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<mods:title id="22F8B9960ABC70D8896558FC0309E0B7">A holistic perspective on species delimitation outperforms all methods based on single data types in freshwater gastropods (Caenogastropoda: Hydrobiidae: Pseudamnicola)</mods:title>
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<mods:title id="C37D1B6BE7145059F52E01162BAE34ED">A holistic perspective on species delimitation outperforms all methods based on single data types in freshwater gastropods (Caenogastropoda: Hydrobiidae: Pseudamnicola)</mods:title>
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<mods:namePart id="81425BB97B01F21EF5524307FCD59BEF">Delicado, Diana</mods:namePart>
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<mods:affiliation id="87D405A5D90D6CD0E48B884BAD28A1A9">Department of Animal Ecology & Systematics, Justus Liebig University Giessen, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany</mods:affiliation>
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<mods:affiliation id="8663765302376392A4425A9A268BC7FB">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:affiliation id="5FC9AB02D0E2DDC1B038A89D718F93AB">Environmental Biomonitoring Laboratory (LR 01 ES 14), Faculty of Sciences of Bizerte, University of Carthage, 7021 Jarzouna, Tunisia</mods:affiliation>
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<mods:affiliation id="899686E43FF749D15B836D821DF23999">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<mods:namePart id="5F9870E70B849736AC26CAA06BB2914F">Hauffe, Torsten</mods:namePart>
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<mods:affiliation id="9459CFA890E1B40ED1CAAD895F0421D4">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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@ -78,9 +81,9 @@ Delicado, Boulaassafer & Hauffe
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<paragraph id="DB79336DFFA0531228E6FCC9F0D1F455" blockId="15.[113,600,857,951]" box="[385,491,857,884]" pageId="15" pageNumber="16">
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(
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<figureCitation id="43FD2FE8FFA0531228ECFCC9F0F3F452" box="[395,457,857,884]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Figs 7</figureCitation>
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<figureCitation id="43FD2FE8FFA0531228ECFCC9F0F3F452" box="[395,457,857,884]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Figs 7</figureCitation>
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,
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<figureCitation id="43FD2FE8FFA0531228B5FCC9F0DAF455" box="[466,480,857,883]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">8</figureCitation>
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<figureCitation id="43FD2FE8FFA0531228B5FCC9F0DAF455" box="[466,480,857,883]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">8</figureCitation>
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)
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</paragraph>
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@ -215,11 +218,11 @@ and M.C.,
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Shell ovate-conic, whorls 4–5, height
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(
|
||||
<figureCitation id="43FD2FE8FFA053122BE0F815F3C9F0BB" box="[647,755,1925,1949]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7A–C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122BE0F815F3C9F0BB" box="[647,755,1925,1949]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7A–C</figureCitation>
|
||||
; Supporting Information, Table S3). Periostracum yellowish. Protoconch ~450 µm wide, whorls 1.5; nucleus ~125 µm wide; protoconch microsculpture pitted (
|
||||
<figureCitation id="43FD2FE8FFA053122DB0FF3FF419F7E1" box="[1239,1315,175,199]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122DB0FF3FF419F7E1" box="[1239,1315,175,199]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7G</figureCitation>
|
||||
). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about two-thirds of total shell length. Aperture slightly oval; inner lip thicker than outer lip; peristome margin simple, straight (
|
||||
<figureCitation id="43FD2FE8FFA053122AEEFEBDF2EFF662" box="[905,981,301,325]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7D</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122AEEFEBDF2EFF662" box="[905,981,301,325]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7D</figureCitation>
|
||||
). Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFA053122A32FEDCF286F642" box="[853,956,332,356]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -234,41 +237,41 @@ Shell ovate-conic, whorls 4–5, height
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFA053122A22FE1BF24AF6C7" blockId="15.[808,1461,144,1609]" pageId="15" pageNumber="16">
|
||||
Operculum oval, brownish, about two and a half whorls; muscle attachment area oval and located near the nucleus (
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFE3AF265F6C7" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7E, F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFE3AF265F6C7" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7E, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA053122A22FE79F404F5BB" blockId="15.[808,1461,144,1609]" pageId="15" pageNumber="16">
|
||||
Radular length intermediate, ~675 µm (~25% of total shell length), with ~65 rows of teeth (
|
||||
<figureCitation id="43FD2FE8FFA053122D15FD98F584F539" box="[1138,1214,520,544]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122D15FD98F584F539" box="[1138,1214,520,544]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7H</figureCitation>
|
||||
). Central tooth formula 4-C-4/1-1 (
|
||||
<figureCitation id="43FD2FE8FFA053122AC1FDB7F2D2F519" box="[934,1000,551,575]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7I</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122AC1FDB7F2D2F519" box="[934,1000,551,575]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7I</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 3-C-3. Inner marginal teeth having 20–23 tapered cusps, shortening towards the base. Outer marginal teeth with ~25 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFA053122D8AFD15F417F5BB" box="[1261,1325,645,669]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Fig. 7J</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122D8AFD15F417F5BB" box="[1261,1325,645,669]" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Fig. 7J</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA053122A22FD35F59DF313" blockId="15.[808,1461,144,1609]" pageId="15" pageNumber="16">
|
||||
Animal darkly pigmented, except for neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFD35F27CF5FA" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFD35F27CF5FA" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8F</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with 17–19 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFCD1F272F45F" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122CEAFCD1F272F45F" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
|
||||
). Stomach slightly longer than wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFA053122A52FC2FF2BAF4F1" box="[821,896,959,983]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122A52FC2FF2BAF4F1" box="[821,896,959,983]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.61), slightly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFA053122D2AFB8DF5ADF313" box="[1101,1175,1053,1077]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122D2AFB8DF5ADF313" box="[1101,1175,1053,1077]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA053122A22FBACF5A9F209" blockId="15.[808,1461,144,1609]" pageId="15" pageNumber="16">
|
||||
Female glandular oviduct approximately four times longer than wide. Albumen gland shorter than capsule gland.
|
||||
<collectingRegion id="1902FD8FFFA053122C1EFBCCF489F355" box="[1401,1459,1116,1139]" country="Turkey" name="Bursa" pageId="15" pageNumber="16">Bursa</collectingRegion>
|
||||
copulatrix pyriform, slightly longer than wide. Bursal duct longer than bursal length. Renal oviduct black-pigmented until the area above the insertion of seminal receptacle, coiled. Seminal receptacle elongate, with a short duct, joining renal oviduct slightly above the insertion point with bursal duct (
|
||||
<figureCitation id="43FD2FE8FFA053122C24FB68F491F229" box="[1347,1451,1272,1296]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122C24FB68F491F229" box="[1347,1451,1272,1296]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8D, E</figureCitation>
|
||||
; Supporting Information, Table S5).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA053122A22FAA7F2A6F16F" blockId="15.[808,1461,144,1609]" pageId="15" pageNumber="16">
|
||||
Male genitalia with a prostate gland approximately twice as long as wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFA053122AC8FA05F2C5F28B" box="[943,1023,1429,1453]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122AC8FA05F2C5F28B" box="[943,1023,1429,1453]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8H</figureCitation>
|
||||
). Penis almost as long as head, gradually tapering, with many folds over the entire surface and a small patch of pigmentation on its distal region (
|
||||
<figureCitation id="43FD2FE8FFA053122DB4FA43F409F2CD" box="[1235,1331,1491,1515]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">Fig. 8F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122DB4FA43F409F2CD" box="[1235,1331,1491,1515]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">Fig. 8F, G</figureCitation>
|
||||
; Supporting Information, Table S6); base moderately wide; attached well behind the right eye; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -350,7 +353,7 @@ populations in Monte Argentario, one from the same area as our studied snails, a
|
|||
</taxonomicName>
|
||||
can potentially occur in other localities in the region.
|
||||
</paragraph>
|
||||
<caption id="8FB963E5FFBF530D29E6FAD6F054F2B0" pageId="16" pageNumber="17" startId="16.[129,194,1350,1374]" targetBox="[129,1473,144,1322]" targetPageId="16" targetType="figure">
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||||
<caption id="8FB963E5FFBF530D29E6FAD6F054F2B0" ID-DOI="http://doi.org/10.5281/zenodo.14774891" ID-Zenodo-Dep="14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="16" pageNumber="17" startId="16.[129,194,1350,1374]" targetBox="[129,1473,144,1322]" targetPageId="16" targetType="figure">
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||||
<paragraph id="DB79336DFFBF530D29E6FAD6F054F2B0" blockId="16.[129,1454,1349,1430]" pageId="16" pageNumber="17">
|
||||
<emphasis id="E9B2EF7FFFBF530D29E6FAD6F1E2F278" bold="true" box="[129,216,1350,1374]" pageId="16" pageNumber="17">Figure 7.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
@ -405,7 +408,7 @@ locality, with the great majority being juveniles. Adult shells differed from th
|
|||
<emphasis id="E9B2EF7FFFBF530D2D33F970F58CF1DE" box="[1108,1206,1760,1784]" italics="true" pageId="16" pageNumber="17">P. lucensis</emphasis>
|
||||
</taxonomicName>
|
||||
according to their generally higher spire and larger aperture (Supporting Information,
|
||||
<figureCitation id="43FD2FE8FFBF530D2A5EF88EF2BFF011" box="[825,901,1822,1847]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="16" pageNumber="17">Fig. S9</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBF530D2A5EF88EF2BFF011" box="[825,901,1822,1847]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="16" pageNumber="17">Fig. S9</figureCitation>
|
||||
).
