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<document ID-CLB-Dataset="297111" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="f1b99209-6e3c-435d-8e27-eefdf36da9dc" ID-ISBN="0-8018-8221-4" checkinTime="1670905265806" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="645E257F11718AD132745D39D60A8027" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Soricomorpha.220-311.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Episoriculus caudatus subsp. sacratus Thomas 1911" docType="treatment" docVersion="3" id="BFF703DCC4E4C98D20FE0AB477F0FE06" lastPageNumber="277" masterDocId="BFF703DCC4E4C98D20FE0AB477F0FE06" masterDocTitle="Order Soricomorpha" masterLastPageNumber="311" masterPageNumber="220" pageNumber="277" updateTime="1716775530109" updateUser="ExternalLinkService">
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<taxonomicName id="F8DF7BC838C2C742DF142B8B86723A97" ID-CoL="5H596" authority="Thomas 1911" authorityName="Thomas" authorityYear="1911" class="Mammalia" family="Soricidae" genus="Episoriculus" kingdom="Animalia" order="Soricomorpha" pageNumber="277" phylum="Chordata" rank="subSpecies" species="caudatus" subSpecies="sacratus">
<emphasis id="B814259E441449A9FFEA43F07CB82231" italics="true" pageNumber="277">Episoriculus caudatus</emphasis>
subsp.
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Thomas 1911
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<emphasis id="5C1D2153CCAC13564F27D6D3B19E0810" inLineHeading="true" pageNumber="277">Synonyms:</emphasis>
<taxonomicName id="4292038DB9F40F2F62477E442A0F5E13" authority="(Gruber 1969)" baseAuthorityName="Gruber" baseAuthorityYear="1969" class="Mammalia" family="Soricidae" genus="Episoriculus" kingdom="Animalia" order="Soricomorpha" pageNumber="277" phylum="Chordata" rank="subSpecies" species="caudatus" subSpecies="soluensis">
<emphasis id="E6E3929C4D46DBDD9E7E0527E2A9AB7C" italics="true" pageNumber="277">Episoriculus caudatus</emphasis>
subsp.
<emphasis id="00B7CDABF8164610A8FB8CCB3F307622" italics="true" pageNumber="277">soluensis</emphasis>
(
<bibRefCitation id="95ABCB8628B4BF4BBB8A960C4FB4556E" author="Gruber, U. F." refId="ref134685" refString="Gruber, U. F. 1969. Tiergeographische, okologische und bionomische Untersuchungen an kleinen Saugetieren in Ost-Nepal. Khumbu Himal, 3: 197 - 312." year="1969">Gruber 1969</bibRefCitation>
)
</taxonomicName>
.
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<heading id="B70087168C40FFD2FF1AFC50FE3BA5C1" bold="true" box="[151,497,906,932]" fontSize="11" level="1" pageId="1" pageNumber="474" reason="1">
<taxonomicName id="2BF74BF98C40FFD2FF1AFC50FE3BA5C1" ID-CoL="8BHWR" authority="Meigen, 1826" authorityName="Meigen" authorityYear="1826" box="[151,497,906,932]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C40FFD2FF1AFC50FE3BA5C1" bold="true" box="[151,497,906,932]" pageId="1" pageNumber="474">
<emphasis id="DE83EC688C40FFD2FF1AFC50FE89A5C1" bold="true" box="[151,323,906,932]" italics="true" pageId="1" pageNumber="474">Chyliza vittata</emphasis>
Meigen, 1826
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(
<figureCitation id="74CC2CFF8C40FFD2FF12FC77FEC3A5A0" box="[159,265,941,965]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1897,1920]" captionStartId-1="3.[151,250,1069,1092]" captionStartId-2="4.[151,250,1132,1155]" captionTargetBox-0="[276,1307,339,1864]" captionTargetBox-1="[170,1429,179,1050]" captionTargetId-0="figure@2.[276,1307,339,1866]" captionTargetId-1="figure@3.[151,1436,173,1063]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Chyliza vittata (= Orchidophaga gastrodiacola) a, right wing (Cz. 1001) in dorsal view. b, male 4 th and 5 th abdominal sternum (Cz. 1001); c, female postabdomen in left lateral view (Cz. 1002). Abbreviations: em: eversible membrane; 4 S, 4 th abdominal sternum; 5 S, 5 th abdominal sternum; 6 S and 6 T: 6 th abdominal sternum and tergum; ST 7 and ST 8: 7 th and 8 th syntergosterna." captionText-1="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." captionText-2="FIGURE 3. Distribution map of Chyliza vittata in Japan." httpUri-0="https://zenodo.org/record/218163/files/figure.png" httpUri-1="https://zenodo.org/record/218164/files/figure.png" pageId="1" pageNumber="474">Figs. 13</figureCitation>
)
</paragraph>
<paragraph id="EC48307A8C40FFD2FF1AFC2FFD67A26E" blockId="1.[151,892,1013,1066]" box="[151,685,1013,1035]" pageId="1" pageNumber="474">
<taxonomicName id="2BF74BF98C40FFD2FF1AFC2FFE3EA26E" authority="Meigen, 1826: 369" authorityName="Meigen" authorityPageNumber="369" authorityYear="1826" box="[151,500,1013,1035]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C40FFD2FF1AFC2FFEE6A26E" box="[151,300,1013,1035]" italics="true" pageId="1" pageNumber="474">Chyliza vittata</emphasis>
Meigen, 1826: 369
</taxonomicName>
; Iwasa 1989: 160.
