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<document id="B1DB5B4327795D8540B30C3A5CA988F4" ID-CLB-Dataset="110554" ID-DOI="10.1649/0010-065X-69.mo4.167" ID-GBIF-Dataset="459316b8-ecf9-4bcb-a14b-100658d73012" ID-ISSN="1938-4394" ID-Zenodo-Dep="4911739" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095475750" checkinUser="carolina" docAuthor="Flores, Gustavo E. &amp; Aballay, Fernando H." docDate="2015" docId="182B9F50FFECFFDDFD22FB66FDDAFB7B" docLanguage="en" docName="TheColeopBull.69.mo14.167-179.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Thinobatis Eschscholtz 1831" docType="treatment" docVersion="7" lastPageNumber="172" masterDocId="E412E728FFE8FFD8FFDCFFAFFFCCFFEA" masterDocTitle="Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson" masterLastPageNumber="179" masterPageNumber="167" pageNumber="171" updateTime="1699002214655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D3F011750395DBE731776C4C34B77CD3">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:namePart id="7181D3420E46512735637494E59DB0F7">Flores, Gustavo E.</mods:namePart>
<mods:affiliation id="A50692EC7F6B3A3564C47525C39966DD">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:namePart id="F190694FF2546BA792981583E2F03303">Aballay, Fernando H.</mods:namePart>
<mods:affiliation id="082D33DFFF3807620057582B6D52F910">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:title id="F1289D581CBFD258EC6A99696B09B98E">The Coleopterists Bulletin</mods:title>
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<mods:date id="A6B74B8C4FF95C1EC2452E72030D2718">2015</mods:date>
<mods:detail id="70507BA96CC8F6EC712D8268F209FE58" type="series">
<mods:title id="976410245FD8A184BE6129AD7C2B445D">mo 14</mods:title>
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<mods:number id="F585CFC455311DC889BC607CEF8839D9">2015-12-18</mods:number>
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<mods:detail id="CF81F36B71B2B99BBEFA48816D5C5224" type="volume">
<mods:number id="15877CC0C49B1829FBCC801B7BF01C5A">69</mods:number>
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<treatment id="182B9F50FFECFFDDFD22FB66FDDAFB7B" ID-DOI="http://doi.org/10.5281/zenodo.6941554" ID-GBIF-Taxon="198139697" ID-Zenodo-Dep="6941554" LSID="urn:lsid:plazi:treatment:182B9F50FFECFFDDFD22FB66FDDAFB7B" httpUri="http://treatment.plazi.org/id/182B9F50FFECFFDDFD22FB66FDDAFB7B" lastPageId="5" lastPageNumber="172" pageId="4" pageNumber="171">
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<heading id="CB75992AFFECFFDCFD22FB66FBA2FB34" box="[766,1134,1225,1246]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="171" reason="2">
<emphasis id="A2F6F254FFECFFDCFD22FB66FBA2FB34" bold="true" box="[766,1134,1225,1246]" pageId="4" pageNumber="171">
Genus
<taxonomicName id="578255C5FFECFFDCFC9AFB66FBA2FB34" authority="Eschscholtz, 1831" authorityName="Eschscholtz" authorityYear="1831" box="[838,1134,1225,1246]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="A2F6F254FFECFFDCFC9AFB66FC63FB34" bold="true" box="[838,943,1225,1246]" italics="true" pageId="4" pageNumber="171">Thinobatis</emphasis>
<bibRefCitation id="F41353B7FFECFFDCFC64FB66FBA2FB34" author="Eschscholtz" box="[952,1134,1225,1246]" firstAuthor="Eschscholtz" pageId="4" pageNumber="171" refId="ref8671" refString="Eschscholtz, J. F. 1831. Zoologischer Atlas, enthaltend Abbildungen und Beschreibungen neuer Theirarten, wahrend des Flottcapitains von Kotzebue zweiter Reise um die Welt, auf der Russisch-Kaiserlichen Kriegsschlupp Predpriaetie in the Jahren 1823 - 1826. Viertes Heft, G. Reimer, Berlin, Germany." type="book" year="1831">Eschscholtz, 1831</bibRefCitation>
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<paragraph id="903D2E46FFECFFDCFC8FFB4CFBD5FB12" blockId="4.[766,1134,1225,1272]" box="[851,1049,1251,1272]" pageId="4" pageNumber="171">
(
<figureCitation id="08B932C3FFECFFDCFC86FB4CFC53FB12" box="[858,927,1251,1272]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="4" pageNumber="171">Figs. 1</figureCitation>
,
<figureCitation id="08B932C3FFECFFDCFC70FB4CFC18FB12" box="[940,980,1251,1272]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="4" pageNumber="171">5, 7</figureCitation>
,
<figureCitation id="08B932C3FFECFFDCFC3CFB4CFC22FB12" box="[992,1006,1251,1272]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="4" pageNumber="171">9</figureCitation>
, 11)
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFECFFDCFD1FFAA1FC83FA37" pageId="4" pageNumber="171" type="reference_group">
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<treatmentCitationGroup id="B0920968FFECFFDCFD1FFAA1FC83FA37" pageId="4" pageNumber="171">
<treatmentCitation id="11230857FFECFFDCFD1FFAA1FC39FAC9" author="Eschscholtz" box="[707,1013,1294,1315]" page="8" pageId="4" pageNumber="171" year="1831">
<taxonomicName id="578255C5FFECFFDCFD1FFAA1FC39FAC9" authority="Eschscholtz 1831: 8" authorityName="Eschscholtz" authorityPageNumber="8" authorityYear="1831" box="[707,1013,1294,1315]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="A2F6F254FFECFFDCFD1FFAA1FCE5FAC9" box="[707,809,1294,1315]" italics="true" pageId="4" pageNumber="171">Thinobatis</emphasis>
<bibRefCitation id="F41353B7FFECFFDCFCECFAA1FC39FAC9" author="Eschscholtz" box="[816,1013,1294,1315]" firstAuthor="Eschscholtz" page="8" pageId="4" pageNumber="171" refId="ref8671" refString="Eschscholtz, J. F. 1831. Zoologischer Atlas, enthaltend Abbildungen und Beschreibungen neuer Theirarten, wahrend des Flottcapitains von Kotzebue zweiter Reise um die Welt, auf der Russisch-Kaiserlichen Kriegsschlupp Predpriaetie in the Jahren 1823 - 1826. Viertes Heft, G. Reimer, Berlin, Germany." type="book" year="1831">Eschscholtz 1831: 8</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="11230857FFECFFDCFBDDFAA1FB69FAC9" author="Solier" box="[1025,1189,1294,1315]" page="407" pageId="4" pageNumber="171" year="1835">
<bibRefCitation id="F41353B7FFECFFDCFBDDFAA1FB69FAC9" author="Solier" box="[1025,1189,1294,1315]" firstAuthor="Solier" page="407" pageId="4" pageNumber="171" pagination="249 - 419" refId="ref9577" refString="Solier, A. J. J. 1835. Essai sur les collapterides (suite). Annales de la Societe Entomologique de France 4: 249 - 419." type="journal article" year="1835">Solier 1835: 407</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFECFFDCFD00FA87FC42FAD7" author="Laporte &amp; (Comte de Castelnau &amp; Historie Naturelle des Insectes" box="[732,910,1320,1341]" page="196" pageId="4" pageNumber="171" year="1840">
<bibRefCitation id="F41353B7FFECFFDCFD00FA87FC42FAD7" author="Laporte" box="[732,910,1320,1341]" firstAuthor="Laporte" page="196" pageId="4" pageNumber="171" refId="ref9362" refString="Laporte, F. L. N. C. (Comte de Castelnau). 1840. Historie Naturelle des Insectes, vol. 2. P. Dumenil, Paris, France." type="book" year="1840">Laporte 1840: 196</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFECFFDCFC44FA87FBF5FAD7" author="Solier &amp; Fauna chilena &amp; Coleopteros &amp; In &amp; Historia Fisica &amp; Politica de Chile" box="[920,1081,1320,1341]" page="126" pageId="4" pageNumber="171" year="1851">
<bibRefCitation id="F41353B7FFECFFDCFC44FA87FBF5FAD7" author="Solier" box="[920,1081,1320,1341]" firstAuthor="Solier" page="126" pageId="4" pageNumber="171" pagination="124 - 285" refId="ref9608" refString="Solier, A. J. J. 1851. Fauna chilena: Coleopteros [pp. 124 - 285]. In: Historia Fisica y Politica de Chile. Zoologia, Vol. 5. (C. Gay, editor). Imprenta de Maulde et Renou, Paris, France." type="journal article" year="1851">Solier 1851: 126</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFECFFDCFB9FFA87FCF4FAB2" author="Lacordaire &amp; Histoire Naturelle des Insectes" page="65" pageId="4" pageNumber="171" year="1859">
<bibRefCitation id="F41353B7FFECFFDCFB9FFA87FCF4FAB2" author="Lacordaire" firstAuthor="Lacordaire" page="65" pageId="4" pageNumber="171" pagination="401 - 750" refId="ref9206" refString="Lacordaire, J. T. 1859. Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu' ici dans cet ordre d' insectes. Tome cinquieme. Contenant les familles des Tenebrionides, Cistelides, Nilionides, Pythides, Melandryides, Lagriides, Pedilides, Anthicides, Pyrochroides, Mordellides, Rhipiphorides, Stylopides, Meloides et Oedemerides. Premiere partie ([1] + pp. 1 - 400), Deuxieme partie ([1] + pp. 401 - 750). Librairie Encyclopedique de Roret, Paris, France." type="journal article" year="1859">Lacordaire 1859: 65</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFECFFDCFC98FAECFB64FAB2" author="Gemminger &amp; Harold &amp; Catalogus Coleopterorum" box="[836,1192,1347,1368]" page="1836" pageId="4" pageNumber="171" year="1870">
<bibRefCitation id="F41353B7FFECFFDCFC98FAECFB64FAB2" author="Gemminger and Harold" box="[836,1192,1347,1368]" firstAuthor="Gemminger" page="1836" pageId="4" pageNumber="171" pagination="1801 - 2180" refId="ref9076" refString="Gemminger, M., and E. Harold. 1870. Catalogus Coleopterorum hujusque descriptorum synonymiscus et sistematicus, 7: 1801 - 2180. S. Gummi, Monachii." type="journal article" year="1870">Gemminger and Harold 1870: 1836</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFECFFDCFC9CFAF1FC34FA99" author="Philippi" box="[832,1016,1374,1395]" page="722" pageId="4" pageNumber="171" year="1887">
<bibRefCitation id="F41353B7FFECFFDCFC9CFAF1FC34FA99" author="Philippi" box="[832,1016,1374,1395]" firstAuthor="Philippi" page="722" pageId="4" pageNumber="171" pagination="619 - 806" refId="ref9550" refString="Philippi, F. H. 1887. Catalogo de los coleopteros de Chile. Anales de la Universidad de Chile 71: 619 - 806." type="journal article" year="1887">Philippi 1887: 722</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFECFFDCFBB8FAF1FCE7FA67" author="Gebien &amp; Tenebrionidae II &amp; In &amp; Coleopterorum Catalogus XVII" page="19" pageId="4" pageNumber="171" year="1910">
<bibRefCitation id="F41353B7FFECFFDCFBB8FAF1FCE7FA67" author="Gebien" firstAuthor="Gebien" page="19" pageId="4" pageNumber="171" pagination="167 - 354" refId="ref9004" refString="Gebien, H. 1910. Tenebrionidae II [pp. 167 - 354]. In: Coleopterorum Catalogus XVII. Pars 22. (W. Junk and S. Schenkling, editors). W. Junk, Berlin, Germany." type="journal article" year="1910">Gebien 1910: 19</bibRefCitation>
</treatmentCitation>
(catalog), 1937: 588 (catalog);
<treatmentCitation id="11230857FFECFFDCFBE6FAD7FC8FFA42" author="Blackwelder" page="513" pageId="4" pageNumber="171" year="1945">
<bibRefCitation id="F41353B7FFECFFDCFBE6FAD7FC8FFA42" author="Blackwelder" firstAuthor="Blackwelder" page="513" pageId="4" pageNumber="171" pagination="343 - 550" refId="ref8449" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 513</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFECFFDCFC69FA3CFBA5FA42" author="Kulzer" box="[949,1129,1427,1448]" page="903" pageId="4" pageNumber="171" year="1956">
<bibRefCitation id="F41353B7FFECFFDCFC69FA3CFBA5FA42" author="Kulzer" box="[949,1129,1427,1448]" firstAuthor="Kulzer" page="903" pageId="4" pageNumber="171" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer 1956: 903</bibRefCitation>
</treatmentCitation>
(revision);
<treatmentCitation id="11230857FFECFFDCFCCBFA02FC7FFA28" author="Freude" box="[791,947,1453,1474]" page="24" pageId="4" pageNumber="171" year="1960">
<bibRefCitation id="F41353B7FFECFFDCFCCBFA02FC7FFA28" author="Freude" box="[791,947,1453,1474]" firstAuthor="Freude" page="24" pageId="4" pageNumber="171" pagination="24 - 34" refId="ref8978" refString="Freude, H. 1960. Revision der Thinobatini (Col. Tenebrionidae). Mitteilungen der Munchner Entomologischen Gesellschaft 50: 24 - 34." type="journal article" year="1960">Freude 1960: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFECFFDCFC61FA02FB83FA28" author="Pena" box="[957,1103,1453,1474]" page="405" pageId="4" pageNumber="171" year="1966">
<bibRefCitation id="F41353B7FFECFFDCFC61FA02FB83FA28" author="Pena" box="[957,1103,1453,1474]" firstAuthor="Pena" page="405" pageId="4" pageNumber="171" pagination="397 - 453" refId="ref9448" refString="Pena, L. E. 1966. Catalogo de los Tenebrionidae (Coleoptera) de Chile. Entomologische Arbeiten aus dem Museum George Frey 17: 397 - 453." type="journal article" year="1966">Peña 1966: 405</bibRefCitation>
</treatmentCitation>
(catalog), 1974b: 243.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFECFFDCFC88FA67FBDBFA1D" pageId="4" pageNumber="171" type="type_taxon">
<paragraph id="903D2E46FFECFFDCFC88FA67FBDBFA1D" blockId="4.[707,1194,1294,1527]" pageId="4" pageNumber="171">
<typeStatus id="4F3990E4FFECFFDCFC88FA67FC4FFA37" box="[852,899,1480,1501]" pageId="4" pageNumber="171">Type</typeStatus>
species:
<taxonomicName id="578255C5FFECFFDCFC05FA67FC45FA1D" authority="Eschscholtz, 1831" authorityName="Eschscholtz" authorityYear="1831" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="A2F6F254FFECFFDCFC05FA67FB65FA37" box="[985,1193,1480,1501]" italics="true" pageId="4" pageNumber="171">Thinobatis ferruginea</emphasis>
<bibRefCitation id="F41353B7FFECFFDCFD00FA4DFC45FA1D" author="Eschscholtz" box="[732,905,1506,1527]" firstAuthor="Eschscholtz" pageId="4" pageNumber="171" refId="ref8671" refString="Eschscholtz, J. F. 1831. Zoologischer Atlas, enthaltend Abbildungen und Beschreibungen neuer Theirarten, wahrend des Flottcapitains von Kotzebue zweiter Reise um die Welt, auf der Russisch-Kaiserlichen Kriegsschlupp Predpriaetie in the Jahren 1823 - 1826. Viertes Heft, G. Reimer, Berlin, Germany." type="book" year="1831">Eschscholtz, 1831</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFECFFDDFD07F9A2FDB3FD59" lastPageId="5" lastPageNumber="172" pageId="4" pageNumber="171" type="description">
<paragraph id="903D2E46FFECFFDDFD07F9A2FDB3FD59" blockId="4.[707,1193,1549,1729]" lastBlockId="5.[151,639,194,1169]" lastPageId="5" lastPageNumber="172" pageId="4" pageNumber="171">
<emphasis id="A2F6F254FFECFFDCFD07F9A2FC9BF9C8" bold="true" box="[731,855,1549,1570]" pageId="4" pageNumber="171">Description.</emphasis>
In addition to the characters stated above: Mandibles thick, base 2X wider than apex, both mandibles with dorsal cusp on basal half, shorter on right than on left mandible (
<figureCitation id="08B932C3FFECFFDCFBF3F9F2FBA4F998" box="[1071,1128,1629,1650]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="4" pageNumber="171">Fig. 5</figureCitation>
), lacking ventral fossa; labrum exposed, entirely sclerotized; clypeus short, not covering labrum, anterior margin straight or concave, nearly at same level with antennal insertions, lacking row of teeth (
<figureCitation id="08B932C3FFEDFFDDFDE8FF6DFDBCFF3D" box="[564,624,194,215]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 5</figureCitation>
); prementum partially sclerotized, base concealed beneath mentum; anterior margin of mentum narrowly notched (
<figureCitation id="08B932C3FFEDFFDDFEF3FEBEFEA5FECC" box="[303,361,273,294]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 7</figureCitation>
); eyes small, not protruding, upper margin lacking carina; length of epicanthus 1.5X length of eye (
<figureCitation id="08B932C3FFEDFFDDFE8BFEE9FE5DFEB1" box="[343,401,326,347]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 5</figureCitation>
); antennae reaching posterior margin of pronotum, antennomere 1 as long as 3, emerging from distal half of antennal insertion (
<figureCitation id="08B932C3FFEDFFDDFF16FE3AFEC8FE40" box="[202,260,405,426]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 7</figureCitation>
), antennomere 3 longer than 2 (
<figureCitation id="08B932C3FFEDFFDDFDEBFE3AFDBDFE40" box="[567,625,405,426]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 5</figureCitation>
), antennomeres 9 and 10 triangular, strongly expanded outwards, wider than long, antennomere 11 equal to or longer than 10 (
<figureCitation id="08B932C3FFEDFFDDFE93FE4AFE47FE10" box="[335,395,485,506]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">Fig. 9</figureCitation>
). Pronotum with anterior angles acute, directed forwards, or rounded sloping towards venter (
<figureCitation id="08B932C3FFEDFFDDFEECFDB5FEABFDC5" box="[304,359,538,559]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 1</figureCitation>
). Metasternum short, as long as mesosternum. Metacoxae separated by less than half metacoxal width; metacoxal width 3X metacoxal length; distance between meso– and metacoxae equal to metacoxal length (
<figureCitation id="08B932C3FFEDFFDDFE2CFD2CFDF7FD72" box="[496,571,643,664]" captionStart="Figs" captionStartId="4.[731,771,973,992]" captionTargetBox="[707,1193,194,950]" captionTargetId="figure-325@4.[707,1194,193,950]" captionTargetPageId="4" captionText="Figs. 11–12. Meso– and metasterna. 11) Thinobatis rufipes rufipes; 12) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.4911745" httpUri="https://zenodo.org/record/4911745/files/figure.png" pageId="5" pageNumber="172">Fig. 11</figureCitation>
). First tarsomere of hind legs longer than 2+3 combined.
</paragraph>
</subSubSection>
<caption id="C4FD7ECEFFECFFDCFF19F939FE61F92B" ID-DOI="http://doi.org/10.5281/zenodo.6941556" ID-Zenodo-Dep="6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="4" pageNumber="171" startId="4.[197,237,1686,1705]" targetBox="[173,659,923,1663]" targetPageId="4">
<paragraph id="903D2E46FFECFFDCFF19F939FE61F92B" blockId="4.[173,659,1686,1729]" pageId="4" pageNumber="171">
<emphasis id="A2F6F254FFECFFDCFF19F939FEE6F943" bold="true" box="[197,298,1686,1705]" pageId="4" pageNumber="171">Figs. 9–10.</emphasis>
Antennomeres 9–11.
