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<document id="ED983D4755D29094E0E2D14F421116FE" ID-DOI="10.11646/zootaxa.5503.1.1" ID-ISSN="1175-5334" ID-ZooBank="7385E4AA-D7AD-4E79-A05C-684319E0843B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051594773" checkinUser="plazi" docAuthor="Hotèkpo, Sourou Joseph, Schön, Isa &amp; Martens, Koen" docDate="2024" docId="FA1F073A4507FF82FF411E84FB1329F0" docLanguage="en" docName="zootaxa.5503.1.1.pdf" docOrigin="Zootaxa 5503 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5503.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Benincandona trapezoides Hotèkpo, Schön &amp; Martens, 2024, gen. et spec. nov." docType="treatment" docUuid="D8C20638-F2AA-4F8D-A762-31CB4CB6C241" docUuidSource="ZooBank" docVersion="1" lastPageNumber="63" masterDocId="06267F424535FFBCFFD61B4EFF8E2D35" masterDocTitle="An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="51" updateTime="1726051738210" updateUser="GgImagineBatch">
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<mods:title id="CEBC9E7C0D35163F1D4C1189D7A97ED1">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="5DB51AA4F81CCD297E8E3AC738B1E7CC">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="0F5931705EE89C6653E9D419B0403C6F">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="051E6CC7992D9DADECCD7B243166D106">Schön, Isa</mods:namePart>
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<mods:affiliation id="1B6D92B969F4DB19E6D3B34E3B56228E">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="A62A9D582BB64C688B6F9DA1C2020D36">Martens, Koen</mods:namePart>
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<mods:affiliation id="F6586EBB81249F6AB301CD3B8CA79554">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
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<mods:title id="E7C7438AF7A943746BDC94AF531D3E61">Zootaxa</mods:title>
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<mods:date id="EF4D577F955A98AD0D9A087511AC5332">2024</mods:date>
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<mods:number id="0271147EA23E022E533A17F396516388">5503</mods:number>
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<paragraph id="7209B62C4507FF8EFF411E84FC5D2B3C" blockId="50.[151,979,1482,1581]" pageId="50" pageNumber="51">
<heading id="294101404507FF8EFF411E84FC3D28D0" bold="true" box="[151,947,1482,1509]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
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<taxonomicName id="B5B6CDAF4507FF8EFF411E84FE5928D0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,471,1482,1509]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4507FF8EFF411E84FE5928D0" bold="true" box="[151,471,1482,1509]" italics="true" pageId="50" pageNumber="51">Benincandona trapezoides</emphasis>
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Hotèkpo &amp; Martens
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<subSubSection id="3AACE5A74507FF8EFF411D5DFE242B18" box="[151,426,1555,1581]" pageId="50" pageNumber="51" type="description">
<paragraph id="7209B62C4507FF8EFF411D5DFE242B18" blockId="50.[151,979,1482,1581]" box="[151,426,1555,1581]" pageId="50" pageNumber="51">
(
<figureCitation id="EA8DAAA94507FF8EFF491D5DFECA2B18" box="[159,324,1555,1581]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStart-3="FIGURE 35" captionStart-4="FIGURE 36" captionStart-5="FIGURE 37" captionStart-6="FIGURE 38" captionStart-7="FIGURE 39" captionStartId-0="51.[151,250,1943,1967]" captionStartId-1="52.[151,250,1520,1544]" captionStartId-2="54.[151,250,1520,1544]" captionStartId-3="55.[151,250,1943,1967]" captionStartId-4="56.[151,250,1943,1967]" captionStartId-5="57.[151,250,1981,2005]" captionStartId-6="58.[151,250,1943,1967]" captionStartId-7="59.[151,250,1960,1984]" captionTargetBox-0="[179,1408,181,1920]" captionTargetBox-1="[151,1436,181,1496]" captionTargetBox-2="[151,1436,181,1496]" captionTargetBox-3="[186,1401,185,1915]" captionTargetBox-4="[166,1418,185,1915]" captionTargetBox-5="[223,1388,222,1895]" captionTargetBox-6="[189,1403,188,1912]" captionTargetBox-7="[175,1409,202,1932]" captionTargetId-0="figure-17@51.[179,1408,181,1920]" captionTargetId-1="figure-31@52.[151,1436,181,1496]" captionTargetId-2="figure-31@54.[151,1436,181,1496]" captionTargetId-3="figure-17@55.[182,1405,181,1919]" captionTargetId-4="figure-31@56.[166,1422,181,1919]" captionTargetId-5="figure-17@57.[166,1422,181,1957]" captionTargetId-6="figure-31@58.[179,1408,181,1919]" captionTargetId-7="figure-17@59.[175,1413,198,1936]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="54" captionTargetPageId-3="55" captionTargetPageId-4="56" captionTargetPageId-5="57" captionTargetPageId-6="58" captionTargetPageId-7="59" captionText-0="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." captionText-1="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-2="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." captionText-3="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." captionText-4="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." captionText-5="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." captionText-6="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="50" pageNumber="51">
Figures 32
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,
<figureCitation id="EA8DAAA94507FF8EFE861D5DFE2C2B18" box="[336,418,1555,1581]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="50" pageNumber="51">41OR</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF411D15FB332BAC" box="[151,1213,1627,1689]" pageId="50" pageNumber="51" type="etymology">
<paragraph id="7209B62C4507FF8EFF411D15FE962B40" blockId="50.[151,1213,1627,1689]" box="[151,280,1627,1653]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF411D15FE962B40" bold="true" box="[151,280,1627,1653]" pageId="50" pageNumber="51">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF411D31FB332BAC" blockId="50.[151,1213,1627,1689]" box="[151,1213,1663,1689]" pageId="50" pageNumber="51">The species is named after the shape of the carapace in lateral view, which is largely trapezoid.</paragraph>
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<subSubSection id="3AACE5A74507FF8EFF411D8EFBA12ACF" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="7209B62C4507FF8EFF411D8EFE842BEF" blockId="50.[151,1437,1728,2043]" box="[151,266,1728,1754]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF411D8EFE842BEF" bold="true" box="[151,266,1728,1754]" pageId="50" pageNumber="51">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF411DAAFBA12ACF" blockId="50.[151,1437,1728,2043]" pageId="50" pageNumber="51">
CpRl relatively high (L/H = 2.0), dorsal margin straight over
<emphasis id="40C26A3E4507FF8EFCEA1DABFCFB2BCB" box="[828,885,1765,1790]" italics="true" pageId="50" pageNumber="51">circa</emphasis>
one third of the entire length, with greatest height in the middle; LV dorsally extending beyond RV over the entire length of the straight part of the dorsal margin, anterior margin broadly rounded, posterior margin running straight towards the ventral side, the latter almost straight, valve surface set with ridges in most of the central part. CpD and CpV relatively wide, with greatest width in the middle and lateral sides straight in the middle, anterior extremity sharply and posterior side more bluntly pointed. A2 with seta X long, reaching well beyond the tip of the aesthetasc Y, the latter distally not bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E4507FF8EFE721CF3FE3E2AE3" box="[420,432,1981,2006]" italics="true" pageId="50" pageNumber="51">c</emphasis>
. 1.5x basal W. T3 with seta d2 absent. Hp with ls and ms with overlapping positions, lobe ls broad and rounded, extending beyond lobe ms with more than half of its length.
</paragraph>
</subSubSection>
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<paragraph id="7209B62C4506FF8FFF411CD9FD892ACD" blockId="51.[151,1436,1943,2040]" pageId="51" pageNumber="52">
<emphasis id="40C26A3E4506FF8FFF411CD9FE912A9A" bold="true" box="[151,287,1943,1967]" pageId="51" pageNumber="52">FIGURE 32.</emphasis>
<taxonomicName id="B5B6CDAF4506FF8FFEF21CD6FDBF2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,561,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4506FF8FFEF21CD6FDBF2A85" box="[292,561,1944,1968]" italics="true" pageId="51" pageNumber="52">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454506FF8FFDE01CD6FD6A2A9A" box="[566,740,1944,1968]" pageId="51" pageNumber="52" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, GK ♂; DF ♀).
<emphasis id="40C26A3E4506FF8FFBF01CD9FBFC2A85" bold="true" box="[1062,1138,1943,1968]" pageId="51" pageNumber="52">AF, H</emphasis>
. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279).
<emphasis id="40C26A3E4506FF8FFE1D1CF5FD9E2AE1" bold="true" box="[459,528,1979,2004]" pageId="51" pageNumber="52">G, I, J</emphasis>
. INV.159279. CpV.
<emphasis id="40C26A3E4506FF8FFD371CF2FD7B2AE1" bold="true" box="[737,757,1980,2004]" pageId="51" pageNumber="52">K</emphasis>
. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm.
</paragraph>
</caption>
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<paragraph id="7209B62C4501FF88FF411EBEFBFC2B18" blockId="52.[151,1437,1520,1581]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="52" pageNumber="53">FIGURE 33.</emphasis>
<taxonomicName id="B5B6CDAF4501FF88FEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4501FF88FEF81EBEFDCE2B3D" box="[302,576,1520,1544]" italics="true" pageId="52" pageNumber="53">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454501FF88FD9C1EBFFC862B3D" box="[586,776,1521,1545]" pageId="52" pageNumber="53" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E4501FF88FC431EBEFB9C2B3D" bold="true" box="[917,1042,1520,1544]" pageId="52" pageNumber="53">A, C, D, G</emphasis>
. LVi (INV.159263).
<emphasis id="40C26A3E4501FF88FB261EBEFAEB2B3D" bold="true" box="[1264,1381,1520,1544]" pageId="52" pageNumber="53">B, E, F, H</emphasis>
. RVi (INV.159263).
<emphasis id="40C26A3E4501FF88FEE71D5AFEE52B19" bold="true" box="[305,363,1556,1580]" pageId="52" pageNumber="53">G, H.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A74501FF89FF411D30FD7D2C98" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="7209B62C4501FF88FF411D30FEB02BAD" blockId="52.[151,318,1662,1688]" box="[151,318,1662,1688]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411D30FEB02BAD" bold="true" box="[151,318,1662,1688]" pageId="52" pageNumber="53">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411D88FE8A2BD5" blockId="52.[151,1282,1734,1797]" box="[151,260,1734,1760]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411D88FE8A2BD5" bold="true" box="[151,260,1734,1760]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411D88FE8A2BD5" box="[151,260,1734,1760]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411DA4FA8C2A30" blockId="52.[151,1282,1734,1797]" box="[151,1282,1770,1797]" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411DA4FA8C2A30" box="[151,1282,1770,1797]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA54501FF88FF411DA4FF4D2A30" box="[151,195,1770,1797]" count="1" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE94501FF88FBB01DA4FB182A31" box="[1126,1174,1770,1796]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159263).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4501FF88FF411C7CFF722A79" blockId="52.[151,736,1842,1905]" box="[151,252,1842,1868]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411C7CFF722A79" bold="true" box="[151,252,1842,1868]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411C7CFF722A79" box="[151,252,1842,1868]" pageId="52" pageNumber="53" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF88FF411C18FD6E2A44" blockId="52.[151,736,1842,1905]" box="[151,736,1878,1905]" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411C18FD6E2A44" box="[151,736,1878,1905]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA54501FF88FF411C18FF4F2A44" box="[151,193,1878,1905]" count="1" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE94501FF88FD921C18FDF72A44" box="[580,633,1878,1905]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159264).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4501FF88FF411CD0FE812A8D" blockId="52.[151,1436,1950,2013]" box="[151,271,1950,1976]" pageId="52" pageNumber="53">
<emphasis id="40C26A3E4501FF88FF411CD0FE812A8D" bold="true" box="[151,271,1950,1976]" pageId="52" pageNumber="53">
<typeStatus id="AD0D088E4501FF88FF411CD0FE812A8D" box="[151,271,1950,1976]" pageId="52" pageNumber="53" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4501FF89FF411C8CFC282DCC" blockId="52.[151,1436,1950,2013]" lastBlockId="53.[151,1436,150,249]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<materialsCitation id="C2DEBC714501FF88FF411C8CFC442AE8" box="[151,970,1986,2013]" collectionCode="INV" pageId="52" pageNumber="53" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="64B07DA54501FF88FF411C8CFF572AE8" box="[151,217,1986,2013]" count="2" pageId="52" pageNumber="53" type="male">2 ♂♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E4501FF88FDF21C8CFD082AE9" box="[548,646,1986,2012]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94501FF88FD4E1C8CFD422AE8" box="[664,716,1986,2013]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="52" pageNumber="53" type="Herbarium">INV</collectionCode>
.159265, INV.159266)
</materialsCitation>
;
<materialsCitation id="C2DEBC714501FF89FC0F1C8CFEA32DE0" collectionCode="INV" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" specimenCount="9" specimenCount-female="9" typeStatus="paratype">
<specimenCount id="64B07DA54501FF88FC0F1C8CFB952AE8" box="[985,1051,1986,2013]" count="9" pageId="52" pageNumber="53" type="female">9 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE94500FF89FF491BD8FF5D2D84" box="[159,211,150,177]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159267, INV.159268, INV.159269, INV.159270, INV.159271, INV.159272, INV.159273, INV.159274, INV.159275)
</materialsCitation>
;
<materialsCitation id="C2DEBC714500FF89FEEE1BF5FA162DE0" box="[312,1432,186,213]" collectionCode="INV" pageId="53" pageNumber="54" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54500FF89FEEE1BF5FEF92DE0" box="[312,375,187,213]" count="3" pageId="53" pageNumber="54" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE94500FF89FC051BF4FB892DE0" box="[979,1031,186,213]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159276, INV.159277, INV.159278)
</materialsCitation>
;
<materialsCitation id="C2DEBC714500FF89FF411B91FC2C2DCC" box="[151,930,222,249]" collectionCode="INV" pageId="53" pageNumber="54" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA54500FF89FF411B91FF4F2DCC" box="[151,193,223,249]" count="1" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE94500FF89FCDD1B90FCCE2DCC" box="[779,832,222,249]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="53" pageNumber="54" type="Herbarium">INV</collectionCode>
.159279)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C4500FF89FF411A69FEBE2C75" blockId="53.[151,1436,294,429]" box="[151,304,294,321]" pageId="53" pageNumber="54">
<emphasis id="40C26A3E4500FF89FF411A69FEBE2C75" bold="true" box="[151,304,294,321]" pageId="53" pageNumber="54">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4500FF89FF411A05FD7D2C98" blockId="53.[151,1436,294,429]" pageId="53" pageNumber="54">
<materialsCitation id="C2DEBC714500FF89FF411A05FE2D2CBC" country="Benin" location="Parakou District" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC4500FF89FF411A05FF542C50" box="[151,218,331,357]" name="Benin" pageId="53" pageNumber="54">Benin</collectingCountry>
,
<collectingRegion id="B07278CE4500FF89FF3E1A05FE442C50" box="[232,458,331,357]" country="Benin" name="Borgou" pageId="53" pageNumber="54">Borgou Department</collectingRegion>
,
<location id="7769E0F74500FF89FE001A05FD1E2C50" LSID="urn:lsid:plazi:treatment:FA1F073A4507FF82FF411E84FB1329F0:7769E0F74500FF89FE001A05FD1E2C50" box="[470,656,331,357]" country="Benin" name="Parakou District" pageId="53" pageNumber="54" stateProvince="Borgou">Parakou District</location>
, town Bapérou located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE4500FF89FF411A21FF662CBC" box="[151,232,367,393]" country="Benin" name="Oueme" pageId="53" pageNumber="54">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB4500FF89FDE81A21FD4A2CBC" box="[574,708,367,393]" degrees="9" direction="north" minutes="16.380" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="9.273">9°16.380N</geoCoordinate>
,
<geoCoordinate id="1782D0EB4500FF89FD191A21FCD82CBC" box="[719,854,367,393]" degrees="2" direction="east" minutes="34.913" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="2.5818834">2°34.913 E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC4500FF89FA561A20FEC22C98" pageId="53" pageNumber="54" value="2022-11-05">05 November 2022</date>
(Locality code A
<quantity id="B54E1BC94500FF89FDC21ADDFDB52C98" box="[532,571,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="53" pageNumber="54" unit="in" value="1.0">1 in</quantity>
<tableCitation id="3F3483974500FF89FD971ADDFD102C98" box="[577,670,403,429]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="53" pageNumber="54">Tables 1</tableCitation>
and
<tableCitation id="3F3483974500FF89FD031ADDFD682C98" box="[725,742,403,429]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="53" pageNumber="54">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74500FF82FF411A94FD9E2FF8" lastPageId="62" lastPageNumber="63" pageId="53" pageNumber="54" type="description">
<paragraph id="7209B62C4500FF89FF411A94FE0F2CC0" blockId="53.[151,1437,474,1905]" box="[151,385,474,501]" pageId="53" pageNumber="54">
<emphasis id="40C26A3E4500FF89FF411A94FE0F2CC0" bold="true" box="[151,385,474,501]" pageId="53" pageNumber="54">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4500FF89FF411AB1FCD82FF8" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
CpRl (
<figureCitation id="EA8DAAA94500FF89FF361AB1FE0D2F2C" box="[224,387,511,537]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32AC</figureCitation>
) with trapezoidal shape (L/H ratio=2.0); dorsal margin straight in the middle 1/3, with greatest height in the middle, anterior margin broadly rounded, posterior margin running straight towards ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV along the entire length of the straight part; valve surface set with ridges in the middle part of the valves (
<figureCitation id="EA8DAAA94500FF89FA871925FF6B2F9C" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32HJ</figureCitation>
). CpD and CpV (
<figureCitation id="EA8DAAA94500FF89FE7019C1FDC32F9C" box="[422,589,655,681]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="53" pageNumber="54">Figure 32G, K</figureCitation>
) relatively wide, pointed sharply anteriorly and bluntly posteriorly, greatest width in the middle, lateral margins straight in the middle part.
</paragraph>
<paragraph id="7209B62C4500FF89FF111999FB412E0C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
LVi (
<figureCitation id="EA8DAAA94500FF89FF281999FE6F2FC4" box="[254,481,727,753]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin straight in the middle third, with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FC9F1851FC452E0C" box="[841,971,799,825]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33G</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FB961851FBFE2E0C" box="[1088,1136,799,825]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF11180DFBF02E90" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
RVi (
<figureCitation id="EA8DAAA94500FF89FED4180DFE682E68" box="[258,486,835,861]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33B, E, F, H</figureCitation>
) elongated, with dorsal margin straight in the middle third and with greatest height in the middle, anterior margin broadly rounded; posterior margin sloping towards ventral margin, the latter almost straight; ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94500FF89FD2118C5FCF42E90" box="[759,890,907,933]" captionStart="FIGURE 33" captionStartId="52.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@52.[151,1436,181,1496]" captionTargetPageId="52" captionText="FIGURE 33. Benincandona trapezoides gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159263). B, E, F, H. RVi (INV.159263). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="53" pageNumber="54">Figure 33H</figureCitation>
) as in the
<typeStatus id="AD0D088E4500FF89FC3818C5FB902E90" box="[1006,1054,907,933]" pageId="53" pageNumber="54">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4500FF89FF1118E1FB0A2838" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A1 (
<figureCitation id="EA8DAAA94500FF89FF2D18E1FEF12EFC" box="[251,383,942,969]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35B</figureCitation>
) with seven segments, one basal and six distal. Basal segment with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer one, and two shorter setae on the mid-dorsal side (one not illustrated here), situated some distance from each other. First distal segment about 2.5 times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio, carrying one short apical setae on each side. Fourth distal segment about three times as long as wide, and set with one short ventro-apical seta and two long dorso-apical. Fifth distal segment about five time as long as wide, with two long dorso-apical setae and one short ventro-apical seta clearly exceeding the next segment. Sixth distal segment narrow, at least five times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long and one short setae (about one third the length of Ya).
</paragraph>
<paragraph id="7209B62C4500FF89FF111E59FA9A2A1C" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
A2 (
<figureCitation id="EA8DAAA94500FF89FF2D1E59FE0D2804" box="[251,387,1302,1329]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E4500FF89FBD41E75FB9B2861" box="[1026,1045,1339,1364]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
four times the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E4500FF89FCFA1E11FCB1284D" box="[812,831,1375,1400]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout, five times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y, and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E4500FF89FDAD1EA1FD002B3D" box="[635,654,1519,1544]" italics="true" pageId="53" pageNumber="54">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a slender aesthetasc y1; t2 and t3 setae hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA94500FF89FF491D31FE9F2BAC" box="[159,273,1663,1689]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="53" pageNumber="54">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into one short seta about equal length as y2; centrally with one short plumose seta z3; dorso-apically, z1 claw-like about 1.5x as long as penultimate segment. Claw G2 and G1 about 2.5 times as long as z1; seta z2 transformed into long claw, reaching tip of G1. Fourth, terminal, endopodal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C4500FF89FF111C7DFAA52A44" blockId="53.[151,1437,474,1905]" pageId="53" pageNumber="54">
MdCoxa (
<figureCitation id="EA8DAAA94500FF89FEED1C7DFE4E2A78" box="[315,448,1842,1869]" captionStart="FIGURE 35" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[186,1401,185,1915]" captionTargetId="figure-17@55.[182,1405,181,1919]" captionTargetPageId="55" captionText="FIGURE 35. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. A2 (INV.159280). B. A1 (INV.159265). C. MdCoxa (INV.159263). Scale bar = 50 µm." pageId="53" pageNumber="54">Figure 35C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<caption id="26C9E6A44503FF8AFF411EBEFBFC2B18" pageId="54" pageNumber="55" startId="54.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="54" targetType="figure">
<paragraph id="7209B62C4503FF8AFF411EBEFBFC2B18" blockId="54.[151,1437,1520,1581]" pageId="54" pageNumber="55">
<emphasis id="40C26A3E4503FF8AFF411EBEFEAA2B3D" bold="true" box="[151,292,1520,1544]" pageId="54" pageNumber="55">FIGURE 34.</emphasis>
<taxonomicName id="B5B6CDAF4503FF8AFEF81EBEFDCE2B3D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1520,1544]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4503FF8AFEF81EBEFDCE2B3D" box="[302,576,1520,1544]" italics="true" pageId="54" pageNumber="55">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454503FF8AFD9C1EBFFC862B3D" box="[586,776,1521,1545]" pageId="54" pageNumber="55" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4503FF8AFC431EBEFB9C2B3D" bold="true" box="[917,1042,1520,1544]" pageId="54" pageNumber="55">A, C, D, G</emphasis>
. LVi (INV.159264).
<emphasis id="40C26A3E4503FF8AFB261EBEFAEB2B3D" bold="true" box="[1264,1381,1520,1544]" pageId="54" pageNumber="55">B, E, F, H</emphasis>
. RVi (INV.159264).
<emphasis id="40C26A3E4503FF8AFEE71D5AFEE52B19" bold="true" box="[305,363,1556,1580]" pageId="54" pageNumber="55">G, H.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4503FF8AFF111D30FEF82A8D" blockId="54.[151,1437,1662,1976]" pageId="54" pageNumber="55">
MdPalp (
<figureCitation id="EA8DAAA94503FF8AFEF91D30FE3C2BAD" box="[303,434,1662,1688]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="54" pageNumber="55">Figure 36C</figureCitation>
) with four segments. Basal segment ventro-subapically carrying two hirsute setae (S1 &amp; S2), S1 long, reaching tip of terminal segment and S2 short and stout, length about one fourth of the length of S1; in between with the short flagellum-like alpha seta (thin and as short as S2), and one smooth and medium size (slightly shorter than S1) more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, tips slightly shorter than the fused claw.
</paragraph>
<caption id="26C9E6A44502FF8BFF411CD9FE7C2AE1" pageId="55" pageNumber="56" startId="55.[151,250,1943,1967]" targetBox="[186,1401,185,1915]" targetPageId="55" targetType="figure">
<paragraph id="7209B62C4502FF8BFF411CD9FE7C2AE1" blockId="55.[151,1437,1943,2004]" pageId="55" pageNumber="56">
<emphasis id="40C26A3E4502FF8BFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="55" pageNumber="56">FIGURE 35.</emphasis>
<taxonomicName id="B5B6CDAF4502FF8BFEF11CD6FDBB2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E4502FF8BFEF11CD6FDBB2A85" box="[295,565,1944,1968]" italics="true" pageId="55" pageNumber="56">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454502FF8BFDEA1CD6FD612A9A" box="[572,751,1944,1968]" pageId="55" pageNumber="56" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4502FF8BFCBF1CD9FCF52A9A" bold="true" box="[873,891,1943,1967]" pageId="55" pageNumber="56">A</emphasis>
. A2 (INV.159280).
<emphasis id="40C26A3E4502FF8BFB911CD6FBD62A85" bold="true" box="[1095,1112,1944,1968]" pageId="55" pageNumber="56">B</emphasis>
. A1 (INV.159265).
<emphasis id="40C26A3E4502FF8BFAF21CD9FAB32A9A" bold="true" box="[1316,1341,1943,1967]" pageId="55" pageNumber="56">C.</emphasis>
MdCoxa (INV.159263). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450DFF84FF411CD9FB492AE1" pageId="56" pageNumber="57" startId="56.[151,250,1943,1967]" targetBox="[166,1418,185,1915]" targetPageId="56" targetType="figure">
<paragraph id="7209B62C450DFF84FF411CD9FB492AE1" blockId="56.[151,1436,1943,2004]" pageId="56" pageNumber="57">
<emphasis id="40C26A3E450DFF84FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="56" pageNumber="57">FIGURE 36.</emphasis>
<taxonomicName id="B5B6CDAF450DFF84FEF91CD6FDCC2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,578,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450DFF84FEF91CD6FDCC2A85" box="[303,578,1944,1968]" italics="true" pageId="56" pageNumber="57">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450DFF84FD9A1CD6FC822A9A" box="[588,780,1944,1968]" pageId="56" pageNumber="57" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450DFF84FC441CD9FC2A2A9A" bold="true" box="[914,932,1943,1967]" pageId="56" pageNumber="57">A</emphasis>
. Mx1, respiratory plate (INV.159263).
<emphasis id="40C26A3E450DFF84FA9C1CD6FAD52A85" bold="true" box="[1354,1371,1944,1968]" pageId="56" pageNumber="57">B</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159263).
<emphasis id="40C26A3E450DFF84FDE41CF5FDCA2AE6" bold="true" box="[562,580,1979,2003]" pageId="56" pageNumber="57">C</emphasis>
. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450CFF85FF411CF3FD7D2ACF" pageId="57" pageNumber="58" startId="57.[151,250,1981,2005]" targetBox="[223,1388,222,1895]" targetPageId="57" targetType="figure">
<paragraph id="7209B62C450CFF85FF411CF3FD7D2ACF" blockId="57.[151,1436,1981,2042]" pageId="57" pageNumber="58">
<emphasis id="40C26A3E450CFF85FF411CF3FEAA2AE0" bold="true" box="[151,292,1981,2005]" pageId="57" pageNumber="58">FIGURE 37.</emphasis>
<taxonomicName id="B5B6CDAF450CFF85FEF81CF3FDCE2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,576,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450CFF85FEF81CF3FDCE2AE0" box="[302,576,1981,2005]" italics="true" pageId="57" pageNumber="58">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450CFF85FD9C1CF0FC892AE0" box="[586,775,1982,2006]" pageId="57" pageNumber="58" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450CFF85FC5A1CF3FC102AE0" bold="true" box="[908,926,1981,2005]" pageId="57" pageNumber="58">A</emphasis>
. T2 (INV.159263).
<emphasis id="40C26A3E450CFF85FBA41CF0FB0D2AE3" bold="true" box="[1138,1155,1982,2006]" pageId="57" pageNumber="58">B</emphasis>
. T3 (INV.159263).
<emphasis id="40C26A3E450CFF85FA811CF3FAE72AE0" bold="true" box="[1367,1385,1981,2005]" pageId="57" pageNumber="58">C</emphasis>
. CR (INV.159280).
