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<document id="EC672A16C8F14E2B770AF007D3E3CBC9" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8151" ID-PMC="PMC4867695" ID-Pensoft-Pub="1314-2828-4-8151" ID-PubMed="27226759" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8151" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" checkinTime="1461288934892" checkinUser="pensoft" docAuthor="Broad, Gavin R., Shaw, Mark R. &amp; Godfray, H. Charles J." docDate="2016" docId="DD7A0EAB29F326EAF9B2404948C4742C" docLanguage="en" docName="BiodivDatJour 4: e8151" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8151" docTitle="Chorebus ovalis Marshall 1896" docType="treatment" docVersion="5" lastPageNumber="8151" masterDocId="EC41FFB1FFD9782CFFAEFFF87E2EFF92" masterDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" masterLastPageNumber="8151" masterPageNumber="8151" pageNumber="8151" updateTime="1668123791244" updateUser="ExternalLinkService">
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<mods:title id="BE4239009A9489D3D6DD132CFE99A9C9">Checklist of British and Irish Hymenoptera - Braconidae</mods:title>
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<mods:namePart id="C5CB56CBB934E5B6BCCB27ECA8EAE06B">Broad, Gavin R.</mods:namePart>
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<mods:namePart id="83C864C9D3628938EEE139133BCDAF45">Shaw, Mark R.</mods:namePart>
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<treatment id="DD7A0EAB29F326EAF9B2404948C4742C" LSID="urn:lsid:plazi:treatment:DD7A0EAB29F326EAF9B2404948C4742C" httpUri="http://treatment.plazi.org/id/DD7A0EAB29F326EAF9B2404948C4742C" lastPageNumber="8151" pageId="0" pageNumber="8151">
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<taxonomicName id="ABF17CE4843FF293BCEE451AD9BEE073" ID-CoL="5YBWD" ID-ENA="2578160" authority="Marshall, 1896" authorityName="Marshall" authorityYear="1896" class="Insecta" family="Braconidae" genus="Chorebus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chorebus ovalis" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="ovalis">Chorebus ovalis (Marshall, 1896)</taxonomicName>
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<paragraph id="0E6E77802709CE621D895E1D0F5ED5DF" pageId="0" pageNumber="8151">
<taxonomicName id="F84F380C78149038F19B7453F538BC70" class="Insecta" family="Braconidae" genus="Dacnusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dacnusa ovalis" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="ovalis">Dacnusa ovalis</taxonomicName>
Marshall, 1896
</paragraph>
</subSubSection>
<subSubSection id="5B8B169DB03C1282773273E452103312" pageId="0" pageNumber="8151" type="distribution">
<paragraph id="4FECA81E74026CC4C1B30B62BD43E491" pageId="0" pageNumber="8151">Distribution</paragraph>
<paragraph id="8977E4A4CFD0354E464433B7B145D159" pageId="0" pageNumber="8151">England, Ireland</paragraph>
</subSubSection>
</treatment>
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<mods:title id="EE03486FC514D541C529E6B52101A353">New species and records of ptyctimous mites (Acari, Oribatida) from Cuba</mods:title>
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<heading id="0E241D69FFEDFFDEFF0C44C0934DF945" bold="true" box="[151,777,1753,1780]" fontSize="11" level="1" pageId="3" pageNumber="138" reason="1">
<emphasis id="67A77617FFEDFFDEFF0C44C0934DF945" bold="true" box="[151,777,1753,1780]" pageId="3" pageNumber="138">
<taxonomicName id="92D3D186FFEDFFDEFF0C44C092E0F945" authority="Niedbala" authorityName="Niedbala" box="[151,676,1753,1780]" class="Arachnida" family="Phthiracaridae" genus="Protophthiracarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="paratripartitus" status="sp. nov.">
<emphasis id="67A77617FFEDFFDEFF0C44C09269F942" bold="true" box="[151,557,1753,1779]" italics="true" pageId="3" pageNumber="138">Protophthiracarus paratripartitus</emphasis>
Niedbała
</taxonomicName>
<taxonomicNameLabel id="7C94CB6CFFEDFFDEFD3744C3934DF945" box="[684,777,1754,1780]" pageId="3" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="556CAA05FFEDFFDEFF0C44E59165F8A4" blockId="3.[151,777,1753,1813]" box="[151,289,1788,1813]" pageId="3" pageNumber="138">
(
<figureCitation id="CDE8B680FFEDFFDEFF0444E5915DF8A4" box="[159,281,1788,1813]" captionStart="FIGURE 3 A – G" captionStartId="5.[151,250,1892,1914]" captionTargetBox="[196,1390,193,1870]" captionTargetId="figure@5.[196,1390,193,1870]" captionTargetPageId="5" captionText="FIGURE 3 A – G. Protophthiracarus paratripartitus Niedbała sp. nov. (holotype); measurements: prodorsum: length 333, width 252, height 136; notogaster: length 737, width 454, height 394. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, mentum of subcapitulum; E, left genitoaggenital plate, F, left anoadanal plate; G, trochanter and femur of leg I." httpUri="https://zenodo.org/record/232411/files/figure.png" pageId="3" pageNumber="138">Fig. 3A–G</figureCitation>
)
</paragraph>
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<paragraph id="556CAA05FFEDFFDEFF0C455D937CF819" blockId="3.[151,1436,1860,1996]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF0C455D916DF8EC" bold="true" box="[151,297,1860,1885]" pageId="3" pageNumber="138">Description.</emphasis>
<emphasis id="67A77617FFEDFFDEFEAB455F9196F8EC" box="[304,466,1862,1885]" italics="true" pageId="3" pageNumber="138">Measurements</emphasis>
of
<typeStatus id="8A6814A7FFEDFFDEFE61455D9224F8EC" box="[506,608,1860,1885]" pageId="3" pageNumber="138" type="holotype">holotype</typeStatus>
. Prodorsum: length 333, width 252, height 136; setae:
<emphasis id="67A77617FFEDFFDEFB57455F94A6F8EC" box="[1228,1250,1862,1885]" italics="true" pageId="3" pageNumber="138">ss</emphasis>
58,
<emphasis id="67A77617FFEDFFDEFA8F455F956EF8EC" box="[1300,1322,1862,1885]" italics="true" pageId="3" pageNumber="138">in</emphasis>
66,
<emphasis id="67A77617FFEDFFDEFAC6455C9535F8EC" box="[1373,1393,1861,1885]" italics="true" pageId="3" pageNumber="138">le</emphasis>
30,
<emphasis id="67A77617FFEDFFDEFF0C457090EAF831" box="[151,174,1897,1920]" italics="true" pageId="3" pageNumber="138">ro</emphasis>
63,
<emphasis id="67A77617FFEDFFDEFF7545709142F831" box="[238,262,1897,1920]" italics="true" pageId="3" pageNumber="138">ex</emphasis>
25; notogaster: length 737, width 454, height 394; setae:
<emphasis id="67A77617FFEDFFDEFC4F457093A4F831" box="[980,992,1897,1920]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1 144,
<emphasis id="67A77617FFEDFFDEFBAE45709405F831" box="[1077,1089,1897,1920]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1/
<emphasis id="67A77617FFEDFFDEFBCA45709419F831" box="[1105,1117,1897,1920]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1–
<emphasis id="67A77617FFEDFFDEFBE8457094C5F830" box="[1139,1153,1897,1921]" italics="true" pageId="3" pageNumber="138">d</emphasis>
1=0.65,
<emphasis id="67A77617FFEDFFDEFB44457094A9F830" box="[1247,1261,1897,1921]" italics="true" pageId="3" pageNumber="138">h</emphasis>
1 139,
<emphasis id="67A77617FFEDFFDEFAD94570951FF831" box="[1346,1371,1897,1920]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
1 134; genitoaggenital plate 182 × 151; anoadanal plate 247 × 177.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C45AD9455F87D" blockId="3.[151,1436,1860,1996]" box="[199,1041,1971,1996]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C45AD9101F87A" box="[199,325,1972,1995]" italics="true" pageId="3" pageNumber="138">Integument</emphasis>
. Colour brown. Surface of body covered with deep concavities.
</paragraph>
<caption id="01ACFA8DFFEAFFD9FF0C45769537F873" httpUri="https://zenodo.org/record/232410/files/figure.png" pageId="4" pageNumber="139" targetBox="[208,1378,193,1881]" targetPageId="4">
<paragraph id="556CAA05FFEAFFD9FF0C45769537F873" blockId="4.[151,1436,1903,1986]" pageId="4" pageNumber="139">
<emphasis id="67A77617FFEAFFD9FF0C45769179F834" bold="true" box="[151,317,1903,1925]" pageId="4" pageNumber="139">FIGURE 2A–G</emphasis>
.
<taxonomicName id="92D3D186FFEAFFD9FED0457692B6F834" authority="Niedbala" authorityName="Niedbala" box="[331,754,1903,1925]" class="Arachnida" family="Phthiracaridae" genus="Hoplophthiracarus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="vinalesensis" status="sp. nov.">
<emphasis id="67A77617FFEAFFD9FED0457692CBF834" box="[331,655,1903,1925]" italics="true" pageId="4" pageNumber="139">Hoplophthiracarus vinalesensis</emphasis>
Niedbała
</taxonomicName>
<emphasis id="67A77617FFEAFFD9FD614576930FF835" bold="true" box="[762,843,1903,1924]" pageId="4" pageNumber="139">
<taxonomicNameLabel id="7C94CB6CFFEAFFD9FD614576930FF835" box="[762,843,1903,1924]" pageId="4" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); measurements: prodorsum: length 159, width 114, height 66; notogaster: length 283, width 202, height 182. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, left genitoaggenital plate, E, left anoadanal plate; F, trochanter and femur of leg I, G, tibia IV.
</paragraph>
</caption>
<caption id="01ACFA8DFFEBFFD8FF0C457D916DF867" httpUri="https://zenodo.org/record/232411/files/figure.png" pageId="5" pageNumber="140" targetBox="[196,1390,193,1870]" targetPageId="5">
<paragraph id="556CAA05FFEBFFD8FF0C457D916DF867" blockId="5.[151,1436,1892,2006]" pageId="5" pageNumber="140">
<emphasis id="67A77617FFEBFFD8FF0C457D917FF8CB" bold="true" box="[151,315,1892,1914]" pageId="5" pageNumber="140">FIGURE 3A–G</emphasis>
.
<taxonomicName id="92D3D186FFEBFFD8FEDC457D9344F8CB" authority="Niedbala" authorityName="Niedbala" box="[327,768,1892,1914]" class="Arachnida" family="Phthiracaridae" genus="Protophthiracarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="paratripartitus" status="sp. nov.">
<emphasis id="67A77617FFEBFFD8FEDC457D92E4F8CB" box="[327,672,1892,1914]" italics="true" pageId="5" pageNumber="140">Protophthiracarus paratripartitus</emphasis>
Niedbała
</taxonomicName>
<emphasis id="67A77617FFEBFFD8FC9D457D9312F8C8" bold="true" box="[774,854,1892,1913]" pageId="5" pageNumber="140">
<taxonomicNameLabel id="7C94CB6CFFEBFFD8FC9D457D9312F8C8" box="[774,854,1892,1913]" pageId="5" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); measurements: prodorsum: length 333, width 252, height 136; notogaster: length 737, width 454, height 394. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, mentum of subcapitulum; E, left genitoaggenital plate, F, left anoadanal plate; G, trochanter and femur of leg I.
</paragraph>
</caption>
<paragraph id="556CAA05FFE8FFDBFF5C4281918FFED5" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C42819100FF01" box="[199,324,152,176]" italics="true" pageId="6" pageNumber="141">Prodorsum</emphasis>
with long, strong humped median crista and two weaker lateral cristae. Lateral carinae distinct but short. Posterior furrows well marked. Sigillar fields visible only from dorsal view; median narrow, enlarged towards rostral setae, laterals shorter than median field. Sensilli club-like, fairly long, with long, thick stalk and dilated head, covered with small spines. Interlamellar setae rather short, procumbent, spiniform, rough. Lamellar setae short spiniform, smooth. Rostral setae thick, rough, directed inwards. Exobothridial setae shortest. Relation of lengths:
<emphasis id="67A77617FFE8FFDBFE8E4354916FFED5" box="[277,299,333,356]" italics="true" pageId="6" pageNumber="141">in</emphasis>
&gt;
<emphasis id="67A77617FFE8FFDBFEA143549115FED5" box="[314,337,333,356]" italics="true" pageId="6" pageNumber="141">ro</emphasis>
&gt;
<emphasis id="67A77617FFE8FFDBFEF94354913CFED5" box="[354,376,333,356]" italics="true" pageId="6" pageNumber="141">ss</emphasis>
&gt;
<emphasis id="67A77617FFE8FFDBFE1C435591DFFED5" box="[391,411,332,356]" italics="true" pageId="6" pageNumber="141">le</emphasis>
&gt;
<emphasis id="67A77617FFE8FFDBFE3743549181FED5" box="[428,453,333,356]" italics="true" pageId="6" pageNumber="141">ex</emphasis>
.
</paragraph>
<paragraph id="556CAA05FFE8FFDBFF5C4368953BFE4D" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C43689100FE39" box="[199,324,369,392]" italics="true" pageId="6" pageNumber="141">Notogaster</emphasis>
with 15 pairs of rigid, rather, thick, short (
<emphasis id="67A77617FFE8FFDBFCB74368937CFE39" box="[812,824,369,392]" italics="true" pageId="6" pageNumber="141">c</emphasis>
1&lt;
<emphasis id="67A77617FFE8FFDBFCD44368931FFE39" box="[847,859,369,392]" italics="true" pageId="6" pageNumber="141">c</emphasis>
1–
<emphasis id="67A77617FFE8FFDBFCE9436993C4FE39" box="[882,896,368,392]" italics="true" pageId="6" pageNumber="141">d</emphasis>
1) setae, covered with small spines in distal half and rather obtuse distally, except
<emphasis id="67A77617FFE8FFDBFD8F43819264FE1E" box="[532,544,408,431]" italics="true" pageId="6" pageNumber="141">c</emphasis>
3 and
<emphasis id="67A77617FFE8FFDBFDF94381923EFE1E" box="[610,634,408,431]" italics="true" pageId="6" pageNumber="141">ps</emphasis>
4 shortest, spiniform, smooth. Setae
<emphasis id="67A77617FFE8FFDBFB8E43819465FE1E" box="[1045,1057,408,431]" italics="true" pageId="6" pageNumber="141">c</emphasis>
3 situated near anterior margin,
<emphasis id="67A77617FFE8FFDBFA13438195D0FE1E" box="[1416,1428,408,431]" italics="true" pageId="6" pageNumber="141">c</emphasis>
1 slightly remote,
<emphasis id="67A77617FFE8FFDBFEC343A79120FE64" box="[344,356,446,469]" italics="true" pageId="6" pageNumber="141">c</emphasis>
2 far from margin. Vestigial setae
<emphasis id="67A77617FFE8FFDBFD6243A79345FE67" box="[761,769,446,470]" italics="true" pageId="6" pageNumber="141">f</emphasis>
1 slightly posterior of setae
<emphasis id="67A77617FFE8FFDBFBD743A4941EFE64" box="[1100,1114,445,469]" italics="true" pageId="6" pageNumber="141">h</emphasis>
1. Setae
<emphasis id="67A77617FFE8FFDBFB2643A79481FE67" box="[1213,1221,446,470]" italics="true" pageId="6" pageNumber="141">f</emphasis>
2 on left side well developed, similar to setae
<emphasis id="67A77617FFE8FFDBFE5343FC9190FE4D" box="[456,468,485,508]" italics="true" pageId="6" pageNumber="141">c</emphasis>
3 and
<emphasis id="67A77617FFE8FFDBFD8843FC9268FE4D" box="[531,556,485,508]" italics="true" pageId="6" pageNumber="141">ps</emphasis>
4; on right side
<emphasis id="67A77617FFE8FFDBFD4E43FD9299FE4D" box="[725,733,484,508]" italics="true" pageId="6" pageNumber="141">f</emphasis>
2 is vestigial. Four pairs of lyrifissures
<emphasis id="67A77617FFE8FFDBFB1643FC94E7FE4D" box="[1165,1187,485,508]" italics="true" pageId="6" pageNumber="141">ia</emphasis>
,
<emphasis id="67A77617FFE8FFDBFB2A43FC9489FE4D" box="[1201,1229,485,508]" italics="true" pageId="6" pageNumber="141">im</emphasis>
,
<emphasis id="67A77617FFE8FFDBFB4043FC94B5FE4D" box="[1243,1265,485,508]" italics="true" pageId="6" pageNumber="141">ip</emphasis>
,
<emphasis id="67A77617FFE8FFDBFB6443FC9564FE4D" box="[1279,1312,485,508]" italics="true" pageId="6" pageNumber="141">ips</emphasis>
present.
</paragraph>
<paragraph id="556CAA05FFE8FFDBFF5C401294DFFDDF" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C4012912EFD92" box="[199,362,523,547]" italics="true" pageId="6" pageNumber="141">Ventral region</emphasis>
. Setae
<emphasis id="67A77617FFE8FFDBFE5940129194FD92" box="[450,464,523,547]" italics="true" pageId="6" pageNumber="141">h</emphasis>
of mentum considerably longer than distance between them. Genitoaggenital plates with 9 pairs of setae with formula: 4:5. Anoadanal plates each with 5 setae; setae
<emphasis id="67A77617FFE8FFDBFBB84036947BFDF6" box="[1059,1087,559,583]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
1 and anal setae long, attenuate,
<emphasis id="67A77617FFE8FFDBFF0C404F90F7FDDF" box="[151,179,598,622]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
1 longest; setae
<emphasis id="67A77617FFE8FFDBFEFB404F9138FDDF" box="[352,380,598,622]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
2 and
<emphasis id="67A77617FFE8FFDBFE21404F9192FDDF" box="[442,470,598,622]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
3 short, smooth, similar in length to notogastral setae
<emphasis id="67A77617FFE8FFDBFBB2404E9471FDDF" box="[1065,1077,599,622]" italics="true" pageId="6" pageNumber="141">c</emphasis>
3 and
<emphasis id="67A77617FFE8FFDBFBEF404E94C9FDDF" box="[1140,1165,599,622]" italics="true" pageId="6" pageNumber="141">ps</emphasis>
4.
