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<mods:namePart id="1BD1DB6AF687165DE5BC0D7D4B67AFAF">López-Soriano, Joaquín</mods:namePart>
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<mods:namePart id="CFCA0F9636DC5821C4D7AE6E159B610D">Quiñonero-Salgado, Sergio</mods:namePart>
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<mods:namePart id="3132CBB2EF98381A5E4D9C79668EA75B">Suárez, Álvaro Alonso</mods:namePart>
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<mods:namePart id="98DAE2D4A66310B78DF2EE55C2C9ECB6">Rolán, Emilio</mods:namePart>
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<mods:namePart id="34593283C69C5B9492CA8ECCB9D10D30">Glöer, Peter</mods:namePart>
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<heading id="D0D11B3BE664FFE0FEF6FE195D82FE76" box="[258,480,477,501]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="162" reason="5">
Genus
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<emphasis id="B9527045E664FFE0FEB4FE1B5DE4FE71" bold="true" box="[320,390,479,501]" italics="true" pageId="2" pageNumber="162">Vilertia</emphasis>
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<typeStatus id="549D12F5E664FFE0FD19FF5F5F79FF34" box="[749,795,155,176]" pageId="2" pageNumber="162">Type</typeStatus>
species:
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<emphasis id="B9527045E664FFE0FC99FF595F92FF37" box="[877,1008,157,179]" italics="true" pageId="2" pageNumber="162">Vilertia galeata</emphasis>
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<subSubSection id="C33CFFDCE664FFE0FD58FF1F5F54FE17" pageId="2" pageNumber="162" type="description">
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<emphasis id="B9527045E664FFE0FD58FF1F5F47FF75" bold="true" box="[684,805,219,241]" pageId="2" pageNumber="162">Description:</emphasis>
The most evident characteristic that differentiates this new species from other known valvatiform species on the Iberian Peninsula is the reflexed outer lip, a characteristic which let us introduce a new genus for this new species. The umbilicus is deep and wide.
</paragraph>
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<subSubSection id="C33CFFDCE664FFE1FD58FE595D8DFF35" lastPageId="3" lastPageNumber="163" pageId="2" pageNumber="162" type="etymology">
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<emphasis id="B9527045E664FFE0FD58FE595F79FE36" bold="true" box="[684,795,413,434]" pageId="2" pageNumber="162">Etymology:</emphasis>
The name is a
<emphasis id="B9527045E664FFE0FC52FE5B585BFE31" box="[934,1081,415,437]" italics="true" pageId="2" pageNumber="162">derivatio nominis</emphasis>
referring to the village of Vilert, in the municipality of Esponellà (Pla de l’Estany, Girona province), where the first shells were found. Gender of genus is feminine.
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<paragraph id="8B99AC57E664FFE0FF77F9F65FEDF9C5" blockId="2.[131,911,1586,1602]" box="[131,911,1586,1602]" pageId="2" pageNumber="162">
<emphasis id="B9527045E664FFE0FF77F9F65CB8F9C5" bold="true" box="[131,218,1586,1601]" pageId="2" pageNumber="162">FIGURE 2. A:</emphasis>
Holotype of
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gen.nov.sp. nov.
<emphasis id="B9527045E664FFE0FDFEF9F65E4BF9C5" bold="true" box="[522,553,1586,1601]" pageId="2" pageNumber="162">B-D:</emphasis>
Paratypes photographed under ethanol.Scale:1 mm.
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<paragraph id="8B99AC57E664FFE0FF77F99D5F76F9EC" blockId="2.[131,788,1625,1641]" box="[131,788,1625,1641]" pageId="2" pageNumber="162">
<emphasis id="B9527045E664FFE0FF77F99D5CF0F9EC" bold="true" box="[131,146,1625,1640]" pageId="2" pageNumber="162">A:</emphasis>
holotip de
<emphasis id="B9527045E664FFE0FF2FF99E5D22F9ED" box="[219,320,1626,1641]" italics="true" pageId="2" pageNumber="162">Vilertia galeata</emphasis>
gen.nov.sp. nov.
<emphasis id="B9527045E664FFE0FE4CF99D5DB5F9EC" bold="true" box="[440,471,1625,1640]" pageId="2" pageNumber="162">B-D:</emphasis>
paratips fotografiats sota etanol. Escala:1 mm.
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<emphasis id="B9527045E665FFE1FF7CFF7F5C87FF54" bold="true" box="[136,229,187,208]" pageId="3" pageNumber="163">Remarks:</emphasis>
Only known for the
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species. See comparison with other genera and main characteristics thereafter.
