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<mods:titleInfo>
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<mods:title>Annelids of the eastern Australian abyss collected by the 2017 RV ' Investigator' voyage</mods:title>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>1020</mods:number>
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<mods:start>1</mods:start>
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<treatment ID-GBIF-Taxon="179910784" LSID="urn:lsid:plazi:treatment:DDF05FF569307E04212367A45C10AF8F" httpUri="http://treatment.plazi.org/id/DDF05FF569307E04212367A45C10AF8F" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName authority="Malm, 1874" authorityName="Malm" authorityYear="1874" class="Polychaeta" family="Nephtyidae" genus="Nephtys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephtys paradoxa" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="paradoxa">Nephtys cf. paradoxa Malm, 1874</taxonomicName>
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Acoetidae, Chrysopetalidae, Nephtyidae, Oweniidae A Acoetidae, Panthalis sp., dorsal view (AM W. 50321) B Chrysopetalidae, Dysponetus cf. caecus, dorsal view C Dysponetus cf. caecus, ventral view D Nephtyidae, Aglaophamus sp. pre- and post-acicular chaetae E Nephtyidae, Nephtys cf. paradoxa (AM W. 52750) F Nephtyidae, Aglaophamus sp. G Oweniidae, Myriowenia sp. in tube (AM W. 51842). Scale bars: 2 mm (A); 1 mm (B, C); 50 µm (D); 5 mm (E, G); 3 mm (F)." figureDoi="10.3897/zookeys.1020.57921.figure2" httpUri="https://binary.pensoft.net/fig/513270" pageId="0" pageNumber="1">Fig. 2E</figureCitation>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
Body stout, wider anteriorly and tapering from middle chaetigers to posterior. Prostomium subrectangular, eyes not visible, pharynx when everted with ten pairs of terminal bifid papillae, and 22 rows of subterminal papillae (rows with 4-6 similar conical papillae) extending only one third of pharyngeal length, median dorsal and ventral papillae not elongated, proximal region smooth. Antennae and palps conical, short, nuchal organs conspicuous, rounded. Parapodia biramous with well-separated rami. Interramal branchiae present, somewhat recurved, from chaetiger 11, becoming foliaceous from ~ chaetiger 16 and appearing membranous from chaetiger 14-16, ciliated and fully developed with membranous expansions (or 'foliaceous
|
||||
<normalizedToken originalValue="lamellae’">lamellae'</normalizedToken>
|
||||
) ~ chaetiger 20, becoming small and rudimentary from ~ chaetiger 38 to posterior chaetigers. Acicular lobes obliquely rounded, notopodia with rudimentary preacicular and low postacicular lobes; neuropodia with rudimentary preacicular lobe, postacicular lamella longer than acicular lobe. Chaetae short and
|
||||
<normalizedToken originalValue="‘spiky’">'spiky'</normalizedToken>
|
||||
, of three kinds: barred chaetae in preacicular position, spinulose chaetae in postacicular position and capillary chaetae present in neuropodia of chaetiger 1.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
This species has previously been recorded from off the east Australian coast by
|
||||
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.29.1974.226" author="Paxton, H" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="197 - 208" refId="B331" refString="Paxton, H, 1974. Contribution to the study of the Australian Nephtyidae (Polychaeta). Records of the Australian Museum 29: 197 - 208, DOI: https://doi.org/10.3853/j.0067-1975.29.1974.226" title="Contribution to the study of the Australian Nephtyidae (Polychaeta)." url="https://doi.org/10.3853/j.0067-1975.29.1974.226" volume="29" year="1974">Paxton (1974)</bibRefCitation>
|
||||
, collected in 1912 from 860 m, and has also been collected from 200 m northeast of Coffs Harbour, NSW in 1993 (unpublished AM records). However, it is a deep-water species widely distributed in the Arctic, the northern Atlantic Ocean, the Mediterranean Sea and the Pacific Ocean (
