinoderes hakaiensis</emphasis>
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Herranz, Yangel & Leander, 2017
|
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4506AFFA216BDFAF5FE740921" blockId="60.[189,426,1323,1349]" box="[189,426,1323,1349]" pageId="60" pageNumber="61">
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<heading id="251C81D8506AFFA216BDFAF5FE740921" bold="true" box="[189,426,1323,1349]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
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<emphasis id="4C9FEAA6506AFFA216BDFAF5FE740921" bold="true" box="[189,426,1323,1349]" pageId="60" pageNumber="61">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4506AFFA216BDFA83FBDD0981" blockId="60.[189,1399,1370,1509]" pageId="60" pageNumber="61">
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<materialsCitation id="CE833CE9506AFFA216BDFA83FBDD0981" ID-GBIF-Occurrence="2611490306" pageId="60" pageNumber="61">
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<collectingCountry id="06FC7624506AFFA216BDFA83FD840912" box="[189,602,1372,1399]" name="United States of America" pageId="60" pageNumber="61">UNITED STATES OF AMERICA</collectingCountry>
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:
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♂,
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<collectionCode id="18FAAE71506AFFA214E7FA82FCD20912" box="[743,780,1372,1398]" country="South Africa" name="University of Stellenbosch" pageId="60" pageNumber="61">US</collectionCode>
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West Coast,
|
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<collectingRegion id="BC2FF856506AFFA215AAFA82FBFA0912" box="[938,1060,1372,1398]" country="United States of America" name="California" pageId="60" pageNumber="61">California</collectingRegion>
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, off Monterey,
|
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<geoCoordinate id="1BDF5073506AFFA212DBFA84FAAC0913" box="[1243,1394,1370,1399]" degrees="36" direction="north" minutes="40" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="52" value="36.681114">36°40′52″ N</geoCoordinate>
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,
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<geoCoordinate id="1BDF5073506AFFA216BDFAA1FEB809F8" box="[189,358,1407,1436]" degrees="122" direction="west" minutes="49" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="37" value="-122.82694">122°49′37″ W</geoCoordinate>
|
||||
, St. 6, 2719 m deep,
|
||||
<date id="0A551074506AFFA21465FA5FFCD809FF" box="[613,774,1409,1435]" pageId="60" pageNumber="61" value="2008-09-24">24 Sep. 2008</date>
|
||||
, mounted in Fluoromount G (NHMD-223924); 1 ♂, same collecting data as for preceding, mounted for SEM and stored in the first author’s personal reference collection. See
|
||||
<figureCitation id="E6D02A31506AFFA21771FA15FE680981" box="[369,438,1483,1509]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="60" pageNumber="61">Fig. 1</figureCitation>
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for localities and
|
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<tableCitation id="3369030F506AFFA21489FA15FD3E0981" box="[649,736,1483,1509]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="60" pageNumber="61" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
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for detailed station data.
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||||
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<heading id="251C81D8506AFFA216BDF9D1FBF70A4D" bold="true" box="[189,1065,1551,1577]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
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<emphasis id="4C9FEAA6506AFFA216BDF9D1FBF70A4D" bold="true" box="[189,1065,1551,1577]" pageId="60" pageNumber="61">
|
||||
Notes on distribution, morphology and comparison with
|
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<typeStatus id="A1508816506AFFA21582F9D1FC660A4D" box="[898,952,1551,1577]" pageId="60" pageNumber="61">type</typeStatus>
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material
|
||||
