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<mods:title id="3828B68C621D06185843E62EBE37F4A3">Pilosella bulgarica (Asteraceae) a new hybridogenous species from Bulgaria</mods:title>
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<mods:namePart id="7B1EC232A81641D4629CD959524CAD5A">Szeląg, Zbigniew</mods:namePart>
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<taxonomicName id="4C2A4D39D8341649FF61F97AC237EA99" ID-CoL="6VK73" authority="Szelag &amp; Vladimirov" authorityName="Szelag &amp; Vladimirov" authorityYear="2020" box="[136,600,1684,1711]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="0" pageNumber="290" phylum="Tracheophyta" rank="species" species="bulgarica" status="sp. nov.">
<emphasis id="B95EEAA8D8341649FF61F97AC10EEA98" bold="true" box="[136,353,1684,1711]" italics="true" pageId="0" pageNumber="290">Pilosella bulgarica</emphasis>
Szeląg &amp; Vladimirov
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,
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(
<figureCitation id="13112A3FD8341649FD20F97AC35CEA99" box="[713,819,1684,1710]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[136,229,2058,2080]" captionStartId-1="2.[136,229,1044,1066]" captionStartId-2="3.[136,229,1969,1991]" captionTargetBox-0="[235,1353,602,2030]" captionTargetBox-1="[151,1435,190,1020]" captionTargetBox-2="[216,1371,207,1946]" captionTargetId-0="figure-238@1.[235,1353,602,2030]" captionTargetId-1="figure-17@2.[151,1435,190,1020]" captionTargetId-2="figure-32@3.[216,1371,207,1946]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Holotype of Pilosella bulgarica (SOM)." captionText-1="FIGURE 2. Details of the holotype of Pilosella bulgarica: rosette leaves (left) and involucres (right)." captionText-2="FIGURE 3. Pilosella bulgarica in the locus classicus, on July 21, 2018." figureDoi-0="http://doi.org/10.5281/zenodo.13875843" figureDoi-1="http://doi.org/10.5281/zenodo.13875845" figureDoi-2="http://doi.org/10.5281/zenodo.13875847" httpUri-0="https://zenodo.org/record/13875843/files/figure.png" httpUri-1="https://zenodo.org/record/13875845/files/figure.png" httpUri-2="https://zenodo.org/record/13875847/files/figure.png" pageId="0" pageNumber="290">Figs. 13</figureCitation>
)
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</paragraph>
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<emphasis id="B95EEAA8D8341649FF54F956C16EEAE5" bold="true" box="[189,257,1720,1746]" pageId="0" pageNumber="290">Type:</emphasis>
<materialsCitation id="3B423CE7D8341649FEF4F956C305EB2D" collectingDate="2018-07-21" collectionCode="Z, V, SOM, KRAM" country="Bulgaria" county="Mt. Triglav" elevation="1620" location="Central Stara Planina Mountains" municipality="Kurudere" pageId="0" pageNumber="290" specimenCount="2" typeStatus="holotype">
<collectingCountry id="F33D762AD8341649FEF4F956C1C1EAE5" box="[285,430,1720,1746]" name="Bulgaria" pageId="0" pageNumber="290">BULGARIA</collectingCountry>
.
<location id="8EF56061D8341649FE52F956C348EAE5" LSID="urn:lsid:plazi:treatment:038387ACD834164BFF61F97AC2BDE42C:8EF56061D8341649FE52F956C348EAE5" box="[443,807,1720,1746]" country="Bulgaria" county="Mt. Triglav" municipality="Kurudere" name="Central Stara Planina Mountains" pageId="0" pageNumber="290">Central Stara Planina Mountains</location>
,
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massif,
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gorge, subalpine grasslands and pastures on calcareous slopes, along a tourist path to
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a.s.l.
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,
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,
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. Vladimirov
</emphasis>
(
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<collectionCode id="ED3BAE7FD8341649FF13F8EEC155EB2D" box="[250,314,1792,1818]" country="Bulgaria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12731" lsid="urn:lsid:biocol.org:col:12731" name="Bulgarian Academy of Sciences" pageId="0" pageNumber="290" type="Herbarium">SOM</collectionCode>
;
<typeStatus id="54918818D8341649FEAFF8EEC1CDEB2D" box="[326,418,1792,1818]" pageId="0" pageNumber="290" type="isotype">isotypes</typeStatus>
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,
<collectionCode id="ED3BAE7FD8341649FE1AF8EEC225EB2D" box="[499,586,1792,1818]" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15714" lsid="urn:lsid:biocol.org:col:15714" name="Polish Academy of Sciences" pageId="0" pageNumber="290" type="Herbarium">KRAM</collectionCode>
, Herb. Hierac.
<collectionCode id="ED3BAE7FD8341649FD1DF8EFC367EB2C" box="[756,776,1793,1819]" country="Switzerland" name="Universität Zürich" pageId="0" pageNumber="290" type="Herbarium">Z</collectionCode>
. Szeląg)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3306531D834164BFF54F8CAC448E844" lastPageId="2" lastPageNumber="292" pageId="0" pageNumber="290" type="description">
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<emphasis id="B95EEAA8D8341649FF54F8CAC13EEB09" bold="true" box="[189,337,1828,1854]" pageId="0" pageNumber="290">Description:</emphasis>
―Phyllopodous. Stem
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high, in the lower third glabrous or with few pale simple hairs
<quantity id="4CD29B5FD8341649FA74F8CAC0DBEB54" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="290" unit="mm" value="3.0">3 mm</quantity>
long; in middle part with scattered microglands and few stellate hairs; within synflorescence with scattered dark-based simple hairs
<quantity id="4CD29B5FD8341649FE8BF882C1AEEBB0" box="[354,449,1900,1927]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="0" pageNumber="290" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, sparse to numerous microglands and numerous stellate hairs. Synflorescence with 35 erect capitula (and usually 12 capitula aborted). Synflorescence branches 13, grouped on top of the stem, up to
<quantity id="4CD29B5FD8341649FF4FF85AC0B0EBF8" box="[166,223,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="290" unit="cm" value="2.0">2 cm</quantity>
long, with 12 capitula. Acladium up to
<quantity id="4CD29B5FD8341649FD5FF85AC280EBF8" box="[694,751,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="290" unit="cm" value="1.0">1 cm</quantity>
long. Rosette leaves 512, up to
<quantity id="4CD29B5FD8341649FB87F85AC4C8EBF8" box="[1134,1191,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="0" pageNumber="290" unit="cm" value="4.0">4 cm</quantity>
long and up to
<quantity id="4CD29B5FD8341649FAB7F85AC5C4EBF8" box="[1374,1451,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="290" unit="cm" value="0.5">0.5 cm</quantity>
wide, sessile, lanceolate to oblanceolate, entire, acute at apex (the outer leaves shorter and rounded at apex); on upper surface glabrous or with sparse pale brown-based simple hairs
<quantity id="4CD29B5FD8341649FCA4F812C3C5E420" box="[845,938,2044,2071]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="0" pageNumber="290" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
long; on lower surface with protruding white midrib, glabrous, only on midrib and on margins with scattered to numerous pale brown-based simple hairs 46(8) mm long; on margins also with few microglands and occasionally with stellate hairs. Cauline leaves 12, up to
<quantity id="4CD29B5FD8351648FA9AFF2CC5C4ECEA" box="[1395,1451,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="291" unit="cm" value="2.0">2 cm</quantity>
long, lanceolate or subulate, sessile, rapidly reduced in size upwards, acute at apex, covered by indumentum like the rosette leaves. Peduncles with scattered to numerous pale, black in the lower third, simple hairs 13(4) mm long, scattered to numerous stellate hairs, and few to numerous blackish glandular hairs
<quantity id="4CD29B5FD8351648FBC4FEC0C4DAED7E" box="[1069,1205,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="291" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
long. Bracteoles 12, blackish green,
<quantity id="4CD29B5FD8351648FEDFFEBCC1FDED5A" box="[310,402,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="291" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, with dark-based simple hairs and stellate hairs. Involucres
<quantity id="4CD29B5FD8351648FBB5FEBCC4D7ED5A" box="[1116,1208,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="291" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
long, globose at base, with quite dense indumentum. Involucral bracts in two rows, acute at apex, up to
<quantity id="4CD29B5FD8351648FBFAFE98C406EDA6" box="[1043,1129,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="291" unit="mm" value="0.7">0.7 mm</quantity>
wide at base, blackish-green (the inner bracts with pale margins and far less dense indumentum), with numerous to dense pale dark-based simple hairs
<quantity id="4CD29B5FD8351648FF2CFE50C159EDEE" box="[197,310,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="291" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 mm</quantity>
long, scattered dark glandular hairs
<quantity id="4CD29B5FD8351648FD22FE50C33DEDEE" box="[715,850,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="291" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 mm</quantity>
long (ratio of simple hairs to glandular hairs 3:1), and sparse stellate hairs (more numerous at base of outer bracts). Ligules yellow, a little bit blackish at apex. Styles pure yellow. Achenes black,
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long. Pappus white. Pollen in anthers quite numerous, spherical and of varying size. Flowering: July.
</paragraph>
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<emphasis id="B95EEAA8D8351648FF61F7E4C091E428" bold="true" box="[136,254,2058,2080]" pageId="1" pageNumber="291">FIGURE 1.</emphasis>
Holotype of
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<emphasis id="B95EEAA8D8351648FE94F7E4C25AE428" box="[381,565,2058,2079]" italics="true" pageId="1" pageNumber="291">Pilosella bulgarica</emphasis>
(SOM)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF556632D836164BFF61FBFAC436E81D" ID-DOI="http://doi.org/10.5281/zenodo.13875845" ID-Zenodo-Dep="13875845" box="[136,1113,1044,1066]" httpUri="https://zenodo.org/record/13875845/files/figure.png" pageId="2" pageNumber="292" startId="2.[136,229,1044,1066]" targetBox="[151,1435,190,1020]" targetPageId="2" targetType="figure">
<paragraph id="8B9536BAD836164BFF61FBFAC436E81D" blockId="2.[136,1113,1044,1066]" box="[136,1113,1044,1066]" pageId="2" pageNumber="292">
<emphasis id="B95EEAA8D836164BFF61FBFAC091E81E" bold="true" box="[136,254,1044,1066]" pageId="2" pageNumber="292">FIGURE 2.</emphasis>
Details of the holotype of
<taxonomicName id="4C2A4D39D836164BFE16FBFAC2D8E81E" box="[511,695,1044,1065]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="bulgarica">
<emphasis id="B95EEAA8D836164BFE16FBFAC2D8E81E" box="[511,695,1044,1065]" italics="true" pageId="2" pageNumber="292">Pilosella bulgarica</emphasis>
</taxonomicName>
: rosette leaves (left) and involucres (right).
</paragraph>
</caption>
<paragraph id="8B9536BAD836164BFF54FBB6C448E844" blockId="2.[136,1452,1112,2075]" box="[189,1063,1112,1139]" pageId="2" pageNumber="292">
<emphasis id="B95EEAA8D836164BFF54FBB6C2E1E845" bold="true" box="[189,654,1112,1139]" pageId="2" pageNumber="292">Ploidy level and mode of reproduction:</emphasis>
―2n ~ 5x ~ 45, probably apomictic.
</paragraph>
</subSubSection>
<subSubSection id="C3306531D836164BFF54FB92C2BDE42C" pageId="2" pageNumber="292" type="distribution">
<paragraph id="8B9536BAD836164BFF54FB92C4F9E910" blockId="2.[136,1452,1112,2075]" pageId="2" pageNumber="292">
<emphasis id="B95EEAA8D836164BFF54FB92C197E8A1" bold="true" box="[189,504,1148,1175]" pageId="2" pageNumber="292">Distribution and habitats:</emphasis>
―Endemic to the Triglav massif in the Central Stara Planina Mountains,
<collectingCountry id="F33D762AD836164BFAAAFB93C5C8E8A0" box="[1347,1447,1149,1175]" name="Bulgaria" pageId="2" pageNumber="292">Bulgaria</collectingCountry>
; additional localities in the Kaloferska Planina Mountains are expected. The populations of
<taxonomicName id="4C2A4D39D836164BFB27FB4FC5C4E88C" box="[1230,1451,1185,1211]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="bulgarica">
<emphasis id="B95EEAA8D836164BFB27FB4FC5C4E88C" box="[1230,1451,1185,1211]" italics="true" pageId="2" pageNumber="292">Pilosella bulgarica</emphasis>
</taxonomicName>
comprised hundreds of plants spread along a tourist path from the Tazha (Тъжа) village to Mt. Triglav in the upper part of the Kurudere gorge, at
<quantity id="4CD29B5FD836164BFE47FB06C225E934" box="[430,586,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.585" metricValueMax="1.65" metricValueMin="1.52" pageId="2" pageNumber="292" unit="m" value="1585.0" valueMax="1650.0" valueMin="1520.0">15201650 m</quantity>
a.s.l. They were growing in subalpine grasslands and pastures on southeastfacing calcareous slopes together with
<taxonomicName id="4C2A4D39D836164BFDD2FAE3C2AEE910" authorityName="Szelag" authorityYear="2008" baseAuthorityName="Grisebach" box="[571,705,1293,1319]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="rhodopea">
<emphasis id="B95EEAA8D836164BFDD2FAE3C2AEE910" box="[571,705,1293,1319]" italics="true" pageId="2" pageNumber="292">P. rhodopea</emphasis>
</taxonomicName>
, which is frequent in the Kurudere gorge.
</paragraph>
<paragraph id="8B9536BAD836164BFF54FADEC181E9C8" blockId="2.[136,1452,1112,2075]" pageId="2" pageNumber="292">
<emphasis id="B95EEAA8D836164BFF54FADEC14BE97D" bold="true" box="[189,292,1328,1354]" pageId="2" pageNumber="292">Affinity:</emphasis>
<taxonomicName id="4C2A4D39D836164BFEA9FADFC277E97C" box="[320,536,1329,1355]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="bulgarica">
<emphasis id="B95EEAA8D836164BFEA9FADFC277E97C" box="[320,536,1329,1355]" italics="true" pageId="2" pageNumber="292">Pilosella bulgarica</emphasis>
</taxonomicName>
shows a combination of morphological characters of
<taxonomicName id="4C2A4D39D836164BFB9BFADFC497E97C" authorityName="Szelag" authorityYear="2008" baseAuthorityName="Grisebach" box="[1138,1272,1329,1355]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="rhodopea">
<emphasis id="B95EEAA8D836164BFB9BFADFC497E97C" box="[1138,1272,1329,1355]" italics="true" pageId="2" pageNumber="292">P. rhodopea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D39D836164BFAC6FADFC5CBE97C" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[1327,1444,1329,1355]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFAC6FADFC5CBE97C" box="[1327,1444,1329,1355]" italics="true" pageId="2" pageNumber="292">P. pavichii</emphasis>
</taxonomicName>
. The characters from
<taxonomicName id="4C2A4D39D836164BFE9CFABBC192E958" authorityName="Szelag" authorityYear="2008" baseAuthorityName="Grisebach" box="[373,509,1365,1391]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="rhodopea">
<emphasis id="B95EEAA8D836164BFE9CFABBC192E958" box="[373,509,1365,1391]" italics="true" pageId="2" pageNumber="292">P. rhodopea</emphasis>
</taxonomicName>
, its supposed maternal parent, are (
<emphasis id="B95EEAA8D836164BFC7DFABBC3CDE958" box="[916,930,1365,1391]" italics="true" pageId="2" pageNumber="292">1</emphasis>
) the few-headed synflorescence with capitula grouped at the top of the stem, (
<emphasis id="B95EEAA8D836164BFE10FA97C268E9A4" box="[505,519,1401,1427]" italics="true" pageId="2" pageNumber="292">2</emphasis>
) the reduced cauline leaves, (
<emphasis id="B95EEAA8D836164BFCB5FA97C305E9A4" box="[860,874,1401,1427]" italics="true" pageId="2" pageNumber="292">3</emphasis>
) the numerous long simple hairs on the involucral bracts, and (
<emphasis id="B95EEAA8D836164BFEF1FA73C149E980" box="[280,294,1437,1463]" italics="true" pageId="2" pageNumber="292">4</emphasis>
) the stellate hairs on the stem. The characters from
<taxonomicName id="4C2A4D39D836164BFC61FA73C46EE980" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[904,1025,1437,1463]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFC61FA73C46EE980" box="[904,1025,1437,1463]" italics="true" pageId="2" pageNumber="292">P. pavichii</emphasis>
</taxonomicName>
are (
<emphasis id="B95EEAA8D836164BFBA8FA73C420E980" box="[1089,1103,1437,1463]" italics="true" pageId="2" pageNumber="292">1</emphasis>
) the smaller capitula with the turbinate involucres, (
<emphasis id="B95EEAA8D836164BFE68FA2FC1E0E9EC" box="[385,399,1473,1499]" italics="true" pageId="2" pageNumber="292">2</emphasis>
) the rosette leaves acute at apex, on lower surface with protruding white midrib, and (
<emphasis id="B95EEAA8D836164BFA80FA2FC518E9EC" box="[1385,1399,1473,1499]" italics="true" pageId="2" pageNumber="292">3</emphasis>
) the sparse indumentum of the stem.
</paragraph>
<paragraph id="8B9536BAD836164BFF54F9E7C080EAB8" blockId="2.[136,1452,1112,2075]" pageId="2" pageNumber="292">
<taxonomicName id="4C2A4D39D836164BFF54F9E7C1F9EA14" authorityName="Szelag" authorityYear="2008" baseAuthorityName="Grisebach" box="[189,406,1545,1571]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="rhodopea">
<emphasis id="B95EEAA8D836164BFF54F9E7C1F9EA14" box="[189,406,1545,1571]" italics="true" pageId="2" pageNumber="292">Pilosella rhodopea</emphasis>
</taxonomicName>
is a common species in the subalpine habitats of the Central Stara Planina Mountains; its population above the Tazha hut (хижа Тъжа), separated by
<emphasis id="B95EEAA8D836164BFCF2F9C3C35AEA71" box="[795,821,1581,1606]" italics="true" pageId="2" pageNumber="292">ca</emphasis>
<quantity id="4CD29B5FD836164BFCD5F9C3C3EDEA70" box="[828,898,1581,1607]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="292" unit="km" value="10.0">10 km</quantity>
from the Kurudere gorge, is composed of diploid, triploid and tetraploid plants (
<bibRefCitation id="EFBB4B4BD836164BFE3BF9BCC2BAEA5C" author="Singliarova, B. &amp; Hodalova, I. &amp; Mraz, P." box="[466,725,1617,1645]" pageId="2" pageNumber="292" pagination="450 - 470" refId="ref2238" refString="Singliarova, B., Hodalova, I. &amp; Mraz, P. (2011) Biosystematic study of the diploid-polyploid Pilosella alpicola group with variation in breeding system: Patterns and processes. Taxon 60: 450 - 470. https: // doi. org / 10.1002 / tax. 602014" type="journal article" year="2011">
Šingliarová
<emphasis id="B95EEAA8D836164BFDB2F9BFC2E3EA5C" box="[603,652,1617,1643]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. 2011
</bibRefCitation>
). Cytogenetic studies of
<taxonomicName id="4C2A4D39D836164BFC0FF9BFC403EA5C" authorityName="Szelag" authorityYear="2008" baseAuthorityName="Grisebach" box="[998,1132,1617,1643]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="rhodopea">
<emphasis id="B95EEAA8D836164BFC0FF9BFC403EA5C" box="[998,1132,1617,1643]" italics="true" pageId="2" pageNumber="292">P. rhodopea</emphasis>
</taxonomicName>
in the Kurudere gorge are in progress.
</paragraph>
<paragraph id="8B9536BAD836164BFF54F977C44AEB98" blockId="2.[136,1452,1112,2075]" pageId="2" pageNumber="292">
<taxonomicName id="4C2A4D39D836164BFF54F977C1EEEA84" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[189,385,1689,1715]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFF54F977C1EEEA84" box="[189,385,1689,1715]" italics="true" pageId="2" pageNumber="292">Pilosella pavichii</emphasis>
</taxonomicName>
, the second putative parental species of
<taxonomicName id="4C2A4D39D836164BFCD2F977C3ACEA84" box="[827,963,1689,1715]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="bulgarica">
<emphasis id="B95EEAA8D836164BFCD2F977C3ACEA84" box="[827,963,1689,1715]" italics="true" pageId="2" pageNumber="292">P. bulgarica</emphasis>
</taxonomicName>
, grows in the subalpine belt and in clearings in the mountain forest; it is known as diploid in
<collectingCountry id="F33D762AD836164BFD70F953C293EAE0" box="[665,764,1725,1751]" name="Bulgaria" pageId="2" pageNumber="292">Bulgaria</collectingCountry>
,
<collectingCountry id="F33D762AD836164BFCEEF953C336EAE0" box="[775,857,1725,1751]" name="Greece" pageId="2" pageNumber="292">Greece</collectingCountry>
,
<collectingCountry id="F33D762AD836164BFC8AF953C380EAE0" box="[867,1007,1725,1751]" name="Serbia" pageId="2" pageNumber="292">Montenegro</collectingCountry>
, North
<collectingCountry id="F33D762AD836164BFBA8F953C4AFEAE0" box="[1089,1216,1725,1751]" name="Macedonia" pageId="2" pageNumber="292">Macedonia</collectingCountry>
,
<collectingCountry id="F33D762AD836164BFB23F953C540EAE0" box="[1226,1327,1725,1751]" name="Romania" pageId="2" pageNumber="292">Romania</collectingCountry>
and
<collectingCountry id="F33D762AD836164BFA8DF953C5C3EAE0" box="[1380,1452,1725,1751]" name="Serbia" pageId="2" pageNumber="292">Serbia</collectingCountry>
(
<bibRefCitation id="EFBB4B4BD836164BFF79F90FC1CFEACC" author="Christov, M. &amp; Popov, A." box="[144,416,1761,1787]" pageId="2" pageNumber="292" pagination="440 - 447" refId="ref1642" refString="Christov, M. &amp; Popov, A. (1933) Cytologische Studien uber die Gattung Hieracium. Planta 20: 440 - 447. https: // doi. org / 10.1007 / BF 01910640" type="journal article" year="1933">Christov &amp; Popov 1933</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFE42F90FC2FBEACC" author="Strid, A &amp; Frazen, R." box="[427,660,1761,1787]" pageId="2" pageNumber="292" pagination="829 - 861" refId="ref1942" refString="Strid, A, &amp; Frazen, R. (1981) Report on Hieracium pavichii. In: Love, A. (Ed.) Chromosome number reports LXXIII. Taxon 30: 829 - 861. https: // doi. org / 10.1002 / j. 1996 - 8175.1981. tb 04309. x" type="journal article" year="1981">Strid &amp; Frazén 1981</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFD49F90FC30BEACC" author="Vladimirov, V." box="[672,868,1761,1787]" pageId="2" pageNumber="292" refId="ref2339" refString="Vladimirov, V. (2000) Diploid species of the genus Hieracium s. l. in Bulgaria. Abhandlungen und Berichte des Naturkundemuseums Gorlitz 72 (Suppl.): 16." type="journal volume" year="2000">Vladimirov 2000</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFC86F90FC433EACC" author="Mraz, P. &amp; Szelag, Z." box="[879,1116,1761,1787]" pageId="2" pageNumber="292" pagination="405 - 414" refId="ref1901" refString="Mraz, P. &amp; Szelag, Z. (2004) Chromosome numbers and reproductive systems in selected species of the genera Hieracium and Pilosella (Asteraceae) from Romania. Annales Botanici Fennici 41: 405 - 414." type="journal article" year="2004">Mráz &amp; Szeląg 2004</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFB8CF90FC0ABEB29" author="Yurukova-Grancharova, P. &amp; Robeva-Davidova, P. &amp; Vladimirov, V." pageId="2" pageNumber="292" pagination="668 - 675" refId="ref2373" refString="Yurukova-Grancharova, P., Robeva-Davidova, P. &amp; Vladimirov, V. (2006) On the embryology and mode of reproduction of selected diploid species of Hieracium s. l. (Asteraceae) from Bulgaria. Flora 201: 668 - 675. https: // doi. org / 10.1016 / j. flora. 2006.01.003" type="journal article" year="2006">
Yurukova-Grancharova
<emphasis id="B95EEAA8D836164BFA9AF90FC5CAEACC" box="[1395,1445,1761,1787]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFF27F8EBC1F6EB28" author="Szelag, Z. &amp; Ilnicki, T. &amp; Niketic, M. &amp; Tomovic, G." box="[206,409,1797,1823]" pageId="2" pageNumber="292" pagination="119 - 121" refId="ref2130" refString="Szelag, Z., Ilnicki, T., Niketic, M. &amp; Tomovic, G. (2007) Diploid chromosome numbers in five Hieracium species from Serbia and Montenegro. Acta Biologica Cracoviensia Series Botanica 49 (1): 119 - 121." type="journal article" year="2007">
Szeląg
<emphasis id="B95EEAA8D836164BFEC9F8EBC13FEB28" box="[288,336,1797,1823]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFBB4B4BD836164BFE4DF8EBC2F2EB28" author="Szelag, Z. &amp; Ilnicki, T." box="[420,669,1797,1823]" pageId="2" pageNumber="292" pagination="124 - 126" refId="ref2072" refString="Szelag, Z. &amp; Ilnicki, T. (2011) Diploid chromosome numbers in Hieracium and Pilosella (Asteraceae) from Macedonia and Montenegro. Acta Biologica Cracoviensia Series Botanica 53 (2): 124 - 126. https: // doi. org / 10.2478 / v 10182 - 011 - 0023 - 2" type="journal article" year="2011">Szeląg &amp; Ilnicki 2011</bibRefCitation>
). Recent studies have revealed also a tetraploid cytotype of
<taxonomicName id="4C2A4D39D836164BFADFF8EBC5C4EB28" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[1334,1451,1797,1823]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFADFF8EBC5C4EB28" box="[1334,1451,1797,1823]" italics="true" pageId="2" pageNumber="292">P. pavichii</emphasis>
</taxonomicName>
growing together with diploid plants in the Pirin Mountains in
<collectingCountry id="F33D762AD836164BFCD5F8C7C3F3EB74" box="[828,924,1833,1859]" name="Bulgaria" pageId="2" pageNumber="292">Bulgaria</collectingCountry>
(
<bibRefCitation id="EFBB4B4BD836164BFC43F8C7C4C3EB74" author="Krahulcova, A. &amp; Vladimirov, V. &amp; Krahulec, F. &amp; Brautigam, S." box="[938,1196,1833,1859]" pageId="2" pageNumber="292" pagination="377 - 384" refId="ref1735" refString="Krahulcova, A., Vladimirov, V., Krahulec, F. &amp; Brautigam, S. (2009) The agamic complex of Pilosella (Asteraceae) in Bulgaria and SW Romania: variation in ploidy level and breeding systems. Phytologia Balcanica 15: 377 - 384." type="journal article" year="2009">
Krahulcová
<emphasis id="B95EEAA8D836164BFBDDF8C7C40BEB74" box="[1076,1124,1833,1859]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. 2009
</bibRefCitation>
,
<bibRefCitation id="EFBB4B4BD836164BFB5FF8C6C49CEB75" author="Krahulcova, A. &amp; Vladimirov, V. &amp; Krahulec, F. &amp; Brautigam, S." box="[1206,1267,1832,1858]" pageId="2" pageNumber="292" pagination="39 - 62" refId="ref1787" refString="Krahulcova, A., Vladimirov, V., Krahulec, F. &amp; Brautigam, S. (2016) The agamic complex of Pilosella (Asteraceae) in Bulgaria and SW Romania: variation in ploidy levels and breeding systems. Part 2. Phytologia Balcanica 22: 39 - 62." type="journal article" year="2016">2016</bibRefCitation>
). The pentaploid and hexaploid cytotypes of
<taxonomicName id="4C2A4D39D836164BFE28F8A3C258EB50" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[449,567,1869,1895]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFE28F8A3C258EB50" box="[449,567,1869,1895]" italics="true" pageId="2" pageNumber="292">P. pavichii</emphasis>
</taxonomicName>
published by
<bibRefCitation id="EFBB4B4BD836164BFD31F8A3C381EB51" author="Krahulcova, A. &amp; Vladimirov, V. &amp; Krahulec, F. &amp; Brautigam, S." box="[728,1006,1868,1895]" pageId="2" pageNumber="292" pagination="39 - 62" refId="ref1787" refString="Krahulcova, A., Vladimirov, V., Krahulec, F. &amp; Brautigam, S. (2016) The agamic complex of Pilosella (Asteraceae) in Bulgaria and SW Romania: variation in ploidy levels and breeding systems. Part 2. Phytologia Balcanica 22: 39 - 62." type="journal article" year="2016">
Krahulcová
<emphasis id="B95EEAA8D836164BFC8AF8A3C3FAEB50" box="[867,917,1869,1895]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. (2016)
</bibRefCitation>
are a bit surprising and may have been based on misidentified specimens of hybrids between
<taxonomicName id="4C2A4D39D836164BFD09F89FC33AEBBC" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[736,853,1905,1931]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFD09F89FC33AEBBC" box="[736,853,1905,1931]" italics="true" pageId="2" pageNumber="292">P. pavichii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D39D836164BFC63F89FC45CEBBC" box="[906,1075,1905,1931]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="piloselloides">
<emphasis id="B95EEAA8D836164BFC63F89FC45CEBBC" box="[906,1075,1905,1931]" italics="true" pageId="2" pageNumber="292">P. piloselloides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D57D3D836164BFBD0F89FC419EBBC" box="[1081,1142,1905,1931]" pageId="2" pageNumber="292" sensu="lato">s. lat.</taxonomicNameLabel>
, which are quite frequent in the anthropogenic disturbed habitats at lower altitudes in the Bulgarian mountains.