<emphasis id="40C26A3E450CFF85FEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="57" pageNumber="58">D</emphasis>
. CRAtt (INV.159280). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450FFF86FF411CD9FDB32ACD" pageId="58" pageNumber="59" startId="58.[151,250,1943,1967]" targetBox="[189,1403,188,1912]" targetPageId="58" targetType="figure">
<paragraph id="7209B62C450FFF86FF411CD9FDB32ACD" blockId="58.[151,1437,1943,2040]" pageId="58" pageNumber="59">
<emphasis id="40C26A3E450FFF86FF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="58" pageNumber="59">FIGURE 38.</emphasis>
<taxonomicName id="B5B6CDAF450FFF86FEF01CD9FDBD2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,563,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450FFF86FEF01CD9FDBD2A9A" box="[294,563,1943,1967]" italics="true" pageId="58" pageNumber="59">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450FFF86FDEF1CD6FD642A9A" box="[569,746,1944,1968]" pageId="58" pageNumber="59" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E450FFF86FCB41CD9FCFA2A9A" bold="true" box="[866,884,1943,1967]" pageId="58" pageNumber="59">A</emphasis>
. Zenker organ (INV.159263).
<emphasis id="40C26A3E450FFF86FB711CD6FB592A85" bold="true" box="[1191,1239,1944,1968]" pageId="58" pageNumber="59">BD</emphasis>
. Hp (INV.159263).
<emphasis id="40C26A3E450FFF86FF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="58" pageNumber="59">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E450FFF86FE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="58" pageNumber="59">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E450FFF86FBEE1CF2FBC42AE1" bold="true" box="[1080,1098,1980,2004]" pageId="58" pageNumber="59">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4450EFF87FF411CE6FA9F2AD0" pageId="59" pageNumber="60" startId="59.[151,250,1960,1984]" targetBox="[175,1409,202,1932]" targetPageId="59" targetType="figure">
<paragraph id="7209B62C450EFF87FF411CE6FA9F2AD0" blockId="59.[151,1436,1960,2021]" pageId="59" pageNumber="60">
<emphasis id="40C26A3E450EFF87FF411CE6FEAE2AF4" bold="true" box="[151,288,1960,1985]" pageId="59" pageNumber="60">FIGURE 39.</emphasis>
<taxonomicName id="B5B6CDAF450EFF87FEF11CE7FDBB2AF4" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,565,1961,1985]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E450EFF87FEF11CE7FDBB2AF4" box="[295,565,1961,1985]" italics="true" pageId="59" pageNumber="60">Benincandona trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745450EFF87FDEA1CE7FD612AF5" box="[572,751,1961,1985]" pageId="59" pageNumber="60" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♀ and CE ♂).
<emphasis id="40C26A3E450EFF87FBCF1CE6FBA52AF5" bold="true" box="[1049,1067,1960,1984]" pageId="59" pageNumber="60">A</emphasis>
. T1, Lpp (INV.159280).
<emphasis id="40C26A3E450EFF87FAFD1CE7FAB22AF4" bold="true" box="[1323,1340,1961,1985]" pageId="59" pageNumber="60">B</emphasis>
. T1, Rpp (INV.159280).
<emphasis id="40C26A3E450EFF87FEE71C82FECD2AD1" bold="true" box="[305,323,1996,2020]" pageId="59" pageNumber="60">C</emphasis>
. A2 (INV.159264, INV.159269).
<emphasis id="40C26A3E450EFF87FD401C83FD262AD0" bold="true" box="[662,680,1997,2021]" pageId="59" pageNumber="60">D</emphasis>
. T1 (INV.159268).
<emphasis id="40C26A3E450EFF87FCA71C83FC0C2AD0" bold="true" box="[881,898,1997,2021]" pageId="59" pageNumber="60">E</emphasis>
. CR (INV.159264). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44509FF80FF411EA2FD1F2B79" pageId="60" pageNumber="61" startId="60.[151,250,1516,1540]" targetBox="[151,1435,181,1492]" targetPageId="60" targetType="figure">
<paragraph id="7209B62C4509FF80FF411EA2FD1F2B79" blockId="60.[151,1437,1516,1612]" pageId="60" pageNumber="61">
<emphasis id="40C26A3E4509FF80FF411EA2FEAD2B31" bold="true" box="[151,291,1516,1540]" pageId="60" pageNumber="61">FIGURE 40.</emphasis>
Diagrams of apical views of the A2 of
<taxonomicName id="B5B6CDAF4509FF80FD071EA2FCED2B31" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[721,867,1516,1540]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E4509FF80FD071EA2FCED2B31" box="[721,867,1516,1540]" italics="true" pageId="60" pageNumber="61">Benincandona</emphasis>
</taxonomicName>
, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures).
</paragraph>
</caption>
<paragraph id="7209B62C4509FF80FF111DD0FCE42AA4" blockId="60.[151,1437,1694,1937]" pageId="60" pageNumber="61">
Mx1 (
<figureCitation id="EA8DAAA94509FF80FED91DD0FE322B8D" box="[271,444,1694,1720]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA94509FF80FC5C1D8CFB802BE9" box="[906,1038,1730,1756]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36A</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E4509FF80FB291D8DFA9C2BE9" box="[1279,1298,1731,1756]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
18 plumose respiratory rays, two short distal setae and a group of shorter proximal setae (not illustrated). Palp (
<figureCitation id="EA8DAAA94509FF80FAC61DA8FA1A2A35" box="[1296,1428,1766,1792]" captionStart="FIGURE 36" captionStartId="56.[151,250,1943,1967]" captionTargetBox="[166,1418,185,1915]" captionTargetId="figure-31@56.[166,1422,181,1919]" captionTargetPageId="56" captionText="FIGURE 36. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Mx1, respiratory plate (INV.159263). B. Mx1 (Chaetotaxy incomplete) (INV.159263). C. MdPalp (INV.159263). Scale bar. 1 (A), 2 (C), 3 (B) = 50 µm." pageId="60" pageNumber="61">Figure 36B</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E4509FF80FE0F1C1CFDE92A59" box="[473,615,1874,1900]" italics="true" pageId="60" pageNumber="61">Zahnborsten</emphasis>
), basally with one lateral seta; sideways directed bristles near base of first endite, one long and one short,
<emphasis id="40C26A3E4509FF80FE211C39FD842AA5" box="[503,522,1911,1936]" italics="true" pageId="60" pageNumber="61">c.</emphasis>
half the length of the long one.
</paragraph>
<caption id="26C9E6A44508FF81FF411F17FB2529A3" pageId="61" pageNumber="62" startId="61.[151,250,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="61" targetType="figure">
<paragraph id="7209B62C4508FF81FF411F17FB2529A3" blockId="61.[151,1437,1113,1174]" pageId="61" pageNumber="62">
<emphasis id="40C26A3E4508FF81FF411F17FEAF2944" bold="true" box="[151,289,1113,1137]" pageId="61" pageNumber="62">FIGURE 41.</emphasis>
<taxonomicName id="B5B6CDAF4508FF81FEFE1F17FE342944" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,442,1113,1137]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E4508FF81FEFE1F17FE342944" box="[296,442,1113,1137]" italics="true" pageId="61" pageNumber="62">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454508FF81FE141F14FDF82944" box="[450,630,1114,1138]" pageId="61" pageNumber="62" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀).
<emphasis id="40C26A3E4508FF81FA9E1F17FAF72944" bold="true" box="[1352,1401,1113,1137]" pageId="61" pageNumber="62">AD</emphasis>
.
<emphasis id="40C26A3E4508FF81FA501F14FF6729A0" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FA501F14FF6B29A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="martini">B. martini</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FF381F30FE9229A3" bold="true" box="[238,284,1150,1174]" pageId="61" pageNumber="62">EF</emphasis>
.
<emphasis id="40C26A3E4508FF81FEFE1F30FE3F29A0" box="[296,433,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FEFE1F30FE2329A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,429,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ibikounlei">B. ibikounlei</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FE601F33FE6D29A3" bold="true" box="[438,483,1149,1174]" pageId="61" pageNumber="62">GJ</emphasis>
.
<emphasis id="40C26A3E4508FF81FE261F30FDDF29A0" box="[496,593,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FE261F30FDC329A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[496,589,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="sakitii">B. sakitii</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FD811F30FD0729A3" bold="true" box="[599,649,1150,1174]" pageId="61" pageNumber="62">KN</emphasis>
.
<emphasis id="40C26A3E4508FF81FD401F30FD7429A0" box="[662,762,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FD401F30FD7829A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[662,758,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="cebios">B. cebios</taxonomicName>
.
</emphasis>
<emphasis id="40C26A3E4508FF81FCD61F33FCBC29A3" bold="true" box="[768,818,1149,1174]" pageId="61" pageNumber="62">OR</emphasis>
.
<emphasis id="40C26A3E4508FF81FCE91F30FC5929A0" box="[831,983,1149,1173]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="B5B6CDAF4508FF81FCE91F30FC5D29A0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[831,979,1149,1173]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="trapezoides">B. trapezoides</taxonomicName>
.
</emphasis>
Scale bar = 500 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4508FF81FF111FA6FDA328A7" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T1 (
<figureCitation id="EA8DAAA94508FF81FF201FA6FE152837" box="[246,411,1256,1282]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites not shown). Protopodite (
<figureCitation id="EA8DAAA94508FF81FEFB1E42FE3C2813" box="[301,434,1292,1318]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical Rpp (
<figureCitation id="EA8DAAA94508FF81FBD11E7EFB03287F" box="[1031,1165,1328,1354]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94508FF81FEDB1E1AFE1B285B" box="[269,405,1364,1390]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 39A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111ED2FB4E2B5F" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T2 (
<figureCitation id="EA8DAAA94508FF81FF231ED2FEF62883" box="[245,376,1436,1462]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length more than four times basal width), without seta. Second segment (segment 2A) with length
<emphasis id="40C26A3E4508FF81FCF81EAAFCCF28C8" box="[814,833,1508,1533]" italics="true" pageId="61" pageNumber="62">c.</emphasis>
4x width with one short ventro-apical seta f. Third segment (segment 2B) about the same length/width ratio as the previous segment, set with one ventro-subapical seta g. Terminal segment (segment 3) almost as long as width, apically with two short setae (h1 and h3), h1 (ventral) and twice the length of h3 (dorsal), and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4508FF81FF111D3AFF642A77" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
T3 (
<figureCitation id="EA8DAAA94508FF81FF231D3AFEF82BBB" box="[245,374,1652,1678]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37B</figureCitation>
) with first segment carrying only one subapical seta d1, of intermediate length. Second (knee-) segment without seta d2. Third segment about four times as long as wide, not carrying any seta. Fourth segment narrower and at least 8x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, about twice as long as basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 and h2, the later slightly exceeding of the former.
</paragraph>
<paragraph id="7209B62C4508FF81FF111C02FC862A9B" blockId="61.[151,1437,1256,2038]" pageId="61" pageNumber="62">
CR (
<figureCitation id="EA8DAAA94508FF81FF2C1C02FEF52A53" box="[250,379,1868,1894]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37C</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4508FF81FF111CF6FA802AE7" blockId="61.[151,1437,1256,2038]" box="[199,1294,1976,2002]" pageId="61" pageNumber="62">
CRAtt (
<figureCitation id="EA8DAAA94508FF81FEC91CF6FE2C2AE7" box="[287,418,1976,2002]" captionStart="FIGURE 37" captionStartId="57.[151,250,1981,2005]" captionTargetBox="[223,1388,222,1895]" captionTargetId="figure-17@57.[166,1422,181,1957]" captionTargetPageId="57" captionText="FIGURE 37. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. T2 (INV.159263). B. T3 (INV.159263). C. CR (INV.159280). D. CRAtt (INV.159280). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 37D</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4508FF82FF111C92FD4E2DE0" blockId="61.[151,1437,1256,2038]" lastBlockId="62.[151,1436,151,249]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Hp (
<figureCitation id="EA8DAAA94508FF81FF211C92FE142AC3" box="[247,410,2012,2038]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="61" pageNumber="62">Figure 38BD</figureCitation>
) with robust and rounded lobe ls, reaching well beyond the asymmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA9450BFF82FB611BD9FAD52D84" box="[1207,1371,151,177]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C450BFF82FF111B91FC382DCC" blockId="62.[151,1436,151,249]" box="[199,950,223,249]" pageId="62" pageNumber="63">
Zenker organ (
<figureCitation id="EA8DAAA9450BFF82FEBD1B91FE602DCC" box="[363,494,223,249]" captionStart="FIGURE 38" captionStartId="58.[151,250,1943,1967]" captionTargetBox="[189,1403,188,1912]" captionTargetId="figure-31@58.[179,1408,181,1919]" captionTargetPageId="58" captionText="FIGURE 38. Benincandona trapezoides gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159263). BD. Hp (INV.159263). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 38A</figureCitation>
) elongated with
<emphasis id="40C26A3E450BFF82FD7C1B91FD332DCD" box="[682,701,223,248]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
seven spinous whirls.
</paragraph>
<paragraph id="7209B62C450BFF82FF411A68FE182C75" blockId="62.[151,1437,294,825]" box="[151,406,294,320]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF411A68FE182C75" bold="true" box="[151,406,294,320]" pageId="62" pageNumber="63">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF411A05FB2E2C50" blockId="62.[151,1437,294,825]" box="[151,1184,331,357]" pageId="62" pageNumber="63">
Cp and valves (
<figureCitation id="EA8DAAA9450BFF82FE921A05FE7E2C50" box="[324,496,331,357]" captionStart="FIGURE 32" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@51.[179,1408,181,1920]" captionTargetPageId="51" captionText="FIGURE 32. Benincandona trapezoides gen. et spec. nov., valves (AC, GK ♂; DF ♀). AF, H. CpRl (AC, H. INV.159276, DF. INV.1592 INV.159279). G, I, J. INV.159279. CpV. K. INV.159277. CpD. Scale bars. 1 (B, E, G, K) = 500 µm; 2 (A, C, D, F, I) = 200 µm, 3 (H, J) = 40 µm." pageId="62" pageNumber="63">Figures 32DF</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FE2D1A05FDC02C50" box="[507,590,331,357]" captionStart="FIGURE 34" captionStartId="54.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-31@54.[151,1436,181,1496]" captionTargetPageId="54" captionText="FIGURE 34. Benincandona trapezoides gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159264). B, E, F, H. RVi (INV.159264). G, H. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (GH) = 50 µm." pageId="62" pageNumber="63">34AH</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C450BFF82FF111A21FB072CBC" blockId="62.[151,1437,294,825]" box="[199,1161,367,393]" pageId="62" pageNumber="63">
Limbs (
<figureCitation id="EA8DAAA9450BFF82FECB1A21FE312CBC" box="[285,447,367,393]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C450BFF82FF111ADDFDCA2F08" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
A2 (
<figureCitation id="EA8DAAA9450BFF82FF2F1ADDFEF22C98" box="[249,380,403,429]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA9450BFF82FDA91AF9FD7C2CE4" box="[639,754,439,465]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="62" pageNumber="63">Figure 40</figureCitation>
) with z1claw-like, z2 and z3 being both setae, z1 about 3x as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male and the tip of Gm slightly longer than the tip of y3. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C450BFF82FF111909FB952F9C" blockId="62.[151,1437,294,825]" pageId="62" pageNumber="63">
T1 (
<figureCitation id="EA8DAAA9450BFF82FF221909FEF92F54" box="[244,375,583,609]" captionStart="FIGURE 39" captionStartId="59.[151,250,1960,1984]" captionTargetBox="[175,1409,202,1932]" captionTargetId="figure-17@59.[175,1413,198,1936]" captionTargetPageId="59" captionText="FIGURE 39. Benincandona trapezoides gen. et spec. nov., limbs (AB ♀ and CE ♂). A. T1, Lpp (INV.159280). B. T1, Rpp (INV.159280). C. A2 (INV.159264, INV.159269). D. T1 (INV.159268). E. CR (INV.159264). Scale bar = 50 µm." pageId="62" pageNumber="63">Figure 39D</figureCitation>
) with basal part with
<emphasis id="40C26A3E450BFF82FDB01909FDF42F55" box="[614,634,583,608]" italics="true" pageId="62" pageNumber="63">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long rays. Endopodite set with a short and narrow palp, with three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C450BFF82FF1119FDFD9E2FF8" blockId="62.[151,1437,294,825]" box="[199,528,691,717]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF1119FDFE0E2FF8" bold="true" box="[199,384,691,717]" pageId="62" pageNumber="63">Measurements:</emphasis>
See
<tableCitation id="3F348397450BFF82FE6019FDFD842FF8" box="[438,522,691,717]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="62" pageNumber="63">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7450BFF82FF111998FDEA2FC4" box="[199,612,726,753]" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="7209B62C450BFF82FF111998FDEA2FC4" blockId="62.[151,1437,294,825]" box="[199,612,726,753]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF111998FE5B2FC5" bold="true" box="[199,469,726,752]" pageId="62" pageNumber="63">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F348397450BFF82FDDC1999FDEE2FC4" box="[522,608,727,753]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="62" pageNumber="63">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7450BFF82FF1119B5FB1329F0" pageId="62" pageNumber="63" type="biology_ecology">
<paragraph id="7209B62C450BFF82FF1119B5FE612E21" blockId="62.[151,1437,294,825]" box="[199,495,762,789]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF1119B5FE612E21" bold="true" box="[199,495,762,789]" pageId="62" pageNumber="63">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF111851FC432E0C" blockId="62.[151,1437,294,825]" box="[199,973,799,825]" pageId="62" pageNumber="63">
This subterranean species was thus far only found in its
<typeStatus id="AD0D088E450BFF82FCEA1851FCE22E0C" box="[828,876,799,825]" pageId="62" pageNumber="63">type</typeStatus>
locality.
</paragraph>
<paragraph id="7209B62C450BFF82FF4118C4FD6D2E91" blockId="62.[151,739,906,969]" box="[151,739,906,933]" pageId="62" pageNumber="63">
<emphasis id="40C26A3E450BFF82FF4118C4FD6D2E91" bold="true" box="[151,739,906,933]" pageId="62" pageNumber="63">
Other candonid species from the wells in
<collectingCountry id="0AA1F6BC450BFF82FD4C18C4FD6D2E91" box="[666,739,906,932]" name="Benin" pageId="62" pageNumber="63">Benin</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="7209B62C450BFF82FF4118E1FE152EFC" blockId="62.[151,739,906,969]" box="[151,411,943,969]" pageId="62" pageNumber="63">
(
<figureCitation id="EA8DAAA9450BFF82FF4918E1FE952EFC" box="[159,283,943,969]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
,
<figureCitation id="EA8DAAA9450BFF82FEF018E1FED52EFC" box="[294,347,943,969]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[160,1427,181,1920]" captionTargetId="figure-17@65.[160,1427,181,1920]" captionTargetPageId="65" captionText="FIGURE 43. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). (Continued PAD). Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">43 A</figureCitation>
<collectingRegion id="B07278CE450BFF82FEBC18E1FE1D2EFC" box="[362,403,943,969]" country="Serbia" name="Ada" pageId="62" pageNumber="63">AD</collectingRegion>
)
</paragraph>
<paragraph id="7209B62C450BFF82FF4118B9FB1329F0" blockId="62.[151,1437,1015,1221]" pageId="62" pageNumber="63">
Thirty further candonid species, in addition to the five new species described above, have been found in the samples from the wells in
<collectingCountry id="0AA1F6BC450BFF82FE8F1F55FE2E2900" box="[345,416,1051,1077]" name="Benin" pageId="62" pageNumber="63">Benin</collectingCountry>
. Most of these are also (sub-) triangular in lateral view, with strong (
<figureCitation id="EA8DAAA9450BFF82FB441F55FA852900" box="[1170,1291,1051,1077]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43E, N, AB) or weak (
<figureCitation id="EA8DAAA9450BFF82FF291F71FEF6296C" box="[255,376,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43K, M, Z) external lateral ridges; others have different shapes (
<figureCitation id="EA8DAAA9450BFF82FB9B1F71FB4B296C" box="[1101,1221,1087,1113]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43F, G, O, V, AC). Some species (
<figureCitation id="EA8DAAA9450BFF82FEE81F2DFD892948" box="[318,519,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figure 42A, D, N</figureCitation>
) are much larger than most, and some are much smaller (
<figureCitation id="EA8DAAA9450BFF82FB5A1F2DFA882948" box="[1164,1286,1123,1149]" captionStart="FIGURE 42" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[163,1424,181,1920]" captionTargetId="figure-31@64.[163,1424,181,1920]" captionTargetPageId="64" captionText="FIGURE 42. Scanning electron microscope illustration of fifteen of thirty further undescribed candonid species, (CpRl). Numbered: AO. Scale bar (for all individuals) = 500 µm." pageId="62" pageNumber="63">Figures 42</figureCitation>
/43O, X, AA, AD). Based on preliminary analyses of the soft parts of these species, it is at present most likely that all of these species belong to
<taxonomicName id="B5B6CDAF450BFF82FE891FE5FD6529F0" authority="Hotekpo &amp; Martens" authorityName="Hotekpo &amp; Martens" box="[351,747,1195,1221]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E450BFF82FE891FE5FD8F29F0" box="[351,513,1195,1221]" italics="true" pageId="62" pageNumber="63">Benincandona</emphasis>
Hotèkpo &amp; Martens
</taxonomicName>
, but this requires further confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,522 @@
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<mods:title id="ACA403E50B2C9DF0F1BD0CBD28B463C4">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="7C4A4CDC326A9673CAA81DC6EEFC1E49">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:namePart id="E608BAABC342BA6149AE7066F029391C">Schön, Isa</mods:namePart>
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<mods:affiliation id="55F17097A04677CDAAFF8B0F271814C9">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="8C1FB5ECED6480767B75801876DAF119">Martens, Koen</mods:namePart>
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<mods:affiliation id="176ED5BBAB148F7EAC3EAE71C54C2307">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
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<taxonomicName id="B5B6CDAF451DFF94FF411AB0FE152F2C" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,411,510,537]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451DFF94FF411AB0FE152F2C" bold="true" box="[151,411,510,537]" italics="true" pageId="40" pageNumber="41">Benincandona cebios</emphasis>
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Hotèkpo &amp; Martens
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<subSubSection id="3AACE5A7451DFF94FF411909FE252F54" box="[151,427,583,609]" pageId="40" pageNumber="41" type="description">
<paragraph id="7209B62C451DFF94FF411909FE252F54" blockId="40.[151,985,510,609]" box="[151,427,583,609]" pageId="40" pageNumber="41">
(
<figureCitation id="EA8DAAA9451DFF94FF491909FECA2F54" box="[159,324,583,609]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStart-4="FIGURE 28" captionStart-5="FIGURE 29" captionStart-6="FIGURE 30" captionStart-7="FIGURE 31" captionStartId-0="41.[151,250,1944,1968]" captionStartId-1="42.[151,250,1673,1697]" captionStartId-2="44.[151,250,1673,1697]" captionStartId-3="45.[151,250,1943,1967]" captionStartId-4="46.[151,250,1943,1967]" captionStartId-5="47.[151,250,1981,2005]" captionStartId-6="48.[151,250,1943,1967]" captionStartId-7="49.[151,250,1943,1967]" captionTargetBox-0="[170,1417,181,1921]" captionTargetBox-1="[151,1436,181,1649]" captionTargetBox-2="[151,1436,181,1649]" captionTargetBox-3="[173,1414,183,1915]" captionTargetBox-4="[188,1401,185,1915]" captionTargetBox-5="[197,1421,196,1942]" captionTargetBox-6="[194,1393,191,1910]" captionTargetBox-7="[184,1406,184,1915]" captionTargetId-0="figure-17@41.[170,1417,181,1921]" captionTargetId-1="figure-31@42.[151,1436,181,1649]" captionTargetId-2="figure-31@44.[151,1436,181,1649]" captionTargetId-3="figure-17@45.[171,1417,181,1919]" captionTargetId-4="figure-31@46.[184,1403,181,1919]" captionTargetId-5="figure-17@47.[166,1422,181,1957]" captionTargetId-6="figure-31@48.[184,1403,181,1920]" captionTargetId-7="figure-17@49.[180,1407,181,1919]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionTargetPageId-2="44" captionTargetPageId-3="45" captionTargetPageId-4="46" captionTargetPageId-5="47" captionTargetPageId-6="48" captionTargetPageId-7="49" captionText-0="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." captionText-1="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." captionText-4="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." captionText-5="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." captionText-6="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="40" pageNumber="41">Figures 2431</figureCitation>
,
<figureCitation id="EA8DAAA9451DFF94FE861909FE2D2F54" box="[336,419,583,609]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="40" pageNumber="41">41KN</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF4119C1FC1F2E20" pageId="40" pageNumber="41" type="etymology">
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<emphasis id="40C26A3E451DFF94FF4119C1FE962F9C" bold="true" box="[151,280,655,681]" pageId="40" pageNumber="41">Etymology</emphasis>
</paragraph>
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The species is named after CEBioS, the Belgian project “Capacities for Biodiversity and Sustainable Development”, which funded various field trips in
<collectingCountry id="0AA1F6BC451DFF94FDC91999FDEB2FC4" box="[543,613,727,753]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
, the first visits of SJH to Brussels, and provided a microscope for work in
<collectingCountry id="0AA1F6BC451DFF94FF4119B5FF542E20" box="[151,218,763,789]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
through the GTI program, the “Global Taxonomic Initiative”.
</paragraph>
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<subSubSection id="3AACE5A7451DFF94FF41180CFD732948" pageId="40" pageNumber="41" type="diagnosis">
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<emphasis id="40C26A3E451DFF94FF41180CFE842E69" bold="true" box="[151,266,834,860]" pageId="40" pageNumber="41">Diagnosis</emphasis>
</paragraph>
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CpRl elongated (L/H = 2.2), sub-triangular, with greatest width slightly behind the middle; LV dorsally extending beyond RV, anterior margin bluntly pointed and with anteroventral part straight, posterior margin asymmetrically rounded towards the ventral side, valve surface smooth in the anterior and posterior part, with slight ridges in the middle section. CpD and CpV narrow, with greatest width in the middle and evenly rounded lateral sides, posterior extremities bluntly rounded compared with anterior end pointed. A2 with seta X short, just reaching base of aesthetasc Y. This aesthetasc Y distally asymmetrically bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E451DFF94FA5F1F55FA1B2901" box="[1417,1429,1051,1076]" italics="true" pageId="40" pageNumber="41">c</emphasis>
. 2.1x basal W. T3 with seta d2 short. Hp with ls and ms with overlapping positions, lobe ls broad, distally rounded, extending beyond lobe ms with
<emphasis id="40C26A3E451DFF94FE281F2DFDB92949" box="[510,567,1123,1148]" italics="true" pageId="40" pageNumber="41">circa</emphasis>
half of its length.
</paragraph>
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<emphasis id="40C26A3E451DFF94FF411FE5FEB029F1" bold="true" box="[151,318,1194,1221]" pageId="40" pageNumber="41">Type material</emphasis>
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<emphasis id="40C26A3E451DFF94FF411FBDFE8A2838" bold="true" box="[151,260,1267,1293]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411FBDFE8A2838" box="[151,260,1267,1293]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
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<materialsCitation id="C2DEBC71451DFF94FF411E59FA8C2804" box="[151,1282,1302,1329]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5451DFF94FF411E59FF4D2804" box="[151,195,1303,1329]" count="1" pageId="40" pageNumber="41" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
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.159244).
