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<mods:title id="B7E5DAEFF24CE63FD0C67D0515657282">Chelonodontops bengalensis Habib et al. 2018, a junior synonym of Tetrodon patoca Hamilton 1822, with a type species designation for Chelonodon Müller 1841</mods:title>
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In his monograph on “the fishes of the River Ganges and its branches”,
<bibRefCitation id="EFB046F909534A23FBDFFADEFB255235" author="Hamilton, F." box="[1036,1236,1367,1393]" pageId="2" pageNumber="563" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamilton (1822)</bibRefCitation>
treated 272 fish species, which he encountered during his two sojourns in
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, 17941805 and 18071815, in the Brahmaputra and Ganges drainages (
<bibRefCitation id="EFB046F909534A23FEABFA16FDF552FD" author="Britz, R." box="[376,516,1439,1465]" pageId="2" pageNumber="563" refId="ref5222" refString="Britz, R. (2019) Francis Hamilton's Gangetic Fishes in colour. A new edition of the 1822 monograph, with reproductions of unpublished coloured illustrations. Ray Society, London, 48 + vii + 405 + vii pp., 228 pls." type="book" year="2019">Britz, 2019</bibRefCitation>
). Among them were six new species of
<taxonomicName id="4C21408B09534A23FC04FA16FB7D52FD" box="[983,1164,1439,1465]" family="Tetraodontidae" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="family">Tetraodontidae</taxonomicName>
, all of which he placed into
<taxonomicName id="4C21408B09534A23FF1FFA4AFEC55299" box="[204,308,1475,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09534A23FF1FFA4AFEC55299" box="[204,308,1475,1501]" italics="true" pageId="2" pageNumber="563">Tetrodon</emphasis>
</taxonomicName>
:
<taxonomicName id="4C21408B09534A23FE97FA4DFE205299" authorityName="Hamilton" authorityYear="1822" box="[324,465,1475,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="fluviatilis">
<emphasis id="B955E71A09534A23FE97FA4DFE205299" box="[324,465,1475,1501]" italics="true" pageId="2" pageNumber="563">T. fluviatilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21408B09534A23FE33FA4DFDBD5299" box="[480,588,1476,1501]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FE33FA4DFDBD5299" box="[480,588,1476,1501]" italics="true" pageId="2" pageNumber="563">T. patoca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21408B09534A23FD88FA4DFD275299" authorityName="Hamilton" authorityYear="1822" box="[603,726,1476,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="cutcutia">
<emphasis id="B955E71A09534A23FD88FA4DFD275299" box="[603,726,1476,1501]" italics="true" pageId="2" pageNumber="563">T. cutcutia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21408B09534A23FD36FA4DFCCE5299" authorityName="Hamilton" authorityYear="1822" box="[741,831,1476,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="caria">
<emphasis id="B955E71A09534A23FD36FA4DFCCE5299" box="[741,831,1476,1501]" italics="true" pageId="2" pageNumber="563">T. caria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21408B09534A23FC9DFA4DFC4E5299" authorityName="Hamilton" authorityYear="1822" box="[846,959,1475,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="gularis">
<emphasis id="B955E71A09534A23FC9DFA4DFC4E5299" box="[846,959,1475,1501]" italics="true" pageId="2" pageNumber="563">T. gularis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C21408B09534A23FC2DFA4DFBBD5299" authorityName="Hamilton" authorityYear="1822" box="[1022,1100,1476,1501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="tepa">
<emphasis id="B955E71A09534A23FC2DFA4DFBBD5299" box="[1022,1100,1476,1501]" italics="true" pageId="2" pageNumber="563">T. tepa</emphasis>
</taxonomicName>
. Of these only the first three are currently considered valid (
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), as
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<emphasis id="B955E71A09534A23FD28FA6EFBC35145" box="[763,1074,1511,1537]" italics="true" pageId="2" pageNumber="563">Dichotomyctere fluviatilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21408B09534A23FB97FA6EFA915145" box="[1092,1376,1511,1537]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FB97FA6EFA915145" box="[1092,1376,1511,1537]" italics="true" pageId="2" pageNumber="563">Chelonodontops patoca</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C21408B09534A23FF44F982FEAE5161" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[151,351,1547,1573]" class="Elasmobranchii" family="Tetraodontidae" genus="Leiodon" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="cutcutia">
<emphasis id="B955E71A09534A23FF44F982FEAE5161" box="[151,351,1547,1573]" italics="true" pageId="2" pageNumber="563">Leiodon cutcutia</emphasis>
</taxonomicName>
, while
<taxonomicName id="4C21408B09534A23FE6BF985FDE55161" authorityName="Hamilton" authorityYear="1822" box="[440,532,1548,1573]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="caria">
<emphasis id="B955E71A09534A23FE6BF985FDE55161" box="[440,532,1548,1573]" italics="true" pageId="2" pageNumber="563">T. caria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C21408B09534A23FD9DF985FD4E5161" authorityName="Hamilton" authorityYear="1822" box="[590,703,1547,1573]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="gularis">
<emphasis id="B955E71A09534A23FD9DF985FD4E5161" box="[590,703,1547,1573]" italics="true" pageId="2" pageNumber="563">T. gularis</emphasis>
</taxonomicName>
are considered synonyms of the latter, and
<taxonomicName id="4C21408B09534A23FB17F985FAE55161" authorityName="Hamilton" authorityYear="1822" box="[1220,1300,1548,1573]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="tepa">
<emphasis id="B955E71A09534A23FB17F985FAE55161" box="[1220,1300,1548,1573]" italics="true" pageId="2" pageNumber="563">T. tepa</emphasis>
</taxonomicName>
a synonym of
<taxonomicName id="4C21408B09534A23FF6BF9A6FE48510D" baseAuthorityName="Bloch &amp; Schneider" baseAuthorityYear="1801" box="[184,441,1583,1609]" family="Tetraodontidae" genus="Lagocephalus" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="lunaris">
<emphasis id="B955E71A09534A23FF6BF9A6FE48510D" box="[184,441,1583,1609]" italics="true" pageId="2" pageNumber="563">Lagocephalus lunaris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9E3B0809534A23FF14F9DAFD71519D" blockId="2.[151,1437,1367,2042]" pageId="2" pageNumber="563">
<bibRefCitation id="EFB046F909534A23FF14F9DAFE565129" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[199,423,1619,1645]" pageId="2" pageNumber="563" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09534A23FEC5F9DDFEA35129" box="[278,338,1619,1645]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
(2018)
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described
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<emphasis id="B955E71A09534A23FDF4F9DAFC9C5129" box="[551,877,1619,1645]" italics="true" pageId="2" pageNumber="563">Chelonodontops bengalensis</emphasis>
</taxonomicName>
based on
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, 234 and
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SL, collected in
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near
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, south of the Sundarbans. The species has since been recorded from other localities in the
<collectingMunicipality id="6BFAA17209534A23FE9FF912FDAB51F1" box="[332,602,1691,1718]" pageId="2" pageNumber="563">northern Bay of Bengal</collectingMunicipality>
, off
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in
<collectingCountry id="F3367B9809534A23FC0CF912FBE851F1" box="[991,1049,1691,1717]" name="India" pageId="2" pageNumber="563">India</collectingCountry>
(
<bibRefCitation id="EFB046F909534A23FBFBF912FAD751F1" author="Mohapatra, A. &amp; Mohanty, S. R. &amp; Mishra S. S. &amp; Tudu, P. C." box="[1064,1318,1691,1717]" pageId="2" pageNumber="563" pagination="37 - 41" refId="ref6385" refString="Mohapatra, A., Mohanty, S. R. Mishra S. S. &amp; Tudu, P. C. (2020) First report of Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae) from Indian coast. FishTaxa, 16, 37 - 41." type="journal article" year="2020">
Mohapatra
<emphasis id="B955E71A09534A23FB78F915FB1551F1" box="[1195,1252,1691,1717]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
2020
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) and from the east coast off
<collectingRegion id="49E5F5EA09534A23FE88F936FE5B519D" box="[347,426,1727,1753]" country="India" name="Odisha" pageId="2" pageNumber="563">Odisha</collectingRegion>
(
<bibRefCitation id="EFB046F909534A23FE6AF936FD85519D" author="Roul, S. K. &amp; Ghosh, S. &amp; Rohit, P." box="[441,628,1727,1753]" pageId="2" pageNumber="563" pagination="363 - 367" refId="ref6860" refString="Roul, S. K., Ghosh, S. &amp; Rohit, P. (2022) First record of Bengal reticulated puffer Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae) from Odisha coast, north-western Bay of Bengal with taxonomic notes. Indian Journal of Geo Marine Sciences, 51 (4), 363 - 367. https: // doi. org / 10.56042 / ijms. v 51 i 04.34695" type="journal article" year="2022">
Roul
<emphasis id="B955E71A09534A23FE2BF949FDC1519D" box="[504,560,1727,1753]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
2022
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).
</paragraph>
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For the comparison of their new species
<taxonomicName id="4C21408B09534A23FD43F96AFC2151B9" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[656,976,1763,1789]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09534A23FD43F96AFC2151B9" box="[656,976,1763,1789]" italics="true" pageId="2" pageNumber="563">Chelonodontops bengalensis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C21408B09534A23FBC2F96DFA9351B9" authority=", Habib et al. (2018)" authorityName="Habib" authorityYear="2018" box="[1041,1378,1763,1790]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FBC2F96DFB8C51B9" box="[1041,1149,1764,1790]" italics="true" pageId="2" pageNumber="563">C. patoca</emphasis>
,
<bibRefCitation id="EFB046F909534A23FB5FF96AFA9351B9" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[1164,1378,1763,1789]" pageId="2" pageNumber="563" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09534A23FB0BF96DFAE151B9" box="[1240,1296,1763,1789]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
(2018)
</bibRefCitation>
</taxonomicName>
used the information for the latter provided in
<bibRefCitation id="EFB046F909534A23FDBEF88EFC4D5066" author="Sujatha, K. &amp; Padmavathi, P." box="[621,956,1799,1826]" pageId="2" pageNumber="563" pagination="95 - 104" refId="ref7152" refString="Sujatha, K. &amp; Padmavathi, P. (2015) Taxonomy of puffer fish (Pisces: Tetraodontidae) represented in the catches of Visakhapatnam, central eastern coast of India. Journal of the Marine Biological Association of India, 57 (2), 95 - 104." type="journal article" year="2015">Sujatha &amp; Padmavathi (2015)</bibRefCitation>
based on specimens from Visakhapatnam,
<collectingRegion id="49E5F5EA09534A23FF44F8A2FEBB5001" box="[151,330,1835,1861]" country="India" name="Andhra Pradesh" pageId="2" pageNumber="563">Andhra Pradesh</collectingRegion>
, and in
<bibRefCitation id="EFB046F909534A23FE72F8A5FD915001" author="Han, K. - H. &amp; Baek, J. - I. &amp; Shin, L. - S. &amp; Kim, H. - J. &amp; Yoon, B. - I. &amp; Hwang, J. - H. &amp; Lee, S. - H." box="[417,608,1835,1862]" pageId="2" pageNumber="563" pagination="77 - 84" refId="ref5747" refString="Han, K. - H., Baek, J. - I., Shin, L. - S., Kim, H. - J., Yoon, B. - I., Hwang, J. - H. &amp; Lee, S. - H. (2017) Morphological description of three species of pufferfishes (Tetraodontidae) from India. Korean Journal of Fisheries and Aquatic Sciences, 50 (1), 77 - 84. https: // doi. org / 10.5657 / KFAS. 2017.0077" type="journal article" year="2017">
Han
<emphasis id="B955E71A09534A23FE05F8A5FDFF5001" box="[470,526,1835,1861]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
(2017)
</bibRefCitation>
, based on specimens from the fish market in Chennai. Both these locations are ca. 800 and
<quantity id="4CD996ED09534A23FE9AF8C6FE5A502D" box="[329,427,1871,1897]" metricMagnitude="6" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="563" unit="km" value="1400.0">1400 km</quantity>
south of Baruipur and Lakshmipur, respectively, from where
<bibRefCitation id="EFB046F909534A23FB8FF8C6FAD6502D" author="Hamilton, F." box="[1116,1319,1871,1897]" pageId="2" pageNumber="563" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
illustrated
<typeStatus id="549A85AA09534A23FF44F8FDFF3750CA" box="[151,198,1908,1934]" pageId="2" pageNumber="563">type</typeStatus>
specimen of
<taxonomicName id="4C21408B09534A23FE8DF8FAFDE350C9" authorityName="Hamilton" authorityYear="1822" box="[350,530,1907,1933]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FE8DF8FAFDE350C9" box="[350,530,1907,1933]" italics="true" pageId="2" pageNumber="563">Tetrodon patoca</emphasis>
</taxonomicName>
originated.
