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<document id="011A363D4405172F6CFAB29F4831ADDE" ID-DOI="10.3897/zookeys.1223.138220" ID-ZooBank="3557A991-63DF-42D8-B8B1-F310CD52FD00" ID-publisher-id="138220" URI-arpha="7352E8C9-65CA-546F-BE46-EF55E52C51A6" URI-zoobank="http://zoobank.org/3557A991-63DF-42D8-B8B1-F310CD52FD00" article-type="research-article" checkinTime="1736187739607" checkinUser="pensoft" docAuthor="Hendrich, Lars &amp; Brancucci, Michel" docDate="2025" docId="DDD59582F95157999813838156C6EE25" docLanguage="en" docName="ZooKeys 1223: 87-147" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/138220/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sandracottus bakewellii subsp. guttatus Sharp 1882, stat. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="147" masterDocId="7352E8C965CA546FBE46EF55E52C51A6" masterDocTitle="Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)" masterLastPageNumber="147" masterPageNumber="87" pageNumber="87" updateTime="1736188416300" updateUser="ExternalLinkService">
<mods:mods id="934167199E6130A368F42FEF4CF149C9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C3B0A2EA58D9791B8085E2935E772201">
<mods:title id="B91B133B47AE8AF081A867B93BB4801D">Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)</mods:title>
</mods:titleInfo>
<mods:name id="DB429F877D66DA0353BDEFD1DFB8705F" type="personal">
<mods:role id="FA87BC5DC2D6C4C2A960FF5CD23F8244">
<mods:roleTerm id="2C4A7F2682C37B642652E23140F710E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC98B3256BE075E7EEF28C824E557B7B">Hendrich, Lars</mods:namePart>
<mods:nameIdentifier id="71C4F36FE22CEAB5F71E1FF8FD2EC33D" type="ORCID">0000-0001-8366-0749</mods:nameIdentifier>
<mods:affiliation id="82F9BCF7B6ED59088E92EE92225979BC">SNSB - Zoologische Staatssammlung München, Münchhausenstraße 21, D – 81247 München, Germany</mods:affiliation>
</mods:name>
<mods:name id="0EDB02E284D2C35D9915A038D019C74C" type="personal">
<mods:role id="13BE0EBC7BF6A7A799021326C11F96FA">
<mods:roleTerm id="E71C5D555FCE1BFD3305AA9B6350A270">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E38B84FF737B7F91A16027644A62D78">Brancucci, Michel</mods:namePart>
<mods:affiliation id="E62C03A3353B1C10631269A9CEDABE33">Naturhistorisches Museum Basel, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9DC815ACEF35F989177118B270B20425">text</mods:typeOfResource>
<mods:relatedItem id="1FFD77880B7176DF7EFCD47C8E146D9B" type="host">
<mods:titleInfo id="B1BF1AC5B16FADBA008A9156D249A022">
<mods:title id="B292E8BF2B64C8C5AA1EB4EECA03260E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="633C8F1BD6ACF91268F3C96FFD192846">
<mods:date id="D23D1448B72C13CBBA43CF3A1A0AF8CA">2025</mods:date>
<mods:detail id="32118DF9FFF62A6525EADDD427BD87C3" type="pubDate">
<mods:number id="A963CAE829A6D899242A70091C0599F1">2025-01-06</mods:number>
</mods:detail>
<mods:detail id="C1A7E074FD3354CA28258A52A7EBBCDE" type="volume">
<mods:number id="F4CA53DA504888B368E1E60184C28228">1223</mods:number>
</mods:detail>
<mods:extent id="7EEE06140E5D9969AAE96ED00F7E80B7" unit="page">
<mods:start id="856220B5900B4F18BCD8ABBB89E3F7CC">87</mods:start>
<mods:end id="D8480134306EBE1991631F60EB6FB527">147</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="109B878A86846C26E1B94734C6270CBA">journal article</mods:classification>
<mods:identifier id="90823C17A6DEEDDC0B40A45065A745AF" type="DOI">10.3897/zookeys.1223.138220</mods:identifier>
<mods:identifier id="F836782B5816A1DB6C0C23B1F90EDDA8" type="ZooBank">3557A991-63DF-42D8-B8B1-F310CD52FD00</mods:identifier>
</mods:mods>
<treatment id="DDD59582F95157999813838156C6EE25" ID-DOI="http://doi.org/10.5281/zenodo.14606317" ID-Zenodo-Dep="14606317" ID-arpha="DDD59582-F951-5799-9813-838156C6EE25" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25" httpUri="http://treatment.plazi.org/id/DDD59582F95157999813838156C6EE25">
<subSubSection id="8F0B05E4091CEADF9CC6D08FC267526C" type="nomenclature">
<paragraph id="E372CCCB047BD24AACF1F0674AFC5F6A">
<taxonomicName id="72E74B900C0EAD2B110A4A6BCA294A6D" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" baseAuthorityName="Sharp Det. C. Watts" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" status="stat. nov." subSpecies="guttatus">
<emphasis id="150546AB4077E1F93A653950F44704D7" italics="true">Sandracottus bakewellii guttatus</emphasis>
<bibRefCitation id="4BAD02AB55E4B348538C1F6577888D43" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="597C930F664DE9BAAAB6158F04359DD8">stat. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3440683968BE35505D6E6BEE186043C5" type="description">
<paragraph id="BD3D5E57667BE77B465738527C4EF258">
<figureCitation id="967DB700F1BEAF2074C9AC437BEB125E" captionStart="Figure 23" captionStartId="F7" captionText="Figure 23. Distribution of Sandracottus bakewellii bakewellii [black dots] and S. bakewellii guttatus [red squares] in Australia." figureDoi="10.3897/zookeys.1223.138220.figure23" httpUri="https://binary.pensoft.net/fig/1214414">Figs 23</figureCitation>
,
<figureCitation id="CBF4E6EE36C2C75F5F89F0F4F1439541" captionStart="Figures 31–36" captionStartId="F15" captionText="Figures 31–36. Habitats of Sandracottus 31 Australia, Northern Territory, 50 km S Adelaide River, creek at Scenic Route (NT 9) and 32 South Queensland, Winfield, Winfield Road, rest pool (QLD 54), habitats of S. bakewellii bakewellii (photos: L. Hendrich) 33 Northern Territory, Ormiston Gorge, habitat of S. bakewellii guttatus (photo: M. Baehr) 34 Pakistan, Sistan va Baluchestan Prov., Pir Sohrab env., rest pool in wadi, habitat of S. dejeanii (photo: J. Hájek) 35 India, southern Madhya Pradesh, Hoshangabad District, Pagara – Pachmarhi road, ca 5 km NNE Panchmarhi, Panar Pani, habitat of S. dejeanii and S. festivus (photo: M. Jäch) 36 Indonesia, Irian Jaya, Nabire, Nabire-Ilaga track, km 62, habitat of S. femoralis (photo: L. Hendrich)." figureDoi="10.3897/zookeys.1223.138220.figures31-36" httpUri="https://binary.pensoft.net/fig/1214422">33</figureCitation>
