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<mods:title id="DCDD137AB72886AA2AA8E109248A6E46">A taxonomic revision of the genus Phrynoglossus in Indochina with the description of a new species and comments on the classification within Occidozyginae (Amphibia, Anura, Dicroglossidae)</mods:title>
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<mods:namePart id="D3F336CBFE8DD570BA1553DA253AAA05">Than, Ni Lar</mods:namePart>
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<mods:namePart id="C72D82F9B955FA2C86782FF5112F7760">Schell, Tilman</mods:namePart>
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<paragraph id="8A3942026341FFC4FF60F8C9FDF3F86F" blockId="6.[136,549,1904,1934]" box="[136,549,1904,1934]" pageId="6" pageNumber="7">
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<taxonomicName id="4D8639816341FFC4FF60F8C9FDF3F86F" ID-CoL="6PCG" authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[136,549,1904,1934]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B8F29E106341FFC4FF60F8C9FDF3F86F" bold="true" box="[136,549,1904,1934]" pageId="6" pageNumber="7">
<emphasis id="B8F29E106341FFC4FF60F8C9FEB4F86F" bold="true" box="[136,354,1904,1934]" italics="true" pageId="6" pageNumber="7">Phrynoglossus</emphasis>
<bibRefCitation id="EE173FF36341FFC4FE82F8C9FDF3F86F" author="Peters WCH" box="[362,549,1904,1934]" pageId="6" pageNumber="7" pagination="13 - 37" refId="ref13326" refString="Peters WCH (1867) Herpetologische Notizen. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1867: 13 - 37." type="journal article" year="1867">Peters, 1867</bibRefCitation>
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species.
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<taxonomicName id="4D8639816341FFC4FECAF80DFD63F82D" authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[290,693,1972,1996]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E106341FFC4FECAF80DFDFDF82D" box="[290,555,1972,1996]" italics="true" pageId="6" pageNumber="7">Phrynoglossus martensii</emphasis>
<bibRefCitation id="EE173FF36341FFC4FDD9F80CFD63F82D" author="Peters WCH" box="[561,693,1972,1996]" pageId="6" pageNumber="7" pagination="13 - 37" refId="ref13326" refString="Peters WCH (1867) Herpetologische Notizen. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1867: 13 - 37." type="journal article" year="1867">Peters, 1867</bibRefCitation>
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<emphasis id="B8F29E106341FFC4FF60F84DFF2BF7ED" bold="true" box="[136,253,2036,2060]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
A genus of Asian frogs of the subfamily
<taxonomicName id="4D8639816341FFC4FD45F84DFDDDF7CD" authority="Fei, Ye, and Huang, 1990" authorityName="Fei, Ye, and Huang" authorityYear="1990" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="subFamily" subFamily="Occidozyginae">Occidozyginae Fei, Ye, and Huang, 1990</taxonomicName>
of the family
<taxonomicName id="4D8639816341FFC4FD4BF7ADFE4BF7AD" authority="Anderson, 1871" authorityName="Anderson" authorityYear="1871" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="family">
Dicroglossidae
<bibRefCitation id="EE173FF36341FFC4FF18F78DFE4BF7AD" author="Anderson J" box="[240,413,2100,2124]" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="6" pageNumber="7" pagination="12 - 39" part="40" refId="ref11030" refString="Anderson J (1871) A list of the reptilian accession to the Indian Museum, Calcutta from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal 40: 12 - 39." title="A list of the reptilian accession to the Indian Museum, Calcutta from 1865 to 1870, with a description of some new species" type="journal article" year="1871">Anderson, 1871</bibRefCitation>
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, that differs from all other genera of its subfamily by having the following combination of characters: (1); tongue fleshy, swollen; (2) vomerine teeth absent; (3) tips of fingers and toes slightly swollen; (4) tympanum moderately distinct; (5) skin covered by extensive mucous, feels slimy to touch in life frogs; (6) throat lining uniformly grey; (7) life style semiaquatic; (8) amplexus inguinal.
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<emphasis id="B8F29E106341FFC4FCE6F80DFCA7F82D" bold="true" box="[782,881,1972,1996]" pageId="6" pageNumber="7">Content.</emphasis>
<taxonomicName id="4D8639816341FFC4FC97F80DFABDF82D" authority="(Boulenger, 1896)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[895,1387,1972,1996]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B8F29E106341FFC4FC97F80DFB46F82D" box="[895,1168,1972,1996]" italics="true" pageId="6" pageNumber="7">Phrynoglossus baluensis</emphasis>
(
<bibRefCitation id="EE173FF36341FFC4FB4DF80DFAB5F82D" author="Boulenger GA" box="[1189,1379,1972,1996]" pageId="6" pageNumber="7" pagination="401 - 406" refId="ref11272" refString="Boulenger GA (1896) Descriptions of new batrachians in the British Museum. Annals and Magazine of Natural History, Series 6, 17: 401 - 406." type="journal article" year="1896">Boulenger, 1896</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D8639816341FFC4FCE6F86DFB12F80D" authority="(Smith, 1927)" baseAuthorityName="Smith" baseAuthorityYear="1927" box="[782,1220,2004,2028]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="celebensis">
<emphasis id="B8F29E106341FFC4FCE6F86DFBF5F80D" box="[782,1059,2004,2028]" italics="true" pageId="6" pageNumber="7">Phrynoglossus celebensis</emphasis>
(
<bibRefCitation id="EE173FF36341FFC4FBDBF86DFB6BF80D" author="Smith MA" box="[1075,1213,2004,2028]" pageId="6" pageNumber="7" pagination="199 - 225" refId="ref13753" refString="Smith MA (1927) Contributions to the herpetology of the Indo Australian region. Proceedings of the Zoological Society of London 1927: 199 - 225." type="book chapter" year="1927">Smith, 1927</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D8639816341FFC4FB3AF86DFBEBF7ED" authority="(Taylor, 1922)" baseAuthorityName="Taylor" baseAuthorityYear="1922" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="diminutivus">
<emphasis id="B8F29E106341FFC4FB3AF86DFC5CF7ED" italics="true" pageId="6" pageNumber="7">Phrynoglossus diminutivus</emphasis>
(
<bibRefCitation id="EE173FF36341FFC4FC49F84DFBE3F7ED" author="Taylor EH" box="[929,1077,2036,2060]" journalOrPublisher="Philippine Journal of Science" pageId="6" pageNumber="7" pagination="257 - 303" part="21" refId="ref13839" refString="Taylor EH (1922) Additions to the herpetological fauna of the Philippine Islands. II. Philippine Journal of Science 21: 257 - 303." title="Additions to the herpetological fauna of the Philippine Islands. II" type="journal article" year="1922">Taylor, 1922</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D8639816341FFC4FBA7F84DFC69F7CD" authority="(Mertens, 1927)" baseAuthorityName="Mertens" baseAuthorityYear="1927" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="floresianus">
<emphasis id="B8F29E106341FFC4FBA7F84DFAB9F7ED" box="[1103,1391,2036,2060]" italics="true" pageId="6" pageNumber="7">Phrynoglossus floresianus</emphasis>
(
<bibRefCitation id="EE173FF36341FFC4FCFEF7ADFC6EF7CD" author="Mertens R" box="[790,952,2068,2092]" pageId="6" pageNumber="7" pagination="234 - 242" refId="ref13012" refString="Mertens R (1927) Neue Amphibien und Reptilien aus dem Indo-Australischen Archipel, gesammelt wahrend der Sunda-Expedition Rensch. Senckenbergiana biologica 9: 234 - 242." type="journal article" year="1927">Mertens, 1927</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D8639816341FFC4FC25F7ADFABDF7CD" authority="(Gunther, 1859)" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[973,1387,2068,2092]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B8F29E106341FFC4FC25F7ADFB67F7CD" box="[973,1201,2068,2092]" italics="true" pageId="6" pageNumber="7">Phrynoglossus laevis</emphasis>
(Günther, 1859)
</taxonomicName>
,
<taxonomicName id="4D8639816341FFCBFCE6F78DFED3F80D" authority="(Taylor and Elbel, 1958)" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" lastPageId="9" lastPageNumber="10" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E106341FFC4FCE6F78DFBACF7AD" box="[782,1146,2100,2124]" italics="true" pageId="6" pageNumber="7">Phrynoglossus magnapustulosus</emphasis>
(
<bibRefCitation id="EE173FF36341FFCBFB7FF78DFF28F80D" author="Taylor EH &amp; Elbel RE" journalOrPublisher="University of Kansas Science Bulletin" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" pagination="1033 - 1189" part="38" refId="ref13888" refString="Taylor EH, Elbel RE (1958) Contribution to the herpetology of Thailand. University of Kansas Science Bulletin 38: 1033 - 1189." title="Contribution to the herpetology of Thailand" type="journal article" year="1958">Taylor and Elbel, 1958</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D863981634EFFCBFEFEF86DFD6BF80D" authority="Peters, 1867" authorityName="Peters" authorityYear="1867" box="[278,701,2004,2028]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634EFFCBFEFEF86DFDF3F80D" box="[278,549,2004,2028]" italics="true" pageId="9" pageNumber="10">Phrynoglossus martensii</emphasis>
<bibRefCitation id="EE173FF3634EFFCBFDD9F86CFD6BF80D" author="Peters WCH" box="[561,701,2004,2028]" pageId="9" pageNumber="10" pagination="13 - 37" refId="ref13326" refString="Peters WCH (1867) Herpetologische Notizen. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1867: 13 - 37." type="journal article" year="1867">Peters, 1867</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4D863981634EFFCBFD26F86DFDA6F7ED" authority="(Smith, 1927)" baseAuthorityName="Smith" baseAuthorityYear="1927" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="semipalmatus">
<emphasis id="B8F29E10634EFFCBFD26F86DFE6CF7ED" italics="true" pageId="9" pageNumber="10">Phrynoglossus semipalmatus</emphasis>
(
<bibRefCitation id="EE173FF3634EFFCBFE3CF84DFDBFF7ED" author="Smith MA" box="[468,617,2036,2060]" pageId="9" pageNumber="10" pagination="199 - 225" refId="ref13753" refString="Smith MA (1927) Contributions to the herpetology of the Indo Australian region. Proceedings of the Zoological Society of London 1927: 199 - 225." type="book chapter" year="1927">Smith, 1927</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D863981634EFFCBFD60F84DFE20F7CD" authority="(Peters, 1877)" baseAuthorityName="Peters" baseAuthorityYear="1877" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="sumatranus">
<emphasis id="B8F29E10634EFFCBFD60F84DFE97F7CD" italics="true" pageId="9" pageNumber="10">Phrynoglossus sumatranus</emphasis>
(
<bibRefCitation id="EE173FF3634EFFCBFEB2F7ADFE38F7CD" author="Peters WCH" box="[346,494,2068,2092]" pageId="9" pageNumber="10" pagination="415 - 423" refId="ref13349" refString="Peters WCH (1877) Herpetologische Notizen. II. Bemerkungen uber neue oder weniger bekannte Amphibien. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1877: 415 - 423" type="journal article" year="1877">Peters, 1877</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4D863981634EFFCBFDE4F7ADFD46F7AD" authority="(Iskandar, Arifin, and Rachmanasah, 2011)" baseAuthorityName="Iskandar, Arifin, and Rachmanasah" baseAuthorityYear="2011" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="tompotika">
<emphasis id="B8F29E10634EFFCBFDE4F7ADFCF3F7CD" box="[524,805,2068,2092]" italics="true" pageId="9" pageNumber="10">Phrynoglossus tompotika</emphasis>
(
<bibRefCitation id="EE173FF3634EFFCBFF24F78DFD5FF7AD" author="Iskandar DT &amp; Arifin U &amp; Rachmanasah A." box="[204,649,2100,2124]" pageId="9" pageNumber="10" pagination="219 - 228" refId="ref12281" refString="Iskandar DT, Arifin U, Rachmanasah A. (2011) A new frog (Anura, Dicroglossidae), related to Occidozyga semipalmata Smith, 1927, from the eastern peninsula of Sulawesi, Indonesia. Raffles Bulletin of Zoology. Singapore 59: 219 - 228." type="journal article" year="2011">Iskandar, Arifin, and Rachmanasah, 2011</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DEF9128A6340FFC5FF2DFF6FFF26FEEA" pageId="7" pageNumber="8" startId="7.[197,253,214,235]" targetBox="[250,1425,301,514]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="8A3942026340FFC5FF2DFF6FFF26FEEA" blockId="7.[197,1452,214,267]" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF2DFF6FFEC0FF0A" bold="true" box="[197,278,214,235]" pageId="7" pageNumber="8">Table 1.</emphasis>
Uncorrected pairwise distances for the
<taxonomicName id="4D8639816340FFC5FD7DFF6FFCF3FF0D" authorityName="Fei, Ye, and Huang" authorityYear="1990" box="[661,805,214,236]" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="7" pageNumber="7" phylum="Chordata" rank="subFamily" subFamily="Occidozyginae">Occidozyginae</taxonomicName>
plus
<taxonomicName id="4D8639816340FFC5FCB1FF6EFC04FF0D" authorityName="Fitzinger" authorityYear="1843" box="[857,978,215,236]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B8F29E106340FFC5FCB1FF6EFC04FF0D" box="[857,978,215,236]" italics="true" pageId="7" pageNumber="8">Limnonectes</emphasis>
</taxonomicName>
(outgroup) included in this study. For details see text.
</paragraph>
</caption>
<paragraph id="8A3942026340FFC5FE18FE94FAA7FDE0" pageId="7" pageNumber="8">
<table id="F886B0A26340003DFF12FE94FA47FDE3" box="[250,1425,301,514]" gridcols="7" gridrows="7" pageId="7" pageNumber="8">
<tr id="34B640406340003DFF12FE94FA47FEA1" box="[250,1425,301,320]" gridrow="0" pageId="7" pageNumber="8" rowspan-0="1">
<emphasis id="B8F29E106340FFC5FE18FE94FA47FEA1" bold="true" box="[496,1425,301,320]" pageId="7" pageNumber="8">
<th id="7767293C6340003DFE18FE94FD95FEA1" box="[496,579,301,320]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FE18FE94FD95FEA1" authorityName="Dubois" authorityYear="1987" box="[496,579,301,320]" class="Amphibia" family="Dicroglossidae" genus="Ingerana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Ingerana</taxonomicName>
</th>
<th id="7767293C6340003DFD63FE94FD22FEA1" box="[651,756,301,320]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FD63FE94FD22FEA1" authorityName="Kuhl &amp; van Hasselt" authorityYear="1822" box="[651,756,301,320]" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Occidozyga</taxonomicName>
</th>
<taxonomicName id="4D8639816340FFC5FCC4FE94FBEEFEA1" box="[812,1080,301,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
<th id="7767293C6340003DFCC4FE94FC76FEA1" box="[812,928,301,320]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">Limnonectes</th>
<th id="7767293C6340003DFC09FE94FBEEFEA1" box="[993,1080,301,320]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">martensii</th>
</taxonomicName>
<th id="7767293C6340003DFB9FFE94FB3BFEA1" box="[1143,1261,301,320]" gridcol="5" gridrow="0" pageId="7" pageNumber="8">
cf.