<bibRefCitation id="EFB046F909534A23FD4BF8FAFC9C50C9" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[664,877,1907,1933]" pageId="2" pageNumber="563" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09534A23FD37F8FDFCED50C9" box="[740,796,1907,1933]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
(2018)
</bibRefCitation>
do not seem to have consulted
<bibRefCitation id="EFB046F909534A23FB02F8FAFA6D50C9" author="Hamilton, F." box="[1233,1436,1907,1933]" pageId="2" pageNumber="563" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
original description of
<taxonomicName id="4C21408B09534A23FE41F811FE0950F5" authorityName="Hamilton" authorityYear="1822" box="[402,504,1944,1969]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FE41F811FE0950F5" box="[402,504,1944,1969]" italics="true" pageId="2" pageNumber="563">T. patoca</emphasis>
</taxonomicName>
: they do not cite his work. This is surprising given that
<bibRefCitation id="EFB046F909534A23FB8CF81EFAD850F5" author="Hamilton, F." box="[1119,1321,1943,1969]" pageId="2" pageNumber="563" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
collecting locality for the specimen of his
<taxonomicName id="4C21408B09534A23FE25F835FDAD5091" authorityName="Hamilton" authorityYear="1822" box="[502,604,1980,2005]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09534A23FE25F835FDAD5091" box="[502,604,1980,2005]" italics="true" pageId="2" pageNumber="563">T. patoca</emphasis>
</taxonomicName>
illustrated on Plate 18 is only
<quantity id="4CD996ED09534A23FC62F832FBF55091" box="[945,1028,1979,2005]" metricMagnitude="5" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="563" unit="km" value="140.0">140 km</quantity>
(Baruipur) or
<quantity id="4CD996ED09534A23FB75F832FB0B5091" box="[1190,1274,1979,2005]" metricMagnitude="5" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="563" unit="km" value="190.0">190 km</quantity>
(Lakshmipur), respectively, from the
<typeStatus id="549A85AA09534A23FE52F869FE5F50BE" box="[385,430,2016,2042]" pageId="2" pageNumber="563">type</typeStatus>
locality of Habib
<emphasis id="B955E71A09534A23FDBEF869FD5050BD" box="[621,673,2015,2041]" italics="true" pageId="2" pageNumber="563">et al.</emphasis>
s (2018)
<taxonomicName id="4C21408B09534A23FCD6F869FC5050BD" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[773,929,2015,2042]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="563" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09534A23FCD6F869FC5050BD" box="[773,929,2015,2042]" italics="true" pageId="2" pageNumber="563">C. bengalensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9E3B0809524A22FF14FF1EFCF0555D" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="564">
<bibRefCitation id="EFB046F909524A22FF14FF1EFE6957F5" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[199,408,151,177]" pageId="3" pageNumber="564" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09524A22FEC0FF1EFEBD57F5" box="[275,332,151,177]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
(2018
</bibRefCitation>
, pp. 4, 67) concluded that their
<taxonomicName id="4C21408B09524A22FCDDFF1EFBA057F5" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[782,1105,151,177]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FCDDFF1EFBA057F5" box="[782,1105,151,177]" italics="true" pageId="3" pageNumber="564">Chelonodontops bengalensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C21408B09524A22FB34FF1EFAA457F4" authorityName="Hamilton" authorityYear="1822" box="[1255,1365,151,177]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FB34FF1EFAA457F4" box="[1255,1365,151,177]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
in the following characters: “
<taxonomicName id="4C21408B09524A22FE44FF32FDCA5791" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[407,571,187,213]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FE44FF32FDCA5791" box="[407,571,187,213]" italics="true" pageId="3" pageNumber="564">C. bengalensis</emphasis>
</taxonomicName>
have both back and belly with spines (vs belly only with spines in
<taxonomicName id="4C21408B09524A22FAF5FF32FA625790" authorityName="Hamilton" authorityYear="1822" box="[1318,1427,187,213]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FAF5FF32FA625790" box="[1318,1427,187,213]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
) … in dorsal rays (12 vs 10), anal rays (10 vs 8), and pectoral rays (19 vs 1516).
<taxonomicName id="4C21408B09524A22FBF7FF56FB3857BD" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[1060,1225,223,249]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FBF7FF56FB3857BD" box="[1060,1225,223,249]" italics="true" pageId="3" pageNumber="564">C. bengalensis</emphasis>
</taxonomicName>
has spinules on its lateral sides (vs. a patch of spinules on the back from behind the interorbital nearly to the dorsal fin and another on the throat and abdomen but absent on the lateral sides in
<taxonomicName id="4C21408B09524A22FCCBFEAEFC745604" authorityName="Hamilton" authorityYear="1822" box="[792,901,295,321]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FCCBFEAEFC745604" box="[792,901,295,321]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
)…Nasal organ not covered by a small sac with two nostrils (vs in
<taxonomicName id="4C21408B09524A22FEBDFEC2FE2D5620" authorityName="Hamilton" authorityYear="1822" box="[366,476,331,357]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FEBDFEC2FE2D5620" box="[366,476,331,357]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
expanded before and behind into a pair of elongate flaps).
<taxonomicName id="4C21408B09524A22FB50FEC2FADA5621" box="[1155,1323,331,357]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bangalensis">
<emphasis id="B955E71A09524A22FB50FEC2FADA5621" box="[1155,1323,331,357]" italics="true" pageId="3" pageNumber="564">C. bangalensis</emphasis>
</taxonomicName>
(sic!) has dorsal and upper half of lateral sides of the body with spiral shaped whitish spots (vs. greenishgray to brown on the back with large round to ovate white spots in
<taxonomicName id="4C21408B09524A22FD42FE1AFD0C56E8" authorityName="Hamilton" authorityYear="1822" box="[657,765,403,429]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FD42FE1AFD0C56E8" box="[657,765,403,429]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
).
<taxonomicName id="4C21408B09524A22FCC7FE1AFC4956E9" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[788,952,403,429]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FCC7FE1AFC4956E9" box="[788,952,403,429]" italics="true" pageId="3" pageNumber="564">C. bengalensis</emphasis>
</taxonomicName>
have blackish grey color on the sides (vs. a broad yellow band running from chin to lower caudal fin base in
<taxonomicName id="4C21408B09524A22FCBAFE3EFC275694" authorityName="Hamilton" authorityYear="1822" box="[873,982,439,465]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FCBAFE3EFC275694" box="[873,982,439,465]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
)…. Genetic distance from 6.0% to 7.6% for COI and 3% to 4% for 16S gene between
<taxonomicName id="4C21408B09524A22FD42FE52FCC456B1" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[657,821,475,501]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FD42FE52FCC456B1" box="[657,821,475,501]" italics="true" pageId="3" pageNumber="564">C. bengalensis</emphasis>
</taxonomicName>
and the closest related species
<taxonomicName id="4C21408B09524A22FB43FE52FB0C56B0" authorityName="Hamilton" authorityYear="1822" box="[1168,1277,475,501]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FB43FE52FB0C56B0" box="[1168,1277,475,501]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
are consistent with usual congeneric differentiation value of&gt; 2% ...”
</paragraph>
<paragraph id="8B9E3B0809524A22FF14FDAAFB2C55B5" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="564">
<bibRefCitation id="EFB046F909524A22FF14FDAAFE645579" author="Hamilton, F." box="[199,405,547,573]" pageId="3" pageNumber="564" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
description of
<taxonomicName id="4C21408B09524A22FDEDFDAAFD055578" authorityName="Hamilton" authorityYear="1822" box="[574,756,547,573]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FDEDFDAAFD055578" box="[574,756,547,573]" italics="true" pageId="3" pageNumber="564">Tetrodon patoca</emphasis>
</taxonomicName>
mentions: “The colour of the
<taxonomicName id="4C21408B09524A22FB96FDAAFB655578" authorityName="Hamilton" authorityYear="1822" box="[1093,1172,547,572]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FB96FDAAFB655578" box="[1093,1172,547,572]" italics="true" pageId="3" pageNumber="564">Patoca</emphasis>
</taxonomicName>
above is blackish, with numerous large yellow spots on the back: below it is white. The fins are inclined to olive. The back and belly are armed with simple retractable prickles…The back fin is rounded, and contains eleven rays, of which the first two are undivided, and the first is the shortest. The pectoral fins are sharpish above. Each contains about sixteen rays. The anal fin is rounded, and contains ten rays, of which the first is the shortest, and, together with the second, is undivided. The tail fin is rounded, and contains ten rays, of which one on each side is undivided.”
</paragraph>
<paragraph id="8B9E3B0809524A22FF14FD72FC69548D" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="564">
<bibRefCitation id="EFB046F909524A22FF14FD72FE655451" author="Hamilton, F." box="[199,404,763,789]" pageId="3" pageNumber="564" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
description of fin-ray number (D11, A10, P16) and colour pattern (blackish above, numerous large yellow spots on the back, white below), as well as spination (back and belly with prickles) matches the description of these characters in Habib
<emphasis id="B955E71A09524A22FD86FCCAFD7C5419" box="[597,653,835,861]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
s (2018) new species
<taxonomicName id="4C21408B09524A22FC51FCCAFB355419" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[898,1220,835,861]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FC51FCCAFB355419" box="[898,1220,835,861]" italics="true" pageId="3" pageNumber="564">Chelonodontops bengalensis</emphasis>
</taxonomicName>
. In addition, Habib
<emphasis id="B955E71A09524A22FF44FCEEFF2054C5" box="[151,209,871,897]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
s (2018:
<figureCitation id="131A278D09524A22FEEEFCEEFE7154C5" box="[317,384,871,897]" captionStart="FIGURE 3" captionStartId="3.[152,255,1449,1473]" captionTargetBox="[183,1404,993,1426]" captionTargetId="figure-459@3.[183,1404,993,1426]" captionTargetPageId="3" captionText="FIGURE 3. Chelonodon patoca, in life, not preserved, ca. 200 mm. Note 11 dorsal-fin rays. Photo: B. Mahatvaraj." figureDoi="http://doi.org/10.5281/zenodo.13916267" httpUri="https://zenodo.org/record/13916267/files/figure.png" pageId="3" pageNumber="564">Fig. 3</figureCitation>
) illustration of the
<typeStatus id="549A85AA09524A22FD8EFCEEFD6954C5" box="[605,664,871,897]" pageId="3" pageNumber="564">types</typeStatus>
of
<taxonomicName id="4C21408B09524A22FD13FCEEFC9754C5" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[704,870,871,897]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09524A22FD13FCEEFC9754C5" box="[704,870,871,897]" italics="true" pageId="3" pageNumber="564">C. bengalensis</emphasis>
</taxonomicName>
shows that they closely match
<bibRefCitation id="EFB046F909524A22FB1EFCEEFA6D54C5" author="Hamilton, F." box="[1229,1436,871,897]" pageId="3" pageNumber="564" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
illustration of
<taxonomicName id="4C21408B09524A22FEE6FC02FE1A54E0" authorityName="Hamilton" authorityYear="1822" box="[309,491,907,933]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FEE6FC02FE1A54E0" box="[309,491,907,933]" italics="true" pageId="3" pageNumber="564">Tetrodon patoca</emphasis>
</taxonomicName>
(
<figureCitation id="131A278D09524A22FE29FC02FDCA54E1" box="[506,571,907,933]" captionStart="FIGURE 1" captionStartId="1.[152,255,770,794]" captionTargetBox="[318,1272,186,705]" captionTargetId="figure-18@1.[271,1316,179,744]" captionTargetPageId="1" captionText="FIGURE 1. Haludars illustration of Tetrodon patoca. From the archive of the British Library (Mss Eur E72, p. 47), photo: Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.13916263" httpUri="https://zenodo.org/record/13916263/files/figure.png" pageId="3" pageNumber="564">Fig. 1</figureCitation>
), reproduced here from
<bibRefCitation id="EFB046F909524A22FC9FFC02FC2854E1" author="Britz, R." box="[844,985,907,933]" pageId="3" pageNumber="564" refId="ref5222" refString="Britz, R. (2019) Francis Hamilton's Gangetic Fishes in colour. A new edition of the 1822 monograph, with reproductions of unpublished coloured illustrations. Ray Society, London, 48 + vii + 405 + vii pp., 228 pls." type="book" year="2019">Britz (2019)</bibRefCitation>
. It is clear, therefore, that the two names refer to the same species, making the former a synonym of the latter.
