Add updates up until 2025-02-18 16:32:23

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<paragraph id="8B8136A4FFFFFFEFFDAAFADD8D90FABF" blockId="10.[354,1235,1244,1341]" box="[597,991,1315,1341]" pageId="10" pageNumber="118">
<figureCitation id="13052A21FFFFFFEFFDAAFADD8CF2FABF" box="[597,701,1315,1341]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="11.[189,232,1928,1954]" captionStartId-1="12.[189,232,1928,1954]" captionStartId-2="15.[189,232,1963,1989]" captionStartId-3="16.[189,232,1963,1989]" captionStartId-4="17.[187,230,1963,1989]" captionStartId-5="18.[189,232,1998,2024]" captionStartId-6="19.[189,232,1998,2024]" captionStartId-7="20.[189,232,1998,2024]" captionTargetBox-0="[307,1280,265,1862]" captionTargetBox-1="[293,1299,268,1854]" captionTargetBox-2="[191,1397,830,1925]" captionTargetBox-3="[206,1356,1134,1881]" captionTargetBox-4="[244,1359,1159,1901]" captionTargetBox-5="[208,1379,1009,1960]" captionTargetBox-6="[223,1373,956,1947]" captionTargetBox-7="[189,1398,1256,1960]" captionTargetId-0="figure-38@11.[299,1289,262,1867]" captionTargetId-1="figure-37@12.[284,1303,262,1858]" captionTargetId-2="figure-184@15.[191,1397,830,1925]" captionTargetId-3="figure-186@16.[199,1398,1087,1922]" captionTargetId-4="figure-301@17.[195,1392,1132,1925]" captionTargetId-5="figure-265@18.[208,1381,1009,1961]" captionTargetId-6="figure-243@19.[195,1395,935,1963]" captionTargetId-7="figure-366@20.[186,1398,1256,1960]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionTargetPageId-6="19" captionTargetPageId-7="20" captionText-0="Fig. 1. Bayesian Maximum Likelihood tree based on partial COI gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML)." captionText-1="Fig. 2. Bayesian Maximum Likelihood tree based on Cytb gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML)." captionText-2="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." captionText-3="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." captionText-4="Fig. 5. Dorsolateral (above), dorsal (middle) and ventral (below) views of male paratype IEBR R.5259 (left) and female paratype IEBR R.5255 (right) of Acanthosaura cuongi sp. nov. in preservative." captionText-5="Fig. 6. Color variations of live paratypes of Acanthosaura cuongi sp. nov." captionText-6="Fig. 7. Distribution of Acanthosaura cuongi sp. nov. in Vietnam (violet circles)." captionText-7="Fig. 8. Forest habitat of Acanthosaura cuongi sp. nov in Phu Yen Province, Vietnam." pageId="10" pageNumber="118">Figs 18</figureCitation>
<figureCitation id="13052A21FFFFFFEFFDAAFADD8CF2FABF" box="[597,701,1315,1341]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="11.[189,232,1928,1954]" captionStartId-1="12.[189,232,1928,1954]" captionStartId-2="15.[189,232,1963,1989]" captionStartId-3="16.[189,232,1963,1989]" captionStartId-4="17.[187,230,1963,1989]" captionStartId-5="18.[189,232,1998,2024]" captionStartId-6="19.[189,232,1998,2024]" captionStartId-7="20.[189,232,1998,2024]" captionTargetBox-0="[307,1280,265,1862]" captionTargetBox-1="[293,1299,268,1854]" captionTargetBox-2="[191,1397,830,1925]" captionTargetBox-3="[206,1356,1134,1881]" captionTargetBox-4="[244,1359,1159,1901]" captionTargetBox-5="[208,1379,1009,1960]" captionTargetBox-6="[223,1373,956,1947]" captionTargetBox-7="[189,1398,1256,1960]" captionTargetId-0="figure-38@11.[299,1289,262,1867]" captionTargetId-1="figure-37@12.[284,1303,262,1858]" captionTargetId-2="figure-184@15.[191,1397,830,1925]" captionTargetId-3="figure-186@16.[199,1398,1087,1922]" captionTargetId-4="figure-301@17.[195,1392,1132,1925]" captionTargetId-5="figure-265@18.[208,1381,1009,1961]" captionTargetId-6="figure-243@19.[195,1395,935,1963]" captionTargetId-7="figure-366@20.