<taxonomicName id="4D8639816340FFC5FB64FE94FB3BFEA1" box="[1164,1261,301,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">_martensii</taxonomicName>
</th>
<th id="7767293C6340003DFAC8FE94FA47FEA1" box="[1312,1425,301,320]" gridcol="6" gridrow="0" pageId="7" pageNumber="8">„Myanmar“</th>
</emphasis>
</tr>
<tr id="34B640406340003DFF12FEF4FA47FE81" box="[250,1425,333,352]" gridrow="1" pageId="7" pageNumber="8" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="7767293C6340003DFF12FEF4FE5BFE81" box="[250,397,333,352]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FF12FEF4FEB6FE81" authorityName="Kuhl &amp; van Hasselt" authorityYear="1822" box="[250,352,333,352]" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Occidozyga</taxonomicName>
</th>
<td id="7767293C6340003DFE18FEF4FD95FE81" box="[496,579,333,352]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">0.189</td>
</tr>
<tr id="34B640406340003DFF12FED4FA47FE60" box="[250,1425,365,385]" gridrow="2" pageId="7" pageNumber="8" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="7767293C6340003DFF12FED4FE5BFE60" box="[250,397,365,385]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FF12FED4FEBEFE61" authorityName="Fitzinger" authorityYear="1843" box="[250,360,365,384]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Limnonectes</taxonomicName>
</th>
<td id="7767293C6340003DFE18FED4FD95FE60" box="[496,579,365,385]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">0.192</td>
<td id="7767293C6340003DFD63FED4FD22FE60" box="[651,756,365,385]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">0.193</td>
</tr>
<tr id="34B640406340003DFF12FE37FA47FE40" box="[250,1425,398,417]" gridrow="3" pageId="7" pageNumber="8" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="7767293C6340003DFF12FE37FE5BFE40" box="[250,397,398,417]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">martensii</th>
<td id="7767293C6340003DFE18FE37FD95FE40" box="[496,579,398,417]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">0.198</td>
<td id="7767293C6340003DFD63FE37FD22FE40" box="[651,756,398,417]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">0.132</td>
<td id="7767293C6340003DFCC4FE37FC76FE40" box="[812,928,398,417]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">0.203</td>
</tr>
<tr id="34B640406340003DFF12FE17FA47FE20" box="[250,1425,430,449]" gridrow="4" pageId="7" pageNumber="8" rowspan-5="1" rowspan-6="1">
<th id="7767293C6340003DFF12FE17FE5BFE20" box="[250,397,430,449]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">
cf.
<taxonomicName id="4D8639816340FFC5FEE6FE17FEBCFE20" box="[270,362,430,449]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">_martensii</taxonomicName>
</th>
<td id="7767293C6340003DFE18FE17FD95FE20" box="[496,579,430,449]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">0.202</td>
<td id="7767293C6340003DFD63FE17FD22FE20" box="[651,756,430,449]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">0.125</td>
<td id="7767293C6340003DFCC4FE17FC76FE20" box="[812,928,430,449]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">0.213</td>
<td id="7767293C6340003DFC09FE17FBEEFE20" box="[993,1080,430,449]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">0.035</td>
</tr>
<tr id="34B640406340003DFF12FE77FA47FE00" box="[250,1425,462,481]" gridrow="5" pageId="7" pageNumber="8" rowspan-6="1">
<th id="7767293C6340003DFF12FE77FE5BFE00" box="[250,397,462,481]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">„Myanmar“</th>
<td id="7767293C6340003DFE18FE77FD95FE00" box="[496,579,462,481]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">0.203</td>
<td id="7767293C6340003DFD63FE77FD22FE00" box="[651,756,462,481]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">0.139</td>
<td id="7767293C6340003DFCC4FE77FC76FE00" box="[812,928,462,481]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">0.209</td>
<td id="7767293C6340003DFC09FE77FBEEFE00" box="[993,1080,462,481]" gridcol="4" gridrow="5" pageId="7" pageNumber="8">0.054</td>
<td id="7767293C6340003DFB9FFE77FB3BFE00" box="[1143,1261,462,481]" gridcol="5" gridrow="5" pageId="7" pageNumber="8">0.051</td>
</tr>
<tr id="34B640406340003DFF12FE57FA47FDE3" box="[250,1425,494,514]" gridrow="6" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF12FE57FE5BFDE3" box="[250,397,494,514]" gridcol="0" gridrow="6" pageId="7" pageNumber="8">magnapustulosus</th>
<td id="7767293C6340003DFE18FE57FD95FDE3" box="[496,579,494,514]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">0.208</td>
<td id="7767293C6340003DFD63FE57FD22FDE3" box="[651,756,494,514]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">0.127</td>
<td id="7767293C6340003DFCC4FE57FC76FDE3" box="[812,928,494,514]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">0.207</td>
<td id="7767293C6340003DFC09FE57FBEEFDE3" box="[993,1080,494,514]" gridcol="4" gridrow="6" pageId="7" pageNumber="8">0.045</td>
<td id="7767293C6340003DFB9FFE57FB3BFDE3" box="[1143,1261,494,514]" gridcol="5" gridrow="6" pageId="7" pageNumber="8">0.038</td>
<td id="7767293C6340003DFAC8FE57FA47FDE3" box="[1312,1425,494,514]" gridcol="6" gridrow="6" pageId="7" pageNumber="8">0.059</td>
</tr>
</table>
</paragraph>
<caption id="DEF9128A6340FFC5FF2DFDEFFBC2FD6A" pageId="7" pageNumber="8" startId="7.[197,253,598,619]" targetBox="[220,1426,685,1375]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="8A3942026340FFC5FF2DFDEFFBC2FD6A" blockId="7.[197,1452,598,652]" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF2DFDEFFEC0FD8A" bold="true" box="[197,278,598,619]" pageId="7" pageNumber="8">Table 2.</emphasis>
Selected bioacoustic parameters of the species related to
<taxonomicName id="4D8639816340FFC5FCABFDEFFBE4FD8D" authorityName="Peters" authorityYear="1867" box="[835,1074,598,620]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E106340FFC5FCABFDEFFBE4FD8D" box="[835,1074,598,620]" italics="true" pageId="7" pageNumber="8">Phrynoglossus martensii</emphasis>
</taxonomicName>
. Range is followed by mean value and standard deviation in parentheses. Dom. Freq. = dominant frequency; Freg. = frequency.
</paragraph>
</caption>
<paragraph id="8A3942026340FFC5FDF0FD03FA5BFABE" pageId="7" pageNumber="8">
<table id="F886B0A26340003DFF34FD14FA44FABE" box="[220,1426,685,1375]" gridcols="7" gridrows="12" pageId="7" pageNumber="8">
<tr id="34B640406340003DFF34FD14FA44FD3B" box="[220,1426,685,730]" gridrow="0" pageId="7" pageNumber="8" rowspan-0="1">
<emphasis id="B8F29E106340FFC5FDF0FD03FCE3FD21" bold="true" box="[536,821,685,717]" pageId="7" pageNumber="8">
<th id="7767293C6340003DFE1EFD14FD59FD3B" box="[502,655,685,730]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">Taxon ID</th>
<th id="7767293C6340003DFD53FD14FCE3FD3B" box="[699,821,685,730]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">Call duration (sec)</th>
</emphasis>
<th id="7767293C6340003DFC80FD14FC1BFD3B" box="[872,973,685,730]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FC82FD14FC1DFD21" bold="true" box="[874,971,685,704]" pageId="7" pageNumber="8">Dom. Freq</emphasis>
(Hz)
</th>
<th id="7767293C6340003DFC1CFD14FBB9FD3B" box="[1012,1135,685,730]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FC1CFD14FBB9FD21" bold="true" box="[1012,1135,685,704]" pageId="7" pageNumber="8">Gap duration</emphasis>
(sec)
</th>
<th id="7767293C6340003DFB7FFD14FB2DFD3B" box="[1175,1275,685,730]" gridcol="5" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FB49FD14FB27FD21" bold="true" box="[1185,1265,685,704]" pageId="7" pageNumber="8">Freq 5%</emphasis>
(Hz)
</th>
<th id="7767293C6340003DFAC6FD14FA44FD3B" box="[1326,1426,685,730]" gridcol="6" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FADAFD14FA58FD21" bold="true" box="[1330,1422,685,704]" pageId="7" pageNumber="8">Freq 95%</emphasis>
(Hz)
</th>
</tr>
<tr id="34B640406340003DFF34FD51FA44FCF4" box="[220,1426,744,789]" gridrow="1" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FD51FE05FCF4" box="[220,467,744,789]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FD51FE7EFCF5" bold="true" pageId="7" pageNumber="8">GK-7475_0544 (n=23 calls) Roi Et Prov., Thailand</emphasis>
</th>
<td id="7767293C6340003DFE1EFD51FD59FCF4" box="[502,655,744,789]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FE11FD4CFD5BFCE9" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[505,653,757,776]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E106340FFC5FE11FD4CFD5BFCE9" box="[505,653,757,776]" italics="true" pageId="7" pageNumber="8">magnapustulosus</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FD51FCE3FCF4" box="[699,821,744,789]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">0.2430.354 (0.291±0.024)</td>
<td id="7767293C6340003DFC80FD51FC1BFCF4" box="[872,973,744,789]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">31873682 (3447±175)</td>
<td id="7767293C6340003DFC1CFD51FBB9FCF4" box="[1012,1135,744,789]" gridcol="4" gridrow="1" pageId="7" pageNumber="8">2.254.28 (3.15±0.58)</td>
<td id="7767293C6340003DFB7FFD51FB2DFCF4" box="[1175,1275,744,789]" gridcol="5" gridrow="1" pageId="7" pageNumber="8">16801938 (1790±74)</td>
<td id="7767293C6340003DFAC6FD51FA44FCF4" box="[1326,1426,744,789]" gridcol="6" gridrow="1" pageId="7" pageNumber="8">41995383 (4823±388)</td>
</tr>
<tr id="34B640406340003DFF34FC9BFA44FCAE" box="[220,1426,802,847]" gridrow="2" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FC9BFE05FCAE" box="[220,467,802,847]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FC9BFE7EFCAE" bold="true" pageId="7" pageNumber="8">GK-7465_0543 (n=15 calls) Roi Et Prov., Thailand</emphasis>
</th>
<td id="7767293C6340003DFE1EFC9BFD59FCAE" box="[502,655,802,847]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FE11FC96FD5BFCA3" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[505,653,815,834]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E106340FFC5FE11FC96FD5BFCA3" box="[505,653,815,834]" italics="true" pageId="7" pageNumber="8">magnapustulosus</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FC9BFCE3FCAE" box="[699,821,802,847]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">0.3220.437 (0.387±0.031)</td>
<td id="7767293C6340003DFC80FC9BFC1BFCAE" box="[872,973,802,847]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">37253790 (3761±20)</td>
<td id="7767293C6340003DFC1CFC9BFBB9FCAE" box="[1012,1135,802,847]" gridcol="4" gridrow="2" pageId="7" pageNumber="8">2.675.31 (3.92±0.67)</td>
<td id="7767293C6340003DFB7FFC9BFB2DFCAE" box="[1175,1275,802,847]" gridcol="5" gridrow="2" pageId="7" pageNumber="8">17442131 (1894±94)</td>
<td id="7767293C6340003DFAC6FC9BFA44FCAE" box="[1326,1426,802,847]" gridcol="6" gridrow="2" pageId="7" pageNumber="8">40914177 (4150±29)</td>
</tr>
<tr id="34B640406340003DFF34FCE4FA44FC6B" box="[220,1426,861,906]" gridrow="3" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FCE4FE05FC6B" box="[220,467,861,906]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FCE4FE7EFC68" bold="true" pageId="7" pageNumber="8">GK-7476_0546 (n=9 calls) Roi Et Prov., Thailand</emphasis>
</th>
<td id="7767293C6340003DFE1EFCE4FD59FC6B" box="[502,655,861,906]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FE11FCD3FD5BFC9C" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[505,653,874,893]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E106340FFC5FE11FCD3FD5BFC9C" box="[505,653,874,893]" italics="true" pageId="7" pageNumber="8">magnapustulosus</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FCE4FCE3FC6B" box="[699,821,861,906]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">0.2620.287 (0.277±0.008)</td>
<td id="7767293C6340003DFC80FCE4FC1BFC6B" box="[872,973,861,906]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">37503750 (3750±0)</td>
<td id="7767293C6340003DFC1CFCE4FBB9FC6B" box="[1012,1135,861,906]" gridcol="4" gridrow="3" pageId="7" pageNumber="8">2.533.20 (2.77±0.19)</td>
<td id="7767293C6340003DFB7FFCE4FB2DFC6B" box="[1175,1275,861,906]" gridcol="5" gridrow="3" pageId="7" pageNumber="8">16401734 (1708±32)</td>
<td id="7767293C6340003DFAC6FCE4FA44FC6B" box="[1326,1426,861,906]" gridcol="6" gridrow="3" pageId="7" pageNumber="8">44064453 (4427±23)</td>
</tr>
<tr id="34B640406340003DFF34FC2EFA44FC25" box="[220,1426,919,964]" gridrow="4" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FC2EFE05FC25" box="[220,467,919,964]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FC21FEB8FC25" bold="true" pageId="7" pageNumber="8">GK-6727_0183 (n=7 calls) EYU, Myanmar</emphasis>
</th>
<td id="7767293C6340003DFE1EFC2EFD59FC25" box="[502,655,919,964]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A3C1236B6340FFC5FE1EFC1CFDF3FC59" box="[502,549,933,952]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
“Myanmar”
</td>
<td id="7767293C6340003DFD53FC2EFCE3FC25" box="[699,821,919,964]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">0.0850.114 (0.104±0.009)</td>
<td id="7767293C6340003DFC80FC2EFC1BFC25" box="[872,973,919,964]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">24542476 (2467±11)</td>
<td id="7767293C6340003DFC1CFC2EFBB9FC25" box="[1012,1135,919,964]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">2.342.90 (2.66±0.23)</td>
<td id="7767293C6340003DFB7FFC2EFB2DFC25" box="[1175,1275,919,964]" gridcol="5" gridrow="4" pageId="7" pageNumber="8">13571809 (1529±156)</td>
<td id="7767293C6340003DFAC6FC2EFA44FC25" box="[1326,1426,919,964]" gridcol="6" gridrow="4" pageId="7" pageNumber="8">28202863 (2839±18)</td>
</tr>
<tr id="34B640406340003DFF34FC6BFA44FC1E" box="[220,1426,978,1023]" gridrow="5" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FC6BFE05FC1E" box="[220,467,978,1023]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FC6BFEB8FC1E" bold="true" pageId="7" pageNumber="8">GK-6926_0008 (n=21 calls) EYU, Myanmar</emphasis>
</th>
<td id="7767293C6340003DFE1EFC6BFD59FC1E" box="[502,655,978,1023]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A3C1236B6340FFC5FE1EFC59FDF3FC12" box="[502,549,992,1011]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
“Myanmar”
</td>
<td id="7767293C6340003DFD53FC6BFCE3FC1E" box="[699,821,978,1023]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">0.0790.109 (0.098±0.008)</td>
<td id="7767293C6340003DFC80FC6BFC1BFC1E" box="[872,973,978,1023]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">27562885 (2839±41)</td>
<td id="7767293C6340003DFC1CFC6BFBB9FC1E" box="[1012,1135,978,1023]" gridcol="4" gridrow="5" pageId="7" pageNumber="8">2.474.78 (3.31±0.67)</td>
<td id="7767293C6340003DFB7FFC6BFB2DFC1E" box="[1175,1275,978,1023]" gridcol="5" gridrow="5" pageId="7" pageNumber="8">17662347 (2119±168)</td>
<td id="7767293C6340003DFAC6FC6BFA44FC1E" box="[1326,1426,978,1023]" gridcol="6" gridrow="5" pageId="7" pageNumber="8">31873316 (3265±37)</td>
</tr>
<tr id="34B640406340003DFF34FBB4FA44FBDB" box="[220,1426,1037,1082]" gridrow="6" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FBB4FE05FBDB" box="[220,467,1037,1082]" gridcol="0" gridrow="6" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FBB4FE42FBDB" bold="true" pageId="7" pageNumber="8">Not collected (n=26 calls) Thanlyin, Myanmar</emphasis>
</th>
<td id="7767293C6340003DFE1EFBB4FD59FBDB" box="[502,655,1037,1082]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A3C1236B6340FFC5FE1EFBA3FDF3FBCC" box="[502,549,1050,1069]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
“Myanmar”
</td>
<td id="7767293C6340003DFD53FBB4FCE3FBDB" box="[699,821,1037,1082]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">0.0820.102 (0.090±0.005)</td>
<td id="7767293C6340003DFC80FBB4FC1BFBDB" box="[872,973,1037,1082]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">25412584 (2566±18)</td>
<td id="7767293C6340003DFC1CFBB4FBB9FBDB" box="[1012,1135,1037,1082]" gridcol="4" gridrow="6" pageId="7" pageNumber="8">1.842.80 (2.18±0.32)</td>
<td id="7767293C6340003DFB7FFBB4FB2DFBDB" box="[1175,1275,1037,1082]" gridcol="5" gridrow="6" pageId="7" pageNumber="8">13572110 (1831±238)</td>
<td id="7767293C6340003DFAC6FBB4FA44FBDB" box="[1326,1426,1037,1082]" gridcol="6" gridrow="6" pageId="7" pageNumber="8">29293015 (2975±26)</td>
</tr>
<tr id="34B640406340003DFF34FBFEFA44FB95" box="[220,1426,1095,1140]" gridrow="7" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FBFEFE05FB95" box="[220,467,1095,1140]" gridcol="0" gridrow="7" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FBFEFEC9FB95" bold="true" pageId="7" pageNumber="8">Goutte_9357 (n=24 calls) Taiwan</emphasis>
</th>
<td id="7767293C6340003DFE1EFBFEFD59FB95" box="[502,655,1095,1140]" gridcol="1" gridrow="7" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FDE5FBEDFDAFFB86" authorityName="Peters" authorityYear="1867" box="[525,633,1108,1127]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
cf.