</paragraph>
<paragraph id="556CAA05FFE8FFDBFF5C406491FBFD08" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C406490B8FD25" box="[199,252,637,660]" italics="true" pageId="6" pageNumber="141">Legs</emphasis>
. Chaetome of setae and solenidia of “complete
<typeStatus id="8A6814A7FFE8FFDBFCAB40649321FD24" box="[816,869,637,661]" pageId="6" pageNumber="141">type</typeStatus>
”. Setae
<emphasis id="67A77617FFE8FFDBFC5F40649396FD24" box="[964,978,637,661]" italics="true" pageId="6" pageNumber="141">d</emphasis>
on femora I remote from distal end of article and curved distally.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFE8FFDBFF5C40DD9299FD6C" box="[199,733,708,733]" pageId="6" pageNumber="141" type="materials_examined">
<paragraph id="556CAA05FFE8FFDBFF5C40DD9299FD6C" blockId="6.[151,1437,151,988]" box="[199,733,708,733]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C40DD91F4FD6C" bold="true" box="[199,432,708,733]" pageId="6" pageNumber="141">Material examined.</emphasis>
<typeStatus id="8A6814A7FFE8FFDBFE2C40DD925BFD6C" box="[439,543,708,733]" pageId="6" pageNumber="141" type="holotype">Holotype</typeStatus>
from Locality 2.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFE8FFDBFF5C40F194D5FC6D" pageId="6" pageNumber="141" type="etymology">
<paragraph id="556CAA05FFE8FFDBFF5C40F19343FC94" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C40F1910AFCB0" bold="true" box="[199,334,744,769]" pageId="6" pageNumber="141">Etymology.</emphasis>
The prefix
<emphasis id="67A77617FFE8FFDBFE7940F09252FCB1" box="[482,534,745,768]" italics="true" pageId="6" pageNumber="141">para</emphasis>
is Latin meaning “near” and refers to the similarity of the new species to
<taxonomicName id="92D3D186FFE8FFDBFF0C41149346FC94" authority="Niedbala &amp; Schatz, 1996" authorityName="Niedbala &amp; Schatz" authorityYear="1996" box="[151,770,780,805]" class="Arachnida" family="Phthiracaridae" genus="Protophthiracarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="67A77617FFE8FFDBFF0C4114919BFC95" box="[151,479,781,805]" italics="true" pageId="6" pageNumber="141">Protophthiracarus tripartitus</emphasis>
<bibRefCitation id="3142D7F4FFE8FFDBFE7D41159346FC94" author="Niedbala" box="[486,770,780,805]" pageId="6" pageNumber="141" refString="Niedbala, W. &amp; Schatz, H. (1996) Euptyctimous mites from the Galapagos Islands, Cocos Islands, and Central America (Acari: Oribatida). Genus, 7 (2), 239 - 317." type="journal article" year="1996">Niedbała &amp; Schatz, 1996</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="556CAA05FFE8FFDBFF5C412994D5FC6D" blockId="6.[151,1437,151,988]" pageId="6" pageNumber="141">
<emphasis id="67A77617FFE8FFDBFF5C41299126FCF8" bold="true" box="[199,354,816,841]" pageId="6" pageNumber="141">Comparison.</emphasis>
The new species is morphologically similar to
<taxonomicName id="92D3D186FFE8FFDBFCE84128908BFCDC" authority="Niedbala &amp; Schatz, 1996" authorityName="Niedbala &amp; Schatz" authorityYear="1996" class="Arachnida" family="Phthiracaridae" genus="Protophthiracarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="tripartitus">
<emphasis id="67A77617FFE8FFDBFCE8412894FFFCF9" box="[883,1211,817,841]" italics="true" pageId="6" pageNumber="141">Protophthiracarus tripartitus</emphasis>
<bibRefCitation id="3142D7F4FFE8FFDBFB5A4129908BFCDC" author="Niedbala" pageId="6" pageNumber="141" refString="Niedbala, W. &amp; Schatz, H. (1996) Euptyctimous mites from the Galapagos Islands, Cocos Islands, and Central America (Acari: Oribatida). Genus, 7 (2), 239 - 317." type="journal article" year="1996">Niedbała &amp; Schatz, 1996</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="2DC4EA95FFE8FFDBFE88414D911FFCDC" box="[275,347,852,877]" name="Belize" pageId="6" pageNumber="141">Belize</collectingCountry>
by the presence of three lobes of dorsal cristae of prodorsum, similarity of shape of interlamellar and lamellar setae and setae
<emphasis id="67A77617FFE8FFDBFE43416091B0FC20" box="[472,500,889,913]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
2 and
<emphasis id="67A77617FFE8FFDBFDA84160920BFC20" box="[563,591,889,913]" italics="true" pageId="6" pageNumber="141">ad</emphasis>
3 being the shortest on anoadanal plates and some similarity of sensilli. It is distinguishable by the shape of longer notogastral setae, mostly rigid, thick and covered with spines (versus majority of setae short, spiniform); different formula of genital setae 4:5 (versus 6(4+2):3).
</paragraph>
</subSubSection>
</treatment>
</document>

213
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<treatment id="DD7A1B13FFEDFFDEFF0C40C39382F936" ID-DOI="http://doi.org/10.5281/zenodo.6103402" ID-GBIF-Taxon="119645823" ID-Zenodo-Dep="6103402" LSID="urn:lsid:plazi:treatment:DD7A1B13FFEDFFDEFF0C40C39382F936" httpUri="http://treatment.plazi.org/id/DD7A1B13FFEDFFDEFF0C40C39382F936" lastPageNumber="138" pageId="3" pageNumber="138">
<subSubSection id="1DC9F98EFFEDFFDEFF0C40C39166FCA7" pageId="3" pageNumber="138" type="nomenclature">
<paragraph id="556CAA05FFEDFFDEFF0C40C392B7FD44" blockId="3.[151,755,730,790]" box="[151,755,730,757]" pageId="3" pageNumber="138">
<heading id="0E241D69FFEDFFDEFF0C40C392B7FD44" bold="true" box="[151,755,730,757]" fontSize="11" level="1" pageId="3" pageNumber="138" reason="1">
<emphasis id="67A77617FFEDFFDEFF0C40C392B7FD44" bold="true" box="[151,755,730,757]" pageId="3" pageNumber="138">
<taxonomicName id="92D3D186FFEDFFDEFF0C40C392C9FD44" ID-CoL="7VFDM" authority="Niedbala" authorityName="Niedbala" box="[151,653,730,757]" class="Arachnida" family="Phthiracaridae" genus="Hoplophthiracarus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="vinalesensis" status="sp. nov.">
<emphasis id="67A77617FFEDFFDEFF0C40C39253FD45" bold="true" box="[151,535,730,756]" italics="true" pageId="3" pageNumber="138">Hoplophthiracarus vinalesensis</emphasis>
Niedbała
</taxonomicName>
<taxonomicNameLabel id="7C94CB6CFFEDFFDEFD0E40C292B7FD44" box="[661,755,731,757]" pageId="3" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="556CAA05FFEDFFDEFF0C40E49166FCA7" blockId="3.[151,755,730,790]" box="[151,290,765,790]" pageId="3" pageNumber="138">
(
<figureCitation id="CDE8B680FFEDFFDEFF0440E4915EFCA7" box="[159,282,765,790]" captionStart="FIGURE 2 A – G" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[208,1378,193,1881]" captionTargetId="figure@4.[208,1379,193,1882]" captionTargetPageId="4" captionText="FIGURE 2 A – G. Hoplophthiracarus vinalesensis Niedbała sp. nov. (holotype); measurements: prodorsum: length 159, width 114, height 66; notogaster: length 283, width 202, height 182. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, left genitoaggenital plate, E, left anoadanal plate; F, trochanter and femur of leg I, G, tibia IV." httpUri="https://zenodo.org/record/232410/files/figure.png" pageId="3" pageNumber="138">Fig. 2A–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEDFFDEFF0C415C9258FA39" pageId="3" pageNumber="138" type="description">
<paragraph id="556CAA05FFEDFFDEFF0C415C930FFC7C" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF0C415C916DFCEF" bold="true" box="[151,297,837,862]" pageId="3" pageNumber="138">Description.</emphasis>
<emphasis id="67A77617FFEDFFDEFEB5415F9194FCEC" box="[302,464,838,861]" italics="true" pageId="3" pageNumber="138">Measurements</emphasis>
of
<typeStatus id="8A6814A7FFEDFFDEFE6E415C921FFCEF" box="[501,603,837,862]" pageId="3" pageNumber="138" type="holotype">holotype</typeStatus>
. Prodorsum: length 159, width 114, height 66; setae:
<emphasis id="67A77617FFEDFFDEFB32415F94FBFCEC" box="[1193,1215,838,861]" italics="true" pageId="3" pageNumber="138">ss</emphasis>
and
<emphasis id="67A77617FFEDFFDEFB6F415F954EFCEC" box="[1268,1290,838,861]" italics="true" pageId="3" pageNumber="138">in</emphasis>
45,
<emphasis id="67A77617FFEDFFDEFAA1415C950BFCEC" box="[1338,1359,837,861]" italics="true" pageId="3" pageNumber="138">le</emphasis>
and
<emphasis id="67A77617FFEDFFDEFA18415F95D8FCEC" box="[1411,1436,838,861]" italics="true" pageId="3" pageNumber="138">ex</emphasis>
7,
<emphasis id="67A77617FFEDFFDEFF284173908EFC30" box="[179,202,874,897]" italics="true" pageId="3" pageNumber="138">ro</emphasis>
20; notogaster: length 283, width 202, height 182; setae:
<emphasis id="67A77617FFEDFFDEFCD64173931DFC30" box="[845,857,874,897]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1 48,
<emphasis id="67A77617FFEDFFDEFC0A417393D9FC30" box="[913,925,874,897]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1/
<emphasis id="67A77617FFEDFFDEFC36417393FDFC30" box="[941,953,874,897]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1–
<emphasis id="67A77617FFEDFFDEFC5441709399FC30" box="[975,989,873,897]" italics="true" pageId="3" pageNumber="138">d</emphasis>
1 = 0.6,
<emphasis id="67A77617FFEDFFDEFBA84173947BFC30" box="[1075,1087,874,897]" italics="true" pageId="3" pageNumber="138">c</emphasis>
3 and
<emphasis id="67A77617FFEDFFDEFBE6417394DCFC30" box="[1149,1176,874,897]" italics="true" pageId="3" pageNumber="138">cp</emphasis>
7,
<emphasis id="67A77617FFEDFFDEFB214170948CFC30" box="[1210,1224,873,897]" italics="true" pageId="3" pageNumber="138">h</emphasis>
1 48,
<emphasis id="67A77617FFEDFFDEFA9B4173955DFC30" box="[1280,1305,874,897]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
1 40,
<emphasis id="67A77617FFEDFFDEFACA4173952DFC30" box="[1361,1385,874,897]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
3 13; genitoaggenital plate 114×61, anoadanal plate 126×76. Measurements of
<typeStatus id="8A6814A7FFEDFFDEFC4241889479FC18" box="[985,1085,913,937]" pageId="3" pageNumber="138" type="paratype">paratype</typeStatus>
. Prodorsum: length 156, width 110, height 63; notogaster: length 313, width 208, height 192.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C41C092B0FC41" blockId="3.[151,1437,837,1671]" box="[199,756,984,1009]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C41C09101FC41" box="[199,325,985,1008]" italics="true" pageId="3" pageNumber="138">Integument</emphasis>
. Colour whitish, integument punctate.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C41E591F2FB30" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C41E59100FBA5" box="[199,324,1020,1044]" italics="true" pageId="3" pageNumber="138">Prodorsum</emphasis>
with feeble lateral carinae. Sigillar fields feeble, median very short, shorter than lateral fields. Sensilli long, with fusiform head covered with small spines. Interlamellar setae erect, curved posteriorly, ciliate in distal half. Lamellar setae short, spiniform. Rostral setae erect, rough, situated near each other (
<emphasis id="67A77617FFEDFFDEFB74465C956DFBED" box="[1263,1321,1093,1116]" italics="true" pageId="3" pageNumber="138">in–in</emphasis>
/
<emphasis id="67A77617FFEDFFDEFAAB465C9529FBED" box="[1328,1389,1093,1116]" italics="true" pageId="3" pageNumber="138">ro–ro</emphasis>
=3). Exobothridial setae short.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C469494BAFB40" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C46949106FB15" box="[199,322,1165,1188]" italics="true" pageId="3" pageNumber="138">Notogaster</emphasis>
. Notogaster with short (
<emphasis id="67A77617FFEDFFDEFDCD46949226FB15" box="[598,610,1165,1188]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1&lt;
<emphasis id="67A77617FFEDFFDEFDE1469492C2FB15" box="[634,646,1165,1188]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1–
<emphasis id="67A77617FFEDFFDEFD07469592EEFB15" box="[668,682,1164,1188]" italics="true" pageId="3" pageNumber="138">d</emphasis>
1) setae, setae
<emphasis id="67A77617FFEDFFDEFCD146949327FB15" box="[842,867,1165,1188]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
3 and
<emphasis id="67A77617FFEDFFDEFC38469493F8FB15" box="[931,956,1165,1188]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
4 shorter, setae
<emphasis id="67A77617FFEDFFDEFBFC46949437FB15" box="[1127,1139,1165,1188]" italics="true" pageId="3" pageNumber="138">c</emphasis>
3 and
<emphasis id="67A77617FFEDFFDEFB2F4694948AFB15" box="[1204,1230,1165,1188]" italics="true" pageId="3" pageNumber="138">cp</emphasis>
shortest; majority of setae with distal half curved anteriorly and ciliated; setae
<emphasis id="67A77617FFEDFFDEFCC946AD931AFB7A" box="[850,862,1204,1227]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1–3 remote from anterior border, setae
<emphasis id="67A77617FFEDFFDEFA9046AD9553FB7A" box="[1291,1303,1204,1227]" italics="true" pageId="3" pageNumber="138">c</emphasis>
2 more than setae
<emphasis id="67A77617FFEDFFDEFF4D46C390A6FB40" box="[214,226,1242,1265]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1 and
<emphasis id="67A77617FFEDFFDEFEBB46C39168FB40" box="[288,300,1242,1265]" italics="true" pageId="3" pageNumber="138">c</emphasis>
3. Vestigial setae
<emphasis id="67A77617FFEDFFDEFE7146C391B6FB43" box="[490,498,1242,1266]" italics="true" pageId="3" pageNumber="138">f</emphasis>
1 slightly posterior of setae
<emphasis id="67A77617FFEDFFDEFCBF46C39376FB43" box="[804,818,1242,1266]" italics="true" pageId="3" pageNumber="138">h</emphasis>
1. Two pairs of lyrifissures
<emphasis id="67A77617FFEDFFDEFBC446C39431FB40" box="[1119,1141,1242,1265]" italics="true" pageId="3" pageNumber="138">ia</emphasis>
,
<emphasis id="67A77617FFEDFFDEFB1846C394DBFB40" box="[1155,1183,1242,1265]" italics="true" pageId="3" pageNumber="138">im</emphasis>
present.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C471894BEFA8C" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C4718912DFAA9" box="[199,361,1281,1305]" italics="true" pageId="3" pageNumber="138">Ventral region</emphasis>
. Mentum of subcapitulum with setae
<emphasis id="67A77617FFEDFFDEFC864718936FFAA8" box="[797,811,1281,1305]" italics="true" pageId="3" pageNumber="138">h</emphasis>
shorter than distance between them; 9 pairs of genital setae present with formula 6(4+2):3; anal setae
<emphasis id="67A77617FFEDFFDEFD37473C9294FAF0" box="[684,720,1317,1345]" italics="true" pageId="3" pageNumber="138">ad1</emphasis>
and
<emphasis id="67A77617FFEDFFDEFC9D473C9366FA8C" box="[774,802,1317,1341]" italics="true" pageId="3" pageNumber="138">ad</emphasis>
2 longer than anal setae, setae
<emphasis id="67A77617FFEDFFDEFBEA473C94C9FA8C" box="[1137,1165,1317,1341]" italics="true" pageId="3" pageNumber="138">ad</emphasis>
3 shortest.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C47559258FA39" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C475590B8FAD2" box="[199,252,1356,1379]" italics="true" pageId="3" pageNumber="138">Legs</emphasis>
. Chaetome of legs incomplete, setae
<emphasis id="67A77617FFEDFFDEFD04475592EFFAD2" box="[671,683,1356,1379]" italics="true" pageId="3" pageNumber="138">v</emphasis>
’ on femora I and setae
<emphasis id="67A77617FFEDFFDEFC2F475293F8FAD2" box="[948,956,1355,1379]" italics="true" pageId="3" pageNumber="138">l</emphasis>
’ on genua IV are absent. Setae
<emphasis id="67A77617FFEDFFDEFABA4752956BFAD2" box="[1313,1327,1355,1379]" italics="true" pageId="3" pageNumber="138">d</emphasis>
on femur I situated near distal end of article.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEDFFDEFF5C478A93E7FA1D" box="[199,931,1427,1452]" pageId="3" pageNumber="138" type="materials_examined">
<paragraph id="556CAA05FFEDFFDEFF5C478A93E7FA1D" blockId="3.[151,1437,837,1671]" box="[199,931,1427,1452]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C478A91F4FA1D" bold="true" box="[199,432,1427,1452]" pageId="3" pageNumber="138">Material examined.</emphasis>
<typeStatus id="8A6814A7FFEDFFDEFE2C478A925BFA1D" box="[439,543,1427,1452]" pageId="3" pageNumber="138" type="holotype">Holotype</typeStatus>
and one
<typeStatus id="8A6814A7FFEDFFDEFD1E478D92A1FA1D" box="[645,741,1428,1452]" pageId="3" pageNumber="138" type="paratype">paratype</typeStatus>
from Locality 2.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEDFFDEFF5C47AE9382F936" pageId="3" pageNumber="138" type="etymology">
<paragraph id="556CAA05FFEDFFDEFF5C47AE9466FA61" blockId="3.[151,1437,837,1671]" box="[199,1058,1463,1488]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C47AE910AFA61" bold="true" box="[199,334,1463,1488]" pageId="3" pageNumber="138">Etymology.</emphasis>
The specific name refers to the locality—Viñales National Park.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C47C29382F936" blockId="3.[151,1437,837,1671]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C47C29126FA45" bold="true" box="[199,354,1499,1524]" pageId="3" pageNumber="138">Comparison.</emphasis>
The new species is similar to
<taxonomicName id="92D3D186FFEDFFDEFD4F47C294A9FA45" authority="Ewing, 1909" authorityName="Ewing" authorityYear="1909" box="[724,1261,1499,1524]" class="Arachnida" family="Phthiracaridae" genus="Hoplophthiracarus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="illinoisensis">
<emphasis id="67A77617FFEDFFDEFD4F47C29478FA42" box="[724,1084,1499,1523]" italics="true" pageId="3" pageNumber="138">Hoplophthiracarus illinoisensis</emphasis>
(
<bibRefCitation id="3142D7F4FFEDFFDEFBD447C294A2FA45" author="Ewing" box="[1103,1254,1499,1524]" pageId="3" pageNumber="138" refString="Ewing, H. E. (1909) New American Oribatoidea. Journal of the New York Entomological Society, 17 (3), 116 - 136." type="journal article" year="1909">Ewing, 1909</bibRefCitation>
)
</taxonomicName>
by the similar shape of sensilli and dorsal setae of notogaster and erect rostral setae but is distinguishable by the presence of very short median sigillar field (versus absence of field), rostral setae are situated near each other,
<emphasis id="67A77617FFEDFFDEFB3C443D94A5F98A" box="[1191,1249,1572,1595]" italics="true" pageId="3" pageNumber="138">in–in</emphasis>
/
<emphasis id="67A77617FFEDFFDEFB72443D9562F98A" box="[1257,1318,1572,1595]" italics="true" pageId="3" pageNumber="138">ro–ro</emphasis>
=3 (versus 2.2); setae
<emphasis id="67A77617FFEDFFDEFE8F44519165F9EE" box="[276,289,1608,1631]" italics="true" pageId="3" pageNumber="138">c</emphasis>
3,
<emphasis id="67A77617FFEDFFDEFEA344519116F9EE" box="[312,338,1608,1631]" italics="true" pageId="3" pageNumber="138">cp</emphasis>
,
<emphasis id="67A77617FFEDFFDEFEF94451913FF9EE" box="[354,379,1608,1631]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
3,
<emphasis id="67A77617FFEDFFDEFE09445191EFF9EE" box="[402,427,1608,1631]" italics="true" pageId="3" pageNumber="138">ps</emphasis>
4 shorter than others (versus similar in length), different genital formula 6:3 (versus 7:2), setae
<emphasis id="67A77617FFEDFFDEFF4D447690A8F937" box="[214,236,1647,1670]" italics="true" pageId="3" pageNumber="138">v’</emphasis>
on femora I and setae
<emphasis id="67A77617FFEDFFDEFE70447791B8F937" box="[491,508,1646,1670]" italics="true" pageId="3" pageNumber="138">l’</emphasis>
on genua IV are absent (versus present).