[186,1398,1256,1960]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionTargetPageId-6="19" captionTargetPageId-7="20" captionText-0="Fig. 1. Bayesian Maximum Likelihood tree based on partial COI gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML)." captionText-1="Fig. 2. Bayesian Maximum Likelihood tree based on Cytb gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML)." captionText-2="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." captionText-3="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." captionText-4="Fig. 5. Dorsolateral (above), dorsal (middle) and ventral (below) views of male paratype IEBR R.5259 (left) and female paratype IEBR R.5255 (right) of Acanthosaura cuongi sp. nov. in preservative." captionText-5="Fig. 6. Color variations of live paratypes of Acanthosaura cuongi sp. nov." captionText-6="Fig. 7. Distribution of Acanthosaura cuongi sp. nov. in Vietnam (violet circles)." captionText-7="Fig. 8. Forest habitat of Acanthosaura cuongi sp. nov in Phu Yen Province, Vietnam." figureDoi-0="http://doi.org/10.5281/zenodo.14888565" figureDoi-1="http://doi.org/10.5281/zenodo.14888569" figureDoi-2="http://doi.org/10.5281/zenodo.14888573" figureDoi-3="http://doi.org/10.5281/zenodo.14888575" figureDoi-4="http://doi.org/10.5281/zenodo.14888577" figureDoi-5="http://doi.org/10.5281/zenodo.14888579" figureDoi-6="http://doi.org/10.5281/zenodo.14888581" figureDoi-7="http://doi.org/10.5281/zenodo.14888583" httpUri-0="https://zenodo.org/record/14888565/files/figure.png" httpUri-1="https://zenodo.org/record/14888569/files/figure.png" httpUri-2="https://zenodo.org/record/14888573/files/figure.png" httpUri-3="https://zenodo.org/record/14888575/files/figure.png" httpUri-4="https://zenodo.org/record/14888577/files/figure.png" httpUri-5="https://zenodo.org/record/14888579/files/figure.png" httpUri-6="https://zenodo.org/record/14888581/files/figure.png" httpUri-7="https://zenodo.org/record/14888583/files/figure.png" pageId="10" pageNumber="118">Figs 18</figureCitation>
;
<tableCitation id="C6BC031FFFFFFFEFFD36FADD8D03FABF" box="[713,844,1315,1341]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,265,291]" captionStartId-1="14.[189,258,265,291]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Table 3. Genetic distances (uncorrected p-distances) among samples and between species based on partial COI gene sequences." captionText-1="Table 4. Genetic distances (uncorrected p-distances) among samples and between species based on partial Cytb gene sequences." pageId="10" pageNumber="118">Tables 34</tableCitation>
<tableCitation id="C6BC031FFFFFFFEFFD36FADD8D03FABF" box="[713,844,1315,1341]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,265,291]" captionStartId-1="14.[189,258,265,291]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Table 3. Genetic distances (uncorrected p-distances) among samples and between species based on partial COI gene sequences." captionText-1="Table 4. Genetic distances (uncorrected p-distances) among samples and between species based on partial Cytb gene sequences." httpUri-0="http://table.plazi.org/id/DF41662CFFF8FFE8FF42FEF78C48FEC5" httpUri-1="http://table.plazi.org/id/DF41662CFFFBFFEBFF42FEF78C42FEC5" pageId="10" pageNumber="118" tableUuid-0="DF41662CFFF8FFE8FF42FEF78C48FEC5" tableUuid-1="DF41662CFFFBFFEBFF42FEF78C42FEC5">Tables 34</tableCitation>
; Supp. file 1
</paragraph>
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@ -152,7 +155,7 @@ in females); head triangular, relatively long (HL/SVL 0.270.31, HW/SVL 0.18
males and
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females), postorbital and occipital spines present; gular pouch undeveloped; 810 scales between nasals; four scales bordering mental scale; 1013 supralabials; 1013 infralabials; 57 scales bordering rostral scale; nuchal crest composed of 79 spines relatively high and one spine short; dorsal crest present, undeveloped; double rows of vertebral scales followed the dorsal crest; nuchal and dorsal crests continuous, filled with small spine scales; lateral scales small, intermixed with large, keeled scales, keels directed backward and back-upward; number of subdigital lamellae 1417 on fourth finger and 1923 on fourth toe (
<tableCitation id="C6BC031FFFFFFFEFFBEEF9538A3FF94A" box="[1041,1136,1709,1736]" captionStart="Table 3" captionStartId="13.[189,258,265,291]" captionTargetPageId="13" captionText="Table 3. Genetic distances (uncorrected p-distances) among samples and between species based on partial COI gene sequences." pageId="10" pageNumber="118">Table 3</tableCitation>
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); black nuchal collar present; dorsum with black bands; a black patch extending from eye posteriorly to the anterior edge of tympanum.