</paragraph>
</subSubSection>
<subSubSection id="A4ED63F18C40FFD0FF1AFBCEFEB4A0A8" lastPageId="3" lastPageNumber="476" pageId="1" pageNumber="474" type="description">
<paragraph id="EC48307A8C40FFD2FF1AFBCEFCB6A24C" blockId="1.[151,892,1013,1066]" box="[151,892,1044,1066]" pageId="1" pageNumber="474">
<taxonomicName id="2BF74BF98C40FFD2FF1AFBCEFDF9A24F" authority="Kato, 2006" authorityName="Kato" authorityYear="2006" box="[151,563,1044,1066]" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C40FFD2FF1AFBCEFE76A24F" box="[151,444,1044,1066]" italics="true" pageId="1" pageNumber="474">Orchidophaga gastrodiacola</emphasis>
Kato,
<quantity id="2B0F9D9F8C40FFD2FD8DFBCEFDF9A24F" box="[512,563,1044,1066]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="1" pageNumber="474" unit="in" value="2006.0">2006</quantity>
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in Kato
<emphasis id="DE83EC688C40FFD2FD07FBCFFD7CA24F" box="[650,694,1044,1066]" italics="true" pageId="1" pageNumber="474">et al</emphasis>
. 2006: 856.
<emphasis id="DE83EC688C40FFD2FCBBFBCEFCB6A24C" bold="true" box="[822,892,1044,1065]" pageId="1" pageNumber="474">syn. n.</emphasis>
</paragraph>
<paragraph id="EC48307A8C40FFD2FF1AFB8DFEB2A2F7" blockId="1.[151,1437,1111,2038]" pageId="1" pageNumber="474">
Male. Body yellow to orange in ground color, with black markings. Body length
<quantity id="2B0F9D9F8C40FFD2FBA1FB8DFB7EA20A" box="[1068,1204,1111,1135]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.5" metricValueMin="5.3" pageId="1" pageNumber="474" unit="mm" value="5.9" valueMax="6.5" valueMin="5.3">5.36.5 mm</quantity>
(n=18), wing length
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(n=21).
</paragraph>
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Head yellow, with dark brown marking on posterior side of occiput, 4/3 as high as long; frons yellow, with sparse, fine, yellow setae; face whitish yellow, without dark markings; gena yellow, covered with yellow setae; genal seta long and black. Head chaetotaxy: 1
<emphasis id="DE83EC688C40FFD2FD31FB32FD1DA29A" box="[700,727,1256,1279]" italics="true" pageId="1" pageNumber="474">oc</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FC8DFB32FCE3A29A" box="[768,809,1256,1279]" italics="true" pageId="1" pageNumber="474">poc</emphasis>
, 2
<emphasis id="DE83EC688C40FFD2FCDEFB32FCA1A29A" box="[851,875,1256,1279]" italics="true" pageId="1" pageNumber="474">or</emphasis>
, 2
<emphasis id="DE83EC688C40FFD2FC19FB32FC7AA29A" box="[916,944,1256,1279]" italics="true" pageId="1" pageNumber="474">ivt</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FC56FB32FC37A29A" box="[987,1021,1256,1279]" italics="true" pageId="1" pageNumber="474">ovt</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FBAAFB32FB83A29A" box="[1063,1097,1256,1279]" italics="true" pageId="1" pageNumber="474">pvt</emphasis>
setae; orbital setae reclinate. Antenna yellow; scape and pedicel with fine black setae; postpedicel with fine yellow setae; arista dark brown with short pubescence. Proboscis yellow; palpus dark brown to black, with distinct black setae, as long as labium.