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</treatment>
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<emphasis id="B9527045E665FFE1FEF5FEFA5DEDFED0" bold="true" box="[257,399,318,340]" italics="true" pageId="3" pageNumber="163">Vilertia galeata</emphasis>
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<figureCitation id="131DB0D2E665FFE1FEBBFE985DF0FEF5" box="[335,402,348,370]" captionStart-0="FIGURE 1" captionStart-1="FIGURE2" captionStart-2="FIGURE 3" captionStartId-0="1.[131,184,1562,1577]" captionStartId-1="2.[131,184,1586,1601]" captionStartId-2="3.[130,183,1586,1601]" captionTargetBox-0="[114,1171,152,1653]" captionTargetBox-1="[114,1171,530,1653]" captionTargetBox-2="[114,1170,747,1653]" captionTargetId-0="figure-70@1.[129,1155,164,1547]" captionTargetId-1="figure-189@2.[129,1156,545,1570]" captionTargetId-2="figure-307@3.[129,1156,763,1571]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. SEM pictures of VilertiO gOleOtO gen. nov. sp. nov. A: Different views of the shell; B: Protoconch; C-D: Microsculpture of the protoconch; E: Operculum." captionText-1="FIGURE2. A:Holotype of VilertiO gOleOtO gen.nov.sp.nov.B-D:Paratypes photographed under ethanol.Scale:1 mm." captionText-2="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.12170408" figureDoi-1="http://doi.org/10.5281/zenodo.12170410" figureDoi-2="http://doi.org/10.5281/zenodo.12170412" httpUri-0="https://zenodo.org/record/12170408/files/figure.png" httpUri-1="https://zenodo.org/record/12170410/files/figure.png" httpUri-2="https://zenodo.org/record/12170412/files/figure.png" pageId="3" pageNumber="163">Fig. 1-3</figureCitation>
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<emphasis id="B9527045E665FFE1FF7CFE595D7EFE36" bold="true" box="[136,284,412,434]" pageId="3" pageNumber="163">Type material:</emphasis>
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in
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Paris, #MNHN- IM-2000-38095
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.
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<typeStatus id="549D12F5E665FFE1FEFFFE795D0BFE56" box="[267,361,445,466]" pageId="3" pageNumber="163" type="paratype">Paratypes</typeStatus>
: 2 shells in
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Paris, #MNHN-IM-2000-38096, 2 shells in PG coll., 10 shells in SQS coll
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.
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<typeStatus id="549D12F5E665FFE1FF7CFDDA5CDAFDB7" box="[136,184,542,563]" pageId="3" pageNumber="163">Type</typeStatus>
locality:
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d’en
<location id="8EF9FA8CE665FFE1FE8EFDDA5DD0FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FE8EFDDA5DD0FDB7" box="[378,434,542,563]" county="Coordinates" municipality="Fluvia River" name="Roure" pageId="3" pageNumber="163">Roure</location>
(=
<location id="8EF9FA8CE665FFE1FE38FDDA5D92FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FE38FDDA5D92FDB7" box="[460,496,542,563]" county="Coordinates" municipality="Fluvia River" name="Deu" pageId="3" pageNumber="163">Deu</location>
d’en
<location id="8EF9FA8CE665FFE1FDDEFDD95E04FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FDDEFDD95E04FDB7" box="[554,614,541,563]" county="Coordinates" municipality="Fluvia River" name="Costei" pageId="3" pageNumber="163">Costei</location>
),
<location id="8EF9FA8CE665FFE1FF85FDFA5CA8FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FF85FDFA5CA8FDD0" box="[113,202,574,596]" county="Coordinates" municipality="Fluvia River" name="Esponella" pageId="3" pageNumber="163">Esponellà</location>
(
<location id="8EF9FA8CE665FFE1FF2FFDFA5D12FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FF2FFDFA5D12FDD0" box="[219,368,574,596]" county="Coordinates" municipality="Fluvia River" name="Pla de l'Estany" pageId="3" pageNumber="163">Pla de l’Estany</location>
,
<location id="8EF9FA8CE665FFE1FE8AFDFA5E79FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE1FEF5FEFA5E0DFD30:8EF9FA8CE665FFE1FE8AFDFA5E79FDD0" box="[382,539,574,596]" county="Coordinates" municipality="Fluvia River" name="Girona province" pageId="3" pageNumber="163">Girona province</location>
) located in the
<collectingMunicipality id="6BFD362DE665FFE1FF4EFD9A5D4DFDF0" box="[186,303,606,628]" pageId="3" pageNumber="163">Fluvià River</collectingMunicipality>
basin (
<figureCitation id="131DB0D2E665FFE1FE8AFD9A5DD5FDF7" box="[382,439,606,628]" captionStart="FIGURE4" captionStartId="4.[131,184,1586,1601]" captionTargetBox="[114,1170,1074,1654]" captionTargetId="figure-524@4.[129,1156,1090,1570]" captionTargetPageId="4" captionText="FIGURE4. Map of Spain showing the localities where VilertiO gOleOtO gen.nov.sp.nov.were found.A:Deu d’en Roure;B: Fontde Beu i Tapa." figureDoi="http://doi.org/10.5281/zenodo.12170414" httpUri="https://zenodo.org/record/12170414/files/figure.png" pageId="3" pageNumber="163">Fig. 4</figureCitation>
&amp;