|
||||
<taxonomicName id="1CC648EEFFBF530D2AC6F88EF5FEF010" ID-CoL="C86QQ" authorityName="Delicado, Boulaassafer & Hauffe" authorityYear="2025" box="[929,1220,1822,1846]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="argentariensis">
|
||||
<emphasis id="E9B2EF7FFFBF530D2AC6F88EF5FEF010" box="[929,1220,1822,1846]" italics="true" pageId="16" pageNumber="17">Pseudamnicola argentariensis</emphasis>
|
||||
|
|
@ -413,7 +416,7 @@ according to their generally higher spire and larger aperture (Supporting Inform
|
|||
differs anatomically by having a larger bursa copulatrix, more slender penis, longer radula, and fewer cusps on the lateral radular teeth (
|
||||
<bibRefCitation id="BF574E9CFFBF530D2C24F8CDF481F053" author="Giusti F" box="[1347,1467,1885,1909]" pageId="16" pageNumber="17" pagination="99 - 355" refId="ref21518" refString="Giusti F. Notulae Malacologicae XXIII. I molluschi terrestri, salmastri e di acqua dolce dell'Elba, Giannutri e scogli minori dell'Arcipelago Toscano. Biogeographia - The Journal of Integrative Biogeography 1976; 5: 99 - 355." type="journal article" year="1976">Giusti 1976</bibRefCitation>
|
||||
; Supporting Information,
|
||||
<figureCitation id="43FD2FE8FFBF530D2D22F8ECF5A7F0B3" box="[1093,1181,1916,1941]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="16" pageNumber="17">Fig. S10</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBF530D2D22F8ECF5A7F0B3" box="[1093,1181,1916,1941]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="16" pageNumber="17">Fig. S10</figureCitation>
|
||||
). Based on these morphological differences, previous studies identified these snails as a
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBE530C2916FFDAF078F779" blockId="17.[113,322,74,96]" box="[113,322,74,96]" pageId="17" pageNumber="18">
|
||||
|
|
@ -421,7 +424,7 @@ differs anatomically by having a larger bursa copulatrix, more slender penis, lo
|
|||
Delicado
|
||||
<emphasis id="E9B2EF7FFFBE530C287EFFDBF078F779" box="[281,322,74,96]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
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</paragraph>
|
||||
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<paragraph id="DB79336DFFBE530C2916FAFAF47EF2B8" blockId="17.[113,1425,1386,1439]" pageId="17" pageNumber="18">
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<emphasis id="E9B2EF7FFFBE530C2916FAFAF1F3F2A4" bold="true" box="[113,201,1386,1410]" pageId="17" pageNumber="18">Figure 8.</emphasis>
|
||||
Anatomy of
|
||||
|
|
|
|||
|
|
@ -1,68 +1,71 @@
|
|||
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<mods:affiliation id="C7DE07E7984A9702DF0A06ED34131B57">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:url id="4CBA4151164545635E278C56F4D66BB0">https://doi.org/10.1093/zoolinnean/zlae010</mods:url>
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<mods:location id="61501B2841654AFC56C8390C3E1EECF1">
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<mods:url id="B200C48F7BE94AC7CED764E21FEE00EC">https://doi.org/10.1093/zoolinnean/zlae010</mods:url>
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<mods:classification id="EE8B1FDF25555FC589223741BCD8C81C">journal article</mods:classification>
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<mods:identifier id="8EAE30EDCB307D7190724AAA0872F76A" type="DOI">10.1093/zoolinnean/zlae010</mods:identifier>
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<mods:identifier id="C5EBB475225DD7A497E9FAEF5C2C5D58" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="A51C3B3B8A7D05783E8D8FC1632E9D6D" type="ZooBank">D1C88C6-B4B9-46D0-94AD-BA7439E6DF22</mods:identifier>
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<mods:classification id="5FDAE85B1A858FADD9D6019784B7C087">journal article</mods:classification>
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<mods:identifier id="BCD92975834B7061756A581A3DCAB8CE" type="CLB-Dataset">307941</mods:identifier>
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<mods:identifier id="86649C57ED221789710F9C561D329D12" type="DOI">10.1093/zoolinnean/zlae010</mods:identifier>
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<mods:identifier id="2B84D2BDBFFC192E60358BAD1680028C" type="Zenodo-Dep">14774861</mods:identifier>
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<mods:identifier id="EDC31A5CA0A90B7A86AC0D50305E1F68" type="ZooBank">D1C88C6-B4B9-46D0-94AD-BA7439E6DF22</mods:identifier>
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<treatment id="536F827BFFA353122AE1FB7AF3B2F5FA" LSID="urn:lsid:plazi:treatment:536F827BFFA353122AE1FB7AF3B2F5FA" httpUri="http://treatment.plazi.org/id/536F827BFFA353122AE1FB7AF3B2F5FA" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
|
||||
<treatment id="536F827BFFA353122AE1FB7AF3B2F5FA" ID-DOI="http://doi.org/10.5281/zenodo.14774863" ID-Zenodo-Dep="14774863" LSID="urn:lsid:plazi:treatment:536F827BFFA353122AE1FB7AF3B2F5FA" httpUri="http://treatment.plazi.org/id/536F827BFFA353122AE1FB7AF3B2F5FA" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda">
|
||||
<subSubSection id="93DC60E6FFA353112AE1FB7AF5F0F202" pageId="12" pageNumber="13" type="nomenclature">
|
||||
<paragraph id="DB79336DFFA353112AE1FB7AF5F0F202" blockId="12.[902,1397,1258,1316]" pageId="12" pageNumber="13">
|
||||
<emphasis id="E9B2EF7FFFA353112AE1FB7AF5F0F202" bold="true" pageId="12" pageNumber="13">
|
||||
|
|
@ -78,9 +81,9 @@ Delicado, Khalloufi & Hauffe
|
|||
<paragraph id="DB79336DFFA353112D2FFAA4F589F268" blockId="12.[1096,1203,1332,1358]" box="[1096,1203,1332,1358]" pageId="12" pageNumber="13">
|
||||
<emphasis id="E9B2EF7FFFA353112D2FFAA4F589F268" bold="true" box="[1096,1203,1332,1358]" pageId="12" pageNumber="13">
|
||||
(
|
||||
<figureCitation id="43FD2FE8FFA353112D34FAA4F5ABF268" box="[1107,1169,1332,1358]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="12" pageNumber="13">Figs 5</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA353112D34FAA4F5ABF268" box="[1107,1169,1332,1358]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="12" pageNumber="13">Figs 5</figureCitation>
|
||||
,
|
||||
<figureCitation id="43FD2FE8FFA353112DFDFAA4F592F268" box="[1178,1192,1332,1358]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="12" pageNumber="13">6</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA353112DFDFAA4F592F268" box="[1178,1192,1332,1358]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="12" pageNumber="13">6</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -164,7 +167,7 @@ collection (
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="8FB963E5FFA253102916FAD6F192F2B0" pageId="13" pageNumber="14" startId="13.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="13" targetType="figure">
|
||||
<caption id="8FB963E5FFA253102916FAD6F192F2B0" ID-DOI="http://doi.org/10.5281/zenodo.14774885" ID-Zenodo-Dep="14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14" startId="13.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="13" targetType="figure">
|
||||
<paragraph id="DB79336DFFA253102916FAD6F192F2B0" blockId="13.[113,1458,1350,1430]" pageId="13" pageNumber="14">
|
||||
<emphasis id="E9B2EF7FFFA253102916FAD6F1F3F278" bold="true" box="[113,201,1350,1374]" pageId="13" pageNumber="14">Figure 5.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
@ -190,11 +193,11 @@ Shells, operculum, and radulae of
|
|||
Shell ovate-conic, whorls 4–4.5, height
|
||||
<quantity id="1C3E9E88FFA253102B6BF96FF34FF031" box="[524,629,1791,1815]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
|
||||
(
|
||||
<figureCitation id="43FD2FE8FFA253102BE0F96FF3C9F031" box="[647,755,1791,1815]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5A–C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102BE0F96FF3C9F031" box="[647,755,1791,1815]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5A–C</figureCitation>
|
||||
; Supporting Information, Table S3). Periostracum yellowish. Protoconch ~450 µm wide, whorls 1.5; nucleus ~150 µm wide; protoconch microsculpture pitted (
|
||||
<figureCitation id="43FD2FE8FFA253102B78F8CDF351F053" box="[543,619,1885,1909]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102B78F8CDF351F053" box="[543,619,1885,1909]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5G</figureCitation>
|
||||
). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about three-quarters of total shell length. Aperture longer than wide, slightly oval; inner lip thicker than outer lip; peristome margin simple, straight (
|
||||
<figureCitation id="43FD2FE8FFA253102DBAFA76F411F2DB" box="[1245,1323,1509,1534]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102DBAFA76F411F2DB" box="[1245,1323,1509,1534]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
|
||||
). Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFA253102DCEF995F42AF13B" box="[1193,1296,1541,1565]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -209,30 +212,30 @@ Shell ovate-conic, whorls 4–4.5, height
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFA253102A22F9D4F290F1BC" blockId="13.[810,1460,1478,1972]" pageId="13" pageNumber="14">
|
||||
Operculum oval, yellowish, about two and a half whorls; muscle attachment area oval and located near the nucleus (
|
||||
<figureCitation id="43FD2FE8FFA253102A52F912F2A3F1BC" box="[821,921,1666,1690]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5E, F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102A52F912F2A3F1BC" box="[821,921,1666,1690]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5E, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA253102A22F932F404F070" blockId="13.[810,1460,1478,1972]" pageId="13" pageNumber="14">
|
||||
Radular length intermediate, ~650 µm (~25% of total shell length), with ~55 rows of teeth (
|
||||
<figureCitation id="43FD2FE8FFA253102D15F951F584F1FF" box="[1138,1214,1729,1753]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102D15F951F584F1FF" box="[1138,1214,1729,1753]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5H</figureCitation>
|
||||
). Central tooth formula 4-C-4/1-1 (
|
||||
<figureCitation id="43FD2FE8FFA253102AC1F970F2D2F1DE" box="[934,1000,1760,1784]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5I</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102AC1F970F2D2F1DE" box="[934,1000,1760,1784]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5I</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 3-C-3. Inner marginal teeth having ~20 tapered cusps, shortening towards the base. Outer marginal teeth with ~25 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFA253102D8AF8AEF417F070" box="[1261,1325,1854,1878]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." pageId="13" pageNumber="14">Fig. 5J</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102D8AF8AEF417F070" box="[1261,1325,1854,1878]" captionStart="Figure 5" captionStartId="13.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-280@13.[114,1458,144,1322]" captionTargetPageId="13" captionText="Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774885" httpUri="https://zenodo.org/record/14774885/files/figure.png" pageId="13" pageNumber="14">Fig. 5J</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA253132A22F8CEF0C4F027" blockId="13.[810,1460,1478,1972]" lastBlockId="14.[129,780,1487,1981]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||||
Animal darkly pigmented, except for neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFA253102CEAF8CEF25CF0B3" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="13" pageNumber="14">Fig. 6F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA253102CEAF8CEF25CF0B3" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="13" pageNumber="14">Fig. 6F, G</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with 17–19 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFA153132B84F99DF1A4F163" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA153132B84F99DF1A4F163" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6A</figureCitation>
|
||||
). Stomach slightly longer than wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFA1531329ECF91BF1ECF185" box="[139,214,1675,1699]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA1531329ECF91BF1ECF185" box="[139,214,1675,1699]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.60), darkly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFA1531328C4F979F0D7F027" box="[419,493,1769,1793]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA1531328C4F979F0D7F027" box="[419,493,1769,1793]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="8FB963E5FFA1531329E6FAFBF5FCF2B9" pageId="14" pageNumber="15" startId="14.[129,194,1387,1411]" targetBox="[129,1473,144,1359]" targetPageId="14" targetType="figure">
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||||
<caption id="8FB963E5FFA1531329E6FAFBF5FCF2B9" ID-DOI="http://doi.org/10.5281/zenodo.14774887" ID-Zenodo-Dep="14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,1387,1411]" targetBox="[129,1473,144,1359]" targetPageId="14" targetType="figure">
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||||
<paragraph id="DB79336DFFA1531329E6FAFBF5FCF2B9" blockId="14.[129,1456,1386,1439]" pageId="14" pageNumber="15">
|
||||
<emphasis id="E9B2EF7FFFA1531329E6FAFBF1E2F2A5" bold="true" box="[129,216,1387,1411]" pageId="14" pageNumber="15">Figure 6.</emphasis>
|
||||
Anatomy of
|
||||
|
|
@ -246,14 +249,14 @@ Anatomy of
|
|||
Female glandular oviduct approximately three times as long as wide. Albumen gland shorter than capsule gland.