</paragraph>
<paragraph id="8B9536BAD836164BFF54F857C2BDE42C" blockId="2.[136,1452,1112,2075]" pageId="2" pageNumber="292">
<emphasis id="B95EEAA8D836164BFF54F857C090EBE4" bold="true" box="[189,255,1977,2003]" pageId="2" pageNumber="292">Note:</emphasis>
<taxonomicName id="4C2A4D39D836164BFEF2F857C18DEBE4" authorityName="Holub" authorityYear="1977" baseAuthorityName="Heuffel" box="[283,482,1977,2003]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFEF2F857C18DEBE4" box="[283,482,1977,2003]" italics="true" pageId="2" pageNumber="292">Pilosella pavichii</emphasis>
</taxonomicName>
, originally described as
<taxonomicName id="4C2A4D39D836164BFCE8F857C4A1EBE5" authority="Heuffel (1853: 618)" authorityName="Heuffel" authorityPageNumber="618" authorityYear="1853" box="[769,1230,1976,2003]" class="Magnoliopsida" family="Asteraceae" genus="Hieracium" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="species" species="pavichii">
<emphasis id="B95EEAA8D836164BFCE8F857C3B3EBE4" box="[769,988,1977,2003]" italics="true" pageId="2" pageNumber="292">Hieracium pavichii</emphasis>
<bibRefCitation id="EFBB4B4BD836164BFC0EF857C4A1EBE5" author="Heuffel, J." box="[999,1230,1976,2003]" pageId="2" pageNumber="292" pagination="617 - 627" refId="ref1679" refString="Heuffel, J. (1853) Sertum plantarum novarum aut minus rite cognitarum. Flora 36: 617 - 627." type="journal article" year="1853">Heuffel (1853: 618)</bibRefCitation>
</taxonomicName>
, was transferred to the genus
<taxonomicName id="4C2A4D39D836164BFF12F833C226EBC0" authority="Vaillant (1754: 703)" authorityName="Vaillant" authorityPageNumber="703" authorityYear="1754" box="[251,585,2012,2039]" class="Magnoliopsida" family="Asteraceae" genus="Pilosella" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="292" phylum="Tracheophyta" rank="genus">
<emphasis id="B95EEAA8D836164BFF12F833C130EBC0" box="[251,351,2013,2039]" italics="true" pageId="2" pageNumber="292">Pilosella</emphasis>
<bibRefCitation id="EFBB4B4BD836164BFE8FF833C226EBC0" author="Vaillant, S." box="[358,585,2012,2039]" pageId="2" pageNumber="292" pagination="697 - 750" refId="ref2292" refString="Vaillant, S. (1754) Fortsetzung der neuen Kennzeichen der Pflanzen mit zusammengesetzten Blumen. Dritte Classe. Von den Wegwartartigen (Cichoraceis). Der Koniglichen Akademie der Wissenschaften in Paris. Anatomische, Chymische und Botanische Abhandlungen 5: 697 - 750." type="journal article" year="1754">Vaillant (1754: 703)</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFBB4B4BD836164BFD9CF833C328EBC1" author="Holub, J." box="[629,839,2012,2039]" pageId="2" pageNumber="292" pagination="293 - 311" refId="ref1701" refString="Holub, J. (1977) New names in Phanerogamae 5. Folia Geobotanica et Phytotaxonomica 12: 293 - 311. https: // doi. org / 10.1007 / BF 02855880" type="journal article" year="1977">Holub (1977: 306)</bibRefCitation>
(see Szeląg 2011), and not by
<bibRefCitation id="EFBB4B4BD836164BFB4AF833C5E3EBC0" author="Arvet-Touvet, C." box="[1187,1420,2013,2039]" pageId="2" pageNumber="292" pagination="278 - 292" refId="ref1596" refString="Arvet-Touvet, C. (1880) Essai de classification sur les genres Pilosella et Hieracium principalement. Pour les especes et les formes de la region Sud-Quest de l'Europe. Bulletin de la Societe Dauphinoise pour l'Echange de Plantes 7: 278 - 292." type="journal article" year="1880">Arvet-Touvet (1880)</bibRefCitation>
as considered by
<bibRefCitation id="EFBB4B4BD836164BFEC2F7EFC257E42C" author="Krahulcova, A. &amp; Vladimirov, V. &amp; Krahulec, F. &amp; Brautigam, S." box="[299,568,2049,2075]" pageId="2" pageNumber="292" pagination="377 - 384" refId="ref1735" refString="Krahulcova, A., Vladimirov, V., Krahulec, F. &amp; Brautigam, S. (2009) The agamic complex of Pilosella (Asteraceae) in Bulgaria and SW Romania: variation in ploidy level and breeding systems. Phytologia Balcanica 15: 377 - 384." type="journal article" year="2009">
Krahulcová
<emphasis id="B95EEAA8D836164BFE5CF7EFC189E42C" box="[437,486,2049,2075]" italics="true" pageId="2" pageNumber="292">et al</emphasis>
. (2009
</bibRefCitation>
,
<bibRefCitation id="EFBB4B4BD836164BFDAAF7EEC211E42D" author="Krahulcova, A. &amp; Vladimirov, V. &amp; Krahulec, F. &amp; Brautigam, S." box="[579,638,2048,2074]" pageId="2" pageNumber="292" pagination="39 - 62" refId="ref1787" refString="Krahulcova, A., Vladimirov, V., Krahulec, F. &amp; Brautigam, S. (2016) The agamic complex of Pilosella (Asteraceae) in Bulgaria and SW Romania: variation in ploidy levels and breeding systems. Part 2. Phytologia Balcanica 22: 39 - 62." type="journal article" year="2016">2016</bibRefCitation>
,
<bibRefCitation id="EFBB4B4BD836164BFD60F7EFC2A9E42C" author="Krahulcova, A. &amp; Krahulec, F. &amp; Vladimirov, V." box="[649,710,2049,2075]" pageId="2" pageNumber="292" pagination="345 - 362" refId="ref1842" refString="Krahulcova, A., Krahulec, F. &amp; Vladimirov, V. (2018) Impact of interspecific hybridization within a polyploid agamic complex of Pilosella (Asteraceae, Cichorieae) in Bulgaria compared with Central Europe. Willdenowia 48: 345 - 362. https: // doi. org / 10.3372 / wi. 48.48302" type="journal article" year="2018">2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,193 @@
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<mods:title id="88135DCA44722628A895602295AA6AEC">Multi-locus phylogeny reveals Phaeodothis mori sp. nov. (Didymosphaeriaceae, Pleosporales) from dead leaves of Morus australis</mods:title>
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<mods:affiliation id="09E1263B9B72987D4C33E0DEE2FD186F">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan. &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China.</mods:affiliation>
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<mods:affiliation id="DAC9812CEC1EA9C5FF7FDF8011D73C87">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan.</mods:affiliation>
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<mods:affiliation id="2B4D2E0A6FB7B1249B3DEB9BE0F21169">School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China. &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand.</mods:affiliation>
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<treatment id="038C043AF96A8304FF3C64C3FE36CA7B" LSID="urn:lsid:plazi:treatment:038C043AF96A8304FF3C64C3FE36CA7B" httpUri="http://treatment.plazi.org/id/038C043AF96A8304FF3C64C3FE36CA7B" lastPageId="8" lastPageNumber="248" pageId="7" pageNumber="248">
<subSubSection id="C33FE6A7F96A830BFF3C64C3FC29CEEF" pageId="7" pageNumber="248" type="nomenclature">
<paragraph id="8B9AB52CF96A830BFF3C64C3FC22CEC2" blockId="7.[136,932,1466,1493]" box="[136,932,1466,1493]" pageId="7" pageNumber="248">
<heading id="D0D20240F96A830BFF3C64C3FC22CEC2" box="[136,932,1466,1493]" fontSize="11" level="2" pageId="7" pageNumber="248" reason="3">
<taxonomicName id="4C25CEAFF96A830BFF3C64C3FD16CEC2" ID-CoL="4FMYY" authority="Syd. &amp; P. Syd., Annls" authorityName="Syd. &amp; P. Syd., Annls" box="[136,656,1466,1493]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFF3C64C3FE10CEC2" bold="true" box="[136,406,1466,1493]" italics="true" pageId="7" pageNumber="248">Phaeodothis tricuspidis</emphasis>
Syd. &amp; P. Syd., Annls
</taxonomicName>
mycol. 2(2): 166 (1904)
</heading>
</paragraph>
<paragraph id="8B9AB52CF96A830BFF3C649BFC29CEEF" blockId="7.[136,943,1506,1528]" box="[136,943,1506,1528]" pageId="7" pageNumber="248">
Index Fungorum Number: IF 185110; Facesoffungi number: FoF 00060,
<figureCitation id="131EA9A9F96A830BFCF3649AFC29CEEF" box="[839,943,1507,1528]" captionStart="FIGURE 3" captionStartId="8.[136,229,1439,1461]" captionTargetBox="[154,1433,389,1408]" captionTargetId="figure-114@8.[151,1436,386,1411]" captionTargetPageId="8" captionText="FIGURE 3. Phaeodothis tricuspidis (SF125876, holotype) a. Herbarium packet and specimen. b. Ascomata on host. c. Close up of ascoma. d, e. Sections through ascomata. f. Peridium comprising hyaline compressed cells. gj. Mature and immature asci surrounded by pseudoparaphyses. kn. ascospores. Scale bars: c = 1000 µm, d, e = 100 µm, f = 20 µm, gj = 30 µm, kn = 5 µm." pageId="7" pageNumber="248">FIGURE 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33FE6A7F96A8304FF3C675EFE36CA7B" lastPageId="8" lastPageNumber="249" pageId="7" pageNumber="248" type="description">
<paragraph id="8B9AB52CF96A830BFF3C675EFB47CC76" blockId="7.[136,1452,1573,2069]" pageId="7" pageNumber="248">
<emphasis id="B951693EF96A830BFF3C675EFF6ACD56" box="[136,236,1575,1601]" italics="true" pageId="7" pageNumber="248">Saprobic</emphasis>
on dead wood or grasses.
<emphasis id="B951693EF96A830BFD80675FFD5DCD56" bold="true" box="[564,731,1574,1601]" italics="true" pageId="7" pageNumber="248">Sexual morph</emphasis>
:
<emphasis id="B951693EF96A830BFD44675EFCD8CD56" box="[752,862,1575,1601]" italics="true" pageId="7" pageNumber="248">Ascomata</emphasis>
220300×120180 μm (
<emphasis id="B951693EF96A830BFB3E675CFB11CD2C" box="[1162,1175,1573,1595]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 255×137 μm, n=10), immersed to nearly superficial under clypeus, scattered to gregarious, subglobose to hemispherical, dark brown to black, papillate.
<emphasis id="B951693EF96A830BFEF16716FE11CD9E" box="[325,407,1647,1673]" italics="true" pageId="7" pageNumber="248">Papilla</emphasis>
black, with pore-like ostioles, ostiolar canal filled with periphyses.
<taxonomicName id="4C25CEAFF96A830BFB106716FA8ACD9E" box="[1188,1292,1647,1673]" class="Magnoliopsida" family="Peraceae" genus="Peridium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="genus">
<emphasis id="B951693EF96A830BFB106716FA8ACD9E" box="[1188,1292,1647,1673]" italics="true" pageId="7" pageNumber="248">Peridium</emphasis>
</taxonomicName>
1525μm (
<emphasis id="B951693EF96A830BFA2C6714FA23CD94" box="[1432,1445,1645,1667]" italics="true" pageId="7" pageNumber="248">x</emphasis>
=23μm, n=20) wide, comprising several layers of hyaline to light brown, compressed cells of
<emphasis id="B951693EF96A830BFB2367EDFADECDBA" box="[1175,1368,1683,1709]" italics="true" pageId="7" pageNumber="248">
textura
<taxonomicName id="4C25CEAFF96A830BFB5967EAFADECDBA" box="[1261,1368,1683,1709]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Paraphaeosphaeria" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="angularis">angularis</taxonomicName>
</emphasis>
, fusing and indistinguishable from the host tissues.
<emphasis id="B951693EF96A830BFD3167CEFCA4CDC6" box="[645,802,1719,1745]" italics="true" pageId="7" pageNumber="248">Hamathecium</emphasis>
of 24 μm wide (
<emphasis id="B951693EF96A830BFC4A67CCFB8DCDDC" box="[1022,1035,1717,1739]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 2.6 μm, n=20), aseptate, cellular pseudoparaphyses often embedded in a gelatinous matrix.
<emphasis id="B951693EF96A830BFCBB67A2FCB9CDE2" box="[783,831,1755,1781]" italics="true" pageId="7" pageNumber="248">Asci</emphasis>
50 80 × 712 μm (
<emphasis id="B951693EF96A830BFB9867A0FBBFCDF8" box="[1068,1081,1753,1775]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 69 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, short pedicellate, apically rounded with a minute ocular chamber.
<emphasis id="B951693EF96A830BFA996786FA2ACC0E" box="[1325,1452,1791,1817]" italics="true" pageId="7" pageNumber="248">Ascospores</emphasis>
1417 × 45μm (
<emphasis id="B951693EF96A830BFEE16658FEE4CC20" box="[341,354,1825,1847]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 15.8 × 4.5 μm, n=30), unit to biseriate and partially overlapping, fusiform with rounded ends, olivaceous brown, 1-septate, constricted at the septa, smooth-walled.
<emphasis id="B951693EF96A830BFC26663FFBC4CC76" bold="true" box="[914,1090,1862,1889]" italics="true" pageId="7" pageNumber="248">Asexual morph</emphasis>
: unknown.
</paragraph>
<paragraph id="8B9AB52CF96A830BFF096612FD49CCBE" blockId="7.[136,1452,1573,2069]" pageId="7" pageNumber="248">
Material examined:
<emphasis id="B951693EF96A830BFE2A6612FE3CCC92" bold="true" box="[414,442,1899,1925]" pageId="7" pageNumber="248"></emphasis>
<collectingCountry id="F332F5BCF96A830BFE0E6612FDE6CC92" box="[442,608,1899,1925]" name="Argentina" pageId="7" pageNumber="248">ARGENTINA</collectingCountry>
, San José,
<collectingRegion id="49E17BCEF96A830BFD5F6612FCA0CC92" box="[747,806,1899,1925]" country="Argentina" name="Salta" pageId="7" pageNumber="248">Salta</collectingRegion>
, on leaves of
<taxonomicName id="4C25CEAFF96A830BFC626615FA87CC92" authority="Griseb." authorityName="Griseb." box="[982,1281,1899,1925]" class="Magnoliopsida" family="Elaeocarpaceae" genus="Tricuspis" kingdom="Plantae" order="Oxalidales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B951693EF96A830BFC626615FB25CC92" box="[982,1187,1899,1925]" italics="true" pageId="7" pageNumber="248">Tricuspis latifolia</emphasis>
Griseb.
</taxonomicName>
(
<taxonomicName id="4C25CEAFF96A830BFAA16612FAF1CC92" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1301,1399,1899,1925]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
),
<date id="FF9B93ECF96A830BFA3B6612FEA9CCBE" pageId="7" pageNumber="248" value="1873-02-12">12 February 1873</date>
, P.G. Lorentz (SF125876,
<emphasis id="B951693EF96A830BFDEC66F6FD39CCBE" bold="true" box="[600,703,1935,1961]" pageId="7" pageNumber="248">
<typeStatus id="549E0B8EF96A830BFDEC66F6FD39CCBE" box="[600,703,1935,1961]" pageId="7" pageNumber="248" type="holotype">holotype</typeStatus>
</emphasis>
).
</paragraph>
<paragraph id="8B9AB52CF96A8304FF0966CAFE36CA7B" blockId="7.[136,1452,1573,2069]" lastBlockId="8.[136,1452,158,365]" lastPageId="8" lastPageNumber="249" pageId="7" pageNumber="248">
Notes:—
<taxonomicName id="4C25CEAFF96A830BFE9666CAFDA8CCDA" box="[290,558,1971,1997]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFE9666CAFDA8CCDA" box="[290,558,1971,1997]" italics="true" pageId="7" pageNumber="248">Phaeodothis tricuspidis</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation id="EFB4C8DDF96A830BFCA566CAFBBBCCDA" author="Sydow, H. &amp; Sydow, P." box="[785,1085,1971,1997]" pageId="7" pageNumber="248" pagination="162 - 174" refId="ref10677" refString="Sydow, H. &amp; Sydow, P. (1904) Novae fungorum species. Ann. Mycol. 2: 162 - 174." type="journal article" year="1904">Sydow and Sydow (1904)</bibRefCitation>
, from living leaves of
<taxonomicName id="4C25CEAFF96A830BFAF166CDFEB8CCE6" authority="Griseb." authorityName="Griseb." class="Magnoliopsida" family="Elaeocarpaceae" genus="Tricuspis" kingdom="Plantae" order="Oxalidales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B951693EF96A830BFAF166CDFF67CCE6" italics="true" pageId="7" pageNumber="248">Tricuspis latifolia</emphasis>
Griseb.
</taxonomicName>
(
<taxonomicName id="4C25CEAFF96A830BFEFB66AEFE28CCE6" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[335,430,2007,2033]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) and they assumed it to be very close to those of
<taxonomicName id="4C25CEAFF96A830BFBB666AEFBFCCCE6" box="[1026,1146,2007,2033]" class="Dothideomycetes" family="Polystomellaceae" genus="Dothidella" kingdom="Fungi" order="Asterinales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="genus">
<emphasis id="B951693EF96A830BFBB666AEFBFCCCE6" box="[1026,1146,2007,2033]" italics="true" pageId="7" pageNumber="248">Dothidella</emphasis>
</taxonomicName>
, but differ in having dark ascospores (
<bibRefCitation id="EFB4C8DDF96A830BFEA76682FDAAC302" author="Malloch, D. &amp; Mallik, A." box="[275,556,2043,2069]" pageId="7" pageNumber="248" pagination="1265 - 1275" refId="ref8890" refString="Malloch, D. &amp; Mallik, A. (1998) Taxonomy of Orphnodactylis kalmiae gen. et sp. nov. influenced by the hyperparasite Didymosphaeria kalmiae. Canadian journal of botany 76: 1265 - 1275. https: // doi. org / 10.1139 / b 98 - 147" type="journal article" year="1998">Malloch &amp; Mallik 1998</bibRefCitation>
).
<taxonomicName id="4C25CEAFF96A830BFDF66682FC8BC302" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[578,781,2043,2069]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96A830BFDF66682FC8BC302" box="[578,781,2043,2069]" italics="true" pageId="7" pageNumber="248">Phaeodothis mori</emphasis>
</taxonomicName>
can be distinguished from the
<typeStatus id="549E0B8EF96A830BFBC36682FB21C302" box="[1143,1191,2043,2069]" pageId="7" pageNumber="248">type</typeStatus>
species,
<taxonomicName id="4C25CEAFF96A830BFAA76685FA2DC302" box="[1299,1451,2043,2069]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFAA76685FAA2C302" box="[1299,1316,2044,2069]" italics="true" pageId="7" pageNumber="248">P</emphasis>
.
<emphasis id="B951693EF96A830BFA816682FA2DC302" box="[1333,1451,2043,2069]" italics="true" pageId="7" pageNumber="248">tricuspidis</emphasis>
</taxonomicName>
in having smaller ascomata (100150 × 80130 μm) and yellowish brown, ascospores (10.512.5 × 34 μm) with mucilaginous sheath, whereas
<taxonomicName id="4C25CEAFF9658304FE5861BAFD03CBCB" box="[492,645,194,221]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF9658304FE5861BAFE7BCBCB" box="[492,509,195,220]" italics="true" pageId="8" pageNumber="249">P</emphasis>
.
<emphasis id="B951693EF9658304FDBB61BBFD03CBCB" box="[527,645,194,220]" italics="true" pageId="8" pageNumber="249">tricuspidis</emphasis>
</taxonomicName>
has larger ascomata (220300×120180 μm) and olivaceous brown, ascospores (1417 × 45μm) lacking a mucilaginous sheath. As well as,
<taxonomicName id="4C25CEAFF9658304FC7F619FFB53CA17" box="[971,1237,230,256]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF9658304FC7F619FFB53CA17" box="[971,1237,230,256]" italics="true" pageId="8" pageNumber="249">Phaeodothis tricuspidis</emphasis>
</taxonomicName>
has been recorded from dead grass stems (
<taxonomicName id="4C25CEAFF9658304FE2E6073FE7ACA33" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[410,508,266,292]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), which is a monocot; whereas
<taxonomicName id="4C25CEAFF9658304FCDE6072FC47CA33" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[874,961,267,293]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF9658304FCDE6072FCFDCA33" box="[874,891,267,292]" italics="true" pageId="8" pageNumber="249">P</emphasis>
.
<emphasis id="B951693EF9658304FC386072FC47CA33" box="[908,961,267,292]" italics="true" pageId="8" pageNumber="249">mori</emphasis>
</taxonomicName>
was recorded from the decaying leaves of
<taxonomicName id="4C25CEAFF9658304FF3C6056FEBCCA5F" box="[136,314,302,328]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF9658304FF3C6056FEBCCA5F" box="[136,314,302,328]" italics="true" pageId="8" pageNumber="249">Morus australis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C25CEAFF9658304FEFD6057FE38CA5F" box="[329,446,302,328]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
), which is a dicot. However, molecular data are essentially needed to clarify phylogenetic placement of
<typeStatus id="549E0B8EF9658304FE94602AFED6CA7A" box="[288,336,339,365]" pageId="8" pageNumber="249">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="BB2FA06A8BE346FECCA083D68EC209A9">Multi-locus phylogeny reveals Phaeodothis mori sp. nov. (Didymosphaeriaceae, Pleosporales) from dead leaves of Morus australis</mods:title>
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<mods:namePart id="9A59231148368199530CF7C102B7EE26">Tennakoon, Danushka S.</mods:namePart>
<mods:affiliation id="B3E56B991C2896EAD63C3662169239E2">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan. &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China.</mods:affiliation>
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<mods:namePart id="537D0E42B580E8958A297C058A4A6776">Kuo, Chang-Hsin</mods:namePart>
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<mods:namePart id="6A43DF556854638399146D6EA4DD7F41">Hyde, Kevin D.</mods:namePart>
<mods:affiliation id="D3FC6C486BEE3AEFE90ED051B6315642">School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China. &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand.</mods:affiliation>
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<mods:title id="568F302F704548E686ADEC51E23E9FCE">Phytotaxa</mods:title>
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<mods:part id="2D3E212D6C4106AAFFCFCD46FC02AF26">
<mods:date id="1518E49A1ACDC21071C4FD8F3EED1AFF">2020</mods:date>
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<mods:number id="4E03F198ABC5D367FDFED979706F2BE9">2020-01-14</mods:number>
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<mods:number id="6F3AEAE9E731A03101C114D7A0005266">428</mods:number>
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<mods:number id="E09CDC6BBFE93057DCDE36402111F7FA">3</mods:number>
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<mods:url id="053F2DCA39A57C3D5138363FB8FC43E5">http://dx.doi.org/10.11646/phytotaxa.428.3.5</mods:url>
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<taxonomicName id="4C25CEAFF96B830AFF3C636DFD97C939" ID-CoL="6MGJ" ID-ENA="100039" authority="Syd. &amp; P. Syd., Annls" authorityName="Syd. &amp; P. Syd., Annls" box="[136,529,532,559]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="genus">
<emphasis id="B951693EF96B830AFF3C636DFE91C938" bold="true" box="[136,279,532,559]" italics="true" pageId="6" pageNumber="247">Phaeodothis</emphasis>
Syd. &amp; P. Syd., Annls
</taxonomicName>
mycol. 2(2): 166 (1904)
</heading>
</paragraph>
<paragraph id="8B9AB52CF96B830AFF3C6345FC9EC945" blockId="6.[136,792,572,594]" box="[136,792,572,594]" pageId="6" pageNumber="247">Index Fungorum Number: IF3914; Facesoffungi number: FoF00059</paragraph>
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<paragraph id="8B9AB52CF96B830AFF3C63F9FE9BC811" blockId="6.[136,1452,640,775]" pageId="6" pageNumber="247">
Notes:—The genus
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<emphasis id="B951693EF96B830AFED963F9FE7FC98D" box="[365,505,640,666]" italics="true" pageId="6" pageNumber="247">Phaeodothis</emphasis>
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was introduced by
<bibRefCitation id="EFB4C8DDF96B830AFD6A63F9FB8CC98D" author="Sydow, H. &amp; Sydow, P." box="[734,1034,640,666]" pageId="6" pageNumber="247" pagination="162 - 174" refId="ref10677" refString="Sydow, H. &amp; Sydow, P. (1904) Novae fungorum species. Ann. Mycol. 2: 162 - 174." type="journal article" year="1904">Sydow and Sydow (1904)</bibRefCitation>
to accommodate
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<emphasis id="B951693EF96B830AFB6363F8FB6EC98D" box="[1239,1256,641,666]" italics="true" pageId="6" pageNumber="247">P</emphasis>
.