</paragraph>
<caption id="DF5E6B8009524A22FF4BFA20FADA5286" ID-DOI="http://doi.org/10.5281/zenodo.13916267" ID-Zenodo-Dep="13916267" box="[152,1323,1449,1474]" httpUri="https://zenodo.org/record/13916267/files/figure.png" pageId="3" pageNumber="564" startId="3.[152,255,1449,1473]" targetBox="[183,1404,993,1426]" targetPageId="3" targetType="figure">
<paragraph id="8B9E3B0809524A22FF4BFA20FADA5286" blockId="3.[152,1323,1449,1474]" box="[152,1323,1449,1474]" pageId="3" pageNumber="564">
<emphasis id="B955E71A09524A22FF4BFA20FEE95285" bold="true" box="[152,280,1449,1473]" pageId="3" pageNumber="564">FIGURE 3.</emphasis>
<taxonomicName id="4C21408B09524A22FECCFA23FE175285" box="[287,486,1450,1474]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FECCFA23FE175285" box="[287,486,1450,1474]" italics="true" pageId="3" pageNumber="564">Chelonodon patoca</emphasis>
</taxonomicName>
, in life, not preserved, ca. 200 mm. Note 11 dorsal-fin rays. Photo: B. Mahatvaraj.
</paragraph>
</caption>
<paragraph id="8B9E3B0809524A22FF14F985FA72502E" blockId="3.[151,1437,1548,2042]" pageId="3" pageNumber="564">
The species that
<bibRefCitation id="EFB046F909524A22FE53F985FDA05162" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[384,593,1548,1574]" pageId="3" pageNumber="564" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09524A22FE19F984FDF25162" box="[458,515,1548,1574]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
(2018)
</bibRefCitation>
mistook for
<taxonomicName id="4C21408B09524A22FD32F985FC475162" box="[737,950,1548,1574]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FD32F985FC475162" box="[737,950,1548,1574]" italics="true" pageId="3" pageNumber="564">Chelonodon patoca</emphasis>
</taxonomicName>
is a widespread Indo-Pacific pufferfish, first mentioned by
<bibRefCitation id="EFB046F909524A22FEE9F9B9FE2C510E" author="Russell, P." box="[314,477,1584,1610]" pageId="3" pageNumber="564" refId="ref6973" refString="Russell, P. (1803) Descriptions and figures of two hundred fishes: collected at Vizagapatam on the coast of Coromandel. Vols. 1 - 2. Printed by W. Bulmer and Co., Shakspeare Press, Cleveland-Row, for G. and W. Nicol, booksellers to his majesty, Pall-Mall, London, vii + 78 pp. &amp; 85 pp., 197 pls. https: // doi. org / 10.5962 / bhl. title. 140747" type="book" year="1803">Russell (1803)</bibRefCitation>
from Vizagapatam (todays Visakhapatnam in Andhra Pradesh) under the local name Kappa and illustrated in Plate 18, Fig. 25. In his description,
<bibRefCitation id="EFB046F909524A22FCECF9DDFC2A512A" author="Russell, P." box="[831,987,1620,1646]" pageId="3" pageNumber="564" refId="ref6973" refString="Russell, P. (1803) Descriptions and figures of two hundred fishes: collected at Vizagapatam on the coast of Coromandel. Vols. 1 - 2. Printed by W. Bulmer and Co., Shakspeare Press, Cleveland-Row, for G. and W. Nicol, booksellers to his majesty, Pall-Mall, London, vii + 78 pp. &amp; 85 pp., 197 pls. https: // doi. org / 10.5962 / bhl. title. 140747" type="book" year="1803">Russell (1803</bibRefCitation>
: p. 18) mentioned also the characteristic colouration of this species: “The colour of the upper part a dark purplish, beautifully variegated with dusky whitish, or greenish spots, and two or three dark transverse bands; the throat and belly white, with a streak of yellow on each side. The dorsal fin dark, the caudal almost black; the anal of a light colour.” The streak of yellow, in particular, seems to be unique to this species. The literature on the fish fauna of the Western Indo-Pacific suggests that the species of
<taxonomicName id="4C21408B09524A22FEC1F881FE6A5066" authorityName="Muller" authorityYear="1841" box="[274,411,1800,1826]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09524A22FEC1F881FE6A5066" box="[274,411,1800,1826]" italics="true" pageId="3" pageNumber="564">Chelonodon</emphasis>
</taxonomicName>
with a yellow lateral streak is widely distributed from the coast of
<collectingCountry id="F3367B9809524A22FB76F881FAF55066" box="[1189,1284,1800,1826]" name="Pakistan" pageId="3" pageNumber="564">Pakistan</collectingCountry>
(
<bibRefCitation id="EFB046F909524A22FAC7F881FEE45003" author="Psomadakis, P. N. &amp; Osmany, H. B. &amp; Moazzam M." pageId="3" pageNumber="564" refId="ref6608" refString="Psomadakis, P. N., Osmany, H. B. &amp; Moazzam M. (2015) Field identification guide to the living marine resources of Pakistan. FAO species identification guide for fishery purposes. Food and Agriculture Organization of the United Nations, Rome, x + 386 pp., 42 pls .." type="book" year="2015">
Psomadakis
<emphasis id="B955E71A09524A22FF44F8A4FF205002" box="[151,209,1836,1862]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2015
</bibRefCitation>
),
<collectingCountry id="F3367B9809524A22FEF9F8A5FE955002" box="[298,356,1836,1862]" name="India" pageId="3" pageNumber="564">India</collectingCountry>
(
<bibRefCitation id="EFB046F909524A22FEA0F8A5FDD85002" author="Han, K. - H. &amp; Baek, J. - I. &amp; Shin, L. - S. &amp; Kim, H. - J. &amp; Yoon, B. - I. &amp; Hwang, J. - H. &amp; Lee, S. - H." box="[371,553,1836,1862]" pageId="3" pageNumber="564" pagination="77 - 84" refId="ref5747" refString="Han, K. - H., Baek, J. - I., Shin, L. - S., Kim, H. - J., Yoon, B. - I., Hwang, J. - H. &amp; Lee, S. - H. (2017) Morphological description of three species of pufferfishes (Tetraodontidae) from India. Korean Journal of Fisheries and Aquatic Sciences, 50 (1), 77 - 84. https: // doi. org / 10.5657 / KFAS. 2017.0077" type="journal article" year="2017">
Han
<emphasis id="B955E71A09524A22FE7FF8A4FE175002" box="[428,486,1836,1862]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EFB046F909524A22FDE5F8A5FCA15002" author="Kaleshkumar, K. &amp; Rajaram, R. &amp; Purushothaman, P. &amp; Arun, G." box="[566,848,1836,1862]" pageId="3" pageNumber="564" pagination="12726 - 12737" refId="ref5919" refString="Kaleshkumar, K., Rajaram, R. Purushothaman, P. &amp; Arun, G. (2018) Morphological variations in marine pufferfish and porcupinefish (Teleostei: Tetraodontiformes) from Tamil Nadu, southeastern coast of India. Journal of Threatened Taxa, 10 (13), 12726 - 12737. https: // doi. org / 10.11609 / jott. 4028.10.13.12726 - 12737" type="journal article" year="2018">
Kaleshkumar
<emphasis id="B955E71A09524A22FD00F8A4FCFC5002" box="[723,781,1836,1862]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2018
</bibRefCitation>
;
<bibRefCitation id="EFB046F909524A22FC8EF8A5FB6E5003" author="Sujatha, K. &amp; Padmavathi, P." box="[861,1183,1836,1863]" pageId="3" pageNumber="564" pagination="95 - 104" refId="ref7152" refString="Sujatha, K. &amp; Padmavathi, P. (2015) Taxonomy of puffer fish (Pisces: Tetraodontidae) represented in the catches of Visakhapatnam, central eastern coast of India. Journal of the Marine Biological Association of India, 57 (2), 95 - 104." type="journal article" year="2015">Sujatha &amp; Padmavathi 2015</bibRefCitation>
;
<bibRefCitation id="EFB046F909524A22FB7FF8A5FA615002" author="Veeruraj, A. &amp; Arumugam, M. &amp; Ajithkumar, T. &amp; Balasubramanian, T." box="[1196,1424,1836,1862]" pageId="3" pageNumber="564" pagination="1 - 12" refId="ref7343" refString="Veeruraj, A., Arumugam, M., Ajithkumar, T. &amp; Balasubramanian, T. (2011) Distribution of Tetraodontiformes (Family: Tetraodontidae) along the Parangipettai Coast, Southeast coast of India. Zootaxa, 3015 (1), 1 - 12. https: // doi. org / 10.11646 / zootaxa. 3015.1.1" type="journal article" year="2011">
Veeruraj
<emphasis id="B955E71A09524A22FAC0F8A4FABC5002" box="[1299,1357,1836,1862]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2011
</bibRefCitation>
),
<collectingCountry id="F3367B9809524A22FF44F8D9FEF5502E" box="[151,260,1872,1898]" name="Myanmar" pageId="3" pageNumber="564">Myanmar</collectingCountry>
(
<bibRefCitation id="EFB046F909524A22FEC1F8D9FDED502E" author="Psomadakis, P. &amp; Thein, H. &amp; Russell, B. C. &amp; Tun, M. T." box="[274,540,1872,1898]" pageId="3" pageNumber="564" refId="ref6667" refString="Psomadakis, P., Thein, H., Russell, B. C. &amp; Tun, M. T. (2020) Field identification guide to the living marine resources of Myanmar. FAO species identification guide for fishery purposes. Food and Agriculture Organization of the United Nations, and Department of Fisheries, Ministry of Agriculture, Livestock and Irrigation, Republic of the Union of Myanmar, Rome, xvii + 694 pp., 58 pls." type="book" year="2020">
Psomadakis
<emphasis id="B955E71A09524A22FE72F8D8FE28502E" box="[417,473,1872,1898]" italics="true" pageId="3" pageNumber="564">et al.</emphasis>
2020
</bibRefCitation>
), eastwards to
<collectingCountry id="F3367B9809524A22FD11F8D9FCDE502E" box="[706,815,1872,1898]" name="Indonesia" pageId="3" pageNumber="564">Indonesia</collectingCountry>
and
<collectingCountry id="F3367B9809524A22FCB6F8D9FC54502E" box="[869,933,1872,1898]" name="Japan" pageId="3" pageNumber="564">Japan</collectingCountry>
(
<bibRefCitation id="EFB046F909524A22FC60F8D9FB93502E" author="Matsuura, K." box="[947,1122,1872,1898]" pageId="3" pageNumber="564" pagination="173 - 178" refId="ref6328" refString="Matsuura, K. (2002) A review of two morphologically similar puffers, Chelonodon laticeps and C. patoca. Proceedings of the 3 rd and 4 th Symposia on Collection Building and Natural History Studies in Asia and the Pacific Rim. National Science Museum Tokyo Monographs, 22, 173 - 178." type="journal article" year="2002">Matsuura 2002</bibRefCitation>
;
<bibRefCitation id="EFB046F909524A22FBBEF8D9FA89502E" author="Okamura, K. &amp; Endo, H." box="[1133,1400,1872,1898]" pageId="3" pageNumber="564" pagination="15 - 18" refId="ref6473" refString="Okamura, K. &amp; Endo, H. (2022) First record of Chelonodontops patoca from Shikoku Island, Japan. I chthy, Natural History of Fishes of Japan, 34, 15 - 18." type="journal article" year="2022">Okamura &amp; Endo 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B688309524A22FF14F8FDFC2150CA" box="[199,976,1907,1934]" pageId="3" pageNumber="564" type="nomenclature">
<paragraph id="8B9E3B0809524A22FF14F8FDFC2150CA" blockId="3.[151,1437,1548,2042]" box="[199,976,1907,1934]" pageId="3" pageNumber="564">
<emphasis id="B955E71A09524A22FF14F8FDFC2150CA" bold="true" box="[199,976,1907,1934]" pageId="3" pageNumber="564">
The origin of confusion regarding the name
<taxonomicName id="4C21408B09524A22FD35F8FAFC2150CA" ID-CoL="TVKJ" authority="(Hamilton, 1822)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[742,976,1907,1934]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FD35F8FAFC3850CA" bold="true" box="[742,969,1907,1934]" italics="true" pageId="3" pageNumber="564">Chelonodon patoca</emphasis>
.