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2682.1.1" author="Ravara, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 68" refId="B360" refString="Ravara, A, Cunha, MR, Pleijel, F, 2010a. Nephtyidae (Annelida, Polychaeta) from southern Europe. Zootaxa 2682: 1 - 68, DOI: https://doi.org/10.11646/zootaxa.2682.1.1" title="Nephtyidae (Annelida, Polychaeta) from southern Europe." url="https://doi.org/10.11646/zootaxa.2682.1.1" volume="2682" year="2010 a">Ravara et al. 2010a</bibRefCitation>
|
||||
), originally described from the North Sea. It is surmised that it may be a species complex and that examination of global material as well as molecular analyses can only resolve its taxonomic status (
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.2010.00424.x" author="Ravara, A" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="1" pagination="394 - 405" refId="B361" refString="Ravara, A, Wiklund, H, Cunha, MR, Pleijel, F, 2010b. Phylogenetic relationships within Nephtyidae (Polychaeta: Annelida). Zoologica Scripta 39: 394 - 405, DOI: https://doi.org/10.1111/j.1463-6409.2010.00424.x" title="Phylogenetic relationships within Nephtyidae (Polychaeta: Annelida)." url="https://doi.org/10.1111/j.1463-6409.2010.00424.x" volume="39" year="2010 b">Ravara et al. 2010b</bibRefCitation>
|
||||
). Therefore, this is a tentative identification.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="records">
|
||||
<paragraph pageId="0" pageNumber="1">Records.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">2 specimens Suppl. material 1: ops. 80, 101 (AM).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
303
data/DD/F0/8F/DDF08FDD855057AA91172BE92C1C8119.xml
Normal file
303
data/DD/F0/8F/DDF08FDD855057AA91172BE92C1C8119.xml
Normal file
|
|
@ -0,0 +1,303 @@
|
|||
<document ID-DOI="http://dx.doi.org/10.3897/alpento.4.50703" ID-GBIF-Dataset="a9e0336c-85fe-4d7c-8998-fadb8b1e4a29" ID-GBIF-Taxon="163737593" ID-Pensoft-Pub="2535-0889-4-39" ID-Pensoft-UUID="73E3CBB8841D53E98BB78BA05DDC69C9" ID-ZooBank="D9502F1AAEC04B0F845C87D86632AF17" LSID="urn:lsid:zoobank.org:act:CE0385B3-C608-48F3-9DEA-67824816B44E" ModsDocID="2535-0889-4-39" checkinTime="1587713590471" checkinUser="pensoft" docAuthor="Huemer, Peter" docDate="2020" docId="DDF08FDD855057AA91172BE92C1C8119" docLanguage="en" docName="AlpEntomol 4: 39-63" docOrigin="Alpine Entomology 4" docSource="http://dx.doi.org/10.3897/alpento.4.50703" docTitle="Caryocolum lamai Huemer 2020, sp. nov." docType="treatment" docUuid="CE0385B3-C608-48F3-9DEA-67824816B44E" docUuidSource="ZooBank" docVersion="4" id="73E3CBB8841D53E98BB78BA05DDC69C9" lastPageNumber="39" masterDocId="73E3CBB8841D53E98BB78BA05DDC69C9" masterDocTitle="Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)" masterLastPageNumber="63" masterPageNumber="39" pageId="0" pageNumber="39" updateTime="1643439336999" updateUser="ExternalLinkService">
|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Huemer, Peter</mods:namePart>
|
||||
<mods:affiliation>Tiroler Landesmuseen Betriebgsges. m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria</mods:affiliation>
|
||||
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-0630-545X</mods:nameIdentifier>
|
||||
<mods:nameIdentifier type="email">p.huemer@tiroler-landesmuseen.at</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Alpine Entomology</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2020</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>4</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>39</mods:start>
|
||||
<mods:end>63</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/alpento.4.50703</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.4.50703</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">2535-0889-4-39</mods:identifier>
|
||||
<mods:identifier type="ZooBank">D9502F1AAEC04B0F845C87D86632AF17</mods:identifier>
|
||||
<mods:identifier type="Pensoft-UUID">73E3CBB8841D53E98BB78BA05DDC69C9</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-GBIF-Taxon="163737593" LSID="urn:lsid:zoobank.org:act:CE0385B3-C608-48F3-9DEA-67824816B44E" httpUri="http://treatment.plazi.org/id/DDF08FDD855057AA91172BE92C1C8119" lastPageNumber="39" pageId="0" pageNumber="39">