</paragraph>
</subSubSection>
</treatment>
</document>

218
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<treatment id="DD7A1B13FFEFFFDEFF0C446291D0FD36" ID-DOI="http://doi.org/10.5281/zenodo.6103400" ID-GBIF-Taxon="119645820" ID-Zenodo-Dep="6103400" LSID="urn:lsid:plazi:treatment:DD7A1B13FFEFFFDEFF0C446291D0FD36" httpUri="http://treatment.plazi.org/id/DD7A1B13FFEFFFDEFF0C446291D0FD36" lastPageId="3" lastPageNumber="138" pageId="1" pageNumber="136">
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<paragraph id="556CAA05FFEFFFDCFF0C446292E2F927" blockId="1.[151,678,1659,1719]" box="[151,678,1659,1686]" pageId="1" pageNumber="136">
<heading id="0E241D69FFEFFFDCFF0C446292E2F927" bold="true" box="[151,678,1659,1686]" fontSize="11" level="1" pageId="1" pageNumber="136" reason="1">
<emphasis id="67A77617FFEFFFDCFF0C446292E2F927" bold="true" box="[151,678,1659,1686]" pageId="1" pageNumber="136">
<taxonomicName id="92D3D186FFEFFFDCFF0C44629205F927" ID-CoL="7WMNY" authority="Niedbala" authorityName="Niedbala" box="[151,577,1659,1686]" class="Arachnida" family="Protoplophoridae" genus="Prototritia" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="triangularibus" status="sp. nov.">
<emphasis id="67A77617FFEFFFDCFF0C4462918EF924" bold="true" box="[151,458,1659,1685]" italics="true" pageId="1" pageNumber="136">Prototritia triangularibus</emphasis>
Niedbała
</taxonomicName>
<taxonomicNameLabel id="7C94CB6CFFEFFFDCFDD2446592E2F927" box="[585,678,1660,1686]" pageId="1" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="556CAA05FFEFFFDCFF0C4487915AF906" blockId="1.[151,678,1659,1719]" box="[151,286,1694,1719]" pageId="1" pageNumber="136">
(
<figureCitation id="CDE8B680FFEFFFDCFF0444879152F906" box="[159,278,1694,1719]" captionStart="FIGURE 1 A – E" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[210,1376,193,1702]" captionTargetId="figure@2.[210,1377,193,1702]" captionTargetPageId="2" captionText="FIGURE 1 A – E. Prototritia triangularibus Niedbała sp. nov. (holotype); measurements: prodorsum: length 172, width 131, height 76; notogaster: length 240, width 220, height 177. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, opisthosoma, posterior view; E, ventral side, partial view of posterior part." httpUri="https://zenodo.org/record/232409/files/figure.png" pageId="1" pageNumber="136">Fig. 1A–E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEFFFDEFF0C44FF936FFED6" lastPageId="3" lastPageNumber="138" pageId="1" pageNumber="136" type="description">
<paragraph id="556CAA05FFEFFFDCFF0C44FF924FF8C0" blockId="1.[151,1437,1766,1905]" pageId="1" pageNumber="136">
<emphasis id="67A77617FFEFFFDCFF0C44FF916DF94E" bold="true" box="[151,297,1766,1791]" pageId="1" pageNumber="136">Description.</emphasis>
<emphasis id="67A77617FFEFFFDCFEB444F19195F94E" box="[303,465,1768,1791]" italics="true" pageId="1" pageNumber="136">Measurements</emphasis>
of
<typeStatus id="8A6814A7FFEFFFDCFE6E44FF921FF94E" box="[501,603,1766,1791]" pageId="1" pageNumber="136" type="holotype">holotype</typeStatus>
. Prodorsum: length 172, width 131, height 76; setae:
<emphasis id="67A77617FFEFFFDCFB3744F19486F94E" box="[1196,1218,1768,1791]" italics="true" pageId="1" pageNumber="136">ss</emphasis>
30,
<emphasis id="67A77617FFEFFFDCFB6944F1954CF94E" box="[1266,1288,1768,1791]" italics="true" pageId="1" pageNumber="136">in</emphasis>
56,
<emphasis id="67A77617FFEFFFDCFAAC44FE950FF94E" box="[1335,1355,1767,1791]" italics="true" pageId="1" pageNumber="136">le</emphasis>
30,
<emphasis id="67A77617FFEFFFDCFAE044F195D0F94E" box="[1403,1428,1768,1791]" italics="true" pageId="1" pageNumber="136">ex</emphasis>
1 13,
<emphasis id="67A77617FFEFFFDCFF5A4517909EF894" box="[193,218,1806,1829]" italics="true" pageId="1" pageNumber="136">ex</emphasis>
2 7,
<emphasis id="67A77617FFEFFFDCFE9A4517915CF894" box="[257,280,1806,1829]" italics="true" pageId="1" pageNumber="136">ro</emphasis>
38; notogaster: length 240, width 220, height 177; setae:
<emphasis id="67A77617FFEFFFDCFC07451793ECF894" box="[924,936,1806,1829]" italics="true" pageId="1" pageNumber="136">c</emphasis>
1 38,
<emphasis id="67A77617FFEFFFDCFC7A451793B8F894" box="[993,1020,1806,1829]" italics="true" pageId="1" pageNumber="136">cp</emphasis>
33,
<emphasis id="67A77617FFEFFFDCFBB74517947CF894" box="[1068,1080,1806,1829]" italics="true" pageId="1" pageNumber="136">e</emphasis>
1 51,
<emphasis id="67A77617FFEFFFDCFBEA4517943DF897" box="[1137,1145,1806,1830]" italics="true" pageId="1" pageNumber="136">f</emphasis>
1 66; genitoaggenital plate 63×38, anoadanal plate 88×15. Measurements of
<typeStatus id="8A6814A7FFEFFFDCFD5B452C936BF8FC" box="[704,815,1845,1869]" pageId="1" pageNumber="136" type="paratype">paratypes</typeStatus>
. Prodorsum: length 156–162, height 63–66; notogaster: length 240–244, height 198–202.
</paragraph>
<footnote id="36C8B60BFFEFFFDCFF0C45B8921EF840" pageId="1" pageNumber="136">
<paragraph id="556CAA05FFEFFFDCFF0C45B8921EF840" blockId="1.[151,1436,1953,2033]" pageId="1" pageNumber="136">
1.
<taxonomicName id="92D3D186FFEFFFDCFF5745B892FAF806" authority="Balogh, 1958" authorityName="Balogh" authorityYear="1958" box="[204,702,1953,1975]" class="Arachnida" family="Phthiracaridae" genus="Atropacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="andrei" subGenus="Hoplophorella">
<emphasis id="67A77617FFEFFFDCFF5745B89107F807" box="[204,323,1953,1974]" italics="true" pageId="1" pageNumber="136">Atropacarus</emphasis>
(
<emphasis id="67A77617FFEFFFDCFECA45B89199F807" box="[337,477,1953,1974]" italics="true" pageId="1" pageNumber="136">Hoplophorella</emphasis>
)
<emphasis id="67A77617FFEFFFDCFE7045B8926CF807" box="[491,552,1953,1974]" italics="true" pageId="1" pageNumber="136">andrei</emphasis>
(Balogh, 1958)
</taxonomicName>
was wrongly synonymized with
<taxonomicName id="92D3D186FFEFFFDCFC6245B89566F806" authority="Berlese, 1913" authorityName="Berlese" authorityYear="1913" box="[1017,1314,1953,1975]" class="Arachnida" family="Phthiracaridae" genus="Atropacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="vitrinus" subGenus="Hoplophorella">
<emphasis id="67A77617FFEFFFDCFC6245B8944CF807" box="[1017,1032,1953,1974]" italics="true" pageId="1" pageNumber="136">A</emphasis>
. (
<emphasis id="67A77617FFEFFFDCFB8745B89469F807" box="[1052,1069,1953,1974]" italics="true" pageId="1" pageNumber="136">H</emphasis>
.)
<emphasis id="67A77617FFEFFFDCFBDB45B894CDF807" box="[1088,1161,1953,1974]" italics="true" pageId="1" pageNumber="136">vitrinus</emphasis>
(Berlese, 1913)
</taxonomicName>
by Niedbała (2000). Liu &amp; Oconnor (2015) showed differences between the two species and essential difference is presence of median band in dorsal part of notogaster in
<taxonomicName id="92D3D186FFEFFFDCFE5445C59210F840" box="[463,596,2011,2033]" class="Arachnida" family="Phthiracaridae" genus="Atropacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="andrei" subGenus="Hoplophorella">
<emphasis id="67A77617FFEFFFDCFE5445C5919AF840" box="[463,478,2012,2033]" italics="true" pageId="1" pageNumber="136">A</emphasis>
. (
<emphasis id="67A77617FFEFFFDCFE6945C59247F840" box="[498,515,2012,2033]" italics="true" pageId="1" pageNumber="136">H</emphasis>
.)
<emphasis id="67A77617FFEFFFDCFD8C45C59210F840" box="[535,596,2012,2033]" italics="true" pageId="1" pageNumber="136">andrei</emphasis>
</taxonomicName>
.
</paragraph>
</footnote>
<caption id="01ACFA8DFFECFFDFFF0C44A594FDF8BE" httpUri="https://zenodo.org/record/232409/files/figure.png" pageId="2" pageNumber="137" targetBox="[210,1376,193,1702]" targetPageId="2">
<paragraph id="556CAA05FFECFFDFFF0C44A594FDF8BE" blockId="2.[151,1435,1724,1807]" pageId="2" pageNumber="137">
<emphasis id="67A77617FFECFFDFFF0C44A5917AF960" bold="true" box="[151,318,1724,1746]" pageId="2" pageNumber="137">FIGURE 1A–E</emphasis>
.
<taxonomicName id="92D3D186FFECFFDFFED644A492F3F963" authority="Niedbala" authorityName="Niedbala" box="[333,695,1724,1746]" class="Arachnida" family="Protoplophoridae" genus="Prototritia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="triangularibus" status="sp. nov.">
<emphasis id="67A77617FFECFFDFFED644A49210F963" box="[333,596,1724,1746]" italics="true" pageId="2" pageNumber="137">Prototritia triangularibus</emphasis>
Niedbała
</taxonomicName>
<emphasis id="67A77617FFECFFDFFD2444A59355F960" bold="true" box="[703,785,1724,1745]" pageId="2" pageNumber="137">
<taxonomicNameLabel id="7C94CB6CFFECFFDFFD2444A59355F960" box="[703,785,1724,1745]" pageId="2" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); measurements: prodorsum: length 172, width 131, height 76; notogaster: length 240, width 220, height 177. A, prodorsum, dorsal view; B, prodorsum, lateral view; C, opisthosoma, lateral view; D, opisthosoma, posterior view; E, ventral side, partial view of posterior part.
</paragraph>
</caption>
<paragraph id="556CAA05FFECFFDFFF5C452E92A5F8FF" blockId="2.[151,1436,1846,2019]" box="[199,737,1846,1871]" pageId="2" pageNumber="137">
<emphasis id="67A77617FFECFFDFFF5C452E9101F8FF" box="[199,325,1847,1870]" italics="true" pageId="2" pageNumber="137">Integument</emphasis>
. Colour yellowish. Cuticle punctate.
</paragraph>
<paragraph id="556CAA05FFECFFDFFF5C4543953CF86F" blockId="2.[151,1436,1846,2019]" pageId="2" pageNumber="137">
<emphasis id="67A77617FFECFFDFFF5C45439100F8C3" box="[199,324,1882,1906]" italics="true" pageId="2" pageNumber="137">Prodorsum</emphasis>
with large median triangular protuberance. Lateral carinae well discernible, long. Sensilli with short stalk and spindle-shaped, flattened head, pointed distally and covered with slender cilia. Interlamellar setae elongated, fusiform, covered with long spines. Lamellar and exobothridial setae needleform, smooth. Rostral setae long, needleform, directed inwards and covered with some long cilia. Relation of lengths:
<emphasis id="67A77617FFECFFDFFB1145DE94E4F86F" box="[1162,1184,1991,2014]" italics="true" pageId="2" pageNumber="137">in</emphasis>
&gt;
<emphasis id="67A77617FFECFFDFFB3445DE9482F86F" box="[1199,1222,1991,2014]" italics="true" pageId="2" pageNumber="137">ro</emphasis>
&gt;
<emphasis id="67A77617FFECFFDFFB4C45DE94A9F86F" box="[1239,1261,1991,2014]" italics="true" pageId="2" pageNumber="137">ss</emphasis>
=
<emphasis id="67A77617FFECFFDFFB6745DF9554F86F" box="[1276,1296,1990,2014]" italics="true" pageId="2" pageNumber="137">le</emphasis>
&gt;
<emphasis id="67A77617FFECFFDFFABB45DE957DF86F" box="[1312,1337,1991,2014]" italics="true" pageId="2" pageNumber="137">ex</emphasis>
1&gt;
<emphasis id="67A77617FFECFFDFFACA45DE952EF86F" box="[1361,1386,1991,2014]" italics="true" pageId="2" pageNumber="137">ex</emphasis>
2.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C42809317FF65" blockId="3.[151,1436,151,647]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C42809106FF01" box="[199,322,153,176]" italics="true" pageId="3" pageNumber="138">Notogaster</emphasis>
. “Notogastral” setae needleform, attenuate; setae of rows
<emphasis id="67A77617FFEDFFDEFC544280939FFF01" box="[975,987,153,176]" italics="true" pageId="3" pageNumber="138">e</emphasis>
and
<emphasis id="67A77617FFEDFFDEFB894281945EFF01" box="[1042,1050,152,176]" italics="true" pageId="3" pageNumber="138">f</emphasis>
longer than setae of rows
<emphasis id="67A77617FFEDFFDEFAD84280950BFF01" box="[1347,1359,153,176]" italics="true" pageId="3" pageNumber="138">c</emphasis>
and
<emphasis id="67A77617FFEDFFDEFA1D428195D0FF01" box="[1414,1428,152,176]" italics="true" pageId="3" pageNumber="138">h</emphasis>
; setae
<emphasis id="67A77617FFEDFFDEFF4D42A490A6FF65" box="[214,226,189,212]" italics="true" pageId="3" pageNumber="138">c</emphasis>
3 more remote from anterior margin than setae
<emphasis id="67A77617FFEDFFDEFD7642A492BDFF65" box="[749,761,189,212]" italics="true" pageId="3" pageNumber="138">c</emphasis>
1 and
<emphasis id="67A77617FFEDFFDEFCA342A49300FF65" box="[824,836,189,212]" italics="true" pageId="3" pageNumber="138">c</emphasis>
2.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C42FA9444FEF2" blockId="3.[151,1436,151,647]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C42FA912EFF4B" box="[199,362,227,251]" italics="true" pageId="3" pageNumber="138">Ventral region</emphasis>
. Genitoaggenital plates with distinct longitudinal furrows, separating 5 pairs of genital setae from 3 pairs of aggenital setae. Anoadanal plates narrow, with 8 pairs of setae (only right plate of
<typeStatus id="8A6814A7FFEDFFDEFB70431F9509FEAE" box="[1259,1357,262,287]" pageId="3" pageNumber="138" type="holotype">holotype</typeStatus>
with 7 setae) arranged in longitudinal row. All setae of ventral region relatively long.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C4349936FFED6" blockId="3.[151,1436,151,647]" box="[199,811,334,359]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C434990B8FED6" box="[199,252,336,359]" italics="true" pageId="3" pageNumber="138">Legs</emphasis>
. Tarsi heterotridactylous, claws shorter than tarsi.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEDFFDEFF5C436B93F4FE3A" box="[199,944,370,395]" pageId="3" pageNumber="138" type="materials_examined">
<paragraph id="556CAA05FFEDFFDEFF5C436B93F4FE3A" blockId="3.[151,1436,151,647]" box="[199,944,370,395]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C436B91F4FE3A" bold="true" box="[199,432,370,395]" pageId="3" pageNumber="138">Material examined.</emphasis>
<typeStatus id="8A6814A7FFEDFFDEFE2C436B925BFE3A" box="[439,543,370,395]" pageId="3" pageNumber="138" type="holotype">Holotype</typeStatus>
and two
<typeStatus id="8A6814A7FFEDFFDEFD1C436A92B6FE3A" box="[647,754,371,395]" pageId="3" pageNumber="138" type="paratype">paratypes</typeStatus>
from Locality 2.
</paragraph>
</subSubSection>
<subSubSection id="1DC9F98EFFEDFFDEFF5C438F91D0FD36" pageId="3" pageNumber="138" type="etymology">
<paragraph id="556CAA05FFEDFFDEFF5C438F91E5FE62" blockId="3.[151,1436,151,647]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C438F910AFE1E" bold="true" box="[199,334,406,431]" pageId="3" pageNumber="138">Etymology.</emphasis>
The specific name
<taxonomicName id="92D3D186FFEDFFDEFDAF438E9292FE1E" box="[564,726,407,431]" class="Arachnida" family="Protoplophoridae" genus="Prototritia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="triangularibus">
<emphasis id="67A77617FFEDFFDEFDAF438E9292FE1E" box="[564,726,407,431]" italics="true" pageId="3" pageNumber="138">triangularibus</emphasis>
</taxonomicName>
is Latin for “triangular” and refers to the triangular shape of prodorsal protuberance.
</paragraph>
<paragraph id="556CAA05FFEDFFDEFF5C43C791D0FD36" blockId="3.[151,1436,151,647]" pageId="3" pageNumber="138">
<emphasis id="67A77617FFEDFFDEFF5C43C79125FE46" bold="true" box="[199,353,478,503]" pageId="3" pageNumber="138">Comparison.</emphasis>
The new species is morphologically similar to
<taxonomicName id="92D3D186FFEDFFDEFC2C43F99559FE46" authority="Mahunka" authorityName="Mahunka" box="[951,1309,478,503]" class="Arachnida" family="Protoplophoridae" genus="Prototritia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="palaciosi">
<emphasis id="67A77617FFEDFFDEFC2C43F994E0FE46" box="[951,1188,479,503]" italics="true" pageId="3" pageNumber="138">Prototritia palaciosi</emphasis>
Mahunka
</taxonomicName>
&amp; Mejia- Recamier, 1998 from
<collectingCountry id="2DC4EA95FFEDFFDEFE0B401B91A1FDAA" box="[400,485,514,539]" name="Mexico" pageId="3" pageNumber="138">Mexico</collectingCountry>
by the shape of rostral setae and number and arrangement of “notogastral”, genital and anal setae. It is distinguishable by the presence of prodorsal median triangular protuberance, different shape of sensilli, interlamellar, lamellar and exobothridial setae and especially by presence of longitudinal furrows on genitoaggenital plates.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America</mods:title>
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<mods:namePart>Nestor-Arriola, Jorge Ismael</mods:namePart>
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<mods:namePart>Toledo-Hernandez, Victor Hugo</mods:namePart>
<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
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<mods:namePart>Solis, Angel</mods:namePart>
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<mods:role>
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<mods:namePart>Gonzalez, Guillermo</mods:namePart>
<mods:affiliation>La Reina, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vetrovec, Jaroslav</mods:namePart>
<mods:affiliation>Buzulucka, Hradec Kralove, Ceska Republika</mods:affiliation>
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<treatment ID-GBIF-Taxon="180585848" LSID="urn:lsid:plazi:treatment:DD7A5258E0C15EBB8E87A67545FD179A" httpUri="http://treatment.plazi.org/id/DD7A5258E0C15EBB8E87A67545FD179A" lastPageNumber="157" pageId="0" pageNumber="157">
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<taxonomicName LSID="DD7A5258-E0C1-5EBB-8E87-A67545FD179A" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha erythrura" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="erythrura">Brachiacantha erythrura Mulsant</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="reference_group">
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha erythrura" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="erythrura">Brachyacantha erythrura</taxonomicName>
Mulsant, 1850: 530.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8975" author="Crotch, GR" journalOrPublisher="Janson EW, London" pageId="0" pageNumber="157" refId="B9" refString="Crotch, GR, 1874. A Revision of the Coleopterous Family Coccinellidae. Janson EW, London, DOI: https://doi.org/10.5962/bhl.title.8975" title="A Revision of the Coleopterous Family Coccinellidae." url="https://doi.org/10.5962/bhl.title.8975" year="1874">Crotch 1874</bibRefCitation>
: 211.
<bibRefCitation author="Gorham, HS" journalOrPublisher="Biologia Centrali-Americana, Insecta, Coleoptera" pageId="0" pageNumber="157" pagination="177 - 208" refId="B17" refString="Gorham, HS, 1894. Coccinellidae. Biologia Centrali-Americana, Insecta, Coleoptera 7: 177 - 208" title="Coccinellidae." volume="7" year="1894">Gorham 1894</bibRefCitation>
: 187.