|
||||
<collectingRegion id="1902FD8FFFA153132BA8F8B8F233F019" box="[719,777,1832,1855]" country="Turkey" name="Bursa" pageId="14" pageNumber="15">Bursa</collectingRegion>
|
||||
copulatrix large, pyriform, almost as long as wide. Bursal duct longer than bursal length. Renal oviduct black-pigmented until the insertion of seminal receptacle, making one or two loops. Seminal receptacle elongate, with a short duct, joining renal oviduct at the insertion point with bursal duct (
|
||||
<figureCitation id="43FD2FE8FFA153132C2CFA5FF480F2C0" box="[1355,1466,1487,1511]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA153132C2CFA5FF480F2C0" box="[1355,1466,1487,1511]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6D, E</figureCitation>
|
||||
; Supporting Information, Table S5).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFA153132A32F99DF49EF027" blockId="14.[823,1474,1487,1793]" pageId="14" pageNumber="15">
|
||||
Male genitalia with a prostate gland approximately twice as long as wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFA153132A1CF9FBF2F0F1A5" box="[891,970,1643,1667]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA153132A1CF9FBF2F0F1A5" box="[891,970,1643,1667]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6H</figureCitation>
|
||||
). Penis as long as head, gradually tapering, with many folds over the entire surface and a small patch of pigmentation on its distal region (
|
||||
<figureCitation id="43FD2FE8FFA153132D37F93AF58CF1E4" box="[1104,1206,1706,1730]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="14" pageNumber="15">Fig. 6F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA153132D37F93AF58CF1E4" box="[1104,1206,1706,1730]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="14" pageNumber="15">Fig. 6F, G</figureCitation>
|
||||
; Supporting Information, Table S6); base medium wide; attached well behind the right eye; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -301,8 +304,8 @@ According to our DNA analyses, this species is more closely related to its conge
|
|||
from
|
||||
<collectingCountry id="A3D173FDFFA053122BC9FE1BF3C0F685" box="[686,762,395,419]" name="Tunisia" pageId="15" pageNumber="16">Tunisia</collectingCountry>
|
||||
in shell and penis features (Supporting Information,
|
||||
<figureCitation id="43FD2FE8FFA053122BB7FE3AF1E8F6C7" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." pageId="15" pageNumber="16">Figs S7F–L, S</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA0531229B5FE59F02FF6C7" box="[210,277,457,481]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." pageId="15" pageNumber="16">8O–Q</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122BB7FE3AF1E8F6C7" captionStart="Figure 7" captionStartId="16.[129,194,1350,1374]" captionTargetBox="[129,1473,144,1322]" captionTargetId="figure-321@16.[129,1473,144,1322]" captionTargetPageId="16" captionText="Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774891" httpUri="https://zenodo.org/record/14774891/files/figure.png" pageId="15" pageNumber="16">Figs S7F–L, S</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA0531229B5FE59F02FF6C7" box="[210,277,457,481]" captionStart="Figure 8" captionStartId="17.[113,178,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-159@17.[114,1458,144,1358]" captionTargetPageId="17" captionText="Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774893" httpUri="https://zenodo.org/record/14774893/files/figure.png" pageId="15" pageNumber="16">8O–Q</figureCitation>
|
||||
). Otherwise, it differs in having a larger bursa copulatrix and a
|
||||
<emphasis id="E9B2EF7FFFA05312284AFE79F063F526" box="[301,345,489,512]" italics="true" pageId="15" pageNumber="16">COI</emphasis>
|
||||
average sequence divergence of 7.4%. The shell and aperture shapes of
|
||||
|
|
@ -341,7 +344,7 @@ Glöer
|
|||
2010
|
||||
</bibRefCitation>
|
||||
: figs 6–20, for comparison;
|
||||
<figureCitation id="43FD2FE8FFA053122B58FD54F34DF5FA" box="[575,631,708,732]" captionStart="Figure 2" captionStartId="4.[129,194,1781,1805]" captionTargetBox="[129,1473,144,1753]" captionTargetPageId="4" captionText="Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section." pageId="15" pageNumber="16">Fig. 2</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFA053122B58FD54F34DF5FA" box="[575,631,708,732]" captionStart="Figure 2" captionStartId="4.[129,194,1781,1805]" captionTargetBox="[129,1473,144,1753]" captionTargetPageId="4" captionText="Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section." figureDoi="http://doi.org/10.5281/zenodo.14774875" httpUri="https://zenodo.org/record/14774875/files/figure.png" pageId="15" pageNumber="16">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,68 +1,71 @@
|
|||
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<mods:namePart id="6ACB9456CCCFA1938EAF47AE6C903C44">Delicado, Diana</mods:namePart>
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<mods:affiliation id="7475B3811A6C74FDE5E932C2E200B7EA">Department of Animal Ecology & Systematics, Justus Liebig University Giessen, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany</mods:affiliation>
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<mods:nameIdentifier id="FA21A5B6E4DA5284918460F1EE98B31A" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:namePart id="75F1656D27BA16A41885E78B1903E7D7">Delicado, Diana</mods:namePart>
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<mods:affiliation id="BAEEFC7A978FB0D6C4DCA61B8A21E604">Department of Animal Ecology & Systematics, Justus Liebig University Giessen, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany</mods:affiliation>
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<mods:nameIdentifier id="6EA1F035C6537200FA510E98DBCAF105" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6C83742E8FB276892B98B2A40ED114B6">Boulaassafer, Khadija</mods:namePart>
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<mods:affiliation id="9F30C9B3422D65A90D16F6919B04ECC0">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:namePart id="523557B07046481CB2540D531E7A4D47">Boulaassafer, Khadija</mods:namePart>
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<mods:affiliation id="85CE524754804932FF453991ACD6BF5A">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:affiliation id="3CB5C186F2DB76927E2AA67DE5793478">Environmental Biomonitoring Laboratory (LR 01 ES 14), Faculty of Sciences of Bizerte, University of Carthage, 7021 Jarzouna, Tunisia</mods:affiliation>
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<mods:affiliation id="ABDCFC77CBB72921E7B7D6BF6D4A2CF4">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<mods:affiliation id="BBBB871703D330F5637CD3FA8DB72233">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<treatment id="536F827BFFB6530629F3F88FF1FCF7E1" LSID="urn:lsid:plazi:treatment:536F827BFFB6530629F3F88FF1FCF7E1" httpUri="http://treatment.plazi.org/id/536F827BFFB6530629F3F88FF1FCF7E1" lastPageId="27" lastPageNumber="27" pageId="25" pageNumber="26">
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<treatment id="536F827BFFB6530629F3F88FF1FCF7E1" ID-DOI="http://doi.org/10.5281/zenodo.14774877" ID-Zenodo-Dep="14774877" LSID="urn:lsid:plazi:treatment:536F827BFFB6530629F3F88FF1FCF7E1" httpUri="http://treatment.plazi.org/id/536F827BFFB6530629F3F88FF1FCF7E1" lastPageId="27" lastPageNumber="27" pageId="25" pageNumber="26" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda">
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<emphasis id="E9B2EF7FFFB6530429F3F88FF338F07F" bold="true" pageId="25" pageNumber="26">
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|
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@ -78,9 +81,9 @@ Delicado, Boulaassafer & Hauffe
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<paragraph id="DB79336DFFB653042814F8E5F0C3F0A9" blockId="25.[113,600,1909,2004]" box="[371,505,1909,1936]" pageId="25" pageNumber="26">
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<emphasis id="E9B2EF7FFFB653042814F8E5F0C3F0A9" bold="true" box="[371,505,1909,1936]" pageId="25" pageNumber="26">
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(
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<figureCitation id="43FD2FE8FFB65304281AF8E5F0F3F0B6" box="[381,457,1909,1936]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="25" pageNumber="26">Figs 15</figureCitation>
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<figureCitation id="43FD2FE8FFB65304281AF8E5F0F3F0B6" box="[381,457,1909,1936]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="25" pageNumber="26">Figs 15</figureCitation>
|
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,
|
||||
<figureCitation id="43FD2FE8FFB6530428B5F8E6F0D4F0B6" box="[466,494,1910,1936]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="25" pageNumber="26">16</figureCitation>
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<figureCitation id="43FD2FE8FFB6530428B5F8E6F0D4F0B6" box="[466,494,1910,1936]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="25" pageNumber="26">16</figureCitation>
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)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -210,9 +213,9 @@ sa
|
|||
Shell ovate-conic, whorls 4, height 2.0–
|
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<quantity id="1C3E9E88FFB653042DA8F80AF427F094" box="[1231,1309,1946,1970]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="26" unit="mm" value="2.5">2.5 mm</quantity>
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(
|
||||
<figureCitation id="43FD2FE8FFB653042C56F80BF496F094" box="[1329,1452,1946,1971]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="25" pageNumber="26">Fig. 15A, B</figureCitation>
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<figureCitation id="43FD2FE8FFB653042C56F80BF496F094" box="[1329,1452,1946,1971]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="25" pageNumber="26">Fig. 15A, B</figureCitation>
|
||||
; Supporting Information, Table S6). Periostracum yellowish. Protoconch ~400 µm wide, whorls 1.5; nucleus ~150 µm wide; protoconch microsculpture pitted (
|
||||
<figureCitation id="43FD2FE8FFB553072B6AFB11F34AF3BF" box="[525,624,1153,1177]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="26" pageNumber="27">Fig. 15C</figureCitation>
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||||
<figureCitation id="43FD2FE8FFB553072B6AFB11F34AF3BF" box="[525,624,1153,1177]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="26" pageNumber="27">Fig. 15C</figureCitation>
|
||||
). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about three-quarters of total shell length. Aperture longer than wide, slightly oval; inner lip thicker than outer lip; peristome margin simple, straight. Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFB553072898FA8EF35CF210" box="[511,614,1310,1334]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -225,7 +228,7 @@ Shell ovate-conic, whorls 4, height 2.0–
|
|||
<quantity id="1C3E9E88FFB553072B5DFAADF3A2F273" box="[570,664,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="26" pageNumber="27" unit="mm" value="1.03">1.03 mm</quantity>
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||||
.