,
<figureCitation id="327704222CE64470FFA3F7B00BFA77E7" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9989B1343B43215C1269585F9E9F166" type="reference_group">
<paragraph id="F397137C68898828EC9C77F53F24CE44">
<treatmentCitationGroup id="7D289C0FDACD3ED2C90D7AC3F4B30927">
<treatmentCitation id="0C94EDFB7D1173480AD127FB110D8D44" author="Sharp D" page="688" year="1882">
<taxonomicName id="9A87571FB0FDBB04C96040373CB56D16" authority="Sharp, 1882: 688" authorityName="Sharp" authorityPageNumber="688" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="AB6BE84142222BFE8039CA0A90A7D849" italics="true">Sandracottus guttatus</emphasis>
<comment id="AD748D09D6D235023CF4681452FF82D7">
<bibRefCitation id="D404AC1B80C09F1D1E0A5F5DB07D7D8B" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882: 688</bibRefCitation>
(
<typeStatus id="D5693260389DF8DE6E219FB8655907F7">type</typeStatus>
locality: Adelaide (?) [most probably mislabelled],
<collectingRegion id="119C26B5EF1B6BDD6025388BA59CC6C2" country="Australia" name="South Australia">South Australia</collectingRegion>
,
<collectingCountry id="63687D841D2CDF17F92D2127281E10EA" name="Australia">Australia</collectingCountry>
);
<treatmentCitation id="8E333A4D8E07E1E952E2F33B4E7334A8" author="Régimbart M" page="337" year="1899">
<bibRefCitation id="2D3A801115F24EA2BA7E7F1DEA2040BB" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" firstAuthor="Regimbart" journalOrPublisher="Annales de la Société Entomologique de France" page="337" pagination="186-367" refId="B42" refString="Régimbart M (1899) Révision des Dytiscidae de la région Indo-Sino-Malaise. Annales de la Société Entomologique de France 68: 186–367. https://doi.org/10.1080/21686351.1899.12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart 1899: 337</bibRefCitation>
</treatmentCitation>
(descr.);
<treatmentCitation id="4AA7628D3DBEB23BE38E204B773A9388" author="Zimmermann A" page="234" year="1920">
<bibRefCitation id="F73F48887D73A071D63E29038B754985" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="234" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 234</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="02BE03C34DEE448473CD773A5CD1F81D" author="Watts CHS" page="148" year="1978">
<bibRefCitation id="BEA3ED984F2141F90C1B2C01C0ABE5D9" DOI="10.1071/AJZS057" author="Watts CHS" firstAuthor="Watts" journalOrPublisher="Australian Journal of Zoology, Supplement Series" page="148" pagination="1-166" refId="B55" refString="Watts CHS (1978) A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology, Supplement Series 57: 1–166. https://doi.org/10.1071/AJZS057" title="A revision of the Australian Dytiscidae (Coleoptera)." volume="57" year="1978">Watts 1978: 148</bibRefCitation>
</treatmentCitation>
(syn.); Hájek and
<treatmentCitation id="E53105C43FE8D45C3EAE842408DBE2AB" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="BD03AE9F5EE28E38150BD5A937BC0A00" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B8F659A647774906607C567FB4F18036">
<treatmentCitationGroup id="C1B9BA0AAEA8CE02939BB21D8780E994">
<treatmentCitation id="0D1B9B54B48C10C00B2F838D7B2F26FE">
<emphasis id="24DB32DDA79641CAB29C16DB21FE114E" italics="true">
<taxonomicName id="1B2F5F082B4CFBF5F7C5C8808C9401C4" ID-CoL="79HNN" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">Sandracottus rotundus</taxonomicName>
ab.
</emphasis>
reductus
<comment id="464EF7B96125657998DB33C13CE88305">
<treatmentCitation id="895362B303990C15D5292251A1F71C15" author="Zimmermann A" page="97" year="1926">
<bibRefCitation id="F0781B48C3F88A01B630494D75AD4F97" author="Zimmermann A" issue="2" journalOrPublisher="Treubia" pagination="97-99" refId="B61" refString="Zimmermann A (1926) Fauna Buruana. Coleoptera, Fam. Dytiscidae. Treubia 7 (2): 97 - 99." title="Fauna Buruana. Coleoptera, Fam. Dytiscidae." volume="7" year="1926">Zimmermann, 1926: 97</bibRefCitation>
</treatmentCitation>
(misidentification, infrasubspecific name).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4ABG" type="type material">
<paragraph id="A1A1AAAFB99F3C5BBB777D434DC1E4C5">
<heading id="9A40B44808AD96BB38ECB02172EEDFFA" reason="title">Type material.</heading>
</paragraph>
<paragraph id="B50816D5D3D39A18E881FD1638DAE59D">
<materialsCitation id="3062261C20E4E45EF0F171B1A34FDE8F" collectionCode="NHMUK" country="Australia" location="Male" specimenCount="2" typeStatus="lectotype">
<emphasis id="BFF70448FB51496EC7B451F1C19A0DB3" bold="true" italics="true">
<typeStatus id="9E4E853342E0614E5159EB9312B897DF" type="lectotype">Lectotype</typeStatus>
</emphasis>
:
<location id="E66774F0F5E14270CCDF898632CF83D4" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:E66774F0F5E14270CCDF898632CF83D4" country="Australia" name="Male">Male</location>
, “
<collectingCountry id="FF406F6BEC584CC951D33D89C3DDAA27" name="Australia">Australia</collectingCountry>
? Adelaide 984
<taxonomicName id="B6C57B1470DDEA84DFC3F9D7F1E1084C" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="E2A37692CA6AE58A4C8F9C5DC3F4B753" italics="true">guttatus</emphasis>
</taxonomicName>
”, “
<typeStatus id="794A02D18E6102B1945FFF60F3D0934C" type="lectotype">Lectotype</typeStatus>
”, “ Sharp Coll. 1905-313. ”, “
<taxonomicName id="6CAD52FB257CDDFC7EF88DE400C151FA" authority="Sharp Det. C. Watts 1979" authorityName="Sharp Det. C. Watts" authorityYear="1979" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="853DF9C22BA51F5370B71B47D78CE1E2" italics="true">Sandracottus guttatus</emphasis>
Sharp Det. C. Watts 1979
</taxonomicName>
” (
<abbrev id="ABBRID0E2BBG" xlink_title="Natural History Museum, London">
<collectionCode id="09F0F3EF31C5E6E452EE747DD35237EB" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
)
</materialsCitation>
.