<bibRefCitation author="Leng, CW" journalOrPublisher="Bulletin American Museum of Natural History" pageId="0" pageNumber="157" pagination="279 - 333" refId="B19" refString="Leng, CW, 1911. The species of Brachyacantha of North and South America. Bulletin American Museum of Natural History 30: 279 - 333" title="The species of Brachyacantha of North and South America." volume="30" year="1911">Leng 1911</bibRefCitation>
: 302.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4701.6.3" author="Nestor-Arriola, JI" journalOrPublisher="Zootaxa" pageId="0" pageNumber="157" pagination="537 - 552" refId="B25" refString="Nestor-Arriola, JI, Toledo-Hernandez, VH, 2019. Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae). Zootaxa 4701 (6): 537 - 552, DOI: https://doi.org/10.11646/zootaxa.4701.6.3" title="Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae)." url="https://doi.org/10.11646/zootaxa.4701.6.3" volume="4701" year="2019">
Nestor-Arriola and
<normalizedToken originalValue="Toledo-Hernández">Toledo-Hernandez</normalizedToken>
2019
</bibRefCitation>
: 545.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="materials_examined">
<paragraph pageId="0" pageNumber="157">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="157">
Material listed in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4701.6.3" author="Nestor-Arriola, JI" journalOrPublisher="Zootaxa" pageId="0" pageNumber="157" pagination="537 - 552" refId="B25" refString="Nestor-Arriola, JI, Toledo-Hernandez, VH, 2019. Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae). Zootaxa 4701 (6): 537 - 552, DOI: https://doi.org/10.11646/zootaxa.4701.6.3" title="Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae)." url="https://doi.org/10.11646/zootaxa.4701.6.3" volume="4701" year="2019">
Nestor-Arriola and
<normalizedToken originalValue="Toledo-Hernández">Toledo-Hernandez</normalizedToken>
(2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="diagnosis">
<paragraph pageId="0" pageNumber="157">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="157">
Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="3.9" metricValueMin="3.8" unit="mm" value="3.8499999999999996" valueMax="3.9" valueMin="3.8">3.8-3.9 mm</quantity>
, width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8-2.9 mm</quantity>
. Dorsally black, head and posterior margin of the elytron red, orange or yellow. Ventral surface light brown to orange. Third ventrite of the male with small cusps. Penis guide of the male genitalia asymmetrical, without apical hook.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="distribution">
<paragraph pageId="0" pageNumber="157">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="157">
<normalizedToken originalValue="México">
<collectingCountry name="Mexico">Mexico</collectingCountry>
</normalizedToken>
and Central America.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="discussion">
<paragraph pageId="0" pageNumber="157">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="157">
This species can be confused with some specimens of
<taxonomicName lsidName="B. quadrillum" pageId="0" pageNumber="157" rank="species" species="quadrillum">
<emphasis italics="true" pageId="0" pageNumber="157">B. quadrillum</emphasis>
</taxonomicName>
LeConte, but the third ventrite cusps of
<taxonomicName lsidName="B. erythrura" pageId="0" pageNumber="157" rank="species" species="erythrura">
<emphasis italics="true" pageId="0" pageNumber="157">B. erythrura</emphasis>
</taxonomicName>
are much smaller and the male genitalia lack the apical hook.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="10.11646/phytotaxa.589.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="7755059" checkinTime="1679382675456" checkinUser="plazi" docAuthor="Leeratiwong, Charan, Karapan, Sunate, Satthaphorn, Jiratthi &amp; Johnson, David M." docDate="2023" docId="DD7A87FBFFD2FF89FF2714EEFB6DC15B" docLanguage="en" docName="phytotaxa.589.1.7.pdf" docOrigin="Phytotaxa 589 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.589.1.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Friesodielsia betongensis Leerat. 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="79" masterDocId="2143FF83FFD2FF8FFFAF135BFF87C406" masterDocTitle="Two new species of Friesodielsia (Annonaceae) from Peninsular Thailand" masterLastPageNumber="82" masterPageNumber="73" pageNumber="73" updateTime="1679427965726" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Two new species of Friesodielsia (Annonaceae) from Peninsular Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Leeratiwong, Charan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0752-4275</mods:nameIdentifier>
<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand &amp; charan. leeratiwong @ gmail. com; https: // orcid. org / 0000 - 0002 - 0752 - 4275</mods:affiliation>
<mods:nameIdentifier type="email">charan.leeratiwong@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karapan, Sunate</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2284-3039</mods:nameIdentifier>
<mods:affiliation>Hala-Bala Wildlife Research Station, Hala-Bala Wildlife Sanctuary, Waeng, Narathiwat, 96160, Thailand &amp; karapann @ gmail. com; https: // orcid. org / 0000 - 0002 - 2284 - 3039</mods:affiliation>
<mods:nameIdentifier type="email">karapann@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Satthaphorn, Jiratthi</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7994-9056</mods:nameIdentifier>
<mods:affiliation>Department of Biology School of Science, Walailak University, Thasala, Nakhon Si Thammarat, 80161, Thailand &amp; jiratthi. sa @ wu. ac. th; https: // orcid. org / 0000 - 0001 - 7994 - 9056</mods:affiliation>
<mods:nameIdentifier type="email">jiratthi.sa@wu.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johnson, David M.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2896-7419</mods:nameIdentifier>
<mods:affiliation>Department of Biological Science, Ohio Wesleyan University, Delaware, Ohio 43015, USA &amp; dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
<mods:nameIdentifier type="email">dmjohnso@owu.edu</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>589</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>73</mods:start>
<mods:end>82</mods:end>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.589.1.7</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.589.1.7</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:DD7A87FBFFD2FF89FF2714EEFB6DC15B" httpUri="http://treatment.plazi.org/id/DD7A87FBFFD2FF89FF2714EEFB6DC15B" lastPageId="6" lastPageNumber="79" pageId="0" pageNumber="73">
<subSubSection box="[136,663,1973,1995]" pageId="0" pageNumber="73" type="nomenclature">
<paragraph blockId="0.[136,1452,1973,2064]" box="[136,663,1973,1995]" pageId="0" pageNumber="73">
<taxonomicName authority="Leerat." authorityName="Leerat." authorityYear="2023" box="[136,459,1973,1995]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="0" pageNumber="73" phylum="Tracheophyta" rank="species" species="betongensis" status="sp. nov.">
<emphasis bold="true" box="[136,386,1973,1995]" italics="true" pageId="0" pageNumber="73">Friesodielsia betongensis</emphasis>
Leerat.
</taxonomicName>
,
<taxonomicNameLabel box="[471,545,1973,1995]" pageId="0" pageNumber="73" rank="species">
<emphasis box="[471,540,1973,1995]" italics="true" pageId="0" pageNumber="73">sp. nov</emphasis>
.
</taxonomicNameLabel>
<emphasis bold="true" box="[551,559,1973,1995]" pageId="0" pageNumber="73">(</emphasis>
Figs 1,3,5)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="materials_examined">
<paragraph blockId="0.[136,1452,1973,2064]" pageId="0" pageNumber="73">
<materialsCitation collectingDate="2022-05-22" country="Thailand" county="Type" elevation="1100" location="Hala-Bala Wildlife Sanctuary" municipality="Betong District" pageId="0" pageNumber="73" specimenCount="1" stateProvince="Yala" typeStatus="holotype">
<collectingCounty box="[136,185,2008,2029]" pageId="0" pageNumber="73">Type</collectingCounty>
:—
<collectingCountry box="[213,339,2008,2029]" name="Thailand" pageId="0" pageNumber="73">THAILAND</collectingCountry>
.
<collectingRegion box="[353,400,2008,2030]" country="Thailand" name="Yala" pageId="0" pageNumber="73">Yala</collectingRegion>
:
<collectingMunicipality box="[412,566,2008,2030]" pageId="0" pageNumber="73">Betong District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DD7A87FBFFD2FF89FF2714EEFB6DC15B:500C6036FFD2FF8FFDEC1483FCE0C3EB" box="[579,871,2008,2030]" country="Thailand" county="Type" municipality="Betong District" name="Hala-Bala Wildlife Sanctuary" pageId="0" pageNumber="73" stateProvince="Yala">Hala-Bala Wildlife Sanctuary</location>
,
<quantity box="[884,1020,2008,2029]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="0" pageNumber="73" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation box="[884,1020,2008,2029]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="0" pageNumber="73" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000–1200 m</elevation>
</quantity>
elev.,
<date box="[1090,1227,2008,2029]" pageId="0" pageNumber="73" value="2022-05-22">
<collectingDate box="[1090,1227,2008,2029]" pageId="0" pageNumber="73" value="2022-05-22">22 May 2022</collectingDate>
</date>
,
<emphasis box="[1240,1452,2008,2029]" italics="true" pageId="0" pageNumber="73">Leeratiwong 22-1854</emphasis>
(
<typeStatus box="[196,285,2042,2064]" pageId="0" pageNumber="73">holotype</typeStatus>
: BKF; isotype: PSU).
</materialsCitation>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7755061" ID-Zenodo-Dep="7755061" httpUri="https://zenodo.org/record/7755061/files/figure.png" pageId="1" pageNumber="74" startId="1.[136,229,1935,1957]" targetBox="[191,1390,200,1900]" targetPageId="1">
<paragraph blockId="1.[136,1452,1935,2030]" pageId="1" pageNumber="74">
<emphasis bold="true" box="[136,255,1935,1957]" pageId="1" pageNumber="74">FIGURE 1.</emphasis>
<taxonomicName box="[262,509,1936,1957]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="74" phylum="Tracheophyta" rank="species" species="betongensis">
<emphasis box="[262,509,1936,1957]" italics="true" pageId="1" pageNumber="74">Friesodielsia betongensis</emphasis>
</taxonomicName>
. A. Flowering branch. B. Flower, C. Sepal, abaxial view. D. Sepal, adaxial view. E. Outer petal, abaxial view. F. Outer petal, adaxial view. G. Inner petal, abaxial view. H. Inner petal, adaxial view. I. Stamen, abaxial view. J. Stamen, adaxial view. K., L. Carpels. M. Fruiting branch. Drawn by A. Somphrom.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7755063" ID-Zenodo-Dep="7755063" httpUri="https://zenodo.org/record/7755063/files/figure.png" pageId="2" pageNumber="75" startId="2.[136,229,1953,1975]" targetBox="[192,1357,203,1927]" targetPageId="2">
<paragraph blockId="2.[136,1452,1953,2047]" pageId="2" pageNumber="75">
<emphasis bold="true" box="[136,255,1953,1975]" pageId="2" pageNumber="75">FIGURE 2.</emphasis>
<taxonomicName box="[261,524,1953,1974]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="2" pageNumber="75" phylum="Tracheophyta" rank="species" species="chalermglinii">
<emphasis box="[261,524,1953,1974]" italics="true" pageId="2" pageNumber="75">Friesodielsia chalermglinii</emphasis>
</taxonomicName>
. A. Flowering branch. B. Flowers, side view. C. Bract, abaxial view. D. Bract, adaxial view. E. Sepal, abaxial view. F. Sepal, adaxial view. G. Outer petal, abaxial view. H. Outer petal, adaxial view. I. Inner petal, abaxial view. J. Inner petal, adaxial view. K. Stamen, abaxial view. L. Stamen, adaxial view. M., N. Carpels. Drawn by A. Somphrom.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7755065" ID-Zenodo-Dep="7755065" httpUri="https://zenodo.org/record/7755065/files/figure.png" pageId="3" pageNumber="76" startId="3.[136,229,1953,1975]" targetBox="[208,1379,190,1929]" targetPageId="3">
<paragraph blockId="3.[136,1452,1953,2011]" pageId="3" pageNumber="76">
<emphasis bold="true" box="[136,253,1953,1975]" pageId="3" pageNumber="76">FIGURE 3.</emphasis>
<taxonomicName box="[259,504,1953,1974]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="76" phylum="Tracheophyta" rank="species" species="betongensis">
<emphasis box="[259,504,1953,1974]" italics="true" pageId="3" pageNumber="76">Friesodielsia betongensis</emphasis>
</taxonomicName>
. A. Fruiting branch. B. Young flower, side view. C., D. Mature flower, side view. F. Mature flower, apical view. F. Nearly mature fruits. Photographs by C. Leeratiwong
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7755067" ID-Zenodo-Dep="7755067" httpUri="https://zenodo.org/record/7755067/files/figure.png" pageId="4" pageNumber="77" startId="4.[136,229,1953,1975]" targetBox="[209,1378,190,1927]" targetPageId="4">
<paragraph blockId="4.[136,1451,1953,2011]" pageId="4" pageNumber="77">
<emphasis bold="true" box="[136,253,1953,1975]" pageId="4" pageNumber="77">FIGURE 4.</emphasis>
<taxonomicName box="[259,520,1953,1974]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="77" phylum="Tracheophyta" rank="species" species="chalermglinii">
<emphasis box="[259,520,1953,1974]" italics="true" pageId="4" pageNumber="77">Friesodielsia chalermglinii</emphasis>
</taxonomicName>
. A. Flowering branch. B. Young flower, side view. C. Nearly mature flower, side view. D. Flower showing sepals and a bract, basal view. E., F. Mature flowers, side view. Photographs by C. Leeratiwong.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7755069" ID-Zenodo-Dep="7755069" httpUri="https://zenodo.org/record/7755069/files/figure.png" pageId="5" pageNumber="78" startId="5.[136,229,1918,1940]" targetBox="[288,1300,190,1894]" targetPageId="5">
<paragraph blockId="5.[136,1452,1918,1976]" pageId="5" pageNumber="78">
<emphasis bold="true" box="[136,255,1918,1940]" pageId="5" pageNumber="78">FIGURE 5.</emphasis>
Distribution of
<taxonomicName box="[415,662,1918,1939]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="betongensis">
<emphasis box="[415,662,1918,1939]" italics="true" pageId="5" pageNumber="78">Friesodielsia betongensis</emphasis>
</taxonomicName>
(●) and
<taxonomicName box="[751,905,1918,1940]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="chalermglinii">
<emphasis box="[751,905,1918,1940]" italics="true" pageId="5" pageNumber="78">F. chalermglinii</emphasis>
</taxonomicName>
(▲) in Thailand. The map and marked locations were constructed in Rstudio v.3.6.3 (R
<bibRefCitation author="Core Team" box="[458,626,1954,1976]" pageId="5" pageNumber="78" refId="ref4346" refString="R Core Team. (2020) R: a language and environment for statistical computing. Vienna: R. Foundation for Statistical Computing. Available at: https: // www. R- project. org / (Accessed 20 November 2022)." type="url" year="2020">Core Team, 2020</bibRefCitation>
) using ggplots package (
<bibRefCitation author="Wickham, H." box="[863,1018,1954,1976]" pageId="5" pageNumber="78" refId="ref4581" refString="Wickham, H. (2016) ggplot 2: elegant graphics for data analysis. Springer-Verlag New York. Available at: https: // https: // ggplot 2. tidyverse. org (Accessed 20 November 2022). https: // doi. org / 10.1007 / 978 - 3 - 319 - 24277 - 4" type="url" year="2016">Wickham, 2016</bibRefCitation>
) and its dependencies.
</paragraph>
</caption>
<subSubSection lastPageId="6" lastPageNumber="79" pageId="5" pageNumber="78" type="description">
<paragraph blockId="5.[136,1451,2023,2085]" lastBlockId="6.[136,1452,158,1373]" lastPageId="6" lastPageNumber="79" pageId="5" pageNumber="78">
The new species is similar to
<taxonomicName authorityName="Steenis" authorityYear="1964" baseAuthorityName="Sinclair" baseAuthorityYear="1955" box="[477,705,2023,2049]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="78" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis box="[477,705,2023,2049]" italics="true" pageId="5" pageNumber="78">Friesodielsia alpina</emphasis>
</taxonomicName>
but differs in having hairy twigs (vs. glabrous), leaves with hairs covering the abaxial surface (vs with hairs only midrib), glabrous pedicels (vs. hairy), shorter bracts (1.5–3.0 mm vs. ca. 4.0 mm), sepals with an obtuse (vs. acute) apex, longer inner petals (10.0–17.0 mm vs 7.0 mm) that are greater than half the length of the outer petals (vs. less than half the length of the outer petals) and hairy monocarps (vs. glabrous) with longer stipes (5.0–7.0 mm vs. ca. 4.0 mm).
</paragraph>
<paragraph blockId="6.[136,1452,158,1373]" pageId="6" pageNumber="79">
Woody climbers and scandent shrubs up to
<quantity box="[696,756,266,293]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="79" unit="m" value="10.0">10 m</quantity>
high. Twigs with dense, brown erect pubescence, becoming subglabrate. Leaves not glaucous below, narrowly oblanceolate, elliptic-oblanceolate or lanceolate, blades 7–15 Í
<quantity box="[136,223,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="79" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
, base rounded, apex acute to acuminate, acumen
<quantity box="[784,878,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="6" pageNumber="79" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 mm</quantity>
long, glabrate above, with denser hairs on midrib, moderately to sparsely appressed-pubescent below, secondary veins 9–12 per side; petioles
<quantity box="[1184,1279,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="79" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
long, flattened above with dense, brown pubescence. Inflorescences supra-axillary to leaf-opposed or terminal, 1(–2)-flowered, pedicel
<quantity box="[136,259,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="79" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
long, densely pubescent, with a lanceolate bract 1.5–3.0 mm long, attached at or just below the pedicel midpoint. Sepals 3, green, valvate, basally connate, coriaceous, broadly ovate, 2.5–4.5 Í 2.5–5.0 mm, apex obtuse, venation not evident, densely pubescent outside, glabrous except hairy just inside the edge. Petals in two whorls of 3, yellowish cream with pale green base, valvate, free; outer petals appressed in bud, spreading widely at maturity, lanceolate to ovate-lanceolate, 15.0–23.0 Í 4.0–
<quantity box="[687,791,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="79" unit="mm" value="10.5">10.5 mm</quantity>
, triquetrous near apex, obtuse, slightly concave on inner base, finely pubescent outside, sparsely hairy and glabrous at base inside; inner petals erect and coherent at margins for about half of their length apically, lanceolate, 10.0–17.0 Í
<quantity box="[851,991,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.5" metricValueMin="3.5" pageId="6" pageNumber="79" unit="mm" value="5.0" valueMax="6.5" valueMin="3.5">3.5–6.5 mm</quantity>
, obtuse to acute, sparsely hairy outside, glabrous inside. Stamens clavate,
<quantity box="[526,663,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="6" pageNumber="79" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2–1.6 mm</quantity>
long, anther dehiscence extrorse, apex of anther connective convex. Carpels 40–60, oblong, 1.2–2.0 mm long, densely pubescent, stigmas falcate-capitate, cleft down the side. Fruits of 1–3 monocarps, pedicels
<quantity box="[458,580,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="6" pageNumber="79" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
long, with persistent bracts and deciduous sepals. Monocarps green (mature not seen), ellipsoid to ellipsoid-oblongoid, 12–15 Í
<quantity box="[729,826,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="6" pageNumber="79" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
, smooth, pubescent, apex apiculate,
<quantity box="[1248,1385,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="6" pageNumber="79" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
long, stipe 5.0–7.0 Í 2.0–
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. Seeds 1 per monocarp, attached basally, broadly ellipsoid, 10–12 Í
<quantity box="[1220,1316,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="79" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
, endosperm rumination lamellate.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="79" type="distribution">
<paragraph blockId="6.[136,1452,158,1373]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,518,914,940]" pageId="6" pageNumber="79">Distribution and ecology:—</emphasis>
<collectingRegion box="[519,677,914,940]" country="Thailand" name="Yala" pageId="6" pageNumber="79">Yala Province</collectingRegion>
, Peninsular
<collectingCountry box="[811,911,914,940]" name="Thailand" pageId="6" pageNumber="79">Thailand</collectingCountry>
(
<figureCitation box="[925,992,914,941]" captionStart="FIGURE 5" captionStartId="5.[136,229,1918,1940]" captionTargetBox="[288,1300,190,1894]" captionTargetId="figure-19@5.[288,1300,190,1894]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of Friesodielsia betongensis (●) and F. chalermglinii (▲) in Thailand. The map and marked locations were constructed in Rstudio v.3.6.3 (R Core Team, 2020) using ggplots package (Wickham, 2016) and its dependencies." figureDoi="http://doi.org/10.5281/zenodo.7755069" httpUri="https://zenodo.org/record/7755069/files/figure.png" pageId="6" pageNumber="79">Fig. 5</figureCitation>
), on hill edges in evergreen forest,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="79" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000– 1200 m</quantity>
elevation.