</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4506AFFA216BDF9E1FAB90B5C" blockId="60.[189,1398,1599,1848]" pageId="60" pageNumber="61">
|
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The two recorded specimens show a very close resemblance with
|
||||
<taxonomicName id="B9EB4D37506AFFA215DFF99EFBA10A3D" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[991,1151,1599,1625]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||||
<emphasis id="4C9FEAA6506AFFA215DFF99EFBA10A3D" box="[991,1151,1599,1625]" italics="true" pageId="60" pageNumber="61">E. hakaiensis</emphasis>
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||||
</taxonomicName>
|
||||
, a species described from the coast of
|
||||
<collectingRegion id="BC2FF856506AFFA21788F9BAFD880A1A" box="[392,598,1636,1662]" country="Canada" name="British Columbia" pageId="60" pageNumber="61">British Columbia</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="06FC7624506AFFA21460F9BAFD660A1A" box="[608,696,1636,1662]" name="Canada" pageId="60" pageNumber="61">Canada</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="1A7A4B45506AFFA214C6F9BAFC730A1A" author="Herranz M. & Yangel E. & Leander B." box="[710,941,1636,1662]" pageId="60" pageNumber="61" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
|
||||
Herranz
|
||||
<emphasis id="4C9FEAA6506AFFA2152DF9BBFCB90A1A" box="[813,871,1636,1662]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
). The single specimen mounted for LM has a trunk length of
|
||||
<quantity id="B9139B51506AFFA217BAF954FDC60AC0" box="[442,536,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="60" pageNumber="61" unit="mm" value="320.0">320 µm</quantity>
|
||||
, which is very close to the mean length of the type specimens (
|
||||
<quantity id="B9139B51506AFFA21308F954FAB70AC0" box="[1288,1385,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="3.2399999999999998" pageId="60" pageNumber="61" unit="mm" value="324.0">324 µm</quantity>
|
||||
). Moreover, spine and segment lengths are within the range of the type material. The distribution of spines, tubes, sensory spots and glandular cell outlets correspond in detail to the original description given by
|
||||
<bibRefCitation id="1A7A4B45506AFFA21730F927FDEF0B77" author="Herranz M. & Yangel E. & Leander B." box="[304,561,1785,1811]" pageId="60" pageNumber="61" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
|
||||
Herranz
|
||||
<emphasis id="4C9FEAA6506AFFA21798F924FE110B77" box="[408,463,1785,1811]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
|
||||
. (2018)
|
||||
</bibRefCitation>
|
||||
, and re-examination of
|
||||
<typeStatus id="A1508816506AFFA21553F927FC1D0B77" box="[851,963,1785,1811]" pageId="60" pageNumber="61" type="paratype">paratypes</typeStatus>
|
||||
deposited at
|
||||
<collectionCode id="18FAAE71506AFFA21267F927FB1F0B77" box="[1127,1217,1785,1811]" pageId="60" pageNumber="61">NHMD</collectionCode>
|
||||
confirmed this compliance. The specimens are not illustrated herein, since no new or different structures were found.
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4506AFFA216BDF8BFFBA80B8F" blockId="60.[189,1399,1889,2027]" pageId="60" pageNumber="61">
|
||||
The only considerable difference regards the habitat.
|
||||
<taxonomicName id="B9EB4D37506AFFA2154BF8BFFBBA0B1F" authorityName="Herranz, Yangel & Leander" authorityYear="2017" box="[843,1124,1889,1915]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="60" pageNumber="61" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||||
<emphasis id="4C9FEAA6506AFFA2154BF8BFFBBA0B1F" box="[843,1124,1889,1915]" italics="true" pageId="60" pageNumber="61">Echinoderes hakaiensis</emphasis>
|
||||
</taxonomicName>
|
||||
was described from a depth of
|
||||
<quantity id="B9139B51506AFFA21724F859FE420BC5" box="[292,412,1927,1953]" metricMagnitude="2" metricUnit="m" metricValue="1.14" metricValueMax="1.4" metricValueMin="0.88" pageId="60" pageNumber="61" unit="m" value="114.0" valueMax="140.0" valueMin="88.0">88–140 m</quantity>
|
||||
, and is thus a part of the shelf or sublittoral fauna. The recorded specimens originate from the bathyal fauna, which suggests that the species is not limited by depth preferences, but can be found along the American west coast, throughout the sublittoral to bathyal zones.