</paragraph>
<paragraph id="EC48307A8C40FFD2FF44FA88FAF3A39F" blockId="1.[151,1437,1111,2038]" pageId="1" pageNumber="474">
Thorax yellow to brown, with dark brown to black markings on tergites and pleura; anterior margin of scutum, propleural lobe, anepisternum, katepisternum at middle, anepimeron, laterotergite, mediotergite, and dorsal half of meron dark brown; scutum yellow in ground color, with distinct dark marking on anterior 1/8 of scutum and indistinct longitudinal dark marking on posterior half of scutum anterior to
<emphasis id="DE83EC688C40FFD2FBA3FA65FB82A3B2" box="[1070,1096,1471,1495]" italics="true" pageId="1" pageNumber="474">dc</emphasis>
; scutellum yellow. Thoracic chaetotaxy: 1
<emphasis id="DE83EC688C40FFD2FEB9FA3EFE92A39E" box="[308,344,1508,1531]" italics="true" pageId="1" pageNumber="474">acr</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FEF6FA39FE5CA39E" box="[379,406,1507,1531]" italics="true" pageId="1" pageNumber="474">dc</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FE35FA3EFE1BA39E" box="[440,465,1508,1531]" italics="true" pageId="1" pageNumber="474">sa</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FE79FA3EFDDAA39E" box="[500,528,1508,1531]" italics="true" pageId="1" pageNumber="474">pa</emphasis>
, 1
<emphasis id="DE83EC688C40FFD2FDBEFA39FD9DA39E" box="[563,599,1507,1531]" italics="true" pageId="1" pageNumber="474">npl</emphasis>
, 23
<emphasis id="DE83EC688C40FFD2FD1BFA39FD77A39E" box="[662,701,1507,1531]" italics="true" pageId="1" pageNumber="474">sctl</emphasis>
;
<emphasis id="DE83EC688C40FFD2FD41FA3EFD3BA39E" box="[716,753,1508,1531]" italics="true" pageId="1" pageNumber="474">acr</emphasis>
and
<emphasis id="DE83EC688C40FFD2FCAAFA39FC88A39E" box="[807,834,1507,1531]" italics="true" pageId="1" pageNumber="474">dc</emphasis>
positioned just before scuto-scutellar suture.
</paragraph>
<paragraph id="EC48307A8C40FFD2FF44F9DDFE1EA020" blockId="1.[151,1437,1111,2038]" pageId="1" pageNumber="474">
Wing (
<figureCitation id="74CC2CFF8C40FFD2FE98F9DDFE99A07A" box="[277,339,1543,1567]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1920]" captionTargetBox="[276,1307,339,1864]" captionTargetId="figure@2.[276,1307,339,1866]" captionTargetPageId="2" captionText="FIGURE 1. Chyliza vittata (= Orchidophaga gastrodiacola) a, right wing (Cz. 1001) in dorsal view. b, male 4 th and 5 th abdominal sternum (Cz. 1001); c, female postabdomen in left lateral view (Cz. 1002). Abbreviations: em: eversible membrane; 4 S, 4 th abdominal sternum; 5 S, 5 th abdominal sternum; 6 S and 6 T: 6 th abdominal sternum and tergum; ST 7 and ST 8: 7 th and 8 th syntergosterna." httpUri="https://zenodo.org/record/218163/files/figure.png" pageId="1" pageNumber="474">Fig. 1</figureCitation>
a) hyaline, with dark brown marking on wing apex from apex of cell r1 to apex of cell m and along vein M to crossvein dM-Cu.