<figureCitation id="131DB0D2E665FFE1FE29FDA45D8AFDF2" box="[477,488,608,630]" captionStart="FIGURE 5" captionStartId="5.[132,185,1585,1600]" captionTargetBox="[114,1171,398,1653]" captionTargetId="figure-171@5.[129,1155,414,1569]" captionTargetPageId="5" captionText="FIGURE 5.VilertiO gOleOtO gen.nov.sp.nov.habitat.A-B:Deu d’en Roure;C:Font de Beu i Tapa.D-E:live specimens." figureDoi="http://doi.org/10.5281/zenodo.12170416" httpUri="https://zenodo.org/record/12170416/files/figure.png" pageId="3" pageNumber="163">5</figureCitation>
)
<collectingCounty id="62F8D4DBE665FFE1FE0FFD9A5E0FFDF0" box="[507,621,606,628]" pageId="3" pageNumber="163">Coordinates</collectingCounty>
: 31TDG
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<quantity id="4CDE01B2E665FFE1FF44FDBA5D26FD10" box="[176,324,638,660]" metricMagnitude="7" metricUnit="m" metricValue="8.205470011499997" pageId="3" pageNumber="163" unit="m" value="8.2054700115E7">82054700. 115 m</quantity>
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<paragraph id="8B99AC57E665FFE1FF7CFD5A5E0DFD30" blockId="3.[113,625,412,692]" box="[136,623,670,692]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FF7CFD5A5D4AFD30" bold="true" box="[136,296,670,692]" pageId="3" pageNumber="163">Other localities:</emphasis>
Font de Beu i Tapa, Vilert (Pla de
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<emphasis id="B9527045E665FFE1FEF5FEFA5DEDFED0" bold="true" box="[257,399,318,340]" italics="true" pageId="3" pageNumber="163">Vilertia galeata</emphasis>
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<paragraph id="8B99AC57E665FFE1FEBBFE985DF0FEF5" blockId="3.[257,480,316,370]" box="[335,402,348,370]" pageId="3" pageNumber="163">
<heading id="D0D11B3BE665FFE1FEBBFE985DF0FEF5" box="[335,402,348,370]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="163" reason="5">
<figureCitation id="131DB0D2E665FFE1FEBBFE985DF0FEF5" box="[335,402,348,370]" captionStart-0="FIGURE 1" captionStart-1="FIGURE2" captionStart-2="FIGURE 3" captionStartId-0="1.[131,184,1562,1577]" captionStartId-1="2.[131,184,1586,1601]" captionStartId-2="3.[130,183,1586,1601]" captionTargetBox-0="[113,1171,148,1544]" captionTargetBox-1="[113,1171,525,1571]" captionTargetBox-2="[123,1168,751,1574]" captionTargetId-0="figure-70@1.[129,1155,164,1547]" captionTargetId-1="figure-189@2.[129,1156,545,1570]" captionTargetId-2="figure-307@3.[129,1156,763,1571]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. SEM pictures of VilertiO gOleOtO gen. nov. sp. nov. A: Different views of the shell; B: Protoconch; C-D: Microsculpture of the protoconch; E: Operculum. Fotografies SEM de VilertiO gOleOtO gen.nov.sp.nov.A:diferents vistes de la closca;B:protoconquilla;C-D:microescultura de la protoconquilla;E:opercle." captionText-1="FIGURE2. A:Holotype of VilertiO gOleOtO gen.nov.sp.nov.B-D:Paratypes photographed under ethanol.Scale:1 mm. A:holotip de VilertiO gOleOtO gen.nov.sp.nov.B-D:paratips fotografiats sota etanol. Escala:1 mm." captionText-2="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm VilertiO gOleOtO gen.nov.sp.nov.A:conquilla (paratip);B: òrgancopulador del mascle;C:gir intestinal;D-E:opercle (costatintern).Escala:1mm" figureDoi-0="http://doi.org/10.5281/zenodo.12170408" figureDoi-1="http://doi.org/10.5281/zenodo.12170410" figureDoi-2="http://doi.org/10.5281/zenodo.12170412" httpUri-0="https://zenodo.org/record/12170408/files/figure.png" httpUri-1="https://zenodo.org/record/12170410/files/figure.png" httpUri-2="https://zenodo.org/record/12170412/files/figure.png" pageId="3" pageNumber="163">Fig. 1-3</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE665FFE1FF7CFE595D14FD10" pageId="3" pageNumber="163" type="materials_examined">
<paragraph id="8B99AC57E665FFE1FF7CFE595CA3FD97" blockId="3.[113,625,412,692]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FF7CFE595D7EFE36" bold="true" box="[136,284,412,434]" pageId="3" pageNumber="163">Type material:</emphasis>
<materialsCitation id="3B4EA60AE665FFE1FED1FE595C9EFE57" collectionCode="MNHN" pageId="3" pageNumber="163" specimenCount="1" typeStatus="holotype">
<typeStatus id="549D12F5E665FFE1FED1FE595D18FE37" box="[293,378,413,435]" pageId="3" pageNumber="163" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED373492E665FFE1FE69FE595DBEFE36" box="[413,476,413,434]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="163">MNHN</collectionCode>
Paris, #MNHN-IM-2000-38095
</materialsCitation>
.
<materialsCitation id="3B4EA60AE665FFE1FEFFFE795CDCFD97" collectionCode="MNHN" pageId="3" pageNumber="163" specimenCount="1" typeStatus="paratype">
<typeStatus id="549D12F5E665FFE1FEFFFE795D0BFE56" box="[267,361,445,466]" pageId="3" pageNumber="163" type="paratype">Paratypes</typeStatus>
: 2 shells in
<collectionCode id="ED373492E665FFE1FE05FE795E52FE56" box="[497,560,445,466]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="163">MNHN</collectionCode>
Paris, #MNHN-IM-2000-38096, 2 shells in PG coll., 10 shells in SQS coll
</materialsCitation>
.