<emphasis id="A2F6F254FFECFFDCFE36F939FE32F943" bold="true" box="[490,510,1686,1705]" pageId="4" pageNumber="171">9)</emphasis>
<taxonomicName id="578255C5FFECFFDCFDDDF939FF2AF92A" authority="rufipes" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="rufipes" subSpecies="rufipes">
<emphasis id="A2F6F254FFECFFDCFDDDF939FF2AF92A" italics="true" pageId="4" pageNumber="171">Thinobatis rufipes rufipes</emphasis>
</taxonomicName>
;
<emphasis id="A2F6F254FFECFFDCFF2EF901FEDCF92B" bold="true" box="[242,272,1710,1729]" pageId="4" pageNumber="171">10)</emphasis>
<taxonomicName id="578255C5FFECFFDCFECAF901FE64F92A" box="[278,424,1710,1728]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="profana">
<emphasis id="A2F6F254FFECFFDCFECAF901FE64F92A" box="[278,424,1710,1728]" italics="true" pageId="4" pageNumber="171">Vaniosus profana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="D8987DCDFFEDFFDDFF73FD17FEE8FC56" pageId="5" pageNumber="172" type="discussion">
<paragraph id="903D2E46FFEDFFDDFF73FD17FEE8FC56" blockId="5.[151,639,194,1169]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFF73FD17FEBFFD27" bold="true" box="[175,371,696,717]" pageId="5" pageNumber="172">Species Included.</emphasis>
<emphasis id="A2F6F254FFEDFFDDFE5DFD16FDB2FD24" box="[385,638,697,718]" italics="true" pageId="5" pageNumber="172">T h i n ob a t i s f e r r u g i ne a</emphasis>
Eschscholtz,
<taxonomicName id="578255C5FFEDFFDDFEFEFD7CFDB6FD02" authority="rufipes Solier" authorityName="Solier" box="[290,634,723,744]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="subSpecies" species="rufipes" subSpecies="rufipes">
<emphasis id="A2F6F254FFEDFFDDFEFEFD7CFDFCFD02" box="[290,560,723,744]" italics="true" pageId="5" pageNumber="172">Thinobatis rufipes rufipes</emphasis>
Solier
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFF4BFD41FE31FCE8" authority="Freude" authorityName="Freude" box="[151,509,749,771]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="subSpecies" species="rufipes" subSpecies="penai">
<emphasis id="A2F6F254FFEDFFDDFF4BFD41FE68FCE9" box="[151,420,750,771]" italics="true" pageId="5" pageNumber="172">Thinobatis rufipes penai</emphasis>
Freude
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFDD2FD41FE5FFCF7" authority="Waterhouse" authorityName="Waterhouse" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="A2F6F254FFEDFFDDFDD2FD41FED8FCF7" italics="true" pageId="5" pageNumber="172">Thinobatis rotundicollis</emphasis>
Waterhouse
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFE7CFCA7FEAAFCD2" authority="Philippi and Philippi" authorityName="Philippi and Philippi" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="A2F6F254FFEDFFDDFE7CFCA7FDB2FCF7" box="[416,638,776,797]" italics="true" pageId="5" pageNumber="172">Thinobatis intermedia</emphasis>
Philippi and Philippi
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFEB3FC8CFDB6FCD2" authority="Kulzer" authorityName="Kulzer" box="[367,634,803,824]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="A2F6F254FFEDFFDDFEB3FC8CFDFCFCD2" box="[367,560,803,824]" italics="true" pageId="5" pageNumber="172">Thinobatis kuscheli</emphasis>
Kulzer
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFF4BFC92FE66FCB8" authority="Kulzer" authorityName="Kulzer" box="[151,426,829,850]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="brevicollis">
<emphasis id="A2F6F254FFEDFFDDFF4BFC92FEAEFCB8" box="[151,354,829,850]" italics="true" pageId="5" pageNumber="172">Thinobatis brevicollis</emphasis>
Kulzer
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFE68FC92FF11FC87" authority="Kulzer" authorityName="Kulzer" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="calderana">
<emphasis id="A2F6F254FFEDFFDDFE68FC92FDB1FCB8" box="[436,637,829,850]" italics="true" pageId="5" pageNumber="172">Thinobatis calderana</emphasis>
Kulzer
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFF36FCF7FDCAFC87" authority="Freude" authorityName="Freude" box="[234,518,856,877]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="melcheri">
<emphasis id="A2F6F254FFEDFFDDFF36FCF7FE79FC87" box="[234,437,856,877]" italics="true" pageId="5" pageNumber="172">Thinobatis melcheri</emphasis>
Freude
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFDCEFCF7FED9FC6D" authority="Pena" authorityName="Pena" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A2F6F254FFEDFFDDFDCEFCF7FF12FC62" italics="true" pageId="5" pageNumber="172">Thinobatis simplex</emphasis>
Peña
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFEC3FCDCFDDCFC6D" authority="Pena" authorityName="Pena" box="[287,528,882,904]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="A2F6F254FFEDFFDDFEC3FCDCFE16FC62" box="[287,474,883,904]" italics="true" pageId="5" pageNumber="172">Thinobatis punctata</emphasis>
Peña
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFDC6FCDCFEE6FC48" authority="Pena" authorityName="Pena" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="A2F6F254FFEDFFDDFDC6FCDCFF26FC48" italics="true" pageId="5" pageNumber="172">Thinobatis confusa</emphasis>
Peña
</taxonomicName>
, and
<taxonomicName id="578255C5FFEDFFDDFEB4FC22FED3FC56" authority="Pena (Pena 1974 b)" authorityName="Pena (Pena" authorityYear="1974" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="A2F6F254FFEDFFDDFEB4FC22FD8CFC48" box="[360,576,909,930]" italics="true" pageId="5" pageNumber="172">Thinobatis arenaria</emphasis>
Peña (
<bibRefCitation id="F41353B7FFEDFFDDFF42FC08FED4FC56" author="Pena" box="[158,280,935,956]" firstAuthor="Pena" pageId="5" pageNumber="172" pagination="243 - 252" refId="ref9514" refString="Pena, L. E. 1974 b. Los tenebrionidos del genero Thinobatis Esch. (Coleoptera: Tenebrionidae). Boletin de la Sociedad de Biologia de Concepcion 48: 243 - 252." type="journal article" year="1974" yearSuffix="b">Peña 1974b</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFF73FC6DFDDAFB7B" pageId="5" pageNumber="172" type="materials_examined">
<paragraph id="903D2E46FFEDFFDDFF73FC6DFDDAFB7B" blockId="5.[151,639,194,1169]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFF73FC6DFEB1FC3D" bold="true" box="[175,381,962,983]" pageId="5" pageNumber="172">Material Examined.</emphasis>
<taxonomicName id="578255C5FFEDFFDDFE57FC6DFDBBFC3D" box="[395,631,962,983]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="A2F6F254FFEDFFDDFE57FC6DFDBBFC3D" box="[395,631,962,983]" italics="true" pageId="5" pageNumber="172">Thinobatis intermedia</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFF4BFC72FEC8FC18" box="[151,260,989,1010]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="melcheri">
<emphasis id="A2F6F254FFEDFFDDFF4BFC72FEC8FC18" box="[151,260,989,1010]" italics="true" pageId="5" pageNumber="172">T. melcheri</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEDFFDDFECCFC72FE7FFC18" authority="rufipes" box="[272,435,989,1010]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="subSpecies" species="rufipes" subSpecies="rufipes">
<emphasis id="A2F6F254FFEDFFDDFECCFC72FE7FFC18" box="[272,435,989,1010]" italics="true" pageId="5" pageNumber="172">T. rufipes rufipes</emphasis>
</taxonomicName>
, and
<taxonomicName id="578255C5FFEDFFDDFE34FC72FF30FBE6" authority="(IADIZA)" baseAuthorityName="IADIZA" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="subSpecies" species="rufipes" subSpecies="penai">
<emphasis id="A2F6F254FFEDFFDDFE34FC72FDB2FC18" box="[488,638,989,1010]" italics="true" pageId="5" pageNumber="172">T. rufipes penai</emphasis>
(IADIZA)
</taxonomicName>
. We also examined
<taxonomicName id="578255C5FFEDFFDDFE63FC58FD9DFBE6" box="[447,593,1015,1036]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="A2F6F254FFEDFFDDFE63FC58FD9DFBE6" box="[447,593,1015,1036]" italics="true" pageId="5" pageNumber="172">T. rotundicollis</emphasis>
</taxonomicName>
, but it should be classified in a distinct genus because of the shape and proportional length and width of the pronotum, rounded pronotal posterior angles, lack of a dorsal cusp on both mandibles, and clypeus and frons with protuberances.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6FD3F33372FD8A7302935C660E0D5282">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:namePart id="25F37C6F52ED942899BFE46BA9214CF8">Flores, Gustavo E.</mods:namePart>
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<emphasis id="A2F6F254FFEDFFDDFD24FAB1FC0CFAD9" bold="true" box="[760,960,1310,1331]" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
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(
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<emphasis id="A2F6F254FFEDFFDDFD19FA27FCE2FA77" bold="true" box="[709,814,1416,1437]" pageId="5" pageNumber="172">Diagnosis.</emphasis>
See
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.
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<emphasis id="A2F6F254FFEDFFDDFD19FA0CFC91FA52" bold="true" box="[709,861,1443,1464]" pageId="5" pageNumber="172">Type Material.</emphasis>
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(
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<figureCitation id="08B932C3FFEDFFDDFCA7FA77FC14FA07" box="[891,984,1495,1517]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 3–4</figureCitation>
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<emphasis id="A2F6F254FFEDFFDDFD19FA5DFC8FF9ED" bold="true" box="[709,835,1522,1543]" pageId="5" pageNumber="172">Distribution.</emphasis>
Known only from the
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locality in
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Genus
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<paragraph id="903D2E46FFEDFFDDFEDEFB51FDDFFAF9" blockId="5.[246,544,1251,1299]" box="[258,531,1277,1299]" pageId="5" pageNumber="172">
(
<figureCitation id="08B932C3FFEDFFDDFED5FB51FEA4FAF9" box="[265,360,1277,1299]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 2–4</figureCitation>
,
<figureCitation id="08B932C3FFEDFFDDFEA8FB51FE57FAF9" box="[372,411,1278,1299]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">6, 8</figureCitation>
,
<figureCitation id="08B932C3FFEDFFDDFE74FB51FE0EFAF9" box="[424,450,1278,1299]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">10</figureCitation>
, 12, 21)
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFF4BFA86FE52FAD7" box="[151,414,1320,1342]" pageId="5" pageNumber="172" type="reference_group">
<paragraph id="903D2E46FFEDFFDDFF4BFA86FE52FAD7" blockId="5.[151,638,1320,1368]" box="[151,414,1320,1342]" pageId="5" pageNumber="172">
<treatmentCitationGroup id="B0920968FFEDFFDDFF4BFA86FE52FAD7" box="[151,414,1320,1342]" pageId="5" pageNumber="172">
<treatmentCitation id="11230857FFEDFFDDFF4BFA86FE56FAD7" author="Kulzer" box="[151,410,1320,1342]" page="896" pageId="5" pageNumber="172" year="1956">
<taxonomicName id="578255C5FFEDFFDDFF4BFA86FE56FAD7" authority="Kulzer 1956: 896" authorityName="Kulzer" authorityPageNumber="896" authorityYear="1956" box="[151,410,1320,1342]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="A2F6F254FFEDFFDDFF4BFA86FF20FAD4" box="[151,236,1321,1342]" italics="true" pageId="5" pageNumber="172">Vaniosus</emphasis>
<bibRefCitation id="F41353B7FFEDFFDDFF2FFA87FE56FAD7" author="Kulzer" box="[243,410,1320,1341]" firstAuthor="Kulzer" page="896" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer 1956: 896</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFE78FA87FDE4FAB2" pageId="5" pageNumber="172" type="type_taxon">
<paragraph id="903D2E46FFEDFFDDFE78FA87FDE4FAB2" blockId="5.[151,638,1320,1368]" pageId="5" pageNumber="172">
<typeStatus id="4F3990E4FFEDFFDDFE78FA87FE1FFAD7" box="[420,467,1320,1341]" pageId="5" pageNumber="172">Type</typeStatus>
species:
<taxonomicName id="578255C5FFEDFFDDFDF4FA86FE54FAB2" authority="Kulzer, 1956" authorityName="Kulzer" authorityYear="1956" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="A2F6F254FFEDFFDDFDF4FA86FED8FAB2" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
<bibRefCitation id="F41353B7FFEDFFDDFEC0FAECFE54FAB2" author="Kulzer" box="[284,408,1347,1368]" firstAuthor="Kulzer" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer, 1956</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFF73FAC1FBD2FD00" pageId="5" pageNumber="172" type="description">
<paragraph id="903D2E46FFEDFFDDFF73FAC1FC77FDDA" blockId="5.[151,638,1390,1729]" lastBlockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFF73FAC1FEE7FA69" bold="true" box="[175,299,1390,1411]" pageId="5" pageNumber="172">Description.</emphasis>
In addition to the characters stated above: Mandibles finer, acute, base 1.5X wider than the apex, both mandibles with a dorsal cusp on basal half longer on the right than on the left mandible (
<figureCitation id="08B932C3FFEDFFDDFEDDFA77FEF3FA07" box="[257,319,1496,1517]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
), both mandibles with a ventral fossa (
<figureCitation id="08B932C3FFEDFFDDFF07FA5DFED5F9ED" box="[219,281,1522,1543]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
); labrum exposed, partially sclerotized; clypeus short, not covering the labrum, anterior margin triangular nearly at same level with antennal insertions, armed with a row of teeth directed downwards, in the apex with a large tooth prolonged anterad (
<figureCitation id="08B932C3FFEDFFDDFEBCF9D8FE6CF966" box="[352,416,1655,1676]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave (
<figureCitation id="08B932C3FFEDFFDDFD37FF6DFCE5FF3D" box="[747,809,194,215]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
); eyes small, protruding out, upper margin lacking carina; length of epicanthus 2.5x the length of eye (
<figureCitation id="08B932C3FFEDFFDDFC86FF58FC58FEE6" box="[858,916,247,268]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 6</figureCitation>
); antennae surpassing posterior margin of pronotum, antennomere 1 sticks out distal half of antennal insertion (
<figureCitation id="08B932C3FFEDFFDDFC33FE83FBE7FEAB" box="[1007,1067,300,321]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
), antennomeres 9 and 10 cylindrical, not expanded outwards, longer than wide, antennomere 11 equal in length to 10 (
<figureCitation id="08B932C3FFEDFFDDFC8CFED3FC56FE7B" box="[848,922,380,401]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">Fig. 10</figureCitation>
). Pronotum with anterior angles obtuse, slopeing towards venter (
<figureCitation id="08B932C3FFEDFFDDFBF0FE38FB44FE46" box="[1068,1160,406,428]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 2–4</figureCitation>
). Metasternum twice the length of mesosternum. Metacoxae separated by less than half metacoxal width; metacoxal width 3X metacoxal length; distance between meso– and metacoxae twice the metacoxal length (
<figureCitation id="08B932C3FFEDFFDDFCB9FDB4FC63FDDA" box="[869,943,539,560]" captionStart="Figs" captionStartId="4.[731,771,973,992]" captionTargetBox="[707,1193,194,950]" captionTargetId="figure-325@4.[707,1194,193,950]" captionTargetPageId="4" captionText="Figs. 11–12. Meso– and metasterna. 11) Thinobatis rufipes rufipes; 12) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.4911745" httpUri="https://zenodo.org/record/4911745/files/figure.png" pageId="5" pageNumber="172">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="903D2E46FFEDFFDDFD19FD99FBD2FD00" blockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
Female genitalia (
<figureCitation id="08B932C3FFEDFFDDFCA1FD99FC06FDA1" box="[893,970,566,587]" captionStart="Figs" captionStartId="9.[175,215,1662,1681]" captionTargetBox="[151,1171,1074,1639]" captionTargetId="figure-36@9.[151,1173,1074,1640]" captionTargetPageId="9" captionText="Figs. 21–22. Ovipositor in ventral view, spiculum and internal female reproductive tract. 21) Vaniosus profana; 22) Achanius (Ambigatus) rufonitens. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911753" httpUri="https://zenodo.org/record/4911753/files/figure.png" pageId="5" pageNumber="172">Fig. 21</figureCitation>
). Spiculum without arms. Paraprocts long (2.0 &lt;P/C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, with annulate duct. Spermatheca single, compact, locular, at the end of vagina, near the duct of spermathecal accessory gland.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFD19FD5FFB4FFC8B" pageId="5" pageNumber="172" type="discussion">
<paragraph id="903D2E46FFEDFFDDFD19FD5FFC13FCF5" blockId="5.[685,1173,194,800]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFD19FD5FFCB9FCEF" bold="true" box="[709,885,752,773]" pageId="5" pageNumber="172">Species included.</emphasis>
<taxonomicName id="578255C5FFEDFFDDFC5EFD5FFB58FCEF" authority="Kulzer" authorityName="Kulzer" authorityYear="1956" box="[898,1172,752,773]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="A2F6F254FFEDFFDDFC5EFD5FFB86FCEF" box="[898,1098,752,773]" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
Kulzer
</taxonomicName>
and
<taxonomicName id="578255C5FFEDFFDDFD04FCA4FC17FCF5" authority="(Kulzer)" baseAuthorityName="Kulzer" box="[728,987,778,800]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana">
<emphasis id="A2F6F254FFEDFFDDFD04FCA4FC4DFCCA" box="[728,897,779,800]" italics="true" pageId="5" pageNumber="172">Vaniosus profana</emphasis>
(Kulzer)
</taxonomicName>
.
</paragraph>
<paragraph id="903D2E46FFEDFFDDFD62FCE4FB4FFC8B" blockId="5.[702,1155,843,867]" box="[702,1155,843,867]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFD62FCE4FC71FC8B" bold="true" box="[702,957,843,866]" pageId="5" pageNumber="172">KEY TO THE SPECIES OF</emphasis>
<taxonomicName id="578255C5FFEDFFDDFC19FCE4FB4FFC8B" authority="KULZER" authorityName="Kulzer, Evaniosomini. Doyen" authorityYear="1994" box="[965,1155,843,867]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="A2F6F254FFEDFFDDFC19FCE4FBE9FC8A" bold="true" box="[965,1061,843,867]" italics="true" pageId="5" pageNumber="172">VANIOSUS</emphasis>
<emphasis id="A2F6F254FFEDFFDDFBF3FCE4FB4FFC8B" bold="true" box="[1071,1155,843,866]" pageId="5" pageNumber="172">KULZER</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172" type="key">
<keyStep id="2B7636E3FFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172">
<key id="E4137977FFEDFFDDFD71FCD2FB58FB19" pageId="5" pageNumber="172">
<paragraph id="903D2E46FFEDFFDDFD71FCD2FB5FFBDB" blockId="5.[685,1172,893,1267]" pageId="5" pageNumber="172">
<keyLead id="2B738D73FFEDFFDDFD71FCD2FB5FFBDB" pageId="5" pageNumber="172">
1. Head widest at epicanthus; antennomere 1 longer than 3, antennomere 2 longer than 3; posterior angles of pronotum very protruding, pointed outwards (
<figureCitation id="08B932C3FFEDFFDDFC7DFC63FC2EFC0B" box="[929,994,972,993]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 4</figureCitation>
); humeri not elevated; hind wings absent; first tarsomere of hind legs longer than 2–4 combined (
<figureCitation id="08B932C3FFEDFFDDFB8DFBADFB47FBFD" box="[1105,1163,1026,1047]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 3</figureCitation>
) .........................
<taxonomicName id="578255C5FFEDFFDDFCA2FBB3FB5FFBDB" authority="Kulzer" authorityName="Kulzer" authorityYear="1956" box="[894,1171,1052,1073]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="A2F6F254FFEDFFDDFCA2FBB3FB5FFBDB" bold="true" box="[894,1171,1052,1073]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFCA2FBB3FB8AFBDB" bold="true" box="[894,1094,1052,1073]" italics="true" pageId="5" pageNumber="172">Vaniosus paradoxus</emphasis>
Kulzer
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="903D2E46FFEDFFDDFD71FB90FB58FB19" blockId="5.[685,1172,893,1267]" pageId="5" pageNumber="172">
<keyLead id="2B738D73FFEDFFDDFD71FB90FB58FB19" pageId="5" pageNumber="172">
1′. Head widest at eyes; antennomere 1 as long as 3, antennomere 2 shorter than 3 (
<figureCitation id="08B932C3FFEDFFDDFB96FBF6FB4AFB84" box="[1098,1158,1113,1134]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">Fig. 8</figureCitation>
); posterior angles of pronotum obtuse, not protruding; humeri elevated; hind wings present, reduced, as long as elytra, not folded; first tarsomere of hind legs as long as 2–3 combined (
<figureCitation id="08B932C3FFEDFFDDFCF8FB71FC97FB19" box="[804,859,1246,1267]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Fig. 2</figureCitation>
).......