<emphasis id="9B8204020774AF553FB62CA9373EF248" bold="true" italics="true">
<typeStatus id="6C99CE548941E015FEC2C8726E05A9B5" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
: Female, “ Carpentaria 984 ”, “
<typeStatus id="A81F5E9E0196B13E4EA43A0EB5C1458C" type="paralectotype">Paralectotype</typeStatus>
”, “ Sharp Coll. 1905-313. ”, “
<taxonomicName id="B3EC3743242DC3A63BC290433DFFB4DF" authority="Sharp Det. C. Watts 1979" authorityName="Sharp Det. C. Watts" authorityYear="1979" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="6D4B80C9D126ED565D375CF717D28229" italics="true">Sandracottus guttatus</emphasis>
Sharp Det. C. Watts 1979
</taxonomicName>
” (
<abbrev id="ABBRID0EMCBG" xlink_title="Natural History Museum, London">
<collectionCode id="9E2AFF63A6F53CF88348A13105BEAE65" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
). Examined.
</paragraph>
<paragraph id="44FF3398B68F6FADD124DD164C6CB877">
<emphasis id="5F16E82CAED3C8610303E0969844844A" italics="true">
<taxonomicName id="C13890E6EF16438157F086F7AD76554B" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">Sandracottus rotundus</taxonomicName>
ab.
</emphasis>
reductus
<bibRefCitation id="6007F798B5E19429FE5B1FAF75F2AB3F" author="Zimmermann A" issue="2" journalOrPublisher="Treubia" pagination="97-99" refId="B61" refString="Zimmermann A (1926) Fauna Buruana. Coleoptera, Fam. Dytiscidae. Treubia 7 (2): 97 - 99." title="Fauna Buruana. Coleoptera, Fam. Dytiscidae." volume="7" year="1926">
Zimmermann, 1926:
<specimenCount id="237BCBBAC88661B7F3A5EC8DC530B38E" count="1" type="female">1 female</specimenCount>
</bibRefCitation>
, “ Burg Station I
<date id="E2768BEBE38029318DDAAF48379606C5" value="1921-02-10" valueMax="1921-03-16" valueMin="1921-02-10">10. II. - 16. III. 1921</date>
, L. J. Toxopeus ”, “
<typeStatus id="F3D603A43BD91C78190E6FBAF134D59D">Type</typeStatus>
”, “ ab. reductus Zimmerm ” [handwritten by Zimmermann] (
<abbrev id="ABBRID0EDDBG" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="ABE9119C15EB2522B780C0D63706B99D" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
). This specimen clearly belongs to
<taxonomicName id="A5D3CDCA1F5CCDAA0F64A691E955407C" authorityName="Sharp" authorityYear="1882" baseAuthorityName="Sharp Det. C. Watts" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="guttatus">
<emphasis id="56460E6F6D7413F2A797A9DF4DE6B232" italics="true">S. bakewellii guttatus</emphasis>
</taxonomicName>
. Toxopeus has collected it at a cattle station [Burg Station], somewhere in Central
<collectingCountry id="8A9B19A0A54F8584129EE0579E23BB09" name="Australia">Australia</collectingCountry>
. Examined.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWDBG" type="material">
<paragraph id="2D4CECEE16CE5EDB6054D7F359CDDFE2">
<heading id="518241C0030630A085FA738BFA50E68F" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="15AA528EB873B8E0BE4CDF9DD127AEBE">
<emphasis id="EFC5D7CFB4F42F9A3F2FAD5274E194B1" bold="true">
(
<specimenCount id="015E9396C758B9275F2F789F421B3062" count="63" type="generic">63 specimens</specimenCount>
):
<collectingCountry id="C8738DF2F304C0F5713A7092A0E84696" name="Australia">Australia</collectingCountry>
.
</emphasis>
•
<materialsCitation id="55EC80C648979DBEB88564BCB6FA34CB" collectingDate="1897-07" collectingDateMax="2006-05-24" collectingDateMin="1897-07" collectionCode="DEI, NHMUK, MNHN, ZSM, CLH, VIC, ANIC, NTM, SAMA" collectorName="N. Territory &amp; S. Aust. &amp; Coll. Kraatz &amp; M. Baehr &amp; C. W. Brazenov &amp; Hart Range C. Barrett &amp; F. E. Wilson &amp; T. Weir &amp; M. Malipatil &amp; J. Hawkins &amp; G. Griffin &amp; D. Nelson &amp; N. T. M. B. D. Nelson &amp; I. Archibald &amp; C. H. S. Watts &amp; P. Ranford &amp; M. S. Upton" country="Australia" county="Alice Springs" determinerName="Zimmermann" latitude="-24.05" location="Moreton Bay" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" specimenCount="57" stateProvince="Northern Territory">
<emphasis id="93E88A131A0A9EE7BE286ACDBA26AAE0" bold="true">
<collectingRegion id="FD9E5367E96BDA0E485D8BD2D75FDE1F" country="Australia" name="Northern Territory">Northern Territory</collectingRegion>
</emphasis>
:
<specimenCount id="9C307F7A5C509CCACBC4F2B184D397A2" count="1" type="generic">1 ex.</specimenCount>
, “
<collectorName id="0C5DC61132D60E1CCCFC2891AD076CEF">N. Territory</collectorName>
,
<collectorName id="269CFD1CE12DBCAA66995864873E8483">S. Aust.</collectorName>
”, “
<collectorName id="2115EB30B0306B70DB028BFA66AAC0EA">Coll. Kraatz</collectorName>
” “
<determinerName id="AA02ADFFFD846E9955C25CFB7C645935">Zimmermann</determinerName>
det. ” (
<abbrev id="ABBRID0EAEBG" xlink_title="Deutsches Entomologisches Institut">
<collectionCode id="C6907C81AEA6794E163874F655D3EF15" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" type="Other type of research institution/biorepository">DEI</collectionCode>
</abbrev>
);
<specimenCount id="AA5491D41EC14FCAC6E1754A4F15DB76" count="2" type="generic">2 exs.</specimenCount>
, “
<location id="2968F5FAB4B80FC5ED80553540BC0FC6" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:2968F5FAB4B80FC5ED80553540BC0FC6" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Moreton Bay" stateProvince="Northern Territory">Moreton Bay</location>
” [doubtful record] (
<abbrev id="ABBRID0EEEBG" xlink_title="Natural History Museum, London">
<collectionCode id="E9A9ADF11D59D6FAE71A500D19999492" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
);
<specimenCount id="8027C1D7608302DF47004CC87686F073" count="1" type="generic">1 exs.</specimenCount>