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</paragraph>
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<emphasis id="40C26A3E451DFF94FF411E10FF72284D" bold="true" box="[151,252,1374,1400]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411E10FF72284D" box="[151,252,1374,1400]" pageId="40" pageNumber="41" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
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<materialsCitation id="C2DEBC71451DFF94FF411ECDFD6E28A8" box="[151,736,1410,1437]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5451DFF94FF411ECDFF4F28A8" box="[151,193,1411,1437]" count="1" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE9451DFF94FD931ECCFDF428A8" box="[581,634,1410,1437]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159245).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C451DFF94FF411E85FE8128D0" blockId="40.[151,1436,1483,1653]" box="[151,271,1483,1509]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411E85FE8128D0" bold="true" box="[151,271,1483,1509]" pageId="40" pageNumber="41">
<typeStatus id="AD0D088E451DFF94FF411E85FE8128D0" box="[151,271,1483,1509]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411EA0FDF62B40" blockId="40.[151,1436,1483,1653]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411EA0FA162B3C" box="[151,1432,1518,1545]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FF411EA0FF572B3C" box="[151,217,1518,1545]" count="5" pageId="40" pageNumber="41" type="male">5 ♂♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E451DFF94FDF51EA1FD0B2B3C" box="[547,645,1519,1545]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9451DFF94FD411EA0FD452B3C" box="[663,715,1518,1545]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159246, INV.159247, INV.159248, INV.159249, INV.159250)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FF411D5DFB5A2B18" box="[151,1236,1554,1581]" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FF411D5DFF542B18" box="[151,218,1555,1581]" count="4" pageId="40" pageNumber="41" type="female">4 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE9451DFF94FDBB1D5CFD2C2B18" box="[621,674,1554,1581]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159251, INV.159252, INV.159253, INV.159254)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FB351D5DFBD92B64" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FB351D5DFAAB2B18" box="[1251,1317,1555,1581]" count="3" pageId="40" pageNumber="41" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE9451DFF94FD5D1D78FD312B64" box="[651,703,1590,1617]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159255, INV.159256, INV.159257)
</materialsCitation>
;
<materialsCitation id="C2DEBC71451DFF94FBB31D79FDFA2B40" collectionCode="INV" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA5451DFF94FBB31D79FB1C2B64" box="[1125,1170,1591,1617]" count="1" pageId="40" pageNumber="41" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE9451DFF94FE0A1D14FD9F2B40" box="[476,529,1626,1653]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="40" pageNumber="41" type="Herbarium">INV</collectionCode>
.159258)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C451DFF94FF411DEDFEBE2B89" blockId="40.[151,1436,1698,1833]" box="[151,304,1698,1725]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411DEDFEBE2B89" bold="true" box="[151,304,1698,1725]" pageId="40" pageNumber="41">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411D89FBAC2A1C" blockId="40.[151,1436,1698,1833]" pageId="40" pageNumber="41">
<materialsCitation id="C2DEBC71451DFF94FF411D89FE652A30" country="Benin" location="Tchaourou District" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC451DFF94FF411D89FF542BD4" box="[151,218,1735,1761]" name="Benin" pageId="40" pageNumber="41">Benin</collectingCountry>
<emphasis id="40C26A3E451DFF94FF371D89FF652BD4" bold="true" box="[225,235,1735,1761]" pageId="40" pageNumber="41"></emphasis>
<collectingRegion id="B07278CE451DFF94FF271D89FE5C2BD4" box="[241,466,1735,1761]" country="Benin" name="Borgou" pageId="40" pageNumber="41">Borgou Department</collectingRegion>
,
<location id="7769E0F7451DFF94FE0A1D89FD3F2BD4" LSID="urn:lsid:plazi:treatment:FA1F073A451DFF8EFF411AB0FEE8284C:7769E0F7451DFF94FE0A1D89FD3F2BD4" box="[476,689,1735,1761]" country="Benin" name="Tchaourou District" pageId="40" pageNumber="41" stateProvince="Borgou">Tchaourou District</location>
, town Badékparou, located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE451DFF94FF091DA5FEBE2A30" box="[223,304,1771,1797]" country="Benin" name="Oueme" pageId="40" pageNumber="41">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB451DFF94FD501DA5FC832A30" box="[646,781,1771,1797]" degrees="9" direction="north" minutes="13.297" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="9.221617">9°13.297N</geoCoordinate>
,
<geoCoordinate id="1782D0EB451DFF94FCCE1DA4FC152A30" box="[792,923,1770,1797]" degrees="2" direction="east" minutes="33.745" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="2.5624166">2°33.745E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC451DFF94FF6C1C41FDCB2A1C" box="[186,581,1807,1833]" pageId="40" pageNumber="41" value="2022-10-11" valueMax="2022-11-06" valueMin="2022-10-11">11 October and 06 November 2022</date>
(Locality code BEN065 in
<tableCitation id="3F348397451DFF94FCAD1C41FC432A1C" box="[891,973,1807,1833]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="40" pageNumber="41">Table 1</tableCitation>
and
<tableCitation id="3F348397451DFF94FBD21C41FB9B2A1C" box="[1028,1045,1807,1833]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="40" pageNumber="41">2</tableCitation>
).
</paragraph>
<paragraph id="7209B62C451DFF94FF411C18FE6E2A45" blockId="40.[151,978,1878,1941]" box="[151,480,1878,1904]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF411C18FE6E2A45" bold="true" box="[151,480,1878,1904]" pageId="40" pageNumber="41">Other material investigated</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF411C35FC5C2AA0" blockId="40.[151,978,1878,1941]" box="[151,978,1915,1941]" pageId="40" pageNumber="41">
See
<tableCitation id="3F348397451DFF94FF101C35FE972AA0" box="[198,281,1915,1941]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="40" pageNumber="41">Table 1</tableCitation>
for a list of localities where the present species was collected.
</paragraph>
</subSubSection>
<caption id="26C9E6A4451CFF95FF411CD6FA052AE0" pageId="41" pageNumber="42" startId="41.[151,250,1944,1968]" targetBox="[170,1417,181,1921]" targetPageId="41" targetType="figure">
<paragraph id="7209B62C451CFF95FF411CD6FA052AE0" blockId="41.[151,1437,1944,2005]" pageId="41" pageNumber="42">
<emphasis id="40C26A3E451CFF95FF411CD6FEAD2A85" bold="true" box="[151,291,1944,1968]" pageId="41" pageNumber="42">FIGURE 24.</emphasis>
<taxonomicName id="B5B6CDAF451CFF95FEFB1CD7FD842A84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1945,1969]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451CFF95FEFB1CD7FD842A84" box="[301,522,1945,1969]" italics="true" pageId="41" pageNumber="42">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451CFF95FDC51CD7FD412A85" box="[531,719,1945,1969]" pageId="41" pageNumber="42" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, JK ♂; DI ♀).
<emphasis id="40C26A3E451CFF95FBF41CD6FBC42A85" bold="true" box="[1058,1098,1944,1968]" pageId="41" pageNumber="42">AI</emphasis>
. CpRl (AC. INV.159255, DI. INV.159258).
<emphasis id="40C26A3E451CFF95FEFE1CF3FEBB2AE0" bold="true" box="[296,309,1981,2005]" pageId="41" pageNumber="42">J</emphasis>
. INV.159256. CpD.
<emphasis id="40C26A3E451CFF95FDD11CF3FD952AE0" bold="true" box="[519,539,1981,2005]" pageId="41" pageNumber="42">K</emphasis>
. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451FFF96FF411DC7FC172BF3" pageId="42" pageNumber="43" startId="42.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="42" targetType="figure">
<paragraph id="7209B62C451FFF96FF411DC7FC172BF3" blockId="42.[151,1437,1673,1734]" pageId="42" pageNumber="43">
<emphasis id="40C26A3E451FFF96FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="42" pageNumber="43">FIGURE 25.</emphasis>
<taxonomicName id="B5B6CDAF451FFF96FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451FFF96FEF11DC7FD8F2B94" box="[295,513,1673,1697]" italics="true" pageId="42" pageNumber="43">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451FFF96FDDE1DC4FD352B94" box="[520,699,1674,1698]" pageId="42" pageNumber="43" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E451FFF96FCEB1DC7FC3E2B94" bold="true" box="[829,944,1673,1698]" pageId="42" pageNumber="43">A, C, D, G</emphasis>
. LVi (INV.159244).
<emphasis id="40C26A3E451FFF96FB551DC4FB402B97" bold="true" box="[1155,1230,1674,1698]" pageId="42" pageNumber="43">B, E, F</emphasis>
. RVi (INV.159244).
<emphasis id="40C26A3E451FFF96FF411DE3FF3F2BF3" bold="true" box="[151,177,1709,1734]" pageId="42" pageNumber="43">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A7451FFF8EFF411C59FD9E29DC" lastPageId="50" lastPageNumber="51" pageId="42" pageNumber="43" type="description">
<paragraph id="7209B62C451FFF96FF411C59FE0F2A07" blockId="42.[151,1436,1815,2022]" box="[151,385,1815,1842]" pageId="42" pageNumber="43">
<emphasis id="40C26A3E451FFF96FF411C59FE0F2A07" bold="true" box="[151,385,1815,1842]" pageId="42" pageNumber="43">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C451FFF97FF411C72FDA32DE0" blockId="42.[151,1436,1815,2022]" lastBlockId="43.[151,1437,151,2013]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
CpRl (
<figureCitation id="EA8DAAA9451FFF96FF351C72FE042A63" box="[227,394,1852,1878]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="42" pageNumber="43">Figure 24AC</figureCitation>
) elongated (L/H ratio=2.0) and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight, posterior part of dorsal margin sloping to ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valve surface smooth in the anterior and posterior parts, with slight ridges in the middle section. CpD and CpV (
<figureCitation id="EA8DAAA9451EFF97FE7E1BD9FDC62D84" box="[424,584,151,177]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="43" pageNumber="44">Figure 24J, K</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins rounded.
</paragraph>
<paragraph id="7209B62C451EFF97FF111B91FEA92CBC" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
LVi (
<figureCitation id="EA8DAAA9451EFF97FED71B91FE7F2DCC" box="[257,497,222,249]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin bluntly pointed, with greatest height slightly behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin straight; posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; one ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA9451EFF97FB711A05FAA42C50" box="[1191,1322,330,357]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25G</figureCitation>
) as in the
<typeStatus id="AD0D088E451EFF97FF411A21FF492CBC" box="[151,199,367,393]" pageId="43" pageNumber="44">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C451EFF97FF111ADDFC882F2C" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
RVi (
<figureCitation id="EA8DAAA9451EFF97FED41ADDFE4A2C98" box="[258,452,402,429]" captionStart="FIGURE 25" captionStartId="42.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@42.[151,1436,181,1649]" captionTargetPageId="42" captionText="FIGURE 25. Benincandona cebios gen. et spec. nov., valves (♂). A, C, D, G. LVi (INV.159244). B, E, F. RVi (INV.159244). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="43" pageNumber="44">Figure 25B, E, F</figureCitation>
) elongated, with dorsal margin partly straight and with greatest height well behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle and with anteroventral part straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C451EFF97FF11196DFC8B2E90" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A1 (
<figureCitation id="EA8DAAA9451EFF97FF2C196DFEF02F08" box="[250,382,547,573]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment with two ventro-subapical setae, one long and one shorter, the latter about one third of the length of the longer seta, and two shorter setae on the mid-dorsal side, situated some distance from each other. First distal segment about three times as long as the basal width, with one short dorso-subapical seta. Second distal segment with about the same length/ width ratio as the previous segment, set with one short dorso-apical seta. Third distal segment also with about the same length/width ratio as the previous segment, carrying one short apical setae on each side. Fourth distal segment more than three times as long as wide, and set with two short setae as on the previous segment. Fifth distal segment more than five times as long as wide, set with one short ventro-apical seta and two long dorso-apical plumose setae. Sixth distal segment narrow, more than four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long but unequal length plumose setae, about twice and 1.5x the length of Ya respectively, and one short plumose seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C451EFF97FF1118E1FA9A28D0" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
A2 (
<figureCitation id="EA8DAAA9451EFF97FF2D18E1FE0C2EFC" box="[251,386,943,969]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27A</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E451EFF97FBD2189DFB992ED9" box="[1028,1047,979,1004]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
five times the length of the more apical one (plumose). Second segment (second protopodite) large and
<emphasis id="40C26A3E451EFF97FC1E18B9FC522925" box="[968,988,1015,1040]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
twice as long as basal width, set with one short and plumose ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout,
<emphasis id="40C26A3E451EFF97FB911F71FBD4296D" box="[1095,1114,1087,1112]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
three times as long as basal width, carrying one long mid-ventral aesthetasc Y (tip clearly exceeding half of segment) and two ventro-apical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E451EFF97FA931FC9FAD62995" box="[1349,1368,1159,1184]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical plumose seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 respectively ventro and centro-apical, hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and about 1.5x as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9451EFF97FB101E59FAB72804" box="[1222,1337,1303,1329]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="43" pageNumber="44">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into short seta of almost equal length; centrally with one short plumose seta z3; dorso-apically, z1 claw-like, length about 1.7x as long as penultimate segment. Claw G2 about 2.5 times as long as z1 with G1 slightly shorter; z2 seta transformed into long claw, reaching tip of G1. Fourth, terminal segment (segment 3) with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a seta g (slightly exceeding tip of aesthetasc y3).
</paragraph>
<paragraph id="7209B62C451EFF97FF111EA1FAA52B18" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdCoxa (
<figureCitation id="EA8DAAA9451EFF97FEED1EA1FE4E2B3C" box="[315,448,1519,1545]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[173,1414,183,1915]" captionTargetId="figure-17@45.[171,1417,181,1919]" captionTargetPageId="45" captionText="FIGURE 27. Benincandona cebios gen. et spec. nov., limbs (♂). A. A2 (INV.159244). B. A1 (INV.159244). C. MdCoxa (INV.159248). Scale bar = 50 µm." pageId="43" pageNumber="44">Figure 27C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
</paragraph>
<paragraph id="7209B62C451EFF97FF111D79FAEA2A78" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
MdPalp (
<figureCitation id="EA8DAAA9451EFF97FEF91D79FE3F2B64" box="[303,433,1591,1617]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28C</figureCitation>
) with four segments. Basal segment ventro-apically with two hirsute setae (S1 &amp; S2); S1 long, reaching tip of terminal segment and S2 short and stout, length about one fifth of the length of S1; in between with the short flagellum-like alpha seta (thin and about twice the length of S2), and one smooth and medium size, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length; ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with one seta of medium length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, slightly shorter than fused claw.
</paragraph>
<paragraph id="7209B62C451EFF97FF111C19FF6B2AE8" blockId="43.[151,1437,151,2013]" pageId="43" pageNumber="44">
Mx1 (
<figureCitation id="EA8DAAA9451EFF97FEDC1C19FE232A44" box="[266,429,1879,1905]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28A, B</figureCitation>
) with first palp segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like. Respiratory plate (
<figureCitation id="EA8DAAA9451EFF97FBE61C35FB3D2AA0" box="[1072,1203,1915,1941]" captionStart="FIGURE 28" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[188,1401,185,1915]" captionTargetId="figure-31@46.[184,1403,181,1919]" captionTargetPageId="46" captionText="FIGURE 28. Benincandona cebios gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159244). B. Mx1, respiratory plate (INV.159248). C. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm." pageId="43" pageNumber="44">Figure 28B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E451EFF97FF411CD1FF242A8D" box="[151,170,1951,1976]" italics="true" pageId="43" pageNumber="44">c.</emphasis>
16 plumose respiratory rays, two short distal setae and a group of about seven shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44519FF90FF411DC7FC172BF3" pageId="44" pageNumber="45" startId="44.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="44" targetType="figure">
<paragraph id="7209B62C4519FF90FF411DC7FC172BF3" blockId="44.[151,1437,1673,1734]" pageId="44" pageNumber="45">
<emphasis id="40C26A3E4519FF90FF411DC7FEAE2B94" bold="true" box="[151,288,1673,1697]" pageId="44" pageNumber="45">FIGURE 26.</emphasis>
<taxonomicName id="B5B6CDAF4519FF90FEF11DC7FD8F2B94" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1673,1697]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4519FF90FEF11DC7FD8F2B94" box="[295,513,1673,1697]" italics="true" pageId="44" pageNumber="45">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454519FF90FDDE1DC4FD352B94" box="[520,699,1674,1698]" pageId="44" pageNumber="45" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4519FF90FCEB1DC7FC3E2B94" bold="true" box="[829,944,1673,1698]" pageId="44" pageNumber="45">A, C, D, G</emphasis>
. LVi (INV.159253).
<emphasis id="40C26A3E4519FF90FB551DC4FB402B97" bold="true" box="[1155,1230,1674,1698]" pageId="44" pageNumber="45">B, E, F</emphasis>
. RVi (INV.159253).
<emphasis id="40C26A3E4519FF90FF411DE3FF3F2BF3" bold="true" box="[151,177,1709,1734]" pageId="44" pageNumber="45">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4519FF90FF111C56FB2B2AF7" blockId="44.[151,1436,1816,1986]" pageId="44" pageNumber="45">
T1 (
<figureCitation id="EA8DAAA94519FF90FF231C56FE142A07" box="[245,410,1816,1842]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented by three plumose setae, two short and one medium length. Protopodite (
<figureCitation id="EA8DAAA94519FF90FC011C72FBD42A63" box="[983,1114,1852,1878]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
) distally set with almost 10 plumose setae of different morphology and length; more proximally set with d, d and b setae. Prehensile palps asymmetrical with Rpp (
<figureCitation id="EA8DAAA94519FF90FE711CCAFDA92AAB" box="[423,551,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94519FF90FC4A1CCAFBAE2AAB" box="[924,1056,1924,1950]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="44" pageNumber="45">Figure 31A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<caption id="26C9E6A44518FF91FF411CD9FE7C2AE1" pageId="45" pageNumber="46" startId="45.[151,250,1943,1967]" targetBox="[173,1414,183,1915]" targetPageId="45" targetType="figure">
<paragraph id="7209B62C4518FF91FF411CD9FE7C2AE1" blockId="45.[151,1437,1943,2004]" pageId="45" pageNumber="46">
<emphasis id="40C26A3E4518FF91FF411CD9FEAA2A9A" bold="true" box="[151,292,1943,1967]" pageId="45" pageNumber="46">FIGURE 27.</emphasis>
<taxonomicName id="B5B6CDAF4518FF91FEFB1CD6FD842A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,522,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4518FF91FEFB1CD6FD842A85" box="[301,522,1944,1968]" italics="true" pageId="45" pageNumber="46">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454518FF91FDC21CD6FD5E2A9A" box="[532,720,1944,1968]" pageId="45" pageNumber="46" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4518FF91FC821CD9FCE82A9A" bold="true" box="[852,870,1943,1967]" pageId="45" pageNumber="46">A</emphasis>
. A2 (INV.159244).
<emphasis id="40C26A3E4518FF91FBED1CD6FBC22A85" bold="true" box="[1083,1100,1944,1968]" pageId="45" pageNumber="46">B</emphasis>
. A1 (INV.159244).
<emphasis id="40C26A3E4518FF91FAF71CD9FABD2A9A" bold="true" box="[1313,1331,1943,1967]" pageId="45" pageNumber="46">C</emphasis>
. MdCoxa (INV.159248). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451BFF92FF411CD9FC5F2AE1" pageId="46" pageNumber="47" startId="46.[151,250,1943,1967]" targetBox="[188,1401,185,1915]" targetPageId="46" targetType="figure">
<paragraph id="7209B62C451BFF92FF411CD9FC5F2AE1" blockId="46.[151,1436,1943,2004]" pageId="46" pageNumber="47">
<emphasis id="40C26A3E451BFF92FF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="46" pageNumber="47">FIGURE 28.</emphasis>
<taxonomicName id="B5B6CDAF451BFF92FEF91CD6FD802A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[303,526,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451BFF92FEF91CD6FD802A85" box="[303,526,1944,1968]" italics="true" pageId="46" pageNumber="47">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451BFF92FDCE1CD6FD562A9A" box="[536,728,1944,1968]" pageId="46" pageNumber="47" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E451BFF92FC881CD9FCFE2A9A" bold="true" box="[862,880,1943,1967]" pageId="46" pageNumber="47">A</emphasis>
. Mx1-palp (INV.159244).
<emphasis id="40C26A3E451BFF92FB591CD6FB2E2A85" bold="true" box="[1167,1184,1944,1968]" pageId="46" pageNumber="47">B</emphasis>
. Mx1, respiratory plate (INV.159248).
<emphasis id="40C26A3E451BFF92FEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="46" pageNumber="47">C</emphasis>
. MdPalp (INV.159244). Scale bars. 1 (B), 2 (A), 3 (C) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4451AFF93FF411CF3FD5D2ACF" pageId="47" pageNumber="48" startId="47.[151,250,1981,2005]" targetBox="[197,1421,196,1942]" targetPageId="47" targetType="figure">
<paragraph id="7209B62C451AFF93FF411CF3FD5D2ACF" blockId="47.[151,1436,1981,2042]" pageId="47" pageNumber="48">
<emphasis id="40C26A3E451AFF93FF411CF3FEAB2AE0" bold="true" box="[151,293,1981,2005]" pageId="47" pageNumber="48">FIGURE 29.</emphasis>
<taxonomicName id="B5B6CDAF451AFF93FEE61CF3FD812AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[304,527,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E451AFF93FEE61CF3FD812AE0" box="[304,527,1981,2005]" italics="true" pageId="47" pageNumber="48">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745451AFF93FDCC1CF0FD552AE0" box="[538,731,1982,2006]" pageId="47" pageNumber="48" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E451AFF93FCB51CF3FCFB2AE0" bold="true" box="[867,885,1981,2005]" pageId="47" pageNumber="48">A</emphasis>
. T3 (INV.159244).
<emphasis id="40C26A3E451AFF93FB9B1CF0FBD02AE3" bold="true" box="[1101,1118,1982,2006]" pageId="47" pageNumber="48">B</emphasis>
. T2 (INV.159244).
<emphasis id="40C26A3E451AFF93FAE01CF3FAC62AE0" bold="true" box="[1334,1352,1981,2005]" pageId="47" pageNumber="48">C</emphasis>
. CRAtt (INV.159244).
<emphasis id="40C26A3E451AFF93FEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="47" pageNumber="48">D</emphasis>
. CR (INV.159244). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44505FF8CFF411CD9FDB32ACD" pageId="48" pageNumber="49" startId="48.[151,250,1943,1967]" targetBox="[194,1393,191,1910]" targetPageId="48" targetType="figure">
<paragraph id="7209B62C4505FF8CFF411CD9FDB32ACD" blockId="48.[151,1437,1943,2040]" pageId="48" pageNumber="49">
<emphasis id="40C26A3E4505FF8CFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="48" pageNumber="49">FIGURE 30.</emphasis>
<taxonomicName id="B5B6CDAF4505FF8CFEFA1CD6FD872A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[300,521,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4505FF8CFEFA1CD6FD872A85" box="[300,521,1944,1968]" italics="true" pageId="48" pageNumber="49">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454505FF8CFDC41CD6FD432A9A" box="[530,717,1944,1968]" pageId="48" pageNumber="49" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4505FF8CFC981CD9FCEE2A9A" bold="true" box="[846,864,1943,1967]" pageId="48" pageNumber="49">A</emphasis>
. Zenker organ (INV.159247).
<emphasis id="40C26A3E4505FF8CFB771CD6FB5F2A85" bold="true" box="[1185,1233,1944,1968]" pageId="48" pageNumber="49">BD</emphasis>
. Hp (INV.159244).
<emphasis id="40C26A3E4505FF8CFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="48" pageNumber="49">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4505FF8CFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="48" pageNumber="49">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E4505FF8CFBEE1CF2FBC42AE1" bold="true" box="[1080,1098,1980,2004]" pageId="48" pageNumber="49">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44504FF8DFF411CD9FBDC2AE1" pageId="49" pageNumber="50" startId="49.[151,250,1943,1967]" targetBox="[184,1406,184,1915]" targetPageId="49" targetType="figure">
<paragraph id="7209B62C4504FF8DFF411CD9FBDC2AE1" blockId="49.[151,1437,1943,2004]" pageId="49" pageNumber="50">
<emphasis id="40C26A3E4504FF8DFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="49" pageNumber="50">FIGURE 31.</emphasis>
<taxonomicName id="B5B6CDAF4504FF8DFEF11CD6FD8F2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E4504FF8DFEF11CD6FD8F2A85" box="[295,513,1944,1968]" italics="true" pageId="49" pageNumber="50">Benincandona cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454504FF8DFDD11CD6FD342A9A" box="[519,698,1944,1968]" pageId="49" pageNumber="50" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4504FF8DFC351CD9FC7B2A9A" bold="true" box="[995,1013,1943,1967]" pageId="49" pageNumber="50">C</emphasis>
. A2 (INV.159245).
<emphasis id="40C26A3E4504FF8DFB171CD6FB5D2A85" bold="true" box="[1217,1235,1944,1968]" pageId="49" pageNumber="50">D</emphasis>
. CR (INV.159245).
<emphasis id="40C26A3E4504FF8DFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="49" pageNumber="50">E</emphasis>
. T1 (INV.159245).
<emphasis id="40C26A3E4504FF8DFEA61CF5FE0C2AE6" bold="true" box="[368,386,1979,2003]" pageId="49" pageNumber="50">A</emphasis>
. T1, Lpp (INV.159244).