</paragraph>
<paragraph id="EC48307A8C40FFD2FF44F988FDBBA0D4" blockId="1.[151,1437,1111,2038]" pageId="1" pageNumber="474">Legs entirely yellow, dark brown marking on femora variable, distinct or indistinct on apices of mid and hind femora; fore coxa without distinct black setae; fore and mid femora without distinct setae; fore tibia anteroventrally with 1020 distinct, short spine-like setae.</paragraph>
<paragraph id="EC48307A8C40FFD2FF44F967FE32A1EC" blockId="1.[151,1437,1111,2038]" pageId="1" pageNumber="474">
Abdomen entirely dark brown, covered with dark brown setae; 5th abdominal sternite divided into 3 sclerites, median sclerite with short spine-like setae on posterior half, 5th abdominal sternite consisting of lateral and median sclerites (
<figureCitation id="74CC2CFF8C40FFD2FE89F8DFFE8DA178" box="[260,327,1797,1821]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1920]" captionTargetBox="[276,1307,339,1864]" captionTargetId="figure@2.[276,1307,339,1866]" captionTargetPageId="2" captionText="FIGURE 1. Chyliza vittata (= Orchidophaga gastrodiacola) a, right wing (Cz. 1001) in dorsal view. b, male 4 th and 5 th abdominal sternum (Cz. 1001); c, female postabdomen in left lateral view (Cz. 1002). Abbreviations: em: eversible membrane; 4 S, 4 th abdominal sternum; 5 S, 5 th abdominal sternum; 6 S and 6 T: 6 th abdominal sternum and tergum; ST 7 and ST 8: 7 th and 8 th syntergosterna." httpUri="https://zenodo.org/record/218163/files/figure.png" pageId="1" pageNumber="474">Fig. 1</figureCitation>
b: 5S); epandrium (
<figureCitation id="74CC2CFF8C40FFD2FDA7F8DFFDBAA178" box="[554,624,1797,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="1" pageNumber="474">Fig. 2</figureCitation>
a, b: ep) dark brown; surstylus (
<figureCitation id="74CC2CFF8C40FFD2FC6EF8DFFBE3A178" box="[995,1065,1797,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="1" pageNumber="474">Fig. 2</figureCitation>
a, b: s) yellow to yellow brown, united with epandrium at base, tapered to apex; apical part of surstylus directed outward; hypandrium (
<figureCitation id="74CC2CFF8C40FFD2FAAAF8F3FAA1A124" box="[1319,1387,1833,1857]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="1" pageNumber="474">Fig. 2</figureCitation>
c, d: h) oval in ventral view, with lateral apodeme; distiphallus (
<figureCitation id="74CC2CFF8C40FFD2FCA8F897FCADA100" box="[805,871,1869,1893]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="1" pageNumber="474">Fig. 2</figureCitation>
c, e, f: dp) bifurcated, outer lateral side sclerotized, inner lateral side, membranous.
</paragraph>
<paragraph id="EC48307A8C40FFD1FF44F84FFA92A779" blockId="1.[151,1437,1111,2038]" lastBlockId="2.[151,1436,151,284]" lastPageId="2" lastPageNumber="475" pageId="1" pageNumber="474">
Female (body length
<quantity id="2B0F9D9F8C40FFD2FE3AF84FFD8BA1C8" box="[439,577,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="6.699999999999999" metricValueMax="6.8" metricValueMin="6.6" pageId="1" pageNumber="474" unit="mm" value="6.699999999999999" valueMax="6.8" valueMin="6.6">6.66.8 mm</quantity>
, n=3; wing length
<quantity id="2B0F9D9F8C40FFD2FC9CF84FFC53A1C8" box="[785,921,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="5.6" metricValueMin="5.3" pageId="1" pageNumber="474" unit="mm" value="5.449999999999999" valueMax="5.6" valueMin="5.3">5.35.6 mm</quantity>
, n=4) similar to male but differing as follows: scutum yellow in ground color, with longitudinal median dark brown marking from anterior margin of scutum to transverse suture, lateral dark marking on line of
<emphasis id="DE83EC688C40FFD2FD4FF804FD16A193" box="[706,732,2014,2038]" italics="true" pageId="1" pageNumber="474">dc</emphasis>
seta from posterior margin of propleural lobe to
<emphasis id="DE83EC688C40FFD2FA88F804FAEAA193" box="[1285,1312,2014,2038]" italics="true" pageId="1" pageNumber="474">dc</emphasis>
seta; these dark markings fused and forming broad dark marking on scutum in some specimens; no distinct setae on fore tibia; syntergosternum 7 (
<figureCitation id="74CC2CFF8C43FFD1FEF0FF66FE0BA6B1" box="[381,449,188,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1897,1920]" captionTargetBox="[276,1307,339,1864]" captionTargetId="figure@2.[276,1307,339,1866]" captionTargetPageId="2" captionText="FIGURE 1. Chyliza vittata (= Orchidophaga gastrodiacola) a, right wing (Cz. 1001) in dorsal view. b, male 4 th and 5 th abdominal sternum (Cz. 1001); c, female postabdomen in left lateral view (Cz. 1002). Abbreviations: em: eversible membrane; 4 S, 4 th abdominal sternum; 5 S, 5 th abdominal sternum; 6 S and 6 T: 6 th abdominal sternum and tergum; ST 7 and ST 8: 7 th and 8 th syntergosterna." httpUri="https://zenodo.org/record/218163/files/figure.png" pageId="2" pageNumber="475">Fig. 1</figureCitation>
c: ST7) dark brown, with rod-like apodeme on anterior margin; eversible membrane shorter than syntergosternum 7; syntergosternum 8 longer than syntergosternum 7; 9th abdominal tergum (
<figureCitation id="74CC2CFF8C43FFD1FACFFF05FA4FA692" box="[1346,1413,223,247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="2" pageNumber="475">
Fig.