|
||||
</paragraph>
|
||||
<caption id="8FB963E5FFB5530729E6FC6EF235F314" pageId="26" pageNumber="27" startId="26.[129,194,1022,1046]" targetBox="[129,1473,144,994]" targetPageId="26" targetType="figure">
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<paragraph id="DB79336DFFB5530729E6FC6EF235F314" blockId="26.[129,1424,1022,1075]" pageId="26" pageNumber="27">
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<emphasis id="E9B2EF7FFFB5530729E6FC6EF1DFF330" bold="true" box="[129,229,1022,1046]" pageId="26" pageNumber="27">Figure 16.</emphasis>
|
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Anatomy of
|
||||
|
|
@ -238,30 +241,30 @@ Anatomy of
|
|||
<paragraph id="DB79336DFFB5530729FBFACDF3F1F2B2" blockId="26.[127,778,1122,1961]" pageId="26" pageNumber="27">Operculum oval, yellowish, about two and a half whorls; muscle attachment area oval and located near the nucleus.</paragraph>
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<paragraph id="DB79336DFFB5530729FBFA0BF23EF176" blockId="26.[127,778,1122,1961]" pageId="26" pageNumber="27">
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Radular length intermediate, ~500 µm (~25% of total shell length), with ~45 rows of teeth (
|
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<figureCitation id="43FD2FE8FFB5530728A5FA2BF322F2F4" box="[450,536,1466,1491]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="26" pageNumber="27">Fig.15D</figureCitation>
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<figureCitation id="43FD2FE8FFB5530728A5FA2BF322F2F4" box="[450,536,1466,1491]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="26" pageNumber="27">Fig.15D</figureCitation>
|
||||
). Central tooth formula (5)4-C-4(5)/1-1 (
|
||||
<figureCitation id="43FD2FE8FFB55307282EFA4AF09EF2D4" box="[329,420,1498,1522]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="26" pageNumber="27">Fig. 15E</figureCitation>
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||||
<figureCitation id="43FD2FE8FFB55307282EFA4AF09EF2D4" box="[329,420,1498,1522]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="26" pageNumber="27">Fig. 15E</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 4-C-4. Inner marginal teeth having 20–25 tapered cusps, shortening towards the base. Outer marginal teeth with ~25 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFB553072BC6F9A8F3CEF176" box="[673,756,1592,1616]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." pageId="26" pageNumber="27">Fig. 15F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072BC6F9A8F3CEF176" box="[673,756,1592,1616]" captionStart="Figure 15" captionStartId="25.[113,178,1025,1049]" captionTargetBox="[114,1458,146,998]" captionTargetId="figure-389@25.[114,1458,146,998]" captionTargetPageId="25" captionText="Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="26" pageNumber="27">Fig. 15F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFB5530729FBF9C7F5F9F3BF" blockId="26.[127,778,1122,1961]" lastBlockId="26.[825,1476,1122,1491]" pageId="26" pageNumber="27">
|
||||
Animal darkly pigmented, except for neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFB553072B84F9C7F194F1A9" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16D</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072B84F9C7F194F1A9" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16D</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with ~15 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFB553072BF8F964F3C3F02A" box="[671,761,1780,1804]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072BF8F964F3C3F02A" box="[671,761,1780,1804]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16A</figureCitation>
|
||||
). Stomach slightly longer than wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFB5530729ECF8E1F1D8F0AF" box="[139,226,1905,1929]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB5530729ECF8E1F1D8F0AF" box="[139,226,1905,1929]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.64), darkly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFB553072D3BFB11F589F3BF" box="[1116,1203,1153,1177]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072D3BFB11F589F3BF" box="[1116,1203,1153,1177]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFB553072A32FB31F55DF39F" blockId="26.[825,1476,1122,1491]" box="[853,1127,1185,1209]" pageId="26" pageNumber="27">Female genitalia unknown.</paragraph>
|
||||
<paragraph id="DB79336DFFB553072A32FB50F2C4F2F5" blockId="26.[825,1476,1122,1491]" pageId="26" pageNumber="27">
|
||||
Male genitalia with a prostate gland approximately twice longer than wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFB553072D79FA8EF54DF210" box="[1054,1143,1310,1334]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072D79FA8EF54DF210" box="[1054,1143,1310,1334]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16F</figureCitation>
|
||||
). Penis as long as head, gradually tapering, with many folds over the entire surface and a small patch of pigmentation on its distal region (
|
||||
<figureCitation id="43FD2FE8FFB553072C25FACDF480F252" box="[1346,1466,1373,1397]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." pageId="26" pageNumber="27">Fig. 16D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB553072C25FACDF480F252" box="[1346,1466,1373,1397]" captionStart="Figure 16" captionStartId="26.[129,194,1022,1046]" captionTargetBox="[129,1473,144,994]" captionTargetId="figure-484@26.[129,1473,144,994]" captionTargetPageId="26" captionText="Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774911" httpUri="https://zenodo.org/record/14774911/files/figure.png" pageId="26" pageNumber="27">Fig. 16D, E</figureCitation>
|
||||
; Supporting Information, Table S6); base moderately wide; attachment area central; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,68 +1,71 @@
|
|||
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<mods:affiliation id="54AB884C4364D7EB1D702135831CC331">Department of Animal Ecology & Systematics, Justus Liebig University Giessen, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany</mods:affiliation>
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<mods:nameIdentifier id="24821D797A4AD0C14C62E6CFA40E6C4C" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:nameIdentifier id="8673C3007446E0982ACB22D5A08049E7" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:namePart id="38947257926A321E7632DB000E23CE14">Boulaassafer, Khadija</mods:namePart>
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<mods:affiliation id="801D77BDBBB199C46606283D25AE740E">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:namePart id="E7F7865746DB25B5B364A564B371A225">Boulaassafer, Khadija</mods:namePart>
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<mods:namePart id="AFC8A2190CFBA2F65247788D1C148D59">Khalloufi, Noureddine</mods:namePart>
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<mods:affiliation id="41F8E61995F78800178F77E3FEE0D826">Environmental Biomonitoring Laboratory (LR 01 ES 14), Faculty of Sciences of Bizerte, University of Carthage, 7021 Jarzouna, Tunisia</mods:affiliation>
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<mods:namePart id="9B80B156101972199B90ADD72FA7911B">Khalloufi, Noureddine</mods:namePart>
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<mods:namePart id="54D7F1D004D7912588E0BA7D60B4517E">Hauffe, Torsten</mods:namePart>
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<mods:affiliation id="2C68157120F7B34B67952BB0481FEFFA">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<mods:namePart id="09631592DAE874E354E1936D01491775">Hauffe, Torsten</mods:namePart>
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<mods:affiliation id="1B78505529B33EF117395BAB5828F565">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<treatment id="536F827BFFB95304299BF913F372F1D0" LSID="urn:lsid:plazi:treatment:536F827BFFB95304299BF913F372F1D0" httpUri="http://treatment.plazi.org/id/536F827BFFB95304299BF913F372F1D0" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
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<treatment id="536F827BFFB95304299BF913F372F1D0" ID-DOI="http://doi.org/10.5281/zenodo.14774873" ID-Zenodo-Dep="14774873" LSID="urn:lsid:plazi:treatment:536F827BFFB95304299BF913F372F1D0" httpUri="http://treatment.plazi.org/id/536F827BFFB95304299BF913F372F1D0" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda">
|
||||
<subSubSection id="93DC60E6FFB9530B299BF913F35EF19B" pageId="22" pageNumber="23" type="nomenclature">
|
||||
<paragraph id="DB79336DFFB9530B299BF913F35EF19B" blockId="22.[252,654,1667,1725]" pageId="22" pageNumber="23">
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||||
<emphasis id="E9B2EF7FFFB9530B299BF913F35EF19B" bold="true" pageId="22" pageNumber="23">