</paragraph>
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@ -174,19 +177,19 @@ provinces and his great contributions to herpetological research in
.
</paragraph>
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<caption id="DF41662CFFFEFFEEFF42F8768D20F86A" ID-DOI="http://doi.org/10.5281/zenodo.14888565" ID-Zenodo-Dep="14888565" httpUri="https://zenodo.org/record/14888565/files/figure.png" pageId="11" pageNumber="119" startId="11.[189,232,1928,1954]" targetBox="[307,1280,265,1862]" targetPageId="11">
<paragraph id="8B8136A4FFFEFFEEFF42F8768D20F86A" blockId="11.[188,1399,1928,2024]" pageId="11" pageNumber="119">
<emphasis id="B94AEAB6FFFEFFEEFF42F8768F45F820" bold="true" box="[189,266,1928,1954]" pageId="11" pageNumber="119">Fig. 1.</emphasis>
Bayesian Maximum Likelihood tree based on partial COI gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML).
</paragraph>
</caption>
<caption id="DF41662CFFF9FFE9FF42F8768D62F86A" pageId="12" pageNumber="120" startId="12.[189,232,1928,1954]" targetBox="[293,1299,268,1854]" targetPageId="12">
<caption id="DF41662CFFF9FFE9FF42F8768D62F86A" ID-DOI="http://doi.org/10.5281/zenodo.14888569" ID-Zenodo-Dep="14888569" httpUri="https://zenodo.org/record/14888569/files/figure.png" pageId="12" pageNumber="120" startId="12.[189,232,1928,1954]" targetBox="[293,1299,268,1854]" targetPageId="12">
<paragraph id="8B8136A4FFF9FFE9FF42F8768D62F86A" blockId="12.[188,1399,1928,2024]" pageId="12" pageNumber="120">
<emphasis id="B94AEAB6FFF9FFE9FF42F8768F44F820" bold="true" box="[189,267,1928,1954]" pageId="12" pageNumber="120">Fig. 2.</emphasis>
Bayesian Maximum Likelihood tree based on Cytb gene sequences of specimens used in this study. Values at each node indicate Bayesian Posterior Probabilities and Ultrafast Bootstrap Values from Maximum Likelihood analysis respectively (BI/ML).
</paragraph>
</caption>
<caption id="DF41662CFFF8FFE8FF42FEF78C48FEC5" pageId="13" pageNumber="121" startId="13.[189,258,265,291]" targetBox="[200,1367,369,1193]" targetIsTable="true" targetPageId="13">
<caption id="DF41662CFFF8FFE8FF42FEF78C48FEC5" ID-Table-UUID="DF41662CFFF8FFE8FF42FEF78C48FEC5" httpUri="http://table.plazi.org/id/DF41662CFFF8FFE8FF42FEF78C48FEC5" pageId="13" pageNumber="121" startId="13.[189,258,265,291]" targetBox="[200,1367,369,1193]" targetIsTable="true" targetPageId="13">
<paragraph id="8B8136A4FFF8FFE8FF42FEF78C48FEC5" blockId="13.[189,1398,265,327]" pageId="13" pageNumber="121">
<emphasis id="B94AEAB6FFF8FFE8FF42FEF78F6CFEA1" bold="true" box="[189,291,265,291]" pageId="13" pageNumber="121">Table 3.</emphasis>
Genetic distances (uncorrected p-distances) among samples and between species based on partial COI gene sequences.