<emphasis id="B8F29E106340FFC5FDC0FBECFDAFFB86" box="[552,633,1109,1127]" italics="true" pageId="7" pageNumber="8">martensii</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FBFEFCE3FB95" box="[699,821,1095,1140]" gridcol="2" gridrow="7" pageId="7" pageNumber="8">0.0410.051 (0.046±0.003)</td>
<td id="7767293C6340003DFC80FBFEFC1BFB95" box="[872,973,1095,1140]" gridcol="3" gridrow="7" pageId="7" pageNumber="8">38763941 (3914±17)</td>
<td id="7767293C6340003DFC1CFBFEFBB9FB95" box="[1012,1135,1095,1140]" gridcol="4" gridrow="7" pageId="7" pageNumber="8">2.122.99 (2.39±0.32)</td>
<td id="7767293C6340003DFB7FFBFEFB2DFB95" box="[1175,1275,1095,1140]" gridcol="5" gridrow="7" pageId="7" pageNumber="8">23043488 (3027±414)</td>
<td id="7767293C6340003DFAC6FBFEFA44FB95" box="[1326,1426,1095,1140]" gridcol="6" gridrow="7" pageId="7" pageNumber="8">42644737 (4493±128)</td>
</tr>
<tr id="34B640406340003DFF34FB3BFA44FB4E" box="[220,1426,1154,1199]" gridrow="8" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FB3BFE05FB4E" box="[220,467,1154,1199]" gridcol="0" gridrow="8" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FB3BFEC9FB4E" bold="true" pageId="7" pageNumber="8">Goutte_9358 (n=28 calls) Taiwan</emphasis>
</th>
<td id="7767293C6340003DFE1EFB3BFD59FB4E" box="[502,655,1154,1199]" gridcol="1" gridrow="8" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FDE5FB36FDAFFB43" authorityName="Peters" authorityYear="1867" box="[525,633,1167,1186]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
cf.
<emphasis id="B8F29E106340FFC5FDC0FB29FDAFFB43" box="[552,633,1168,1186]" italics="true" pageId="7" pageNumber="8">martensii</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FB3BFCE3FB4E" box="[699,821,1154,1199]" gridcol="2" gridrow="8" pageId="7" pageNumber="8">0.0380.052 (0.043±0.003)</td>
<td id="7767293C6340003DFC80FB3BFC1BFB4E" box="[872,973,1154,1199]" gridcol="3" gridrow="8" pageId="7" pageNumber="8">39193962 (3934±15)</td>
<td id="7767293C6340003DFC1CFB3BFBB9FB4E" box="[1012,1135,1154,1199]" gridcol="4" gridrow="8" pageId="7" pageNumber="8">1.942.78 (2.37±0.65)</td>
<td id="7767293C6340003DFB7FFB3BFB2DFB4E" box="[1175,1275,1154,1199]" gridcol="5" gridrow="8" pageId="7" pageNumber="8">28643553 (3356±414)</td>
<td id="7767293C6340003DFAC6FB3BFA44FB4E" box="[1326,1426,1154,1199]" gridcol="6" gridrow="8" pageId="7" pageNumber="8">42644371 (4300±24)</td>
</tr>
<tr id="34B640406340003DFF34FB04FA44FB0B" box="[220,1426,1213,1258]" gridrow="9" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FB04FE05FB0B" box="[220,467,1213,1258]" gridcol="0" gridrow="9" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FB04FEFFFB08" bold="true" pageId="7" pageNumber="8">Ziegler TZ-473 (n=24 calls) Vietnam</emphasis>
</th>
<td id="7767293C6340003DFE1EFB04FD59FB0B" box="[502,655,1213,1258]" gridcol="1" gridrow="9" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FDE5FB73FDAFFB3D" authorityName="Peters" authorityYear="1867" box="[525,633,1226,1245]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
cf.
<emphasis id="B8F29E106340FFC5FDC0FB73FDAFFB3D" box="[552,633,1226,1244]" italics="true" pageId="7" pageNumber="8">martensii</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FB04FCE3FB0B" box="[699,821,1213,1258]" gridcol="2" gridrow="9" pageId="7" pageNumber="8">0.0320.046 (0.037±0.004)</td>
<td id="7767293C6340003DFC80FB04FC1BFB0B" box="[872,973,1213,1258]" gridcol="3" gridrow="9" pageId="7" pageNumber="8">33383553 (3444±70)</td>
<td id="7767293C6340003DFC1CFB04FBB9FB0B" box="[1012,1135,1213,1258]" gridcol="4" gridrow="9" pageId="7" pageNumber="8">0.530.90 (0.72±0.09)</td>
<td id="7767293C6340003DFB7FFB04FB2DFB0B" box="[1175,1275,1213,1258]" gridcol="5" gridrow="9" pageId="7" pageNumber="8">18733144 (2874±275)</td>
<td id="7767293C6340003DFAC6FB04FA44FB0B" box="[1326,1426,1213,1258]" gridcol="6" gridrow="9" pageId="7" pageNumber="8">37254027 (3830±76)</td>
</tr>
<tr id="34B640406340003DFF34FB4EFA44FAC5" box="[220,1426,1271,1316]" gridrow="10" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FB4EFE05FAC5" box="[220,467,1271,1316]" gridcol="0" gridrow="10" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FB4EFEF9FAC5" bold="true" pageId="7" pageNumber="8">Chiang Mai (n=10 calls) Thailand</emphasis>
</th>
<td id="7767293C6340003DFE1EFB4EFD59FAC5" box="[502,655,1271,1316]" gridcol="1" gridrow="10" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FDE5FABDFDAFFAF6" authorityName="Peters" authorityYear="1867" box="[525,633,1284,1303]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" isUncertain="true" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
cf.
<emphasis id="B8F29E106340FFC5FDC0FABCFDAFFAF6" box="[552,633,1285,1303]" italics="true" pageId="7" pageNumber="8">martensii</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FB4EFCE3FAC5" box="[699,821,1271,1316]" gridcol="2" gridrow="10" pageId="7" pageNumber="8">0.0330.043 (0.037±0.003)</td>
<td id="7767293C6340003DFC80FB4EFC1BFAC5" box="[872,973,1271,1316]" gridcol="3" gridrow="10" pageId="7" pageNumber="8">36603704 (3689±14)</td>
<td id="7767293C6340003DFC1CFB4EFBB9FAC5" box="[1012,1135,1271,1316]" gridcol="4" gridrow="10" pageId="7" pageNumber="8">1.912.56 (2.19±0.20)</td>
<td id="7767293C6340003DFB7FFB4EFB2DFAC5" box="[1175,1275,1271,1316]" gridcol="5" gridrow="10" pageId="7" pageNumber="8">26703165 (2922±130)</td>
<td id="7767293C6340003DFAC6FB4EFA44FAC5" box="[1326,1426,1271,1316]" gridcol="6" gridrow="10" pageId="7" pageNumber="8">41344435 (4254±111)</td>
</tr>
<tr id="34B640406340003DFF34FA8BFA44FABE" box="[220,1426,1330,1375]" gridrow="11" pageId="7" pageNumber="8">
<th id="7767293C6340003DFF34FA8BFE05FABE" box="[220,467,1330,1375]" gridcol="0" gridrow="11" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF34FA8BFEF9FABF" bold="true" pageId="7" pageNumber="8">Bangkok (n=14 calls) Thailand</emphasis>
</th>
<td id="7767293C6340003DFE1EFA8BFD59FABE" box="[502,655,1330,1375]" gridcol="1" gridrow="11" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FDF2FA86FDBDFAB0" authorityName="Peters" authorityYear="1867" box="[538,619,1343,1361]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E106340FFC5FDF2FA86FDBDFAB0" box="[538,619,1343,1361]" italics="true" pageId="7" pageNumber="8">martensii</emphasis>
</taxonomicName>
</td>
<td id="7767293C6340003DFD53FA8BFCE3FABE" box="[699,821,1330,1375]" gridcol="2" gridrow="11" pageId="7" pageNumber="8">0.0400.048 (0.043±0.002)</td>
<td id="7767293C6340003DFC80FA8BFC1BFABE" box="[872,973,1330,1375]" gridcol="3" gridrow="11" pageId="7" pageNumber="8">34673596 (3544±37)</td>
<td id="7767293C6340003DFC1CFA8BFBB9FABE" box="[1012,1135,1330,1375]" gridcol="4" gridrow="11" pageId="7" pageNumber="8">2.803.86 (3.42±0.34)</td>
<td id="7767293C6340003DFB7FFA8BFB2DFABE" box="[1175,1275,1330,1375]" gridcol="5" gridrow="11" pageId="7" pageNumber="8">22613165 (2689±257)</td>
<td id="7767293C6340003DFAC6FA8BFA44FABE" box="[1326,1426,1330,1375]" gridcol="6" gridrow="11" pageId="7" pageNumber="8">39194155 (4028±69)</td>
</tr>
</table>
</paragraph>
<caption id="DEF9128A6340FFC5FF2DFA0FFD61FA0A" pageId="7" pageNumber="8" startId="7.[197,253,1462,1483]" targetBox="[257,1428,1549,2108]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="8A3942026340FFC5FF2DFA0FFD61FA0A" blockId="7.[197,1451,1462,1516]" pageId="7" pageNumber="8">
<emphasis id="B8F29E106340FFC5FF2DFA0FFEC0FA2A" bold="true" box="[197,278,1462,1483]" pageId="7" pageNumber="8">Table 3.</emphasis>
Selected measurements and proportions of the species of
<taxonomicName id="4D8639816340FFC5FCA2FA0FFC01FA2A" authorityName="Peters" authorityYear="1867" box="[842,983,1462,1483]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B8F29E106340FFC5FCA2FA0FFC01FA2A" box="[842,983,1462,1483]" italics="true" pageId="7" pageNumber="8">Phrynoglossus</emphasis>
</taxonomicName>
. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text.