|
||||
|
|
@ -78,9 +81,9 @@ Delicado, Boulaassafer & Hauffe
|
|||
<paragraph id="DB79336DFFB9530B28E5F971F332F1DD" blockId="22.[129,724,1761,1886]" box="[386,520,1761,1787]" pageId="22" pageNumber="23">
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<emphasis id="E9B2EF7FFFB9530B28E5F971F332F1DD" bold="true" box="[386,520,1761,1787]" pageId="22" pageNumber="23">
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(
|
||||
<figureCitation id="43FD2FE8FFB9530B28EBF971F0E2F1DD" box="[396,472,1761,1787]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="22" pageNumber="23">Figs 13</figureCitation>
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||||
<figureCitation id="43FD2FE8FFB9530B28EBF971F0E2F1DD" box="[396,472,1761,1787]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="22" pageNumber="23">Figs 13</figureCitation>
|
||||
,
|
||||
<figureCitation id="43FD2FE8FFB9530B2885F971F0C4F1DD" box="[482,510,1761,1787]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="22" pageNumber="23">14</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB9530B2885F971F0C4F1DD" box="[482,510,1761,1787]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="22" pageNumber="23">14</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -206,7 +209,7 @@ A spring discharging to the
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="8FB963E5FFB8530A2916FAD6F313F2B0" pageId="23" pageNumber="24" startId="23.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="23" targetType="figure">
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||||
<caption id="8FB963E5FFB8530A2916FAD6F313F2B0" ID-DOI="http://doi.org/10.5281/zenodo.14774905" ID-Zenodo-Dep="14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24" startId="23.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="23" targetType="figure">
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||||
<paragraph id="DB79336DFFB8530A2916FAD6F313F2B0" blockId="23.[113,1455,1349,1430]" pageId="23" pageNumber="24">
|
||||
<emphasis id="E9B2EF7FFFB8530A2916FAD6F1ECF278" bold="true" box="[113,214,1350,1374]" pageId="23" pageNumber="24">Figure 13.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
@ -236,11 +239,11 @@ population); protoconch microsculpture pitted; central radular tooth formula (4)
|
|||
Shell ovate-conic, whorls 3.5–4, height 2.00–
|
||||
<quantity id="1C3E9E88FFB8530A2B38F88EF386F010" box="[607,700,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="23" pageNumber="24" unit="mm" value="3.25">3.25 mm</quantity>
|
||||
, with a fine net-like surface sculpture (in the type population) (
|
||||
<figureCitation id="43FD2FE8FFB8530A2BB3F8ADF186F053" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13A–C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2BB3F8ADF186F053" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13A–C</figureCitation>
|
||||
; Supporting Information, Table S3). Periostracum yellowish to whitish. Protoconch ~500 µm wide, eroded in the type population, whorls 1.5; nucleus ~150 µm wide; protoconch microsculpture pitted (
|
||||
<figureCitation id="43FD2FE8FFB8530A2D75FA56F552F2F8" box="[1042,1128,1478,1502]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig.13G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2D75FA56F552F2F8" box="[1042,1128,1478,1502]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig.13G</figureCitation>
|
||||
). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about three-quarters of total shell length. Aperture longer than wide, slightly oval; inner lip thicker than outer lip; peristome margin simple, straight (
|
||||
<figureCitation id="43FD2FE8FFB8530A2A82F9D3F57EF17D" box="[997,1092,1603,1627]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13D</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2A82F9D3F57EF17D" box="[997,1092,1603,1627]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13D</figureCitation>
|
||||
). Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFB8530A2A96F9F2F562F15C" box="[1009,1112,1634,1658]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -255,19 +258,19 @@ Shell ovate-conic, whorls 3.5–4, height 2.00–
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFB8530A2A22F931F247F1DE" blockId="23.[808,1460,1478,1972]" pageId="23" pageNumber="24">
|
||||
Operculum oval, yellowish, about two and a half whorls; muscle attachment area oval and located near the nucleus (
|
||||
<figureCitation id="43FD2FE8FFB8530A2CEAF950F256F1D1" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13E, F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2CEAF950F256F1D1" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13E, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFB8530A2A22F96FF440F092" blockId="23.[808,1460,1478,1972]" pageId="23" pageNumber="24">
|
||||
Radular length intermediate, ~600 µm (~25% of total shell length), with ~55 rows of teeth (
|
||||
<figureCitation id="43FD2FE8FFB8530A2D0CF88EF5F9F011" box="[1131,1219,1822,1847]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2D0CF88EF5F9F011" box="[1131,1219,1822,1847]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13H</figureCitation>
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||||
). Central tooth formula (4)3-C-3(4)/1-1 (
|
||||
<figureCitation id="43FD2FE8FFB8530A2A8FF8AEF50EF070" box="[1000,1076,1854,1878]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13I</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2A8FF8AEF50EF070" box="[1000,1076,1854,1878]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13I</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 3-C-3. Inner marginal teeth having 15–20 tapered cusps, shortening towards the base. Outer marginal teeth with ~20 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFB8530A2C7AF80CF453F092" box="[1309,1385,1948,1972]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." pageId="23" pageNumber="24">Fig. 13J</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB8530A2C7AF80CF453F092" box="[1309,1385,1948,1972]" captionStart="Figure 13" captionStartId="23.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-286@23.[114,1458,144,1322]" captionTargetPageId="23" captionText="Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.14774905" httpUri="https://zenodo.org/record/14774905/files/figure.png" pageId="23" pageNumber="24">Fig. 13J</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="8FB963E5FFB7530529E6FAFAF469F2B8" pageId="24" pageNumber="25" startId="24.[129,194,1386,1410]" targetBox="[129,1473,144,1358]" targetPageId="24" targetType="figure">
|
||||
<caption id="8FB963E5FFB7530529E6FAFAF469F2B8" ID-DOI="http://doi.org/10.5281/zenodo.14774907" ID-Zenodo-Dep="14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25" startId="24.[129,194,1386,1410]" targetBox="[129,1473,144,1358]" targetPageId="24" targetType="figure">
|
||||
<paragraph id="DB79336DFFB7530529E6FAFAF469F2B8" blockId="24.[129,1408,1386,1439]" pageId="24" pageNumber="25">
|
||||
<emphasis id="E9B2EF7FFFB7530529E6FAFAF1DFF2A4" bold="true" box="[129,229,1386,1410]" pageId="24" pageNumber="25">Figure 14.</emphasis>
|
||||
Anatomy of
|
||||
|
|
@ -279,27 +282,27 @@ Anatomy of
|
|||
</caption>
|
||||
<paragraph id="DB79336DFFB7530529FBFA5EF3DCF078" blockId="24.[128,778,1486,2012]" pageId="24" pageNumber="25">
|
||||
Animal darkly pigmented, with pigmentation lighter on neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFB753052837FA7EF09FF120" box="[336,421,1518,1542]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052837FA7EF09FF120" box="[336,421,1518,1542]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14F</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with ~20 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFB753052879F91AF043F184" box="[286,377,1674,1698]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052879F91AF043F184" box="[286,377,1674,1698]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14A</figureCitation>
|
||||
). Stomach slightly longer than wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFB753052813F978F0F2F026" box="[372,456,1768,1792]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052813F978F0F2F026" box="[372,456,1768,1792]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.52), darkly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFB753052BE7F8D6F3ECF078" box="[640,726,1862,1886]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052BE7F8D6F3ECF078" box="[640,726,1862,1886]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFB7530529FBF8F6F292F142" blockId="24.[128,778,1486,2012]" lastBlockId="24.[824,1474,1486,1918]" pageId="24" pageNumber="25">
|
||||
Female glandular oviduct approximately three times as long as wide. Albumen gland shorter than capsule gland.