</paragraph>
<paragraph id="8B99AC57E665FFE1FF7CFDDA5D14FD10" blockId="3.[113,625,412,692]" pageId="3" pageNumber="163">
<materialsCitation id="3B4EA60AE665FFE1FF7CFDDA5D14FD10" county="Coordinates" elevation="82054700" location="Deu" municipality="Fluvia River" pageId="3" pageNumber="163" specimenCount="1" typeStatus="holotype">
<emphasis id="B9527045E665FFE1FF7CFDDA5D71FDB7" bold="true" box="[136,275,541,563]" pageId="3" pageNumber="163">
<typeStatus id="549D12F5E665FFE1FF7CFDDA5CDAFDB7" box="[136,184,542,563]" pageId="3" pageNumber="163">Type</typeStatus>
locality:
</emphasis>
<location id="8EF9FA8CE665FFE1FEE8FDDA5D22FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FEE8FDDA5D22FDB7" box="[284,320,542,563]" county="Coordinates" municipality="Fluvia River" name="Deu" pageId="3" pageNumber="163">Deu</location>
d’en
<location id="8EF9FA8CE665FFE1FE8EFDDA5DD0FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FE8EFDDA5DD0FDB7" box="[378,434,542,563]" county="Coordinates" municipality="Fluvia River" name="Roure" pageId="3" pageNumber="163">Roure</location>
(=
<location id="8EF9FA8CE665FFE1FE38FDDA5D92FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FE38FDDA5D92FDB7" box="[460,496,542,563]" county="Coordinates" municipality="Fluvia River" name="Deu" pageId="3" pageNumber="163">Deu</location>
d’en
<location id="8EF9FA8CE665FFE1FDDEFDD95E04FDB7" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FDDEFDD95E04FDB7" box="[554,614,541,563]" county="Coordinates" municipality="Fluvia River" name="Costei" pageId="3" pageNumber="163">Costei</location>
),
<location id="8EF9FA8CE665FFE1FF85FDFA5CA8FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FF85FDFA5CA8FDD0" box="[113,202,574,596]" county="Coordinates" municipality="Fluvia River" name="Esponella" pageId="3" pageNumber="163">Esponellà</location>
(
<location id="8EF9FA8CE665FFE1FF2FFDFA5D12FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FF2FFDFA5D12FDD0" box="[219,368,574,596]" county="Coordinates" municipality="Fluvia River" name="Pla de l'Estany" pageId="3" pageNumber="163">Pla de l’Estany</location>
,
<location id="8EF9FA8CE665FFE1FE8AFDFA5E79FDD0" LSID="urn:lsid:plazi:treatment:038F1D41E665FFE6FEF5FEFA5841FE76:8EF9FA8CE665FFE1FE8AFDFA5E79FDD0" box="[382,539,574,596]" county="Coordinates" municipality="Fluvia River" name="Girona province" pageId="3" pageNumber="163">Girona province</location>
) located in the
<collectingMunicipality id="6BFD362DE665FFE1FF4EFD9A5D4DFDF0" box="[186,303,606,628]" pageId="3" pageNumber="163">Fluvià River</collectingMunicipality>
basin (
<figureCitation id="131DB0D2E665FFE1FE8AFD9A5DD5FDF7" box="[382,439,606,628]" captionStart="FIGURE4" captionStartId="4.[131,184,1586,1601]" captionTargetBox="[121,1163,1083,1575]" captionTargetId="figure-524@4.[129,1156,1090,1570]" captionTargetPageId="4" captionText="FIGURE4. Map of Spain showing the localities where VilertiO gOleOtO gen.nov.sp.nov.were found.A:Deu d’en Roure;B: Fontde Beu i Tapa. Mapa d’Espanya mostrant les localitats on es va trobar VilertiO gOleOtO gen.nov.sp.nov.A:deu d’en Roure;B:font de Beu i Tapa." figureDoi="http://doi.org/10.5281/zenodo.12170414" httpUri="https://zenodo.org/record/12170414/files/figure.png" pageId="3" pageNumber="163">Fig. 4</figureCitation>
&amp;
<figureCitation id="131DB0D2E665FFE1FE29FDA45D8AFDF2" box="[477,488,608,630]" captionStart="FIGURE 5" captionStartId="5.[132,185,1585,1600]" captionTargetBox="[122,1165,404,1575]" captionTargetId="figure-171@5.[129,1155,414,1569]" captionTargetPageId="5" captionText="FIGURE 5.VilertiO gOleOtO gen.nov.sp.nov.habitat.A-B:Deu d’en Roure;C:Font de Beu i Tapa.D-E:live specimens. Hàbitatde VilertiO gOleOtO gen.nov.sp.nov.A-B:deud’en Roure;C:font de Beu i Tapa.D-E:exemplars vius." figureDoi="http://doi.org/10.5281/zenodo.12170416" httpUri="https://zenodo.org/record/12170416/files/figure.png" pageId="3" pageNumber="163">5</figureCitation>
)
<collectingCounty id="62F8D4DBE665FFE1FE0FFD9A5E0FFDF0" box="[507,621,606,628]" pageId="3" pageNumber="163">Coordinates</collectingCounty>
: 31TDG
<elevation id="000B4B64E665FFE1FF44FDBA5D14FD10" box="[176,374,638,660]" metricMagnitude="7" metricUnit="m" metricValue="8.205470011499997" pageId="3" pageNumber="163" unit="m" value="8.2054700115E7">
