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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.145.2274" ID-GBIF-Dataset="df8a64b6-e49c-4a22-96a2-63e990890a41" ID-PMC="PMC3267457" ID-Pensoft-Pub="1313-2970-145-79" ID-Pensoft-UUID="1433FFB9FFFD8D6FF52A994B6A73FFA5" ID-PubMed="22287885" ID-Zenodo-Dep="577349" ID-ZooBank="94F032BD93F24652B7A3E914EAB8BB92" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-145-79" ModsDocOrigin="ZooKeys 145" ModsDocTitle="Rainforest understory beetles of the Neotropics, Mizotrechus Bates 1872, a generic synopsis with descriptions of new species from Central America and northern South America (Coleoptera, Carabidae, Perigonini)" checkinTime="1451249702697" checkinUser="pensoft" docAuthor="Erwin, Terry L." docDate="2011" docId="59F67FD8DF941BFB78EBFDD807C3B4B8" docLanguage="en" docName="ZooKeys 145: 79-128" docOrigin="ZooKeys 145" docSource="http://dx.doi.org/10.3897/zookeys.145.2274" docTitle="Mizotrechus gorgona Erwin 2011, sp. n." docType="treatment" docUuid="14F5ED42-8D4B-4720-829E-5F1145C10EA9" docUuidSource="ZooBank" docVersion="5" id="1433FFB9FFFD8D6FF52A994B6A73FFA5" lastPageNumber="94" masterDocId="1433FFB9FFFD8D6FF52A994B6A73FFA5" masterDocTitle="Rainforest understory beetles of the Neotropics, Mizotrechus Bates 1872, a generic synopsis with descriptions of new species from Central America and northern South America (Coleoptera, Carabidae, Perigonini)" masterLastPageNumber="128" masterPageNumber="79" pageNumber="94" updateTime="1668152588380" updateUser="ExternalLinkService">
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<mods:title>Rainforest understory beetles of the Neotropics, Mizotrechus Bates 1872, a generic synopsis with descriptions of new species from Central America and northern South America (Coleoptera, Carabidae, Perigonini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian, Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
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<mods:number>145</mods:number>
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<mods:start>79</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.145.2274</mods:url>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="152031893" LSID="urn:lsid:zoobank.org:act:14F5ED42-8D4B-4720-829E-5F1145C10EA9" httpUri="http://treatment.plazi.org/id/59F67FD8DF941BFB78EBFDD807C3B4B8" lastPageNumber="94" pageId="15" pageNumber="94">
<subSubSection pageId="15" pageNumber="94" type="nomenclature">
<paragraph pageId="15" pageNumber="94">
Isla Gorgona trough beetle
<taxonomicName LSID="59F67FD8-DF94-1BFB-78EB-FDD807C3B4B8" authority="Erwin, 2011" authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Mizotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mizotrechus gorgona" order="Coleoptera" pageId="15" pageNumber="94" phylum="Arthropoda" rank="species" species="gorgona" status="sp. n.">Mizotrechus gorgona</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="94">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Plate 3" captionStartId="F3" captionText="Plate 3. Digital Photo-illustrations, habitus, dorsal aspect: 9 Mizotrechus fortunensi s sp. n., ABL = 8.7 mm, male holotype, ADP 124968; type locality. 10 Mizotrechus gorgona. sp. n., ABL = 8.4 mm, female Holotype, ADP 128622; type locality. 11 Mizotrechus grossus sp. n., dorsal aspect, ABL = 8.9 mm, male paratype, ADP 127169; type locality. 12 Mizotrechus jefe sp. n., ABL = 6.6 mm, male paratype, ADP 011173; type locality." httpUri="https://binary.pensoft.net/fig/19344" pageId="15" pageNumber="94">Figs 10</figureCitation>
<figureCitation captionStart="Plate 6" captionStartId="F6" captionText="Plate 6. Digital Photo-illustrations, anterior femur, ventral aspect: 19 Mizotrechus minutus sp. n., ADP 124966; type locality. 20 Mizotrechus dalensi sp. n., ADP 124896; type locality. 21 Mizotrechus chontalesensis sp. n., ADP 127181; type locality. 22 Mizotrechus edithpiafae sp. n., ADP 124948; type locality. 23 Mizotrechus brulei sp. n., ADP 129205; type locality. 24 Mizotrechus gorgona. sp. n., ADP 128622; type locality." httpUri="https://binary.pensoft.net/fig/19350" pageId="15" pageNumber="94">24</figureCitation>