|
||||
<subSubSection pageId="0" pageNumber="39" type="nomenclature">
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<taxonomicName LSID="DDF08FDD-8550-57AA-9117-2BE92C1C8119" authority="Huemer, 2020" authorityName="Huemer" authorityYear="2020" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum lamai" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="lamai" status="sp. nov.">Caryocolum lamai</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="39">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<materialsCitation ID-GBIF-Occurrence="2609799208" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="TLMF" collectorName="Huemer & P. Huemer" country="Italy" elevation="0" latitude="45.06056" location="PN del Gran Bosco di / Salbertrand, 2 km SE Colle / dell'Assieta" longitude="6.97889" municipality="Torino" specimenCode="TLMF 2019-026, GEL 1283" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
</emphasis>
|
||||
<specimenCount type="male">♂</specimenCount>
|
||||
, "
|
||||
<collectingCountry>ITALIA</collectingCountry>
|
||||
sept., prov.
|
||||
<collectingMunicipality>Torino</collectingMunicipality>
|
||||
/
|
||||
<location LSID="urn:lsid:plazi:treatment:DDF08FDD855057AA91172BE92C1C8119:D7FC612A6669993E22A0CF2C040C1207" country="Italy" municipality="Torino" name="PN del Gran Bosco di / Salbertrand, 2 km SE Colle / dell'Assieta">
|
||||
PN del Gran Bosco di / Salbertrand, 2 km SE Colle /
|
||||
<normalizedToken originalValue="dell´Assieta">dell'Assieta</normalizedToken>
|
||||
</location>
|
||||
,
|
||||
<elevation>2240 m</elevation>
|
||||
/
|
||||
<geoCoordinate>6°58'44"E</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate>45°3'38"N</geoCoordinate>
|
||||
/
|
||||
<collectingDate>25.7.2019</collectingDate>
|
||||
, leg.
|
||||
<collectorName>Huemer</collectorName>
|
||||
/
|
||||
<specimenCode>TLMF 2019-026</specimenCode>
|
||||
" "DNA Barcode / TLMF Lep 27645" "
|
||||
<collectorName>P. Huemer</collectorName>
|
||||
/
|
||||
<specimenCode>GEL 1283</specimenCode>
|
||||
♂" (
|
||||
<collectionCode>TLMF</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">Paratypes</emphasis>
|
||||
: Italy: 7 ♂♂, same data as holotype, but DNA Barcodes 27646, 27647 (TLMF); 21 ♂♂, 1 ♀, Cuneo province, Val Traversagn, Casteldelfino W, 2050,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="34" orientation="latitude" precision="15" seconds="00" value="44.566666">44°34'00"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="6" direction="east" minutes="58" orientation="longitude" precision="15" seconds="30" value="6.975">6°58'30"E</geoCoordinate>
|
||||
, 21 Jul. 2001, leg. Huemer, gen. slides GEL 1258 ♂, GEL 1262 ♂, GEL 1286 ♀ (TLMF); 12 ♂♂, same data, but leg. Mayr (RCTM); 1 ♂, same data, but S. Anna, 21-22 Jul. 2001, leg. Huemer (TLMF); 9 ♂♂, Cuneo province, Demonte NW, Colle Fauniera env., 2480-2500 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="23" orientation="latitude" precision="15" seconds="08" value="44.38556">44°23'08"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="7" direction="east" minutes="7" orientation="longitude" precision="15" seconds="19" value="7.1219444">7°7'19"E</geoCoordinate>
|
||||
, 3 Aug. 2008, leg. Huemer, DNA Barcodes TLMF Lep 00107, 00108, 03747, 03748, 03749, 2 gen. slides in glycerin (TLMF); 2 ♂, same data, but 19 Jul. 2016, leg. Schmid, DNA Barcodes TLMF Lep 22437, 22438 (RCJS); 5 ♂♂, Cuneo province, Gias Valcavera, 2050 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="22,6" orientation="latitude" precision="92" value="44.376667">44°22,6'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="07" direction="east" minutes="06,2" orientation="longitude" precision="92" value="7.1033335">07°06,2'E</geoCoordinate>
|
||||