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33B688309524A24FC0AF8FDFB8C50BD" lastPageId="5" lastPageNumber="566" pageId="3" pageNumber="564" type="discussion">
<paragraph id="8B9E3B0809524A25FC0AF8FDFB2056CD" blockId="3.[151,1437,1548,2042]" lastBlockId="4.[151,1437,151,1905]" lastPageId="4" lastPageNumber="565" pageId="3" pageNumber="564">
<bibRefCitation id="EFB046F909524A22FC0AF8FDFB7550CB" author="Bleeker, P." box="[985,1156,1908,1935]" pageId="3" pageNumber="564" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker (1850)</bibRefCitation>
may have been the first to assume a wide distribution for
<taxonomicName id="4C21408B09524A22FDF4F811FD6950F6" box="[551,664,1944,1970]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="3" pageNumber="564" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09524A22FDF4F811FD6950F6" box="[551,664,1944,1970]" italics="true" pageId="3" pageNumber="564">C. patoca</emphasis>
</taxonomicName>
, mentioning this species from the Malay Archipelago (“indischen Archipel”) and also “
<collectingCountry id="F3367B9809524A22FE56F835FE055092" box="[389,500,1980,2006]" name="Bangladesh" pageId="3" pageNumber="564">Bengalen</collectingCountry>
,
<collectingCountry id="F3367B9809524A22FE2CF835FDB45092" box="[511,581,1980,2006]" name="China" pageId="3" pageNumber="564">China</collectingCountry>
,
<collectingCountry id="F3367B9809524A22FD83F835FD645092" box="[592,661,1980,2006]" name="Japan" pageId="3" pageNumber="564">Japan</collectingCountry>
”. This was reiterated in
<bibRefCitation id="EFB046F909524A22FC76F835FB975092" author="Gunther, A." box="[933,1126,1980,2006]" pageId="3" pageNumber="564" refId="ref5537" refString="Gunther, A. (1870) Catalogue of the fishes in the British Museum. Catalogue of the Physostomi, containing the families Gymnotidae, Symbranchidae, Muraenidae, Pegasidae, and of the Lophobranchii, Plectognathi, Dipnoi, ... [thru] ... Leptocardii, in the British Museum. Vol. 8. Department of Zoology, British Museum (Natural History), London, xxv + 549 pp." type="book" year="1870">Günthers (1870)</bibRefCitation>
entry for this species in his influential Catalog, based on material from Bengal,
<collectingCountry id="F3367B9809524A22FD32F869FCA450BE" box="[737,853,2016,2042]" name="Singapore" pageId="3" pageNumber="564">Singapore</collectingCountry>
, Penang, Amboina, the East-Indian archipelago and
<collectingCountry id="F3367B9809554A25FF44FF1EFF2F57F5" box="[151,222,151,177]" name="China" pageId="4" pageNumber="565">China</collectingCountry>
. This identity and distribution subsequentlycame to be widely adopted, as evidenced in many treatises on the fish fauna of South and Southeast Asia (see e.g.,
<bibRefCitation id="EFB046F909554A25FD39FF32FBD05791" author="de Beaufort, L. F. &amp; Briggs, J. C." box="[746,1057,187,213]" pageId="4" pageNumber="565" pagination="1 - 481" refId="ref5269" refString="de Beaufort, L. F. &amp; Briggs, J. C. (1962) Scleroparei, Hypostomides, Pediculati, Plectognathi, Opisthomi, Discocephali, Xenopterygii. In: Weber, M. &amp; Beaufort, L. F. de, The fishes of the Indo-Australian Archipelago. Vol. 11. E. J. Brill, Leiden, pp. i - xi + 1 - 481. https: // doi. org / 10.1163 / 9789004590953" type="book chapter" year="1962">de Beaufort &amp; Briggs 1962</bibRefCitation>
; Matsuura in Masuda
<emphasis id="B955E71A09554A25FACFFF32FAA45791" box="[1308,1365,187,213]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
1984;
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Kottelat
<emphasis id="B955E71A09554A25FF29FF56FEC257BD" box="[250,307,223,249]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
1993
</bibRefCitation>
; Talwar &amp; Jhinghran 1992;
<bibRefCitation id="EFB046F909554A25FD66FF56FC7957BD" author="Vidthayanon, C." box="[693,904,223,249]" pageId="4" pageNumber="565" refId="ref7456" refString="Vidthayanon, C. (2008) Field guide to fishes of the Mekong Delta. Mekong River Commission, Vientiane, 288 pp." type="book" year="2008">Vidthayanon 2008</bibRefCitation>
;
<bibRefCitation id="EFB046F909554A25FC46FF56FB7057BD" author="Rainboth, W. J. &amp; Vidthayanon, C. &amp; Yen, M. D." box="[917,1153,223,249]" pageId="4" pageNumber="565" pagination="1 - 121" refId="ref6799" refString="Rainboth, W. J., Vidthayanon, C. &amp; Yen, M. D. (2012) Fishes of the greater Mekong ecosystem with species list and photographic atlas. Miscellaneous Publications, Museum of Zoology, University of Michigan, 201, i - xvi + 1 - 173, pls. 1 - 121." type="journal article" year="2012">
Rainboth
<emphasis id="B955E71A09554A25FBD7FF56FBCC57BD" box="[1028,1085,223,249]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
2012
</bibRefCitation>
;
<bibRefCitation id="EFB046F909554A25FB5EFF56FA6957BD" author="Psomadakis, P. N. &amp; Osmany, H. B. &amp; Moazzam M." box="[1165,1432,222,249]" pageId="4" pageNumber="565" refId="ref6608" refString="Psomadakis, P. N., Osmany, H. B. &amp; Moazzam M. (2015) Field identification guide to the living marine resources of Pakistan. FAO species identification guide for fishery purposes. Food and Agriculture Organization of the United Nations, Rome, x + 386 pp., 42 pls .." type="book" year="2015">
Psomadakis
<emphasis id="B955E71A09554A25FACFFF56FAA457BD" box="[1308,1365,223,249]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
2015
</bibRefCitation>
,
<bibRefCitation id="EFB046F909554A25FF44FE8AFF225659" author="Psomadakis, P. &amp; Thein, H. &amp; Russell, B. C. &amp; Tun, M. T." box="[151,211,259,285]" pageId="4" pageNumber="565" refId="ref6667" refString="Psomadakis, P., Thein, H., Russell, B. C. &amp; Tun, M. T. (2020) Field identification guide to the living marine resources of Myanmar. FAO species identification guide for fishery purposes. Food and Agriculture Organization of the United Nations, and Department of Fisheries, Ministry of Agriculture, Livestock and Irrigation, Republic of the Union of Myanmar, Rome, xvii + 694 pp., 58 pls." type="book" year="2020">2020</bibRefCitation>
;
<bibRefCitation id="EFB046F909554A25FF0EFE8AFD115659" author="Nagao Natural Environment Foundation" box="[221,736,259,285]" pageId="4" pageNumber="565" refId="ref6512" refString="Nagao Natural Environment Foundation. (2021). Fishes of the Indochinese Mekong. Nagao Natural Environment Foundation, Tokyo, xii + 546 pp. [https: // www. nagaofoundation. or. jp / e / publication / ebook / mekongfishes / book / index. html # target / page _ no = 1]" type="book" year="2021">Nagao Natural Environment Foundation 2021</bibRefCitation>
). As we have shown above, however, most of these distribution records concern the species with the yellow lateral streak, not
<bibRefCitation id="EFB046F909554A25FCB6FEAEFBC75605" author="Hamilton, F." box="[869,1078,295,321]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FBECFEAEFAEC5604" box="[1087,1309,295,321]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FBECFEAEFAEC5604" box="[1087,1309,295,321]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
. So far,
<taxonomicName id="4C21408B09554A25FA50FEAEFF125620" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FA50FEAEFF125620" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
seems to be known with certainty only from brackish and marine waters in the northern part of the Bay of Bengal from
<collectingCountry id="F3367B9809554A25FEF8FEE6FE9456CD" box="[299,357,367,393]" name="India" pageId="4" pageNumber="565">India</collectingCountry>
and
<collectingCountry id="F3367B9809554A25FE48FEE6FDEF56CD" box="[411,542,367,393]" name="Bangladesh" pageId="4" pageNumber="565">Bangladesh</collectingCountry>
(
<bibRefCitation id="EFB046F909554A25FDFFFEE6FD0256CD" author="Habib, K. A. &amp; Neogi, A. K. &amp; Oh, J. &amp; Lee, Y. - H. &amp; Kim, C. - G." box="[556,755,367,393]" pageId="4" pageNumber="565" pagination="1 - 8" refId="ref5612" refString="Habib, K. A., Neogi, A. K., Oh, J., Lee, Y. - H. &amp; Kim, C. - G. (2018) Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae): a new species of puffer fish from the Northern Bay of Bengal based on morphology and DNA barcode. Ocean Science Journal, 54, 1 - 8. https: // doi. org / 10.1007 / s 12601 - 018 - 0054 - 7" type="journal article" year="2018">
Habib
<emphasis id="B955E71A09554A25FDAAFEE6FD4356CD" box="[633,690,367,393]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
2018
</bibRefCitation>
;
<bibRefCitation id="EFB046F909554A25FD2CFEE6FBF056CD" author="Mohapatra, A. &amp; Mohanty, S. R. &amp; Mishra S. S. &amp; Tudu, P. C." box="[767,1025,367,393]" pageId="4" pageNumber="565" pagination="37 - 41" refId="ref6385" refString="Mohapatra, A., Mohanty, S. R. Mishra S. S. &amp; Tudu, P. C. (2020) First report of Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae) from Indian coast. FishTaxa, 16, 37 - 41." type="journal article" year="2020">
Mohapatra
<emphasis id="B955E71A09554A25FCACFEE6FC4956CD" box="[895,952,367,393]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
, 2020
</bibRefCitation>
;
<bibRefCitation id="EFB046F909554A25FBDFFEE6FB3756CD" author="Roul, S. K. &amp; Ghosh, S. &amp; Rohit, P." box="[1036,1222,367,393]" pageId="4" pageNumber="565" pagination="363 - 367" refId="ref6860" refString="Roul, S. K., Ghosh, S. &amp; Rohit, P. (2022) First record of Bengal reticulated puffer Chelonodontops bengalensis (Tetraodontiformes: Tetraodontidae) from Odisha coast, north-western Bay of Bengal with taxonomic notes. Indian Journal of Geo Marine Sciences, 51 (4), 363 - 367. https: // doi. org / 10.56042 / ijms. v 51 i 04.34695" type="journal article" year="2022">
Roul
<emphasis id="B955E71A09554A25FB99FEE6FB7356CD" box="[1098,1154,367,393]" italics="true" pageId="4" pageNumber="565">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9E3B0809554A25FF14FE1AFAED5579" blockId="4.[151,1437,151,1905]" pageId="4" pageNumber="565">
<emphasis id="B955E71A09554A25FF14FE1AFA6D56E9" bold="true" box="[199,1436,402,429]" pageId="4" pageNumber="565">
The correct name of the species of
<taxonomicName id="4C21408B09554A25FD88FE1BFD1856E9" authorityName="Muller" authorityYear="1841" box="[603,745,402,429]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FD88FE1BFD1856E9" bold="true" box="[603,745,402,429]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
with the yellow streak, previously confused with
<taxonomicName id="4C21408B09554A25FAF9FE1AFA6456E9" box="[1322,1429,403,429]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FAF9FE1AFA6456E9" bold="true" box="[1322,1429,403,429]" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
.