<taxonomicName id="578255C5FFEDFFDDFC52FB71FB58FB19" authority="(Kulzer)" baseAuthorityName="Kulzer" box="[910,1172,1246,1267]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana">
<emphasis id="A2F6F254FFEDFFDDFC52FB71FB58FB19" bold="true" box="[910,1172,1246,1267]" pageId="5" pageNumber="172">
<emphasis id="A2F6F254FFEDFFDDFC52FB71FBFAFB19" bold="true" box="[910,1078,1246,1267]" italics="true" pageId="5" pageNumber="172">Vaniosus profana</emphasis>
(Kulzer)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>

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<document id="9A1E61F945815771E18402CF9A93B5F0" ID-CLB-Dataset="110554" ID-DOI="10.1649/0010-065X-69.mo4.167" ID-GBIF-Dataset="459316b8-ecf9-4bcb-a14b-100658d73012" ID-ISSN="1938-4394" ID-Zenodo-Dep="4911739" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095475750" checkinUser="carolina" docAuthor="Flores, Gustavo E. &amp; Aballay, Fernando H." docDate="2015" docId="182B9F50FFEDFFDEFD71F9E2FC6CFBE5" docLanguage="en" docName="TheColeopBull.69.mo14.167-179.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Vaniosus profana, new combination" docType="treatment" docVersion="9" lastPageNumber="173" masterDocId="E412E728FFE8FFD8FFDCFFAFFFCCFFEA" masterDocTitle="Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson" masterLastPageNumber="179" masterPageNumber="167" pageNumber="172" updateTime="1699002214655" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75C28ADDFD8E26B094877E7D3706EBE3">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:affiliation id="06A6A1FCE5FFB0AE74584EEBD243F440">CONICET, Laboratorio de Entomología Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT-CONICET Mendoza) Casilla de correo 507, 5500 Mendoza, ARGENTINA</mods:affiliation>
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<mods:namePart id="4B1B05C9186520D34FBF07D037D33E98">Aballay, Fernando H.</mods:namePart>
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<mods:title id="74B311839DBC24B59AB76451B1896263">The Coleopterists Bulletin</mods:title>
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<mods:part id="78EEA476FD8588529568CDE7A366B458">
<mods:date id="065538BEA151D8EEDEA8D056885335F6">2015</mods:date>
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<mods:title id="95C5DFE739B8876AFAD1381EC423FB4E">mo 14</mods:title>
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<treatment id="182B9F50FFEDFFDEFD71F9E2FC6CFBE5" ID-DOI="http://doi.org/10.5281/zenodo.6941563" ID-GBIF-Taxon="198139694" ID-Zenodo-Dep="6941563" LSID="urn:lsid:plazi:treatment:182B9F50FFEDFFDEFD71F9E2FC6CFBE5" httpUri="http://treatment.plazi.org/id/182B9F50FFEDFFDEFD71F9E2FC6CFBE5" lastPageId="6" lastPageNumber="173" pageId="5" pageNumber="172">
<subSubSection id="D8987DCDFFEDFFDDFD71F9E2FB58F988" box="[685,1172,1613,1634]" pageId="5" pageNumber="172" type="nomenclature">
<paragraph id="903D2E46FFEDFFDDFD71F9E2FB58F988" blockId="5.[685,1172,1613,1660]" box="[685,1172,1613,1634]" pageId="5" pageNumber="172">
<heading id="CB75992AFFEDFFDDFD71F9E2FB58F988" box="[685,1172,1613,1634]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="172" reason="2">
<emphasis id="A2F6F254FFEDFFDDFD71F9E2FB58F988" bold="true" box="[685,1172,1613,1634]" pageId="5" pageNumber="172">
<taxonomicName id="578255C5FFEDFFDDFD71F9E2FC28F988" authority="(Kulzer, 1956)" baseAuthorityName="Kulzer" baseAuthorityYear="1956" box="[685,996,1613,1634]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana" status="comb. nov.">
<emphasis id="A2F6F254FFEDFFDDFD71F9E2FC9EF988" bold="true" box="[685,850,1613,1634]" italics="true" pageId="5" pageNumber="172">Vaniosus profana</emphasis>
(
<bibRefCitation id="F41353B7FFEDFFDDFC81F9E2FC11F988" author="Kulzer" box="[861,989,1613,1634]" firstAuthor="Kulzer" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer, 1956</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="B9C54F2FFFEDFFDDFC32F9E2FB58F988" box="[1006,1172,1613,1634]" pageId="5" pageNumber="172" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEDFFDDFCEEF9C8FBC3F996" box="[818,1039,1639,1660]" pageId="5" pageNumber="172" type="description">
<paragraph id="903D2E46FFEDFFDDFCEEF9C8FBC3F996" blockId="5.[685,1172,1613,1660]" box="[818,1039,1639,1660]" pageId="5" pageNumber="172">
(
<figureCitation id="08B932C3FFEDFFDDFCE5F9C8FCB5F996" box="[825,889,1639,1660]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="5" pageNumber="172">Figs. 2</figureCitation>
,
<figureCitation id="08B932C3FFEDFFDDFC5EF9C8FC69F996" box="[898,933,1639,1660]" captionStart="Figs" captionStartId="3.[175,215,1662,1681]" captionTargetBox="[151,1171,864,1639]" captionTargetId="figure-317@3.[151,1173,863,1639]" captionTargetPageId="3" captionText="Figs. 5–8. Head. 5) Thinobatis rufipes rufipes, frontal view; 6) Vaniosus profana, frontal view; 7) Thinobatis rufipes rufipes, mouthparts, ventral view; 8) Vaniosus profana, mouthparts, ventral view, white arrow indicates ventral fossa on mandible." figureDoi="http://doi.org/10.5281/zenodo.4911743" httpUri="https://zenodo.org/record/4911743/files/figure.png" pageId="5" pageNumber="172">6, 8</figureCitation>
,
<figureCitation id="08B932C3FFEDFFDDFC72F9C8FC0BF996" box="[942,967,1639,1660]" captionStart="Figs" captionStartId="4.[197,237,1686,1705]" captionTargetBox="[173,659,923,1663]" captionTargetId="figure-306@4.[172,659,923,1663]" captionText="Figs. 9–10. Antennomeres 9–11.9) Thinobatis rufipes rufipes; 10) Vaniosus profana." figureDoi="http://doi.org/10.5281/zenodo.6941556" httpUri="https://zenodo.org/record/6941556/files/figure.png" pageId="5" pageNumber="172">10</figureCitation>
, 12, 21)
</paragraph>
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<subSubSection id="D8987DCDFFEDFFDDFD71F93DFC1BF92B" pageId="5" pageNumber="172" type="reference_group">
<paragraph id="903D2E46FFEDFFDDFD71F93DFC1BF92B" blockId="5.[685,1172,1682,1729]" pageId="5" pageNumber="172">
<treatmentCitationGroup id="B0920968FFEDFFDDFD71F93DFC1BF92B" pageId="5" pageNumber="172">
<treatmentCitation id="11230857FFEDFFDDFD71F93DFB8CF94D" author="Kulzer" box="[685,1088,1682,1703]" page="906" pageId="5" pageNumber="172" year="1956">
<taxonomicName id="578255C5FFEDFFDDFD71F93DFB8CF94D" authority="Kulzer 1956: 906" authorityName="Kulzer" authorityPageNumber="906" authorityYear="1956" box="[685,1088,1682,1703]" class="Insecta" family="Tenebrionidae" genus="Thinobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="profana">
<emphasis id="A2F6F254FFEDFFDDFD71F93DFCBAF94D" box="[685,886,1682,1703]" italics="true" pageId="5" pageNumber="172">Thinobatis profana</emphasis>
<bibRefCitation id="F41353B7FFEDFFDDFC5DF93DFB8CF94D" author="Kulzer" box="[897,1088,1682,1703]" firstAuthor="Kulzer" page="906" pageId="5" pageNumber="172" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer 1956: 906</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="11230857FFEDFFDDFB91F93DFCD3F92B" author="Freude" page="31" pageId="5" pageNumber="172" year="1960">
<bibRefCitation id="F41353B7FFEDFFDDFB91F93DFCD3F92B" author="Freude" firstAuthor="Freude" page="31" pageId="5" pageNumber="172" pagination="24 - 34" refId="ref8978" refString="Freude, H. 1960. Revision der Thinobatini (Col. Tenebrionidae). Mitteilungen der Munchner Entomologischen Gesellschaft 50: 24 - 34." type="journal article" year="1960">Freude 1960: 31</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFEDFFDDFCF0F903FC1FF92B" author="Pena" box="[812,979,1708,1729]" page="244" pageId="5" pageNumber="172" year="1974">
<bibRefCitation id="F41353B7FFEDFFDDFCF0F903FC1FF92B" author="Pena" box="[812,979,1708,1729]" firstAuthor="Pena" page="244" pageId="5" pageNumber="172" pagination="243 - 252" refId="ref9514" refString="Pena, L. E. 1974 b. Los tenebrionidos del genero Thinobatis Esch. (Coleoptera: Tenebrionidae). Boletin de la Sociedad de Biologia de Concepcion 48: 243 - 252." type="journal article" year="1974" yearSuffix="b">Peña 1974b: 244</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEEFFDEFF19FF6DFE3FFF3D" box="[197,499,194,215]" pageId="6" pageNumber="173" type="diagnosis">
<paragraph id="903D2E46FFEEFFDEFF19FF6DFE3FFF3D" blockId="6.[173,660,194,1729]" box="[197,499,194,215]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFF19FF6DFEE1FF3D" bold="true" box="[197,301,194,215]" pageId="6" pageNumber="173">Diagnosis.</emphasis>
See
<bibRefCitation id="F41353B7FFEEFFDEFEB9FF6DFE23FF3D" author="Kulzer" box="[357,495,194,215]" firstAuthor="Kulzer" pageId="6" pageNumber="173" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer (1956)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEEFFDEFF19FF72FE75F998" pageId="6" pageNumber="173" type="materials_examined">
<paragraph id="903D2E46FFEEFFDEFF19FF72FE39FE9D" blockId="6.[173,660,194,1729]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFF19FF72FE90FF18" bold="true" box="[197,348,221,242]" pageId="6" pageNumber="173">Type Material.</emphasis>
<materialsCitation id="20EA241BFFEEFFDEFEB5FF72FDECFEE6" ID-GBIF-Occurrence="3872861365" collectingDate="1934-12" collectionCode="NHMB" location="Forres" municipality="Forres" pageId="6" pageNumber="173" specimenCount="1" stateProvince="Santiago del Estero" typeStatus="holotype">
<emphasis id="A2F6F254FFEEFFDEFEB5FF72FE09FF18" bold="true" box="[361,453,221,242]" pageId="6" pageNumber="173">
<typeStatus id="4F3990E4FFEEFFDEFEB5FF72FE09FF18" box="[361,453,221,242]" pageId="6" pageNumber="173" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingRegion id="5246E0A4FFEEFFDEFE0FFF72FD5CFF18" box="[467,656,221,242]" country="Argentina" name="Santiago del Estero" pageId="6" pageNumber="173">Santiago del Estero</collectingRegion>
,
<collectingMunicipality id="7059B43CFFEEFFDEFF71FF58FF3CFEE6" box="[173,240,247,268]" name="Forres" pageId="6" pageNumber="173" stateProvince="Santiago del Estero">Forres</collectingMunicipality>
,
<date id="E43C0886FFEEFFDEFF22FF58FE97FEE6" box="[254,347,247,268]" pageId="6" pageNumber="173" value="1934-12">
<collectingDate id="F478F16EFFEEFFDEFF22FF58FE97FEE6" box="[254,347,247,268]" pageId="6" pageNumber="173" value="1934-12">XII-1934</collectingDate>
</date>
(
<collectionCode id="F693B683FFEEFFDEFEB0FF58FE75FEE6" box="[364,441,247,268]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="6" pageNumber="173">NHMB</collectionCode>
) (
<figureCitation id="08B932C3FFEEFFDEFE0EFF58FDD8FEE6" box="[466,532,247,268]" captionStart="Figs" captionStartId="2.[197,237,1581,1600]" captionTargetBox="[185,1193,205,1559]" captionTargetId="figure-1@2.[172,1194,183,1574]" captionTargetPageId="2" captionText="Figs. 1–4. Dorsal habitus. 1) Thinobatis rufipes rufipes; 2) Vaniosus profana, holotype, (Image courtesy of Eva Sprecher, NHMB); 3) Vaniosus paradoxus, holotype (Image courtesy of Isabelle Züercher, NHMB); 4) V. paradoxus, holotype, pronotum and head (Image courtesy of Isabelle Züercher, NHMB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911741" httpUri="https://zenodo.org/record/4911741/files/figure.png" pageId="6" pageNumber="173">Fig. 2</figureCitation>
).
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFDF6FF58FD58FECD" ID-GBIF-Occurrence="3872861340" collectingDate="2021-03-12" collectionCode="NHMB" location="Caspinchango" municipality="Caspinchango" pageId="6" pageNumber="173" specimenCount="1" stateProvince="Catamarca" typeStatus="paratype">
<emphasis id="A2F6F254FFEEFFDEFDF6FF58FD40FEE6" bold="true" box="[554,652,247,268]" pageId="6" pageNumber="173">
<typeStatus id="4F3990E4FFEEFFDEFDF6FF58FD40FEE6" box="[554,652,247,268]" pageId="6" pageNumber="173" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectingRegion id="5246E0A4FFEEFFDEFF71FEBDFED0FECD" box="[173,284,274,295]" country="Argentina" name="Catamarca" pageId="6" pageNumber="173">Catamarca</collectingRegion>
,
<collectingMunicipality id="7059B43CFFEEFFDEFEF6FEBDFE72FECD" box="[298,446,274,295]" country="Argentina" municipality="These" name="Caspinchango" pageId="6" pageNumber="173" stateProvince="Catamarca">Caspinchango</collectingMunicipality>
, 12.III.21 (
<collectionCode id="F693B683FFEEFFDEFDE4FEBDFD45FECD" box="[568,649,274,295]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="6" pageNumber="173">NHMB</collectionCode>
).
</materialsCitation>
<collectingMunicipality id="7059B43CFFEEFFDEFF71FE83FF2BFEAB" box="[173,231,300,321]" pageId="6" pageNumber="173">These</collectingMunicipality>
and all localities mentioned in the original description correspond to
<collectingCountry id="E8956ED6FFEEFFDEFE67FEE8FDD2FEB6" box="[443,542,327,348]" name="Argentina" pageId="6" pageNumber="173">Argentina</collectingCountry>
and not to
<collectingCountry id="E8956ED6FFEEFFDEFF71FECDFF10FE9D" box="[173,220,354,375]" name="Peru" pageId="6" pageNumber="173">Peru</collectingCountry>
, as stated by
<bibRefCitation id="F41353B7FFEEFFDEFEB8FECDFE3DFE9D" author="Kulzer" box="[356,497,354,375]" firstAuthor="Kulzer" pageId="6" pageNumber="173" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer (1956)</bibRefCitation>
.
</paragraph>
<paragraph id="903D2E46FFEEFFDEFF19FED3FE75F998" blockId="6.[173,660,194,1729]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFF19FED3FE1BFE7B" bold="true" box="[197,471,380,401]" pageId="6" pageNumber="173">Other Material Examined.</emphasis>
<specimenCount id="8684E5CFFFEEFFDEFE38FED3FDBBFE7B" box="[484,631,380,401]" pageId="6" pageNumber="173" type="generic">237 specimens</specimenCount>
in IADIZA.
<materialsCitation id="20EA241BFFEEFFDEFECCFE38FEF0FE0B" ID-GBIF-Occurrence="3872861342" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFECCFE38FE7CFE46" bold="true" box="[272,432,407,428]" pageId="6" pageNumber="173">
<collectingCountry id="E8956ED6FFEEFFDEFECCFE38FE67FE46" box="[272,427,407,428]" name="Argentina" pageId="6" pageNumber="173">ARGENTINA</collectingCountry>
:
</emphasis>
<collectingRegion id="5246E0A4FFEEFFDEFE66FE38FE23FE46" box="[442,495,407,428]" country="Argentina" name="Salta" pageId="6" pageNumber="173">Salta</collectingRegion>
: Dto. Chicoana:
<quantity id="577A83A3FFEEFFDEFF71FE1EFF2CFE2C" box="[173,224,433,454]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="173" unit="km" value="2.0">2 km</quantity>
E Escoipe,
<quantity id="577A83A3FFEEFFDEFE86FE1EFE64FE2C" box="[346,424,433,454]" metricMagnitude="3" metricUnit="m" metricValue="1.8559999999999999" pageId="6" pageNumber="173" unit="m" value="1856.0">1856 m</quantity>
,
<date id="E43C0886FFEEFFDEFE68FE1EFDD7FE2C" box="[436,539,433,454]" pageId="6" pageNumber="173" value="2004-11-07">7-XI-2004</date>
, G. Flores - G. Zalazar, 70;
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE98FE63FE68FE11" ID-GBIF-Occurrence="3872861361" pageId="6" pageNumber="173">
Dto. Cafayate: Tolombón,
<quantity id="577A83A3FFEEFFDEFD9FFE63FD5CFE0B" box="[579,656,460,481]" metricMagnitude="3" metricUnit="m" metricValue="1.8119999999999998" pageId="6" pageNumber="173" unit="m" value="1812.0">1812 m</quantity>
,
<date id="E43C0886FFEEFFDEFF71FE49FED7FE11" box="[173,283,486,507]" pageId="6" pageNumber="173" value="2003-12-08">8-XII-2003</date>
, G. Flores, 4.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE70FE49FF07FDDA" ID-GBIF-Occurrence="3872861309" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFE70FE49FDC5FE11" box="[428,521,486,507]" country="Argentina" name="Tucuman" pageId="6" pageNumber="173">Tucumán</collectingRegion>
: Dto. Tafí del Valle: Ruinas de Quilmes,
<date id="E43C0886FFEEFFDEFE1EFDAEFDFDFDFC" box="[450,561,513,534]" pageId="6" pageNumber="173" value="1997-10-17">17-X-1997</date>
, S. Roig, 16.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF0FFDB4FE87FDA1" ID-GBIF-Occurrence="3872861312" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFF0FFDB4FEF6FDDA" box="[211,314,539,560]" country="Argentina" name="Catamarca" pageId="6" pageNumber="173">Catamarca</collectingRegion>
:
<quantity id="577A83A3FFEEFFDEFE99FDB4FEB4FDDA" box="[325,376,539,560]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="173" unit="km" value="2.0">2 km</quantity>
N Villa Vil,
<quantity id="577A83A3FFEEFFDEFE27FDB4FD84FDDA" box="[507,584,539,560]" metricMagnitude="3" metricUnit="m" metricValue="2.358" pageId="6" pageNumber="173" unit="m" value="2358.0">2358 m</quantity>
,
<date id="E43C0886FFEEFFDEFD88FDB4FF13FDA1" pageId="6" pageNumber="173" value="2009-11-16">16-XI- 2009</date>
, G. Flores,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE8FFD99FE85FD8C" ID-GBIF-Occurrence="3872861306" pageId="6" pageNumber="173">
1; Punta de Balasto,
<date id="E43C0886FFEEFFDEFDFCFD99FD43FDA1" box="[544,655,566,587]" pageId="6" pageNumber="173" value="2001-02-03" valueMax="2001-02-05" valueMin="2001-02-03">3-5-II-2001</date>
, G. Arriagada, 1.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE8DFDFEFDEEFD6A" ID-GBIF-Occurrence="3872861366" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFE8DFDFEFE65FD8C" box="[337,425,593,614]" country="Argentina" name="La Rioja" pageId="6" pageNumber="173">La Rioja</collectingRegion>
: Dto. Famatina:
<quantity id="577A83A3FFEEFFDEFD96FDFEFDB1FD8C" box="[586,637,593,614]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="173" unit="km" value="2.0">2 km</quantity>
S Campanas,
<date id="E43C0886FFEEFFDEFEC3FDC4FE55FD6A" box="[287,409,619,640]" pageId="6" pageNumber="173" value="2003-12-10">10-XII-2003</date>
, G. Flores,
<quantity id="577A83A3FFEEFFDEFDCCFDC4FDEEFD6A" box="[528,546,619,640]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="173" unit="km" value="4.1">4,</quantity>
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFDF7FDC4FE01FD71" ID-GBIF-Occurrence="3872861321" pageId="6" pageNumber="173">
10 km NE Pituil,
<date id="E43C0886FFEEFFDEFF36FD29FE9FFD71" box="[234,339,646,667]" pageId="6" pageNumber="173" value="1997-10-15">15-X-1997</date>
, S. Roig, 11;
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE0FFD29FE23FD5F" ID-GBIF-Occurrence="3872861382" pageId="6" pageNumber="173">
Dto. Gral. Belgrano:
<quantity id="577A83A3FFEEFFDEFF71FD0FFF28FD5F" box="[173,228,672,693]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="173" unit="km" value="20.0">20km</quantity>
N Olta,
<date id="E43C0886FFEEFFDEFEF5FD0FFE42FD5F" box="[297,398,672,693]" pageId="6" pageNumber="173" value="1997-10-21">21-X-1997</date>
,S.Roig,3.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE2DFD0FFEDDFD00" ID-GBIF-Occurrence="3872861354" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFE2DFD0FFD8EFD5F" box="[497,578,672,693]" country="Argentina" name="Cordoba" pageId="6" pageNumber="173">Córdoba</collectingRegion>
:
<quantity id="577A83A3FFEEFFDEFD94FD0FFDB3FD5F" box="[584,639,672,693]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="173" unit="km" value="15.0">15km</quantity>
N Capilla del Monte,
<date id="E43C0886FFEEFFDEFEB1FD14FE25FD3A" box="[365,489,699,720]" pageId="6" pageNumber="173" value="2000-12-11">11-XII-2000</date>
, C. Domínguez - S. Roig, 2.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFECAFD7AFE94FCEF" ID-GBIF-Occurrence="3872861373" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFECAFD7AFEA1FD00" box="[278,365,725,746]" country="Argentina" name="San Juan" pageId="6" pageNumber="173">San Juan</collectingRegion>
: Dto. Valle Fértil: Astica,
<date id="E43C0886FFEEFFDEFD81FD7AFF2DFCEF" pageId="6" pageNumber="173" value="1979-05-13">13-V- 1979</date>
, S. Roig,
<quantity id="577A83A3FFEEFFDEFE9AFD5FFE94FCEF" box="[326,344,752,773]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="173" unit="km" value="1.1">1,</quantity>
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEBDFD5FFEDBFCF5" ID-GBIF-Occurrence="3872861381" pageId="6" pageNumber="173">
10 km E Marayes,
<date id="E43C0886FFEEFFDEFDFDFD5FFD43FCEF" box="[545,655,752,773]" pageId="6" pageNumber="173" value="1997-10-23">23-X-1997</date>
, S. Roig, 1.