<quantity id="4CDE01B2E665FFE1FF44FDBA5D26FD10" box="[176,324,638,660]" metricMagnitude="7" metricUnit="m" metricValue="8.205470011499997" pageId="3" pageNumber="163" unit="m" value="8.2054700115E7">82054700. 115 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE665FFE1FF7CFD5A5FABFF55" pageId="3" pageNumber="163" type="distribution">
<paragraph id="8B99AC57E665FFE1FF7CFD5A5FABFF55" blockId="3.[113,625,412,692]" lastBlockId="3.[659,1171,155,692]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FF7CFD5A5D4AFD30" bold="true" box="[136,296,670,692]" pageId="3" pageNumber="163">Other localities:</emphasis>
Font de Beu i Tapa, Vilert (Pla de l’Estany, Girona province), also in the Fluvià River basin. 31TDG
<quantity id="4CDE01B2E665FFE1FCFAFF7F5FF5FF54" box="[782,919,187,209]" metricMagnitude="7" metricUnit="m" metricValue="8.53416919" pageId="3" pageNumber="163" unit="m" value="8.53416919E7">85341691. 90 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE665FFE6FD5EFF1F5D88FF75" lastPageId="4" lastPageNumber="164" pageId="3" pageNumber="163" type="description">
<paragraph id="8B99AC57E665FFE1FD5EFF1F586CFDF0" blockId="3.[659,1171,155,692]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FD5EFF1F5F41FF75" bold="true" box="[682,803,219,241]" pageId="3" pageNumber="163">Description:</emphasis>
Small shell, with valvatiform shape,
<quantity id="4CDE01B2E665FFE1FD67FF3F5F70FE95" box="[659,786,251,273]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.4" metricValueMin="5.1" pageId="3" pageNumber="163" unit="mm" value="0.625" valueMax="0.74" valueMin="0.51">0.51 - 0.74 mm</quantity>
in height and
<quantity id="4CDE01B2E665FFE1FC5BFF3F584FFE95" box="[943,1069,251,273]" metricMagnitude="-3" metricUnit="m" metricValue="1.045" metricValueMax="1.13" metricValueMin="0.96" pageId="3" pageNumber="163" unit="mm" value="1.045" valueMax="1.13" valueMin="0.96">0.96 -1.13 mm</quantity>
diameter, with 3 whorls, deep sutures, and translucid coloration in freshly-collected specimens (
<figureCitation id="131DB0D2E665FFE1FC4DFEF85F8EFED5" box="[953,1004,316,338]" captionStart="FIGURE2" captionStartId="2.[131,184,1586,1601]" captionTargetBox="[113,1171,525,1571]" captionTargetId="figure-189@2.[129,1156,545,1570]" captionTargetPageId="2" captionText="FIGURE2. A:Holotype of VilertiO gOleOtO gen.nov.sp.nov.B-D:Paratypes photographed under ethanol.Scale:1 mm. A:holotip de VilertiO gOleOtO gen.nov.sp.nov.B-D:paratips fotografiats sota etanol. Escala:1 mm." figureDoi="http://doi.org/10.5281/zenodo.12170410" httpUri="https://zenodo.org/record/12170410/files/figure.png" pageId="3" pageNumber="163">Fig. 2</figureCitation>
,
<figureCitation id="131DB0D2E665FFE1FC01FEF85863FED5" box="[1013,1025,316,337]" captionStart="FIGURE 3" captionStartId="3.[130,183,1586,1601]" captionTargetBox="[123,1168,751,1574]" captionTargetId="figure-307@3.[129,1156,763,1571]" captionTargetPageId="3" captionText="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm VilertiO gOleOtO gen.nov.sp.nov.A:conquilla (paratip);B: òrgancopulador del mascle;C:gir intestinal;D-E:opercle (costatintern).Escala:1mm" figureDoi="http://doi.org/10.5281/zenodo.12170412" httpUri="https://zenodo.org/record/12170412/files/figure.png" pageId="3" pageNumber="163">3</figureCitation>
,
<figureCitation id="131DB0D2E665FFE1FBFEFEF85874FED6" box="[1034,1046,316,338]" captionStart="FIGURE 6" captionStartId="6.[131,184,691,706]" captionTargetBox="[120,1162,164,680]" captionTargetId="figure-281@6.[130,1156,169,675]" captionTargetPageId="6" captionText="FIGURE 6.VilertiO gOleOtO gen.nov.sp.nov.shell profile (lef) and measurements from the different specimens studied (right). Aspecte de la conquilla de VilertiO gOleOtO gen.nov.sp.nov.(esquerra) i mesures dels exemplars estudiats (dreta)." figureDoi="http://doi.org/10.5281/zenodo.12170418" httpUri="https://zenodo.org/record/12170418/files/figure.png" pageId="3" pageNumber="163">6</figureCitation>
). Protoconch microsculpture is formed by irregularly arranged microdepressions (
<figureCitation id="131DB0D2E665FFE1FCBAFEB85FE5FE16" box="[846,903,380,402]" captionStart="FIGURE 1" captionStartId="1.[131,184,1562,1577]" captionTargetBox="[113,1171,148,1544]" captionTargetId="figure-70@1.[129,1155,164,1547]" captionTargetPageId="1" captionText="FIGURE 1. SEM pictures of VilertiO gOleOtO gen. nov. sp. nov. A: Different views of the shell; B: Protoconch; C-D: Microsculpture of the protoconch; E: Operculum. Fotografies SEM de VilertiO gOleOtO gen.nov.sp.nov.A:diferents vistes de la closca;B:protoconquilla;C-D:microescultura de la protoconquilla;E:opercle." figureDoi="http://doi.org/10.5281/zenodo.12170408" httpUri="https://zenodo.org/record/12170408/files/figure.png" pageId="3" pageNumber="163">Fig. 1</figureCitation>