|
||||
</paragraph>
|
||||
<treatment id="F64287A2506BFFDF1795FED5FE900AF9" ID-DOI="http://doi.org/10.5281/zenodo.3818834" ID-GBIF-Taxon="163993847" ID-Zenodo-Dep="3818834" LSID="urn:lsid:plazi:treatment:F64287A2506BFFDF1795FED5FE900AF9" httpUri="http://treatment.plazi.org/id/F64287A2506BFFDF1795FED5FE900AF9" lastPageId="65" lastPageNumber="66" pageId="61" pageNumber="62" scope_class="Cyclorhagida" scope_family="Echinoderidae" scope_order="Echinorhagata" scope_phylum="Kinorhyncha">
|
||||
<subSubSection id="36F1653F506BFFA31795FED5FC6F0D2C" pageId="61" pageNumber="62" type="nomenclature">
|
||||
<paragraph id="7E5436B4506BFFA31795FED5FC6F0D2C" blockId="61.[405,1183,267,329]" pageId="61" pageNumber="62">
|
||||
<taxonomicName id="B9EB4D37506BFFA31795FED5FB410D41" authority="Yamasaki, Neuhaus & George, 2018" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[405,1183,267,294]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="61" pageNumber="62" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||||
<taxonomicName id="B9EB4D37506BFFA31795FED5FB410D41" authority="Yamasaki, Neuhaus & George, 2018" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[405,1183,267,294]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" isUncertain="true" kingdom="Animalia" order="Cyclorhagida" pageId="61" pageNumber="62" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||||
<emphasis id="4C9FEAA6506BFFA31795FED5FDF30D42" bold="true" box="[405,557,267,294]" italics="true" pageId="61" pageNumber="62">Echinoderes</emphasis>
|
||||
cf.
|
||||
<emphasis id="4C9FEAA6506BFFA3145AFED5FD350D41" bold="true" box="[602,747,267,293]" italics="true" pageId="61" pageNumber="62">unispinosus</emphasis>
|
||||
|
|
@ -153,6 +67,8 @@ Yamasaki, Neuhaus & George, 2018
|
|||
,
|
||||
<tableCitation id="3369030F506BFFA31514FEF0FC6F0D2C" box="[788,945,302,328]" captionStart-0="Table 16" captionStart-1="Table 17" captionStartId-0="64.[189,258,268,294]" captionStartId-1="65.[189,258,268,294]" captionTargetBox-1="[194,1382,621,1067]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionText-0="Table 16. Measurements from light microscopy of Echinoderes cf. unispinosus (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 17. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes cf. unispinosus. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; (♀) = female and (♂) = male conditions of sexually dimorphic characters; * = marks characters differing in presence/absence from the original species description of Yamasaki et al. (2018b); (*) = marks characters also present in the original species description of Yamasaki et al. (2018b), but in a slightly different position." httpUri-1="http://table.plazi.org/id/2A94663C5017FFDF16BDFED2FBC30E5B" pageId="61" pageNumber="62" tableUuid-1="2A94663C5017FFDF16BDFED2FBC30E5B">Tables 16–17</tableCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F506BFFA316BDFEAEFBF70EAC" pageId="61" pageNumber="62" type="materials_examined">
|
||||
<paragraph id="7E5436B4506BFFA316BDFEAEFE740DEE" blockId="61.[189,426,368,394]" box="[189,426,368,394]" pageId="61" pageNumber="62">
|
||||
<heading id="251C81D8506BFFA316BDFEAEFE740DEE" bold="true" box="[189,426,368,394]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||||
<emphasis id="4C9FEAA6506BFFA316BDFEAEFE740DEE" bold="true" box="[189,426,368,394]" pageId="61" pageNumber="62">Material examined</emphasis>
|
||||
|
|
@ -185,8 +101,11 @@ for detailed station data
|
|||
<emphasis id="4C9FEAA6506BFFA316EAFDF8FE780E24" bold="true" box="[234,422,550,576]" pageId="61" pageNumber="62">Other material</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4506BFFA316BDFD94FDA10EE3" blockId="61.[189,1399,550,647]" pageId="61" pageNumber="62">
|
||||
<materialsCitation id="CE833CE9506BFFA316BDFD94FDA50EE3" pageId="61" pageNumber="62">
|
||||
<collectingCountry id="06FC7624506BFFA316BDFD94FD880E00" box="[189,598,586,612]" name="United States of America" pageId="61" pageNumber="62">UNITED STATES OF AMERICA</collectingCountry>
|
||||
: 3 ♂♂, St. 3; 1 ♂, St. 5. All mounted for SEM and stored in the first author’s personal reference collection.