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEC3FCA5FE54FCD0" ID-GBIF-Occurrence="3872861357" pageId="6" pageNumber="173">
<collectingRegion id="5246E0A4FFEEFFDEFEC3FCA5FEB2FCF5" box="[287,382,778,799]" country="Argentina" name="Mendoza" pageId="6" pageNumber="173">Mendoza</collectingRegion>
: Dto. La Paz: Desaguadero,
<date id="E43C0886FFEEFFDEFF71FC8AFEECFCD0" box="[173,288,805,826]" pageId="6" pageNumber="173" value="1997-11-16">16-XI-1997</date>
, S. Roig,
<quantity id="577A83A3FFEEFFDEFE5AFC8AFE54FCD0" box="[390,408,805,826]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="173" unit="km" value="2.2">2,</quantity>
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE7DFC8AFEA1FCBF" ID-GBIF-Occurrence="3872861378" pageId="6" pageNumber="173">
20 km E La Paz,
<date id="E43C0886FFEEFFDEFD8EFC8AFF2EFCBF" pageId="6" pageNumber="173" value="1997-11-16">16-XI- 1997</date>
, S. Roig, 1;
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEA7FCEFFD5FFC85" ID-GBIF-Occurrence="3872861329" pageId="6" pageNumber="173">
Dto. Santa Rosa: Reserva Ñacuñán,
<quantity id="577A83A3FFEEFFDEFECEFCF5FE9EFC85" box="[274,338,858,879]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="173" unit="m" value="560.0">560 m</quantity>
, pitfall:
<date id="E43C0886FFEEFFDEFE7FFCF5FDDFFC85" box="[419,531,858,879]" pageId="6" pageNumber="173" value="1997-09-13">13-IX-1997</date>
, S. Lagos, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF71FCDAFD94FC60" ID-GBIF-Occurrence="3872861304" box="[173,600,885,906]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF71FCDAFEECFC60" box="[173,288,885,906]" pageId="6" pageNumber="173" value="1997-10-20">20-X-1997</date>
to
<date id="E43C0886FFEEFFDEFE94FCDAFE0BFC60" box="[328,455,885,906]" pageId="6" pageNumber="173" value="1997-11-22">22-XI-1997</date>
, S. Lagos, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFDBFFCDAFD99FC4E" ID-GBIF-Occurrence="3872861316" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFDBFFCDAFF13FC4E" pageId="6" pageNumber="173" value="1998-02-07">7-II- 1998</date>
, S. Lagos, 1,
<date id="E43C0886FFEEFFDEFEBBFC20FE03FC4E" box="[359,463,911,932]" pageId="6" pageNumber="173" value="1998-05-15">15-V-1998</date>
, S. Claver, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFD81FC20FEBEFC55" ID-GBIF-Occurrence="3872861311" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFD81FC20FF2FFC55" pageId="6" pageNumber="173" value="1998-05-31">31-V- 1998</date>
, S. Lagos, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEA0FC05FD5EFC55" ID-GBIF-Occurrence="3872861379" box="[380,658,938,959]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFEA0FC05FDCFFC55" box="[380,515,938,959]" pageId="6" pageNumber="173" value="1998-12-12">12-XII-1998</date>
, S. Lagos, 4,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF71FC6BFE65FC33" ID-GBIF-Occurrence="3872861383" box="[173,425,964,985]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF71FC6BFEC6FC33" box="[173,266,964,985]" pageId="6" pageNumber="173" value="1999-06">VI-1999</date>
, S. Lagos, 2,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE6AFC6BFEBCFC1E" ID-GBIF-Occurrence="3872861324" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE6AFC6BFDD7FC33" box="[438,539,964,985]" pageId="6" pageNumber="173" value="1999-05-09">9-V-1999</date>
to
<date id="E43C0886FFEEFFDEFD90FC6BFF2DFC1E" pageId="6" pageNumber="173" value="1999-06-11">11-VI- 1999</date>
, S. Lagos, 4,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEA1FC70FE89FBE5" ID-GBIF-Occurrence="3872861315" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFEA1FC70FE3CFC1E" box="[381,496,991,1012]" pageId="6" pageNumber="173" value="2010-03-24">24-III-2010</date>
to
<date id="E43C0886FFEEFFDEFDC6FC70FD43FC1E" box="[538,655,991,1012]" pageId="6" pageNumber="173" value="2010-04-25">25-IV-2010</date>
, G. Arriagada, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE90FC55FD85FBE5" ID-GBIF-Occurrence="3872861326" box="[332,585,1018,1039]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE90FC55FE75FBE5" box="[332,441,1018,1039]" pageId="6" pageNumber="173" value="2011-03-22">22-III-2011</date>
, F. Ocampo, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFD93FC55FDC9FBC3" ID-GBIF-Occurrence="3872861317" pageId="6" pageNumber="173">
Campo Quemado,
<date id="E43C0886FFEEFFDEFEC0FBBBFEB1FBC3" box="[284,381,1044,1065]" pageId="6" pageNumber="173" value="1998-02-07">7-II-1998</date>
, S. Lagos, 1;
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFDD2FBBBFE72FBB4" ID-GBIF-Occurrence="3872861369" pageId="6" pageNumber="173">
Dto. Lavalle, Reserva Telteca,
<quantity id="577A83A3FFEEFFDEFEB6FB80FE7DFBAE" box="[362,433,1071,1092]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="6" pageNumber="173" unit="m" value="548.0">548 m</quantity>
, pitfall:
<date id="E43C0886FFEEFFDEFDCEFB80FD5FFBAE" box="[530,659,1071,1092]" pageId="6" pageNumber="173" value="1994-08-09">9-VIII-1994</date>
to
<date id="E43C0886FFEEFFDEFF14FBE6FEFEFBB4" box="[200,306,1097,1118]" pageId="6" pageNumber="173" value="1994-09-06">6-IX-1994</date>
, G. Flores, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE1AFBE6FEB8FB93" ID-GBIF-Occurrence="3872861385" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE1AFBE6FD8EFBB4" box="[454,578,1097,1118]" pageId="6" pageNumber="173" value="1994-12-15">15-XII-1994</date>
to
<date id="E43C0886FFEEFFDEFDBAFBE6FF2FFB93" pageId="6" pageNumber="173" value="1995-02-03">3-II- 1995</date>
, G. Flores, 2,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEA2FBCBFEE2FB7E" ID-GBIF-Occurrence="3872861368" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFEA2FBCBFE22FB93" box="[382,494,1124,1145]" pageId="6" pageNumber="173" value="1996-02-15">15-II-1996</date>
to
<date id="E43C0886FFEEFFDEFDC8FBCBFD42FB93" box="[532,654,1124,1145]" pageId="6" pageNumber="173" value="1996-03-25">25-III-1996</date>
, G. Flores, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFEE4FBD0FD8EFB7E" ID-GBIF-Occurrence="3872861308" box="[312,578,1151,1172]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFEE4FBD0FE63FB7E" box="[312,431,1151,1172]" pageId="6" pageNumber="173" value="2008-04-15">15-IV-2008</date>
, L. Muñoz, 2,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFD97FBD0FE14FB44" ID-GBIF-Occurrence="3872861331" pageId="6" pageNumber="173">
Campo Norte,
<date id="E43C0886FFEEFFDEFF2CFB36FEAFFB44" box="[240,355,1177,1198]" pageId="6" pageNumber="173" value="1999-11-17">17-XI-1999</date>
, S. Roig, 2;
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE3CFB36FF73FB14" ID-GBIF-Occurrence="3872861371" pageId="6" pageNumber="173">
Dto. Capital: CCT CONICET,
<quantity id="577A83A3FFEEFFDEFEFCFB1BFEADFB23" box="[288,353,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="6" pageNumber="173" unit="m" value="839.0">839 m</quantity>
,
<specimenCount id="8684E5CFFFEEFFDEFEB1FB1BFE3CFB23" box="[365,496,1204,1225]" pageId="6" pageNumber="173" type="generic">91 specimens</specimenCount>
collected on pig carcasses (
<emphasis id="A2F6F254FFEEFFDEFECBFB61FEB0FB09" box="[279,380,1230,1251]" italics="true" pageId="6" pageNumber="173">Sus scrofa</emphasis>
L.):
<date id="E43C0886FFEEFFDEFE6CFB61FDD7FB09" box="[432,539,1230,1251]" pageId="6" pageNumber="173" value="2007-05-15">15-V-2007</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF19FB46FE79FB14" ID-GBIF-Occurrence="3872861377" box="[197,437,1257,1278]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF19FB46FEE1FB14" box="[197,301,1257,1278]" pageId="6" pageNumber="173" value="2007-05-17">17-V-2007</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE60FB46FF72FAF2" ID-GBIF-Occurrence="3872861339" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE60FB46FDEFFB14" box="[444,547,1257,1278]" pageId="6" pageNumber="173" value="2007-05-18">18-V-2007</date>
, F. Aballay, 2,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF18FAACFE67FAF2" ID-GBIF-Occurrence="3872861370" box="[196,427,1283,1304]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF18FAACFEE5FAF2" box="[196,297,1283,1304]" pageId="6" pageNumber="173" value="2007-05-19">19-V-2007</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE6CFAACFF73FAD9" ID-GBIF-Occurrence="3872861363" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE6CFAACFDEAFAF2" box="[432,550,1283,1304]" pageId="6" pageNumber="173" value="2007-07-24">24-VII-2007</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF19FAB1FE63FAD9" ID-GBIF-Occurrence="3872861360" box="[197,431,1310,1331]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF19FAB1FEEBFAD9" box="[197,295,1310,1331]" pageId="6" pageNumber="173" value="2008-04-08">8-IV-2008</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE69FAB1FF72FAA7" ID-GBIF-Occurrence="3872861380" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE69FAB1FDE8FAD9" box="[437,548,1310,1331]" pageId="6" pageNumber="173" value="2008-04-12">12-IV-2008</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF18FA97FE78FAA7" ID-GBIF-Occurrence="3872861328" box="[196,436,1336,1357]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF18FA97FEFCFAA7" box="[196,304,1336,1357]" pageId="6" pageNumber="173" value="2008-04-18">18-IV-2008</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE66FA97FF72FA82" ID-GBIF-Occurrence="3872861352" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE66FA97FDEAFAA7" box="[442,550,1336,1357]" pageId="6" pageNumber="173" value="2008-04-21">21-IV-2008</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF18FAFCFE78FA82" ID-GBIF-Occurrence="3872861334" box="[196,436,1363,1384]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF18FAFCFEFCFA82" box="[196,304,1363,1384]" pageId="6" pageNumber="173" value="2008-04-24">24-IV-2008</date>
, F. Aballay, 5,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE66FAFCFF06FA69" ID-GBIF-Occurrence="3872861358" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE66FAFCFDEAFA82" box="[442,550,1363,1384]" pageId="6" pageNumber="173" value="2008-04-28">28-IV-2008</date>
, F. Aballay, 12,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF0DFAC1FE0FFA69" ID-GBIF-Occurrence="3872861310" box="[209,451,1390,1411]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF0DFAC1FEF1FA69" box="[209,317,1390,1411]" pageId="6" pageNumber="173" value="2008-04-30">30-IV-2008</date>
, F. Aballay, 5,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE17FAC1FF73FA77" ID-GBIF-Occurrence="3872861349" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE17FAC1FDE8FA69" box="[459,548,1390,1411]" pageId="6" pageNumber="173" value="2008-05-08">8-V-2008</date>
, F. Aballay, 5,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF19FA27FE79FA77" ID-GBIF-Occurrence="3872861346" box="[197,437,1416,1437]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF19FA27FEE0FA77" box="[197,300,1416,1437]" pageId="6" pageNumber="173" value="2008-05-15">15-V-2008</date>
, F. Aballay, 6,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE60FA27FF72FA52" ID-GBIF-Occurrence="3872861384" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE60FA27FDEEFA77" box="[444,546,1416,1437]" pageId="6" pageNumber="173" value="2008-05-30">30-V-2008</date>
, F. Aballay, 4,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF19FA0CFE78FA52" ID-GBIF-Occurrence="3872861376" box="[197,436,1443,1464]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF19FA0CFEE1FA52" box="[197,301,1443,1464]" pageId="6" pageNumber="173" value="2008-10-30">30-X-2008</date>
, F. Aballay, 9,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE67FA0CFF72FA38" ID-GBIF-Occurrence="3872861307" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE67FA0CFDEFFA52" box="[443,547,1443,1464]" pageId="6" pageNumber="173" value="2008-10-31">31-X-2008</date>
, F. Aballay, 9,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF1AFA12FE7AFA38" ID-GBIF-Occurrence="3872861353" box="[198,438,1469,1490]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF1AFA12FEE6FA38" box="[198,298,1469,1490]" pageId="6" pageNumber="173" value="2008-11-01">1-XI-2008</date>
, F. Aballay, 3,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFE61FA12FF73FA07" ID-GBIF-Occurrence="3872861302" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFE61FA12FDEDFA38" box="[445,545,1469,1490]" pageId="6" pageNumber="173" value="2008-11-02">2-XI-2008</date>
, F. Aballay, 1,
</materialsCitation>
<materialsCitation id="20EA241BFFEEFFDEFF1AFA77FE7AFA07" ID-GBIF-Occurrence="3872861319" box="[198,438,1496,1517]" pageId="6" pageNumber="173">
<date id="E43C0886FFEEFFDEFF1AFA77FEE7FA07" box="[198,299,1496,1517]" pageId="6" pageNumber="173" value="2008-11-03">3-XI-2008</date>
, F. Aballay, 1,
</materialsCitation>
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, F. Aballay, 4,
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<date id="E43C0886FFEEFFDEFF18FA5DFEE4F9ED" box="[196,296,1522,1543]" pageId="6" pageNumber="173" value="2008-11-07">7-XI-2008</date>
, F. Aballay, 2,
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<date id="E43C0886FFEEFFDEFE68FA5DFDE8F9ED" box="[436,548,1522,1543]" pageId="6" pageNumber="173" value="2008-11-10">10-XI-2008</date>
, F. Aballay, 1,
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<date id="E43C0886FFEEFFDEFF12F9A2FE83F9C8" box="[206,335,1549,1570]" pageId="6" pageNumber="173" value="2008-11-11">11-XI-2008</date>
, F. Aballay, 4,
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, F. Aballay, 2,
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, F. Aballay, 3,
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<date id="E43C0886FFEEFFDEFD8DF988FF2EF9BD" pageId="6" pageNumber="173" value="2008-11-16">16-XI- 2008</date>
, F. Aballay,
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2,
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, F. Aballay, 1,
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<date id="E43C0886FFEEFFDEFF71F9F2FEE5F998" box="[173,297,1629,1650]" pageId="6" pageNumber="173" value="2008-12-17">17-XII-2008</date>
, F. Aballay, 2.
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</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEEFFDEFF19F9D8FE35F92B" pageId="6" pageNumber="173" type="distribution">
<paragraph id="903D2E46FFEEFFDEFF19F9D8FE35F92B" blockId="6.[173,660,194,1729]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFF19F9D8FE9AF966" bold="true" box="[197,342,1655,1676]" pageId="6" pageNumber="173">Distribution.</emphasis>
<collectingCountry id="E8956ED6FFEEFFDEFEB8F9D8FE1FF966" box="[356,467,1655,1676]" name="Argentina" pageId="6" pageNumber="173">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFEEFFDEFE3EF9D8FDD6F966" box="[482,538,1655,1676]" country="Argentina" name="Salta" pageId="6" pageNumber="173">Salta</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFDF6F9D8FD43F966" box="[554,655,1655,1676]" country="Argentina" name="Tucuman" pageId="6" pageNumber="173">Tucumán</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFF71F93DFEDEF94D" box="[173,274,1682,1703]" country="Argentina" name="Catamarca" pageId="6" pageNumber="173">Catamarca</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFEC0F93DFEBCF94D" box="[284,368,1682,1703]" country="Argentina" name="La Rioja" pageId="6" pageNumber="173">La Rioja</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFEA6F93DFDFEF94D" box="[378,562,1682,1703]" country="Argentina" name="Santiago del Estero" pageId="6" pageNumber="173">Santiago del Estero</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFDE1F93DFD5CF94D" box="[573,656,1682,1703]" country="Argentina" name="Cordoba" pageId="6" pageNumber="173">Córdoba</collectingRegion>
,
<collectingRegion id="5246E0A4FFEEFFDEFF71F903FEC9F92B" box="[173,261,1708,1729]" country="Argentina" name="San Juan" pageId="6" pageNumber="173">San Juan</collectingRegion>
, and
<collectingRegion id="5246E0A4FFEEFFDEFEE6F903FE5DF92B" box="[314,401,1708,1729]" country="Argentina" name="Mendoza" pageId="6" pageNumber="173">Mendoza</collectingRegion>
provinces.
</paragraph>
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<paragraph id="903D2E46FFEEFFDEFD07FF6DFC18FEB6" blockId="6.[707,1194,194,1039]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFD07FF6DFCE3FF3D" bold="true" box="[731,815,194,215]" pageId="6" pageNumber="173">Habitat.</emphasis>
This species occurs in xerophilous warm steppes and dry forests east of the Andes, in the biogeographic provinces of Monte and
<collectingRegion id="5246E0A4FFEEFFDEFBB1FF58FB65FEE6" box="[1133,1193,247,268]" country="Argentina" name="Chaco" pageId="6" pageNumber="173">Chaco</collectingRegion>
(
<bibRefCitation id="F41353B7FFEEFFDEFD15FEBDFC91FECD" author="Morrone" box="[713,861,274,295]" firstAuthor="Morrone" pageId="6" pageNumber="173" pagination="1 - 110" refId="ref9397" refString="Morrone, J. J. 2014. Biogeographical regionalisation of the Neotropical region. Zootaxa 3782 (1): 1 - 110." type="journal article" year="2014">Morrone 2014</bibRefCitation>
) at altitudes of 150 to
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. Specimens were collected throughout the year, even in the coldest months.
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<paragraph id="903D2E46FFEEFFDEFD07FECEFCA2FD71" blockId="6.[707,1194,194,1039]" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFD07FECEFCE0FE9C" bold="true" box="[731,812,353,374]" pageId="6" pageNumber="173">Biology.</emphasis>
Specimens collected in
<collectingRegion id="5246E0A4FFEEFFDEFBEDFECEFB41FE9C" box="[1073,1165,353,374]" country="Argentina" name="Mendoza" pageId="6" pageNumber="173">Mendoza</collectingRegion>
at the Science and Technology
<collectingRegion id="5246E0A4FFEEFFDEFC0DFED3FBC3FE7B" box="[977,1039,380,401]" country="France" name="Centre" pageId="6" pageNumber="173">Center</collectingRegion>
(91 individuals) were found feeding on decomposing pig carcasses, mainly on the carcass in direct sunlight (88 individuals) and very few on the carcass in the shade (3 individuals). These specimens were recorded in greatest abundance in autumn (46 individuals) and spring (44 individuals), with only one record in the winter and none in the summer. This species was associated with the Advanced Decay and Dry Remains stages of decomposition defined by
<bibRefCitation id="F41353B7FFEEFFDEFD3FFD29FCA5FD71" author="Payne" box="[739,873,646,667]" firstAuthor="Payne" pageId="6" pageNumber="173" pagination="592 - 602" refId="ref9422" refString="Payne, J. A. 1965. A summer carrion study of the baby pig Sus scrofa. Ecology 46: 592 - 602." type="journal article" year="1965">Payne (1965)</bibRefCitation>
.
</paragraph>
<paragraph id="903D2E46FFEEFFDEFD07FD0FFB58FC85" blockId="6.[707,1194,194,1039]" pageId="6" pageNumber="173">Because of these three behavioral characteristics, 1) preference for carcasses under direct sun, 2) seasonality (autumn and spring), and 3) association with specific stages of decomposition (Advanced Decay and Dry Remains), this species should be considered for potential forensic importance for estimating time of death (PMI = Post Mortem Interval) in bodies found in arid environments.</paragraph>
<paragraph id="903D2E46FFEEFFDEFD07FCDAFC6CFBE5" blockId="6.[707,1194,194,1039]" pageId="6" pageNumber="173">
Specimens that were collected using pitfall traps partly filled with water and propylene glycol in two reserves of
<collectingRegion id="5246E0A4FFEEFFDEFCB5FC05FBE4FC55" box="[873,1064,938,959]" country="Argentina" name="Mendoza" pageId="6" pageNumber="173">Mendoza province</collectingRegion>
, Telteca and Ñacuñán, could have been attracted by the smell of decomposition of propylene glycol, similar to the smell of a carcass.
</paragraph>
</subSubSection>
</treatment>
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(
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,
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.