</emphasis>
Several names have been treated as synonyms of
<taxonomicName id="4C21408B09554A25FD13FE3EFC6B5694" box="[704,922,439,465]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FD13FE3EFC6B5694" box="[704,922,439,465]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
:
<taxonomicName id="4C21408B09554A25FC7AFE3EFAEE5695" authority="Cantor 1849" authorityName="Cantor" authorityYear="1849" box="[937,1311,439,465]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="dissutidens">
<emphasis id="B955E71A09554A25FC7AFE3EFB7D5695" box="[937,1164,439,465]" italics="true" pageId="4" pageNumber="565">Tetrodon dissutidens</emphasis>
<bibRefCitation id="EFB046F909554A25FB47FE3EFAEE5695" author="Cantor, T. E." box="[1172,1311,439,465]" pageId="4" pageNumber="565" pagination="1 - 14" refId="ref5354" refString="Cantor, T. E. (1849) Catalogue of Malayan fishes. Journal of the Asiatic Society of Bengal, 18 (2), i - xii + 983 - 1443, pls. 1 - 14." type="journal article" year="1849">Cantor 1849</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C21408B09554A25FAFFFE3EFE7356B1" authority="Bleeker 1850" authorityName="Bleeker" authorityYear="1850" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FAFFFE3EFF2A56B1" italics="true" pageId="4" pageNumber="565">Tetraodon kappa</emphasis>
<bibRefCitation id="EFB046F909554A25FF35FE52FE7356B1" author="Bleeker, P." box="[230,386,474,501]" pageId="4" pageNumber="565" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker 1850</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C21408B09554A25FE43FE52FCC156B1" authority="Bianconi 1854" authorityName="Bianconi" authorityYear="1854" box="[400,816,474,501]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="hartlaubii">
<emphasis id="B955E71A09554A25FE43FE52FD8D56B1" box="[400,636,475,501]" italics="true" pageId="4" pageNumber="565">Tetraodon hartlaubii</emphasis>
<bibRefCitation id="EFB046F909554A25FD55FE52FCC156B1" author="Bianconi, G. G." box="[646,816,474,501]" pageId="4" pageNumber="565" pagination="68 - 69" refId="ref5025" refString="Bianconi, G. G. (1854) [Specimina zoologica mosambicana, Fasciculus VIII: Pisces novi Mozambicani.]. Rendiconto delle Sessioni dell'Accademia delle Scienze dell'Instituto dei Bologna, 1853 - 1854, 68 - 69." type="book chapter" year="1854">Bianconi 1854</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C21408B09554A25FCA2FE52FAE856B1" authority="Swainson 1839" authorityName="Swainson" authorityYear="1839" box="[881,1305,475,501]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B955E71A09554A25FCA2FE52FB9156B1" box="[881,1120,475,501]" italics="true" pageId="4" pageNumber="565">Tetraodon maculatus</emphasis>
<bibRefCitation id="EFB046F909554A25FBB9FE52FAE856B1" author="Swainson, W." box="[1130,1305,475,501]" pageId="4" pageNumber="565" refId="ref7204" refString="Swainson, W. (1839) On the natural history and classification of fishes, amphibians, &amp; reptiles, or monocardian animals. Vol. 2. Spottiswoode &amp; Co., London, vi + 452 pp. https: // doi. org / 10.5962 / bhl. title. 62140" type="book" year="1839">Swainson 1839</bibRefCitation>
</taxonomicName>
(the last is an unneeded replacement name for
<bibRefCitation id="EFB046F909554A25FDFDFE76FD0F555D" author="Hamilton, F." box="[558,766,511,537]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons [1822]</bibRefCitation>
<taxonomicName id="4C21408B09554A25FCD4FE76FC4F555C" authorityName="Hamilton" authorityYear="1822" box="[775,958,511,537]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FCD4FE76FC4F555C" box="[775,958,511,537]" italics="true" pageId="4" pageNumber="565">Tetrodon patoca</emphasis>
</taxonomicName>
). Of these,
<bibRefCitation id="EFB046F909554A25FB94FE76FB0B555D" author="Cantor, T. E." box="[1095,1274,511,537]" pageId="4" pageNumber="565" pagination="1 - 14" refId="ref5354" refString="Cantor, T. E. (1849) Catalogue of Malayan fishes. Journal of the Asiatic Society of Bengal, 18 (2), i - xii + 983 - 1443, pls. 1 - 14." type="journal article" year="1849">Cantors (1849)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FAD0FE76FA6D555D" authorityName="Cantor" authorityYear="1849" box="[1283,1436,511,537]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="dissutidens">
<emphasis id="B955E71A09554A25FAD0FE76FA6D555D" box="[1283,1436,511,537]" italics="true" pageId="4" pageNumber="565">T. dissutidens</emphasis>
</taxonomicName>
would appear to be the oldest available name for the wide-spread species with the yellow lateral streak.
</paragraph>
<paragraph id="8B9E3B0809554A25FF14FDCEFD9B548D" blockId="4.[151,1437,151,1905]" pageId="4" pageNumber="565">
In his description of
<taxonomicName id="4C21408B09554A25FE63FDCEFD975524" authorityName="Hamilton" authorityYear="1822" box="[432,614,583,609]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FE63FDCEFD975524" box="[432,614,583,609]" italics="true" pageId="4" pageNumber="565">Tetrodon patoca</emphasis>
</taxonomicName>
, however,
<bibRefCitation id="EFB046F909554A25FD31FDCEFC4B5525" author="Hamilton, F." box="[738,954,583,609]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamilton (1822: 8)</bibRefCitation>
stated: “The
<taxonomicName id="4C21408B09554A25FB82FDCEFB0E5525" box="[1105,1279,583,609]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FB82FDCEFB0E5525" box="[1105,1279,583,609]" italics="true" pageId="4" pageNumber="565">Tetrodon kappa</emphasis>
</taxonomicName>
of Dr Russell (Indian Fishes, Vol. I No. 25) has also a considerable affinity to the Patoca, but is less prominent on the belly, and more so between the eyes, while the spots on the back are pale blue its belly yellow, and its back only rough, and devoid of visible prickles.” Hamilton listed
<taxonomicName id="4C21408B09554A25FDACFD3AFCDC5589" box="[639,813,691,717]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FDACFD3AFCDC5589" box="[639,813,691,717]" italics="true" pageId="4" pageNumber="565">Tetrodon kappa</emphasis>
</taxonomicName>
in the index together with the other
<taxonomicName id="4C21408B09554A25FB12FD3AFA8A5589" box="[1217,1403,691,717]" pageId="4" pageNumber="565">
<emphasis id="B955E71A09554A25FB12FD3AFAD55589" box="[1217,1316,691,717]" italics="true" pageId="4" pageNumber="565">Tetrodon</emphasis>
species
</taxonomicName>
he described.
<bibRefCitation id="EFB046F909554A25FEC3FD5EFE2C55B5" author="Hamilton, F." box="[272,477,727,753]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
note and listing in the index provide the conditions for
<taxonomicName id="4C21408B09554A25FB91FD5EFB0155B5" box="[1090,1264,727,753]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FB91FD5EFB0155B5" box="[1090,1264,727,753]" italics="true" pageId="4" pageNumber="565">Tetrodon kappa</emphasis>
</taxonomicName>
to be treated as available for the species of
<taxonomicName id="4C21408B09554A25FE1DFD72FDA65451" authorityName="Muller" authorityYear="1841" box="[462,599,763,789]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FE1DFD72FDA65451" box="[462,599,763,789]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
with the yellow lateral streak. The author of the name is
<bibRefCitation id="EFB046F909554A25FB33FD72FA6D5451" author="Hamilton, F." box="[1248,1436,763,789]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamilton (1822)</bibRefCitation>
by indication to
<bibRefCitation id="EFB046F909554A25FE85FC96FDF1547D" author="Russell, P." box="[342,512,799,825]" pageId="4" pageNumber="565" refId="ref6973" refString="Russell, P. (1803) Descriptions and figures of two hundred fishes: collected at Vizagapatam on the coast of Coromandel. Vols. 1 - 2. Printed by W. Bulmer and Co., Shakspeare Press, Cleveland-Row, for G. and W. Nicol, booksellers to his majesty, Pall-Mall, London, vii + 78 pp. &amp; 85 pp., 197 pls. https: // doi. org / 10.5962 / bhl. title. 140747" type="book" year="1803">Russell (1803)</bibRefCitation>
.
<bibRefCitation id="EFB046F909554A25FDDEFC96FD45547D" author="Russell, P." box="[525,692,799,825]" pageId="4" pageNumber="565" refId="ref6973" refString="Russell, P. (1803) Descriptions and figures of two hundred fishes: collected at Vizagapatam on the coast of Coromandel. Vols. 1 - 2. Printed by W. Bulmer and Co., Shakspeare Press, Cleveland-Row, for G. and W. Nicol, booksellers to his majesty, Pall-Mall, London, vii + 78 pp. &amp; 85 pp., 197 pls. https: // doi. org / 10.5962 / bhl. title. 140747" type="book" year="1803">Russell (1803)</bibRefCitation>
referenced only a single specimen, the putative
<typeStatus id="549A85AA09554A25FB36FC96FABB547D" box="[1253,1354,799,825]" pageId="4" pageNumber="565" type="holotype">holotype</typeStatus>
, which he illustrated on Plate 18, fig. 25 and mentioned as measuring
<quantity id="4CD996ED09554A25FCB2FCCAFC315419" box="[865,960,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="565" unit="in" value="3.0">3 inches</quantity>
and 5 lines. Accepting
<bibRefCitation id="EFB046F909554A25FB1EFCCAFA6D5419" author="Hamilton, F." box="[1229,1436,835,861]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FF44FCEEFEA454C5" authorityName="Bleeker" authorityYear="1850" box="[151,341,871,897]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FF44FCEEFEA454C5" box="[151,341,871,897]" italics="true" pageId="4" pageNumber="565">Tetraodon kappa</emphasis>
</taxonomicName>
as an available name renders
<bibRefCitation id="EFB046F909554A25FD7EFCEEFC9B54C5" author="Bleeker, P." box="[685,874,870,897]" pageId="4" pageNumber="565" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleekers (1850)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FCA0FCEEFBC054C5" authorityName="Bleeker" authorityYear="1850" box="[883,1073,871,897]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FCA0FCEEFBC054C5" box="[883,1073,871,897]" italics="true" pageId="4" pageNumber="565">Tetraodon kappa</emphasis>
</taxonomicName>
a junior primary homonym and thus unavailable, and renders
<taxonomicName id="4C21408B09554A25FE23FC02FC8254E1" authority="Cantor 1849" authorityName="Cantor" authorityYear="1849" box="[496,883,907,933]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="dissutidens">
<emphasis id="B955E71A09554A25FE23FC02FD2954E1" box="[496,728,907,933]" italics="true" pageId="4" pageNumber="565">Tetrodon dissutidens</emphasis>
<bibRefCitation id="EFB046F909554A25FD30FC02FC8254E1" author="Cantor, T. E." box="[739,883,907,933]" pageId="4" pageNumber="565" pagination="1 - 14" refId="ref5354" refString="Cantor, T. E. (1849) Catalogue of Malayan fishes. Journal of the Asiatic Society of Bengal, 18 (2), i - xii + 983 - 1443, pls. 1 - 14." type="journal article" year="1849">Cantor 1849</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C21408B09554A25FC60FC02FAA154E1" authority="Bianconi 1854" authorityName="Bianconi" authorityYear="1854" box="[947,1360,906,933]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="hartlaubii">
<emphasis id="B955E71A09554A25FC60FC02FB6E54E1" box="[947,1183,907,933]" italics="true" pageId="4" pageNumber="565">Tetraodon hartlaubii</emphasis>
<bibRefCitation id="EFB046F909554A25FB7AFC02FAA154E1" author="Bianconi, G. G." box="[1193,1360,906,933]" pageId="4" pageNumber="565" pagination="68 - 69" refId="ref5025" refString="Bianconi, G. G. (1854) [Specimina zoologica mosambicana, Fasciculus VIII: Pisces novi Mozambicani.]. Rendiconto delle Sessioni dell'Accademia delle Scienze dell'Instituto dei Bologna, 1853 - 1854, 68 - 69." type="book chapter" year="1854">Bianconi 1854</bibRefCitation>
</taxonomicName>
junior synonyms of
<bibRefCitation id="EFB046F909554A25FEFDFC26FE0D548D" author="Hamilton, F." box="[302,508,943,969]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FDD0FC26FD92548D" authorityName="Bleeker" authorityYear="1850" box="[515,611,943,969]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FDD0FC26FD92548D" box="[515,611,943,969]" italics="true" pageId="4" pageNumber="565">T. kappa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9E3B0809554A25FF14FC5AFB2C52D9" blockId="4.[151,1437,151,1905]" pageId="4" pageNumber="565">
The easiest way by which to distinguish
<taxonomicName id="4C21408B09554A25FD40FC5AFC8154A8" box="[659,880,979,1005]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FD40FC5AFC8154A8" box="[659,880,979,1005]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
from
<taxonomicName id="4C21408B09554A25FC65FC5AFBEA54A9" authorityName="Bleeker" authorityYear="1850" box="[950,1051,979,1005]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FC65FC5AFBEA54A9" box="[950,1051,979,1005]" italics="true" pageId="4" pageNumber="565">C. kappa</emphasis>
</taxonomicName>
is the colour pattern (cf. Figs. 2,3 with Figs. 4,5) and the number of fin rays
<taxonomicName id="4C21408B09554A25FDAAFC7EFCA45354" box="[633,853,1015,1041]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FDAAFC7EFCA45354" box="[633,853,1015,1041]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
lacks the yellow lateral streak along the side of the body and the dark saddle-like blotches, both typical of
<taxonomicName id="4C21408B09554A25FCC0FB92FC8B5371" authorityName="Bleeker" authorityYear="1850" box="[787,890,1051,1077]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FCC0FB92FC8B5371" box="[787,890,1051,1077]" italics="true" pageId="4" pageNumber="565">C. kappa</emphasis>
</taxonomicName>
. Instead it shows an overall dark dorsal half of the body covered in small white to cream spots and elongate blotches (
<figureCitation id="131A278D09554A25FC10FBB6FBC0531D" box="[963,1073,1087,1113]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[152,255,770,794]" captionStartId-1="1.[152,255,1937,1961]" captionStartId-2="3.[152,255,1449,1473]" captionTargetBox-0="[318,1272,186,705]" captionTargetBox-1="[151,1436,890,1913]" captionTargetBox-2="[183,1404,993,1426]" captionTargetId-0="figure-18@1.[271,1316,179,744]" captionTargetId-1="figure-46@1.[151,1436,890,1913]" captionTargetId-2="figure-459@3.[183,1404,993,1426]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="3" captionText-0="FIGURE 1. Haludars illustration of Tetrodon patoca. From the archive of the British Library (Mss Eur E72, p. 47), photo: Kevin Webb." captionText-1="FIGURE 2. Chelonodon patoca, ZRC66125, 165 mm SL specimen. a, in life. b, shortly after preservation." captionText-2="FIGURE 3. Chelonodon patoca, in life, not preserved, ca. 200 mm. Note 11 dorsal-fin rays. Photo: B. Mahatvaraj." figureDoi-0="http://doi.org/10.5281/zenodo.13916263" figureDoi-1="http://doi.org/10.5281/zenodo.13916265" figureDoi-2="http://doi.org/10.5281/zenodo.13916267" httpUri-0="https://zenodo.org/record/13916263/files/figure.png" httpUri-1="https://zenodo.org/record/13916265/files/figure.png" httpUri-2="https://zenodo.org/record/13916267/files/figure.png" pageId="4" pageNumber="565">Figs. 13</figureCitation>
).