, “
<location id="2B9418D2E90C0883B75BF9E965B9E38E" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:2B9418D2E90C0883B75BF9E965B9E38E" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Moreton Bay" stateProvince="Northern Territory">Moreton Bay</location>
”,
<taxonomicName id="241B7DFE0F7F7F1498B1C85510837F94" authority="Shp" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
“
<emphasis id="B6FD2C0DCC4174227FD4317503FE82A5" italics="true">guttatus</emphasis>
Shp
</taxonomicName>
” [handwritten label by Régimbart but doubtful record] (
<abbrev id="ABBRID0EREBG" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="D94A3395B8790C221EB73E7100052D3D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="15F9695079F9E4A1C911770D330AA374" count="2" type="generic">2 exs.</specimenCount>
, “
<location id="1F99694395C9C4E054AFB971DC57CB87" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:1F99694395C9C4E054AFB971DC57CB87" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Ormiston Gorge" stateProvince="Northern Territory">Ormiston Gorge</location>
,
<date id="BE263FB343BBF07489401AC3888DDC36" value="1972-10">
<collectingDate id="90C8558E275BD97B445ECEA1A2788B36" value="1972-10">X. 1972</collectingDate>
</date>
,
<collectorName id="B77D3E423C7A142225C8AFB02DD31C86">M. Baehr</collectorName>
leg. ” (
<abbrev id="ABBRID0EVEBG" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="CCF5A0A96F415382C715326DFAD9C814" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EZEBG" xlink_title="Collection Lars Hendrich">
<collectionCode id="995DB644384183C2ECC20D292CBAB581">CLH</collectionCode>
</abbrev>
);
<specimenCount id="192A2A4FD2E3B3404E96B491EA5994BF" count="6" type="generic">6 exs.</specimenCount>
, “ Tallipatta Gorge,
<date id="E88E439E5168645AA1958695341A9C3E" value="1947-07-20">
<collectingDate id="084EEBFE4919BC9F79FF05D643FC30A6" value="1947-07-20">20. VII. 1947</collectingDate>
</date>
,
<collectorName id="7F52D84393AE0C06045D17F534C8E1B6">C. W. Brazenov</collectorName>
leg. ” (
<collectionCode id="0FD403A368BFC3B4E0777781A5F9C98D" country="Brazil" lsid="urn:lsid:biocol.org:col:15317" name="Universidade Federal de Viçosa" type="Herbarium">VIC</collectionCode>
);
<specimenCount id="76410EC640125A76A84490F9DDF99157" count="2" type="generic">2 exs.</specimenCount>
, “
<collectorName id="87A10F69361F77E1145F0E1F21B65103">Hart Range C. Barrett</collectorName>
leg.,
<collectorName id="75A5BE1D10FE4F21E945667F3AA98F51">F. E. Wilson</collectorName>
coll. ” (
<collectionCode id="951B4D15EF1B7B2A519C97FD0310377C" country="Brazil" lsid="urn:lsid:biocol.org:col:15317" name="Universidade Federal de Viçosa" type="Herbarium">VIC</collectionCode>
);
<specimenCount id="3B4A8352DDC2BE389A67D51A979E5D83" count="3" type="generic">3 exs.</specimenCount>
, “ Central
<collectingCountry id="DB81ACCBED5C8EBD2CC69C78D194FC42" name="Australia">Australia</collectingCountry>
Collection Horn Expedition,
<date id="EDD87E55DD95FD0276E57BD9762BEFE8" value="1897-07">
<collectingDate id="0FE71F56EF90EC573FC44E5167383CA1" value="1897-07">VII. 1897</collectingDate>
</date>
” (
<collectionCode id="11ACC6E1875BA7E05A1A5AF472FEA6A3" country="Brazil" lsid="urn:lsid:biocol.org:col:15317" name="Universidade Federal de Viçosa" type="Herbarium">VIC</collectionCode>
);
<specimenCount id="B1FA6124E792C956B8E61AC04CBAB9F1" count="3" type="generic">3 exs.</specimenCount>
, “
<location id="FA9C4F15D6AF4CD63105B583C3242FC8" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:FA9C4F15D6AF4CD63105B583C3242FC8" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Illamurta Springs Conservation Reserve" stateProvince="Northern Territory">Illamurta Springs Conservation Reserve</location>
, small temporary rock pool, sandy bottom, S
<geoCoordinate id="64B9BB67C0AF3B4588D66D3260DAE976" degrees="24.19" direction="east" orientation="longitude" precision="555" value="24.19">24.19 E</geoCoordinate>
132.41,
<date id="B7112D014B0F764734D37DE9C69DCA18" value="1995-03-16">
<collectingDate id="A410A1D88CAC5FDF22686F334F342A6D" value="1995-03-16">16. III. 1995</collectingDate>
</date>
,
<collectorName id="A87FD61A75CC0B3267FB64096932F4EE">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0E4EBG" xlink_title="Australian National Insect Collection">
<collectionCode id="3D2E1A1289870BD9715D1964FC198314" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="B36D5E629E9DDF8F82F00C8F55139A5E" count="1" type="generic">1 ex.</specimenCount>
, “ Finke Gorge NP, temporary pools above old
<location id="042DD428C22A640954B7D413724C79E0" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:042DD428C22A640954B7D413724C79E0" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Ranger Station" stateProvince="Northern Territory">Ranger Station</location>
,
<date id="4DE53817F5D0CE1848FC2286778D773A" value="1995-03-12">
<collectingDate id="EC5A3EB938A750A7BCDF84911C250303" value="1995-03-12">12. III. 1995</collectingDate>
</date>
,
<collectorName id="BAAF60DF31BB705DE6202F31762578E7">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0EBFBG" xlink_title="Australian National Insect Collection">
<collectionCode id="FA33B357316C01F41B692251406EC067" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="2B5E24178457C30794A7C1834B1283DE" count="2" type="generic">2 exs.</specimenCount>
, “ Finke Gorge NP, gorge
<locationDeviation id="0E43EC83975FE696608FCA65055D65D0" location="Finke River">W of Finke River</locationDeviation>
, permanent and temporary rock pools, algal growth and detritus, S