C-D). Teleoconch showing growing lines. Deep and wide umbilicus, ovoid aperture, about
<quantity id="4CDE01B2E665FFE1FCF4FE795F1CFE56" box="[768,894,445,467]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="163" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60 -0.80 mm</quantity>
high and
<quantity id="4CDE01B2E665FFE1FC28FE785838FE56" box="[988,1114,444,467]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.6" metricValueMin="5.1" pageId="3" pageNumber="163" unit="mm" value="0.585" valueMax="0.66" valueMin="0.51">0.51 -0.66 mm</quantity>
wide, slightly oblique, and having a sharp peristome, slightly reflected towards the columella, and strongly towards the outer lip. Last whorl represents 90 % of total height. Corneum operculum, with orange to yellow coloration, convex in the inner side (see
<figureCitation id="131DB0D2E665FFE1FC56FD9A5F82FDF7" box="[930,992,606,628]" captionStart="FIGURE 1" captionStartId="1.[131,184,1562,1577]" captionTargetBox="[113,1171,148,1544]" captionTargetId="figure-70@1.[129,1155,164,1547]" captionTargetPageId="1" captionText="FIGURE 1. SEM pictures of VilertiO gOleOtO gen. nov. sp. nov. A: Different views of the shell; B: Protoconch; C-D: Microsculpture of the protoconch; E: Operculum. Fotografies SEM de VilertiO gOleOtO gen.nov.sp.nov.A:diferents vistes de la closca;B:protoconquilla;C-D:microescultura de la protoconquilla;E:opercle." figureDoi="http://doi.org/10.5281/zenodo.12170408" httpUri="https://zenodo.org/record/12170408/files/figure.png" pageId="3" pageNumber="163">Fig. 1E</figureCitation>
,
<figureCitation id="131DB0D2E665FFE1FC1DFD9A5866FDF0" box="[1001,1028,606,628]" captionStart="FIGURE 3" captionStartId="3.[130,183,1586,1601]" captionTargetBox="[123,1168,751,1574]" captionTargetId="figure-307@3.[129,1156,763,1571]" captionTargetPageId="3" captionText="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm VilertiO gOleOtO gen.nov.sp.nov.A:conquilla (paratip);B: òrgancopulador del mascle;C:gir intestinal;D-E:opercle (costatintern).Escala:1mm" figureDoi="http://doi.org/10.5281/zenodo.12170412" httpUri="https://zenodo.org/record/12170412/files/figure.png" pageId="3" pageNumber="163">3D</figureCitation>
).
</paragraph>
<paragraph id="8B99AC57E665FFE6FD5EFDBA5D88FF75" blockId="3.[659,1171,155,692]" lastBlockId="4.[113,624,155,499]" lastPageId="4" lastPageNumber="164" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FD5EFDBA5E9AFD10" bold="true" box="[682,760,638,660]" pageId="3" pageNumber="163">Animal:</emphasis>
The animal is colorless, and the eyes are missing. The penis is broad and rounded at the tip with a penial lobe of the same length (
<figureCitation id="131DB0D2E662FFE6FE30FF5F5E6BFF35" box="[452,521,155,177]" captionStart="FIGURE 3" captionStartId="3.[130,183,1586,1601]" captionTargetBox="[123,1168,751,1574]" captionTargetId="figure-307@3.[129,1156,763,1571]" captionTargetPageId="3" captionText="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm VilertiO gOleOtO gen.nov.sp.nov.A:conquilla (paratip);B: òrgancopulador del mascle;C:gir intestinal;D-E:opercle (costatintern).Escala:1mm" figureDoi="http://doi.org/10.5281/zenodo.12170412" httpUri="https://zenodo.org/record/12170412/files/figure.png" pageId="4" pageNumber="164">Fig. 3B</figureCitation>
). Intestine with one loop (
<figureCitation id="131DB0D2E662FFE6FF09FF7F5D5DFF55" box="[253,319,187,209]" captionStart="FIGURE 3" captionStartId="3.[130,183,1586,1601]" captionTargetBox="[123,1168,751,1574]" captionTargetId="figure-307@3.[129,1156,763,1571]" captionTargetPageId="3" captionText="FIGURE 3. VilertiO gOleOtO gen.nov.sp.nov.A:shell (paratype);B: male copulatory organ;C:intestinal loop;D-E:operculum (inner side).Scale:1 mm VilertiO gOleOtO gen.nov.sp.nov.A:conquilla (paratip);B: òrgancopulador del mascle;C:gir intestinal;D-E:opercle (costatintern).Escala:1mm" figureDoi="http://doi.org/10.5281/zenodo.12170412" httpUri="https://zenodo.org/record/12170412/files/figure.png" pageId="4" pageNumber="164">Fig. 3C</figureCitation>
). Female sex tract with spherical bursa, receptaculum could not be found.