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</figureCitation>
g) and sternum distinct, as long as cercus; cercus (
<figureCitation id="74CC2CFF8C43FFD1FD27FEDEFD25A779" box="[682,751,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1069,1092]" captionTargetBox="[170,1429,179,1050]" captionTargetId="figure@3.[151,1436,173,1063]" captionTargetPageId="3" captionText="FIGURE 2. Chyliza vittata (= Orchidophaga gastrodiacola) a d, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz. 1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz. 1001). d, ventral view of hypandrial complex (Cz. 1001). e, f, phallus (Cz. 1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz. 1002). Scale bars: a d, 0.3 mm; e g, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8 ST: 8 th syntergosterna; 9 T and 9 S, 9 th abdominal tergum and sternum; c, cercus." httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="2" pageNumber="475">
Fig.
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</figureCitation>
g: c) round, with tuft of short setae, as long as epiproct.
</paragraph>
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<paragraph id="EC48307A8C43FFD1FF1AF8B3FE92A1BE" blockId="2.[151,1436,1897,2011]" pageId="2" pageNumber="475">
<emphasis id="DE83EC688C43FFD1FF1AF8B3FEDDA11B" bold="true" box="[151,279,1897,1920]" pageId="2" pageNumber="475">FIGURE 1.</emphasis>
<taxonomicName id="2BF74BF98C43FFD1FEACF8B3FE73A11A" box="[289,441,1897,1919]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C43FFD1FEACF8B3FE73A11A" box="[289,441,1897,1919]" italics="true" pageId="2" pageNumber="475">Chyliza vittata</emphasis>
</taxonomicName>
(=
<taxonomicName id="2BF74BF98C43FFD1FE69F8B3FCC7A11A" box="[484,781,1897,1919]" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C43FFD1FE69F8B3FCC7A11A" box="[484,781,1897,1919]" italics="true" pageId="2" pageNumber="475">Orchidophaga gastrodiacola</emphasis>
</taxonomicName>
) a, right wing (Cz.1001) in dorsal view. b, male 4th and 5th abdominal sternum (Cz. 1001); c, female postabdomen in left lateral view (Cz.1002). Abbreviations: em: eversible membrane; 4S, 4th abdominal sternum; 5S, 5th abdominal sternum; 6S and 6T: 6th abdominal sternum and tergum; ST7 and ST8: 7th and 8th syntergosterna.
</paragraph>
</caption>
<caption id="B88860F28C42FFD0FF1AFBF7FB3BA2B8" httpUri="https://zenodo.org/record/218164/files/figure.png" pageId="3" pageNumber="476" targetBox="[170,1429,179,1050]" targetPageId="3">
<paragraph id="EC48307A8C42FFD0FF1AFBF7FB3BA2B8" blockId="3.[151,1437,1069,1245]" pageId="3" pageNumber="476">
<emphasis id="DE83EC688C42FFD0FF1AFBF7FEDEA226" bold="true" box="[151,276,1069,1092]" pageId="3" pageNumber="476">FIGURE 2.</emphasis>
<taxonomicName id="2BF74BF98C42FFD0FE91FBF4FE79A221" box="[284,435,1070,1092]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C42FFD0FE91FBF4FE79A221" box="[284,435,1070,1092]" italics="true" pageId="3" pageNumber="476">Chyliza vittata</emphasis>
</taxonomicName>
(=
<taxonomicName id="2BF74BF98C42FFD0FE57FBF4FCCBA221" box="[474,769,1070,1092]" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C42FFD0FE57FBF4FCCBA221" box="[474,769,1070,1092]" italics="true" pageId="3" pageNumber="476">Orchidophaga gastrodiacola</emphasis>
</taxonomicName>
) ad, male. a, b, male epandrial complex (epandrium, surstylus, and proctiger) (Cz.1001). a, poserior view; b, ventral view; c, left lateral view of hypandrial complex (hypandrium and phallapodeme) and phallus (Cz.1001). d, ventral view of hypandrial complex (Cz.1001). e, f, phallus (Cz.1001). e, posterior view; f, ventral view. g, female proctiger in left lateral view (Cz.1002). Scale bars: ad, 0.3 mm; eg, 0.4 mm. Abbreviations: bp, basiphallus; dp, distiphallus; ep, epandrium; h, hypandrium; la, lateral apodeme of hypandrium; p, proctiger; pa, phallapodeme; s, surstylus; 8ST: 8th syntergosterna; 9T and 9S, 9th abdominal tergum and sternum; c, cercus.