<geoCoordinate id="79CFF41A88217A0A7E0B1FCE44211CEB" degrees="24.08" direction="east" orientation="longitude" precision="555" value="24.08">24.08 E</geoCoordinate>
132.51,
<date id="F8401E2F58846F6AF376AF7F6E18A016" value="1995-03-15">
<collectingDate id="7FE1C247A1FAB2075C5F3464130315BF" value="1995-03-15">15. III. 1995</collectingDate>
</date>
,
<collectorName id="CA909910AACE1D914B336FE1B0008857">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0EFFBG" xlink_title="Australian National Insect Collection">
<collectionCode id="B7D8F57111530DB2A81D35C483698A44" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="0D3C18DCC5DDADA1A36340133DBF0FCD" count="2" type="generic">2 exs.</specimenCount>
, “ Finke Gorge NP,
<location id="409360D2FD690AA65EE967566F9D22EC" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:409360D2FD690AA65EE967566F9D22EC" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Palm Valley" stateProvince="Northern Territory">Palm Valley</location>
, small temp. pools, rocky, some sandy base, algal growth, detritus, S
<geoCoordinate id="666AF8D873A4576CDE7A876EC77B86A2" degrees="24.03" direction="east" orientation="longitude" precision="555" value="24.03">24.03 E</geoCoordinate>
132.43,
<date id="7338369D329298B4D8ABDDD5B6C2D5F8" value="1995-03-14">
<collectingDate id="C0DCDF11E3CD2ACE75CE615C3AA3DE08" value="1995-03-14">14. III. 1995</collectingDate>
</date>
,
<collectorName id="490628091B2AE07D1593421D3D53A12B">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0EJFBG" xlink_title="Australian National Insect Collection">
<collectionCode id="A0338EA66A7ED64C6B782732FC77D52E" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="2BD2A12D87728410F75346BB09CCB51B" count="3" type="generic">3 exs.</specimenCount>
, “
<locationDeviation id="47010F0259B963D322B98ADC830FBA50" location="Alice Springs">
<quantity id="FF42479B013C5E463A55B02CACF01D63" metricMagnitude="4" metricUnit="m" metricValue="4.5" unit="km" value="45.0">45 km</quantity>
W of Alice Springs
</locationDeviation>
,
<location id="E0E2237C74CB71640A40E62B74309A9D" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:E0E2237C74CB71640A40E62B74309A9D" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Standley Chasm" stateProvince="Northern Territory">Standley Chasm</location>
S
<geoCoordinate id="2A585B3C520932D72A0E1ECB49B8C83E" degrees="23.43" direction="east" orientation="longitude" precision="555" value="23.43">23.43 E</geoCoordinate>
133.28
<date id="68CC1A189EE739CBB734C342D755FA3B" value="1979-11-05">
<collectingDate id="19FBFA31BBA327790974C169A40C23B5" value="1979-11-05">5. XI. 1979</collectingDate>
</date>
,
<collectorName id="F5022956AC99AD51112DA740BC37BC32">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0ENFBG" xlink_title="Australian National Insect Collection">
<collectionCode id="7DE18F64F8AA577BD77B2D23D5E6018F" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="460CE17766E617F5E6F5C422C851FEDE" count="6" type="generic">6 exs.</specimenCount>
, “
<locationDeviation id="8F894A788AD55623D250069E4781A099" location="Alice Springs">
<quantity id="7CBB05FA5C771F71A1FE58624D9549E9" metricMagnitude="4" metricUnit="m" metricValue="3.8" unit="km" value="38.0">38 km</quantity>
SSE of Alice Springs
</locationDeviation>
S
<geoCoordinate id="6F86332061C56BC8DD49BD7E7F11FAD2" degrees="24.01" direction="east" orientation="longitude" precision="555" value="24.01">24.01 E</geoCoordinate>
134.01
<date id="C19663BD8F84F17C812338A2BA72519E" value="1979-11-07">
<collectingDate id="36D19551F317B73CF71EFDCEEF9F2BD8" value="1979-11-07">7. XI. 1979</collectingDate>
</date>
,
<collectorName id="F54AD2B4B42EEDDD0F6E46209F4FE0A5">T. Weir</collectorName>
leg. ” (
<abbrev id="ABBRID0ERFBG" xlink_title="Australian National Insect Collection">
<collectionCode id="9E50272A2533692D10728B62FAB63BC5" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="B3C33F7F960575CAF5C0F922AF5011E9" count="1" type="generic">1 ex.</specimenCount>
, “
<locationDeviation id="89C5BA1FBC6CEA6331BA372FD9E68B37" location="Alice Springs Ooraminna">
<quantity id="E1D01A833CF24E8528DA9F3F0328A190" metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
S of Alice Springs Ooraminna
</locationDeviation>
rockhole S
<geoCoordinate id="4C3CBA2AFE4892120A6A0CC943411E64" degrees="24.05" direction="east" orientation="longitude" precision="555" value="24.05">24.05 E</geoCoordinate>
134.00
<date id="1FB76358232CD9ECA2CF642BD837F222" value="1981-04-09">
<collectingDate id="7B69314AF22E9A3EADC923DB66600829" value="1981-04-09">9. IV. 1981</collectingDate>
</date>
<collectorName id="43CA5505BCAEB25F813412533569F899">M. Malipatil</collectorName>
&amp;
<collectorName id="1DA7DBFAB12EC0DB9802E8B73C63B8F9">J. Hawkins</collectorName>
leg. ” (
<abbrev id="ABBRID0EVFBG" xlink_title="Northern Territory Museum">
<collectionCode id="9A1CC222EADFF6A1152E338FE1CE1D4B" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="49188424B5A6868B1BE3507F3DC3CAAB" count="1" type="generic">1 ex.</specimenCount>
, “
<locationDeviation id="EF87D66444929DCB8D0E7319CFEF6B51" location="Alice Springs">
<quantity id="6BF8C5077CF94BF3B0F1A27CEA9ABAAB" metricMagnitude="4" metricUnit="m" metricValue="6.0" unit="km" value="60.0">60 km</quantity>
S of Alice Springs
</locationDeviation>
,
<location id="AE82B2C2E9D43F78281E93A5D93F2291" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:AE82B2C2E9D43F78281E93A5D93F2291" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Ooraminna" stateProvince="Northern Territory">Ooraminna</location>
rockhole,