<emphasis id="B951693EF96B830AFB4D63F9FAE9C98D" box="[1273,1391,640,666]" italics="true" pageId="6" pageNumber="247">tricuspidis</emphasis>
Syd. &amp; P. Syd.
</taxonomicName>
as the
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species.
<taxonomicName id="4C25CEAFF96B830AFE5163DDFD4DC9A9" box="[485,715,676,702]" pageId="6" pageNumber="247">
<emphasis id="B951693EF96B830AFE5163DDFDF7C9A9" box="[485,625,676,702]" italics="true" pageId="6" pageNumber="247">Phaeodothis</emphasis>
species
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are characterized by immersed to nearly superficial ascomata, a sparse hamathecium consisting of cellular pseudoparaphyses and 1-septate ascospores (
<bibRefCitation id="EFB4C8DDF96B830AFBD763B1FB79C9F4" author="Aptroot, A." box="[1123,1279,712,739]" pageId="6" pageNumber="247" pagination="325 - 379" refId="ref6381" refString="Aptroot, A. (1995) Redisposition of some species excluded from Didymosphaen'a (Ascomycotina). Nova Hedwigia 60: 325 - 379." type="journal article" year="1995">Aptroot 1995</bibRefCitation>
,
<bibRefCitation id="EFB4C8DDF96B830AFABD63B1FF75C811" author="Ariyawansa, H. A. &amp; Camporesi, E. &amp; Thambugala, K. M. &amp; Mapook, A. &amp; Kang, J. C. &amp; Alias, S. A. &amp; Chukeatirote, E. &amp; Thines, M. &amp; Mckenzie, E. H. &amp; Hyde, K. D." pageId="6" pageNumber="247" pagination="102 - 119" refId="ref6407" refString="Ariyawansa, H. A., Camporesi, E., Thambugala, K. M., Mapook, A., Kang, J. C., Alias, S. A., Chukeatirote, E., Thines, M., Mckenzie, E. H. &amp; Hyde, K. D. (2014 a) Confusion surrounding Didymosphaeria - phylogenetic and morphological evidence suggest Didymosphaeriaceae is not a distinct family. Phytotaxa 176: 102 - 119. https: // doi. org / 10.11646 / phytotaxa. 176.1.12" type="journal article" year="2014">
Ariyawansa
<emphasis id="B951693EF96B830AFA2363B0FF18C811" italics="true" pageId="6" pageNumber="247">et al</emphasis>
. 2014a
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, b).
</paragraph>
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<mods:namePart id="EA173389AC7619DB2EB88E956831908D">Kuo, Chang-Hsin</mods:namePart>
<mods:affiliation id="1AF35367EBA5BDE73019D8FA66F61691">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan.</mods:affiliation>
</mods:name>
<mods:name id="D674608261021213C189687960FA4246" type="personal">
<mods:role id="351772FD421FF8363C84C7B6A6B21F0D">
<mods:roleTerm id="6ECEDE30AB1D038488F5124468DE9F7A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFE893F9D4CC52C5FE2D2E9F3B54E540">Hyde, Kevin D.</mods:namePart>
<mods:affiliation id="98623AE9E27248089B1A7CC0E1A5E5A8">School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China. &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0C5DAAB0E74CD84697A73974A5F512AB">text</mods:typeOfResource>
<mods:relatedItem id="44DEC57FE63542A8DD8E0D9A8076DA0C" type="host">
<mods:titleInfo id="20231035B80B7C7BCAF41FF4E457DD0B">
<mods:title id="891DFE9EF7BF83FDBD9687431D17BC8D">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="6DE2A40F4D2B1982496729B6C59636D0">
<mods:date id="252F5C1FB20D13AEA9E5EE08084A1D06">2020</mods:date>
<mods:detail id="53420A6396D7A0B8F181A1BD06785872" type="pubDate">
<mods:number id="56A3C34A1F224BF8B6DF1F5FFA53CFE0">2020-01-14</mods:number>
</mods:detail>
<mods:detail id="05B40B9607F88C3283D40EB966647F93" type="volume">
<mods:number id="0E0D5F102C4EF458165B105EEA9F7F01">428</mods:number>
</mods:detail>
<mods:detail id="97025A60064EDDC997EC450125FC00F9" type="issue">
<mods:number id="E0DEB73BFE5C88384BEFE874618F940E">3</mods:number>
</mods:detail>
<mods:extent id="7AA4454FD2ED38BCDA392742A05B0A5C" unit="page">
<mods:start id="50742D8CE69ED6C146BF6D4C156B7906">241</mods:start>
<mods:end id="44D318C8CC1FE2DDDC002AD8EFF5B105">254</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3F7645D3F9E168B1C9CA74EAE3746C24">
<mods:url id="9C62367529A5BA61FE4FF4ECD0212A41">http://dx.doi.org/10.11646/phytotaxa.428.3.5</mods:url>
</mods:location>
<mods:classification id="AE6A5DF78A445954CBC452F5761D6428">journal article</mods:classification>
<mods:identifier id="9863272594666D3740F2A5322C404AB0" type="DOI">10.11646/phytotaxa.428.3.5</mods:identifier>
<mods:identifier id="189FBAF204F48F83B76E763E2965C1E3" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="038C043AF96B830BFF3C624DFC23CA17" LSID="urn:lsid:plazi:treatment:038C043AF96B830BFF3C624DFC23CA17" httpUri="http://treatment.plazi.org/id/038C043AF96B830BFF3C624DFC23CA17" lastPageId="7" lastPageNumber="247" pageId="6" pageNumber="247">
<subSubSection id="C33FE6A7F96B830AFF3C624DFC2CC866" pageId="6" pageNumber="247" type="nomenclature">
<paragraph id="8B9AB52CF96B830AFF3C624DFCD5C859" blockId="6.[136,851,820,847]" box="[136,851,820,847]" pageId="6" pageNumber="247">
<heading id="D0D20240F96B830AFF3C624DFCD5C859" box="[136,851,820,847]" fontSize="11" level="2" pageId="6" pageNumber="247" reason="3">
<taxonomicName id="4C25CEAFF96B830AFF3C624DFD77C859" ID-CoL="9ZD84" authority="Tennakoon, C.H. Kuo &amp; K.D. Hyde" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[136,753,820,847]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori" status="sp. nov.">
<emphasis id="B951693EF96B830AFF3C624DFED2C859" bold="true" box="[136,340,820,847]" italics="true" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
Tennakoon, C.H. Kuo &amp; K.D. Hyde
</taxonomicName>
,
<emphasis id="B951693EF96B830AFD49624DFCD5C859" bold="true" box="[765,851,820,846]" italics="true" pageId="6" pageNumber="247">
<taxonomicNameLabel id="A262D445F96B830AFD49624DFCD5C859" box="[765,851,820,846]" pageId="6" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9AB52CF96B830AFF3C6225FC2CC866" blockId="6.[136,938,860,882]" box="[136,938,860,882]" pageId="6" pageNumber="247">
Index Fungorum Number: IF 556761; Facesoffungi number: FoF06268,
<figureCitation id="131EA9A9F96B830AFCF66225FC2CC866" box="[834,938,860,881]" captionStart="FIGURE 2" captionStartId="7.[136,229,1326,1348]" captionTargetBox="[156,1431,278,1296]" captionTargetId="figure-68@7.[151,1436,278,1299]" captionTargetPageId="7" captionText="FIGURE 2. Phaeodothis mori (holotype, MFLU 18-2612) a, b. Appearance of ascomata on the host. c. Section of ascoma. d. Section of peridium. e. Pseudoparaphyses. fi. Asci. jl. Ascospores. m. Ascospores stained in Indian ink showing mucilaginous sheath. n. Germinated ascospore. o. Colony from below. p. Colony from above. Scale bars: c = 50 µm, d = 10 µm, ei = 20 µm, jn = 5 µm." pageId="6" pageNumber="247">FIGURE 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33FE6A7F96B830BFF3C62D9FC23CA17" lastPageId="7" lastPageNumber="248" pageId="6" pageNumber="247" type="etymology">
<paragraph id="8B9AB52CF96B830AFF3C62D9FB23C8AD" blockId="6.[136,1452,928,2071]" box="[136,1189,928,955]" pageId="6" pageNumber="247">
Etymology:
<emphasis id="B951693EF96B830AFEB862D9FEAEC8AD" bold="true" box="[268,296,928,954]" pageId="6" pageNumber="247"></emphasis>
Name reflects the host
<taxonomicName id="4C25CEAFF96B830AFD9E62D8FD5BC8AD" box="[554,733,928,954]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="247" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF96B830AFD9E62D8FD5BC8AD" box="[554,733,928,954]" italics="true" pageId="6" pageNumber="247">Morus australis</emphasis>
</taxonomicName>
, from which the
<typeStatus id="549E0B8EF96B830AFC2962D9FC79C8AD" box="[925,1023,928,954]" pageId="6" pageNumber="247" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
<paragraph id="8B9AB52CF96B830AFF0962BDFE7AC8C9" blockId="6.[136,1452,928,2071]" box="[189,508,964,991]" pageId="6" pageNumber="247">
<typeStatus id="549E0B8EF96B830AFF0962BDFEACC8C9" box="[189,298,964,990]" pageId="6" pageNumber="247" type="holotype">Holotype</typeStatus>
:
<emphasis id="B951693EF96B830AFE9962BDFECFC8C9" bold="true" box="[301,329,964,990]" pageId="6" pageNumber="247"></emphasis>
MFLU 18-2612
</paragraph>
<paragraph id="8B9AB52CF96B830AFF096291FC40CE7D" blockId="6.[136,1452,928,2071]" pageId="6" pageNumber="247">
<emphasis id="B951693EF96B830AFF096291FEA7CF15" box="[189,289,1000,1026]" italics="true" pageId="6" pageNumber="247">Saprobic</emphasis>
on decaying leaves of
<taxonomicName id="4C25CEAFF96B830AFD9F6290FCF2CF14" authority="Poir. Sexual" authorityName="Poir. Sexual" box="[555,884,1000,1027]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="247" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF96B830AFD9F6290FD58CF15" box="[555,734,1000,1026]" italics="true" pageId="6" pageNumber="247">Morus australis</emphasis>
Poir.
<emphasis id="B951693EF96B830AFC916291FCF2CF14" bold="true" box="[805,884,1000,1027]" italics="true" pageId="6" pageNumber="247">Sexual</emphasis>
</taxonomicName>
<emphasis id="B951693EF96B830AFCC96291FC4FCF14" bold="true" box="[893,969,1000,1027]" italics="true" pageId="6" pageNumber="247">morph</emphasis>
:
<emphasis id="B951693EF96B830AFC6E6291FBCECF15" box="[986,1096,1000,1026]" italics="true" pageId="6" pageNumber="247">Ascomata</emphasis>
100150 μm high, 80130 μm diam., scattered to gregarious, immersed, papilla erumpent through host surface, coriaceous, dark brown to black, globose to subglobose, ostiolate.
<taxonomicName id="4C25CEAFF96B830AFDB16549FDEBCF5D" box="[517,621,1072,1098]" class="Magnoliopsida" family="Peraceae" genus="Peridium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="247" phylum="Tracheophyta" rank="genus">
<emphasis id="B951693EF96B830AFDB16549FDEBCF5D" box="[517,621,1072,1098]" italics="true" pageId="6" pageNumber="247">Peridium</emphasis>
</taxonomicName>
1520 μm wide, composing several layers of irregular cells arranged in a
<emphasis id="B951693EF96B830AFF2A652CFEE4CF79" box="[158,354,1108,1134]" italics="true" pageId="6" pageNumber="247">
textura
<taxonomicName id="4C25CEAFF96B830AFF43652DFEE4CF79" box="[247,354,1108,1134]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Paraphaeosphaeria" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="angularis">angularis</taxonomicName>
</emphasis>
, cells towards the inside hyaline, at the outside, light brown, thick-walled, somewhat flattened, fusing and indistinguishable from the host tissues.
<emphasis id="B951693EF96B830AFD7C6501FCE3CF85" box="[712,869,1144,1170]" italics="true" pageId="6" pageNumber="247">Hamathecium</emphasis>
of 23 µm wide, cylindrical cellular, anastomosed, septate, unbranched, pseudoparaphyses.
<emphasis id="B951693EF96B830AFDE765E5FD05CFA1" box="[595,643,1180,1206]" italics="true" pageId="6" pageNumber="247">Asci</emphasis>
(43.5) 4565 × (5.3) 5.87 (7.3) μm (
<emphasis id="B951693EF96B830AFBDC65E3FBF3CFA7" box="[1128,1141,1178,1200]" italics="true" pageId="6" pageNumber="247">x</emphasis>
= 51.6 × 6.5 μm, n = 20), 8-spored, bitunicate, cylindrical to cylindric-clavate, short pedicellate with furcate to obtuse ends, apically rounded with an indistinct ocular chamber.
<emphasis id="B951693EF96B830AFDBD659DFD0ECFE9" box="[521,648,1252,1278]" italics="true" pageId="6" pageNumber="247">Ascospores</emphasis>
10.512.5 × 34 μm (
<emphasis id="B951693EF96B830AFC37659BFC16CFEF" box="[899,912,1250,1272]" italics="true" pageId="6" pageNumber="247">x</emphasis>
= 11.5 × 3.6 μm, n = 25), overlapping biseriate, yellowish brown, subfusoid, septate at the centre, strongly constricted at the septum, straight or slightly curved, enlarged near the septum at the upper cell, surrounded by thin mucilaginous sheath, spreading up to 56 μm at both ends (with
<taxonomicName id="4C25CEAFF96B830AFF3C6429FF56CE7D" box="[136,208,1360,1386]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="6" pageNumber="252" phylum="Chordata" rank="genus">Indian</taxonomicName>
ink), smooth-walled with guttules.
<emphasis id="B951693EF96B830AFDEB6429FC89CE7C" bold="true" box="[607,783,1360,1387]" italics="true" pageId="6" pageNumber="247">Asexual morph</emphasis>
: Undetermined.
</paragraph>
<paragraph id="8B9AB52CF96B830AFF09640DFB72CEED" blockId="6.[136,1452,928,2071]" pageId="6" pageNumber="247">
Culture characteristics:
<emphasis id="B951693EF96B830AFE76640DFE58CE99" bold="true" box="[450,478,1396,1422]" pageId="6" pageNumber="247"></emphasis>
Colonies on PDA,
<quantity id="4CDD18C9F96B830AFD01640DFCB7CE98" box="[693,817,1396,1423]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="6" pageNumber="247" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">2025 mm</quantity>
, diam. after 3 weeks, colonies medium dense, irregular, flat, slightly raised, surface smooth with crenate edge, fluffy to velvety with smooth aspects, zonate with different sector light brown to yellowish at the margin, dark brown at the middle, white to yellowish at centre; reverse light brown to yellowish at margin, dark brown to black at the centre, no pigmentation produced in media.
</paragraph>
<paragraph id="8B9AB52CF96B830AFF09677DFD3FCD9D" blockId="6.[136,1452,928,2071]" pageId="6" pageNumber="247">
Material examined:
<emphasis id="B951693EF96B830AFE28677DFE3ECD09" bold="true" box="[412,440,1540,1566]" pageId="6" pageNumber="247"></emphasis>
<collectingCountry id="F332F5BCF96B830AFE0C677DFD8BCD09" box="[440,525,1540,1566]" name="Taiwan" pageId="6" pageNumber="247">Taiwan</collectingCountry>
,
<collectingRegion id="49E17BCEF96B830AFDAF677DFDEECD09" box="[539,616,1540,1566]" country="Taiwan" name="Chiayi" pageId="6" pageNumber="247">Chiayi</collectingRegion>
, Fanlu Township area, Dahu forest, decaying leaves of
<taxonomicName id="4C25CEAFF96B830AFB42677CFF3BCD55" authority="Poir." authorityName="Poir." class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="247" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF96B830AFB42677CFA2DCD09" box="[1270,1451,1540,1566]" italics="true" pageId="6" pageNumber="247">Morus australis</emphasis>
Poir.
</taxonomicName>
(
<taxonomicName id="4C25CEAFF96B830AFF7A6751FEC5CD55" box="[206,323,1576,1602]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="247" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
),
<date id="FF9B93ECF96B830AFEEC6751FDB1CD55" box="[344,567,1576,1602]" pageId="6" pageNumber="247" value="2018-09-18">18 September 2018</date>
(
<geoCoordinate id="EE11D3EBF96B830AFDFD6751FD5ACD55" box="[585,732,1576,1602]" degrees="23" direction="north" minutes="27.422" orientation="latitude" pageId="6" pageNumber="247" precision="1" value="23.457033">23°27.422N</geoCoordinate>
<geoCoordinate id="EE11D3EBF96B830AFD526751FC00CD54" box="[742,902,1576,1603]" degrees="120" direction="east" minutes="36.385" orientation="longitude" pageId="6" pageNumber="247" precision="1" value="120.606415">120°36.385E</geoCoordinate>
), D.S. Tennakoon, TAP037-A (MFLU 18-2612,
<emphasis id="B951693EF96B830AFF3C6735FF69CD71" bold="true" box="[136,239,1612,1638]" pageId="6" pageNumber="247">
<typeStatus id="549E0B8EF96B830AFF3C6735FF69CD71" box="[136,239,1612,1638]" pageId="6" pageNumber="247" type="holotype">holotype</typeStatus>
</emphasis>
), ex-type living culture, MFLUCC 18-1634;
<emphasis id="B951693EF96B830AFD546735FC8ACD71" box="[736,780,1612,1638]" italics="true" pageId="6" pageNumber="247">ibid</emphasis>
.
<date id="FF9B93ECF96B830AFCAC6735FC68CD71" box="[792,1006,1612,1638]" pageId="6" pageNumber="247" value="2018-09-20">20 September 2018</date>
(
<geoCoordinate id="EE11D3EBF96B830AFC4F6735FB09CD70" box="[1019,1167,1612,1639]" degrees="23" direction="north" minutes="27.405" orientation="latitude" pageId="6" pageNumber="247" precision="1" value="23.45675">23°27.405N</geoCoordinate>
<geoCoordinate id="EE11D3EBF96B830AFB206735FAB2CD70" box="[1172,1332,1612,1639]" degrees="120" direction="east" minutes="36.568" orientation="longitude" pageId="6" pageNumber="247" precision="1" value="120.60947">120°36.568E</geoCoordinate>
), TAP037- B (MFLU 19-2334,
<emphasis id="B951693EF96B830AFEDF6709FE50CD9D" bold="true" box="[363,470,1648,1674]" pageId="6" pageNumber="247">
<typeStatus id="549E0B8EF96B830AFEDF6709FE50CD9D" box="[363,470,1648,1674]" pageId="6" pageNumber="247" type="paratype">paratype</typeStatus>
</emphasis>
), KUMCC19-0136.
</paragraph>
<paragraph id="8B9AB52CF96B830BFF0967ECFC23CA17" blockId="6.[136,1452,928,2071]" lastBlockId="7.[136,1451,158,256]" lastPageId="7" lastPageNumber="248" pageId="6" pageNumber="247">
Notes:
<emphasis id="B951693EF96B830AFEB267EDFEA4CDB9" bold="true" box="[262,290,1684,1710]" pageId="6" pageNumber="247"></emphasis>
Fresh collections of
<taxonomicName id="4C25CEAFF96B830AFDBD67EDFD54CDB9" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[521,722,1684,1710]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96B830AFDBD67EDFD54CDB9" box="[521,722,1684,1710]" italics="true" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
</taxonomicName>
were made from different places in Dahu forest,
<collectingRegion id="49E17BCEF96B830AFAB467EDFACBCDB9" box="[1280,1357,1684,1710]" country="Taiwan" name="Chiayi" pageId="6" pageNumber="247">Chiayi</collectingRegion>
,
<collectingCountry id="F332F5BCF96B830AFAED67EDFA2DCDB9" box="[1369,1451,1684,1710]" name="Taiwan" pageId="6" pageNumber="247">Taiwan</collectingCountry>
(
<geoCoordinate id="EE11D3EBF96B830AFF2467C1FEA5CDC5" box="[144,291,1720,1746]" degrees="23" direction="north" minutes="27.422" orientation="latitude" pageId="6" pageNumber="247" precision="1" value="23.457033">23°27.422N</geoCoordinate>
<geoCoordinate id="EE11D3EBF96B830AFE9A67C1FE4DCDC4" box="[302,459,1720,1747]" degrees="120" direction="east" minutes="36.385" orientation="longitude" pageId="6" pageNumber="247" precision="1" value="120.606415">120°36.385E</geoCoordinate>
and
<geoCoordinate id="EE11D3EBF96B830AFDBD67C1FD1ACDC4" box="[521,668,1720,1747]" degrees="23" direction="north" minutes="27.405" orientation="latitude" pageId="6" pageNumber="247" precision="1" value="23.45675">23°27.405N</geoCoordinate>
<geoCoordinate id="EE11D3EBF96B830AFD1267C1FCCECDC4" box="[678,840,1720,1747]" degrees="120" direction="east" minutes="36.568" orientation="longitude" pageId="6" pageNumber="247" precision="1" value="120.60947">120°36.568E</geoCoordinate>
). According to the multi-gene phylogenetic analyses (ITS, LSU, SSU and
<emphasis id="B951693EF96B830AFECB67A5FE46CDE1" box="[383,448,1756,1782]" italics="true" pageId="6" pageNumber="247">tef1-α</emphasis>
),
<taxonomicName id="4C25CEAFF96B830AFE6E67A5FD23CDE1" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[474,677,1756,1782]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96B830AFE6E67A5FD23CDE1" box="[474,677,1756,1782]" italics="true" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
</taxonomicName>
nested with the
<taxonomicName id="4C25CEAFF96B830AFCD167A5FA8CCDE1" authority="(Niessl) Aptroot" authorityName="Aptroot" baseAuthorityName="Niessl" box="[869,1290,1756,1782]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="winteri">
<emphasis id="B951693EF96B830AFCD167A5FBCFCDE1" box="[869,1097,1756,1782]" italics="true" pageId="6" pageNumber="247">Phaeodothis winteri</emphasis>
(Niessl) Aptroot
</taxonomicName>
species (CBS 182.58 and AFTOL-ID 1590) in moderate bootstrap support (72% ML, 85% MP, 0.90 BYPP). The morphological characteristics of
<taxonomicName id="4C25CEAFF96B830AFEE4665DFD9FCC29" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[336,537,1828,1854]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96B830AFEE4665DFD9FCC29" box="[336,537,1828,1854]" italics="true" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
</taxonomicName>
fits with the generic morphology of
<taxonomicName id="4C25CEAFF96B830AFC09665DFBCFCC29" box="[957,1097,1828,1854]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="genus">
<emphasis id="B951693EF96B830AFC09665DFBCFCC29" box="[957,1097,1828,1854]" italics="true" pageId="6" pageNumber="247">Phaeodothis</emphasis>
</taxonomicName>
by having, immersed to nearly superficial ascomata, a sparse hamathecium consisting of cellular pseudoparaphyses, bitunicate, fissitunicate, cylindrical asci and fusiform, rounded end, 1-septate ascospores (
<bibRefCitation id="EFB4C8DDF96B830AFCD86615FB8ECC90" author="Aptroot, A." box="[876,1032,1900,1927]" pageId="6" pageNumber="247" pagination="325 - 379" refId="ref6381" refString="Aptroot, A. (1995) Redisposition of some species excluded from Didymosphaen'a (Ascomycotina). Nova Hedwigia 60: 325 - 379." type="journal article" year="1995">Aptroot 1995</bibRefCitation>
,
<bibRefCitation id="EFB4C8DDF96B830AFBA66615FAADCC91" author="Ariyawansa, H. A. &amp; Tanaka, K. &amp; Thambugala, K. M. &amp; Phookamsak, R. &amp; Tian, Q. &amp; Camporesi, E. &amp; Hongsanan, S. &amp; Monkai, J. &amp; Wanasinghe, D. N. &amp; Mapook, A. &amp; Chukeatirote, E." box="[1042,1323,1900,1927]" pageId="6" pageNumber="247" pagination="69 - 104" refId="ref6507" refString="Ariyawansa, H. A., Tanaka, K., Thambugala, K. M., Phookamsak, R., Tian, Q., Camporesi, E., Hongsanan, S., Monkai, J., Wanasinghe, D. N., Mapook, A. &amp; Chukeatirote, E. (2014 b) A molecular phylogenetic reappraisal of the Didymosphaeriaceae (= Montagnulaceae). Fungal Diversity 68: 69 - 104. https: // doi. org / 10.1007 / s 13225 - 014 - 0305 - 6" type="journal article" year="2014">
Ariyawansa
<emphasis id="B951693EF96B830AFB146614FB54CC91" box="[1184,1234,1900,1926]" italics="true" pageId="6" pageNumber="247">et al</emphasis>
. 2014b
</bibRefCitation>
). However, our novel species,
<taxonomicName id="4C25CEAFF96B830AFED566E9FDAACCBD" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[353,556,1936,1962]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96B830AFED566E9FDAACCBD" box="[353,556,1936,1962]" italics="true" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C25CEAFF96B830AFCDF66E8FC5ACCBD" box="[875,988,1937,1963]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="winteri">
<emphasis id="B951693EF96B830AFCDF66E8FCFACCBD" box="[875,892,1937,1962]" italics="true" pageId="6" pageNumber="247">P</emphasis>
.
<emphasis id="B951693EF96B830AFC3966E8FC5ACCBD" box="[909,988,1937,1962]" italics="true" pageId="6" pageNumber="247">winteri</emphasis>
</taxonomicName>
(a synonym of
<taxonomicName id="4C25CEAFF96B830AFB2366E9FF56CCD9" authority="Niessl" authorityName="Niessl" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Didymosphaeria" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="winteri">
<emphasis id="B951693EF96B830AFB2366E9FA2DCCBD" box="[1175,1451,1936,1962]" italics="true" pageId="6" pageNumber="247">Didymosphaeria winteri</emphasis>
Niessl
</taxonomicName>
) in having smaller asci (4565 × 5.87 μm), ascospores (10.512.5 × 34 μm) with mucilaginous sheath, whereas
<taxonomicName id="4C25CEAFF96B830AFF5866A0FEDDCCE5" box="[236,347,2009,2035]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Didymosphaeria" kingdom="Fungi" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="winteri">
<emphasis id="B951693EF96B830AFF5866A0FF7BCCE5" box="[236,253,2009,2034]" italics="true" pageId="6" pageNumber="247">P</emphasis>
.