</paragraph>
</caption>
<paragraph id="8A3942026340FFC5FD59F9B7FA42F7DD" pageId="7" pageNumber="8">
<table id="F886B0A26340003DFEE9F9B4FA42F7DD" box="[257,1428,1549,2108]" colsContinueIn="8.[197,1365,272,655]" gridcols="5" gridrows="15" pageId="7" pageNumber="8">
<tr id="34B640406340003DFEE9F9B4FA42F9DA" box="[257,1428,1549,1595]" gridrow="0" pageId="7" pageNumber="8" rowspan-0="1" rowspan-1="1">
<th id="7767293C6340003DFD98F9B4FC8CF9DA" box="[624,858,1549,1595]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FD59F9B7FCCFF9C0" authorityName="Peters" authorityYear="1867" box="[689,793,1550,1569]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E106340FFC5FD59F9B7FCCFF9C0" bold="true" box="[689,793,1550,1569]" italics="true" pageId="7" pageNumber="8">P. martensii</emphasis>
</taxonomicName>
♂ 7 | ♀ 8
</th>
<th id="7767293C6340003DFC64F9B4FBA0F9DA" box="[908,1142,1549,1595]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FC41F9B7FB8EF9C1" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[937,1112,1549,1569]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E106340FFC5FC41F9B7FB8EF9C1" bold="true" box="[937,1112,1549,1569]" italics="true" pageId="7" pageNumber="8">P. magnapustulosus</emphasis>
</taxonomicName>
♂ 8 | ♀ 12
</th>
<th id="7767293C6340003DFB4CF9B4FA42F9DA" box="[1188,1428,1549,1595]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="4D8639816340FFC5FB36F9B7FA8FF9C1" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[1246,1369,1549,1569]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E106340FFC5FB36F9B7FA8FF9C1" bold="true" box="[1246,1369,1549,1569]" italics="true" pageId="7" pageNumber="8">P. myanhessei</emphasis>
</taxonomicName>
♂ 9 | ♀ 6
</th>
</tr>
<tr id="34B640406340003DFEE9F9F0FA42F9BD" box="[257,1428,1609,1628]" gridrow="1" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F9F0FEB7F9BD" box="[257,353,1609,1628]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">SVL</th>
<td id="7767293C6340003DFE30F9F0FDCCF9BD" box="[472,538,1609,1628]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F9F0FC8CF9BD" box="[624,858,1609,1628]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">19.7723.31 (21.42±1.28)</td>
<td id="7767293C6340003DFC64F9F0FBA0F9BD" box="[908,1142,1609,1628]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">17.2119.68 (17.91±0.804)</td>
<td id="7767293C6340003DFB4CF9F0FA42F9BD" box="[1188,1428,1609,1628]" gridcol="4" gridrow="1" pageId="7" pageNumber="8">22.0027.35 (24.06±1.65)</td>
</tr>
<tr id="34B640406340003DFEE9F9D7FA42F960" box="[257,1428,1646,1665]" gridrow="2" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F9D7FDCCF960" box="[472,538,1646,1665]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F9D7FC8CF960" box="[624,858,1646,1665]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">25.5530.35 (27.60±2.061)</td>
<td id="7767293C6340003DFC64F9D7FBA0F960" box="[908,1142,1646,1665]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">16.9024.30 (21.66±2.26)</td>
<td id="7767293C6340003DFB4CF9D7FA42F960" box="[1188,1428,1646,1665]" gridcol="4" gridrow="2" pageId="7" pageNumber="8">28.3831.27 (29.67±1.15)</td>
</tr>
<tr id="34B640406340003DFEE9F92AFA42F947" box="[257,1428,1683,1702]" gridrow="3" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F92AFEB7F947" box="[257,353,1683,1702]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">SHL / SVL</th>
<td id="7767293C6340003DFE30F92AFDCCF947" box="[472,538,1683,1702]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F92AFC8CF947" box="[624,858,1683,1702]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">0.4410.509 (0.479±0.021)</td>
<td id="7767293C6340003DFC64F92AFBA0F947" box="[908,1142,1683,1702]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">0.4420.557 (0.509±0.036)</td>
<td id="7767293C6340003DFB4CF92AFA42F947" box="[1188,1428,1683,1702]" gridcol="4" gridrow="3" pageId="7" pageNumber="8">0.4570.571 (0.506±0.032)</td>
</tr>
<tr id="34B640406340003DFEE9F901FA42F92A" box="[257,1428,1720,1739]" gridrow="4" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F901FDCCF92A" box="[472,538,1720,1739]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F901FC8CF92A" box="[624,858,1720,1739]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">0.4360.479 (0.462±0.016)</td>
<td id="7767293C6340003DFC64F901FBA0F92A" box="[908,1142,1720,1739]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">0.4020.548 (0.478±0.041)</td>
<td id="7767293C6340003DFB4CF901FA42F92A" box="[1188,1428,1720,1739]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">0.4660.592 (0.500±0.047)</td>
</tr>
<tr id="34B640406340003DFEE9F965FA42F911" box="[257,1428,1756,1776]" gridrow="5" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F965FEB7F911" box="[257,353,1756,1776]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">FL / SVL</th>
<td id="7767293C6340003DFE30F965FDCCF911" box="[472,538,1756,1776]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F965FC8CF911" box="[624,858,1756,1776]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">0.4710.535 (0.491±0.024)</td>
<td id="7767293C6340003DFC64F965FBA0F911" box="[908,1142,1756,1776]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">0.4730.564 (0.527±0.031)</td>
<td id="7767293C6340003DFB4CF965FA42F911" box="[1188,1428,1756,1776]" gridcol="4" gridrow="5" pageId="7" pageNumber="8">0.4330.535 (0.476±0.037)</td>
</tr>
<tr id="34B640406340003DFEE9F8B8FA42F8F4" box="[257,1428,1793,1813]" gridrow="6" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F8B8FDCCF8F4" box="[472,538,1793,1813]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F8B8FC8CF8F4" box="[624,858,1793,1813]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">0.4320.488 (0.468±0.021)</td>
<td id="7767293C6340003DFC64F8B8FBA0F8F4" box="[908,1142,1793,1813]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">0.4390.567 (0.482±0.045)</td>
<td id="7767293C6340003DFB4CF8B8FA42F8F4" box="[1188,1428,1793,1813]" gridcol="4" gridrow="6" pageId="7" pageNumber="8">0.4070.491 (0.449±0.033)</td>
</tr>
<tr id="34B640406340003DFEE9F89FFA42F8DB" box="[257,1428,1830,1850]" gridrow="7" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F89FFEB7F8DB" box="[257,353,1830,1850]" gridcol="0" gridrow="7" pageId="7" pageNumber="8">HL / SVL</th>
<td id="7767293C6340003DFE30F89FFDCCF8DB" box="[472,538,1830,1850]" gridcol="1" gridrow="7" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F89FFC8CF8DB" box="[624,858,1830,1850]" gridcol="2" gridrow="7" pageId="7" pageNumber="8">0.3410.383 (0.359±0.015)</td>
<td id="7767293C6340003DFC64F89FFBA0F8DB" box="[908,1142,1830,1850]" gridcol="3" gridrow="7" pageId="7" pageNumber="8">0.3130.377 (0.341±0.024)</td>
<td id="7767293C6340003DFB4CF89FFA42F8DB" box="[1188,1428,1830,1850]" gridcol="4" gridrow="7" pageId="7" pageNumber="8">0.2680.378 (0.330±0.031)</td>
</tr>
<tr id="34B640406340003DFEE9F8F2FA42F8BF" box="[257,1428,1867,1886]" gridrow="8" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F8F2FDCCF8BF" box="[472,538,1867,1886]" gridcol="1" gridrow="8" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F8F2FC8CF8BF" box="[624,858,1867,1886]" gridcol="2" gridrow="8" pageId="7" pageNumber="8">0.3110.383 (0.344±0.022)</td>
<td id="7767293C6340003DFC64F8F2FBA0F8BF" box="[908,1142,1867,1886]" gridcol="3" gridrow="8" pageId="7" pageNumber="8">0.2720.347 (0.304±0.028)</td>
<td id="7767293C6340003DFB4CF8F2FA42F8BF" box="[1188,1428,1867,1886]" gridcol="4" gridrow="8" pageId="7" pageNumber="8">0.2720.304 (0.289±0.011)</td>
</tr>
<tr id="34B640406340003DFEE9F8C9FA42F865" box="[257,1428,1904,1924]" gridrow="9" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F8C9FEB7F865" box="[257,353,1904,1924]" gridcol="0" gridrow="9" pageId="7" pageNumber="8">HW / SVL</th>
<td id="7767293C6340003DFE30F8C9FDCCF865" box="[472,538,1904,1924]" gridcol="1" gridrow="9" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F8C9FC8CF865" box="[624,858,1904,1924]" gridcol="2" gridrow="9" pageId="7" pageNumber="8">0.3360.401 (0.357±0.023)</td>
<td id="7767293C6340003DFC64F8C9FBA0F865" box="[908,1142,1904,1924]" gridcol="3" gridrow="9" pageId="7" pageNumber="8">0.3350.394 (0.364±0.021)</td>
<td id="7767293C6340003DFB4CF8C9FA42F865" box="[1188,1428,1904,1924]" gridcol="4" gridrow="9" pageId="7" pageNumber="8">0.2860.369 (0.338±0.029)</td>
</tr>
<tr id="34B640406340003DFEE9F82CFA42F849" box="[257,1428,1941,1960]" gridrow="10" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F82CFDCCF849" box="[472,538,1941,1960]" gridcol="1" gridrow="10" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F82CFC8CF849" box="[624,858,1941,1960]" gridcol="2" gridrow="10" pageId="7" pageNumber="8">0.3380.388 (0.351±0.017)</td>
<td id="7767293C6340003DFC64F82CFBA0F849" box="[908,1142,1941,1960]" gridcol="3" gridrow="10" pageId="7" pageNumber="8">0.2950.360 (0.336±0.019)</td>
<td id="7767293C6340003DFB4CF82CFA42F849" box="[1188,1428,1941,1960]" gridcol="4" gridrow="10" pageId="7" pageNumber="8">0.2830.354 (0.312±0.025)</td>
</tr>
<tr id="34B640406340003DFEE9F803FA42F82C" box="[257,1428,1978,1997]" gridrow="11" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F803FEB7F82C" box="[257,353,1978,1997]" gridcol="0" gridrow="11" pageId="7" pageNumber="8">HL / HW</th>
<td id="7767293C6340003DFE30F803FDCCF82C" box="[472,538,1978,1997]" gridcol="1" gridrow="11" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F803FC8CF82C" box="[624,858,1978,1997]" gridcol="2" gridrow="11" pageId="7" pageNumber="8">0.9091.077 (1.011±0.063)</td>
<td id="7767293C6340003DFC64F803FBA0F82C" box="[908,1142,1978,1997]" gridcol="3" gridrow="11" pageId="7" pageNumber="8">0.8551.030 (0.937±0.058)</td>
<td id="7767293C6340003DFB4CF803FA42F82C" box="[1188,1428,1978,1997]" gridcol="4" gridrow="11" pageId="7" pageNumber="8">0.8971.191 (0.980±0.092)</td>
</tr>
<tr id="34B640406340003DFEE9F866FA42F813" box="[257,1428,2015,2034]" gridrow="12" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F866FDCCF813" box="[472,538,2015,2034]" gridcol="1" gridrow="12" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F866FC8CF813" box="[624,858,2015,2034]" gridcol="2" gridrow="12" pageId="7" pageNumber="8">0.9201.020 (0.980±0.031)</td>
<td id="7767293C6340003DFC64F866FBA0F813" box="[908,1142,2015,2034]" gridcol="3" gridrow="12" pageId="7" pageNumber="8">0.7791.057 (0.907±0.098)</td>
<td id="7767293C6340003DFB4CF866FA42F813" box="[1188,1428,2015,2034]" gridcol="4" gridrow="12" pageId="7" pageNumber="8">0.8081.043 (0.929±0.093)</td>
</tr>
<tr id="34B640406340003DFEE9F7BDFA42F7F6" box="[257,1428,2052,2071]" gridrow="13" pageId="7" pageNumber="8">
<th id="7767293C6340003DFEE9F7BDFEB7F7F6" box="[257,353,2052,2071]" gridcol="0" gridrow="13" pageId="7" pageNumber="8">IOD / SVL</th>
<td id="7767293C6340003DFE30F7BDFDCCF7F6" box="[472,538,2052,2071]" gridcol="1" gridrow="13" pageId="7" pageNumber="8">males</td>
<td id="7767293C6340003DFD98F7BDFC8CF7F6" box="[624,858,2052,2071]" gridcol="2" gridrow="13" pageId="7" pageNumber="8">0.0710.091 (0.086±0.007)</td>
<td id="7767293C6340003DFC64F7BDFBA0F7F6" box="[908,1142,2052,2071]" gridcol="3" gridrow="13" pageId="7" pageNumber="8">0.0930.135 (0.112±0.015)</td>
<td id="7767293C6340003DFB4CF7BDFA42F7F6" box="[1188,1428,2052,2071]" gridcol="4" gridrow="13" pageId="7" pageNumber="8">0.0900.132 (0.107±0.013)</td>
</tr>
<tr id="34B640406340003DFEE9F790FA42F7DD" box="[257,1428,2089,2108]" gridrow="14" pageId="7" pageNumber="8" rowspan-0="1">
<td id="7767293C6340003DFE30F790FDCCF7DD" box="[472,538,2089,2108]" gridcol="1" gridrow="14" pageId="7" pageNumber="8">females</td>
<td id="7767293C6340003DFD98F790FC8CF7DD" box="[624,858,2089,2108]" gridcol="2" gridrow="14" pageId="7" pageNumber="8">0.0730.084 (0.077±0.003)</td>
<td id="7767293C6340003DFC64F790FBA0F7DD" box="[908,1142,2089,2108]" gridcol="3" gridrow="14" pageId="7" pageNumber="8">0.0840.137 (0.106±0.018)</td>
<td id="7767293C6340003DFB4CF790FA42F7DD" box="[1188,1428,2089,2108]" gridcol="4" gridrow="14" pageId="7" pageNumber="8">0.0770.163 (0.104±0.032)</td>
</tr>
</table>
</paragraph>
<caption id="DEF9128A634FFFCAFF60FF6FFEE8FF0D" box="[136,318,214,236]" isContinuationCaption="true" pageId="8" pageNumber="9" startId="8.[136,192,214,235]" targetType="table">
<paragraph id="8A394202634FFFCAFF60FF6FFEE8FF0D" blockId="8.[136,318,214,236]" box="[136,318,214,236]" pageId="8" pageNumber="9">
<emphasis id="B8F29E10634FFFCAFF60FF6FFF05FF0A" bold="true" box="[136,211,214,235]" pageId="8" pageNumber="9">Table 3</emphasis>
continued.