<geoCoordinate id="9F3875995B55B1CE04C154B2FA8BD945" degrees="24.05" direction="south" orientation="latitude" precision="555" value="-24.05">24.05 S</geoCoordinate>
<geoCoordinate id="7B2DFACAE990F1526CBB4CE284C95980" degrees="134.00" direction="east" orientation="longitude" precision="555" value="134.0">134.00 E</geoCoordinate>
,
<date id="117CD4587541B8AACDC82589BDE48BF8" value="1976-07-25">
<collectingDate id="25562DC7ED493C4A011193BB7985F08D" value="1976-07-25">25. VII. 1976</collectingDate>
</date>
,
<collectorName id="24D45C8A9B2FED5BA323BE0D294F1958">G. Griffin</collectorName>
leg. ” (
<abbrev id="ABBRID0EZFBG" xlink_title="Northern Territory Museum">
<collectionCode id="0B2DB7BE4156DBEB1B4AA1F45F2EB3C3" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="4F7CD02399ED14390B6F3C049A56151A" count="2" type="generic">2 exs.</specimenCount>
, “ 80 E of
<location id="F46C876734FF6FE11F0D3E5A011B2C47" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:F46C876734FF6FE11F0D3E5A011B2C47" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Alice Springs" stateProvince="Northern Territory">Alice Springs</location>
,
<location id="A07A82383E5CD078BFFEE3EEB3551E27" LSID="urn:lsid:plazi:treatment:DDD59582F95157999813838156C6EE25:A07A82383E5CD078BFFEE3EEB3551E27" country="Australia" county="Alice Springs" latitude="-24.05" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" name="Standley Chasm" stateProvince="Northern Territory">Standley Chasm</location>
<date id="D9B9696BAF4FCCEE315F886A3D2554B0" value="1979-03-26">
<collectingDate id="9E9810E79DAAB87426620F9C823F169B" value="1979-03-26">26. III. 1979</collectingDate>
</date>
<collectorName id="EA24E5B51081D5FC3186E5207907B6FC">G. Griffin</collectorName>
leg. ” (
<abbrev id="ABBRID0E4FBG" xlink_title="Northern Territory Museum">
<collectionCode id="4E9782CF728382F523C4DA11892427C2" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="C79EC6053382FD5B4708B38DBCA2B63B" count="5" type="generic">5 exs.</specimenCount>
, “
<collectingMunicipality id="5B0E614A8459688173839956B2AF8124">Alice Springs</collectingMunicipality>
Old Huckitta Homestead,
<date id="AB4E90C85F28B658B0BA4D89096D2BAC" value="1970-07-20">
<collectingDate id="434E4059FA7EC0E3DEFE26A3008ED53D" value="1970-07-20">20. VII. 1970</collectingDate>
</date>
,
<collectorName id="DEB7115C6CB58A1B8CFA18D97076B6B3">D. Nelson</collectorName>
leg. ” (
<abbrev id="ABBRID0EBGBG" xlink_title="Northern Territory Museum">
<collectionCode id="FC2DA0A505956928FC2BB4314A1CB1D9" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="860D4BF2188779B5E950254D521E4815" count="2" type="generic">2 exs.</specimenCount>
, “
<collectingCounty id="76B0B67F61B20436E832343E9A224A0A">Alice Springs</collectingCounty>
, Valley of the Eagles,
<date id="EA32A37C917C3423E20B8D5B8EBC1962" value="1971-02-14">
<collectingDate id="4B4DBD9527FA18E1587F434F21C3CF13" value="1971-02-14">14. II. 1971</collectingDate>
</date>
<collectorName id="4F6D9B0E54FE2C194FF022197F22FF1D">N. T. M. B. D. Nelson</collectorName>
leg. ” (
<abbrev id="ABBRID0EFGBG" xlink_title="Northern Territory Museum">
<collectionCode id="98C20C6FE0508F16E4000DF843C8443E" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="982236697BE1B1A126211D9DACEDF56B" count="1" type="generic">1 ex.</specimenCount>
, “ Tallaputta Gorge,
<date id="96BCD0FE9D2367C6E856D47797E71FD4" value="1958-09-07">
<collectingDate id="122009A083050A6083D2F52497FEA206" value="1958-09-07">7. IX. 1958</collectingDate>
</date>
” (
<abbrev id="ABBRID0EJGBG" xlink_title="Northern Territory Museum">
<collectionCode id="9DF8E4BEE326028B361D89A3257C768F" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="E4EF6800E63CAA4997C995F69ABF4892" count="1" type="generic">1 ex.</specimenCount>
, “ Kings Canyon George Gill Range
<date id="B058CE5744D163437AFCCEFA5FED798A" value="1983-03-25" valueMax="1983-03-26" valueMin="1983-03-25">
<collectingDate id="8B99FEE7D5A84B34D16A57CDBD868904" value="1983-03-25" valueMax="1983-03-26" valueMin="1983-03-25">25. - 26. III. 1983</collectingDate>
</date>
at light
<collectorName id="BFBF15E2F80F3295D6E6867D6A397869">I. Archibald</collectorName>
leg. ” (
<abbrev id="ABBRID0ENGBG" xlink_title="Northern Territory Museum">
<collectionCode id="4A9AB104145725E7EE577D177E6D6FD9" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" type="">NTM</collectionCode>
</abbrev>
);
<specimenCount id="4D26750E0953D1B233C4A2975ADB8D0E" count="3" type="generic">3 exs.</specimenCount>
, “ Kings Canyon, George Gill Range
<date id="F43CA7F911FCA3398700BA08ED62F8A8" value="2006-05-24">
<collectingDate id="9FE3070CE48BDAE85BFB0324F27800E1" value="2006-05-24">24. V. 2006</collectingDate>
</date>
<collectorName id="4B56EBEED420AC28E38C25AA2079F123">C. H. S. Watts</collectorName>
leg. ” (
<abbrev id="ABBRID0ERGBG" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="A659202D55DD609DD361E658CB1F35AA" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EVGBG" xlink_title="South Australian Museum">
<collectionCode id="5484B7A2E0F7ADD07240AB379FF10495" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" type="Museum">SAMA</collectionCode>
</abbrev>
);
<specimenCount id="7205B641F5EAE6496EC60EE74A4BF2F1" count="2" type="generic">2 exs.</specimenCount>
, “ Northern Territory, nr. Reedy Rockhole Amadeus Basin
<date id="BAA43FEF9F96F73E508CF8002892D5AD" value="1962-09-12">
<collectingDate id="AA0009AE8A7A77F34F7C1AF727B08E07" value="1962-09-12">12. IX. 1962</collectingDate>
</date>
25-018565,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3GBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[131.583300,-24.333330]}">