, 23 Jul. 2009, leg. Mayr; 32 ♂♂, same data, but 27 Jul. 2009; 1 ♂, same data, but 17 Aug. 2012; 3 ♂♂, same data, but 15 Aug. 2019 (all RCTM); 3 ♂♂, same data, but 23 Jul. 2009, leg. Huemer, DNA Barcode TLMF Lep 00533 (TLMF); 6 ♂♂, Cuneo province, Colle Valcavera, 2450 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="22,9" orientation="latitude" precision="92" value="44.381668">44°22,9'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="07" direction="east" minutes="07,2" orientation="longitude" precision="92" value="7.12">07°07,2'E</geoCoordinate>
|
||||
, 28 Jul. 2015, leg. Mayr (RCTM); 1 ♂, same data, but 2400-2450 m, 5 Aug. 2008, leg. Huemer; 12 ♂♂, Colle Valcavera NE, 2450 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="23" orientation="latitude" precision="15" seconds="04" value="44.384445">44°23'04"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="07" direction="east" minutes="06" orientation="longitude" precision="15" seconds="23" value="7.1063886">07°06'23"E</geoCoordinate>
|
||||
, 17 Aug. 2012, leg. Huemer (TLMF); 9 ♂♂, Cuneo prov., N Colle della Lombarda, 2370 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="12.12" orientation="latitude" precision="9" value="44.202">44°12.12'N</geoCoordinate>
|
||||
, 7°08.64' E, 18-19 Aug. 2012, leg. Huemer, gen. slide in glycerin (TLMF); 9 ♂♂, Torino province, PN Orsiera -
|
||||
<normalizedToken originalValue="Rocciavrè">Rocciavre</normalizedToken>
|
||||
, Usseaux, Colle delle Finestre N, 2180 m,
|
||||
<geoCoordinate degrees="45" direction="north" minutes="4" orientation="latitude" precision="15" seconds="21" value="45.0725">45°4'21"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="7" direction="east" minutes="3" orientation="longitude" precision="15" seconds="11" value="7.053056">7°3'11"E</geoCoordinate>
|
||||
, 24 Jul. 2019, leg. Huemer (TLMF); 3 ♂♂, 1 ♀, Torino province, Via Colle delle Finestre, Pequerel NE, 1840 m,
|
||||
<geoCoordinate degrees="45" direction="north" minutes="3" orientation="latitude" precision="1" seconds="7.65" value="45.052124">45°3'7.65"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="7" direction="east" minutes="4" orientation="longitude" precision="1" seconds="16.27" value="7.071186">7°4'16.27"E</geoCoordinate>
|
||||
, 26 Aug. 2019, leg. Wieser (LMK); 28 ♂♂, Torino province, Testa dell Assietta, 2530 m,
|
||||
<geoCoordinate degrees="45" direction="north" minutes="3" orientation="latitude" precision="1" seconds="49.56" value="45.063766">45°3'49.56"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="6" direction="east" minutes="57" orientation="longitude" precision="1" seconds="2.79" value="6.9507747">6°57'2.79"E</geoCoordinate>
|
||||
, 25 Aug. 2019, leg. Wieser (LMK); 1 ♂, Torino prov., Meana di Susa, Piano del Tiraculo, 2000 m, 25 Jul. 2007, leg. Baldizzone (TLMF). France: 22 ♂♂, Dep. Alpes-Maritimes, Col de la Lombarde, 2350 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="12" orientation="latitude" precision="15" seconds="8" value="44.202225">44°12'8"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="7" direction="east" minutes="9" orientation="longitude" precision="925" value="7.15">7°9'E</geoCoordinate>
|
||||
, 18-19 Aug. 2012, leg. Huemer, gen. slide in glycerine; 2 ♂♂, Dep. Alpes-Maritimes, PN Mercantour, N Col de la Cayolle, Col de la Boucharde N, 1930-1950 m,
|
||||
<geoCoordinate degrees="44" direction="north" minutes="17" orientation="latitude" precision="925" value="44.283333">44°17'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="6" direction="east" minutes="44" orientation="longitude" precision="15" seconds="36" value="6.743334">6°44'36"E</geoCoordinate>
|
||||
, 26 Jul. 2009, leg. Huemer; 1 ♂, Alpes-de-Haute-Provence, Col
|
||||
<normalizedToken originalValue="d´Allos">d'Allos</normalizedToken>
|
||||
, 2400 m, 21 Jul. 1999, leg. J. Nel, gen. slide 9458 ♂ J. Nel (all TLMF).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/alpento.4.50703.figures43-46" httpUri="https://binary.pensoft.net/fig/403199" pageId="0" pageNumber="39" start="Figures 43–46" startId="F14">
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">Figures 43-46.</emphasis>
|
||||
Female genitalia (signum).