</paragraph>
<paragraph blockId="6.[136,1452,158,1373]" box="[189,698,986,1013]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,351,986,1012]" pageId="6" pageNumber="79">Phenology:—</emphasis>
Flowering and fruiting in May.
</paragraph>
</subSubSection>
<subSubSection box="[189,686,1022,1048]" pageId="6" pageNumber="79" type="vernacular_names">
<paragraph blockId="6.[136,1452,158,1373]" box="[189,686,1022,1048]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,366,1022,1048]" pageId="6" pageNumber="79">Local name:—</emphasis>
<emphasis box="[366,601,1022,1048]" italics="true" pageId="6" pageNumber="79">Bu nga soeng betong</emphasis>
(Thai).
</paragraph>
</subSubSection>
<subSubSection box="[189,814,1058,1085]" pageId="6" pageNumber="79" type="etymology">
<paragraph blockId="6.[136,1452,158,1373]" box="[189,814,1058,1085]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,356,1058,1084]" pageId="6" pageNumber="79">Etymology:—</emphasis>
From the Betong District,
<collectingRegion box="[650,810,1058,1084]" country="Thailand" name="Yala" pageId="6" pageNumber="79">Yala Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="79" type="conservation">
<paragraph blockId="6.[136,1452,158,1373]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,467,1094,1120]" pageId="6" pageNumber="79">Conservation status:—</emphasis>
This species is known only from gatherings in 2022, when it was collected from the narrow range of hills in evergreen forest, Hala-Bala Wildlife Sanctuary. The forest is well protected, however, and the larger part of it is still unexplored. To clarify the number of individuals/populations and the full distribution, further field studies are needed. Therefore, we have preliminarily assessed the species as data deficient (DD, IUCN Red List Criteria 2019).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="79" type="discussion">
<paragraph blockId="6.[136,1452,158,1373]" pageId="6" pageNumber="79">
<emphasis bold="true" box="[189,294,1274,1300]" pageId="6" pageNumber="79">Notes:—</emphasis>
<taxonomicName box="[294,582,1274,1300]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="betongensis">
<emphasis box="[294,582,1274,1300]" italics="true" pageId="6" pageNumber="79">Friesodielsia betongensis</emphasis>
</taxonomicName>
is characterised by its densely erect-hairy stems, long pedicels (
<quantity box="[1312,1440,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="79" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
), flowers with pale green petal bases, triquetrous outer petals and inner petals longer than half the length of outer petals. This species is one of only two Thai
<taxonomicName box="[548,783,1346,1372]" pageId="6" pageNumber="79">
<emphasis box="[548,696,1346,1372]" italics="true" pageId="6" pageNumber="79">Friesodielsia</emphasis>
species
</taxonomicName>
occurring at elevations above 900 metres.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Two new species of Friesodielsia (Annonaceae) from Peninsular Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Leeratiwong, Charan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0752-4275</mods:nameIdentifier>
<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand &amp; charan. leeratiwong @ gmail. com; https: // orcid. org / 0000 - 0002 - 0752 - 4275</mods:affiliation>
<mods:nameIdentifier type="email">charan.leeratiwong@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karapan, Sunate</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2284-3039</mods:nameIdentifier>
<mods:affiliation>Hala-Bala Wildlife Research Station, Hala-Bala Wildlife Sanctuary, Waeng, Narathiwat, 96160, Thailand &amp; karapann @ gmail. com; https: // orcid. org / 0000 - 0002 - 2284 - 3039</mods:affiliation>
<mods:nameIdentifier type="email">karapann@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Satthaphorn, Jiratthi</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7994-9056</mods:nameIdentifier>
<mods:affiliation>Department of Biology School of Science, Walailak University, Thasala, Nakhon Si Thammarat, 80161, Thailand &amp; jiratthi. sa @ wu. ac. th; https: // orcid. org / 0000 - 0001 - 7994 - 9056</mods:affiliation>
<mods:nameIdentifier type="email">jiratthi.sa@wu.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johnson, David M.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2896-7419</mods:nameIdentifier>
<mods:affiliation>Department of Biological Science, Ohio Wesleyan University, Delaware, Ohio 43015, USA &amp; dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-21</mods:number>
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<mods:number>589</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<subSubSection box="[136,711,1421,1444]" pageId="6" pageNumber="79" type="nomenclature">
<paragraph blockId="6.[136,1452,1421,1516]" box="[136,711,1421,1444]" pageId="6" pageNumber="79">
<taxonomicName authority="Leerat." authorityName="Leerat." authorityYear="2023" box="[136,507,1421,1443]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="chalermgliniana" status="sp. nov.">
<emphasis bold="true" box="[136,434,1421,1443]" italics="true" pageId="6" pageNumber="79">Friesodielsia chalermgliniana</emphasis>
Leerat.
</taxonomicName>
,
<emphasis box="[519,593,1422,1443]" italics="true" pageId="6" pageNumber="79">
<taxonomicNameLabel box="[519,593,1422,1443]" pageId="6" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs 2,4,5)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="79" type="materials_examined">
<paragraph blockId="6.[136,1452,1421,1516]" pageId="6" pageNumber="79">
<materialsCitation collectingDate="2022-05-27" country="Thailand" county="Type" elevation="500" location="Hala-Bala Wildlife Sanctuary" municipality="Betong District" pageId="6" pageNumber="79" specimenCount="1" stateProvince="Yala" typeStatus="holotype">
<collectingCounty box="[136,184,1458,1479]" pageId="6" pageNumber="79">Type</collectingCounty>
:
<emphasis bold="true" box="[188,215,1458,1479]" pageId="6" pageNumber="79">—</emphasis>
<collectingCountry box="[216,340,1458,1479]" name="Thailand" pageId="6" pageNumber="79">THAILAND</collectingCountry>
.
<collectingRegion box="[350,393,1458,1480]" country="Thailand" name="Yala" pageId="6" pageNumber="79">Yala</collectingRegion>
:
<collectingMunicipality box="[406,555,1458,1480]" pageId="6" pageNumber="79">Betong District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DD7A87FBFFD4FF88FF2716D6FE0EC0AE:500C6036FFD4FF89FD9C16E9FCCAC1C1" box="[563,845,1458,1480]" country="Thailand" county="Type" municipality="Betong District" name="Hala-Bala Wildlife Sanctuary" pageId="6" pageNumber="79" stateProvince="Yala">Hala-Bala Wildlife Sanctuary</location>
, ca.
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<elevation box="[885,945,1458,1480]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="79" unit="m" value="500.0">500 m</elevation>
</quantity>
elev.,
<date box="[1006,1133,1458,1479]" pageId="6" pageNumber="79" value="2022-05-27">
<collectingDate box="[1006,1133,1458,1479]" pageId="6" pageNumber="79" value="2022-05-27">27 May 2022</collectingDate>
</date>
,
<emphasis box="[1142,1348,1458,1479]" italics="true" pageId="6" pageNumber="79">Leeratiwong 22-1855</emphasis>
(
<typeStatus box="[1359,1448,1458,1480]" pageId="6" pageNumber="79">holotype</typeStatus>
: BKF; isotype: PSU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="80" pageId="6" pageNumber="79" type="description">
<paragraph blockId="6.[136,1452,1562,2093]" pageId="6" pageNumber="79">
The new species is most similar to
<taxonomicName authorityName="Das" authorityYear="1963" baseAuthorityName="Hooker &amp; Thomson" baseAuthorityYear="1872" box="[548,649,1562,1588]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis box="[548,649,1562,1588]" italics="true" pageId="6" pageNumber="79">F. affinis</emphasis>
</taxonomicName>
, but differs in its sparsely pubescent stem, abaxially shiny leaves (vs abaxially glaucous leaves), longer flower pedicels (
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vs.
<quantity box="[907,1021,1598,1624]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="6" pageNumber="79" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
), broadly ovate bracts,
<quantity box="[1294,1390,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="6" pageNumber="79" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
(vs. cordate to reniform,
<quantity box="[378,508,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="11.2" metricValueMin="5.0" pageId="6" pageNumber="79" unit="mm" value="8.1" valueMax="11.2" valueMin="5.0">5–11.2 mm</quantity>
long), shorter sepals (7–11.0 mm vs.
<quantity box="[957,1082,1634,1660]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="79" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
) with 5–7 indistinct veins (vs. distinctly 5-veins), outer petals without a raised midrib on the abaxial surface and a greater number of carpels (ca. 30 vs. 7–15).
</paragraph>
<paragraph blockId="6.[136,1452,1562,2093]" pageId="6" pageNumber="79">
This species also resembles
<taxonomicName authorityName="Steenis" authorityYear="1964" baseAuthorityName="Blume" baseAuthorityYear="1830" box="[504,804,1742,1768]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="79" phylum="Tracheophyta" rank="species" species="biglandulosa">
<emphasis box="[504,804,1742,1768]" italics="true" pageId="6" pageNumber="79">Friesodielsia biglandulosa</emphasis>
</taxonomicName>
but is distinguished by its abaxially not golden-pubescent (vs. golden-pubescent) leaves, the broadly ovate bract 5.0–8.0 mm long (vs. 1.5–4.0 mm and linear), ovate sepals 5.0–10.0 mm wide (vs. lanceolate sepals 2.0–
<quantity box="[651,741,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="79" unit="mm" value="4.5">4.5 mm</quantity>
), longer outer petals (35.0–57.0 mm vs. 25.0–32.0 mm) widest at base (vs. widest at midpoint), larger inner petals (23.0–31.0 Í 4.5–7.0 mm vs 5.0–11.0 Í
<quantity box="[1178,1319,1850,1877]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="6" pageNumber="79" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
), which are lanceolate and greater than half the length of outer petals (vs. ovate and less than half the length of outer petals) and a hairy stigma (vs. glabrous).
</paragraph>
<paragraph blockId="6.[136,1452,1562,2093]" lastBlockId="7.[136,1452,158,1193]" lastPageId="7" lastPageNumber="80" pageId="6" pageNumber="79">
Woody climbers up to
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high. Twigs with sparse, brown, appressed pubescence, becoming glabrate. Leaves shiny below, broadly oblanceolate, oblong-oblanceolate or obovate, blades 12–24 Í
<quantity box="[1101,1204,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="6" pageNumber="79" unit="cm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 cm</quantity>
, base rounded, rarely broadly cuneate, apex obtuse or acute to acuminate, acumen
<quantity box="[832,941,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="6" pageNumber="79" unit="mm" value="9.0" valueMax="15.0" valueMin="3.0">3–15 mm</quantity>
long, glabrous except sparsely pubescent on adaxial midrib, appressed-pubescent abaxially, midrib sunken above, secondary veins 8–16 per side; petioles
<quantity lastPageId="7" lastPageNumber="80" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="6" pageNumber="79" unit="mm" value="9.5" valueMax="13.0" valueMin="6.0">6–13 mm</quantity>
long, sunken above, sparsely brown-pubescent to subglabrous. Inflorescences internodal to leaf-opposed, 1- or 2-flowered, pedicel
<quantity box="[364,486,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="80" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
long, densely short-pubescent, bract broadly ovate,
<quantity box="[1081,1175,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="7" pageNumber="80" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
long, attached at or just below the pedicel midpoint, pubescent. Sepals 3, green, valvate, basally connate, ovate, 7.0–11.0 Í 5–1.0 mm, apex acute-obtuse, 5–7-veined, the veins obscure, densely pubescent abaxially, sparsely hairy and glabrous at base adaxially. Petals in two whorls of 3, yellow, valvate, free; outer petals appressed in bud, slightly spreading at maturity, narrowly lanceolate to linear-lanceolate, 35.0–57.0 Í 5.0–11.0 mm, widest basally, triquetrous apically, narrowly obtuse, concave on inner base, finely pubescent abaxially, glabrous, except sparsely pubescent at edge inside; inner petals erect, and coherent apically for about half of their length, narrowly lanceolate, 23.0–31.0 Í 4.5–7.0 mm, narrowly obtuse to slightly acute, sparsely hairy outside, glabrous inside. Stamens clavate, 1.3–2.0 mm long, anther dehiscence extrorse, apex of anther connective flattened, slightly oblique. Carpels ca. 30, oblong to elliptic-oblong,
<quantity box="[1246,1384,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="7" pageNumber="80" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 mm</quantity>
long, densely pubescent, stigmas falcate-capitate, cleft down the side, hairy. Fruit and seed unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="80" type="distribution">
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,520,554,580]" pageId="7" pageNumber="80">Distribution and ecology:—</emphasis>
Endemic to
<collectingRegion box="[655,705,554,580]" country="Thailand" name="Yala" pageId="7" pageNumber="80">Yala</collectingRegion>
and
<collectingRegion box="[760,884,554,580]" country="Thailand" name="Narathiwat" pageId="7" pageNumber="80">Narathiwat</collectingRegion>
Provinces (
<figureCitation box="[1017,1084,554,581]" captionStart="FIGURE 5" captionStartId="5.[136,229,1918,1940]" captionTargetBox="[288,1300,190,1894]" captionTargetId="figure-19@5.[288,1300,190,1894]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of Friesodielsia betongensis (●) and F. chalermglinii (▲) in Thailand. The map and marked locations were constructed in Rstudio v.3.6.3 (R Core Team, 2020) using ggplots package (Wickham, 2016) and its dependencies." figureDoi="http://doi.org/10.5281/zenodo.7755069" httpUri="https://zenodo.org/record/7755069/files/figure.png" pageId="7" pageNumber="80">Fig. 5</figureCitation>
), in shaded areas of tropical rain forest, ca.
<quantity box="[253,324,590,617]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="80" unit="m" value="500.0">500 m</quantity>
elev.
</paragraph>
<paragraph blockId="7.[136,1452,158,1193]" box="[189,642,626,653]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,351,626,652]" pageId="7" pageNumber="80">Phenology:—</emphasis>
Flowering April and May.
</paragraph>
</subSubSection>
<subSubSection box="[189,660,662,688]" pageId="7" pageNumber="80" type="vernacular_names">
<paragraph blockId="7.[136,1452,158,1193]" box="[189,660,662,688]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,366,662,688]" pageId="7" pageNumber="80">Local name:—</emphasis>
<emphasis box="[366,575,662,688]" italics="true" pageId="7" pageNumber="80">Bu nga soeng hala</emphasis>
(Thai).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="80" type="etymology">
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,356,698,724]" pageId="7" pageNumber="80">Etymology:—</emphasis>
In honour of Piya Chalermglin for his outstanding contributions to the taxonomy of
<taxonomicName box="[1285,1423,698,724]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
in
<collectingCountry box="[136,239,734,760]" name="Thailand" pageId="7" pageNumber="80">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="80" type="conservation">
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,465,770,796]" pageId="7" pageNumber="80">Conservation status:—</emphasis>
This species is currently known only from two collections in the tropical rain forest in the Hala-Bala Wildlife Sanctuary. It has a small population with few (&lt;50) individuals in the natural habitat (area of occupancy &lt;
<quantity box="[279,350,842,868]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="80" unit="km" value="10.0">10 km</quantity>
<superScript attach="left" box="[350,358,842,856]" fontSize="6" pageId="7" pageNumber="80">2</superScript>
), which makes it critically endangered (CR, B2a + b(ii, iii)+C2ai+D, IUCN 2012). Although this area was formerly well-protected, it is near agricultural areas, mainly rubber plantations. This may affect the area of occupancy of the species because of agricultural expansion.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="80" type="materials_examined">
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,596,950,976]" pageId="7" pageNumber="80">Additional specimen examined:—</emphasis>
<materialsCitation collectingDate="1972-04-16" country="Thailand" location="Waeng" pageId="7" pageNumber="80" specimenCount="1" stateProvince="Narathiwat">
<collectingCountry box="[596,744,950,976]" name="Thailand" pageId="7" pageNumber="80">THAILAND</collectingCountry>
.
<collectingRegion box="[755,882,950,976]" country="Thailand" name="Narathiwat" pageId="7" pageNumber="80">Narathiwat</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DD7A87FBFFD4FF88FF2716D6FE0EC0AE:500C6036FFD5FF88FCD110ECFC49C7D7" box="[894,974,951,977]" country="Thailand" name="Waeng" pageId="7" pageNumber="80" stateProvince="Narathiwat">Waeng</location>
,
<date box="[986,1131,950,976]" pageId="7" pageNumber="80" value="1972-04-16">
<collectingDate box="[986,1131,950,976]" pageId="7" pageNumber="80" value="1972-04-16">16 Apr 1972</collectingDate>
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,
<emphasis box="[1143,1451,950,976]" italics="true" pageId="7" pageNumber="80">Sangkhanchand et al. 1040</emphasis>
(BKF)
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.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="80" type="discussion">
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
<emphasis bold="true" box="[189,294,1022,1048]" pageId="7" pageNumber="80">Notes:—</emphasis>
The distinctive characters of
<taxonomicName box="[618,830,1022,1048]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="chalermgliniana">
<emphasis box="[618,830,1022,1048]" italics="true" pageId="7" pageNumber="80">F. chalermgliniana</emphasis>
</taxonomicName>
are broadly ovate bracts, flowers with triquetrous outer petals and inner petals longer than the half length of outer petals.
</paragraph>
<paragraph blockId="7.[136,1452,158,1193]" pageId="7" pageNumber="80">
Both new species belong to a species group within
<taxonomicName authorityName="Steenis" authorityYear="1948" box="[742,890,1094,1120]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="genus">
<emphasis box="[742,890,1094,1120]" italics="true" pageId="7" pageNumber="80">Friesodielsia</emphasis>
</taxonomicName>
marked by apically triquetrous outer petals pressed together until the flowers are near anthesis (
<bibRefCitation author="Leeratiwong, C. &amp; Chalermglin, P. &amp; Satthaphorn, J. &amp; Aongyong, K. &amp; Johnson, D. M." box="[637,915,1130,1157]" pageId="7" pageNumber="80" pagination="212 - 230" refId="ref4207" refString="Leeratiwong, C., Chalermglin, P., Satthaphorn, J., Aongyong, K. &amp; Johnson, D. M. (2021) New species and new records for the climber genus Friesodielsia (Annonaceae) in the flora of Thailand. Thai Forest Bulletin (Botany) 49: 212 - 230. https: // doi. org / 10.20531 / tfb. 2021.49.2.09" type="journal article" year="2021">
Leeratiwong
<emphasis box="[789,840,1130,1156]" italics="true" pageId="7" pageNumber="80">et al</emphasis>
. 2021
</bibRefCitation>
). The two new species raise Thai
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<emphasis box="[1304,1452,1130,1156]" italics="true" pageId="7" pageNumber="80">Friesodielsia</emphasis>
species
</taxonomicName>
diversity to 18.
</paragraph>
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</treatment>
</document>

418
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Key to the
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species
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of
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(
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(modified from
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Leeratiwong
<emphasis id="67A7EAFFFFD5FF88FC4D1644FB9DC13E" box="[994,1050,1310,1336]" italics="true" pageId="7" pageNumber="80">et al.</emphasis>
, 2022
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<keyLead id="EE2295D8FFD5FF88FF271638FA2CC17E" box="[136,1451,1379,1401]" pageId="7" pageNumber="80">1. Outer petals appressed until anthesis, then separating slightly, triquetrous apically..........................................................................2</keyLead>
</paragraph>
<paragraph id="556C36EDFFD5FF88FF3B16DBFA2CC193" blockId="7.[136,1452,1379,2091]" box="[148,1451,1407,1429]" pageId="7" pageNumber="80">
<keyLead id="EE2295D8FFD5FF88FF3B16DBFA2CC193" box="[148,1451,1407,1429]" pageId="7" pageNumber="80">- Outer petals separating widely in early bud stage, flat apically .........................................................................................................9</keyLead>
</paragraph>
</keyStep>
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<keyLead id="EE2295D8FFD5FF88FF2716C7FA2CC1B7" box="[136,1451,1436,1458]" pageId="7" pageNumber="80">
2. Twigs persistently hispid, pedicels
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long, sepals with 1 vein, leaf base irregularly cordate .................................