</paragraph>
</caption>
<paragraph id="8A394202634FFFCAFD9DFEA9FA83FD6E" pageId="8" pageNumber="9">
<table id="F886B0A2634F003DFF2DFEA9FA83FD6E" box="[197,1365,272,655]" colsContinueFrom="7.[257,1428,1549,2108]" gridcols="5" gridrows="9" pageId="8" pageNumber="9">
<tr id="34B64040634F003DFF2DFEA9FA83FEDC" box="[197,1365,272,317]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1" rowspan-1="1">
<th id="7767293C634F003DFDDCFEA9FCC8FEDC" box="[564,798,272,317]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
<taxonomicName id="4D863981634FFFCAFD9DFEA9FD0BFEC2" authorityName="Peters" authorityYear="1867" box="[629,733,272,291]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634FFFCAFD9DFEA9FD0BFEC2" bold="true" box="[629,733,272,291]" italics="true" pageId="8" pageNumber="9">P. martensii</emphasis>
</taxonomicName>
♂ 7 | ♀ 8
</th>
<th id="7767293C634F003DFCA7FEA9FBEFFEDC" box="[847,1081,272,317]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">
<taxonomicName id="4D863981634FFFCAFC84FEA9FBCDFEC2" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[876,1051,272,291]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634FFFCAFC84FEA9FBCDFEC2" bold="true" box="[876,1051,272,291]" italics="true" pageId="8" pageNumber="9">P. magnapustulosus</emphasis>
</taxonomicName>
♂ 8 | ♀ 12
</th>
<th id="7767293C634F003DFB83FEA9FA83FEDC" box="[1131,1365,272,317]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">
<taxonomicName id="4D863981634FFFCAFB4AFEA9FACBFEC2" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[1186,1309,272,291]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634FFFCAFB4AFEA9FACBFEC2" bold="true" box="[1186,1309,272,291]" italics="true" pageId="8" pageNumber="9">P. myanhessei</emphasis>
</taxonomicName>
♂ 9 | ♀ 6
</th>
</tr>
<tr id="34B64040634F003DFF2DFEF2FA83FEBE" box="[197,1365,331,351]" gridrow="1" pageId="8" pageNumber="9">
<th id="7767293C634F003DFF2DFEF2FEFBFEBE" box="[197,301,331,351]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">TYD / SVL</th>
<td id="7767293C634F003DFE73FEF2FE0BFEBE" box="[411,477,331,351]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">males</td>
<td id="7767293C634F003DFDDCFEF2FCC8FEBE" box="[564,798,331,351]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">0.0420.068 (0.052±0.010)</td>
<td id="7767293C634F003DFCA7FEF2FBEFFEBE" box="[847,1081,331,351]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">0.0550.099 (0.082±0.014)</td>
<td id="7767293C634F003DFB83FEF2FA83FEBE" box="[1131,1365,331,351]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">0.0290.066 (0.050±0.012)</td>
</tr>
<tr id="34B64040634F003DFF2DFECEFA83FE6B" box="[197,1365,375,394]" gridrow="2" pageId="8" pageNumber="9" rowspan-0="1">
<td id="7767293C634F003DFE73FECEFE0BFE6B" box="[411,477,375,394]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">females</td>
<td id="7767293C634F003DFDDCFECEFCC8FE6B" box="[564,798,375,394]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">0.0350.052 (0.043±0.006)</td>
<td id="7767293C634F003DFCA7FECEFBEFFE6B" box="[847,1081,375,394]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">0.0620.102 (0.080±0.012)</td>
<td id="7767293C634F003DFB83FECEFA83FE6B" box="[1131,1365,375,394]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">0.0340.064 (0.051±0.010)</td>
</tr>
<tr id="34B64040634F003DFF2DFE1BFA83FE57" box="[197,1365,418,438]" gridrow="3" pageId="8" pageNumber="9">
<th id="7767293C634F003DFF2DFE1BFEFBFE57" box="[197,301,418,438]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">EYD / SVL</th>
<td id="7767293C634F003DFE73FE1BFE0BFE57" box="[411,477,418,438]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">males</td>
<td id="7767293C634F003DFDDCFE1BFCC8FE57" box="[564,798,418,438]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">0.1090.138 (0.122±0.010)</td>
<td id="7767293C634F003DFCA7FE1BFBEFFE57" box="[847,1081,418,438]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">0.0910.143 (0.116±0.017)</td>
<td id="7767293C634F003DFB83FE1BFA83FE57" box="[1131,1365,418,438]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">0.0940.140 (0.112±0.015)</td>
</tr>
<tr id="34B64040634F003DFF2DFE77FA83FE00" box="[197,1365,462,481]" gridrow="4" pageId="8" pageNumber="9" rowspan-0="1">
<td id="7767293C634F003DFE73FE77FE0BFE00" box="[411,477,462,481]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">females</td>
<td id="7767293C634F003DFDDCFE77FCC8FE00" box="[564,798,462,481]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">0.0950.119 (0.104±0.008)</td>
<td id="7767293C634F003DFCA7FE77FBEFFE00" box="[847,1081,462,481]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">0.0820.118 (0.100±0.012)</td>
<td id="7767293C634F003DFB83FE77FA83FE00" box="[1131,1365,462,481]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">0.0770.132 (0.096±0.019)</td>
</tr>
<tr id="34B64040634F003DFF2DFE43FA83FDEC" box="[197,1365,506,525]" gridrow="5" pageId="8" pageNumber="9">
<th id="7767293C634F003DFF2DFE43FEFBFDEC" box="[197,301,506,525]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">NED / SVL</th>
<td id="7767293C634F003DFE73FE43FE0BFDEC" box="[411,477,506,525]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">males</td>
<td id="7767293C634F003DFDDCFE43FCC8FDEC" box="[564,798,506,525]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">0.0560.071 (0.064±0.006)</td>
<td id="7767293C634F003DFCA7FE43FBEFFDEC" box="[847,1081,506,525]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">0.0640.123 (0.086±0.018)</td>
<td id="7767293C634F003DFB83FE43FA83FDEC" box="[1131,1365,506,525]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">0.0610.101 (0.083±0.016)</td>
</tr>
<tr id="34B64040634F003DFF2DFD9CFA83FDD9" box="[197,1365,549,568]" gridrow="6" pageId="8" pageNumber="9" rowspan-0="1">
<td id="7767293C634F003DFE73FD9CFE0BFDD9" box="[411,477,549,568]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">females</td>
<td id="7767293C634F003DFDDCFD9CFCC8FDD9" box="[564,798,549,568]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">0.0550.066 (0.060±0.004)</td>
<td id="7767293C634F003DFCA7FD9CFBEFFDD9" box="[847,1081,549,568]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">0.0580.086 (0.074±0.009)</td>
<td id="7767293C634F003DFB83FD9CFA83FDD9" box="[1131,1365,549,568]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">0.0600.145 (0.090±0.033)</td>
</tr>
<tr id="34B64040634F003DFF2DFDE8FA83FD85" box="[197,1365,593,612]" gridrow="7" pageId="8" pageNumber="9">
<th id="7767293C634F003DFF2DFDE8FEFBFD85" box="[197,301,593,612]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">TYD / EYD</th>
<td id="7767293C634F003DFE73FDE8FE0BFD85" box="[411,477,593,612]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">males</td>
<td id="7767293C634F003DFDDCFDE8FCC8FD85" box="[564,798,593,612]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">0.3570.512 (0.425±0.063)</td>
<td id="7767293C634F003DFCA7FDE8FBEFFD85" box="[847,1081,593,612]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">0.4820.920 (0.722±0.147)</td>
<td id="7767293C634F003DFB83FDE8FA83FD85" box="[1131,1365,593,612]" gridcol="4" gridrow="7" pageId="8" pageNumber="9">0.2910.653 (0.460±0.146)</td>
</tr>
<tr id="34B64040634F003DFF2DFDC5FA83FD6E" box="[197,1365,636,655]" gridrow="8" pageId="8" pageNumber="9" rowspan-0="1">
<td id="7767293C634F003DFE73FDC5FE0BFD6E" box="[411,477,636,655]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">females</td>
<td id="7767293C634F003DFDDCFDC5FCC8FD6E" box="[564,798,636,655]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">0.3280.515 (0.411±0.060)</td>
<td id="7767293C634F003DFCA7FDC5FBEFFD6E" box="[847,1081,636,655]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">0.5910.962 (0.807±0.120)</td>
<td id="7767293C634F003DFB83FDC5FA83FD6E" box="[1131,1365,636,655]" gridcol="4" gridrow="8" pageId="8" pageNumber="9">0.3850.727 (0.550±0.143)</td>
</tr>
</table>
</paragraph>
<caption id="DEF9128A634FFFCAFF60F7AFFCA8F7AD" pageId="8" pageNumber="9" startId="8.[136,204,2070,2092]" targetBox="[136,1391,742,2050]" targetPageId="8" targetType="figure">
<paragraph id="8A394202634FFFCAFF60F7AFFCA8F7AD" blockId="8.[136,1391,2070,2124]" pageId="8" pageNumber="9">
<emphasis id="B8F29E10634FFFCAFF60F7AFFF32F7CA" bold="true" box="[136,228,2070,2092]" pageId="8" pageNumber="9">Figure 4.</emphasis>
Advertisement calls of male
<taxonomicName id="4D863981634FFFCAFE17F7AFFD5AF7CA" authorityName="Peters" authorityYear="1867" box="[511,652,2070,2091]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B8F29E10634FFFCAFE17F7AFFD5AF7CA" box="[511,652,2070,2091]" italics="true" pageId="8" pageNumber="9">Phrynoglossus</emphasis>
</taxonomicName>
. (A)
<taxonomicName id="4D863981634FFFCAFD28F7AEFC56F7CA" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[704,896,2070,2092]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634FFFCAFD28F7AEFC56F7CA" box="[704,896,2070,2092]" italics="true" pageId="8" pageNumber="9">P. magnapustulosus</emphasis>
</taxonomicName>
, GK-7395; (B)
<taxonomicName id="4D863981634FFFCAFBF2F7AEFB77F7CA" authority="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti, 2021" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[1050,1185,2070,2092]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="myanhessei" status="n. sp.">
<emphasis id="B8F29E10634FFFCAFBF2F7AEFB77F7CA" box="[1050,1185,2070,2092]" italics="true" pageId="8" pageNumber="9">P. myanhessei</emphasis>
</taxonomicName>
<emphasis id="B8F29E10634FFFCAFB4FF7AFFB0BF7CA" bold="true" box="[1191,1245,2070,2091]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A3C1236B634FFFCAFB4FF7AFFB0BF7CA" box="[1191,1245,2070,2091]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SMF 103799; (C)
<emphasis id="B8F29E10634FFFCAFF46F78EFEF1F7AD" box="[174,295,2103,2124]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4D863981634FFFCAFF46F78EFEF5F7AD" authorityName="Peters" authorityYear="1867" box="[174,291,2103,2124]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="martensii">P. martensii</taxonomicName>
,
</emphasis>
PT-2634 (Bangkok); (D)
<emphasis id="B8F29E10634FFFCAFDF7F78EFD41F7AD" box="[543,663,2103,2124]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4D863981634FFFCAFDF7F78EFD45F7AD" authorityName="Peters" authorityYear="1867" box="[543,659,2103,2124]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="martensii">P. martensii</taxonomicName>
,
</emphasis>
PT-2076 (Chiang Mai).
</paragraph>
</caption>
<caption id="DEF9128A634EFFCBFF2CF8E0FA91F86E" pageId="9" pageNumber="10" startId="9.[196,264,1881,1903]" targetBox="[202,1449,205,1861]" targetPageId="9" targetType="figure">
<paragraph id="8A394202634EFFCBFF2CF8E0FA91F86E" blockId="9.[196,1452,1881,1935]" pageId="9" pageNumber="10">
<emphasis id="B8F29E10634EFFCBFF2CF8E0FEA4F88F" bold="true" box="[196,370,1881,1903]" pageId="9" pageNumber="10">Figure 5 part 1.</emphasis>
Scatter plots illustrating morphological variation in the species of
<taxonomicName id="4D863981634EFFCBFC0EF8E0FBA5F88F" authorityName="Peters" authorityYear="1867" box="[998,1139,1881,1902]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B8F29E10634EFFCBFC0EF8E0FBA5F88F" box="[998,1139,1881,1902]" italics="true" pageId="9" pageNumber="10">Phrynoglossus</emphasis>
</taxonomicName>
. magnap. =
<taxonomicName id="4D863981634EFFCBFB00F8E3FA73F88F" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[1256,1445,1881,1903]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634EFFCBFB00F8E3FA73F88F" box="[1256,1445,1881,1903]" italics="true" pageId="9" pageNumber="10">P. magnapustulosus</emphasis>
</taxonomicName>
; m = male; f = female. For abbreviations of morphological characters see text. Abbreviations of taxon names as in Fig. 1.
</paragraph>
</caption>
<subSubSection id="C29C1189634EFFC8FC81F86DFBF8F7AD" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="conservation">
<paragraph id="8A394202634EFFC8FC81F86DFBF8F7AD" blockId="9.[843,1452,2004,2124]" lastBlockId="10.[782,1392,2004,2124]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
This published work and the nomenclatural acts it contains have been registered in ZooBank, the online registration system for the International Commission on Zoological Nomenclature (
<collectionCode id="EC97DAC7634EFFCBFC04F78DFBF9F7AD" box="[1004,1071,2100,2124]" pageId="9" pageNumber="10">ICZN</collectionCode>
). The ZooBank LSIDs (Life Science Identifiers) can be resolved and the associated information can be viewed through any standard web browser by appending the LSID to the prefix http://zoobank.org. The LSID for this publication is as follows: urn:lsid:zoobank. org:pub:A978D105-6D7F-4858-BA27-8639B76A6039. The LSID registration and any associated information can be viewed in a web browser by adding the LSID to the prefix “http://zoobank.org.”
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,132 +0,0 @@
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<mods:title id="0AF89BF52DA1D7A627343CA08BD88AA6">A taxonomic revision of the genus Phrynoglossus in Indochina with the description of a new species and comments on the classification within Occidozyginae (Amphibia, Anura, Dicroglossidae)</mods:title>
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<mods:namePart id="B6901BA9BF25806A4624E4B5D3FACEC5">Schell, Tilman</mods:namePart>
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<paragraph id="8A3942026342FFC7FCA3F969FAADF90F" blockId="5.[843,1403,1744,1774]" box="[843,1403,1744,1774]" pageId="5" pageNumber="6">
<heading id="D171F56E6342FFC7FCA3F969FAADF90F" bold="true" box="[843,1403,1744,1774]" fontSize="12" level="2" pageId="5" pageNumber="6" reason="0">
<taxonomicName id="4D8639816342FFC7FCA3F969FAADF90F" ID-CoL="66CF" authority="Kuhl &amp; van Hasselt, 1822" authorityName="Kuhl &amp; van Hasselt" authorityYear="1822" box="[843,1403,1744,1774]" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B8F29E106342FFC7FCA3F969FAADF90F" bold="true" box="[843,1403,1744,1774]" pageId="5" pageNumber="6">
<emphasis id="B8F29E106342FFC7FCA3F969FC2AF90F" bold="true" box="[843,1020,1744,1774]" italics="true" pageId="5" pageNumber="6">Occidozyga</emphasis>
<bibRefCitation id="EE173FF36342FFC7FBECF969FAADF90F" author="Kuhl H van &amp; Hasselt JC" box="[1028,1403,1744,1774]" pageId="5" pageNumber="6" pagination="99 - 104" refId="ref12788" refString="Kuhl H van, Hasselt JC (1822) Uittreksels uit breieven van de Heeren Kuhl en van Hasselt, aan de Heeren C. J. Temminck, Th. van Swinderen en W. de Haan. Algemeene Konst-En Letter-Bode 7: 99 - 104." type="journal article" year="1822">Kuhl &amp; van Hasselt, 1822</bibRefCitation>
</emphasis>
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<emphasis id="B8F29E106342FFC7FCA3F8ADFC09F8CD" bold="true" box="[843,991,1812,1836]" pageId="5" pageNumber="6">
<typeStatus id="553DFCA06342FFC7FCA3F8ADFC55F8CD" box="[843,899,1812,1836]" pageId="5" pageNumber="6">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4D8639816342FFC7FC0DF8ACFAF2F8CD" authority="Gravenhorst, 1829" authorityName="Gravenhorst" authorityYear="1829" box="[997,1316,1812,1836]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="lima">
<emphasis id="B8F29E106342FFC7FC0DF8ACFB85F8CD" box="[997,1107,1812,1836]" italics="true" pageId="5" pageNumber="6">Rana lima</emphasis>
<bibRefCitation id="EE173FF36342FFC7FBB2F8ADFAF2F8CD" author="Gravenhorst JLC" box="[1114,1316,1812,1836]" pageId="5" pageNumber="6" refId="ref11956" refString="Gravenhorst JLC (1829) Deliciae Musei Zoologici Vratislaviensis. Fasciculus primus. Chelonios et Batrachia. Leopold Voss, Leipzig." type="book" year="1829">Gravenhorst, 1829</bibRefCitation>
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.
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<paragraph id="8A3942026342FFC4FCA3F8EDFE1BF90D" blockId="5.[843,1452,1876,2124]" lastBlockId="6.[136,744,1716,1772]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B8F29E106342FFC7FCA3F8EDFC16F88D" bold="true" box="[843,960,1876,1900]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
A genus of Asian frogs of the subfamily
<taxonomicName id="4D8639816342FFC7FA98F8EDFB18F86D" authority="Fei, Ye, and Huang, 1990" authorityName="Fei, Ye, and Huang" authorityYear="1990" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="subFamily" subFamily="Occidozyginae">Occidozyginae Fei, Ye, and Huang, 1990</taxonomicName>
of the family
<taxonomicName id="4D8639816342FFC7FA8EF8CDFBB6F84D" authority="Anderson, 1871" authorityName="Anderson" authorityYear="1871" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="family">
Dicroglossidae
<bibRefCitation id="EE173FF36342FFC7FC5BF82DFBB6F84D" author="Anderson J" box="[947,1120,1940,1964]" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="5" pageNumber="6" pagination="12 - 39" part="40" refId="ref11030" refString="Anderson J (1871) A list of the reptilian accession to the Indian Museum, Calcutta from 1865 to 1870, with a description of some new species. Journal of the Asiatic Society of Bengal 40: 12 - 39." title="A list of the reptilian accession to the Indian Museum, Calcutta from 1865 to 1870, with a description of some new species" type="journal article" year="1871">Anderson, 1871</bibRefCitation>
</taxonomicName>
, that differs from all other genera of its subfamily by having the following combination of characters: (1); tongue slender, worm-like; (2) vomerine teeth absent; (3) tips of fingers and toes pointed; (4) tympanum hidden; (5) skin not covered by extensive mucous, feels dry to touch in life frogs; (6) throat lining whitish with longitudinal brown stripe; (7) life style fully aquatic; (8) amplexus axillary.
</paragraph>
</subSubSection>
<caption id="DEF9128A6341FFC4FF60F98FFAC8F98A" pageId="6" pageNumber="7" startId="6.[136,204,1590,1612]" targetBox="[134,1506,257,1512]" targetPageId="6" targetType="figure">
<paragraph id="8A3942026341FFC4FF60F98FFAC8F98A" blockId="6.[136,1391,1590,1644]" pageId="6" pageNumber="7">
<emphasis id="B8F29E106341FFC4FF60F98FFF31F9AA" bold="true" box="[136,231,1590,1612]" pageId="6" pageNumber="7">Figure 3.</emphasis>
Species delimitation analysis of frogs of the subfamily
<taxonomicName id="4D8639816341FFC4FCF1F98FFC7FF9AD" authorityName="Fei, Ye, and Huang" authorityYear="1990" box="[793,937,1590,1612]" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="subFamily" subFamily="Occidozyginae">Occidozyginae</taxonomicName>
using the Automatic Barcode Gap Discovery (ABGD) approach; initial partition with prior maximal distance at 0.00167. See text for details. Abbreviations as in Fig. 1.