<emphasis id="B951693EF96B830AFEB866A0FEDDCCE5" box="[268,347,2009,2034]" italics="true" pageId="6" pageNumber="247">winteri</emphasis>
</taxonomicName>
has larger asci (6080 × 1012 μm) and ascospores (1114 × 4.55 μm) without a mucilaginous sheath (
<bibRefCitation id="EFB4C8DDF96B830AFF546685FEEAC300" author="Niessl, G." box="[224,364,2044,2071]" pageId="6" pageNumber="247" pagination="165 - 166" refId="ref9218" refString="Niessl, G. (1875) Neue Kernpilze. I. Serie. Oesterreichische botanische Zeitschrift 25: 165 - 166. https: // doi. org / 10.1007 / BF 01614606" type="journal article" year="1875">Niessl 1875</bibRefCitation>
). In addition, a comparison of the 851 nucleotides across the
<emphasis id="B951693EF96B830AFB9C6685FBECC301" box="[1064,1130,2044,2070]" italics="true" pageId="6" pageNumber="247">tef1-α</emphasis>
gene region of
<taxonomicName id="4C25CEAFF96B830BFAAB6685FF3BCBAF" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" lastPageId="7" lastPageNumber="248" order="Pleosporales" pageId="6" pageNumber="247" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96B830BFAAB6685FF3BCBAF" italics="true" lastPageId="7" lastPageNumber="248" pageId="6" pageNumber="247">Phaeodothis mori</emphasis>
</taxonomicName>
(MFLUCC 18-1634) and
<taxonomicName id="4C25CEAFF96A830BFE5661E6FDD6CBAF" box="[482,592,159,185]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Didymosphaeria" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="winteri">
<emphasis id="B951693EF96A830BFE5661E6FE75CBAF" box="[482,499,159,184]" italics="true" pageId="7" pageNumber="248">P</emphasis>
.
<emphasis id="B951693EF96A830BFDB561E6FDD6CBAF" box="[513,592,159,184]" italics="true" pageId="7" pageNumber="248">winteri</emphasis>
</taxonomicName>
(CBS 182.58) reveals 29 base pair differences (3.4%) and therefore provides further evidence to introduce
<taxonomicName id="4C25CEAFF96A830BFE6461BAFDA5CBCB" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[464,547,195,221]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96A830BFE6461BAFE67CBCB" box="[464,481,195,220]" italics="true" pageId="7" pageNumber="248">P</emphasis>
.
<emphasis id="B951693EF96A830BFE5A61BAFDA5CBCB" box="[494,547,195,220]" italics="true" pageId="7" pageNumber="248">mori</emphasis>
</taxonomicName>
as a new species as in the guidelines of
<bibRefCitation id="EFB4C8DDF96A830BFC6A61BAFB63CBCB" author="Jeewon, R. &amp; Hyde, K. D." box="[990,1253,194,221]" pageId="7" pageNumber="248" pagination="1669 - 1677" refId="ref8488" refString="Jeewon, R. &amp; Hyde, K. D. (2016) Establishing species boundaries and new taxa among fungi: recommendations to resolve taxonomic ambiguities. Mycosphere 7: 1669 - 1677. https: // doi. org / 10.5943 / mycosphere / 7 / 11 / 4" type="journal article" year="2016">Jeewon &amp; Hyde (2016)</bibRefCitation>
. Remarkably, this is the first
<taxonomicName id="4C25CEAFF96A830BFF4B619FFE65CA17" box="[255,483,230,256]" pageId="7" pageNumber="248">
<emphasis id="B951693EF96A830BFF4B619FFE0DCA17" box="[255,395,230,256]" italics="true" pageId="7" pageNumber="248">Phaeodothis</emphasis>
species
</taxonomicName>
record from
<taxonomicName id="4C25CEAFF96A830BFDC2619EFCAECA17" box="[630,808,230,256]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF96A830BFDC2619EFCAECA17" box="[630,808,230,256]" italics="true" pageId="7" pageNumber="248">Morus australis</emphasis>
</taxonomicName>
in
<collectingCountry id="F332F5BCF96A830BFCFF619FFC27CA17" box="[843,929,230,256]" name="Taiwan" pageId="7" pageNumber="248">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,407 @@
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<taxonomicName id="4C364D40754FBA791387550AFDACF84A" ID-CoL="3PFJN" authority="Lei Jiang &amp; K.W.Xu" authorityName="Lei Jiang &amp; K. W. Xu" authorityYear="2020" box="[136,596,1908,1935]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="jiangmenensis" status="sp. nov.">
<emphasis id="B942EAD1754FBA791387550AFE9AF84B" bold="true" box="[136,354,1908,1935]" italics="true" pageId="0" pageNumber="153">Ilex jiangmenensis</emphasis>
Lei Jiang &amp; K.W.Xu
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,
<emphasis id="B942EAD1754FBA791150550BFD4CF84B" box="[607,692,1909,1934]" italics="true" pageId="0" pageNumber="153">
<taxonomicNameLabel id="A27157AA754FBA791150550BFD4CF84B" box="[607,692,1909,1934]" pageId="0" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="130D2A46754FBA7911CC550AFCEAF84B" box="[707,786,1908,1934]" captionStart="FIGURE 2" captionStartId="2.[136,229,1345,1366]" captionTargetBox="[170,1416,190,1320]" captionTargetId="figure-30@2.[170,1416,190,1320]" captionTargetPageId="2" captionText="FIGURE 2. Ilex jiangmenensis sp. nov. —A. Branches. —B. Old branches. —C. Current years branch —D.Abaxial view of leaf blade. —E. Adaxial view of leaf blade. —F. Branches with mature fruit. —G. Staminate inflorescences. —H. Female flower. —I. Male flower. —J. Infructescences and fruits. —K. Fruits and pyrenes. —L. Mature fruit." figureDoi="http://doi.org/10.5281/zenodo.13875821" httpUri="https://zenodo.org/record/13875821/files/figure.png" pageId="0" pageNumber="153">Figs. 2</figureCitation>
,
<figureCitation id="130D2A46754FBA791011550AFCD4F84B" box="[798,812,1908,1934]" captionStart="FIGURE 3" captionStartId="3.[136,229,1311,1333]" captionTargetBox="[138,1448,190,1286]" captionTargetId="figure-19@3.[138,1448,190,1287]" captionTargetPageId="3" captionText="FIGURE 3. Ilex jiangmenensis sp. nov. —A, B. Habitat and vegetation. —C. Habit." figureDoi="http://doi.org/10.5281/zenodo.13875823" httpUri="https://zenodo.org/record/13875823/files/figure.png" pageId="0" pageNumber="153">3</figureCitation>
&amp;
<figureCitation id="130D2A46754FBA79105F550AFC8EF84B" box="[848,886,1908,1934]" captionStart="FIGURE 4" captionStartId="4.[136,229,2033,2055]" captionTargetBox="[161,1427,1108,2009]" captionTargetId="figure-203@4.[161,1427,1108,2009]" captionTargetPageId="4" captionText="FIGURE 4. Type specimens of Ilex jiangmenensis and I. pingheensis. —A. Holotype of I. jiangmenensis (Jiang et al. GDS00201, IBSC!). —B. Holotype of I. pingheensis (He 0116, FJSI-016556!)." figureDoi="http://doi.org/10.5281/zenodo.13875825" httpUri="https://zenodo.org/record/13875825/files/figure.png" pageId="0" pageNumber="153">4A</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
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<paragraph id="8B8936C3754FBA7813B255E6FC03FF19" blockId="0.[136,1452,1944,2079]" lastBlockId="1.[136,1451,158,220]" lastPageId="1" lastPageNumber="154" pageId="0" pageNumber="153">
<emphasis id="B942EAD1754FBA7913B255E6FEAEF877" bold="true" box="[189,342,1944,1970]" pageId="0" pageNumber="153">Diagnosis:—</emphasis>
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<emphasis id="B942EAD1754FBA79125955E6FDD1F877" box="[342,553,1944,1970]" italics="true" pageId="0" pageNumber="153">Ilex jiangmenensis</emphasis>
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is most similar to
<taxonomicName id="4C364D40754FBA79100E55E6FB57F877" authority="C. J. Tseng (1989: 29)" authorityName="C. J. Tseng" authorityPageNumber="29" authorityYear="1989" box="[769,1199,1944,1970]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="pingheensis">
<emphasis id="B942EAD1754FBA79100E55E6FC40F877" box="[769,952,1944,1970]" italics="true" pageId="0" pageNumber="153">Ilex pingheensis</emphasis>
C.J.
<bibRefCitation id="EFA74B32754FBA7910E955E6FB57F877" author="Tseng, C. - J." box="[998,1199,1944,1970]" pageId="0" pageNumber="153" pagination="29 - 30" refId="ref3310" refString="Tseng, C. - J. (1989) New taxa of Ilex from Fujian. Bulletin of Botanical Research 9 (4): 29 - 30." type="journal article" year="1989">Tseng (1989: 29)</bibRefCitation>
</taxonomicName>
in having leave blade thickly leathery, blade margin slightly or deeply undulate, sparsely black serrulate, reticulate veins conspicuous abaxially, inflorescences fasciculate, axillary, pyrenes 4(5), but
<taxonomicName id="4C364D40754FBA79105D559FFBF0F83F" authorityName="Lei Jiang &amp; K. W. Xu" authorityYear="2020" box="[850,1032,2017,2042]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="0" pageNumber="153" phylum="Tracheophyta" rank="species" species="jiangmenensis">
<emphasis id="B942EAD1754FBA79105D559FFBF0F83F" box="[850,1032,2017,2042]" italics="true" pageId="0" pageNumber="153">I. jiangmenensis</emphasis>
</taxonomicName>
has leave blade (1.2)2.53.5(4.5) × (0.6)1.21.5(2) cm, apex acute or short acuminate, fascicles of pistillate inflorescences 12(4)-cymes, and fascicles of staminate inflorescence (1)-3cymes, while
<taxonomicName id="4C364D40754EBA78119652E1FCCDFF7D" authorityName="C. J. Tseng" authorityYear="1989" box="[665,821,158,184]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="1" pageNumber="154" phylum="Tracheophyta" rank="species" species="pingheensis">
<emphasis id="B942EAD1754EBA78119652E1FCCDFF7D" box="[665,821,158,184]" italics="true" pageId="1" pageNumber="154">I. pingheensis</emphasis>
</taxonomicName>
has leave blade 3.56 ×
<quantity id="4CCE9B26754EBA78175F52E0FB29FF7C" box="[1104,1233,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="1" pageNumber="154" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 cm</quantity>
, apex acuminate or long acuminate, fascicles of pistillate and staminate inflorescences 4-flowered.
</paragraph>
</subSubSection>
<caption id="DF49664B754EBA78138756B7FC9FFB1A" ID-DOI="http://doi.org/10.5281/zenodo.13875819" ID-Zenodo-Dep="13875819" box="[136,871,1225,1247]" httpUri="https://zenodo.org/record/13875819/files/figure.png" pageId="1" pageNumber="154" startId="1.[136,229,1225,1247]" targetBox="[181,1405,253,1187]" targetPageId="1" targetType="figure">
<paragraph id="8B8936C3754EBA78138756B7FC9FFB1A" blockId="1.[136,871,1225,1247]" box="[136,871,1225,1247]" pageId="1" pageNumber="154">
<emphasis id="B942EAD1754EBA78138756B7FF06FB1B" bold="true" box="[136,254,1225,1247]" pageId="1" pageNumber="154">FIGURE 1.</emphasis>
Type locality of
<taxonomicName id="4C364D40754EBA7812AE56B7FDACFB1B" authorityName="Lei Jiang &amp; K. W. Xu" authorityYear="2020" box="[417,596,1225,1246]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="1" pageNumber="154" phylum="Tracheophyta" rank="species" species="jiangmenensis">
<emphasis id="B942EAD1754EBA7812AE56B7FDACFB1B" box="[417,596,1225,1246]" italics="true" pageId="1" pageNumber="154">Ilex jiangmenensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C364D40754EBA78118C56B7FCF0FB1B" authorityName="C. J. Tseng" authorityYear="1989" box="[643,776,1225,1246]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="1" pageNumber="154" phylum="Tracheophyta" rank="species" species="pingheensis">
<emphasis id="B942EAD1754EBA78118C56B7FCF0FB1B" box="[643,776,1225,1246]" italics="true" pageId="1" pageNumber="154">I. pingheensis</emphasis>
</taxonomicName>
in China.
</paragraph>
</caption>
<subSubSection id="C32C6548754EBA7B13B25773FBC5F97A" lastPageId="2" lastPageNumber="155" pageId="1" pageNumber="154" type="materials_examined">
<paragraph id="8B8936C3754EBA7813B25773FC64FAAA" blockId="1.[136,1452,1293,2075]" pageId="1" pageNumber="154">
<emphasis id="B942EAD1754EBA7813B25773FEE6FAE2" bold="true" box="[189,286,1293,1319]" pageId="1" pageNumber="154">Type:—</emphasis>
<materialsCitation id="3B5E3C9E754EBA7812125773FC6FFAAA" collectingDate="2011-03-27" collectionCode="IBSC, KUN" country="China" county="Jiangmen City" elevation="505" latitude="22.208838" location="Huogaishan" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" pageId="1" pageNumber="154" specimenCount="2" stateProvince="Guangdong" typeStatus="holotype">
<collectingCountry id="F3217653754EBA7812125773FE81FAE2" box="[285,377,1293,1319]" name="China" pageId="1" pageNumber="154">CHINA</collectingCountry>
.
<collectingRegion id="49F2F821754EBA7812805773FDEEFAE2" box="[399,534,1293,1319]" country="China" name="Guangdong" pageId="1" pageNumber="154">Guangdong</collectingRegion>
:
<collectingCounty id="62E84E4F754EBA7811235773FD22FAE2" box="[556,730,1293,1319]" pageId="1" pageNumber="154">Jiangmen City</collectingCounty>
,
<collectingMunicipality id="6BEDACB9754EBA7811FE5773FB3CFAED" box="[753,1220,1293,1320]" pageId="1" pageNumber="154">Gudoushan Provincial Nature Reserve</collectingMunicipality>
, elev. ca.
<quantity id="4CCE9B26754EBA78165D5773FA5FFAED" box="[1362,1447,1293,1320]" metricMagnitude="2" metricUnit="m" metricValue="5.05" pageId="1" pageNumber="154" unit="m" value="505.0">
<elevation id="001BD1F0754EBA78165D5773FA5FFAED" box="[1362,1447,1293,1320]" metricMagnitude="2" metricUnit="m" metricValue="5.05" pageId="1" pageNumber="154" unit="m" value="505.0">505 m</elevation>
</quantity>
,
<geoCoordinate id="EE025004754EBA781387574FFECFFA8E" box="[136,311,1329,1355]" degrees="22" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="154" precision="1" seconds="31.81" value="22.208838">22°1231.81”N</geoCoordinate>
,
<geoCoordinate id="EE025004754EBA78124A574FFE05FA89" box="[325,509,1329,1356]" degrees="112" direction="east" minutes="56" orientation="longitude" pageId="1" pageNumber="154" precision="1" seconds="28.26" value="112.941185">112°5628.26”E</geoCoordinate>
, on the way from
<location id="8EE96018754EBA7811D9574FFCA4FA8E" LSID="urn:lsid:plazi:treatment:039F87D5754FBA7B1387550AFCE9F7ED:8EE96018754EBA7811D9574FFCA4FA8E" box="[726,860,1329,1355]" country="China" county="Jiangmen City" latitude="22.208838" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" name="Huogaishan" pageId="1" pageNumber="154" stateProvince="Guangdong">Huogaishan</location>
to
<location id="8EE96018754EBA78108A574FFC0BFA8E" LSID="urn:lsid:plazi:treatment:039F87D5754FBA7B1387550AFCE9F7ED:8EE96018754EBA78108A574FFC0BFA8E" box="[901,1011,1329,1355]" country="China" county="Jiangmen City" latitude="22.208838" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" name="Banyuzui" pageId="1" pageNumber="154" stateProvince="Guangdong">Banyuzui</location>
, in evergreen broad-leaved forests,
<date id="FF881003754EBA781680574FFEEAFAAA" pageId="1" pageNumber="154" value="2011-03-27">
<collectingDate id="EFCCE9EB754EBA781680574FFEEAFAAA" pageId="1" pageNumber="154" value="2011-03-27">27 March 2011</collectingDate>
</date>
,
<emphasis id="B942EAD1754EBA7812125728FDE6FAAA" box="[285,542,1365,1391]" italics="true" pageId="1" pageNumber="154">Jiang et al. GDS00201</emphasis>
(
<typeStatus id="548D8861754EBA781123572BFD68FAAA" box="[556,656,1365,1391]" pageId="1" pageNumber="154" type="holotype">holotype</typeStatus>
<collectionCode id="ED27AE06754EBA781198572BFD26FAAA" box="[663,734,1365,1391]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="1" pageNumber="154" type="Herbarium">IBSC</collectionCode>
!;
<typeStatus id="548D8861754EBA7811E2572BFCC6FAAA" box="[749,830,1365,1391]" pageId="1" pageNumber="154" type="isotype">isotype</typeStatus>
<collectionCode id="ED27AE06754EBA78104A572BFC74FAAA" box="[837,908,1365,1391]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15842" lsid="urn:lsid:biocol.org:col:15842" name="Kunming Institute of Botany, Chinese Academy of Sciences" pageId="1" pageNumber="154" type="Herbarium">KUN</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8936C3754EBA7813B25707FEDCF86A" blockId="1.[136,1452,1293,2075]" pageId="1" pageNumber="154">
Shrubs, 1.02.0 m tall, stem
<quantity id="4CCE9B26754EBA7812FB5707FDBDFA51" box="[500,581,1401,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="154" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">12 cm</quantity>
in diam. Bark grey.Young branchlets yellow-green to green, terete, puberulent; second years branchlets yellow-brown to gray brown, with slightly raised circular or elliptic leaf scars and lenticels. Leaves persist on young branchlets. Stipules deciduous, minute; petiole abaxially terete, adaxially sulcate,
<quantity id="4CCE9B26754EBA78165857BFFF4EF9C5" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="1" pageNumber="154" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.54.5 mm</quantity>
, glabrous. Leaf blade pale green to yellow green abaxially, green to dark green adaxially, oval or elliptic to lanceolate, (1.2)2.53.5(4.5) × (0.6)1.21.5(2) cm, leathery, abaxially glabrous, midvein impressed adaxially and raised abaxially, lateral veins 47 pairs, abaxially conspicuous, green, flat or slightly raised, anastomosing near margin, adaxially slightly impressed or indistinct, reticulate veins conspicuous, green, flat or slightly raised abaxially, obscure adaxially, blade base obtuse or cuneate, margin sparsely serrulate, apex acute or short acuminate. Pistillate inflorescences: 1-flowered cymes, solitary or fasciculate, fascicles 12(4)-flowered, axillary on current years branchlets; pedicels ca.
<quantity id="4CCE9B26754EBA7812AE54C3FE18F91D" box="[417,480,1725,1752]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="154" unit="cm" value="1.0">1 cm</quantity>
, puberulent; bracteoles 2, yellow-brown, small, subulate. Staminate inflorescences: individual branches of the fascicles (1)3-flowered, fasciculate, axillary. Infructescences 1-fruited cymes, solitary or fasciculate, fascicles 12(4)-fruited, axillary on current years branchlets. Fruit red, globose or slightly depressed globose, 68(10) mm in diam.; fruit pedicel
<quantity id="4CCE9B26754EBA78119B5557FCFAF881" box="[660,770,1833,1860]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="1" pageNumber="154" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">410 mm</quantity>
, puberulent, usually purple in the upper part of the pedicel; calyx persistent, explanate, ca.
<quantity id="4CCE9B26754EBA7812FB5533FDC1F8AD" box="[500,569,1869,1896]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="154" unit="mm" value="1.0">1 mm</quantity>
in diam., glabrous, 4-lobed, lobes half-orbicular; stigma persistent, discoid; pyrenes 4(5), ellipsoidal,
<quantity id="4CCE9B26754EBA7812BA550FFDEAF849" box="[437,530,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="154" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
in length, ca.
<quantity id="4CCE9B26754EBA7811BD550FFD0BF849" box="[690,755,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="154" unit="mm" value="4.0">4 mm</quantity>
in width, abaxially thinly slightly striate and sulcate, laterally rugose-pitted.
</paragraph>
<paragraph id="8B8936C3754EBA7813B255C7FF1BF7DE" blockId="1.[136,1452,1293,2075]" pageId="1" pageNumber="154">
<emphasis id="B942EAD1754EBA7813B255C7FE0CF816" bold="true" box="[189,500,1977,2003]" pageId="1" pageNumber="154">Geographical distribution</emphasis>
:—
<taxonomicName id="4C364D40754EBA78111855C7FD1FF816" authorityName="Lei Jiang &amp; K. W. Xu" authorityYear="2020" box="[535,743,1977,2003]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="1" pageNumber="154" phylum="Tracheophyta" rank="species" species="jiangmenensis">
<emphasis id="B942EAD1754EBA78111855C7FD1FF816" box="[535,743,1977,2003]" italics="true" pageId="1" pageNumber="154">Ilex jiangmenensis</emphasis>
</taxonomicName>
is currently only known from Taishan City and Xinhui District, both belonging to Jiangmen City,
<collectingRegion id="49F2F821754EBA78110D55A3FD09F832" box="[514,753,2013,2039]" country="China" name="Guangdong" pageId="1" pageNumber="154">Guangdong Province</collectingRegion>
,
<collectingCountry id="F3217653754EBA7811F355A3FCBBF832" box="[764,835,2013,2039]" name="China" pageId="1" pageNumber="154">China</collectingCountry>
. This species is likely endemic to
<collectingRegion id="49F2F821754EBA7817B155A3FA54F832" box="[1214,1452,2013,2039]" country="China" name="Guangdong" pageId="1" pageNumber="154">Guangdong Province</collectingRegion>
(
<figureCitation id="130D2A46754EBA78139F5A7FFF2AF7DE" box="[144,210,2049,2075]" captionStart="FIGURE 1" captionStartId="1.[136,229,1225,1247]" captionTargetBox="[181,1405,253,1187]" captionTargetId="figure-45@1.[171,1417,242,1196]" captionTargetPageId="1" captionText="FIGURE 1. Type locality of Ilex jiangmenensis and I. pingheensis in China." figureDoi="http://doi.org/10.5281/zenodo.13875819" httpUri="https://zenodo.org/record/13875819/files/figure.png" pageId="1" pageNumber="154">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF49664B754DBA7B1387573FFCA1FA5A" ID-DOI="http://doi.org/10.5281/zenodo.13875821" ID-Zenodo-Dep="13875821" httpUri="https://zenodo.org/record/13875821/files/figure.png" pageId="2" pageNumber="155" startId="2.[136,229,1345,1366]" targetBox="[170,1416,190,1320]" targetPageId="2" targetType="figure">
<paragraph id="8B8936C3754DBA7B1387573FFCA1FA5A" blockId="2.[136,1453,1345,1439]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B1387573FFF06FA93" bold="true" box="[136,254,1345,1366]" pageId="2" pageNumber="155">FIGURE 2.</emphasis>
<taxonomicName id="4C364D40754DBA7B120B573FFE4FFA92" authorityName="Jiang &amp; Wu &amp; Huang &amp; Yu &amp; He &amp; Xu" authorityYear="2020" box="[260,439,1345,1367]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="jiangmenensis" status="sp. nov.">
<emphasis id="B942EAD1754DBA7B120B573FFE4FFA92" box="[260,439,1345,1367]" italics="true" pageId="2" pageNumber="155">Ilex jiangmenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157AA754DBA7B12B2573CFDF0FA92" box="[445,520,1346,1367]" pageId="2" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
—A. Branches. —B. Old branches. —C. Current years branch —D. Abaxial view of leaf blade. —E. Adaxial view of leaf blade. —F. Branches with mature fruit. —G. Staminate inflorescences. —H. Female flower. —I. Male flower. —J. Infructescences and fruits. —K. Fruits and pyrenes. —L. Mature fruit.
</paragraph>
</caption>
<paragraph id="8B8936C3754DBA7B13B257B3FE69F959" blockId="2.[136,1452,1485,2088]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B13B257B3FEDBFA22" bold="true" box="[189,291,1485,1511]" pageId="2" pageNumber="155">Habitat:</emphasis>
—The new species was observed to grow in rock crevices or beside the evergreen broad-leaved forests in the mountains of Gudoushan and Nanfengshan at elevation range of
<quantity id="4CCE9B26754DBA7B1067578FFC21F9C9" box="[872,985,1521,1548]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="5.5" metricValueMin="0.5" pageId="2" pageNumber="155" unit="m" value="300.0" valueMax="550.0" valueMin="50.0">
50
<specimenCount id="9D30FD4A754DBA7B109D578FFC21F9C9" box="[914,985,1521,1548]" count="550" pageId="2" pageNumber="155" type="generic">550 m</specimenCount>
</quantity>
.The dominant woody (arboreal)species of the community include
<taxonomicName id="4C364D40754DBA7B1281546BFC6EF9F5" authority="(Champ. ex Benth.) Masam." authorityName="Masam." baseAuthorityName="Benth." box="[398,918,1557,1584]" class="Magnoliopsida" family="Rubiaceae" genus="Aidia" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="canthioides">
<emphasis id="B942EAD1754DBA7B1281546BFDAAF9EA" box="[398,594,1557,1583]" italics="true" pageId="2" pageNumber="155">Aidia canthioides</emphasis>
(Champ. ex Benth.) Masam.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B10AF5468FB44F9EA" authority="Lel." authorityName="Lel." box="[928,1212,1557,1583]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" kingdom="Plantae" order="Buxales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="megistophylla">
<emphasis id="B942EAD1754DBA7B10AF5468FB7FF9EA" box="[928,1159,1557,1583]" italics="true" pageId="2" pageNumber="155">Buxus megistophylla</emphasis>
Lél.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B17C8546BFDD2F991" authority="(Chun) Chun ex Y. C. Hsu &amp; H. W. Jen." authorityName="Y. C. Hsu &amp; H. W. Jen." baseAuthorityName="Chun" class="Magnoliopsida" family="Fagaceae" genus="Cyclobalanopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="hui">
<emphasis id="B942EAD1754DBA7B17C8546BFA54F9EA" box="[1223,1452,1557,1583]" italics="true" pageId="2" pageNumber="155">Cyclobalanopsis hui</emphasis>
(Chun) Chun ex Y.C.Hsu &amp; H.W.Jen.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B113A5447FC42F991" authority="(Lindl.) Masam." authorityName="Masam." baseAuthorityName="Lindl." box="[565,954,1593,1620]" class="Magnoliopsida" family="Rubiaceae" genus="Diplospora" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="dubia">
<emphasis id="B942EAD1754DBA7B113A5447FD03F996" box="[565,763,1593,1619]" italics="true" pageId="2" pageNumber="155">Diplospora dubia</emphasis>
(Lindl.) Masam.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B10CB5444FAEDF996" authority="Lindl." authorityName="Lindl." box="[964,1301,1593,1619]" class="Magnoliopsida" family="Ebenaceae" genus="Diospyros" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="vaccinioides">
<emphasis id="B942EAD1754DBA7B10CB5444FB30F996" box="[964,1224,1593,1619]" italics="true" pageId="2" pageNumber="155">Diospyros vaccinioides</emphasis>
Lindl.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B16105447FE2AF9B2" authority="Gardn. &amp; Champ." authorityName="Gardn. &amp; Champ." class="Magnoliopsida" family="Pentaphylacaceae" genus="Pentaphylax" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="euryoides">
<emphasis id="B942EAD1754DBA7B16105447FF0DF9B2" italics="true" pageId="2" pageNumber="155">Pentaphylax euryoides</emphasis>
Gardn. &amp; Champ.