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- Twigs tomentose, pubescent or glabrous, pedicels less than
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long, sepals obscurely to strongely 5–7 veined or without visible veins, leaf base regularly cordate, rounded or broadly cuneate..............................................................................................3
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<keyLead id="EE2295D8FFD5FF88FF2716A8FA2CC20E" box="[136,1451,1523,1545]" pageId="7" pageNumber="80">3. Sepals 7.0–17.0 mm long ...................................................................................................................................................................4</keyLead>
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<keyLead id="EE2295D8FFD5FF88FF27154BFA2CC223" box="[136,1451,1551,1573]" pageId="7" pageNumber="80">- Sepals 2.5–6.0 mm long .....................................................................................................................................................................6</keyLead>
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4. Pedicellar bract apical, leaf apex rounded or truncate-retuse, rarely with a short mucro ca.
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long., outer petals lanceolate, inner half the length of the outer petals .......................................................................................................................................
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<keyLead id="EE2295D8FFD5FF88FF27153DFA2CC29E" pageId="7" pageNumber="80">- Pedicellar bract at pedicel midpoint or basal, leaf apex acute, acuminate, ormucronate, outer petals linear-oblong or linearlanceolate, inner petal greater than half the length of the outer petals ...............................................................................................5</keyLead>
</paragraph>
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<keyLead id="EE2295D8FFD5FF88FF2715C4FA2CC2D7" pageId="7" pageNumber="80">
5. Stem densely pubescent, leaves glaucous abaxially, floral pedicels
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long, with cordate to reniform bracts, sepals with 5 distinct veins, outer petals with a raised abaxial midrib, carpels 7–15 ..................................................................................
<taxonomicName id="92D34D6EFFD5FF88FAF815E6FA2CC2D7" ID-CoL="6JQ3Q" authorityName="Das" authorityYear="1963" baseAuthorityName="Hooker &amp; Thomson" baseAuthorityYear="1872" box="[1367,1451,1724,1746]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="67A7EAFFFFD5FF88FAF815E6FA2CC2D7" box="[1367,1451,1724,1746]" italics="true" pageId="7" pageNumber="80">F. affinis</emphasis>
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<keyLead id="EE2295D8FFD5FF88FF271582FA2CC30D" pageId="7" pageNumber="80">
- Stem sparsely pubescent, leaves shiny abaxially, floral pedicels
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long with broadly ovate bracts, sepals with 5–7 obscure veins, outer petals without raised abaxial midrib, carpels ca. 30 ..........................................................................
<taxonomicName id="92D34D6EFFD5FF88FB5A15ADFA2CC30D" box="[1269,1451,1782,1803]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="chalermgliniana">
<emphasis id="67A7EAFFFFD5FF88FB5A15ADFA2CC30D" box="[1269,1451,1782,1803]" italics="true" pageId="7" pageNumber="80">F. chalermgliniana</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
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<paragraph id="556C36EDFFD5FF88FF271448FA2CC32E" blockId="7.[136,1452,1379,2091]" box="[136,1451,1811,1833]" pageId="7" pageNumber="80">
<keyLead id="EE2295D8FFD5FF88FF271448FA2CC32E" box="[136,1451,1811,1833]" pageId="7" pageNumber="80">
6. Pedicels of flower/fruit ≥
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long ................................................................................................................................................7
</keyLead>
</paragraph>
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- Pedicels of flower/fruit &lt;
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long ................................................................................................................................................8
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</paragraph>
</keyStep>
<keyStep id="EE272E48FFD5FF88FF271416FA2DC3BE" pageId="7" pageNumber="80">
<paragraph id="556C36EDFFD5FF88FF271416FA2DC378" blockId="7.[136,1452,1379,2091]" pageId="7" pageNumber="80">
<keyLead id="EE2295D8FFD5FF88FF271416FA2DC378" pageId="7" pageNumber="80">
7. Twigs glabrous or sparsely hairy, leaves
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wide, outer petals linear, inner petal length greater than half the length of the outer petals.............................................................................................................................................................................
<taxonomicName id="92D34D6EFFD5FF88FAFD1431FA2DC378" ID-CoL="6JQ44" box="[1362,1450,1897,1919]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="glauca">
<emphasis id="67A7EAFFFFD5FF88FAFD1431FA2DC378" box="[1362,1450,1897,1919]" italics="true" pageId="7" pageNumber="80">F. glauca</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFD5FF88FF2714DDFA2DC3BE" blockId="7.[136,1452,1379,2091]" pageId="7" pageNumber="80">
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- Twigs densely hairy, leaves less than
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wide, outer petals lanceolate to ovate-lanceolate, inner petal length greater than half the length of the outer petals ...............................................................................................................................................
<taxonomicName id="92D34D6EFFD5FF88FA8D14F8FA2DC3BE" box="[1314,1450,1955,1976]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="betongensis">
<emphasis id="67A7EAFFFFD5FF88FA8D14F8FA2DC3BE" box="[1314,1450,1955,1976]" italics="true" pageId="7" pageNumber="80">F. betongensis</emphasis>
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</paragraph>
</keyStep>
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<keyLead id="EE2295D8FFD5FF88FF27149BFA2DC3F7" pageId="7" pageNumber="80">
8. Outer petals linear-lanceolate, acute, sepal apex acute/acuminate, leaf blades
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wide, apex acuminate/caudate, acumen (3–)
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long ...................................................................................................................................................
<taxonomicName id="92D34D6EFFD5FF88FB511486FA2DC3F7" box="[1278,1450,2012,2034]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="khaoluangensis">
<emphasis id="67A7EAFFFFD5FF88FB511486FA2DC3F7" box="[1278,1450,2012,2034]" italics="true" pageId="7" pageNumber="80">F. khaoluangensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFD5FF88FF2114A2FA2CCC2D" blockId="7.[136,1452,1379,2091]" pageId="7" pageNumber="80">
<keyLead id="EE2295D8FFD5FF88FF2114A2FA2CCC2D" pageId="7" pageNumber="80">
- Outer petals lanceolate, obtuse, sepal apex rounded, leaf blades 7.5–15.0 cm wide, apex obtuse to rounded, occasionally emarginate or mucronate........................................................................................................................................................................
<taxonomicName id="92D34D6EFFD5FF88FAE81B4DFA2CCC2D" ID-CoL="6JQ3G" box="[1351,1451,2070,2091]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="80" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="67A7EAFFFFD5FF88FAE81B4DFA2CCC2D" box="[1351,1451,2070,2091]" italics="true" pageId="7" pageNumber="80">F. latifolia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EE272E48FFDAFF87FF2713C4FA2CC4E8" pageId="8" pageNumber="81">
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<keyLead id="EE2295D8FFDAFF87FF2713C4FA2CC4B2" box="[136,1451,159,181]" pageId="8" pageNumber="81">9. Twig hairs persistently appressed, often glabrate proximal to first fully expanded leaves on specimen.........................................10</keyLead>
</paragraph>
<paragraph id="556C36EDFFDAFF87FF2713E7FA2CC4E8" blockId="8.[136,1452,159,843]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF2713E7FA2CC4E8" pageId="8" pageNumber="81">- Twig hairs at first appressed, soon becoming erect to spreading, twigs always pubescent proximal to first fully expanded leaves on a specimen ........................................................................................................................................................................................12</keyLead>
</paragraph>
</keyStep>
<keyStep id="EE272E48FFDAFF87FF2713ADFA2CC521" pageId="8" pageNumber="81">
<paragraph id="556C36EDFFDAFF87FF2713ADFA2CC50C" blockId="8.[136,1452,159,843]" box="[136,1451,245,267]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF2713ADFA2CC50C" box="[136,1451,245,267]" pageId="8" pageNumber="81">
10. Bracts
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long, shorter than pedicel, monocarp stipes
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long...................................................................
<taxonomicName id="92D34D6EFFDAFF87FAEB13ADFA2CC50C" ID-CoL="6JQ3T" box="[1348,1451,245,267]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="67A7EAFFFFDAFF87FAEB13ADFA2CC50C" box="[1348,1451,245,267]" italics="true" pageId="8" pageNumber="81">F. discolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFDAFF87FF271248FA2CC521" blockId="8.[136,1452,159,843]" box="[136,1451,274,296]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271248FA2CC521" box="[136,1451,274,296]" pageId="8" pageNumber="81">
- Bracts 4.0–9.0 mm long, longer than pedicel, monocarp stipes
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long ................................................................................11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EE272E48FFDAFF87FF271274FA2CC567" pageId="8" pageNumber="81">
<paragraph id="556C36EDFFDAFF87FF271274FA2CC542" blockId="8.[136,1452,159,843]" box="[136,1451,303,325]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271274FA2CC542" box="[136,1451,303,325]" pageId="8" pageNumber="81">
11. Leaves glaucous abaxially, inner petals lobed at midpoint on both sides .......................................................................
<taxonomicName id="92D34D6EFFDAFF87FA9F126BFA2CC542" box="[1328,1451,303,325]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="unonifolia">
<emphasis id="67A7EAFFFFDAFF87FA9F126BFA2CC542" box="[1328,1451,303,325]" italics="true" pageId="8" pageNumber="81">F. unonifolia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFDAFF87FF271217FA2CC567" blockId="8.[136,1452,159,843]" box="[136,1451,332,354]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271217FA2CC567" box="[136,1451,332,354]" pageId="8" pageNumber="81">
- Leaves shiny abaxially, inner petals without lobes at midpoint ............................................................................................
<taxonomicName id="92D34D6EFFDAFF87FAFC1216FA2CC567" box="[1363,1451,332,354]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="67A7EAFFFFDAFF87FAFC1216FA2CC567" box="[1363,1451,332,354]" italics="true" pageId="8" pageNumber="81">F. glabra</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EE272E48FFDAFF87FF271232FA2CC59D" pageId="8" pageNumber="81">
<paragraph id="556C36EDFFDAFF87FF271232FA2CC578" blockId="8.[136,1452,159,843]" box="[136,1451,361,383]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271232FA2CC578" box="[136,1451,361,383]" pageId="8" pageNumber="81">12. Leaves spreading-pubescent below ..................................................................................................................................................13</keyLead>
</paragraph>
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<keyLead id="EE2295D8FFDAFF87FF2712DDFA2CC59D" box="[136,1451,389,411]" pageId="8" pageNumber="81">- Leaves appressed-pubescent to glabrate below................................................................................................................................14</keyLead>
</paragraph>
</keyStep>
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<paragraph id="556C36EDFFDAFF87FF2712F9FA2CC5D2" blockId="8.[136,1452,159,843]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF2712F9FA2CC5D2" pageId="8" pageNumber="81">
13. Monocarps ellipsoid to oblongoid, (8.0–)10.0–14.0 mm wide, sepals (5.0–)6.5–10.0 mm long, outer petals 40–70 Í
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.... .................................................................................................................................................................................................
<taxonomicName id="92D34D6EFFDAFF87FAF2129BFA2CC5D2" ID-CoL="6JPRH" box="[1373,1451,447,469]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="kingii">
<emphasis id="67A7EAFFFFDAFF87FAF2129BFA2CC5D2" box="[1373,1451,447,469]" italics="true" pageId="8" pageNumber="81">F. kingii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFDAFF87FF271287FA2CC5F7" blockId="8.[136,1452,159,843]" box="[136,1451,476,499]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271287FA2CC5F7" box="[136,1451,476,499]" pageId="8" pageNumber="81">
- Monocarps ovoid, 6.5–7.0 mm wide, sepals
<quantity id="922B9B08FFDAFF87FDD31287FD4CC5F7" box="[636,715,476,498]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="81" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, outer petals (19–)27–31 Í
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....................................
<taxonomicName id="92D34D6EFFDAFF87FA801286FA2CC5F7" ID-CoL="6K246" box="[1327,1451,476,498]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="desmoides">
<emphasis id="67A7EAFFFFDAFF87FA801286FA2CC5F7" box="[1327,1451,476,498]" italics="true" pageId="8" pageNumber="81">F. desmoides</emphasis>
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<keyStep id="EE272E48FFDAFF87FF2712A2FA2CC62D" pageId="8" pageNumber="81">
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14. Sepals ≤
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long .........................................................................................................................................................................15
</keyLead>
</paragraph>
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- Sepals&gt;
<quantity id="922B9B08FFDAFF87FE8A114DFEEFC62D" box="[293,360,534,555]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="81" unit="mm" value="10.0">10 mm</quantity>
long .........................................................................................................................................................................16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EE272E48FFDAFF87FF271169FA2DC687" pageId="8" pageNumber="81">
<paragraph id="556C36EDFFDAFF87FF271169FA2CC641" blockId="8.[136,1452,159,843]" box="[136,1451,562,584]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF271169FA2CC641" box="[136,1451,562,584]" pageId="8" pageNumber="81">
15. Inner petals
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long, ovate-lanceolate to ovate, with marginal lobes, monocarp stipes
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long............
<taxonomicName id="92D34D6EFFDAFF87FABB1168FA2CC641" box="[1300,1451,562,584]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="brevistipitata">
<emphasis id="67A7EAFFFFDAFF87FABB1168FA2CC641" box="[1300,1451,562,584]" italics="true" pageId="8" pageNumber="81">F. brevistipitata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="556C36EDFFDAFF87FF27110BFA2DC687" blockId="8.[136,1452,159,843]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF27110BFA2DC687" pageId="8" pageNumber="81">
- Inner petals (11–)
<quantity id="922B9B08FFDAFF87FED41114FE64C663" box="[379,483,591,613]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="8" pageNumber="81" unit="mm" value="20.0" valueMax="25.0" valueMin="15.0">15–25 mm</quantity>
long, narrowly lanceolate to lanceolate, lacking marginal lobes, monocarp stipes
<quantity id="922B9B08FFDAFF87FB5C1114FAC9C663" box="[1267,1358,591,613]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="81" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long....... ........................................................................................................................................................................................
<taxonomicName id="92D34D6EFFDAFF87FA891136FA2DC687" box="[1318,1450,620,642]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="longipetala">
<emphasis id="67A7EAFFFFDAFF87FA891136FA2DC687" box="[1318,1450,620,642]" italics="true" pageId="8" pageNumber="81">F. longipetala</emphasis>
</taxonomicName>
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<keyStep id="EE272E48FFDAFF87FF2711D2FA2CC6D1" pageId="8" pageNumber="81">
<paragraph id="556C36EDFFDAFF87FF2711D2FA2CC6BC" blockId="8.[136,1452,159,843]" pageId="8" pageNumber="81">
<keyLead id="EE2295D8FFDAFF87FF2711D2FA2CC6BC" pageId="8" pageNumber="81">
16. Sepals gradually tapering to a blunt-acuminate apex, outer petals
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wide, inner petals
<quantity id="922B9B08FFDAFF87FBC111D2FB4DC698" box="[1134,1226,649,670]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="8" pageNumber="81" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long, petioles grooved adaxially ....................................................................................................................................................................
<taxonomicName id="92D34D6EFFDAFF87FAA211FDFA2CC6BC" box="[1293,1451,677,699]" class="Magnoliopsida" family="Annonaceae" genus="Friesodielsia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="81" phylum="Tracheophyta" rank="species" species="songkhlaensis">
<emphasis id="67A7EAFFFFDAFF87FAA211FDFA2CC6BC" box="[1293,1451,677,699]" italics="true" pageId="8" pageNumber="81">F. songkhlaensis</emphasis>
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- Sepals rounded to obtuse or acute apex, outer petals
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wide, inner petals
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long, petioles flat adaxially.........17
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17. Monocarp stipes
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long, swollen apically .......................................................................................................
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- Monocarp stipes
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long, uniformly thick ............................................................................................................................18
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18. Outer petals (50–)
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long, carpels 10–14, monocarps sparsely pubescent.....................................................
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- Outer petals
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long, carpels 14–24, monocarps densely pubescent .............................................................
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<paragraph id="E43F39D041AB77191C886D91A266B052" pageId="0" pageNumber="15050">New record for Kazakhstan.</paragraph>
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Art ISFS: 179300 Checklist: 1020560
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<heading id="1EDE07603CB2D63BC210A890233892A2">Zusammenfassung</heading>
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Artbeschreibung
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(nach
<bibRefCitation id="CFBBC8962689FD60A975E747847F5D41" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
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:
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hoch, unverzweigt, an den Knoten behaart, sonst kahl, duftend.
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zu 6-8
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,
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lang,
<emphasis id="BEB8568D0BB2C7F340298E06DCE5609C" bold="true">stachelspitzig</emphasis>
, am Rand und auf dem Mittelnerv rau.
<normalizedToken id="199DD55886672FF446EDB5500A3DCAD2" originalValue="Blütenstand">Bluetenstand</normalizedToken>
locker doldenrispig, mit
<normalizedToken id="79C5B89DA7FF8A5041E1C276D859549D" originalValue="wenigblütigen">wenigbluetigen</normalizedToken>
<normalizedToken id="90A9CB9BA6B4E0A4A4389BE13A95198D" originalValue="Teilblütenständen">Teilbluetenstaenden</normalizedToken>
.
<emphasis id="762F55B1891CE66CA057FDBFDE99C2F9" bold="true">
Krone weiss,
<normalizedToken id="D42A7FA6E2D644F0D7446697F6EB7027" originalValue="weit-trichterförmig">weit-trichterfoermig</normalizedToken>
, mit ca.
<quantity id="CDB7ABDD0321FE821042A21F951EFA60" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
langer
<normalizedToken id="00455FF44456068621A4CD5ED2E0CFF8" originalValue="Röhre">Roehre</normalizedToken>
und spitzen Zipfeln
</emphasis>
, Durchmesser
<quantity id="245101572DC7A0F69FEF465FEF6B2AB0" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4-6 mm</quantity>
.