|
||||
<collectingRegion id="1902FD8FFFB7530529E6F835F184F09A" box="[129,190,1957,1980]" country="Turkey" name="Bursa" pageId="24" pageNumber="25">Bursa</collectingRegion>
|
||||
copulatrix small, pyriform, longer than wide. Bursal duct slightly longer than bursal length. Renal oviduct black-pigmented until the area above the insertion of seminal receptacle, relatively straight. Seminal receptacle elongate, with a short duct, joining renal oviduct at the insertion point with bursal duct (
|
||||
<figureCitation id="43FD2FE8FFB753052D75F9BCF5A3F162" box="[1042,1177,1580,1604]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052D75F9BCF5A3F162" box="[1042,1177,1580,1604]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14D, E</figureCitation>
|
||||
; Supporting Information, Table S5).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFB753052A32F9FBF561F058" blockId="24.[824,1474,1486,1918]" pageId="24" pageNumber="25">
|
||||
Male genitalia with a prostate gland approximately twice longer than wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFB753052D7BF959F540F1C7" box="[1052,1146,1737,1761]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052D7BF959F540F1C7" box="[1052,1146,1737,1761]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14H</figureCitation>
|
||||
). Penis as long as head, gradually tapering, with many folds over the entire surface and a small patch of pigmentation on its distal region (
|
||||
<figureCitation id="43FD2FE8FFB753052C24F898F480F006" box="[1347,1466,1800,1824]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." pageId="24" pageNumber="25">Fig. 14F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFB753052C24F898F480F006" box="[1347,1466,1800,1824]" captionStart="Figure 14" captionStartId="24.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-301@24.[129,1473,144,1358]" captionTargetPageId="24" captionText="Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774907" httpUri="https://zenodo.org/record/14774907/files/figure.png" pageId="24" pageNumber="25">Fig. 14F, G</figureCitation>
|
||||
; Supporting Information, Table S6); base moderately wide; attached well behind the right eye; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -309,7 +312,7 @@ Male genitalia with a prostate gland approximately twice longer than wide, bean-
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFB753052A5EF854F5ABF0FA" blockId="24.[825,1169,1988,2012]" box="[825,1169,1988,2012]" pageId="24" pageNumber="25">Endemic to a small coastal spring.</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="8FB963E5FFB653042916FB91F5DBF313" pageId="25" pageNumber="26" startId="25.[113,178,1025,1049]" targetBox="[114,1458,146,998]" targetPageId="25" targetType="figure">
|
||||
<caption id="8FB963E5FFB653042916FB91F5DBF313" ID-DOI="http://doi.org/10.5281/zenodo.14774909" ID-Zenodo-Dep="14774909" httpUri="https://zenodo.org/record/14774909/files/figure.png" pageId="25" pageNumber="26" startId="25.[113,178,1025,1049]" targetBox="[114,1458,146,998]" targetPageId="25" targetType="figure">
|
||||
<paragraph id="DB79336DFFB653042916FB91F5DBF313" blockId="25.[113,1410,1025,1078]" pageId="25" pageNumber="26">
|
||||
<emphasis id="E9B2EF7FFFB653042916FB91F1ECF33F" bold="true" box="[113,214,1025,1049]" pageId="25" pageNumber="26">Figure 15.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
|
|||
|
|
@ -1,68 +1,71 @@
|
|||
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<mods:namePart id="87191C72F243CDC8A3581472B4D0776A">Delicado, Diana</mods:namePart>
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<mods:affiliation id="D025FFD701F764182A4D90B207C10E3B">Department of Animal Ecology & Systematics, Justus Liebig University Giessen, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany</mods:affiliation>
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<mods:nameIdentifier id="8229B9DC4D97EB0DCD381F659DB98F56" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:affiliation id="AE236F72B074C94BEDB1CCBFFBCD3EDF">Muséum d’Histoire Naturelle de Marrakech, Cadi Ayyad University, Prince Moulay Abdellah Boulevard, B. P. 2390, 40000 Marrakesh, Morocco</mods:affiliation>
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<mods:affiliation id="6CEF813273B9B9D63C7DFB269AEF248C">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<mods:affiliation id="59B20CBF029C24F894FB89FEF97A77BE">Department of Biology, University of Fribourg, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland & Swiss Institute of Bioinformatics, Chemin du Musée 10, CH- 1700 Fribourg, Switzerland</mods:affiliation>
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<treatment id="536F827BFFBB530B2995FC42F3CBF145" LSID="urn:lsid:plazi:treatment:536F827BFFBB530B2995FC42F3CBF145" httpUri="http://treatment.plazi.org/id/536F827BFFBB530B2995FC42F3CBF145" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<treatment id="536F827BFFBB530B2995FC42F3CBF145" ID-DOI="http://doi.org/10.5281/zenodo.14774869" ID-Zenodo-Dep="14774869" LSID="urn:lsid:plazi:treatment:536F827BFFBB530B2995FC42F3CBF145" httpUri="http://treatment.plazi.org/id/536F827BFFBB530B2995FC42F3CBF145" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda">
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<subSubSection id="93DC60E6FFBB53092995FC42F368F32A" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="DB79336DFFBB53092995FC42F368F32A" blockId="20.[242,664,978,1036]" pageId="20" pageNumber="21">
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<emphasis id="E9B2EF7FFFBB53092995FC42F368F32A" bold="true" pageId="20" pageNumber="21">
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|
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@ -78,9 +81,9 @@ Delicado, Khalloufi & Hauffe
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<paragraph id="DB79336DFFBB530928E5FBB0F332F31C" blockId="20.[129,644,1056,1182]" box="[386,520,1056,1083]" pageId="20" pageNumber="21">
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<emphasis id="E9B2EF7FFFBB530928E5FBB0F332F31C" bold="true" box="[386,520,1056,1083]" pageId="20" pageNumber="21">
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(
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<figureCitation id="43FD2FE8FFBB530928EBFBB1F0E2F31D" box="[396,472,1057,1083]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Figs 11</figureCitation>
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||||
<figureCitation id="43FD2FE8FFBB530928EBFBB1F0E2F31D" box="[396,472,1057,1083]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
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||||
,
|
||||
<figureCitation id="43FD2FE8FFBB53092885FBB1F0C4F31D" box="[482,510,1057,1083]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">12</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092885FBB1F0C4F31D" box="[482,510,1057,1083]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">12</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -232,11 +235,11 @@ and
|
|||
Shell ovate-conic, whorls 4–4.5, height
|
||||
<quantity id="1C3E9E88FFBB53092DBEFEBDF415F662" box="[1241,1327,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="20" pageNumber="21" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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(
|
||||
<figureCitation id="43FD2FE8FFBB53092C25FEBDF276F645" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11A, C, D</figureCitation>
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||||
<figureCitation id="43FD2FE8FFBB53092C25FEBDF276F645" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11A, C, D</figureCitation>
|
||||
; Supporting Information, Table S3). Periostracum yellowish. Protoconch ~450 µm wide, whorls 1.5; nucleus ~120 µm wide; protoconch microsculpture pitted (
|
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<figureCitation id="43FD2FE8FFBB53092DAFFE1BF412F685" box="[1224,1320,395,419]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11F</figureCitation>
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<figureCitation id="43FD2FE8FFBB53092DAFFE1BF412F685" box="[1224,1320,395,419]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11F</figureCitation>
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). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about three-quarters of total shell length. Aperture almost as wide as long, slightly oval; inner lip thicker than outer lip; peristome margin simple, straight (
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<figureCitation id="43FD2FE8FFBB53092D84FD98F406F506" box="[1251,1340,520,544]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092D84FD98F406F506" box="[1251,1340,520,544]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11B</figureCitation>
|
||||
). Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFBB53092DDFFDB7F425F519" box="[1208,1311,551,575]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -251,41 +254,41 @@ Shell ovate-conic, whorls 4–4.5, height
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB53092A32FDF6F296F59B" blockId="20.[824,1476,263,1829]" pageId="20" pageNumber="21">
|
||||
Operculum oval, brownish, about two and a half whorls; muscle attachment area oval and located near the nucleus (
|
||||
<figureCitation id="43FD2FE8FFBB53092A23FD35F2A1F59B" box="[836,923,677,701]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092A23FD35F2A1F59B" box="[836,923,677,701]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB53092A32FD54F5D5F4BE" blockId="20.[824,1476,263,1829]" pageId="20" pageNumber="21">
|
||||
Radular length intermediate, ~750 µm (~25% of total shell length), with ~55 rows of teeth (
|
||||
<figureCitation id="43FD2FE8FFBB53092DD4FD73F429F5DD" box="[1203,1299,739,763]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092DD4FD73F429F5DD" box="[1203,1299,739,763]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11G</figureCitation>
|
||||
). Central tooth formula (4)3-C-3(4)/1-1 (
|
||||
<figureCitation id="43FD2FE8FFBB53092D3FFC93F58FF43D" box="[1112,1205,771,795]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092D3FFC93F58FF43D" box="[1112,1205,771,795]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11H</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 2-C-2 (rarely with three lateral cusps). Inner marginal teeth having 16–17 tapered cusps, shortening towards the base. Outer marginal teeth with ~22 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFBB53092DF6FC10F5E5F4BE" box="[1169,1247,896,920]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." pageId="20" pageNumber="21">Fig. 11I</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092DF6FC10F5E5F4BE" box="[1169,1247,896,920]" captionStart="Figure 11" captionStartId="21.[113,178,1350,1374]" captionTargetBox="[114,1458,144,1322]" captionTargetId="figure-329@21.[114,1458,144,1322]" captionTargetPageId="21" captionText="Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P)." figureDoi="http://doi.org/10.5281/zenodo.14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="20" pageNumber="21">Fig. 11I</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB53092A32FC0FF54CF209" blockId="20.[824,1476,263,1829]" pageId="20" pageNumber="21">
|
||||
Animal black-pigmented except for neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFBB53092CFBFC0FF25BF4F1" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092CFBFC0FF25BF4F1" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12F</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with 16–19 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFBB53092C30FBACF48BF372" box="[1367,1457,1084,1108]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092C30FBACF48BF372" box="[1367,1457,1084,1108]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12A</figureCitation>
|
||||
). Stomach as long as wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFBB53092C03FB0AF481F394" box="[1380,1467,1178,1202]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092C03FB0AF481F394" box="[1380,1467,1178,1202]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.52), darkly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFBB53092D69FA87F55FF209" box="[1038,1125,1303,1327]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092D69FA87F55FF209" box="[1038,1125,1303,1327]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB53092A32FAA7F598F10C" blockId="20.[824,1476,263,1829]" pageId="20" pageNumber="21">
|
||||
Female glandular oviduct approximately three times as long as wide. Albumen gland shorter than capsule gland.
|
||||
<collectingRegion id="1902FD8FFFBB53092CEFFAC6F4F8F24B" box="[1416,1474,1366,1389]" country="Turkey" name="Bursa" pageId="20" pageNumber="21">Bursa</collectingRegion>
|
||||
copulatrix pyriform, longer than wide. Bursal duct longer than bursal length. Renal oviduct black-pigmented until the insertion of seminal receptacle or the area above, relatively straight. Seminal receptacle elongate, with a short duct, joining renal oviduct at the insertion point with bursal duct (
|
||||
<figureCitation id="43FD2FE8FFBB53092C27FA63F480F12C" box="[1344,1466,1523,1547]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092C27FA63F480F12C" box="[1344,1466,1523,1547]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12D, E</figureCitation>
|
||||
; Supporting Information, Table S5).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB53092A32F9A1F44FF003" blockId="20.[824,1476,263,1829]" pageId="20" pageNumber="21">
|
||||
Male genitalia with a prostate gland approximately three times as long as wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFBB53092A93F91FF56AF181" box="[1012,1104,1679,1703]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092A93F91FF56AF181" box="[1012,1104,1679,1703]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12H</figureCitation>
|
||||
). Penis longer than head, gradually tapering, unpigmented, with many folds over the entire surface (
|
||||
<figureCitation id="43FD2FE8FFBB53092A23F95EF28EF1C0" box="[836,948,1742,1766]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 12F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092A23F95EF28EF1C0" box="[836,948,1742,1766]" captionStart="Figure 12" captionStartId="22.[129,194,1386,1410]" captionTargetBox="[129,1473,144,1358]" captionTargetId="figure-177@22.[129,1473,144,1358]" captionTargetPageId="22" captionText="Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774903" httpUri="https://zenodo.org/record/14774903/files/figure.png" pageId="20" pageNumber="21">Fig. 12F, G</figureCitation>
|
||||
; Supporting Information, Table S6); base narrower than central area of penis; attached well behind the right eye; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -335,7 +338,7 @@ sp.