</paragraph>
</caption>
<paragraph id="EC48307A8C42FFD0FF44FADEFEB4A0A8" blockId="3.[151,1436,1284,2029]" pageId="3" pageNumber="476">
<emphasis id="DE83EC688C42FFD0FF44FADEFE02A378" bold="true" box="[201,456,1284,1309]" pageId="3" pageNumber="476">Specimens examined.</emphasis>
<taxonomicName id="2BF74BF98C42FFD0FE42FADFFCD9A378" box="[463,787,1285,1309]" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C42FFD0FE42FADFFCD9A378" box="[463,787,1285,1309]" italics="true" pageId="3" pageNumber="476">Orchidophaga gastrodiacola</emphasis>
</taxonomicName>
<typeStatus id="334C8ED88C42FFD0FCA2FADFFC5DA378" box="[815,919,1285,1309]" pageId="3" pageNumber="476" type="holotype">Holotype</typeStatus>
(3): “
<typeStatus id="334C8ED88C42FFD0FC53FADFFB82A378" box="[990,1096,1285,1309]" pageId="3" pageNumber="476" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="2BF74BF98C42FFD0FBD5FADFFED9A325" authority="Kato, 2006" authorityName="Kato" authorityYear="2006" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C42FFD0FBD5FADFFA56A378" box="[1112,1436,1285,1309]" italics="true" pageId="3" pageNumber="476">Orchidophaga gastrodiacola</emphasis>
Kato, 2006
</taxonomicName>
/ 3 Aug. 2003, Kiso-Koma-Kogen, Kiso-cho,/ Nagano Pref.,
<collectingCountry id="94E070EA8C42FFD0FC49FAF3FBCDA325" box="[964,1031,1321,1344]" name="Japan" pageId="3" pageNumber="476">Japan</collectingCountry>
. collected by M. Kato,/ reared from flower shoot of
<taxonomicName id="2BF74BF98C42FFD0FEC4FA97FE32A300" box="[329,504,1357,1381]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="476" phylum="Tracheophyta" rank="species" species="elata">
<emphasis id="DE83EC688C42FFD0FEC4FA97FE32A300" box="[329,504,1357,1381]" italics="true" pageId="3" pageNumber="476">Gastrodia elata</emphasis>
</taxonomicName>
/”, “
<typeStatus id="334C8ED88C42FFD0FDA9FA97FD5BA300" box="[548,657,1357,1381]" pageId="3" pageNumber="476" type="holotype">Holotype</typeStatus>
” [hand-written], “03.8.3. KSO/ Oninoyagara [in Japanese characters] FR/ v.
<geoCoordinate id="89C356BD8C42FFD0FF51FAAAFEECA3ED" box="[220,294,1392,1416]" direction="east" orientation="longitude" pageId="3" pageNumber="476" precision="555" value="4.1">4.10 e</geoCoordinate>
” [hand-written], “photographed in 2007 [in Japanese characters]/ 1129156”. Other specimens: 13, same locality and date as
<typeStatus id="334C8ED88C42FFD0FE4BFA4FFDE2A3C8" box="[454,552,1429,1453]" pageId="3" pageNumber="476" type="holotype">holotype</typeStatus>
(Cz.1001) (KUM): 1Ƥ, same locality,
<date id="984916BA8C42FFD0FC63FA4FFBB8A3C8" box="[1006,1138,1429,1453]" pageId="3" pageNumber="476" value="2003-08-01">1. viii.2003</date>
(Cz.1002) (KUM). Other specimens:
<taxonomicName id="2BF74BF98C42FFD0FE96FA63FE75A3B4" box="[283,447,1465,1489]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C42FFD0FE96FA63FE75A3B4" box="[283,447,1465,1489]" italics="true" pageId="3" pageNumber="476">Chyliza vittata</emphasis>
</taxonomicName>
[HOKKAIDO] 83, Hamanaka, Rebun T,
<date id="984916BA8C42FFD0FC3FFA62FBF9A3B5" box="[946,1075,1464,1488]" pageId="3" pageNumber="476" value="2011-07-25">25.vii.2011</date>
, N Sugiura leg. (Cz.