<figureCitation captionStart="Figure 39" captionStartId="F13" captionText="Figure 39. Distribution symbol map for known localities of Mizotrechus chontalesensis, Mizotrechus costaricensis, Mizotrechus fortunensis, Mizotrechus gorgona, Mizotrechus jefe, Mizotrechus woldai, spp. n." httpUri="https://binary.pensoft.net/fig/19359" pageId="15" pageNumber="94">39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="holotype">
<paragraph pageId="15" pageNumber="94">Holotype.</paragraph>
<paragraph pageId="15" pageNumber="94">
COLOMBIA, Cacua, Isla Gorgona Mancora, 60 m,
<geoCoordinate degrees="2.96" direction="north" orientation="latitude" precision="555" value="2.96">2.96°N</geoCoordinate>
,
<geoCoordinate degrees="78.18" direction="west" orientation="longitude" precision="555" value="-78.18">78.18°W</geoCoordinate>
, 26 June - 18 July 2000 (H Torres)(UASM: ADP128622, female).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="derivation of specific epithet">
<paragraph pageId="15" pageNumber="94">Derivation of specific epithet.</paragraph>
<paragraph pageId="15" pageNumber="94">
The epithet
<normalizedToken originalValue="“gorgona”">
&quot;
<emphasis italics="true" pageId="15" pageNumber="94">gorgona</emphasis>
&quot;
</normalizedToken>
is a Latinized noun in apposition and is the name of the island off Colombia where these beetles are found.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="proposed english vernacular name">
<paragraph pageId="15" pageNumber="94">Proposed English vernacular name.</paragraph>
<paragraph pageId="15" pageNumber="94">Isla Gorgona trough beetle.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="diagnosis">
<paragraph pageId="15" pageNumber="94">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="94">With the attributes of the genus as described above and moderately large sized for the genus as it is presently understood; adults have castaneous integument, except anterior parts of mandible, baso-lateral corner of labrum, and clypeal suture piceous. Frons shallowly rugose above and behind eye in an arc, not extended on occiput, and moderately punctulate. Occiput moderately punctulate. Pronotum longer than wide with lateral margins narrowly explanate and basally with sharp, not denticulate, hind angles; base smooth, without regular microrugose. Elytra moderately narrow and elongate, apex prolonged, wider than the width of pronotum across anterior third, and with 7 well-impressed irregularly punctulate interneurs, intervals not convex; margins behind humeri serrate, intervals between microsetae dentate. Foreleg femur with markedly serrate postero-ventral margin.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="description">
<paragraph pageId="15" pageNumber="94">Description.</paragraph>
<paragraph pageId="15" pageNumber="94">
(
<figureCitation captionStart="Plate 3" captionStartId="F3" captionText="Plate 3. Digital Photo-illustrations, habitus, dorsal aspect: 9 Mizotrechus fortunensi s sp. n., ABL = 8.7 mm, male holotype, ADP 124968; type locality. 10 Mizotrechus gorgona. sp. n., ABL = 8.4 mm, female Holotype, ADP 128622; type locality. 11 Mizotrechus grossus sp. n., dorsal aspect, ABL = 8.9 mm, male paratype, ADP 127169; type locality. 12 Mizotrechus jefe sp. n., ABL = 6.6 mm, male paratype, ADP 011173; type locality." httpUri="https://binary.pensoft.net/fig/19344" pageId="15" pageNumber="94">Figs 10</figureCitation>
,
<figureCitation captionStart="Plate 6" captionStartId="F6" captionText="Plate 6. Digital Photo-illustrations, anterior femur, ventral aspect: 19 Mizotrechus minutus sp. n., ADP 124966; type locality. 20 Mizotrechus dalensi sp. n., ADP 124896; type locality. 21 Mizotrechus chontalesensis sp. n., ADP 127181; type locality. 22 Mizotrechus edithpiafae sp. n., ADP 124948; type locality. 23 Mizotrechus brulei sp. n., ADP 129205; type locality. 24 Mizotrechus gorgona. sp. n., ADP 128622; type locality." httpUri="https://binary.pensoft.net/fig/19350" pageId="15" pageNumber="94">24</figureCitation>
).