|
||||
: 3 ♂♂, St. 3; 1 ♂, St. 5. All mounted for SEM and stored in the first author’s personal reference collection
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B4506BFFA316BDFD70FBF70EAC" blockId="61.[189,1065,686,712]" box="[189,1065,686,712]" pageId="61" pageNumber="62">
|
||||
<heading id="251C81D8506BFFA316BDFD70FBF70EAC" bold="true" box="[189,1065,686,712]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||||
|
|
@ -197,6 +116,8 @@ material
|
|||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="36F1653F506BFFDF16BDFD03FE900AF9" lastPageId="65" lastPageNumber="66" pageId="61" pageNumber="62" type="discussion">
|
||||
<paragraph id="7E5436B4506BFFA316BDFD03FABB0F7E" blockId="61.[189,1398,733,794]" pageId="61" pageNumber="62">
|
||||
A complete overview of measurements and dimensions is presented in
|
||||
<tableCitation id="3369030F506BFFA31215FD03FBA10E93" box="[1045,1151,733,759]" captionStart="Table 16" captionStartId="64.[189,258,268,294]" captionTargetPageId="64" captionText="Table 16. Measurements from light microscopy of Echinoderes cf. unispinosus (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." pageId="61" pageNumber="62">Table 16</tableCitation>
|
||||
|
|
@ -298,7 +219,7 @@ Yamasaki, Neuhaus & George, 2018
|
|||
</taxonomicName>
|
||||
, ♂ (A, H) (NHMD-227016),
|
||||
<specimenCount id="68EDFD3D506BFFA3127AF948FB4E0AD4" box="[1146,1168,1686,1712]" pageId="61" pageNumber="62" type="female">♀</specimenCount>
|
||||
(B–E, G) (NHMD- 227020) and
|
||||
(B–E, G) (NHMD-227020) and
|
||||
<specimenCount id="68EDFD3D506BFFA31755F967FEB50AB7" box="[341,363,1721,1747]" pageId="61" pageNumber="62" type="female">♀</specimenCount>
|
||||
(F) (NHMD-227019).
|
||||
<emphasis id="4C9FEAA6506BFFA31476F967FD550AB7" bold="true" box="[630,651,1721,1747]" pageId="61" pageNumber="62">A</emphasis>
|
||||
|
|
@ -796,13 +717,6 @@ locality of
|
|||
</taxonomicName>
|
||||
, but ideally the problem should be approached by a molecular comparison of the two populations.
|
||||
</paragraph>
|
||||
<paragraph id="7E5436B45017FFDF1408F916FBF20A86" blockId="65.[520,1068,1736,1762]" box="[520,1068,1736,1762]" pageId="65" pageNumber="66">
|
||||
Genus
|
||||
<taxonomicName id="B9EB4D375017FFDF145BF916FBF20A86" authority="Neuhaus & Blasche, 2006" authorityName="Neuhaus & Blasche" authorityYear="2006" box="[603,1068,1736,1762]" class="Kinorhyncha" family="Echinoderidae" genus="Fissuroderes" kingdom="Animalia" order="Cyclorhagida" pageId="65" pageNumber="66" phylum="Cephalorhyncha" rank="genus">
|
||||
<emphasis id="4C9FEAA65017FFDF145BF916FD2C0A86" box="[603,754,1736,1762]" italics="true" pageId="65" pageNumber="66">Fissuroderes</emphasis>
|
||||
<bibRefCitation id="1A7A4B455017FFDF14F9F916FBF20A86" author="Neuhaus B. & Blasche T." box="[761,1068,1736,1762]" pageId="65" pageNumber="66" pagination="19 - 52" refId="ref37069" refString="Neuhaus B. & Blasche T. 2006. Fissuroderes, a new genus of Kinorhyncha (Cyclorhagida) from the deep sea and continental shelf of New Zealand and from the continental shelf of Costa Rica. Zoologischer Anzeiger 245: 19 - 52. https: // doi. org / 10.1016 / j. jcz. 2006.03.003" type="journal article" year="2006">Neuhaus & Blasche, 2006</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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