@ -483,7 +486,7 @@ were recorded for each specimen and can be shared upon request to the authors to
<paragraph id="8B8136A4FFF8FFE8FF15F9098FB8F893" blockId="13.[186,1400,1783,2025]" box="[234,503,1783,1809]" pageId="13" pageNumber="121">
<emphasis id="B94AEAB6FFF8FFE8FF15F9098F3AF893" bold="true" box="[234,373,1783,1809]" pageId="13" pageNumber="121">Adult male</emphasis>
(
<figureCitation id="13052A21FFF8FFE8FE7AF9098FA2F893" box="[389,493,1783,1809]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1963,1989]" captionStartId-1="16.[189,232,1963,1989]" captionTargetBox-0="[191,1397,830,1925]" captionTargetBox-1="[206,1356,1134,1881]" captionTargetId-0="figure-184@15.[191,1397,830,1925]" captionTargetId-1="figure-186@16.[199,1398,1087,1922]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." captionText-1="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." pageId="13" pageNumber="121">Figs 34</figureCitation>
<figureCitation id="13052A21FFF8FFE8FE7AF9098FA2F893" box="[389,493,1783,1809]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1963,1989]" captionStartId-1="16.[189,232,1963,1989]" captionTargetBox-0="[191,1397,830,1925]" captionTargetBox-1="[206,1356,1134,1881]" captionTargetId-0="figure-184@15.[191,1397,830,1925]" captionTargetId-1="figure-186@16.[199,1398,1087,1922]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." captionText-1="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." figureDoi-0="http://doi.org/10.5281/zenodo.14888573" figureDoi-1="http://doi.org/10.5281/zenodo.14888575" httpUri-0="https://zenodo.org/record/14888573/files/figure.png" httpUri-1="https://zenodo.org/record/14888575/files/figure.png" pageId="13" pageNumber="121">Figs 34</figureCitation>
)
</paragraph>
<paragraph id="8B8136A4FFF8FFEBFF42F8E08F07F88C" blockId="13.[186,1400,1783,2025]" lastBlockId="14.[187,1399,1252,1806]" lastPageId="14" lastPageNumber="122" pageId="13" pageNumber="121">
@ -503,7 +506,7 @@ in height (HH/HL 0.55); snout moderately long (SL
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) at base of nape with four low continuous spines; dorsal crest composed of three single and four of two rows of slightly low dagger-like spines; followed by vertebral crest, composed of two rows of enlarged, keeled, pointed scales, arranged as a zipper line, ending at base of tail.
</paragraph>
<caption id="DF41662CFFFBFFEBFF42FEF78C42FEC5" pageId="14" pageNumber="122" startId="14.[189,258,265,291]" targetBox="[200,1379,369,1193]" targetIsTable="true" targetPageId="14">
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<paragraph id="8B8136A4FFFBFFEBFF42FEF78C42FEC5" blockId="14.[189,1398,265,327]" pageId="14" pageNumber="122">
<emphasis id="B94AEAB6FFFBFFEBFF42FEF78F6CFEA1" bold="true" box="[189,291,265,291]" pageId="14" pageNumber="122">Table 4.</emphasis>
Genetic distances (uncorrected p-distances) among samples and between species based on partial Cytb gene sequences.