<geoCoordinate id="35F659C796C457ED6E88FD5978881BD7" degrees="24.33333" direction="south" orientation="latitude" precision="1" value="-24.33333">- 24.33333</geoCoordinate>
,
<geoCoordinate id="FE0B8AF301990EB9C997B8D464D069D1" degrees="131.5833" direction="east" orientation="longitude" precision="5" value="131.5833">131.5833</geoCoordinate>
</named-content>
,
<collectorName id="10D9B3503DC470C1D01555E351B13393">P. Ranford</collectorName>
leg. ” (
<abbrev id="ABBRID0EBHBG" xlink_title="Australian National Insect Collection">
<collectionCode id="2EEDEA8723AF6DDC9AA0EF3DA8FF51A4" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
);
<specimenCount id="678C756337288C566E632F5F86D01FCB" count="5" type="generic">5 exs.</specimenCount>
, “ Northern Territory Standley Chasm
<quantity id="109FE096DF82F45F49668FA3E74CD506" metricMagnitude="4" metricUnit="m" metricValue="4.3" unit="km" value="43.0">43 km</quantity>
W by
<locationDeviation id="1ADE6DC71E5AF3C20DF33DCA640ECAC0" location="Alice Springs">S of Alice Springs</locationDeviation>
,
<date id="BDCFA8374D04B351E6B3E4FA119E504B" value="1972-10-11">
<collectingDate id="2BA40B8D090B4AE1C1BB6950FF35E66D" value="1972-10-11">11. X. 1972</collectingDate>
</date>
, 25-018569,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIHBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[133.466700,-23.716670]}">
<geoCoordinate id="89D75EAF15CA94337145F0C8DE64A822" degrees="23.71667" direction="south" orientation="latitude" precision="1" value="-23.71667">- 23.71667</geoCoordinate>
,
<geoCoordinate id="1E7FD811D2C4BA1082F1875F603CF0FC" degrees="133.4667" direction="east" orientation="longitude" precision="5" value="133.4667">133.4667</geoCoordinate>
</named-content>
<collectorName id="B49979C8C886E830606EECC1962F6F65">M. S. Upton</collectorName>
leg. ” (
<abbrev id="ABBRID0ENHBG" xlink_title="Australian National Insect Collection">
<collectionCode id="C1DC2F8429ED3BEA2036199A079320AF" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
)
</materialsCitation>
. •
<materialsCitation id="94552580C92C72DA7A8B8C16D8150308" collectingDate="1967-07-22" collectingDateMax="2006-05-24" collectingDateMin="1897-07" collectionCode="WADA" collectorName="K. J. Richards" country="Australia" county="Alice Springs" determinerName="Zimmermann" isTailEnumeration="true" latitude="-24.05" location="Moreton Bay" longLatPrecision="752" longitude="24.19" municipality="Alice Springs" specimenCount="12" stateProvince="Western Australia">
<emphasis id="1ADE61A8A8B80F9B278387F62235C1CC" bold="true">
<collectingRegion id="C7638B8B39870CFAA8EA6E82374299C8" country="Australia" name="Western Australia">Western Australia</collectingRegion>
</emphasis>
:
<specimenCount id="59B223D72907D425AC55390EAB142C4D" count="12" type="generic">12 exs.</specimenCount>
, “ Rawlinson Range
<date id="286D4B49A29CA33289E4D7165B91A92D" value="1967-07-22">
<collectingDate id="2BEE9714F25B59FEC84C2EB8976726B9" value="1967-07-22">22. VII. 1967</collectingDate>
</date>
<collectorName id="25AC90DD663623912E24C58C764D2202">K. J. Richards</collectorName>
leg. ” (
<abbrev id="ABBRID0ETHBG" xlink_title="Department of Agriculture, Western Australia">
<collectionCode id="2F939254329513035E6BAA78C725B54D" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" type="">WADA</collectionCode>
</abbrev>
)
</materialsCitation>
.
</paragraph>
<caption id="F060D426B5DA23F99FE734126FEB753B" ID-DOI="10.3897/zookeys.1223.138220.figures16-19" ID-arpha="EF33FBC7-83B1-5DFB-BBA8-0364A2D99356" httpUri="https://binary.pensoft.net/fig/1214412" startId="F5">
<paragraph id="DAF776F7F4362B0DD0E108F30EDAA984">
<label id="CFD2EFD9735D7B031AEC79FB8A09BC09">Figures 16–19.</label>
</paragraph>
<paragraph id="EC26C157B1A0CCAD8929F1BE92F90B65">
Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b):
<emphasis id="CC78BA5496237D6591C9F984377EBC90" bold="true">16</emphasis>
<taxonomicName id="EF43868F9223DE95492F7F3AF388021C" baseAuthorityName="Illiger" baseAuthorityYear="1802" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="EB6E05DC43F51B677D849400284BAE77" italics="true">Sandracottus festivus</emphasis>
</taxonomicName>
<emphasis id="476C7CB5331776572491C0C48ECE1DAF" bold="true">17</emphasis>
<taxonomicName id="155F23D0109D44D2B91DAEEC8AF4AF9A" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="8355E4149DD745252C2FF8B93ED0261D" italics="true">S. femoralis</emphasis>
</taxonomicName>
<emphasis id="80FC2C316AA4042EC9EA8E9940CE37C5" bold="true">18</emphasis>
<taxonomicName id="D773EC35A413821422DCD50C5ABC618B" authorityName="Hendrich &amp; Brancucci" authorityYear="2025" baseAuthorityName="Crotch" baseAuthorityYear="1872" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hunteri">
<emphasis id="6046FEA81A17F2B935E0949ACABB9F0E" italics="true">S. hunteri</emphasis>
</taxonomicName>
and
<emphasis id="85E08E1262B1C8F3DC354799A1B47386" bold="true">19</emphasis>
<taxonomicName id="3B59C6D9D5561B383CED2CDC6699F312" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="4CF87AF9C6AFAF18CA214EEFC4C9AD01" italics="true">S. insignis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EYJBG" type="comments">
<paragraph id="FCA6E461A24230DD2DCA434CD8366F8E">
<heading id="1488CBB0C578F5C5305AE4F2893CBCAA" reason="title">Comments on classification.</heading>
</paragraph>
<paragraph id="AC00DF656EAEB8AC1C74191D3F1E8682">
Specimens with reduced yellow basal, subbasal and apical elytral markings or almost completely black elytra and pronotum (Fig.