<emphasis id="40C26A3E4504FF8DFD561CF2FD1F2AE1" bold="true" box="[640,657,1980,2004]" pageId="49" pageNumber="50">B</emphasis>
. T1, Rpp (INV.159244). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4507FF8EFF111BD9FC692C50" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T2 (
<figureCitation id="EA8DAAA94507FF8EFF231BD9FEF92D84" box="[245,375,151,177]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29B</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment without setae d1 and d2. First segment of endopodite elongated (length almost 4x the basal width), without seta. Second endopodal segment (segment 2A), with length
<emphasis id="40C26A3E4507FF8EFCEC1B91FCC32DCD" box="[826,845,223,248]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
3x basal width, set with one short ventro-apical seta f. Third endopodal segment (segment 2B) of about the same length/width ratio as the previous segment, ventro-subapically set with a g-seta. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111A21FF642F08" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T3 (
<figureCitation id="EA8DAAA94507FF8EFF211A21FEF02CBC" box="[247,382,367,393]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29A</figureCitation>
) with first segment carrying only one subapical seta d1, of medium length. Second (knee-) segment with short seta d2. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower and at least 5x basal width, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, length about twice as long as basal width, carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two short ventrally directed and curved setae h1 &amp; h2, the latter slightly exceeding tip of the former.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111909FC862F9C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
CR (
<figureCitation id="EA8DAAA94507FF8EFF2C1909FEF32F54" box="[250,381,583,609]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29D</figureCitation>
) with stout ramus; claw Ga well-developed, stout with length slightly exceeding half the length of ramus, setae Sp, Sa and claw Gp reduced to small setae; seta Sp situated about at one-fifth length of ramus from distal end, Sa dorso-subapical and Gp ventro-subapical.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1119FDFA832FF8" blockId="50.[151,1437,151,1401]" box="[199,1293,691,717]" pageId="50" pageNumber="51">
CRAtt (
<figureCitation id="EA8DAAA94507FF8EFEC919FDFE2F2FF8" box="[287,417,691,717]" captionStart="FIGURE 29" captionStartId="47.[151,250,1981,2005]" captionTargetBox="[197,1421,196,1942]" captionTargetId="figure-17@47.[166,1422,181,1957]" captionTargetPageId="47" captionText="FIGURE 29. Benincandona cebios gen. et spec. nov., limbs (♂). A. T3 (INV.159244). B. T2 (INV.159244). C. CRAtt (INV.159244). D. CR (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 29C</figureCitation>
) long and stout, distally bifurcated, subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111999FD772E0C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
Hp (
<figureCitation id="EA8DAAA94507FF8EFF2C1999FE2E2FC4" box="[250,416,727,753]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30BD</figureCitation>
) with robust lobe ls, apically rounded, reaching well beyond symmetrically rounded lobe ms, both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94507FF8EFB3819B5FA1A2E20" box="[1262,1428,763,789]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11180DFBDD2E68" blockId="50.[151,1437,151,1401]" box="[199,1107,835,861]" pageId="50" pageNumber="51">
Zenker organ (
<figureCitation id="EA8DAAA94507FF8EFEBD180DFE612E68" box="[363,495,835,861]" captionStart="FIGURE 30" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[194,1393,191,1910]" captionTargetId="figure-31@48.[184,1403,181,1920]" captionTargetPageId="48" captionText="FIGURE 30. Benincandona cebios gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159247). BD. Hp (INV.159244). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 30A</figureCitation>
) long and slightly broad, with
<emphasis id="40C26A3E4507FF8EFC91180DFCD52E69" box="[839,859,835,860]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
seven spinous whirls.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111828FE482EB5" blockId="50.[151,1437,151,1401]" box="[199,454,870,896]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111828FE482EB5" bold="true" box="[199,454,870,896]" pageId="50" pageNumber="51">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118C5FB422E90" blockId="50.[151,1437,151,1401]" box="[199,1228,907,933]" pageId="50" pageNumber="51">
Cp and valves (
<figureCitation id="EA8DAAA94507FF8EFEA218C5FD952E90" box="[372,539,907,933]" captionStart="FIGURE 24" captionStartId="41.[151,250,1944,1968]" captionTargetBox="[170,1417,181,1921]" captionTargetId="figure-17@41.[170,1417,181,1921]" captionTargetPageId="41" captionText="FIGURE 24. Benincandona cebios gen. et spec. nov., valves (AC, JK ♂; DI ♀). AI. CpRl (AC. INV.159255, DI. INV.159258). J. INV.159256. CpD. K. INV.159257. CpV. Scale bars. 1 (B, E, J, K) = 500 µm; 2 (A, C, D, F, H, I) = 200 µm." pageId="50" pageNumber="51">Figures 24DI</figureCitation>
,
<figureCitation id="EA8DAAA94507FF8EFDF118C5FDF72E90" box="[551,633,907,933]" captionStart="FIGURE 26" captionStartId="44.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-31@44.[151,1436,181,1649]" captionTargetPageId="44" captionText="FIGURE 26. Benincandona cebios gen. et spec. nov., valves (♀). A, C, D, G. LVi (INV.159253). B, E, F. RVi (INV.159253). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="50" pageNumber="51">26AG</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C4507FF8EFF1118E1FB072EFC" blockId="50.[151,1437,151,1401]" box="[199,1161,943,969]" pageId="50" pageNumber="51">
Limbs (
<figureCitation id="EA8DAAA94507FF8EFECB18E1FE302EFC" box="[285,446,943,969]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4507FF8EFF11189DFB53296C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
A2 (
<figureCitation id="EA8DAAA94507FF8EFF2F189DFEF22ED8" box="[249,380,979,1005]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31C</figureCitation>
) with second endopodal segment undivided, with t1-seta (long, narrow and plumose) and t2- seta (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94507FF8EFDA418B9FD6C2924" box="[626,738,1015,1041]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="50" pageNumber="51">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 about twice as long as z2, and z3 slightly shorter than z2; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4507FF8EFF111F2DFBDF29F0" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
T1 (
<figureCitation id="EA8DAAA94507FF8EFF221F2DFEFA2948" box="[244,372,1123,1149]" captionStart="FIGURE 31" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[184,1406,184,1915]" captionTargetId="figure-17@49.[180,1407,181,1919]" captionTargetPageId="49" captionText="FIGURE 31. Benincandona cebios gen. et spec. nov., limbs (AB ♂ and CE ♀). C. A2 (INV.159245). D. CR (INV.159245). E. T1 (INV.159245). A. T1, Lpp (INV.159244). B. T1, Rpp (INV.159244). Scale bar = 50 µm." pageId="50" pageNumber="51">Figure 31E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4507FF8EFDB21F2DFDF62949" box="[612,632,1123,1148]" italics="true" pageId="50" pageNumber="51">c.</emphasis>
10 distal setae of unequal length and morphology, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate represented by two long and one short rays. Endopodite with a broad palp, carrying three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C4507FF8EFF111F81FD9E29DC" blockId="50.[151,1437,151,1401]" box="[199,528,1231,1257]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111F81FE0E29DC" bold="true" box="[199,384,1231,1257]" pageId="50" pageNumber="51">Measurements:</emphasis>
See
<tableCitation id="3F3483974507FF8EFE601F81FD8429DC" box="[438,522,1231,1257]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="50" pageNumber="51">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF111FBCFDEA2838" box="[199,612,1266,1293]" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="7209B62C4507FF8EFF111FBCFDEA2838" blockId="50.[151,1437,151,1401]" box="[199,612,1266,1293]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111FBCFE5B2839" bold="true" box="[199,469,1266,1292]" pageId="50" pageNumber="51">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974507FF8EFDDC1FBDFDEE2838" box="[522,608,1267,1293]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="50" pageNumber="51">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74507FF8EFF111E59FEE8284C" pageId="50" pageNumber="51" type="biology_ecology">
<paragraph id="7209B62C4507FF8EFF111E59FE612805" blockId="50.[151,1437,151,1401]" box="[199,495,1302,1329]" pageId="50" pageNumber="51">
<emphasis id="40C26A3E4507FF8EFF111E59FE612805" bold="true" box="[199,495,1302,1329]" pageId="50" pageNumber="51">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4507FF8EFF111E75FEE8284C" blockId="50.[151,1437,151,1401]" pageId="50" pageNumber="51">
This subterranean species is presently known from its
<typeStatus id="AD0D088E4507FF8EFCFE1E75FCD62860" box="[808,856,1339,1365]" pageId="50" pageNumber="51">type</typeStatus>
locality and from a second, closely situated station, only (see
<tableCitation id="3F3483974507FF8EFED21E11FEDB284C" box="[260,341,1375,1401]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="50" pageNumber="51">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F4A5C34D890A7CC3606D137ED0E0596E">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<taxonomicName id="B5B6CDAF4520FFA9FF41190AFE442F6A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,458,580,607]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FF41190AFE442F6A" bold="true" box="[151,458,580,607]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
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Hotèkpo &amp; Martens
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</subSubSection>
<subSubSection id="3AACE5A74520FFA9FF4119C3FE2D2F92" box="[151,419,653,680]" pageId="21" pageNumber="22" type="description">
<paragraph id="7209B62C4520FFA9FF4119C3FE2D2F92" blockId="21.[151,980,580,680]" box="[151,419,653,680]" pageId="21" pageNumber="22">
(
<figureCitation id="EA8DAAA94520FFA9FF4919C3FECA2F9D" box="[159,324,653,680]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStartId-0="21.[151,250,1855,1879]" captionStartId-1="22.[151,250,1943,1967]" captionStartId-2="25.[151,250,1981,2005]" captionStartId-3="26.[151,250,1943,1967]" captionStartId-4="27.[151,250,1943,1967]" captionStartId-5="28.[151,250,1943,1967]" captionTargetBox-0="[151,1436,883,1832]" captionTargetBox-1="[183,1405,185,1915]" captionTargetBox-2="[166,1422,183,1955]" captionTargetBox-3="[191,1395,185,1916]" captionTargetBox-4="[229,1359,191,1909]" captionTargetBox-5="[176,1407,181,1915]" captionTargetId-0="figure-159@21.[151,1436,883,1832]" captionTargetId-1="figure-31@22.[179,1407,181,1919]" captionTargetId-2="figure-17@25.[166,1422,181,1957]" captionTargetId-3="figure-31@26.[187,1399,181,1919]" captionTargetId-4="figure-17@27.[219,1369,181,1919]" captionTargetId-5="figure-31@28.[176,1411,181,1919]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionTargetPageId-5="28" captionText-0="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." captionText-1="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." captionText-2="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-3="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." captionText-4="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-5="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="21" pageNumber="22">
Figures 10
<geoCoordinate id="1782D0EB4520FFA9FEFF19C3FECA2F9D" box="[297,324,653,680]" degrees="15, 41" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="15.41">15</geoCoordinate>
</figureCitation>
,
<figureCitation id="EA8DAAA94520FFA9FE8619C3FE122F92" box="[336,412,653,679]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="21" pageNumber="22">41E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74520FFA9FF41199BFEB92E6E" pageId="21" pageNumber="22" type="etymology">
<paragraph id="7209B62C4520FFA9FF41199BFE962FDA" blockId="21.[151,1436,725,859]" box="[151,280,725,751]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF41199BFE962FDA" bold="true" box="[151,280,725,751]" pageId="21" pageNumber="22">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4520FFA9FF4119B7FEB92E6E" blockId="21.[151,1436,725,859]" pageId="21" pageNumber="22">
The species is named after Prof. Moudachirou Ibikounlé (Université dAbomey Calavi,
<collectingCountry id="0AA1F6BC4520FFA9FB5E19B7FB432E26" box="[1160,1229,761,787]" name="Benin" pageId="21" pageNumber="22">Bénin</collectingCountry>
) in recognition of his contribution to the advancement of scientific research in
<collectingCountry id="0AA1F6BC4520FFA9FCEF1853FCF12E02" box="[825,895,797,823]" name="Benin" pageId="21" pageNumber="22">Benin</collectingCountry>
, and in gratitude for his continued support to the career of SJH.
</paragraph>
</subSubSection>
<caption id="26C9E6A44520FFA9FF411C71FCDB2A49" pageId="21" pageNumber="22" startId="21.[151,250,1855,1879]" targetBox="[151,1436,883,1832]" targetPageId="21" targetType="figure">
<paragraph id="7209B62C4520FFA9FF411C71FCDB2A49" blockId="21.[151,1437,1855,1916]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF411C71FEAE2A62" bold="true" box="[151,288,1855,1879]" pageId="21" pageNumber="22">FIGURE 10.</emphasis>
<taxonomicName id="B5B6CDAF4520FFA9FEF01C71FDAD2A62" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,547,1855,1879]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FEF01C71FDAD2A62" box="[294,547,1855,1879]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454520FFA9FDFC1C0EFD552A62" box="[554,731,1856,1880]" pageId="21" pageNumber="22" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves.
<emphasis id="40C26A3E4520FFA9FCE41C71FCCA2A62" bold="true" box="[818,836,1855,1879]" pageId="21" pageNumber="22">A</emphasis>
. INV.159222. CpRl.
<emphasis id="40C26A3E4520FFA9FBCA1C71FBA32A62" bold="true" box="[1052,1069,1855,1879]" pageId="21" pageNumber="22">B</emphasis>
. INV.159222. CpV.
<emphasis id="40C26A3E4520FFA9FB2D1C71FA832A62" bold="true" box="[1275,1293,1855,1879]" pageId="21" pageNumber="22">C</emphasis>
. INV.159223. CpD.
<emphasis id="40C26A3E4520FFA9FF021C2DFF682A4E" bold="true" box="[212,230,1891,1915]" pageId="21" pageNumber="22">D</emphasis>
. INV.159224. LVi.
<emphasis id="40C26A3E4520FFA9FE7B1C2DFE302A4E" bold="true" box="[429,446,1891,1915]" pageId="21" pageNumber="22">E</emphasis>
. INV.159224. RVi. Scale bar = 500 µm.
</paragraph>
</caption>
<caption id="26C9E6A44523FFAAFF411CD9FC362AE1" pageId="22" pageNumber="23" startId="22.[151,250,1943,1967]" targetBox="[183,1405,185,1915]" targetPageId="22" targetType="figure">
<paragraph id="7209B62C4523FFAAFF411CD9FC362AE1" blockId="22.[151,1436,1943,2004]" pageId="22" pageNumber="23">
<emphasis id="40C26A3E4523FFAAFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="22" pageNumber="23">FIGURE 11.</emphasis>
<taxonomicName id="B5B6CDAF4523FFAAFEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4523FFAAFEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="22" pageNumber="23">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454523FFAAFDE71CD6FD662A9A" box="[561,744,1944,1968]" pageId="22" pageNumber="23" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4523FFAAFCB11CD9FCF72A9A" bold="true" box="[871,889,1943,1967]" pageId="22" pageNumber="23">A</emphasis>
. A2 (INV.159219, INV.159217, INV.159218).
<emphasis id="40C26A3E4523FFAAFA891CD6FAFE2A85" bold="true" box="[1375,1392,1944,1968]" pageId="22" pageNumber="23">B</emphasis>
. A1 (INV.159219, INV.159218).
<emphasis id="40C26A3E4523FFAAFE6C1CF5FE652AE1" bold="true" box="[442,491,1979,2004]" pageId="22" pageNumber="23">C, D</emphasis>
. MdCoxa, (INV.159218). Scale bar = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A74522FFABFF411BD8FD132CBC" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="7209B62C4522FFABFF411BD8FE842D85" blockId="23.[151,1436,150,393]" box="[151,266,150,176]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411BD8FE842D85" bold="true" box="[151,266,150,176]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411BF5FD132CBC" blockId="23.[151,1436,150,393]" pageId="23" pageNumber="24">CpRl elongated (L/H = 2.4), sub-triangular, with greatest height slightly in front of the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, valve surface smooth. CpD and CpV narrow, with greatest width in the middle and angular lateral sides, anterior and posterior extremities pointed. A2 with seta X short, not reaching base of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with seta d2 absent. Hp with ls and ms with slightly diverging positions, narrower lobe ls on Hp with skewed rectangular distal margin, extending well beyond lobe ms.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74522FFABFF411AF9FC4E296C" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="7209B62C4522FFABFF411AF9FEB02CE5" blockId="23.[151,318,438,465]" box="[151,318,438,465]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AF9FEB02CE5" bold="true" box="[151,318,438,465]" pageId="23" pageNumber="24">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411AB1FE8A2F2C" blockId="23.[151,1394,511,573]" box="[151,260,511,537]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AB1FE8A2F2C" bold="true" box="[151,260,511,537]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411AB1FE8A2F2C" box="[151,260,511,537]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF41196DFAFC2F08" blockId="23.[151,1394,511,573]" box="[151,1394,546,573]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF41196DFAFC2F08" box="[151,1394,546,573]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA54522FFABFF41196DFF4D2F08" box="[151,195,547,573]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, specimen with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE94522FFABFB01196DFA892F08" box="[1239,1287,547,573]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159215).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF411924FF722FB1" blockId="23.[151,1436,618,717]" box="[151,252,618,644]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411924FF722FB1" bold="true" box="[151,252,618,644]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411924FF722FB1" box="[151,252,618,644]" pageId="23" pageNumber="24" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119C1FEE92FF8" blockId="23.[151,1436,618,717]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4119C1FEE92FF8" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA54522FFABFF4119C1FF4F2F9C" box="[151,193,655,681]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, soft part dissected as the male (
<collectionCode id="14A72EE94522FFABFDFA19C0FDEF2F9C" box="[556,609,654,681]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159216). Valves lost: the decalcified valves could not be preserved during the manipulations.
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119B5FE812E20" blockId="23.[151,1436,763,933]" box="[151,271,763,789]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF4119B5FE812E20" bold="true" box="[151,271,763,789]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF4119B5FE812E20" box="[151,271,763,789]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411851FEBF2E68" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF411851FC4A2E0C" box="[151,964,798,825]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFF411851FF582E0C" box="[151,214,799,825]" count="3" pageId="23" pageNumber="24" type="male">3 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E4522FFABFE591851FE7F2E0C" box="[399,497,799,825]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94522FFABFE291850FDBD2E0C" box="[511,563,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159217, INV.159218, INV.159219)
</materialsCitation>
;
<materialsCitation id="C2DEBC714522FFABFC181851FEA32E68" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFC181851FB832E0C" box="[974,1037,799,825]" count="2" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE94522FFABFADC1850FAB02E0C" box="[1290,1342,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159220, INV.159221)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C4522FFABFF111829FE872E90" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
The
<typeStatus id="AD0D088E4522FFABFF291829FEEF2EB4" box="[255,353,871,897]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
male comprises both valves and dissected soft parts, but no illustrations are based on this specimen.
</paragraph>
<paragraph id="7209B62C4522FFABFF41189DFEBE2ED9" blockId="23.[151,1436,978,1113]" box="[151,304,978,1005]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF41189DFEBE2ED9" bold="true" box="[151,304,978,1005]" pageId="23" pageNumber="24">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4118B9FC4E296C" blockId="23.[151,1436,978,1113]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4118B9FC32296C" collectingDate="2016-08-09" collectorName="Leg. &amp; S. J. Hotekpo &amp; M. Lagnika &amp; P. Martin" country="Benin" county="Colline Department" latitude="7.8160834" location="Tchetti" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Zou">
<collectingCountry id="0AA1F6BC4522FFABFF4118B9FF542924" box="[151,218,1015,1041]" name="Benin" pageId="23" pageNumber="24">Benin</collectingCountry>
,
<collectingCounty id="9B68CEA04522FFABFF3C18B9FE422924" box="[234,460,1015,1041]" pageId="23" pageNumber="24">Colline Department</collectingCounty>
,
<collectingMunicipality id="926D2C564522FFABFE0F18B9FD1C2924" box="[473,658,1015,1041]" pageId="23" pageNumber="24">Savalou District</collectingMunicipality>
, town of
<location id="7769E0F74522FFABFD2918B9FCC12924" LSID="urn:lsid:plazi:treatment:FA1F073A4520FFA1FF41190AFB072994:7769E0F74522FFABFD2918B9FCC12924" box="[767,847,1015,1041]" country="Benin" county="Colline Department" latitude="7.8160834" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" name="Tchetti" pageId="23" pageNumber="24" stateProvince="Zou">Tchetti</location>
, located in the
<collectingRegion id="B07278CE4522FFABFC2918B9FBA22924" box="[1023,1068,1015,1041]" country="Benin" name="Zou" pageId="23" pageNumber="24">Zou</collectingRegion>
catchment area (tributary of the
<collectingRegion id="B07278CE4522FFABFF411F55FF662900" box="[151,232,1051,1077]" country="Benin" name="Oueme" pageId="23" pageNumber="24">Ouémé</collectingRegion>
catchment area), coordinates:
<geoCoordinate id="1782D0EB4522FFABFD9A1F55FD532900" box="[588,733,1050,1077]" degrees="7" direction="north" minutes="48.965" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="7.8160834">7° 48.965N</geoCoordinate>
,
<geoCoordinate id="1782D0EB4522FFABFD3D1F55FCF92900" box="[747,887,1051,1077]" degrees="1" direction="east" minutes="40.311" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="1.67185">1° 40.311E</geoCoordinate>
,
<collectorName id="DF43D3FA4522FFABFC501F55FC332900" box="[902,957,1051,1077]" pageId="23" pageNumber="24">Leg.</collectorName>
:
<collectorName id="DF43D3FA4522FFABFC1D1F55FBEF2900" box="[971,1121,1051,1077]" pageId="23" pageNumber="24">S.J. Hotèkpo</collectorName>
,
<collectorName id="DF43D3FA4522FFABFBB91F55FB782900" box="[1135,1270,1051,1077]" pageId="23" pageNumber="24">M. Lagnika</collectorName>
and
<collectorName id="DF43D3FA4522FFABFAE41F55FA122900" box="[1330,1436,1051,1077]" pageId="23" pageNumber="24">P. Martin</collectorName>
collected on
<date id="060890EC4522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">
<collectingDate id="164C69044522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">09 August 2016</collectingDate>
</date>
(Locality code BEN072 in
<tableCitation id="3F3483974522FFABFCDB1F71FCE5296C" box="[781,875,1087,1113]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="23" pageNumber="24">Tables 1</tableCitation>
and
<tableCitation id="3F3483974522FFABFC771F71FC3C296C" box="[929,946,1087,1113]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="23" pageNumber="24">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74522FFA1FF411FC8FD9E2900" lastPageId="29" lastPageNumber="30" pageId="23" pageNumber="24" type="description">
<paragraph id="7209B62C4522FFABFF411FC8FE0F2994" blockId="23.[151,1437,1158,2013]" box="[151,385,1158,1185]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411FC8FE0F2994" bold="true" box="[151,385,1158,1185]" pageId="23" pageNumber="24">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411FE5FC9F28A8" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
CpRl (
<figureCitation id="EA8DAAA94522FFABFF331FE5FEE229F0" box="[229,364,1195,1221]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10A</figureCitation>
) elongated (L/H ratio = 2.4) and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height slightly in front of the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valves externally smooth. CpV (
<figureCitation id="EA8DAAA94522FFABFCF11E75FC272860" box="[807,937,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10B</figureCitation>
) and CpD (
<figureCitation id="EA8DAAA94522FFABFBF81E75FB3F2860" box="[1070,1201,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10C</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins angular in the middle; CpV with LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C4522FFABFF111EE9FBEA2B18" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
LVi (
<figureCitation id="EA8DAAA94522FFABFF291EE9FE0A28F4" box="[255,388,1447,1473]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10D</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle showing an overlap, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; one ventral il and antero-ventral tooth.
</paragraph>
<paragraph id="7209B62C4522FFABFF111D79FE062B88" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
RVi (
<figureCitation id="EA8DAAA94522FFABFED71D79FE0C2B64" box="[257,386,1591,1617]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." pageId="23" pageNumber="24">Figure 10E</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C4522FFA4FF111D89FE0D2C74" blockId="23.[151,1437,1158,2013]" lastBlockId="24.[151,1437,151,1905]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
A1 (
<figureCitation id="EA8DAAA94522FFABFF2E1D89FEF62BD4" box="[248,376,1735,1761]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="23" pageNumber="24">Figure 11B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment large with
<emphasis id="40C26A3E4522FFABFA5F1D89FA122BD5" box="[1417,1436,1735,1760]" italics="true" pageId="23" pageNumber="24">c.</emphasis>
2.5 times as long as basal width, supporting two ventro-subapical setae, more proximal about three times as long as second one; and two setae on the dorsal side situated some distance from each other with the more proximal one slightly longer than the second one. First distal segment length about twice the basal width, with one short dorso-apical seta, no ventral apical seta present. Next (second distal segment) about twice as long as wide, with one dorso-apical seta. Third distal segment slightly longer than the previous one, with one short apical seta on each side. Fourth distal segment also slightly longer than the previous one (more than three time as long as wide) with one short ventro-apical seta and one long dorso-apical seta, reaching beyond tip of sixth distal segment with almost half the total length. Fifth distal segment more than four times as long as wide, with two long dorso-apical setae (almost five times longer than the length of the segment) and one short ventro-apical seta slightly exceeding the middle of the next segment. Sixth distal (terminal) segment, narrow,
<emphasis id="40C26A3E452DFFA4FCFC1B91FCB32DCD" box="[810,829,223,248]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as wide, bearing one short and two longer setae and one aesthetasc (Ya). Both long setae about 2.5x the length of Ya, length of the short one is about half the length of Ya.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111A05FBED2E68" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
A2 (
<figureCitation id="EA8DAAA9452DFFA4FF2E1A05FEF52C50" box="[248,379,331,357]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 11A</figureCitation>
) with a basal segment broad and rounded, mid-ventrally with one seta of medium length, two subapical setae of unequal length, proximal one
<emphasis id="40C26A3E452DFFA4FD671A21FD4A2CBD" box="[689,708,367,392]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E452DFFA4FE4D1ADDFE202C99" box="[411,430,403,428]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as basal width, set with one short ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments with the penultimate segment divided. First endopodal segment long
<emphasis id="40C26A3E452DFFA4FEEF1A95FEC22CC1" box="[313,332,475,500]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width, carrying one long mid-ventral aesthetasc (Y) (reaching tip of segment) and two ventro-apical setae, one short and one long, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E452DFFA4FEEF196DFEC32F09" box="[313,333,547,572]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 (ventro-apical) and t3 (centro-apical) hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and almost as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9452DFFA4FACA19C1FA1E2F9C" box="[1308,1424,655,681]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="24" pageNumber="25">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of almost equal length; centrally with one short seta z3; dorso-apically, with z1 claw-like, length twice as long as penultimate segment. Claw G2 about 2.5 times as long as z1; z2 seta transformed into long claw, slightly shorter than G1. Fourth endopodal, terminal, segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and seta g slightly longer than aesthetasc y3.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111829FC302EFC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdCoxa (
<figureCitation id="EA8DAAA9452DFFA4FEEE1829FE522EB4" box="[312,476,871,897]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 11CD</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta. Remark: asymmetry illustrated possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C452DFFA4FF11189DFAEE29DC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdPalp (
<figureCitation id="EA8DAAA9452DFFA4FEE7189DFE3B2ED8" box="[305,437,979,1005]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12C</figureCitation>
) consisting of four segments. Basal segment ventro-apically with two hirsute setae, S1 (long, reaching tip of terminal segment) and S2 (short and stout, length about one third of the length of S1), in between with the short flagellum-like alpha seta (thin and about as long as S2), and one smooth and medium length, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with 2 setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, about as long as the fused claw.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111FBDFC2028D0" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
Mx1 (
<figureCitation id="EA8DAAA9452DFFA4FED91FBDFE322838" box="[271,444,1267,1293]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA9452DFFA4FC5D1E59FB802804" box="[907,1038,1303,1329]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Figure 12B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E452DFFA4FB281E59FA9C2805" box="[1278,1298,1303,1328]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
13 plumose respiratory rays, two short distal setae and a group of four shorter proximal setae of unequal length. Palp (
<figureCitation id="EA8DAAA9452DFFA4FAE31E75FA1A2860" box="[1333,1428,1339,1365]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="24" pageNumber="25">Fig 12A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying also 4 distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E452DFFA4FDF11EE9FD3028F4" box="[551,702,1447,1473]" italics="true" pageId="24" pageNumber="25">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E452DFFA4FDED1E85FDC028D1" box="[571,590,1483,1508]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
half the length of the long one.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111EA1FF582BAC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T1 (
<figureCitation id="EA8DAAA9452DFFA4FF251EA1FE1A2B3C" box="[243,404,1518,1545]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) basal part (protopodite) carrying respiratory plates (exopodites, not shown), distally set with 4+7 setae of different morphology and length; more proximally with setae d and b of medium length. Prehensile palps (
<figureCitation id="EA8DAAA9452DFFA4FF351D79FE052B64" box="[227,395,1590,1617]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA9452DFFA4FCA11D15FC722B40" box="[887,1020,1626,1653]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA9452DFFA4FA871D15FF472BAC" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 15B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C452DFFA4FF111DEDFB5C2A44" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T2 (
<figureCitation id="EA8DAAA9452DFFA4FF231DEDFEF72B88" box="[245,377,1699,1725]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="24" pageNumber="25">Figure 13D</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), elongated, without basal setae. Setae d1 and d2 absent. First endopodal segment without any setae, length
<emphasis id="40C26A3E452DFFA4FC2B1D89FB9E2BD5" box="[1021,1040,1735,1760]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width. Second endopodal segment (2A) set with one short ventro-apical seta f, length
<emphasis id="40C26A3E452DFFA4FBC21DA5FBA92A31" box="[1044,1063,1771,1796]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
3x basal width. Third endopodal segment (2B) of about the same length/width ratio as the previous segment and with one ventro-subapical seta g, reaching about 1/3 of following segment. Terminal segment about 1.5 times as long as basal width, apically two short setae, h1 ventral and h3 dorsal; and a long slender claw (h2), distally set with short setulae.
</paragraph>
<caption id="26C9E6A4452CFFA5FF411CF3FBFF2ACF" pageId="25" pageNumber="26" startId="25.[151,250,1981,2005]" targetBox="[166,1422,183,1955]" targetPageId="25" targetType="figure">
<paragraph id="7209B62C452CFFA5FF411CF3FBFF2ACF" blockId="25.[151,1436,1981,2042]" pageId="25" pageNumber="26">
<emphasis id="40C26A3E452CFFA5FF411CF3FEAE2AE0" bold="true" box="[151,288,1981,2005]" pageId="25" pageNumber="26">FIGURE 12.</emphasis>
<taxonomicName id="B5B6CDAF452CFFA5FEF11CF3FDAA2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,548,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452CFFA5FEF11CF3FDAA2AE0" box="[295,548,1981,2005]" italics="true" pageId="25" pageNumber="26">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452CFFA5FDFD1CF0FD532AE0" box="[555,733,1982,2006]" pageId="25" pageNumber="26" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452CFFA5FC811CF3FCE72AE0" bold="true" box="[855,873,1981,2005]" pageId="25" pageNumber="26">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159218).
<emphasis id="40C26A3E452CFFA5FA9E1CF0FAD72AE3" bold="true" box="[1352,1369,1982,2006]" pageId="25" pageNumber="26">B</emphasis>
. Mx1, respiratory plate (INV.159218).