@ -706,10 +709,10 @@ Coloration of live
</paragraph>
<paragraph id="8B8136A4FFFAFFEAFF42FE248B43FC8F" blockId="15.[187,1401,474,782]" pageId="15" pageNumber="123">
Dorsal surface of head brown-green with a short transverse band light-green between orbital regions; occipital surface light-green; a black patch extending from behind the eye backward and slightly upward to anterior margin of tympanum; upper and lower lips green; lateral side of neck grey-white in most parts; nuchal crest spines pastel-orange (third, fifth and sixth spines) and green (remaining spines); iris yellowbrown; tongue blueish gray; dorsal surface of body light-green with seven brownish blackblured stripes, oriented obliquely forward from both sides of the vertebral region and one on shoulder; dorsal surface of limbs light-green, light spots on elbows and knees present; gular region and ventral side yellow-white in most parts; ventral sides of limbs yellow-green in the upper half part and white in the remaining part; black and light-green rings on tail, the rings become indistinct on posterior part of tail (
<figureCitation id="13052A21FFFAFFEAFB49FD0D8AB3FC8F" box="[1206,1276,755,781]" captionStart="Fig" captionStartId="15.[189,232,1963,1989]" captionTargetBox="[191,1397,830,1925]" captionTargetId="figure-184@15.[191,1397,830,1925]" captionTargetPageId="15" captionText="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." pageId="15" pageNumber="123">Fig. 3</figureCitation>
<figureCitation id="13052A21FFFAFFEAFB49FD0D8AB3FC8F" box="[1206,1276,755,781]" captionStart="Fig" captionStartId="15.[189,232,1963,1989]" captionTargetBox="[191,1397,830,1925]" captionTargetId="figure-184@15.[191,1397,830,1925]" captionTargetPageId="15" captionText="Fig. 3. Live male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250). A. Dorsolateral view. B. Dorsal view. C. Cloacal region. D. Head, lateral view. E. Head, dorsal view. F. Head, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14888573" httpUri="https://zenodo.org/record/14888573/files/figure.png" pageId="15" pageNumber="123">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF41662CFFFAFFEAFF42F8558B27F86A" pageId="15" pageNumber="123" startId="15.[189,232,1963,1989]" targetBox="[191,1397,830,1925]" targetPageId="15">
<caption id="DF41662CFFFAFFEAFF42F8558B27F86A" ID-DOI="http://doi.org/10.5281/zenodo.14888573" ID-Zenodo-Dep="14888573" httpUri="https://zenodo.org/record/14888573/files/figure.png" pageId="15" pageNumber="123" startId="15.[189,232,1963,1989]" targetBox="[191,1397,830,1925]" targetPageId="15">
<paragraph id="8B8136A4FFFAFFEAFF42F8558B27F86A" blockId="15.[189,1399,1963,2025]" pageId="15" pageNumber="123">
<emphasis id="B94AEAB6FFFAFFEAFF42F8558F41F847" bold="true" box="[189,270,1963,1989]" pageId="15" pageNumber="123">Fig. 3.</emphasis>
Live male holotype of
@ -743,7 +746,7 @@ in preservative
</paragraph>
<paragraph id="8B8136A4FFE5FFF5FF42FEBA8A77FE03" blockId="16.[188,1398,323,385]" pageId="16" pageNumber="124">
In preservative, the light-green on the dorsal surface of the body, dorsal and ventral surface of limbs and rings on tail faded to dark gray but the pattern remained unchanged (
<figureCitation id="13052A21FFE5FFF5FC1DFE988A64FE03" box="[994,1067,358,385]" captionStart="Fig" captionStartId="16.[189,232,1963,1989]" captionTargetBox="[206,1356,1134,1881]" captionTargetId="figure-186@16.[199,1398,1087,1922]" captionTargetPageId="16" captionText="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." pageId="16" pageNumber="124">Fig. 4</figureCitation>
<figureCitation id="13052A21FFE5FFF5FC1DFE988A64FE03" box="[994,1067,358,385]" captionStart="Fig" captionStartId="16.[189,232,1963,1989]" captionTargetBox="[206,1356,1134,1881]" captionTargetId="figure-186@16.[199,1398,1087,1922]" captionTargetPageId="16" captionText="Fig. 4. Dorsolateral (above) and ventral (below) views of the male holotype of Acanthosaura cuongi sp. nov. (IEBR R.5250) in preservative." figureDoi="http://doi.org/10.5281/zenodo.14888575" httpUri="https://zenodo.org/record/14888575/files/figure.png" pageId="16" pageNumber="124">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B8136A4FFE5FFF5FF42FE4A8F71FE4C" blockId="16.[189,318,436,462]" box="[189,318,436,462]" pageId="16" pageNumber="124">
@ -757,7 +760,7 @@ Morphometric and meristic data for the type series are provided in Supp. file 1.
