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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.361.6448" ID-GBIF-Dataset="4f6ecf86-d77f-4c6a-803c-1bbc5ec78679" ID-PMC="PMC3867118" ID-Pensoft-Pub="1313-2970-361-27" ID-PubMed="24363595" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-361-27" ModsDocOrigin="ZooKeys 361" ModsDocTitle="Reassessment of Paleotachina Townsend and Electrotachina Townsend and their removal from the Tachinidae (Diptera)" checkinTime="1451246575341" checkinUser="pensoft" docAuthor="O'Hara, James E., Raper, Christopher M., Pont, Adrian C. &amp; Whitmore, Daniel" docDate="2013" docId="DDE9FDFBD6AD84AE70F92275ADC4574C" docLanguage="en" docName="ZooKeys 361: 27-36" docOrigin="ZooKeys 361" docSource="http://dx.doi.org/10.3897/zookeys.361.6448" docTitle="Dolichotachina smithii Townsend 1938, comb. n." docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="E345FFA7B630D07991610A24205FFFFC" masterDocTitle="Reassessment of Paleotachina Townsend and Electrotachina Townsend and their removal from the Tachinidae (Diptera)" masterLastPageNumber="36" masterPageNumber="27" pageNumber="31" updateTime="1668157156992" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Reassessment of Paleotachina Townsend and Electrotachina Townsend and their removal from the Tachinidae (Diptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>O'Hara, James E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raper, Christopher M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pont, Adrian C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Whitmore, Daniel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>361</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>36</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.361.6448</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.361.6448</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-361-27</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152050070" LSID="urn:lsid:plazi:treatment:DDE9FDFBD6AD84AE70F92275ADC4574C" httpUri="http://treatment.plazi.org/id/DDE9FDFBD6AD84AE70F92275ADC4574C" lastPageId="6" lastPageNumber="33" pageId="4" pageNumber="31">
<subSubSection pageId="4" pageNumber="31" type="nomenclature">
<paragraph pageId="4" pageNumber="31">
<taxonomicName LSID="http://species-id.net/wiki/Dolichotachina_smithii" authority="Townsend, 1938" authorityName="Townsend" authorityYear="1938" class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina smithii" order="Diptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="smithii">Dolichotachina smithii (Townsend, 1938)</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="31">comb. n.</taxonomicNameLabel>
Figs 7-13
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="31" type="reference_group">
<paragraph pageId="4" pageNumber="31">
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Electrotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electrotachina smithii" order="Diptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="smithii">Electrotachina smithii</taxonomicName>
Townsend, 1938: 166. Holotype female, in copal (NHM, No. 58551). Type locality: East Africa (
<bibRefCitation author="Grimaldi, DA" journalOrPublisher="Curator" pageId="7" pageNumber="34" pagination="251 - 274" title="Forgeries of fossils in &quot; amber &quot;: history, identification and case studies." url="10.1111/j.2151-6952.1994.tb01023.x" volume="37" year="1994">Grimaldi et al. 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="33" pageId="4" pageNumber="31" type="remarks">
<paragraph pageId="4" pageNumber="31">Remarks.</paragraph>
<paragraph pageId="4" pageNumber="31">
<bibRefCitation author="Townsend, CHT" journalOrPublisher="Entomological News" pageId="8" pageNumber="35" pagination="166 - 167" title="Five new genera of fossil Oestromuscaria. (Diptera)." volume="49" year="1938">Townsend (1938</bibRefCitation>
: 166) began his description of
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Electrotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electrotachina" order="Diptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Electrotachina</taxonomicName>
with: &quot;Genotype,
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Electrotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electrotachina smithii" order="Diptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="smithii">Electrotachina smithii</taxonomicName>
sp. nov. For new genus
<taxonomicName family="Muscidae" lsidName="" pageId="4" pageNumber="31" rank="family">Muscidae</taxonomicName>
aff. Tachina sp. F. Smith, Quart. Jn. Sc., V, 184, pl. 18, fig. 5 (
<bibRefCitation author="Smith, F" journalOrPublisher="Quarterly Journal of Science" pageId="8" pageNumber="35" pagination="183 - 184" title="Explanation of the plate of organic remains found in amber." volume="5" year="1868">1868</bibRefCitation>
). Fly Lower Oligocene of Baltic amber&quot;. A brief description followed, ending with the statement &quot;probably exoristid or tachinid stock&quot;. There is no indication that Townsend examined the specimen and his description is consistent with the drawing of a fly in fig. 5 in
<bibRefCitation author="Smith, F" journalOrPublisher="Quarterly Journal of Science" pageId="8" pageNumber="35" pagination="183 - 184" title="Explanation of the plate of organic remains found in amber." volume="5" year="1868">Smith (1868)</bibRefCitation>
. As with
<taxonomicName class="Insecta" family="Muscidae" genus="Paleotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paleotachina" order="Diptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Paleotachina</taxonomicName>
,
<bibRefCitation author="Townsend, CHT" journalOrPublisher="Itaquaquecetuba, Sao Paulo" pageId="8" pageNumber="35" title="Manual of myiology in twelve parts. Part XII. General consideration of the Oestromuscaria. Geologic history &amp; geographic distribution-environment &amp; response-relations to man-hosts &amp; flowers-bibliography &amp; plates." year="1942">Townsend (1942</bibRefCitation>
: 12) later provided a brief description of the genus, presumably from fig. 5 in
<bibRefCitation author="Smith, F" journalOrPublisher="Quarterly Journal of Science" pageId="8" pageNumber="35" pagination="183 - 184" title="Explanation of the plate of organic remains found in amber." volume="5" year="1868">Smith (1868)</bibRefCitation>
, and referred to the genus as &quot;almost certainly exoristid stock&quot;.