<treatmentCitation id="11230857FFEEFFDEFC38FB26FB69FB74" author="Lacordaire &amp; Histoire Naturelle des Insectes" box="[996,1189,1161,1182]" page="82" pageId="6" pageNumber="173" year="1859">
<bibRefCitation id="F41353B7FFEEFFDEFC38FB26FB69FB74" author="Lacordaire" box="[996,1189,1161,1182]" firstAuthor="Lacordaire" page="82" pageId="6" pageNumber="173" pagination="401 - 750" refId="ref9206" refString="Lacordaire, J. T. 1859. Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu' ici dans cet ordre d' insectes. Tome cinquieme. Contenant les familles des Tenebrionides, Cistelides, Nilionides, Pythides, Melandryides, Lagriides, Pedilides, Anthicides, Pyrochroides, Mordellides, Rhipiphorides, Stylopides, Meloides et Oedemerides. Premiere partie ([1] + pp. 1 - 400), Deuxieme partie ([1] + pp. 401 - 750). Librairie Encyclopedique de Roret, Paris, France." type="journal article" year="1859">Lacordaire 1859: 82</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="11230857FFEEFFDEFD00FB0BFBAEFB53" author="Gemminger &amp; Harold &amp; Catalogus Coleopterorum" box="[732,1122,1188,1209]" page="1842" pageId="6" pageNumber="173" year="1870">
<bibRefCitation id="F41353B7FFEEFFDEFD00FB0BFBAEFB53" author="Gemminger and Harold" box="[732,1122,1188,1209]" firstAuthor="Gemminger" page="1842" pageId="6" pageNumber="173" pagination="1801 - 2180" refId="ref9076" refString="Gemminger, M., and E. Harold. 1870. Catalogus Coleopterorum hujusque descriptorum synonymiscus et sistematicus, 7: 1801 - 2180. S. Gummi, Monachii." type="journal article" year="1870">Gemminger and Harold 1870: 1842</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEEFFDEFCD2FB11FC60FB39" author="Gebien &amp; Tenebrionidae II &amp; In &amp; Coleopterorum Catalogus XVII" box="[782,940,1214,1235]" page="30" pageId="6" pageNumber="173" year="1910">
<bibRefCitation id="F41353B7FFEEFFDEFCD2FB11FC60FB39" author="Gebien" box="[782,940,1214,1235]" firstAuthor="Gebien" page="30" pageId="6" pageNumber="173" pagination="167 - 354" refId="ref9004" refString="Gebien, H. 1910. Tenebrionidae II [pp. 167 - 354]. In: Coleopterorum Catalogus XVII. Pars 22. (W. Junk and S. Schenkling, editors). W. Junk, Berlin, Germany." type="journal article" year="1910">Gebien 1910: 30</bibRefCitation>
</treatmentCitation>
(catalog), 1937: 579 (catalog);
<treatmentCitation id="11230857FFEEFFDEFCCFFB76FC35FB04" author="Blackwelder" box="[787,1017,1241,1262]" page="511" pageId="6" pageNumber="173" year="1945">
<bibRefCitation id="F41353B7FFEEFFDEFCCFFB76FC35FB04" author="Blackwelder" box="[787,1017,1241,1262]" firstAuthor="Blackwelder" page="511" pageId="6" pageNumber="173" pagination="343 - 550" refId="ref8449" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 511</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEEFFDEFBB4FB76FCFFFAE2" author="Kulzer" page="30" pageId="6" pageNumber="173" year="1950">
<bibRefCitation id="F41353B7FFEEFFDEFBB4FB76FCFFFAE2" author="Kulzer" firstAuthor="Kulzer" page="30" pageId="6" pageNumber="173" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950: 30</bibRefCitation>
</treatmentCitation>
(revision), 1956: 898 (revision);
<treatmentCitation id="11230857FFEEFFDEFBA7FB5CFC8FFAC9" author="Pena" page="404" pageId="6" pageNumber="173" year="1966">
<bibRefCitation id="F41353B7FFEEFFDEFBA7FB5CFC8FFAC9" author="Pena" firstAuthor="Pena" page="404" pageId="6" pageNumber="173" pagination="397 - 453" refId="ref9448" refString="Pena, L. E. 1966. Catalogo de los Tenebrionidae (Coleoptera) de Chile. Entomologische Arbeiten aus dem Museum George Frey 17: 397 - 453." type="journal article" year="1966">Peña 1966: 404</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEEFFDEFC69FAA1FBA7FAC9" author="Doyen" box="[949,1131,1294,1315]" page="500" pageId="6" pageNumber="173" year="1994">
<bibRefCitation id="F41353B7FFEEFFDEFC69FAA1FBA7FAC9" author="Doyen" box="[949,1131,1294,1315]" firstAuthor="Doyen" page="500" pageId="6" pageNumber="173" pagination="443 - 514" refId="ref8578" refString="Doyen, J. T. 1994. Cladistic relationships among Pimeliinae Tenebrionidae (Coleoptera). Journal of the New York Entomological Society 101: 443 - 514." type="journal article" year="1994">Doyen 1994: 500</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEEFFDEFBA4FAA1FCA9FAB2" pageId="6" pageNumber="173" type="type_taxon">
<paragraph id="903D2E46FFEEFFDEFBA4FAA1FCA9FAB2" blockId="6.[707,1193,1161,1368]" pageId="6" pageNumber="173">
<typeStatus id="4F3990E4FFEEFFDEFBA4FAA1FB65FAC9" box="[1144,1193,1294,1315]" pageId="6" pageNumber="173">Type</typeStatus>
species:
<taxonomicName id="578255C5FFEEFFDEFCEEFA86FB69FAD7" authority="Erichson, 1847" authorityName="Erichson. For" authorityYear="1847" box="[818,1189,1320,1342]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="A2F6F254FFEEFFDEFCEEFA86FBC9FAD4" box="[818,1029,1321,1342]" italics="true" pageId="6" pageNumber="173">Achanius anthicoides</emphasis>
<bibRefCitation id="F41353B7FFEEFFDEFBD1FA87FB69FAD7" author="Erichson" box="[1037,1189,1320,1341]" firstAuthor="Erichson" pageId="6" pageNumber="173" pagination="67 - 185" refId="ref8638" refString="Erichson, W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte Berlin 13 (1 - 2): 67 - 185." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEEFFDFFD07FAC1FDADFC83" lastPageId="7" lastPageNumber="174" pageId="6" pageNumber="173" type="description">
<paragraph id="903D2E46FFEEFFDFFD07FAC1FDADFC83" blockId="6.[707,1193,1390,1729]" lastBlockId="7.[151,639,194,1296]" lastPageId="7" lastPageNumber="174" pageId="6" pageNumber="173">
<emphasis id="A2F6F254FFEEFFDEFD07FAC1FC9BFA69" bold="true" box="[731,855,1390,1411]" pageId="6" pageNumber="173">Description.</emphasis>
In addition to the characters stated above: Mandibles finer, acute, base 1.5X wider than apex, with dorsal cusp only on right mandible on basal half, left mandible with sharp, raised dorsal edge (
<figureCitation id="08B932C3FFEEFFDEFD23FA77FC85FA07" box="[767,841,1496,1517]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="6" pageNumber="173">Fig. 17</figureCitation>
), both mandibles with ventral fossa (
<figureCitation id="08B932C3FFEEFFDEFD16FA5DFCD9F9ED" box="[714,789,1522,1543]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="6" pageNumber="173">Fig. 18</figureCitation>
); labrum not exposed, entirely membranous; clypeus oval, produced anteriorad of antennal insertions, covering labrum, anterior margin triangular or concave and armed with row of teeth directed downwards, apically with a large tooth prolonged anteriorad, genoclypeal suture wellmarked (
<figureCitation id="08B932C3FFEEFFDEFCC9F93DFC95F94D" box="[789,857,1682,1703]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="6" pageNumber="173">Fig. 17</figureCitation>
), epicanthus separated from clypeus (
<figureCitation id="08B932C3FFEEFFDEFD16F903FCDEF92B" box="[714,786,1708,1729]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="6" pageNumber="173">Fig. 19</figureCitation>
); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave (
<figureCitation id="08B932C3FFEFFFDFFE26FF73FD8CFF1B" box="[506,576,220,241]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 18</figureCitation>
); eyes not protruding, upper margin with slightly or wellmarked carina always present, starting posterior to epicanthus and ending in posterior margin of eye; length of epicanthus equal to length of eye, head widest at eyes, postgenal margin inconspicuous (
<figureCitation id="08B932C3FFEFFFDFFF42FED5FF2BFE65" box="[158,231,378,399]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 19</figureCitation>
); antennae surpassing posterior margin of pronotum, all antennomeres longer than wide, antennomere 1 longer than 3, emerging from distal ¾ of antennal insertion (
<figureCitation id="08B932C3FFEFFFDFFE52FE66FE14FE34" box="[398,472,457,478]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 19</figureCitation>
), antennomere 2 equal to or longer than 3 or shorter than 3 but almost equal (proportion 2/3 = 17/19) (
<figureCitation id="08B932C3FFEFFFDFFE13FE51FDDBFDF9" box="[463,535,510,531]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 19</figureCitation>
); antennomeres 9 and 10 triangular, expanded outwards, longer than wide (
<figureCitation id="08B932C3FFEFFFDFFE88FD9CFE1EFDA2" box="[340,466,563,584]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Figs. 13, 14</figureCitation>
). Pronotum with anterior angles rounded, sloping towards venter. Metacoxae transverse, separated by less than third of metacoxal width (
<figureCitation id="08B932C3FFEFFFDFFEB6FD2DFE7BFD7D" box="[362,439,642,663]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 20</figureCitation>
). First tarsomere of hind legs longer than 2 + 3 combined (
<figureCitation id="08B932C3FFEFFFDFFDFAFD33FF7CFD21" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Figs. 14, 16</figureCitation>
). Female genitalia as in
<figureCitation id="08B932C3FFEFFFDFFE52FD19FE1EFD21" box="[398,466,694,715]" captionStart="Figs" captionStartId="9.[175,215,1662,1681]" captionTargetBox="[151,1171,1074,1639]" captionTargetId="figure-36@9.[151,1173,1074,1640]" captionTargetPageId="9" captionText="Figs. 21–22. Ovipositor in ventral view, spiculum and internal female reproductive tract. 21) Vaniosus profana; 22) Achanius (Ambigatus) rufonitens. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911753" httpUri="https://zenodo.org/record/4911753/files/figure.png" pageId="7" pageNumber="174">Fig. 22</figureCitation>
. Spiculum without arms. Paraprocts long (2.0 &lt;P/C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, duct not annulate. Spermatheca single, compact, locular, emptying at end of vagina, spermathecal accessory gland emptying inside spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFF73FCC0FDB5FAFA" pageId="7" pageNumber="174" type="materials_examined">
<paragraph id="903D2E46FFEFFFDFFF73FCC0FDB5FAFA" blockId="7.[151,639,194,1296]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFF73FCC0FEB1FC6E" bold="true" box="[175,381,879,900]" pageId="7" pageNumber="174">Material Examined.</emphasis>
<materialsCitation id="20EA241BFFEFFFDFFE57FCC0FE16FC74" ID-GBIF-Occurrence="3872861303" pageId="7" pageNumber="174" typeStatus="syntype">
<typeStatus id="4F3990E4FFEFFFDFFE57FCC0FE15FC6E" box="[395,473,879,900]" pageId="7" pageNumber="174" type="syntype">Syntype</typeStatus>
of
<taxonomicName id="578255C5FFEFFFDFFE26FCC0FEB9FC74" authority="(MNHUB)" baseAuthorityName="MNHUB" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="anthicoides" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFE26FCC0FDDCFC6E" box="[506,528,879,900]" italics="true" pageId="7" pageNumber="174">A.</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFDC2FCC0FDBAFC6E" box="[542,630,879,900]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFF4BFC25FEC8FC75" box="[151,260,906,927]" italics="true" pageId="7" pageNumber="174">anthicoides</emphasis>
(MNHUB)
</taxonomicName>
(
<figureCitation id="08B932C3FFEFFFDFFE59FC26FE02FC74" box="[389,462,905,926]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Fig. 13</figureCitation>
);
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFE3FFC26FEB5FC39" ID-GBIF-Occurrence="3872861347" collectionCode="NHMB, IADIZA" pageId="7" pageNumber="174" specimenCount="3">
three specimens of
<taxonomicName id="578255C5FFEFFFDFFF69FC0BFDB1FC53" authority="Fairmaire" authorityName="Fairmaire" box="[181,637,932,953]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bembidioides" subGenus="Achanius">
<originalDetermination id="B39C96BAFFEFFFDFFF69FC0BFDB1FC53" box="[181,637,932,953]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFF69FC0BFEDDFC53" box="[181,273,932,953]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFEFFFC0BFEB3FC53" box="[291,383,932,953]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFE4DFC0BFDD9FC53" box="[401,533,932,953]" italics="true" pageId="7" pageNumber="174">bembidioides</emphasis>
Fairmaire
</originalDetermination>
</taxonomicName>
(
<specimenCount id="8684E5CFFFEFFFDFFF7DFC11FF67FC39" box="[161,171,958,979]" pageId="7" pageNumber="174">1</specimenCount>
<collectionCode id="F693B683FFEFFFDFFF6EFC11FF30FC39" box="[178,252,958,979]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="174">NHMB</collectionCode>
,
<specimenCount id="8684E5CFFFEFFFDFFEDAFC11FEDEFC39" box="[262,274,958,979]" pageId="7" pageNumber="174">2</specimenCount>
<collectionCode id="F693B683FFEFFFDFFEC5FC11FEA3FC39" box="[281,367,958,979]" pageId="7" pageNumber="174">IADIZA</collectionCode>
);
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFEA3FC11FDD1FC04" ID-GBIF-Occurrence="3872861343" pageId="7" pageNumber="174">
<specimenCount id="8684E5CFFFEFFFDFFEA3FC11FDC8FC39" box="[383,516,958,979]" pageId="7" pageNumber="174" type="generic">six specimens</specimenCount>
of
<taxonomicName id="578255C5FFEFFFDFFDF9FC11FDD6FC04" authority="Kulzer (IADIZA)" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="minutus" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFDF9FC11FDB1FC39" box="[549,637,958,979]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFF7CFC76FF30FC04" box="[160,252,985,1006]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFED2FC76FE90FC04" box="[270,348,985,1006]" italics="true" pageId="7" pageNumber="174">minutus</emphasis>
Kulzer (IADIZA)
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFDFBFC76FF22FBE2" ID-GBIF-Occurrence="3872861327" collectionCode="MNHN" pageId="7" pageNumber="174" specimenCount="1" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFDFBFC76FDB2FC04" box="[551,638,985,1006]" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
(
<collectionCode id="F693B683FFEFFFDFFF42FC5CFF24FBE2" box="[158,232,1011,1032]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="174">MNHN</collectionCode>
)
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFF28FC5CFE02FBC8" ID-GBIF-Occurrence="3872861332" pageId="7" pageNumber="174">
and
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(
<collectionCode id="F693B683FFEFFFDFFE6CFC5CFDCFFBE2" box="[432,515,1011,1032]" pageId="7" pageNumber="174">IADIZA</collectionCode>
) of
<taxonomicName id="578255C5FFEFFFDFFDFBFC5CFE06FBC8" authority="Fairmaire" authorityName="Fairmaire" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rufonitens" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFDFBFC5CFDB2FBE2" box="[551,638,1011,1032]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFF43FBA1FECCFBC9" box="[159,256,1038,1059]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFED2FBA1FEA6FBC9" box="[270,362,1038,1059]" italics="true" pageId="7" pageNumber="174">rufonitens</emphasis>
Fairmaire
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFE0FFBA2FDB1FBC8" ID-GBIF-Occurrence="3872861336" box="[467,637,1037,1058]" collectionCode="NHMB" originalDetermination="Achanius (Ambigatus) rufonitens Fairmaire" pageId="7" pageNumber="174" specimenCount="1" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFE0FFBA2FDEFFBC8" box="[467,547,1037,1058]" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
(
<collectionCode id="F693B683FFEFFFDFFDF3FBA2FDBBFBC8" box="[559,631,1037,1058]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="174">NHMB</collectionCode>
)
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFF4BFB87FEC4FBBD" ID-GBIF-Occurrence="3872861362" pageId="7" pageNumber="174">
and
<specimenCount id="8684E5CFFFEFFFDFFF1EFB87FE90FBD7" box="[194,348,1064,1085]" pageId="7" pageNumber="174" type="generic">three specimens</specimenCount>
(
<collectionCode id="F693B683FFEFFFDFFEB7FB87FE72FBD7" box="[363,446,1064,1085]" pageId="7" pageNumber="174">IADIZA</collectionCode>
) of
<taxonomicName id="578255C5FFEFFFDFFE36FB87FECDFBBD" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="stricticollis" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFE36FB87FE33FBD7" box="[490,511,1064,1085]" italics="true" pageId="7" pageNumber="174">A.</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFDD3FB87FDB9FBD7" box="[527,629,1064,1085]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFF4BFBEDFECDFBBD" box="[151,257,1090,1111]" italics="true" pageId="7" pageNumber="174">stricticollis</emphasis>
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFECDFBEDFE09FBBD" ID-GBIF-Occurrence="3872861314" box="[273,453,1090,1111]" collectionCode="NHMB" originalDetermination="A. (Ambigatus) stricticollis" pageId="7" pageNumber="174" specimenCount="1" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFECDFBEDFEA8FBBD" box="[273,356,1090,1111]" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
(
<collectionCode id="F693B683FFEFFFDFFEA8FBEDFE72FBBD" box="[372,446,1090,1111]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="174">NHMB</collectionCode>
)
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFE12FBEDFDB2FB98" ID-GBIF-Occurrence="3872861335" pageId="7" pageNumber="174">
and
<specimenCount id="8684E5CFFFEFFFDFFE26FBEDFDB2FBBD" box="[506,638,1090,1111]" pageId="7" pageNumber="174" type="generic">39 specimens</specimenCount>
(
<collectionCode id="F693B683FFEFFFDFFF42FBF2FF3DFB98" box="[158,241,1117,1138]" pageId="7" pageNumber="174">IADIZA</collectionCode>
) of
<taxonomicName id="578255C5FFEFFFDFFEC4FBF2FDB6FB98" authority="Kulzer" authorityName="Kulzer" box="[280,634,1117,1138]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bicolor" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFEC4FBF2FEA3FB98" box="[280,367,1117,1138]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFEA1FBF2FE2CFB98" box="[381,480,1117,1138]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFE33FBF2FDFEFB98" box="[495,562,1117,1138]" italics="true" pageId="7" pageNumber="174">bicolor</emphasis>
Kulzer
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFF4BFBD8FEF5FB4C" ID-GBIF-Occurrence="3872861364" collectionCode="NHMB" pageId="7" pageNumber="174" specimenCount="1" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFF4BFBD8FF26FB66" box="[151,234,1143,1164]" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
of
<taxonomicName id="578255C5FFEFFFDFFECDFBD8FEF9FB4C" authority="Kulzer (NHMB)" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rhinosomoides" subGenus="Ambigatus">
<originalDetermination id="B39C96BAFFEFFFDFFECDFBD8FF1AFB4C" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFECDFBD8FEA6FB66" box="[273,362,1143,1164]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFEA7FBD8FE2DFB66" box="[379,481,1143,1164]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFE2EFBD8FDB2FB66" box="[498,638,1143,1164]" italics="true" pageId="7" pageNumber="174">rhinosomoides</emphasis>
Kulzer
</originalDetermination>
(
<collectionCode id="F693B683FFEFFFDFFF3FFB3EFEE2FB4C" box="[227,302,1169,1190]" pageId="7" pageNumber="174">NHMB</collectionCode>
)
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFEE3FB3EFE55FB2B" ID-GBIF-Occurrence="3872861337" pageId="7" pageNumber="174" typeStatus="paratype">
<typeStatus id="4F3990E4FFEFFFDFFEE3FB3EFE43FB4C" box="[319,399,1169,1190]" pageId="7" pageNumber="174" type="paratype">paratype</typeStatus>
of
<taxonomicName id="578255C5FFEFFFDFFE72FB3DFE59FB2B" authority="Kulzer (IADIZA)" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="piceus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFE72FB3DFDC9FB4D" box="[430,517,1170,1191]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFDCEFB3DFDB9FB4D" box="[530,629,1170,1191]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFF4BFB03FF1AFB2B" box="[151,214,1196,1217]" italics="true" pageId="7" pageNumber="174">piceus</emphasis>
Kulzer (IADIZA)
</taxonomicName>
;
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFE7FFB03FD85FB31" ID-GBIF-Occurrence="3872861344" collectionCode="NHMB" pageId="7" pageNumber="174" specimenCount="1" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFE7FFB03FE36FB2B" box="[419,506,1196,1217]" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
of
<taxonomicName id="578255C5FFEFFFDFFDFEFB03FE3DFB31" authority="Kulzer (NHMB)" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri" subGenus="Ambigatus">
<originalDetermination id="B39C96BAFFEFFFDFFDFEFB03FE55FB31" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFDFEFB03FDB2FB2B" box="[546,638,1196,1217]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFF43FB68FECDFB36" box="[159,257,1223,1244]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFED0FB68FE94FB36" box="[268,344,1223,1244]" italics="true" pageId="7" pageNumber="174">wittmeri</emphasis>
Kulzer
</originalDetermination>
(
<collectionCode id="F693B683FFEFFFDFFE7FFB69FE27FB31" box="[419,491,1222,1243]" pageId="7" pageNumber="174">NHMB</collectionCode>
)
</taxonomicName>
(
<figureCitation id="08B932C3FFEFFFDFFE20FB69FDF2FB31" box="[508,574,1222,1243]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Fig. 16</figureCitation>
);
</materialsCitation>
<materialsCitation id="20EA241BFFEFFFDFFD91FB69FDB5FAFA" ID-GBIF-Occurrence="3872861375" pageId="7" pageNumber="174" typeStatus="holotype">
<typeStatus id="4F3990E4FFEFFFDFFD91FB69FF0DFB1C" pageId="7" pageNumber="174" type="holotype">holotype</typeStatus>
,
<typeStatus id="4F3990E4FFEFFFDFFF10FB4EFEDBFB1C" box="[204,279,1249,1270]" pageId="7" pageNumber="174" type="allotype">allotype</typeStatus>
, and
<specimenCount id="8684E5CFFFEFFFDFFE96FB4EFE1CFB1C" box="[330,464,1249,1270]" pageId="7" pageNumber="174" type="generic" typeStatus="paratypes">nine paratypes</specimenCount>
of
<taxonomicName id="578255C5FFEFFFDFFE2EFB4EFEE2FAFA" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="antofagastensis" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFE2EFB4EFDCBFB1C" box="[498,519,1249,1270]" italics="true" pageId="7" pageNumber="174">A.</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFDC8FB4EFDBAFB1C" box="[532,630,1249,1270]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFF4BFB54FEE2FAFA" box="[151,302,1275,1296]" italics="true" pageId="7" pageNumber="174">antofagastensis</emphasis>
</taxonomicName>
<collectingCountry id="E8956ED6FFEFFFDFFEEAFB54FEBEFAFA" box="[310,370,1275,1296]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
and Aballay,
<taxonomicNameLabel id="B9C54F2FFFEFFFDFFE20FB54FDB9FAFA" box="[508,629,1275,1296]" pageId="7" pageNumber="174" rank="species">new species</taxonomicNameLabel>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

442
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<mods:title id="A3FA640C36E7173B20F00F475BC311B8">Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson</mods:title>
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<mods:name id="EE078C05E4A85793BD38ECC58DB943E1" type="personal">
<mods:role id="D5255D88C31F1C7BC626F9DE072D3C9F">
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<mods:namePart id="3E1244497FD40A2D6888295D53B1265A">Flores, Gustavo E.</mods:namePart>
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<mods:namePart id="34D1D29B1BDB9EC3C94399F293FB5C62">Aballay, Fernando H.</mods:namePart>
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<mods:date id="6A105818E8C7C9C950B232EB3E2A091C">2015</mods:date>
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<mods:title id="79B9545B9906ECA9583E5DB6681FACB4">mo 14</mods:title>
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<treatment id="182B9F50FFEFFFD2FD60FB11FCE6FE46" ID-DOI="http://doi.org/10.5281/zenodo.6950038" ID-GBIF-Taxon="198139691" ID-Zenodo-Dep="6950038" LSID="urn:lsid:plazi:treatment:182B9F50FFEFFFD2FD60FB11FCE6FE46" httpUri="http://treatment.plazi.org/id/182B9F50FFEFFFD2FD60FB11FCE6FE46" lastPageId="10" lastPageNumber="177" pageId="7" pageNumber="174">
<subSubSection id="D8987DCDFFEFFFDFFD60FB11FBEDFB04" pageId="7" pageNumber="174" type="nomenclature">
<paragraph id="903D2E46FFEFFFDFFD60FB11FBEDFB04" blockId="7.[700,1157,1214,1289]" pageId="7" pageNumber="174">
<heading id="CB75992AFFEFFFDFFD60FB11FBEDFB04" centered="true" fontSize="10" level="2" pageId="7" pageNumber="174" reason="2">
<emphasis id="A2F6F254FFEFFFDFFD60FB11FBEDFB04" bold="true" pageId="7" pageNumber="174">
<taxonomicName id="578255C5FFEFFFDFFD60FB11FC53FB04" authority="Flores and Aballay" authorityName="Flores and Aballay" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="antofagastensis" status="sp. nov." subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFD60FB11FCD6FB39" bold="true" box="[700,794,1214,1235]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCF0FB11FC5AFB39" bold="true" box="[812,918,1214,1235]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFC7BFB11FBF2FB39" bold="true" box="[935,1086,1214,1235]" italics="true" pageId="7" pageNumber="174">antofagastensis</emphasis>
<collectingCountry id="E8956ED6FFEFFFDFFB9BFB11FB49FB39" box="[1095,1157,1214,1235]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
and Aballay
</taxonomicName>
,
<taxonomicNameLabel id="B9C54F2FFFEFFFDFFC77FB76FBEDFB04" box="[939,1057,1241,1262]" pageId="7" pageNumber="174" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="D8987DCDFFEFFFDFFC83FB5CFC2EFAE2" box="[863,994,1267,1289]" pageId="7" pageNumber="174" type="description">
<paragraph id="903D2E46FFEFFFDFFC83FB5CFC2EFAE2" blockId="7.[700,1157,1214,1289]" box="[863,994,1267,1289]" pageId="7" pageNumber="174">
(
<figureCitation id="08B932C3FFEFFFDFFCBAFB5CFC17FAE2" box="[870,987,1267,1289]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Figs. 14–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFD2FD19FAB1FE05FF18" lastPageId="10" lastPageNumber="177" pageId="7" pageNumber="174" type="diagnosis">
<paragraph id="903D2E46FFEFFFDFFD19FAB1FC99FA38" blockId="7.[685,1172,1310,1729]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FAB1FCE4FAD9" bold="true" box="[709,808,1310,1331]" pageId="7" pageNumber="174">Diagnosis.</emphasis>
Antennomere 2 longer than 3, antennomeres 1–5 subcylindrical, antennomeres 6–10 triangular, longer than wide, antennomere 11 equal to 10, pear-shaped; apical maxillary palpomere ovoid, acuminate; prothorax longer than wide, anterior margin of pronotum exceeding width of posterior margin (
<figureCitation id="08B932C3FFEFFFDFFCDCFA12FC85FA38" box="[768,841,1469,1490]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="903D2E46FFEFFFD2FD19FA77FE05FF18" blockId="7.[685,1172,1310,1729]" lastBlockId="10.[173,660,194,1729]" lastPageId="10" lastPageNumber="177" pageId="7" pageNumber="174">
<taxonomicName id="578255C5FFEFFFDFFD19FA77FC0FFA07" box="[709,963,1496,1517]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="antofagastensis">
<emphasis id="A2F6F254FFEFFFDFFD19FA77FC0FFA07" box="[709,963,1496,1517]" italics="true" pageId="7" pageNumber="174">Achanius antofagastensis</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="578255C5FFEFFFDFFD01FA5CFCF3F9E2" box="[733,831,1523,1544]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A2F6F254FFEFFFDFFD01FA5CFCF3F9E2" box="[733,831,1523,1544]" italics="true" pageId="7" pageNumber="174">A. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="578255C5FFEFFFDFFCAFFA5CFC13F9E2" box="[883,991,1523,1544]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="A2F6F254FFEFFFDFFCAFFA5CFC13F9E2" box="[883,991,1523,1544]" italics="true" pageId="7" pageNumber="174">A. wittmeri</emphasis>
</taxonomicName>
(
<figureCitation id="08B932C3FFEFFFDFFC33FA5DFBFBF9ED" box="[1007,1079,1522,1543]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Fig. 16</figureCitation>
) in color pattern. It differs from these species by having reduced wings, antennomere 2 longer than 3 and antennomere 11 equal to 10, whereas
<taxonomicName id="578255C5FFEFFFDFFBF3F9EDFB58F9BD" box="[1071,1172,1602,1623]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A2F6F254FFEFFFDFFBF3F9EDFB58F9BD" box="[1071,1172,1602,1623]" italics="true" pageId="7" pageNumber="174">A. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="578255C5FFEFFFDFFD08F9F2FCF6F998" box="[724,826,1629,1650]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="A2F6F254FFEFFFDFFD08F9F2FCF6F998" box="[724,826,1629,1650]" italics="true" pageId="7" pageNumber="174">A. wittmeri</emphasis>
</taxonomicName>
have well-developed wings, antennomere 2 shorter than 3, and antennomere 11 longer than 10. In addition, in
<taxonomicName id="578255C5FFEFFFDFFC40F93DFBC4F94D" box="[924,1032,1682,1703]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="A2F6F254FFEFFFDFFC40F93DFBC4F94D" box="[924,1032,1682,1703]" italics="true" pageId="7" pageNumber="174">A. wittmeri</emphasis>
</taxonomicName>
the prothorax is wider than long, and the elytra have an additional triangular dark brown spot pointed backwards between the humeri (
<figureCitation id="08B932C3FFE2FFD2FEAAFF72FE72FF18" box="[374,446,221,242]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="10" pageNumber="177">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="C4FD7ECEFFE0FFD0FF19F9D1FB42F92B" ID-DOI="http://doi.org/10.5281/zenodo.4911747" ID-Zenodo-Dep="4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="8" pageNumber="175" startId="8.[197,238,1662,1681]" targetBox="[208,1149,201,1640]" targetPageId="8">
<paragraph id="903D2E46FFE0FFD0FF19F9D1FB42F92B" blockId="8.[173,1194,1662,1729]" pageId="8" pageNumber="175">
<emphasis id="A2F6F254FFE0FFD0FF19F9D1FEF4F97B" bold="true" box="[197,312,1662,1681]" pageId="8" pageNumber="175">Figs. 13–16.</emphasis>
Dorsal habitus.