</paragraph>
</subSubSection>
<caption id="DF59FCDFE665FFE1FF76F9F6585DF9EC" pageId="3" pageNumber="163" startId="3.[130,183,1586,1601]" targetBox="[123,1168,751,1574]" targetPageId="3">
<paragraph id="8B99AC57E665FFE1FF76F9F65802F9C5" blockId="3.[130,1120,1586,1602]" box="[130,1120,1586,1602]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FF76F9F65CAAF9C5" bold="true" box="[130,200,1586,1601]" pageId="3" pageNumber="163">FIGURE 3.</emphasis>
<emphasis id="B9527045E665FFE1FF3FF9F75D4DF9C6" box="[203,303,1587,1602]" italics="true" pageId="3" pageNumber="163">Vilertia galeata</emphasis>
gen.nov.sp.nov.
<emphasis id="B9527045E665FFE1FE5CF9F65DD5F9C5" bold="true" box="[424,439,1586,1601]" pageId="3" pageNumber="163">A:</emphasis>
shell (paratype);
<emphasis id="B9527045E665FFE1FDDCF9F65E55F9C5" bold="true" box="[552,567,1586,1601]" pageId="3" pageNumber="163">B:</emphasis>
male copulatory organ;
<emphasis id="B9527045E665FFE1FD2CF9F65E85F9C5" bold="true" box="[728,743,1586,1601]" pageId="3" pageNumber="163">C:</emphasis>
intestinal loop;
<emphasis id="B9527045E665FFE1FCA4F9F65F0DF9C5" bold="true" box="[848,879,1586,1601]" pageId="3" pageNumber="163">D-E:</emphasis>
operculum (inner side).Scale:1 mm
</paragraph>
<paragraph id="8B99AC57E665FFE1FF77F99E585DF9EC" blockId="3.[131,1087,1625,1641]" box="[131,1087,1625,1641]" pageId="3" pageNumber="163">
<emphasis id="B9527045E665FFE1FF77F99E5C85F9ED" box="[131,231,1626,1641]" italics="true" pageId="3" pageNumber="163">Vilertia galeata</emphasis>
gen.nov.sp.nov.
<emphasis id="B9527045E665FFE1FE94F99D5D0DF9EC" bold="true" box="[352,367,1625,1640]" pageId="3" pageNumber="163">A:</emphasis>
conquilla (paratip);
<emphasis id="B9527045E665FFE1FE07F99D5E60F9EC" bold="true" box="[499,514,1625,1640]" pageId="3" pageNumber="163">B:</emphasis>
òrgan copulador del mascle;
<emphasis id="B9527045E665FFE1FD31F99D5EB6F9EC" bold="true" box="[709,724,1625,1640]" pageId="3" pageNumber="163">C:</emphasis>
gir intestinal;
<emphasis id="B9527045E665FFE1FCC4F99D5F2DF9EC" bold="true" box="[816,847,1625,1640]" pageId="3" pageNumber="163">D-E:</emphasis>
opercle (costat intern). Escala:1mm
</paragraph>
</caption>
<subSubSection id="C33CFFDCE662FFE6FF7CFF3F5D5DFED5" pageId="4" pageNumber="164" type="etymology">
<paragraph id="8B99AC57E662FFE6FF7CFF3F5D5DFED5" blockId="4.[113,624,155,499]" pageId="4" pageNumber="164">
<emphasis id="B9527045E662FFE6FF7CFF3F5C95FE94" bold="true" box="[136,247,251,272]" pageId="4" pageNumber="164">Etymology:</emphasis>
The specific name refers to the helmetshape of the shell, resembling the M15 Adrian French model of World War I.
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE662FFE6FF7CFE985C85FE56" pageId="4" pageNumber="164" type="description">
<paragraph id="8B99AC57E662FFE6FF7CFE985C85FE56" blockId="4.[113,624,155,499]" pageId="4" pageNumber="164">
<emphasis id="B9527045E662FFE6FF7CFE985D60FEF6" bold="true" box="[136,258,348,370]" pageId="4" pageNumber="164">Dimensions:</emphasis>
The
<typeStatus id="549D12F5E662FFE6FECCFE985DE9FEF6" box="[312,395,348,370]" pageId="4" pageNumber="164" type="holotype">holotype</typeStatus>
is
<quantity id="4CDE01B2E662FFE6FE5FFE985D95FEF6" box="[427,503,348,370]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="164" unit="mm" value="1.05">1.05 mm</quantity>
in diameter and
<quantity id="4CDE01B2E662FFE6FF5DFEB85C97FE16" box="[169,245,380,402]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="164" unit="mm" value="0.68">0.68 mm</quantity>
in height.
<typeStatus id="549D12F5E662FFE6FE73FEB95D81FE16" box="[391,483,381,402]" pageId="4" pageNumber="164" type="paratype">Paratypes</typeStatus>
range from
<quantity id="4CDE01B2E662FFE6FF85FE585C92FE36" box="[113,240,412,434]" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.27" metricValueMin="0.96" pageId="4" pageNumber="164" unit="mm" value="1.115" valueMax="1.27" valueMin="0.96">0.96 - 1.27 mm</quantity>
in diameter and
<quantity id="4CDE01B2E662FFE6FE60FE585E71FE36" box="[404,531,412,434]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.4" metricValueMin="5.1" pageId="4" pageNumber="164" unit="mm" value="0.625" valueMax="0.74" valueMin="0.51">0.51 - 0.74 mm</quantity>
in height (see Table I).