<taxonomicName id="4C21408B09554A25FB94FBB6FAD4531C" box="[1095,1317,1087,1113]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FB94FBB6FAD4531C" box="[1095,1317,1087,1113]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
according to
<bibRefCitation id="EFB046F909554A25FF64FBEAFE845339" author="Hamilton, F." box="[183,373,1123,1149]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamilton (1822)</bibRefCitation>
has 11 dorsal- and 10 anal-fin rays. Habib
<emphasis id="B955E71A09554A25FCA1FBEAFC575339" box="[882,934,1123,1149]" italics="true" pageId="4" pageNumber="565">et al</emphasis>
.s (2018) specimens of
<taxonomicName id="4C21408B09554A25FB6EFBEAFA6D5338" box="[1213,1436,1123,1149]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FB6EFBEAFA6D5338" box="[1213,1436,1123,1149]" italics="true" pageId="4" pageNumber="565">Chelonodon patoca</emphasis>
</taxonomicName>
(
<typeStatus id="549A85AA09554A25FF4CFB0EFEF253E5" box="[159,259,1159,1185]" pageId="4" pageNumber="565" type="holotype">holotype</typeStatus>
and
<typeStatus id="549A85AA09554A25FEE5FB0EFE6753E5" box="[310,406,1159,1185]" pageId="4" pageNumber="565" type="paratype">paratype</typeStatus>
of their
<taxonomicName id="4C21408B09554A25FE20FB0EFD6753E5" authorityName="Habib, Neogi, Og, Lee &amp; Kim" authorityYear="2018" box="[499,662,1159,1185]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="bengalensis">
<emphasis id="B955E71A09554A25FE20FB0EFD6753E5" box="[499,662,1159,1185]" italics="true" pageId="4" pageNumber="565">C. bengalensis</emphasis>
</taxonomicName>
) were recorded with 12 dorsal- and 10 anal-fin rays. This matches our counts of dorsal- and anal-fin rays in ZRC66125, and BMNH 1889.2.1.4116. However, BMNH 1858.8.15.107 has D 12 and A 11, and another individual of
<taxonomicName id="4C21408B09554A25FDB0FB46FD3F53AC" box="[611,718,1231,1257]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FDB0FB46FD3F53AC" box="[611,718,1231,1257]" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
photographed by Beta Mahatvaraj (
<figureCitation id="131A278D09554A25FB8EFB46FB6F53AD" box="[1117,1182,1231,1257]" captionStart="FIGURE 3" captionStartId="3.[152,255,1449,1473]" captionTargetBox="[183,1404,993,1426]" captionTargetId="figure-459@3.[183,1404,993,1426]" captionTargetPageId="3" captionText="FIGURE 3. Chelonodon patoca, in life, not preserved, ca. 200 mm. Note 11 dorsal-fin rays. Photo: B. Mahatvaraj." figureDoi="http://doi.org/10.5281/zenodo.13916267" httpUri="https://zenodo.org/record/13916267/files/figure.png" pageId="4" pageNumber="565">Fig. 3</figureCitation>
) also shows 11 dorsal-fin rays, matching the count provided by
<bibRefCitation id="EFB046F909554A25FDB5FB7AFCD55249" author="Hamilton, F." box="[614,804,1267,1293]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamilton (1822)</bibRefCitation>
. Based on these examples, the dorsal-fin ray count of
<taxonomicName id="4C21408B09554A25FA50FB7AFF125274" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FA50FB7AFF125274" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
is thus D 1112 and the anal-fin ray count A 1011.
<bibRefCitation id="EFB046F909554A25FCFCFA9EFC235275" author="Russell, P." box="[815,978,1303,1329]" pageId="4" pageNumber="565" refId="ref6973" refString="Russell, P. (1803) Descriptions and figures of two hundred fishes: collected at Vizagapatam on the coast of Coromandel. Vols. 1 - 2. Printed by W. Bulmer and Co., Shakspeare Press, Cleveland-Row, for G. and W. Nicol, booksellers to his majesty, Pall-Mall, London, vii + 78 pp. &amp; 85 pp., 197 pls. https: // doi. org / 10.5962 / bhl. title. 140747" type="book" year="1803">Russell (1803)</bibRefCitation>
counted 10 dorsal- and 8 anal-fin rays in
<taxonomicName id="4C21408B09554A25FF44FAB2FF0D5211" authorityName="Bleeker" authorityYear="1850" box="[151,252,1339,1365]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FF44FAB2FF0D5211" box="[151,252,1339,1365]" italics="true" pageId="4" pageNumber="565">C. kappa</emphasis>
</taxonomicName>
and
<bibRefCitation id="EFB046F909554A25FEE0FAB2FD755211" author="Sujatha, K. &amp; Padmavathi, P." box="[307,644,1338,1365]" pageId="4" pageNumber="565" pagination="95 - 104" refId="ref7152" refString="Sujatha, K. &amp; Padmavathi, P. (2015) Taxonomy of puffer fish (Pisces: Tetraodontidae) represented in the catches of Visakhapatnam, central eastern coast of India. Journal of the Marine Biological Association of India, 57 (2), 95 - 104." type="journal article" year="2015">Sujatha &amp; Padmavathi (2015)</bibRefCitation>
, who collected
<specimenCount id="9D27F08109554A25FCE0FAB2FC2B5211" box="[819,986,1339,1365]" count="178" pageId="4" pageNumber="565" type="generic">178 specimens</specimenCount>
of
<taxonomicName id="4C21408B09554A25FC2CFAB2FB955211" authorityName="Bleeker" authorityYear="1850" box="[1023,1124,1339,1365]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FC2CFAB2FB955211" box="[1023,1124,1339,1365]" italics="true" pageId="4" pageNumber="565">C. kappa</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C21408B09554A25FB40FAB2FAF15210" box="[1171,1280,1339,1365]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FB40FAB2FAF15210" box="[1171,1280,1339,1365]" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
) from around its type locality, provide a range of 910 dorsal- and 8 anal-fin rays for this species. We conclude that in addition to the colour pattern, the number of dorsal- and anal-fin rays distinguishes
<taxonomicName id="4C21408B09554A25FC13FA0AFBDC52D8" box="[960,1069,1411,1437]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FC13FA0AFBDC52D8" box="[960,1069,1411,1437]" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
</taxonomicName>
from
<taxonomicName id="4C21408B09554A25FBA2FA0AFB2752D9" authorityName="Bleeker" authorityYear="1850" box="[1137,1238,1411,1437]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FBA2FA0AFB2752D9" box="[1137,1238,1411,1437]" italics="true" pageId="4" pageNumber="565">C. kappa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9E3B0809554A25FF14FA2FFA805169" blockId="4.[151,1437,151,1905]" pageId="4" pageNumber="565">
<emphasis id="B955E71A09554A25FF14FA2FFDBC5285" bold="true" box="[199,589,1446,1473]" pageId="4" pageNumber="565">
<taxonomicName id="4C21408B09554A25FF14FA2FFEA45285" authorityName="Muller" authorityYear="1841" box="[199,341,1446,1473]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FF14FA2FFEA45285" bold="true" box="[199,341,1446,1473]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
vs.
<taxonomicName id="4C21408B09554A25FE59FA2FFDB75285" authorityName="Smith" authorityYear="1958" box="[394,582,1446,1473]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FE59FA2FFDB75285" bold="true" box="[394,582,1446,1473]" italics="true" pageId="4" pageNumber="565">Chelonodontops</emphasis>
</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFB046F909554A25FD84FA2EFCD85285" author="Hamilton, F." box="[599,809,1447,1473]" pageId="4" pageNumber="565" refId="ref5703" refString="Hamilton, F. (1822) An account of the fishes found in the River Ganges and its branches. Constable, Edinburgh, 3 + 405 pp. https: // doi. org / 10.5962 / bhl. title. 59540" type="book" year="1822">Hamiltons (1822)</bibRefCitation>
<taxonomicName id="4C21408B09554A25FCE0FA2EFC1C5284" authorityName="Hamilton" authorityYear="1822" box="[819,1005,1447,1473]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FCE0FA2EFC1C5284" box="[819,1005,1447,1473]" italics="true" pageId="4" pageNumber="565">Tetrodon patoca</emphasis>
</taxonomicName>
has long been included in the genus
<taxonomicName id="4C21408B09554A25FF44FA42FE8552A1" authority="Muller" authorityName="Muller" authorityYear="1841" box="[151,372,1483,1509]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FF44FA42FED152A1" box="[151,288,1483,1509]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
Müller
</taxonomicName>
(see
<bibRefCitation id="EFB046F909554A25FE62FA42FDA652A1" author="Gunther, A." box="[433,599,1483,1509]" pageId="4" pageNumber="565" refId="ref5537" refString="Gunther, A. (1870) Catalogue of the fishes in the British Museum. Catalogue of the Physostomi, containing the families Gymnotidae, Symbranchidae, Muraenidae, Pegasidae, and of the Lophobranchii, Plectognathi, Dipnoi, ... [thru] ... Leptocardii, in the British Museum. Vol. 8. Department of Zoology, British Museum (Natural History), London, xxv + 549 pp." type="book" year="1870">Günther, 1870</bibRefCitation>
).