</paragraph>
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<paragraph id="8A3942026341FFC4FF60F8ADFD53F8CD" blockId="6.[136,645,1812,1836]" box="[136,645,1812,1836]" pageId="6" pageNumber="7">
<emphasis id="B8F29E106341FFC4FF60F8ADFF3DF8CD" bold="true" box="[136,235,1812,1836]" pageId="6" pageNumber="7">Content.</emphasis>
<taxonomicName id="4D8639816341FFC4FF19F8ADFD53F8CD" authority="(Gravenhorst, 1829)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[241,645,1812,1836]" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lima">
<emphasis id="B8F29E106341FFC4FF19F8ADFE73F8CD" box="[241,421,1812,1836]" italics="true" pageId="6" pageNumber="7">Occidozyga lima</emphasis>
(
<bibRefCitation id="EE173FF36341FFC4FE5BF8ADFDABF8CD" author="Gravenhorst JLC" box="[435,637,1812,1836]" pageId="6" pageNumber="7" refId="ref11956" refString="Gravenhorst JLC (1829) Deliciae Musei Zoologici Vratislaviensis. Fasciculus primus. Chelonios et Batrachia. Leopold Voss, Leipzig." type="book" year="1829">Gravenhorst, 1829</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,439 +0,0 @@
<document id="3E4804D25AAA2D0B4FF3D29C55F698C1" ID-DOI="10.3897/vz.71.e60312" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1737914713964" checkinUser="juliana" docAuthor="Köhler, Gunther, Vargas, Joseph, Than, Ni Lar, Schell, Tilman, Janke, Axel, Pauls, Steffen U. &amp; Thammachoti, Panupong" docDate="2021" docId="022FF314634BFFCFFF60FF69FC46F7AD" docLanguage="en" docName="VertZool.71.1-26.pdf" docOrigin="Vertebrate Zoology 71" docSource="https://doi.org/10.3897/vz.71.e60312" docTitle="Phrynoglossus myanhessei Kohler, Vargas, Than &amp; Thammachoti 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="14" masterDocId="FE168B6C6347FFC2FFE8FFB9FFD6FFE1" masterDocTitle="A taxonomic revision of the genus Phrynoglossus in Indochina with the description of a new species and comments on the classification within Occidozyginae (Amphibia, Anura, Dicroglossidae)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="13" updateTime="1737914716862" updateUser="juliana">
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<mods:title id="29D2EB5622A76A3A2283DA71C7E93B7F">A taxonomic revision of the genus Phrynoglossus in Indochina with the description of a new species and comments on the classification within Occidozyginae (Amphibia, Anura, Dicroglossidae)</mods:title>
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<mods:namePart id="1E46142136A8B51D7AC3AF0F5E13D974">Köhler, Gunther</mods:namePart>
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<mods:namePart id="36AD023160924D1A7CC669713C5079A6">Vargas, Joseph</mods:namePart>
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<mods:namePart id="5FE5E0BADEA47EA31171849CF0BA4396">Than, Ni Lar</mods:namePart>
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<mods:roleTerm id="70FA1FF91C121AAC7267BF24356FF58F">Author</mods:roleTerm>
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<mods:namePart id="8B55794BAFE29ED0564FF777A0428C62">Schell, Tilman</mods:namePart>
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<mods:namePart id="70422FB912B58256E202C5588D788362">Janke, Axel</mods:namePart>
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<mods:role id="59BFE3F6D313289192955B4C8B9BE001">
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<mods:namePart id="5E1D3B09FC8EF98529B99B8B284933BC">Pauls, Steffen U.</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="9FDA9B5A0A87D13E879CFBDC26C15B05">Thammachoti, Panupong</mods:namePart>
</mods:name>
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<mods:date id="F8A1DAC373ECBF74BD4E798D4BF8DCC5">2021</mods:date>
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<treatment id="022FF314634BFFCFFF60FF69FC46F7AD" LSID="urn:lsid:plazi:treatment:022FF314634BFFCFFF60FF69FC46F7AD" httpUri="http://treatment.plazi.org/id/022FF314634BFFCFFF60FF69FC46F7AD" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C29C1189634BFFCEFF60FF69FD43FEF5" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8A394202634BFFCEFF60FF69FD43FEF5" blockId="12.[136,725,208,276]" pageId="12" pageNumber="13">
<heading id="D171F56E634BFFCEFF60FF69FD43FEF5" bold="true" fontSize="12" level="2" pageId="12" pageNumber="13" reason="0">
<emphasis id="B8F29E10634BFFCEFF60FF69FD43FEF5" bold="true" pageId="12" pageNumber="13">
<taxonomicName id="4D863981634BFFCEFF60FF69FDCFFEF5" authority="Kohler, Vargas, Than &amp; Thammachoti" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei" status="sp. nov.">
<emphasis id="B8F29E10634BFFCEFF60FF69FDCFFF0F" bold="true" box="[136,537,208,238]" italics="true" pageId="12" pageNumber="13">Phrynoglossus myanhessei</emphasis>
Köhler, Vargas, Than &amp; Thammachoti
</taxonomicName>
<taxonomicNameLabel id="A3C1236B634BFFCEFDC9FF4FFD43FEF5" box="[545,661,246,276]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C29C1189634BFFCEFF60FE8FFF38FE6A" pageId="12" pageNumber="13" type="description">
<paragraph id="8A394202634BFFCEFF60FE8FFD3FFEAD" blockId="12.[136,745,310,332]" box="[136,745,310,332]" pageId="12" pageNumber="13">http://zoobank.org/4D51EF6E-E61D-4A91-90EC-56B160400126</paragraph>
<paragraph id="8A394202634BFFCEFF60FECFFF38FE6A" blockId="12.[136,238,374,396]" box="[136,238,374,396]" pageId="12" pageNumber="13">
<figureCitation id="12BD5E87634BFFCEFF60FECFFF38FE6A" box="[136,238,374,396]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStartId-0="13.[196,264,1494,1516]" captionStartId-1="14.[136,204,1494,1516]" captionStartId-2="15.[196,264,1046,1068]" captionTargetBox-0="[196,1451,198,1474]" captionTargetBox-1="[136,1391,198,1474]" captionTargetBox-2="[196,1451,198,1026]" captionTargetId-0="figure-238@13.[196,1451,198,1474]" captionTargetId-1="figure-245@14.[136,1391,198,1474]" captionTargetId-2="figure-566@15.[196,1451,198,1026]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figure 8. Holotype of Phrynoglossus myanhessei n. sp. (SMF 103841). Scale bars equal 10.0 mm (A, B) and 5 mm (CF). Photos by G.K." captionText-1="Figure 9. Holotype of Phrynoglossus myanhessei sp. n. (SMF 103841). Scale bars equal 5.0 mm. Photos by G.K." captionText-2="Figure 10. Phrynoglossus myanhessei sp. n. in life. (A) Male holotype, SMF 103841; (B) male paratype, SMF 103840; (C) male paratype, SMF 103799; (D) female paratype, SMF 103798. Photos by G.K." pageId="12" pageNumber="13">Figs. 810</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634BFFCEFF60FE01FE89FD70" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8A394202634BFFCEFF60FE01FEF0FDFA" blockId="12.[136,746,440,539]" pageId="12" pageNumber="13">
<materialsCitation id="3AEE485F634BFFCEFF60FE01FEF4FDFA" collectingDate="2017-07-06" collectionCode="SMF" collectorName="Gunther Kohler &amp; Ni Lar Than" country="Myanmar" elevation="17" latitude="16.77737" location="East Yangon University" longLatPrecision="1" longitude="96.24065" municipality="Thanlyin" pageId="12" pageNumber="13" specimenCode="SMF 103841" specimenCount="1" specimenCount-male="1" stateProvince="Yangon" typeStatus="holotype">
<emphasis id="B8F29E10634BFFCEFF60FE01FF37FE2A" bold="true" box="[136,225,440,459]" pageId="12" pageNumber="13">
<typeStatus id="553DFCA0634BFFCEFF60FE01FF0BFE2A" box="[136,221,440,459]" pageId="12" pageNumber="13">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DA20EA79634BFFCEFF0FFE01FE8DFE2A" box="[231,347,440,459]" pageId="12" pageNumber="13">SMF 103841</specimenCode>
, an adult
<specimenCount id="9C80898B634BFFCEFE5BFE01FE0AFE2A" box="[435,476,440,459]" pageId="12" pageNumber="13" type="male">male</specimenCount>
collected at
<location id="8F5914D9634BFFCEFDA6FE00FF14FE07" LSID="urn:lsid:plazi:treatment:022FF314634BFFCFFF60FF69FC46F7AD:8F5914D9634BFFCEFDA6FE00FF14FE07" country="Myanmar" latitude="16.77737" longLatPrecision="1" longitude="96.24065" municipality="Thanlyin" name="East Yangon University" pageId="12" pageNumber="13" stateProvince="Yangon">East Yangon University</location>
(
<geoCoordinate id="EFB224C5634BFFCEFF25FE6AFEF6FE07" box="[205,288,467,486]" degrees="16.77737" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="16.77737">16.77737</geoCoordinate>
,
<geoCoordinate id="EFB224C5634BFFCEFEC1FE6AFEAAFE07" box="[297,380,467,486]" degrees="96.24065" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="96.24065">96.24065</geoCoordinate>
;
<quantity id="4D7EEFE7634BFFCEFE6DFE6AFE66FE07" box="[389,432,467,486]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="m" value="17.0">
<elevation id="01ABA531634BFFCEFE6DFE6AFE66FE07" box="[389,432,467,486]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="m" value="17.0">17 m</elevation>
</quantity>
a.s.l.),
<collectingMunicipality id="6A5DD878634BFFCEFE06FE6AFDEBFE07" box="[494,573,467,486]" pageId="12" pageNumber="13">Thanlyin</collectingMunicipality>
,
<collectingRegion id="48428CE0634BFFCEFDADFE6AFD5FFE07" box="[581,649,467,486]" pageId="12" pageNumber="13">Yangon</collectingRegion>
,
<collectingCountry id="F2910292634BFFCEFD79FE6AFD33FE07" box="[657,741,467,486]" name="Myanmar" pageId="12" pageNumber="13">Myanmar</collectingCountry>
, collected
<date id="FE3864C2634BFFCEFF35FE57FE93FDE0" box="[221,325,494,513]" pageId="12" pageNumber="13" value="2017-07-06">
<collectingDate id="EE7C9D2A634BFFCEFF35FE57FE93FDE0" box="[221,325,494,513]" pageId="12" pageNumber="13" value="2017-07-06">6 July 2017</collectingDate>
</date>
by
<collectorName id="277327D4634BFFCEFE81FE57FE24FDE0" box="[361,498,494,513]" pageId="12" pageNumber="13">Gunther Köhler</collectorName>
and
<collectorName id="277327D4634BFFCEFDC8FE57FD47FDE0" box="[544,657,494,513]" pageId="12" pageNumber="13">Ni Lar Than</collectorName>
. Field tag number GK-6728
</materialsCitation>
.
</paragraph>
<paragraph id="8A394202634BFFCEFF60FD97FE89FD70" blockId="12.[136,745,558,658]" pageId="12" pageNumber="13">
<materialsCitation id="3AEE485F634BFFCEFF60FD97FD78FDA0" box="[136,686,558,578]" collectingDate="2017-07-06" collectionCode="SMF" collectorName="Gunther Kohler &amp; Ni Lar Than" country="Myanmar" elevation="17" latitude="16.77737" location="East Yangon University" longLatPrecision="1" longitude="96.24065" municipality="Thanlyin" pageId="12" pageNumber="13" specimenCode="SMF 103840" specimenCount="1" stateProvince="Yangon" typeStatus="paratype">
<emphasis id="B8F29E10634BFFCEFF60FD97FF3FFDA0" bold="true" box="[136,233,558,577]" pageId="12" pageNumber="13">
<typeStatus id="553DFCA0634BFFCEFF60FD97FF33FDA0" box="[136,229,558,577]" pageId="12" pageNumber="13">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DA20EA79634BFFCEFF1DFD97FEBBFDA0" box="[245,365,558,577]" pageId="12" pageNumber="13">SMF 103840</specimenCode>
, same collecting data as holotype
</materialsCitation>
;
<materialsCitation id="3AEE485F634BFFCEFD56FD97FE8AFD70" collectingDate="2018-06-15" collectionCode="SMF" collectorName="Gunther Kohler &amp; Ni Lar Than" country="Myanmar" elevation="17" latitude="16.77737" location="East Yangon University" longLatPrecision="1" longitude="96.24065" municipality="Thanlyin" pageId="12" pageNumber="13" specimenCode="SMF 103797-99" specimenCount="1" stateProvince="Yangon" typeStatus="paratype">
<specimenCode id="DA20EA79634BFFCEFD56FD97FF3CFDBD" pageId="12" pageNumber="13">SMF 10379799</specimenCode>
, same locality data and collectors as holotype but collected
<date id="FE3864C2634BFFCEFF60FDDAFED7FD96" box="[136,257,611,631]" pageId="12" pageNumber="13" value="2018-06-15">
<collectingDate id="EE7C9D2A634BFFCEFF60FDDAFED7FD96" box="[136,257,611,631]" pageId="12" pageNumber="13" value="2018-06-15">15 June 2018</collectingDate>
</date>
. All paratypes are adult males except for SMF 103798 which is an adult female
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634BFFCEFF60FD0DFBBCFE8D" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8A394202634BFFCEFF60FD0DFF6DFB8D" blockId="12.[135,745,692,1132]" pageId="12" pageNumber="13">
<emphasis id="B8F29E10634BFFCEFF60FD0DFF28FD2D" bold="true" box="[136,254,692,716]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
A species of the genus
<taxonomicName id="4D863981634BFFCEFDCEFD0DFD13FD2D" authorityName="Peters" authorityYear="1867" box="[550,709,692,716]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B8F29E10634BFFCEFDCEFD0DFD13FD2D" box="[550,709,692,716]" italics="true" pageId="12" pageNumber="13">Phrynoglossus</emphasis>
</taxonomicName>
as defined above that differs from all mainland Southeast Asian congeners by having (1) a large body size (males 22.0
<quantity id="4D7EEFE7634BFFCEFF2DFCADFEFAFCCD" box="[197,300,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="12" pageNumber="13" unit="mm" value="27.4">27.4 mm</quantity>
; females
<quantity id="4D7EEFE7634BFFCEFE73FCADFD96FCCD" box="[411,576,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="2.985" metricValueMax="3.13" metricValueMin="2.84" pageId="12" pageNumber="13" unit="mm" value="29.85" valueMax="31.3" valueMin="28.4">28.431.3 mm</quantity>
); (2) relatively small tympanum (ratio
<collectionCode id="EC97DAC7634BFFCEFE7BFC8DFE1CFCAD" box="[403,458,820,844]" pageId="12" pageNumber="13">TYD</collectionCode>
/
<collectionCode id="EC97DAC7634BFFCEFE39FC8DFDD5FCAD" box="[465,515,820,844]" pageId="12" pageNumber="13">SVL</collectionCode>
0.480.92); and (3) call duration of male advertisement call 85114 ms.