IEBR R.5258 lacks patches on the back;
<typeStatus id="54858806FFE5FFF5FDB0FDC88CFBFDD2" box="[591,692,566,592]" pageId="16" pageNumber="124" type="paratype">paratype</typeStatus>
IEBR R.5256 has brown upper and lower lips and the nuchal crest is grayish black, green and yellow (
<figureCitation id="13052A21FFE5FFF5FD64FDA68D48FDF1" box="[667,775,600,627]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="17.[187,230,1963,1989]" captionStartId-1="18.[189,232,1998,2024]" captionTargetBox-0="[244,1359,1159,1901]" captionTargetBox-1="[208,1379,1009,1960]" captionTargetId-0="figure-301@17.[195,1392,1132,1925]" captionTargetId-1="figure-265@18.[208,1381,1009,1961]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Fig. 5. Dorsolateral (above), dorsal (middle) and ventral (below) views of male paratype IEBR R.5259 (left) and female paratype IEBR R.5255 (right) of Acanthosaura cuongi sp. nov. in preservative." captionText-1="Fig. 6. Color variations of live paratypes of Acanthosaura cuongi sp. nov." pageId="16" pageNumber="124">Figs 56</figureCitation>
<figureCitation id="13052A21FFE5FFF5FD64FDA68D48FDF1" box="[667,775,600,627]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="17.[187,230,1963,1989]" captionStartId-1="18.[189,232,1998,2024]" captionTargetBox-0="[244,1359,1159,1901]" captionTargetBox-1="[208,1379,1009,1960]" captionTargetId-0="figure-301@17.[195,1392,1132,1925]" captionTargetId-1="figure-265@18.[208,1381,1009,1961]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Fig. 5. Dorsolateral (above), dorsal (middle) and ventral (below) views of male paratype IEBR R.5259 (left) and female paratype IEBR R.5255 (right) of Acanthosaura cuongi sp. nov. in preservative." captionText-1="Fig. 6. Color variations of live paratypes of Acanthosaura cuongi sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.14888577" figureDoi-1="http://doi.org/10.5281/zenodo.14888579" httpUri-0="https://zenodo.org/record/14888577/files/figure.png" httpUri-1="https://zenodo.org/record/14888579/files/figure.png" pageId="16" pageNumber="124">Figs 56</figureCitation>
).
</paragraph>
<paragraph id="8B8136A4FFE5FFF5FF42FD588FAAFD42" blockId="16.[189,485,678,704]" box="[189,485,678,704]" pageId="16" pageNumber="124">
@ -798,10 +801,10 @@ and Tay Hoa District of
, with findings recorded from nearby the coastline, south-central
<collectingCountry id="F3297634FFE5FFF5FEA2FC088F8FFB92" box="[349,448,1014,1040]" name="Vietnam" pageId="16" pageNumber="124">Vietnam</collectingCountry>
(
<figureCitation id="13052A21FFE5FFF5FE2FFC088C57FB92" box="[464,536,1014,1040]" captionStart="Fig" captionStartId="19.[189,232,1998,2024]" captionTargetBox="[223,1373,956,1947]" captionTargetId="figure-243@19.[195,1395,935,1963]" captionTargetPageId="19" captionText="Fig. 7. Distribution of Acanthosaura cuongi sp. nov. in Vietnam (violet circles)." pageId="16" pageNumber="124">Fig. 7</figureCitation>
<figureCitation id="13052A21FFE5FFF5FE2FFC088C57FB92" box="[464,536,1014,1040]" captionStart="Fig" captionStartId="19.[189,232,1998,2024]" captionTargetBox="[223,1373,956,1947]" captionTargetId="figure-243@19.[195,1395,935,1963]" captionTargetPageId="19" captionText="Fig. 7. Distribution of Acanthosaura cuongi sp. nov. in Vietnam (violet circles)." figureDoi="http://doi.org/10.5281/zenodo.14888581" httpUri="https://zenodo.org/record/14888581/files/figure.png" pageId="16" pageNumber="124">Fig. 7</figureCitation>
).