<emphasis id="A2F6F254FFE0FFD0FE00F9D1FE37F97B" bold="true" box="[476,507,1662,1681]" pageId="8" pageNumber="175">13)</emphasis>
<taxonomicName id="578255C5FFE0FFD0FDDFF9D1FCEBF97A" authorityName="Erichson. For" authorityYear="1847" box="[515,807,1662,1681]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="anthicoides" subGenus="Achanius">
<emphasis id="A2F6F254FFE0FFD0FDDFF9D1FD9FF97A" box="[515,595,1662,1680]" italics="true" pageId="8" pageNumber="175">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE0FFD0FDBFF9D1FD7FF97A" box="[611,691,1662,1680]" italics="true" pageId="8" pageNumber="175">Achanius</emphasis>
)
<emphasis id="A2F6F254FFE0FFD0FD1FF9D1FCEBF97A" box="[707,807,1662,1680]" italics="true" pageId="8" pageNumber="175">anthicoides</emphasis>
</taxonomicName>
, syntype (Image courtesy of Bernd Jaeger, MNHUB);
<emphasis id="A2F6F254FFE0FFD0FED1F939FEE7F943" bold="true" box="[269,299,1686,1705]" pageId="8" pageNumber="175">14)</emphasis>
<taxonomicName id="578255C5FFE0FFD0FEEDF939FDBCF942" box="[305,624,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="antofagastensis" subGenus="Ambigatus">
<emphasis id="A2F6F254FFE0FFD0FEEDF939FEB2F942" box="[305,382,1686,1704]" italics="true" pageId="8" pageNumber="175">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE0FFD0FE56F939FE2EF942" box="[394,482,1686,1704]" italics="true" pageId="8" pageNumber="175">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFE0FFD0FE2CF939FDBCF942" box="[496,624,1686,1704]" italics="true" pageId="8" pageNumber="175">antofagastensis</emphasis>
</taxonomicName>
, holotype;
<emphasis id="A2F6F254FFE0FFD0FD13F939FD21F943" bold="true" box="[719,749,1686,1705]" pageId="8" pageNumber="175">15)</emphasis>
<taxonomicName id="578255C5FFE0FFD0FD2FF939FC47F942" box="[755,907,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="antofagastensis">
<emphasis id="A2F6F254FFE0FFD0FD2FF939FCCCF942" box="[755,768,1686,1704]" italics="true" pageId="8" pageNumber="175">A</emphasis>
.
<emphasis id="A2F6F254FFE0FFD0FCD7F939FC47F942" box="[779,907,1686,1704]" italics="true" pageId="8" pageNumber="175">antofagastensis</emphasis>
</taxonomicName>
, paratype showing reduced wings (white);
<emphasis id="A2F6F254FFE0FFD0FF28F901FEDEF92B" bold="true" box="[244,274,1710,1729]" pageId="8" pageNumber="175">16)</emphasis>
<taxonomicName id="578255C5FFE0FFD0FEC5F901FDD2F92A" box="[281,542,1710,1729]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="wittmeri" subGenus="Ambigatus">
<emphasis id="A2F6F254FFE0FFD0FEC5F901FEAAF92A" box="[281,358,1710,1728]" italics="true" pageId="8" pageNumber="175">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE0FFD0FEAFF901FE07F92A" box="[371,459,1710,1728]" italics="true" pageId="8" pageNumber="175">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFE0FFD0FE06F901FDD2F92A" box="[474,542,1710,1728]" italics="true" pageId="8" pageNumber="175">wittmeri</emphasis>
</taxonomicName>
, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="C4FD7ECEFFE1FFD1FF73FC7AFC04FBEA" ID-DOI="http://doi.org/10.5281/zenodo.4911749" ID-Zenodo-Dep="4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="9" pageNumber="176" startId="9.[175,216,981,1000]" targetBox="[151,1171,194,957]" targetPageId="9">
<paragraph id="903D2E46FFE1FFD1FF73FC7AFC04FBEA" blockId="9.[151,1171,980,1024]" pageId="9" pageNumber="176">
<emphasis id="A2F6F254FFE1FFD1FF73FC7AFEEEFC02" bold="true" box="[175,290,980,1000]" pageId="9" pageNumber="176">Figs. 17–20.</emphasis>
<taxonomicName id="578255C5FFE1FFD1FEE4FC7AFDF2FC0D" box="[312,574,981,1000]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="bicolor" subGenus="Ambigatus">
<emphasis id="A2F6F254FFE1FFD1FEE4FC7AFE4BFC0D" box="[312,391,981,999]" italics="true" pageId="9" pageNumber="176">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE1FFD1FE4BFC7AFE3EFC0D" box="[407,498,981,999]" italics="true" pageId="9" pageNumber="176">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFE1FFD1FDDDFC7AFDF2FC0D" box="[513,574,981,999]" italics="true" pageId="9" pageNumber="176">bicolor</emphasis>
</taxonomicName>
.
<emphasis id="A2F6F254FFE1FFD1FD97FC7AFDA6FC02" bold="true" box="[587,618,981,1000]" pageId="9" pageNumber="176">17)</emphasis>
Mouthparts, dorsal view;
<emphasis id="A2F6F254FFE1FFD1FC89FC7AFCB8FC02" bold="true" box="[853,884,981,1000]" pageId="9" pageNumber="176">18)</emphasis>
Mouthparts, ventral view, white arrow indicates ventral fossa on mandible;
<emphasis id="A2F6F254FFE1FFD1FDD0FC42FDE7FBEA" bold="true" box="[524,555,1005,1024]" pageId="9" pageNumber="176">19)</emphasis>
Head, frontal view;
<emphasis id="A2F6F254FFE1FFD1FD03FC42FD31FBEA" bold="true" box="[735,765,1005,1024]" pageId="9" pageNumber="176">20)</emphasis>
Meso- and metasterna.
</paragraph>
</caption>
<caption id="C4FD7ECEFFE1FFD1FF73F9D1FCF9F92B" ID-DOI="http://doi.org/10.5281/zenodo.4911753" ID-Zenodo-Dep="4911753" httpUri="https://zenodo.org/record/4911753/files/figure.png" pageId="9" pageNumber="176" startId="9.[175,215,1662,1681]" targetBox="[151,1171,1074,1639]" targetPageId="9">
<paragraph id="903D2E46FFE1FFD1FF73F9D1FCF9F92B" blockId="9.[151,1172,1662,1729]" pageId="9" pageNumber="176">
<emphasis id="A2F6F254FFE1FFD1FF73F9D1FEEDF97B" bold="true" box="[175,289,1662,1681]" pageId="9" pageNumber="176">Figs. 21–22.</emphasis>
Ovipositor in ventral view, spiculum and internal female reproductive tract.
<emphasis id="A2F6F254FFE1FFD1FC0DF9D1FC23F97B" bold="true" box="[977,1007,1662,1681]" pageId="9" pageNumber="176">21)</emphasis>
<taxonomicName id="578255C5FFE1FFD1FC2BF9D0FB42F97B" box="[1015,1166,1663,1681]" class="Insecta" family="Tenebrionidae" genus="Vaniosus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="profana">
<emphasis id="A2F6F254FFE1FFD1FC2BF9D0FB42F97B" box="[1015,1166,1663,1681]" italics="true" pageId="9" pageNumber="176">Vaniosus profana</emphasis>
</taxonomicName>
;
<emphasis id="A2F6F254FFE1FFD1FF4BF939FF79F943" bold="true" box="[151,181,1686,1705]" pageId="9" pageNumber="176">22)</emphasis>
<taxonomicName id="578255C5FFE1FFD1FF61F939FE15F942" box="[189,473,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="176" phylum="Arthropoda" rank="species" species="rufonitens" subGenus="Ambigatus">
<emphasis id="A2F6F254FFE1FFD1FF61F939FEC0F942" box="[189,268,1686,1704]" italics="true" pageId="9" pageNumber="176">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE1FFD1FEC7F939FEB9F942" box="[283,373,1686,1704]" italics="true" pageId="9" pageNumber="176">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFE1FFD1FE58F939FE15F942" box="[388,473,1686,1704]" italics="true" pageId="9" pageNumber="176">rufonitens</emphasis>
</taxonomicName>
. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="D8987DCDFFE2FFD2FF19FF58FD8FFD6A" pageId="10" pageNumber="177" type="description">
<paragraph id="903D2E46FFE2FFD2FF19FF58FD8FFD6A" blockId="10.[173,660,194,1729]" pageId="10" pageNumber="177">
<emphasis id="A2F6F254FFE2FFD2FF19FF58FE8DFEE6" bold="true" box="[197,321,247,268]" pageId="10" pageNumber="177">Description.</emphasis>
Length 3.6–4.7 mm. Head and prothorax dark brown, elytra light brown on anterior half and posterior quarter, with dark brown spot on penultimate quarter extending along suture towards apex; antennae, legs, and mouthparts light brown as elytra. One specimen (
<figureCitation id="08B932C3FFE2FFD2FDDBFED3FD98FE7B" box="[519,596,380,401]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="10" pageNumber="177">Fig. 15</figureCitation>
) with same color pattern but elytra with spot outlined in dark brown. One specimen (
<collectingRegion id="5246E0A4FFE2FFD2FE14FE1EFDFEFE2C" box="[456,562,433,454]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
,
<quantity id="577A83A3FFE2FFD2FDE3FE1EFDBFFE2C" box="[575,627,433,454]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</quantity>
W
<collectingRegion id="5246E0A4FFE2FFD2FF71FE63FEEFFE0B" box="[173,291,460,481]" country="Chile" name="Antofagasta" pageId="10" pageNumber="177">Antofagasta</collectingRegion>
de la Sierra (
<quantity id="577A83A3FFE2FFD2FE6EFE63FE3FFE0B" box="[434,499,460,481]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="177" unit="km" value="16.0">16 km</quantity>
W of the
<typeStatus id="4F3990E4FFE2FFD2FD83FE63FF1BFE11" pageId="10" pageNumber="177" type="holotype">holotype</typeStatus>
locality)) with similar color pattern but head, prothorax, legs, and elytral spot very dark brown. Humeri elevated, wings present, reduced, narrow, reaching 3/4 length of elytra, not folded (
<figureCitation id="08B932C3FFE2FFD2FDE2FD99FD4AFDA1" box="[574,646,566,587]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="10" pageNumber="177">Fig. 15</figureCitation>
); metasternum 3X length of mesosternum; metacoxal width 3X metacoxal length (as in
<figureCitation id="08B932C3FFE2FFD2FE2EFDC4FDF4FD6A" box="[498,568,619,640]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="10" pageNumber="177">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFE2FFD2FF19FD29FEA3FD3A" pageId="10" pageNumber="177" type="etymology">
<paragraph id="903D2E46FFE2FFD2FF19FD29FEA3FD3A" blockId="10.[173,660,194,1729]" pageId="10" pageNumber="177">
<emphasis id="A2F6F254FFE2FFD2FF19FD29FEFBFD71" bold="true" box="[197,311,646,667]" pageId="10" pageNumber="177">Etymology.</emphasis>
Named “antofagastensis” after the collection site of Antofagasta de la Sierra,
<collectingRegion id="5246E0A4FFE2FFD2FDEEFD0FFECDFD3A" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca province</collectingRegion>
,
<collectingCountry id="E8956ED6FFE2FFD2FED7FD14FEA7FD3A" box="[267,363,699,720]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFE2FFD2FF19FD7AFEB0F9ED" pageId="10" pageNumber="177" type="materials_examined">
<paragraph id="903D2E46FFE2FFD2FF19FD7AFEB0F9ED" blockId="10.[173,660,194,1729]" pageId="10" pageNumber="177">
<emphasis id="A2F6F254FFE2FFD2FF19FD7AFEADFD00" bold="true" box="[197,353,725,746]" pageId="10" pageNumber="177">Type Material.</emphasis>
<materialsCitation id="20EA241BFFE2FFD2FEB3FD7AFE1FFC60" ID-GBIF-Occurrence="3872861325" collectingDate="2009-11-17" collectingDateMax="2009-11-20" collectingDateMin="2009-11-17" collectionCode="IADIZA" collectorName="G. Flores" country="Argentina" elevation="3593" latitude="-26.027649" location="Punta de la" longitude="-67.344108" municipality="Antofagasta de la Sierra" pageId="10" pageNumber="177" specimenCount="1" specimenCount-male="1" stateProvince="Catamarca" typeStatus="holotype">
<emphasis id="A2F6F254FFE2FFD2FEB3FD7AFE12FD00" bold="true" box="[367,478,725,746]" pageId="10" pageNumber="177">
<typeStatus id="4F3990E4FFE2FFD2FEB3FD7AFE15FD00" box="[367,473,725,746]" pageId="10" pageNumber="177" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="8684E5CFFFE2FFD2FE36FD7AFDD3FD00" box="[490,543,725,746]" pageId="10" pageNumber="177" type="male">Male</specimenCount>
(
<figureCitation id="08B932C3FFE2FFD2FDEEFD7AFD4AFD00" box="[562,646,725,746]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="10" pageNumber="177">Fig. 14</figureCitation>
): [
<collectingCountry id="E8956ED6FFE2FFD2FF6EFD5FFEDEFCEF" box="[178,274,752,773]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFE2FFD2FEC7FD5FFEB2FCEF" box="[283,382,752,773]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
,
<locationDeviation id="818F3764FFE2FFD2FE54FD5FFE12FCEF" bearing="45.0" box="[392,478,752,773]" distance="8000.0" pageId="10" pageNumber="177">
<quantity id="577A83A3FFE2FFD2FE54FD5FFE75FCEF" box="[392,441,752,773]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</quantity>
NE
</locationDeviation>
/
<collectingMunicipality id="7059B43CFFE2FFD2FE2EFD5FFF2BFCF5" pageId="10" pageNumber="177">Antofagasta de la Sierra</collectingMunicipality>
,
<location id="955D789DFFE2FFD2FF2EFCA5FE92FCF5" LSID="urn:lsid:plazi:treatment:182B9F50FFEFFFD2FD60FB11FCE6FE46:955D789DFFE2FFD2FF2EFCA5FE92FCF5" box="[242,350,778,799]" country="Argentina" latitude="-26.027649" municipality="Antofagasta de la Sierra" name="Punta de la" pageId="10" pageNumber="177" stateProvince="Catamarca">Punta de la</location>
/
<location id="955D789DFFE2FFD2FEA8FCA5FE6EFCF5" LSID="urn:lsid:plazi:treatment:182B9F50FFEFFFD2FD60FB11FCE6FE46:955D789DFFE2FFD2FEA8FCA5FE6EFCF5" box="[372,418,778,799]" country="Argentina" latitude="-26.027649" municipality="Antofagasta de la Sierra" name="Pena" pageId="10" pageNumber="177" stateProvince="Catamarca">Peña</location>
<quantity id="577A83A3FFE2FFD2FE76FCA5FE3FFCF5" box="[426,499,778,799]" metricMagnitude="3" metricUnit="m" metricValue="3.593" pageId="10" pageNumber="177" unit="m" value="3593.0">
<elevation id="1BAFC975FFE2FFD2FE76FCA5FE3FFCF5" box="[426,499,778,799]" metricMagnitude="3" metricUnit="m" metricValue="3.593" pageId="10" pageNumber="177" unit="m" value="3593.0">3593 m</elevation>
</quantity>
26°01′39.54″ S, 67° / 20′38.79″ W,
<date id="E43C0886FFE2FFD2FEB9FC8AFE3EFCD0" box="[357,498,805,826]" pageId="10" pageNumber="177" value="2009-11-17" valueMax="2009-11-20" valueMin="2009-11-17">
<collectingDate id="F478F16EFFE2FFD2FEB9FC8AFE3EFCD0" box="[357,498,805,826]" pageId="10" pageNumber="177" value="2009-11-17" valueMax="2009-11-20" valueMin="2009-11-17">17-20-XI-2009</collectingDate>
</date>
/ coll.
<collectorName id="3D774B90FFE2FFD2FDE9FC8AFD43FCD0" box="[565,655,805,826]" pageId="10" pageNumber="177">
G.