<taxonomicName id="4D863981634BFFCEFF60FCCDFE7DFC6D" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" box="[136,427,884,908]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634BFFCEFF60FCCDFE7DFC6D" box="[136,427,884,908]" italics="true" pageId="12" pageNumber="13">Phrynoglossus myanhessei</emphasis>
</taxonomicName>
differs from its congeners in Indochina (i.e.,
<taxonomicName id="4D863981634BFFCEFEDDFC2CFE60FC4D" authorityName="Peters" authorityYear="1867" box="[309,438,917,940]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFEDDFC2CFE60FC4D" box="[309,438,917,940]" italics="true" pageId="12" pageNumber="13">P. martensii</emphasis>
</taxonomicName>
and
<taxonomicName id="4D863981634BFFCEFE06FC2CFD14FC4D" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[494,706,916,940]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFE06FC2CFD14FC4D" box="[494,706,916,940]" italics="true" pageId="12" pageNumber="13">P. magnapustulosus</emphasis>
</taxonomicName>
) in the male advertisement call, most obvious in call duration (85114 ms in
<taxonomicName id="4D863981634BFFCEFE89FC6CFE2FFC0D" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" box="[353,505,980,1004]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634BFFCEFE89FC6CFE2FFC0D" box="[353,505,980,1004]" italics="true" pageId="12" pageNumber="13">P. myanhessei</emphasis>
</taxonomicName>
, 3252 ms in
<taxonomicName id="4D863981634BFFCEFD72FC6CFF15FBED" authorityName="Peters" authorityYear="1867" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFD72FC6CFF15FBED" italics="true" pageId="12" pageNumber="13">P. martensii</emphasis>
</taxonomicName>
, 243437 ms in
<taxonomicName id="4D863981634BFFCEFE9AFC4CFD90FBED" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[370,582,1012,1036]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFE9AFC4CFD90FBED" box="[370,582,1012,1036]" italics="true" pageId="12" pageNumber="13">P. magnapustulosus</emphasis>
</taxonomicName>
) and dominant frequency (24542885 Hz in
<taxonomicName id="4D863981634BFFCEFE21FBACFDB7FBCD" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" box="[457,609,1044,1068]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634BFFCEFE21FBACFDB7FBCD" box="[457,609,1044,1068]" italics="true" pageId="12" pageNumber="13">P. myanhessei</emphasis>
</taxonomicName>
, 33383962 Hz in
<taxonomicName id="4D863981634BFFCEFF27FB8CFE87FBAD" authorityName="Peters" authorityYear="1867" box="[207,337,1077,1100]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFF27FB8CFE87FBAD" box="[207,337,1077,1100]" italics="true" pageId="12" pageNumber="13">P. martensii</emphasis>
</taxonomicName>
, 31873790 Hz in
<taxonomicName id="4D863981634BFFCEFDC2FB8CFF7CFB8D" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFDC2FB8CFF7CFB8D" italics="true" pageId="12" pageNumber="13">P. magnapustulosus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8A394202634BFFCEFF60FB2DFBBCFE8D" blockId="12.[136,745,1172,2125]" lastBlockId="12.[782,1391,212,364]" pageId="12" pageNumber="13">
<emphasis id="B8F29E10634BFFCEFF60FB2DFEFBFB4D" bold="true" box="[136,301,1172,1196]" pageId="12" pageNumber="13">Comparisons.</emphasis>
<taxonomicName id="4D863981634BFFCEFEDBFB2DFD8BFB4D" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" box="[307,605,1172,1196]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634BFFCEFEDBFB2DFD8BFB4D" box="[307,605,1172,1196]" italics="true" pageId="12" pageNumber="13">Phrynoglossus myanhessei</emphasis>
</taxonomicName>
differs from its congeners as follows (condition for
<taxonomicName id="4D863981634BFFCEFDA0FB0CFD3FFB2D" authorityName="Kohler, Vargas, Than &amp; Thammachoti" authorityYear="2021" box="[584,745,1204,1228]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634BFFCEFDA0FB0CFD8EFB2D" box="[584,600,1205,1228]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFD81FB0DFD3FFB2D" box="[617,745,1204,1228]" italics="true" pageId="12" pageNumber="13">myanhessei</emphasis>
</taxonomicName>
in parentheses).
<taxonomicName id="4D863981634BFFCEFEA3FB6DFDA1FB0D" baseAuthorityName="Mertens" baseAuthorityYear="1927" box="[331,631,1236,1260]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="floresianus">
<emphasis id="B8F29E10634BFFCEFEA3FB6DFDA1FB0D" box="[331,631,1236,1260]" italics="true" pageId="12" pageNumber="13">Phrynoglossus floresianus</emphasis>
</taxonomicName>
,
<taxonomicName id="4D863981634BFFCEFD63FB6CFF38FAED" baseAuthorityName="Smith" baseAuthorityYear="1927" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="semipalmatus">
<emphasis id="B8F29E10634BFFCEFD63FB6CFF38FAED" italics="true" pageId="12" pageNumber="13">P. semipalmatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4D863981634BFFCEFEC2FB4CFE62FAED" baseAuthorityName="Iskandar, Arifin, and Rachmanasah" baseAuthorityYear="2011" box="[298,436,1268,1292]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="tompotika">
<emphasis id="B8F29E10634BFFCEFEC2FB4CFE62FAED" box="[298,436,1268,1292]" italics="true" pageId="12" pageNumber="13">P. tompotika</emphasis>
</taxonomicName>
all have distinctly enlarged flattened toe and finger disks (tips of toes rounded, only slightly expanded into discs, not distinctly flattened);
<taxonomicName id="4D863981634BFFCEFF0BFAECFEA9FA8D" baseAuthorityName="Mertens" baseAuthorityYear="1927" box="[227,383,1364,1388]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="floresianus">
<emphasis id="B8F29E10634BFFCEFF0BFAECFEA9FA8D" box="[227,383,1364,1388]" italics="true" pageId="12" pageNumber="13">P. floresianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4D863981634BFFCEFE28FAECFDF4FA8D" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[448,546,1364,1388]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B8F29E10634BFFCEFE28FAECFDF4FA8D" box="[448,546,1364,1388]" italics="true" pageId="12" pageNumber="13">P. laevis</emphasis>
</taxonomicName>
are much larger frogs,
<quantity id="4D7EEFE7634BFFCEFF38FACDFE90FA6D" box="[208,326,1396,1420]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="12" pageNumber="13" unit="mm" value="36.0" valueMax="37.0" valueMin="35.0">3537 mm</quantity>
in males,
<quantity id="4D7EEFE7634BFFCEFE51FACDFDF9FA6D" box="[441,559,1396,1420]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.1" metricValueMin="4.0" pageId="12" pageNumber="13" unit="mm" value="45.5" valueMax="51.0" valueMin="40.0">4051 mm</quantity>
in adult females in
<taxonomicName id="4D863981634BFFCEFF40FA2CFE95FA4D" baseAuthorityName="Mertens" baseAuthorityYear="1927" box="[168,323,1428,1452]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="floresianus">
<emphasis id="B8F29E10634BFFCEFF40FA2CFE95FA4D" box="[168,323,1428,1452]" italics="true" pageId="12" pageNumber="13">P. floresianus</emphasis>
</taxonomicName>
,
<quantity id="4D7EEFE7634BFFCEFEBDFA2DFE06FA4D" box="[341,464,1428,1452]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="4.2" metricValueMin="2.6" pageId="12" pageNumber="13" unit="mm" value="34.0" valueMax="42.0" valueMin="26.0">2642 mm</quantity>
in males,
<quantity id="4D7EEFE7634BFFCEFDA6FA2DFD1EFA4D" box="[590,712,1428,1452]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="6.2" metricValueMin="3.5" pageId="12" pageNumber="13" unit="mm" value="48.5" valueMax="62.0" valueMin="35.0">3562 mm</quantity>
in females in
<taxonomicName id="4D863981634BFFCEFEE7FA0CFEA4FA2D" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[271,370,1460,1484]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B8F29E10634BFFCEFEE7FA0CFEA4FA2D" box="[271,370,1460,1484]" italics="true" pageId="12" pageNumber="13">P. laevis</emphasis>
</taxonomicName>
(males 22.0
<quantity id="4D7EEFE7634BFFCEFDFDFA0DFD57FA2D" box="[533,641,1460,1484]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="12" pageNumber="13" unit="mm" value="27.4">27.4 mm</quantity>
; females
<quantity id="4D7EEFE7634BFFCEFF61FA6DFEE0FA0D" box="[137,310,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="2.985" metricValueMax="3.13" metricValueMin="2.84" pageId="12" pageNumber="13" unit="mm" value="29.85" valueMax="31.3" valueMin="28.4">28.431.3 mm</quantity>
); furthermore, the eyes are directed dorsolaterally in
<taxonomicName id="4D863981634BFFCEFEB8FA4CFE67F9ED" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[336,433,1524,1548]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B8F29E10634BFFCEFEB8FA4CFE67F9ED" box="[336,433,1524,1548]" italics="true" pageId="12" pageNumber="13">P. laevis</emphasis>
</taxonomicName>
and
<taxonomicName id="4D863981634BFFCEFE19FA4CFD52F9ED" baseAuthorityName="Smith" baseAuthorityYear="1927" box="[497,644,1524,1548]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="celebensis">
<emphasis id="B8F29E10634BFFCEFE19FA4CFD52F9ED" box="[497,644,1524,1548]" italics="true" pageId="12" pageNumber="13">P. celebensis</emphasis>
</taxonomicName>
(laterally);
<taxonomicName id="4D863981634BFFCEFF5EF9ACFEE9F9CD" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[182,319,1556,1580]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B8F29E10634BFFCEFF5EF9ACFF10F9CD" box="[182,198,1557,1580]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFF3EF9ADFEE9F9CD" box="[214,319,1556,1580]" italics="true" pageId="12" pageNumber="13">baluensis</emphasis>
</taxonomicName>
usually has a large inverted U-shaped ridge on the dorsum (no such ridge present) and reduced toe webbing with at least two phalanges free of webbing along fourth toe (feet almost fully webbed, less than one phalange free of webbing along fourth toe);
<taxonomicName id="4D863981634BFFCEFF2FF90CFEBBF92D" baseAuthorityName="Peters" baseAuthorityYear="1877" box="[199,365,1717,1740]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="sumatranus">
<emphasis id="B8F29E10634BFFCEFF2FF90CFF01F92D" box="[199,215,1717,1740]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFF03F90CFEBBF92D" box="[235,365,1717,1740]" italics="true" pageId="12" pageNumber="13">sumatranus</emphasis>
</taxonomicName>
has a dark brown band on each side of the cloaca (such band absent), diamond shaped pupil (ovoid), and its eyes are oriented dorsolaterally (laterally);
<taxonomicName id="4D863981634BFFCEFECEF8ACFE1EF8CD" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[294,456,1812,1836]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="diminutivus">
<emphasis id="B8F29E10634BFFCEFECEF8ACFEE0F8CD" box="[294,310,1813,1836]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFEAEF8ADFE1EF8CD" box="[326,456,1812,1836]" italics="true" pageId="12" pageNumber="13">diminutivus</emphasis>
</taxonomicName>
has reduced toe webbing with two phalanges free of webbing along fourth toe (feet almost fully webbed, less than one phalange free of webbing along fourth toe);
<taxonomicName id="4D863981634BFFCEFE3DF8CCFD8BF86D" authorityName="Peters" authorityYear="1867" box="[469,605,1909,1932]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFE3DF8CCFE33F86D" box="[469,485,1909,1932]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFE1CF8CCFD8BF86D" box="[500,605,1909,1932]" italics="true" pageId="12" pageNumber="13">martensii</emphasis>
</taxonomicName>
and
<taxonomicName id="4D863981634BFFCEFD79F8CCFECCF84D" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFD79F8CCFD77F86D" box="[657,673,1909,1932]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFD59F8CCFECCF84D" italics="true" pageId="12" pageNumber="13">magnapustulosus</emphasis>
</taxonomicName>
are smaller frogs,
<quantity id="4D7EEFE7634BFFCEFE13F82DFDA3F84D" box="[507,629,1940,1964]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="12" pageNumber="13" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">2023 mm</quantity>
in males,
<quantity id="4D7EEFE7634BFFCEFF61F80DFF29F82D" box="[137,255,1972,1996]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="12" pageNumber="13" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">2630 mm</quantity>
in adult females in
<taxonomicName id="4D863981634BFFCEFE34F80CFDB2F82D" authorityName="Peters" authorityYear="1867" box="[476,612,1973,1996]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFE34F80CFE3AF82D" box="[476,492,1973,1996]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFE13F80CFDB2F82D" box="[507,612,1973,1996]" italics="true" pageId="12" pageNumber="13">martensii</emphasis>
</taxonomicName>
,
<quantity id="4D7EEFE7634BFFCEFD9AF80DFD31F82D" box="[626,743,1972,1996]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="12" pageNumber="13" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 mm</quantity>
in males,
<quantity id="4D7EEFE7634BFFCEFF1CF86DFEBCF80D" box="[244,362,2004,2028]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="12" pageNumber="13" unit="mm" value="20.5" valueMax="24.0" valueMin="17.0">1724 mm</quantity>
in females in
<taxonomicName id="4D863981634BFFCEFDE1F86CFD3FF80D" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[521,745,2004,2028]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFDE1F86CFDCFF80D" box="[521,537,2005,2028]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFDC0F86DFD3FF80D" box="[552,745,2004,2028]" italics="true" pageId="12" pageNumber="13">magnapustulosus</emphasis>
</taxonomicName>
(males 22.0
<quantity id="4D7EEFE7634BFFCEFEF5F84DFE5EF7ED" box="[285,392,2036,2060]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="12" pageNumber="13" unit="mm" value="27.4">27.4 mm</quantity>
; females
<quantity id="4D7EEFE7634BFFCEFE14F84DFD70F7ED" box="[508,678,2036,2060]" metricMagnitude="-2" metricUnit="m" metricValue="2.985" metricValueMax="3.13" metricValueMin="2.84" pageId="12" pageNumber="13" unit="mm" value="29.85" valueMax="31.3" valueMin="28.4">28.431.3 mm</quantity>
); furthermore,
<taxonomicName id="4D863981634BFFCEFF16F7ACFE34F7CD" baseAuthorityName="Taylor and Elbel" baseAuthorityYear="1958" box="[254,482,2068,2092]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="magnapustulosus">
<emphasis id="B8F29E10634BFFCEFF16F7ACFED8F7CD" box="[254,270,2069,2092]" italics="true" pageId="12" pageNumber="13">P</emphasis>
.