<emphasis id="40C26A3E452CFFA5FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="25" pageNumber="26">C</emphasis>
. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452FFFA6FF411CD9FD432AE1" pageId="26" pageNumber="27" startId="26.[151,250,1943,1967]" targetBox="[191,1395,185,1916]" targetPageId="26" targetType="figure">
<paragraph id="7209B62C452FFFA6FF411CD9FD432AE1" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="27">
<emphasis id="40C26A3E452FFFA6FF411CD9FEAC2A9A" bold="true" box="[151,290,1943,1967]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
<taxonomicName id="B5B6CDAF452FFFA6FEFD1CD6FDA22A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[299,556,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452FFFA6FEFD1CD6FDA22A85" box="[299,556,1944,1968]" italics="true" pageId="26" pageNumber="27">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452FFFA6FDE31CD6FD602A9A" box="[565,750,1944,1968]" pageId="26" pageNumber="27" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452FFFA6FCB91CD9FC0F2A9A" bold="true" box="[879,897,1943,1967]" pageId="26" pageNumber="27">A</emphasis>
. CR (INV.159219).
<emphasis id="40C26A3E452FFFA6FB811CD6FBE62A85" bold="true" box="[1111,1128,1944,1968]" pageId="26" pageNumber="27">B</emphasis>
. CRAtt (INV.159219).
<emphasis id="40C26A3E452FFFA6FA891CD9FAFF2A9A" bold="true" box="[1375,1393,1943,1967]" pageId="26" pageNumber="27">C</emphasis>
. T3 (INV.159219).
<emphasis id="40C26A3E452FFFA6FEE71CF2FECD2AE1" bold="true" box="[305,323,1980,2004]" pageId="26" pageNumber="27">D</emphasis>
. T2 (INV.159217). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452EFFA7FF411CD9FDB32ACD" pageId="27" pageNumber="28" startId="27.[151,250,1943,1967]" targetBox="[229,1359,191,1909]" targetPageId="27" targetType="figure">
<paragraph id="7209B62C452EFFA7FF411CD9FDB32ACD" blockId="27.[151,1437,1943,2040]" pageId="27" pageNumber="28">
<emphasis id="40C26A3E452EFFA7FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="27" pageNumber="28">FIGURE 14.</emphasis>
<taxonomicName id="B5B6CDAF452EFFA7FEFE1CD9FDA82A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,550,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452EFFA7FEFE1CD9FDA82A9A" box="[296,550,1943,1967]" italics="true" pageId="27" pageNumber="28">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452EFFA7FDFB1CD6FD6F2A9A" box="[557,737,1944,1968]" pageId="27" pageNumber="28" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452EFFA7FC8A1CD9FCE02A9A" bold="true" box="[860,878,1943,1967]" pageId="27" pageNumber="28">A</emphasis>
. Zenker organ (INV.159219).
<emphasis id="40C26A3E452EFFA7FB701CD9FB5B2A9A" bold="true" box="[1190,1237,1943,1967]" pageId="27" pageNumber="28">BD</emphasis>
. Hp (INV.159219).
<emphasis id="40C26A3E452EFFA7FF411CF5FF262AE6" bold="true" box="[151,168,1979,2003]" pageId="27" pageNumber="28">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E452EFFA7FE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="27" pageNumber="28">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E452EFFA7FBEE1CF5FBC42AE6" bold="true" box="[1080,1098,1979,2003]" pageId="27" pageNumber="28">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44529FFA0FF411CD9FB072AE1" pageId="28" pageNumber="29" startId="28.[151,250,1943,1967]" targetBox="[176,1407,181,1915]" targetPageId="28" targetType="figure">
<paragraph id="7209B62C4529FFA0FF411CD9FB072AE1" blockId="28.[151,1436,1943,2004]" pageId="28" pageNumber="29">
<emphasis id="40C26A3E4529FFA0FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
<taxonomicName id="B5B6CDAF4529FFA0FEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4529FFA0FEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="28" pageNumber="29">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454529FFA0FDE61CD6FD6B2A9A" box="[560,741,1944,1968]" pageId="28" pageNumber="29" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4529FFA0FBC21CD9FBA82A9A" bold="true" box="[1044,1062,1943,1967]" pageId="28" pageNumber="29">A</emphasis>
. T1, Lpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FAFC1CD6FAB52A85" bold="true" box="[1322,1339,1944,1968]" pageId="28" pageNumber="29">B</emphasis>
. T1, Rpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="28" pageNumber="29">C</emphasis>
. A2 (INV.159216).
<emphasis id="40C26A3E4529FFA0FDDB1CF2FD912AE1" bold="true" box="[525,543,1980,2004]" pageId="28" pageNumber="29">D</emphasis>
. CR (INV.159216).
<emphasis id="40C26A3E4529FFA0FD381CF2FD712AE1" bold="true" box="[750,767,1980,2004]" pageId="28" pageNumber="29">E</emphasis>
. T1 (INV.159221). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4528FFA1FF111BD9FE902C50" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
T3 (
<figureCitation id="EA8DAAA94528FFA1FF231BD9FEF82D84" box="[245,374,151,177]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13C</figureCitation>
) first segment with only one medium length seta d1. Second (knee-) segment without any seta. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower,
<emphasis id="40C26A3E4528FFA1FB701BF5FB372DE1" box="[1190,1209,187,212]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
7x basal width and slightly longer than the previous segment, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4528FFA1FE241A4DFD8B2C29" box="[498,517,259,284]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), clearly exceeding half of the third segment and two ventrally directed short setae h1 and h2, the former
<emphasis id="40C26A3E4528FFA1FB0B1A69FB7E2C75" box="[1245,1264,295,320]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A21FB082C98" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
CR (
<figureCitation id="EA8DAAA94528FFA1FF2D1A21FEF32CBC" box="[251,381,367,393]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13A</figureCitation>
) with ramus stout; claw Ga well-developed, stout with length ratio ramus/claw Ga = 1.5, setae Sp (posterior seta), Sa (anterior seta) and claw Gp (posterior claw) reduced to small setae.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111AF9FABB2CE4" blockId="29.[151,1437,151,645]" box="[199,1333,439,465]" pageId="29" pageNumber="30">
CRAtt (
<figureCitation id="EA8DAAA94528FFA1FEC91AF9FE2F2CE4" box="[287,417,439,465]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 13B</figureCitation>
) long and stout, distally bifurcated and subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A95FD772F08" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Hp (
<figureCitation id="EA8DAAA94528FFA1FF2F1A95FE102CC0" box="[249,414,475,501]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14BD</figureCitation>
) of medium size, with (ls) relatively narrow, with skewed rectangular distal part, reaching well beyond the asymmetrically rounded ms; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94528FFA1FB241AB1FA1A2F2C" box="[1266,1428,511,537]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111909FCDD2FB0" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Zenker organ (
<figureCitation id="EA8DAAA94528FFA1FEA61909FE782F54" box="[368,502,583,609]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). BD. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 14A</figureCitation>
) with
<emphasis id="40C26A3E4528FFA1FD951909FDD92F55" box="[579,599,583,608]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
seven spinous whirls, relatively long and with an elongated central tube joining posterior funnel-shaped end plate to anterior end plate.
</paragraph>
<paragraph id="7209B62C4528FFA1FF4119FCFE182FF9" blockId="29.[151,1436,690,1185]" box="[151,406,690,716]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF4119FCFE182FF9" bold="true" box="[151,406,690,716]" pageId="29" pageNumber="30">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411999FB6F2FC4" blockId="29.[151,1436,690,1185]" box="[151,1249,727,753]" pageId="29" pageNumber="30">Cp and valves most likely as in the male, without obvious sexual dimorphism (see remark above).</paragraph>
<paragraph id="7209B62C4528FFA1FF1119B5FB072E20" blockId="29.[151,1436,690,1185]" box="[199,1161,762,789]" pageId="29" pageNumber="30">
Limbs (
<figureCitation id="EA8DAAA94528FFA1FECB19B5FE302E20" box="[285,446,762,789]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111851FC232E90" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
A2 (
<figureCitation id="EA8DAAA94528FFA1FF2F1851FEF22E0C" box="[249,380,798,825]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15C</figureCitation>
) with second endopodal segment undivided, with two t-setae, t1 (long, narrow and plumose) and t2 (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94528FFA1FD47180DFC8F2E68" box="[657,769,835,861]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="29" pageNumber="30">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4528FFA1FF1118E1FCFA2924" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
T1 (
<figureCitation id="EA8DAAA94528FFA1FF2E18E1FEF52EFC" box="[248,379,942,969]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." pageId="29" pageNumber="30">Figure 15E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4528FFA1FDA918E1FD1C2EFD" box="[639,658,943,968]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
10 distal setae of unequal length, setae b and d present, with long additional seta (d) present close to seta d; respiratory plate hidden on the dissected specimens. Endopodite a broad and short palp, with three subequal but short apical setae (h1-h3).
</paragraph>
<paragraph id="7209B62C4528FFA1FF111F55FD9E2900" blockId="29.[151,1436,690,1185]" box="[199,528,1051,1077]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F55FE0E2900" bold="true" box="[199,384,1051,1077]" pageId="29" pageNumber="30">Measurements:</emphasis>
See
<tableCitation id="3F3483974528FFA1FE601F55FD842900" box="[438,522,1051,1077]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="29" pageNumber="30">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F70FDEA296C" box="[199,612,1086,1113]" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7209B62C4528FFA1FF111F70FDEA296C" blockId="29.[151,1436,690,1185]" box="[199,612,1086,1113]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F70FE5B296D" bold="true" box="[199,469,1086,1112]" pageId="29" pageNumber="30">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974528FFA1FDDC1F71FDEE296C" box="[522,608,1087,1113]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="29" pageNumber="30">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F2DFB072994" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="7209B62C4528FFA1FF111F2DFE612949" blockId="29.[151,1436,690,1185]" box="[199,495,1122,1149]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F2DFE612949" bold="true" box="[199,495,1122,1149]" pageId="29" pageNumber="30">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF111FC9FB072994" blockId="29.[151,1436,690,1185]" box="[199,1161,1159,1185]" pageId="29" pageNumber="30">
This subterranean species is presently known from its
<typeStatus id="AD0D088E4528FFA1FCF11FC9FCD92994" box="[807,855,1159,1185]" pageId="29" pageNumber="30">type</typeStatus>
locality only (see
<tableCitation id="3F3483974528FFA1FBF01FC9FBF62994" box="[1062,1144,1159,1185]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="29" pageNumber="30">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E8DC3C9BB380F01983B76F4F04BC17E8">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="355D8BC12C8E209632FBED134354AAFA">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="2E7056142F919CBD10743945687018BB">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="EE7B131E97EEAEDE52029649FC7F4F2F">Schön, Isa</mods:namePart>
<mods:nameIdentifier id="5BA76AEBFE0374770C96BA8C8329E899" type="ORCID">0000-0001-9269-6487</mods:nameIdentifier>
<mods:affiliation id="B763117B4FDE59E02CDFB35F97666BD1">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="FC4E8969106A933DB394F4370EB5FC4F" type="email">ischoen@naturalsciences.be</mods:nameIdentifier>
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<mods:name id="C21950B283F65E0467C75787C7E26A49" type="personal">
<mods:role id="1286BE8B341CB342BEEFD46C09DCD12E">
<mods:roleTerm id="85FAAE6792D22F72081BFE7CD408BE2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1088AC7095505FB6AF77CDA391DE013E">Martens, Koen</mods:namePart>
<mods:nameIdentifier id="628EE4E3C3C4137A4D414D154BDB6E50" type="ORCID">0000-0001-8680-973X</mods:nameIdentifier>
<mods:affiliation id="42520D1326FE68C099CCD9B9E0251F81">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="CE30DA05B18908B84E4389AA056A3C9A" type="email">kmartens@naturalsciences.be</mods:nameIdentifier>
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<mods:typeOfResource id="29575DE43DEC715370322F9BDAD6E725">text</mods:typeOfResource>
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<mods:title id="3F98D86425B2826DB18FB1BE7891D65D">Zootaxa</mods:title>
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<mods:part id="D39E3F6FD7C4A173D66BC717ECD2B160">
<mods:date id="1F43DBFB52F5038AD9D8E471F50D10E7">2024</mods:date>
<mods:detail id="2CD9DE0740278474F569BD78E052CC2B" type="pubDate">
<mods:number id="70C565D9BE43995B36BA7E13D5170411">2024-09-04</mods:number>
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<mods:detail id="EBE42A52B47C05069772B9D21D37F548" type="volume">
<mods:number id="26A4DD54E032CA705F69B929CBAB9C09">5503</mods:number>
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<mods:number id="DF30B63167309D2C1100195E6DF7DD13">1</mods:number>
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<mods:identifier id="34784655C6BB44F49604563D6D47BEAD" type="DOI">10.11646/zootaxa.5503.1.1</mods:identifier>
<mods:identifier id="9A6142849FF1B42F4167A099158DF3AD" type="ISSN">1175-5334</mods:identifier>
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<treatment id="FA1F073A4528FF94FF411FBCFEA02C98" LSID="urn:lsid:plazi:treatment:FA1F073A4528FF94FF411FBCFEA02C98" httpUri="http://treatment.plazi.org/id/FA1F073A4528FF94FF411FBCFEA02C98" lastPageId="40" lastPageNumber="41" pageId="29" pageNumber="30">
<subSubSection id="3AACE5A74528FFA1FF411FBCFC5D2804" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="7209B62C4528FFA1FF411FBCFC5D2804" blockId="29.[151,979,1266,1365]" pageId="29" pageNumber="30">
<heading id="294101404528FFA1FF411FBCFCFB2838" bold="true" box="[151,885,1266,1293]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
<emphasis id="40C26A3E4528FFA1FF411FBCFCFB2838" bold="true" box="[151,885,1266,1293]" pageId="29" pageNumber="30">
<taxonomicName id="B5B6CDAF4528FFA1FF411FBCFE172838" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,409,1266,1293]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4528FFA1FF411FBCFE172838" bold="true" box="[151,409,1266,1293]" italics="true" pageId="29" pageNumber="30">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454528FFA1FE771FBDFDFF2838" box="[417,625,1267,1293]" pageId="29" pageNumber="30" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF411E75FE2C2860" box="[151,418,1339,1365]" pageId="29" pageNumber="30" type="description">
<paragraph id="7209B62C4528FFA1FF411E75FE2C2860" blockId="29.[151,979,1266,1365]" box="[151,418,1339,1365]" pageId="29" pageNumber="30">
(
<figureCitation id="EA8DAAA94528FFA1FF491E75FECA2860" box="[159,324,1339,1365]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStart-5="FIGURE 21" captionStart-6="FIGURE 22" captionStart-7="FIGURE 23" captionStartId-0="30.[151,250,1942,1966]" captionStartId-1="31.[151,250,1630,1654]" captionStartId-2="33.[151,250,1798,1822]" captionStartId-3="34.[151,250,1943,1967]" captionStartId-4="35.[151,250,1943,1967]" captionStartId-5="37.[151,250,1981,2005]" captionStartId-6="38.[151,250,1943,1967]" captionStartId-7="39.[151,250,1943,1967]" captionTargetBox-0="[169,1418,181,1919]" captionTargetBox-1="[151,1436,181,1606]" captionTargetBox-2="[151,1436,345,1775]" captionTargetBox-3="[156,1433,185,1915]" captionTargetBox-4="[185,1402,185,1915]" captionTargetBox-5="[177,1420,212,1949]" captionTargetBox-6="[195,1394,191,1912]" captionTargetBox-7="[173,1413,185,1915]" captionTargetId-0="figure-31@30.[169,1418,181,1919]" captionTargetId-1="figure-17@31.[151,1436,181,1606]" captionTargetId-2="figure-109@33.[151,1436,345,1775]" captionTargetId-3="figure-31@34.[152,1436,181,1919]" captionTargetId-4="figure-17@35.[181,1406,181,1919]" captionTargetId-5="figure-17@37.[166,1422,181,1957]" captionTargetId-6="figure-31@38.[185,1402,181,1919]" captionTargetId-7="figure-17@39.[170,1417,181,1919]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="37" captionTargetPageId-6="38" captionTargetPageId-7="39" captionText-0="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." captionText-1="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-2="FIGURE 18. Benincandona sakitii gen. et spec. nov., valves (♀). A, C, D. LVi (A, C, D. INV.159226). B, E, F, G. RVi (INV.159226). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." captionText-3="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." captionText-4="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-5="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." captionText-6="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." captionText-7="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="29" pageNumber="30">Figures 1623</figureCitation>
,
<figureCitation id="EA8DAAA94528FFA1FE861E75FE142860" box="[336,410,1339,1365]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="29" pageNumber="30">41GJ</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF411ECDFB6728D0" pageId="29" pageNumber="30" type="etymology">
<paragraph id="7209B62C4528FFA1FF411ECDFE9628A8" blockId="29.[151,1436,1411,1509]" box="[151,280,1411,1437]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF411ECDFE9628A8" bold="true" box="[151,280,1411,1437]" pageId="29" pageNumber="30">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411EE9FB6728D0" blockId="29.[151,1436,1411,1509]" pageId="29" pageNumber="30">
The species is named after Prof. Nestor G. Sakiti who initiated, in collaboration with Prof. Claude Boutin, subterranean biology research in
<collectingCountry id="0AA1F6BC4528FFA1FDDD1E85FDDF28D0" box="[523,593,1483,1509]" name="Benin" pageId="29" pageNumber="30">Benin</collectingCountry>
, in gratitude for his continued support to the career of SJH.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF411D5CFDE92A44" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7209B62C4528FFA1FF411D5CFE842B19" blockId="29.[151,1437,1554,1905]" box="[151,266,1554,1580]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF411D5CFE842B19" bold="true" box="[151,266,1554,1580]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411D79FDE92A44" blockId="29.[151,1437,1554,1905]" pageId="29" pageNumber="30">
CpRl elongated (L/H = 2.0), sub-triangular, with greatest height slightly behind the middle; LV dorsally extending beyond RV, anterior margin bluntly pointed and with anteroventral part almost straight, posterior margin asymmetrically rounded towards the almost straight ventral side, valve surface set with polygonal ridges in midventral and central part. CpD and CpV narrow, with greatest width in the middle and evenly rounded lateral sides, anterior margin sharply and posterior margin bluntly pointed. A2 with seta X (medium length) reaching beyond middle of aesthetasc Y in males, beyond tip of aesthetasc Y in females (long). This aesthetasc Y distally asymmetrically bifurcated. Second palp segment of Mx1 with L
<emphasis id="40C26A3E4528FFA1FD7B1C41FD372A1D" box="[685,697,1807,1832]" italics="true" pageId="29" pageNumber="30">c</emphasis>
. 1.5x basal W. T3 with seta d2 about 2/3 of L of seta d1. Hp with ls and ms with overlapping positions, narrower lobe ls on Hp with skewed rectangular distal margin, extending beyond lobe ms with more than half of its length.
</paragraph>
</subSubSection>
<caption id="26C9E6A4452BFFA2FF411CD8FAB22AE6" pageId="30" pageNumber="31" startId="30.[151,250,1942,1966]" targetBox="[169,1418,181,1919]" targetPageId="30" targetType="figure">
<paragraph id="7209B62C452BFFA2FF411CD8FAB22AE6" blockId="30.[151,1436,1942,2003]" pageId="30" pageNumber="31">
<emphasis id="40C26A3E452BFFA2FF411CD8FEAE2A9B" bold="true" box="[151,288,1942,1966]" pageId="30" pageNumber="31">FIGURE 16.</emphasis>
<taxonomicName id="B5B6CDAF452BFFA2FEF01CD9FE732A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,509,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E452BFFA2FEF01CD9FE732A9A" box="[294,509,1943,1967]" italics="true" pageId="30" pageNumber="31">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452BFFA2FDD21CD9FD382A9B" box="[516,694,1943,1967]" pageId="30" pageNumber="31" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AC, G, IK ♂; DF, H ♀).
<emphasis id="40C26A3E452BFFA2FBE51CD8FBEB2A9B" bold="true" box="[1075,1125,1942,1966]" pageId="30" pageNumber="31">AH</emphasis>
. CpRl, (AC, G: INV.159235, DF, H: INV.159237).
<emphasis id="40C26A3E452BFFA2FE561CF5FE242AE6" bold="true" box="[384,426,1979,2003]" pageId="30" pageNumber="31">IK</emphasis>
. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452AFFA3FF411D10FB172BAE" pageId="31" pageNumber="32" startId="31.[151,250,1630,1654]" targetBox="[151,1436,181,1606]" targetPageId="31" targetType="figure">
<paragraph id="7209B62C452AFFA3FF411D10FB172BAE" blockId="31.[151,1436,1630,1691]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411D10FEAE2B43" bold="true" box="[151,288,1630,1654]" pageId="31" pageNumber="32">FIGURE 17.</emphasis>
<taxonomicName id="B5B6CDAF452AFFA3FEF11D10FE702B43" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,510,1630,1654]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E452AFFA3FEF11D10FE702B43" box="[295,510,1630,1654]" italics="true" pageId="31" pageNumber="32">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452AFFA3FDD31D11FD392B43" box="[517,695,1631,1655]" pageId="31" pageNumber="32" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♂).
<emphasis id="40C26A3E452AFFA3FCEC1D10FC222B43" bold="true" box="[826,940,1630,1655]" pageId="31" pageNumber="32">A, C, D, G</emphasis>
. LVi (A, C, D. INV.159225; G. INV.159227).
<emphasis id="40C26A3E452AFFA3FA531D11FF4A2BAE" bold="true" pageId="31" pageNumber="32">B, E, F</emphasis>
. RVi (INV.159225).
<emphasis id="40C26A3E452AFFA3FE4E1DCCFE3F2BAE" bold="true" box="[408,433,1666,1691]" pageId="31" pageNumber="32">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A7452AFF9CFF411DA3FC5C2F08" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="7209B62C452AFFA3FF411DA3FEB02A33" blockId="31.[151,318,1772,1799]" box="[151,318,1772,1799]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411DA3FEB02A33" bold="true" box="[151,318,1772,1799]" pageId="31" pageNumber="32">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C7BFE8A2A7A" blockId="31.[151,1282,1845,1907]" box="[151,260,1845,1871]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411C7BFE8A2A7A" bold="true" box="[151,260,1845,1871]" pageId="31" pageNumber="32">
<typeStatus id="AD0D088E452AFFA3FF411C7BFE8A2A7A" box="[151,260,1845,1871]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C17FA8C2A46" blockId="31.[151,1282,1845,1907]" box="[151,1282,1880,1907]" pageId="31" pageNumber="32">
<materialsCitation id="C2DEBC71452AFFA3FF411C17FA8C2A46" box="[151,1282,1880,1907]" collectionCode="INV" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5452AFFA3FF411C17FF4D2A46" box="[151,195,1881,1907]" count="1" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9452AFFA3FBB01C17FB182A46" box="[1126,1174,1881,1907]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="31" pageNumber="32" type="Herbarium">INV</collectionCode>
.159225).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411CEEFF722A8F" blockId="31.[151,736,1952,2015]" box="[151,252,1952,1978]" pageId="31" pageNumber="32">
<emphasis id="40C26A3E452AFFA3FF411CEEFF722A8F" bold="true" box="[151,252,1952,1978]" pageId="31" pageNumber="32">
<typeStatus id="AD0D088E452AFFA3FF411CEEFF722A8F" box="[151,252,1952,1978]" pageId="31" pageNumber="32" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C452AFFA3FF411C8BFD6E2AEA" blockId="31.[151,736,1952,2015]" box="[151,736,1988,2015]" pageId="31" pageNumber="32">
<materialsCitation id="C2DEBC71452AFFA3FF411C8BFD6E2AEA" box="[151,736,1988,2015]" collectionCode="INV" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5452AFFA3FF411C8BFF4F2AEA" box="[151,193,1989,2015]" count="1" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
, dissected and stored as the male (
<collectionCode id="14A72EE9452AFFA3FD921C8AFDF72AEA" box="[580,633,1988,2015]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="31" pageNumber="32" type="Herbarium">INV</collectionCode>
.159226).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411BD9FE812D84" blockId="32.[151,1436,151,285]" box="[151,271,151,177]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411BD9FE812D84" bold="true" box="[151,271,151,177]" pageId="32" pageNumber="33">
<typeStatus id="AD0D088E4515FF9CFF411BD9FE812D84" box="[151,271,151,177]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411BF5FADB2C28" blockId="32.[151,1436,151,285]" pageId="32" pageNumber="33">
<materialsCitation id="C2DEBC714515FF9CFF411BF5FBD12DE0" box="[151,1119,186,213]" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFF411BF5FF542DE0" box="[151,218,187,213]" count="3" pageId="32" pageNumber="33" type="male">3 ♂♂</specimenCount>
, dissected and stored as the
<typeStatus id="AD0D088E4515FF9CFDF51BF5FD0B2DE0" box="[547,645,187,213]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94515FF9CFD431BF4FD472DE0" box="[661,713,186,213]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159227, INV.159228, INV.159229)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFBBB1BF5FD7D2DCC" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFBBB1BF5FB202DE0" box="[1133,1198,187,213]" count="3" pageId="32" pageNumber="33" type="female">3 ♀♀</specimenCount>
dissected and stored as the male (
<collectionCode id="14A72EE94515FF9CFEFF1B90FED32DCC" box="[297,349,222,249]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159230, INV.159231, INV.159232)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFCD61B91FDCD2C28" collectionCode="JSH" pageId="32" pageNumber="33" specimenCode="JSH0344" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFCD61B91FCCF2DCC" box="[768,833,223,249]" count="3" pageId="32" pageNumber="33" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<specimenCode id="22101E574515FF9CFF491A4DFE842C28" box="[159,266,259,285]" collectionCode="JSH" pageId="32" pageNumber="33">JSH0344</specimenCode>
, INV.159235, INV.159236)
</materialsCitation>
;
<materialsCitation id="C2DEBC714515FF9CFD981A4DFADB2C28" box="[590,1365,258,285]" collectionCode="INV" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA54515FF9CFD981A4DFDF62C28" box="[590,632,259,285]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE94515FF9CFB171A4CFB7B2C28" box="[1217,1269,258,285]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="32" pageNumber="33" type="Herbarium">INV</collectionCode>
.159227)
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411A05FEBE2C51" blockId="32.[151,1436,330,465]" box="[151,304,330,357]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411A05FEBE2C51" bold="true" box="[151,304,330,357]" pageId="32" pageNumber="33">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF411A21FCD32CE4" blockId="32.[151,1436,330,465]" pageId="32" pageNumber="33">
<materialsCitation id="C2DEBC714515FF9CFF411A21FE632C98" country="Benin" location="Tchaourou District" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Borgou">
<collectingCountry id="0AA1F6BC4515FF9CFF411A21FF542CBC" box="[151,218,367,393]" name="Benin" pageId="32" pageNumber="33">Benin</collectingCountry>
,
<collectingRegion id="B07278CE4515FF9CFF311A21FE492CBC" box="[231,455,367,393]" country="Benin" name="Borgou" pageId="32" pageNumber="33">Borgou Department</collectingRegion>
,
<location id="7769E0F74515FF9CFE061A21FD2A2CBC" LSID="urn:lsid:plazi:treatment:FA1F073A4528FF94FF411FBCFEA02C98:7769E0F74515FF9CFE061A21FD2A2CBC" box="[464,676,367,393]" country="Benin" name="Tchaourou District" pageId="32" pageNumber="33" stateProvince="Borgou">Tchaourou District</location>
, town of Badékparou located in the Okpara catchment area (tributary of the
<collectingRegion id="B07278CE4515FF9CFF361ADDFEBF2C98" box="[224,305,403,429]" country="Benin" name="Oueme" pageId="32" pageNumber="33">Ouémé</collectingRegion>
catchment area)
</materialsCitation>
, coordinates:
<geoCoordinate id="1782D0EB4515FF9CFD5F1ADDFC832C98" box="[649,781,403,429]" degrees="9" direction="north" minutes="13.314" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="9.2219">9°13.314N</geoCoordinate>
<geoCoordinate id="1782D0EB4515FF9CFCC21ADDFC162C98" box="[788,920,403,429]" degrees="2" direction="east" minutes="33.790" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="2.5631666">2°33.790E</geoCoordinate>
, Leg.: S.J. Hotèkpo and M. Lagnika collected on
<date id="060890EC4515FF9CFF6C1AF9FEFB2CE4" box="[186,373,439,465]" pageId="32" pageNumber="33" value="2022-10-11">11 October 2022</date>
(Locality code BEN066 in
<tableCitation id="3F3483974515FF9CFD7D1AF9FC872CE4" box="[683,777,439,465]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="32" pageNumber="33">Tables 1</tableCitation>
and
<tableCitation id="3F3483974515FF9CFCE91AF9FCDE2CE4" box="[831,848,439,465]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="32" pageNumber="33">2</tableCitation>
).