<normalizedToken id="C43771D0239AF96B9CD938FCF51FD711" originalValue="Früchte">Fruechte</normalizedToken>
<quantity id="379C8C2D727AA6D3BBB86196713A1B81" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
hoch,
<emphasis id="924E9C639F4E7E61256D2DBD46038F51" bold="true">mit hakigen Borsten</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="7DA031EE05502D5E71F39A11A5D7DAFD" type="biology_ecology">
<paragraph id="03A71637F7268EE472010BE4D3E66C4B">
<normalizedToken id="2BCDC44AC9E597B9DF74EC765D9F221C" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="E8B17A306B12FFBA5F72FABF1E09D507" italics="true">
(nach
<bibRefCitation id="3F3FDFA0651A85A1ED81202082986B24" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: 4-6
</paragraph>
<paragraph id="BF7548F57FC9C715BAF1137F372AD68E">
Standort und Verbreitung in der Schweiz
<emphasis id="5F4AD0857CB82C3ACA5C3DA09540F787" italics="true">
(nach
<bibRefCitation id="88A0DBD2C9C3534C132B20B6F0B6DC51" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="B342253B65F6BDEDC544FC84226B4211" originalValue="Wälder">Waelder</normalizedToken>
, besonders Buchenhaine / kollin-montan(-subalpin) / CH (fehlt im Engadin)
</paragraph>
</subSubSection>
<subSubSection id="344F0E0B9F8E3F29596F87927FAA8B63" type="distribution">
<paragraph id="0490D023C3E7CE6677005D43C3559A6A">
Verbreitung global
<emphasis id="1FA4219F548DF0C35BDA2FCC4C0A1D85" italics="true">
(nach
<bibRefCitation id="C601001AD6C3DF945FEAE46978D0E6F0" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Eurosibirisch
</paragraph>
</subSubSection>
<subSubSection id="5C1FA489DD1CEFF961887F43DC06D905" type="biology_ecology">
<paragraph id="D96713CF7D2184D164852BEB7298C9DC">
<normalizedToken id="B9B25B6EFD0E017571C3BE540BC08277" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte
<emphasis id="2C6B4D0F280CC90AD190BFBA93CDA6C7" italics="true">
(nach
<bibRefCitation id="CCA576BC619C53480F3F79E002B9E537" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. 2010</bibRefCitation>
)
</emphasis>
3w33-13 + 2.g.2n=44
</paragraph>
</subSubSection>
<subSubSection id="85B828FF42FBC50CD09F7A84458C4579" type="multiple">
<paragraph id="6837974847B2481C7055B80B97D7CFCF">Status</paragraph>
</subSubSection>
<subSubSection id="1B56824FD0B292DC763AD7AB68B20F11" type="conservation">
<paragraph id="470467538F9AADF67D4014AA1520211E">
<heading id="CB5BE2603E070B22D27EEF6C77BD227F">Status IUCN</heading>
: Nicht
<normalizedToken id="DA30CD0E5F1A6D901D641A698C0BC365" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="0FD2A65BAB87F9AEAD41124329D7BD55" type="multiple">
<paragraph id="7856FB8CB2E4FF429C926DA12B7BB21C">
<heading id="06EEDD412F01B60ED6C038782800B5E7">
<normalizedToken id="6539E726807C328331569DE7485D679C" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="287D03E9CDF44D3335C6BAE6A55F03C4">Lebensform Geophyt</paragraph>
<paragraph id="6B3CE25C497014FEED47B34B09ED4FA8">
<heading id="81A103F931E7A936FC6609FCEBB8937E">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="FF9E682141050637A058CC4DEA951BB8" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="8FCF1ACE57996424AC2657E02CFEE2CF">
<table id="2A74589C653B44CC783261827E7F162B" class="table" inLine="true">
<tbody id="140D8D56BA2232E8D2C36ADB2B172203">
<tr id="AB712D672C3C823D648E03F530C0D13D">
<td id="77754DE9BAEF1541A8670CEA86B5D7C6">
6.1.4 - Hartholz-Auenwald (
<emphasis id="1467EA31A72C4A578995AB37E826CEDA" italics="true">Fraxinion</emphasis>
)
</td>
</tr>
<tr id="9A02F90547A6B7D1F870CD8DC65F443A">
<td id="D297CEE43B604B2A4945E81FC4B4FE45">
6.2 -
<normalizedToken id="2499CE0EFEECB542FDB10ABE6698006C" originalValue="Buchenwälder">Buchenwaelder</normalizedToken>
</td>
</tr>
<tr id="5FBD103552F7CA90ACD7914566BC2C8D">
<td id="B75D88FA5512A3771B3E762A8428970E">
6.2.1 - Orchideen-Buchenwald (
<emphasis id="13E01BADA5D2D6543729BE307096C4DC" italics="true">Cephalanthero-Fagenion</emphasis>
)
</td>
</tr>
<tr id="8D3D41EE79A319DA9AB98B95F18F1396">
<td id="50ED2C22FBFDFBE169312634340B7D98">
6.2.2 - Hainsimsen-Buchenwald (
<emphasis id="6DD7B7DBFED61DB6D925AD74C89AC4A5" italics="true">Luzulo-Fagenion</emphasis>
)
</td>
</tr>
<tr id="011133AB8A6E948B81703806BC55CF59">
<td id="0688AB0183E6A1AB8B39DCBBC918AEC4">
<emphasis id="A0D45FFE4B31F36E3F667F06A5CB52AD" bold="true">
6.2.3 - Waldmeister-Buchenwald (
<emphasis id="03D82C6FE34F0534023B40E115B86C7B" italics="true">Galio-Fagenion</emphasis>
)
</emphasis>
</td>
</tr>
<tr id="4DFEBE4BF1F301D6B4112B3A743061BB">
<td id="84170772A5EAC1419A5DF514200878B7">
<emphasis id="943FC3835DF6174B686ED10058A49AAD" bold="true">
6.2.4 - Zahnwurz-Buchenwald (
<emphasis id="BC4AA9FEBBF1AEE3BBA1CFA5D49B2080" italics="true">Lonicero-Fagenion</emphasis>
)
</emphasis>
</td>
</tr>
<tr id="F097BE1DAA631A2662F6B57098391FFD">
<td id="E8543105BCE7950F163A18A6584F9910">
6.3.1 - Ahorn-Schluchtwald (
<emphasis id="448676463E0535BAD52E3C75042FB386" italics="true">Lunario-Acerion</emphasis>
)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="DF697DC52F61EA75037864BCE7BAF72D">
<tableNote id="B1FC81897B65B3187A2C4D9C34D30976">
<emphasis id="E5C3790899BAF86634390925A9B804E8" bold="true">fett</emphasis>
<emphasis id="99160EE9F24BC160EF29C49177FCF00D" italics="true">
Dominante Art, welche das Aussehen des Lebensraumes
<normalizedToken id="9497E95A5C9ECBD615D4DDF3862532A1" originalValue="mitprägt">mitpraegt</normalizedToken>
</emphasis>
<emphasis id="B08F307233A63AA98F144252A4382930" italics="true">Charakterart</emphasis>
<emphasis id="F15581359888A16E9692437B51AD206A" italics="true">Weniger strikt an den Lebensraum gebundene Art</emphasis>
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="7208E73CA8D0B2809A2DF41D419012D7" type="biology_ecology">
<paragraph id="0D2A0A189F1BB9AC32A99FF4DFCE7CB9">
<normalizedToken id="530476E20F3F5AF3B29A2508B8F97B43" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="C5BFD1A1B160C22472B7CA53CBF7356E" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="83B1C56E79948D3AF70CAF527B683E15">
<table id="8046BD890BEA40C944C3174F0AC8382D" class="table" inLine="true">
<tbody id="2C75CCB7F71102E84461A2685182872E">
<tr id="F7C5ABA581DDAE51D8C7FDA494803F0F">
<th id="6337623442730CBE674AB29D44FCD86B" colspan="2">
<emphasis id="77A602CF27D92CABE9B48684B98666A5" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="5175DA45192C540746FA4795D4962F31" colspan="2">
<emphasis id="1B8F99C8CFC706869C03FF3AF2188A4C" bold="true">Klimafaktoren</emphasis>
</th>
<th id="441C4BF01C08F38C51D03E6963C0E6CC" colspan="2">
<emphasis id="ED74590E569264F25AC7A10B3A22A3D9" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="9E1F7A0B5A2FF51CB511B47716DA34A9">
<td id="FAFB3CA704B6E8125C3F243F4A34AB9B">Feuchtezahl F</td>
<td id="E6D8A6E3E78F80A9289C6FEC8D265B55" style="text-align:left;width:100px">
<normalizedToken id="BB189AD982C20FDEB6F373633E9DB5A8" originalValue="mässig">maessig</normalizedToken>
feucht; Feuchtigkeit
<normalizedToken id="6D2AD7531FA949F657A5BB032951F167" originalValue="mässig">maessig</normalizedToken>
wechselnd (
<normalizedToken id="5CF1D4E222B2A2023051F3C57DB04988" originalValue="±">+/-</normalizedToken>
1-2 Stufen)
</td>
<td id="38664A4F75C409DD2F77C6F0D9A3D924">Lichtzahl L</td>
<td id="D90BD6997F3879C62DCC1818EE213130" style="text-align:left;width:60px">sehr schattig</td>
<td id="3FFA29E3F37E10249D34EB8890DDC60D">Salzzeichen</td>
<td id="DDB10A06B1DA6F1671873254CE933C4D" style="text-align:left;width:60px">--</td>
</tr>
<tr id="22D9BAA5C6C93AEA83A3589AE0AB6D33">
<td id="EFAAA3F5A10741938B76B9CBBF41B4EB">Reaktionszahl R</td>
<td id="D7B48EA18046E0AAF1F20DF866C7BD67" style="text-align:left">schwach sauer bis neutral (pH 4.5-7.5)</td>
<td id="58286E7326FFB4BE28167A6ECECF826A">Temperaturzahl T</td>
<td id="0EEB80AEE30579CAE4902F39FDB8BC35" style="text-align:left">unter-montan und ober-kollin</td>
</tr>
<tr id="B906764312AA4A37A59662449B3F37C8">
<td id="3F4D7D4366AF16C318F7082FE23BE4C2">
<normalizedToken id="6041404A434C4710988FE21DD8068A43" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="96915DB6CD08EFE2BD8C570C3E4B256B" style="text-align:left">
<normalizedToken id="B3B86BEB6024F12F29421673CA4BED46" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="469D5AF66BE29B920674B141CACED94A" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
bis
<normalizedToken id="4E35E756ECE631CA724E2D7D8C4750B8" originalValue="mässig">maessig</normalizedToken>
<normalizedToken id="E88CDCB50C6BB844F077180E78EB5A82" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="A6B11AD3BC2209F449F8BF7F28C63E8D">
<normalizedToken id="EE74AA0780E216E3BAE211C453FD0A64" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="5AE3AFF5008A86411EED0E44DFD4D4BE" style="text-align:left">subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter)</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="5CFCC07751B4B2E4E172DCBF0D2DEC6B" type="reference_group">
<paragraph id="760ED9CC7E1E1E7AD18C60D025845887">
<heading id="45A8568E22F104E29F399115E1B13106">Nomenklatur</heading>
</paragraph>
<paragraph id="4D1A43049C38AE66B2B7F04C2BE45FFC">
<treatmentCitation id="69E28D93A3A41AE8B5C63B1A4EC40D00">
<heading id="BAF2CECCB3377CD3439445F0CCAE4D37">
<normalizedToken id="C8E806F9598274A9111D1B18090ECEFB" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="C4E72ABC33705C0CCA87A3C6E6314FD1" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="3727A01A94EE30D3E21B8FA835C07C0E" ID-CoL="3F66X" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">
<emphasis id="37F25AC662CBC4B6FE18D31FCA622462" italics="true">Galium odoratum</emphasis>
(L.) Scop.
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="AF0178AB02F90D7A269AEB898D1EC6ED" type="vernacular_names">
<paragraph id="0028CB39CDC658D633037C5A05E2D4A8">
Volksname Deutscher Name:
<vernacularName id="C90E205771BA14D1560CEE4F866FF1A5" language="deu">Echter Waldmeister</vernacularName>
Nom
<normalizedToken id="CA1934D038A3713D4014B081BAB2AEC2" originalValue="français">francais</normalizedToken>
:
<vernacularName id="F2D1DF61BD8F78D3C59B4A57E33AE769" language="fra">Gaillet odorant</vernacularName>
Nome italiano:
<vernacularName id="FE57DA924F972D3E6DC39CFB91BB885D" language="ita">Caglio odoroso</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="4CB46EBC17999E8550A20283FDF78591" type="reference_group">
<paragraph id="AE2840F4F941CB8AF9AFEFF7CF73AE6F">
<normalizedToken id="A5B0B6B9120356A396B75ED486D2E132" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="4072B19BAAF5BE82980589C587CBD717">
<table id="FFC3E1336C95D8DF0AB577B223026490" class="table table-condensed" inLine="true">
<tbody id="7BF38D28A5EDC1A9626C6755E53E91D4">
<tr id="5624842682B84EB51BCD6CF2DFCC6157">
<th id="C7BACFE51E595D1C9D7D7AF694037313" width="80px">Relation</th>
<th id="9E42E9C27BDDFAD301959D84AA2EF4B6">Nom</th>
<th id="BADE6E748D578424FDB36843A26A780C">Referenzwerke</th>
<th id="3C7FBAEC9F444A13F42874E33C235A46" width="90px">No</th>
</tr>
<tr id="54886F67AED13BDD54E7584567525FF3">
<td id="752CE6D7EC3776F1D14C5ACC3FCEF780" class="taxon-relation">=</td>
<td id="266D354D575BC940AF35D4E72EEB43A8">
<taxonomicName id="721D38E53528936636F109F434E893CC" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
</td>
<td id="09108ED14F23C8BA3F6E3FA25F46D483">
<treatmentCitation id="01C9C24C8B5FCDA72D839E0C75921994">
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<td id="938C8AB066ACDF73E036981298524F3C">179300</td>
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<tr id="D376755DA4070AA0763216CFF8772D4B">
<td id="7E469442BC3DC78972029EFDCEABEAD4" class="taxon-relation">=</td>
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<taxonomicName id="811D3C6F0712E43CF81A0F400579BEC8" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<treatmentCitation id="ED72506B1032D6AB7E6E3958999D6A6A">
<bibRefCitation id="8BDA196490B7AD4E8E7D55789B23816B">Flora Helvetica 2001</bibRefCitation>
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<td id="71E0E5D6ABF061A341596810A0442673">1939</td>
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<tr id="57FB1FC9BB107124AEE445555F6C2E62">
<td id="B004CCA83394585155E4D74CCE5956DF" class="taxon-relation">=</td>
<td id="0B18E9A17E1FAC97EA5C265206C239A2">
<taxonomicName id="5928D38E5F0B47FED48851C65B17AB19" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="B361BE45662339875E040772BDC667C1">
<treatmentCitation id="D63DCBDF1C3130816A99AF1BCB25C685">
<bibRefCitation id="F835E0A47C3C7B708D7A517C7C01181C">Flora Helvetica 2012</bibRefCitation>
</treatmentCitation>
</td>
<td id="07A97BD366C6C5F5E975032822C71AD0">1443</td>
</tr>
<tr id="4AC253C1FE71C19E86ADE60808CD36EA">
<td id="4DA91966E0EB8BD8F88937ADBBDC09F3" class="taxon-relation">=</td>
<td id="C0682907DABDE5DAEF70AEA8E92335E8">
<taxonomicName id="29BB06B4F4353A87439D6B4FE7AE45E3" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="181C2459C72EAC64A8C3893497E07898">
<treatmentCitation id="B5ED4E375ABD31A65F640D2BAC47ACB1" httpUri="http://treatment.plazi.org/id/5F3ADAE75FB9E5C97C0D950B1D9E0320">
<bibRefCitation id="FFBC5D58665754FEF5D5B9A6B179A2BA">Flora Helvetica 2018</bibRefCitation>
</treatmentCitation>
</td>
<td id="F08160FCAF2743737C188F78A8CD0705">1443</td>
</tr>
<tr id="ECAA5C6806F59960C834EE9B61E7F245">
<td id="DF390332084D119ABE0EC5E39CF6F8E2" class="taxon-relation">=</td>
<td id="290ED7BE344F9336B8E8CC9CB1E3024D">
<taxonomicName id="17A26B023E1A9A70CED2D9C491F94599" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="0505F650CA61FF4740E87CB5DCBBEB64">
<treatmentCitation id="08223E2A819098434BCFB3BFF135008D">
<bibRefCitation id="6AAC5CDC1B5889A1C344191874C48691">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="75A7C3CE698D2A1C94BF0439A5AF70CB">179300</td>
</tr>
<tr id="A100082BEFF660192F50F97FD0F516CA">
<td id="8BB7FF5F275FB53FB155DAC18867C23E" class="taxon-relation">=</td>
<td id="B1F4794CCA2845DCA08F2F68EFA6C247">
<taxonomicName id="A73DDF8388EEC84FDFF2506D8DC2FD42" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="2F9531B3E921EAA90B7B769099A6B605">
<treatmentCitation id="7B5C2159DAC4346C98D27E1DA7C44AAA">
<bibRefCitation id="656147C8BB4864D08F6246A5AC390C4B">Landolt 1977</bibRefCitation>
</treatmentCitation>
</td>
<td id="1042E2253337FA232D1132F8E76DD424">2785</td>
</tr>
<tr id="77C106038B95071657523722ED5FD8FF">
<td id="B3689D19B99D6DDD8241E3686E2D719C" class="taxon-relation">=</td>
<td id="51115337628142D129799D3F1BD7AD35">
<taxonomicName id="66B88A6D49D105F8CF5D4931B4226FB3" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="28DD1D5BA0C97AA70260CEA604C9AF18">
<treatmentCitation id="1EB7A588A343D8F126BBA5EFB597948F">
<bibRefCitation id="3A96EF2F9E2ADD11AFEEC60ED38EC336">Landolt 1991</bibRefCitation>
</treatmentCitation>
</td>
<td id="2BA53E828E63BB31432410DC46ED2E9B">2265</td>
</tr>
<tr id="F7DD82B2D6FD0881E93B9F34F7FC9EDB">
<td id="CA10F2AA3CA2AD38C3FC62891FC482D3" class="taxon-relation">=</td>
<td id="679F424A96AF7A32C64ADC896AE7399F">
<taxonomicName id="D8521661B30C15BB3248D5D81BBDA2D6" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<treatmentCitation id="C576A6491E925FBE988D765C7DF17D86">
<bibRefCitation id="4DA899DE783D8336950B694324152481">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="278815494E9293A4BFB63FE1983ACA87">179300</td>
</tr>
<tr id="29710609AEA6A6A30A0261E49351F306">
<td id="71324AB67460EF016B3531A598672539" class="taxon-relation">=</td>
<td id="E465CFD35F70BE6FDF70DD28DC8D8877">
<taxonomicName id="9C9B2FAC885219A668FCB147F10832E4" authority="(L.) Scop." authorityName="Scop." baseAuthorityName="L." class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="odoratum">Galium odoratum (L.) Scop.</taxonomicName>
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<td id="034EAF034D6A46A90A7560EE0BFB0E20">
<treatmentCitation id="994C3418810CCED8766123C3E02C6543">
<bibRefCitation id="54D351A0BE39B19D678231A4FD02767B">Welten &amp; Sutter 1982</bibRefCitation>
</treatmentCitation>
</td>
<td id="9A4B120244CA9FCF9C76D81BDAC64A46">1608</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="E00C9021C2916B1E08D1991FBBD2B81C">
<tableNote id="7A5439AF63E26751E98C2DF08DE64B55">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="D254117AEF6488158C5C024B8550BEE7" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="908239235F72D34A8473736DC3FD4D04" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="7CAFDA5DEC608D9B8A131FC5ED994A64" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="A5EF86E2E11F3B8E196721B48C061058" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
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)
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</paragraph>
</subSubSection>
<subSubSection id="DCAF41863302B2C56D1590C48E8625A7" type="conservation">
<paragraph id="14D260974F11FE623FFBDE8982C61A2A">
<heading id="D3757079498FFB74E6FE71418BA4EFD5">Status Indigenat</heading>
: Indigen
</paragraph>
<paragraph id="D33944972BCF093C4A3B70F6A9ABF37E">
<emphasis id="44EDF6A1CA87E6354D4739D200219BE6" bold="true">
<heading id="F610878C5CCBDFD4C7D7EF20D3A80CC1">
Liste der
<normalizedToken id="393B0212CB17B5C059E91F590DF6AFF5" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="1CD4898C77FC2E6BDAF468BA405588C7" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
<paragraph id="F48073511663605FDFC5B533D508FDBB">
<heading id="7A426419FB7820A1D6F8F3B05DBFD750">Status Rote Liste national 2016</heading>
</paragraph>
<paragraph id="D4496DE93CADDBF388502C17B3AA1AE2">
<heading id="9CC35697D46B5DE8574C6A75EBE63A61">Status IUCN</heading>
: Nicht
<normalizedToken id="A14481F8FA5FAE0785B94529F94CCAA3" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
<paragraph id="6F9E762DD675387AD72AB11932DC9373">
<heading id="EE2C93B4B7F9ECF962946E3DDC4E55BF">
<normalizedToken id="2DF066280469DBF5136C96033C851570" originalValue="Zusätzliche">Zusaetzliche</normalizedToken>
Informationen
</heading>
Kriterien IUCN: --
</paragraph>
<paragraph id="5D0BE5DE23E81F3B5811BC58C3EC0DCE">
<heading id="40681DC3CB758880BF5FA36842FBB88C">Status Rote Liste regional 2019</heading>
</paragraph>
<paragraph id="E3AFEC2B273AA5A3D3ACB955375F78D4">
<table id="3EC2007191A92786559A7672C49D8CE9" class="table table-condensed" inLine="true">
<tbody id="686ECAA3B6379DD0156D39CF0314E23E">
<tr id="8E0CCD8F38E4C7040DDE711EEEBB4919">
<th id="502CA6F03DE769A60F9DB036650B4E22">Biogeografische Regionen</th>
<th id="54AA2C917729C87440EEDBCB4FE5B16B">Status</th>
<th id="D6A030A4AF6E48DBF112B21551610BDD">Kriterien IUCN</th>
</tr>
<tr id="BD26DAA4254CD03BC37BCB7E02E6B511">
<td id="49E205766A56B19602EBA4961FF06501" class="regional_status">Jura (JU)</td>
<td id="C0B9BEBCE514C2BE010361DDC35B5E1A">
nicht
<normalizedToken id="823DD66986BA552AC2DEC52F34242598" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="F2A06921BFB15BFBDFC33BD79CF2F13C">