</taxonomicName>
,
<taxonomicName id="4C364D40754DBA7B12EF5423FC27F9BD" authority="(Lour.) Spreng." authorityName="Spreng." baseAuthorityName="Lour." box="[480,991,1629,1656]" class="Magnoliopsida" family="Phyllanthaceae" genus="Phyllanthus" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="cochinchinensis">
<emphasis id="B942EAD1754DBA7B12EF5423FCDBF9B2" box="[480,803,1629,1655]" italics="true" pageId="2" pageNumber="155">Phyllanthus cochinchinensis</emphasis>
(Lour.) Spreng.
</taxonomicName>
, and
<taxonomicName id="4C364D40754DBA7B172F5423FED7F959" authority="(Roxb. ex Ker Gawler) Sweet" authorityName="Sweet" baseAuthorityName="Ker Gawler" class="Magnoliopsida" family="Theaceae" genus="Polyspora" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="B942EAD1754DBA7B172F5423FB03F9B2" box="[1056,1275,1629,1655]" italics="true" pageId="2" pageNumber="155">Polyspora axillaris</emphasis>
(Roxb. ex Ker Gawler) Sweet
</taxonomicName>
(
<figureCitation id="130D2A46754DBA7B123054FFFE7BF959" box="[319,387,1665,1692]" captionStart="FIGURE 3" captionStartId="3.[136,229,1311,1333]" captionTargetBox="[138,1448,190,1286]" captionTargetId="figure-19@3.[138,1448,190,1287]" captionTargetPageId="3" captionText="FIGURE 3. Ilex jiangmenensis sp. nov. —A, B. Habitat and vegetation. —C. Habit." figureDoi="http://doi.org/10.5281/zenodo.13875823" httpUri="https://zenodo.org/record/13875823/files/figure.png" pageId="2" pageNumber="155">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B8936C3754DBA7B13B254D8FBC5F97A" blockId="2.[136,1452,1485,2088]" box="[189,1085,1701,1728]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B13B254D8FEC1F905" bold="true" box="[189,313,1702,1728]" pageId="2" pageNumber="155">Phenology</emphasis>
:—Flowering from March to April; fruiting from April to November.
</paragraph>
</subSubSection>
<subSubSection id="C32C6548754DBA7B13B254B4FCE9F7ED" pageId="2" pageNumber="155" type="etymology">
<paragraph id="8B8936C3754DBA7B13B254B4FDD2F8C2" blockId="2.[136,1452,1485,2088]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B13B254B4FEC6F921" bold="true" box="[189,318,1738,1764]" pageId="2" pageNumber="155">Etymology</emphasis>
:—The species epithet is based on the city name, Jiangmen City, and the Latin suffix, -ensis, of origin, where the new species was collected.
</paragraph>
<paragraph id="8B8936C3754DBA7B13B2556CFEC7F88A" blockId="2.[136,1452,1485,2088]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B13B2556CFE49F8E9" bold="true" box="[189,433,1810,1836]" pageId="2" pageNumber="155">Vernacular name:—</emphasis>
We propose a Chinese name, jiāngmén dōngqīng (
<emphasis id="B942EAD1754DBA7B10ED556DFBAAF8EF" bold="true" box="[994,1106,1811,1834]" pageId="2" pageNumber="155">江Ṅḳff</emphasis>
), to reflect the
<typeStatus id="548D8861754DBA7B160C556CFACBF8E9" box="[1283,1331,1810,1836]" pageId="2" pageNumber="155">type</typeStatus>
locality of the new species.
</paragraph>
<paragraph id="8B8936C3754DBA7B13B25527FCE9F7ED" blockId="2.[136,1452,1485,2088]" pageId="2" pageNumber="155">
<emphasis id="B942EAD1754DBA7B13B25527FDA2F8B6" bold="true" box="[189,602,1881,1907]" pageId="2" pageNumber="155">Additional specimens examined:—</emphasis>
<materialsCitation id="3B5E3C9E754DBA7B11555524FC7BF879" collectingDate="2019-03-27" collectionCode="IBSC, SYS" collectorName="Jiang" country="China" county="Jiangmen City" elevation="461" latitude="22.208838" location="Huogaishan" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" pageId="2" pageNumber="155" specimenCount="1" stateProvince="Guangdong">
<collectingCountry id="F3217653754DBA7B11555524FD4EF8B1" box="[602,694,1882,1908]" name="China" pageId="2" pageNumber="155">CHINA</collectingCountry>
.
<collectingRegion id="49F2F821754DBA7B11B05527FCBEF8B6" box="[703,838,1881,1907]" country="China" name="Guangdong" pageId="2" pageNumber="155">Guangdong</collectingRegion>
:
<collectingCounty id="62E84E4F754DBA7B10405527FC09F8B6" box="[847,1009,1881,1907]" pageId="2" pageNumber="155">Jiangmen City</collectingCounty>
,
<collectingMunicipality id="6BEDACB9754DBA7B10F45527FA5FF8B1" box="[1019,1447,1881,1908]" pageId="2" pageNumber="155">Gudoushan Provincial Nature Reserve</collectingMunicipality>
, on the way from
<location id="8EE96018754DBA7B12445503FE29F852" LSID="urn:lsid:plazi:treatment:039F87D5754FBA7B1387550AFCE9F7ED:8EE96018754DBA7B12445503FE29F852" box="[331,465,1917,1943]" country="China" county="Jiangmen City" latitude="22.208838" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" name="Huogaishan" pageId="2" pageNumber="155" stateProvince="Guangdong">Huogaishan</location>
to
<location id="8EE96018754DBA7B12F95503FD9CF852" LSID="urn:lsid:plazi:treatment:039F87D5754FBA7B1387550AFCE9F7ED:8EE96018754DBA7B12F95503FD9CF852" box="[502,612,1917,1943]" country="China" county="Jiangmen City" latitude="22.208838" longLatPrecision="1" longitude="112.941185" municipality="Gudoushan Provincial Nature Reserve" name="Banyuzui" pageId="2" pageNumber="155" stateProvince="Guangdong">Banyuzui</location>
, in rock crevices along the stream, elev. ca.
<quantity id="4CCE9B26754DBA7B17575503FB5AF85D" box="[1112,1186,1917,1944]" metricMagnitude="2" metricUnit="m" metricValue="4.61" pageId="2" pageNumber="155" unit="m" value="461.0">
<elevation id="001BD1F0754DBA7B17575503FB5AF85D" box="[1112,1186,1917,1944]" metricMagnitude="2" metricUnit="m" metricValue="4.61" pageId="2" pageNumber="155" unit="m" value="461.0">461 m</elevation>
</quantity>
,
<geoCoordinate id="EE025004754DBA7B17A15500FA93F85D" box="[1198,1387,1918,1944]" degrees="22" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="155" precision="1" seconds="31.81" value="22.208838">22° 12 31.81”N</geoCoordinate>
,
<geoCoordinate id="EE025004754DBA7B16785500FEE9F87E" degrees="112" direction="east" minutes="56" orientation="longitude" pageId="2" pageNumber="155" precision="1" seconds="28.26" value="112.941185">112° 56 28.26”E</geoCoordinate>
,
<date id="FF881003754DBA7B121355DCFE32F879" box="[284,458,1953,1980]" pageId="2" pageNumber="155" value="2019-03-27">
<collectingDate id="EFCCE9EB754DBA7B121355DCFE32F879" box="[284,458,1953,1980]" pageId="2" pageNumber="155" value="2019-03-27">27 March 2019</collectingDate>
</date>
,
<emphasis id="B942EAD1754DBA7B12DA55DCFD2DF879" box="[469,725,1953,1980]" italics="true" pageId="2" pageNumber="155">
<collectorName id="26C35315754DBA7B12DA55DCFDEBF87E" box="[469,531,1954,1979]" pageId="2" pageNumber="155">Jiang</collectorName>
et al. GDS00222
</emphasis>
(
<collectionCode id="ED27AE06754DBA7B11EB55DCFCD3F879" box="[740,811,1954,1980]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="155" type="Herbarium">IBSC</collectionCode>
!,
<collectionCode id="ED27AE06754DBA7B103555DCFC8EF879" box="[826,886,1954,1980]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" lsid="urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="2" pageNumber="155" type="Herbarium">SYS</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B5E3C9E754DBA7B108055DFFD1BF825" collectingDate="2019-03-27" collectionCode="IBSC, NF" collectorName="Jiang" country="China" elevation="500" latitude="22.209522" longLatPrecision="1" longitude="112.94007" pageId="2" pageNumber="155" specimenCount="1">
elev.
<quantity id="4CCE9B26754DBA7B10C655DFFBEAF879" box="[969,1042,1953,1980]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="155" unit="m" value="500.0">
<elevation id="001BD1F0754DBA7B10C655DFFBEAF879" box="[969,1042,1953,1980]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="155" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<geoCoordinate id="EE025004754DBA7B171255DCFB2FF879" box="[1053,1239,1954,1980]" degrees="22" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="155" precision="1" seconds="34.28" value="22.209522">22° 12 34.28”N</geoCoordinate>
,
<geoCoordinate id="EE025004754DBA7B17EC55DCFA5FF879" box="[1251,1447,1953,1980]" degrees="112" direction="east" minutes="56" orientation="longitude" pageId="2" pageNumber="155" precision="1" seconds="24.25" value="112.94007">112° 56 24.25”E</geoCoordinate>
,
<date id="FF881003754DBA7B138755B8FECFF825" box="[136,311,1989,2016]" pageId="2" pageNumber="155" value="2019-03-27">
<collectingDate id="EFCCE9EB754DBA7B138755B8FECFF825" box="[136,311,1989,2016]" pageId="2" pageNumber="155" value="2019-03-27">27 March 2019</collectingDate>
</date>
,
<emphasis id="B942EAD1754DBA7B124D55B8FDBCF825" box="[322,580,1989,2016]" italics="true" pageId="2" pageNumber="155">
<collectorName id="26C35315754DBA7B124D55B8FE78F81A" box="[322,384,1990,2015]" pageId="2" pageNumber="155">Jiang</collectorName>
et al. GDS00217
</emphasis>
(
<collectionCode id="ED27AE06754DBA7B115C55B8FD62F825" box="[595,666,1990,2016]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="155" type="Herbarium">IBSC</collectionCode>
!,
<collectionCode id="ED27AE06754DBA7B11A555B8FD2EF825" box="[682,726,1990,2016]" country="China" lsid="urn:lsid:biocol.org:col:13689" name="Nanjing Forestry University" pageId="2" pageNumber="155" type="Herbarium">NF</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B5E3C9E754DBA7B11FF55BBFD94F7C1" collectingDate="2019-03-27" collectionCode="IBSC, NF" collectorName="Jiang" country="China" elevation="521" latitude="22.210155" longLatPrecision="1" longitude="112.941605" pageId="2" pageNumber="155" specimenCount="1">
elev. ca.
<quantity id="4CCE9B26754DBA7B105D55BBFC63F825" box="[850,923,1989,2016]" metricMagnitude="2" metricUnit="m" metricValue="5.21" pageId="2" pageNumber="155" unit="m" value="521.0">
<elevation id="001BD1F0754DBA7B105D55BBFC63F825" box="[850,923,1989,2016]" metricMagnitude="2" metricUnit="m" metricValue="5.21" pageId="2" pageNumber="155" unit="m" value="521.0">521 m</elevation>
</quantity>
,
<geoCoordinate id="EE025004754DBA7B10A855B8FB9AF825" box="[935,1122,1989,2016]" degrees="22" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="155" precision="1" seconds="36.56" value="22.210155">22° 12 36.56”N</geoCoordinate>
,
<geoCoordinate id="EE025004754DBA7B176155B8FACBF81A" box="[1134,1331,1989,2016]" degrees="112" direction="east" minutes="56" orientation="longitude" pageId="2" pageNumber="155" precision="1" seconds="29.76" value="112.941605">112° 56 29.76”E</geoCoordinate>
,
<date id="FF881003754DBA7B163055B8FF3BF7C1" pageId="2" pageNumber="155" value="2019-03-27">
<collectingDate id="EFCCE9EB754DBA7B163055B8FF3BF7C1" pageId="2" pageNumber="155" value="2019-03-27">27 March 2019</collectingDate>
</date>
,
<emphasis id="B942EAD1754DBA7B13C15594FE36F7C6" box="[206,462,2025,2051]" italics="true" pageId="2" pageNumber="155">
<collectorName id="26C35315754DBA7B13C15594FEF4F7C6" box="[206,268,2026,2051]" pageId="2" pageNumber="155">Jiang</collectorName>
et al. GDS00167
</emphasis>
(
<collectionCode id="ED27AE06754DBA7B12D25594FDDCF7C1" box="[477,548,2026,2052]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="155" type="Herbarium">IBSC</collectionCode>
!,
<collectionCode id="ED27AE06754DBA7B113D5594FDA6F7C1" box="[562,606,2026,2052]" country="China" lsid="urn:lsid:biocol.org:col:13689" name="Nanjing Forestry University" pageId="2" pageNumber="155" type="Herbarium">NF</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B5E3C9E754DBA7B11785597FCF5F7ED" collectingDate="2019-07-04" collectionCode="SYS, NF" collectorName="Zhong" country="China" elevation="110" latitude="22.230476" location="Maji Reservoir" longLatPrecision="1" longitude="113.00271" municipality="Xinhui District" pageId="2" pageNumber="155" specimenCount="1">
<collectingMunicipality id="6BEDACB9754DBA7B11785597FCDBF7C6" box="[631,803,2025,2051]" pageId="2" pageNumber="155">Xinhui District</collectingMunicipality>
,
<location id="8EE96018754DBA7B10225597FC20F7C6" LSID="urn:lsid:plazi:treatment:039F87D5754FBA7B1387550AFCE9F7ED:8EE96018754DBA7B10225597FC20F7C6" box="[813,984,2025,2051]" country="China" latitude="22.230476" longLatPrecision="1" longitude="113.00271" municipality="Xinhui District" name="Maji Reservoir" pageId="2" pageNumber="155">Maji Reservoir</location>
, elev. ca.
<quantity id="4CCE9B26754DBA7B174C5594FB73F7C1" box="[1091,1163,2026,2052]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="155" unit="m" value="110.0">
<elevation id="001BD1F0754DBA7B174C5594FB73F7C1" box="[1091,1163,2026,2052]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="155" unit="m" value="110.0">110 m</elevation>
</quantity>
,
<geoCoordinate id="EE025004754DBA7B17995594FAA9F7C1" box="[1174,1361,2026,2052]" degrees="22" direction="north" minutes="13" orientation="latitude" pageId="2" pageNumber="155" precision="1" seconds="49.71" value="22.230476">22° 13 49.71”N</geoCoordinate>
,
<geoCoordinate id="EE025004754DBA7B16545594FF21F7E2" degrees="113" direction="east" minutes="0" orientation="longitude" pageId="2" pageNumber="155" precision="1" seconds="9.76" value="113.00271">113° 0 9.76”E</geoCoordinate>
,
<date id="FF881003754DBA7B13EA5A70FE9FF7ED" box="[229,359,2061,2088]" pageId="2" pageNumber="155" value="2019-07-04">
<collectingDate id="EFCCE9EB754DBA7B13EA5A70FE9FF7ED" box="[229,359,2061,2088]" pageId="2" pageNumber="155" value="2019-07-04">4 July 2019</collectingDate>
</date>
<emphasis id="B942EAD1754DBA7B12615A73FD80F7ED" box="[366,632,2061,2088]" italics="true" pageId="2" pageNumber="155">
<collectorName id="26C35315754DBA7B12615A73FE4EF7E2" box="[366,438,2061,2087]" pageId="2" pageNumber="155">Zhong</collectorName>
et al. GDS00739
</emphasis>
(
<collectionCode id="ED27AE06754DBA7B11875A70FD3BF7ED" box="[648,707,2062,2088]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" lsid="urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="2" pageNumber="155" type="Herbarium">SYS</collectionCode>
!,
<collectionCode id="ED27AE06754DBA7B11DC5A70FCF8F7ED" box="[723,768,2062,2088]" country="China" lsid="urn:lsid:biocol.org:col:13689" name="Nanjing Forestry University" pageId="2" pageNumber="155" type="Herbarium">NF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,175 @@
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<mods:title id="BC3C2B5A28A4AC314C16DD87D68E76B2">Cleisostoma hainanense, a new species (Orchidaceae: Epidendroideae) from Hainan, China: evidence from morphological and DNA analyses</mods:title>
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<heading id="D0F281D30B50FF87FF7DFC7AFB7CFC13" box="[136,1177,948,975]" fontSize="11" level="2" pageId="6" pageNumber="269" reason="3">
<taxonomicName id="4C054D3C0B50FF87FF7DFC7AFCCDFC13" ID-CoL="9V4W5" authority="M.Z.Huang, D.K.Liu &amp; G.S.Yang" authorityName="M. Z. Huang, D. K. Liu &amp; G. S. Yang Huang" authorityYear="2020" box="[136,808,948,975]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="269" phylum="Tracheophyta" rank="species" species="hainanense" status="sp. nov.">
<emphasis id="B971EAAD0B50FF87FF7DFC7AFE44FC12" bold="true" box="[136,417,948,975]" italics="true" pageId="6" pageNumber="269">Cleisostoma hainanense</emphasis>
M.Z.Huang, D.K.Liu &amp; G.S.Yang
</taxonomicName>
,
<emphasis id="B971EAAD0B50FF87FCC6FC7BFC6DFC13" box="[819,904,949,974]" italics="true" pageId="6" pageNumber="269">
<taxonomicNameLabel id="A24257D60B50FF87FCC6FC7BFC6DFC13" box="[819,904,949,974]" pageId="6" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(海#ḃfi兰; Fig. 2, 3)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F65340B50FF87FF7DFC12FC3AF98B" pageId="6" pageNumber="269" type="materials_examined">
<paragraph id="8BBA36BF0B50FF87FF7DFC12FEF6FBC8" blockId="6.[136,1453,988,1045]" pageId="6" pageNumber="269">
<emphasis id="B971EAAD0B50FF87FF7DFC12FF3FFC2C" bold="true" box="[136,218,988,1009]" pageId="6" pageNumber="269">Type:—</emphasis>
<collectingCountry id="F312762F0B50FF87FF2FFC12FEF3FC2F" box="[218,278,988,1010]" name="China" pageId="6" pageNumber="269">China</collectingCountry>
.
<collectingRegion id="49C1F85D0B50FF87FED5FC12FE8CFC2F" box="[288,361,988,1010]" country="China" name="Hainan" pageId="6" pageNumber="269">Hainan</collectingRegion>
: Lingshui City, Mount Diaoluoshan, tree trunks,
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,
<date id="FFBB107F0B50FF87FC7FFC12FBFBFC2C" box="[906,1054,988,1010]" pageId="6" pageNumber="269" value="2011-03-16">16 March 2011</date>
,
<emphasis id="B971EAAD0B50FF87FBDCFC12FB35FC2C" box="[1065,1232,988,1009]" italics="true" pageId="6" pageNumber="269">Huang 13070201</emphasis>
(
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&amp;
<typeStatus id="54BE881D0B50FF87FAA7FC12FA4DFC2C" box="[1362,1448,988,1009]" pageId="6" pageNumber="269" type="paratype">paratype</typeStatus>
:
<collectionCode id="ED14AE7A0B50FF87FF48FBCEFEE0FBC8" box="[189,261,1024,1045]" pageId="6" pageNumber="269">ATCH</collectionCode>
!).
</paragraph>
<paragraph id="8BBA36BF0B50FF87FF7DFB8AFB40FB5F" blockId="6.[136,1452,1092,1910]" pageId="6" pageNumber="269">
This new species is similar to
<taxonomicName id="4C054D3C0B50FF87FE21FB8AFC83FB83" authority="(Ker Gawler) Garay" authorityName="Garay Z. J. Liu" authorityYear="2400" baseAuthorityName="Ker Gawler" box="[468,870,1092,1118]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="269" phylum="Tracheophyta" rank="species" species="paniculatum">
<emphasis id="B971EAAD0B50FF87FE21FB8AFD65FB83" box="[468,640,1092,1118]" italics="true" pageId="6" pageNumber="269">C. paniculatum</emphasis>
(Ker Gawler) Garay
</taxonomicName>
, but differs in smaller plant; tauren-shaped mid-lobe with truncated tip; conical spur with inconspicuous septum; and arch-shaped back wall callus.
</paragraph>
<paragraph id="8BBA36BF0B50FF87FF48FB42FC3AF98B" blockId="6.[136,1452,1092,1910]" pageId="6" pageNumber="269">
Epiphytic herb. Stem up to
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, ca.
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in diameter, erect, producing numerous elongate roots ca.
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in diameter. Leaf-blade leathery, straight or slightly recurved, narrowly oblong, 613 ×
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, ligulate unequally bilobed, sometimes mucronate in sinus; sheath
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long. Inflorescences axillary, up to
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, 23 branched, many flowered; floral bracts ovate-triangular, ca. 1.6 ×
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. Flowers ca.
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in diameter; sepals and petals brown, margins and midvein yellowish green, lip creamy yellow; pedicel and ovary ca.
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. Dorsal sepal elliptic, concave, 3.13.5 × 2.0
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, obtuse; lateral sepals obliquely oblong, 3.84.0 ×
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, base adnate to column foot. Petals sub-oblong, ca. 3.4 ×
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; lip with lateral lobes erect, deltoid, ca. 1.6 × 1.0 mm, at the proximal horizontal edge a rather conspicuous flap adnate to column foot, apex obtuse, proximal margin with a thickened-callose; mid-lobe fleshy, tauren-shaped, at a right angle to the spur, ca. 2.0 mm long, apex truncate, with 2 basal backward subulate lobules, adaxially with a divergent ridge connected with tip and two subulate lobules, and then lowering into spur; spur straight, conical, ca. 2.0 mm, apex obtuse, interior inconspicuously septate, with a back wall callus; callus arc-shaped, slightly bilobed, top smooth and the remaining densely papillate-hairy. Column stout; anther cap truncate, incised; stipe short, broad, geniculate-folded at base; viscidium saddle-shaped, large.
</paragraph>
</subSubSection>
<subSubSection id="C31F65340B50FF87FF48F9AEFD47F91F" pageId="6" pageNumber="269" type="etymology">
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<emphasis id="B971EAAD0B50FF87FF48F9AEFEADF9A7" bold="true" box="[189,328,1632,1658]" pageId="6" pageNumber="269">Etymology:</emphasis>
—The specific epithet refers to the locality of
<collectingRegion id="49C1F85D0B50FF87FC9DF9AEFBC2F9A7" box="[872,1063,1632,1658]" country="China" name="Hainan" pageId="6" pageNumber="269">Hainan Province</collectingRegion>
where this new species has been found.
</paragraph>
<paragraph id="8BBA36BF0B50FF87FF48F966FD47F91F" blockId="6.[136,1452,1092,1910]" box="[189,674,1704,1731]" pageId="6" pageNumber="269">
<emphasis id="B971EAAD0B50FF87FF48F966FEA6F91F" bold="true" box="[189,323,1704,1730]" pageId="6" pageNumber="269">Phenology:</emphasis>
—Flowering from May to July.
</paragraph>
</subSubSection>
<subSubSection id="C31F65340B50FF87FF48F902FDF7F8AB" pageId="6" pageNumber="269" type="distribution">
<paragraph id="8BBA36BF0B50FF87FF48F902FF36F8D7" blockId="6.[136,1452,1092,1910]" pageId="6" pageNumber="269">
<emphasis id="B971EAAD0B50FF87FF48F902FEBCF93B" bold="true" box="[189,345,1740,1766]" pageId="6" pageNumber="269">Distribution:</emphasis>
—Known thus far only from Mount Wuzhishan and Mount Diaoluoshan on
<collectingRegion id="49C1F85D0B50FF87FB11F902FA42F93B" box="[1252,1447,1740,1766]" country="China" name="Hainan" pageId="6" pageNumber="269">Hainan Province</collectingRegion>
,
<collectingCountry id="F312762F0B50FF87FF7DF93EFF2AF8D7" box="[136,207,1776,1802]" name="China" pageId="6" pageNumber="269">China</collectingCountry>
.
</paragraph>
<paragraph id="8BBA36BF0B50FF87FF48F8DAFC2CF8F3" blockId="6.[136,1452,1092,1910]" box="[189,969,1812,1839]" pageId="6" pageNumber="269">
<emphasis id="B971EAAD0B50FF87FF48F8DAFEC6F8F3" bold="true" box="[189,291,1812,1838]" pageId="6" pageNumber="269">Habitat:</emphasis>
—Epiphyte on tree trunks in broad-leaved evergreen forests.
</paragraph>
<paragraph id="8BBA36BF0B50FF87FF48F8F6FDF7F8AB" blockId="6.[136,1452,1092,1910]" pageId="6" pageNumber="269">
<emphasis id="B971EAAD0B50FF87FF48F8F6FE51F88F" bold="true" box="[189,436,1848,1874]" pageId="6" pageNumber="269">Conservation status:</emphasis>
—We found only several populations around
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. It is an endemic species, known only from
<collectingRegion id="49C1F85D0B50FF87FF0BF892FE59F8AB" box="[254,444,1884,1910]" country="China" name="Hainan" pageId="6" pageNumber="269">Hainan Province</collectingRegion>
,
<collectingCountry id="F312762F0B50FF87FE3DF892FDEAF8AB" box="[456,527,1884,1910]" name="China" pageId="6" pageNumber="269">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,50 +1,51 @@
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<figureCitation id="13451DE1FFCCFFCF943A3B2B6586EE6B" box="[617,694,1854,1876]" captionStart="FIGURE 23" captionStartId="28.[136,229,1849,1871]" captionTargetBox="[226,1361,190,1825]" captionTargetId="figure-21@28.[226,1361,190,1825]" captionTargetPageId="28" captionText="FIGURE 23. Lectotype of Maranta ruiziana (Poeppig 1682 [G00302578]). Image reproduced with the consent of the Conservatoire et Jardin Botaniques de la Ville de Genève." figureDoi="http://doi.org/10.5281/zenodo.13875690" httpUri="https://zenodo.org/record/13875690/files/figure.png" pageId="25" pageNumber="158">Fig. 23</figureCitation>
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@ -112,7 +113,7 @@ on protologue) (
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:—Peruvia: Ad Chicoplaya anno 1798 leg. Pavon (Herb. Berolin.). In Peruvia subandina Versurae leg. Poeppig n. 1682 (Herb. Berolin., Vindob., Hort. et Acad. Petrop. Turczaninow).
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<emphasis id="B90ADD76FFCFFFCC96DB3BBA6639EEFC" bold="true" box="[136,265,1967,1989]" pageId="26" pageNumber="159">FIGURE 21.</emphasis>
Lectotype of
@ -124,7 +125,7 @@ Lectotype of
[BR0000013326272]). Image reproduced with the consent of the Botanic Garden Meise.