</paragraph>
<caption id="DF41662CFFE5FFF5FF42F8558CDEF86A" pageId="16" pageNumber="124" startId="16.[189,232,1963,1989]" targetBox="[206,1356,1134,1881]" targetPageId="16">
<caption id="DF41662CFFE5FFF5FF42F8558CDEF86A" ID-DOI="http://doi.org/10.5281/zenodo.14888575" ID-Zenodo-Dep="14888575" httpUri="https://zenodo.org/record/14888575/files/figure.png" pageId="16" pageNumber="124" startId="16.[189,232,1963,1989]" targetBox="[206,1356,1134,1881]" targetPageId="16">
<paragraph id="8B8136A4FFE5FFF5FF42F8558CDEF86A" blockId="16.[189,1401,1963,2025]" pageId="16" pageNumber="124">
<emphasis id="B94AEAB6FFE5FFF5FF42F8558F42F847" bold="true" box="[189,269,1963,1989]" pageId="16" pageNumber="124">Fig. 4.</emphasis>
Dorsolateral (above) and ventral (below) views of the male holotype of
@ -825,7 +828,7 @@ a.s.l. in evergreen forests, belonging to the southeast Truong Son Mountain Rang
and
<collectingRegion id="49FAF846FFE4FFF4FC8DFEA28DB7FEF5" box="[882,1016,348,375]" country="Vietnam" name="Khanh Hoa" pageId="17" pageNumber="125">Khanh Hoa</collectingRegion>
provinces (
<figureCitation id="13052A21FFE4FFF4FB7EFEA28A88FEF4" box="[1153,1223,348,374]" captionStart="Fig" captionStartId="20.[189,232,1998,2024]" captionTargetBox="[189,1398,1256,1960]" captionTargetId="figure-366@20.[186,1398,1256,1960]" captionTargetPageId="20" captionText="Fig. 8. Forest habitat of Acanthosaura cuongi sp. nov in Phu Yen Province, Vietnam." pageId="17" pageNumber="125">Fig. 8</figureCitation>
<figureCitation id="13052A21FFE4FFF4FB7EFEA28A88FEF4" box="[1153,1223,348,374]" captionStart="Fig" captionStartId="20.[189,232,1998,2024]" captionTargetBox="[189,1398,1256,1960]" captionTargetId="figure-366@20.[186,1398,1256,1960]" captionTargetPageId="20" captionText="Fig. 8. Forest habitat of Acanthosaura cuongi sp. nov in Phu Yen Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.14888583" httpUri="https://zenodo.org/record/14888583/files/figure.png" pageId="17" pageNumber="125">Fig. 8</figureCitation>
). Lizards were observed during night surveys, clinging to, and sleeping in shrubs, thin trunks of small trees, beneath dense forest canopy. The height of occupied perches ranged from
<quantity id="4CC69B41FFE4FFF4FC41FE5D8A7BFE3F" box="[958,1076,419,445]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="17" pageNumber="125" unit="m" value="0.95" valueMax="1.8" valueMin="0.1">0.11.8 m</quantity>
above the ground. The new species occurs in sympatry with
@ -1011,7 +1014,7 @@ by having a shorter body length (max snout-vent length: 104.61 vs
<quantity id="4CC69B41FFE4FFF4FB04FBDD8B3DFBBC" box="[1275,1394,1059,1086]" metricMagnitude="-1" metricUnit="m" metricValue="1.184" pageId="17" pageNumber="125" unit="mm" value="118.4">118.4 mm</quantity>
, respectively), fewer subdigital lamellae on the fourth finger (1417 vs 23, 2021, respectively) and on the fourth toe (1923 vs 2628, 2327, respectively), fewer canthus rostralis scales (911 vs 1415, 1415, respectively), and the presence of light knee spot (vs absence) (Supp. file 2).