</paragraph>
<paragraph pageId="5" pageNumber="32">
<pageBreakToken pageId="5" pageNumber="32" start="start">As</pageBreakToken>
with
<taxonomicName class="Insecta" family="Muscidae" genus="Paleotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paleotachina smithii" order="Diptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="smithii">Paleotachina smithii</taxonomicName>
, certain liberties were taken in the depiction of NHM specimen #58551 (holotype of
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Electrotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electrotachina smithii" order="Diptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="smithii">Electrotachina smithii</taxonomicName>
) in fig. 5 in
<bibRefCitation author="Smith, F" journalOrPublisher="Quarterly Journal of Science" pageId="8" pageNumber="35" pagination="183 - 184" title="Explanation of the plate of organic remains found in amber." volume="5" year="1868">Smith (1868)</bibRefCitation>
, and it may also have been shown as a mirror image of the original specimen; cf. Figs 7, 13.
</paragraph>
<paragraph pageId="6" pageNumber="33">
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Electrotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electrotachina smithii" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="smithii">
<pageBreakToken pageId="6" pageNumber="33" start="start">Electrotachina</pageBreakToken>
smithii
</taxonomicName>
belongs to the family
<taxonomicName family="Sarcophagidae" lsidName="" pageId="6" pageNumber="33" rank="family">Sarcophagidae</taxonomicName>
, subfamily
<taxonomicName lsidName="" pageId="6" pageNumber="33" rank="subFamily" subFamily="Miltogramminae">Miltogramminae</taxonomicName>
. The holotype female has a body length of about 7 mm and is preserved in a small block of copal (approx. 15
<normalizedToken originalValue="×">x</normalizedToken>
10
<normalizedToken originalValue="×">x</normalizedToken>
7 mm) together with two other adult dipterans: a small female
<taxonomicName family="Agromyzidae" lsidName="" pageId="6" pageNumber="33" rank="family">Agromyzidae</taxonomicName>
and a
<taxonomicName family="Cecidomyiidae" lsidName="" pageId="6" pageNumber="33" rank="family">Cecidomyiidae</taxonomicName>
. The specimen is in very good condition except for the lack of its right fore tarsus. Antennae, wings and chaetotaxy are all in excellent condition. The specimen can be confidently assigned to the genus
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">Dolichotachina</taxonomicName>
based on the following combination of external character states: arista short pubescent, thickened on approximately basal 1/5; eye bare; parafacial with an uneven row of setae anteriorly; proepisternum bare; katepisternum with two, widely separated, setae; mid tibia with one anterodorsal seta; wing cell r4+5 open at wing margin.
</paragraph>
<paragraph pageId="6" pageNumber="33">
In addition to the above features,
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina smithii" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="smithii">Dolichotachina smithii</taxonomicName>
is characterized by an elongated postpedicel (about 3 times length of pedicel), relatively short vibrissa, and short proboscis (about twice as long as wide).
</paragraph>
<paragraph pageId="6" pageNumber="33">
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">Dolichotachina</taxonomicName>
is a mainly Afrotropical genus with 12 species previously known from this region (
<bibRefCitation author="Pape, T" journalOrPublisher="Memoirs on Entomology, International" pageId="7" pageNumber="34" pagination="1 - 558" title="Catalogue of the Sarcophagidae of the world (Insecta: Diptera)." volume="8" year="1996">Pape 1996</bibRefCitation>
). Material recently collected in Burundi and Namibia has demonstrated that the Afrotropical diversity of
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">Dolichotachina</taxonomicName>
is probably greatly underestimated (Whitmore &amp; Pape, unpublished). The condition of the holotype and the difficulty of identifying
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">Dolichotachina</taxonomicName>
females have not allowed us to verify whether this specimen is conspecific with any of the other described species. Lacking any strong indication to the contrary, we consider
<taxonomicName class="Insecta" family="Sarcophagidae" genus="Dolichotachina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichotachina smithii" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="smithii">Dolichotachina smithii</taxonomicName>
to be a valid, probably extant, species from East Africa.
</paragraph>
</subSubSection>
</treatment>
</document>