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<emphasis id="B90ADD76FFCEFFCD96DB3B8E6636EE88" bold="true" box="[136,262,1947,1969]" pageId="27" pageNumber="160">FIGURE 22.</emphasis>
Lectotype of
@ -136,7 +137,7 @@ Lectotype of
[U0003919]). Image reproduced with the consent of the Nationaal natuurhistorisch museum Naturalis.
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<figureCitation id="13451DE1FFD4FFD7940E3BFA65A6E13D" box="[605,662,2031,2053]" captionStart="FIGURE 5" captionStartId="6.[136,229,1972,1994]" captionTargetBox="[238,1350,190,1943]" captionTargetId="figure-21@6.[238,1350,190,1947]" captionTargetPageId="6" captionText="FIGURE 5. Lectotype of Maranta cristata (Wied-Neuwied s.n. [BR5185863]). Image reproduced with the consent of the Botanic Garden Meise." figureDoi="http://doi.org/10.5281/zenodo.13875642" httpUri="https://zenodo.org/record/13875642/files/figure.png" pageId="1" pageNumber="134">Fig. 5</figureCitation>
)
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@ -106,7 +107,7 @@ a)
(Ker-Gawler) Steudel
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<caption id="DF0151ECFFD7FFD496DB3AE06520EE16" ID-DOI="http://doi.org/10.5281/zenodo.13875630" ID-Zenodo-Dep="13875630" httpUri="https://zenodo.org/record/13875630/files/figure.png" pageId="2" pageNumber="135" startId="2.[136,229,1781,1803]" targetBox="[239,1349,205,1750]" targetPageId="2" targetType="figure">
<paragraph id="8BC10164FFD7FFD496DB3AE06520EE16" blockId="2.[136,1451,1781,1839]" pageId="2" pageNumber="135">
<emphasis id="B90ADD76FFD7FFD496DB3AE06630EE32" bold="true" box="[136,256,1781,1803]" pageId="2" pageNumber="135">FIGURE 1.</emphasis>
Lectotype of
@ -118,7 +119,7 @@ Lectotype of
[G00168760]). Image reproduced with the consent of the Conservatoire et Jardin Botaniques de la Ville de Genève.
</paragraph>
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<caption id="DF0151ECFFD6FFD596DB3BBB67CAEED1" pageId="3" pageNumber="136" startId="3.[136,229,1966,1988]" targetBox="[229,1358,190,1933]" targetPageId="3" targetType="figure">
<caption id="DF0151ECFFD6FFD596DB3BBB67CAEED1" ID-DOI="http://doi.org/10.5281/zenodo.13875634" ID-Zenodo-Dep="13875634" httpUri="https://zenodo.org/record/13875634/files/figure.png" pageId="3" pageNumber="136" startId="3.[136,229,1966,1988]" targetBox="[229,1358,190,1933]" targetPageId="3" targetType="figure">
<paragraph id="8BC10164FFD6FFD596DB3BBB67CAEED1" blockId="3.[136,1451,1966,2025]" pageId="3" pageNumber="136">
<emphasis id="B90ADD76FFD6FFD596DB3BBB6631EEFC" bold="true" box="[136,257,1966,1988]" pageId="3" pageNumber="136">FIGURE 2.</emphasis>
Lectotype of
@ -130,7 +131,7 @@ Lectotype of
[C10014449]). Image reproduced with the consent of the Københavns Universitet.
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<caption id="DF0151ECFFD1FFD296DB3BB96627EEDE" pageId="4" pageNumber="137" startId="4.[136,229,1964,1986]" targetBox="[209,1378,190,1938]" targetPageId="4" targetType="figure">
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<paragraph id="8BC10164FFD1FFD296DB3BB96627EEDE" blockId="4.[136,1452,1964,2023]" pageId="4" pageNumber="137">
<emphasis id="B90ADD76FFD1FFD296DB3BB96630EEF9" bold="true" box="[136,256,1964,1986]" pageId="4" pageNumber="137">FIGURE 3.</emphasis>
Lectotype of
@ -142,7 +143,7 @@ Lectotype of
[K00056892]). Image reproduced with the consent of the Royal Botanic Gardens, Kew.
</paragraph>
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<caption id="DF0151ECFFD0FFD396DB3BA4665BEED3" pageId="5" pageNumber="138" startId="5.[136,229,1969,1991]" targetBox="[228,1359,190,1943]" targetPageId="5" targetType="figure">
<caption id="DF0151ECFFD0FFD396DB3BA4665BEED3" ID-DOI="http://doi.org/10.5281/zenodo.13875638" ID-Zenodo-Dep="13875638" httpUri="https://zenodo.org/record/13875638/files/figure.png" pageId="5" pageNumber="138" startId="5.[136,229,1969,1991]" targetBox="[228,1359,190,1943]" targetPageId="5" targetType="figure">
<paragraph id="8BC10164FFD0FFD396DB3BA4665BEED3" blockId="5.[136,1451,1969,2027]" pageId="5" pageNumber="138">
<emphasis id="B90ADD76FFD0FFD396DB3BA46632EEFE" bold="true" box="[136,258,1969,1991]" pageId="5" pageNumber="138">FIGURE 4.</emphasis>
Lectotype of
@ -154,7 +155,7 @@ Lectotype of
[S09-21806]). Image reproduced with the consent of the Naturhistoriska riksmuseet, Stockholm.
</paragraph>
</caption>
<caption id="DF0151ECFFD3FFD096DB3BA167FAEED6" pageId="6" pageNumber="139" startId="6.[136,229,1972,1994]" targetBox="[238,1350,190,1943]" targetPageId="6" targetType="figure">
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<paragraph id="8BC10164FFD3FFD096DB3BA167FAEED6" blockId="6.[136,1452,1972,2030]" pageId="6" pageNumber="139">
<emphasis id="B90ADD76FFD3FFD096DB3BA167CDEEF1" bold="true" box="[136,253,1972,1994]" pageId="6" pageNumber="139">FIGURE 5.</emphasis>
Lectotype of
@ -180,7 +181,7 @@ Lectotype of
[illustration] published in
<bibRefCitation id="EFEF7C95FFD2FFD1947D3CF265D2E9C4" author="Ker-Gawler, J. B." box="[558,738,231,253]" pageId="7" pageNumber="140" refId="ref7077" refString="Ker-Gawler, J. B. (1824) Maranta bicolor. Edwards's Botanical Register 10: 786." type="journal volume" year="1824">Ker-Gawler (1824</bibRefCitation>
: tab 786). (
<figureCitation id="13451DE1FFD2FFD1951E3CF264B7E9C5" box="[845,903,231,253]" captionStart="FIGURE 6" captionStartId="7.[136,229,2048,2070]" captionTargetBox="[243,1343,1359,2023]" captionTargetId="figure-505@7.[243,1344,1359,2024]" captionTargetPageId="7" captionText="FIGURE 6. Image of the holotype of Maranta bicolor published in Ker-Gawler (1824: Tab. 786 [illustration])." pageId="7" pageNumber="140">Fig. 6</figureCitation>
<figureCitation id="13451DE1FFD2FFD1951E3CF264B7E9C5" box="[845,903,231,253]" captionStart="FIGURE 6" captionStartId="7.[136,229,2048,2070]" captionTargetBox="[243,1343,1359,2023]" captionTargetId="figure-505@7.[243,1344,1359,2024]" captionTargetPageId="7" captionText="FIGURE 6. Image of the holotype of Maranta bicolor published in Ker-Gawler (1824: Tab. 786 [illustration])." figureDoi="http://doi.org/10.5281/zenodo.13875646" httpUri="https://zenodo.org/record/13875646/files/figure.png" pageId="7" pageNumber="140">Fig. 6</figureCitation>
)
<emphasis id="B90ADD76FFD2FFD196DB3D1E667AE819" bold="true" box="[136,330,267,289]" pageId="7" pageNumber="140">Protologue citation</emphasis>
:—“
@ -225,7 +226,7 @@ Pellegrini
), and published in
<bibRefCitation id="EFEF7C95FFD2FFD1921D3DAA63E8E8EC" author="Vellozo, J. M. C." box="[1102,1240,447,469]" pageId="7" pageNumber="140" refId="ref8429" refString="Vellozo, J. M. C. (1831 [1827]) Petro nomine ac imperio primo brasiliensis imperii perpetuo defensore imo fundatore scientiarum artium litterarumque patrono et cultore Jubente florae fluminensis icones nunc primo eduntur. Vol. 1. Lithogr. Senefelder, Parisiis [Paris], 153 pp. https: // doi. org / 10.5962 / bhl. title. 70380" type="book" year="1831">Vellozo (1831</bibRefCitation>
, Fl. Flumin. Icones 1: tab. 16). (
<figureCitation id="13451DE1FFD2FFD197493DF66664E8C0" box="[282,340,483,505]" captionStart="FIGURE 7" captionStartId="8.[136,229,1937,1959]" captionTargetBox="[235,1348,190,1909]" captionTargetId="figure-21@8.[235,1353,190,1912]" captionTargetPageId="8" captionText="FIGURE 7. Image of the lectotype of Thalia colorata published in Vellozo (1831: Tab 16 [illustration])." pageId="7" pageNumber="140">Fig. 7</figureCitation>
<figureCitation id="13451DE1FFD2FFD197493DF66664E8C0" box="[282,340,483,505]" captionStart="FIGURE 7" captionStartId="8.[136,229,1937,1959]" captionTargetBox="[235,1348,190,1909]" captionTargetId="figure-21@8.[235,1353,190,1912]" captionTargetPageId="8" captionText="FIGURE 7. Image of the lectotype of Thalia colorata published in Vellozo (1831: Tab 16 [illustration])." figureDoi="http://doi.org/10.5281/zenodo.13875648" httpUri="https://zenodo.org/record/13875648/files/figure.png" pageId="7" pageNumber="140">Fig. 7</figureCitation>
)
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@ -115,7 +116,7 @@ s.n.
(
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@ -89,7 +90,7 @@ s.n.
,
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<figureCitation id="13451DE1FFDCFFDF94503B746579EE4E" box="[515,585,1889,1911]" captionStart="FIGURE 12" captionStartId="14.[136,229,1935,1957]" captionTargetBox="[232,1355,190,1904]" captionTargetId="figure-21@14.[232,1355,190,1910]" captionTargetPageId="14" captionText="FIGURE 12. Lectotype of Maranta friedrichsthaliana (Ørsted s.n. [C10014454]). Image reproduced with the consent of the Københavns Universitet." figureDoi="http://doi.org/10.5281/zenodo.13875662" httpUri="https://zenodo.org/record/13875662/files/figure.png" pageId="9" pageNumber="142">Fig. 12</figureCitation>
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@ -110,7 +111,7 @@ Remaining
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in monte Aquacate leg. Friedrichsthal n. l269 (Herb. Vindob.), et Oersted (Herb. Oersted).”
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<emphasis id="B90ADD76FFDFFFDC96DB3B8567CEEE9D" bold="true" box="[136,254,1936,1958]" pageId="10" pageNumber="143">FIGURE 8.</emphasis>
Lectotype of
@ -122,7 +123,7 @@ Lectotype of
[K000586896]). Image reproduced with the consent of the Royal Botanic Gardens, Kew.
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<emphasis id="B90ADD76FFDEFFDD96DB3B1367CDEE23" bold="true" box="[136,253,1798,1820]" pageId="11" pageNumber="144">FIGURE 9.</emphasis>
Image of the lectotype of
@ -134,7 +135,7 @@ published in
: Tab. 27 [illustration]).
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<paragraph id="8BC10164FFD9FFDA96DB3AE863FBEE2B" blockId="12.[136,1227,1789,1811]" box="[136,1227,1789,1811]" pageId="12" pageNumber="145">
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Image of the lectotype of
@ -146,7 +147,7 @@ published in
: Tab. 28 [illustration]).
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<caption id="DF0151ECFFD8FFDB96DB3B826670EEEA" ID-DOI="http://doi.org/10.5281/zenodo.13875658" ID-Zenodo-Dep="13875658" httpUri="https://zenodo.org/record/13875658/files/figure.png" pageId="13" pageNumber="146" startId="13.[136,229,1943,1965]" targetBox="[242,1345,190,1919]" targetPageId="13" targetType="figure">
<paragraph id="8BC10164FFD8FFDB96DB3B826670EEEA" blockId="13.[136,1452,1943,2002]" pageId="13" pageNumber="146">
<emphasis id="B90ADD76FFD8FFDB96DB3B82663CEE95" bold="true" box="[136,268,1943,1965]" pageId="13" pageNumber="146">FIGURE 11.</emphasis>
Lectotype of
@ -158,7 +159,7 @@ Lectotype of
[LE01041379]). Image reproduced with the consent of the V. L. Komarov Botanical Institute.
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<paragraph id="8BC10164FFDBFFD896DB3B9A67CAEEF1" blockId="14.[136,1451,1934,1993]" pageId="14" pageNumber="147">
<emphasis id="B90ADD76FFDBFFD896DB3B9A6639EE9C" bold="true" box="[136,265,1935,1957]" pageId="14" pageNumber="147">FIGURE 12.</emphasis>
Lectotype of

View file

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Phylogenetic tree obtained by maximum-likelihood analysis of the combined matrix. Numbers near the nodes are bootstrap percentages and Bayesian posterior probabilities (BP, BP, PP). A dash () indicates a node is inconsistent between the topology of the ML MP ML/MP and the Bayesian trees. An asterisk (*) indicates a node is 100 bootstrap percentage or 1.00 posterior probability.

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<mods:affiliation id="23A690D971679F0CA8B5A8339A108337">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica de Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="3EB0B88AE5F517086DF650EAB47E2F1C" type="personal">
<mods:role id="DC2F8773DC8DF3829430D5CF2A51DA4C">
<mods:roleTerm id="0420D93311DB120B7D200746543FD704">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9495DF5D80D15655F0C00520147128D9">Persson, Claes</mods:namePart>
<mods:affiliation id="D0E309FC5FC7EA7051226D23191FE9E7">Department of Biological and Environmental Sciences, University of Gothenburg, P. O. Box 461, 40530 Göteborg, Sweden</mods:affiliation>
</mods:name>
<mods:name id="D39AB126799D9FE8AF55F25E4DD158D4" type="personal">
<mods:role id="E2EF7CD469BFEC0AE81DC2FD4E533388">
<mods:roleTerm id="475543697C3A46B8535C51C74C9CE3B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC6AFA8B353068983D6CEF720121CB8D">Zapata, Nicolás</mods:namePart>
<mods:affiliation id="447A613BCD5F028655D91F6CEB60B218">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica de Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="DF343D5C43920D947B26BAD25EA4E85B" type="personal">
<mods:role id="E4646208FD23C008CF94C3B5953E4105">
<mods:roleTerm id="ABB5376551F08913CF2441D94AB85E46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAE627E6CA5D323C3338A19A2BC92A43">Mashburn, Brock</mods:namePart>
<mods:affiliation id="8BE92A1548556ABECC8CD4A1C1018813">Division of Biology and Biomedical Sciences, Washington University in St. Louis, St. Louis, Missouri 63130, USA</mods:affiliation>
</mods:name>
<mods:name id="5CD99960CE5A8E452082E38F577F6C58" type="personal">
<mods:role id="33BAB38A1A166DDA82DCB6CDD11E95CD">
<mods:roleTerm id="30775BABD2013EDD99719C2633F10F6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F9336EB980A9964B37C75DF46FD7EF5">Vázquez-García, J. Antonio</mods:namePart>
</mods:name>
<mods:typeOfResource id="4073D25DED5B9C5E227B8F6188840401">text</mods:typeOfResource>
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<mods:titleInfo id="8AD8A988D275F65E96BE39BBFEDE5876">
<mods:title id="D316E7D701D5439DA1EC3BA3B033F30C">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="84AD6F5AE4B47BE1B944E9E407F22269">
<mods:date id="5D43FD29551D650EDD7AF2C051DB484E">2020</mods:date>
<mods:detail id="CB5A021FFD82CF19CC3D023460B5B8E4" type="pubDate">
<mods:number id="7189F949F51B5F3ADF917FB64B88B944">2020-01-14</mods:number>
</mods:detail>
<mods:detail id="3CD1D7CAFF6751BFB746B6CA67701C65" type="volume">
<mods:number id="4B27C2DD2F50283B68C577CEA289D5A7">428</mods:number>
</mods:detail>
<mods:detail id="AF8263E0987A770185ABDD6BB2BDA9B5" type="issue">
<mods:number id="B9048591ABA5AA3C25CF2A956168DF86">3</mods:number>
</mods:detail>
<mods:extent id="C30620E5D549A7FF45BBA4079B267496" unit="page">
<mods:start id="BC28A322F7D4AA2436BEDF9ED13AE0C0">255</mods:start>
<mods:end id="829F0233651287E0920F4301A863E741">262</mods:end>
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<mods:location id="EC74274B0CC3EC511CA95281E7114BCD">
<mods:url id="D267534FB069A47B4DE4001C134483C0">http://dx.doi.org/10.11646/phytotaxa.428.3.6</mods:url>
</mods:location>
<mods:classification id="BC9D63B20EB7446CB6E562053698A4DB">journal article</mods:classification>
<mods:identifier id="E05A49A42347E08F5D6F1729CD3AE410" type="DOI">10.11646/phytotaxa.428.3.6</mods:identifier>
<mods:identifier id="F66E98C6C9707D4D38B92787FACCF045" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="1210CDC34B6E2498633A5D60BE815624" type="Zenodo-Dep">13875827</mods:identifier>
</mods:mods>
<treatment id="1874C705FFD37603FF40FDEAFC0EFC9B" LSID="urn:lsid:plazi:treatment:1874C705FFD37603FF40FDEAFC0EFC9B" httpUri="http://treatment.plazi.org/id/1874C705FFD37603FF40FDEAFC0EFC9B" lastPageId="6" lastPageNumber="261" pageId="3" pageNumber="258">
<subSubSection id="D8C72598FFD37606FF40FDEAFD1CFDC8" box="[136,761,564,591]" pageId="3" pageNumber="258" type="nomenclature">
<paragraph id="90627613FFD37606FF40FDEAFD1CFDC8" blockId="3.[136,761,564,591]" box="[136,761,564,591]" pageId="3" pageNumber="258">
<heading id="CB2AC17FFFD37606FF40FDEAFD1CFDC8" box="[136,761,564,591]" fontSize="11" level="2" pageId="3" pageNumber="258" reason="3">
<taxonomicName id="57DD0D90FFD37606FF40FDEAFD11FDC8" ID-CoL="6QTJW" authority="Vazquez &amp; Castro-Arce (2012: 114)" authorityName="Vazquez &amp; Castro-Arce" authorityPageNumber="114" authorityYear="2012" box="[136,756,564,591]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD37606FF40FDEAFEB6FDC9" bold="true" box="[136,339,564,591]" italics="true" pageId="3" pageNumber="258">Magnolia lozanoi</emphasis>
Vázquez &amp; Castro-Arce (2012: 114)
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8C72598FFD37606FF75FD86FD01F900" pageId="3" pageNumber="258" type="materials_examined">
<paragraph id="90627613FFD37606FF75FD86FEDBFD58" blockId="3.[136,1452,600,2067]" pageId="3" pageNumber="258">
<materialsCitation id="20B57C4EFFD37606FF75FD86FEDFFD58" collectingDate="1993-07-25" collectionCode="WIS, MO, F, QCNE" collectorName="Reserva Biologica El Quimi" country="Ecuador" elevation="2090" latitude="-3.45" location="Gualaquiza" longLatPrecision="1307" longitude="-78.36667" municipality="Cordillera del Condor" pageId="3" pageNumber="258" specimenCount="2" stateProvince="Morona-Santiago" typeStatus="holotype">
Type:—
<collectingCountry id="E8CA3683FFD37606FED0FD86FE41FDF4" box="[280,420,600,626]" name="Ecuador" pageId="3" pageNumber="258">ECUADOR</collectingCountry>
.
<collectingRegion id="5219B8F1FFD37606FE7DFD86FD9BFDF4" box="[437,638,600,626]" country="Ecuador" name="Morona-Santiago" pageId="3" pageNumber="258">Morona-Santiago</collectingRegion>
: Gualaquiza Cantón, Campamento Achupalla (
<collectorName id="3D2813C5FFD37606FB65FD86FF31FD10" pageId="3" pageNumber="258">Reserva Biológica El Quimi</collectorName>
),
<collectingMunicipality id="7006EC69FFD37606FF2EFDA2FE39FD10" box="[230,476,636,662]" pageId="3" pageNumber="258">Cordillera del Cóndor</collectingMunicipality>
,
<quantity id="5725DBF6FFD37606FE2FFDA2FDC8FD10" box="[487,557,636,663]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="258" unit="km" value="15.0">15 km</quantity>
east of
<location id="950220C8FFD37606FD4AFDA2FCE1FD10" LSID="urn:lsid:plazi:treatment:1874C705FFD37603FF40FDEAFC0EFC9B:950220C8FFD37606FD4AFDA2FCE1FD10" box="[642,772,636,662]" country="Ecuador" latitude="-3.45" longLatPrecision="1307" longitude="-78.36667" municipality="Cordillera del Condor" name="Gualaquiza" pageId="3" pageNumber="258" stateProvince="Morona-Santiago">Gualaquiza</location>
, open bromeliad sward with scattered shrubs and small trees,
<quantity id="5725DBF6FFD37606FF40FD7EFF3AFD3D" box="[136,223,672,699]" metricMagnitude="3" metricUnit="m" metricValue="2.09" pageId="3" pageNumber="258" unit="m" value="2090.0">
<elevation id="1BF09120FFD37606FF40FD7EFF3AFD3D" box="[136,223,672,699]" metricMagnitude="3" metricUnit="m" metricValue="2.09" pageId="3" pageNumber="258" unit="m" value="2090.0">2090 m</elevation>
</quantity>
,
<geoCoordinate id="F5E910D4FFD37606FF22FD7EFEDFFD3C" box="[234,314,672,698]" degrees="3" direction="south" minutes="27" orientation="latitude" pageId="3" pageNumber="258" precision="925" value="-3.45">3°27´S</geoCoordinate>
,
<geoCoordinate id="F5E910D4FFD37606FE8EFD7EFE48FD3C" box="[326,429,672,698]" degrees="78" direction="west" minutes="22" orientation="longitude" pageId="3" pageNumber="258" precision="925" value="-78.36667">78°22´W</geoCoordinate>
,
<date id="E46350D3FFD37606FE70FD7EFDDCFD3C" box="[440,569,672,699]" pageId="3" pageNumber="258" value="1993-07-25">
<collectingDate id="F427A93BFFD37606FE70FD7EFDDCFD3C" box="[440,569,672,699]" pageId="3" pageNumber="258" value="1993-07-25">25 Jul 1993</collectingDate>
</date>
(fl bud),
<emphasis id="A2A9AA01FFD37606FD6AFD7EFCD9FD3C" box="[674,828,672,698]" italics="true" pageId="3" pageNumber="258">Gentry 80443</emphasis>
(
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:
<collectionCode id="F6CCEED6FFD37606FC74FD7EFC1DFD3C" box="[956,1016,672,698]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15628" lsid="urn:lsid:biocol.org:col:15628" name="University of Wisconsin" pageId="3" pageNumber="258" type="Herbarium">WIS</collectionCode>
!;
<typeStatus id="4F66C8B1FFD37606FBCFFD7EFB82FD3C" box="[1031,1127,672,698]" pageId="3" pageNumber="258" type="isotype">isotypes</typeStatus>
:
<collectionCode id="F6CCEED6FFD37606FBB9FD7EFB44FD3D" box="[1137,1185,672,699]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="258" type="Herbarium">MO</collectionCode>
!-6143527,
<collectionCode id="F6CCEED6FFD37606FAE8FD7EFAD4FD3C" box="[1312,1329,672,698]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="3" pageNumber="258" type="Herbarium">F</collectionCode>
!-2140379,
<collectionCode id="F6CCEED6FFD37606FF40FD1AFF3EFD58" box="[136,219,708,734]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="3" pageNumber="258" type="Herbarium">QCNE</collectionCode>
!-92267)
</materialsCitation>
.
</paragraph>
<paragraph id="90627613FFD37606FF75FD37FF57FB7D" blockId="3.[136,1452,600,2067]" pageId="3" pageNumber="258">
Trees
<quantity id="5725DBF6FFD37606FEC9FD36FEB3FC85" box="[257,342,744,771]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="3" pageNumber="258" unit="m" value="7.5" valueMax="12.0" valueMin="3.0">312 m</quantity>
tall, up to
<quantity id="5725DBF6FFD37606FE05FD36FDF7FC85" box="[461,530,744,771]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="258" unit="cm" value="12.0">12 cm</quantity>
dbh; bark gray with fine longitudinal fissures; twig internodes
<quantity id="5725DBF6FFD37606FB1BFD36FA89FC85" box="[1235,1388,744,771]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="11.0" metricValueMin="3.5" pageId="3" pageNumber="258" unit="cm" value="0.725" valueMax="1.1" valueMin="0.35">0.351.10 cm</quantity>
long,
<quantity id="5725DBF6FFD37606FF40FCD2FEC6FCA1" box="[136,291,780,807]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="3" pageNumber="258" unit="cm" value="0.55" valueMax="0.8" valueMin="0.3">0.300.80 cm</quantity>
in diameter, densely creamy yellowish pubescent. Leaves with petioles 0.71.3 ×
<quantity id="5725DBF6FFD37606FB1FFCD2FABCFCA1" box="[1239,1369,780,807]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="258" unit="cm" value="0.2" valueMax="0.3" valueMin="0.1">0.10.3 cm</quantity>
, terete, pubescent to glabrous, blades elliptic to obovate, 4.59.5 × 2.55.0 cm, apex slightly acuminate, base cuneate, margin entire, adaxially with hairs along the midvein, abaxially appressed-pilose, densely creamy yellowish pubescent, mature leaves with black spots or completely black below caused by a fungus, lateral veins 1518 on each side of midvein, inconspicuous, midvein impressed above, prominent beneath, stipules 3.07.0 ×
<quantity id="5725DBF6FFD37606FBCBFC42FB67FC31" box="[1027,1154,924,951]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="258" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 cm</quantity>
, densely creamy yellowish pubescent, darker hairs on stem. Flowers 5.06.0 cm in diam., terminal, solitary, fragrant, peduncle
<quantity id="5725DBF6FFD37606FB25FC1EFA8EFC5D" box="[1261,1387,960,987]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="258" unit="cm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 cm</quantity>
long, bud ellipsoid, 1.92.7 ×
<quantity id="5725DBF6FFD37606FE69FC3AFDC1FC79" box="[417,548,996,1023]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="3" pageNumber="258" unit="cm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 cm</quantity>
, hypsophylls 3, persistent, ovate-oblong, whitish green, abaxially covered with creamy yellowish pubescence, 2.32.7 × 1.0
<quantity id="5725DBF6FFD37606FD4AFBD6FD2AFBA5" box="[642,719,1032,1059]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="258" unit="cm" value="1.3">1.3 cm</quantity>
, sepals 3, white, ovate-oblong and gradually narrower towards the base, glabrous, apex obtuse to rounded, base rounded, 2.73.0 ×
<quantity id="5725DBF6FFD37606FCABFBF2FC01FBC1" box="[867,996,1068,1095]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="258" unit="cm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 cm</quantity>
, petals 6, white, navicular, ovate-oblong and gradually narrower towards the base, the three outer ones 2.62.8 ×
<quantity id="5725DBF6FFD37606FC7DFB8EFBD0FBED" box="[949,1077,1104,1131]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="258" unit="cm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 cm</quantity>
, inner petals 2.42.6 ×
<quantity id="5725DBF6FFD37606FAF4FB8EFA42FBED" box="[1340,1447,1104,1131]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="258" unit="cm" value="1.05" valueMax="1.1" valueMin="1.0">11.1 cm</quantity>
, stamens 5072, ca.