</paragraph>
<paragraph id="7209B62C4515FF9CFF411AB0FE6E2F2D" blockId="32.[151,978,510,573]" box="[151,480,510,536]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411AB0FE6E2F2D" bold="true" box="[151,480,510,536]" pageId="32" pageNumber="33">Other material investigated</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF41196DFC5C2F08" blockId="32.[151,978,510,573]" box="[151,978,547,573]" pageId="32" pageNumber="33">
See
<tableCitation id="3F3483974515FF9CFF10196DFE972F08" box="[198,281,547,573]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="32" pageNumber="33">Table 2</tableCitation>
for a list of localities where the present species was collected.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74515FF94FF411924FD862C28" lastPageId="40" lastPageNumber="41" pageId="32" pageNumber="33" type="description">
<paragraph id="7209B62C4515FF9CFF411924FE0F2FB0" blockId="32.[151,1437,618,2013]" box="[151,385,618,645]" pageId="32" pageNumber="33">
<emphasis id="40C26A3E4515FF9CFF411924FE0F2FB0" bold="true" box="[151,385,618,645]" pageId="32" pageNumber="33">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4515FF9CFF4119C1FDC42EB4" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
CpRl (
<figureCitation id="EA8DAAA94515FF9CFF3619C1FE2C2F9C" box="[224,418,655,681]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="32" pageNumber="33">Figure 16AC, G</figureCitation>
) elongated (L/H ratio = 2.0) and sub-triangular, dorsal margin bluntly pointed, with greatest height behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part of anterior margin almost straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; LV overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valve surface set with polygonal ridges in midventral and central part. CpD (
<figureCitation id="EA8DAAA94515FF9CFF49180DFEB62E68" box="[159,312,835,861]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="32" pageNumber="33">Figure 16IK</figureCitation>
) narrow and sharply pointed at anterior margin, more bluntly pointed at posterior side, greatest width in the middle, lateral margins rounded.
</paragraph>
<paragraph id="7209B62C4515FF9CFF1118C5FF7E2900" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
LVi (
<figureCitation id="EA8DAAA94515FF9CFED418C5FE762E90" box="[258,504,907,933]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17A, C, D, G</figureCitation>
) elongated and sub-triangular, dorsal margin bluntly pointed in the centre, with greatest height behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part almost straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; one ventral il without antero-ventral tooth. CMS (
<figureCitation id="EA8DAAA94515FF9CFBB318B9FB642924" box="[1125,1258,1015,1041]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17G</figureCitation>
) as in the
<typeStatus id="AD0D088E4515FF9CFABA18B9FA122924" box="[1388,1436,1015,1041]" pageId="32" pageNumber="33">type</typeStatus>
species.
</paragraph>
<paragraph id="7209B62C4515FF9CFF111F71FE0629F0" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
RVi (
<figureCitation id="EA8DAAA94515FF9CFED41F71FE4B296C" box="[258,453,1087,1113]" captionStart="FIGURE 17" captionStartId="31.[151,250,1630,1654]" captionTargetBox="[151,1436,181,1606]" captionTargetId="figure-17@31.[151,1436,181,1606]" captionTargetPageId="31" captionText="FIGURE 17. Benincandona sakitii gen. et spec. nov., valves (♂). A, C, D, G. LVi (A, C, D. INV.159225; G. INV.159227). B, E, F. RVi (INV.159225). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="32" pageNumber="33">Figure 17B, E, F</figureCitation>
) elongated, dorsal margin with greatest height well behind the middle, anterior part of dorsal margin sloping almost straight to anterior margin, passing into it with a blunt angle, anteroventral part straight, posterior part of dorsal margin sloping towards ventral margin, the latter almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C4515FF9CFF111F81FB4C2B64" blockId="32.[151,1437,618,2013]" pageId="32" pageNumber="33">
A1 (
<figureCitation id="EA8DAAA94515FF9CFF211F81FEF929DC" box="[247,375,1231,1257]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="32" pageNumber="33">Figure 19B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment
<emphasis id="40C26A3E4515FF9CFADE1F81FA9529DD" box="[1288,1307,1231,1256]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
three times as long as basal width, supporting two long ventro-subapical setae with the more apical slightly shorter than the other one; and two medium length setae on the dorsal side (one illustrated here), situated some distance from each other. First distal segment about twice the basal width, with one short dorso-apical seta. Second distal segment about the same length/width ratio as the previous segment with one very small dorso-apical seta. Third distal segment about three times as long as basal width, set with one very small apical seta in each side. Fourth distal segment about twice the basal width, with one short ventro-apical seta and one medium length dorso-apical seta, reaching beyond tip of sixth segment with more than the half the total length. Fifth distal segment about four times as long as wide, with two long dorso-apical setae and one short ventro-apical seta clearly exceeding the next segment. Sixth distal segment narrow,
<emphasis id="40C26A3E4515FF9CFE801D5DFEE72B19" box="[342,361,1555,1580]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long setae about twice the length of Ya and one short seta, about one third the length of Ya.
</paragraph>
<paragraph id="7209B62C4515FF9DFF111D15FE042C74" blockId="32.[151,1437,618,2013]" lastBlockId="33.[151,1437,151,321]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
A2 (
<figureCitation id="EA8DAAA94515FF9CFF2C1D15FEF12B40" box="[250,383,1627,1653]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="32" pageNumber="33">Figure 19A</figureCitation>
) with a basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, two subapical setae of unequal length, proximal one
<emphasis id="40C26A3E4515FF9CFC621D31FC492BAD" box="[948,967,1663,1688]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice the length of the more apical one. Second protopodite large and stout,
<emphasis id="40C26A3E4515FF9CFDF91DEDFDCC2B89" box="[559,578,1699,1724]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments with the penultimate segment divided. First endopodal segment large, more than five times as long as basal width, carrying one long aesthetasc Y (distally asymmetrically bifurcated) on the mid-ventral side (slightly under tip of segment) and two ventro-apical setae, one short and one large, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E4515FF9CFB5D1C7DFB102A79" box="[1163,1182,1843,1868]" italics="true" pageId="32" pageNumber="33">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender aesthetasc y1; setae t2 and t3 hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and almost as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA94515FF9CFD451C8DFC862AE8" box="[659,776,1987,2013]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="32" pageNumber="33">Figure 40</figureCitation>
): ventro-apically with one slender aesthetasc y2 and claw G3 transformed into short seta of almost equal length; centrally with one short seta z3; dorso-apically with z1 claw-like, length about twice as long as penultimate segment. Claw G2 and G1 about 2.5 times as long as z1, with z2 seta transformed into long claw, reaching tip of G2. Fourth, terminal, endopodal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and seta g almost as long as aesthetasc y3.
</paragraph>
<caption id="26C9E6A44514FF9DFF411C48FBBD2A76" pageId="33" pageNumber="34" startId="33.[151,250,1798,1822]" targetBox="[151,1436,345,1775]" targetPageId="33" targetType="figure">
<paragraph id="7209B62C4514FF9DFF411C48FBBD2A76" blockId="33.[151,1437,1798,1859]" pageId="33" pageNumber="34">
<emphasis id="40C26A3E4514FF9DFF411C48FEAD2A2B" bold="true" box="[151,291,1798,1822]" pageId="33" pageNumber="34">FIGURE 18.</emphasis>
<taxonomicName id="B5B6CDAF4514FF9DFEFB1C49FD892A2A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,519,1799,1823]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4514FF9DFEFB1C49FD892A2A" box="[301,519,1799,1823]" italics="true" pageId="33" pageNumber="34">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454514FF9DFDC71C49FD432A2B" box="[529,717,1799,1823]" pageId="33" pageNumber="34" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (♀).
<emphasis id="40C26A3E4514FF9DFC8F1C48FC3E2A2A" bold="true" box="[857,944,1798,1823]" pageId="33" pageNumber="34">A, C, D</emphasis>
. LVi (A, C, D. INV.159226).
<emphasis id="40C26A3E4514FF9DFB241C49FAE82A2B" bold="true" box="[1266,1382,1798,1823]" pageId="33" pageNumber="34">B, E, F, G</emphasis>
. RVi (INV.159226).
<emphasis id="40C26A3E4514FF9DFEE71C64FEC52A76" bold="true" box="[305,331,1834,1859]" pageId="33" pageNumber="34">G.</emphasis>
CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44517FF9EFF411CD9FE7C2AE1" pageId="34" pageNumber="35" startId="34.[151,250,1943,1967]" targetBox="[156,1433,185,1915]" targetPageId="34" targetType="figure">
<paragraph id="7209B62C4517FF9EFF411CD9FE7C2AE1" blockId="34.[151,1437,1943,2004]" pageId="34" pageNumber="35">
<emphasis id="40C26A3E4517FF9EFF411CD9FEAC2A9A" bold="true" box="[151,290,1943,1967]" pageId="34" pageNumber="35">FIGURE 19.</emphasis>
<taxonomicName id="B5B6CDAF4517FF9EFEFC1CD6FD8C2A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[298,514,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4517FF9EFEFC1CD6FD8C2A85" box="[298,514,1944,1968]" italics="true" pageId="34" pageNumber="35">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454517FF9EFDDC1CD6FD4F2A9A" box="[522,705,1944,1968]" pageId="34" pageNumber="35" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4517FF9EFCE91CD9FCDF2A9A" bold="true" box="[831,849,1943,1967]" pageId="34" pageNumber="35">A</emphasis>
. A2 (INV.159225).
<emphasis id="40C26A3E4517FF9EFBF71CD6FBBC2A85" bold="true" box="[1057,1074,1944,1968]" pageId="34" pageNumber="35">B</emphasis>
. A1 (INV.159225).
<emphasis id="40C26A3E4517FF9EFAD41CD9FABB2A85" bold="true" box="[1282,1333,1943,1968]" pageId="34" pageNumber="35">C, D</emphasis>
. MdCoxa (INV.159228). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44516FF9FFF411CD9FC482AE1" pageId="35" pageNumber="36" startId="35.[151,250,1943,1967]" targetBox="[185,1402,185,1915]" targetPageId="35" targetType="figure">
<paragraph id="7209B62C4516FF9FFF411CD9FC482AE1" blockId="35.[151,1436,1943,2004]" pageId="35" pageNumber="36">
<emphasis id="40C26A3E4516FF9FFF411CD9FEAB2A9A" bold="true" box="[151,293,1943,1967]" pageId="35" pageNumber="36">FIGURE 20.</emphasis>
<taxonomicName id="B5B6CDAF4516FF9FFEE61CD6FD852A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[304,523,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4516FF9FFEE61CD6FD852A85" box="[304,523,1944,1968]" italics="true" pageId="35" pageNumber="36">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454516FF9FFDC01CD6FD582A9A" box="[534,726,1944,1968]" pageId="35" pageNumber="36" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4516FF9FFC8B1CD9FCE12A9A" bold="true" box="[861,879,1943,1967]" pageId="35" pageNumber="36">A</emphasis>
. Mx1-palp (INV.159225).
<emphasis id="40C26A3E4516FF9FFB581CD6FB112A85" bold="true" box="[1166,1183,1944,1968]" pageId="35" pageNumber="36">B</emphasis>
. Mx1, respiratory plate (INV.159227).
<emphasis id="40C26A3E4516FF9FFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="35" pageNumber="36">C</emphasis>
. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4511FF98FF111BD9FC472DCC" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
MdCoxa (
<figureCitation id="EA8DAAA94511FF98FEEE1BD9FE532D84" box="[312,477,151,177]" captionStart="FIGURE 19" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[156,1433,185,1915]" captionTargetId="figure-31@34.[152,1436,181,1919]" captionTargetPageId="34" captionText="FIGURE 19. Benincandona sakitii gen. et spec. nov., limbs (♂). A. A2 (INV.159225). B. A1 (INV.159225). C, D. MdCoxa (INV.159228). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 19CD</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
<emphasis id="40C26A3E4511FF98FAEB1BF5FA1A2DE0" box="[1341,1428,187,213]" italics="true" pageId="36" pageNumber="37">Remark</emphasis>
: asymmetry illustrated possibly resulting from different positions in slide.
</paragraph>
<paragraph id="7209B62C4511FF98FF111A4DFAAB2F2C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
MdPalp (
<figureCitation id="EA8DAAA94511FF98FEE61A4DFE3D2C28" box="[304,435,259,285]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20C</figureCitation>
) consisting of four segments. Basal segment ventro-apically with two hirsute setae: one long (S1), reaching tip of terminal segment and one short and stout (S2), length about one fifth of the length of S1; in between with a short flagellum-like alpha seta (thin and about 1.5x length of S2); more proximally, one smooth, medium-length seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four setae (long and unequal size). Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta, and ventro-apically with three setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, slightly shorter than the fused claw.
</paragraph>
<paragraph id="7209B62C4511FF98FF11196DFF6B2F9C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Mx1 (
<figureCitation id="EA8DAAA94511FF98FEDB196DFE382F08" box="[269,438,547,573]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20A, B</figureCitation>
) palp with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying also four distal setae, some slightly claw-like. Respiratory plate (
<figureCitation id="EA8DAAA94511FF98FB401909FA962F54" box="[1174,1304,583,609]" captionStart="FIGURE 20" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[185,1402,185,1915]" captionTargetId="figure-17@35.[181,1406,181,1919]" captionTargetPageId="35" captionText="FIGURE 20. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Mx1-palp (INV.159225). B. Mx1, respiratory plate (INV.159227). C. MdPalp (INV.159227). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="36" pageNumber="37">Figure 20B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E4511FF98FF2D1925FE802FB1" box="[251,270,619,644]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
15 plumose respiratory rays, two short distal setae and a group of four shorter proximal setae of unequal length.
</paragraph>
<paragraph id="7209B62C4511FF98FF1119FDFD712E68" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T1 (
<figureCitation id="EA8DAAA94511FF98FF2119FDFE132FF8" box="[247,413,691,717]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23A, B</figureCitation>
) with basal part (protopodite) carrying respiratory plates (exopodites not shown), distally set with 3+7 setae of different morphology and length including two plumose; more proximally set with d and d setae, length of the setae d being 1.5x the length of d. Prehensile palps clearly asymmetrical with Rpp (
<figureCitation id="EA8DAAA94511FF98FAC519B5FA1A2E20" box="[1299,1428,763,789]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23B</figureCitation>
) slightly more swollen than Lpp (
<figureCitation id="EA8DAAA94511FF98FE281851FD0D2E0C" box="[510,643,799,825]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23A</figureCitation>
); one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111829FAFB2900" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T2 (
<figureCitation id="EA8DAAA94511FF98FF251829FEFC2EB4" box="[243,370,871,897]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21B</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), elongated, without basal setae (d1 and d2). First endopodal segment without any setae, length
<emphasis id="40C26A3E4511FF98FCE318C5FCC62E91" box="[821,840,907,932]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
four times as long as basal width. Second endopodal segment with length
<emphasis id="40C26A3E4511FF98FE5418E1FE1B2EFD" box="[386,405,943,968]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
4x basal width, set with one short ventro-apical seta f. Third endopodal segment with about the same length/width ratio as the previous segment and with one ventro-subapical seta g, about 1/3 the length of following segment. Terminal segment about 1.5 times as long as basal width, apically two short setae, h1 ventral and h3 (length being the half of the length of h1) dorsal and a long slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111F71FE1B2838" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
T3 (
<figureCitation id="EA8DAAA94511FF98FF231F71FEF6296C" box="[245,376,1087,1113]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21A</figureCitation>
) with first segment carrying only one medium length seta d1. Second (knee-) segment set with d2-seta. Third segment about 4x as long as wide, not carrying any seta. Fourth segment narrower,
<emphasis id="40C26A3E4511FF98FB0F1F2DFB622949" box="[1241,1260,1123,1148]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
7x basal width, slightly longer than the previous segment, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, length about twice basal width and carrying three setae: one long and reflexed (h3), tip clearly reaching base of the third segment, and two ventrally directed and curved setae h1 &amp; h2, the former
<emphasis id="40C26A3E4511FF98FA801F81FAE729DD" box="[1366,1385,1231,1256]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4511FF98FF111E59FEB9284C" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
CR (
<figureCitation id="EA8DAAA94511FF98FF2D1E59FEF32804" box="[251,381,1303,1329]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21C</figureCitation>
) with stout ramus; claw Ga sigmoid, well-developed and stout, length ratio of ramus and claw Ga approximately 1.6; setae Sp (ventro-subapical), Sa (dorso-apical) and claw Gp (first ventro-subapical) reduced to small setae.
</paragraph>
<paragraph id="7209B62C4511FF98FF111ECDFE1D28F4" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
CRAtt (
<figureCitation id="EA8DAAA94511FF98FEC91ECDFE2D28A8" box="[287,419,1411,1437]" captionStart="FIGURE 21" captionStartId="37.[151,250,1981,2005]" captionTargetBox="[177,1420,212,1949]" captionTargetId="figure-17@37.[166,1422,181,1957]" captionTargetPageId="37" captionText="FIGURE 21. Benincandona sakitii et spec. nov., limbs (♂). A. T3 (INV.159225). B. T2 (INV.159225). C. CR (INV.159225). D. CRAtt (INV.15922). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 21D</figureCitation>
) long and stout, distally bifurcated and subapically with additional lateral branch; rounded fist-shaped apical end.
</paragraph>
<paragraph id="7209B62C4511FF98FF111E85FE032B64" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Hp (
<figureCitation id="EA8DAAA94511FF98FF2A1E85FE2A28D0" box="[252,420,1483,1509]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22BD</figureCitation>
) with elongated lobe ls, apically rounded and with a pronounced hump-shaped ventral protrusion, reaching well beyond asymmetrically rounded medium shield (ms), both lobes overlapping, not diverging; lobe h shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94511FF98FC571D5DFBA82B18" box="[897,1062,1555,1581]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4511FF98FF111D15FD732BAC" blockId="36.[151,1437,151,1689]" pageId="36" pageNumber="37">
Zenker organ (
<figureCitation id="EA8DAAA94511FF98FEA41D15FE772B40" box="[370,505,1627,1653]" captionStart="FIGURE 22" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[195,1394,191,1912]" captionTargetId="figure-31@38.[185,1402,181,1919]" captionTargetPageId="38" captionText="FIGURE 22. Benincandona sakitii gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159227). BD. Hp (INV.159225). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 22A</figureCitation>
) elongated, with
<emphasis id="40C26A3E4511FF98FD131D15FD562B41" box="[709,728,1627,1652]" italics="true" pageId="36" pageNumber="37">c.</emphasis>
seven spinous whirls, with an elongated central tube joining posterior funnel-shaped end plate to anterior end plate.
</paragraph>
<paragraph id="7209B62C4511FF98FF411D88FE182BD5" blockId="36.[151,1436,1734,1977]" box="[151,406,1734,1760]" pageId="36" pageNumber="37">
<emphasis id="40C26A3E4511FF98FF411D88FE182BD5" bold="true" box="[151,406,1734,1760]" pageId="36" pageNumber="37">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4511FF98FF411DA5FB4D2A30" blockId="36.[151,1436,1734,1977]" box="[151,1219,1771,1797]" pageId="36" pageNumber="37">
Cp and valves (
<figureCitation id="EA8DAAA94511FF98FE921DA5FD9C2A30" box="[324,530,1771,1797]" captionStart="FIGURE 16" captionStartId="30.[151,250,1942,1966]" captionTargetBox="[169,1418,181,1919]" captionTargetId="figure-31@30.[169,1418,181,1919]" captionTargetPageId="30" captionText="FIGURE 16. Benincandona sakitii gen. et spec. nov., valves (AC, G, IK ♂; DF, H ♀). AH. CpRl, (AC, G: INV.159235, DF, H: INV.159237). IK. INV.159236, CpD. Scale bars. 1 (B, E, I) = 500 µm; 2 (A, C, D, F, G, H, J, K) = 200 µm." pageId="36" pageNumber="37">Figures 16DF, H</figureCitation>
;
<figureCitation id="EA8DAAA94511FF98FDC81DA5FDFE2A30" box="[542,624,1771,1797]" captionStart="FIGURE 18" captionStartId="33.[151,250,1798,1822]" captionTargetBox="[151,1436,345,1775]" captionTargetId="figure-109@33.[151,1436,345,1775]" captionTargetPageId="33" captionText="FIGURE 18. Benincandona sakitii gen. et spec. nov., valves (♀). A, C, D. LVi (A, C, D. INV.159226). B, E, F, G. RVi (INV.159226). G. CMS. Scale bars. 1 (A, B) = 500 µm; 2 (CF) = 200 µm; 3 (G) = 50 µm." pageId="36" pageNumber="37">18AG</figureCitation>
) as in the male, without obvious sexual dimorphism.
</paragraph>
<paragraph id="7209B62C4511FF98FF111C41FB072A1C" blockId="36.[151,1436,1734,1977]" box="[199,1161,1807,1833]" pageId="36" pageNumber="37">
Limbs (
<figureCitation id="EA8DAAA94511FF98FECB1C41FE312A1C" box="[285,447,1807,1833]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4511FF98FF111C7DFC232A8C" blockId="36.[151,1436,1734,1977]" pageId="36" pageNumber="37">
A2 (
<figureCitation id="EA8DAAA94511FF98FF2F1C7DFEF22A78" box="[249,380,1843,1869]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="36" pageNumber="37">Figure 23C</figureCitation>
) with second endopodal segment undivided, with two t-setae, t1 (long, narrow and plumose) and t2 (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94511FF98FD451C19FC8A2A44" box="[659,772,1879,1905]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="36" pageNumber="37">Figure 40</figureCitation>
) with z1claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<caption id="26C9E6A44510FF99FF411CF3FDA02ACF" pageId="37" pageNumber="38" startId="37.[151,250,1981,2005]" targetBox="[177,1420,212,1949]" targetPageId="37" targetType="figure">
<paragraph id="7209B62C4510FF99FF411CF3FDA02ACF" blockId="37.[151,1436,1981,2042]" pageId="37" pageNumber="38">
<emphasis id="40C26A3E4510FF99FF411CF3FE912AE0" bold="true" box="[151,287,1981,2005]" pageId="37" pageNumber="38">FIGURE 21.</emphasis>
<taxonomicName id="B5B6CDAF4510FF99FEF31CF3FE752AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[293,507,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sakitii">
<emphasis id="40C26A3E4510FF99FEF31CF3FE752AE0" box="[293,507,1981,2005]" italics="true" pageId="37" pageNumber="38">Benincandona sakitii</emphasis>
</taxonomicName>
et spec. nov., limbs (♂).
<emphasis id="40C26A3E4510FF99FD231CF3FC892AE0" bold="true" box="[757,775,1981,2005]" pageId="37" pageNumber="38">A</emphasis>
. T3 (INV.159225).
<emphasis id="40C26A3E4510FF99FC181CF0FC512AE3" bold="true" box="[974,991,1982,2006]" pageId="37" pageNumber="38">B</emphasis>
. T2 (INV.159225).
<emphasis id="40C26A3E4510FF99FB731CF3FB392AE0" bold="true" box="[1189,1207,1981,2005]" pageId="37" pageNumber="38">C</emphasis>
. CR (INV.159225).
<emphasis id="40C26A3E4510FF99FA521CF0FA182AE3" bold="true" box="[1412,1430,1982,2006]" pageId="37" pageNumber="38">D</emphasis>
. CRAtt (INV.15922). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44513FF9AFF411CD9FDB32ACD" pageId="38" pageNumber="39" startId="38.[151,250,1943,1967]" targetBox="[195,1394,191,1912]" targetPageId="38" targetType="figure">
<paragraph id="7209B62C4513FF9AFF411CD9FDB32ACD" blockId="38.[151,1437,1943,2040]" pageId="38" pageNumber="39">
<emphasis id="40C26A3E4513FF9AFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="38" pageNumber="39">FIGURE 22.</emphasis>
<taxonomicName id="B5B6CDAF4513FF9AFEFB1CD9FD882A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[301,518,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4513FF9AFEFB1CD9FD882A9A" box="[301,518,1943,1967]" italics="true" pageId="38" pageNumber="39">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454513FF9AFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="38" pageNumber="39" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4513FF9AFC9B1CD9FCD12A9A" bold="true" box="[845,863,1943,1967]" pageId="38" pageNumber="39">A</emphasis>
. Zenker organ (INV.159227).
<emphasis id="40C26A3E4513FF9AFB771CD6FB5E2A85" bold="true" box="[1185,1232,1944,1968]" pageId="38" pageNumber="39">BD</emphasis>
. Hp (INV.159225).
<emphasis id="40C26A3E4513FF9AFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="38" pageNumber="39">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4513FF9AFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="38" pageNumber="39">C</emphasis>
. Internal view (photograph with interference contrast).
<emphasis id="40C26A3E4513FF9AFBCE1CF2FBA42AE1" bold="true" box="[1048,1066,1980,2004]" pageId="38" pageNumber="39">D</emphasis>
. Internal view (photograph without interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44512FF9BFF411CD9FA9F2AE1" pageId="39" pageNumber="40" startId="39.[151,250,1943,1967]" targetBox="[173,1413,185,1915]" targetPageId="39" targetType="figure">
<paragraph id="7209B62C4512FF9BFF411CD9FA9F2AE1" blockId="39.[151,1436,1943,2004]" pageId="39" pageNumber="40">
<emphasis id="40C26A3E4512FF9BFF411CD9FEAD2A9A" bold="true" box="[151,291,1943,1967]" pageId="39" pageNumber="40">FIGURE 23.</emphasis>
<taxonomicName id="B5B6CDAF4512FF9BFEFA1CD6FD882A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[300,518,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E4512FF9BFEFA1CD6FD882A85" box="[300,518,1944,1968]" italics="true" pageId="39" pageNumber="40">Benincandona sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454512FF9BFDD91CD6FD442A9A" box="[527,714,1944,1968]" pageId="39" pageNumber="40" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (A, B ♂ and CE ♀).
<emphasis id="40C26A3E4512FF9BFBDF1CD9FB952A9A" bold="true" box="[1033,1051,1943,1967]" pageId="39" pageNumber="40">A</emphasis>
. T1, Lpp (INV.159228).
<emphasis id="40C26A3E4512FF9BFAF01CD6FAB92A85" bold="true" box="[1318,1335,1944,1968]" pageId="39" pageNumber="40">B</emphasis>
. T1, Rpp (INV.159228).
<emphasis id="40C26A3E4512FF9BFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="39" pageNumber="40">C</emphasis>
. A2 (INV.159226, INV.159231).
<emphasis id="40C26A3E4512FF9BFD401CF2FD262AE1" bold="true" box="[662,680,1980,2004]" pageId="39" pageNumber="40">D</emphasis>
. CR (INV.159232).