<td id="2F4272F24057FDDB6308038E6FE1FAB9" class="regional_status">Mittelland (MP)</td>
<td id="0530C3669DA22EC1B6A62407264CDA77">
nicht
<normalizedToken id="D3201F0A837AB2C0183761AE82A1E478" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="01B7F5DAB8F71874DE355DC87882438C">
<td id="8542D93CDCDCCEB0DFC9CF2765A6381E" class="regional_status">Alpennordflanke (NA)</td>
<td id="6C13C76E6AFC766FA5901DFF5451713F">
nicht
<normalizedToken id="D530759D4D5BF74CA5FEB4B49656EFC0" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="CB19FD05F644A9EBB46B069425E7447B">
<td id="66B2B2C1C2992118F5DAA4E2DFBF4425" class="regional_status">
<normalizedToken id="4EB97D472E1DB78C230DB9A3D8567C3E" originalValue="Alpensüdflanke">Alpensuedflanke</normalizedToken>
(SA)
</td>
<td id="2039AD9F357F0DA69B82E703C437CE01">
nicht
<normalizedToken id="8CED3DDC1C610F73D153D962A9A69561" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="048C5969AFA4F0DDE229D144D827AB83">
<td id="8E613F2206B3CAEAC5B809B97097652F" class="regional_status">
<normalizedToken id="10B8E05DB8992A49567127059EF46738" originalValue="Östliche">Oestliche</normalizedToken>
Zentralalpen (EA)
</td>
<td id="A3ED1B51741AECA79D5FC88D04F6CBD1">
nicht
<normalizedToken id="B7441281EC193F89365FF05DBB139D44" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="BDC3EE181403709D562631C2BBA61B92">
<td id="D1AD475EBA1784CAF7C97FD5823A9BE9" class="regional_status">Westliche Zentralalpen (WA)</td>
<td id="3D7299AFA7A63865FBB603EE274211F1">
nicht
<normalizedToken id="211E57A691BC205B8E4F9B5646B97C67" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="7BA0AE564ECC2139009D7AC08E6B7F89">
<heading id="B77393D0F83C565359CEFD4BE6F57133">
Status nationale
<normalizedToken id="7A8B794DD901D0864C67D421C8AFFBD3" originalValue="Priorität">Prioritaet</normalizedToken>
/Verantwortung
</heading>
</paragraph>
<paragraph id="E17EDCAF4C9B4949E667A2F8740B4678">
<table id="7E550043FC23040D47C29500732541AF" class="table" inLine="true">
<tr id="6B8C42EF39489D21E6F79D05009151F3">
<td id="840AEE309156729B6F9CE2D7AD9A11D2">
Keine nationale
<normalizedToken id="5938D29614AEA8F313255738E8804F26" originalValue="Priorität">Prioritaet</normalizedToken>
oder internationale Verantwortung
</td>
</tr>
</table>
</paragraph>
<paragraph id="D071B6AC395D8A59384E86E4D170B7F5">
<heading id="69756F6DB19724F9D1E38C6F233EE4D5">Schutzstatus</heading>
</paragraph>
<paragraph id="0F9B74B7095A99A549C87016D31B8FEB">
<table id="FFE6E764607E81A0D9855BBE70D1D0E2" class="table" inLine="true">
<tr id="BE3F49F120EB2F8C5DBFC8AE3F43277A">
<td id="4C779FE322C354A32E4B33D17331F775">Kein internationaler, nationaler oder kantonaler Schutz</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

189
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<mods:namePart id="719932B426F7C89D45F8827E2E2E16F7">Aspoeck, Horst</mods:namePart>
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<mods:namePart id="170F3323B3CB0E7BE00FC5EDCCEAB8D7">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="B9088C3022CFBDBE0CD79EC6BC280006">Aspoeck, Ulrike</mods:namePart>
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<mods:title id="CBD7B91E7EE076F6FD0EAEB52FC56884">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="4A9EA30DBEFA39065D6B97F260C2A518">
<mods:date id="CA108ED7865AF7226BE3979FC75196B4">2014</mods:date>
<mods:detail id="717E04D2153E40E2B5CB7825B50C5A9C" type="volume">
<mods:number id="18BC1FBA773DFB3CA080AB2DC9B9C8A7">61</mods:number>
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<mods:url id="D34EC823CE020A4F62736AF4119BB946">http://dx.doi.org/10.3897/dez.61.8793</mods:url>
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<mods:classification id="8222EE1F3953E48ADDFA925CB565D324">journal article</mods:classification>
<mods:identifier id="115D2E3403C815C0997C710FA237C172" type="DOI">http://dx.doi.org/10.3897/dez.61.8793</mods:identifier>
<mods:identifier id="341CEE36B3BAD290B78B8327C577B4E8" type="Pensoft-Pub">1860-1324-2-141</mods:identifier>
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<treatment id="DD7AFEFAC42483A6E4040F3502C6864A" ID-GBIF-Taxon="154366013" LSID="urn:lsid:plazi:treatment:DD7AFEFAC42483A6E4040F3502C6864A" httpUri="http://treatment.plazi.org/id/DD7AFEFAC42483A6E4040F3502C6864A" lastPageNumber="142" pageId="1" pageNumber="142">
<subSubSection id="9D52D2531084C72B120AB0B9BF254C45" pageId="1" pageNumber="142" type="nomenclature">
<paragraph id="FEFB3C267B6C0F4C0716C34BBEFF2EEC" pageId="1" pageNumber="142">
<taxonomicName id="DC2CBAF7D80AE0289122140A6B1BC791" ID-CoL="3657Y" authority="McLachlan, 1883" authorityName="McLachlan" authorityYear="1883" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">
<pageBreakToken id="CDBC1A9D9B398B924F34EFD2EC0B1AED" pageId="1" pageNumber="142" start="start">Dilar</pageBreakToken>
japonicus McLachlan, 1883
</taxonomicName>
Figs 1, 7-12
</paragraph>
</subSubSection>
<subSubSection id="4046173966DD862D4F1FBBFD19A408BC" pageId="1" pageNumber="142" type="reference_group">
<paragraph id="EE02694151C1C4D12D5957592CE9BA00" pageId="1" pageNumber="142">
<taxonomicName id="38B20187C52C56EB34E2C6C6CC4B3D81" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
McLachlan, 1883: 220. Type locality: Japan (Fukushima).
</paragraph>
<paragraph id="8D788D8678E29EDB1F970F44FB530127" pageId="1" pageNumber="142">
<taxonomicName id="529D3CE0155FF9098D65DA7399F333F0" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar nohirae" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="nohirae">Dilar nohirae</taxonomicName>
Nakahara, 1914: 297. Type locality: Japan (Nara).
</paragraph>
<paragraph id="6BB4ADF7C07905D9983783F611581FE8" pageId="1" pageNumber="142">
<taxonomicName id="21160D5F2D2A68BC30C433956EF1EF53" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
Dilar japonicus var. gracilis Kuwayama, 1921: 71. Type locality: Japan (Fukushima).
</paragraph>
</subSubSection>
<subSubSection id="BBCC5DA17AB5CF6A73EBE37A48C8AEEA" pageId="1" pageNumber="142" type="diagnosis">
<paragraph id="B4187604C577C5F3948D4DB5CD633ECA" pageId="1" pageNumber="142">Diagnosis.</paragraph>
<paragraph id="A3D52CB637ADF99A4D92F12760E648E3" pageId="1" pageNumber="142">This species is characterized by the forewings with numerous pale brown markings, and the male ectoproct in dorsal view with three projections at tip which is sparsely haired.</paragraph>
</subSubSection>
<subSubSection id="705190DCCFE56157A175A4430B3E135F" pageId="1" pageNumber="142" type="description">
<paragraph id="A79AF155135E23E8A50EDAC9BA6C0130" pageId="1" pageNumber="142">Description.</paragraph>
<paragraph id="452D32E540919FA700C8B49830360225" pageId="1" pageNumber="142">Male. Body length 4.3-5.3 mm; forewing length 10.0-11.4 mm, hindwing length 8.8-9.4 mm.</paragraph>
<paragraph id="36EF50DA7350500F68A79162FFEC6324" pageId="1" pageNumber="142">Head pale yellowish brown, with pale yellow setose tubercles. Compound eyes blackish brown. Antenna with ca. 29 segments, pale yellowish brown, pedicel with brown annular stripes, flagellum unipectinate on most flagellomeres, medial branches much longer than those on both ends, longest branch nearly 3.0 times as long as relevant flagellomere, but branch of 1st flagellomere short and dentate, distal seven flagellomeres simple.</paragraph>
<paragraph id="5F848D27476E65FF68254E63C94673A6" pageId="1" pageNumber="142">Prothorax pale yellow, pronotum pale yellowish brown, with anterior margin and posterolateral corners pale yellow, medially with a pair of ovoid markings; mesothorax pale yellowish brown, mesonotum dark brown on anterior and lateral margins; metanotum pale yellowish brown, slightly darker on lateral margins. Legs pale yellowish brown, femora blackish brown at tip. Wings hyaline, slightly smoky brown, with numerous brown spots. Forewing ~2.1 times as long as wide, densely spotted, with markings on proximal half much darker, arranging as transversely arcuate pattern, a broad immaculate area present on either side of median nygma; two nygmata present on proximal and median portion of forewing, median one much larger than proximal one. Hindwing ~2.0 times as long as wide, slightly paler than forewing, with similar marking pattern; one nygma present at middle. Veins pale brown, crossveins much paler than longitudinal veins. Forewing with trichosors present along wing margin between R and CuA; Rs with three main branches; MP with two main branches; two gradate crossveins present at middle. Hindwing with trichosors present along wing margin between R and CuA; Rs with three main branches.</paragraph>
<paragraph id="12F714E8C70D27A11EE493061D4EB561" pageId="1" pageNumber="142">Abdomen pale yellow, pregenital segments dorsally yellowish brown. Ninth tergite in dorsal view with an arcuate anterior incision and a deeply V-shaped posterior incision, leaving a narrow median portion and a pair of subtriangular hemitergites, which are obtuse distally and densely haired; in lateral view broad, with straight ventral margin and arcuate posterior margin. Ninth sternite obviously shorter than ninth tergite, truncately posteriad. Ectoproct in dorsal view with three projections at tip which is sparsely haired, posteroventrally with a pair of subsemicircular and flattened projections, a pair of bifid unguiform projections and a pair of short, feebly sclerotized, digitiform projections. Ninth gonocoxite with anterior half broadly spoon-shaped and with posterior half slenderly elongate and curved posteroventrally; tenth gonocoxite slenderly elongate, much longer ninth gonocoxite, with incurved base and spinous tip, medially with a lobe connecting to ninth gonocoxite; gonarcus beam-shaped, slightly expanded on both ends, laterally connecting to base of ninth gonocoxites. Hypandrium internum nearly trapezoidal, with lateral margins slightly arcuate.</paragraph>
<paragraph id="4AB11E67B16D6D15CB35D17EBA638B20" pageId="1" pageNumber="142">Female. Body length 6.5 mm; forewing length 14.0 mm, hindwing length 12.7 mm.</paragraph>
<paragraph id="0DC8C58D675BCEC83F6CE0500C4F9BE0" pageId="1" pageNumber="142">Seventh sternite (Fig. 11) in lateral view subtrapezoidal, in ventral view nearly rectangular and slightly convex on posterior margin. Eighth abdominal segment ventrally without subgenitale. Bursa copulatrix (Fig. 11) with colleterial gland tubular and elongate, sinuate medially; basal part of bursa copulatrix sac-like in lateral view and nearly ovoid in ventral view, posteriorly slightly sclerotized on both sides; bursal accessory gland not observed. Ectoproct rather small, ovoid.</paragraph>
</subSubSection>
<subSubSection id="CAA72BCA0A345A809F1051D81D6F5F0C" pageId="1" pageNumber="142" type="materials examined">
<paragraph id="9EFEA0719A74E41B00DBEEBF65F486FF" pageId="1" pageNumber="142">Materials examined.</paragraph>
<paragraph id="CB11D68522E14704D28EF86226333EAB" pageId="1" pageNumber="142">
Holotype ♂, &quot;Japan, Fukushima [
<geoCoordinate id="44BC6F95E0669F8BA434CE8B975C0D3A" direction="north" orientation="latitude" precision="925" value="37.733334">37°44'N</geoCoordinate>
,
<geoCoordinate id="CAF935BD03298AD0A3E5157877633FFD" direction="east" orientation="longitude" precision="925" value="140.46666">140°28'E</geoCoordinate>
]/28.7.[18]81/Type/japonicus McL[achlan]&quot; (BMNH). JAPAN: 1f#, Aomori, Tsuta-onsen [
<geoCoordinate id="9D7A3D11F8C307719FC41482EC0F0A9E" direction="north" orientation="latitude" precision="925" value="40.216667">40°13'N</geoCoordinate>
,
<geoCoordinate id="7FB9F51520AD0BB849A3654E5007FABC" direction="east" orientation="longitude" precision="925" value="140.5">140°30'E</geoCoordinate>
], 6.VIII.1953, K. Sadanaga (NIAES); 1♀, Gunma, Kuni-mura [
<geoCoordinate id="957A976F583A8188D9414D570DD8EA4B" direction="north" orientation="latitude" precision="925" value="36.633335">36°38'N</geoCoordinate>
,
<geoCoordinate id="56C62F8B7DAB22E0D354B8ACDCB4A6D3" direction="east" orientation="longitude" precision="925" value="138.63333">138°38'E</geoCoordinate>
], 10.VIII.1992, K. Konishi (NIAES); 1♂, Gunma [
<geoCoordinate id="C14211B331E5C91EE4669EA3FA4E2DED" direction="north" orientation="latitude" precision="925" value="36.383335">36°23'N</geoCoordinate>
,
<geoCoordinate id="C3D4869BA5B343EB6C823577CECA33E7" direction="east" orientation="longitude" precision="925" value="139.05">139°03'E</geoCoordinate>
], 3.VIII.1960, K. Fujimoto (NIAES); 1♂, Shikoku, Mt. Sara [
<geoCoordinate id="219DB05D7F9C0CDE98BF99ECA7BBE9A6" direction="north" orientation="latitude" precision="925" value="33.75">33°45'N</geoCoordinate>
,
<geoCoordinate id="4949EDF027526CD92791CFD1773C36B6" direction="east" orientation="longitude" precision="925" value="133.5">133°30'E</geoCoordinate>
], 27.VII.1955, Toshiro Yano (NIAES); 1♂, Yamanashi, Shojiko [
<geoCoordinate id="DDAF420E03AFE8113500408CC419DC92" direction="north" orientation="latitude" precision="925" value="35.483334">35°29'N</geoCoordinate>
,
<geoCoordinate id="ECCE828E784A05D2AB9E1C883809381E" direction="east" orientation="longitude" precision="925" value="138.6">138°36'E</geoCoordinate>
], 23.VII.1969, P. H. Hasegawa (NIAES); 1f#, Fukushima, Yokomuki Onsen [
<geoCoordinate id="12105536AF9FC8EF376344A079089656" direction="north" orientation="latitude" precision="925" value="37.883335">37°53'N</geoCoordinate>
,
<geoCoordinate id="00F13EFB7ECC6DF0D753A07F3AB5512C" direction="east" orientation="longitude" precision="925" value="140.55">140°33'E</geoCoordinate>
], 26.VII.1964, H. Kobayashi (NIAES).
</paragraph>
</subSubSection>
<subSubSection id="D9EF95DE75ECC2A463495939DBD5A08F" pageId="1" pageNumber="142" type="distribution">
<paragraph id="C03E66D32A7CB93F2F946B601490742A" pageId="1" pageNumber="142">Distribution.</paragraph>
<paragraph id="506B8D9D9A213B71180889A781458C64" pageId="1" pageNumber="142">Japan (Honshu, Kyushu, Shikoku).</paragraph>
</subSubSection>
<subSubSection id="AD2A39EC035C5DCE747393EDDA46DAE4" pageId="1" pageNumber="142" type="remarks">
<paragraph id="B2DFFA2E45EEC0C42C3CC80B77F7A3F0" pageId="1" pageNumber="142">Remarks.</paragraph>
<paragraph id="0D272748107F023140FE92A5FC254CBF" pageId="1" pageNumber="142">
This species appears to be very closely related to
<taxonomicName id="0DC1E3BA10E405B170F06DEA3C6CA2FF" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
Nakahara in having similar wing marking patterns and male genitalia. The only difference between these two Japanese species refers to the male ectoproct. In
<taxonomicName id="7C5EA122643E34D16197353AB428B1E2" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
, the male ectoproct distally bears three projections which are sparsely haired. However, in
<taxonomicName id="3EF8553B38C87FD69A2A00C5920C27A5" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
the ectoproct has a pair of parallel projections which are truncate and serrate at tip. Primarily we even suspected
<taxonomicName id="AC0AE6148D04786B868D1B37F6B99062" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
to be conspecific with
<taxonomicName id="FC077DD0A59304B010B317CBF434DC38" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
due to the remarkable similarity of the external morphological and genitalic characters. Nevertheless, we examined several males of both species, and the difference of the male ectoproct mentioned above is stable. Therefore, we consider
<taxonomicName id="3FFB1607549BD2461E41A7E00E2FD161" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
and
<taxonomicName id="5C64C48BD4A8CBE5E4E1885FC844BF5A" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
to be different species.
<taxonomicName id="273C807345FCB892144E1E0131297A24" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
is a widely distributed species in Honshu, Kyushu and Shikoku, while
<taxonomicName id="083756831D596B601511975E99CFE495" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
seems to be endemic to Kyushu, and more exactly to the Mt. Hikosan.
</paragraph>
<caption id="BA3C43E513392AC8DAEFABD5E78090BB" pageId="1" pageNumber="142">
<paragraph id="2D813F326D99A4274C77D67EDDB28B68" pageId="1" pageNumber="142">
Figures 1-6. Adults of
<taxonomicName id="2ED86FBED14BAA18B02CA2F071351AE6" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="genus">Dilar</taxonomicName>
spp. 1.
<taxonomicName id="6528C3582B00D74B00C5B54A65076765" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
McLachlan, holotype male; 2.
<taxonomicName id="01BDD3848B12A180B7BDEC1CC301C168" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar hikosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="hikosanus">Dilar hikosanus</taxonomicName>
Nakahara, male; 3.
<taxonomicName id="B92811C2E8CCBCA1BDC21CE5E1C12347" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar taiwanensis" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="taiwanensis">Dilar taiwanensis</taxonomicName>
Banks, holotype male; 4.
<taxonomicName id="3EEA1573205A0391427E02BB22F10441" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar formosanus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="formosanus">Dilar formosanus</taxonomicName>
(Okamoto &amp; Kuwayama), holotype female, fore- and hindwing; 5.
<taxonomicName id="8952CFBF7E0B3F4C67183280E9A1C726" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar pallidus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="pallidus">Dilar pallidus</taxonomicName>
Nakahara, holotype male; 6.
<taxonomicName id="66E0DD664CC4E296EF199F5EBE0F658C" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar insularis" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="insularis">Dilar insularis</taxonomicName>
sp. n., holotype male. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="82E67CDF12DFDB3A93047A12FA69CF23" pageId="1" pageNumber="142">
<paragraph id="AAF5AE9B7A2759F8C75DD575E4EAF3D4" pageId="1" pageNumber="142">
Figures 7-12.
<taxonomicName id="EBE632E5CFCD9219FF8A5307BB185966" class="Insecta" family="Dilaridae" genus="Dilar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dilar japonicus" order="Neuroptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="japonicus">Dilar japonicus</taxonomicName>
McLachlan. 7. Male genitalia, dorsal view; 8. Male genitalia, ventral view; 9. Male genitalia, lateral view; 10. Male ectoproct, caudal view; 11. Female genitalia, lateral view; 12. Female genitalia, ventral view. bc: bursa copulatrix; cg: colleterial gland; e: ectoproct; g: gonarcus = eleventh gonocoxites; gx9: ninth gonocoxite; gx10: tenth gonocoxite; hi: hypandrium internum; S7-9: sternite 7-9; T7-9: tergite 7-9. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>