|
|||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="8FB963E5FFBA53082916FAD6F33AF294" pageId="21" pageNumber="22" startId="21.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="21" targetType="figure">
|
||||
<caption id="8FB963E5FFBA53082916FAD6F33AF294" ID-DOI="http://doi.org/10.5281/zenodo.14774901" ID-Zenodo-Dep="14774901" httpUri="https://zenodo.org/record/14774901/files/figure.png" pageId="21" pageNumber="22" startId="21.[113,178,1350,1374]" targetBox="[114,1458,144,1322]" targetPageId="21" targetType="figure">
|
||||
<paragraph id="DB79336DFFBA53082916FAD6F33AF294" blockId="21.[113,1451,1349,1458]" pageId="21" pageNumber="22">
|
||||
<emphasis id="E9B2EF7FFFBA53082916FAD6F1ECF278" bold="true" box="[113,214,1350,1374]" pageId="21" pageNumber="22">Figure 11.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
@ -424,10 +427,10 @@ Boulaassafer
|
|||
2020
|
||||
</bibRefCitation>
|
||||
) and by all our phylogenetic analyses (
|
||||
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<figureCitation id="43FD2FE8FFBA53082D90F808F40BF097" box="[1271,1329,1944,1969]" captionStart="Figure 2" captionStartId="4.[129,194,1781,1805]" captionTargetBox="[129,1473,144,1753]" captionTargetPageId="4" captionText="Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section." figureDoi="http://doi.org/10.5281/zenodo.14774875" httpUri="https://zenodo.org/record/14774875/files/figure.png" pageId="21" pageNumber="22">Fig. 2</figureCitation>
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; Supporting Information,
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<figureCitation id="43FD2FE8FFBA53082AD6F828F52CF0F6" box="[945,1046,1976,2000]" captionStart="Figure 4" captionStartId="9.[113,178,1788,1812]" captionTargetBox="[114,1457,149,1760]" captionTargetPageId="9" captionText="Figure 4. Species validation by IBPP based on the species-tree topology from DELINEATE as the guide tree. Posterior probabilities of classification support are shown as three coloured boxes above branches for analyses with morphological linear measurements alone (left box), with mitochondrial and nuclear gene sequences (middle box), and with all linear measurements and gene sequences (right box). Tips group the populations belonging to a particular species and are connected to our proposed taxonomic name.These are followed by the number of DNA-based methods that support each depicted species grouping.Pie charts on the right provide the percentage of individuals classified within a delimited species according to the canonical variate analysis of shell shape, shell linear measurements, and measurements of the female and male genitalia.According to DNA-sequence data, populations marked with an asterisk were lumped into a single species, but given their high morphological differentiation, they are now considered three separate species.Abbreviation: ND, no data available." pageId="21" pageNumber="22">Figs S4–S</figureCitation>
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<figureCitation id="43FD2FE8FFBA53082D71F828F519F0F6" box="[1046,1059,1976,2000]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." pageId="21" pageNumber="22">6</figureCitation>
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<figureCitation id="43FD2FE8FFBA53082AD6F828F52CF0F6" box="[945,1046,1976,2000]" captionStart="Figure 4" captionStartId="9.[113,178,1788,1812]" captionTargetBox="[114,1457,149,1760]" captionTargetPageId="9" captionText="Figure 4. Species validation by IBPP based on the species-tree topology from DELINEATE as the guide tree. Posterior probabilities of classification support are shown as three coloured boxes above branches for analyses with morphological linear measurements alone (left box), with mitochondrial and nuclear gene sequences (middle box), and with all linear measurements and gene sequences (right box). Tips group the populations belonging to a particular species and are connected to our proposed taxonomic name.These are followed by the number of DNA-based methods that support each depicted species grouping.Pie charts on the right provide the percentage of individuals classified within a delimited species according to the canonical variate analysis of shell shape, shell linear measurements, and measurements of the female and male genitalia.According to DNA-sequence data, populations marked with an asterisk were lumped into a single species, but given their high morphological differentiation, they are now considered three separate species.Abbreviation: ND, no data available." figureDoi="http://doi.org/10.5281/zenodo.14774883" httpUri="https://zenodo.org/record/14774883/files/figure.png" pageId="21" pageNumber="22">Figs S4–S</figureCitation>
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<figureCitation id="43FD2FE8FFBA53082D71F828F519F0F6" box="[1046,1059,1976,2000]" captionStart="Figure 6" captionStartId="14.[129,194,1387,1411]" captionTargetBox="[129,1473,144,1359]" captionTargetId="figure-304@14.[129,1473,144,1359]" captionTargetPageId="14" captionText="Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774887" httpUri="https://zenodo.org/record/14774887/files/figure.png" pageId="21" pageNumber="22">6</figureCitation>
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). However, this relationship has always
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<treatment id="536F827BFFBE53092866F9EAF0E1F494" ID-DOI="http://doi.org/10.5281/zenodo.14774867" ID-Zenodo-Dep="14774867" LSID="urn:lsid:plazi:treatment:536F827BFFBE53092866F9EAF0E1F494" httpUri="http://treatment.plazi.org/id/536F827BFFBE53092866F9EAF0E1F494" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" scope_family="Hydrobiidae" scope_subClass="Caenogastropoda">
|
||||
<subSubSection id="93DC60E6FFBE530C2866F9EAF0C8F1D4" pageId="17" pageNumber="18" type="nomenclature">
|
||||
<paragraph id="DB79336DFFBE530C2866F9EAF36EF192" blockId="17.[257,619,1658,1716]" pageId="17" pageNumber="18">
|
||||
<emphasis id="E9B2EF7FFFBE530C2866F9EAF36EF192" bold="true" pageId="17" pageNumber="18">
|
||||
|
|
@ -76,9 +79,9 @@ Delicado, Boulaassafer & Hauffe
|
|||
<paragraph id="DB79336DFFBE530C281DF948F0C8F1D4" blockId="17.[113,600,1752,1878]" box="[378,498,1752,1779]" pageId="17" pageNumber="18">
|
||||
<emphasis id="E9B2EF7FFFBE530C281DF948F0C8F1D4" bold="true" box="[378,498,1752,1779]" pageId="17" pageNumber="18">
|
||||
(
|
||||
<figureCitation id="43FD2FE8FFBE530C28E3F948F0F8F1D4" box="[388,450,1752,1778]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="17" pageNumber="18">Figs 9</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBE530C28E3F948F0F8F1D4" box="[388,450,1752,1778]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="17" pageNumber="18">Figs 9</figureCitation>
|
||||
,
|
||||
<figureCitation id="43FD2FE8FFBE530C28ABF948F0D2F1D4" box="[460,488,1752,1778]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="17" pageNumber="18">10</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBE530C28ABF948F0D2F1D4" box="[460,488,1752,1778]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="17" pageNumber="18">10</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -254,11 +257,11 @@ A stream near Sperlonga,
|
|||
Shell ovate-conic, whorls 3–3.5, height
|
||||
<quantity id="1C3E9E88FFBD530F2DB3FF26F407F7E8" box="[1236,1341,182,206]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="18" pageNumber="19" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
|
||||
(
|
||||
<figureCitation id="43FD2FE8FFBD530F2C28FF26F481F7E8" box="[1359,1467,182,206]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="18" pageNumber="19">Fig. 9A–C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBD530F2C28FF26F481F7E8" box="[1359,1467,182,206]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="18" pageNumber="19">Fig. 9A–C</figureCitation>
|
||||
; Supporting Information, Table S3). Periostracum yellowish. Protoconch ~400 µm wide, whorls 1.5; nucleus ~125 µm wide; protoconch microsculpture pitted (
|
||||
<figureCitation id="43FD2FE8FFBD530F2D80FE84F409F60A" box="[1255,1331,276,300]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="18" pageNumber="19">Fig. 9G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBD530F2D80FE84F409F60A" box="[1255,1331,276,300]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="18" pageNumber="19">Fig. 9G</figureCitation>
|
||||
). Teleoconch whorls moderately convex, with deep sutures; body whorl large, occupying about three-quarters of total shell length. Aperture longer than wide, slightly oval; inner lip thicker than outer lip; peristome margin simple, straight (
|
||||
<figureCitation id="43FD2FE8FFBD530F2D8EFE02F401F68C" box="[1257,1339,402,426]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="18" pageNumber="19">Fig. 9D</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBD530F2D8EFE02F401F68C" box="[1257,1339,402,426]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="18" pageNumber="19">Fig. 9D</figureCitation>
|
||||
). Umbilicus narrow, not covered by the inner lip.