<emphasis id="40C26A3E4512FF9BFCA11CF2FC062AE1" bold="true" box="[887,904,1980,2004]" pageId="39" pageNumber="40">E</emphasis>
. T1 (INV.159226). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C451DFF94FF111BD9FC112DCC" blockId="40.[151,1436,151,429]" pageId="40" pageNumber="41">
T1 (
<figureCitation id="EA8DAAA9451DFF94FF231BD9FEF82D84" box="[245,374,151,177]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[173,1413,185,1915]" captionTargetId="figure-17@39.[170,1417,181,1919]" captionTargetPageId="39" captionText="FIGURE 23. Benincandona sakitii gen. et spec. nov., limbs (A, B ♂ and CE ♀). A. T1, Lpp (INV.159228). B. T1, Rpp (INV.159228). C. A2 (INV.159226, INV.159231). D. CR (INV.159232). E. T1 (INV.159226). Scale bar = 50 µm." pageId="40" pageNumber="41">Figure 23E</figureCitation>
) with basal part with
<emphasis id="40C26A3E451DFF94FDBB1BD9FD0E2D85" box="[621,640,151,176]" italics="true" pageId="40" pageNumber="41">c.</emphasis>
10 distal setae of unequal length; d, d and b setae present; respiratory plate reduced to about three rays, two short, one medium size about twice as long as others. Endopodite with a broad and short palp, set with three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C451DFF94FF111A4DFD862C28" blockId="40.[151,1436,151,429]" box="[199,520,259,285]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A4DFE0E2C28" bold="true" box="[199,384,259,285]" pageId="40" pageNumber="41">Measurements:</emphasis>
See
<tableCitation id="3F348397451DFF94FE601A4DFD862C28" box="[438,520,259,285]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="40" pageNumber="41">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF111A68FDD32C74" box="[199,605,294,321]" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="7209B62C451DFF94FF111A68FDD32C74" blockId="40.[151,1436,151,429]" box="[199,605,294,321]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A68FE5B2C75" bold="true" box="[199,469,294,320]" pageId="40" pageNumber="41">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F348397451DFF94FDDC1A69FDD32C74" box="[522,605,295,321]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="40" pageNumber="41">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7451DFF94FF111A05FEA02C98" pageId="40" pageNumber="41" type="biology_ecology">
<paragraph id="7209B62C451DFF94FF111A05FE612C51" blockId="40.[151,1436,151,429]" box="[199,495,330,357]" pageId="40" pageNumber="41">
<emphasis id="40C26A3E451DFF94FF111A05FE612C51" bold="true" box="[199,495,330,357]" pageId="40" pageNumber="41">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C451DFF94FF111A21FEA02C98" blockId="40.[151,1436,151,429]" pageId="40" pageNumber="41">
This subterranean species is presently known from its
<typeStatus id="AD0D088E451DFF94FCFC1A21FCD42CBC" box="[810,858,367,393]" pageId="40" pageNumber="41">type</typeStatus>
locality and a second, closely situated station, only (see
<tableCitation id="3F348397451DFF94FF1D1ADDFEAF2C98" box="[203,289,403,429]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="40" pageNumber="41">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,193 @@
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<mods:title id="13280BAFB0ED69F5758D1EF7104CFA79">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="78C4943CC6350CE0C543E3E3273FBBF9">Hotèkpo, Sourou Joseph</mods:namePart>
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<mods:affiliation id="EB197509940B553C6EE1171E285BD38E">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université dAbomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
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<mods:namePart id="00494C7D52625320DF2C136794A87093">Schön, Isa</mods:namePart>
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<mods:namePart id="A45DE0746F7512F5268C483760A1BB5D">Martens, Koen</mods:namePart>
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<paragraph id="7209B62C453DFFB4FF411E74FC45284C" blockId="8.[151,971,1338,1401]" pageId="8" pageNumber="9">
<heading id="29410140453DFFB4FF411E74FC2C2860" bold="true" box="[151,930,1338,1365]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="3">
<emphasis id="40C26A3E453DFFB4FF411E74FC2C2860" bold="true" box="[151,930,1338,1365]" pageId="8" pageNumber="9">
Subfamily
<taxonomicName id="B5B6CDAF453DFFB4FEC91E74FE762861" authority="Hotekpo &amp; Martens" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[287,504,1338,1364]" class="Ostracoda" family="Candonidae" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Benincandoninae">Benincandoninae</taxonomicName>
<taxonomicNameLabel id="5BF1D745453DFFB4FE291E74FD102860" box="[511,670,1338,1365]" pageId="8" pageNumber="9" rank="subFamily">subfam. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
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genus:
<taxonomicName id="B5B6CDAF453DFFB4FEF81EE8FE5628F4" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[302,472,1446,1473]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="40C26A3E453DFFB4FEF81EE8FE5628F4" bold="true" box="[302,472,1446,1473]" italics="true" pageId="8" pageNumber="9">Benincandona</emphasis>
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Hotèkpo &amp; Martens
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<emphasis id="40C26A3E453DFFB4FF411D5CFE9E2B19" bold="true" box="[151,272,1554,1580]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453DFFB4FF411D15FDBE2AA0" blockId="8.[151,1437,1627,1941]" pageId="8" pageNumber="9">Cp and valves whitish, opaque, in general laterally flattened; in some species with external linear ornamentation. LV overlapping RV on all sides, but especially dorsally. Both valves without obvious marginal structures, only with ventral remnants of an il. A1 seven-segmented, without fused segments, but with reduced chaetotaxy on some segments. A2 with aesthetasc Y long and slender, sometimes distally bifurcated. MdPalp with seta s2 much shorter than seta s1; one large end claw fused with terminal segment. Mx1-palp with terminal segment quadrate to rectangular. T1 with setae d and d subequal to unequal in length; male prehensile palps subequal. T2 without d1, d2 and e setae, setae f and g present. T3 with seta dp absent, seta d1 long, seta d2 short, long or absent. CR with well-developed ramus, one claw Ga and three small setae (Gp, Sa and Sp). CRatt simple and linear, with few short side branches. Hp with three distal lobes.</paragraph>
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<paragraph id="7209B62C453CFFB5FF411F16FDF129A1" blockId="9.[151,1436,1112,1173]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF411F16FE982945" bold="true" box="[151,278,1112,1136]" pageId="9" pageNumber="10">FIGURE 2.</emphasis>
Map of the Ouémé catchment (and its position in Benin — right inlets), indicating localities where species of
<taxonomicName id="B5B6CDAF453CFFB5FF411F32FEA729A1" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,297,1148,1172]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FF411F32FEA729A1" box="[151,297,1148,1172]" italics="true" pageId="9" pageNumber="10">Benincandona</emphasis>
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were collected.
</paragraph>
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<emphasis id="40C26A3E453CFFB5FF411FA8FE2D2835" bold="true" box="[151,419,1254,1280]" pageId="9" pageNumber="10">Differential diagnosis</emphasis>
</paragraph>
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The new subfamily can be distinguished from the other subfamilies in the
<taxonomicName id="B5B6CDAF453CFFB5FC181E61FBDA287C" authorityName="Kaufmann" authorityYear="1900" box="[974,1108,1327,1353]" class="Ostracoda" family="Candonidae" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Candonidae</taxonomicName>
by a unique set of characters, including the seven-segmented A1 and the CR with a single distal claw. See Discussion for further details and analysis.
</paragraph>
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<emphasis id="40C26A3E453CFFB5FF411EACFC9728C8" bold="true" box="[151,793,1506,1533]" pageId="9" pageNumber="10">
Genus
<taxonomicName id="B5B6CDAF453CFFB5FF381EACFE2E28C8" authority="Hotekpo &amp; Martens" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[238,416,1506,1533]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="40C26A3E453CFFB5FF381EACFE2E28C8" bold="true" box="[238,416,1506,1533]" italics="true" pageId="9" pageNumber="10">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FE711EADFD9B28C8" box="[423,533,1507,1533]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
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<emphasis id="40C26A3E453CFFB5FF411D01FEBB2B5D" bold="true" box="[151,309,1614,1641]" pageId="9" pageNumber="10">
<typeStatus id="AD0D088E453CFFB5FF411D01FF5C2B5C" box="[151,210,1615,1641]" pageId="9" pageNumber="10">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="B5B6CDAF453CFFB5FEEA1D01FDB82B5D" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[316,566,1615,1641]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FEEA1D01FDB82B5D" box="[316,566,1615,1641]" italics="true" pageId="9" pageNumber="10">Benincandona martini</emphasis>
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Hotèkpo &amp; Martens
<emphasis id="40C26A3E453CFFB5FC3A1D00FC7B2B5D" bold="true" box="[1004,1013,1614,1640]" pageId="9" pageNumber="10">(</emphasis>
here designated)
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<emphasis id="40C26A3E453CFFB5FF111D3CFEFA2BB9" bold="true" box="[199,372,1650,1676]" pageId="9" pageNumber="10">Other species:</emphasis>
<taxonomicName id="B5B6CDAF453CFFB5FEAB1D3DFD812BB8" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[381,527,1651,1677]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov">
<emphasis id="40C26A3E453CFFB5FEAB1D3DFD812BB8" box="[381,527,1651,1677]" italics="true" pageId="9" pageNumber="10">B. ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FDCE1D3DFD522BB8" box="[536,732,1651,1677]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FC0D1D3DFBCE2BB8" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[987,1088,1651,1677]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sakitii" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FC0D1D3DFBCE2BB8" box="[987,1088,1651,1677]" italics="true" pageId="9" pageNumber="10">B. sakitii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FB9D1D3DFA9D2BB8" box="[1099,1299,1651,1677]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FED71DD9FEE62B84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[257,360,1687,1713]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cebios" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FED71DD9FEE62B84" box="[257,360,1687,1713]" italics="true" pageId="9" pageNumber="10">B. cebios</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FEB91DD9FDBE2B84" box="[367,560,1687,1713]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens,
<taxonomicName id="B5B6CDAF453CFFB5FCF21DD9FC4A2B84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[804,964,1687,1713]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trapezoides" status="gen. et spec. nov.">
<emphasis id="40C26A3E453CFFB5FCF21DD9FC4A2B84" box="[804,964,1687,1713]" italics="true" pageId="9" pageNumber="10">B. trapezoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453CFFB5FC1D1DD9FB022B84" box="[971,1164,1687,1713]" pageId="9" pageNumber="10" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453CFFB5FF111DF5FE9E2A28" pageId="9" pageNumber="10" type="etymology">
<paragraph id="7209B62C453CFFB5FF111DF5FE9E2A28" blockId="9.[151,1436,1614,1893]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111DF5FEDC2BE0" bold="true" box="[199,338,1723,1749]" pageId="9" pageNumber="10">Etymology:</emphasis>
The prefix of the genus name is derived from the country from where the genus is described,
<collectingCountry id="0AA1F6BC453CFFB5FF411D91FF542BCC" box="[151,218,1759,1785]" name="Benin" pageId="9" pageNumber="10">Benin</collectingCountry>
in West Africa. The suffix is derived from the genus
<taxonomicName id="B5B6CDAF453CFFB5FC961D91FC292BCC" box="[832,935,1759,1785]" class="Ostracoda" family="Candonidae" genus="Candona" kingdom="Animalia" order="Podocopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="40C26A3E453CFFB5FC961D91FC292BCC" box="[832,935,1759,1785]" italics="true" pageId="9" pageNumber="10">Candona</emphasis>
</taxonomicName>
, thus indicating the taxonomic affinity with this genus.
</paragraph>
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<subSubSection id="3AACE5A7453CFFB5FF111C68FC6F2A50" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="7209B62C453CFFB5FF111C68FD6D2A74" blockId="9.[151,1436,1614,1893]" box="[199,739,1830,1857]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111C68FECA2A75" bold="true" box="[199,324,1830,1856]" pageId="9" pageNumber="10">Diagnosis:</emphasis>
As for the subfamily, see discussion.
</paragraph>
<paragraph id="7209B62C453CFFB5FF111C04FC6F2A50" blockId="9.[151,1436,1614,1893]" box="[199,993,1866,1893]" pageId="9" pageNumber="10">
<emphasis id="40C26A3E453CFFB5FF111C04FE5B2A51" bold="true" box="[199,469,1866,1892]" pageId="9" pageNumber="10">Differential Diagnosis:</emphasis>
The new subfamily is at present monogeneric.
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,639 @@
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<mods:title id="F09BBAEAB105CB39E41C8097174221C9">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<emphasis id="40C26A3E453FFFB6FF411BD9FE242D84" bold="true" box="[151,426,151,178]" italics="true" pageId="10" pageNumber="11">Benincandona martini</emphasis>
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Hotèkpo &amp; Martens
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(
<figureCitation id="EA8DAAA9453FFFB6FF491B91FEA92DCC" box="[159,295,223,249]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStartId-0="11.[151,250,1933,1957]" captionStartId-1="12.[151,250,1944,1968]" captionStartId-2="13.[151,250,1943,1967]" captionStartId-3="14.[151,250,1981,2005]" captionStartId-4="16.[151,250,1981,2005]" captionStartId-5="17.[151,250,1943,1967]" captionStartId-6="18.[151,250,1943,1967]" captionTargetBox-0="[172,1415,181,1910]" captionTargetBox-1="[157,1430,181,1920]" captionTargetBox-2="[170,1415,185,1915]" captionTargetBox-3="[169,1422,182,1942]" captionTargetBox-4="[186,1404,198,1941]" captionTargetBox-5="[196,1391,191,1914]" captionTargetBox-6="[172,1415,185,1915]" captionTargetId-0="figure-17@11.[172,1415,178,1910]" captionTargetId-1="figure-31@12.[157,1430,181,1920]" captionTargetId-2="figure-17@13.[167,1419,181,1919]" captionTargetId-3="figure-31@14.[166,1422,181,1957]" captionTargetId-4="figure-31@16.[166,1422,181,1957]" captionTargetId-5="figure-17@17.[186,1401,181,1919]" captionTargetId-6="figure-31@18.[168,1419,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." captionText-1="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." captionText-2="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." captionText-3="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-4="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." captionText-5="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." captionText-6="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="10" pageNumber="11">Figures 39</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FEE21B91FE092DCC" box="[308,391,223,249]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). AD. B. martini. EF. B. ibikounlei. GJ. B. sakitii. KN. B. cebios. OR. B. trapezoides. Scale bar = 500 µm." pageId="10" pageNumber="11">41AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A69FBD22C98" pageId="10" pageNumber="11" type="etymology">
<paragraph id="7209B62C453FFFB6FF411A69FE962C74" blockId="10.[151,1436,295,430]" box="[151,280,295,321]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A69FE962C74" bold="true" box="[151,280,295,321]" pageId="10" pageNumber="11">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411A05FBD22C98" blockId="10.[151,1436,295,430]" pageId="10" pageNumber="11">
The species is named after Dr Patrick Martin (RBINS,
<collectingRegion id="B07278CE453FFFB6FCDD1A05FCE32C50" box="[779,877,331,357]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="10" pageNumber="11">Brussels</collectingRegion>
,
<collectingCountry id="0AA1F6BC453FFFB6FCAF1A05FC552C50" box="[889,987,331,357]" name="Belgium" pageId="10" pageNumber="11">Belgium</collectingCountry>
) in acknowledgement of his significant contributions to groundwater biodiversity research in
<collectingCountry id="0AA1F6BC453FFFB6FD2A1A21FCCC2CBC" box="[764,834,367,393]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
, and with gratitude for his continuous support of the career of SJH and in recognition of decades of collegiality with IS and KM at RBINS.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A9AFE552FF2" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="7209B62C453FFFB6FF411A9AFE842CDB" blockId="10.[151,1437,468,711]" box="[151,266,468,494]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A9AFE842CDB" bold="true" box="[151,266,468,494]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411AB6FE552FF2" blockId="10.[151,1437,468,711]" pageId="10" pageNumber="11">CpRl elongated (L/H = 2.0), sub-triangular, with greatest height in the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, surface with ridges and polygonal ornamentation at the mid-ventral side. CpD and CpV narrow, with greatest width in the middle, lateral sides evenly rounded, anterior and posterior extremities pointed. A2 with seta X long, reaching beyond tip of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with short seta d2. Hp with ls and ms with diverging positions, ls relatively short, reaching slightly beyond ms.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF4119A0FF742864" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="7209B62C453FFFB6FF4119A0FEB02E32" blockId="10.[151,318,749,776]" box="[151,318,749,776]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4119A0FEB02E32" bold="true" box="[151,318,749,776]" pageId="10" pageNumber="11">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411861FE8A2E7C" blockId="10.[151,1282,815,878]" box="[151,260,815,841]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411861FE8A2E7C" bold="true" box="[151,260,815,841]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF411861FE8A2E7C" box="[151,260,815,841]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF41181DFA8C2E5B" blockId="10.[151,1282,815,878]" box="[151,1282,851,878]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF41181DFA8C2E5B" box="[151,1282,851,878]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5453FFFB6FF41181DFF4D2E5B" box="[151,195,851,878]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9453FFFB6FBB0181DFB182E58" box="[1126,1174,851,877]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159200).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118DAFF722E9B" blockId="10.[151,780,916,979]" box="[151,252,916,942]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118DAFF722E9B" bold="true" box="[151,252,916,942]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118DAFF722E9B" box="[151,252,916,942]" pageId="10" pageNumber="11" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118F7FC822EE6" blockId="10.[151,780,916,979]" box="[151,780,952,979]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF4118F7FC822EE6" box="[151,780,952,979]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5453FFFB6FF4118F7FF4F2EE6" box="[151,193,953,979]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
, dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FE2918F6FDEF2EE7" box="[511,609,952,978]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FDA618F6FD282EE6" box="[624,678,952,979]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159201).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118B4FE812921" blockId="10.[151,1436,1018,1189]" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118B4FE812921" bold="true" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118B4FE812921" box="[151,271,1018,1044]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F50FDF92990" blockId="10.[151,1436,1018,1189]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411F50FCC1290C" box="[151,847,1054,1081]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FF411F50FF45290D" box="[151,203,1054,1080]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FDEB1F50FD11290D" box="[573,671,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FD611F50FD62290C" box="[695,748,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159202)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FCB21F50FEA32968" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FCB21F50FC23290D" box="[868,941,1054,1080]" count="2" pageId="10" pageNumber="11" type="male">2 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E453FFFB6FB461F50FB7C290D" box="[1168,1266,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FADC1F50FAB0290C" box="[1290,1342,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159203, INV.159204)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FEE91F0CFBF72968" box="[319,1145,1090,1117]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FEE91F0CFE0A2969" box="[319,388,1090,1116]" count="3" pageId="10" pageNumber="11" type="female">3 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE9453FFFB6FD701F0CFD542968" box="[678,730,1090,1117]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159205, INV.159206, INV.159207)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB5D1F0CFBBE29B4" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB5D1F0CFB5E2969" box="[1163,1232,1090,1116]" count="3" pageId="10" pageNumber="11" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE9453FFFB6FD811F28FD0529B4" box="[599,651,1126,1153]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159208, INV.159209, INV.159210)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB931F28FDFD2990" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB931F28FBF629B5" box="[1093,1144,1126,1152]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE9453FFFB6FE0A1FC4FD9F2990" box="[476,529,1162,1189]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159211)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F85FEBE29D0" blockId="10.[151,1437,1227,1361]" box="[151,304,1227,1253]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411F85FEBE29D0" bold="true" box="[151,304,1227,1253]" pageId="10" pageNumber="11">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411FA1FF742864" blockId="10.[151,1437,1227,1361]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411FA1FB89283C" box="[151,1031,1263,1290]" country="Benin" location="Adjohoun District" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingCountry id="0AA1F6BC453FFFB6FF411FA1FF54283C" box="[151,218,1263,1289]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
,
<collectingRegion id="B07278CE453FFFB6FF301FA1FE4C283F" box="[230,450,1263,1290]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé Department</collectingRegion>
,
<location id="7769E0F7453FFFB6FE1F1FA1FD1B283C" LSID="urn:lsid:plazi:treatment:FA1F073A453FFFAFFF411BD9FA1E2AD6:7769E0F7453FFFB6FE1F1FA1FD1B283C" box="[457,661,1263,1289]" country="Benin" name="Adjohoun District" pageId="10" pageNumber="11" stateProvince="Oueme">Adjohoun District</location>
, town of Azowlissé, located in the
</materialsCitation>
<materialsCitation id="C2DEBC71453FFFB6FBDA1FA1FF782864" collectingDate="2016-08-27" collectingDateMax="2016-08-29" collectingDateMin="2016-08-27" collectorName="Leg. &amp; S. J. Hotekpo" country="Benin" latitude="6.66605" location="Oueme" longLatPrecision="1" longitude="2.4920833" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingRegion id="B07278CE453FFFB6FBDA1FA1FBD3283C" box="[1036,1117,1263,1289]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé</collectingRegion>
catchment area, coordinates:
<geoCoordinate id="1782D0EB453FFFB6FF411E5DFEAA2818" box="[151,292,1299,1325]" degrees="6" direction="north" minutes="39.963" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="6.66605">6° 39.963N</geoCoordinate>
,
<geoCoordinate id="1782D0EB453FFFB6FEF91E5DFE36281B" box="[303,440,1299,1326]" degrees="2" direction="east" minutes="29.525" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="2.4920833">2° 29.525E</geoCoordinate>
,
<collectorName id="DF43D3FA453FFFB6FE151E5AFE74281B" box="[451,506,1300,1326]" pageId="10" pageNumber="11">Leg.</collectorName>
:
<collectorName id="DF43D3FA453FFFB6FDD21E5DFD182818" box="[516,662,1299,1325]" pageId="10" pageNumber="11">S.J. Hotèkpo</collectorName>
, collected on
<date id="060890EC453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">
<collectingDate id="164C6904453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">27 and 29 August 2016</collectingDate>
</date>
(Locality code BVS
<quantity id="B54E1BC9453FFFB6FAC11E5DFAC02818" box="[1303,1358,1299,1325]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="10" pageNumber="11" unit="in" value="19.0">19 in</quantity>
<tableCitation id="3F348397453FFFB6FA851E5DFF2B2864" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="10" pageNumber="11">Tables 1</tableCitation>
and
<tableCitation id="3F348397453FFFB6FF0A1E79FF632864" box="[220,237,1335,1361]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="10" pageNumber="11">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFAFFF411E36FD8A28A8" lastPageId="19" lastPageNumber="20" pageId="10" pageNumber="11" type="description">
<paragraph id="7209B62C453FFFB6FF411E36FE0F28A6" blockId="10.[151,1437,1400,2039]" box="[151,385,1400,1427]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411E36FE0F28A6" bold="true" box="[151,385,1400,1427]" pageId="10" pageNumber="11">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411ED2FB592BBA" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
CpRl (
<figureCitation id="EA8DAAA9453FFFB6FF091ED2FEFC2882" box="[223,370,1436,1463]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3A, C</figureCitation>
) elongated and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; external ornamentation consisting of linear ridges (
<figureCitation id="EA8DAAA9453FFFB6FE061D62FDE42B72" box="[464,618,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3AD</figureCitation>
) and polygonal structures (
<figureCitation id="EA8DAAA9453FFFB6FC741D62FB802B72" box="[930,1038,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3I</figureCitation>
) in mid-ventral position of valves. CpD (
<figureCitation id="EA8DAAA9453FFFB6FF0C1D1EFEC52B5E" box="[218,331,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3E</figureCitation>
) and CpV (
<figureCitation id="EA8DAAA9453FFFB6FE1C1D1EFDB42B5E" box="[458,570,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figure 3F</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins evenly rounded; LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111DD6FB8C2A77" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
LVi (
<figureCitation id="EA8DAAA9453FFFB6FED71DD6FE0D2B86" box="[257,387,1688,1715]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3G</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE421DD7FE552B86" box="[404,475,1689,1715]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4A, C</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, overlap present at that site, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; CMS (
<figureCitation id="EA8DAAA9453FFFB6FC261C4AFBED2A2A" box="[1008,1123,1796,1823]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">Figure 4C</figureCitation>
) consisting of six scars: one large dorsal, three anterior and two posterior ones; one short antero-ventral il.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111C02FCDE2AC3" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
RVi (
<figureCitation id="EA8DAAA9453FFFB6FF291C02FE182A52" box="[255,406,1868,1895]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3H, J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE771C03FE6D2A52" box="[417,483,1869,1895]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4B, D</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest with height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin but with a blunt angle in the middle, posterior part sloping almost straight to posterior margin, ventral margin almost straight. CMS (
<figureCitation id="EA8DAAA9453FFFB6FE441CF6FD862AE6" box="[402,520,1976,2003]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="10" pageNumber="11">Figures 3J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FDC51CF7FDB92AE6" box="[531,567,1977,2003]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="10" pageNumber="11">4D</figureCitation>
) consisting of six scars: one large dorsal (1), three anterior (24) and two (5,6) posterior ones; il running submarginally along ventral margin.
</paragraph>
<caption id="26C9E6A4453EFFB7FF411CC3FC722ADB" pageId="11" pageNumber="12" startId="11.[151,250,1933,1957]" targetBox="[172,1415,181,1910]" targetPageId="11" targetType="figure">
<paragraph id="7209B62C453EFFB7FF411CC3FC722ADB" blockId="11.[151,1437,1933,2030]" pageId="11" pageNumber="12">
<emphasis id="40C26A3E453EFFB7FF411CC3FE802A90" bold="true" box="[151,270,1933,1957]" pageId="11" pageNumber="12">FIGURE 3</emphasis>
.
<taxonomicName id="B5B6CDAF453EFFB7FECD1CC3FD8E2A90" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[283,512,1933,1957]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453EFFB7FECD1CC3FD8E2A90" box="[283,512,1933,1957]" italics="true" pageId="11" pageNumber="12">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453EFFB7FDD11CC0FD352A90" box="[519,699,1934,1958]" pageId="11" pageNumber="12" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (A, C, EH, J ♂; B, D, I ♀).
<emphasis id="40C26A3E453EFFB7FBEF1CC3FB0C2A93" bold="true" box="[1081,1154,1933,1958]" pageId="11" pageNumber="12">AD, I</emphasis>
. CpRl, (A, C. INV.159208; B, D, I. INV.159211).
<emphasis id="40C26A3E453EFFB7FEAB1CFCFE002AFF" bold="true" box="[381,398,1970,1994]" pageId="11" pageNumber="12">E</emphasis>
. INV.159209. CpD.
<emphasis id="40C26A3E453EFFB7FDB41CFCFDFF2AFF" bold="true" box="[610,625,1970,1994]" pageId="11" pageNumber="12">F</emphasis>
. INV.159210. CpV.
<emphasis id="40C26A3E453EFFB7FC951CFFFCD92AFC" bold="true" box="[835,855,1969,1993]" pageId="11" pageNumber="12">G</emphasis>
. INV.159200. LVi.
<emphasis id="40C26A3E453EFFB7FBF61CFCFBBA2AFF" bold="true" box="[1056,1076,1970,1994]" pageId="11" pageNumber="12">H</emphasis>
. INV.159200. RVi.
<emphasis id="40C26A3E453EFFB7FAD61CFCFA832AFF" bold="true" box="[1280,1293,1970,1994]" pageId="11" pageNumber="12">J</emphasis>
. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44539FFB0FF411CD6FC792AE0" pageId="12" pageNumber="13" startId="12.[151,250,1944,1968]" targetBox="[157,1430,181,1920]" targetPageId="12" targetType="figure">
<paragraph id="7209B62C4539FFB0FF411CD6FC792AE0" blockId="12.[151,1436,1944,2005]" pageId="12" pageNumber="13">
<emphasis id="40C26A3E4539FFB0FF411CD6FE9D2A85" bold="true" box="[151,275,1944,1968]" pageId="12" pageNumber="13">FIGURE 4.</emphasis>
<taxonomicName id="B5B6CDAF4539FFB0FECC1CD6FE702A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[282,510,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4539FFB0FECC1CD6FE702A85" box="[282,510,1944,1968]" italics="true" pageId="12" pageNumber="13">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454539FFB0FDD31CD7FD392A85" box="[517,695,1945,1969]" pageId="12" pageNumber="13" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (AD: INV.159202 ♂ and EH. INV.159201 ♀)
<emphasis id="40C26A3E4539FFB0FB2F1CD6FAA72A85" bold="true" box="[1273,1321,1944,1968]" pageId="12" pageNumber="13">A, E</emphasis>
. LVi.
<emphasis id="40C26A3E4539FFB0FABE1CD6FA1B2A85" bold="true" box="[1384,1429,1944,1968]" pageId="12" pageNumber="13">B, F</emphasis>
. RVi,
<emphasis id="40C26A3E4539FFB0FF1D1CF2FEB32AE1" bold="true" box="[203,317,1980,2004]" pageId="12" pageNumber="13">C, D, G, H</emphasis>
. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44538FFB1FF411CD9FD0F2AE1" pageId="13" pageNumber="14" startId="13.[151,250,1943,1967]" targetBox="[170,1415,185,1915]" targetPageId="13" targetType="figure">
<paragraph id="7209B62C4538FFB1FF411CD9FD0F2AE1" blockId="13.[151,1436,1943,2004]" pageId="13" pageNumber="14">
<emphasis id="40C26A3E4538FFB1FF411CD9FE9A2A85" bold="true" box="[151,276,1943,1968]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
<taxonomicName id="B5B6CDAF4538FFB1FECA1CD6FD8F2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[284,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4538FFB1FECA1CD6FD8F2A9A" box="[284,513,1944,1968]" italics="true" pageId="13" pageNumber="14">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454538FFB1FDDF1CD6FD312A9A" box="[521,703,1944,1968]" pageId="13" pageNumber="14" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4538FFB1FCEA1CD9FCC02A9A" bold="true" box="[828,846,1943,1967]" pageId="13" pageNumber="14">A</emphasis>
. A1 (INV.159203).