<emphasis italics="true" pageId="15" pageNumber="94">Size</emphasis>
: See Appendix 1. Moderately large for genus, ABL = 8.4 mm, SBL = 7.15 mm, TW = 2.86 mm.
<emphasis italics="true" pageId="15" pageNumber="94">Color</emphasis>
: see diagnosis, above.
<emphasis italics="true" pageId="15" pageNumber="94">Luster</emphasis>
: Head, pronotum, and legs shiny; elytra matte.
<emphasis italics="true" pageId="15" pageNumber="94">Head</emphasis>
: Labrum quadrate, apico-medially barely emarginate. Eye small and barely convex. Gena straight. Frons, occiput, and gena glabrous.
<emphasis italics="true" pageId="15" pageNumber="94">Prothorax</emphasis>
: Moderately broad, narrowed slightly toward base, margin narrowly explanate, slightly wider before hind angle; surface punctulate, punctures widespread, very fine, glabrous.
<emphasis italics="true" pageId="15" pageNumber="94">Pterothorax</emphasis>
: Elytron barely convex, intervals nearly flat, all interneurs with well-impressed irregularly-spaced punctures, apex prolonged, slightly rounded at extreme sutural apex.
<emphasis italics="true" pageId="15" pageNumber="94">Legs</emphasis>
: Normal in female; foreleg femur (
<figureCitation captionStart="Plate 6" captionStartId="F6" captionText="Plate 6. Digital Photo-illustrations, anterior femur, ventral aspect: 19 Mizotrechus minutus sp. n., ADP 124966; type locality. 20 Mizotrechus dalensi sp. n., ADP 124896; type locality. 21 Mizotrechus chontalesensis sp. n., ADP 127181; type locality. 22 Mizotrechus edithpiafae sp. n., ADP 124948; type locality. 23 Mizotrechus brulei sp. n., ADP 129205; type locality. 24 Mizotrechus gorgona. sp. n., ADP 128622; type locality." httpUri="https://binary.pensoft.net/fig/19350" pageId="15" pageNumber="94">Fig. 24</figureCitation>
) with markedly serrate postero-ventral margin, not dentate; posterior trochanter tapered to rounded point in female, length half that of femur.
<emphasis italics="true" pageId="15" pageNumber="94">Abdomen</emphasis>
: Abdominal sterna moderately setiferous, densely medially on II and III; sternum VII barely medially notched in female.
<emphasis italics="true" pageId="15" pageNumber="94">Male genitalia</emphasis>
: Unknown.
<emphasis italics="true" pageId="15" pageNumber="94">Female genitalia</emphasis>
: Not investigated; however, it is likely similar to that illustrated on
<figureCitation captionStart="Plate 11" captionStartId="F11" captionText="Plate 11. Figure 37. Digital Photo-illustration, female genitalia: Mizotrechus dalensi sp. n., ADP 127161; Petite Montague Tortue, Guyane. A. Dorsal aspect. Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal; lt, laterotergite; gc 1, gonocoxite 1; gc 2, gonocoxite 2. B. Gonocoxite 2, dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; ves, ventral ensiform setae." httpUri="https://binary.pensoft.net/fig/19357" pageId="15" pageNumber="94">Plate 11</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="dispersal potential">
<paragraph pageId="15" pageNumber="94">Dispersal potential.</paragraph>
<paragraph pageId="15" pageNumber="94">These beetles are macropterous and capable of flight. They are moderately swift and agile runners.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="way of life">
<paragraph pageId="15" pageNumber="94">Way of life.</paragraph>
<paragraph pageId="15" pageNumber="94">
The type specimen was collected in a Malaise trap in the rainforest understory at the type locality. Adults are active in late June, or early July during the
<normalizedToken originalValue="“relatively”">&quot;relatively&quot;</normalizedToken>
dry season.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="other specimens examined">
<paragraph pageId="15" pageNumber="94">Other specimens examined.</paragraph>
<paragraph pageId="15" pageNumber="94">None.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="94" type="geographic distribution">
<paragraph pageId="15" pageNumber="94">Geographic distribution.</paragraph>
<paragraph pageId="15" pageNumber="94">
(
<figureCitation captionStart="Figure 39" captionStartId="F13" captionText="Figure 39. Distribution symbol map for known localities of Mizotrechus chontalesensis, Mizotrechus costaricensis, Mizotrechus fortunensis, Mizotrechus gorgona, Mizotrechus jefe, Mizotrechus woldai, spp. n." httpUri="https://binary.pensoft.net/fig/19359" pageId="15" pageNumber="94">Fig. 39</figureCitation>