<quantity id="5725DBF6FFD37606FEADFBAAFE54FB09" box="[357,433,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="cm" value="0.8">0.8 cm</quantity>
long, the connective
<quantity id="5725DBF6FFD37606FD6AFBAAFD0BFB09" box="[674,750,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="258" unit="cm" value="0.4">0.4 cm</quantity>
long, brittle and entangled, gynoecium, 1.01.2 ×
<quantity id="5725DBF6FFD37606FAEFFBAAFA42FB09" box="[1319,1447,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="258" unit="cm" value="0.6" valueMax="0.7" valueMin="0.5">0.50.7 cm</quantity>
, elliptic, densely tomentose, stigma
<quantity id="5725DBF6FFD37606FDD0FB46FD80FB35" box="[536,613,1176,1203]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="258" unit="cm" value="0.2">0.2 cm</quantity>
long, whitish green. Fruits globose, green with densely creamy yellowish pubescent, 2.02.5 ×
<quantity id="5725DBF6FFD37606FE48FB62FDE7FB51" box="[384,514,1212,1239]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="3" pageNumber="258" unit="cm" value="1.9" valueMax="2.3" valueMin="1.5">1.52.3 cm</quantity>
, with 28 carpels, seeds 12 per carpel, sub-prismatic, angled, 0.70.8 ×
<quantity id="5725DBF6FFD37606FA9FFB62FF4BFB7D" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="258" unit="cm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 cm</quantity>
.
</paragraph>
<paragraph id="90627613FFD37606FF75FADAFD01F900" blockId="3.[136,1452,600,2067]" pageId="3" pageNumber="258">
<emphasis id="A2A9AA01FFD37606FF75FADAFD45FA98" bold="true" box="[189,672,1284,1310]" pageId="3" pageNumber="258">Eponymy, discovery and ethnobotany:—</emphasis>
This species is named after Gustavo Lozano-Contreras, an outstanding Colombian scholar and leading authority for decades in the systematics of Neotropical
<taxonomicName id="57DD0D90FFD37606FBA3FAF6FAE9FAC4" authorityName="de Jussieu" authorityYear="1789" box="[1131,1292,1320,1346]" class="Magnoliopsida" family="Magnoliaceae" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="family">Magnoliaceae</taxonomicName>
. In his career, he described 34 new taxa in
<taxonomicName id="57DD0D90FFD37606FE18FA92FD88FAE0" authorityName="de Jussieu" authorityYear="1789" box="[464,621,1356,1382]" class="Magnoliopsida" family="Magnoliaceae" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="family">Magnoliaceae</taxonomicName>
(
<bibRefCitation id="F44C0BE2FFD37606FDB4FA92FC69FAE0" author="Lozano-Contreras, G." box="[636,908,1356,1382]" pageId="3" pageNumber="258" refId="ref3820" refString="Lozano-Contreras, G. (1994) Dugandiodendron y Talauma (Magnoliaceae) en el Neotropico. Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, Bogota, 147 pp." type="book" year="1994">Lozano-Contreras 1994</bibRefCitation>
,
<bibRefCitation id="F44C0BE2FFD37606FC50FA92FBA3FAE0" author="Gonzalez, F." box="[920,1094,1356,1382]" pageId="3" pageNumber="258" pagination="343 - 344" refId="ref3547" refString="Gonzalez, F. (2001) Gustavo Lozano-Contreras Bogota, 22 de Mayo de 1938 - Bogota, 10 de Julio de 2000. Caldasia 23: 343 - 344." type="journal article" year="2001">González 2001</bibRefCitation>
,
<bibRefCitation id="F44C0BE2FFD37606FB9AFA92FB13FAE0" author="Parra-O., C." box="[1106,1270,1356,1382]" pageId="3" pageNumber="258" pagination="345 - 350" refId="ref4173" refString="Parra-O., C. (2001) Publicaciones de Gustavo Lozano-Contreras (1938 - 2000) durante su carrera como botanico en el Instituto de Ciencias Naturales de la Universidad Nacional de Colombia. Caldasia 23: 345 - 350." type="journal article" year="2001">Parra-O, 2001</bibRefCitation>
). The discovery of
<taxonomicName id="57DD0D90FFD37606FF6EFAAFFEFBFA0C" box="[166,286,1392,1418]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD37606FF6EFAAFFEFBFA0C" box="[166,286,1392,1418]" italics="true" pageId="3" pageNumber="258">M. lozanoi</emphasis>
</taxonomicName>
occurred in 1993 during the first scientific exploration to the Cordillera del Cóndor by researchers from Conservation International and Escuela Politécnica Nacional del
<collectingCountry id="E8CA3683FFD37606FCB0FA4AFC32FA28" box="[888,983,1428,1454]" name="Ecuador" pageId="3" pageNumber="258">Ecuador</collectingCountry>
, arriving by helicopter, with the support from the Ecuadorian Army (
<bibRefCitation id="F44C0BE2FFD37606FE04FA66FCF8FA54" author="Schulenberg, T. S. &amp; Awbrey, K." box="[460,797,1464,1490]" pageId="3" pageNumber="258" refId="ref4525" refString="Schulenberg, T. S. &amp; Awbrey, K. (1997) The Cordillera del Condor region of Ecuador and Peru: a biological assessment. RAP Working Papers 7. Conservation International, Washington, DC, 231 pp." type="book" year="1997">Schulenberg &amp; Awbrey 1997</bibRefCitation>
). Al Gentry, the leading botanist of the expedition, made two collections of
<taxonomicName id="57DD0D90FFD37606FE95FA03FE33FA70" box="[349,470,1500,1526]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD37606FE95FA03FE33FA70" box="[349,470,1500,1526]" italics="true" pageId="3" pageNumber="258">M. lozanoi</emphasis>
</taxonomicName>
. In the 26 years since these first collections, the species was not collected again until recently by the first author and colleagues (
<figureCitation id="08E66A96FFD37606FDA1F9DEFD53F99C" box="[617,694,1536,1562]" captionStart="FIGURE 1" captionStartId="1.[136,229,1953,1975]" captionTargetBox="[178,1409,187,1926]" captionTargetId="figure-19@1.[178,1410,187,1926]" captionTargetPageId="1" captionText="FIGURE 1. Magnolia lozanoi. A. Habit. B. Branch showing the underside of the leaves covered by a black mycelium.C. Flower in female phase. D. Flower in male phase. E. Leaf beetle (Chrysomelidae, Alticinae) foraging on the stamens and stigmas. F. Fruits. (A, B, C and F from Pérez et al. 11419, D from Pérez et al. 11421, E from Pérez et al. 11461, all QCA). Photographs by Álvaro J. Pérez." figureDoi="http://doi.org/10.5281/zenodo.13875829" httpUri="https://zenodo.org/record/13875829/files/figure.png" pageId="3" pageNumber="258">Figs. 1</figureCitation>
,
<figureCitation id="08E66A96FFD37606FD0BF9DEFD37F99C" box="[707,722,1536,1562]" captionStart="FIGURE 3" captionStartId="4.[136,229,1499,1521]" captionTargetBox="[157,1430,197,1470]" captionTargetId="figure-17@4.[151,1435,190,1474]" captionTargetPageId="4" captionText="FIGURE 3. Distribution of Magnolia lozanoi in the Cordillera del Cóndor, Morona-Santiago, southeastern Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13875835" httpUri="https://zenodo.org/record/13875835/files/figure.png" pageId="3" pageNumber="258">3</figureCitation>
,
<figureCitation id="08E66A96FFD37606FD17F9DEFD15F99C" box="[735,752,1536,1562]" captionStart="FIGURE 4" captionStartId="5.[136,229,1953,1975]" captionTargetBox="[181,1405,190,1923]" captionTargetId="figure-19@5.[181,1406,190,1929]" captionTargetPageId="5" captionText="FIGURE 4. Habitat of Magnolia lozanoi in the Cordillera del Cóndor, El Quimi Biological Reserve. A, B and C. Broad plateau of the Hollín sandstone formation, the so-called Andean “tepui”. B and C. Open tepui-like bromeliad sward and elfin forest. A and B from the photographic archive of the Escuela Politécnica Nacional, RAP 1993. C by Álvaro J. Pérez." figureDoi="http://doi.org/10.5281/zenodo.13875837" httpUri="https://zenodo.org/record/13875837/files/figure.png" pageId="3" pageNumber="258">4</figureCitation>
).
<taxonomicName id="57DD0D90FFD37606FCCBF9DEFC2CF99C" box="[771,969,1536,1562]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD37606FCCBF9DEFC2CF99C" box="[771,969,1536,1562]" italics="true" pageId="3" pageNumber="258">Magnolia lozanoi</emphasis>
</taxonomicName>
was first described by the second author of this article six years ago based on Gentrys specimens. This fact illustrates the importance of herbaria as repositories of undescribed species, particularly in connection with field exploration through RAP (
<bibRefCitation id="F44C0BE2FFD37606FB9FF996FAC9F9E4" author="Alonso, L. E. &amp; Deichmann, J. L. &amp; McKenna, S. A. &amp; Naskrecki, P. &amp; Richards, S. J." box="[1111,1324,1608,1634]" pageId="3" pageNumber="258" refId="ref3121" refString="Alonso, L. E., Deichmann, J. L., McKenna, S. A., Naskrecki, P. &amp; Richards, S. J. (Eds.) (2011) Still counting ...: biodiversity exploration for conservation - the first 20 years of the rapid assessment program. Conservation International, Arlington, 316 pp." type="book" year="2011">
Alonso
<emphasis id="A2A9AA01FFD37606FB79F997FB0CF9E4" box="[1201,1257,1608,1634]" italics="true" pageId="3" pageNumber="258">et al.</emphasis>
2011
</bibRefCitation>
). No use or vernacular names have been recorded for this species.
</paragraph>
</subSubSection>
<subSubSection id="D8C72598FFD37603FF75F94EFC0EFC9B" lastPageId="6" lastPageNumber="261" pageId="3" pageNumber="258" type="distribution">
<paragraph id="90627613FFD37606FF75F94EFA7EF794" blockId="3.[136,1452,600,2067]" pageId="3" pageNumber="258">
<emphasis id="A2A9AA01FFD37606FF75F94EFD91F92C" bold="true" box="[189,628,1680,1706]" pageId="3" pageNumber="258">Distribution, habitat and ecology:—</emphasis>
Endemic to the Cordillera del Cóndor,
<collectingCountry id="E8CA3683FFD37606FBFFF94EFB73F92C" box="[1079,1174,1680,1706]" name="Ecuador" pageId="3" pageNumber="258">Ecuador</collectingCountry>
, in El Quimi Biological Reserve in
<collectingRegion id="5219B8F1FFD37606FEC4F96AFDDBF948" box="[268,574,1716,1742]" country="Ecuador" name="Morona-Santiago" pageId="3" pageNumber="258">Morona-Santiago Province</collectingRegion>
and known only from the
<typeStatus id="4F66C8B1FFD37606FCB1F96BFC4CF949" box="[889,937,1717,1743]" pageId="3" pageNumber="258">type</typeStatus>
locality and surrounding areas (
<figureCitation id="08E66A96FFD37606FAE9F96AFA8DF948" box="[1313,1384,1716,1742]" captionStart="FIGURE 3" captionStartId="4.[136,229,1499,1521]" captionTargetBox="[157,1430,197,1470]" captionTargetId="figure-17@4.[151,1435,190,1474]" captionTargetPageId="4" captionText="FIGURE 3. Distribution of Magnolia lozanoi in the Cordillera del Cóndor, Morona-Santiago, southeastern Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13875835" httpUri="https://zenodo.org/record/13875835/files/figure.png" pageId="3" pageNumber="258">Fig. 3</figureCitation>
). The locality is a broad plateau with quartzitic sandstones of the Hollín Formation, referred to in recent literature as an Andean tepui (
<bibRefCitation id="F44C0BE2FFD37606FEE3F922FE01F890" author="Neill, D. A. &amp; Rios Paredes, M. &amp; Torres Montenengro, L. A. &amp; Mori Vargas, T. J. &amp; Vriesendorp, C." box="[299,484,1788,1815]" pageId="3" pageNumber="258" pagination="98 - 119" refId="ref4021" refString="Neill, D. A., Rios Paredes, M., Torres Montenengro, L. A., Mori Vargas, T. J. &amp; Vriesendorp, C. (2014) Vegetacion y flora / Vegetation and flora. In: Pitman, N., Vriesendorp, C., Alvira, D., Markel, J. A., Johnston, M., Ruelas Inzunza, E., Lancha Pizango, A., Sarmiento Valenzuela, G., Alvarez-Loayza, P., Homan, J., Wachter, T., del Campo, A., Stotz, D. F. &amp; Heilpern, S. (Eds.) Peru: Cordillera Escalera-Loreto. Rapid Biological and Social Inventories Report 26. The Field Museum, Chicago, pp. 98 - 119." type="book chapter" year="2014">
Neill
<emphasis id="A2A9AA01FFD37606FEA2F923FE7FF890" box="[362,410,1788,1814]" italics="true" pageId="3" pageNumber="258">et al</emphasis>
. 2014
</bibRefCitation>
). This is in reference to its similarity with the tepuis of the Guiana Shield (
<bibRefCitation id="F44C0BE2FFD37606FADDF922FA7AF891" author="Huber, O." box="[1301,1439,1788,1815]" pageId="3" pageNumber="258" refId="ref3637" refString="Huber, O. (1995) Geographical and physical features. In: Berry, P. E., Holst, B. K. &amp; Yatskievych, K. (Eds.) Flora of the Venezuelan Guayana, vol. 1: introduction. Missouri Botanical Garden, St. Louis, 62 pp." type="book" year="1995">Huber 1995</bibRefCitation>
). The habitat is an open tepui-like bromeliad sward and elfin forest at
<quantity id="5725DBF6FFD37606FC5EF8FEFBD7F8BD" box="[918,1074,1824,1851]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" pageNumber="258" unit="m" value="2050.0" valueMax="2200.0" valueMin="1900.0">19002200 m</quantity>
(
<figureCitation id="08E66A96FFD37606FB8BF8FEFB6CF8BC" box="[1091,1161,1824,1850]" captionStart="FIGURE 4" captionStartId="5.[136,229,1953,1975]" captionTargetBox="[181,1405,190,1923]" captionTargetId="figure-19@5.[181,1406,190,1929]" captionTargetPageId="5" captionText="FIGURE 4. Habitat of Magnolia lozanoi in the Cordillera del Cóndor, El Quimi Biological Reserve. A, B and C. Broad plateau of the Hollín sandstone formation, the so-called Andean “tepui”. B and C. Open tepui-like bromeliad sward and elfin forest. A and B from the photographic archive of the Escuela Politécnica Nacional, RAP 1993. C by Álvaro J. Pérez." figureDoi="http://doi.org/10.5281/zenodo.13875837" httpUri="https://zenodo.org/record/13875837/files/figure.png" pageId="3" pageNumber="258">Fig. 4</figureCitation>
). Approximately
<specimenCount id="86DBBD9AFFD37606FA87F8FEFA4EF8BC" box="[1359,1451,1824,1850]" count="20" pageId="3" pageNumber="258" type="adult">20 adult</specimenCount>
individuals were observed but no seedlings or juveniles. According to the Ministerio del Ambiente de
<collectingCountry id="E8CA3683FFD37606FB3EF89AFAB6F8D8" box="[1270,1363,1860,1886]" name="Ecuador" pageId="3" pageNumber="258">Ecuador</collectingCountry>
(2013), this locality lies within a much larger zone dominated by evergreen mountain forest of sandstone plains of Cordillera del Cóndor (bosque siempreverde montano sobre mesetas de arenisca de la Cordillera del
<taxonomicName id="57DD0D90FFD37606FBABF852FAEFF820" authority=", BsMa" authorityName="BsMa" box="[1123,1290,1932,1958]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="condor">Cóndor, BsMa</taxonomicName>
01). Regarding animal interactions, leaf beetles (
<taxonomicName id="57DD0D90FFD37606FE34F86EFD4FF84C" authorityName="Latreille" authorityYear="1802" box="[508,682,1968,1994]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
,
<taxonomicName id="57DD0D90FFD37606FD7CF86EFCF8F84C" box="[692,797,1968,1994]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="subFamily" subFamily="Alticinae">Alticinae</taxonomicName>
) were seen foraging on the stamens and stigmas, possibly acting as pollinators (
<figureCitation id="08E66A96FFD37606FEB6F80AFE27F868" box="[382,450,2004,2030]" captionStart="FIGURE 1" captionStartId="1.[136,229,1953,1975]" captionTargetBox="[178,1409,187,1926]" captionTargetId="figure-19@1.[178,1410,187,1926]" captionTargetPageId="1" captionText="FIGURE 1. Magnolia lozanoi. A. Habit. B. Branch showing the underside of the leaves covered by a black mycelium.C. Flower in female phase. D. Flower in male phase. E. Leaf beetle (Chrysomelidae, Alticinae) foraging on the stamens and stigmas. F. Fruits. (A, B, C and F from Pérez et al. 11419, D from Pérez et al. 11421, E from Pérez et al. 11461, all QCA). Photographs by Álvaro J. Pérez." figureDoi="http://doi.org/10.5281/zenodo.13875829" httpUri="https://zenodo.org/record/13875829/files/figure.png" pageId="3" pageNumber="258">Fig. 1</figureCitation>
). Another finding was the identification of a fungus (Basidiomycota) that grows on the pubescence on the underside of mature leaves, making them black, a character mentioned in the protologue (
<figureCitation id="08E66A96FFD37606FA81F826FA6FF794" box="[1353,1418,2040,2066]" captionStart="FIGURE 1" captionStartId="1.[136,229,1953,1975]" captionTargetBox="[178,1409,187,1926]" captionTargetId="figure-19@1.[178,1410,187,1926]" captionTargetPageId="1" captionText="FIGURE 1. Magnolia lozanoi. A. Habit. B. Branch showing the underside of the leaves covered by a black mycelium.C. Flower in female phase. D. Flower in male phase. E. Leaf beetle (Chrysomelidae, Alticinae) foraging on the stamens and stigmas. F. Fruits. (A, B, C and F from Pérez et al. 11419, D from Pérez et al. 11421, E from Pérez et al. 11461, all QCA). Photographs by Álvaro J. Pérez." figureDoi="http://doi.org/10.5281/zenodo.13875829" httpUri="https://zenodo.org/record/13875829/files/figure.png" pageId="3" pageNumber="258">Fig. 1</figureCitation>
).
</paragraph>
<caption id="C4A2269BFFD47601FF40FA05FB0CFA77" ID-DOI="http://doi.org/10.5281/zenodo.13875835" ID-Zenodo-Dep="13875835" box="[136,1257,1499,1521]" httpUri="https://zenodo.org/record/13875835/files/figure.png" pageId="4" pageNumber="259" startId="4.[136,229,1499,1521]" targetBox="[157,1430,197,1470]" targetPageId="4" targetType="figure">
<paragraph id="90627613FFD47601FF40FA05FB0CFA77" blockId="4.[136,1257,1499,1521]" box="[136,1257,1499,1521]" pageId="4" pageNumber="259">
<emphasis id="A2A9AA01FFD47601FF40FA05FF12FA76" bold="true" box="[136,247,1499,1521]" pageId="4" pageNumber="259">FIGURE 3</emphasis>
. Distribution of
<taxonomicName id="57DD0D90FFD47601FE5FFA05FDA7FA76" box="[407,578,1499,1520]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="259" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD47601FE5FFA05FDA7FA76" box="[407,578,1499,1520]" italics="true" pageId="4" pageNumber="259">Magnolia lozanoi</emphasis>
</taxonomicName>
in the Cordillera del
<taxonomicName id="57DD0D90FFD47601FCC6FA05FBE9FA77" authority=", Morona-Santiago" authorityName="Morona-Santiago" box="[782,1036,1499,1521]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="259" phylum="Tracheophyta" rank="species" species="condor">Cóndor, Morona-Santiago</taxonomicName>
, southeastern Ecuador.
</paragraph>
</caption>
<paragraph id="90627613FFD47601FF75F9FEFCA0F9BC" blockId="4.[136,1452,1568,2062]" box="[189,837,1568,1594]" pageId="4" pageNumber="259">
<emphasis id="A2A9AA01FFD47601FF75F9FEFEBAF9BC" bold="true" box="[189,351,1568,1594]" pageId="4" pageNumber="259">Phenology:—</emphasis>
Flowering and fruiting throughout the year.
</paragraph>
<paragraph id="90627613FFD47601FF75F99AFB88F8DC" blockId="4.[136,1452,1568,2062]" pageId="4" pageNumber="259">
<emphasis id="A2A9AA01FFD47601FF75F99AFE34F9D8" bold="true" box="[189,465,1604,1630]" pageId="4" pageNumber="259">Conservation status:—</emphasis>
According to
<bibRefCitation id="F44C0BE2FFD47601FDA7F99AFCB0F9D8" author="Rivers, M. &amp; Beech, E. &amp; Murphy, L. &amp; Oldfield, S." box="[623,853,1604,1630]" pageId="4" pageNumber="259" refId="ref4297" refString="Rivers, M., Beech, E., Murphy, L. &amp; Oldfield, S. (2016 a) The Red List of Magnoliaceae - revised and extended. BGCI, Richmond, 60 pp." type="book" year="2016">
Rivers
<emphasis id="A2A9AA01FFD47601FD09F99BFD11F9D8" box="[705,756,1604,1630]" italics="true" pageId="4" pageNumber="259">et al</emphasis>
. (2016a
</bibRefCitation>
,
<bibRefCitation id="F44C0BE2FFD47601FCA9F99AFC4EF9D8" author="Rivers, M. C. &amp; Wheeler, L. &amp; Perez Castaneda, A." box="[865,939,1604,1630]" pageId="4" pageNumber="259" refId="ref4337" refString="Rivers, M. C., Wheeler, L. &amp; Perez Castaneda, A. (2016 b) Magnolia lozanoi. The IUCN Red List of Threatened Species 2016: e. T 67513550 A 67513748. Available from: https: // doi. org / 10.2305 / IUCN. UK. 2016 - 1. RLTS. T 67513550 A 67513748. en. (accessed 27 June 2019)." type="book" year="2016">2016b</bibRefCitation>
), this species was originally assessed as data deficient (DD;
<bibRefCitation id="F44C0BE2FFD47601FEFAF9B6FE5EF904" author="IUCN, International Union &amp; Conservation of Nature &amp; Cambridge, IUCN" box="[306,443,1640,1666]" pageId="4" pageNumber="259" refId="ref3696" refString="IUCN, International Union for Conservation of Nature (2012) IUCN Red List categories and criteria, version 3.1, second edition. Gland and Cambridge, IUCN, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
). Based on these new findings, as well as the imminent threat of nearby mining activities, and following
<bibRefCitation id="F44C0BE2FFD47601FEE6F952FE23F920" author="IUCN" box="[302,454,1676,1702]" pageId="4" pageNumber="259" refId="ref3731" refString="IUCN (2017) Guidelines for using the IUCN red list categories and criteria, version 13. Prepared by the Standards and Petitions Subcommittee. Available from: http: // www. iucnredlist. org / documents / redlistGuidelines. pdf (accessed 27 June 2019)" type="book" year="2017">IUCN (2017)</bibRefCitation>
criteria, we suggest raising its status to critically endangered (CR). This extremely rare species is known only from the
<typeStatus id="4F66C8B1FFD47601FE3CF96EFDC1F94C" box="[500,548,1712,1738]" pageId="4" pageNumber="259">type</typeStatus>
locality and surrounding areas in El Quimi Biological Reserve and is currently threatened by an ongoing open-pit copper mine from the Mirador project, which has already caused deforestation of an area of about 1300 hectares (
<bibRefCitation id="F44C0BE2FFD47601FE23F926FD0AF894" author="Mazabanda, C. &amp; Kemper, R. &amp; Thieme, A. &amp; Hettler, B. &amp; Finer, M." box="[491,751,1784,1810]" pageId="4" pageNumber="259" refId="ref3906" refString="Mazabanda, C., Kemper, R., Thieme, A., Hettler, B. &amp; Finer, M. (2018) Impacts of mining project &quot; Mirador &quot; in the Ecuadorian Amazon. Available from: https: // maaproject. org / mirador-ecuador / (accessed 2 July 2019)" type="url" year="2018">
Mazabanda
<emphasis id="A2A9AA01FFD47601FDBDF927FD42F894" box="[629,679,1784,1810]" italics="true" pageId="4" pageNumber="259">et al</emphasis>
. 2018
</bibRefCitation>
,
<bibRefCitation id="F44C0BE2FFD47601FD32F926FC16F894" author="Vandegrift, R. &amp; Thomas, D. C. &amp; Roy, B. A. &amp; Levy, M." box="[762,1011,1784,1810]" pageId="4" pageNumber="259" refId="ref4568" refString="Vandegrift, R., Thomas, D. C., Roy, B. A. &amp; Levy, M. (2018) Alcance de las concesiones mineras recientes en Ecuador. Rainforest Information Center, Nimbin, New South Wales, Australia, 7 pp. Available from: https: // ecuadorendangered. com / wp-content / uploads / 2018 / 01 / RIC-Mapping-Report-v 1.1 - 20180117 - esp. pdf (accessed 2 July 2019)" type="book" year="2018">
Vandegrift
<emphasis id="A2A9AA01FFD47601FCBFF927FC4AF894" box="[887,943,1784,1810]" italics="true" pageId="4" pageNumber="259">et al.</emphasis>
2018
</bibRefCitation>
). Urgent awareness more generally and in the Ministerio del Ambiente de
<collectingCountry id="E8CA3683FFD47601FDC6F8C2FD8EF8B0" box="[526,619,1820,1846]" name="Ecuador" pageId="4" pageNumber="259">Ecuador</collectingCountry>
is needed. Financial support for a continuous protection, monitoring, and conservation programme is urgently needed for this area of great conservation relevance.