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">43</emphasis>
|
||||
<taxonomicName baseAuthorityName="Christoph" baseAuthorityYear="1872" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum schleichi" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="schleichi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">Caryocolum schleichi</emphasis>
|
||||
</taxonomicName>
|
||||
, Syria, slide GU 86/277 P. Huemer;
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">44</emphasis>
|
||||
<taxonomicName genus="C." lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
|
||||
</taxonomicName>
|
||||
, Spain, slide GEL 1285 P. Huemer;
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">45</emphasis>
|
||||
<taxonomicName genus="C." lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
|
||||
</taxonomicName>
|
||||
, Italy, slide GEL 1291 P. Huemer;
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">46</emphasis>
|
||||
<taxonomicName genus="C." lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
|
||||
</taxonomicName>
|
||||
, Sweden, slide GEL 1292 P. Huemer.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption doi="10.3897/alpento.4.50703.figures47-49" httpUri="https://binary.pensoft.net/fig/403200" pageId="0" pageNumber="39" start="Figures 47–49" startId="F15">
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">Figures 47-49.</emphasis>
|
||||
Female genitalia (signum).
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">47</emphasis>
|
||||
<taxonomicName authorityName="Huemer" authorityYear="2020" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum messneri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="messneri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">Caryocolum messneri</emphasis>
|
||||
</taxonomicName>
|
||||
, paratype, Italy, slide GEL 1295 P. Huemer;
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">48</emphasis>
|
||||
<taxonomicName genus="C." lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
|
||||
</taxonomicName>
|
||||
, paratype, Italy, slide GEL 1286 P. Huemer;
|
||||
<emphasis bold="true" pageId="0" pageNumber="39">49</emphasis>
|
||||
<taxonomicName genus="C." lsidName="C. habeleri" pageId="0" pageNumber="39" rank="species" species="habeleri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. habeleri</emphasis>
|
||||
</taxonomicName>
|
||||
, paratype, Germany, slide M 1703.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="39">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
<taxonomicName genus="C." lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
|
||||
</taxonomicName>
|
||||
is, together with
|
||||
<taxonomicName genus="C." lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
|
||||
</taxonomicName>
|
||||
, the smallest and darkest species of the
|
||||
<taxonomicName genus="C." lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||||
</taxonomicName>
|
||||
species group with reduced white markings and brown scales. It differs from
|
||||
<taxonomicName genus="C." lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||||
</taxonomicName>
|
||||
by the brown rather than white head, thorax and tegulae but cannot be reliably separated from the other species of the complex by external characters of the forewings. However, the male genitalia are characterized by the shape of the valva, particularly the long, sharply pointed dorsal process, only shared with
|
||||
<taxonomicName genus="C." lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName genus="C." lsidName="C. messneri" pageId="0" pageNumber="39" rank="species" species="messneri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. messneri</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="C." lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
|
||||
</taxonomicName>
|
||||
. This process is more slender and longer than in
|
||||
<taxonomicName genus="C." lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="C." lsidName="C. messneri" pageId="0" pageNumber="39" rank="species" species="messneri">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. messneri</emphasis>
|
||||
</taxonomicName>
|
||||
, whereas the male genitalia differ from
|
||||
<taxonomicName genus="C." lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
|
||||
</taxonomicName>
|
||||
in particular in the more slender sacculus and the broader and the digitate medial processes of the vinculum. The female genitalia are hardly discernible from other species of the