<collectingCountry id="E8956ED6FFE2FFD2FD8FFC8AFD43FCD0" box="[595,655,805,826]" name="Indonesia" pageId="10" pageNumber="177">Flores</collectingCountry>
</collectorName>
, F. Aballay] [
<taxonomicName id="578255C5FFE2FFD2FEF9FCEFFE3FFCBF" authority="" authorityName="Fairmaire" authorityYear="1892" box="[293,499,832,853]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFE2FFD2FEF9FCEFFEB0FCBF" box="[293,380,832,853]" italics="true" pageId="10" pageNumber="177">Achanius</emphasis>
(
<emphasis id="A2F6F254FFE2FFD2FE56FCEFFE21FCBF" box="[394,493,832,853]" italics="true" pageId="10" pageNumber="177">Ambigatus</emphasis>
)
</taxonomicName>
/
<taxonomicName id="578255C5FFE2FFD2FDDEFCEFFD5FFCBF" box="[514,659,832,853]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="177" phylum="Arthropoda" rank="species" species="antofagastensis" status="sp. nov.">
<emphasis id="A2F6F254FFE2FFD2FDDEFCEFFD5FFCBF" box="[514,659,832,853]" italics="true" pageId="10" pageNumber="177">antofagastensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9C54F2FFFE2FFD2FF71FCF5FF28FC85" box="[173,228,858,879]" pageId="10" pageNumber="177" rank="species">n. sp.</taxonomicNameLabel>
/
<typeStatus id="4F3990E4FFE2FFD2FF29FCF5FE4AFC85" box="[245,390,858,879]" pageId="10" pageNumber="177" type="holotype">HOLOTYPUS</typeStatus>
male/ Det. G.
<collectingCountry id="E8956ED6FFE2FFD2FDFEFCF5FD93FC85" box="[546,607,858,879]" name="Indonesia" pageId="10" pageNumber="177">Flores</collectingCountry>
and/ F. Aballay 2015] (
<collectionCode id="F693B683FFE2FFD2FEB0FCDAFE04FC60" box="[364,456,885,906]" pageId="10" pageNumber="177">IADIZA</collectionCode>
).
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FE07FCDAFEC2FC4E" ID-GBIF-Occurrence="3872861351" collectingDate="2009-11-17" collectingDateMax="2009-11-20" collectingDateMin="2009-11-17" collectionCode="IADIZA" collectorName="G. Flores" country="Argentina" elevation="3593" latitude="-26.027649" location="Punta de la" longitude="-67.344108" municipality="Antofagasta de la Sierra" pageId="10" pageNumber="177" specimenCount="1" specimenCount-female="1" stateProvince="Catamarca" typeStatus="allotype">
<emphasis id="A2F6F254FFE2FFD2FE07FCDAFD8DFC60" bold="true" box="[475,577,885,906]" pageId="10" pageNumber="177">
<typeStatus id="4F3990E4FFE2FFD2FE07FCDAFDF7FC60" box="[475,571,885,906]" pageId="10" pageNumber="177" type="allotype">Allotype</typeStatus>
:
</emphasis>
Female (
<collectionCode id="F693B683FFE2FFD2FF6FFC20FEC4FC4E" box="[179,264,911,932]" pageId="10" pageNumber="177">IADIZA</collectionCode>
)
</materialsCitation>
and
<specimenCount id="8684E5CFFFE2FFD2FE9FFC20FE03FC4E" box="[323,463,911,932]" pageId="10" pageNumber="177" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
with the same data as
<typeStatus id="4F3990E4FFE2FFD2FF1AFC05FED7FC55" box="[198,283,938,959]" pageId="10" pageNumber="177" type="holotype">holotype</typeStatus>
:
<materialsCitation id="20EA241BFFE2FFD2FEFFFC05FE16FC55" ID-GBIF-Occurrence="3872861322" box="[291,474,938,959]" collectingDate="2009-11-17" collectingDateMax="2009-11-20" collectingDateMin="2009-11-17" collectionCode="NHMB" collectorName="G. Flores" country="Argentina" elevation="3593" latitude="-26.027649" location="Punta de la" longitude="-67.344108" municipality="Antofagasta de la Sierra" pageId="10" pageNumber="177" specimenCount="1" specimenCount-male="1" stateProvince="Catamarca" typeStatus="paratype">
<specimenCount id="8684E5CFFFE2FFD2FEFFFC05FEB4FC55" box="[291,376,938,959]" pageId="10" pageNumber="177" type="male">one male</specimenCount>
(
<collectionCode id="F693B683FFE2FFD2FE59FC05FE1CFC55" box="[389,464,938,959]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="10" pageNumber="177">NHMB</collectionCode>
),
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FE3CFC05FD5FFC55" ID-GBIF-Occurrence="3872861374" box="[480,659,938,959]" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FE3CFC05FDF8FC55" box="[480,564,938,959]" pageId="10" pageNumber="177" type="male">one male</specimenCount>
(
<collectionCode id="F693B683FFE2FFD2FD9DFC05FD45FC55" box="[577,649,938,959]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="177" type="Museum">FMNH</collectionCode>
),
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FF71FC6BFE40FC33" ID-GBIF-Occurrence="3872861345" box="[173,396,964,985]" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FF71FC6BFED3FC33" box="[173,287,964,985]" pageId="10" pageNumber="177" type="female">one female</specimenCount>
(
<collectionCode id="F693B683FFE2FFD2FEECFC6BFE4DFC33" box="[304,385,964,985]" collectionName="Argentina, Buenos Aires, Museo Argentina de Ciencias Naturales" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="10" pageNumber="177" type="Museum">MACN</collectionCode>
),
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FE49FC6BFD58FC33" ID-GBIF-Occurrence="3872861356" box="[405,660,964,985]" pageId="10" pageNumber="177">
one not sexed (
<collectionCode id="F693B683FFE2FFD2FDEBFC6BFD45FC33" box="[567,649,964,985]" collectionName="Argentina, Buenos Aires, Museo Argentina de Ciencias Naturales" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="10" pageNumber="177" type="Museum">MACN</collectionCode>
).
</materialsCitation>
<emphasis id="A2F6F254FFE2FFD2FF71FC70FEDBFC1E" bold="true" box="[173,279,991,1012]" pageId="10" pageNumber="177">
<typeStatus id="4F3990E4FFE2FFD2FF71FC70FEDFFC1E" box="[173,275,991,1012]" pageId="10" pageNumber="177" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<materialsCitation id="20EA241BFFE2FFD2FEC0FC70FED6FBB4" ID-GBIF-Occurrence="3872861318" collectingDate="2009-11-21" collectionCode="MEKRB" collectorName="F. Aballay" country="Argentina" elevation="0" latitude="-26.025944" location="NE Antofagasta de la Sierra" longLatPrecision="1" longitude="-67.345139" municipality="Antofagasta de la Sierra" pageId="10" pageNumber="177" specimenCount="1" specimenCount-male="1" stateProvince="Catamarca">
One
<specimenCount id="8684E5CFFFE2FFD2FE94FC70FEB4FC1E" box="[328,376,991,1012]" pageId="10" pageNumber="177" type="male">male</specimenCount>
: [
<collectingCountry id="E8956ED6FFE2FFD2FE59FC70FE2AFC1E" box="[389,486,991,1012]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFE2FFD2FE32FC70FD83FC1E" box="[494,591,991,1012]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
/ (Dto.)
<collectingMunicipality id="7059B43CFFE2FFD2FF71FC55FE43FBE5" box="[173,399,1018,1039]" pageId="10" pageNumber="177">Antofagasta de la Sierra</collectingMunicipality>
,
<quantity id="577A83A3FFE2FFD2FE46FC55FE00FBE5" box="[410,460,1018,1039]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">
<locationDeviation id="818F3764FFE2FFD2FE46FC55FE00FBE5" box="[410,460,1018,1039]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</locationDeviation>
</quantity>
/
<location id="955D789DFFE2FFD2FE3DFC55FF30FBC3" LSID="urn:lsid:plazi:treatment:182B9F50FFEFFFD2FD60FB11FCE6FE46:955D789DFFE2FFD2FE3DFC55FF30FBC3" country="Argentina" latitude="-26.025944" longLatPrecision="1" longitude="-67.34514" municipality="Antofagasta de la Sierra" name="NE Antofagasta de la Sierra" pageId="10" pageNumber="177" stateProvince="Catamarca">NE Antofagasta de la Sierra</location>
/ llama M-4 P-3, 3585 m / 26°01′33.4″ S/ 67°20′42.5″ W,
<date id="E43C0886FFE2FFD2FE8AFB80FE02FBAE" box="[342,462,1071,1092]" pageId="10" pageNumber="177" value="2009-11-21">
<collectingDate id="F478F16EFFE2FFD2FE8AFB80FE02FBAE" box="[342,462,1071,1092]" pageId="10" pageNumber="177" value="2009-11-21">21-XI-2009</collectingDate>
</date>
/ coll.
<collectorName id="3D774B90FFE2FFD2FDC2FB80FD41FBAE" box="[542,653,1071,1092]" pageId="10" pageNumber="177">F. Aballay</collectorName>
] (
<collectionCode id="F693B683FFE2FFD2FF6FFBE6FEC3FBB4" box="[179,271,1097,1118]" pageId="10" pageNumber="177">MEKRB</collectionCode>
);
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FEFEFBE6FD5FFB93" ID-GBIF-Occurrence="3872861341" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FEFEFBE6FE5CFBB4" box="[290,400,1097,1118]" pageId="10" pageNumber="177" type="female">one female</specimenCount>
with same label information except M-11,
<date id="E43C0886FFE2FFD2FE81FBCBFE07FB93" box="[349,459,1124,1145]" pageId="10" pageNumber="177" value="2009-11-28">28-XI-2009</date>
(
<collectionCode id="F693B683FFE2FFD2FE05FBCBFDE0FB93" box="[473,556,1124,1145]" pageId="10" pageNumber="177">IADIZA</collectionCode>
) (
<figureCitation id="08B932C3FFE2FFD2FD9CFBCBFD4BFB93" box="[576,647,1124,1145]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="10" pageNumber="177">Fig. 15</figureCitation>
);
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FF71FBD0FE7CFB14" ID-GBIF-Occurrence="3872861338" latitude="-26.025639" longitude="-67.343472" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FF71FBD0FE8FFB7E" box="[173,323,1151,1172]" pageId="10" pageNumber="177" type="generic">one specimen</specimenCount>
: [
<collectingCountry id="E8956ED6FFE2FFD2FE8AFBD0FE0AFB7E" box="[342,454,1151,1172]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFE2FFD2FE0FFBD0FD8EFB7E" box="[467,578,1151,1172]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
/ (Dto.) Antofagasta de la Sierra,
<quantity id="577A83A3FFE2FFD2FE77FB36FE12FB44" box="[427,478,1177,1198]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</quantity>
/ NE Antofagasta de la Sierra / cerdo al sol M-11 P-
<quantity id="577A83A3FFE2FFD2FDF4FB1BFD5EFB23" box="[552,658,1204,1225]" metricMagnitude="0" metricUnit="m" metricValue="3.3578" pageId="10" pageNumber="177" unit="m" value="3.3578">3, 3578 m</quantity>
26°01′32.3″ S, 67° / 20′36.5″ W,
<date id="E43C0886FFE2FFD2FDD3FB61FD4DFB09" box="[527,641,1230,1251]" pageId="10" pageNumber="177" value="2009-11-27">27-XI-2009</date>
/ coll. F. Aballay] (
<collectionCode id="F693B683FFE2FFD2FE83FB46FE6AFB14" box="[351,422,1257,1278]" pageId="10" pageNumber="177">MLPA</collectionCode>
);
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FE65FB46FDFCFA82" ID-GBIF-Occurrence="3872861313" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FE65FB46FDDBFB14" box="[441,535,1257,1278]" pageId="10" pageNumber="177" type="male">one male</specimenCount>
: [
<collectingCountry id="E8956ED6FFE2FFD2FDF5FB46FD5CFB14" box="[553,656,1257,1278]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFE2FFD2FF71FAACFEDFFAF2" box="[173,275,1283,1304]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
/ (Dto.) Antofagasta de la Sierra,
<quantity id="577A83A3FFE2FFD2FD89FAACFD4BFAF2" box="[597,647,1283,1304]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</quantity>
/ NE Antofagasta de la Sierra / cerdo sombra M-17 P-
<quantity id="577A83A3FFE2FFD2FF14FA97FEE2FAA7" box="[200,302,1336,1357]" metricMagnitude="0" metricUnit="m" metricValue="1.3595" pageId="10" pageNumber="177" unit="m" value="1.3595">1, 3595 m</quantity>
26°01′38.2″ S, 67° / 20′31.6″ W,
<date id="E43C0886FFE2FFD2FF71FAFCFEECFA82" box="[173,288,1363,1384]" pageId="10" pageNumber="177" value="2009-12-03">03-XII-2009</date>
/ coll. F. Aballay] (
<collectionCode id="F693B683FFE2FFD2FE0EFAFCFDE9FA82" box="[466,549,1363,1384]" pageId="10" pageNumber="177">IADIZA</collectionCode>
),
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FDE9FAFCFEDAFA52" ID-GBIF-Occurrence="3872861330" pageId="10" pageNumber="177" typeStatus="paratype">
<specimenCount id="8684E5CFFFE2FFD2FDE9FAFCFF1BFA69" pageId="10" pageNumber="177" type="generic">one specimen</specimenCount>
with same label information except M-20,
<date id="E43C0886FFE2FFD2FF71FA27FEEAFA77" box="[173,294,1416,1437]" pageId="10" pageNumber="177" value="2009-12-06">06-XII-2009</date>
without head, not labelled
<typeStatus id="4F3990E4FFE2FFD2FD9CFA27FD5FFA77" box="[576,659,1416,1437]" pageId="10" pageNumber="177" type="paratype">paratype</typeStatus>
(
<collectionCode id="F693B683FFE2FFD2FF6FFA0CFEC7FA52" box="[179,267,1443,1464]" pageId="10" pageNumber="177">IADIZA</collectionCode>
);
</materialsCitation>
<materialsCitation id="20EA241BFFE2FFD2FEC3FA0CFEB0F9ED" ID-GBIF-Occurrence="3872861372" latitude="-26.027278" longitude="-67.342111" pageId="10" pageNumber="177">
<specimenCount id="8684E5CFFFE2FFD2FEC3FA0CFE65FA52" box="[287,425,1443,1464]" pageId="10" pageNumber="177" type="generic">one specimen</specimenCount>
: [
<collectingCountry id="E8956ED6FFE2FFD2FE66FA0CFDECFA52" box="[442,544,1443,1464]" name="Argentina" pageId="10" pageNumber="177">Argentina</collectingCountry>
:
<collectingRegion id="5246E0A4FFE2FFD2FDF0FA0CFD5FFA52" box="[556,659,1443,1464]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca</collectingRegion>
<quantity id="577A83A3FFE2FFD2FF71FA12FF11FA38" box="[173,221,1469,1490]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="177" unit="km" value="8.0">8 km</quantity>
W / Antofagasta de la Sierra, Salar
<quantity id="577A83A3FFE2FFD2FDE0FA12FD49FA38" box="[572,645,1469,1490]" metricMagnitude="3" metricUnit="m" metricValue="3.4339999999999997" pageId="10" pageNumber="177" unit="m" value="3434.0">3434 m</quantity>
/ 26°01′54.6″ S, 67°27′07.7″ W /
<date id="E43C0886FFE2FFD2FE3DFA77FD99FA07" box="[481,597,1496,1517]" pageId="10" pageNumber="177" value="2009-12-11">11-XII-2009</date>
/ coll. G.
<collectingCountry id="E8956ED6FFE2FFD2FF10FA5DFEC4F9ED" box="[204,264,1522,1543]" name="Indonesia" pageId="10" pageNumber="177">Flores</collectingCountry>
] (
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).
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</paragraph>
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<subSubSection id="D8987DCDFFE2FFD2FF19F9A2FCE6FE46" pageId="10" pageNumber="177" type="biology_ecology">
<paragraph id="903D2E46FFE2FFD2FF19F9A2FCE6FE46" blockId="10.[173,660,194,1729]" lastBlockId="10.[707,1193,194,428]" pageId="10" pageNumber="177">
<emphasis id="A2F6F254FFE2FFD2FF19F9A2FEDAF9C8" bold="true" box="[197,278,1549,1570]" pageId="10" pageNumber="177">Biology.</emphasis>
Specimens collected in the spring at an altitude of over
<quantity id="577A83A3FFE2FFD2FEB2F988FE0CF9D6" box="[366,448,1575,1596]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="177" unit="m" value="3.5">3,500 m</quantity>
in
<collectingRegion id="5246E0A4FFE2FFD2FE39F988FD95F9D6" box="[485,601,1575,1596]" country="Chile" name="Antofagasta" pageId="10" pageNumber="177">Antofagasta</collectingRegion>
de la Sierra,
<collectingRegion id="5246E0A4FFE2FFD2FF2BF9EDFE0DF9BD" box="[247,449,1602,1623]" country="Argentina" name="Catamarca" pageId="10" pageNumber="177">Catamarca province</collectingRegion>
, were found feeding on decomposing pig carcasses (12 individuals) and on llama (
<taxonomicName id="578255C5FFE2FFD2FE98F9D8FDE4F966" authority="Linnaeus" authorityName="Linnaeus" box="[324,552,1655,1676]" class="Mammalia" family="Camelidae" genus="Lama" kingdom="Animalia" order="Artiodactyla" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="glama">
<emphasis id="A2F6F254FFE2FFD2FE98F9D8FE0EF966" box="[324,450,1655,1676]" italics="true" pageId="10" pageNumber="177">Lama glama</emphasis>
Linnaeus
</taxonomicName>
) carcasses (four individuals). These specimens showed a preference for pig carcasses, nine of which were recorded on the pig carcass in the shade and three under direct sun. This species was associated with the intermediate stages of decomposition (Active Decay and Advanced Decay) defined by
<bibRefCitation id="F41353B7FFE2FFD2FBB1FEBDFCC4FEAB" author="Payne" firstAuthor="Payne" pageId="10" pageNumber="177" pagination="592 - 602" refId="ref9422" refString="Payne, J. A. 1965. A summer carrion study of the baby pig Sus scrofa. Ecology 46: 592 - 602." type="journal article" year="1965">Payne (1965)</bibRefCitation>
. Because of these three behavioral characteristics, this species should be considered for potential forensic importance for estimating time of death (PMI) in arid high environments during the spring.
</paragraph>
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</treatment>
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208
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Subgenus
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<emphasis id="A2F6F254FFEFFFDFFCCFFE61FCA2FE09" bold="true" box="[787,878,462,483]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCA2FE61FC28FE09" bold="true" box="[894,996,462,483]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
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<bibRefCitation id="F41353B7FFEFFFDFFC28FE61FB58FE09" author="Fairmaire" box="[1012,1172,462,483]" firstAuthor="Fairmaire" pageId="7" pageNumber="174" pagination="242 - 253" refId="ref8723" refString="Fairmaire, L. 1892. Description de quelques Coleopteres argentins. Bulletin de la Societe Entomologique de Belgique 36 (3): 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
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<paragraph id="903D2E46FFEFFFDFFC93FE46FC3FFE14" blockId="7.[685,1172,462,510]" box="[847,1011,488,510]" pageId="7" pageNumber="174">
(
<figureCitation id="08B932C3FFEFFFDFFC8AFE46FC04FE14" box="[854,968,488,510]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="8.[197,238,1662,1681]" captionStartId-1="9.[175,216,981,1000]" captionTargetBox-0="[208,1149,201,1640]" captionTargetBox-1="[151,1171,194,957]" captionTargetId-0="figure-1@8.[190,1176,183,1645]" captionTargetId-1="figure-1@9.[151,1173,193,957]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." captionText-1="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi-0="http://doi.org/10.5281/zenodo.4911747" figureDoi-1="http://doi.org/10.5281/zenodo.4911749" httpUri-0="https://zenodo.org/record/4911747/files/figure.png" httpUri-1="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Figs. 14–20</figureCitation>
,
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)
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<emphasis id="A2F6F254FFEFFFDFFD71FDBBFCDFFDC3" box="[685,787,532,553]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
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.
<treatmentCitation id="11230857FFEFFFDFFC2CFDBCFB58FDC2" author="Gebien &amp; Tenebrionidae II &amp; In &amp; Coleopterorum Catalogus XVII" box="[1008,1172,531,552]" page="81" pageId="7" pageNumber="174" year="1910">
<bibRefCitation id="F41353B7FFEFFFDFFC2CFDBCFB58FDC2" author="Gebien" box="[1008,1172,531,552]" firstAuthor="Gebien" page="81" pageId="7" pageNumber="174" pagination="167 - 354" refId="ref9004" refString="Gebien, H. 1910. Tenebrionidae II [pp. 167 - 354]. In: Coleopterorum Catalogus XVII. Pars 22. (W. Junk and S. Schenkling, editors). W. Junk, Berlin, Germany." type="journal article" year="1910">Gebien 1910: 81</bibRefCitation>
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(catalog), 1937: 579 (catalog);
<treatmentCitation id="11230857FFEFFFDFFC2DFD81FB58FDA9" author="Bruch" box="[1009,1172,558,579]" page="264" pageId="7" pageNumber="174" year="1915">
<bibRefCitation id="F41353B7FFEFFFDFFC2DFD81FB58FDA9" author="Bruch" box="[1009,1172,558,579]" firstAuthor="Bruch" page="264" pageId="7" pageNumber="174" pagination="235 - 302" refId="ref8543" refString="Bruch, C. 1915. Catalogo sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata Serie 2 (19): 235 - 302." type="journal article" year="1915">Bruch 1915: 264</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEFFFDFFCEFFDE6FBEBFDB4" author="Blackwelder" box="[819,1063,585,606]" page="511" pageId="7" pageNumber="174" year="1945">
<bibRefCitation id="F41353B7FFEFFFDFFCEFFDE6FBEBFDB4" author="Blackwelder" box="[819,1063,585,606]" firstAuthor="Blackwelder" page="511" pageId="7" pageNumber="174" pagination="343 - 550" refId="ref8449" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 511</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="11230857FFEFFFDFFD1BFDCCFC4EFD92" author="Doyen" box="[711,898,611,632]" page="500" pageId="7" pageNumber="174" year="1994">
<bibRefCitation id="F41353B7FFEFFFDFFD1BFDCCFC4EFD92" author="Doyen" box="[711,898,611,632]" firstAuthor="Doyen" page="500" pageId="7" pageNumber="174" pagination="443 - 514" refId="ref8578" refString="Doyen, J. T. 1994. Cladistic relationships among Pimeliinae Tenebrionidae (Coleoptera). Journal of the New York Entomological Society 101: 443 - 514." type="journal article" year="1994">Doyen 1994: 500</bibRefCitation>
</treatmentCitation>
.