|
||||
<typeStatus id="047D8DCFFFBD530F2DDFFE21F425F6EF" box="[1208,1311,433,457]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||||
: SL =
|
||||
|
|
@ -273,10 +276,10 @@ Shell ovate-conic, whorls 3–3.5, height
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFBD530F2A32FE60F283F560" blockId="18.[825,1476,145,582]" pageId="18" pageNumber="19">
|
||||
Operculum oval, brownish, about two and a half whorls; muscle attachment area oval and located near the nucleus (
|
||||
<figureCitation id="43FD2FE8FFBD530F2A23FDBEF292F560" box="[836,936,558,582]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="18" pageNumber="19">Fig. 9E, F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBD530F2A23FDBEF292F560" box="[836,936,558,582]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="18" pageNumber="19">Fig. 9E, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="8FB963E5FFBD530F29E6F8F8F5A6F09F" pageId="18" pageNumber="19" startId="18.[129,194,1896,1920]" targetBox="[129,1473,690,1868]" targetPageId="18" targetType="figure">
|
||||
<caption id="8FB963E5FFBD530F29E6F8F8F5A6F09F" ID-DOI="http://doi.org/10.5281/zenodo.14774895" ID-Zenodo-Dep="14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="18" pageNumber="19" startId="18.[129,194,1896,1920]" targetBox="[129,1473,690,1868]" targetPageId="18" targetType="figure">
|
||||
<paragraph id="DB79336DFFBD530F29E6F8F8F5A6F09F" blockId="18.[129,1465,1896,1977]" pageId="18" pageNumber="19">
|
||||
<emphasis id="E9B2EF7FFFBD530F29E6F8F8F1E2F0A6" bold="true" box="[129,216,1896,1920]" pageId="18" pageNumber="19">Figure 9.</emphasis>
|
||||
Shells, operculum, and radulae of
|
||||
|
|
@ -288,7 +291,7 @@ Shells, operculum, and radulae of
|
|||
). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="8FB963E5FFBC530E2914FAFAF5C9F2B8" pageId="19" pageNumber="20" startId="19.[115,180,1386,1410]" targetBox="[114,1458,144,1358]" targetPageId="19" targetType="figure">
|
||||
<caption id="8FB963E5FFBC530E2914FAFAF5C9F2B8" ID-DOI="http://doi.org/10.5281/zenodo.14774897" ID-Zenodo-Dep="14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20" startId="19.[115,180,1386,1410]" targetBox="[114,1458,144,1358]" targetPageId="19" targetType="figure">
|
||||
<paragraph id="DB79336DFFBC530E2914FAFAF5C9F2B8" blockId="19.[113,1426,1386,1439]" pageId="19" pageNumber="20">
|
||||
<emphasis id="E9B2EF7FFFBC530E2914FAFAF1EDF2A4" bold="true" box="[115,215,1386,1410]" pageId="19" pageNumber="20">Figure 10.</emphasis>
|
||||
Anatomy of
|
||||
|
|
@ -300,36 +303,36 @@ Anatomy of
|
|||
</caption>
|
||||
<paragraph id="DB79336DFFBC530E29EAFA5EF3BFF1A5" blockId="19.[113,764,1486,1981]" pageId="19" pageNumber="20">
|
||||
Radular length intermediate, ~600 µm (~25% of total shell length), with ~60 rows of teeth (
|
||||
<figureCitation id="43FD2FE8FFBC530E28DEFA7EF33FF120" box="[441,517,1518,1542]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="19" pageNumber="20">Fig. 9H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E28DEFA7EF33FF120" box="[441,517,1518,1542]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="19" pageNumber="20">Fig. 9H</figureCitation>
|
||||
). Central tooth formula 4-C-4/1-1 (
|
||||
<figureCitation id="43FD2FE8FFBC530E298AF99DF015F103" box="[237,303,1549,1573]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="19" pageNumber="20">Fig. 9I</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E298AF99DF015F103" box="[237,303,1549,1573]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="19" pageNumber="20">Fig. 9I</figureCitation>
|
||||
); basal tongue V-shaped, length about equal to lateral margin. Lateral tooth formula 3-C-3. Inner marginal teeth having ~25 tapered cusps, shortening towards the base. Outer marginal teeth with ~20 sharp cusps (
|
||||
<figureCitation id="43FD2FE8FFBC530E2B53F9FBF34FF1A5" box="[564,629,1643,1667]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="19" pageNumber="20">Fig. 9J</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2B53F9FBF34FF1A5" box="[564,629,1643,1667]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="19" pageNumber="20">Fig. 9J</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBC530E29EAF91AF55DF103" blockId="19.[113,764,1486,1981]" lastBlockId="19.[810,1460,1486,1981]" pageId="19" pageNumber="20">
|
||||
Animal darkly pigmented except for neck and tentacles (
|
||||
<figureCitation id="43FD2FE8FFBC530E2BB3F91AF1A3F1E4" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10F</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2BB3F91AF1A3F1E4" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10F</figureCitation>
|
||||
). Snout as long as wide, with medium distal lobation; foot size intermediate, with dorsal pigmentation. Ctenidium with 17–19 well-developed gill filaments, occupying ~50% of pallial cavity length and posteriorly positioned. Osphradium of intermediate width and opposite middle of ctenidium (
|
||||
<figureCitation id="43FD2FE8FFBC530E2BF7F8B7F3D0F019" box="[656,746,1831,1855]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10A</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2BF7F8B7F3D0F019" box="[656,746,1831,1855]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10A</figureCitation>
|
||||
). Stomach as long as wide, with two chambers almost equal in size and a medium-sized posterior caecum; style sac longer than wide, surrounded by an unpigmented intestine (
|
||||
<figureCitation id="43FD2FE8FFBC530E2BFBF815F3C9F0BB" box="[668,755,1925,1949]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10B</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2BFBF815F3C9F0BB" box="[668,755,1925,1949]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10B</figureCitation>
|
||||
; Supporting Information, Table S4). Nervous system elongate (mean RPG ratio =.50), darkly pigmented, darker on ganglia than on connectives and commissures; cerebral ganglia approximately equal in size (
|
||||
<figureCitation id="43FD2FE8FFBC530E2A98F99DF56CF103" box="[1023,1110,1549,1573]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2A98F99DF56CF103" box="[1023,1110,1549,1573]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBC530E2A22F9BCF595F006" blockId="19.[810,1460,1486,1981]" pageId="19" pageNumber="20">
|
||||
Female glandular oviduct approximately four times as long as wide. Albumen gland shorter than capsule gland.
|
||||
<collectingRegion id="1902FD8FFFBC530E2C1EF9DCF489F145" box="[1401,1459,1612,1635]" country="Turkey" name="Bursa" pageId="19" pageNumber="20">Bursa</collectingRegion>
|
||||
copulatrix small, pyriform, longer than wide. Bursal duct longer than bursal length. Renal oviduct black-pigmented until the area above the insertion of seminal receptacle, making one or two loops. Seminal receptacle elongate, with a short duct, joining renal oviduct at the insertion point with bursal duct (
|
||||
<figureCitation id="43FD2FE8FFBC530E2C33F978F203F039" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10D, E</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2C33F978F203F039" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10D, E</figureCitation>
|
||||
; Supporting Information, Table S5).
|
||||
</paragraph>
|
||||
<paragraph id="DB79336DFFBC53092A22F8B7F09EF7C1" blockId="19.[810,1460,1486,1981]" lastBlockId="20.[129,778,144,231]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||||
Male genitalia with a prostate gland approximately three times as long as wide, bean-shaped; seminal duct entering the middle-posterior region; pallial vas deferens emerging close to its anterior edge (
|
||||
<figureCitation id="43FD2FE8FFBC530E2A85F815F507F0BB" box="[994,1085,1925,1949]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="19" pageNumber="20">Fig. 10H</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBC530E2A85F815F507F0BB" box="[994,1085,1925,1949]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="19" pageNumber="20">Fig. 10H</figureCitation>
|
||||
). Penis shorter than head, gradually tapering, unpigmented, with many folds over the entire surface (
|
||||
<figureCitation id="43FD2FE8FFBB530929ECFF00F03EF78E" box="[139,260,144,168]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." pageId="20" pageNumber="21">Fig. 10F, G</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB530929ECFF00F03EF78E" box="[139,260,144,168]" captionStart="Figure 10" captionStartId="19.[115,180,1386,1410]" captionTargetBox="[114,1458,144,1358]" captionTargetId="figure-312@19.[114,1458,144,1358]" captionTargetPageId="19" captionText="Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland." figureDoi="http://doi.org/10.5281/zenodo.14774897" httpUri="https://zenodo.org/record/14774897/files/figure.png" pageId="20" pageNumber="21">Fig. 10F, G</figureCitation>
|
||||
; Supporting Information, Table S6); base wide; attached well behind the right eye; penial duct narrow, curved, coursing close to outer edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -345,7 +348,7 @@ Male genitalia with a prostate gland approximately three times as long as wide,
|
|||
</paragraph>
|
||||
<paragraph id="DB79336DFFBB530929E6FE35F0E1F494" blockId="20.[128,780,383,947]" pageId="20" pageNumber="21">
|
||||
The few individuals available for the study showed little variation in shell features (
|
||||
<figureCitation id="43FD2FE8FFBB53092849FE55F0ACF6FB" box="[302,406,453,477]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." pageId="20" pageNumber="21">Fig. 9A–C</figureCitation>
|
||||
<figureCitation id="43FD2FE8FFBB53092849FE55F0ACF6FB" box="[302,406,453,477]" captionStart="Figure 9" captionStartId="18.[129,194,1896,1920]" captionTargetBox="[129,1473,690,1868]" captionTargetId="figure-231@18.[129,1473,690,1868]" captionTargetPageId="18" captionText="Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988)." figureDoi="http://doi.org/10.5281/zenodo.14774895" httpUri="https://zenodo.org/record/14774895/files/figure.png" pageId="20" pageNumber="21">Fig. 9A–C</figureCitation>
|
||||
). The species was first discovered by DNA-based phylogenetic analyses (
|
||||
<bibRefCitation id="BF574E9CFFBB5309289EFE74F3F7F6DA" author="Delicado D & Machordom A & Ramos MA" box="[505,717,484,508]" pageId="20" pageNumber="21" pagination="403 - 17" refId="ref20723" refString="Delicado D, Machordom A, Ramos MA. Effects of habitat transition on the evolutionary patterns of the microgastropod genus Pseudamnicola (Mollusca, Hydrobiidae). Zoologica Scripta 2015; 44: 403 - 17. https: // doi. org / 10.1111 / zsc. 12104" type="journal article" year="2015">
|
||||
Delicado
|
||||
|
|
|
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Reference in a new issue