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE662FFE6FF7CFE19585CFF35" box="[136,1086,155,499]" pageId="4" pageNumber="164" type="biology_ecology">
<paragraph id="8B99AC57E662FFE6FF7CFE19585CFF35" blockId="4.[113,624,155,499]" box="[136,1086,155,499]" lastBlockId="4.[661,1173,155,501]" pageId="4" pageNumber="164">
<emphasis id="B9527045E662FFE6FF7CFE195CBAFE77" bold="true" box="[136,216,477,499]" pageId="4" pageNumber="164">Habitat:</emphasis>
Stygobiotic, all the shells were collected from sediments at the opening of the springs.
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE662FFE6FD58FF7F5F69FE95" pageId="4" pageNumber="164" type="distribution">
<paragraph id="8B99AC57E662FFE6FD58FF7F5F69FE95" blockId="4.[661,1173,155,501]" pageId="4" pageNumber="164">
<emphasis id="B9527045E662FFE6FD58FF7F5F49FF55" bold="true" box="[684,811,187,209]" pageId="4" pageNumber="164">Distribution:</emphasis>
Only known from two different springs, very close to each other, in the margins of the Fluvià River.
</paragraph>
</subSubSection>
<subSubSection id="C33CFFDCE662FFE6FD58FEDF5841FE76" pageId="4" pageNumber="164" type="diagnosis">
<paragraph id="8B99AC57E662FFE6FD58FEDF5841FE76" blockId="4.[661,1173,155,501]" pageId="4" pageNumber="164">
<emphasis id="B9527045E662FFE6FD58FEDF5FA4FEB5" bold="true" box="[684,966,283,305]" pageId="4" pageNumber="164">Differentiating characters:</emphasis>
<taxonomicName id="4C26D7D4E662FFE6FC29FEDA587FFEB0" authority="López-Soriano &amp; Quiñonero-Salgado &amp; Suárez &amp; Rolán &amp; Glöer, 2022" authorityName="López-Soriano &amp; Quiñonero-Salgado &amp; Suárez &amp; Rolán &amp; Glöer" authorityYear="2022" box="[989,1053,286,308]" class="Gastropoda" family="Hydrobiidae" genus="Vilertia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="164" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9527045E662FFE6FC29FEDA587FFEB0" box="[989,1053,286,308]" italics="true" pageId="4" pageNumber="164">Vilertia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261CD3EE662FFE6FBC4FED858F7FEB5" box="[1072,1173,284,305]" pageId="4" pageNumber="164" rank="genus">gen. nov.</taxonomicNameLabel>
differs from all valvatoid genera by the reflexed peristome. It differs from
<taxonomicName id="4C26D7D4E662FFE6FCAEFE985867FEF5" authority="Ghamizi 2020" authorityName="Ghamizi" authorityYear="2020" box="[858,1029,348,370]" class="Gastropoda" family="Hydrobiidae" genus="Rifia" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="164" phylum="Mollusca" rank="genus">
<emphasis id="B9527045E662FFE6FCAEFE985FE3FEF5" box="[858,897,348,369]" italics="true" pageId="4" pageNumber="164">Rifia</emphasis>
Ghamizi 2020
</taxonomicName>
by the missing outgrowth at the operculum; from
<taxonomicName id="4C26D7D4E662FFE6FC0EFEBB5EA3FE36" authority="Boeters, 1991" authorityName="Boeters" authorityYear="1991" class="Gastropoda" family="Hydrobiidae" genus="Fissuria" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="164" phylum="Mollusca" rank="genus">
<emphasis id="B9527045E662FFE6FC0EFEBB585EFE11" box="[1018,1084,383,405]" italics="true" pageId="4" pageNumber="164">Fissuria</emphasis>
Boeters, 1991
</taxonomicName>
, by the penis, which is papillose but without a penial lobe. In addition,
<taxonomicName id="4C26D7D4E662FFE6FC66FE7B5FB6FE51" authorityName="Boeters" authorityYear="1991" box="[914,980,447,469]" class="Gastropoda" family="Hydrobiidae" genus="Fissuria" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="164" phylum="Mollusca" rank="genus">
<emphasis id="B9527045E662FFE6FC66FE7B5FB6FE51" box="[914,980,447,469]" italics="true" pageId="4" pageNumber="164">Fissuria</emphasis>
</taxonomicName>
has two intestinal loops, while
<taxonomicName id="4C26D7D4E662FFE6FCFDFE1B5F2BFE71" authority="López-Soriano &amp; Quiñonero-Salgado &amp; Suárez &amp; Rolán &amp; Glöer, 2022" authorityName="López-Soriano &amp; Quiñonero-Salgado &amp; Suárez &amp; Rolán &amp; Glöer" authorityYear="2022" box="[777,841,479,501]" class="Gastropoda" family="Hydrobiidae" genus="Vilertia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="164" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9527045E662FFE6FCFDFE1B5F2BFE71" box="[777,841,479,501]" italics="true" pageId="4" pageNumber="164">Vilertia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261CD3EE662FFE6FCBBFE195FC6FE76" box="[847,932,477,498]" pageId="4" pageNumber="164" rank="genus">gen. nov.</taxonomicNameLabel>
has only one.
</paragraph>
</subSubSection>
</treatment>
</document>