<bibRefCitation id="EFB046F909554A25FDB8FA42FCFB52A1" author="Muller, J." box="[619,778,1483,1509]" pageId="4" pageNumber="565" pagination="1 - 5" refId="ref6433" refString="Muller, J. (1841) Vergleichende Anatomie der Myxinoiden. Dritte Fortsetzung. Uber das Gefassystem. Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin, 1839, 175 - 304, pls. 1 - 5." type="journal article" year="1841">Müller (1841)</bibRefCitation>
did not, when establishing the name
<emphasis id="B955E71A09554A25FB61FA42FAB352A1" box="[1202,1346,1483,1509]" italics="true" pageId="4" pageNumber="565">
<taxonomicName id="4C21408B09554A25FB61FA42FACF52A1" authorityName="Muller" authorityYear="1841" box="[1202,1342,1483,1509]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">Chelonodon</taxonomicName>
,
</emphasis>
include any species in this genus. According to
<bibRefCitation id="EFB046F909554A25FDBAFA66FD0A514D" author="Tyler, J. C." box="[617,763,1519,1545]" pageId="4" pageNumber="565" pagination="119 - 148" refId="ref7406" refString="Tyler, J. C. (1964). A diagnosis of the two species of South American puffer fishes (Tetraodontidae, Plectognathi) of the genus Colomesus. Proceedings of the Academy of Natural Sciences of Philadelphia, 116 (3), 119 - 148." type="journal article" year="1964">Tyler (1964)</bibRefCitation>
,
<bibRefCitation id="EFB046F909554A25FCD9FA66FC46514D" author="Ruppell, E." box="[778,951,1518,1545]" pageId="4" pageNumber="565" refId="ref6935" refString="Ruppell, E. (1852) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Vierte Abtheilung: Fische und deren Skelette. Sauerlander, Frankfurt am Main, iv + 40 pp." type="book" year="1852">Rüppell (1852)</bibRefCitation>
was the first to list several species under
<taxonomicName id="4C21408B09554A25FF44F99AFED15169" authorityName="Muller" authorityYear="1841" box="[151,288,1555,1581]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FF44F99AFED15169" box="[151,288,1555,1581]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
:
<taxonomicName id="4C21408B09554A25FEFCF99AFDC15169" authority="(Hamilton)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[303,560,1555,1581]" class="Elasmobranchii" family="Tetraodontidae" genus="Leiodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="cutcutia">
<emphasis id="B955E71A09554A25FEFCF99AFE5B5168" box="[303,426,1555,1581]" italics="true" pageId="4" pageNumber="565">C. cutcutia</emphasis>
(Hamilton)
</taxonomicName>
,
<taxonomicName id="4C21408B09554A25FDE8F99AFC005169" authority="(Bloch &amp; Schneider 1801)" baseAuthorityName="Bloch &amp; Schneider" baseAuthorityYear="1801" box="[571,1009,1555,1581]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="psittacus">
<emphasis id="B955E71A09554A25FDE8F99AFD315168" box="[571,704,1555,1581]" italics="true" pageId="4" pageNumber="565">C. psittacus</emphasis>
(
<bibRefCitation id="EFB046F909554A25FD1DF99AFC195169" author="Bloch, M. E. &amp; Schneider, J. G." box="[718,1000,1555,1581]" pageId="4" pageNumber="565" refId="ref5148" refString="Bloch, M. E. &amp; Schneider, J. G. (1801) M. E. Blochii, Systema Ichthyologiae Iconibus cx Ilustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sanderiano, Berlin, lx + 584 pp., 110 pls. https: // doi. org / 10.5962 / bhl. title. 5750" type="book" year="1801">Bloch &amp; Schneider 1801</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C21408B09554A25FBFFF99AFA9D5169" authority="(Freminville 1813)" baseAuthorityName="Freminville" baseAuthorityYear="1813" box="[1068,1388,1555,1581]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="richei">
<emphasis id="B955E71A09554A25FBFFF99AFB7F5169" box="[1068,1166,1555,1581]" italics="true" pageId="4" pageNumber="565">C. richei</emphasis>
(
<bibRefCitation id="EFB046F909554A25FB4FF99AFA955169" author="Freminville, C. P. P." box="[1180,1380,1555,1581]" pageId="4" pageNumber="565" pagination="249 - 253" refId="ref5492" refString="Freminville, C. P. P. (1813) Description de quelques nouvelles especes de poissons de l'ordre des Branchiosteges. Nouveau Bulletin des Sciences, par la Societe Philomathique, Paris, 3, 249 - 253, pl. 4." type="journal article" year="1813">Fréminville 1813</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B9E3B0809554A25FF14F9BEFB065035" blockId="4.[151,1437,151,1905]" pageId="4" pageNumber="565">
In the same year, but after
<bibRefCitation id="EFB046F909554A25FE39F9BEFD645115" author="Ruppell, E." box="[490,661,1590,1617]" pageId="4" pageNumber="565" refId="ref6935" refString="Ruppell, E. (1852) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Vierte Abtheilung: Fische und deren Skelette. Sauerlander, Frankfurt am Main, iv + 40 pp." type="book" year="1852">Rüppell (1852)</bibRefCitation>
,
<bibRefCitation id="EFB046F909554A25FD4CF9BEFCB65115" author="Bleeker, P." box="[671,839,1590,1617]" pageId="4" pageNumber="565" pagination="1 - 20" refId="ref5114" refString="Bleeker, P. (1852) Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20." type="journal article" year="1852">Bleeker (1852)</bibRefCitation>
mentioned two species under
<taxonomicName id="4C21408B09554A25FB47F9BEFAEC5115" authorityName="Muller" authorityYear="1841" box="[1172,1309,1591,1617]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FB47F9BEFAEC5115" box="[1172,1309,1591,1617]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
:
<taxonomicName id="4C21408B09554A25FAF9F9BEFEC75131" authority="Russell" authorityName="Russell" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09554A25FAF9F9BEFF2A5131" italics="true" pageId="4" pageNumber="565">Tetraodon kappa</emphasis>
Russell
</taxonomicName>
and
<taxonomicName id="4C21408B09554A25FEA3F9D2FD935131" authority="Bleeker" authorityName="Bleeker" authorityYear="1850" box="[368,610,1627,1653]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="kunhardtii">
<emphasis id="B955E71A09554A25FEA3F9D2FDF05131" box="[368,513,1627,1653]" italics="true" pageId="4" pageNumber="565">T. kunhardtii</emphasis>
Bleeker
</taxonomicName>
, and listed another three as belonging to this taxon:
<taxonomicName id="4C21408B09554A25FB65F9D2FEC951DD" authority="Russell" authorityName="Russell" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="subSpecies" species="bondaroo" subSpecies="kappa">
<emphasis id="B955E71A09554A25FB65F9D2FF2A51DD" italics="true" pageId="4" pageNumber="565">Tetraodon bondaroo kappa</emphasis>
Russell
</taxonomicName>
,
<taxonomicName id="4C21408B09554A25FE97F9F6FE0451DD" authority="&quot; Bl." authorityName="Bl." box="[324,501,1663,1689]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="hispidus">
<emphasis id="B955E71A09554A25FE97F9F6FE4C51DD" box="[324,445,1663,1689]" italics="true" pageId="4" pageNumber="565">T. hispidus</emphasis>
“Bl.
</taxonomicName>
”, and
<taxonomicName id="4C21408B09554A25FDEDF9F6FCCB51DD" authority="Solander." authorityName="Solander." box="[574,826,1663,1689]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="meleagris">
<emphasis id="B955E71A09554A25FDEDF9F6FD3951DD" box="[574,712,1663,1689]" italics="true" pageId="4" pageNumber="565">T. meleagris</emphasis>
Solander.
</taxonomicName>
<taxonomicName id="4C21408B09554A25FC92F9F6FB9D51DD" authority="Hamilton" authorityName="Hamilton" authorityYear="1822" box="[833,1132,1663,1689]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FC92F9F6FC0651DC" box="[833,1015,1663,1689]" italics="true" pageId="4" pageNumber="565">Tetrodon patoca</emphasis>
Hamilton
</taxonomicName>
, however, was assigned by
<bibRefCitation id="EFB046F909554A25FF44F92AFECE51F9" author="Bleeker, P." box="[151,319,1698,1725]" pageId="4" pageNumber="565" pagination="1 - 20" refId="ref5114" refString="Bleeker, P. (1852) Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20." type="journal article" year="1852">Bleeker (1852)</bibRefCitation>
to
<bibRefCitation id="EFB046F909554A25FEB3F92AFDE151F9" author="Muller, J." box="[352,528,1699,1725]" pageId="4" pageNumber="565" pagination="1 - 5" refId="ref6433" refString="Muller, J. (1841) Vergleichende Anatomie der Myxinoiden. Dritte Fortsetzung. Uber das Gefassystem. Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin, 1839, 175 - 304, pls. 1 - 5." type="journal article" year="1841">Müllers (1841)</bibRefCitation>
genus
<taxonomicName id="4C21408B09554A25FD8DF92AFD0651F9" authorityName="Müller" authorityYear="1843" box="[606,759,1699,1725]" family="Tetraodontidae" genus="Gastrophysus" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FD8DF92AFD0651F9" box="[606,759,1699,1725]" italics="true" pageId="4" pageNumber="565">Gastrophysus</emphasis>
</taxonomicName>
. As
<bibRefCitation id="EFB046F909554A25FCF4F92AFC2151F9" author="Ruppell, E." box="[807,976,1698,1725]" pageId="4" pageNumber="565" refId="ref6935" refString="Ruppell, E. (1852) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Vierte Abtheilung: Fische und deren Skelette. Sauerlander, Frankfurt am Main, iv + 40 pp." type="book" year="1852">Rüppell (1852)</bibRefCitation>
predates
<bibRefCitation id="EFB046F909554A25FBEAF92AFB1251F9" author="Bleeker, P." box="[1081,1251,1698,1725]" pageId="4" pageNumber="565" pagination="1 - 20" refId="ref5114" refString="Bleeker, P. (1852) Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20." type="journal article" year="1852">Bleeker (1852)</bibRefCitation>
, the
<typeStatus id="549A85AA09554A25FAC6F92AFAB451F9" box="[1301,1349,1699,1725]" pageId="4" pageNumber="565">type</typeStatus>
species for
<taxonomicName id="4C21408B09554A25FF12F94EFEBB51A5" authorityName="Muller" authorityYear="1841" box="[193,330,1735,1761]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FF12F94EFEBB51A5" box="[193,330,1735,1761]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
could be chosen only from among the three species originally included in this genus by
<bibRefCitation id="EFB046F909554A25FA97F94EFF145041" author="Ruppell, E." pageId="4" pageNumber="565" refId="ref6935" refString="Ruppell, E. (1852) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Vierte Abtheilung: Fische und deren Skelette. Sauerlander, Frankfurt am Main, iv + 40 pp." type="book" year="1852">Rüppell (1852)</bibRefCitation>
. This selection was made by
<bibRefCitation id="EFB046F909554A25FDEBF962FD385041" author="Tyler, J. C." box="[568,713,1771,1797]" pageId="4" pageNumber="565" pagination="119 - 148" refId="ref7406" refString="Tyler, J. C. (1964). A diagnosis of the two species of South American puffer fishes (Tetraodontidae, Plectognathi) of the genus Colomesus. Proceedings of the Academy of Natural Sciences of Philadelphia, 116 (3), 119 - 148." type="journal article" year="1964">Tyler (1964)</bibRefCitation>
, who gave
<taxonomicName id="4C21408B09554A25FC9DF962FBC85041" authority="Hamilton" authorityName="Hamilton" box="[846,1081,1771,1797]" class="Elasmobranchii" family="Tetraodontidae" genus="Leiodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="cutcutia">
<emphasis id="B955E71A09554A25FC9DF962FC375040" box="[846,966,1771,1796]" italics="true" pageId="4" pageNumber="565">T. cutcutia</emphasis>
Hamilton
</taxonomicName>
precedence over the other two originally-included species. As a result, the genus name
<taxonomicName id="4C21408B09554A25FCD6F886FC7F506D" authorityName="Muller" authorityYear="1841" box="[773,910,1807,1833]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FCD6F886FC7F506D" box="[773,910,1807,1833]" italics="true" pageId="4" pageNumber="565">Chelonodon</emphasis>
</taxonomicName>
has not been available for the species
<taxonomicName id="4C21408B09554A25FAE6F886FEF05009" authority="Hamilton" authorityName="Hamilton" authorityYear="1822" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FAE6F886FA6D506C" box="[1333,1436,1807,1832]" italics="true" pageId="4" pageNumber="565">T. patoca</emphasis>
Hamilton
</taxonomicName>
and thus the genus name
<taxonomicName id="4C21408B09554A25FDCEF8BAFCAE5009" authority="Smith 1958" authorityName="Smith" authorityYear="1958" box="[541,863,1842,1869]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09554A25FDCEF8BAFD245009" box="[541,725,1843,1869]" italics="true" pageId="4" pageNumber="565">Chelonodontops</emphasis>
<bibRefCitation id="EFB046F909554A25FD08F8BAFCAE5009" author="Smith, J. L. B." box="[731,863,1842,1869]" pageId="4" pageNumber="565" pagination="156 - 160" refId="ref7061" refString="Smith, J. L. B. (1958) Tetraodont fishes from South and East Africa. Annals and Magazine of Natural History, Series 13, 1 (2), 156 - 160. https: // doi. org / 10.1080 / 00222935808650932" type="journal article" year="1958">Smith 1958</bibRefCitation>
</taxonomicName>
, with the
<typeStatus id="549A85AA09554A25FC1AF8BAFC085009" box="[969,1017,1843,1869]" pageId="4" pageNumber="565">type</typeStatus>
species
<taxonomicName id="4C21408B09554A25FB84F8BAFA805009" authority="Smith 1958" authorityName="Smith" authorityYear="1958" box="[1111,1393,1842,1869]" family="Tetraodontidae" genus="Chelonodontops" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="B955E71A09554A25FB84F8BAFB185009" box="[1111,1257,1843,1869]" italics="true" pageId="4" pageNumber="565">C. pulchellus</emphasis>
<bibRefCitation id="EFB046F909554A25FB23F8BAFA805009" author="Smith, J. L. B." box="[1264,1393,1842,1869]" pageId="4" pageNumber="565" pagination="156 - 160" refId="ref7061" refString="Smith, J. L. B. (1958) Tetraodont fishes from South and East Africa. Annals and Magazine of Natural History, Series 13, 1 (2), 156 - 160. https: // doi. org / 10.1080 / 00222935808650932" type="journal article" year="1958">Smith 1958</bibRefCitation>
</taxonomicName>
has been applied recently for this group of pufferfish species, including
<taxonomicName id="4C21408B09554A25FC43F8DEFB035035" authority="(Kottelat, 2013: 475)" baseAuthorityName="Kottelat" baseAuthorityPageNumber="475" baseAuthorityYear="2013" box="[912,1266,1878,1905]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="565" phylum="Chordata" rank="species" species="patoca">
<emphasis id="B955E71A09554A25FC43F8DEFC0C5034" box="[912,1021,1879,1905]" italics="true" pageId="4" pageNumber="565">C. patoca</emphasis>
(
<bibRefCitation id="EFB046F909554A25FBD8F8DEFB1B5035" author="Kottelat, M." box="[1035,1258,1878,1905]" pageId="4" pageNumber="565" pagination="1 - 663" refId="ref6173" refString="Kottelat, M. (2013) The fishes of the inland waters of southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology Supplement, No. 27, 1 - 663." type="book chapter" year="2013">Kottelat, 2013: 475</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="DF5E6B8009544A24FF4BFABDFE7052D1" ID-DOI="http://doi.org/10.5281/zenodo.13916269" ID-Zenodo-Dep="13916269" httpUri="https://zenodo.org/record/13916269/files/figure.png" pageId="5" pageNumber="566" startId="5.[152,255,1332,1356]" targetBox="[151,1436,181,1308]" targetPageId="5" targetType="figure">
<paragraph id="8B9E3B0809544A24FF4BFABDFE7052D1" blockId="5.[151,1436,1332,1429]" pageId="5" pageNumber="566">
<emphasis id="B955E71A09544A24FF4BFABDFEEA5208" bold="true" box="[152,283,1332,1356]" pageId="5" pageNumber="566">FIGURE 4.</emphasis>
<taxonomicName id="4C21408B09544A24FEF7FABDFE195208" authorityName="Hamilton" authorityYear="1822" box="[292,488,1332,1356]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FEF7FABDFE195208" box="[292,488,1332,1356]" italics="true" pageId="5" pageNumber="566">Chelonodon kappa</emphasis>
</taxonomicName>
, in life, not preserved. a, ca. 50 mm SL, Sundarbans, India, photo: B. Mahatvaraj. b, size unknown, Lembeh, Indonesia, photo: Rickard Zerpe, CC BY 2.0 &lt;https://creativecommons.org/licenses/by/2.0&gt;, via Wikimedia Commons.