|
||||
<taxonomicName genus="C." lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
|
||||
</taxonomicName>
|
||||
species group, except for the signum with a slender and long crescent-shaped base, a character only shared with
|
||||
<taxonomicName genus="C." lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
|
||||
</taxonomicName>
|
||||
, from which it differs by the longer apophysis posterior. However, the individual variation of this character is insufficiently documented due to lack of material.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="description">
|
||||
<paragraph pageId="0" pageNumber="39">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
Adult (Figs
|
||||
<figureCitation captionStart="Figures 9–12" captionStartId="F4" captionText="Figures 9 - 12. Adults. 9 Caryocolum lamai, paratype, male, Italy; 10 C. lamai, paratype, male, Italy; 11 C. messneri, paratype, male, Italy; 12 C. habeleri, paratype, male, France." figureDoi="10.3897/alpento.4.50703.figures9-12" httpUri="https://binary.pensoft.net/fig/403189" pageId="0" pageNumber="39">9</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 9–12" captionStartId="F4" captionText="Figures 9 - 12. Adults. 9 Caryocolum lamai, paratype, male, Italy; 10 C. lamai, paratype, male, Italy; 11 C. messneri, paratype, male, Italy; 12 C. habeleri, paratype, male, France." figureDoi="10.3897/alpento.4.50703.figures9-12" httpUri="https://binary.pensoft.net/fig/403189" pageId="0" pageNumber="39">10</figureCitation>
|
||||
). Forewing length ♂ 4.5-5.5 mm, ♀ 4.0 mm. Head dark grey-brown, mottled with light grey, frons greyish white; labial palpus dark brown, second segment cream-white on inner surface, third segment blackish, mottled with white; antenna black, weakly ringed paler. Thorax and tegula dark grey-brown, with few rusty brown scales. Abdomen dark grey on ventral surface. Forewing dark brown with some light mottling, particularly on dorsum; distinct white markings: narrow transverse fascia from fold to costa at one-fifth, irregular white medial spot, separate costal and tornal spots; fringes basally dark brown, distal part lighter. Hindwing light grey.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="variation">
|
||||
<paragraph pageId="0" pageNumber="39">Variation.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">The extension of light mottling of the forewing varies slightly.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="male genitalia">
|
||||
<paragraph pageId="0" pageNumber="39">Male genitalia</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
(Figs
|
||||
<figureCitation captionStart="Figures 17–20" captionStartId="F6" captionText="Figures 17 - 20. Male genitalia. 17 Caryocolum arenariella, Russia, slide GEL 1260 P. Huemer; 18 C. messneri, holotype, Italy, slide GEL 1255; 19 C. lamai, paratype, Italy, slide GEL 1258 P. Huemer; 21 C. habeleri, paratype, France, slide GEL 1289 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures17-20" httpUri="https://binary.pensoft.net/fig/403191" pageId="0" pageNumber="39">19</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 25–28" captionStartId="F8" captionText="Figures 25 - 28. Male genitalia (details vinculum-valva-complex). 25 Caryocolum arenariella, Russia, slide GEL 1260 P. Huemer; 26 C. messneri, holotype, Italy, slide GEL 1255; 27 C. lamai, paratype, Italy, slide GEL 1258 P. Huemer; 28 C. habeleri, paratype, France, slide GEL 1289 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures25-28" httpUri="https://binary.pensoft.net/fig/403193" pageId="0" pageNumber="39">27</figureCitation>
|
||||
). Uncus broadly sub-quadrangular, posterior corners rounded; lateral sclerites of gnathos distinct, medial part with large minutely spined culcitula; tegumen weakly widened anteriorly, with slightly emarginated anterior margin; transtilla sclerotized, longitudinally folded; pedunculus large, sub-triangular, with sclerotized inner edge; valva nearly straight, long, distal part moderately slender, weakly dilated with straight dorsal and ventral edges, apex with two processes, particularly long and pointed dorsal process extended to posterior third of uncus, slender digitate ventral process at about right-angles to dorsal process; sacculus long, slender, with parallel margins, apically pointed; posterior margin of vinculum deeply incised medially, with pair of short digitate processes, pair of latero-medial processes broadly projected; saccus slightly longer than valva, slender, gradually tapered to apex; phallus long, slender, nearly straight, apically with area of small cornuti.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="female genitalia">