<emphasis id="40C26A3E4538FFB1FBCB1CD6FBC02A9A" bold="true" box="[1053,1102,1943,1968]" pageId="13" pageNumber="14">B, C</emphasis>
. MdCoxa, (INV.159202).
<emphasis id="40C26A3E4538FFB1FA881CD6FAFE2A85" bold="true" box="[1374,1392,1944,1968]" pageId="13" pageNumber="14">D</emphasis>
. A2 (INV.159200 INV.159203). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4453BFFB2FF411CF3FA8E2ACF" pageId="14" pageNumber="15" startId="14.[151,250,1981,2005]" targetBox="[169,1422,182,1942]" targetPageId="14" targetType="figure">
<paragraph id="7209B62C453BFFB2FF411CF3FA8E2ACF" blockId="14.[151,1436,1981,2042]" pageId="14" pageNumber="15">
<emphasis id="40C26A3E453BFFB2FF411CF3FE982AE0" bold="true" box="[151,278,1981,2005]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
<taxonomicName id="B5B6CDAF453BFFB2FEC91CF3FD8B2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[287,517,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453BFFB2FEC91CF3FD8B2AE0" box="[287,517,1981,2005]" italics="true" pageId="14" pageNumber="15">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453BFFB2FDD81CF0FD492AE0" box="[526,711,1982,2006]" pageId="14" pageNumber="15" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E453BFFB2FC9E1CF3FCD42AE0" bold="true" box="[840,858,1981,2005]" pageId="14" pageNumber="15">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159200).
<emphasis id="40C26A3E453BFFB2FA901CF0FAD92AE3" bold="true" box="[1350,1367,1982,2006]" pageId="14" pageNumber="15">B</emphasis>
. Mx1, respiratory plate (INV.159202).
<emphasis id="40C26A3E453BFFB2FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="14" pageNumber="15">C</emphasis>
. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C453AFFB3FF111BD9FCD62F2C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A1 (
<figureCitation id="EA8DAAA9453AFFB3FF2E1BD9FEE32D84" box="[248,365,150,177]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5A</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment
<emphasis id="40C26A3E453AFFB3FAD01BD9FA942D85" box="[1286,1306,151,176]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
three times as long as basal width, with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer seta; and two shorter setae on the dorsal side, situated some distance from each other (one sub-apical and one more proximal). First distal segment slightly longer than basal width, with one short dorso-apical seta. Second distal segment about twice as long as wide, set with one short dorso-apical seta. Third distal segment with about the same length/width ratio as the second segment, carrying one short apical seta on each side. Fourth distal segment about three times as long as wide, with one ventro-apical seta, and one long dorso-apical seta, reaching beyond tip of sixth segment with almost one third of total length. Fifth distal segment four times as long as wide, set with two long dorso-apical setae and one short ventro-apical seta. Sixth distal segment narrow,
<emphasis id="40C26A3E453AFFB3FB261AF9FA8D2CE5" box="[1264,1283,439,464]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long setae more than twice the length of Ya and one short seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C453AFFB3FF11196DFCE5296C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A2 (
<figureCitation id="EA8DAAA9453AFFB3FF20196DFEE72F08" box="[246,361,546,573]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5D</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E453AFFB3FCAC1909FC032F55" box="[890,909,583,608]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E453AFFB3FD8A1925FDE12FB1" box="[604,623,619,644]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X, reaching tip of next segment (first endopodal segment). No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout,
<emphasis id="40C26A3E453AFFB3FBB319FDFBFF2FF9" box="[1125,1137,691,716]" italics="true" pageId="15" pageNumber="16">c</emphasis>
four times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y (reaching tip of segment) and two ventro-subapical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E453AFFB3FAFF19B5FAB22E21" box="[1321,1340,763,788]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9453AFFB3FBD518C5FBF82E90" box="[1027,1142,907,933]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="15" pageNumber="16">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of equal length; centrally with one short seta z3; dorso-apically with the seta z1 transformed into claw, length about twice as long as penultimate segment. Claw G1 slightly shorter than G2; seta z2 transformed into long claw, slightly shorter than G1. Fourth, terminal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a long seta g slightly shorter than claw Gm.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F2DFBDC29F0" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdCoxa (
<figureCitation id="EA8DAAA9453AFFB3FEEF1F2DFE5F2948" box="[313,465,1122,1149]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5B, C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (67) of strong teeth, interspaced with setae; subapically with a short seta.
<emphasis id="40C26A3E453AFFB3FAEB1FC9FA1A2994" box="[1341,1428,1159,1185]" italics="true" pageId="15" pageNumber="16">Remark</emphasis>
: asymmetry illustrated in
<figureCitation id="EA8DAAA9453AFFB3FE781FE5FDAE29F0" box="[430,544,1194,1221]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200 INV.159203). Scale bar = 50 µm." pageId="15" pageNumber="16">Figure 5C</figureCitation>
possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F81FE092B64" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdPalp (
<figureCitation id="EA8DAAA9453AFFB3FEF81F81FE2F29DC" box="[302,417,1231,1257]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6C</figureCitation>
) set at about one third of the tip of the coxa and consisting of four segments. Basal segment dorsally carrying the respiratory plate, set with six distal rays (four long, two short), subapically with one long and two short rays, proximally with one long ray (this chaetotaxy approximate); ventro-subapically with one long hirsute seta (S1) and more apically S2 hirsute seta, shorter and stout, length about one third of the length of S1; in between with the short flagellum-like alpha seta, and one smooth and short, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, length of dorsal one (slightly shorter than the fused claw) about 1.5 times length of ventral one.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111D15FCB12A78" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
Mx1 (
<figureCitation id="EA8DAAA9453AFFB3FEDD1D15FE2E2B40" box="[267,416,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Palp (
<figureCitation id="EA8DAAA9453AFFB3FD771D31FC992BAC" box="[673,791,1663,1689]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E453AFFB3FB701D89FAB22BD4" box="[1190,1340,1735,1761]" italics="true" pageId="15" pageNumber="16">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E453AFFB3FB141DA5FB5B2A31" box="[1218,1237,1771,1796]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
half the length of the long one. Respiratory plate (
<figureCitation id="EA8DAAA9453AFFB3FE2F1C41FDE22A1C" box="[505,620,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204 INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." pageId="15" pageNumber="16">Figure 6B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E453AFFB3FC8E1C41FCE52A1D" box="[856,875,1807,1832]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
15 plumose respiratory rays, two short distal setae and a group of five shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44525FFACFF411CF3FD432ACF" pageId="16" pageNumber="17" startId="16.[151,250,1981,2005]" targetBox="[186,1404,198,1941]" targetPageId="16" targetType="figure">
<paragraph id="7209B62C4525FFACFF411CF3FD432ACF" blockId="16.[151,1436,1981,2042]" pageId="16" pageNumber="17">
<emphasis id="40C26A3E4525FFACFF411CF3FE962AE3" bold="true" box="[151,280,1981,2006]" pageId="16" pageNumber="17">FIGURE 7.</emphasis>
<taxonomicName id="B5B6CDAF4525FFACFEF21CF3FD832AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,525,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4525FFACFEF21CF3FD832AE0" box="[292,525,1981,2005]" italics="true" pageId="16" pageNumber="17">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454525FFACFDCE1CF0FD542AE0" box="[536,730,1982,2006]" pageId="16" pageNumber="17" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4525FFACFCB51CF3FCFB2AE0" bold="true" box="[867,885,1981,2005]" pageId="16" pageNumber="17">A</emphasis>
. T2 (INV.159200).
<emphasis id="40C26A3E4525FFACFB9B1CF0FBD02AE3" bold="true" box="[1101,1118,1982,2006]" pageId="16" pageNumber="17">B</emphasis>
. CR (INV.159200).
<emphasis id="40C26A3E4525FFACFAEB1CF3FAC12AE0" bold="true" box="[1341,1359,1981,2005]" pageId="16" pageNumber="17">C</emphasis>
. CRatt (INV.159200).
<emphasis id="40C26A3E4525FFACFEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="16" pageNumber="17">D</emphasis>
. T3 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44524FFADFF411CD9FDB32ACD" pageId="17" pageNumber="18" startId="17.[151,250,1943,1967]" targetBox="[196,1391,191,1914]" targetPageId="17" targetType="figure">
<paragraph id="7209B62C4524FFADFF411CD9FDB32ACD" blockId="17.[151,1437,1943,2040]" pageId="17" pageNumber="18">
<emphasis id="40C26A3E4524FFADFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="17" pageNumber="18">FIGURE 8.</emphasis>
<taxonomicName id="B5B6CDAF4524FFADFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4524FFADFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="17" pageNumber="18">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454524FFADFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="17" pageNumber="18" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4524FFADFC9B1CD9FCD12A9A" bold="true" box="[845,863,1943,1967]" pageId="17" pageNumber="18">A</emphasis>
. Zenker organ (INV.159200).
<emphasis id="40C26A3E4524FFADFB771CD6FB5E2A85" bold="true" box="[1185,1232,1944,1968]" pageId="17" pageNumber="18">BD</emphasis>
. Hp (INV.159200).
<emphasis id="40C26A3E4524FFADFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="17" pageNumber="18">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4524FFADFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="17" pageNumber="18">C</emphasis>
. Internal view (photograph with interference contrast).
<emphasis id="40C26A3E4524FFADFBCE1CF2FBA42AE1" bold="true" box="[1048,1066,1980,2004]" pageId="17" pageNumber="18">D</emphasis>
. Internal view (photograph without interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44527FFAEFF411CD9FA9F2AE1" pageId="18" pageNumber="19" startId="18.[151,250,1943,1967]" targetBox="[172,1415,185,1915]" targetPageId="18" targetType="figure">
<paragraph id="7209B62C4527FFAEFF411CD9FA9F2AE1" blockId="18.[151,1436,1943,2004]" pageId="18" pageNumber="19">
<emphasis id="40C26A3E4527FFAEFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
<taxonomicName id="B5B6CDAF4527FFAEFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4527FFAEFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="18" pageNumber="19">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454527FFAEFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="18" pageNumber="19" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (AB ♂ and CE ♀).
<emphasis id="40C26A3E4527FFAEFBDE1CD9FB942A9A" bold="true" box="[1032,1050,1943,1967]" pageId="18" pageNumber="19">A</emphasis>
. T1, Lpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFAF01CD6FAB92A85" bold="true" box="[1318,1335,1944,1968]" pageId="18" pageNumber="19">B</emphasis>
. T1, Rpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="18" pageNumber="19">C</emphasis>
. A2 (INV.159205, INV.159207).
<emphasis id="40C26A3E4527FFAEFD401CF2FD262AE1" bold="true" box="[662,680,1980,2004]" pageId="18" pageNumber="19">D</emphasis>
. CR (INV.159205).
<emphasis id="40C26A3E4527FFAEFCA11CF2FC062AE1" bold="true" box="[887,904,1980,2004]" pageId="18" pageNumber="19">E</emphasis>
. T1 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFF111BD9FD062C50" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF201BD9FE012D84" box="[246,399,151,177]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented here by one long and one short plumose setae, and slightly asymmetrical prehensile palps (endopodites). Protopodite (
<figureCitation id="EA8DAAA94526FFAFFF491B91FE9E2DCC" box="[159,272,223,249]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9B</figureCitation>
) distally set with 5+2 setae of different morphology and length (not illustrated); more proximally carrying d, d and b setae of medium length. Prehensile palps (
<figureCitation id="EA8DAAA94526FFAFFD341A4DFCFB2C28" box="[738,885,259,285]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps apically set with a short sensory organ, proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA94526FFAFFA871A69FF352C50" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA94526FFAFFDD31A05FDF52C50" box="[517,635,331,357]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111A21FCD12F08" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T2 (
<figureCitation id="EA8DAAA94526FFAFFF201A21FEE32CBC" box="[246,365,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment and knee-segment without setae d1 and d2. First segment of endopodite elongated (length at least 4x the basal width), without setae. Second endopodal segment with length
<emphasis id="40C26A3E4526FFAFFC451AF9FC292CE5" box="[915,935,439,464]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
3x basal width, with one short ventro-apical seta f. Third endopodal segment of about the same length/width ratio as the previous segment and with one ventro-subapical seta g. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111909FBDB2FC4" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T3 (
<figureCitation id="EA8DAAA94526FFAFFF211909FEE12F54" box="[247,367,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7D</figureCitation>
) with first segment carrying only one long subapical seta d1. Second (knee-) segment with a short seta d2. Third segment about 5x as long as wide, not carrying any setae. Fourth segment narrower, but still
<emphasis id="40C26A3E4526FFAFFA5F1925FA122FB1" box="[1417,1436,619,644]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
5x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4526FFAFFF7919FDFF4C2FF9" box="[175,194,691,716]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 &amp; h2, the former
<emphasis id="40C26A3E4526FFAFFCDD1999FC902FC5" box="[779,798,727,752]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4526FFAFFF1119B5FC852E0C" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
CR (
<figureCitation id="EA8DAAA94526FFAFFF2819B5FEFD2E20" box="[254,371,763,789]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7B</figureCitation>
) with ramus stout; claw Ga well-developed, stout and reaching widely beyond half L of the ramus, setae Sp, Sa and claw Gp reduced to small setae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11180DFCA92E68" blockId="19.[151,1437,151,1005]" box="[199,807,835,861]" pageId="19" pageNumber="20">
CRAtt (
<figureCitation id="EA8DAAA94526FFAFFEC9180DFE1D2E68" box="[287,403,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 7C</figureCitation>
) short and stout, distally bifurcated.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111829FD4E2EFC" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
Hp (
<figureCitation id="EA8DAAA94526FFAFFF2F1829FE1E2EB4" box="[249,400,871,897]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8BD</figureCitation>
) relatively small, with bluntly pointed ls, reaching slightly beyond asymmetrically rounded ms; lobe h shorter and sub-rectangular, ls and ms with diverging positions; internal anatomy (
<figureCitation id="EA8DAAA94526FFAFFB1218C5FAD52E90" box="[1220,1371,907,933]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11189DFB892ED8" blockId="19.[151,1437,151,1005]" box="[199,1031,979,1005]" pageId="19" pageNumber="20">
Zenker organ (
<figureCitation id="EA8DAAA94526FFAFFEBD189DFE6F2ED8" box="[363,481,979,1005]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). BD. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 8A</figureCitation>
) short and broad, with eight-nine spinous whirls.
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F54FE182901" blockId="19.[151,1436,1050,1545]" box="[151,406,1050,1076]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF411F54FE182901" bold="true" box="[151,406,1050,1076]" pageId="19" pageNumber="20">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F71FB10296C" blockId="19.[151,1436,1050,1545]" box="[151,1182,1087,1113]" pageId="19" pageNumber="20">
Cp and valves as in the male, without obvious sexual dimorphism (
<figureCitation id="EA8DAAA94526FFAFFC501F71FBCE296C" box="[902,1088,1087,1113]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, EH, J ♂; B, D, I ♀). AD, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." pageId="19" pageNumber="20">Figures 3B, D, I</figureCitation>
;
<figureCitation id="EA8DAAA94526FFAFFB9A1F71FB1C296C" box="[1100,1170,1087,1113]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (AD: INV.159202 ♂ and EH. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." pageId="19" pageNumber="20">4EH</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111F2DFBF52948" blockId="19.[151,1436,1050,1545]" box="[199,1147,1123,1149]" pageId="19" pageNumber="20">
Limbs (
<figureCitation id="EA8DAAA94526FFAFFECB1F2DFE3E2948" box="[285,432,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9CE</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111FC9FC232838" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
A2 (
<figureCitation id="EA8DAAA94526FFAFFF2F1FC9FEE12994" box="[249,367,1159,1185]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9C</figureCitation>
) with second endopodal segment undivided, with two t-setae (one long and plumose (t1), one smooth short and thin (t2)); apical chaetotaxy (
<figureCitation id="EA8DAAA94526FFAFFD771FE5FC9E29F0" box="[673,784,1195,1221]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." pageId="19" pageNumber="20">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111E59FC31284C" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF2F1E59FEE12804" box="[249,367,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (AB ♂ and CE ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." pageId="19" pageNumber="20">Figure 9E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4526FFAFFDA01E59FD072805" box="[630,649,1303,1328]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
10 distal setae of unequal length, setae d and b present, with long additional seta (d) present close to seta d; respiratory plate reduced to about three short rays. Endopodite set with a broad and short palp, with three subequal but short apical setae (h1h3).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111ECDFD8A28A8" blockId="19.[151,1436,1050,1545]" box="[199,516,1411,1437]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111ECDFE0E28A8" bold="true" box="[199,384,1411,1437]" pageId="19" pageNumber="20">Measurements:</emphasis>
see
<tableCitation id="3F3483974526FFAFFE671ECDFD8A28A8" box="[433,516,1411,1437]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="19" pageNumber="20">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111EE8FDD328F4" box="[199,605,1446,1473]" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="7209B62C4526FFAFFF111EE8FDD328F4" blockId="19.[151,1436,1050,1545]" box="[199,605,1446,1473]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111EE8FE5B28F5" bold="true" box="[199,469,1446,1472]" pageId="19" pageNumber="20">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974526FFAFFDDC1EE9FDD328F4" box="[522,605,1447,1473]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="19" pageNumber="20">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111E85FA1E2AD6" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="7209B62C4526FFAFFF111E85FE6128D1" blockId="19.[151,1436,1050,1545]" box="[199,495,1482,1509]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111E85FE6128D1" bold="true" box="[199,495,1482,1509]" pageId="19" pageNumber="20">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF111EA1FBD02B3C" blockId="19.[151,1436,1050,1545]" box="[199,1118,1519,1545]" pageId="19" pageNumber="20">
This subterranean species is presently only known from its
<typeStatus id="AD0D088E4526FFAFFCB61EA1FC1E2B3C" box="[864,912,1519,1545]" pageId="19" pageNumber="20">type</typeStatus>
locality (
<tableCitation id="3F3483974526FFAFFC2C1EA1FBC32B3C" box="[1018,1101,1519,1545]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="19" pageNumber="20">Table 1</tableCitation>
).
</paragraph>
<caption id="26C9E6A44526FFAFFF4E1D76FB332B64" box="[152,1213,1592,1617]" pageId="19" pageNumber="20" startId="19.[152,244,1592,1616]" targetType="table">
<paragraph id="7209B62C4526FFAFFF4E1D76FB332B64" blockId="19.[152,1213,1592,1617]" box="[152,1213,1592,1617]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF4E1D76FE812B65" bold="true" box="[152,271,1592,1616]" pageId="19" pageNumber="20">TABLE 3.</emphasis>
Measurements of illustrated specimens of
<taxonomicName id="B5B6CDAF4526FFAFFD0F1D76FCFD2B65" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[729,883,1592,1616]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFD0F1D76FCFD2B65" box="[729,883,1592,1616]" italics="true" pageId="19" pageNumber="20">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFCAF1D77FBBA2B65" box="[889,1076,1593,1617]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, using SEM.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFE8C1D2EFAF62AA7" pageId="19" pageNumber="20">
<table id="00B6448C45260043FF751D2EFA0B2AA7" box="[163,1413,1632,1938]" gridcols="7" gridrows="8" pageId="19" pageNumber="20">
<tr id="CC86B46E45260043FF751D2EFA0B2B4D" box="[163,1413,1632,1656]" gridrow="0" pageId="19" pageNumber="20">
<th id="8F57DD1245260043FF751D2EFE312B4D" box="[163,447,1632,1656]" gridcol="0" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFE8C1D2EFE272B4D" bold="true" box="[346,425,1632,1656]" pageId="19" pageNumber="20">Species</emphasis>
</th>
<th id="8F57DD1245260043FD4D1D2EFD4F2B4D" box="[667,705,1632,1656]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFD4D1D2EFD4F2B4D" bold="true" box="[667,705,1632,1656]" pageId="19" pageNumber="20">Sex</emphasis>
</th>
<th id="8F57DD1245260043FCD51D2EFCF12B4D" box="[771,895,1632,1656]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFCD91D2EFCFD2B4D" bold="true" box="[783,883,1632,1656]" pageId="19" pageNumber="20">specimen</emphasis>
</th>
<th id="8F57DD1245260043FC611D2EFC7E2B4D" box="[951,1008,1632,1656]" gridcol="3" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFC611D2EFC7E2B4D" bold="true" box="[951,1008,1632,1656]" pageId="19" pageNumber="20">V/Cp</emphasis>
</th>
<th id="8F57DD1245260043FBE71D2EFBF52B4D" box="[1073,1147,1632,1656]" gridcol="4" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFBE71D2EFBF52B4D" bold="true" box="[1073,1147,1632,1656]" pageId="19" pageNumber="20">L (µm)</emphasis>
</th>
<th id="8F57DD1245260043FB641D2EFA8F2B4D" box="[1202,1281,1632,1656]" gridcol="5" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB641D2EFA8F2B4D" bold="true" box="[1202,1281,1632,1656]" pageId="19" pageNumber="20">H (µm)</emphasis>
</th>
<th id="8F57DD1245260043FAE41D2EFA0B2B4D" box="[1330,1413,1632,1656]" gridcol="6" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFAE41D2EFA0B2B4D" bold="true" box="[1330,1413,1632,1656]" pageId="19" pageNumber="20">W (µm)</emphasis>
</th>
</tr>
<tr id="CC86B46E45260043FF751DC6FA0B2B94" box="[163,1413,1672,1697]" gridrow="1" pageId="19" pageNumber="20" rowspan-6="1">
<th id="8F57DD1245260043FF751DC6FE312B94" box="[163,447,1672,1697]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">
<taxonomicName id="B5B6CDAF4526FFAFFF751DC7FE842B95" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[163,266,1673,1696]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFF751DC7FE842B95" box="[163,266,1673,1696]" italics="true" pageId="19" pageNumber="20">B. martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFEC61DC7FE312B95" box="[272,447,1673,1697]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
</th>
<td id="8F57DD1245260043FD4D1DC6FD4F2B94" box="[667,705,1672,1697]" gridcol="1" gridrow="1" pageId="19" pageNumber="20"></td>
<td id="8F57DD1245260043FCD51DC6FCF12B94" box="[771,895,1672,1697]" gridcol="2" gridrow="1" pageId="19" pageNumber="20">INV.159200</td>
<td id="8F57DD1245260043FC611DC6FC7E2B94" box="[951,1008,1672,1697]" gridcol="3" gridrow="1" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71DC6FBF52B94" box="[1073,1147,1672,1697]" gridcol="4" gridrow="1" pageId="19" pageNumber="20">706.7</td>
<td id="8F57DD1245260043FB641DC6FA8F2B94" box="[1202,1281,1672,1697]" gridcol="5" gridrow="1" pageId="19" pageNumber="20">331.7</td>
</tr>
<tr id="CC86B46E45260043FF751DFFFA0B2BFC" box="[163,1413,1713,1737]" gridrow="2" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611DFFFC7E2BFC" box="[951,1008,1713,1737]" gridcol="3" gridrow="2" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71DFFFBF52BFC" box="[1073,1147,1713,1737]" gridcol="4" gridrow="2" pageId="19" pageNumber="20">697.9</td>
<td id="8F57DD1245260043FB641DFFFA8F2BFC" box="[1202,1281,1713,1737]" gridcol="5" gridrow="2" pageId="19" pageNumber="20">310.7</td>
</tr>
<tr id="CC86B46E45260043FF751D97FA0B2BC4" box="[163,1413,1753,1777]" gridrow="3" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51D97FCF12BC4" box="[771,895,1753,1777]" gridcol="2" gridrow="3" pageId="19" pageNumber="20">INV.159202</td>
<td id="8F57DD1245260043FC611D97FC7E2BC4" box="[951,1008,1753,1777]" gridcol="3" gridrow="3" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71D97FBF52BC4" box="[1073,1147,1753,1777]" gridcol="4" gridrow="3" pageId="19" pageNumber="20">667.5</td>
<td id="8F57DD1245260043FB641D97FA8F2BC4" box="[1202,1281,1753,1777]" gridcol="5" gridrow="3" pageId="19" pageNumber="20">317.5</td>
</tr>
<tr id="CC86B46E45260043FF751C4FFA0B2A2C" box="[163,1413,1793,1817]" gridrow="4" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611C4FFC7E2A2C" box="[951,1008,1793,1817]" gridcol="3" gridrow="4" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71C4FFBF52A2C" box="[1073,1147,1793,1817]" gridcol="4" gridrow="4" pageId="19" pageNumber="20">656.7</td>
<td id="8F57DD1245260043FB641C4FFA8F2A2C" box="[1202,1281,1793,1817]" gridcol="5" gridrow="4" pageId="19" pageNumber="20">295.0</td>
</tr>
<tr id="CC86B46E45260043FF751C67FA0B2A77" box="[163,1413,1833,1858]" gridrow="5" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51C67FCF12A77" box="[771,895,1833,1858]" gridcol="2" gridrow="5" pageId="19" pageNumber="20">INV.159208</td>
<td id="8F57DD1245260043FC611C67FC7E2A77" box="[951,1008,1833,1858]" gridcol="3" gridrow="5" pageId="19" pageNumber="20">CpRl</td>
<td id="8F57DD1245260043FBE71C67FBF52A77" box="[1073,1147,1833,1858]" gridcol="4" gridrow="5" pageId="19" pageNumber="20">692.9</td>
<td id="8F57DD1245260043FB641C67FA8F2A77" box="[1202,1281,1833,1858]" gridcol="5" gridrow="5" pageId="19" pageNumber="20">334.3</td>
</tr>
<tr id="CC86B46E45260043FF751C1CFA0B2A5F" box="[163,1413,1874,1898]" gridrow="6" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C1CFCF12A5F" box="[771,895,1874,1898]" gridcol="2" gridrow="6" pageId="19" pageNumber="20">INV.159209</td>
<td id="8F57DD1245260043FC611C1CFC7E2A5F" box="[951,1008,1874,1898]" gridcol="3" gridrow="6" pageId="19" pageNumber="20">CpD</td>
<td id="8F57DD1245260043FBE71C1CFBF52A5F" box="[1073,1147,1874,1898]" gridcol="4" gridrow="6" pageId="19" pageNumber="20">698.6</td>
<td id="8F57DD1245260043FAE41C1CFA0B2A5F" box="[1330,1413,1874,1898]" gridcol="6" gridrow="6" pageId="19" pageNumber="20">185.0</td>
</tr>
<tr id="CC86B46E45260043FF751C34FA0B2AA7" box="[163,1413,1914,1938]" gridrow="7" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C34FCF12AA7" box="[771,895,1914,1938]" gridcol="2" gridrow="7" pageId="19" pageNumber="20">INV.159210</td>
<td id="8F57DD1245260043FC611C34FC7E2AA7" box="[951,1008,1914,1938]" gridcol="3" gridrow="7" pageId="19" pageNumber="20">CpV</td>
<td id="8F57DD1245260043FBE71C34FBF52AA7" box="[1073,1147,1914,1938]" gridcol="4" gridrow="7" pageId="19" pageNumber="20">679.3</td>
<td id="8F57DD1245260043FAE41C34FA0B2AA7" box="[1330,1413,1914,1938]" gridcol="6" gridrow="7" pageId="19" pageNumber="20">190.7</td>
</tr>
</table>
</paragraph>
<paragraph id="7209B62C4526FFAFFB8F1C85FA1E2AD6" blockId="19.[1113,1424,1995,2019]" box="[1113,1424,1995,2019]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB8F1C85FA1E2AD6" box="[1113,1424,1995,2019]" italics="true" pageId="19" pageNumber="20">......continued on the next page</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>