</paragraph>
</caption>
<paragraph id="8B9E3B0809544A24FF14FA6EFB8C50BD" blockId="5.[151,1437,1511,2041]" pageId="5" pageNumber="566">
However,
<bibRefCitation id="EFB046F909544A24FE91FA6EFE065145" author="Kottelat, M." box="[322,503,1511,1537]" pageId="5" pageNumber="566" pagination="11 - 94" refId="ref6122" refString="Kottelat, M. (2011) Pieter Bleeker in the Netherlands East Indies (10 March 1842 - ca. 21 September 1860): new biographical data and a chronology of his zoological publications. Ichthyological Exploration of Freshwaters, 22 (1), 11 - 94." type="journal article" year="2011">Kottelat (2011)</bibRefCitation>
pointed out that a number of Bleekers publications were made available as preprints before the official distribution of the respective journal volume, with these preprints predating the distributed journal sometimes by months or years. The publication year for the preprint that included Bleekers “Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel”, was given by
<bibRefCitation id="EFB046F909544A24FB56F9DAFAC45129" author="Kottelat, M." box="[1157,1333,1619,1645]" pageId="5" pageNumber="566" pagination="11 - 94" refId="ref6122" refString="Kottelat, M. (2011) Pieter Bleeker in the Netherlands East Indies (10 March 1842 - ca. 21 September 1860): new biographical data and a chronology of his zoological publications. Ichthyological Exploration of Freshwaters, 22 (1), 11 - 94." type="journal article" year="2011">Kottelat (2011)</bibRefCitation>
as 1850.
<bibRefCitation id="EFB046F909544A24FF44F9FEFEB451D5" author="Bleeker, P." box="[151,325,1654,1681]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker (1850)</bibRefCitation>
contains reference to Müller`s
<taxonomicName id="4C21408B09544A24FD62F9FEFCCD51D5" authorityName="Muller" authorityYear="1841" box="[689,828,1655,1681]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FD62F9FEFCCD51D5" box="[689,828,1655,1681]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
and the above-mentioned species, and thus predates
<bibRefCitation id="EFB046F909544A24FF44F912FEB351F1" author="Ruppell, E." box="[151,322,1690,1717]" pageId="5" pageNumber="566" refId="ref6935" refString="Ruppell, E. (1852) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Vierte Abtheilung: Fische und deren Skelette. Sauerlander, Frankfurt am Main, iv + 40 pp." type="book" year="1852">Rüppell (1852)</bibRefCitation>
by more than one year. It follows, therefore, that the
<typeStatus id="549A85AA09544A24FC72F912FC2051F1" box="[929,977,1691,1717]" pageId="5" pageNumber="566">type</typeStatus>
species of
<taxonomicName id="4C21408B09544A24FB82F912FB2B51F1" authorityName="Muller" authorityYear="1841" box="[1105,1242,1691,1717]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FB82F912FB2B51F1" box="[1105,1242,1691,1717]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
has to be chosen from the species included by
<bibRefCitation id="EFB046F909544A24FE37F936FD65519D" author="Bleeker, P." box="[484,660,1726,1753]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker (1850)</bibRefCitation>
:
<bibRefCitation id="EFB046F909544A24FD4DF936FCB0519D" author="Tyler, J. C." box="[670,833,1727,1753]" pageId="5" pageNumber="566" pagination="119 - 148" refId="ref7406" refString="Tyler, J. C. (1964). A diagnosis of the two species of South American puffer fishes (Tetraodontidae, Plectognathi) of the genus Colomesus. Proceedings of the Academy of Natural Sciences of Philadelphia, 116 (3), 119 - 148." type="journal article" year="1964">Tylers (1964)</bibRefCitation>
<typeStatus id="549A85AA09544A24FC94F936FC89519D" box="[839,888,1727,1753]" pageId="5" pageNumber="566">type</typeStatus>
species designation is invalid. As a
<typeStatus id="549A85AA09544A24FAC1F936FAB2519D" box="[1298,1347,1727,1753]" pageId="5" pageNumber="566">type</typeStatus>
species for
<taxonomicName id="4C21408B09544A24FF12F96AFEBD51B9" authorityName="Muller" authorityYear="1841" box="[193,332,1763,1789]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FF12F96AFEBD51B9" box="[193,332,1763,1789]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
has not to our knowledge previously been selected from among the species originally referred to this genus by
<bibRefCitation id="EFB046F909544A24FEB3F88EFDE35065" author="Bleeker, P." box="[352,530,1798,1825]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker (1850)</bibRefCitation>
, we here designate
<bibRefCitation id="EFB046F909544A24FD2CF88EFC325065" author="Bleeker, P." box="[767,963,1798,1825]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleekers (1850)</bibRefCitation>
<taxonomicName id="4C21408B09544A24FC1EF88EFB635065" box="[973,1170,1799,1825]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FC1EF88EFB635065" box="[973,1170,1799,1825]" italics="true" pageId="5" pageNumber="566">Tetraodon kappa</emphasis>
</taxonomicName>
as the
<typeStatus id="549A85AA09544A24FB3FF88EFAEC5065" box="[1260,1309,1799,1825]" pageId="5" pageNumber="566">type</typeStatus>
species of
<taxonomicName id="4C21408B09544A24FF44F8A2FED35001" authorityName="Muller" authorityYear="1841" box="[151,290,1835,1861]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FF44F8A2FED35001" box="[151,290,1835,1861]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB046F909544A24FEE3F8A2FE015001" author="Bleeker, P." box="[304,496,1834,1861]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleekers (1850)</bibRefCitation>
<taxonomicName id="4C21408B09544A24FE25F8A2FD465001" box="[502,695,1835,1861]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FE25F8A2FD465001" box="[502,695,1835,1861]" italics="true" pageId="5" pageNumber="566">Tetraodon kappa</emphasis>
</taxonomicName>
is “
<taxonomicName id="4C21408B09544A24FD31F8A2FB505001" authority="Russell Corom. Fish I" authorityName="Russell Corom. Fish I" box="[738,1185,1835,1861]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FD31F8A2FC525001" box="[738,931,1835,1861]" italics="true" pageId="5" pageNumber="566">Tetraodon kappa</emphasis>
Russell Corom. Fish I
</taxonomicName>
p. 18, fig. 25” (which, because Russell did not consistently apply the Principle of Binominal Nomenclature, is considered not to have been validly published: ICZN Art. 11.4), as he explained further on p. 16.
<taxonomicName id="4C21408B09544A24FC7DF8FAFAE550C9" authority="Bleeker 1850" authorityName="Bleeker" authorityYear="1850" box="[942,1300,1906,1933]" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FC7DF8FAFB8050C9" box="[942,1137,1907,1933]" italics="true" pageId="5" pageNumber="566">Tetraodon kappa</emphasis>
<bibRefCitation id="EFB046F909544A24FBABF8FAFAE550C9" author="Bleeker, P." box="[1144,1300,1906,1933]" pageId="5" pageNumber="566" pagination="1 - 20" refId="ref5066" refString="Bleeker, P. (1850) Preprint: Bijdrage tot de kennis der Gymnognathen van den Indischen Archipel. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen, 24, 1 - 20. [published in Volume 1852, but preprint issued in 1850]" type="book chapter" year="1850">Bleeker 1850</bibRefCitation>
</taxonomicName>
, as we have shown above, is a junior primary homonym of
<taxonomicName id="4C21408B09544A24FD73F81EFC4C50F5" authority="Hamilton" authorityName="Hamilton" box="[672,957,1943,1969]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FD73F81EFCBD50F5" box="[672,844,1943,1969]" italics="true" pageId="5" pageNumber="566">Tetrodon kappa</emphasis>
Hamilton
</taxonomicName>
, so that
<taxonomicName id="4C21408B09544A24FBC9F81EFAC450F5" authority="Hamilton" authorityName="Hamilton" box="[1050,1333,1943,1969]" class="Actinopterygii" family="Tetraodontidae" genus="Tetrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="species" species="kappa">
<emphasis id="B955E71A09544A24FBC9F81EFB3450F5" box="[1050,1221,1943,1969]" italics="true" pageId="5" pageNumber="566">Tetrodon kappa</emphasis>
Hamilton
</taxonomicName>
becomes the
<typeStatus id="549A85AA09544A24FF12F832FF015091" box="[193,240,1979,2005]" pageId="5" pageNumber="566">type</typeStatus>
species of
<taxonomicName id="4C21408B09544A24FEBEF832FE025091" authorityName="Muller" authorityYear="1841" box="[365,499,1979,2005]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FEBEF832FE025091" box="[365,499,1979,2005]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
. This selection of the
<typeStatus id="549A85AA09544A24FD38F832FCEB5091" box="[747,794,1979,2005]" pageId="5" pageNumber="566">type</typeStatus>
species also assures nomenclatural stability, as the genus name
<taxonomicName id="4C21408B09544A24FF0FF856FE9650BD" authorityName="Muller" authorityYear="1841" box="[220,359,2015,2041]" family="Tetraodontidae" genus="Chelonodon" kingdom="Animalia" order="Tetraodontiformes" pageId="5" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B955E71A09544A24FF0FF856FE9650BD" box="[220,359,2015,2041]" italics="true" pageId="5" pageNumber="566">Chelonodon</emphasis>
</taxonomicName>
has for more than a century been used for this group of pufferfishes.
</paragraph>
</subSubSection>
</treatment>
</document>