|
||||
<paragraph pageId="0" pageNumber="39">Female genitalia</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
(Figs
|
||||
<figureCitation captionStart="Figures 33–35" captionStartId="F11" captionText="Figures 33 - 35. Female genitalia. 33 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 34 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 35 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures33-35" httpUri="https://binary.pensoft.net/fig/403196" pageId="0" pageNumber="39">34</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 40–42" captionStartId="F13" captionText="Figures 40 - 42. Female genitalia (segment VIII). 40 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 41 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 42 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures40-42" httpUri="https://binary.pensoft.net/fig/403198" pageId="0" pageNumber="39">41</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 47–49" captionStartId="F15" captionText="Figures 47 - 49. Female genitalia (signum). 47 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 48 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 49 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures47-49" httpUri="https://binary.pensoft.net/fig/403200" pageId="0" pageNumber="39">48</figureCitation>
|
||||
). Apophysis posterior about four times length of apophysis anterior; segment VIII without processes, smooth; ostium bursae with short lateral folds; apophysis anterior about length of segment VIII; antrum short, about one-fifth length of apophysis anterior, funnel-shaped; posterior part of ductus bursae with pair of long lateral sclerites extending to about apex of apophysis anterior, membranous part of ductus bursae about length of segment VIII including apophysis anterior; signum on right side of entrance to sub-oval corpus bursae, with long and slender crescent-shaped base and long slender hook.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="molecular data">
|
||||
<paragraph pageId="0" pageNumber="39">Molecular data.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
BIN: BOLD:AAE9478. The intraspecific average distance of the barcode region is 0.04%, the maximum distance 0.15% (p-dist) (n = 7). The minimum distance to the nearest neighbor,
|
||||
<taxonomicName genus="C." lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
|
||||
</taxonomicName>
|
||||
, is 3.05%.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="39">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">The species is named in honour of David Lama (1990-2019), one of the most famous Austrian alpinists, who was tragically killed by an avalanche in Banff National Park (Canada) on the 1st of April 2019. David supported earlier work on landscape conservation in Tyrol with enthusiasm.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="39">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">The species is currently only known from a restricted area in northern Italy (region of Piedmont) and France (Alpes-Maritimes, Alpes-de-Haute-Provence).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="39" type="bionomics">
|
||||
<paragraph pageId="0" pageNumber="39">Bionomics.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="39">
|
||||
Host plant and early stages are unknown but it seems most likely that the species shows a similar behaviour as related taxa with a hostplant restriction to
|
||||
<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="39">Dianthus</emphasis>
|
||||
</taxonomicName>
|
||||
spp. The adults have been found from late July until late August near rock and scree at altitudes of about 1800 to 2600 m on calcareous soil (Fig.
|
||||
<figureCitation captionStart="Figure 50" captionStartId="F16" captionText="Figure 50. Habitat of Caryocolum lamai, Italy, Alpi Cozie, Colle Valcavera." figureDoi="10.3897/alpento.4.50703.figure50" httpUri="https://binary.pensoft.net/fig/403201" pageId="0" pageNumber="39">50</figureCitation>
|
||||
), where they were attracted to artificial light sources.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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