</paragraph>
<paragraph id="90627613FFD47601FF75F8BAFEE5F788" blockId="4.[136,1452,1568,2062]" pageId="4" pageNumber="259">
<emphasis id="A2A9AA01FFD47601FF75F8BAFE60F8F8" bold="true" box="[189,389,1892,1918]" pageId="4" pageNumber="259">Relationships:—</emphasis>
<taxonomicName id="57DD0D90FFD47601FE4DF8BAFDAEF8F8" box="[389,587,1892,1918]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="259" phylum="Tracheophyta" rank="species" species="lozanoi">
<emphasis id="A2A9AA01FFD47601FE4DF8BAFDAEF8F8" box="[389,587,1892,1918]" italics="true" pageId="4" pageNumber="259">Magnolia lozanoi</emphasis>
</taxonomicName>
belongs to subsection
<taxonomicName id="57DD0D90FFD47601FC83F8BAFBF2F8F8" box="[843,1047,1892,1918]" pageId="4" pageNumber="259" rank="subSection" subSection="Dugandiodendron">
<emphasis id="A2A9AA01FFD47601FC83F8BAFBF2F8F8" box="[843,1047,1892,1918]" italics="true" pageId="4" pageNumber="259">Dugandiodendron</emphasis>
</taxonomicName>
. It is morphologically similar to the Colombian species
<taxonomicName id="57DD0D90FFD47601FEACF856FBE1F824" authority="(Lozano-Contreras 1975: 33) Govaerts" authorityName="Govaerts" baseAuthorityName="Lozano-Contreras" baseAuthorityPageNumber="33" baseAuthorityYear="1975" box="[356,1028,1928,1955]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="259" phylum="Tracheophyta" rank="species" species="mahechae">
<emphasis id="A2A9AA01FFD47601FEACF856FDAFF824" box="[356,586,1928,1954]" italics="true" pageId="4" pageNumber="259">Magnolia mahechae</emphasis>
(
<bibRefCitation id="F44C0BE2FFD47601FD90F856FC6AF824" author="Lozano-Contreras, G." box="[600,911,1928,1955]" pageId="4" pageNumber="259" pagination="27 - 50" refId="ref3797" refString="Lozano-Contreras, G. (1975) Contribucion al estudio de las Magnoliaceae de Colombia. Caldasia 11: 27 - 50." type="journal article" year="1975">Lozano-Contreras 1975: 33</bibRefCitation>
) Govaerts
</taxonomicName>
in
<bibRefCitation id="F44C0BE2FFD47601FBE0F856FA99F824" author="Frodin, D. G. &amp; Govaerts, R." box="[1064,1404,1928,1954]" pageId="4" pageNumber="259" refId="ref3429" refString="Frodin, D. G. &amp; Govaerts, R. (1996) World checklist and bibliography of Magnoliaceae. Kew Publishing, Richmond, 79 pp." type="book" year="1996">Frodin &amp; Govaerts (1996: 71)</bibRefCitation>
, but differs from the latter in being shorter (312 vs.
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), having smaller obovate leaf blades (4.59.5 × 2.55.0 cm vs. elliptic 11.8 ×
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), shorter petioles (
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long vs.
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long) and sparse long black hairs on the twigs (vs. none).
</paragraph>
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<emphasis id="A2A9AA01FFD57600FF40F87FFF1CF830" bold="true" box="[136,249,1953,1975]" pageId="5" pageNumber="260">FIGURE 4</emphasis>
. Habitat of
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<emphasis id="A2A9AA01FFD57600FEA7F87FFDFEF830" box="[367,539,1953,1974]" italics="true" pageId="5" pageNumber="260">Magnolia lozanoi</emphasis>
</taxonomicName>
in the Cordillera del
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Biological Reserve. A, B and C. Broad plateau of the Hollín sandstone formation, the so-called Andean “tepui”. B and C. Open tepui-like bromeliad sward and elfin forest. A and B from the photographic archive of the Escuela Politécnica Nacional, RAP 1993. C by Álvaro J. Pérez.
</paragraph>
</caption>
<paragraph id="90627613FFD67603FF75FF40FEF2FEEA" blockId="6.[136,1452,158,797]" pageId="6" pageNumber="261">
<emphasis id="A2A9AA01FFD67603FF75FF40FDBCFF3E" bold="true" box="[189,601,158,184]" pageId="6" pageNumber="261">Additional specimens examined:—</emphasis>
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<collectingCountry id="E8CA3683FFD67603FD91FF40FD01FF3E" box="[601,740,158,184]" name="Ecuador" pageId="6" pageNumber="261">ECUADOR</collectingCountry>
.
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:
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,
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(
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),
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,
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east of
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, transect # 2,
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</quantity>
,
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,
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,
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</date>
,
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80383
</emphasis>
(
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!)
</materialsCitation>
;
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<collectorName id="3D2813C5FFD67603FD4EFF38FC8BFE86" box="[646,878,230,256]" pageId="6" pageNumber="261">Parroquia Bomboiza</collectorName>
,
<collectorName id="3D2813C5FFD67603FCBFFF39FB54FE86" box="[887,1201,230,257]" pageId="6" pageNumber="261">Reserva Biológica El Quimi</collectorName>
, sendero y alrededores entre el campamento
<collectingCounty id="79030E9FFFD67603FEBFFED4FDF1FEA2" box="[375,532,266,292]" pageId="6" pageNumber="261">Río Cristalino</collectingCounty>
y la frontera con
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,
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,
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,
<geoCoordinate id="F5E910D4FFD67603FB9AFED4FB04FEA2" box="[1106,1249,266,292]" degrees="78" direction="west" minutes="23" orientation="longitude" pageId="6" pageNumber="261" precision="15" seconds="28" value="-78.391106">78°2328”W</geoCoordinate>
,
<date id="E46350D3FFD67603FB24FED4FA42FEA2" box="[1260,1447,266,293]" pageId="6" pageNumber="261" value="2019-01-23">
<collectingDate id="F427A93BFFD67603FB24FED4FA42FEA2" box="[1260,1447,266,293]" pageId="6" pageNumber="261" value="2019-01-23">23 January 2019</collectingDate>
</date>
,
<emphasis id="A2A9AA01FFD67603FF40FEF1FEB5FECE" box="[136,336,302,329]" italics="true" pageId="6" pageNumber="261">
<location id="950220C8FFD67603FF40FEF1FF22FECF" LSID="urn:lsid:plazi:treatment:1874C705FFD37603FF40FDEAFC0EFC9B:950220C8FFD67603FF40FEF1FF22FECF" box="[136,199,303,329]" country="Ecuador" county="Rio Cristalino" latitude="-3.5180554" longLatPrecision="21" longitude="-78.391106" municipality="Peru" name="Perez" pageId="6" pageNumber="261">Pérez</location>
et al. 11419
</emphasis>
,
<emphasis id="A2A9AA01FFD67603FE95FEF0FE44FECE" box="[349,417,302,328]" italics="true" pageId="6" pageNumber="261">11421</emphasis>
(
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!)
</materialsCitation>
;
<materialsCitation id="20B57C4EFFD67603FDC5FEF0FBA8FECE" box="[525,1101,302,329]" collectingDate="2019-01-24" collectionCode="QCA" country="Ecuador" location="Perez" pageId="6" pageNumber="261" specimenCount="1">
<date id="E46350D3FFD67603FDC5FEF0FD2DFECE" box="[525,712,302,329]" pageId="6" pageNumber="261" value="2019-01-24">
<collectingDate id="F427A93BFFD67603FDC5FEF0FD2DFECE" box="[525,712,302,329]" pageId="6" pageNumber="261" value="2019-01-24">24 January 2019</collectingDate>
</date>
,
<emphasis id="A2A9AA01FFD67603FD1BFEF1FC7FFECE" box="[723,922,302,329]" italics="true" pageId="6" pageNumber="261">
<location id="950220C8FFD67603FD1BFEF1FCF7FECF" LSID="urn:lsid:plazi:treatment:1874C705FFD37603FF40FDEAFC0EFC9B:950220C8FFD67603FD1BFEF1FCF7FECF" box="[723,786,303,329]" country="Ecuador" name="Perez" pageId="6" pageNumber="261">Pérez</location>
et al. 11440
</emphasis>
,
<emphasis id="A2A9AA01FFD67603FC60FEF0FC09FECE" box="[936,1004,302,328]" italics="true" pageId="6" pageNumber="261">11461</emphasis>
(
<collectionCode id="F6CCEED6FFD67603FC33FEF0FBA5FECE" box="[1019,1088,302,328]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15650" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="6" pageNumber="261" type="Herbarium">QCA</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="20B57C4EFFD67603FB90FEF0FEF1FEEA" collectingDate="2019-01-24" collectionCode="QCA" country="Ecuador" pageId="6" pageNumber="261" specimenCount="1">
<date id="E46350D3FFD67603FB90FEF0FAF6FECE" box="[1112,1299,302,329]" pageId="6" pageNumber="261" value="2019-01-24">
<collectingDate id="F427A93BFFD67603FB90FEF0FAF6FECE" box="[1112,1299,302,329]" pageId="6" pageNumber="261" value="2019-01-24">24 January 2019</collectingDate>
</date>
,
<emphasis id="A2A9AA01FFD67603FAD6FEF1FF57FEEA" italics="true" pageId="6" pageNumber="261">
<taxonomicName id="57DD0D90FFD67603FAD6FEF1FA88FECE" authorityName="Bruner" authorityYear="1904" box="[1310,1389,303,328]" class="Insecta" family="Acrididae" genus="Zapata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">Zapata</taxonomicName>
et al. 551
</emphasis>
(
<collectionCode id="F6CCEED6FFD67603FF0AFE8CFEEDFEEA" box="[194,264,338,364]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15650" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="6" pageNumber="261" type="Herbarium">QCA</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="90627613FFD67603FF75FEA8FC0EFC9B" blockId="6.[136,1452,158,797]" pageId="6" pageNumber="261">
<emphasis id="A2A9AA01FFD67603FF75FEA8FE49FE16" bold="true" box="[189,428,374,400]" pageId="6" pageNumber="261">Acknowledgements.</emphasis>
Fieldwork was granted by the Ministry of the Environment of
<collectingCountry id="E8CA3683FFD67603FB57FEA8FB19FE16" box="[1183,1276,374,400]" name="Ecuador" pageId="6" pageNumber="261">Ecuador</collectingCountry>
(003-2019-IC- FLO-DNB/MAE). AJP,
<collectingCountry id="E8CA3683FFD67603FE55FE45FE27FE33" box="[413,450,411,437]" name="New Zealand" pageId="6" pageNumber="261">NZ</collectingCountry>
and CP express their gratitude to Wilson Paucar (director of El Quimi Reserve), Juan Carlos Fernandez (park ranger of El Quimi Reserve) and the Valle del Quimi community for their support during the fieldwork. We also thank Luis Albuja, Ana Armendariz and Ramiro Barriga from Escuela Politécnica Nacional for their valuable information and photographs of the RAP to the Cordillera del Cóndor in 1993; Robert Gradstein kindly provided image 2A and Françoise Lopez from Muséum National dHistoire naturelle, Paris for editing; Carlos E. Sarmiento editor of
<taxonomicName id="57DD0D90FFD67603FEA4FD90FE34FDEE" box="[364,465,590,616]" class="Magnoliopsida" family="Apiaceae" genus="Caldasia" kingdom="Plantae" order="Apiales" pageId="6" pageNumber="261" phylum="Tracheophyta" rank="genus">
<emphasis id="A2A9AA01FFD67603FEA4FD90FE34FDEE" box="[364,465,590,616]" italics="true" pageId="6" pageNumber="261">Caldasia</emphasis>
</taxonomicName>
kindly authorised use of image 2B; Rubén D. Jarrín E. (ComCienCia) kindly designed figures 1, 2 and 4; staff of Fungario QCAM kindly identified the fungus; Secretaría de Educación Superior, Ciencia, Tecnología e Innovación de la
<collectingCountry id="E8CA3683FFD67603FE20FD48FD0FFD36" box="[488,746,662,688]" name="Ecuador" pageId="6" pageNumber="261">República del Ecuador</collectingCountry>
for financial support (SENESCYT, Arca de Noé Initiative, S. R. Ron and O. TorresCarvajal, Principal Investigators); the Universidad de Guadalajara-CUCBA, CONACyT-SNI and PROMEP-SEP in
<collectingCountry id="E8CA3683FFD67603FE41FD00FE07FD7E" box="[393,482,734,760]" name="Mexico" pageId="6" pageNumber="261">Mexico</collectingCountry>
; the curators of F, MO, QCA, QCNE &amp; WIS herbaria for providing material for this research; anonymous reviewers and Mark Chase, subject editor of
<emphasis id="A2A9AA01FFD67603FCBAFCDCFC01FC9A" box="[882,996,770,796]" italics="true" pageId="6" pageNumber="261">Phytotaxa</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,168 @@
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<mods:classification id="E08B4452A7411B8CDE1D41C5CFA4CE98">journal article</mods:classification>
<mods:identifier id="624A79423EA96992C2CEE13D10EAB3D6" type="DOI">10.11646/phytotaxa.428.2.7</mods:identifier>
<mods:identifier id="669BEEB110CC7ABBF1A511943C964517" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="1C131894D2259738400D197F866226CB" type="Zenodo-Dep">13875795</mods:identifier>
</mods:mods>
<treatment id="9B36878EB71BFFF5CCD76688FBEAC225" ID-DOI="http://doi.org/10.5281/zenodo.13875793" ID-Zenodo-Dep="13875793" LSID="urn:lsid:plazi:treatment:9B36878EB71BFFF5CCD76688FBEAC225" httpUri="http://treatment.plazi.org/id/9B36878EB71BFFF5CCD76688FBEAC225" lastPageNumber="136" pageId="5" pageNumber="136">
<subSubSection id="5B856513B71BFFF5CCD76688FC0DC66C" box="[136,896,244,271]" pageId="5" pageNumber="136" type="nomenclature">
<paragraph id="13203698B71BFFF5CCD76688FC0DC66C" blockId="5.[136,896,244,271]" box="[136,896,244,271]" pageId="5" pageNumber="136">
<heading id="486881F4B71BFFF5CCD76688FC0DC66C" box="[136,896,244,271]" fontSize="11" level="2" pageId="5" pageNumber="136" reason="3">
<taxonomicName id="D49F4D1BB71BFFF5CCD76688FD12C66C" authority="Sheng H. Wu &amp; C.L. Wei" authorityName="Sheng H. Wu &amp; C. L. Wei" authorityYear="2020" box="[136,671,244,271]" class="Agaricomycetes" family="Hericiaceae" genus="Dentipellis" kingdom="Fungi" order="Russulales" pageId="5" pageNumber="136" phylum="Basidiomycota" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="21EBEA8AB71BFFF5CCD76688FEFAC66D" bold="true" box="[136,375,244,271]" italics="true" pageId="5" pageNumber="136">Dentipellis fimbriata</emphasis>
Sheng H. Wu &amp; C.L. Wei
</taxonomicName>
,
<emphasis id="21EBEA8AB71BFFF5CEF56689FD72C66C" box="[682,767,245,270]" italics="true" pageId="5" pageNumber="136">
<taxonomicNameLabel id="3AD857F1B71BFFF5CEF56689FD72C66C" box="[682,767,245,270]" pageId="5" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="8BA42A1DB71BFFF5CF516688FCD0C66C" box="[782,861,244,270]" captionStart="FIGURE 2" captionStartId="3.[136,229,1925,1947]" captionTargetBox="[151,1435,190,1901]" captionTargetId="figure-19@3.[151,1435,190,1901]" captionTargetPageId="3" captionText="FIGURE 2. Basidiomata of Dentipellis fimbriata (holotype, WEI 18-288). A. Basidiome margin with rhizomorphs; B. Odontioid to hydnoid hymenial surface. Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13875799" httpUri="https://zenodo.org/record/13875799/files/figure.png" pageId="5" pageNumber="136">Figs. 2</figureCitation>
,
<figureCitation id="8BA42A1DB71BFFF5CF366688FCFAC66C" box="[873,887,244,270]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[191,1383,203,1895]" captionTargetId="figure-30@4.[179,1409,190,1929]" captionTargetPageId="4" captionText="FIGURE 3. Microscopic structures of Dentipellis fimbriata (holotype, WEI 18-288). A. Profile of basidiome section; B. Basidiome section; C. Basidiome section from trama; D. Gloeocystidia; E. Gloeocystidia; F. Basidia; G. Basidiospores (rightmost: in IKI). Bars: A = 100 μm; BG = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13875801" httpUri="https://zenodo.org/record/13875801/files/figure.png" pageId="5" pageNumber="136">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5B856513B71BFFF5CCE26765FBEAC225" pageId="5" pageNumber="136" type="diagnosis">
<paragraph id="13203698B71BFFF5CCE26765FAB5C634" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCE26765FEBAC650" box="[189,311,281,306]" italics="true" pageId="5" pageNumber="136">Diagnosis.</emphasis>
—This new species is diagnostic by resupinate basidiomes with rhizomorphic margins, having yellowish brown subicular hyphae, and bearing two kinds (tubular and fusoid-cylindrical) of gloeocystidia.
</paragraph>
<paragraph id="13203698B71BFFF5CCE2671CFE4DC618" blockId="5.[136,1452,280,1351]" box="[189,448,352,379]" pageId="5" pageNumber="136">MycoBank no: 831394</paragraph>
<paragraph id="13203698B71BFFF5CCE267F9FAFAC6A0" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCE267F9FF74C6FC" box="[189,249,389,414]" italics="true" pageId="5" pageNumber="136">Type.</emphasis>
<emphasis id="21EBEA8AB71BFFF5CD4A67F8FE0BC6FC" bold="true" box="[277,390,388,414]" pageId="5" pageNumber="136">
<collectingCountry id="6B887608B71BFFF5CD4A67F8FE0BC6FC" box="[277,390,388,414]" name="Taiwan" pageId="5" pageNumber="136">TAIWAN</collectingCountry>
</emphasis>
.
<collectingRegion id="D15BF87AB71BFFF5CDC867F8FDD6C6FC" box="[407,603,388,414]" country="Taiwan" name="Pingtung" pageId="5" pageNumber="136">Pingtung County</collectingRegion>
, Chunjih Township,
<quantity id="D4679B7DB71BFFF5CF1167F8FC14C6FC" box="[846,921,388,414]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="136" unit="km" value="16.0">16 km</quantity>
of Tahanshan Forestry Road,
<geoCoordinate id="76AB505FB71BFFF5C8A467F8FA2AC6FD" box="[1275,1447,388,415]" degrees="120" direction="east" minutes="43" orientation="longitude" pageId="5" pageNumber="136" precision="1" seconds="53.9" value="120.731636">120°4353.9”E</geoCoordinate>
,
<geoCoordinate id="76AB505FB71BFFF5CCD767D4FEA4C6A1" box="[136,297,424,451]" degrees="22" direction="north" minutes="24" orientation="latitude" pageId="5" pageNumber="136" precision="1" seconds="55.2" value="22.415333">22°2455.2”N</geoCoordinate>
,
<quantity id="D4679B7DB71BFFF5CD6B67D4FE04C6A1" box="[308,393,424,451]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="136" unit="m" value="1000.0">1000 m</quantity>
alt., on fallen angiosperm trunk,
<date id="67211058B71BFFF5CEA167D4FC1BC6A0" box="[766,918,424,451]" pageId="5" pageNumber="136" value="2018-06-12">12 June 2018</date>
,
<emphasis id="21EBEA8AB71BFFF5CFFE67D5FBA4C6A0" box="[929,1065,424,451]" italics="true" pageId="5" pageNumber="136">WEI 18-288</emphasis>
(
<typeStatus id="CC24883AB71BFFF5C86767D4FB28C6A0" box="[1080,1189,424,450]" pageId="5" pageNumber="136" type="holotype">Holotype</typeStatus>
, TNM F0032731).
</paragraph>
<paragraph id="13203698B71BFFF5CCD767B0FBFDC684" blockId="5.[136,1452,280,1351]" box="[136,1136,460,487]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCD767B0FE8DC684" box="[136,256,460,486]" italics="true" pageId="5" pageNumber="136">Etymology</emphasis>
.—From
<emphasis id="21EBEA8AB71BFFF5CD3967B0FE54C684" box="[358,473,460,486]" italics="true" pageId="5" pageNumber="136">fimbriatus</emphasis>
(L.), referring to the morphology of the basidiome margin.
</paragraph>
<paragraph id="13203698B71BFFF5CCE2678DFE3CC514" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCE2678DFEC4C568" box="[189,329,497,522]" italics="true" pageId="5" pageNumber="136">Description.</emphasis>
—Basidiomes resupinate, effuse, adnate, membranaceous, 250500 μm thick in section (aculei excluded). Hymenial surface cream to buff, no color change in KOH, odontioid to hydnoid, minutely cracked when dry; margin whitish, with rhizomorphs. Aculei 34 per mm, subulate, separate,
<quantity id="D4679B7DB71BFFF5C85F6444FBFFC531" box="[1024,1138,568,595]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="5" pageNumber="136" unit="mm" value="0.7" valueMax="1.0" valueMin="0.4">0.41 mm</quantity>
long and
<quantity id="D4679B7DB71BFFF5C8BF6444FAEAC531" box="[1248,1383,568,595]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="136" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.10.2 mm</quantity>
wide, apically slightly fimbriate.
</paragraph>
<paragraph id="13203698B71BFFF5CCE264FCFEE0C3F0" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
Hyphal system monomitic; hyphae nodose-septate. Subiculum fairly uniform, with loose texture, with dense texture near substrate and hymenial layer, 220450 μm thick; hyphae interwoven, ± vertical near hymenial layer, ± horizontal near substrate, yellowish brown near substrate, colorless near hymeanial layer, occasionally swollen or with small protuberance, richly ramified, 24 (5) μm diam, with about 0.81 μm thick walls near substrate, thin- to slightly thick-walled elsewhere, finely encrusted near hymenial layer. Trama with dense texture; hyphae somewhat vertical, colorless, richly ramified, finely encrusted, 24 (5) μm diam, thin- to slightly thick-walled. Hymenial layer slightly thickening, with dense texture, 3050 μm thick; hyphae somewhat vertical, colorless, 23 μm diam, thin-walled. Gloeoplerous hyphae mainly present in trama, rare in subiculum, deeply stained by phloxine, 24 μm diam, thin- to slightly thick-walled. Gloeocystidia of two kinds: (1) tubular, originate from trama or subiculum, parallel with trama hyphae, usually curve into hymenium, immersed or slightly projecting, up to 250 μm long, 57 μm diam, thin-walled, SA; (2) fusoid to cylindrical, originate from deep hymenial layer, slightly projecting, usually with narrow apices, occasionally moniliform toward apices, 3065 × 69 μm diam, SA. Basidia subclavate to clavate, 1520 × 3.84 μm, 4-sterigmate. Basidiospores ellipsoid to broadly ellipsoid, asperulate, colorless, thin-walled, amyloid, acyanophilous, mostly 34 × 2.22.8 μm. (3) 3.24 (4.2) × (2.2) 2.32.8 μm, L = 3.58±0.34 μm, W = 2.58±0.18 μm, Q = 1.39 (n = 30) (
<emphasis id="21EBEA8AB71BFFF5CC8A6205FED0C3F0" box="[213,349,1144,1171]" italics="true" pageId="5" pageNumber="136">WEI 18-288</emphasis>
).
</paragraph>
<paragraph id="13203698B71BFFF5CCE262E0FED4C39C" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCE262E0FE26C3D4" box="[189,427,1180,1206]" italics="true" pageId="5" pageNumber="136">Additional specimens</emphasis>
(
<emphasis id="21EBEA8AB71BFFF5CDE662E1FD90C3D4" box="[441,541,1181,1206]" italics="true" pageId="5" pageNumber="136">
<typeStatus id="CC24883AB71BFFF5CDE662E1FD90C3D4" box="[441,541,1181,1206]" pageId="5" pageNumber="136" type="paratype">paratype</typeStatus>
</emphasis>
)
<emphasis id="21EBEA8AB71BFFF5CE7562E0FD18C3D4" box="[554,661,1180,1206]" italics="true" pageId="5" pageNumber="136">examined</emphasis>
.—
<emphasis id="21EBEA8AB71BFFF5CEE762E0FCA4C3D4" bold="true" box="[696,809,1180,1206]" pageId="5" pageNumber="136">
<collectingCountry id="6B887608B71BFFF5CEE762E0FCA4C3D4" box="[696,809,1180,1206]" name="Taiwan" pageId="5" pageNumber="136">TAIWAN</collectingCountry>
</emphasis>
.
<collectingRegion id="D15BF87AB71BFFF5CF6B62E0FC7FC3D4" box="[820,1010,1180,1206]" country="Taiwan" name="Pingtung" pageId="5" pageNumber="136">Pingtung County</collectingRegion>
, Chunjih Township,
<quantity id="D4679B7DB71BFFF5C88E62E0FA98C3D4" box="[1233,1301,1180,1206]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="136" unit="km" value="16.0">16 km</quantity>
of Tahanshan Forestry Road,
<geoCoordinate id="76AB505FB71BFFF5CD6662BCFE68C3B9" box="[313,485,1216,1243]" degrees="120" direction="east" minutes="43" orientation="longitude" pageId="5" pageNumber="136" precision="1" seconds="53.9" value="120.731636">120°4353.9”E</geoCoordinate>
,
<geoCoordinate id="76AB505FB71BFFF5CDAC62BCFD1EC3B9" box="[499,659,1216,1243]" degrees="22" direction="north" minutes="24" orientation="latitude" pageId="5" pageNumber="136" precision="1" seconds="55.2" value="22.415333">22°2455.2”N</geoCoordinate>
,
<quantity id="D4679B7DB71BFFF5CEFE62BCFD75C3B9" box="[673,760,1216,1243]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="136" unit="m" value="1000.0">1000 m</quantity>
alt., on fallen angiosperm trunk,
<date id="67211058B71BFFF5C82762BCFA99C3B8" box="[1144,1300,1216,1243]" pageId="5" pageNumber="136" value="2018-06-12">12 June 2018</date>
,
<emphasis id="21EBEA8AB71BFFF5C97E62BDFA26C3B8" box="[1313,1451,1216,1243]" italics="true" pageId="5" pageNumber="136">WEI 18-305</emphasis>
(TNM F0032732).
</paragraph>
<paragraph id="13203698B71BFFF5CCE26374FBEAC225" blockId="5.[136,1452,280,1351]" pageId="5" pageNumber="136">
<emphasis id="21EBEA8AB71BFFF5CCE26374FE7BC240" box="[189,502,1288,1314]" italics="true" pageId="5" pageNumber="136">Distribution and vegetation</emphasis>
.—This new species is so far known only in southern subtropical
<collectingCountry id="6B887608B71BFFF5C8806374FABCC240" box="[1247,1329,1288,1314]" name="Taiwan" pageId="5" pageNumber="136">Taiwan</collectingCountry>
at
<quantity id="D4679B7DB71BFFF5C90A6374FA26C241" box="[1365,1451,1288,1315]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="136" unit="m" value="1000.0">1000 m</quantity>
alt. The broad-leaved forest of the collecting site has many
<taxonomicName id="D49F4D1BB71BFFF5CF7F6351FC05C225" authorityName="Dumortier" authorityYear="1829" box="[800,904,1325,1351]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="136" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
and
<taxonomicName id="D49F4D1BB71BFFF5CFE06351FBBFC225" box="[959,1074,1325,1351]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="136" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
spp.
</paragraph>
</subSubSection>
</treatment>
</document>

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