<figureCitation id="9C9EC941DE7427402F819662EBB9410E" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">47</figureCitation>
) have been described as
<taxonomicName id="A2D1BEFC43E32140A8749BB39C440579" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="C6737005EBB9ED6CD63F4DCE0126C265" italics="true">Sandracottus guttatus</emphasis>
<bibRefCitation id="CD22152BA9A8FCBCD616141E9625D852" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
but were later synonymised with
<taxonomicName id="C400C62FA401BC3F2270E3C9457C3101" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bakewellii">
<emphasis id="F843D322AC2D5943A422980F4536B2C4" italics="true">S. bakewellii</emphasis>
</taxonomicName>
by
<bibRefCitation id="2E2D3A90AF1A79789E1B908B086383EA" DOI="10.1071/AJZS057" author="Watts CHS" firstAuthor="Watts" journalOrPublisher="Australian Journal of Zoology, Supplement Series" pagination="1-166" refId="B55" refString="Watts CHS (1978) A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology, Supplement Series 57: 1–166. https://doi.org/10.1071/AJZS057" title="A revision of the Australian Dytiscidae (Coleoptera)." volume="57" year="1978">Watts (1978)</bibRefCitation>
. Despite the fact that they are genetically (cox 1) and morphologically identical with specimens from coastal northern and eastern
<collectingCountry id="4D9A16DD1C0CD8E8098898796FAB46CB" name="Australia">Australia</collectingCountry>
, they have a very restricted distribution in the ranges and gorges of Central
<collectingCountry id="E52E9609DDAAE6390A1931D83F11DE17" name="Australia">Australia</collectingCountry>
, and no intermediate forms are known so far. We propose subspecific rank for the population from Central
<collectingCountry id="AAE242BC38F794F7F1BF82EA5A0919AE" name="Australia">Australia</collectingCountry>
. The form described by Zimmermann as “ ab. reductus ” also refers to such a dark specimen with reduced yellow elytral markings.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3KBG" type="distribution">
<paragraph id="477F147575859CDAA367A2C8E52B1452">
<heading id="FF108612CC98F535C37D65320D0CF919" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A40151F8CA90B5354951ED6B782B99A2">
Central
<collectingCountry id="5D492D2A22A0769D736D15DA5950E3DC" name="Australia">Australia</collectingCountry>
(e. g., Macdonnell Ranges, Finke Gorge, Rawlinson Range) (Fig.
<figureCitation id="3EBA9C1239E156B52ECD64C2675F1400" captionStart="Figure 23" captionStartId="F7" captionText="Figure 23. Distribution of Sandracottus bakewellii bakewellii [black dots] and S. bakewellii guttatus [red squares] in Australia." figureDoi="10.3897/zookeys.1223.138220.figure23" httpUri="https://binary.pensoft.net/fig/1214414">23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGLBG" type="Differential diagnosis">
<paragraph id="77CE88E4888FB5A49D983CE5EB0C9A42">
<heading id="9EBDF7706D631B6AFC495784276F0C8D" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="AC5C7455A74AA9941FD5229AD2CDEE36">
The subspecies
<taxonomicName id="C07665F9CC099151E7FCA3E1B139AE05" authorityName="Sharp" authorityYear="1882" baseAuthorityName="Sharp Det. C. Watts" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="guttatus">
<emphasis id="808C65181146DBDE3AE56F601CBCCF39" italics="true">S. bakewellii guttatus</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="7347499C3D9CA49BA8E34FD4034CC68A" baseAuthorityName="Clark" baseAuthorityYear="1864" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="bakewellii">
<emphasis id="851E0293A53811EEF9BCCBBE3160B246" italics="true">S. bakewellii bakewellii</emphasis>
</taxonomicName>
by less expanded and interrupted yellowish antemedian, postmedian and preapical yellow markings on elytra (Figs
<figureCitation id="E0D90C7DB828900DE3B1EE8590707A45" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">46</figureCitation>
,
<figureCitation id="E91BA51ADC939B62A8ED7B06BDD56168" captionStart="Figures 46–49" captionStartId="F18" captionText="Figures 46–49. Colouration and habitus of 46 Sandracottus bakewellii bakewellii (Australia, Northern Queensland, Atherton Tableland, Mareeba) 47 S. bakewellii guttatus (Australia, Northern Territory, Ormiston Gorge) 48 S. bizonatus (Borneo, Sabah, Keningau) 49 S. chevrolati (Indonesia, Sumba Island, Waingapu)." figureDoi="10.3897/zookeys.1223.138220.figures46-49" httpUri="https://binary.pensoft.net/fig/1214425">47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESMBG" type="habitat">
<paragraph id="A42BF30A4898DB12B4B4CC08BFE4ED4A">
<heading id="2A5D4EF6655EC6F6212729700523E244" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="B272D749B61EBB71375694FF79C83ED0">
<taxonomicName id="3D28EB9213432B27AE44730A08189234" authorityName="Sharp" authorityYear="1882" baseAuthorityName="Sharp Det. C. Watts" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="bakewellii" subSpecies="guttatus">
<emphasis id="8544C1196E643467A4B30ABD65F09CB3" italics="true">Sandracottus bakewellii guttatus</emphasis>
</taxonomicName>
inhabit more or less permanent pools of seasonal streams and creeks, and spring fed pools. The adults are generally found in places where the water is shaded. Habitats are often enriched with dead leaves and twigs (Fig.
<figureCitation id="92D96AAC4B9F8FE855B4D52D0CBAEA93" captionStart="Figures 31–36" captionStartId="F15" captionText="Figures 31–36. Habitats of Sandracottus 31 Australia, Northern Territory, 50 km S Adelaide River, creek at Scenic Route (NT 9) and 32 South Queensland, Winfield, Winfield Road, rest pool (QLD 54), habitats of S. bakewellii bakewellii (photos: L. Hendrich) 33 Northern Territory, Ormiston Gorge, habitat of S. bakewellii guttatus (photo: M. Baehr) 34 Pakistan, Sistan va Baluchestan Prov., Pir Sohrab env., rest pool in wadi, habitat of S. dejeanii (photo: J. Hájek) 35 India, southern Madhya Pradesh, Hoshangabad District, Pagara – Pachmarhi road, ca 5 km NNE Panchmarhi, Panar Pani, habitat of S. dejeanii and S. festivus (photo: M. Jäch) 36 Indonesia, Irian Jaya, Nabire, Nabire-Ilaga track, km 62, habitat of S. femoralis (photo: L. Hendrich)." figureDoi="10.3897/zookeys.1223.138220.figures31-36" httpUri="https://binary.pensoft.net/fig/1214422">33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELNBG" type="Conservation">
<paragraph id="D1B695B33E0289959DD1C7AEBB2DE111">
<heading id="61315542BA9CDDE3BE15AD760CA6883A" reason="title">Conservation.</heading>
</paragraph>
<paragraph id="34738F127F8FF4846774FCEF0889DB4A">An isolated subspecies with a very small range which needs special conservation attention as surface water in this area is very limited.</paragraph>
</subSubSection>
</treatment>
</document>