). This species is currently known only from the type locality on an island off the Pacific coast of Colombia.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="DEBB9443656B1AE0FE4CD8463D2A3206" class="Insecta" family="Braconidae" genus="Rogas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rogas meditator" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="meditator">Rogas meditator</taxonomicName>
Haliday, 1836
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<paragraph id="B666B6F407A829F1570D64A344D4E8B2" pageId="0" pageNumber="8151">Distribution</paragraph>
<paragraph id="18E47E926D9DCDF4C828A2462594080E" pageId="0" pageNumber="8151">England, Scotland</paragraph>
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<mods:namePart>Ramahandrison, Andriamirado Tahina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0833-8730</mods:nameIdentifier>
<mods:affiliation>Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar &amp; Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakouth, Bakolimalala</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5710-2006</mods:nameIdentifier>
<mods:affiliation>Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Manuel, Michael</mods:namePart>
<mods:affiliation>Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France</mods:affiliation>
<mods:nameIdentifier type="email">manuel1570@yahoo.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1127</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1127.85737</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1127.85737</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1127-1</mods:identifier>
<mods:identifier type="ZooBank">4759AFC32EFD47A7853F645FB32829BA</mods:identifier>
<mods:identifier type="Pensoft-UUID">D72971CE12A85992AFFD69F186474E85</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:59F6BE32D147525C8F99CE88AFD08254" httpUri="http://treatment.plazi.org/id/59F6BE32D147525C8F99CE88AFD08254" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="59F6BE32-D147-525C-8F99-CE88AFD08254" authority="Bistroem, Nilsson &amp; Bergsten, 2015" authorityName="Bistroem, Nilsson &amp; Bergsten" authorityYear="2015" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus insularum" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="insularum">
Laccophilus insularum
<normalizedToken originalValue="Biström">Bistroem</normalizedToken>
, Nilsson &amp; Bergsten, 2015
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">Madagascar, Ankarafantsika National Park, Mahajanga, Boeny.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<specimenCount type="male">2 ♂♂</specimenCount>
,
<specimenCount type="female">2 ♀♀</specimenCount>
: MAK-5A.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar, widespread (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.542.5975" author="Bistroem, O" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 379" refId="B21" refString="Bistroem, O, Nilsson, AN, Bergsten, J, 2015. Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae). ZooKeys 542: 1 - 379, DOI: https://doi.org/10.3897/zookeys.542.5975" title="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)." url="https://doi.org/10.3897/zookeys.542.5975" volume="542" year="2015">
<normalizedToken originalValue="Biström">Bistroem</normalizedToken>
et al. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat in study area</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei." figureDoi="10.3897/zookeys.1127.85737.figure2" httpUri="https://binary.pensoft.net/fig/764728" pageId="0" pageNumber="1">2F</figureCitation>
).
</emphasis>
This species was encountered only once, in south-central inner Makay. The habitat was a deep pond (dimensions about 30 m
<normalizedToken originalValue="×">x</normalizedToken>
15 m) at the bottom of a small canyon along the course of a stream. This water body was formed as a result of the collapse of downstream canyon walls (natural dam). It was shaded by the canyon walls and surrounded by trees (notably
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Cyathea" kingdom="Plantae" lsidName="Cyathea" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cyathea</emphasis>
</taxonomicName>
tree ferns) and hygrophilous vegetation. The habitat was furthermore characterised by clay bottom, with moderate amount of plant debris, clear water, and absence of marginal vegetation. This site was unaffected by anthropic disturbance.
</paragraph>
</subSubSection>
</treatment>
</document>