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<typeStatus id="4F3990E4FFEFFFDFFC4CFDCCFC0EFD92" box="[912,962,611,632]" pageId="7" pageNumber="174">Type</typeStatus>
species:
<taxonomicName id="578255C5FFEFFFDFFBFBFDCCFC19FD79" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" class="Insecta" family="Tenebrionidae" genus="Ambigatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="stricticollis">
<emphasis id="A2F6F254FFEFFFDFFBFBFDCCFCFDFD79" italics="true" pageId="7" pageNumber="174">Ambigatus stricticollis</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFCE5FDD1FC19FD79" author="Fairmaire" box="[825,981,638,659]" firstAuthor="Fairmaire" pageId="7" pageNumber="174" pagination="242 - 253" refId="ref8723" refString="Fairmaire, L. 1892. Description de quelques Coleopteres argentins. Bulletin de la Societe Entomologique de Belgique 36 (3): 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
</taxonomicName>
.
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<treatmentCitationGroup id="B0920968FFEFFFDFFD71FD36FC58FD22" pageId="7" pageNumber="174">
<taxonomicName id="578255C5FFEFFFDFFD71FD36FC48FD47" authority="" authorityName="Fairmaire" authorityYear="1892" box="[685,900,664,686]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFD71FD36FCC9FD44" box="[685,773,665,686]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCC8FD36FCB6FD44" box="[788,890,665,686]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
</taxonomicName>
:
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<bibRefCitation id="F41353B7FFEFFFDFFC53FD37FBE7FD47" author="Kulzer" box="[911,1067,664,685]" firstAuthor="Kulzer" page="31" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950: 31</bibRefCitation>
</treatmentCitation>
(revision), 1956: 901 (revision).
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</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FD72FC83FC9D" pageId="7" pageNumber="174" type="diagnosis">
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<emphasis id="A2F6F254FFEFFFDFFD19FD72FCF8FD18" bold="true" box="[709,820,733,754]" pageId="7" pageNumber="174">Diagnosis.</emphasis>
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Kulzer (
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</bibRefCitation>
in key) stated the humeri are not elevated and hind wings are present. This study: metasternum 3X length of mesosternum; metacoxal width 3X metacoxal length; distance between meso– and metacoxae 2.5X metacoxal length (
<figureCitation id="08B932C3FFEFFFDFFD25FCCDFC88FC9D" box="[761,836,866,887]" captionStart="Figs" captionStartId="9.[175,216,981,1000]" captionTargetBox="[151,1171,194,957]" captionTargetId="figure-1@9.[151,1173,193,957]" captionTargetPageId="9" captionText="Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna." figureDoi="http://doi.org/10.5281/zenodo.4911749" httpUri="https://zenodo.org/record/4911749/files/figure.png" pageId="7" pageNumber="174">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FCD2FB5EFBFD" pageId="7" pageNumber="174" type="biology_ecology">
<paragraph id="903D2E46FFEFFFDFFD19FCD2FB5EFBFD" blockId="7.[685,1173,733,1179]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FCD2FCDBFC78" bold="true" box="[709,791,893,914]" pageId="7" pageNumber="174">Biology.</emphasis>
Species of
<taxonomicName id="578255C5FFEFFFDFFC48FCD2FBA2FC78" authority="" authorityName="Fairmaire" authorityYear="1892" box="[916,1134,893,914]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFC48FCD2FC22FC78" box="[916,1006,893,914]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFC22FCD2FBA9FC78" box="[1022,1125,893,914]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
</taxonomicName>
are winged, capable of flying, and of crepuscular and nocturnal habits. They were collected at white and UV lights, also using pitfall traps partly filled with water and propylene glycol (
<bibRefCitation id="F41353B7FFEFFFDFFC09FC48FB5CFC16" author="Flores" box="[981,1168,999,1020]" etAl="et al." firstAuthor="Flores" pageId="7" pageNumber="174" pagination="71 - 90" refId="ref8833" refString="Flores, G. E., S. J. Lagos, and S. Roig-Junent. 2004. Artropodos epigeos que viven bajo la copa del algarrobo (Prosopis flexuosa) en la Reserva Telteca (Mendoza, Argentina). Multequina 13: 71 - 90." type="journal article" year="2004">
<collectingCountry id="E8956ED6FFEFFFDFFC09FC48FBD8FC16" box="[981,1044,999,1020]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
<emphasis id="A2F6F254FFEFFFDFFBC1FC48FB87FC16" box="[1053,1099,999,1020]" italics="true" pageId="7" pageNumber="174">et al</emphasis>
. 2004
</bibRefCitation>
;
<bibRefCitation id="F41353B7FFEFFFDFFD71FBADFCE9FBFD" author="Lagos" box="[685,805,1026,1047]" firstAuthor="Lagos" pageId="7" pageNumber="174" refId="ref9312" refString="Lagos, S. J. 2004. Diversidad biologica de las comunidades epigeas de artropodos en areas pastoreadas y no pastoreadas del Monte (Argentina). PhD dissertation, Programa de Postgrado en Ciencias Biologicas (PROBIOL), Universidad Nacional de Cuyo, Mendoza, Argentina." type="book" year="2004">Lagos 2004</bibRefCitation>
), and cattle dung-baited pitfall traps.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FBB3FCE7FB71" pageId="7" pageNumber="174" type="discussion">
<paragraph id="903D2E46FFEFFFDFFD19FBB3FCE7FB71" blockId="7.[685,1173,733,1179]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FBB3FC4AFBDB" bold="true" box="[709,902,1052,1073]" pageId="7" pageNumber="174">Species Included.</emphasis>
<taxonomicName id="578255C5FFEFFFDFFC48FBB3FBA7FBDB" box="[916,1131,1052,1073]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rufonitens">
<emphasis id="A2F6F254FFEFFFDFFC48FBB3FBA7FBDB" box="[916,1131,1052,1073]" italics="true" pageId="7" pageNumber="174">Achanius rufonitens</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBA2FBB3FCD8FBA6" authorityName="Fairmaire" authorityYear="1892" class="Insecta" family="Tenebrionidae" genus="Ambigatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="stricticollis">
<emphasis id="A2F6F254FFEFFFDFFBA2FBB3FCD8FBA6" italics="true" pageId="7" pageNumber="174">A. stricticollis</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFCFDFB98FCB1FBA6" box="[801,893,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A2F6F254FFEFFFDFFCFDFB98FCB1FBA6" box="[801,893,1079,1100]" italics="true" pageId="7" pageNumber="174">A. bicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFC55FB98FBE0FBA6" box="[905,1068,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="rhinosomoides">
<emphasis id="A2F6F254FFEFFFDFFC55FB98FBE0FBA6" box="[905,1068,1079,1100]" italics="true" pageId="7" pageNumber="174">A. rhinosomoides</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBE4FB98FB41FBA6" box="[1080,1165,1079,1100]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="A2F6F254FFEFFFDFFBE4FB98FB41FBA6" box="[1080,1165,1079,1100]" italics="true" pageId="7" pageNumber="174">A. piceus</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFD71FBFDFCD3FB8D" box="[685,799,1106,1127]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="A2F6F254FFEFFFDFFD71FBFDFCD3FB8D" box="[685,799,1106,1127]" italics="true" pageId="7" pageNumber="174">A. wittmeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="F41353B7FFEFFFDFFCF3FBFEFC7EFB8C" author="Kulzer" box="[815,946,1105,1126]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950</bibRefCitation>
,
<bibRefCitation id="F41353B7FFEFFFDFFC1CFBFEFC39FB8C" author="Kulzer" box="[960,1013,1105,1126]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">1956</bibRefCitation>
), and
<taxonomicName id="578255C5FFEFFFDFFBEBFBFDFC0BFB6B" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="antofagastensis" subGenus="Ambigatus">
<emphasis id="A2F6F254FFEFFFDFFBEBFBFDFB58FB8D" box="[1079,1172,1106,1127]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFD69FBC3FCD1FB6B" box="[693,797,1132,1153]" italics="true" pageId="7" pageNumber="174">Ambigatus</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFCF3FBC3FC0BFB6B" box="[815,967,1132,1153]" italics="true" pageId="7" pageNumber="174">antofagastensis</emphasis>
</taxonomicName>
<collectingCountry id="E8956ED6FFEFFFDFFC0CFBC3FBC1FB6B" box="[976,1037,1132,1153]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
and Aballay,
<taxonomicNameLabel id="B9C54F2FFFEFFFDFFD71FB29FCEBFB71" box="[685,807,1158,1179]" pageId="7" pageNumber="174" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>

223
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<emphasis id="A2F6F254FFEFFFDFFF45FA9CFDB0FAA2" bold="true" box="[153,636,1331,1352]" pageId="7" pageNumber="174">
Subgenus
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<emphasis id="A2F6F254FFEFFFDFFEDFFA9CFEADFAA2" bold="true" box="[259,353,1331,1352]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFEAEFA9CFE1CFAA2" bold="true" box="[370,464,1331,1352]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<bibRefCitation id="F41353B7FFEFFFDFFE3EFA9CFDB0FAA2" author="Erichson" box="[482,636,1331,1352]" firstAuthor="Erichson" pageId="7" pageNumber="174" pagination="67 - 185" refId="ref8638" refString="Erichson, W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte Berlin 13 (1 - 2): 67 - 185." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
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(
<figureCitation id="08B932C3FFEFFFDFFEBBFAE1FE62FA89" box="[359,430,1358,1379]" captionStart="Figs" captionStartId="8.[197,238,1662,1681]" captionTargetBox="[208,1149,201,1640]" captionTargetId="figure-1@8.[190,1176,183,1645]" captionTargetPageId="8" captionText="Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4911747" httpUri="https://zenodo.org/record/4911747/files/figure.png" pageId="7" pageNumber="174">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFF4BFAD7FE60FA28" pageId="7" pageNumber="174" type="reference_group">
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<emphasis id="A2F6F254FFEFFFDFFF4BFAD7FF23FA67" box="[151,239,1400,1421]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFF24FAD7FE74FA67" author="Erichson" box="[248,440,1400,1421]" firstAuthor="Erichson" page="118" pageId="7" pageNumber="174" pagination="67 - 185" refId="ref8638" refString="Erichson, W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte Berlin 13 (1 - 2): 67 - 185." type="journal article" year="1847">Erichson 1847: 118</bibRefCitation>
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.
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<paragraph id="903D2E46FFEFFFDFFF4BFA3CFE60FA28" blockId="7.[151,638,1400,1501]" pageId="7" pageNumber="174">
<treatmentCitationGroup id="B0920968FFEFFFDFFF4BFA3CFE60FA28" pageId="7" pageNumber="174">
<taxonomicName id="578255C5FFEFFFDFFF4BFA3CFEAFFA42" authority="(Achanius)" authorityName="Erichson" authorityYear="1847" baseAuthorityName="Achanius" box="[151,355,1427,1448]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFF4BFA3CFF23FA42" box="[151,239,1427,1448]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFEDDFA3CFE95FA42" box="[257,345,1427,1448]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="11230857FFEFFFDFFEADFA3CFDDEFA42" author="Kulzer" box="[369,530,1427,1448]" page="36" pageId="7" pageNumber="174" year="1950">
<bibRefCitation id="F41353B7FFEFFFDFFEADFA3CFDDEFA42" author="Kulzer" box="[369,530,1427,1448]" firstAuthor="Kulzer" page="36" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950: 36</bibRefCitation>
</treatmentCitation>
(revision);
<treatmentCitation id="11230857FFEFFFDFFF6DFA02FE81FA28" author="Kulzer" box="[177,333,1453,1474]" page="898" pageId="7" pageNumber="174" year="1956">
<bibRefCitation id="F41353B7FFEFFFDFFF6DFA02FE81FA28" author="Kulzer" box="[177,333,1453,1474]" firstAuthor="Kulzer" page="898" pageId="7" pageNumber="174" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">Kulzer 1956: 898</bibRefCitation>
</treatmentCitation>
(revision).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFE73FA02FD8DFA37" pageId="7" pageNumber="174" type="type_taxon">
<paragraph id="903D2E46FFEFFFDFFE73FA02FD8DFA37" blockId="7.[151,638,1400,1501]" pageId="7" pageNumber="174">
<typeStatus id="4F3990E4FFEFFFDFFE73FA02FE10FA28" box="[431,476,1453,1474]" pageId="7" pageNumber="174">Type</typeStatus>
species:
<taxonomicName id="578255C5FFEFFFDFFDF6FA02FE7FFA37" authority="Erichson, 1847" authorityName="Erichson. For" authorityYear="1847" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="A2F6F254FFEFFFDFFDF6FA02FED0FA37" italics="true" pageId="7" pageNumber="174">Achanius anthicoides</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFEFFFA67FE7FFA37" author="Erichson" box="[291,435,1480,1501]" firstAuthor="Erichson" pageId="7" pageNumber="174" pagination="67 - 185" refId="ref8638" refString="Erichson, W. F. 1847. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte Berlin 13 (1 - 2): 67 - 185." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFF73FA5DFDB9F966" pageId="7" pageNumber="174" type="diagnosis">
<paragraph id="903D2E46FFEFFFDFFF73FA5DFDB9F966" blockId="7.[151,639,1522,1729]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFF73FA5DFED2F9ED" bold="true" box="[175,286,1522,1543]" pageId="7" pageNumber="174">Diagnosis.</emphasis>
<bibRefCitation id="F41353B7FFEFFFDFFEF1FA5DFE74F9ED" author="Kulzer" box="[301,440,1522,1543]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">
Kulzer (
<quantity id="577A83A3FFEFFFDFFE58FA5DFE74F9ED" box="[388,440,1522,1543]" metricMagnitude="1" metricUnit="m" metricValue="4.953" pageId="7" pageNumber="174" unit="in" value="1950.0">1950</quantity>
</bibRefCitation>
in key) stated the humeri are not elevated and hind wings are absent. This study: metasternum 2X length of mesosternum; metacoxal width twice metacoxal length; distance between meso– and metacoxae exceeding metacoxal length but not longer than 1.5X metacoxal length.
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFF73F93DFBB7FF18" pageId="7" pageNumber="174" type="biology_ecology">
<paragraph id="903D2E46FFEFFFDFFF73F93DFBB7FF18" blockId="7.[151,639,1522,1729]" lastBlockId="7.[685,1173,194,428]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFF73F93DFECDF94D" bold="true" box="[175,257,1682,1703]" pageId="7" pageNumber="174">Biology.</emphasis>
Species of
<taxonomicName id="578255C5FFEFFFDFFE5EF93DFD9AF94D" authority="(Achanius)" authorityName="Erichson" authorityYear="1847" baseAuthorityName="Achanius" box="[386,598,1682,1703]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFE5EF93DFE13F94D" box="[386,479,1682,1703]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFE2DF93DFD82F94D" box="[497,590,1682,1703]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
</taxonomicName>
are apterous and have been collected under stones or using pitfall traps partly filled with water and propylene glycol (
<bibRefCitation id="F41353B7FFEFFFDFFCE2FF72FC23FF18" author="Flores" box="[830,1007,221,242]" etAl="et al." firstAuthor="Flores" pageId="7" pageNumber="174" pagination="71 - 90" refId="ref8833" refString="Flores, G. E., S. J. Lagos, and S. Roig-Junent. 2004. Artropodos epigeos que viven bajo la copa del algarrobo (Prosopis flexuosa) en la Reserva Telteca (Mendoza, Argentina). Multequina 13: 71 - 90." type="journal article" year="2004">
<collectingCountry id="E8956ED6FFEFFFDFFCE2FF72FCB6FF18" box="[830,890,221,242]" name="Indonesia" pageId="7" pageNumber="174">Flores</collectingCountry>
<emphasis id="A2F6F254FFEFFFDFFC5FFF72FC62FF18" box="[899,942,221,242]" italics="true" pageId="7" pageNumber="174">et al</emphasis>
. 2004
</bibRefCitation>
;
<bibRefCitation id="F41353B7FFEFFFDFFC26FF72FBBCFF18" author="Lagos" box="[1018,1136,221,242]" firstAuthor="Lagos" pageId="7" pageNumber="174" refId="ref9312" refString="Lagos, S. J. 2004. Diversidad biologica de las comunidades epigeas de artropodos en areas pastoreadas y no pastoreadas del Monte (Argentina). PhD dissertation, Programa de Postgrado en Ciencias Biologicas (PROBIOL), Universidad Nacional de Cuyo, Mendoza, Argentina." type="book" year="2004">Lagos 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D8987DCDFFEFFFDFFD19FF58FB59FE46" pageId="7" pageNumber="174" type="discussion">
<paragraph id="903D2E46FFEFFFDFFD19FF58FB59FE46" blockId="7.[685,1173,194,428]" pageId="7" pageNumber="174">
<emphasis id="A2F6F254FFEFFFDFFD19FF58FCB7FEE6" bold="true" box="[709,891,247,268]" pageId="7" pageNumber="174">Species included.</emphasis>
<taxonomicName id="578255C5FFEFFFDFFC55FF58FBA0FEE6" authorityName="Erichson. For" authorityYear="1847" box="[905,1132,247,268]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="anthicoides">
<emphasis id="A2F6F254FFEFFFDFFC55FF58FBA0FEE6" box="[905,1132,247,268]" italics="true" pageId="7" pageNumber="174">Achanius anthicoides</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBA2FF58FCE1FECD" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="bembidioides">
<emphasis id="A2F6F254FFEFFFDFFBA2FF58FCE1FECD" italics="true" pageId="7" pageNumber="174">A. bembidioides</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFCE0FEBDFCC9FEAB" authority="Fairmaire" authorityName="Fairmaire" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="castanescens" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFCE0FEBDFC5BFECD" box="[828,919,274,295]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFC74FEBDFBCFFECD" box="[936,1027,274,295]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFBC8FEBDFB58FECD" box="[1044,1172,274,295]" italics="true" pageId="7" pageNumber="174">castanescens</emphasis>
Fairmaire
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFCD1FE82FB5CFEAB" authority="Fairmaire" authorityName="Fairmaire" box="[781,1168,300,322]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="piceofuscus" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFCD1FE82FCADFEA8" box="[781,865,301,322]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCB1FE82FC0CFEA8" box="[877,960,301,322]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFC10FE82FBFFFEA8" box="[972,1075,301,322]" italics="true" pageId="7" pageNumber="174">piceofuscus</emphasis>
Fairmaire
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFD71FEE8FCD9FEB6" box="[685,789,327,348]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="A2F6F254FFEFFFDFFD71FEE8FCD9FEB6" box="[685,789,327,348]" italics="true" pageId="7" pageNumber="174">A. minutus</emphasis>
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFCFDFEE8FB5CFEB6" authority="Kulzer" authorityName="Kulzer" box="[801,1168,327,348]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="obscurus" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFCFDFEE8FCB6FEB6" box="[801,890,327,348]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFC55FEE8FC2EFEB6" box="[905,994,327,348]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFC2CFEE8FB8BFEB6" box="[1008,1095,327,348]" italics="true" pageId="7" pageNumber="174">obscurus</emphasis>
Kulzer
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFD71FECDFBE1FE9D" authority="Kulzer" authorityName="Kulzer" box="[685,1069,354,375]" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="peruensis" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFD71FECDFCCBFE9D" box="[685,775,354,375]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFCC4FECDFCBEFE9D" box="[792,882,354,375]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFC58FECDFC2DFE9D" box="[900,993,354,375]" italics="true" pageId="7" pageNumber="174">peruensis</emphasis>
Kulzer
</taxonomicName>
,
<taxonomicName id="578255C5FFEFFFDFFBE5FECDFC3AFE7B" authority="Kulzer" authorityName="Kulzer" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFBE5FECDFB58FE9D" box="[1081,1172,354,375]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFD6AFED3FCDAFE7B" box="[694,790,380,401]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFCF5FED3FC68FE7B" box="[809,932,380,401]" italics="true" pageId="7" pageNumber="174">puncticollis</emphasis>
Kulzer
</taxonomicName>
, and
<taxonomicName id="578255C5FFEFFFDFFBEFFED3FC02FE46" authority="Kulzer" authorityName="Kulzer" class="Insecta" family="Tenebrionidae" genus="Achanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="174" phylum="Arthropoda" rank="species" species="angusticollis" subGenus="Achanius">
<emphasis id="A2F6F254FFEFFFDFFBEFFED3FB5FFE7B" box="[1075,1171,380,401]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
(
<emphasis id="A2F6F254FFEFFFDFFD69FE38FCC6FE46" box="[693,778,407,428]" italics="true" pageId="7" pageNumber="174">Achanius</emphasis>
)
<emphasis id="A2F6F254FFEFFFDFFCCBFE38FC47FE46" box="[791,907,407,428]" italics="true" pageId="7" pageNumber="174">angusticollis</emphasis>
Kulzer
</taxonomicName>
(
<bibRefCitation id="F41353B7FFEFFFDFFC06FE38FB9CFE46" author="Kulzer" box="[986,1104,407,428]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="9 - 46" refId="ref9146" refString="Kulzer, H. 1950. 3 Beitrag zur Kenntnis der Tenebrioniden. Entomologische Arbeiten aus dem Museum Georg Frey 1: 9 - 46." type="journal article" year="1950">Kulzer 1950</bibRefCitation>
,
<bibRefCitation id="F41353B7FFEFFFDFFB84FE38FB45FE46" author="Kulzer" box="[1112,1161,407,428]" firstAuthor="Kulzer" pageId="7" pageNumber="174" pagination="895 - 962" refId="ref9172" refString="Kulzer, H. 1956. Neue Tenebrioniden aus Sudamerika. (15. Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 7: 895 - 962." type="journal article" year="1956">1956</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>