<emphasis id="B8F29E10634BFFCEFEC9F7ADFE34F7CD" box="[289,482,2068,2092]" italics="true" pageId="12" pageNumber="13">magnapustulosus</emphasis>
</taxonomicName>
has a relatively larger tympanum, ratio tympanum/SVL
<geoCoordinate id="EFB224C5634BFFCEFDE7F78DFD99F7AD" box="[527,591,2100,2124]" degrees="0.055" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.055">0.055</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFDA6F78DFD4BF7AD" box="[590,669,2100,2124]" degrees="0.099" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.099">0.099</geoCoordinate>
, mean 0.082 (
<geoCoordinate id="EFB224C5634BFFCEFC88FF6DFC77FF0D" box="[864,929,212,236]" degrees="0.029" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.029">0.029</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFC49FF6DFC27FF0D" box="[929,1009,212,236]" degrees="0.066" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.066">0.066</geoCoordinate>
, mean 0.050). Also,
<taxonomicName id="4D863981634BFFCEFB0FFF6CFAB9FF0D" authorityName="Peters" authorityYear="1867" box="[1255,1391,213,236]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="martensii">
<emphasis id="B8F29E10634BFFCEFB0FFF6CFAB9FF0D" box="[1255,1391,213,236]" italics="true" pageId="12" pageNumber="13">P. martensii</emphasis>
</taxonomicName>
has a relatively broader head, ratio HW/SVL
<geoCoordinate id="EFB224C5634BFFCEFACBFF4DFAB7FEED" box="[1315,1377,244,268]" degrees="0.336" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.336">0.336</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFA89FF4DFC99FECD" degrees="0.401" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.401"> 0.401</geoCoordinate>
, mean
<quantity id="4D7EEFE7634BFFCEFC77FEADFC2DFECD" box="[927,1019,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="9.067799999999998" pageId="12" pageNumber="13" unit="in" value="0.357">0.357 in</quantity>
males,
<geoCoordinate id="EFB224C5634BFFCEFBBBFEADFB45FECD" box="[1107,1171,276,300]" degrees="0.338" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.338">0.338</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFB7BFEADFB37FECD" box="[1171,1249,276,300]" degrees="0.388" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.388">0.388</geoCoordinate>
, mean
<quantity id="4D7EEFE7634BFFCEFAD9FEADFCF2FEAD" metricMagnitude="-3" metricUnit="m" metricValue="8.9154" pageId="12" pageNumber="13" unit="in" value="0.351">0.351 in</quantity>
females (
<geoCoordinate id="EFB224C5634BFFCEFC7AFE8DFC05FEAD" box="[914,979,308,332]" degrees="0.286" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.286">0.286</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFC3AFE8DFBF4FEAD" box="[978,1058,308,332]" degrees="0.369" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.369">0.369</geoCoordinate>
, mean
<quantity id="4D7EEFE7634BFFCEFB99FE8DFB1AFEAD" box="[1137,1228,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="8.5852" pageId="12" pageNumber="13" unit="in" value="0.338">0.338 in</quantity>
males,
<geoCoordinate id="EFB224C5634BFFCEFACBFE8DFAB7FEAD" box="[1315,1377,308,332]" degrees="0.283" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="0.283">0.283</geoCoordinate>
<geoCoordinate id="EFB224C5634BFFCEFA89FE8DFC99FE8D" degrees="0.354" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="-0.354"> 0.354</geoCoordinate>
, mean
<quantity id="4D7EEFE7634BFFCEFC77FEEDFC2CFE8D" box="[927,1018,340,364]" metricMagnitude="-3" metricUnit="m" metricValue="7.924799999999999" pageId="12" pageNumber="13" unit="in" value="0.312">0.312 in</quantity>
females).
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634BFFCFFCE6FE2DFEF5F86D" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8A394202634BFFCEFCE6FE2DFAF1FA0D" blockId="12.[782,1391,404,2124]" pageId="12" pageNumber="13">
<emphasis id="B8F29E10634BFFCEFCE6FE2DFB94FE4D" bold="true" box="[782,1090,404,428]" pageId="12" pageNumber="13">
Description of the
<typeStatus id="553DFCA0634BFFCEFC08FE2DFB94FE4D" box="[992,1090,404,428]" pageId="12" pageNumber="13">holotype</typeStatus>
</emphasis>
(
<figureCitation id="12BD5E87634BFFCEFBB9FE2DFB4CFE4D" box="[1105,1178,404,428]" captionStart="Figure 8" captionStartId="13.[196,264,1494,1516]" captionTargetBox="[196,1451,198,1474]" captionTargetId="figure-238@13.[196,1451,198,1474]" captionTargetPageId="13" captionText="Figure 8. Holotype of Phrynoglossus myanhessei n. sp. (SMF 103841). Scale bars equal 10.0 mm (A, B) and 5 mm (CF). Photos by G.K." pageId="12" pageNumber="13">Figs. 8</figureCitation>
and
<figureCitation id="12BD5E87634BFFCEFB27FE2DFB09FE4D" box="[1231,1247,404,428]" captionStart="Figure 9" captionStartId="14.[136,204,1494,1516]" captionTargetBox="[136,1391,198,1474]" captionTargetId="figure-245@14.[136,1391,198,1474]" captionTargetPageId="14" captionText="Figure 9. Holotype of Phrynoglossus myanhessei sp. n. (SMF 103841). Scale bars equal 5.0 mm. Photos by G.K." pageId="12" pageNumber="13">9</figureCitation>
). Adult male, as indicated by dark colored throat region and presence of vocal slits; SVL
<quantity id="4D7EEFE7634BFFCEFC31FE6DFB91FE0D" box="[985,1095,468,492]" metricMagnitude="-2" metricUnit="m" metricValue="2.356" pageId="12" pageNumber="13" unit="mm" value="23.56">23.56 mm</quantity>
; habitus robust; head broad, about as wide as long, ratio HL/HW 1.06; snout nearly rounded in dorsal view, projecting beyond lower jaw, rounded in profile; nostril dorsolateral, closer to tip of snout than eye; canthus rounded; ratio EYD/SVL 0.12; IOD (2.12) greater than width of upper eyelid (1.94); tympanum concealed, slightly depressed relative to skin of temporal region, tympanic rim weakly elevated relative to tympanum; ratio TYD/EYD 0.42; vomerine teeth absent; tongue fleshy, rounded, without notch; tips of all four fingers rounded, not expanded into discs; relative finger lengths III&gt;I&gt;IV&gt;II; no webbing; distinct subarticular tubercles, palmar tubercle distinct, bifid; thenar tubercle large, about twice the size of palmar tubercle; tips of toes rounded, slightly expanded into discs; relative toe lengths IV&gt;III&gt;V&gt;II&gt;I; feet almost fully webbed, webbing formula I 0.80.8 II 0.80.8 III 0.80.9 IV 0.90.8 V; a well-developed flap of skin on postaxial side of Toe V from level of outer metatarsal tubercle to distal subarticular tubercle; strong fold on distal one-half of tarsus; large, flap-like inner metatarsal tubercle; outer metatarsal tubercle not differentiated, but rather two tiny tubercles present in that area; skin on top of head and on dorsum and flank smooth; skin on throat and venter shagreen; skin on upper surface of forelimbs and thigh with low, rounded tubercles, that of dorsal surface of shank with scattered keratinized pointed tubercles; indistinct, glandular supratympanic fold from posterior edge of upper eyelid along upper margin of tympanum and then obliquely down to shoulder; no dorsolateral fold. Measurements (mm) of
<typeStatus id="553DFCA0634BFFCEFCE6FA2DFCA6FA4D" box="[782,880,1428,1452]" pageId="12" pageNumber="13">holotype</typeStatus>
: SVL 23.56; HL 8.91; HW 8.44; SL 3.60; EYD 2.81; IOD 2.12; TYD 1.17; TED 0.67; SHL 12.51; THL 12.52; HNL 6.04; FL 12.22; NED 2.37; IND 1.92.
</paragraph>
<paragraph id="8A394202634BFFCEFCC5FA4DFBC8F84D" blockId="12.[782,1391,404,2124]" pageId="12" pageNumber="13">
Coloration in life was recorded as follows (
<figureCitation id="12BD5E87634BFFCEFAFAFA4DFAB4F9ED" box="[1298,1378,1524,1548]" captionStart="Figure 10" captionStartId="15.[196,264,1046,1068]" captionTargetBox="[196,1451,198,1026]" captionTargetId="figure-566@15.[196,1451,198,1026]" captionTargetPageId="15" captionText="Figure 10. Phrynoglossus myanhessei sp. n. in life. (A) Male holotype, SMF 103841; (B) male paratype, SMF 103840; (C) male paratype, SMF 103799; (D) female paratype, SMF 103798. Photos by G.K." pageId="12" pageNumber="13">Fig. 10</figureCitation>
): Dorsal and lateral ground color of head and body Drab-Gray (256) with indistinct Raw Umber (280) blotches and mottling; dorsal surfaces of forelimbs True Cinnamon (260) with Raw Umber (280) speckles; dorsal surfaces of hind limbs Drab-Gray (256) with Raw Umber (280) transverse broken bars; a Drab-Gray (256) oblique bar from anterior corner of eye to snout; throat region heavily suffused with Vandyke Brown (282); venter Pale Neutral Gray (296) with Smoky White (261) stipples; ventral surfaces of forelimbs Medium Fawn Color (257); ventral surfaces of hind limbs Cream White (52); iris Olive Brown (278) with a suffusion of Orange-Rufous (56) above and whitish below.
</paragraph>
<paragraph id="8A394202634BFFCFFCC5F80DFD72F8ED" blockId="12.[782,1391,404,2124]" lastBlockId="13.[197,805,1620,1804]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">Coloration after about three years preservation in 70% ethanol was recorded as follows: Dorsal and lateral ground color of head and body Glaucous (272) with indistinct Sepia (279) blotches and mottling; dorsal surfaces of hind limbs Glaucous (272) with Sepia (279) transverse broken bars; a Drab-Gray (256) oblique bar from anterior corner of eye to snout; supraocular region Dusky Brown (285); throat region Glaucous (289) with Pale Buff (1) stipples; venter Pale Buff (1) with Smoky White (261) stipples; ventral surfaces of forelimbs Glaucous (289); ventral surfaces of hind limbs Light Buff (2).</paragraph>
<caption id="DEF9128A634AFFCFFF2CFA6FFEC4F9EA" pageId="13" pageNumber="14" startId="13.[196,264,1494,1516]" targetBox="[196,1451,198,1474]" targetPageId="13" targetType="figure">
<paragraph id="8A394202634AFFCFFF2CFA6FFEC4F9EA" blockId="13.[196,1451,1494,1548]" pageId="13" pageNumber="14">
<emphasis id="B8F29E10634AFFCFFF2CFA6FFEF6FA0A" bold="true" box="[196,288,1494,1516]" pageId="13" pageNumber="14">Figure 8.</emphasis>
Holotype of
<taxonomicName id="4D863981634AFFCFFE77FA6FFD74FA0A" authority="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti, 2021" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[415,674,1494,1515]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="myanhessei" status="sp. nov.">
<emphasis id="B8F29E10634AFFCFFE77FA6FFD74FA0A" box="[415,674,1494,1515]" italics="true" pageId="13" pageNumber="14">Phrynoglossus myanhessei</emphasis>
</taxonomicName>
<emphasis id="B8F29E10634AFFCFFD4FFA6FFD08FA0A" bold="true" box="[679,734,1494,1515]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A3C1236B634AFFCFFD4FFA6FFD08FA0A" box="[679,734,1494,1515]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(SMF 103841). Scale bars equal 10.0 mm (A, B) and 5 mm (CF). Photos by G.K.
</paragraph>
</caption>
<paragraph id="8A394202634AFFCFFF2DF88DFEF5F86D" blockId="13.[197,805,1844,1932]" pageId="13" pageNumber="14">
<emphasis id="B8F29E10634AFFCFFF2DF88DFEE1F8AD" bold="true" box="[197,311,1844,1868]" pageId="13" pageNumber="14">Variation.</emphasis>
The
<typeStatus id="553DFCA0634AFFCFFE85F88CFE05F8AC" box="[365,467,1845,1869]" pageId="13" pageNumber="14">paratypes</typeStatus>
agree well with the
<typeStatus id="553DFCA0634AFFCFFD44F88DFCDFF8AD" box="[684,777,1844,1868]" pageId="13" pageNumber="14">holotype</typeStatus>
in general appearance; morphometrics and coloration (see
<tableCitation id="C70477B9634AFFCFFF2DF8CDFEC3F86D" box="[197,277,1908,1932]" captionStart="Table 3" captionStartId="7.[197,253,1462,1483]" captionTargetId="graphics-635@7.[197,1450,1540,2116]" captionTargetPageId="7" captionText="Table 3. Selected measurements and proportions of the species of Phrynoglossus. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." pageId="13" pageNumber="14">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634AFFCFFF2DF80DFC3BF94D" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8A394202634AFFCFFF2DF80DFC3BF94D" blockId="13.[197,806,1972,2124]" lastBlockId="13.[843,1451,1620,1708]" pageId="13" pageNumber="14">
<emphasis id="B8F29E10634AFFCFFF2DF80DFE93F82D" bold="true" box="[197,325,1972,1996]" pageId="13" pageNumber="14">Etymology.</emphasis>
“Myan” is Myanmars abbreviated name and was chosen because this species is endemic to Myanmar as far as we know. “hessei” was chosen in recognition of the long-term support and funding of Senckenberg by the German State of Hesse. In combination, the species name
<taxonomicName id="4D863981634AFFCFFCA3F9EDFC10F98D" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[843,966,1620,1644]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634AFFCFFCA3F9EDFC10F98D" box="[843,966,1620,1644]" italics="true" pageId="13" pageNumber="14">myanhessei</emphasis>
</taxonomicName>
reflects the long-term productive collaboration of researchers from Hesse and Myanmar in the field of herpetology.
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634AFFCFFCA3F96DFBF0F88D" pageId="13" pageNumber="14" type="biology_ecology">
<paragraph id="8A394202634AFFCFFCA3F96DFBF0F88D" blockId="13.[843,1452,1748,1901]" pageId="13" pageNumber="14">
<emphasis id="B8F29E10634AFFCFFCA3F96DFB9AF90D" bold="true" box="[843,1100,1748,1772]" pageId="13" pageNumber="14">Natural history notes.</emphasis>
At the
<typeStatus id="553DFCA0634AFFCFFB4FF96CFB03F90C" box="[1191,1237,1749,1773]" pageId="13" pageNumber="14">type</typeStatus>
locality, the specimens were collected at night in a patch of muddy grass area, partly open, partly covered by bushes and low trees. The frogs were sitting at the edge of small shallow temporary water bodies.
</paragraph>
</subSubSection>
<subSubSection id="C29C1189634AFFCFFCA3F82DFC46F7AD" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8A394202634AFFCFFCA3F82DFC46F7AD" blockId="13.[843,1452,1940,2124]" pageId="13" pageNumber="14">
<emphasis id="B8F29E10634AFFCFFCA3F82DFA9AF84D" bold="true" box="[843,1356,1940,1964]" pageId="13" pageNumber="14">Geographic Distribution and Conservation.</emphasis>
As currently known,
<taxonomicName id="4D863981634AFFCFFC02F80DFADDF82D" authorityName="Köhler &amp; Vargas &amp; Than &amp; Schell &amp; Janke &amp; Pauls &amp; Thammachoti" authorityYear="2021" box="[1002,1291,1972,1996]" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="myanhessei">
<emphasis id="B8F29E10634AFFCFFC02F80DFADDF82D" box="[1002,1291,1972,1996]" italics="true" pageId="13" pageNumber="14">Phrynoglossus myanhessei</emphasis>
</taxonomicName>
is restricted to central and lower Myanmar (
<figureCitation id="12BD5E87634AFFCFFB65F86DFB08F80D" box="[1165,1246,2004,2028]" captionStart="Figure 11" captionStartId="16.[136,204,1110,1132]" captionTargetBox="[136,1389,200,1090]" captionTargetId="figure-10@16.[136,1391,198,1091]" captionTargetPageId="16" captionText="Figure 11. Map indicating collecting localities of the Phrynoglossus species occurring in Indochina. Each symbol can represent one or more adjacent localities. Black squares: P. myanhessei n. sp.; black circles: P. martensii; green pentagons: P.cf. martensii; black triangles: P. magnapustulosus; white circles: additional specimens of Phrynoglossus from the FMNH, not examined by authors." pageId="13" pageNumber="14">Fig. 11</figureCitation>
). This species was abundant wherever we found it. Thus, we classify it as Least Concern according to the IUCN categories (
<bibRefCitation id="EE173FF3634AFFCFFA8AF7ADFC53F7AD" author="IUCN" pageId="13" pageNumber="14" refId="ref12331" refString="IUCN (2012) IUCN Red List Categories and Criteria, Version 3. 1. Second edition. https: // portals. ucn. org / library / node / 10315" type="url" year="2012">
<collectionCode id="EC97DAC7634AFFCFFA8AF7ADFA71F7CD" box="[1378,1447,2068,2092]" pageId="13" pageNumber="14">IUCN</collectionCode>
, 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>