<date id="984916BA8C42FFD0FA9BFA62FA5EA3B5" box="[1302,1428,1464,1488]" pageId="3" pageNumber="476" value="2001" valueMax="2008">2001-2008</date>
) (BLKU); 33, same locality,
<date id="984916BA8C42FFD0FE73FA07FDB5A390" box="[510,639,1501,1525]" pageId="3" pageNumber="476" value="2011-07-26">26.vii.2011</date>
, N Sugiura leg. (Cz.
<date id="984916BA8C42FFD0FC0AFA07FBC0A390" box="[903,1034,1501,1525]" pageId="3" pageNumber="476" value="2009" valueMax="2011">20092011</date>
) (BLKU); 432Ƥ, same locality,
<date id="984916BA8C42FFD0FF1AF9DAFED2A07D" box="[151,280,1536,1560]" pageId="3" pageNumber="476" value="2011-07-27">27.vii.2011</date>
, N Sugiura leg. (Cz.
<date id="984916BA8C42FFD0FD88F9DAFD40A07D" box="[517,650,1536,1560]" pageId="3" pageNumber="476" value="2012" valueMax="2017">20122017</date>
) (BLKU); 23, same locality,
<date id="984916BA8C42FFD0FC64F9DAFBA0A07D" box="[1001,1130,1536,1560]" pageId="3" pageNumber="476" value="2011-07-28">28.vii.2011</date>
, N Sugiura leg. (Cz.2018, 2019) (BLKU); 23, Kushuko, Rebun T,
<date id="984916BA8C42FFD0FDE8F9FFFD2CA058" box="[613,742,1573,1597]" pageId="3" pageNumber="476" value="2011-07-29">29.vii.2011</date>
, N Sugiura leg. (Cz.2020, 2021) (BLKU). [Honshu] Iwate P: 13, Mannyokaisawa, Kawai V,
<date id="984916BA8C42FFD0FDABF992FD68A005" box="[550,674,1608,1632]" pageId="3" pageNumber="476" value="1996-08-09">9.viii.1996</date>
, M Sueyoshi leg. (Cz.2022) (BLKU).
<collectingCountry id="94E070EA8C42FFD0FB0CF992FB14A005" box="[1153,1246,1608,1632]" name="Sweden" pageId="3" pageNumber="476">Sweden</collectingCountry>
: 13, Messaure, Jokkmokk kommun,
<date id="984916BA8C42FFD0FE1CF9B7FDD9A0E0" box="[401,531,1645,1669]" pageId="3" pageNumber="476" value="2012-07-12">12.vii.2012</date>
, M Sueyoshi leg. (Cz.2023) (BLKU).
<collectingCountry id="94E070EA8C42FFD0FC6AF9B7FB9EA0E0" box="[999,1108,1645,1669]" name="Denmark" pageId="3" pageNumber="476">Denmark</collectingCountry>
: 13, undetermined locality,
<date id="984916BA8C42FFD0FF1AF94AFEDBA0CD" box="[151,273,1680,1704]" pageId="3" pageNumber="476" value="1976-06-18">18.vi.1976</date>
, collector unknown (Cz3001) (ZMCU); 1Ƥ, undetermined locality,
<date id="984916BA8C42FFD0FBB4F94AFB7EA0CD" box="[1081,1204,1680,1704]" pageId="3" pageNumber="476" value="1906-06-22">22.vi.1906</date>
, collector unknown (CZ.3002) (ZMCU).
</paragraph>
</subSubSection>
<subSubSection id="A4ED63F18C42FFD0FF44F902FBF1A170" pageId="3" pageNumber="476" type="distribution">
<paragraph id="EC48307A8C42FFD0FF44F902FBF1A170" blockId="3.[151,1436,1284,2029]" pageId="3" pageNumber="476">
<emphasis id="DE83EC688C42FFD0FF44F902FEA8A094" bold="true" box="[201,354,1752,1777]" pageId="3" pageNumber="476">Distribution.</emphasis>
Europe (Soós 1984).