</paragraph>
<caption id="DF41662CFFE4FFF4FF44F8558B68F86A" pageId="17" pageNumber="125" startId="17.[187,230,1963,1989]" targetBox="[244,1359,1159,1901]" targetPageId="17">
<caption id="DF41662CFFE4FFF4FF44F8558B68F86A" ID-DOI="http://doi.org/10.5281/zenodo.14888577" ID-Zenodo-Dep="14888577" httpUri="https://zenodo.org/record/14888577/files/figure.png" pageId="17" pageNumber="125" startId="17.[187,230,1963,1989]" targetBox="[244,1359,1159,1901]" targetPageId="17">
<paragraph id="8B8136A4FFE4FFF4FF44F8558B68F86A" blockId="17.[187,1399,1963,2025]" pageId="17" pageNumber="125">
<emphasis id="B94AEAB6FFE4FFF4FF44F8558F44F847" bold="true" box="[187,267,1963,1989]" pageId="17" pageNumber="125">Fig. 5.</emphasis>
Dorsolateral (above), dorsal (middle) and ventral (below) views of male paratype IEBR R.5259 (left) and female paratype IEBR R.5255 (right) of
@ -1083,7 +1086,7 @@ by having a shorter nuchal crest spines (1.632.66 vs
<quantity id="4CC69B41FFE6FFF6FC7FFEB18A5EFEE8" box="[896,1041,335,362]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.8" metricValueMin="3.6" pageId="19" pageNumber="127" unit="mm" value="4.2" valueMax="4.8" valueMin="3.6">3.64.8 mm</quantity>
), fewer canthus rostralis scales (911 vs 12), fewer scales bordering the rostral scale (57 vs 89), fewer scales bordering the mental (4 vs 56), the absence of a diastema between nuchal and dorsal crests (vs presence) (Supp. file 2).
</paragraph>
<caption id="DF41662CFFE7FFF7FF42F8308A50F86B" box="[189,1055,1998,2025]" pageId="18" pageNumber="126" startId="18.[189,232,1998,2024]" targetBox="[208,1379,1009,1960]" targetPageId="18">
<caption id="DF41662CFFE7FFF7FF42F8308A50F86B" ID-DOI="http://doi.org/10.5281/zenodo.14888579" ID-Zenodo-Dep="14888579" box="[189,1055,1998,2025]" httpUri="https://zenodo.org/record/14888579/files/figure.png" pageId="18" pageNumber="126" startId="18.[189,232,1998,2024]" targetBox="[208,1379,1009,1960]" targetPageId="18">
<paragraph id="8B8136A4FFE7FFF7FF42F8308A50F86B" blockId="18.[189,1055,1998,2025]" box="[189,1055,1998,2025]" pageId="18" pageNumber="126">
<emphasis id="B94AEAB6FFE7FFF7FF42F8308F45F86A" bold="true" box="[189,266,1998,2024]" pageId="18" pageNumber="126">Fig. 6.</emphasis>
Color variations of live paratypes of
@ -1126,7 +1129,7 @@ by having a longer body length (79.4 104.61 vs
<quantity id="4CC69B41FFE6FFF6FF1FFCC58FC0FCD4" box="[224,399,827,854]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.59" metricValueMin="1.31" pageId="19" pageNumber="127" unit="mm" value="14.5" valueMax="15.9" valueMin="13.1">13.115.9 mm</quantity>
), a shorter orbit diameter (orbit/HL: 0.270.38 vs 0.40.57), two rows of slightly low dagger-like vertebral spines (vs single row, black eye patch extending backward and slightly upward to the anterior edge of the tympanum vs upward to through occipital spine to reach anterior edge of black nuchal collar) (Supp. file 2).
</paragraph>
<caption id="DF41662CFFE6FFF6FF42F8308A2AF86A" box="[189,1125,1998,2025]" pageId="19" pageNumber="127" startId="19.[189,232,1998,2024]" targetBox="[223,1373,956,1947]" targetPageId="19">
<caption id="DF41662CFFE6FFF6FF42F8308A2AF86A" ID-DOI="http://doi.org/10.5281/zenodo.14888581" ID-Zenodo-Dep="14888581" box="[189,1125,1998,2025]" httpUri="https://zenodo.org/record/14888581/files/figure.png" pageId="19" pageNumber="127" startId="19.[189,232,1998,2024]" targetBox="[223,1373,956,1947]" targetPageId="19">
<paragraph id="8B8136A4FFE6FFF6FF42F8308A2AF86A" blockId="19.[189,1125,1998,2025]" box="[189,1125,1998,2025]" pageId="19" pageNumber="127">
<emphasis id="B94AEAB6FFE6FFF6FF42F8308F44F86A" bold="true" box="[189,267,1998,2024]" pageId="19" pageNumber="127">Fig. 7.</emphasis>
Distribution of