<collectingCountry id="94E070EA8C42FFD0FDDBF903FD50A095" box="[598,666,1753,1776]" name="Japan" pageId="3" pageNumber="476">Japan</collectingCountry>
: Hokkaido including Kunashiri and Rishiri Is. (Iwasa 1989), Honshu (Kato
<emphasis id="DE83EC688C42FFD0FF50F924FEC5A170" box="[221,271,1789,1813]" italics="true" pageId="3" pageNumber="476">et al</emphasis>
. 2006), Kyushu (Iwasa 1989) (
<figureCitation id="74CC2CFF8C42FFD0FDE1F927FD7AA170" box="[620,688,1789,1813]" captionStart="FIGURE 3" captionStartId="4.[151,250,1132,1155]" captionText="FIGURE 3. Distribution map of Chyliza vittata in Japan." pageId="3" pageNumber="476">Fig. 3</figureCitation>
). This is a transpalaearctic species.
</paragraph>
</subSubSection>
<subSubSection id="A4ED63F18C42FFD0FF44F8FBFB1BA188" pageId="3" pageNumber="476" type="discussion">
<paragraph id="EC48307A8C42FFD0FF44F8FBFBA8A1E5" blockId="3.[151,1436,1284,2029]" pageId="3" pageNumber="476">
<emphasis id="DE83EC688C42FFD0FF44F8FBFEF7A15C" bold="true" box="[201,317,1825,1849]" pageId="3" pageNumber="476">Remarks.</emphasis>
<taxonomicName id="2BF74BF98C42FFD0FEC9F8FBFE22A15C" box="[324,488,1825,1849]" class="Insecta" family="Psilidae" genus="Chyliza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="DE83EC688C42FFD0FEC9F8FBFE22A15C" box="[324,488,1825,1849]" italics="true" pageId="3" pageNumber="476">Chyliza vittata</emphasis>
</taxonomicName>
is distinguished from other congeners in having a whitish yellow face, yellow frons, black palpus, dark brown laterotergite, fore coxa without distinct black setae, and yellow femora (Shatalkin 1997). It also has a yellow scutum with median and lateral longitudinal dark brown markings.
</paragraph>
<paragraph id="EC48307A8C42FFD0FF44F857FB1BA188" blockId="3.[151,1436,1284,2029]" pageId="3" pageNumber="476">
A pair of the specimens (Cz.1001, 1002) was reared from
<taxonomicName id="2BF74BF98C42FFD0FCDCF854FC62A1C0" box="[849,936,1933,1957]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="476" phylum="Tracheophyta" rank="species" species="elata">
<emphasis id="DE83EC688C42FFD0FCDCF854FCA8A1C0" box="[849,866,1934,1957]" italics="true" pageId="3" pageNumber="476">G</emphasis>
.
<emphasis id="DE83EC688C42FFD0FCFDF857FC62A1C0" box="[880,936,1933,1957]" italics="true" pageId="3" pageNumber="476">elata</emphasis>
</taxonomicName>
by M. Kato and was examined by him at the time of the description of
<taxonomicName id="2BF74BF98C42FFD0FE43F86BFD45A1AC" box="[462,655,1969,1993]" class="Insecta" family="Scathophagidae" genus="Orchidophaga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="gastrodiacola">
<emphasis id="DE83EC688C42FFD0FE43F86BFE28A1AD" box="[462,482,1969,1992]" italics="true" pageId="3" pageNumber="476">O</emphasis>
.
<emphasis id="DE83EC688C42FFD0FE78F86BFD45A1AC" box="[501,655,1969,1993]" italics="true" pageId="3" pageNumber="476">gastrodiacola</emphasis>
</taxonomicName>
but have not been assigned as
<typeStatus id="334C8ED88C42FFD0FB85F86BFB8DA1AC" box="[1032,1095,1969,1993]" pageId="3" pageNumber="476">types</typeStatus>
. A series of these specimens (Cz.
<date id="984916BA8C42FFD0FF4EF80FFE8BA188" box="[195,321,2005,2029]" pageId="3" pageNumber="476" value="2001" valueMax="2021">2001-2021</date>
) from Hokkaido were collected on flowers of
<taxonomicName id="2BF74BF98C42FFD0FCC5F80CFC55A188" box="[840,927,2005,2029]" class="Liliopsida" family="Orchidaceae" genus="Gastrodia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="476" phylum="Tracheophyta" rank="species" species="elata">
<emphasis id="DE83EC688C42FFD0FCC5F80CFC93A188" box="[840,857,2006,2029]" italics="true" pageId="3" pageNumber="476">G</emphasis>
.
<emphasis id="DE83EC688C42FFD0FCEAF80FFC55A188" box="[871,927,2005,2029]" italics="true" pageId="3" pageNumber="476">elata</emphasis>
</taxonomicName>
(N. Sugiura, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>